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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.653.10252" ID-PMC="PMC5345375" ID-Pensoft-Pub="1313-2970-653-1" ID-Pensoft-UUID="FFE87C63852C5772725FBE55FF95902D" ID-PubMed="28331393" ID-Zenodo-Dep="269679" ID-ZooBank="8ECC0674017A48588BE8DDD05C0D7CF6" ModsDocID="1313-2970-653-1" checkinTime="1486523616090" checkinUser="pensoft" docAuthor="Abdel-Dayem, Mahmoud S., Fad, Hassan H., El-Torkey, Ashraf M., Elgharbawy, Ali A., Aldryhim, Yousif N., Kondratieff, Boris C., Ansi, Amin N. Al &amp; Aldhafer, Hathal M." docDate="2017" docId="5E38000163AB51F9AC5733A16816B3A3" docLanguage="en" docName="ZooKeys 653: 1-78" docOrigin="ZooKeys 653" docPubDate="2017-02-07" docSource="http://dx.doi.org/10.3897/zookeys.653.10252" docTitle="Cybocephalus rufifrons subsp. flaviceps Reitter 1874" docType="treatment" docVersion="3" id="FFE87C63852C5772725FBE55FF95902D" lastPageNumber="44" masterDocId="FFE87C63852C5772725FBE55FF95902D" masterDocTitle="The beetle fauna (Insecta, Coleoptera) of the Rawdhat Khorim National Park, Central Saudi Arabia" masterLastPageNumber="78" masterPageNumber="1" pageNumber="44" updateTime="1668164043815" updateUser="ExternalLinkService">
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<mods:title>The beetle fauna (Insecta, Coleoptera) of the Rawdhat Khorim National Park, Central Saudi Arabia</mods:title>
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<mods:namePart>Abdel-Dayem, Mahmoud S.</mods:namePart>
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<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
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<mods:namePart>Fad, Hassan H.</mods:namePart>
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<mods:namePart>El-Torkey, Ashraf M.</mods:namePart>
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<mods:namePart>Elgharbawy, Ali A.</mods:namePart>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia &amp; Zoology Department, Faculty of Science, Al Azhar University, Nasr City, Cairo, Egypt</mods:affiliation>
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<mods:namePart>Aldryhim, Yousif N.</mods:namePart>
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<mods:namePart>Kondratieff, Boris C.</mods:namePart>
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<mods:namePart>Ansi, Amin N. Al</mods:namePart>
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<mods:namePart>Aldhafer, Hathal M.</mods:namePart>
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<taxonomicName LSID="5E380001-63AB-51F9-AC57-33A16816B3A3" authority="Reitter, 1874" authorityName="Reitter" authorityYear="1874" class="Insecta" family="Cleridae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybocephalus rufifrons subsp. flaviceps" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="subSpecies" species="rufifrons" subSpecies="flaviceps">Cybocephalus rufifrons flaviceps Reitter, 1874</taxonomicName>
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<paragraph pageId="43" pageNumber="44">World distribution.</paragraph>
<paragraph pageId="43" pageNumber="44">
<emphasis bold="true" pageId="43" pageNumber="44">Asia</emphasis>
: IQ, IR, SA, TR.
<emphasis bold="true" pageId="43" pageNumber="44">North Africa</emphasis>
: DZ, EG, MA, TN.
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<paragraph pageId="43" pageNumber="44">General distribution.</paragraph>
<paragraph pageId="43" pageNumber="44">SAR.</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="distribution">
<paragraph pageId="43" pageNumber="44">Local distribution.</paragraph>
<paragraph pageId="43" pageNumber="44">
MK (
<bibRefCitation author="Endroedy-Younga, S" pageId="72" pageNumber="73" refId="B43" refString="Endroedy-Younga, S, 1979. Insects of Saudi Arabia, Coleoptera: Fam. Cybocephalidae. Fauna of Saudi Arabia 1: 228-231." title="Insects of Saudi Arabia, Coleoptera: Fam. Cybocephalidae. Fauna of Saudi Arabia 1: 228 - 231." year="1979">
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1979
</bibRefCitation>
), RI (
<bibRefCitation author="Beccari, F" pageId="70" pageNumber="71" refId="B19" refString="Beccari, F, 1971. Contribution alla conoscenze d'ell' entomofauna dell' Arabia Saudita. Primo elenco di insetti acari e nematode. Rivista di Agricolatura Subtropical et Tropicale 65: 243-258." title="Contribution alla conoscenze d'ell' entomofauna dell' Arabia Saudita. Primo elenco di insetti acari e nematode. Rivista di Agricolatura Subtropical et Tropicale 65: 243 - 258." year="1971">Beccari 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="collecting month and method">
<paragraph pageId="43" pageNumber="44">Collecting month and method.</paragraph>
<paragraph pageId="43" pageNumber="44">
A common species. The specimens were collected by BV, SW and VC on branches of
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia ehrenbergiana" order="Fabales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="ehrenbergiana">
<emphasis italics="true" pageId="43" pageNumber="44">Acacia ehrenbergiana</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia gerrardii" order="Fabales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="species" species="gerrardii">
<emphasis italics="true" pageId="43" pageNumber="44">Acacia gerrardii</emphasis>
</taxonomicName>
,
<taxonomicName class="Dicotyledoneae" family="Apocynaceae" genus="Calotropis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calotropis procera" order="Gentianales" pageId="43" pageNumber="44" phylum="Angiospermae" rank="species" species="procera">
<emphasis italics="true" pageId="43" pageNumber="44">Calotropis procera</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Roem &amp; Schult" authorityYear="1819" class="Dicotyledoneae" family="Solanaceae" genus="Lycium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lycium shawii" order="Tubiflorae" pageId="43" pageNumber="44" phylum="Angiospermae" rank="species" species="shawii">
<emphasis italics="true" pageId="43" pageNumber="44">Lycium shawii</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Decne" authorityYear="1835" class="Dicotyledoneae" family="Apocynaceae" genus="Rhazya" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Rhazya stricta" order="Gentianales" pageId="43" pageNumber="44" phylum="Angiospermae" rank="species" species="stricta">
<emphasis italics="true" pageId="43" pageNumber="44">Rhazya stricta</emphasis>
</taxonomicName>
and
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<emphasis italics="true" pageId="43" pageNumber="44">Ziziphus nummularia</emphasis>
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; and by MT through II, V-IX and XII.
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<taxonomicName id="F65CA5F7DE31B031F0D05C720076EA11" genus="Chalcidoidea" lsidName="Chalcidoidea fasciculatus" pageId="0" pageNumber="8013" rank="species" species="fasciculatus">fasciculatus</taxonomicName>
(
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, 1841,
<taxonomicName id="3B04ED1D50D3D0DBD34EFC303C6B1E2C" class="Insecta" family="Pteromalidae" genus="Pteromalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteromalus" order="Hymenoptera" pageId="0" pageNumber="8013" phylum="Arthropoda" rank="genus">Pteromalus</taxonomicName>
)
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<taxonomicName id="3DFE75D655D838238E20649943A380B9" genus="Chalcidoidea" lsidName="Chalcidoidea saxesenii" pageId="0" pageNumber="8013" rank="species" species="saxesenii">saxesenii</taxonomicName>
(Ratzeburg, 1844,
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<taxonomicName id="D2745B15B277384821562EE29B981DDC" genus="Chalcidoidea" lsidName="Chalcidoidea trochilus" pageId="0" pageNumber="8013" rank="species" species="trochilus">trochilus</taxonomicName>
(Ratzeburg, 1844,
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)
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</subSubSection>
<subSubSection id="6322EAB53450F8C8AA7A86D5B02D3843" pageId="0" pageNumber="8013" type="distribution">
<paragraph id="C201D6506BFFE9D6D5B1690246EA0A84" pageId="0" pageNumber="8013">Distribution</paragraph>
<paragraph id="3AD9AF00EB0B04A0546BBD1E2450A5B9" pageId="0" pageNumber="8013">England</paragraph>
