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<mods:title id="72C5D196EF5049BAB11877868F92C4E1">New material of Paleopsychoda jarzembowskii Azar &amp; Maksoud, 2022 from Bqaatouta amber outcrop, showing the importance of insect fossils in biostratigraphy</mods:title>
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<emphasis id="E4D31E41FFB1FF87FF03FBB023C5C901" bold="true" box="[136,481,1106,1133]" italics="true" pageId="1" pageNumber="9">Paleopsychoda jarzembowskii</emphasis>
<bibRefCitation id="B236BFA2FFB1FF87FE6DFBB020D8C900" author="Maksoud, S. &amp; Granier, B. &amp; Azar, D." box="[486,764,1106,1133]" pageId="1" pageNumber="9" pagination="699 - 743" refId="ref3610" refString="Maksoud, S., Granier, B. &amp; Azar, D. (2022) Palaeoentomological (fossil insects) outcrops in Lebanon. Carnets de Geologie, 22 (16), 699 - 743. https: // doi. org / 10.2110 / carnets. 2022.2216" type="journal article" year="2022">Azar &amp; Maksoud, 2022</bibRefCitation>
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<emphasis id="E4D31E41FFB1FF87FF03FB79230BC9DF" bold="true" box="[136,303,1179,1203]" pageId="1" pageNumber="9">Type material.</emphasis>
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1615A, from Mdeyrij-Hammana, Central Lebanon, Azar collection, deposited in the Natural History Museum of the Lebanese University,
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Faculty of
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II, Fanar,
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.
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<emphasis id="E4D31E41FFB1FF87FF36FAC82379C82D" bold="true" box="[189,349,1321,1346]" pageId="1" pageNumber="9">New material.</emphasis>
Specimen BKT-9A, Maalouf collection, from Bqaatouta, deposited in the Natural History Museum of the Lebanese University, Faculty of Sciences II, Fanar,
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.
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<emphasis id="E4D31E41FFB1FF87FF36FA55239AC8A3" bold="true" box="[189,446,1463,1488]" pageId="1" pageNumber="9">Locality and horizon.</emphasis>
Bqaatouta, Caza (= District) Kesserouan (Central
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), lower Barremian (
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Granier
<emphasis id="E4D31E41FFB1FF87FD7DFA3E22B9CB7A" italics="true" pageId="1" pageNumber="9">et al</emphasis>
., 2016
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;
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Maksoud
<emphasis id="E4D31E41FFB1FF87FED5FA1D23A9CB7A" box="[350,397,1534,1558]" italics="true" pageId="1" pageNumber="9">et al</emphasis>
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, 2022).
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<emphasis id="E4D31E41FFB1FF87FF36F9C0236CCB56" bold="true" box="[189,328,1570,1594]" pageId="1" pageNumber="9">Description.</emphasis>
The specimen of Bqaatouta is a complete sample but somehow with incomplete antennae and it carries all the diagnostic features of
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,
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.
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., eye-bridge incomplete, mouthparts toothed, welldeveloped and of phlebotomine-type, with sharp toothed mandibles and laciniae; maxillary palp four-segmented, with last palpomere twice as long as others and presenting a constriction in its basal third; anterior sub-basal part of costal vein inflated, from either side of end of humeral vein, allowing as such larger surface, with insertions of a basal wing tuft of setae (specific male character); R1 reaching costa beyond level of radial fork; M3 and M4 [CuA1] meeting basally for a very small distance; CuA [CuA2] well-developed, reaching wing margin beyond half of wing; CuP [A] well developed; male genitalia with stylus bearing two apical spines; a specialized long seta strongly curved in its middle part arising between gonocoxite and gonostylus; parameres paired with apically series of four strong setae with incurved apices; surstyli joined basally, bearing apically two strong setae on each. Measurements of the different body parts are very similar to the type specimen, the
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number 1615A.
