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<paragraph id="110B2577268E9539493E2A192DFABBC7" pageId="14" pageNumber="65">Taxon classification Animalia Araneae Agelenidae</paragraph>
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<taxonomicName id="93A360DE91F25F0EC32784E7131654D2" ID-CoL="4XKTS" authority="Dankittipakul &amp; Wang, 2003" authorityName="Dankittipakul &amp; Wang" authorityYear="2003" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes thailandensis" order="Araneae" pageId="14" pageNumber="65" phylum="Arthropoda" rank="species" species="thailandensis">Sinocoelotes thailandensis (Dankittipakul &amp; Wang, 2003)</taxonomicName>
<taxonomicNameLabel id="B36CB6FD43FCFC9709DD92F4811C667A" pageId="14" pageNumber="65">comb. n.</taxonomicNameLabel>
Figs 18, 19, 21
</paragraph>
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<paragraph id="F80C5721016F3AFA954EB00904B27126" pageId="15" pageNumber="66">
<taxonomicName id="E2644EE3CD2AC87D454B2FAB3AC4AF55" class="Arachnida" family="Agelenidae" genus="Coelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelotes thailandensis" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="thailandensis">
<pageBreakToken id="845B1A4F9DB0DB141ECEFE10CCF96A48" pageId="15" pageNumber="66" start="start">Coelotes</pageBreakToken>
thailandensis
</taxonomicName>
<bibRefCitation id="7D00A72E68EA0AC6C19CC4FD8C5C5AF9" author="Dankittipakul, P" journalOrPublisher="Revue Suisse de Zoologie" pageId="18" pageNumber="69" pagination="723 - 737" title="New species of coelotine spiders (Araneae, Amaurobiidae) from northern Thailand I." url="10.5962/bhl.part.80208" volume="110" year="2003">Dankittipakul and Wang 2003</bibRefCitation>
: 735, figs 24-25 (♂ holotype from Thailand, in MHNG, not examined);
<bibRefCitation id="E54FD87724E85706CD96CFFC76C5C960" author="Dankittipakul, P" journalOrPublisher="Zootaxa" pageId="18" pageNumber="69" pagination="1 - 11" title="New and poorly known species of coelotine spiders (Araneae, Amaurobiidae) from Thailand." volume="970" year="2005">Dankittipakul et al. 2005</bibRefCitation>
: 7, figs 9-10 (♂♀);
<bibRefCitation id="C34FFEEADC7F6BB7751A7DE922E0E6F6" author="Wang, XP" journalOrPublisher="Zootaxa" pageId="19" pageNumber="70" pagination="1 - 34" title="On the pseudoterrestris species group of the spider genus Coelotes (Araneae, Amaurobiidae)." volume="2313" year="2009">Wang et al. 2009</bibRefCitation>
: 26, f. 128-142 (♂♀).
</paragraph>
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<subSubSection id="DDB880BDB3966C269C4AA119C2401451" pageId="15" pageNumber="66" type="materials_examined">
<paragraph id="51BAC557C8B710E103CB9907A0A7B91B" pageId="15" pageNumber="66">Material examined.</paragraph>
<paragraph id="5B7984AB71DDACA3D1B4B12B7DDC9629" pageId="15" pageNumber="66">
1♂2♀: Thailand: Chiangmai Province: Mae Cham District, Jeep tract,
<geoCoordinate id="670844FBC12876A4172C5D6F99B0D085" direction="north" orientation="latitude" precision="15" value="18.528055">N18°31'41&quot;</geoCoordinate>
,
<geoCoordinate id="2029D9E54F8D813D487796F3A33A997C" direction="east" orientation="longitude" precision="15" value="98.49944">E98°29'58&quot;</geoCoordinate>
, 1649 m, 14.X.2014, H. Zhao, Y. Li and Z. Chen.
</paragraph>
</subSubSection>
<subSubSection id="582E049ED077B20B78D3E8F106D91D1D" pageId="15" pageNumber="66" type="diagnosis">
<paragraph id="31E92B058AF03F09FBC36CF13A8FAA3A" pageId="15" pageNumber="66">Diagnosis.</paragraph>
<paragraph id="917C60DCFB61F8D10173237FCC6CB8D2" pageId="15" pageNumber="66">
The species is similar to
<taxonomicName id="2335D9715290473642316C54ED0EFD9D" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes hehuaensis" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="hehuaensis">Sinocoelotes hehuaensis</taxonomicName>
sp. n., but male can be easily distinguished by a shorter and broader conductor (about 1/3 length of the conductor in
<taxonomicName id="5F7CCB0A3AFC0EA848CDF08BC7EC1BE2" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes hehuaensis" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="hehuaensis">Sinocoelotes hehuaensis</taxonomicName>
sp. n.), the broad and wedge-shaped dorsal conductor apophysis (cf. Figs 18
<normalizedToken id="31A0731E5F7DDAC74754D980DF62E197" originalValue="A–C">A-C</normalizedToken>
and 7
<normalizedToken id="AE214779AC3E78BB7D3D10929FFF4650" originalValue="A–C">A-C</normalizedToken>
). The female can be distinguished from that of
<taxonomicName id="B2F8C5617DFA6FD5F9B0522BE5490689" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes hehuaensis" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="hehuaensis">Sinocoelotes hehuaensis</taxonomicName>
sp. n. by the broad (almost round) atrium, the broader and shorter copulatory ducts, the shorter spermathecal heads (about 1/3 length of the spermathecal heads
<taxonomicName id="F3F27CF095FCD356413E7D19121144A9" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes hehuaensis" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="hehuaensis">Sinocoelotes hehuaensis</taxonomicName>
sp. n.) (cf. Figs 19
<normalizedToken id="ABF521FCDD44ECFF26FC680F5FFA0F80" originalValue="A–B">A-B</normalizedToken>
and 8
<normalizedToken id="5A50BAF25764F8A131ACE5E53126E9F3" originalValue="A–B">A-B</normalizedToken>
).
</paragraph>
<caption id="8F1A9C88ABC952291FD2E52A7CE36EC2" pageId="15" pageNumber="66">
<paragraph id="25D14BD3CB6C71599B97D475D93731E2" pageId="15" pageNumber="66">
Figure 18. Male palp of
<taxonomicName id="64327C61B9D9B60E941A43FD8AE0E9F1" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes thailandensis" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="thailandensis">Sinocoelotes thailandensis</taxonomicName>
. A Prolateral B Ventral C Retrolateral. Scale bar: equal for A, B and C.
