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<document id="7E6B28CE9AB7453A2AA64280A03EFED5" ID-DOI="10.5281/zenodo.11512244" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512244" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1717719192119" checkinUser="felipe" docAuthor="Zug, George R." docDate="2015" docId="03856357CC41FFE6FFD059C7FEE26423" docLanguage="en" docName="ProcCalifAcadSci.62.5.135-190.pdf" docOrigin="Proceedings of the California Academy of Sciences 62 (5)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Kalophrynus cryptophonus Vassilieva, Galoyan, Gogoleva, and Poyarkov 2014" docType="treatment" docVersion="2" lastPageNumber="158" masterDocId="FFBC1B2FCC57FFF1FFAB5C0AFF8C611C" masterDocTitle="Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs" masterLastPageNumber="190" masterPageNumber="135" pageNumber="157" updateTime="1721167902083" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D4C7363C77E99EB68FAF7B81958115B5">Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs</mods:title>
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<mods:namePart id="D09F6966413ED416E9146FCB0FAB43BE">Zug, George R.</mods:namePart>
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<mods:date id="C0E0AC96853BF61CA80A1805A63BA0EC">2015</mods:date>
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<taxonomicName id="4C2CA9C2CC41FFE7FFD059C7FC7F64F9" ID-CoL="6NJSY" authority="Vassilieva, Galoyan, Gogoleva, and Poyarkov, 2014" authorityName="Vassilieva, Galoyan, Gogoleva, and Poyarkov" authorityYear="2014" box="[123,1011,1485,1509]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="157" phylum="Chordata" rank="species" species="cryptophonus">
<emphasis id="B9580E53CC41FFE7FFD059C7FE2A64F9" bold="true" box="[123,422,1485,1509]" italics="true" pageId="22" pageNumber="157">Kalophrynus cryptophonus</emphasis>
<bibRefCitation id="EFBDAFB0CC41FFE7FE0659C7FC7F64F9" author="VASSILIEVA, A. B. &amp; E. A. GALOYAN &amp; S. S. GOGOLEVA &amp; N. A. POYARKOV, JR." box="[429,1011,1485,1509]" pageId="22" pageNumber="157" pagination="401 - 434" refId="ref38707" refString="VASSILIEVA, A. B., E. A. GALOYAN, S. S. GOGOLEVA, AND N. A. POYARKOV, JR. 2014. Two new species of Kalophrynus Tschudi, 1838 (Anura: Microhylidae) from the Annamite mountains in southern Vietnam. Zootaxa 3796 (3): 401 - 434." type="journal article" year="2014">Vassilieva, Galoyan, Gogoleva, and Poyarkov, 2014</bibRefCitation>
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<paragraph id="8B93D241CC41FFE7FC8F59E7FB026719" blockId="22.[123,1166,1485,1541]" box="[804,1166,1517,1541]" pageId="22" pageNumber="157">
<collectingRegion id="49E81CA3CC41FFE7FC8F59E7FC126719" box="[804,926,1517,1541]" country="Vietnam" name="Lam Dong" pageId="22" pageNumber="157">Lam Dong</collectingRegion>
Bamboo Sticky Frog
</paragraph>
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</emphasis>
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<taxonomicName id="4C2CA9C2CC41FFE7FFD05A11FC0E672C" authority="Vassilieva, Galoyan, Gogoleva, and Poyarkov, 2014" authorityName="Vassilieva, Galoyan, Gogoleva, and Poyarkov" authorityYear="2014" box="[123,898,1563,1585]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="157" phylum="Chordata" rank="species" species="cryptophonus">
<emphasis id="B9580E53CC41FFE7FFD05A11FE0D672C" box="[123,385,1563,1584]" italics="true" pageId="22" pageNumber="157">Kalophrynus cryptophonus</emphasis>
<materialsCitation id="3B44D81CCC41FFE7FE235A11FF776773" country="Vietnam" elevation="800" latitude="11.738056" location="Zootaxa" longLatPrecision="21" longitude="107.70694" pageId="22" pageNumber="157" specimenCount="1" stateProvince="Lam Dong">
<bibRefCitation id="EFBDAFB0CC41FFE7FE235A11FC0E672C" author="VASSILIEVA, A. B. &amp; E. A. GALOYAN &amp; S. S. GOGOLEVA &amp; N. A. POYARKOV, JR." box="[392,898,1563,1585]" pageId="22" pageNumber="157" pagination="401 - 434" refId="ref38707" refString="VASSILIEVA, A. B., E. A. GALOYAN, S. S. GOGOLEVA, AND N. A. POYARKOV, JR. 2014. Two new species of Kalophrynus Tschudi, 1838 (Anura: Microhylidae) from the Annamite mountains in southern Vietnam. Zootaxa 3796 (3): 401 - 434." type="journal article" year="2014">Vassilieva, Galoyan, Gogoleva, and Poyarkov, 2014</bibRefCitation>
,
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3769(3):410 [
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locality: “Loc Bao,
<collectingRegion id="49E81CA3CC41FFE7FEF55A36FDAF674D" box="[350,547,1595,1617]" country="Vietnam" name="Lam Dong" pageId="22" pageNumber="157">Lam Dong Province</collectingRegion>
,
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(coordinates
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,
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elevation
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a.s.l.