</subSubSection>
</treatment>
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Genus firstly recorded from PI in
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<treatment id="5E38C31F9149A184A40937F59B06EDE0" ID-DOI="http://doi.org/10.5281/zenodo.11338849" ID-GBIF-Taxon="231542135" ID-Zenodo-Dep="11338849" LSID="urn:lsid:plazi:treatment:5E38C31F9149A184A40937F59B06EDE0" httpUri="http://treatment.plazi.org/id/5E38C31F9149A184A40937F59B06EDE0" lastPageNumber="166" pageNumber="166" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=12100586">
<subSubSection id="DBE143214DCBD9AEC077CB1E3A5D2E73" pageNumber="166" type="nomenclature">
<paragraph id="865DEC9853BE39E2FCAD415C1C07ED16" pageNumber="166">
<taxonomicName id="CD0447BC19BE2D00E7A7E6462EBF8E7A" ID-CoL="4CKZ5" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="hamadryas">
<emphasis id="356E3ADFF09904B8717D591DF484FBD1" italics="true" pageNumber="166">Papio hamadryas</emphasis>
(Linnaeus 1758)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F615970AF6B1B3C5F474B94CD6F7EF45" pageNumber="166" type="reference_group">
<paragraph id="CE255C4AB996D9726B28EDB96FE73CD8" pageNumber="166">
<treatmentCitation id="E42BD9D51E57C7513EEE8D13DD95239A" ID-DOI="http://doi.org/10.5281/zenodo.3914754" ID-Zenodo-Dep="3914754" LSID="urn:lsid:zoobank.org:act:A828DE68-FC5A-4E6C-9EC6-394233AFB094" author="Linnaeus" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Cercopithecidae" genus="Simia" httpUri="http://treatment.plazi.org/id/F66FA2A7F76C805B3EB16001347E995F" kingdom="Animalia" order="Primates" page="27" pageNumber="166" phylum="Chordata" rank="species" species="hamadryas" title="Simia hamadryas" volumeTitle="Syst. Nat., 10th ed., Vol. 1" year="1758">
<taxonomicName id="8C39BC47ED7E834FB915078DE529400C" ID-CoL="4T3R" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Cercopithecidae" genus="Simia" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="hamadryas">
<emphasis id="4E13FACAEAF2C334DC9773017E3D7990" italics="true" pageNumber="166">[Simia] hamadryas</emphasis>
<bibRefCitation id="C92C574BC3509F6F43410BA6AA8AEC93" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
</taxonomicName>
,
<bibCitation id="67362E02325A5DE411624B25308B1E4A" author="Linnaeus" pageNumber="166" pagination="27" title="Simia hamadryas" volumeTitle="Syst. Nat., 10th ed., Vol. 1" year="1758">Syst. Nat., 10th ed., Vol. 1: 27</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="069CD83FF6ED12DA98D1323BAC9F6E44" pageNumber="166" type="materials_examined">
<paragraph id="32BCB00176D7241B045EB3A491F548F7" pageNumber="166">
<emphasis id="D3CBAC1BB87C721712F17FA06EEAF594" inLineHeading="true" pageNumber="166">Type Locality:</emphasis>
<materialsCitation id="A402BE80185BEF3FB5C5CF65A350399A" ID-GBIF-Occurrence="4867331575" country="Egypt" pageNumber="166">
<collectingCountry id="1DD9BC53EBEE19F3D12E8E622B191CA9" name="Egypt">Egypt</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FFDCA4202A17939DD10107E0F5317C63" pageNumber="166" type="vernacular_names">
<paragraph id="C6BF08CBF242C2865B6E9EA35586CF1F" pageNumber="166">
<emphasis id="748B7138D0E5998CBCFB6BC388E94A39" inLineHeading="true" pageNumber="166">Vernacular Names:</emphasis>
<vernacularName id="5CF4B8A2CEC98D30D00562BF2680B22F" language="eng" pageNumber="166">Hamadryas Baboon</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="79A272A761FD0D12B59323ABA597BF2E" pageNumber="166" type="synonym_list">
<paragraph id="8C822D492827A694C0F6F53500CAB42C" pageNumber="166">
<emphasis id="6FC86241879342271B827F44CB982B2E" inLineHeading="true" pageNumber="166">Synonyms:</emphasis>
<taxonomicName id="DDE9CED89B039FBD592FD0D53B3430A6" authority="(Gray 1870)" baseAuthorityName="Gray" baseAuthorityYear="1870" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="aegyptiaca">
<emphasis id="2097A6838706544B844D3669DE123B8E" italics="true" pageNumber="166">Papio aegyptiaca</emphasis>
(
<bibRefCitation id="26BD193CF7B1D91FE8B3DC5C40E7905A" author="Gray, J. E." refId="ref128843" refString="Gray, J. E. 1870. Catalogue of monkeys, lemurs and fruit eating bats in the collections of the British Museum. British Museum, London, 137 pp." year="1870">Gray 1870</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="9AFFA88F3C2ECAA5EDF219ED01A5E7CE" authority="Thomas 1900" authorityName="Thomas" authorityYear="1900" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="arabicus">
<emphasis id="7CF5AFACC8D72972FB9D40441D0A3BFB" italics="true" pageNumber="166">Papio arabicus</emphasis>
Thomas 1900
</taxonomicName>
;
<taxonomicName id="AB668F57D9E6012962640C95BBD6B5E6" authority="Elliot 1909" authorityName="Elliot" authorityYear="1909" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="brockmani">
<emphasis id="E2AE04C8837A27E97C14E5960C98A7B4" italics="true" pageNumber="166">Papio brockmani</emphasis>
Elliot 1909
</taxonomicName>
;
<taxonomicName id="D0B08D8AE6D728EF57EF390DC6BB803E" authority="(Lesson 1840)" baseAuthorityName="Lesson" baseAuthorityYear="1840" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="chaeropitheus">
<emphasis id="4256E5AA0A168C6588A1433CDA48C820" italics="true" pageNumber="166">Papio chaeropitheus</emphasis>
(
<bibRefCitation id="7BFF603D49BFEF543492F9ABA1FDD2E5" author="Lesson, R. P." refId="ref210535" refString="Lesson, R. P. 1840. Species es mammiferes bimanes et quadrimanes ... Bailliere, London, 292 pp." year="1840">Lesson 1840</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="35BA7C053EF5D8355933F5F31FA3ECBA" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="cynamolgus">
<emphasis id="BE6EA6DF80218D180F64B1E611965DB5" italics="true" pageNumber="166">Papio cynamolgus</emphasis>
(
<bibRefCitation id="E784C793DBF62582DD156C2D3C048775" author="Linnaeus, C." refId="ref213949" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tenth ed. Vol. 1. Laurentii Salvii, Stockholm, 824 pp." year="1758">Linnaeus 1758</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="635DC251072FA7009D86AFCE3A1E3361" authority="(Reichenbach 1863)" baseAuthorityName="Reichenbach" baseAuthorityYear="1863" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="nedjo">
<emphasis id="9CCE2C93F2E5B6E0FF0CC801B0949031" italics="true" pageNumber="166">Papio nedjo</emphasis>
(Reichenbach 1863)
</taxonomicName>
;
<taxonomicName id="6A7A2BC812E21B48A0914BBE02DE8D62" authority="(Agassiz 1828)" baseAuthorityName="Agassiz" baseAuthorityYear="1828" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" pageNumber="166" phylum="Chordata" rank="species" species="wagleri">
<emphasis id="A34B296B19E6318937BEEAD2EDB3A327" italics="true" pageNumber="166">Papio wagleri</emphasis>
(Agassiz 1828)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FC79743CFCEE74A83E0B3498A3FB4F6C" pageNumber="166" type="distribution">
<paragraph id="38B206503F2FD823C07FA3CF3FAD2BE8" pageNumber="166">
<emphasis id="5A17C53C349F215EC7A8453E8C2E2E75" inLineHeading="true" pageNumber="166">Distribution:</emphasis>
Arid zone of
<collectingRegion id="1787EE55EA8762BFE9F433A01CC31644" country="Sudan" name="Al Bahr al Ahmar">Red Sea</collectingRegion>
coast of
<collectingCountry id="A394BB90C40020C6684AB45C534BFD7B" name="Sudan">Sudan</collectingCountry>
,
<collectingCountry id="8E871FEEE774940E04153B48073863C9" name="Eritrea">Eritrea</collectingCountry>
,
<collectingCountry id="FAC774B0120F95BF6C714DBC7DC0D26A" name="Ethiopia">Ethiopia</collectingCountry>
, N
<collectingCountry id="3CD91414D3F1B0C3692C57CDE1FA4824" name="Somalia">Somalia</collectingCountry>
,
<collectingCountry id="D10BC90FE6EF91319296652FADFC8FA3" name="Yemen">Yemen</collectingCountry>
,
<collectingCountry id="793E58710FC04C2FAAB46CAAD2FC1C4F" name="Saudi Arabia">Saudi Arabia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E2F49834107D526C6A1E2F1E7895C5B5" pageNumber="166" type="conservation">
<paragraph id="D93E981F961413E1FA9D51F7E4AAD7B3" pageNumber="166">
<emphasis id="FBA008C55A569C4C48AAF856050D8149" inLineHeading="true" pageNumber="166">Conservation:</emphasis>
<collectionCode id="E14E818BA3ABD4EE8F0C4866506FDF13">CITES</collectionCode>
Appendix II;
<collectionCode id="78019AAE1E22F102E7AA2D16C0F24E5E">IUCN</collectionCode>
Lower Risk (nt).
