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<mods:namePart id="F047345768C4D7508240AACD3E9DFF47">Ortea, Jesús</mods:namePart>
<mods:affiliation id="C02DC3167EC6CE11B36634A050174F05">Departamento de Biología de Organismos y Sistemas, Laboratorio de Zoología Universidad de Oviedo, 33071 Oviedo (Spain) jortea @ sci. cpd. uniovi. es</mods:affiliation>
<mods:nameIdentifier id="E39FE29DCCACB6760F23F8F3A0F32307" type="email">jortea@sci.cpd.uniovi.es</mods:nameIdentifier>
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<mods:typeOfResource id="1CE22FCF9F704C74DAD66EE418956CF1">text</mods:typeOfResource>
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<mods:titleInfo id="BD05617D714C0E0901190D49AF0D070C">
<mods:title id="4A46AAEFF778E4962E4C85CF95AB23AF">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="FB9207C0F0082D5A8A8CEE32D6287AE6">
<mods:date id="C6805CD29863F2867AE457753FF38842">2000</mods:date>
<mods:detail id="28D918DD26A818F94DC9545A5430FBBD" type="volume">
<mods:number id="9C999709417435922D1F797FDA64653D">22</mods:number>
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<mods:number id="19F48F42284E32FD8160027E88805F0B">1</mods:number>
</mods:detail>
<mods:extent id="71FBE124C0AF692D8267D761EDA59EEF" unit="page">
<mods:start id="EE9C27569A3E7C0521EB57D0276CFB73">15</mods:start>
<mods:end id="09F4B721457141FA8765212DB21E5378">31</mods:end>
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<mods:classification id="F7C0ADE8420D0F64D2793B6F10CDEFA3">journal article</mods:classification>
<mods:identifier id="C72E47293BA93CC941B9B9E00497E3AB" type="DOI">10.5281/zenodo.5398985</mods:identifier>
<mods:identifier id="F6374DF0BCE05E5752EA12CF89F54601" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="DA58705C7843F90955627D0570B21F7A" type="Zenodo-Dep">5398985</mods:identifier>
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<treatment id="096B7348DD69FF87FF00FF72FDDCFE4E" LSID="urn:lsid:plazi:treatment:096B7348DD69FF87FF00FF72FDDCFE4E" httpUri="http://treatment.plazi.org/id/096B7348DD69FF87FF00FF72FDDCFE4E" lastPageId="5" lastPageNumber="19" pageId="3" pageNumber="18">
<subSubSection id="C9D891D5DD69FF81FF00FF72FE7DFEF8" pageId="3" pageNumber="18" type="nomenclature">
<paragraph id="817DC25EDD69FF81FF00FF72FDF5FF1B" blockId="3.[194,566,208,265]" box="[194,566,208,234]" pageId="3" pageNumber="18">
<taxonomicName id="46C2B9DDDD69FF81FF00FF72FDF5FF1B" ID-CoL="74J38" authority="(Edmunds, 1966)" baseAuthorityName="Edmunds" baseAuthorityYear="1966" box="[194,566,208,234]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="18" phylum="Mollusca" rank="species" species="digitata">
<emphasis id="B3B61E4CDD69FF81FF00FF72FEADFF1B" bold="true" box="[194,366,208,234]" italics="true" pageId="3" pageNumber="18">Okenia digitata</emphasis>
(
<bibRefCitation id="E553BFAFDD69FF81FEBFFF72FDEDFF1B" author="Edmunds M." box="[381,558,208,234]" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="3" pageNumber="18" pagination="69 - 72" part="37" refId="ref8352" refString="Edmunds M. 1966. - Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana. Proceedings of the Malacological Society of London 37: 69 - 72." title="Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana" type="journal article" year="1966">Edmunds, 1966</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="817DC25EDD69FF81FEF8FF4DFE7DFEF8" blockId="3.[194,566,208,265]" box="[314,446,239,265]" pageId="3" pageNumber="18">