</paragraph>
</caption>
<caption id="B938FBE7C23BDA7327B06830BDBCDCA0" pageId="15" pageNumber="66">
<paragraph id="DB6EA061AB9E6EA66D8A7BA0E0444E6D" pageId="15" pageNumber="66">
Figure 19. Epigyne and habitus of
<taxonomicName id="0BBD9EC4235F7121D55ABF9910AECC30" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes thailandensis" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="thailandensis">Sinocoelotes thailandensis</taxonomicName>
. A Epigyne, ventral B Vulva, dorsal C Male habitus, dorsal D Female habitus, dorsal E Female habitus, ventral. Scale bars: equal for A and B; equal for C, D and E.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="71739417A773D3CB120C9A2DA93FEA31" pageId="15" pageNumber="66" type="description">
<paragraph id="32C369874C5CEBE1BF6EF38C287C961B" pageId="15" pageNumber="66">Description.</paragraph>
<paragraph id="904E0EFE5989772B69C107C3DA202113" pageId="15" pageNumber="66">
Described by
<bibRefCitation id="B6751EA06B4C17A8D9FBC8888D267824" author="Wang, XP" journalOrPublisher="Zootaxa" pageId="19" pageNumber="70" pagination="1 - 34" title="On the pseudoterrestris species group of the spider genus Coelotes (Araneae, Amaurobiidae)." volume="2313" year="2009">Wang et al. (2009)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FF072445253233BFB1CF6C7D708EB9D9" pageId="15" pageNumber="66" type="comments">
<paragraph id="6C432361C95319B3771B549297BBF6D2" pageId="15" pageNumber="66">Comments.</paragraph>
<paragraph id="27F0910A80F2268991E53806869D703E" pageId="15" pageNumber="66">
The species shares a combination of somatic morphology characters with
<taxonomicName id="33863748537BB56509F1AC3262B5B954" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes hehuaensis" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="hehuaensis">Sinocoelotes hehuaensis</taxonomicName>
sp. n., and therefore was assigned to
<taxonomicName id="4D563634E22D6473E0A1A417D4E00679" class="Arachnida" family="Agelenidae" genus="Sinocoelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinocoelotes" order="Araneae" pageId="15" pageNumber="66" phylum="Arthropoda" rank="genus">Sinocoelotes</taxonomicName>
gen. n. The molecular analysis supports this transfer.
</paragraph>
</subSubSection>
<subSubSection id="029B5AA7356100400E8B6CBC5938762C" pageId="15" pageNumber="66" type="distribution">
<paragraph id="754B0CEB9B0A52E524EC15C8E46FC84D" pageId="15" pageNumber="66">Distribution.</paragraph>
<paragraph id="1F95CDC43B53EFF4997959FCAF2C560D" pageId="15" pageNumber="66">China (Yunnan) (Fig. 21).</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName authority="Sseliwanoff, 1884" authorityName="Sseliwanoff" authorityYear="1884" class="Chilopoda" family="Linotaeniidae" genus="Strigamia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strigamia pusilla" order="Lithobiomorpha" pageId="15" pageNumber="234" phylum="Arthropoda" rank="species" species="pusilla">Strigamia pusilla (Sseliwanoff, 1884)</taxonomicName>
</paragraph>
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<paragraph pageId="15" pageNumber="234">
<taxonomicName class="Chilopoda" family="Linotaeniidae" genus="Strigamia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strigamia pusilla" order="Lithobiomorpha" pageId="15" pageNumber="234" phylum="Arthropoda" rank="species" species="pusilla">Strigamia pusilla</taxonomicName>
-
<bibRefCitation pageId="15" pageNumber="234">Nefediev et al. 2017c</bibRefCitation>
: 13;
<bibRefCitation author="Nefediev, PS" journalOrPublisher="Arthropoda Selecta" pageId="29" pageNumber="248" pagination="217 - 224" title="A preliminary review of the centipede fauna of the Altai State Nature Biosphere Reserve, southwestern Siberia, Russia (Chilopoda: Lithobiomorpha, Geophilomorpha)." volume="26" year="2017 d">2017d</bibRefCitation>
: 223, 222: map.
</paragraph>
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<subSubSection pageId="15" pageNumber="234" type="materials_examined">
<paragraph pageId="15" pageNumber="234">Material examined</paragraph>
<paragraph pageId="15" pageNumber="234">(all from Russia, southwestern Siberia, Altai Province, Charysh District, ca. 4.5 km SE of Charyshskoye Village). 1 ♂, 1 juv. (ZMUM), 1 ♂ (ASU), site 1 on N slope, soil sample 5 (0-10 cm deep), 2.06.2016; 1 ♀ (ASU), site 1 on N slope, soil sample 1 (0-10 cm deep), 23.08.2016; 1 juv. (ASU), site 2 on N slope, soil sample 1 (0-10 cm deep), 23.08.2016, all leg. P.N., Kh.N., S.N., V.S.</paragraph>
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<paragraph pageId="15" pageNumber="234">Distribution.</paragraph>
<paragraph pageId="15" pageNumber="234">
Central-Palearctic temperate range: widespread from Central Europe and the Caucasus,
<taxonomicName lsidName="S. pusilla" pageId="15" pageNumber="234" rank="species" species="pusilla">S. pusilla</taxonomicName>
is found in the Urals, SW and central Siberia and N Mongolia (
<bibRefCitation author="Bonato, L" journalOrPublisher="Zootaxa" pageId="27" pageNumber="246" pagination="1 - 39" title="Species diversity of Strigamia Gray, 1843 (Chilopoda: Linotaeniidae): a preliminary synthesis." volume="3593" year="2012">Bonato et al. 2012</bibRefCitation>
;
<bibRefCitation author="Poloczek, A" journalOrPublisher="European Journal of Soil Biology" pageId="30" pageNumber="249" pagination="114 - 120" title="The Chilopoda (Myriapoda) of the Khentey-Mountain Range, Northern Mongolia. Communities of different forest-types under a varying fire regime." url="https://doi.org/10.1016/j.ejsobi.2016.04.004" volume="74" year="2016">Poloczek et al. 2016</bibRefCitation>
;
<bibRefCitation pageId="15" pageNumber="234">Nefediev et al. 2017c</bibRefCitation>
,
<bibRefCitation author="Nefediev, PS" journalOrPublisher="Arthropoda Selecta" pageId="29" pageNumber="248" pagination="217 - 224" title="A preliminary review of the centipede fauna of the Altai State Nature Biosphere Reserve, southwestern Siberia, Russia (Chilopoda: Lithobiomorpha, Geophilomorpha)." volume="26" year="2017 d">d</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="234" type="remarks">
<paragraph pageId="15" pageNumber="234">Remarks.</paragraph>
<paragraph pageId="15" pageNumber="234">In the study area, the species was found rarely and on the northern slope only.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Nahuatlea: a new genus of compositae (Gochnatieae) from North America</mods:title>
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<mods:namePart>Funk, Vicki A.</mods:namePart>
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<mods:namePart>Sancho, Gisela</mods:namePart>
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<mods:namePart>Roque, Nadia</mods:namePart>
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<treatment ID-GBIF-Taxon="139337058" LSID="urn:lsid:plazi:treatment:5EAAEF43924DB62B0FD885FD189CEE6A" httpUri="http://treatment.plazi.org/id/5EAAEF43924DB62B0FD885FD189CEE6A" lastPageId="13" lastPageNumber="116" pageId="10" pageNumber="115">
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<paragraph pageId="10" pageNumber="115">
5.