</paragraph>
</subSubSection>
<subSubSection id="7707567E1E7B97F41DA03D8306BE866D" pageNumber="166" type="discussion">
<paragraph id="B3A49714EDFEA86AAC65FA67A09C8CBD" pageNumber="166">
<emphasis id="E0D077B8D4A191167E398739CF854AE4" inLineHeading="true" pageNumber="166">Discussion:</emphasis>
Includes
<taxonomicName id="EF969C011E6CF576E7C5E900E7ADFFDC" authorityName="Lesson" authorityYear="1827" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="anubis">
<emphasis id="BF49153833530ACE834137C0102CD0B5" italics="true" pageNumber="166">anubis</emphasis>
</taxonomicName>
,
<taxonomicName id="D5302E94979C84C6C0B613DFE7D37E45" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="cynocephalus">
<emphasis id="C9BAF6E9F521861451DBA72407DBED0E" italics="true" pageNumber="166">cynocephalus</emphasis>
</taxonomicName>
,
<taxonomicName id="31497AF5293D8B00667C669FF2749707" authorityName="Desmarest" authorityYear="1820" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="papio">
<emphasis id="437A47DC278DD89E4BD6BFA8B3AA7FEA" italics="true" pageNumber="166">papio</emphasis>
</taxonomicName>
, and
<taxonomicName id="7603403B451BC557D22152C9E6A132D9" authorityName="Kerr" authorityYear="1792" class="Mammalia" family="Cercopithecidae" genus="Papio" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="ursinus">
<emphasis id="E5F77558C3F3F02D5506E9D0E3E9382A" italics="true" pageNumber="166">ursinus</emphasis>
</taxonomicName>
according to
<bibRefCitation id="DF270420492F1DA496FEFF4CDCA7141B" author="Szalay, F. S. &amp; E. Delson" refId="ref345717" refString="Szalay, F. S., and E. Delson. 1979. Evolutionary history of the primates. Academic Press, New York, 580 pp." year="1979">Szalay and Delson (1979:336)</bibRefCitation>
, but see
<bibRefCitation id="F9505480BC1B0E90E3415581613D0BF6" author="Jolly, C. J. &amp; F. L. Brett" refId="ref175177" refString="Jolly, C. J., and F. L. Brett. 1973. Genetic markers and baboon biology. Journal of Medical Primatology, 2 (2): 85 - 99." year="1973">Jolly and Brett (1973:85)</bibRefCitation>
,
<bibRefCitation id="7ECD9E76E016EFB70525A2A15D97CCE9" author="Dandelot, P." refId="ref73122" refString="Dandelot, P. 1974. Order Primates. Part 3. Pp. 1 - 45, in The mammals of Africa: An identification manual (J. Meester and H. W. Setzer, eds.) [issued 10 Sep 1974]. Smithsonian Institution Press, Washington, D. C., not continuously paginated." year="1974">Dandelot (1974:9)</bibRefCitation>
,
<bibRefCitation id="020FCB09D15D62BB18BC0A58DFA0B6EF" author="Corbet, G. B. &amp; J. E. Hill" refId="ref67843" refString="Corbet, G. B., and J. E. Hill. 1980. A world list of mammalian species. British Museum (Natural History), London, 226 pp." year="1980">Corbet and Hill (1980:88)</bibRefCitation>
,
<bibRefCitation id="40CFF6011522F8AC8DF398043C5F7C52" author="Groves, C. P." refId="ref132585" refString="Groves, C. P. 2001 c. Primate taxonomy. Smithsonian Institution Press, Washington, DC, 350 pp." year="2001">
Groves (2001
<emphasis id="DD4A8CC3C9DC29EB02EDF5C0FFC6D604" italics="true" pageNumber="166">c</emphasis>
)
</bibRefCitation>
and others who recognized these as distinct species.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,454 @@
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<mods:title id="0D692D5470B4F0B50536DDA44A9D4F32">Description of a new species of Cephalaeschna (Odonata: Anisoptera: Aeshnidae) from northern Vietnam</mods:title>
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<emphasis id="E4E59D67FF8ADB1EA1A53004FE61F988" bold="true" box="[151,494,1563,1589]" italics="true" pageId="0" pageNumber="334">Cephalaeschna yanagisawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFD6201CFF8ADB1EA0C73004FDDDF988" box="[501,594,1563,1589]" pageId="0" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D62E4175FF8ADB1EA1A53022FE9EF9EB" blockId="0.[151,594,1563,1622]" box="[151,273,1597,1622]" pageId="0" pageNumber="334">
(
<figureCitation id="4EAA5DF0FF8ADB1EA1AD3022FE86F9EB" box="[159,265,1597,1622]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1304,1326]" captionStartId-1="2.[151,250,1935,1957]" captionStartId-2="3.[151,250,1970,1992]" captionTargetBox-0="[151,1336,377,1255]" captionTargetBox-1="[186,1425,537,1800]" captionTargetBox-2="[151,1421,459,1921]" captionTargetId-0="figure@1.[151,1435,373,1282]" captionTargetId-1="figure@2.[151,1437,483,1911]" captionTargetId-2="figure@3.[151,1436,459,1936]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Lateral habitus of C. yanagisawai holotype male (scale bar: 1cm)." captionText-1="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." captionText-2="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri-0="https://zenodo.org/record/1439752/files/figure.png" httpUri-1="https://zenodo.org/record/1439754/files/figure.png" httpUri-2="https://zenodo.org/record/1439756/files/figure.png" pageId="0" pageNumber="334">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9E8B12FEFF8ADB1EA1A53099FA96F893" pageId="0" pageNumber="334" type="materials_examined">
<paragraph id="D62E4175FF8ADB1EA1A53099FD33F95B" blockId="0.[151,1436,1669,2019]" pageId="0" pageNumber="334">
<materialsCitation id="66F94B28FF8ADB1EA1A53099FB20F923" ID-GBIF-Occurrence="1913888926" box="[151,1199,1669,1695]" location="Hoang Lien National Park" pageId="0" pageNumber="334" specimenCount="1" specimenCount-male="1" stateProvince="Lai Chau Province" typeStatus="holotype">
<emphasis id="E4E59D67FF8ADB1EA1A53099FE84F923" bold="true" box="[151,267,1670,1694]" pageId="0" pageNumber="334">
<typeStatus id="092AFFD7FF8ADB1EA1A53099FE89F923" box="[151,262,1670,1694]" pageId="0" pageNumber="334" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="C0978AFCFF8ADB1EA0203098FEA8F923" box="[274,295,1671,1694]" pageId="0" pageNumber="334" type="male"></specimenCount>
(NSWT-I-Od-32335) (
<figureCitation id="4EAA5DF0FF8ADB1EA318309AFDE4F923" box="[554,619,1669,1694]" captionStart="FIGURE 1" captionStartId="1.[151,250,1304,1326]" captionTargetBox="[151,1336,377,1255]" captionTargetId="figure@1.[151,1435,373,1282]" captionTargetPageId="1" captionText="FIGURE 1. Lateral habitus of C. yanagisawai holotype male (scale bar: 1cm)." httpUri="https://zenodo.org/record/1439752/files/figure.png" pageId="0" pageNumber="334">Fig. 1</figureCitation>
),
<location id="D34E17AEFF8ADB1EA3B63098FC22F923" LSID="urn:lsid:plazi:treatment:5E38F063FF8ADB1AA1A53004FBE3F925:D34E17AEFF8ADB1EA3B63098FC22F923" box="[644,941,1669,1695]" name="Hoang Lien National Park" pageId="0" pageNumber="334" stateProvince="Lai Chau Province">Hoang Lien National Park</location>
,
<collectingRegion id="14558F97FF8ADB1EA28A309AFB02F923" box="[952,1165,1669,1694]" country="Vietnam" name="Lai Chau" pageId="0" pageNumber="334">
Lai
<collectingRegion id="14558F97FF8ADB1EA2D7309AFB02F923" box="[997,1165,1669,1694]" inferredBySuffix="true" pageId="0" pageNumber="334">Chau Province</collectingRegion>
</collectingRegion>
, N
</materialsCitation>
.
<materialsCitation id="66F94B28FF8ADB1EA588309AFD37F95B" ID-GBIF-Occurrence="1913888925" collectingDate="2017-06-29" collectorName="Takashi Yanagisawa &amp; The &amp; Science" country="Vietnam" elevation="2000" latitude="22.333334" location="National Museum of Nature" longLatPrecision="1260" longitude="103.75" pageId="0" pageNumber="334" specimenCount="1" stateProvince="Tokyo" typeStatus="holotype">
<collectingCountry id="AE8601E5FF8ADB1EA588309AFA96F923" box="[1210,1305,1669,1694]" name="Vietnam" pageId="0" pageNumber="334">Vietnam</collectingCountry>
(
<geoCoordinate id="B3A527B2FF8ADB1EA41B309AFA17F923" box="[1321,1432,1669,1694]" direction="east" orientation="longitude" pageId="0" pageNumber="334" precision="925" value="103.75">103°45E</geoCoordinate>
,
<geoCoordinate id="B3A527B2FF8ADB1EA1A530B5FF75F97F" box="[151,250,1706,1730]" direction="north" orientation="latitude" pageId="0" pageNumber="334" precision="925" value="22.333334">22°20N</geoCoordinate>
, 1900 ~
<elevation id="5DBCA646FF8ADB1EA05B30B5FE4CF97F" box="[361,451,1706,1730]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="334" unit="m" value="2000.0">
<quantity id="1169EC90FF8ADB1EA05B30B5FE4CF97F" box="[361,451,1706,1730]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="334" unit="m" value="2000.0">2000 m</quantity>
</elevation>
a.l.s.),
<collectingDate id="B26B9E5DFF8ADB1EA32C30B5FD29F97F" box="[542,678,1706,1730]" pageId="0" pageNumber="334" value="2017-06-29">
<date id="A22F67B5FF8ADB1EA32C30B5FD29F97F" box="[542,678,1706,1730]" pageId="0" pageNumber="334" value="2017-06-29">29-VI-2017</date>
</collectingDate>
;
<collectorName id="7B6424A3FF8ADB1EA38430B5FC11F97E" box="[694,926,1706,1731]" pageId="0" pageNumber="334">Takashi Yanagisawa</collectorName>
leg.
