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<document ID-DOI="http://dx.doi.org/10.3897/vz.73.e102475" ID-Pensoft-Pub="2625-8498-73-931" ID-Pensoft-UUID="EE261ADB23925B95AD23FA3F71DF2FBA" ID-ZooBank="6723C6BC034B49B68F26806978782527" ModsDocID="2625-8498-73-931" checkinTime="1696476018141" checkinUser="pensoft" docAuthor="Sheridan, Jennifer A., Phimmachak, Somphouthone, Sivongxay, Niane &amp; Stuart, Bryan L." docDate="2023" docId="5EB04F147E28594F84C4897B2938A2B6" docLanguage="en" docName="VertZool 73: 931-956" docOrigin="Vertebrate Zoology 73" docPubDate="2023-10-04" docSource="http://dx.doi.org/10.3897/vz.73.e102475" docTitle="Amolops attiguus Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart 2023, sp. nov." docType="treatment" docUuid="27659B24-8B32-4D0A-B0B9-AECBEFB954AC" docUuidSource="ZooBank" docVersion="2" id="EE261ADB23925B95AD23FA3F71DF2FBA" lastPageNumber="931" masterDocId="EE261ADB23925B95AD23FA3F71DF2FBA" masterDocTitle="Systematics of the Lao torrent frog, Amolops cremnobatus Inger &amp; Kottelat, 1998 (Anura: Ranidae), with descriptions of four new species" masterLastPageNumber="956" masterPageNumber="931" pageNumber="931" updateTime="1696476320615" updateUser="ExternalLinkService">
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<mods:title id="C453C1155BA54FA18B5DF82994EFE580">Systematics of the Lao torrent frog, Amolops cremnobatus Inger &amp; Kottelat, 1998 (Anura: Ranidae), with descriptions of four new species</mods:title>
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<mods:name id="E08B3C8294010F8B1C64459396EF2230" type="personal">
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<mods:namePart id="5D00E6ACFA82D462C824D94DA1F5AB42">Sheridan, Jennifer A.</mods:namePart>
<mods:affiliation id="1EBE940A450E15A54E1A8866461ED580">Section of Amphibians &amp; Reptiles, Carnegie Museum of Natural History, 4400 Forbes Ave, Pittsburgh, PA 15213 USA</mods:affiliation>
<mods:nameIdentifier id="6AF560B67584299B0078B5377D673A4E" type="email">jasheridan@gmail.com</mods:nameIdentifier>
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<mods:name id="4297FB0A4FD6C0158B1938689B87E4BE" type="personal">
<mods:role id="23A529448BBB2921FBB8A8982BD222EF">
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<mods:namePart id="F1F8F4CC33DE9BE9A351129C463CB3DC">Phimmachak, Somphouthone</mods:namePart>
<mods:affiliation id="7AB3D55C64B648573DA8E449637DC956">Department of Biology, Faculty of Natural Sciences, National University of Laos, Dongdok Campus, P. O. Box 7322, Vientiane, Laos PDR</mods:affiliation>
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<mods:name id="266D398DE142BE8C6F4AEB4FDB236572" type="personal">
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<mods:namePart id="A08EA81EE2494A902640F8AFF2923E3C">Sivongxay, Niane</mods:namePart>
<mods:affiliation id="BF7FF2F35AA04A8FD24385050DB9C342">SEAMEO Regional Centre for Community Education Development, Ministry of Education and Sports, Dongdok, Xaithany District, P. O. Box 67, Vientiane, Lao PDR</mods:affiliation>
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<mods:namePart id="DD1A6AAC7B9CAABAAA0265CBEDE52AE5">Stuart, Bryan L.</mods:namePart>
<mods:affiliation id="3AA44AEDF8CE1866AAAAC6C37A41DEBD">North Carolina Museum of Natural Sciences, 11 West Jones St, Raleigh, NC 27601 USA</mods:affiliation>
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<mods:typeOfResource id="E3B985D9E83BFB0AACD15D0E96F9C1C3">text</mods:typeOfResource>
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<mods:part id="0874649CFF2BB0F46F8C402E4BF1EF84">
<mods:date id="E1B8A9AF307806BDF112C77016DE25BA">2023</mods:date>
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<mods:number id="A0448E62DDAE2C473D15DE00A648E8CD">2023-10-04</mods:number>
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<mods:number id="8CD336692B21902C02B1A127D0606115">73</mods:number>
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<mods:identifier id="607B28D8F9ACA821B737BE2D7307F529" type="DOI">http://dx.doi.org/10.3897/vz.73.e102475</mods:identifier>
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<subSection id="493E9FEEAC042E0BECDDAF2E9EC42E9D" pageId="0" pageNumber="931" type="taxonomy">
<treatment id="5EB04F147E28594F84C4897B2938A2B6" LSID="urn:lsid:zoobank.org:act:27659B24-8B32-4D0A-B0B9-AECBEFB954AC" httpUri="http://treatment.plazi.org/id/5EB04F147E28594F84C4897B2938A2B6" lastPageNumber="931" pageId="0" pageNumber="931">
<subSubSection id="AA1662D3D09DA1DCE4560F4561B51096" pageId="0" pageNumber="931" type="nomenclature">
<paragraph id="4798561F9081B1CE7F39D9054EE5BDA6" pageId="0" pageNumber="931">
<taxonomicName id="C69B469480CB77ABB6F6E9173E31109A" LSID="https://zoobank.org/27659B24-8B32-4D0A-B0B9-AECBEFB954AC" authority="Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart, 2023" authorityName="Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart" authorityYear="2023" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops attiguus" order="Anura" pageId="0" pageNumber="931" phylum="Chordata" rank="species" species="attiguus" status="sp. nov.">Amolops attiguus</taxonomicName>
<taxonomicNameLabel id="E762434F5743CE7B1F086D9C19D14F1C" pageId="0" pageNumber="931">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="CF0491DE9B210B965E564D17D232732E" pageId="0" pageNumber="931" type="description">
<paragraph id="4CF2BFE21BA4F4ADF5C0BBB1B3352613" pageId="0" pageNumber="931">
<figureCitation id="59B1A449EB88D5E24E99727812BC3966" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Amolops attiguus sp. nov. NCSM 80907 in life (top row) and in preservative (bottom row)." figureDoi="10.3897/vz.73.e102475.figure12" httpUri="https://binary.pensoft.net/fig/916375" pageId="0" pageNumber="931">Figure 12 &quot;Clade E&quot;</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FFB795CF5135CA235D1CF3E648AE07C6" pageId="0" pageNumber="931" type="holotype">
<paragraph id="D61659AA633EAAADA4755428A232F950" pageId="0" pageNumber="931">Holotype.</paragraph>
<paragraph id="3600917D4541EA56D5CFE59ADD1EC3D7" pageId="0" pageNumber="931">
NCSM 80907 (field number BLS 15852), adult male, Laos, Bolikhamxay Province, Viengthong District, Nam Kading National Protected Area, Nam Xouang,
<geoCoordinate id="8CF7E6B6E7ED5BD40C514C769183754F" degrees="18.42676" direction="north" orientation="latitude" precision="1" value="18.42676">18.42676°N</geoCoordinate>
,
<geoCoordinate id="54ABE44C31310FAAB12C2B0A8DC8D47A" degrees="104.39136" direction="east" orientation="longitude" precision="1" value="104.39136">104.39136°E</geoCoordinate>
, 343 m elev., coll. 2200 h on wet rock face next to series of waterfalls over exposed sandstone bedrock with potholes on 1 March 2013 by Bryan L. Stuart, Jennifer A. Sheridan, Sengvilay Seateun, and Niane Sivongxay.
</paragraph>
<caption id="B6CE9B09BA728217D6B8D2820D033287" doi="10.3897/vz.73.e102475.figure12" httpUri="https://binary.pensoft.net/fig/916375" pageId="0" pageNumber="931" start="Figure 12" startId="F12">
<paragraph id="ADE1389EB14201465F5D32C6D2A076DD" pageId="0" pageNumber="931">
<emphasis id="072625039CF03644D1C1E9E0D8F0752C" bold="true" pageId="0" pageNumber="931">Figure 12.</emphasis>
<taxonomicName id="98AB7DB48E0F901BB24049C7081B56E5" authorityName="Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart" authorityYear="2023" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops attiguus" order="Caudata" pageId="0" pageNumber="931" phylum="Chordata" rank="species" species="attiguus">
<emphasis id="BB079041A34FF553C55ED546165D91A6" italics="true" pageId="0" pageNumber="931">Amolops attiguus</emphasis>
</taxonomicName>
<emphasis id="2800555232728A5031D56208B143C254" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
NCSM 80907 in life (top row) and in preservative (bottom row).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A87341959DCE0459002E6A34E5182E8C" pageId="0" pageNumber="931" type="paratypes">
<paragraph id="0F30D2DB8C55ADF97B962C459C6F169C" pageId="0" pageNumber="931">Paratypes.</paragraph>
<paragraph id="DC771A9BB129E8DA2FF2A88CB1E2ABB7" pageId="0" pageNumber="931">
Laos, Xieng Khouang, Mork District, tributary of Nam Mork: NCSM 80761 (one adult male),
<geoCoordinate id="434A5EA310A5337873B0D8CC3C8CCEC0" degrees="19.07075" direction="north" orientation="latitude" precision="1" value="19.07075">19.07075°N</geoCoordinate>
,
<geoCoordinate id="DE62B2A74073EC41452129DD432D1E45" degrees="103.95870" direction="east" orientation="longitude" precision="1" value="103.9587">103.95870°E</geoCoordinate>
, 454 m elev., coll. 12 April 2012 by Somphouthone Phimmachak. - Vietnam, Nghe An Province, Anh Son District, Pu Mat National Park, Cao Ven: AMNH A 191846 (one adult male),
<geoCoordinate id="BD0D0FDF1D20C942386F0BFA07AD6CE0" degrees="18.81050" direction="north" orientation="latitude" precision="1" value="18.8105">18.81050°N</geoCoordinate>
,
<geoCoordinate id="EC918D78729F2910FCE6EE044F169929" degrees="104.95150" direction="east" orientation="longitude" precision="1" value="104.9515">104.95150°E</geoCoordinate>
, 273 m elev., coll. 16 May 2010 by David A. Kizirian, Truong Quang Nguyen, and Sinh Van Nguyen; AMNH A 191848 (one adult male), AMNH A 191849, NCSM 79166 (two adult females), coll. 18 May 2010 by David A. Kizirian, Truong Quang Nguyen, and Sinh Van Nguyen.