<taxonomicName LSID="00AA1569-48CD-50FA-8646-ABEBEEE1BC64" authority="(S. F. Blake) V. A. Funk" authorityName="V. A. Funk" authorityYear="2017" baseAuthorityName="S. F. Blake" baseAuthorityYear="2017" class="Magnoliopsida" family="Compositae" genus="Nahuatlea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nahuatlea obtusata" order="Asterales" pageId="10" pageNumber="115" phylum="Tracheophyta" rank="species" species="obtusata" status="comb. nov.">Nahuatlea obtusata (S.F. Blake) V.A.Funk</taxonomicName>
<taxonomicNameLabel pageId="10" pageNumber="115">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Images of four type specimens: A Nahuatlea arborescens (UC holotype) B N. smithii (K lectotype) C N. obtusata (US holotype) D N. hypoleuca (G holotype). [Furnished by the respective herbaria.]" figureDoi="10.3897/phytokeys.91.21340.figure3" httpUri="https://binary.pensoft.net/fig/174646" pageId="10" pageNumber="115">Figs 3D</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cabrera's drawing of Nahuatlea hypoleuca. A Habit B Head C Floret D Style E Anther F Pubescence. [Modified from Cabrera 1971.]" figureDoi="10.3897/phytokeys.91.21340.figure7" httpUri="https://binary.pensoft.net/fig/174650" pageId="10" pageNumber="115">, 7</figureCitation>
</paragraph>
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<paragraph pageId="11" pageNumber="116">
<taxonomicName authorityName="S. F. Blake" authorityYear="1924" class="Magnoliopsida" family="Compositae" genus="Gochnatia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Gochnatia obtusata" order="Asterales" pageId="11" pageNumber="116" phylum="Tracheophyta" rank="species" species="obtusata">
<pageBreakToken pageId="11" pageNumber="116" start="start">Gochnatia</pageBreakToken>
obtusata
</taxonomicName>
S.F. Blake, Contributions from the United States National Herbarium 22: 652. 1924.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="116" type="materials_examined">
<paragraph pageId="11" pageNumber="116">Type.</paragraph>
<paragraph pageId="11" pageNumber="116">
Mexico. Puebla: dry limestone hills at Tehuacan, 20 Dec 1895,
<emphasis italics="true" pageId="11" pageNumber="116">C.G. Pringle 6253</emphasis>
(Holotype: US 00119530 [254705]; Isotypes: A 00008380, BM 000947903, BR 0000005318124, CM 2403, E 00413002, JE 00000693, K 000502539, GH 00008381, MEXU 01220107, MO 1544305, NDG 63791, NY 00169557, P 00703351, P 00703352, PH 00025961, S 10-11705, TEX 00000454, US 01100608). [
<bibRefCitation author="Cabrera, A" journalOrPublisher="Revista del Museo de la Plata, Seccion Botanica" pageId="16" pageNumber="121" pagination="1 - 160" refId="B4" refString="Cabrera, A, 1971. Revision del Genero Gochnatia. Revista del Museo de la Plata, Seccion Botanica 12 (66): 1 - 160" title="Revision del Genero Gochnatia." volume="12" year="1971">Cabrera (1971)</bibRefCitation>
mentions additional specimens at F, G, M, UC, and W but these were not found in JSTOR-GP.]
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="116" type="description">
<paragraph pageId="11" pageNumber="116">Description.</paragraph>
<paragraph pageId="11" pageNumber="116">
<emphasis italics="true" pageId="11" pageNumber="116">Shrub</emphasis>
1.0
<normalizedToken originalValue="–1.5(–">-1.5(-</normalizedToken>
2.6) m tall;
<emphasis italics="true" pageId="11" pageNumber="116">leaves</emphasis>
coriaceous, (4.5)5-8(20)
<normalizedToken originalValue="×">x</normalizedToken>
3-5 mm, broadly elliptic, apex rounded to obtuse, rounded at base, glabrous adaxially, cinereous-tomentose abaxially; heads short-pedunculate (ca. 2 mm long), in clusters of 10-20 heads, at apices of branches;
<emphasis italics="true" pageId="11" pageNumber="116">involucre</emphasis>
narrowly obconic, 6-8 mm tall
<normalizedToken originalValue="×">x</normalizedToken>
2.8-3.2 mm wide, bracts in ca. 5 series, glabrous;
<emphasis italics="true" pageId="11" pageNumber="116">flowers</emphasis>
5-6 per head; corollas white, 6.5-7.5 mm long;
<emphasis italics="true" pageId="11" pageNumber="116">anther</emphasis>
base caudate, tails ca. 2 mm long, fimbrillate;
<emphasis italics="true" pageId="11" pageNumber="116">pappus</emphasis>
6.0-7.5 mm long, bristles biseriate, broadening slightly at the tips.
</paragraph>
<paragraph pageId="11" pageNumber="116">
<bibRefCitation author="Jervis, RN" journalOrPublisher="8. Eudicots-Asterales. Springer-Verlag, Berlin, Heidelberg" pageId="16" pageNumber="121" refId="B12" refString="Jervis, RN, 1954. A summary of the genus Gochnatia including a revision of the West Indian species which comprise the Section Anastraphioides. University of Michigan: unpublished thesis." title="A summary of the genus Gochnatia including a revision of the West Indian species which comprise the Section Anastraphioides. University of Michigan: unpublished thesis." year="1954">Jervis (1954)</bibRefCitation>
said that the leaf length is &quot;rarely up to 2 cm&quot; and the width is &quot;less than 1 cm&quot; and
<bibRefCitation author="Cabrera, A" journalOrPublisher="Revista del Museo de la Plata, Seccion Botanica" pageId="16" pageNumber="121" pagination="1 - 160" refId="B4" refString="Cabrera, A, 1971. Revision del Genero Gochnatia. Revista del Museo de la Plata, Seccion Botanica 12 (66): 1 - 160" title="Revision del Genero Gochnatia." volume="12" year="1971">Cabrera (1971)</bibRefCitation>
lists the length as 4-20
<normalizedToken originalValue="×">x</normalizedToken>
3-13 mm. Both seem to be correct; we found only one specimen that had leaves 20 mm long and all but two were less than 1 cm.
</paragraph>
<caption ID-Zenodo-Dep="1138377" doi="10.3897/phytokeys.91.21340.figure7" httpUri="https://binary.pensoft.net/fig/174650" pageId="11" pageNumber="116" start="Figure 7" startId="F7">
<paragraph pageId="11" pageNumber="116">
<emphasis bold="true" pageId="11" pageNumber="116">Figure 7.</emphasis>
<normalizedToken originalValue="Cabrera’s">Cabrera's</normalizedToken>
drawing of
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<emphasis italics="true" pageId="11" pageNumber="116">Nahuatlea hypoleuca</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="11" pageNumber="116">A</emphasis>
Habit
<emphasis bold="true" pageId="11" pageNumber="116">B</emphasis>
Head
<emphasis bold="true" pageId="11" pageNumber="116">C</emphasis>
Floret
<emphasis bold="true" pageId="11" pageNumber="116">D</emphasis>
Style
<emphasis bold="true" pageId="11" pageNumber="116">E</emphasis>
Anther
<emphasis bold="true" pageId="11" pageNumber="116">F</emphasis>
Pubescence. [Modified from
<bibRefCitation author="Cabrera, A" journalOrPublisher="Revista del Museo de la Plata, Seccion Botanica" pageId="16" pageNumber="121" pagination="1 - 160" refId="B4" refString="Cabrera, A, 1971. Revision del Genero Gochnatia. Revista del Museo de la Plata, Seccion Botanica 12 (66): 1 - 160" title="Revision del Genero Gochnatia." volume="12" year="1971">Cabrera 1971</bibRefCitation>
.]