<collectorName id="7B6424A3FF8ADB1EA2ED30B5FB84F97E" box="[991,1035,1706,1731]" pageId="0" pageNumber="334">The</collectorName>
<typeStatus id="092AFFD7FF8ADB1EA52430B5FBF7F97E" box="[1046,1144,1706,1731]" pageId="0" pageNumber="334" type="holotype">holotype</typeStatus>
will be deposited in the
<location id="D34E17AEFF8ADB1EA1A530D2FE5DF95B" LSID="urn:lsid:plazi:treatment:5E38F063FF8ADB1AA1A53004FBE3F925:D34E17AEFF8ADB1EA1A530D2FE5DF95B" box="[151,466,1741,1766]" country="Vietnam" latitude="22.333334" longLatPrecision="1260" longitude="103.75" name="National Museum of Nature" pageId="0" pageNumber="334" stateProvince="Tokyo">National Museum of Nature</location>
and
<collectorName id="7B6424A3FF8ADB1EA33A30D2FDEDF95B" box="[520,610,1741,1766]" pageId="0" pageNumber="334">Science</collectorName>
,
<collectingRegion id="14558F97FF8ADB1EA35F30D2FD37F95B" box="[621,696,1741,1766]" country="Japan" name="Tokyo" pageId="0" pageNumber="334">Tokyo</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="D62E4175FF8ADB1EA1F530EDFA96F893" blockId="0.[151,1436,1669,2019]" pageId="0" pageNumber="334">
<typeStatus id="092AFFD7FF8ADB1EA1F530EDFEB0F8B7" box="[199,319,1778,1802]" pageId="0" pageNumber="334" type="paratype">
<emphasis id="E4E59D67FF8ADB1EA1F530EDFEB0F8B7" bold="true" box="[199,319,1778,1802]" pageId="0" pageNumber="334">Paratypes</emphasis>
</typeStatus>
.
<specimenCount id="C0978AFCFF8ADB1EA07F30EDFEFEF8B7" box="[333,369,1778,1802]" pageId="0" pageNumber="334" type="male">1♂</specimenCount>
, same place as the
<typeStatus id="092AFFD7FF8ADB1EA37930EDFD3FF8B7" box="[587,688,1778,1802]" pageId="0" pageNumber="334" type="holotype">holotype</typeStatus>
,
<date id="A22F67B5FF8ADB1EA38E30EDFCDAF8B7" box="[700,853,1778,1802]" pageId="0" pageNumber="334" value="2017-08-16">16-VIII-2017</date>
, Kenji Yashiro leg.;
<specimenCount id="C0978AFCFF8ADB1EA50F30EDFBEFF8B7" box="[1085,1120,1778,1802]" pageId="0" pageNumber="334" type="female">1♀</specimenCount>
(
<figureCitation id="4EAA5DF0FF8ADB1EA54230EDFB4DF8B7" box="[1136,1218,1778,1802]" captionStart="FIGURE 3" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[151,1421,459,1921]" captionTargetId="figure@3.[151,1436,459,1936]" captionTargetPageId="3" captionText="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="0" pageNumber="334">Fig. 3a</figureCitation>
), same place as the
<typeStatus id="092AFFD7FF8ADB1EA1A5310AFF73F893" box="[151,252,1813,1838]" pageId="0" pageNumber="334" type="holotype">holotype</typeStatus>
,
<date id="A22F67B5FF8ADB1EA0353109FE0BF893" box="[263,388,1814,1838]" pageId="0" pageNumber="334" value="2017-10-29">29-X-2017</date>
, Kenji Yashiro leg. The
<typeStatus id="092AFFD7FF8ADB1EA3A73108FC8FF892" box="[661,768,1815,1839]" pageId="0" pageNumber="334" type="paratype">paratypes</typeStatus>
are preserved in Mr. Kozo Yashiros collection.
</paragraph>
</subSubSection>
<subSubSection id="9E8B12FEFF8ADB1EA1F53125FEC7F8CB" pageId="0" pageNumber="334" type="etymology">
<paragraph id="D62E4175FF8ADB1EA1F53125FEC7F8CB" blockId="0.[151,1436,1669,2019]" pageId="0" pageNumber="334">
<emphasis id="E4E59D67FF8ADB1EA1F53125FEC9F8EF" bold="true" box="[199,326,1850,1874]" pageId="0" pageNumber="334">Etymology</emphasis>
. The species name, a noun in the genitive case, is dedicated to Mr. Takashi Yanagisawa, collector of the
<typeStatus id="092AFFD7FF8ADB1EA1EC3142FECBF8CB" box="[222,324,1885,1910]" pageId="0" pageNumber="334" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="9E8B12FEFF8ADB1AA1F5319EFADEFD15" lastPageId="4" lastPageNumber="338" pageId="0" pageNumber="334" type="description">
<paragraph id="D62E4175FF8ADB1FA1F5319EFF59FEFD" blockId="0.[151,1436,1669,2019]" lastBlockId="1.[151,1436,151,320]" lastPageId="1" lastPageNumber="335" pageId="0" pageNumber="334">
<emphasis id="E4E59D67FF8ADB1EA1F5319EFD86F827" bold="true" box="[199,521,1921,1946]" pageId="0" pageNumber="334">
Description of the
<typeStatus id="092AFFD7FF8ADB1EA090319DFD86F827" box="[418,521,1922,1946]" pageId="0" pageNumber="334" type="holotype">holotype</typeStatus>
</emphasis>
.
<emphasis id="E4E59D67FF8ADB1EA325319DFDDBF826" box="[535,596,1922,1947]" italics="true" pageId="0" pageNumber="334">Head</emphasis>
(
<figureCitation id="4EAA5DF0FF8ADB1EA351319DFD3BF827" box="[611,692,1922,1947]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="0" pageNumber="334">Fig. 2a</figureCitation>
). Labium brown with partly blackish shades; labrum brown with a large transverse oblong greenish yellow spot, a little split at the center by vertical brown line; genae brown with largely greenish yellow on upper part; ante- and postclypeus brown, with dark brown at the lower border of anteclypeus, and vaguely yellowish in the pits of lower side of postclypeus and on the upper part of the same; antefrons brown, much darker on upper frontal and dorsal sides; dorso-frontal apex of antefrons is obtusely but distinctly angled; vertex and antennae black, partly dark brown. The width of anteforns is about 0.65 times of the distance between the ends of compound eyes. The compound eye in life is of emerald blue, margined with green color.
</paragraph>
<caption id="82EE11FDFF8BDB1FA1A53307FC3AFA93" box="[151,949,1304,1326]" httpUri="https://zenodo.org/record/1439752/files/figure.png" pageId="1" pageNumber="335" startId="1.[151,250,1304,1326]" targetBox="[151,1336,377,1255]" targetPageId="1">
<paragraph id="D62E4175FF8BDB1FA1A53307FC3AFA93" blockId="1.[151,949,1304,1326]" box="[151,949,1304,1326]" pageId="1" pageNumber="335">
<emphasis id="E4E59D67FF8BDB1FA1A53307FE9CFA90" bold="true" box="[151,275,1304,1326]" pageId="1" pageNumber="335">FIGURE 1.</emphasis>
Lateral habitus of
<taxonomicName id="11913AF6FF8BDB1FA0E33306FDE0FA93" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[465,623,1305,1326]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="335" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8BDB1FA0E33306FDE0FA93" box="[465,623,1305,1326]" italics="true" pageId="1" pageNumber="335">C. yanagisawai</emphasis>
</taxonomicName>
holotype male (scale bar: 1cm).
</paragraph>
</caption>
<paragraph id="D62E4175FF8BDB1FA1F53345FD0AF92E" blockId="1.[151,1437,1370,2007]" pageId="1" pageNumber="335">
<emphasis id="E4E59D67FF8BDB1FA1F53345FE90FACE" box="[199,287,1370,1395]" italics="true" pageId="1" pageNumber="335">Thorax.</emphasis>
Prothorax brown, bordered with black. Synthorax dark brown with greenish yellow markings as follows; mesepisternum (ref.
<figureCitation id="4EAA5DF0FF8BDB1FA0D63361FDB9FA2A" box="[484,566,1406,1431]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="1" pageNumber="335">Fig. 2b</figureCitation>
) with antehumeral stripe, turning gently inwards and upwards, and its upper portion abruptly curving outwards; antealar sinus with greenish yellow above the ridge; mesepimeron with broad stripe, slightly constricted at a little above the middle, then narrower in upper portion and changing direction a little anteriorly; metaepisternum with a small inverted triangular spot at the dorsal border and with tiny vague vertical spot at the middle; interalar sclerite greenish; metaepimeron broadly greenish, except for the ventral and anterior margin; mesoinfraepisternum and metainfraepisternum each with a tiny spot at posterior corner; ventral side pale brown. Legs black, coxa, trochanter and basal femur tinted with dark brown; fore-leg with small greenish spots at inner side of trochanter and the basal femur.