</paragraph>
</subSubSection>
<subSubSection id="AEF6720757B2C78BF767E2F1BBC07E8E" pageId="0" pageNumber="931" type="etymology">
<paragraph id="722BA891EDE554DC12D7D44FAC984461" pageId="0" pageNumber="931">Etymology.</paragraph>
<paragraph id="FC7013AE45B41E672697B4C75AB939CE" pageId="0" pageNumber="931">
The specific epithet taken from
<emphasis id="26E5E9266BD3AE46AB62D9444A64F530" italics="true" pageId="0" pageNumber="931">attigua</emphasis>
(L.) for neighboring or adjacent, in reference to the new
<normalizedToken id="ED1D23F383D0BA54C3B200581EA62ED2" originalValue="species">species'</normalizedToken>
nested geographic distribution between its morphologically similar relatives
<taxonomicName id="DCDBC8ADEFF90654FB8368457D07F76E" lsidName="A. tanfuilianae" pageId="0" pageNumber="931" rank="species" species="tanfuilianae">
<emphasis id="343C29A1AE84E59E45FACF26DC14CBFD" italics="true" pageId="0" pageNumber="931">A. tanfuilianae</emphasis>
</taxonomicName>
<emphasis id="0DCF7462B8240ED4E3DA3BE94E92DAB4" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
and
<taxonomicName id="41A251D4F9077FB40E0AE5B0C2416200" lsidName="A. kottelati" pageId="0" pageNumber="931" rank="species" species="kottelati">
<emphasis id="79C3BA9F5FFD48FE3647EBA4C3B61E39" italics="true" pageId="0" pageNumber="931">A. kottelati</emphasis>
</taxonomicName>
<emphasis id="4004B94B4F49FC5403C26F4CEFA17ECB" bold="true" pageId="0" pageNumber="931">sp. nov</emphasis>
. (Fig.
<figureCitation id="92DC412912078DAD1A108225642E829B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of specimen sampling locations of Amolops cremnobatus (orange stars), A. tanfuilianae sp. nov. (blue circles), A. sengae sp. nov. (green diamonds), A. kottelati sp. nov. (red triangles), and A. attiguus sp. nov. (purple squares). Filled symbols are adults and open symbols are tadpoles. Symbols with black dots are sequenced individuals, and symbols without black dots are unsequenced individuals. The sample in Thailand is provisionally referred to A. sengae sp. nov. (see text)." figureDoi="10.3897/vz.73.e102475.figure1" httpUri="https://binary.pensoft.net/fig/916364" pageId="0" pageNumber="931">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="306CEFD68D61B49E43FE51AD650D18E5" pageId="0" pageNumber="931" type="suggested common names">
<paragraph id="0BEFAFC48EB1B1BECE1F3B0128F3544B" pageId="0" pageNumber="931">Suggested Common Names.</paragraph>
<paragraph id="9BFE6F850724B7D9886654188E932ACA" pageId="0" pageNumber="931">
Similar Lao torrent frog (English),
<normalizedToken id="ED38F0D645E38E12D4013299112F7247" originalValue="ຂຽດເກາະຜາລາວແອດທ">ຂຽດເກາະຜາລາວແອດທ</normalizedToken>
ີກຸສ (Khiat Korpha Lao Sumphan; Lao).
</paragraph>
</subSubSection>
<subSubSection id="E0F69276D8E468052EFF26FDB9A7C837" pageId="0" pageNumber="931" type="diagnosis">
<paragraph id="D8D08F060C42CAAEBA463D869852C631" pageId="0" pageNumber="931">Diagnosis.</paragraph>
<paragraph id="A5531AC881A0816206D3677DBEA7B7E5" pageId="0" pageNumber="931">
A member of the
<taxonomicName id="B72CBBEAB46A16508504322E917A8D54" baseAuthorityName="Boulenger" baseAuthorityYear="1899" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops larutensis" order="Caudata" pageId="0" pageNumber="931" phylum="Chordata" rank="species" species="larutensis">
<emphasis id="A3B851C758FDDC64A4F5463F2E6F1B61" italics="true" pageId="0" pageNumber="931">Amolops larutensis</emphasis>
</taxonomicName>
group having the combination of 2-3 vomerine teeth, sometimes faint or absent; relative finger lengths I &lt;IV&lt;II &lt;III; mean
<normalizedToken id="1BED603DAE7C14DDE9527B887FC31B1D" originalValue="±">+/-</normalizedToken>
SE SVL of adult males 30.2
<normalizedToken id="67F2C956E156645B5CD56ABD907F6407" originalValue="±">+/-</normalizedToken>
0.9 mm (range 28.9-31.3 mm; n = 5) and of females 39.2-39.4 mm (n = 2); EYE 4.7
<normalizedToken id="0CADB6F33856DB652AC18371AC9B0D83" originalValue="±">+/-</normalizedToken>
0.3 mm (range 4.1-4.8 mm) in males and 5.3-6.1 mm in females; TEY 0.7
<normalizedToken id="AC14C8998CC7742FF6798238AC4C68B8" originalValue="±">+/-</normalizedToken>
0.1 mm (range 0.5-0.9 mm) in males and 1.1-1.3 mm in females; SNT 4.1
<normalizedToken id="CD00B940B48A6EE5D190511909AADF03" originalValue="±">+/-</normalizedToken>
0.1 mm (range 4.5-4.7 mm) in males and 5.4-5.5 mm in females; and FTL 15.8
<normalizedToken id="586C5378595E83FC6440F35F31E49018" originalValue="±">+/-</normalizedToken>
0.7 mm (range 15.0-16.8 mm) in males and 18.6-18.7mm in females.
</paragraph>
</subSubSection>
<subSubSection id="24415CF1EF961264380BEDAB4F890A2D" pageId="0" pageNumber="931" type="description">
<paragraph id="DC05FE90A82DCB5709E660C6E3205FA3" pageId="0" pageNumber="931">Description of the holotype.</paragraph>
<paragraph id="5FC312B505FA1764EC1FEF72DFF8E1B9" pageId="0" pageNumber="931">Habitus moderately slender. Head length subequal to head width. Snout weakly pointed in dorsal view. Snout projecting slightly beyond lower jaw in lateral view, sloping obliquely back to lip. Nostril lateral, nearer to tip of snout than to eye. Canthus rostralis distinct. Lores obliquely and slightly concave. Eye diameter approximately equal to snout. No pineal body visible. Tympanum distinct, round, approximately one-third of eye diameter, slightly depressed relative to skin of temporal region, tympanic rim elevated relative to tympanum. Rictal glands slightly oval and pronounced. Vomerine teeth obliquely angled, closer to each other than to choanae, and extremely small, with 2-3 teeth each. Tongue cordiform, notched posteriorly. Vocal slit opening near corner of jaw. No gular pouch, but with some extra skin at edge of jaw.</paragraph>
<paragraph id="C7B95BF3D7FD9B76F0DC856C42EEAA2E" pageId="0" pageNumber="931">Forelimb moderately slender. Tips of all four fingers expanded into wide discs with circummarginal grooves. Fingers slender. Relative finger lengths I &lt;IV&lt;II &lt;III. Minimal basal webbing on hands. Subarticular tubercles conspicuous, surfaces rounded, formula 1, 1, 2, 2. One supernumary tubercle at base of fingers 2-4. Irregularly-shaped palmar and oval thenar tubercles present. Well-developed nuptial pad on Finger I, on dorsal surface to level of distal end of subarticular tubercle.</paragraph>
<paragraph id="0810B9FC2139B5334897D353E7F20721" pageId="0" pageNumber="931">Hindlimbs moderately long and slender. Toes slender. Tips of toes expanded into wide discs with circummarginal grooves. Web on all toes to base of disc. Subarticular tubercles conspicuous, surfaces rounded, formula 1, 1, 2, 2, 2. Inner metatarsal tubercle oval. No outer metatarsal tubercle. Discs of toes smaller than that of finger, but larger than or equal to that of outermost finger. Outermost toe disc slightly wider than toe. Widest toe disc narrower than all finger discs except for that of thumb. Subarticular tubercles conspicuous, surfaces rounded; formula 1, 1, 2, 2, 2. Inner metatarsal tubercle oval; outer metatarsal tubercle barely visible, small, round.</paragraph>
<paragraph id="61EE07437B01AFA799D1457BAA4EA933" pageId="0" pageNumber="931">Dorsal surface with weakly spinose glands. No supra-tympanic fold. Glandular dorsolateral fold. Flanks glandular. Spinose glands behind tympanum above arm. Glands around vent. Ventral surfaces weakly granular.</paragraph>
<paragraph id="E1187157103050CDB873B40BD3945A9B" pageId="0" pageNumber="931">Measurements of holotype (mm): SVL 30.4, HDL 10.3, HDW 11.0, SNT 4.7, EYE 4.6, IOD 3.2, IND 3.6, TMP 1.7, TEY 0.8, SHK 19.1, TGH 17.2, HND 9.7, FTL 16.8.</paragraph>
</subSubSection>
<subSubSection id="F6011B5188EC02DFA8F31533BA10A485" pageId="0" pageNumber="931" type="coloration">
<paragraph id="9668C6B9C2EAAB5A9965FB97F7D04B77" pageId="0" pageNumber="931">Coloration.</paragraph>
<paragraph id="B0E394AEFE65B915C10D1449F3893698" pageId="0" pageNumber="931">In preservative, dorsal coloration black-brown with paler markings. Pale bars on dorsal surface of thighs and arms. Posterior surface of thighs with irregular pale mottling or reticulation on dark surface. Ventral surfaces cream with very fine dark mottling. Ventral surface of feet dark grey-brown, ventral surface of hands dark beige.</paragraph>
<paragraph id="2A08AE3C3C4311E9E52E28F08CFC290C" pageId="0" pageNumber="931">
In life (from photos of NCSM 80906, Fig.