(
<figureCitation id="19F9DEDBDD69FF81FE83FF4DFE46FEF8" box="[321,389,239,265]" captionStart="FIG" captionStartId="3.[113,124,1680,1697]" captionTargetBox="[175,1132,857,1653]" captionTargetId="figure-338@3.[161,1150,850,1669]" captionTargetPageId="3" captionText="FIG. 2. — Okenia digitata (MNHN), dorsal view of the living animal. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5398991" httpUri="https://zenodo.org/record/5398991/files/figure.png" pageId="3" pageNumber="18">Figs 2</figureCitation>
;
<figureCitation id="19F9DEDBDD69FF81FE53FF4DFE76FEF8" box="[401,437,239,265]" captionStart="FIG" captionStartId="4.[114,125,1021,1038]" captionTargetBox="[119,1203,204,981]" captionTargetId="figure-378@4.[119,1203,199,981]" captionTargetPageId="4" captionText="FIG. 3. — A, Kaloplocamus ramosus (MNHN), rachis of the radula; B, Kaloplocamus ramosus (MNHN), inner lateral teeth; C, Kaloplocamus ramosus (MNHN), outer lateral teeth; D, Okenia digitata (MNHN), radular teeth morphology. Scale bars: A, C, 100 µm; B, 20 µm; D, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5398995" httpUri="https://zenodo.org/record/5398995/files/figure.png" pageId="3" pageNumber="18">3D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD69FF81FFB3FE8AFD90FEB1" box="[113,595,296,320]" pageId="3" pageNumber="18" type="reference_group">
<paragraph id="817DC25EDD69FF81FFB3FE8AFD90FEB1" blockId="3.[113,647,296,578]" box="[113,595,296,320]" pageId="3" pageNumber="18">
<taxonomicName id="46C2B9DDDD69FF81FFB3FE8AFE2FFEB1" authority="Edmunds, 1966: 69 - 72" authorityName="Edmunds" authorityPageNumber="69 - 72" authorityYear="1966" box="[113,492,296,320]" class="Gastropoda" family="Goniodorididae" genus="Teshia" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="18" phylum="Mollusca" rank="species" species="digitata">
<emphasis id="B3B61E4CDD69FF81FFB3FE8AFF3DFEB1" box="[113,254,296,320]" italics="true" pageId="3" pageNumber="18">Teshia digitata</emphasis>
<bibRefCitation id="E553BFAFDD69FF81FEC6FE8AFE2FFEB1" author="Edmunds M." box="[260,492,296,320]" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="3" pageNumber="18" pagination="69 - 72" part="37" refId="ref8352" refString="Edmunds M. 1966. - Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana. Proceedings of the Malacological Society of London 37: 69 - 72." title="Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana" type="journal article" year="1966">Edmunds, 1966: 69-72</bibRefCitation>
</taxonomicName>
, figs 1-10.
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD69FF81FFB3FEF2FEDAFE16" pageId="3" pageNumber="18" type="materials_examined">
<paragraph id="817DC25EDD69FF81FFB3FEF2FEDAFE16" blockId="3.[113,647,296,578]" pageId="3" pageNumber="18">
<materialsCitation id="31AAC803DD69FF81FFB3FEF2FEC3FE6A" collectionCode="MATERIAL" latitude="-8.583333" location="Sao Tiago" longLatPrecision="1301" longitude="13.35" pageId="3" pageNumber="18" specimenCount="1" stateProvince="Bengo">
<collectionCode id="E7D35A9BDD69FF81FFB3FEF2FF15FE97" box="[113,214,336,359]" pageId="3" pageNumber="18">MATERIAL</collectionCode>
EXAMINED. —
<emphasis id="B3B61E4CDD69FF81FEB1FEEDFDE2FE99" bold="true" box="[371,545,335,360]" pageId="3" pageNumber="18">
<collectingRegion id="43060CBCDD69FF81FEB1FEEDFDDEFE99" box="[371,541,335,360]" country="Angola" name="Bengo" pageId="3" pageNumber="18">Bengo Province</collectingRegion>
.