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)”]
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.
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.—
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: Zoological Museum of the Lomonosov Moscow University
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(
<collectionCode id="ED3D4A84CC41FFE7FF2F5AACFF5B67A1" box="[132,215,1702,1725]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="22" pageNumber="157" type="Museum">ZMMU</collectionCode>
)
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A-4944.
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:
<collectionCode id="ED3D4A84CC41FFE7FE6D5AACFD9567A1" box="[454,537,1702,1725]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="22" pageNumber="157" type="Museum">ZMMU</collectionCode>
A-4858–59. Original designations
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.
</paragraph>
<paragraph id="8B93D241CC40FFE6FF005CD1FCC960A9" blockId="23.[123,1174,219,1344]" pageId="23" pageNumber="158">
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<emphasis id="B9580E53CC40FFE6FF005CD1FEA561ED" bold="true" box="[171,297,219,243]" pageId="23" pageNumber="158">DEFINITION</emphasis>
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.— Small, adult female
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<collectionCode id="ED3D4A84CC40FFE6FD265CD1FD3361EF" box="[653,703,219,243]" pageId="23" pageNumber="158">SVL</collectionCode>
(
<emphasis id="B9580E53CC40FFE6FD655CD6FD7861EF" box="[718,756,220,243]" italics="true" pageId="23" pageNumber="158">n =</emphasis>
1), adult males 27.9– 30.
<quantity id="4CD47FA4CC40FFE6FBA85CD1FBCC61EE" box="[1027,1088,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="158" unit="mm" value="4.0">4 mm</quantity>
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(
<emphasis id="B9580E53CC40FFE6FB235CD6FF01600F" italics="true" pageId="23" pageNumber="158">n =</emphasis>
5); head moderately long 28–32 % HeadL/
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; head wider than long 105–122 % HeadW/HeadL; naris closer to snout than to eye 59–68 % NarEye/SnEye; eye moderately large 35–42 % EyeD/HeadL; tympanum visible and smaller than eye 55–77 % Tymp/EyeD; slender moderately long forelimb NA % Forarm/
<collectionCode id="ED3D4A84CC40FFE6FD955D57FDFC6069" box="[574,624,349,373]" pageId="23" pageNumber="158">SVL</collectionCode>
and forearm to crus length NA % Forarm/CrusL; hindlimb slender and moderate length 129–140 % HndlL/
<collectionCode id="ED3D4A84CC40FFE6FD445D77FCAD6089" box="[751,801,381,405]" pageId="23" pageNumber="158">SVL</collectionCode>
, 40–43 % CrusL/
<collectionCode id="ED3D4A84CC40FFE6FC495D77FB986089" box="[994,1044,381,405]" pageId="23" pageNumber="158">SVL</collectionCode>
, and 89–98 % CrusL/ThghL; hindfoot well developed 77–96 % HndfL/CrusL.