<figureCitation id="0BFFC824E6074FB7981263EA9EA2AFC9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of specimen sampling locations of Amolops cremnobatus (orange stars), A. tanfuilianae sp. nov. (blue circles), A. sengae sp. nov. (green diamonds), A. kottelati sp. nov. (red triangles), and A. attiguus sp. nov. (purple squares). Filled symbols are adults and open symbols are tadpoles. Symbols with black dots are sequenced individuals, and symbols without black dots are unsequenced individuals. The sample in Thailand is provisionally referred to A. sengae sp. nov. (see text)." figureDoi="10.3897/vz.73.e102475.figure1" httpUri="https://binary.pensoft.net/fig/916364" pageId="0" pageNumber="931">1</figureCitation>
), dorsal surface greenish-grey with bright green markings. Dorsal surface of legs paler in color than dorsum. Eye mottled gray on lower one-third, mottled yellowish on upper one-third, with orange on anterior and posterior portions. No ventral photos exist for this specimen in life, but photos of this species recently euthanized (NCSM 80907, Fig.
<figureCitation id="0BB703887C992BDC259AB596AEAAC996" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Amolops attiguus sp. nov. NCSM 80907 in life (top row) and in preservative (bottom row)." figureDoi="10.3897/vz.73.e102475.figure12" httpUri="https://binary.pensoft.net/fig/916375" pageId="0" pageNumber="931">12</figureCitation>
) show chin and chest to be creamy white with dark mottling, belly to be white with very fine mottling, and underside of arms and thighs to be translucent.
</paragraph>
</subSubSection>
<subSubSection id="090CA23FF6FA0F4E86CB430F99B41A16" pageId="0" pageNumber="931" type="sexual dimorphism">
<paragraph id="B98C62C2AE797D05A424B7F439BBEC10" pageId="0" pageNumber="931">Sexual dimorphism.</paragraph>
<paragraph id="3FBE1CEC31FA7359A4B22E290C1EE255" pageId="0" pageNumber="931">
Males (n = 5) differ from females (n = 2) in being smaller (30.2
<normalizedToken id="DA3497FF24D3ACD3A40DC3D7E46833E8" originalValue="±">+/-</normalizedToken>
0.9 mm SVL vs 39.2-39.4 mm; Table
<tableCitation id="0AFC66151E53F07F06391F8FA1C0F9FB" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Mean + / - SD (range) of adult body measurements (mm) of Amolops examined in the present study." httpUri="http://table.plazi.org/id/229B758EC90EA927C14B047ECBBC36DC" pageId="0" pageNumber="931" tableUuid="229B758EC90EA927C14B047ECBBC36DC">1</tableCitation>
), and in possessing paired vocal slits and distinct nuptial pads at base of first finger.
</paragraph>
</subSubSection>
<subSubSection id="FE5F982825F37D5BC0E07F2234A76BCD" pageId="0" pageNumber="931" type="variation">
<paragraph id="874194CEC6330135C727A81316990E1A" pageId="0" pageNumber="931">Variation.</paragraph>
<paragraph id="FE4A02C9C51A2820C9334AAECC9C1355" pageId="0" pageNumber="931">Pineal body distinguishable in NCSM 80761, and clear and obvious in AMNH A 191845, 191846, 191848, and 191849, but not apparent in NCSM 79166. Rictal glands indistinct, bordering on absent in all paratypes. Vomerine teeth totally absent in AMNH A 191847; almost absent in AMNH A 191846 and A 191848; extremely small in NCSM 80761, and a bit more prominent in AMNH A 191845. Females have oval thenar tubercle. Outermost toe disc variable: in some individuals it is barely wider than toe, but in others appears wider.</paragraph>
<paragraph id="6828910E7677551B40A6E03193596526" pageId="0" pageNumber="931">Male AMNH A 191845 very slightly spinose. Dorsal surface of females glandular but not spinose. Glandular dorsolateral fold weak in females and in AMNH A 191845, 191846, and 191848. Spinose glands behind tympanum above arm, and in NCSM 79166 (female), coming down to skin in front of arm, though this feature is absent in AMNH A 191845, 191846, and 191848. In NCSM 79166, the pale dorsal markings are extremely sparse, giving the frog a dark appearance overall with few markings, while in the others, the dorsal surfaces are much more mottled with the paler markings. Pale bars on dorsal surface of thighs and arms less apparent in NCSM 79166 than in other individuals. Ventral coloration of NCSM 79166 creamy on the belly, and white on chin and chest with dark mottling. AMNH specimens A 191845, 19148, and 19149 have white chin and chest with yellow-cream belly (no dark mottling). AMNH A 19146 has white chin and chest with some dark mottling, and yellow-cream belly. Ventral surfaces of feet are dark brown to black, and ventral surfaces of hands are dark beige to black.</paragraph>
</subSubSection>
<subSubSection id="6D480055F3877030708877EFB8BD66DD" pageId="0" pageNumber="931" type="distribution">
<paragraph id="2A43510AE498E023330BE14CAF6F9D71" pageId="0" pageNumber="931">Distribution and natural history.</paragraph>
<paragraph id="4CD478F1385CA67A25204F7700E0966E" pageId="0" pageNumber="931">This species is known from clear streams with torrents in eastern Bolikhamxay and Xieng Khouang Provinces of northern Laos and southern Nghe An Province of northern Vietnam. Larvae of the new species remain unknown.</paragraph>
</subSubSection>
<subSubSection id="CB7D70E328E3E3DEFE2395004B146A83" pageId="0" pageNumber="931" type="comparisons">
<paragraph id="684240653A640233A829FE904768639A" pageId="0" pageNumber="931">Comparisons.</paragraph>
<paragraph id="1913EA17744B60D6FBA5A5796FF9A1A0" pageId="0" pageNumber="931">
<taxonomicName id="ADF3BD79072BCE99AC5875A5FF3FC452" authorityName="Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart" authorityYear="2023" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops attiguus" order="Caudata" pageId="0" pageNumber="931" phylum="Chordata" rank="species" species="attiguus">
<emphasis id="62CDC12DF143A6E8D76CCBE0396F1468" italics="true" pageId="0" pageNumber="931">Amolops attiguus</emphasis>
</taxonomicName>
<emphasis id="FD1151528ED4311221B897B7809D00D7" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
differs from
<taxonomicName id="BB8FA81786A970D67051B2B126223D3D" lsidName="A. tanfuilianae" pageId="0" pageNumber="931" rank="species" species="tanfuilianae">
<emphasis id="2E9AE0E969C6D50986D3AC4FCF244896" italics="true" pageId="0" pageNumber="931">A. tanfuilianae</emphasis>
</taxonomicName>
<emphasis id="40F9C3360A4618230987E30BCE5E66F9" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
and
<taxonomicName id="5DD071FC4161DAD6D15C0018B0B8FF0E" lsidName="A. kottelati" pageId="0" pageNumber="931" rank="species" species="kottelati">
<emphasis id="3FEFB54C72BC035B0D49ED359DA73E28" italics="true" pageId="0" pageNumber="931">A. kottelati</emphasis>
</taxonomicName>
<emphasis id="52C2FC7F235512EDD59F61C766D38818" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
by having a smaller number (2-3) of vomerine teeth (3-4 in
<taxonomicName id="24F1B333E41E613309794DE967FD95B4" lsidName="A. tanfuilianae" pageId="0" pageNumber="931" rank="species" species="tanfuilianae">
<emphasis id="C4A0ED3AC396DDC74B9B65AE47E3A081" italics="true" pageId="0" pageNumber="931">A. tanfuilianae</emphasis>
</taxonomicName>
<emphasis id="7088BF1F2DCF75B55124273A327452C1" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
and
<taxonomicName id="62E416A151428A31013B0E7FC67FBCA6" lsidName="A. kottelati" pageId="0" pageNumber="931" rank="species" species="kottelati">
<emphasis id="729029E0E5B3B3C2C39CD746EDD50614" italics="true" pageId="0" pageNumber="931">A. kottelati</emphasis>
</taxonomicName>
<emphasis id="B96723FCB0B3B5A22E0DB4871FB40D17" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
).