</paragraph>
<paragraph id="D62E4175FF8BDB1FA1F530BFFC27F8FA" blockId="1.[151,1437,1370,2007]" pageId="1" pageNumber="335">
<emphasis id="E4E59D67FF8BDB1FA1F530BFFE9DF90A" box="[199,274,1696,1719]" italics="true" pageId="1" pageNumber="335">Wings.</emphasis>
Hyaline, faintly tinted with pale brown. Nodal index: antenodal veins (An) 21 (right) and 22 (left) in forewing (Fw), 16 (both) in hind wing (Hw); postnodal veins (Pn) 15 (both) in Fw, 18 (right) and 19 (left) in Hw. Pterostigma (Pt) reddish brown, covering 2.5 to 3 cell-length in Fw and
<quantity id="1169EC90FF8BDB1FA2EE30F8FBD9F942" box="[988,1110,1766,1791]" metricMagnitude="-2" metricUnit="m" metricValue="5.715" metricValueMax="6.35" metricValueMin="5.08" pageId="1" pageNumber="335" unit="in" value="2.25" valueMax="2.5" valueMin="2.0">2 to 2.5 in</quantity>
Hw, with an embraced vein situated at or slightly interior to the proximal vein of Pt in both wings. Median space with 4 cross-veins in both wings; triangle 5 cells; anal loop contains 9 cells; anal triangle 4 cells.
</paragraph>
<paragraph id="D62E4175FF8BDB1CA1F5314DFAC5FE82" blockId="1.[151,1437,1370,2007]" lastBlockId="2.[151,1436,151,464]" lastPageId="2" lastPageNumber="336" pageId="1" pageNumber="335">
<emphasis id="E4E59D67FF8BDB1FA1F5314DFEBEF8D6" box="[199,305,1874,1899]" italics="true" pageId="1" pageNumber="335">Abdomen</emphasis>
(
<figureCitation id="4EAA5DF0FF8BDB1FA077314DFE13F8D6" box="[325,412,1874,1899]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="1" pageNumber="335">Fig. 2c</figureCitation>
). Slender cylindrical, most swelling at 2nd segment (S); colored dark brown or black, slightly pale ventrally, with greenish yellow maculation as follows. S1 hairy dorsally, with vertical thin green spot near postero-ventral border; S2 with arrowhead-like spot on the anterior half of dorsum, with a pair of thin median dorsal (MD) markings obliquely continuous to the antero-ventral marking, covering auricle, and with annular postero-dorsal (PD) marking, which is expanded ventrally; MD oblique markings present on S2 to S9, gradually inconspicuous and situating anteriad in segment towards the back; at antero-ventral part of each tergite of S3 to S10, faintly yellow vague tints, connecting with MD marking; PD markings present on S2 to S6, those on S3 and S4 semi-circular, that on S5 almost linear, and that on S6 rudimentary; A pair of inconspicuous yellow spots on distal fourth of dorsa of S3 to S7; S10 without markings, except for dark yellowish at antero-ventral corner.
</paragraph>
<paragraph id="D62E4175FF88DB1CA1F53752FD8DFE6D" blockId="2.[151,1436,151,464]" pageId="2" pageNumber="336">
<emphasis id="E4E59D67FF88DB1CA1F53752FE89FED9" box="[199,262,333,356]" italics="true" pageId="2" pageNumber="336">Penis</emphasis>
(
<figureCitation id="4EAA5DF0FF88DB1CA0263753FEF0FED8" box="[276,383,332,357]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="2" pageNumber="336">Fig. 2fg</figureCitation>
). dark brown with whitish membranous parts; vesicle robust, trapezoidal shape in lateral view; 2nd and 3rd segments are almost similar length, with a dorsal protuberance on antero-dorsum of 2nd; distal segment triangular in lateral view with a pair of long flagelliform projections originating from ventral corner and with ear-like lobules on dorsum.
</paragraph>
<caption id="82EE11FDFF88DB1CA1A53190FCFAF857" httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="2" pageNumber="336" startId="2.[151,250,1935,1957]" targetBox="[186,1425,537,1800]" targetPageId="2">
<paragraph id="D62E4175FF88DB1CA1A53190FCFAF857" blockId="2.[151,1436,1935,2026]" pageId="2" pageNumber="336">
<emphasis id="E4E59D67FF88DB1CA1A53190FE9CF819" bold="true" box="[151,275,1935,1957]" pageId="2" pageNumber="336">FIGURE 2.</emphasis>
Male of
<taxonomicName id="11913AF6FF88DB1CA042318FFD81F818" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[368,526,1936,1957]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF88DB1CA042318FFD81F818" box="[368,526,1936,1957]" italics="true" pageId="2" pageNumber="336">C. yanagisawai</emphasis>
</taxonomicName>
(a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view.
</paragraph>
</caption>
<paragraph id="D62E4175FF89DB1DA1F53688FF5BFED8" blockId="3.[151,1436,151,392]" pageId="3" pageNumber="337">
<emphasis id="E4E59D67FF89DB1DA1F53688FE05FF0D" box="[199,394,151,176]" italics="true" pageId="3" pageNumber="337">Anal appendages</emphasis>
. Dark brown. Cercus (superior appendages), in dorsal view (
<figureCitation id="4EAA5DF0FF89DB1DA5773688FB15FF0D" box="[1093,1178,151,176]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="3" pageNumber="337">Fig. 2d</figureCitation>
), basally narrow, from basal 2/5 expanded inwards, then from its half gradually tapering distad; the apex round; the dorsal longitudinal ridge running from middle to the apex. Cercus, in lateral view (
<figureCitation id="4EAA5DF0FF89DB1DA24136C0FC48FF45" box="[883,967,223,248]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="3" pageNumber="337">Fig. 2e</figureCitation>
), slender and gently warped downwards, with an obtusely shallow triangular ventral protrusion at apical forth. Epiproct (inferior appendage), about 0.72 times as the length of cercus, narrow triangle shape in dorsal view, and bow shape, warped downwards, in lateral view.
</paragraph>
<paragraph id="D62E4175FF89DB1DA1F5376FFC47FE35" blockId="3.[151,1436,151,392]" box="[199,968,367,392]" pageId="3" pageNumber="337">Measurements (mm): Hw 44.0, abdomen plus anal appendages 53.8.</paragraph>
<caption id="82EE11FDFF89DB1DA1A531ADFD83F857" httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="3" pageNumber="337" startId="3.[151,250,1970,1992]" targetBox="[151,1421,459,1921]" targetPageId="3">
<paragraph id="D62E4175FF89DB1DA1A531ADFD83F857" blockId="3.[151,1436,1970,2026]" pageId="3" pageNumber="337">
<emphasis id="E4E59D67FF89DB1DA1A531ADFE9DF87A" bold="true" box="[151,274,1970,1992]" pageId="3" pageNumber="337">FIGURE 3.</emphasis>
Paratype female of
<taxonomicName id="11913AF6FF89DB1DA0EC31ADFDF4F875" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[478,635,1970,1992]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF89DB1DA0EC31ADFDF4F875" box="[478,635,1970,1992]" italics="true" pageId="3" pageNumber="337">C. yanagisawai</emphasis>
</taxonomicName>
. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite.
</paragraph>
</caption>
<paragraph id="D62E4175FF8EDB1AA1F53688FD61FEA0" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="338">
<emphasis id="E4E59D67FF8EDB1AA1F53688FDBEFF0D" bold="true" box="[199,561,151,176]" pageId="4" pageNumber="338">
Variation in a
<typeStatus id="092AFFD7FF8EDB1AA0493687FE69FF0D" box="[379,486,152,176]" pageId="4" pageNumber="338" type="paratype">paratype</typeStatus>
male.
</emphasis>
There are only small variations in a
<typeStatus id="092AFFD7FF8EDB1AA2D73687FBCAFF0D" box="[997,1093,152,176]" pageId="4" pageNumber="338" type="paratype">paratype</typeStatus>
male as follows. Wing nodal index: An
<quantity id="1169EC90FF8EDB1AA03F36A3FEC9FF68" box="[269,326,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="4" pageNumber="338" unit="in" value="24.0">24 in</quantity>
Fw, 17 and
<quantity id="1169EC90FF8EDB1AA0FD36A3FD88FF68" box="[463,519,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="4" pageNumber="338" unit="in" value="18.0">18 in</quantity>
Hw; Pn 15 and
<quantity id="1169EC90FF8EDB1AA38E36A3FD7AFF68" box="[700,757,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="4" pageNumber="338" unit="in" value="16.0">16 in</quantity>
Fw, 17 and
<quantity id="1169EC90FF8EDB1AA24F36A3FC39FF68" box="[893,950,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="4" pageNumber="338" unit="in" value="18.0">18 in</quantity>
Hw; anal loops 10 or 11 cells, anal triangle 3 or 4 cells. Abdominal maculation is similar to the
<typeStatus id="092AFFD7FF8EDB1AA3C836C0FCD0FF45" box="[762,863,223,248]" pageId="4" pageNumber="338" type="holotype">holotype</typeStatus>
, but PD markings on S6 invisible. Measurements (mm): Hw 41.9, abdomen plus anal appendages 50.6.