</paragraph>
</caption>
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<subSubSection pageId="11" pageNumber="116" type="distribution">
<paragraph pageId="11" pageNumber="116">Distribution.</paragraph>
<paragraph pageId="11" pageNumber="116">Mexico: Puebla and Oaxaca.</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="118" pageId="11" pageNumber="116" type="remarks">
<paragraph pageId="11" pageNumber="116">Remarks.</paragraph>
<paragraph lastPageId="13" lastPageNumber="118" pageId="11" pageNumber="116">
<taxonomicName authorityName="S. F. Blake" authorityYear="1924" class="Magnoliopsida" family="Asteraceae" genus="Gochnatia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Gochnatia obtusata" order="Asterales" pageId="11" pageNumber="116" phylum="Tracheophyta" rank="species" species="obtusata">
<emphasis italics="true" pageId="11" pageNumber="116">Gochnatia obtusata</emphasis>
</taxonomicName>
was described by S. F.
<bibRefCitation author="Blake, SF" journalOrPublisher="Contributions from the United States National Herbarium" pageId="16" pageNumber="121" pagination="587 - 661" publicationUrl="https://biodiversitylibrary.org/page/381906" refId="B1" refString="Blake, SF, 1924. New American Asteraceae. Contributions from the United States National Herbarium 22 (8): 587 - 661, https://biodiversitylibrary.org/page/381906" title="New American Asteraceae." url="https://biodiversitylibrary.org/page/381906" volume="22" year="1924">Blake (1924)</bibRefCitation>
and accepted as a good species by
<bibRefCitation author="Jervis, RN" journalOrPublisher="8. Eudicots-Asterales. Springer-Verlag, Berlin, Heidelberg" pageId="16" pageNumber="121" refId="B12" refString="Jervis, RN, 1954. A summary of the genus Gochnatia including a revision of the West Indian species which comprise the Section Anastraphioides. University of Michigan: unpublished thesis." title="A summary of the genus Gochnatia including a revision of the West Indian species which comprise the Section Anastraphioides. University of Michigan: unpublished thesis." year="1954">Jervis (1954)</bibRefCitation>
. Cabrera placed it as a subspecies of
<taxonomicName lsidName="G. hypoleuca" pageId="11" pageNumber="116" rank="species" species="hypoleuca">
<emphasis italics="true" pageId="11" pageNumber="116">G. hypoleuca</emphasis>
</taxonomicName>
as did
<bibRefCitation DOI="https://doi.org/10.2307/3298594" author="Freire, SE" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="16" pageNumber="121" pagination="525 - 550" refId="B7" refString="Freire, SE, Katinas, L, Sancho, G, 2002. Gochnatia (Asteraceae: Mutisieae) and the Gochnatia complex: taxonomic implications from morphology. Annals of the Missouri Botanical Garden 89 (4): 525 - 550, DOI: https://doi.org/10.2307/3298594" title="Gochnatia (Asteraceae: Mutisieae) and the Gochnatia complex: taxonomic implications from morphology." url="https://doi.org/10.2307/3298594" volume="89" year="2002">Freire et al. (2002)</bibRefCitation>
. The difference of opinion is based on the importance one gives to the characters that separate the two entities. Blake stated that
<taxonomicName lsidName="G. hypoleuca" pageId="11" pageNumber="116" rank="species" species="hypoleuca">
<emphasis italics="true" pageId="11" pageNumber="116">G. hypoleuca</emphasis>
</taxonomicName>
could be separated from
<taxonomicName lsidName="G. obtusata" pageId="11" pageNumber="116" rank="species" species="obtusata">
<emphasis italics="true" pageId="11" pageNumber="116">G. obtusata</emphasis>
</taxonomicName>
by the following characters: leaves chiefly elliptic, apex acute or acutish and mucronate, 20-50 mm
<normalizedToken originalValue="×">x</normalizedToken>
8-15 mm; involucre 5-6 mm tall, phyllaries obtuse to acute or obtusely acuminate. We have added some additional characters: shrub or small tree 3-7 m; heads arranged in short clusters or panicles; involucre with
<pageBreakToken pageId="12" pageNumber="117" start="start">bracts</pageBreakToken>
in 3-5 series; corolla 10-12 mm long; style branches 0.7-1 mm long; pappus up to 6.5 mm. In contrast,
<taxonomicName lsidName="G. obtusata" pageId="12" pageNumber="117" rank="species" species="obtusata">
<emphasis italics="true" pageId="12" pageNumber="117">G. obtusata</emphasis>
</taxonomicName>
has the following: leaves broadly elliptic, apex broadly rounded to obtuse, not mucronate, 4.5-20 mm
<normalizedToken originalValue="×">x</normalizedToken>
3-13 mm; involucre 6.5-7.5 mm tall, involucral bracts acute to sharply acuminate; style branches 0.5-0.7 mm long. Additional characters for
<taxonomicName lsidName="G. obtusata" pageId="12" pageNumber="117" rank="species" species="obtusata">
<emphasis italics="true" pageId="12" pageNumber="117">G. obtusata</emphasis>
</taxonomicName>
include: shrub of 1.0-2.6 m; heads glomerate at tips of branches and in clusters of 1-several in the subterminal axils; involucre with bracts in 5-6 series; corollas 6.5-7.5 mm long; pappus up to 7.5 mm long. In the field
<pageBreakToken pageId="13" pageNumber="118" start="start">the</pageBreakToken>
difference is striking with the larger more robust
<taxonomicName lsidName="G. hypoleuca" pageId="13" pageNumber="118" rank="species" species="hypoleuca">
<emphasis italics="true" pageId="13" pageNumber="118">G. hypoleuca</emphasis>
</taxonomicName>
contrasting with the smaller, more compact
<taxonomicName lsidName="G. obtusata" pageId="13" pageNumber="118" rank="species" species="obtusata">
<emphasis italics="true" pageId="13" pageNumber="118">G. obtusata</emphasis>
</taxonomicName>
. Therefore we have chosen to recognize
<taxonomicName lsidName="G. obtusata" pageId="13" pageNumber="118" rank="species" species="obtusata">
<emphasis italics="true" pageId="13" pageNumber="118">G. obtusata</emphasis>
</taxonomicName>
as a separate species.