<taxonomicName id="9A8CDDEA58061A2E4B0B41118FBF89D5" authorityName="Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart" authorityYear="2023" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops attiguus" order="Caudata" pageId="0" pageNumber="931" phylum="Chordata" rank="species" species="attiguus">
<emphasis id="A857CD7DCBA95CD5A437D299A18E306D" italics="true" pageId="0" pageNumber="931">Amolops attiguus</emphasis>
</taxonomicName>
<emphasis id="61ACA496EF0274367F06384A90DE5298" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
differs from
<taxonomicName id="F37D519F98232083A9602CAE22582AF5" lsidName="A. cremnobatus" pageId="0" pageNumber="931" rank="species" species="cremnobatus">
<emphasis id="93B358B129EFC1C668A01BF9BDDCDF22" italics="true" pageId="0" pageNumber="931">A. cremnobatus</emphasis>
</taxonomicName>
and further differs from
<taxonomicName id="11D193C62683B215C44DF2A242BA5779" lsidName="A. tanfuilianae" pageId="0" pageNumber="931" rank="species" species="tanfuilianae">
<emphasis id="612FE5EBB37D864B35320373BD9F8224" italics="true" pageId="0" pageNumber="931">A. tanfuilianae</emphasis>
</taxonomicName>
<emphasis id="BC0F40FFD97B6F4707776F7DDEB91A03" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
by having relative finger lengths I&lt;IV&lt;II&lt;III (I&lt;II&lt;IV&lt;III in
<taxonomicName id="E7A0FE9B894E4CDDE38A6E58F4031600" lsidName="A. cremnobatus" pageId="0" pageNumber="931" rank="species" species="cremnobatus">
<emphasis id="D4BD22F181E4A759308246D5896B572E" italics="true" pageId="0" pageNumber="931">A. cremnobatus</emphasis>
</taxonomicName>
and
<taxonomicName id="55DCF261047AC8F5DF699E4108205C39" lsidName="A. tanfuilianae" pageId="0" pageNumber="931" rank="species" species="tanfuilianae">
<emphasis id="325176829715606D8D7225CE550C12BA" italics="true" pageId="0" pageNumber="931">A. tanfuilianae</emphasis>
</taxonomicName>
<emphasis id="9C9F889286B65A4577AC362D5BFA82F9" bold="true" pageId="0" pageNumber="931">sp. nov.)</emphasis>
.
<taxonomicName id="BE5708C8A8E075E3810EE74B8A959FFC" authorityName="Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart" authorityYear="2023" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops attiguus" order="Caudata" pageId="0" pageNumber="931" phylum="Chordata" rank="species" species="attiguus">
<emphasis id="8B363C890C31EBD18A860DBB1DA82A23" italics="true" pageId="0" pageNumber="931">Amolops attiguus</emphasis>
</taxonomicName>
<emphasis id="65B3F2A18F4483B8D0B76FFE32410ABE" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
further differs from
<taxonomicName id="274B850DE38E8F2DDB5462F7950199DF" lsidName="A. sengae" pageId="0" pageNumber="931" rank="species" species="sengae">
<emphasis id="9C587F31AA7EB95417FD3B60C05F3A40" italics="true" pageId="0" pageNumber="931">A. sengae</emphasis>
</taxonomicName>
<emphasis id="B3B7A82B461C81ACF3EE1294BEB74610" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
by having spinose glands above the arm usually present (absent in
<taxonomicName id="75C04AD87D69C1009C562AEA915D7599" lsidName="A. sengae" pageId="0" pageNumber="931" rank="species" species="sengae">
<emphasis id="2681C3498BC3F7AA60E24C07F0CC126B" italics="true" pageId="0" pageNumber="931">A. sengae</emphasis>
</taxonomicName>
<emphasis id="0C676C60464C66AB517467DC17A20C1F" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
).
<taxonomicName id="70F0EB98A3556C63AD2A93FA6837112F" authorityName="Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart" authorityYear="2023" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops attiguus" order="Caudata" pageId="0" pageNumber="931" phylum="Chordata" rank="species" species="attiguus">
<emphasis id="6F1EC2B112D57DBBAF999E72E86FCF3B" italics="true" pageId="0" pageNumber="931">Amolops attiguus</emphasis>
</taxonomicName>
<emphasis id="892CFAE9A680B01D81BEB37805489CB5" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
further differs from
<taxonomicName id="30CAA4142BBB69F07211311C6A8FD39D" lsidName="A. cremnobatus" pageId="0" pageNumber="931" rank="species" species="cremnobatus">
<emphasis id="99C163FD6203E1B7BF8667A96B288C2E" italics="true" pageId="0" pageNumber="931">A. cremnobatus</emphasis>
</taxonomicName>
and
<taxonomicName id="55137447E80A332A63E248577DDD6E69" lsidName="A. sengae" pageId="0" pageNumber="931" rank="species" species="sengae">
<emphasis id="FC65FA411FECA3E608BC87ECB5C190A8" italics="true" pageId="0" pageNumber="931">A. sengae</emphasis>
</taxonomicName>
<emphasis id="92A4DD8DC0D2FAA65BAEBF2D5117112D" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
by having larger EYE and smaller TEY (Fig.
<figureCitation id="A97C51F4104B2EE55A00B089A7FAF7D0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Box and whisker plots of measurements (mm) of morphological features of males of Amolops cremnobatus (B; n = 12), A. sengae sp. nov. (C; n = 5), and A. attiguus sp. nov. (E; n = 5). Measurements are snout-vent length (SVL), snout length (SNT), eye diameter (EYE), tympanum-eye distance (TEY), tarsus length (SHK), and foot length (FTL). Boxes show interquartile range and median (thick black line). Whiskers represent range up to 1.5 x interquartile range (IQR); dots show range beyond 1.5 x IQR. Clusters with the same symbol (* and †) are not significantly different, as determined by paired two-tailed t-tests following ANOVA." figureDoi="10.3897/vz.73.e102475.figure11" httpUri="https://binary.pensoft.net/fig/916374" pageId="0" pageNumber="931">11</figureCitation>
).
<taxonomicName id="5A37D7F276CB6708E8A5CA599A62C31E" authorityName="Sheridan &amp; Phimmachak &amp; Sivongxay &amp; Stuart" authorityYear="2023" class="Amphibia" family="Ranidae" genus="Amolops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amolops attiguus" order="Caudata" pageId="0" pageNumber="931" phylum="Chordata" rank="species" species="attiguus">
<emphasis id="8902376F88D19481131C48061F184B36" italics="true" pageId="0" pageNumber="931">Amolops attiguus</emphasis>
</taxonomicName>
<emphasis id="A269F9E4492DE9D66227A6565ABEE020" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
further differs from
<taxonomicName id="02D14BD2BEDF7621EA604253F02F485F" lsidName="A. sengae" pageId="0" pageNumber="931" rank="species" species="sengae">
<emphasis id="F6249824937EC7D00C50E4D281E23824" italics="true" pageId="0" pageNumber="931">A. sengae</emphasis>
</taxonomicName>
<emphasis id="3FC2AD156B2061582035EF9F9CF32CCD" bold="true" pageId="0" pageNumber="931">sp. nov.</emphasis>
by having smaller SNT and FTL (Fig.
<figureCitation id="20777E83CBA5E1D09AC53761632C25EC" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Box and whisker plots of measurements (mm) of morphological features of males of Amolops cremnobatus (B; n = 12), A. sengae sp. nov. (C; n = 5), and A. attiguus sp. nov. (E; n = 5). Measurements are snout-vent length (SVL), snout length (SNT), eye diameter (EYE), tympanum-eye distance (TEY), tarsus length (SHK), and foot length (FTL). Boxes show interquartile range and median (thick black line). Whiskers represent range up to 1.5 x interquartile range (IQR); dots show range beyond 1.5 x IQR. Clusters with the same symbol (* and †) are not significantly different, as determined by paired two-tailed t-tests following ANOVA." figureDoi="10.3897/vz.73.e102475.figure11" httpUri="https://binary.pensoft.net/fig/916374" pageId="0" pageNumber="931">11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

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@ -0,0 +1,69 @@
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<treatment id="5EB08823FCE4D456C22DEF7B93B952D8" ID-DOI="http://doi.org/10.5281/zenodo.6287992" ID-GBIF-Taxon="100133488" ID-Zenodo-Dep="6287992" LSID="urn:lsid:plazi:treatment:5EB08823FCE4D456C22DEF7B93B952D8" httpUri="http://treatment.plazi.org/id/5EB08823FCE4D456C22DEF7B93B952D8" lastPageNumber="14" pageNumber="14">
<subSubSection id="52055D3E7369D82C8655F0823043E46F" type="nomenclature">
<paragraph id="28F75E034C9D8DDA73CE8E0C1D518F96" pageNumber="14">
2. Subgenus
<taxonomicName id="178A02824E7E5D932393C869AEDD9C20" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:230390" class="Insecta" family="Formicidae" genus="Anoplolepis" kingdom="Animalia" lsidName="Anoplolepis (Zealleyella) braunsi st. parsonsi Santschi" lsidName-HNS="Anoplolepis (Zealleyella) braunsi st. parsonsi Santschi" order="Hymenoptera" pageNumber="14" phylum="Arthropoda" rank="subGenus" subGenus="Zealleyella">Zealleyella</taxonomicName>
(Arnold) Santschi emend.