</paragraph>
<paragraph id="D62E4175FF8EDB1AA1F53737FADEFD15" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="338">
<emphasis id="E4E59D67FF8EDB1AA1F53737FE91FEFD" bold="true" box="[199,286,296,320]" pageId="4" pageNumber="338">Female</emphasis>
. The body shape is generally thicker than in male. The background color is brownish and maculation is more yellowish than in male, but its pattern is similar to that of male (
<figureCitation id="4EAA5DF0FF8EDB1AA2E73753FBA5FED9" box="[981,1066,332,357]" captionStart="FIGURE 3" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[151,1421,459,1921]" captionTargetId="figure@3.[151,1436,459,1936]" captionTargetPageId="3" captionText="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="4" pageNumber="338">Fig. 3a</figureCitation>
). Head: the yellow markings of labium are thinner and more ambiguous (
<figureCitation id="4EAA5DF0FF8EDB1AA3563770FD39FE35" box="[612,694,367,392]" captionStart="FIGURE 3" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[151,1421,459,1921]" captionTargetId="figure@3.[151,1436,459,1936]" captionTargetPageId="3" captionText="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="4" pageNumber="338">Fig. 3b</figureCitation>
); postcylepus with a pair of two dark yellow spots around the pits; the width of antefrons is about 0.67 times as wide as the distance between the ends of compound eyes. Wing generally tinted with brown and basally yellow; nodal index: An 23 and
<quantity id="1169EC90FF8EDB1AA2F837A8FB8CFE6D" box="[970,1027,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="4" pageNumber="338" unit="in" value="25.0">25 in</quantity>
Fw, 18 and
<quantity id="1169EC90FF8EDB1AA5BD37A7FB47FE6D" box="[1167,1224,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="4" pageNumber="338" unit="in" value="19.0">19 in</quantity>
Hw; Pn 15 and
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Fw, 16 and
<quantity id="1169EC90FF8EDB1AA00A37C3FEFDFE48" box="[312,370,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="4" pageNumber="338" unit="in" value="19.0">19 in</quantity>
Hw; irregular halfway subcostal extension is found in only left Fw; triangle and median space are crossed same as in male; anal loops contain 11 or 13 cells. Abdomen (
<figureCitation id="4EAA5DF0FF8EDB1AA53837E0FBD1FDA5" box="[1034,1118,511,536]" captionStart="FIGURE 3" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[151,1421,459,1921]" captionTargetId="figure@3.[151,1436,459,1936]" captionTargetPageId="3" captionText="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="4" pageNumber="338">Fig. 3a</figureCitation>
) thick cylindrical, and less constricted in S3 than male; ground color dark brown and maculation patterns are almost same as males, but S9 without any markings: the postero-ventral part of tergite of S10 projecting posteriorly, each tip of which on both side is fused (
<figureCitation id="4EAA5DF0FF8EDB1AA0043473FE3DFD38" box="[310,434,620,645]" captionStart="FIGURE 3" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[151,1421,459,1921]" captionTargetId="figure@3.[151,1436,459,1936]" captionTargetPageId="3" captionText="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="4" pageNumber="338">Figs. 3cd</figureCitation>
); semi-membranous structure is formed between each projections. The tip of ovipositor valves slightly beyond the level of dorsum of S10. Measurements (mm): Hw 45.0, abdomen plus cerci 47.0.
</paragraph>
</subSubSection>
<subSubSection id="9E8B12FEFF8EDB1AA1F534ACFBE3F925" pageId="4" pageNumber="338" type="diagnosis">
<paragraph id="D62E4175FF8EDB1AA1F534ACFAD1FCE0" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="338">
<emphasis id="E4E59D67FF8EDB1AA1F534ACFECEFD71" bold="true" box="[199,321,691,716]" pageId="4" pageNumber="338">Diagnosis.</emphasis>
The external morphology of most
<taxonomicName id="11913AF6FF8EDB1AA3F734ABFCFDFD70" authorityName="Selys" authorityYear="1883" box="[709,882,692,717]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="E4E59D67FF8EDB1AA3F734ABFCFDFD70" box="[709,882,692,717]" italics="true" pageId="4" pageNumber="338">Cephalaeschna</emphasis>
</taxonomicName>
species more or less resemble each other, but can be identified by combinations of the following characters in male (
<bibRefCitation id="B2003C84FF8EDB1AA29734C8FBC6FD4D" box="[933,1097,727,752]" pageId="4" pageNumber="338" refId="ref2801" refString="Asahina, S. (1983) Further contributions to the knowledge of Nepalese Cephalaeschna and its allies (Odonata, Aeshnidae). Bulletin of the National Science Museum, Tokyo, Series A, 8 (2), 51 - 67." type="journal article">Asahina 1983</bibRefCitation>
;
<bibRefCitation id="B2003C84FF8EDB1AA56A34C7FB6AFD4D" box="[1112,1253,727,752]" pageId="4" pageNumber="338" refId="ref2846" refString="Fraser, F. C. (1936) The Fauna of British India, including Ceylon and Burma. Vol. 3. Taylor and Francis, Ltd., London, 461 pp." type="book">Fraser 1936</bibRefCitation>
;
<bibRefCitation id="B2003C84FF8EDB1AA5C134C8FF5AFCA9" pageId="4" pageNumber="338" refId="ref2882" refString="Joshi, S. &amp; Kunte, K. (2017) Two new dragonfly species (Odonata: Anisoptera: Aeshnidae) from north-eastern India. Zootaxa, 4300 (2), 259 - 268. https: // doi. org / 10.11646 / zootaxa. 4300.2.7" type="journal article">Joshi &amp; Kunte 2017</bibRefCitation>
): (a) pointedness and width of antefrons, (b) marking on labrum, (c) maculation patterns on synthorax and abdomen, (d) the number of cells of anal triangle (3 or 4 vs ± 5), and (e) morphology of anal appendages, which can roughly be classified into two
<typeStatus id="092AFFD7FF8EDB1AA32A355AFDD9FCE0" box="[536,598,837,861]" pageId="4" pageNumber="338">types</typeStatus>
, i.e. the apex of cercus ending rounded (or obtuse) vs acutely angled.
</paragraph>
<paragraph id="D62E4175FF8EDB1AA1F53578FC46FBAD" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="338">
The degree of acuteness in dorsal antefrons is characteristic in each species. The antefrons of
<taxonomicName id="11913AF6FF8EDB1AA5E73577FA0DFC3D" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[1237,1410,872,896]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8EDB1AA5E73577FA0DFC3D" box="[1237,1410,872,896]" italics="true" pageId="4" pageNumber="338">C. yanagisawai</emphasis>
</taxonomicName>
is obtusely angled (
<figureCitation id="4EAA5DF0FF8EDB1AA06E3593FE23FC19" box="[348,428,908,933]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="4" pageNumber="338">Fig. 2a</figureCitation>
&amp;
<figureCitation id="4EAA5DF0FF8EDB1AA0E73593FE7BFC18" box="[469,500,908,933]" captionStart="FIGURE 3" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[151,1421,459,1921]" captionTargetId="figure@3.[151,1436,459,1936]" captionTargetPageId="3" captionText="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="4" pageNumber="338">3b</figureCitation>
), but neither pointed upwards nor smoothly rounded. This state a little resembles those of
<taxonomicName id="11913AF6FF8EDB1AA1C935AFFDB3FC75" authority="(Fraser, 1922)" authorityName="Fraser" authorityYear="1922" baseAuthorityName="Fraser" baseAuthorityYear="1922" box="[251,572,943,968]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="viridifrons">
<emphasis id="E4E59D67FF8EDB1AA1C935AFFE1CFC75" box="[251,403,943,968]" italics="true" pageId="4" pageNumber="338">C. viridifrons</emphasis>
(Fraser, 1922)
</taxonomicName>
(
<collectingCountry id="AE8601E5FF8EDB1AA37E35AFFD34FC75" box="[588,699,944,968]" name="Myanmar" pageId="4" pageNumber="338">Myanmar</collectingCountry>
to Himalaya),
<taxonomicName id="11913AF6FF8EDB1AA25B35AFFB35FC75" authority="Schmidt, 1961" authorityName="Schmidt" authorityYear="1961" box="[873,1210,943,968]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="E4E59D67FF8EDB1AA25B35AFFB84FC75" box="[873,1035,943,968]" italics="true" pageId="4" pageNumber="338">C. klapperichi</emphasis>
Schmidt, 1961
</taxonomicName>
(
<collectingCountry id="AE8601E5FF8EDB1AA5F835B0FADAFC75" box="[1226,1365,943,968]" name="Afghanistan" pageId="4" pageNumber="338">Afghanistan</collectingCountry>
to W. Himalaya), and
<taxonomicName id="11913AF6FF8EDB1AA07835CAFD05FC51" authority="Needham, 1930" authorityName="Needham" authorityYear="1930" box="[330,650,980,1005]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="patrorum">
<emphasis id="E4E59D67FF8EDB1AA07835CAFE5CFC51" box="[330,467,981,1004]" italics="true" pageId="4" pageNumber="338">C. patrorum</emphasis>
<bibRefCitation id="B2003C84FF8EDB1AA0EB35CBFD05FC51" box="[473,650,980,1005]" pageId="4" pageNumber="338" refId="ref2968" refString="Needham, J. G. (1930) A manual of the dragonflies of China. Zoologia Sinica, 11, 1 - 345, pls. I - XX, index 1 - 11." type="journal article">Needham, 1930</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="14558F97FF8EDB1AA3AA35CBFD63FC50" box="[664,748,980,1005]" country="China" name="Beijing" pageId="4" pageNumber="338">Beijing</collectingRegion>
to
<collectingRegion id="14558F97FF8EDB1AA23D35CBFCE1FC50" box="[783,878,980,1005]" country="China" name="Sichuan" pageId="4" pageNumber="338">Sichuan</collectingRegion>
), as far as we know. However
<taxonomicName id="11913AF6FF8EDB1AA5F335CAFAE1FC51" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[1217,1390,981,1004]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8EDB1AA5F335CAFAE1FC51" box="[1217,1390,981,1004]" italics="true" pageId="4" pageNumber="338">C. yanagisawai</emphasis>
</taxonomicName>
can easily be discriminated from the latters by maculation and other features.