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<mods:identifier id="D47ECD12E57DBF66275F66B7A7757AE4" type="DOI">http://dx.doi.org/10.3897/zookeys.1185.108286</mods:identifier>
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<treatment id="5EAAF589767553BEAC7CC7C05918669D" LSID="urn:lsid:zoobank.org:act:C15B8CD5-0E11-47C4-8626-81D33D718055" httpUri="http://treatment.plazi.org/id/5EAAF589767553BEAC7CC7C05918669D" lastPageNumber="163" pageId="0" pageNumber="163">
<subSubSection id="0B6850D412A7DDAAD7525BEDE327AFE4" pageId="0" pageNumber="163" type="nomenclature">
<paragraph id="E6A3C6DDD3DD18FE88DA27A76A5C9175" pageId="0" pageNumber="163">
<taxonomicName id="E18A024120785B61E7C551CF4A279243" LSID="https://zoobank.org/C15B8CD5-0E11-47C4-8626-81D33D718055" authority="Zehnpfennig &amp; Mahon, 2023" authorityName="Zehnpfennig &amp; Mahon" authorityYear="2023" class="Pycnogonida" family="Callipallenidae" genus="Austropallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austropallene halanychi" order="Pantopoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="halanychi" status="sp. nov.">Austropallene halanychi</taxonomicName>
<taxonomicNameLabel id="9C0228B8464D14FF86428EEEE95BBAAA" pageId="0" pageNumber="163">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="91439C338997D4D9DC9AD2C1B12E67F1" pageId="0" pageNumber="163" type="description">
<paragraph id="6EBBE27FBD73A21935B21F510070ED75" pageId="0" pageNumber="163">
<figureCitation id="762A6A21349AC66A0982DE7BE965854C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">Figs 2</figureCitation>
<figureCitation id="1A496E0AC61B58D46FCBFB3133B134EF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Austropallene halanychi sp. nov., male holotype A dorsal view B dorsal-frontal view; note shape, relative size, and black tips of chela fingers, cephalic spurs, and eye tubercle and eyes; note sharp conical outgrowths at base of fixed and movable fingers of chelifores (red arrow) C eye tubercle, top view D dorsal view of cephalic spurs, chelifore claws, and proboscis; note setose wreath on proboscis E setose wreath of proboscis F dorsal view of chelifore claws; note lack of space between chelifore fingers and small outgrowth (red arrow) G dorsal view of tip of chelifore claw; note presence of bristles along top of immovable finger." figureDoi="10.3897/zookeys.1185.108286.figure3" httpUri="https://binary.pensoft.net/fig/943966" pageId="0" pageNumber="163">, 3</figureCitation>
<figureCitation id="F29DB55DA558F48F06AC083920EFC352" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Austropallene halanychi sp. nov., male holotype A oviger, entire B strigilis, close-up view showing compound spines on segments 7 - 10 C compound spines on segment 9 D compound spines on segment 9 E strigilis, side view, showing compound spines on segments 7 - 10 (some spines on segment 10 are damaged or missing) F segments 8 - 10 with compound spines, and segment 10 with two terminal spines G segment 10, top view, close-up, with two terminal compound spines." figureDoi="10.3897/zookeys.1185.108286.figure4" httpUri="https://binary.pensoft.net/fig/943967" pageId="0" pageNumber="163">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="74D37829D325C17524F260B59E31EFE8" pageId="0" pageNumber="163" type="type locality">
<paragraph id="35159D9A840C9CE5B191666B0CB5E5E8" pageId="0" pageNumber="163">Type locality.</paragraph>
<paragraph id="357276523907B561C057AB150B0B0E8A" pageId="0" pageNumber="163">
Antarctica, Ross Sea, Ross Shelf, 570 m depth,
<geoCoordinate id="1C5BFE6606184BD4D2D8B23C6844277A" degrees="75" direction="south" minutes="19" orientation="latitude" precision="1" seconds="46.7" value="-75.329636">75°19'46.7&quot;S</geoCoordinate>
,
<geoCoordinate id="2B515B978663087889E4A97622E26326" degrees="176" direction="west" minutes="59" orientation="longitude" precision="1" seconds="06.3" value="-176.98509">176°59'06.3&quot;W</geoCoordinate>
, collected via benthic trawl, 31 January 2013, RVIB
<emphasis id="0FFF7553B703A8851A255D0E86E8E71C" italics="true" pageId="0" pageNumber="163">Nathaniel B. Palmer</emphasis>
(NBP12-10), A. Mahon leg.
</paragraph>
</subSubSection>
<subSubSection id="7CD72B75AD17FB75759D36E420D887F3" pageId="0" pageNumber="163" type="type specimen">
<paragraph id="70550AE1F591632E1B982363DC42F7F2" pageId="0" pageNumber="163">Type specimen.</paragraph>
<paragraph id="90411C23699A2601D3DC35F5874B2233" pageId="0" pageNumber="163">
Holotype male preserved in ethanol, original label &quot;Antarctica, Ross Sea, Ross Shelf, 570 m depth,
<geoCoordinate id="9A691354F2EED6EB4CD6D5B9C5744ABF" degrees="75" direction="south" minutes="19" orientation="latitude" precision="1" seconds="46.7" value="-75.329636">75°19'46.7&quot;S</geoCoordinate>
,
<geoCoordinate id="DE874AF5E8A44DF220BECDE9AA565D69" degrees="176" direction="west" minutes="59" orientation="longitude" precision="1" seconds="06.3" value="-176.98509">176°59'06.3&quot;W</geoCoordinate>
, 31 January 2013, J. Zehnpfennig and A. Mahon&quot;, NMNH 1548440&quot;; handwritten label &quot;Ch 226.1E&quot;.
</paragraph>
</subSubSection>
<subSubSection id="FC467675302A199A0FD38C8512ED3612" pageId="0" pageNumber="163" type="diagnosis">
<paragraph id="DAC67713F9C24CDABC7FC97EEE5964CD" pageId="0" pageNumber="163">Diagnosis.</paragraph>
<paragraph id="C841C7D1F89C01BB862DDB714D642D98" pageId="0" pageNumber="163">
The new species can be differentiated from all other described
<taxonomicName id="7B9D57AD414F3BC56041F30E878B016C" class="Pycnogonida" family="Callipallenidae" genus="Austropallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austropallene" order="Pantopoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="E0851626865D0E99D9A879EE9E19F4F5" italics="true" pageId="0" pageNumber="163">Austropallene</emphasis>
</taxonomicName>
species based on its much larger size, as well as its unique and distinctive chelifores. The chelifore claw of
<taxonomicName id="723D3ED06D03F3EC6B43827FCFE3D02D" class="Pycnogonida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. halanychi" order="Pantopoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="halanychi">
<emphasis id="8DA05187FF21DB758006E369D344B1CC" italics="true" pageId="0" pageNumber="163">A. halanychi</emphasis>
</taxonomicName>
sp. nov. closes completely when the fingers converge, with no visible space remaining between the fixed and movable fingers. The movable finger of the chelifore claw is distinctly shorter than the fixed finger, and the chelifore fingers do not contain any denticles or notches on their inner surfaces.