</paragraph>
</subSubSection>
<subSubSection id="51F8804446A3A2C4379997E46A05DB49" type="reference_group">
<paragraph id="90F328570D6F4AE4B7D717AC260621D7" pageNumber="14">Arnold, 1922, Ann. South. Afr'. Mus., p. 579.</paragraph>
</subSubSection>
<subSubSection id="1CDC161557553E0632E8C13443F3A7C8" type="description">
<paragraph id="F4B0BCEED00FF50634EAB82F3775AA37" pageNumber="14">
[[ worker ]]. Polymorphes, [[ male ]]. Aussi grand que la 9. Metanotum plus ou moins distinct chez les [[ worker ]] major. Antennes comme chez le s. - g.
<taxonomicName id="A7A027C6D6FCF3A4664F21CC2830C1BD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2349" class="Insecta" family="Formicidae" genus="Anoplolepis" kingdom="Animalia" lsidName="Anoplolepis Santschi" lsidName-HNS="Anoplolepis Santschi" order="Hymenoptera" pageNumber="14" phylum="Arthropoda" rank="genus">Anoplolepis</taxonomicName>
. Ocelles chez la [[ worker ]] major, manque chez [[ worker ]] minor.
</paragraph>
</subSubSection>
<subSubSection id="D9DFB634B6D96D3570C47AFB049AE122" type="nomenclature">
<paragraph id="D22B336EE77A82A229DB2464FA837329" pageNumber="14">
Type:
<taxonomicName id="3632C2D3F919C04741877AAA7C393389" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:230391" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" lsidName="Uploaded - Formica cutodiens" lsidName-HNS="Uploaded - Formica cutodiens" order="Hymenoptera" pageNumber="14" phylum="Arthropoda" rank="species" species="cutodiens">F. cutodiens Sm</taxonomicName>
. 1858.
</paragraph>
<paragraph id="D045FD30B6EAB1EA657B06237B40F258" pageNumber="14">
Autres especes
<taxonomicName id="8329B3573F1C5EC68992F2BC5FA0B84F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:133781" class="Insecta" family="Formicidae" genus="Plagiolepis" kingdom="Animalia" lsidName="Plagiolepis (Anoplolepis) rufescens Santschi" lsidName-HNS="Plagiolepis (Anoplolepis) rufescens Santschi" order="Hymenoptera" pageNumber="14" phylum="Arthropoda" rank="species" species="rufescens" subGenus="Anoplolepis">P. (A.) rufescens Sants</taxonomicName>
.,
<taxonomicName id="6A3F5DF10793DE3A5C7FD5DBAA2986E7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:133778" class="Insecta" family="Formicidae" genus="Plagiolepis" kingdom="Animalia" lsidName="Plagiolepis (Anoplolepis) nuptialis Santschi" lsidName-HNS="Plagiolepis (Anoplolepis) nuptialis Santschi" order="Hymenoptera" pageNumber="14" phylum="Arthropoda" rank="species" species="nuptialis">nuptialis</taxonomicName>
Sants.,
<taxonomicName id="E28C53DBF8F3D0EE779FB19698E751F9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:180289" class="Insecta" family="Formicidae" genus="Plagiolepis" kingdom="Animalia" lsidName="Plagiolepis steingroveri Forel" lsidName-HNS="Plagiolepis steingroveri Forel" order="Hymenoptera" pageNumber="14" phylum="Arthropoda" rank="species" species="steingroveri">Plagiolepis steingroveri Forel</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,116 @@
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<mods:title id="6376B6318FE3D8B5BB1F3FE1EEAB33AC">Order Soricomorpha</mods:title>
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<taxonomicName id="68B07DACB6A5082997F219DA626D0153" ID-CoL="7WW6P" authority="Bailey 1913" authorityName="Bailey" authorityYear="1913" class="Mammalia" family="Soricidae" genus="Sorex" kingdom="Animalia" order="Soricomorpha" pageNumber="293" phylum="Chordata" rank="species" species="neomexicanus" subGenus="Otisorex">
<emphasis id="820EAE90AE424F57C390C3E212940D57" italics="true" pageNumber="293">Sorex (Otisorex) neomexicanus</emphasis>
Bailey 1913
</taxonomicName>
</paragraph>
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<paragraph id="F9744501DD5B1D13156C70CB99DDC0F0" pageNumber="293">
<treatmentCitation id="0DDB4DBDBFE8B6B95178883BE5ED0FA9" author="Bailey" authority="Bailey 1913" authorityName="Bailey" authorityYear="1913" class="Mammalia" family="Soricidae" genus="Sorex" journal="Proc. Biol. Soc. Wash." kingdom="Animalia" order="Soricomorpha" page="133" pageNumber="293" phylum="Chordata" rank="species" species="neomexicanus" subGenus="Otisorex" title="Sorex (Otisorex) neomexicanus" volume="26" year="1913">
<taxonomicName id="2E5D59B7D5E503067E2BE75177BFF5D6" ID-CoL="7WW6P" authority="Bailey 1913" authorityName="Bailey" authorityYear="1913" class="Mammalia" family="Soricidae" genus="Sorex" kingdom="Animalia" order="Soricomorpha" pageNumber="293" phylum="Chordata" rank="species" species="neomexicanus" subGenus="Otisorex">
<emphasis id="17E73070FCBE6935CB8BC68C4CD7C811" italics="true" pageNumber="293">Sorex (Otisorex) neomexicanus</emphasis>
Bailey 1913
</taxonomicName>
,
<bibCitation id="139947B7559369C3F4608EC28D4F03CF" author="Bailey" journal="Proc. Biol. Soc. Wash." pageNumber="293" pagination="133" title="Sorex (Otisorex) neomexicanus" volume="26" year="1913">Proc. Biol. Soc. Wash., 26: 133</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ADDF5F445824B31CD68115DB7D43AED8" pageNumber="293" type="materials_examined">
<paragraph id="3773187CE4EC730DB596E380B4FEA435" pageNumber="293">
<emphasis id="78BEEE576F3C96F868FCFF2CD0CEF655" inLineHeading="true" pageNumber="293">Type Locality:</emphasis>
<materialsCitation id="B820D335F0B3BCA08023287E70DBF1ED" ID-GBIF-Occurrence="4867338538" country="United States of America" pageNumber="293" stateProvince="New Mexico">
<collectingCountry id="F7CEAB77C6D2C7054D55481C37257C48" name="United States of America">USA</collectingCountry>
,
<collectingRegion id="0093544CCB6D8B26AC55028A1AC7D603" country="United States of America" name="New Mexico">New Mexico</collectingRegion>
, Sacramento Mtns, Cloudcroft.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="063051849DD511F331E219C1916FDD83" pageNumber="293" type="vernacular_names">
<paragraph id="891B27E5F80BC5980F066E07FC3B390C" pageNumber="293">
<emphasis id="D05D04BB43BE67248487E35461579239" inLineHeading="true" pageNumber="293">Vernacular Names:</emphasis>
<vernacularName id="527BD848DB5EBB4C961178ADFD033B3E" language="eng" pageNumber="293">
<collectingRegion id="B0FFB78C911744FE599DBD74405084B4" country="United States of America" name="New Mexico">New Mexico</collectingRegion>
Shrew
</vernacularName>
.
</paragraph>
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<subSubSection id="167E2FC7528A156FDDDA57068D6A533A" pageNumber="293" type="distribution">
<paragraph id="334E2F30DEAA3D54CAF78B71C1DE989E" pageNumber="293">
<emphasis id="3F350A8704AC44F74B9457FA3B676F13" inLineHeading="true" pageNumber="293">Distribution:</emphasis>
<collectingCountry id="730C973B501CB04B7BA46BFC4A46173D" name="United States of America">USA</collectingCountry>
,
<collectingRegion id="D463D87C13230D673118796C82A4A21E" country="United States of America" name="New Mexico">New Mexico</collectingRegion>
, Capitan and Sacramento Mtns.
</paragraph>
</subSubSection>
<subSubSection id="022952E21DB777AE8CDE481AAC8B5BE2" pageNumber="293" type="discussion">
<paragraph id="BDA2069EDF01B9283E6A44ADA4AAFCCF" pageNumber="293">
<emphasis id="250F1757B3293FE9D3749AC9195926DB" inLineHeading="true" pageNumber="293">Discussion:</emphasis>
Subgenus
<taxonomicName id="903D407490C9289DDA852C13A5D13E28" baseAuthorityName="Diersing and Hoffmeister" baseAuthorityYear="1977" class="Mammalia" family="Soricidae" genus="Sorex" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="subGenus" subGenus="Otisorex">
<emphasis id="B249039B5DB29453904547FCCE966B81" italics="true" pageNumber="293">Otisorex</emphasis>
</taxonomicName>
.
<taxonomicName id="065D0CC85D29147C3F42F5445D1614A9" authorityName="Baird" authorityYear="1857" class="Mammalia" family="Soricidae" genus="Sorex" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="vagrans">
<emphasis id="598735811382226DCE8AE5B454BFBC43" italics="true" pageNumber="293">S. vagrans</emphasis>
</taxonomicName>
complex (
<bibRefCitation id="D9409ADEC0CA573D76694217577BDF23" author="Demboski, J. R. &amp; J. A. Cook" refId="ref80509" refString="Demboski, J. R., and J. A. Cook. 2001. Phylogeography of the dusky shrew, Sorex monticolus (Insectivora, Soricidae): Insight into deep and shallow history in northwestern North America. Molecular Ecology, 10: 1277 - 1240." year="2001">Demboski and Cook, 2001</bibRefCitation>
). Formerly included in
<taxonomicName id="131F45276DD4578C823F338FF2EBA45D" authorityName="Merriam" authorityYear="1890" class="Mammalia" family="Soricidae" genus="Sorex" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="monticolus">
<emphasis id="C0AE128AED8979DCC77F3CE271A445A3" italics="true" pageNumber="293">S. monticolus</emphasis>
</taxonomicName>
but given species rank by
<bibRefCitation id="C91DE14C7DEEEB8E363E698338DDFB68" author="Alexander, L. F." refId="ref4136" refString="Alexander, L. F. 1996. A morphometric analysis of geographic variation within Sorex monticolus (Insectivora: Soricidae). University of Kansas Natural History Museum Miscellaneous Publication, 88: 1 - 54." year="1996">Alexander (1996)</bibRefCitation>
based on morphometric differences.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="33C0ABCC0BA0034F17DC03708F8BAC8D" italics="true" pageNumber="196">Crassula tillaea</emphasis>
Lest.