</paragraph>
<paragraph id="8B93D241CC40FFE6FF005DB4FD37630A" blockId="23.[123,1174,219,1344]" pageId="23" pageNumber="158">Vomerine teeth absence; three palatal folds, vomerine one continuous, postorbital short and low, buccal continuous and crenulated; tongue rounded free end. Skin on margin of mandible with longitudinal series of short triangular spines in males.</paragraph>
<paragraph id="8B93D241CC40FFE6FF005E15FC3F6205" blockId="23.[123,1174,219,1344]" pageId="23" pageNumber="158">Fingers rudimentary webbing; lengths 3&gt;2&gt;4&gt;1; tips rounded and not dilated; subarticular subarticular tubercles distinct, round, and one on digits 1, 2, 4, and two on 3; one large oval palmar tubercle on outer half of palm, bordered distally by four small, round tubercles; smooth nuptial excrescences on base of fingers 2 and 3; fine-spined asperities dorsally covering fingers 2, 3, and 4 from base to end of penultimate phalanx. Toes modestly webbed not extending beyond proximal subarticular tubercles of toe 4, lengths 4&gt;3&gt;5&gt;2&gt;1; tips rounded, not dilated; subarticular tubercles moderate, oval and one on digits 1, 2, two on 3, three on 4, two on 5 (only distal one prominent); moderate oval inner and small, round outer metatarsal tubercles.</paragraph>
<paragraph id="8B93D241CC40FFE6FF005F28FC3962C7" blockId="23.[123,1174,219,1344]" pageId="23" pageNumber="158">In life, dorsum ranging from dark brown at night to diurnal pinkish beige with faint darker reticulation from between eyes to end of trunk and reverse Y-mark with base on crown and nape and arms extending to rear of trunk; body; narrow, orangish dorsolateral stripe from tip of snout and edge of canthus rostralis above eye running diagonally to mid inguina, bordered below by dark brown that fades ventrolaterally; small round, black inguinal spot, not light edged; venter pale yellowish-pink, chin to mid chest with dark marking, belly largely immaculate.</paragraph>
</subSubSection>
<subSubSection id="C33681CACC40FFE6FF005FEEFDCD6520" pageId="23" pageNumber="158" type="etymology">
<paragraph id="8B93D241CC40FFE6FF005FEEFDCD6520" blockId="23.[123,1174,219,1344]" pageId="23" pageNumber="158">
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<emphasis id="B9580E53CC40FFE6FF005FEEFEB962E6" bold="true" box="[171,309,996,1020]" pageId="23" pageNumber="158">ETYMOLOGY</emphasis>
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.— Because this species was only discovered by the males’ vocalization from hollow bamboo stems, the authors highlight this behavior with Greek
<emphasis id="B9580E53CC40FFE6FCD8580FFC4E6500" box="[883,962,1029,1052]" italics="true" pageId="23" pageNumber="158">cryptos</emphasis>
for hidden or mystery and
<emphasis id="B9580E53CC40FFE6FF72582FFEAA6521" box="[217,294,1061,1085]" italics="true" pageId="23" pageNumber="158">phonus</emphasis>
, voice, thus hidden voice.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC40FFE6FF00584FFDD865E2" pageId="23" pageNumber="158" type="distribution">
<paragraph id="8B93D241CC40FFE6FF00584FFE896561" blockId="23.[123,1174,219,1344]" pageId="23" pageNumber="158">
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<emphasis id="B9580E53CC40FFE6FF00584FFECF6547" bold="true" box="[171,323,1093,1117]" pageId="23" pageNumber="158">DISTRIBUTION</emphasis>
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.—
<materialsCitation id="3B44D81CCC40FFE6FEC0584FFF726561" country="Vietnam" location="Vietnam" pageId="23" pageNumber="158" specimenCount="1">
<collectingCountry id="F33B92D1CC40FFE6FEC0584FFE496541" box="[363,453,1093,1117]" name="Vietnam" pageId="23" pageNumber="158">Vietnam</collectingCountry>
.
<taxonomicName id="4C2CA9C2CC40FFE6FE78584FFD796541" authorityName="Vassilieva, Galoyan, Gogoleva, and Poyarkov" authorityYear="2014" box="[467,757,1093,1117]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="23" pageNumber="158" phylum="Chordata" rank="species" species="cryptophonus">
<emphasis id="B9580E53CC40FFE6FE78584FFD796541" box="[467,757,1093,1117]" italics="true" pageId="23" pageNumber="158">Kalophrynus cryptophonus</emphasis>
</taxonomicName>
is known only from the vicinity of the
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locality
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.