</paragraph>
</subSubSection>
<subSubSection id="AF29871E5AE659148A0E9EFC6B2FB87C" pageId="0" pageNumber="163" type="description">
<paragraph id="7B012552AC222D9B7645626DF5154150" pageId="0" pageNumber="163">Description of holotype (male).</paragraph>
<paragraph id="EB68F7FA70EC820530A79FB1C4D1A104" pageId="0" pageNumber="163">
Slender appearance, fully segmented trunk; neck distinct, large oblong ovigers attaching laterally; cephalic spurs present (Figs
<figureCitation id="98DE3AA0A3F224B70EDCD14FDEDBDB0A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">2A-C</figureCitation>
,
<figureCitation id="9DAA7F49F14B0B49647BFCCA0A7F57FA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Austropallene halanychi sp. nov., male holotype A dorsal view B dorsal-frontal view; note shape, relative size, and black tips of chela fingers, cephalic spurs, and eye tubercle and eyes; note sharp conical outgrowths at base of fixed and movable fingers of chelifores (red arrow) C eye tubercle, top view D dorsal view of cephalic spurs, chelifore claws, and proboscis; note setose wreath on proboscis E setose wreath of proboscis F dorsal view of chelifore claws; note lack of space between chelifore fingers and small outgrowth (red arrow) G dorsal view of tip of chelifore claw; note presence of bristles along top of immovable finger." figureDoi="10.3897/zookeys.1185.108286.figure3" httpUri="https://binary.pensoft.net/fig/943966" pageId="0" pageNumber="163">3A, B</figureCitation>
). Ocular tubercle (Figs
<figureCitation id="E7842769DC7E4270227F6B0A6E802888" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">2A-C</figureCitation>
,
<figureCitation id="EF1F9F58386E5C2F4698F7B7E09BF9A9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Austropallene halanychi sp. nov., male holotype A dorsal view B dorsal-frontal view; note shape, relative size, and black tips of chela fingers, cephalic spurs, and eye tubercle and eyes; note sharp conical outgrowths at base of fixed and movable fingers of chelifores (red arrow) C eye tubercle, top view D dorsal view of cephalic spurs, chelifore claws, and proboscis; note setose wreath on proboscis E setose wreath of proboscis F dorsal view of chelifore claws; note lack of space between chelifore fingers and small outgrowth (red arrow) G dorsal view of tip of chelifore claw; note presence of bristles along top of immovable finger." figureDoi="10.3897/zookeys.1185.108286.figure3" httpUri="https://binary.pensoft.net/fig/943966" pageId="0" pageNumber="163">3B, C</figureCitation>
) low, rounded, slightly inclined backwards, prominent distal papillae; four darkly pigmented eyes, anterior pair larger than posterior pair (Fig.
<figureCitation id="42CF2C4AB4E067AFFF1376C6FC27D96B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">2A, I</figureCitation>
). Lateral processes long, smooth, without spines. Abdomen short, conical, swollen distally, cleft anal opening. Proboscis (Figs
<figureCitation id="1A8E4D25507BC86B854CA3A493669A43" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">2A-C</figureCitation>
,
<figureCitation id="DD06A6CCB17A948035DF49AB364CD8D2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Austropallene halanychi sp. nov., male holotype A dorsal view B dorsal-frontal view; note shape, relative size, and black tips of chela fingers, cephalic spurs, and eye tubercle and eyes; note sharp conical outgrowths at base of fixed and movable fingers of chelifores (red arrow) C eye tubercle, top view D dorsal view of cephalic spurs, chelifore claws, and proboscis; note setose wreath on proboscis E setose wreath of proboscis F dorsal view of chelifore claws; note lack of space between chelifore fingers and small outgrowth (red arrow) G dorsal view of tip of chelifore claw; note presence of bristles along top of immovable finger." figureDoi="10.3897/zookeys.1185.108286.figure3" httpUri="https://binary.pensoft.net/fig/943966" pageId="0" pageNumber="163">3B, D, E</figureCitation>
) directed ventrally, broad at base, conical, slight mid-point constriction, distally tapering; mouth surrounded by setose wreath (Figs
<figureCitation id="167534A4C26A7C505B27D9E058C5665C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">2G, H</figureCitation>
,
<figureCitation id="E384FE6F73AB9FDA73AA6E2119C82084" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Austropallene halanychi sp. nov., male holotype A dorsal view B dorsal-frontal view; note shape, relative size, and black tips of chela fingers, cephalic spurs, and eye tubercle and eyes; note sharp conical outgrowths at base of fixed and movable fingers of chelifores (red arrow) C eye tubercle, top view D dorsal view of cephalic spurs, chelifore claws, and proboscis; note setose wreath on proboscis E setose wreath of proboscis F dorsal view of chelifore claws; note lack of space between chelifore fingers and small outgrowth (red arrow) G dorsal view of tip of chelifore claw; note presence of bristles along top of immovable finger." figureDoi="10.3897/zookeys.1185.108286.figure3" httpUri="https://binary.pensoft.net/fig/943966" pageId="0" pageNumber="163">3E</figureCitation>
).
</paragraph>
<caption id="D62B2DFC5283AF4BE4F2679F2600E63C" doi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163" start="Figure 2" startId="F2">
<paragraph id="3B73E4D02D9A20617B824E3DCBBD0CFD" pageId="0" pageNumber="163">
<emphasis id="A40F5C802ADBD557799FCC8476972469" bold="true" pageId="0" pageNumber="163">Figure 2.</emphasis>
<taxonomicName id="42CBB0F83FE8697D775EE7DD80CB7A7A" authorityName="Zehnpfennig &amp; Mahon" authorityYear="2023" class="Pycnogonida" family="Callipallenidae" genus="Austropallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austropallene halanychi" order="Pantopoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="halanychi">
<emphasis id="EEB58217B95C2F8A53353BF4424BF417" italics="true" pageId="0" pageNumber="163">Austropallene halanychi</emphasis>
</taxonomicName>
sp. nov., male holotype
<emphasis id="1C26582E3B813A95C4FB3E94710AFF3B" bold="true" pageId="0" pageNumber="163">A</emphasis>
side view (scale)
<emphasis id="CD6F596EF8F4E6627796EA1358134BBC" bold="true" pageId="0" pageNumber="163">B</emphasis>
dorsal view
<emphasis id="C8FD5DB15E13A008D7319934FF5C0299" bold="true" pageId="0" pageNumber="163">C</emphasis>
front view, note cephalic spurs
<emphasis id="19B0C02AE2AA2778594CB752FA9DC309" bold="true" pageId="0" pageNumber="163">D</emphasis>
dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers
<emphasis id="58FE90D60E1E32EF9DEBB08D08B7C920" bold="true" pageId="0" pageNumber="163">E</emphasis>
dorsal view of chelifore claw
<emphasis id="D53D3AB599BD84E69B937B65CD191374" bold="true" pageId="0" pageNumber="163">F</emphasis>
ventral view of chelifore claw; note conical outgrowth
<emphasis id="CC92A0915658394CC6BDD8C7A54CBD5B" bold="true" pageId="0" pageNumber="163">G</emphasis>
dorsal view of proboscis with setose wreath H setose wreath around proboscis
<emphasis id="1159AD2626923034BD210277588E73FF" bold="true" pageId="0" pageNumber="163">I</emphasis>
dorsal view of eyespot
<emphasis id="6E275BBF628F5E53D29BCC0E56E8B2CE" bold="true" pageId="0" pageNumber="163">J</emphasis>
third walking leg
<emphasis id="08B20E9A24E2B5F41AA8CF06DC30C206" bold="true" pageId="0" pageNumber="163">K</emphasis>
first coxa of third walking leg
<emphasis id="D44DA09C53FA4D392EAE167492CA485F" bold="true" pageId="0" pageNumber="163">L</emphasis>
side view of second coxa of third walking leg
<emphasis id="DDC00032B65D60E2FE90E37241339F1A" bold="true" pageId="0" pageNumber="163">M</emphasis>
propodus of walking leg
<emphasis id="DBA149730377D4E06E3F8F62A4A2FDEA" bold="true" pageId="0" pageNumber="163">N</emphasis>
tubercles and spines on walking legs
<emphasis id="F78245C1ADCBDE69604E4586DA420289" bold="true" pageId="0" pageNumber="163">O</emphasis>
ovigerous leg of male
<emphasis id="358FEF9816C6956D12B5498A9FD30DF1" bold="true" pageId="0" pageNumber="163">P</emphasis>
strigilis of ovigerous leg
<emphasis id="39801947F7964582604BC51243BC6F7E" bold="true" pageId="0" pageNumber="163">Q</emphasis>
10th segment of oviger; note terminal spines
<emphasis id="68E49DA90DAA78BC8EA5A25C53DD5C78" bold="true" pageId="0" pageNumber="163">R</emphasis>
serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg.