</emphasis>
<collectingRegion id="43060CBCDD69FF81FDEBFEF2FD98FE99" box="[553,603,336,360]" country="Cape Verde" name="Praia" pageId="3" pageNumber="18">Praia</collectingRegion>
<location id="841D9485DD69FF81FDA1FEF2FF6EFE73" LSID="urn:lsid:plazi:treatment:096B7348DD69FF87FF00FF72FDDCFE4E:841D9485DD69FF81FDA1FEF2FF6EFE73" latitude="-8.583333" longLatPrecision="1301" longitude="13.35" name="Sao Tiago" pageId="3" pageNumber="18" stateProvince="Bengo">São Tiago</location>
,
<geoCoordinate id="E4F6A499DD69FF81FF75FEC8FEC5FE73" box="[183,262,362,386]" degrees="08" direction="south" minutes="35" orientation="latitude" pageId="3" pageNumber="18" precision="925" value="-8.583333">08°35S</geoCoordinate>
,
<geoCoordinate id="E4F6A499DD69FF81FED2FEC8FEA0FE71" box="[272,355,361,386]" degrees="13" direction="east" minutes="21" orientation="longitude" pageId="3" pageNumber="18" precision="925" value="13.35">13°21E</geoCoordinate>
, 1983,
<specimenCount id="97C409D7DD69FF81FE6EFEC8FDDAFE73" box="[428,537,362,386]" pageId="3" pageNumber="18" type="generic">1 specimen</specimenCount>
<quantity id="463A6FBBDD69FF81FDDDFEC8FD9AFE71" box="[543,601,361,386]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="18" unit="mm" value="6.0">6 mm</quantity>
preserved length
</materialsCitation>
;
<specimenCount id="97C409D7DD69FF81FECEFE21FE41FE6A" box="[268,386,387,411]" pageId="3" pageNumber="18" type="generic">1 specimen</specimenCount>
<quantity id="463A6FBBDD69FF81FE49FE21FE09FE68" box="[395,458,386,411]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="18" unit="mm" value="5.0">5 mm</quantity>
preserved length;
<materialsCitation id="31AAC803DD69FF81FFB3FE3EFED7FE16" latitude="-8.783334" location="Cacuaco" longLatPrecision="1300" longitude="13.35" pageId="3" pageNumber="18" specimenCount="2" stateProvince="Bengo">
<specimenCount id="97C409D7DD69FF81FFB3FE3EFF2EFE45" box="[113,237,412,436]" pageId="3" pageNumber="18" type="generic">1 specimen</specimenCount>
<quantity id="463A6FBBDD69FF81FF34FE3EFEF6FE42" box="[246,309,412,436]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="18" unit="mm" value="3.0">3 mm</quantity>
preserved length. —
<location id="841D9485DD69FF81FDE6FE3EFD41FE45" LSID="urn:lsid:plazi:treatment:096B7348DD69FF87FF00FF72FDDCFE4E:841D9485DD69FF81FDE6FE3EFD41FE45" box="[548,642,412,436]" latitude="-8.783334" longLatPrecision="1300" longitude="13.35" name="Cacuaco" pageId="3" pageNumber="18" stateProvince="Bengo">Cacuaco</location>
,
<geoCoordinate id="E4F6A499DD69FF81FFB3FE14FF01FE3F" box="[113,194,438,462]" degrees="08" direction="south" minutes="47" orientation="latitude" pageId="3" pageNumber="18" precision="925" value="-8.783334">08°47S</geoCoordinate>
,
<geoCoordinate id="E4F6A499DD69FF81FF0DFE14FEE0FE3D" box="[207,291,437,462]" degrees="13" direction="east" minutes="21" orientation="longitude" pageId="3" pageNumber="18" precision="925" value="13.35">13°21E</geoCoordinate>
, date unknown,
<specimenCount id="97C409D7DD69FF81FE0DFE14FD82FE3F" box="[463,577,438,462]" pageId="3" pageNumber="18" type="generic">1 specimen</specimenCount>
<quantity id="463A6FBBDD69FF81FD8BFE14FD45FE3D" box="[585,646,437,462]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="18" unit="mm" value="6.0">6 mm</quantity>
preserved length
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD69FF81FFB3FE55FF25FDB3" pageId="3" pageNumber="18" type="distribution">
<paragraph id="817DC25EDD69FF81FFB3FE55FF25FDB3" blockId="3.[113,647,296,578]" pageId="3" pageNumber="18">
DISTRIBUTION. — The specimens from
<collectingCountry id="F9D582CEDD69FF81FDC2FE5AFD86FDE1" box="[512,581,504,528]" name="Angola" pageId="3" pageNumber="18">Angola</collectingCountry>