</paragraph>
<paragraph id="8B93D241CC40FFE6FF00588CFDD865E2" blockId="23.[123,1174,219,1344]" pageId="23" pageNumber="158">
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<emphasis id="B9580E53CC40FFE6FF00588CFE9D6580" bold="true" box="[171,273,1158,1182]" pageId="23" pageNumber="158">NATURAL</emphasis>
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<emphasis id="B9580E53CC40FFE6FEB3588CFEF46580" bold="true" box="[280,376,1158,1182]" pageId="23" pageNumber="158">HISTORY</emphasis>
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.— This sticky frog lives in a mid-montane evergreen tropical forest (secondary) with abundant bamboo clumps. It was discovered in mid April by the males’ vocalization from inside cut bamboo stems. The water inside the bamboo stems serve as egg-deposition and larval development sites (phytotelm breeding).
</paragraph>
</subSubSection>
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<emphasis id="B9580E53CC40FFE6FF00590DFEA46402" bold="true" box="[171,296,1287,1311]" pageId="23" pageNumber="158">COMMENTS</emphasis>
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.— Preceding information extracted from the original description by Vassilieva and colleagues (2014).
</paragraph>
</subSubSection>
</treatment>
</document>

201
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<mods:title id="861C5065F6EF225EDB16EB3690714EB7">Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs</mods:title>
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<emphasis id="B9580E53CC41FFE7FFD05CD1FE0461EF" bold="true" box="[123,392,219,243]" italics="true" pageId="22" pageNumber="157">Kalophrynus calciphilus</emphasis>
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<bibRefCitation id="EFBDAFB0CC41FFE7FE3B5CD1FDA761EF" author="DEHLING, J. M." box="[400,555,219,243]" pageId="22" pageNumber="157" pagination="49 - 60" refId="ref37239" refString="DEHLING, J. M. 2011. A new karst-dwelling species of Kalophrynus (Anura: Microhylidae) from Gunung Mulu National Park, Borneo, Malaysia. Zootaxa 2737: 49 - 60." type="journal article" year="2011">Dehling, 2011</bibRefCitation>
</paragraph>
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<paragraph id="8B93D241CC41FFE7FC265CD1FB0261EF" blockId="22.[123,1166,219,243]" box="[909,1166,219,243]" pageId="22" pageNumber="157">Limestone Sticky Frog</paragraph>
</subSubSection>
</emphasis>
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<emphasis id="B9580E53CC41FFE7FFD05D09FEE46004" box="[123,360,259,280]" italics="true" pageId="22" pageNumber="157">Kalophrynus calciphilus</emphasis>
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,
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2737:51 [
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locality: “close to the northern entrance of
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(
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) on
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, ca.
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a.s.l.
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,
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,
<collectingCountry id="F33B92D1CC41FFE7FF305D4AFF62604A" box="[155,238,320,342]" name="Malaysia" pageId="22" pageNumber="157">Sarawak</collectingCountry>
,
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”].
</paragraph>
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.—
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:
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<collectingRegion id="49E81CA3CC41FFE7FBCB5D60FB19609D" box="[1120,1173,362,385]" country="Switzerland" name="Bern" pageId="22" pageNumber="157">Bern</collectingRegion>
1056261
</materialsCitation>
.
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<smallCapsWord id="8D75449DCC41FFE7FF4F5D80FEDA60BC" baselines="411,411" box="[228,342,394,417]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Paratypes" pageId="22" pageNumber="157">
<typeStatus id="54976CE3CC41FFE7FF4F5D80FEDA60BC" box="[228,342,394,417]" pageId="22" pageNumber="157" type="paratype">PARATYPES</typeStatus>
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:
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.1611–1615,
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171777,
<collectionCode id="ED3D4A84CC41FFE7FC825D80FCFA60BD" box="[809,886,394,417]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="22" pageNumber="157">NMBE</collectionCode>
1056262–263
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. All by original designation.