</paragraph>
</caption>
<caption id="6139CA042C0F76BE70F959FD79000BFC" doi="10.3897/zookeys.1185.108286.figure3" httpUri="https://binary.pensoft.net/fig/943966" pageId="0" pageNumber="163" start="Figure 3" startId="F3">
<paragraph id="9433923784D0E0FFCD5ECE81F99940CD" pageId="0" pageNumber="163">
<emphasis id="AAB1996F3DAD5E3B40DF52707804BB64" bold="true" pageId="0" pageNumber="163">Figure 3.</emphasis>
<taxonomicName id="8AB82307C06841289E490B56FBADD859" authorityName="Zehnpfennig &amp; Mahon" authorityYear="2023" class="Pycnogonida" family="Callipallenidae" genus="Austropallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austropallene halanychi" order="Pantopoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="halanychi">
<emphasis id="EA313299661466B7E52BDCCA8A148999" italics="true" pageId="0" pageNumber="163">Austropallene halanychi</emphasis>
</taxonomicName>
sp. nov., male holotype
<emphasis id="BAF36D4E6CA55FEE3CBFEDECE7CF6B52" bold="true" pageId="0" pageNumber="163">A</emphasis>
dorsal view
<emphasis id="EC0A10CCA34E5EF17653B2D68FEAEC6D" bold="true" pageId="0" pageNumber="163">B</emphasis>
dorsal-frontal view; note shape, relative size, and black tips of chela fingers, cephalic spurs, and eye tubercle and eyes; note sharp conical outgrowths at base of fixed and movable fingers of chelifores (red arrow)
<emphasis id="38E7E0A19E745800F2FEF0A2FC7523C9" bold="true" pageId="0" pageNumber="163">C</emphasis>
eye tubercle, top view
<emphasis id="9AE4A1ECD6C306B9DD741258BF251424" bold="true" pageId="0" pageNumber="163">D</emphasis>
dorsal view of cephalic spurs, chelifore claws, and proboscis; note setose wreath on proboscis
<emphasis id="6D16C2F66C3AAC6A84D4A4DECD263D4C" bold="true" pageId="0" pageNumber="163">E</emphasis>
setose wreath of proboscis
<emphasis id="AEC07B3E5FAE4BE70FD0B5F8DF7B6908" bold="true" pageId="0" pageNumber="163">F</emphasis>
dorsal view of chelifore claws; note lack of space between chelifore fingers and small outgrowth (red arrow)
<emphasis id="E7DA3AC2F287DCEB1FA59D6D91553FCE" bold="true" pageId="0" pageNumber="163">G</emphasis>
dorsal view of tip of chelifore claw; note presence of bristles along top of immovable finger.
</paragraph>
</caption>
<paragraph id="427ECB5AEC24330F0968AE3A4888C17A" pageId="0" pageNumber="163">
Chelifore (Figs
<figureCitation id="2D634F35478BF6C4E15F5A13AB554495" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">2D-F</figureCitation>
,
<figureCitation id="84AFD7CBB74EB8F79159A588C615ACE6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Austropallene halanychi sp. nov., male holotype A dorsal view B dorsal-frontal view; note shape, relative size, and black tips of chela fingers, cephalic spurs, and eye tubercle and eyes; note sharp conical outgrowths at base of fixed and movable fingers of chelifores (red arrow) C eye tubercle, top view D dorsal view of cephalic spurs, chelifore claws, and proboscis; note setose wreath on proboscis E setose wreath of proboscis F dorsal view of chelifore claws; note lack of space between chelifore fingers and small outgrowth (red arrow) G dorsal view of tip of chelifore claw; note presence of bristles along top of immovable finger." figureDoi="10.3897/zookeys.1185.108286.figure3" httpUri="https://binary.pensoft.net/fig/943966" pageId="0" pageNumber="163">3B, D, F, G</figureCitation>
), scape large and oblong, shorter than proboscis, directed slightly anteroventrally, slight tapering towards base; chelae long, 77.5% of scape length; tips of chela fingers blackened; fingers longer than one-half of palm length, fixed finger of chelifore concave with pointed tip. Movable finger 0.73 mm shorter than fixed finger, convex, with rounded tip. Chelifore fingers converge completely, no space present between fingers, denticles not present on inner surface of either finger. Two small conical outgrowths present where fingers of chelifore claw meet (Fig.
<figureCitation id="CFC6A8D880D7DC31984767AA0A9D2052" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Austropallene halanychi sp. nov., male holotype A dorsal view B dorsal-frontal view; note shape, relative size, and black tips of chela fingers, cephalic spurs, and eye tubercle and eyes; note sharp conical outgrowths at base of fixed and movable fingers of chelifores (red arrow) C eye tubercle, top view D dorsal view of cephalic spurs, chelifore claws, and proboscis; note setose wreath on proboscis E setose wreath of proboscis F dorsal view of chelifore claws; note lack of space between chelifore fingers and small outgrowth (red arrow) G dorsal view of tip of chelifore claw; note presence of bristles along top of immovable finger." figureDoi="10.3897/zookeys.1185.108286.figure3" httpUri="https://binary.pensoft.net/fig/943966" pageId="0" pageNumber="163">3B</figureCitation>
, arrow). Chelifore scape and claw contain small setae along dorsal and ventral sides.
</paragraph>
<paragraph id="9BED5B306B7DC865DE8FB3EF02519F4F" pageId="0" pageNumber="163">
Oviger 10-segmented (Figs
<figureCitation id="5CD6FB77E30EA0C83FCEF9BDE33E8C63" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">2O-R</figureCitation>
,
<figureCitation id="857CF123F8B87D4F0808D8DC37C25C83" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Austropallene halanychi sp. nov., male holotype A oviger, entire B strigilis, close-up view showing compound spines on segments 7 - 10 C compound spines on segment 9 D compound spines on segment 9 E strigilis, side view, showing compound spines on segments 7 - 10 (some spines on segment 10 are damaged or missing) F segments 8 - 10 with compound spines, and segment 10 with two terminal spines G segment 10, top view, close-up, with two terminal compound spines." figureDoi="10.3897/zookeys.1185.108286.figure4" httpUri="https://binary.pensoft.net/fig/943967" pageId="0" pageNumber="163">4A-G</figureCitation>
), fifth segment longest, slightly curved, with prominent apophysis distally; oviger compound spines serrated, present on terminal four segments of the strigillis, strigilis formula: 20:20:18:18, with 2 prominent terminal spines. Some of the spines are broken off (Fig.