</taxonomicName>
-Garl.
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<paragraph id="68161BA997B9DBC93DFA144431F2D1D3" pageNumber="196">
Artbeschreibung:
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1-5(-8) cm lang, verzweigt, niederliegend bis aufrecht,
<emphasis id="9112B711AAA31960E1BA798181E7B4AF" bold="true" pageNumber="196">
moosartig. Alle
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<normalizedToken id="BD6B34E6543C522AA2708DB9DC2CE837" originalValue="gegenständig" pageNumber="196">gegenstaendig</normalizedToken>
und
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, oft
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</emphasis>
,
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lang, in den Achseln mit 1-4 winzigen,
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3
<normalizedToken id="C5D4416FC17DBB07D78398B20EE72FF1" originalValue="zähligen" pageNumber="196">zaehligen</normalizedToken>
, sitzenden
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.
</emphasis>
<normalizedToken id="944187C046ABA1E88F7155C21587E4E9" originalValue="Staubblätter" pageNumber="196">Staubblaetter</normalizedToken>
3.
<normalizedToken id="14EB399FB80277C0E114CC2E17EC5C68" originalValue="Kronblätter" pageNumber="196">Kronblaetter</normalizedToken>
<normalizedToken id="D4C527D4C0649930A4DBD68D3A734000" originalValue="kürzer" pageNumber="196">kuerzer</normalizedToken>
als die
<normalizedToken id="C9F5DE66E5BFCDE0282B817255BE82EC" originalValue="Kelchblätter" pageNumber="196">Kelchblaetter</normalizedToken>
, weiss bis hellrosa.
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<paragraph id="C473B0F0A9078B2359339E9F15C29519" pageNumber="196">
<normalizedToken id="CFD004812969F89E85651E6C9359033F" originalValue="Blütezeit" pageNumber="196">Bluetezeit</normalizedToken>
: 4-7
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Standort und Verbreitung in der Schweiz: Sandige, wechselfeuchte Orte,
<normalizedToken id="B23BB30070475F5B30FD368F02BADDCE" originalValue="gestörte" pageNumber="196">gestoerte</normalizedToken>
Wiesen, Ruderalstellen / kollin / TI, GE
</paragraph>
</subSubSection>
<subSubSection id="4CE215E478FD8187990A6C92CEC3859B" pageNumber="196" type="distribution">
<paragraph id="429C363E78D106386CC2480604C54573" pageNumber="196">Verbreitung global: Mediterran-atlantisch</paragraph>
</subSubSection>
<subSubSection id="F08905A5E6AA812255CF967D37755974" pageNumber="196" type="biology_ecology">
<paragraph id="4252D57524DB580DB450BB7063132095" pageNumber="196">
<normalizedToken id="C2E1F4AC9725E6C024E61B0A8FDBA157" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="F7DFE99DC42A3D2095F4F709B5D9F6E2" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." pageNumber="196" publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="3F08E695E0CBEA501CF83D3EA1FDD5F9" pageNumber="196">
<table id="0B5FED63A154AA5F5197ADD9A41F0F1F" class="table" inLine="true" pageNumber="196">
<tr id="2D9D586C697F231ECCB79BA94FCE24FF" pageNumber="196">
<th id="9110E62A06168B91DEAE885E2E910D35" colspan="2" pageNumber="196">
<emphasis id="F6E59091D2A692791C0AF76CF8C386D5" bold="true" pageNumber="196">Bodenfaktoren</emphasis>
</th>
<th id="5AE2655E57C51AEE8B902AEA4391564F" colspan="2" pageNumber="196">
<emphasis id="42B78D9D0F0668A6375F873E70ADADA4" bold="true" pageNumber="196">Klimafaktoren</emphasis>
</th>
<th id="EE9D5EC965804DDF994758E89A2A5C0C" colspan="2" pageNumber="196">
<emphasis id="FE9F02C363AF0B6F06E613B1AE11F3F2" bold="true" pageNumber="196">Salztoleranz</emphasis>
</th>
</tr>
<tr id="267E7B513FC9DD6E2BFB38131454D6CF" pageNumber="196">
<td id="1D54F80204A84B2528D421C332F3B6C9" pageNumber="196">Feuchtezahl F</td>
<td id="B164AF97C4CAB53319CEA0B305708617" pageNumber="196" style="text-align:left;width:100px">--</td>
<td id="CBC5B05C862D4926A43A6405C5E5FC7F" pageNumber="196">Lichtzahl L</td>
<td id="3ABF650530F075EDA42DECAF5760D38D" pageNumber="196" style="text-align:left;width:60px">--</td>
<td id="4D22DDFF4FFB38D93E0BF3DCBA5C9A3F" pageNumber="196">Salzzeichen</td>
<td id="A3E84949602A1F2765D0EF405566894D" pageNumber="196" style="text-align:left;width:60px">--</td>
</tr>
<tr id="22D46549BFAFA4356561A3E945601156" pageNumber="196">
<td id="93FD252A0CAFA8E20A77864C39DFF479" pageNumber="196">Reaktionszahl R</td>
<td id="65692FB5E7411CC9EBE7FC939B949D4A" pageNumber="196" style="text-align:left">--</td>
<td id="E3225BC49863963EA5537667DBD00D0C" pageNumber="196">Temperaturzahl T</td>
<td id="7377CFA81489824EFF89B3BDD5832BAD" pageNumber="196" style="text-align:left">--</td>
</tr>
<tr id="A641F32E2DC9CA2ABB28827119477BF2" pageNumber="196">
<td id="5B70F6258EDF47B41A46B095D75636BA" pageNumber="196">
<normalizedToken id="BB6BE96E446DFBA1B7AC56E68CEBAD00" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="E033477B458627195523EFDFDB8E9866" pageNumber="196" style="text-align:left">--</td>
<td id="A56FFB6437CF5FB9DD42EC8DB8CABBB1" pageNumber="196">
<normalizedToken id="4471A6684D74980A6A88869D386FA2B5" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="1D6DC922DD87C2B8B15CE8A4D238B592" pageNumber="196" style="text-align:left">--</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="61EEFB4EFBD9BA040C08C2776A86C5FA" pageNumber="196" type="vernacular_names">
<paragraph id="7F0B67D0EEA6B6AB661A6BBF41F75975" pageNumber="196">
Volksname Deutscher Name:
<vernacularName id="A28046DA405179C11D027CC895C34810" language="deu" pageNumber="196">Moos-Dickblatt</vernacularName>
Nom
<normalizedToken id="86DDBBA04EB498DB51D9C2E4BF4C6BFA" originalValue="français">francais</normalizedToken>
:
<vernacularName id="FA678A95EDF5D7490CBEBD6D84543608" language="fra" pageNumber="196">Crassule mousse</vernacularName>
Nome italiano:
<vernacularName id="C1702B53BB484EC3141E33F6F3453B23" language="ita" pageNumber="196">Erba grassa muscosa</vernacularName>
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,300 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.243.3990" ID-GBIF-Dataset="443af630-ba06-4b78-8924-9896713e9cf1" ID-PMC="PMC3697044" ID-Pensoft-Pub="1313-2970-243-27" ID-PubMed="23818811" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-243-27" ModsDocOrigin="ZooKeys 243" ModsDocTitle="New neotropical species of Opiinae (Hymenoptera, Braconidae) reared from fruit-infesting and leaf-mining Tephritidae (Diptera) with comments on the Diachasmimorpha mexicana species group and the genera Lorenzopius and Tubiformopius" checkinTime="1451248541764" checkinUser="pensoft" docAuthor="Wharton, Robert, Ward, Lauren &amp; Miko, Istvan" docDate="2012" docId="5EB0C5BEFE3C8419163DBA19E77C9DC0" docLanguage="en" docName="ZooKeys 243: 27-82" docOrigin="ZooKeys 243" docSource="http://dx.doi.org/10.3897/zookeys.243.3990" docTitle="Eurytenes (Stigmatopoea) norrbomi Wharton, sp. n." docType="treatment" docVersion="4" lastPageNumber="59" masterDocId="A334154EFF8AFF9C0E2B8B29FFD7FB06" masterDocTitle="New neotropical species of Opiinae (Hymenoptera, Braconidae) reared from fruit-infesting and leaf-mining Tephritidae (Diptera) with comments on the Diachasmimorpha mexicana species group and the genera Lorenzopius and Tubiformopius" masterLastPageNumber="82" masterPageNumber="27" pageNumber="55" updateTime="1668154739309" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>New neotropical species of Opiinae (Hymenoptera, Braconidae) reared from fruit-infesting and leaf-mining Tephritidae (Diptera) with comments on the Diachasmimorpha mexicana species group and the genera Lorenzopius and Tubiformopius</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wharton, Robert</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ward, Lauren</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miko, Istvan</mods:namePart>
</mods:name>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>243</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>82</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.243.3990</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.243.3990</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-243-27</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152037973" LSID="urn:lsid:zoobank.org:act:FC57AAB3-6290-4076-BB44-6289F8ABF2EC" httpUri="http://treatment.plazi.org/id/5EB0C5BEFE3C8419163DBA19E77C9DC0" lastPageId="32" lastPageNumber="59" pageId="28" pageNumber="55">
<subSubSection pageId="28" pageNumber="55" type="nomenclature">
<paragraph pageId="28" pageNumber="55">
<taxonomicName LSID="urn:lsid:zoobank.org:act:FC57AAB3-6290-4076-BB44-6289F8ABF2EC" authority="Wharton" class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes (Stigmatopoea) norrbomi" order="Hymenoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="norrbomi" subGenus="Stigmatopoea">Eurytenes (Stigmatopoea) norrbomi Wharton</taxonomicName>
<taxonomicNameLabel pageId="28" pageNumber="55">sp. n.</taxonomicNameLabel>
Figs 43, 4547, 4951, 5357
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="55" type="type locality">
<paragraph pageId="28" pageNumber="55">Type locality.</paragraph>
<paragraph pageId="28" pageNumber="55">Mexico, Morelos, Km. 9-10 between Huitzilac and Lago Zempoala.</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="56" type="type material">
<paragraph pageId="29" pageNumber="56">
<pageBreakToken pageId="29" pageNumber="56" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="29" pageNumber="56">Holotype. Female (UNAM), first label, first line: MEXICO: Morelos second line: Km 9-10, btw. Huitzilac third line: &amp; Lago Zempoala fourth line: roadside, 22-24.ix.1991 fifth line: A. L. Norrbom #42</paragraph>
<paragraph pageId="29" pageNumber="56">
Paratypes: Mexico, 4 females, same data as holotype (TAMU, USNM); 1 female, Mexico, Rt. 890, Km 9 area, 6 km W Lago Zempoala 2.x.1991, Norrbom, #43, reared ex.