</paragraph>
<paragraph id="8B93D241CC41FFE7FF005DC0FCAA63BE" blockId="22.[123,1174,361,1445]" pageId="22" pageNumber="157">
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<emphasis id="B9580E53CC41FFE7FF005DC0FEA560FC" bold="true" box="[171,297,458,482]" pageId="22" pageNumber="157">DEFINITION</emphasis>
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.— Small, adult females 35.5–38.
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(
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2), adult males 29.7–30.
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(
<emphasis id="B9580E53CC41FFE7FF625DE0FF7C631D" box="[201,240,490,513]" italics="true" pageId="22" pageNumber="157">n =</emphasis>
2); head moderately long 31 % HeadL/
<collectionCode id="ED3D4A84CC41FFE7FD485DE0FC9A631E" box="[739,790,490,514]" pageId="22" pageNumber="157">SVL</collectionCode>
; head wider than long 119 % HeadW/HeadL; naris closer to snout than to eye NA % NarEye/SnEye; eye moderately large 38 % EyeD/HeadL; tympanum visible and distinctly smaller than eye 54 % Tymp/EyeD; slender moderately long forelimb NA % Forarm/
<collectionCode id="ED3D4A84CC41FFE7FE465E40FD93637E" box="[493,543,586,610]" pageId="22" pageNumber="157">SVL</collectionCode>
and forearm to crus length NA % Forarm/CrusL; hindlimb slender and long 183 % HndlL/
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, 43 % CrusL/
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, and 96 % CrusL/ThghL; hindfoot well developed 84 % HndfL/CrusL. Proportions from
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only.
</paragraph>
<paragraph id="8B93D241CC41FFE7FF005EA0FE3B621E" blockId="22.[123,1174,361,1445]" pageId="22" pageNumber="157">Vomerine teeth absence; vomerine fold paired and crenulated, condition of other palatal fold morphology not reported; tongue broad and not bifurcated. Adult males with series of spiny tubercles on rear half of mandible.</paragraph>
<paragraph id="8B93D241CC41FFE7FF005F01FF2762FF" blockId="22.[123,1174,361,1445]" pageId="22" pageNumber="157">Fingers with slight basal web; lengths 3&gt;2&gt;1&gt;4; tips rounded and not dilated; subarticular tubercles distinct, round, and one on digits 1, 2, 4, and two on 3; one large palmar tubercle on outer half of palm, bordered distally by four small, round tubercles. No nuptial pad or asperities present. Toes modestly webbed not extending to proximal subarticular tubercle of toe 4, lengths 4&gt;3&gt;5&gt;2&gt;1; tips rounded, slightly dilated; subarticular tubercles large, round and one on digits 1, 2, two on 3, three on 4, none on 5; large, elongate inner and small, round outer metatarsal tubercles.</paragraph>
<paragraph id="8B93D241CC41FFE7FF005FE6FC56657F" blockId="22.[123,1174,361,1445]" pageId="22" pageNumber="157">In life and at night, dorsum and sides of head and body black from tip of snout to end of trunk; narrow white oblique stripe from tip of snout, passing above eye and tympanum to lower inguinal area; venter from chin to anterior belly fading to dusky thereafter; chest and anterior belly with scattered white spots. During day, dorsum and sides lighten to a medium brown.</paragraph>
</subSubSection>
<subSubSection id="C33681CACC41FFE7FF005866FF1365D8" pageId="22" pageNumber="157" type="etymology">
<paragraph id="8B93D241CC41FFE7FF005866FF1365D8" blockId="22.[123,1174,361,1445]" pageId="22" pageNumber="157">
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<emphasis id="B9580E53CC41FFE7FF005866FEB9659E" bold="true" box="[171,309,1132,1156]" pageId="22" pageNumber="157">ETYMOLOGY</emphasis>
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.— The specific name derives from the Latin
<emphasis id="B9580E53CC41FFE7FCB35866FCC86598" box="[792,836,1132,1156]" italics="true" pageId="22" pageNumber="157">calx</emphasis>
for limestone and the Latinized Greek suffix
<emphasis id="B9580E53CC41FFE7FEA25886FEC665B8" box="[265,330,1164,1188]" italics="true" pageId="22" pageNumber="157">philus</emphasis>
for “who is attracted to” combined to denote this species affinity to karst forest.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC41FFE7FF0058C6FD8E6498" pageId="22" pageNumber="157" type="distribution">
<paragraph id="8B93D241CC41FFE7FF0058C6FE0C6418" blockId="22.[123,1174,361,1445]" pageId="22" pageNumber="157">
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<emphasis id="B9580E53CC41FFE7FF0058C6FECF65FF" bold="true" box="[171,323,1228,1252]" pageId="22" pageNumber="157">DISTRIBUTION</emphasis>
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.—
<collectingCountry id="F33B92D1CC41FFE7FEC058C6FE4B65F8" box="[363,455,1228,1252]" name="Malaysia" pageId="22" pageNumber="157">Sarawak</collectingCountry>
, Borneo. Known only from the karst forest in Gunung Mulu National Park,
<collectingCountry id="F33B92D1CC41FFE7FEB658E6FEF56418" box="[285,377,1260,1284]" name="Malaysia" pageId="22" pageNumber="157">Sarawak</collectingCountry>
.