</paragraph>
<paragraph id="D62E4175FF8EDB1AA1F53203FF4DFA21" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="338">
The following species have somewhat similar maculation to
<taxonomicName id="11913AF6FF8EDB1AA2423202FB91FB89" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[880,1054,1053,1076]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8EDB1AA2423202FB91FB89" box="[880,1054,1053,1076]" italics="true" pageId="4" pageNumber="338">C. yanagisawai</emphasis>
</taxonomicName>
, sharing with it an obtuse apex of cercus, but can be differentiated as described below:
<taxonomicName id="11913AF6FF8EDB1AA3C3325FFC73FBE5" authority="Asahina, 1982" authorityName="Asahina" authorityYear="1982" box="[753,1020,1087,1112]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="klotsi">
<emphasis id="E4E59D67FF8EDB1AA3C3325FFCDFFBE5" box="[753,848,1087,1112]" italics="true" pageId="4" pageNumber="338">C. klotsi</emphasis>
Asahina, 1982
</taxonomicName>
(
<collectingRegion id="14558F97FF8EDB1AA5393220FBFEFBE5" box="[1035,1137,1087,1112]" country="China" name="Zhejiang" pageId="4" pageNumber="338">Zhejiang</collectingRegion>
to
<collectingRegion id="14558F97FF8EDB1AA5A53220FB74FBE5" box="[1175,1275,1087,1112]" country="China" name="Guangxi" pageId="4" pageNumber="338">Guangxi</collectingRegion>
,
<collectingCountry id="AE8601E5FF8EDB1AA435325FFA1FFBE5" box="[1287,1424,1088,1112]" name="Hong Kong" pageId="4" pageNumber="338">Hong Kong</collectingCountry>
), which has narrow antefrons, i.e. the width of which is less than the half of the head width, exceptional in this genus (
<bibRefCitation id="B2003C84FF8EDB1AA1AC3298FEB3FB1D" box="[158,316,1159,1184]" pageId="4" pageNumber="338" refId="ref2801" refString="Asahina, S. (1983) Further contributions to the knowledge of Nepalese Cephalaeschna and its allies (Odonata, Aeshnidae). Bulletin of the National Science Museum, Tokyo, Series A, 8 (2), 51 - 67." type="journal article">Asahina 1983</bibRefCitation>
<bibRefCitation id="B2003C84FF8EDB1AA0713298FD91FB1D" box="[323,542,1159,1184]" pageId="4" pageNumber="338" refId="ref3070" refString="Wilson, K. D. P. &amp; Xu, Z. (2008) Aeshnidae of Guangdong and Hong Kong (China), with the descriptions of three new Planaeschna species (Anisoptera). Odonatologica, 37 (4), 329 - 360." type="journal article">Wilson &amp; Xu 2008</bibRefCitation>
), whereas the width of antefrons of
<taxonomicName id="11913AF6FF8EDB1AA2813297FBEEFB1D" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[947,1121,1160,1184]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8EDB1AA2813297FBEEFB1D" box="[947,1121,1160,1184]" italics="true" pageId="4" pageNumber="338">C. yanagisawai</emphasis>
</taxonomicName>
is much broader (
<figureCitation id="4EAA5DF0FF8EDB1AA41D3298FAF1FB1D" box="[1327,1406,1159,1184]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="4" pageNumber="338">Fig. 2a</figureCitation>
&amp;
<figureCitation id="4EAA5DF0FF8EDB1AA1A532B3FF39FB78" box="[151,182,1196,1221]" captionStart="FIGURE 3" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[151,1421,459,1921]" captionTargetId="figure@3.[151,1436,459,1936]" captionTargetPageId="3" captionText="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="4" pageNumber="338">3b</figureCitation>
).
<taxonomicName id="11913AF6FF8EDB1AA1E232B3FD1FFB79" authority="(Martin, 1909)" authorityName="Martin" authorityYear="1909" baseAuthorityName="Martin" baseAuthorityYear="1909" box="[208,656,1196,1221]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="masoni">
<emphasis id="E4E59D67FF8EDB1AA1E232B3FE55FB79" box="[208,474,1196,1221]" italics="true" pageId="4" pageNumber="338">Cephalaeschna masoni</emphasis>
(Martin, 1909)
</taxonomicName>
(NE
<collectingCountry id="AE8601E5FF8EDB1AA3EA32B3FC9DFB78" box="[728,786,1196,1221]" name="India" pageId="4" pageNumber="338">India</collectingCountry>
&amp;
<collectingCountry id="AE8601E5FF8EDB1AA27232B3FC0AFB78" box="[832,901,1196,1221]" name="Nepal" pageId="4" pageNumber="338">Nepal</collectingCountry>
) has a “roundly inflated antefrons” (
<bibRefCitation id="B2003C84FF8EDB1AA40C32B3FF51FB55" pageId="4" pageNumber="338" refId="ref2718" refString="Asahina, S. (1981 a) A revision of the Himalayan dragonflies of the genus Cephalaeschna and its allies (Odonata, Aeshnidae). (Part 1). Bulletin of the National Science Museum, Tokyo, Series A, 7 (1), 27 - 49." type="journal article">Asahina 1981a</bibRefCitation>
), not dorsally angled like
<taxonomicName id="11913AF6FF8EDB1AA32832CFFD43FB55" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[538,716,1232,1256]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8EDB1AA32832CFFD43FB55" box="[538,716,1232,1256]" italics="true" pageId="4" pageNumber="338">C. yanagisawai</emphasis>
</taxonomicName>
.
<taxonomicName id="11913AF6FF8EDB1AA3D232D0FB3CFB55" authority="Needham, 1930" authorityName="Needham" authorityYear="1930" box="[736,1203,1231,1256]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="obversa">
<emphasis id="E4E59D67FF8EDB1AA3D232D0FC7DFB55" box="[736,1010,1231,1256]" italics="true" pageId="4" pageNumber="338">Cephalaeschna obversa</emphasis>
<bibRefCitation id="B2003C84FF8EDB1AA2CF32D0FB3CFB55" box="[1021,1203,1231,1256]" pageId="4" pageNumber="338" refId="ref2968" refString="Needham, J. G. (1930) A manual of the dragonflies of China. Zoologia Sinica, 11, 1 - 345, pls. I - XX, index 1 - 11." type="journal article">Needham, 1930</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="14558F97FF8EDB1AA5F532D0FA9FFB55" box="[1223,1296,1231,1256]" country="China" name="Hubei" pageId="4" pageNumber="338">Hubei</collectingRegion>
,
<collectingRegion id="14558F97FF8EDB1AA41232D0FAF5FB55" box="[1312,1402,1231,1256]" country="China" name="Sichuan" pageId="4" pageNumber="338">Sichuan</collectingRegion>
&amp;
<collectingRegion id="14558F97FF8EDB1AA1A532EBFF76FAB0" box="[151,249,1268,1293]" country="China" name="Guizhou" pageId="4" pageNumber="338">Guizhou</collectingRegion>
) and
<taxonomicName id="11913AF6FF8EDB1AA00A32EAFCB3FAB0" authority="Zhang, Cai &amp; Liao, 2013" authorityName="Zhang, Cai &amp; Liao" authorityYear="2013" box="[312,828,1268,1293]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="solitaria">
<emphasis id="E4E59D67FF8EDB1AA00A32EAFE36FAB0" box="[312,441,1268,1293]" italics="true" pageId="4" pageNumber="338">C. solitaria</emphasis>
<bibRefCitation id="B2003C84FF8EDB1AA0F232EBFD52FAB1" box="[448,733,1268,1293]" pageId="4" pageNumber="338" refId="ref3119" refString="Zhang, H. - M., Cai, Q. - H. &amp; Liao, M. - Y. (2013) Three new Cephalaeschna species from central China with descriptions of the hitherto unknown sex of related species (Odonata: Aeshnidae). International Journal of Odonatology, 16 (2), 157 - 176. https: // doi. org / 10.1080 / 13887890.2013.782530" type="journal article">Zhang, Cai &amp; Liao, 2013</bibRefCitation>
(
<collectingRegion id="14558F97FF8EDB1AA3DE32EBFCBBFAB0" box="[748,820,1268,1293]" country="China" name="Hubei" pageId="4" pageNumber="338">Hubei</collectingRegion>
)
</taxonomicName>
have PD markings on S2S8 (Zhang
<emphasis id="E4E59D67FF8EDB1AA5DA32EAFA95FAB0" box="[1256,1306,1268,1293]" italics="true" pageId="4" pageNumber="338">et al</emphasis>
. 2013), but
<taxonomicName id="11913AF6FF8EDB1AA1A53307FECBFA8D" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[151,324,1304,1328]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8EDB1AA1A53307FECBFA8D" box="[151,324,1304,1328]" italics="true" pageId="4" pageNumber="338">C. yanagisawai</emphasis>
</taxonomicName>
has those only on S2S6 (
<figureCitation id="4EAA5DF0FF8EDB1AA35C3308FD4EFA8D" box="[622,705,1303,1328]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="4" pageNumber="338">Fig. 2c</figureCitation>
).