<taxonomicName class="Insecta" family="Tephritidae" genus="Trypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypeta concolor" order="Diptera" pageId="29" pageNumber="56" phylum="Arthropoda" rank="species" species="concolor">Trypeta concolor</taxonomicName>
ex. leafmines on
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Barkleyanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Barkleyanthus salicifolius" order="Asterales" pageId="29" pageNumber="56" phylum="Tracheophyta" rank="species" species="salicifolius">Barkleyanthus salicifolius</taxonomicName>
(91M1D) (TAMU). 3 males, Distrito Federal, Rt. 95 (libre), Km 42-43, 1 km N. La Cima, 20-26.ix.1991 A. L. Norrbom, #41, reared ex.
<taxonomicName class="Insecta" family="Tephritidae" genus="Trypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypeta concolor" order="Diptera" pageId="29" pageNumber="56" phylum="Arthropoda" rank="species" species="concolor">Trypeta concolor</taxonomicName>
ex. leafmines on
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Barkleyanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Barkleyanthus salicifolius" order="Asterales" pageId="29" pageNumber="56" phylum="Tracheophyta" rank="species" species="salicifolius">Barkleyanthus salicifolius</taxonomicName>
(91M1) (TAMU, USNM).
</paragraph>
</subSubSection>
<subSubSection lastPageId="32" lastPageNumber="59" pageId="29" pageNumber="56" type="description">
<paragraph pageId="29" pageNumber="56">Description.</paragraph>
<paragraph lastPageId="30" lastPageNumber="57" pageId="29" pageNumber="56">
Female. Head in dorsal view 1.2-1.3
<normalizedToken originalValue="×">x</normalizedToken>
broader than mesoscutum, 1.75-1.85
<normalizedToken originalValue="×">x</normalizedToken>
broader than face; eye in dorsal view 1.2-1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than temple, temples weakly receding behind eyes. Frons and vertex as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="29" pageNumber="56" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
except vertex and outer part
<pageBreakToken pageId="30" pageNumber="57" start="start">of</pageBreakToken>
occiput densely covered with long, decumbent setae; width of ocellar field 1.20-1.35
<normalizedToken originalValue="×">x</normalizedToken>
distance from ocellar field to eye. Face 1.75-1.85
<normalizedToken originalValue="×">x</normalizedToken>
wider than high; slightly less polished than frons; uniformly setose (as in Figs 51, 53), distinctly punctate, the punctures separated by about 1
<normalizedToken originalValue="×">x</normalizedToken>
their diameter. Frons and face delimited by a slightly more distinct change in sculpture in area between torulus and eye. Malar space about 0.6
<normalizedToken originalValue="×">x</normalizedToken>
basal width of mandible, 0.25
<normalizedToken originalValue="×">x</normalizedToken>
eye height. Clypeus 3.0-3.4
<normalizedToken originalValue="×">x</normalizedToken>
wider than high; protruding in profile. Occipital carina distinctly curved medially at dorsal end, absent mid-dorsally, the space where the carina is absent approximating width of ocellar field. Antenna 1.15-1.30
<normalizedToken originalValue="×longer">xlonger</normalizedToken>
than fore wing, with 31-33 flagellomeres; first flagellomere 1.05-1.10
<normalizedToken originalValue="×">x</normalizedToken>
longer than second, 1.05-1.20
<normalizedToken originalValue="×">x</normalizedToken>
longer than third; flagellomeres 3.1-4.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide basally, 2.3-2.7 longer than wide apically. Head otherwise as described for
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
.
</paragraph>
<paragraph pageId="30" pageNumber="57">
Mesosoma 1.35-1.45
<normalizedToken originalValue="×">x</normalizedToken>
longer than high; 1.8-1.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; 1.3-1.4
<normalizedToken originalValue="×">x</normalizedToken>
higher than wide. Pronotum dorsally as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
but with slightly enlarged pit in middle of crenulate groove consistently present; crenulae extending in shallow groove onto pronotum laterally, covering dorsal 0.2-0.6; groove margined anteriorly as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
. Anterior declivity of mesoscutum completely vertical, densely covered with white, decumbent setae except for bare median band extending posteriorly to midpit; anterior-lateral corners of mesoscutum at upper edge of declivity elevated, rounded, densely setose, the setal pattern extending in broad bands all along notauli and laterally from anterior declivity to tegula; notauli complete, extending from anterior margin to scuto-scutellar sulcus, weakly converging posteriorly alongside but not into tear-drop shaped midpit; crenulate throughout, with sculpture extending laterally around margin to tegula, sculpture largely obscured by dense setae; lateral lobes of mesoscutum bare posterior-medially. Scuto-scutellar sulcus 4-5
<normalizedToken originalValue="×">x</normalizedToken>
wider than midlength, lateral margins difficult to discern due to setal density; with low midridge and indistinct crenulae on either side; otherwise as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
. Scutellum as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
except with long marginal setae extending medially to cover most of posterior 0.5. Propodeum extensively rugulose, obscuring nearly all traces of carinae; pleural carina weak, often indistinct, very short median carina often present basally; transverse carina rarely weakly indicated across middle; propodeum uniformly setose anteriorly, with a few scattered setae posteriorly. Mesopleuron as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
except subalar region densely setose and groove below subalar ridge varying from nearly smooth to weakly rugulose. Precoxal sulcus distinctly impressed, unsculptured. Metapleuron a little more extensively setose but otherwise as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
.
</paragraph>
<paragraph pageId="30" pageNumber="57">
Wings. Fore wing stigma parallel-sided, discrete posteriorly, 6.3-6.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; r1 arising from basal 0.35; 1RS (excluding parastigma) 0.25-0.35
<normalizedToken originalValue="×">x</normalizedToken>
length of 1M; RS+M weakly sinuate; 3RSa 1.05-1.30
<normalizedToken originalValue="×">x</normalizedToken>
longer than 2RS; 2RS 2.6-3.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than r; 2RS and 3RSb straight; 3M variable, but often pigmented and sclerotized for most of its length; 2CUa 0.8-0.9
<normalizedToken originalValue="×">x</normalizedToken>
length of 2cu-a, 2CUb arising slightly above middle of first subdiscal cell; position of m-cu, 1cu-a, and 1-1A, shape of second submarginal cell, and angle between r1 and 2RS as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
. Hind wing as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="30" pageNumber="57" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
.
</paragraph>
<paragraph lastPageId="32" lastPageNumber="59" pageId="30" pageNumber="57">
Metasoma distinctly petiolate; head 3.75-4.10
<normalizedToken originalValue="×">x</normalizedToken>
wider than apex of T1. T1 2.2-2.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; nearly parallel-sided, with apex 1.20-1.35
<normalizedToken originalValue="×">x</normalizedToken>
wider than base; surface granular coriaceous throughout; completely without subapical depre
<pageBreakToken pageId="31" pageNumber="58" start="start">ssions</pageBreakToken>
dorsally; dorsope, laterope, dorsal carinae, dorsal surface of T1 in profile, as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="31" pageNumber="58" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
; lateral carina at least partially present but difficult to distinguish from surrounding sculpture. S1 extending about 0.25-0.30
<normalizedToken originalValue="×">x</normalizedToken>
length of T1; T2 and following without sharp lateral margins; spiracle of second metasomal terga laterally displaced, only partially visible in dorsal view. Ovipositor shorter than mesosoma, base not visible
<pageBreakToken pageId="32" pageNumber="59" start="start">in</pageBreakToken>
type series, but total length approximately 0.6-0.7
<normalizedToken originalValue="×">x</normalizedToken>
length of mesosoma; ovipositor sheath 0.30-0.35
<normalizedToken originalValue="×">x</normalizedToken>
length of mesosoma, with setal pattern as in
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
.