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<emphasis id="B9580E53CC41FFE7FF005907FE9D643F" bold="true" box="[171,273,1293,1317]" pageId="22" pageNumber="157">NATURAL</emphasis>
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<emphasis id="B9580E53CC41FFE7FEBC5907FEF4643F" bold="true" box="[279,376,1293,1317]" pageId="22" pageNumber="157">HISTORY</emphasis>
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.—
<taxonomicName id="4C2CA9C2CC41FFE7FE0B5906FD2A6438" authorityName="Dehling" authorityYear="2011" box="[416,678,1292,1316]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="157" phylum="Chordata" rank="species" species="calciphilus">
<emphasis id="B9580E53CC41FFE7FE0B5906FD2A6438" box="[416,678,1292,1316]" italics="true" pageId="22" pageNumber="157">Kalophrynus calciphilus</emphasis>
</taxonomicName>
is a terrestrial species, apparently confined to karst forest. It calls day and night from the entrance of small burrows in the soil and owing to the absent of standing water in the karst forest, it is hypothesized that eggs are laid in these burrows and possibly “guarded” by the male.
</paragraph>
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<emphasis id="B9580E53CC41FFE7FF005987FEA464BF" bold="true" box="[171,296,1421,1445]" pageId="22" pageNumber="157">COMMENTS</emphasis>
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.— Preceding information extracted from
<bibRefCitation id="EFBDAFB0CC41FFE7FD405987FC0264B9" author="DEHLING, J. M." box="[747,910,1421,1445]" pageId="22" pageNumber="157" pagination="49 - 60" refId="ref37239" refString="DEHLING, J. M. 2011. A new karst-dwelling species of Kalophrynus (Anura: Microhylidae) from Gunung Mulu National Park, Borneo, Malaysia. Zootaxa 2737: 49 - 60." type="journal article" year="2011">Dehling (2011)</bibRefCitation>
.
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</treatment>
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<taxonomicName id="4C2CA9C2CC58FFFEFFD05A1AFEC96734" ID-CoL="6NK3C" authority="Zug, 2015" authorityName="Zug" authorityYear="2015" box="[123,325,1552,1576]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="150" phylum="Chordata" rank="species" species="anya">
<emphasis id="B9580E53CC58FFFEFFD05A1AFEC96734" bold="true" box="[123,325,1552,1576]" italics="true" pageId="15" pageNumber="150">Kalophrynus anya</emphasis>
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<collectingRegion id="49E81CA3CC58FFFEFEE65A1AFEF06734" box="[333,380,1552,1576]" country="Switzerland" name="Zug" pageId="15" pageNumber="150">Zug</collectingRegion>
, sp. nov.
<location id="8EF3849ACC58FFFEFC865A1AFB026734" LSID="urn:lsid:plazi:treatment:03856357CC58FFE2FFD05A1AFDD2657E:8EF3849ACC58FFFEFC865A1AFB026734" box="[813,1166,1552,1576]" name="Northern Burmese Sticky Frog" pageId="15" pageNumber="150" stateProvince="Zug">Northern Burmese Sticky Frog</location>
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, sp. nov.