<figureCitation id="BCD6F242A085FBC0A78066AA6F8A6038" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Austropallene halanychi sp. nov., male holotype A oviger, entire B strigilis, close-up view showing compound spines on segments 7 - 10 C compound spines on segment 9 D compound spines on segment 9 E strigilis, side view, showing compound spines on segments 7 - 10 (some spines on segment 10 are damaged or missing) F segments 8 - 10 with compound spines, and segment 10 with two terminal spines G segment 10, top view, close-up, with two terminal compound spines." figureDoi="10.3897/zookeys.1185.108286.figure4" httpUri="https://binary.pensoft.net/fig/943967" pageId="0" pageNumber="163">4E, F</figureCitation>
).
</paragraph>
<caption id="DEFDB742CBE51113F1062F48B0F62BD7" doi="10.3897/zookeys.1185.108286.figure4" httpUri="https://binary.pensoft.net/fig/943967" pageId="0" pageNumber="163" start="Figure 4" startId="F4">
<paragraph id="2B4B88E664BBD6704DABA3E47FECD3EF" pageId="0" pageNumber="163">
<emphasis id="2E3D17CED966364186B5D58453A7DE1E" bold="true" pageId="0" pageNumber="163">Figure 4.</emphasis>
<taxonomicName id="21CF047776AC545942B600F8122B8CCD" authorityName="Zehnpfennig &amp; Mahon" authorityYear="2023" class="Pycnogonida" family="Callipallenidae" genus="Austropallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austropallene halanychi" order="Pantopoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="halanychi">
<emphasis id="D0D06F0263B1128ED23B7B10652DC48E" italics="true" pageId="0" pageNumber="163">Austropallene halanychi</emphasis>
</taxonomicName>
sp. nov., male holotype
<emphasis id="90F1C98EC9E7DBB7A9311BBBDD7CBFCE" bold="true" pageId="0" pageNumber="163">A</emphasis>
oviger, entire
<emphasis id="8CE4606E24AD56216B9B09B28D8AE7FF" bold="true" pageId="0" pageNumber="163">B</emphasis>
strigilis, close-up view showing compound spines on segments 7-10
<emphasis id="BF8302FA99F797EA3F83B16A26686B4D" bold="true" pageId="0" pageNumber="163">C</emphasis>
compound spines on segment 9
<emphasis id="B7B0E6D7790F484342625D640A004022" bold="true" pageId="0" pageNumber="163">D</emphasis>
compound spines on segment 9
<emphasis id="E8C611A56F86E29FDF5F83C31357BE23" bold="true" pageId="0" pageNumber="163">E</emphasis>
strigilis, side view, showing compound spines on segments 7-10 (some spines on segment 10 are damaged or missing)
<emphasis id="1A42ECF949529B03E3F2F270FD7CC6C1" bold="true" pageId="0" pageNumber="163">F</emphasis>
segments 8-10 with compound spines, and segment 10 with two terminal spines
<emphasis id="18F47958190BEC861FA265E78C2566F4" bold="true" pageId="0" pageNumber="163">G</emphasis>
segment 10, top view, close-up, with two terminal compound spines.
</paragraph>
</caption>
<paragraph id="301C3096671CB79EE0C10E23D96E4FE7" pageId="0" pageNumber="163">
Walking legs (Fig.
<figureCitation id="A209A01D3A2D28152582C1625396BAA6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Austropallene halanychi sp. nov., male holotype A side view (scale) B dorsal view C front view, note cephalic spurs D dorsal view of opened chelifore claw; note absence of denticle and conical outgrowth at base of fixed and movable fingers E dorsal view of chelifore claw F ventral view of chelifore claw; note conical outgrowth G dorsal view of proboscis with setose wreath H setose wreath around proboscis I dorsal view of eyespot J third walking leg K first coxa of third walking leg L side view of second coxa of third walking leg M propodus of walking leg N tubercles and spines on walking legs O ovigerous leg of male P strigilis of ovigerous leg Q 10 th segment of oviger; note terminal spines R serrated ovigerous leg spines (compound spines) on strigilis of ovigerous leg." figureDoi="10.3897/zookeys.1185.108286.figure2" httpUri="https://binary.pensoft.net/fig/943965" pageId="0" pageNumber="163">2J-N</figureCitation>
), long, slender, sparse short spinules dorsally and ventrally on major segments; small conical tubercles and setae on first coxa. Femur shortest leg segment; tibia 2 longest segment; tarsus short, curved, small distal spine in line with heel spines; propodus curved, distinct propodal heel; five heel spines, relatively similar in size; many sole spines present; auxiliary claws absent, main claw extends nearly to heel. Gonopores and cement glands not observed.
</paragraph>
</subSubSection>
<subSubSection id="F10D83FC2611F4D245BB209AC4B608FC" pageId="0" pageNumber="163" type="description">
<paragraph id="4AEFB3C411FDEDD12F32BDF735CFBC0A" pageId="0" pageNumber="163">Measurements (male holotype-mm).</paragraph>
<paragraph id="D224828F11E536F10B5069AC9D9B2765" pageId="0" pageNumber="163">Body length (anterior end of cephalon to posterior end of trunk) = 10.50; body width (right lateral process of leg 1 + width of trunk + left lateral process of leg 1) = 4.02; trunk width = 1.24; abdomen length = 6.97; ocular tubercle height = 0.39; proboscis length = 2.73; chela fingers = 2.61; main claw length = 3.37; scape = 2.39; oviger 5th segment = 4.18; 10th segment = 0.74; terminal oviger spines = 0.10; leg span 63.84mm (distance between terminal claws second pair of walking legs); walking legs 29.78 mm in length (from where first coxa meets lateral process to terminal claw on second pair of walking legs); 3rd walking leg coxa 1 = 1.23, coxa 2 = 2.60, coxa 3 = 1.17, femur = 6.69, tibia 1 = 6.85, tibia 2 = 7.89, tarsus = 0.32, propodus = 1.95, terminal claw = 1.39.</paragraph>
</subSubSection>
<subSubSection id="D1754BB9822400764A9325E8130E9DA6" pageId="0" pageNumber="163" type="etymology">
<paragraph id="1FEBC6228FE908629F14E6D4AA76B1C1" pageId="0" pageNumber="163">Etymology.</paragraph>
<paragraph id="1C1760147C02B20965042FC33B8F8EA2" pageId="0" pageNumber="163">
The species (
<taxonomicName id="B8A56EF738C5AACA213C96B3D1A2E3E3" authorityName="Zehnpfennig &amp; Mahon" authorityYear="2023" class="Pycnogonida" family="Callipallenidae" genus="Austropallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austropallene halanychi" order="Pantopoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="halanychi">
<emphasis id="AF09560830FF0B8054F3D3220E1654C7" italics="true" pageId="0" pageNumber="163">Austropallene halanychi</emphasis>
</taxonomicName>
, male genitive) is dedicated to Dr Kenneth M. Halanych, a mentor, colleague, and prolific marine invertebrate scientist whose commitment and dedication to the benthic marine systems in the Southern Ocean has provided a wealth of information related to biodiversity in the Antarctic system.
</paragraph>
</subSubSection>
</treatment>
</document>