</paragraph>
<paragraph pageId="32" pageNumber="59">Color: Mesosoma, T1, S1, ovipositor sheath, and most of head dark brown to black; antenna yellow basally, apical 0.3 brown; mandibles yellow; palps white; lower gena adjacent malar sulcus brown to brownish red; ventral 0.3-0.4 of clypeus yellow to brownish red. Tegula reddish brown with yellow margin. Legs yellow to pale yellow except most of hind coxa, apical 0.6-0.7 of hind femur, and fifth tarsomere of all legs brown; hind tibia varying from weakly infumate to light brown, basal 0.2 nearly always pale yellow. T2 mostly brownish red with median yellow blotch posteriorly; T3 yellow with anterior and lateral margins brownish red; T4-T6 yellow with anterior and lateral margins dark brown; visible parts of remaining terga yellow.</paragraph>
<paragraph pageId="32" pageNumber="59">
Male. As in female except antenna with 37 flagellomeres; eye in dorsal view 1.55-1.75
<normalizedToken originalValue="×">x</normalizedToken>
longer than temple; width of ocellar field 1.05-1.10
<normalizedToken originalValue="×">x</normalizedToken>
distance from ocellar field to eye. Color same except visible parts of apical terga dark brown.
</paragraph>
<paragraph pageId="32" pageNumber="59">Body length 2.8-3.5 mm; wing length 3.2-3.6 mm.</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="59" type="diagnosis">
<paragraph pageId="32" pageNumber="59">Diagnosis.</paragraph>
<paragraph pageId="32" pageNumber="59">
This species shares with
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
and
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes macrocerus" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="macrocerus">Eurytenes macrocerus</taxonomicName>
the diagnostic features noted above for
<taxonomicName class="Insecta" family="Braconidae" genus="Stigmatopoea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stigmatopoea" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Stigmatopoea</taxonomicName>
.
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes norrbomi" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="norrbomi">Eurytenes norrbomi</taxonomicName>
is most readily differentiated from
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
on the basis of the more densely setose head and body (Figs 43, 47, 49), particularly the vertex, occiput, and mesoscutum, and the more extensively rugose propodeum. It also has a shorter ovipositor than
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
(Fig. 45 vs. Fig. 42). The setal pattern on the mesoscutum also differentiates
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes norrbomi" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="norrbomi">Eurytenes norrbomi</taxonomicName>
from
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes macrocerus" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="macrocerus">Eurytenes macrocerus</taxonomicName>
. The latter has shorter setae that are more sparsely distributed laterally (Fig. 54).
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="59" type="biology">
<paragraph pageId="32" pageNumber="59">Biology.</paragraph>
<paragraph pageId="32" pageNumber="59">
Four of the specimens from the type series were reared from puparia of
<taxonomicName class="Insecta" family="Tephritidae" genus="Trypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypeta concolor" order="Diptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="concolor">Trypeta concolor</taxonomicName>
(Wulp) (
<taxonomicName family="Tephritidae" lsidName="" pageId="32" pageNumber="59" rank="family">Tephritidae</taxonomicName>
) mining leaves of
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Barkleyanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Barkleyanthus salicifolius" order="Asterales" pageId="32" pageNumber="59" phylum="Tracheophyta" rank="species" species="salicifolius">Barkleyanthus salicifolius</taxonomicName>
(H.B.K.) H. Robins &amp; Brett (
<taxonomicName genus="Asteraceae" lsidName="Asteraceae" pageId="32" pageNumber="59" rank="genus">Asteraceae</taxonomicName>
). The remaining specimens were collected from flowers of this same plant together with
<taxonomicName class="Insecta" family="Tephritidae" genus="Trypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypeta concolor" order="Diptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="concolor">Trypeta concolor</taxonomicName>
and
<taxonomicName class="Insecta" family="Tephritidae" genus="Trypeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypeta reducta" order="Diptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="reducta">Trypeta reducta</taxonomicName>
Han and Norrbom. See
<bibRefCitation author="Han, H-Y" journalOrPublisher="Systematic Entomology" pageId="45" pageNumber="72" pagination="208 - 247" title="A systematic revision of the New World species of Trypeta Meigen (Diptera: Tephritidae)." url="10.1111/j.1365-3113.2005.00268.x" volume="30" year="2005">Han and Norrbom (2005)</bibRefCitation>
for details on the hosts and the collecting localities.
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="59" type="etymology">
<paragraph pageId="32" pageNumber="59">Etymology.</paragraph>
<paragraph pageId="32" pageNumber="59">This species is named after the collector, Allen Norrbom, who has provided many valuable host records for tephritid parasitoids.</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="59" type="remarks">
<paragraph pageId="32" pageNumber="59">Remarks.</paragraph>
<paragraph pageId="32" pageNumber="59">
This species attacks leaf-mining tephritids, as does
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes macrocerus" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="macrocerus">Eurytenes macrocerus</taxonomicName>
, while
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes maya" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="maya">Eurytenes maya</taxonomicName>
attacks fruit-infesting tephritids. Despite the difference in host habitat, all three species share many morphological features, and readily fit the characterization of
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Eurytenes</taxonomicName>
(
<taxonomicName class="Insecta" family="Braconidae" genus="Stigmatopoea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stigmatopoea" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Stigmatopoea</taxonomicName>
) as given above.
</paragraph>
<caption pageId="32" pageNumber="59">
<paragraph pageId="32" pageNumber="59">
Figures 50-53.
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Eurytenes</taxonomicName>
(
<taxonomicName class="Insecta" family="Braconidae" genus="Stigmatopoea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stigmatopoea" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Stigmatopoea</taxonomicName>
) spp. 50ES.) maya Wharton sp. n., paratype female, face, frontal view 51ES.) norrbomi Wharton sp. n., holotype female, face, frontal view 52ES.) maya, paratype female, face, slightly deflected 53ES.) norrbomi, holotype female, face, slightly deflected.
</paragraph>
</caption>
<caption pageId="32" pageNumber="59">
<paragraph pageId="32" pageNumber="59">
Figures 54-57.
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Eurytenes</taxonomicName>
(
<taxonomicName class="Insecta" family="Braconidae" genus="Stigmatopoea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stigmatopoea" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Stigmatopoea</taxonomicName>
) and
<taxonomicName class="Insecta" family="Braconidae" genus="Lorenzopius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lorenzopius" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Lorenzopius</taxonomicName>
. 54ES.) macrocerus (Thomson), mesosoma and metasoma, dorsal view 55
<taxonomicName class="Insecta" family="Braconidae" genus="Lorenzopius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lorenzopius calycomyzae" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="calycomyzae">Lorenzopius calycomyzae</taxonomicName>
van Achterberg and Salvo, holotype female, T1, dorsal view 56ES.) maya Wharton sp. n., paratype female, propodeum and T1, dorsal view 57ES.) norrbomi Wharton sp. n., holotype female, mesosoma and metasoma, dorsal view.
</paragraph>
</caption>
<caption pageId="32" pageNumber="59">
<paragraph pageId="32" pageNumber="59">
Figures 58-61.
<taxonomicName class="Insecta" family="Braconidae" genus="Lorenzopius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lorenzopius" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Lorenzopius</taxonomicName>
and
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Eurytenes</taxonomicName>
(
<taxonomicName class="Insecta" family="Braconidae" genus="Stigmatopoea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stigmatopoea" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="genus">Stigmatopoea</taxonomicName>
). 58
<taxonomicName class="Insecta" family="Braconidae" genus="Lorenzopius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lorenzopius calycomyzae" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="calycomyzae">Lorenzopius calycomyzae</taxonomicName>
van Achterberg and Salvo, holotype female, face 59ES.) maya Wharton sp. n., propodeum and T1 60
<taxonomicName class="Insecta" family="Braconidae" genus="Lorenzopius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lorenzopius calycomyzae" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="calycomyzae">Lorenzopius calycomyzae</taxonomicName>
, propodeum 61
<taxonomicName class="Insecta" family="Braconidae" genus="Lorenzopius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lorenzopius calycomyzae" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="calycomyzae">Lorenzopius calycomyzae</taxonomicName>
, head and mesoscutum, dorsal view.
</paragraph>
</caption>
<caption pageId="32" pageNumber="59">
<paragraph pageId="32" pageNumber="59">
Figures 62-65. Opiinae. 62
<taxonomicName class="Insecta" family="Braconidae" genus="Lorenzopius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lorenzopius calycomyzae" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="calycomyzae">Lorenzopius calycomyzae</taxonomicName>
van Achterberg and Salvo, holotype female, wings 63
<taxonomicName class="Insecta" genus="Tubiformopius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tubiformopius tubigaster" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="tubigaster">Tubiformopius tubigaster</taxonomicName>
(Fischer), holotype male, lateral view showing T1, S1, and wing base 64
<taxonomicName class="Insecta" family="Braconidae" genus="Eurytenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurytenes (Stigmatopoea) maya" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="maya" subGenus="Stigmatopoea">Eurytenes (Stigmatopoea) maya</taxonomicName>
Wharton, sp. n., paratype male, fore wing 65
<taxonomicName class="Insecta" genus="Tubiformopius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tubiformopius tubigaster" order="Hymenoptera" pageId="32" pageNumber="59" phylum="Arthropoda" rank="species" species="tubigaster">Tubiformopius tubigaster</taxonomicName>
, holotype male, habitus.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>