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<location id="8EF3849ACC58FFFEFC865A1AFB026734" LSID="urn:lsid:plazi:treatment:03856357CC58FFE2FFD05A1AFDD2657E:8EF3849ACC58FFFEFC865A1AFB026734" box="[813,1166,1552,1576]" name="Northern Burmese Sticky Frog" pageId="15" pageNumber="150" stateProvince="Zug">Northern Burmese Sticky Frog</location>
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<figureCitation id="1317CEC4CC58FFFEFFD05A3AFF41675A" box="[123,205,1584,1606]" captionStart="FIGURE" captionStartId="18.[155,167,1675,1694]" captionText="FIGURE 5 (left). Images of holotype of Kalophrynus anya (USNM 537420, male, 39.7 mm SVL) in (A) dorsal, (B) lateral, and (C) ventral views [photographer, J.A. Poindexter]." pageId="15" pageNumber="150">Figure 5</figureCitation>
<figureCitation id="1317CEC4CC58FFFEFFD05A3AFF41675A" box="[123,205,1584,1606]" captionStart="FIGURE" captionStartId="18.[155,167,1675,1694]" captionTargetId="figure-23@17.[197,1106,216,1731]" captionTargetPageId="17" captionText="FIGURE 5 (left). Images of holotype of Kalophrynus anya (USNM 537420, male, 39.7 mm SVL) in (A) dorsal, (B) lateral, and (C) ventral views [photographer, J.A. Poindexter]." figureDoi="http://doi.org/10.5281/zenodo.12752982" httpUri="https://zenodo.org/record/12752982/files/figure.png" pageId="15" pageNumber="150">Figure 5</figureCitation>
,
<figureCitation id="1317CEC4CC58FFFEFF725A3AFEAF675A" box="[217,291,1584,1606]" captionStart="FIGURE" captionStartId="44.[155,167,1351,1370]" captionTargetBox="[169,1129,216,1336]" captionTargetId="figure-124@44.[169,1129,215,1336]" captionTargetPageId="44" captionText="FIGURE 11. Coloration of living representatives of pleurostigma-interlineatus group. (A) Dorsolateral view of a K. anya (USNM 520321) from the Chatthin Wildlife Sanctuary, Sagaing, Myanmar [photographer, C. Hansen]; (B) dorsolateral view of a K. anya (USNM 523966) from the Chatthin Wildlife Sanctuary, Sagaing, Myanmar [photographer, G.R. Zug]; (C) dorsolateral view of a K. interlineatus (CAS) from the Tanintharyi National Park, Tanintharyi, Myanmar [photograph- er, J.V. Vindum]; (D) dorsolateral view of a K. meizon (FMNH 242796) from Poring Station, Mt. Kinabulu Park, Ranua District, Sabah [photographer, R.F. Inger]; (E) dorsolateral view of a K. sinensis (UK 333148) from Agusan del Norte, Mindanao, Philippines [photographer, R.M. Brown]; (F) dorsolateral view of a K. pleurostigma (UK 333148) from Gunung Kunyit, Jambi, Sumatra [photographer, E.N. Smith]." figureDoi="http://doi.org/10.5281/zenodo.11512262" httpUri="https://zenodo.org/record/11512262/files/figure.png" pageId="15" pageNumber="150">11A–B.</figureCitation>
</paragraph>
@ -430,7 +434,7 @@ to the region of Upper
endemic species.