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<mods:url>http://dx.doi.org/10.3897/dez.69.84080</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.69.84080</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2-151</mods:identifier>
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<subSubSection pageId="0" pageNumber="151" type="nomenclature">
<paragraph pageId="0" pageNumber="151">
<taxonomicName LSID="5E0E8E52-4CA8-5ADB-9481-3CBB5607D40B" authority="Serville, 1823" authorityName="Serville" authorityYear="1823" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="ferrugatus">Dolerus ferrugatus Serville, 1823</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="151" type="reference_group">
<paragraph pageId="0" pageNumber="151">
<taxonomicName authority="Panzer, 1801" authorityName="Serville" authorityYear="1823" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="ferrugatus">Dolerus ferrugatus</taxonomicName>
Serville, 1823: 59. ♀. Syntypes. Type locality: Paris. Lectotype designated below.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<taxonomicName authority="Panzer, 1801" authorityName="Panzer" authorityYear="1801" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus brevicornis" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="brevicornis">Dolerus brevicornis</taxonomicName>
Thomson, 1871: 288. ♀, ♂. Syntypes. Published type locality:
<normalizedToken originalValue="“södra">&quot;soedra</normalizedToken>
Sverige&quot; [southern Sweden]. Lectotype designated below. Primary homonym of
<taxonomicName family="Tenthredinidae" genus="D." higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="D. brevicornis" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="brevicornis">D. brevicornis</taxonomicName>
Zaddach, 1859.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<taxonomicName authority="Panzer, 1801" authorityName="Konow" authorityYear="1887" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus thomsoni subsp. var. var. miricolor" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="variety" species="thomsoni" subSpecies="var." variety="miricolor">Dolerus thomsoni var. miricolor</taxonomicName>
Konow, 1887a: 283. ♂. Syntypes (assumed). Type locality:
<normalizedToken originalValue="Fürstenberg">Fuerstenberg</normalizedToken>
/ Mecklenburg. Lectotype designated below.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="151" type="type material">
<paragraph pageId="0" pageNumber="151">Type material examined and taxonomic notes.</paragraph>
<paragraph pageId="0" pageNumber="151">
<taxonomicName authority="Panzer, 1801" authorityName="Serville" authorityYear="1823" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus ferrugatus</emphasis>
</taxonomicName>
Serville, 1823. Lectotype ♀ hereby designated, labelled:
<normalizedToken originalValue="“Saint-Fargeau”">&quot;Saint-Fargeau&quot;</normalizedToken>
[round],
<normalizedToken originalValue="“Lectotype”">&quot;Lectotype&quot;</normalizedToken>
[red, printed in block letters], &quot;Lectotype
<taxonomicName family="Tenthredinidae" genus="D." kingdom="Animalia" lsidName="D. ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">D. ferrugatus</emphasis>
</taxonomicName>
LEP. ♀ det. A.Haris 1996&quot;, &quot;
<taxonomicName family="Tenthredinidae" genus="D." kingdom="Animalia" lsidName="D. ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">D. ferrugatus</emphasis>
</taxonomicName>
LEP. ♀ det.: A.Haris&quot;, &quot;
<taxonomicName genus="Lectotypus" lsidName="Lectotypus" pageId="0" pageNumber="151" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="151">Lectotypus</emphasis>
</taxonomicName>
[printed]♀
<taxonomicName authority="Panzer, 1801" authorityName="Serville" authorityYear="1823" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus ferrugatus</emphasis>
</taxonomicName>
Serville, 1823 [handwritten in block letters] M.Heidemaa design. [printed]&quot; [red], &quot;
<taxonomicName authority="Panzer, 1801" authorityName="Serville" authorityYear="1823" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus ferrugatus</emphasis>
</taxonomicName>
Serville, 1823 M.Heidemaa det.&quot; [printed]. In good condition (the right flagellum missing). MNHN.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<taxonomicName authority="Panzer, 1801" authorityName="Panzer" authorityYear="1801" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus brevicornis" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus brevicornis</emphasis>
</taxonomicName>
Thomson, 1871. Lectotype ♀ hereby designated (type locality: Sweden,
<normalizedToken originalValue="Öland">Oeland</normalizedToken>
), labelled:
<normalizedToken originalValue="“Ö.”">&quot;Oe.&quot;</normalizedToken>
[
<normalizedToken originalValue="Öland">Oeland</normalizedToken>
], &quot; MZLU 2013 414&quot; [green, printed (a loan record label)], &quot;
<taxonomicName genus="Lectotypus" lsidName="Lectotypus" pageId="0" pageNumber="151" rank="genus">Lectotypus</taxonomicName>
[printed] ♀ 2014
<taxonomicName authority="Panzer, 1801" authorityName="Panzer" authorityYear="1801" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus brevicornis" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus brevicornis</emphasis>
</taxonomicName>
Thomson, 1871 [handwritten in block letters] M.Heidemaa des. [printed]&quot; [red], &quot;
<taxonomicName authority="Panzer, 1801" authorityName="Serville" authorityYear="1823" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus ferrugatus</emphasis>
</taxonomicName>
Serville, 1823 M.Heidemaa det.&quot; [printed]. 2♀ (from
<normalizedToken originalValue="Öland">Oeland</normalizedToken>
and Scania), 1♂ (the melanic form, from Lund) labelled as paralectotypes. All in MZLU.