<taxonomicName id="11913AF6FF8EDB1AA3E73308FC7EFA8D" authorityName="Needham" authorityYear="1930" box="[725,1009,1303,1328]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="patrorum">
<emphasis id="E4E59D67FF8EDB1AA3E73308FC7EFA8D" box="[725,1009,1303,1328]" italics="true" pageId="4" pageNumber="338">Cephalaeschna patrorum</emphasis>
</taxonomicName>
has the cercus, the inner side of distal half of which is almost linear and PD markings on S2 to S8 (
<bibRefCitation id="B2003C84FF8EDB1AA27E3323FC72FAE9" box="[844,1021,1340,1365]" pageId="4" pageNumber="338" refId="ref2968" refString="Needham, J. G. (1930) A manual of the dragonflies of China. Zoologia Sinica, 11, 1 - 345, pls. I - XX, index 1 - 11." type="journal article">Needham 1930</bibRefCitation>
;
<bibRefCitation id="B2003C84FF8EDB1AA53B3323FB39FAE8" box="[1033,1206,1340,1365]" pageId="4" pageNumber="338" refId="ref2771" refString="Asahina, S. (1981 b) A revision of the Chinese dragonflies of the genus Cephalaeschna and its allies. Tombo, 24, 2 - 12." type="journal article">Asahina 1981b</bibRefCitation>
; Zhang
<emphasis id="E4E59D67FF8EDB1AA4213322FACAFAE8" box="[1299,1349,1340,1365]" italics="true" pageId="4" pageNumber="338">et al</emphasis>
. 2013), whereas that of
<taxonomicName id="11913AF6FF8EDB1AA07B337FFE77FAC5" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[329,504,1376,1400]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8EDB1AA07B337FFE77FAC5" box="[329,504,1376,1400]" italics="true" pageId="4" pageNumber="338">C. yanagisawai</emphasis>
</taxonomicName>
is gently concaved inwards (
<figureCitation id="4EAA5DF0FF8EDB1AA20D3340FC1DFAC5" box="[831,914,1375,1400]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="4" pageNumber="338">Fig. 2d</figureCitation>
), and lacking PD markings on S7 and S8 (
<figureCitation id="4EAA5DF0FF8EDB1AA45D3340FF3AFA21" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[186,1425,537,1800]" captionTargetId="figure@2.[151,1437,483,1911]" captionTargetPageId="2" captionText="FIGURE 2. Male of C. yanagisawai (a, cg: holotype, b: paratype). (a) head in frontal view; (b) dorsal view of synthorax; (c) dorsal view of abdomen; (d) anal appendages in dorsal view (dotted line indicates hidden part of epiproct); (e) the same in lateral view; (f) penile organ in dorsal view; (g) the same in lateral view." httpUri="https://zenodo.org/record/1439754/files/figure.png" pageId="4" pageNumber="338">Fig. 2c</figureCitation>
).
</paragraph>
<paragraph id="D62E4175FF8EDB1AA1F533B8FB65F991" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="338">
The female of
<taxonomicName id="11913AF6FF8EDB1AA04233B7FDAFFA7D" authorityName="Sasamoto &amp; Lien" authorityYear="2018" box="[368,544,1448,1472]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="yanagisawai">
<emphasis id="E4E59D67FF8EDB1AA04233B7FDAFFA7D" box="[368,544,1448,1472]" italics="true" pageId="4" pageNumber="338">C. yanagisawai</emphasis>
</taxonomicName>
is peculiar in that postero-ventral part of S10 tergite projects posteriorly (
<figureCitation id="4EAA5DF0FF8EDB1AA45C33B8FF3BFA59" captionStart="FIGURE 3" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[151,1421,459,1921]" captionTargetId="figure@3.[151,1436,459,1936]" captionTargetPageId="3" captionText="FIGURE 3. Paratype female of C. yanagisawai. (a) lateral habitus; (b) head in frontal view; (c) distal abdomen in lateral view; (d) ventral view of tip of S10 tergite." httpUri="https://zenodo.org/record/1439756/files/figure.png" pageId="4" pageNumber="338">Fig. 3c</figureCitation>
). Although there are not many species in the genus for which female morphology is described, the females with such structure are exceptional, only known in
<taxonomicName id="11913AF6FF8EDB1AA3FE33EFFB24F9B5" authority="(Martin, 1909) (Himalaya)" authorityName="Martin" authorityYear="1909" baseAuthorityName="Martin" baseAuthorityYear="1909" box="[716,1195,1519,1544]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="acutifrons">
<emphasis id="E4E59D67FF8EDB1AA3FE33EFFCEBF9B5" box="[716,868,1520,1544]" italics="true" pageId="4" pageNumber="338">C. acutifrons</emphasis>
(Martin, 1909) (Himalaya)
</taxonomicName>
(
<bibRefCitation id="B2003C84FF8EDB1AA5F233F0FAE8F9B5" box="[1216,1383,1519,1544]" pageId="4" pageNumber="338" refId="ref2801" refString="Asahina, S. (1983) Further contributions to the knowledge of Nepalese Cephalaeschna and its allies (Odonata, Aeshnidae). Bulletin of the National Science Museum, Tokyo, Series A, 8 (2), 51 - 67." type="journal article">Asahina 1983</bibRefCitation>
),
<taxonomicName id="11913AF6FF8EDB1AA4B033EFFDD7F991" authority="Zhang &amp; Cai, 2013" authorityName="Zhang &amp; Cai" authorityYear="2013" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="cornifrons">
<emphasis id="E4E59D67FF8EDB1AA4B033EFFE84F991" italics="true" pageId="4" pageNumber="338">C. cornifrons</emphasis>
Zhang &amp; Cai, 2013 (
<collectingRegion id="14558F97FF8EDB1AA0CA300BFDDEF991" box="[504,593,1556,1580]" country="China" name="Yunnan" pageId="4" pageNumber="338">Yunnan</collectingRegion>
)
</taxonomicName>
(Zhang &amp; Cai 2013) and
<taxonomicName id="11913AF6FF8EDB1AA248300AFB69F991" authority="(Zhang et al. 2013)" authorityName="Zhang et al." authorityYear="2013" box="[890,1254,1556,1581]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="patrorum">
<emphasis id="E4E59D67FF8EDB1AA248300AFB8BF991" box="[890,1028,1557,1580]" italics="true" pageId="4" pageNumber="338">C. patrorum</emphasis>
(Zhang
<emphasis id="E4E59D67FF8EDB1AA550300AFB1CF990" box="[1122,1171,1556,1581]" italics="true" pageId="4" pageNumber="338">et al</emphasis>
. 2013)
</taxonomicName>
.
</paragraph>
<paragraph id="D62E4175FF8EDB1AA1F53027FBE3F925" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="338">
<emphasis id="E4E59D67FF8EDB1AA1F53027FE1EF9ED" bold="true" box="[199,401,1591,1616]" pageId="4" pageNumber="338">Notes on habitat.</emphasis>
According to Mr Yanagisawa (
<emphasis id="E4E59D67FF8EDB1AA3DC3026FCE0F9ED" box="[750,879,1593,1616]" italics="true" pageId="4" pageNumber="338">pers. comm</emphasis>
.), he collected the
<typeStatus id="092AFFD7FF8EDB1AA5703028FB2BF9ED" box="[1090,1188,1591,1616]" pageId="4" pageNumber="338" type="holotype">holotype</typeStatus>
male at around
<specimenCount id="C0978AFCFF8EDB1AA4643027FAF1F9ED" box="[1366,1406,1592,1616]" pageId="4" pageNumber="338" type="generic">4 p.</specimenCount>
m. following rainy conditions from morning. The specimen was flying over a wet grassy place near the stream (
<figureCitation id="4EAA5DF0FF8EDB1AA45D3043FF27F925" captionStart="FIGURE 4" captionStartId="5.[151,250,1713,1735]" captionTargetBox="[313,1273,411,1691]" captionTargetId="figure@5.[313,1273,411,1691]" captionTargetPageId="5" captionText="FIGURE 4. The habitat of C. yanagisawai, Hoang Lien National Park, Lai Chau Province, N. Vietnam (photo by T. Yanagisawa)." httpUri="https://zenodo.org/record/1439758/files/figure.png" pageId="4" pageNumber="338">Fig. 4</figureCitation>
). In the same place,
<taxonomicName id="11913AF6FF8EDB1AA0BF3060FC91F925" authority="Karube, 2003" authorityName="Karube" authorityYear="2003" box="[397,798,1663,1688]" class="Insecta" family="Aeshnidae" genus="Cephalaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="338" phylum="Arthropoda" rank="species" species="aritai">
<emphasis id="E4E59D67FF8EDB1AA0BF3060FDF0F925" box="[397,639,1663,1688]" italics="true" pageId="4" pageNumber="338">Cephalaeschna aritai</emphasis>
Karube, 2003
</taxonomicName>
was observed the day before.
</paragraph>
</subSubSection>
</treatment>
</document>