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5 (left). Images of holotype of

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<mods:title id="2BEE1579A1F01924891593EED72AD8AE">Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs</mods:title>
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<taxonomicName id="4C2CA9C2CC5BFFFDFD3A5F6DFCB1629D" ID-CoL="62W7Y" authority="Tschudi, 1838" authorityName="Tschudi" authorityYear="1838" box="[657,829,871,899]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="147" phylum="Chordata" rank="genus">
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<emphasis id="B9580E53CC5BFFFDFD3A5F6DFCB1629D" box="[657,829,871,899]" italics="true" pageId="12" pageNumber="147">KALOPHRYNUS</emphasis>
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Species are arranged alphabetically. The two new species described herein are
<taxonomicName id="4C2CA9C2CC5BFFFDFBA75F94FF2362CA" authorityName="Zug" authorityYear="2015" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="anya">
<emphasis id="B9580E53CC5BFFFDFBA75F94FF2362CA" italics="true" pageId="12" pageNumber="147">Kalophrynus anya</emphasis>
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from northern
<collectingCountry id="F33B92D1CC5BFFFDFEF35FB5FE4C62CB" box="[344,448,959,983]" name="Myanmar" pageId="12" pageNumber="147">Myanmar</collectingCountry>
and
<taxonomicName id="4C2CA9C2CC5BFFFDFE525FB5FDEA62CA" authorityName="Zug" authorityYear="2015" box="[505,614,959,982]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="meizon">
<emphasis id="B9580E53CC5BFFFDFE525FB5FDEA62CA" box="[505,614,959,982]" italics="true" pageId="12" pageNumber="147">K. meizon</emphasis>
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from Borneo. Those two taxa and the members of
<emphasis id="B9580E53CC5BFFFDFFD05FD5FE1A62EB" box="[123,406,991,1015]" italics="true" pageId="12" pageNumber="147">
<taxonomicName id="4C2CA9C2CC5BFFFDFFD05FD5FE8E62EB" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[123,258,991,1015]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="interlineatus">interlineatus</taxonomicName>
-
<taxonomicName id="4C2CA9C2CC5BFFFDFEA15FD5FE1A62EB" box="[266,406,991,1015]" class="Amphibia" family="Microhylidae" genus="Calophrynus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="pleurostigma">pleurostigma</taxonomicName>
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group of populations (i.e.,
<taxonomicName id="4C2CA9C2CC5BFFFDFD1F5FD5FCD562EB" authorityName="Striped Sticky Frog" baseAuthorityName="Blyth" baseAuthorityYear="1855" box="[692,857,991,1015]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC5BFFFDFD1F5FD5FCD562EB" box="[692,857,991,1015]" italics="true" pageId="12" pageNumber="147">K. interlineatus</emphasis>
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,
<taxonomicName id="4C2CA9C2CC5BFFFDFCCE5FD5FB8362EB" box="[869,1039,991,1015]" class="Amphibia" family="Microhylidae" genus="Calophrynus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="pleurostigma">
<emphasis id="B9580E53CC5BFFFDFCCE5FD5FB8362EB" box="[869,1039,991,1015]" italics="true" pageId="12" pageNumber="147">K. pleurostigma</emphasis>
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,
<taxonomicName id="4C2CA9C2CC5BFFFDFBB05FD5FB0162EA" authorityName="Peters" authorityYear="1867" box="[1051,1165,991,1014]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="147" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="B9580E53CC5BFFFDFBB05FD5FB0162EA" box="[1051,1165,991,1014]" italics="true" pageId="12" pageNumber="147">K. sinensis</emphasis>
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;
<figureCitation id="1317CEC4CC5BFFFDFFD05FF5FF35650B" box="[123,185,1023,1047]" captionStart="FIGURE" captionStartId="15.[155,167,1076,1095]" captionTargetBox="[163,1133,216,1060]" captionTargetId="figure-290@15.[163,1133,216,1060]" captionTargetPageId="15" captionText="FIGURE 4. General distribution of the Kalophrynus interlineatus-pleurostigma group. Solid circles denote specimens examined in this study; open circles are literature records and museum records of specimens not seen; stars represent type localities of available names. Source of the locality records are in Appendix Specimens examined section." figureDoi="http://doi.org/10.5281/zenodo.11512250" httpUri="https://zenodo.org/record/11512250/files/figure.png" pageId="12" pageNumber="147">Fig. 4</figureCitation>
) have more detailed descriptive synopses than the other species in order to present the morphological details from the preceding analysis. Morphological details for the remainder (and majority) of the
<taxonomicName id="4C2CA9C2CC5BFFFDFF46584AFE476544" box="[237,459,1088,1112]" pageId="12" pageNumber="147">
<emphasis id="B9580E53CC5BFFFDFF46584AFEFB6544" box="[237,375,1088,1112]" italics="true" pageId="12" pageNumber="147">Kalophrynus</emphasis>
species
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derive entirely from published information, as I examine no
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or other specimens of these taxa. Measurements and proportions are for adults unless noted otherwise. NA indicates that data were not available in any published studies.
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