</paragraph>
<paragraph pageId="0" pageNumber="151">
<taxonomicName authority="Panzer, 1801" authorityName="Konow" authorityYear="1887" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus thomsoni subsp. var. var. miricolor" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="variety" species="thomsoni" subSpecies="var." variety="miricolor">Dolerus thomsoni var. miricolor</taxonomicName>
Konow, 1887. Lectotype ♂ hereby designated, labelled: &quot;f. [
<normalizedToken originalValue="Fürstenberg">Fuerstenberg</normalizedToken>
] 12/4. [18]84.&quot;, &quot;
<taxonomicName authority="Panzer, 1801" authorityName="Panzer" authorityYear="1801" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus</emphasis>
</taxonomicName>
v. [var.]
<taxonomicName authority="Panzer, 1801" authorityName="Panzer" authorityYear="1801" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus miricolor" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="miricolor">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus miricolor</emphasis>
</taxonomicName>
Knw. [Konow] fbg. 12/4. 84. Typus&quot; [with double black frame], &quot;
<taxonomicName authority="Panzer, 1801" authorityName="Panzer" authorityYear="1801" class="Insecta" family="Tenthredinidae" form="miricolor" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus ferrugatus subsp. f." order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="form" species="ferrugatus" subSpecies="f.">Dolerus ferrugatus Lep. f. miricolor</taxonomicName>
Knw. Typus. ♂ det. OConde [Otto Conde] 1943&quot;,
<normalizedToken originalValue="“Typus”">&quot;Typus&quot;</normalizedToken>
[red, printed],
<normalizedToken originalValue="“Holotypus”">&quot;Holotypus&quot;</normalizedToken>
[red, printed], &quot;Coll. Konow&quot; [printed], &quot;GBIF-GISHym 4215&quot; [printed], &quot;
<taxonomicName genus="Lectotypus" lsidName="Lectotypus" pageId="0" pageNumber="151" rank="genus">Lectotypus</taxonomicName>
[printed] ♂
<taxonomicName authority="Panzer, 1801" authorityName="Konow" authorityYear="1887" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus thomsoni subsp. var. var. miricolor" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="variety" species="thomsoni" subSpecies="var." variety="miricolor">Dolerus thomsoni var. miricolor</taxonomicName>
[handwritten in block letters] M.Heidemaa des. [printed]&quot; [red], &quot;
<taxonomicName authority="Panzer, 1801" authorityName="Serville" authorityYear="1823" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolerus ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">Dolerus ferrugatus</emphasis>
</taxonomicName>
Serville, 1823 M.Heidemaa det.&quot; [printed]. Right flagellum missing, penis valves dissected (a microscope slide:
<taxonomicName authority="Panzer, 1801" authorityName="Panzer" authorityYear="1801" class="Insecta" family="Tenthredinidae" genus="Dolerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="subOrder" subOrder="Symphyta">Symphyta</taxonomicName>
coll. Nr: 159). All in SDEI.
</paragraph>
<paragraph pageId="0" pageNumber="151">
The melanic colour form of
<taxonomicName family="Tenthredinidae" genus="D." kingdom="Animalia" lsidName="D. ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">D. ferrugatus</emphasis>
</taxonomicName>
, described by Konow as
<taxonomicName family="Tenthredinidae" genus="D." kingdom="Animalia" lsidName="D. thomsoni subsp. var. var. miricolor" order="Hymenoptera" pageId="0" pageNumber="151" rank="variety" species="thomsoni" subSpecies="var." variety="miricolor">D. thomsoni var. miricolor</taxonomicName>
, has sometimes been mixed up with
<taxonomicName family="Tenthredinidae" genus="D." kingdom="Animalia" lsidName="D. pachycerus" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="pachycerus">
<emphasis italics="true" pageId="0" pageNumber="151">D. pachycerus</emphasis>
</taxonomicName>
Hartig, 1837. Apart from their different penis valves, they can be separated by the shape and the structure of the mesoscutellar appendage: long, concave at the sides, and with a distinct longitudinal keel in
<taxonomicName family="Tenthredinidae" genus="D." kingdom="Animalia" lsidName="D. pachycerus" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="pachycerus">
<emphasis italics="true" pageId="0" pageNumber="151">D. pachycerus</emphasis>
</taxonomicName>
compared with
<taxonomicName family="Tenthredinidae" genus="D." kingdom="Animalia" lsidName="D. ferrugatus" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="ferrugatus">
<emphasis italics="true" pageId="0" pageNumber="151">D. ferrugatus</emphasis>
</taxonomicName>
.
</paragraph>
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</treatment>
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