represent the first reported occurrence of this species outside
<collectingCountry id="F9D582CEDD69FF81FF5FFD88FF21FDB3" box="[157,226,554,578]" name="Ghana" pageId="3" pageNumber="18">Ghana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD69FF81FFB3FDD8FD2DFCE1" pageId="3" pageNumber="18" type="description">
<paragraph id="817DC25EDD69FF81FFB3FDD8FEC1FD63" blockId="3.[113,647,634,784]" box="[113,258,634,660]" pageId="3" pageNumber="18">DESCRIPTION</paragraph>
<paragraph id="817DC25EDD69FF81FFB3FD38FD2DFCE1" blockId="3.[113,647,634,784]" lastBlockId="3.[677,1210,205,784]" pageId="3" pageNumber="18">
The background body colour is white in the living animal. On its lateral border the dorsum has 16 tentacular papillae alternating between large and small ones (
<figureCitation id="19F9DEDBDD69FF81FEF7FD54FEBCFCE1" box="[309,383,758,784]" captionStart="FIG" captionStartId="3.[113,124,1680,1697]" captionTargetBox="[175,1132,857,1653]" captionTargetId="figure-338@3.[161,1150,850,1669]" captionTargetPageId="3" captionText="FIG. 2. — Okenia digitata (MNHN), dorsal view of the living animal. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5398991" httpUri="https://zenodo.org/record/5398991/files/figure.png" pageId="3" pageNumber="18">Fig. 2</figureCitation>
). Almost every lateral papilla has, at the same height, a yellow ring, and some black spots. The whole dorsum has small black spots that are more abundant in the centre of the dorsum, being dispersed in the sides. There are two oral tentacles, which are white and similar in form to the lateral papillae. The rhinophores are white as well, with very few black spots. The gill emerges from the body and divides into three branches. Each one has seven branchial leaves. The branchial leaves are unipinnate, white with very little black and have yellow spots dispersed all over. The tail is white with black spots. In each half-row of the radula there is one inner lateral and one outer lateral tooth. The inner lateral tooth (
<figureCitation id="19F9DEDBDD69FF81FCE2FDD9FCBBFD64" box="[800,888,635,661]" captionStart="FIG" captionStartId="4.[114,125,1021,1038]" captionTargetBox="[119,1203,204,981]" captionTargetId="figure-378@4.[119,1203,199,981]" captionTargetPageId="4" captionText="FIG. 3. — A, Kaloplocamus ramosus (MNHN), rachis of the radula; B, Kaloplocamus ramosus (MNHN), inner lateral teeth; C, Kaloplocamus ramosus (MNHN), outer lateral teeth; D, Okenia digitata (MNHN), radular teeth morphology. Scale bars: A, C, 100 µm; B, 20 µm; D, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5398995" httpUri="https://zenodo.org/record/5398995/files/figure.png" pageId="3" pageNumber="18">Fig. 3D</figureCitation>
) is flat and wide at the base. The large cusp on the inner side has ten strong denticles. The outer lateral tooth is thin and the apex is folded, being smaller than the inner lateral.
</paragraph>
</subSubSection>
<caption id="D5BD92D6DD69FF81FFB3F932FCEBF950" ID-DOI="http://doi.org/10.5281/zenodo.5398991" ID-Zenodo-Dep="5398991" box="[113,808,1680,1697]" httpUri="https://zenodo.org/record/5398991/files/figure.png" pageId="3" pageNumber="18" startId="3.[113,124,1680,1697]" targetBox="[175,1132,857,1653]" targetPageId="3">
<paragraph id="817DC25EDD69FF81FFB3F932FCEBF950" blockId="3.[113,808,1680,1697]" box="[113,808,1680,1697]" pageId="3" pageNumber="18">
FIG. 2. —
<taxonomicName id="46C2B9DDDD69FF81FF03F932FE4BF950" authority="(MNHN)" baseAuthorityName="MNHN" box="[193,392,1680,1697]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="18" phylum="Mollusca" rank="species" species="digitata">
<emphasis id="B3B61E4CDD69FF81FF03F932FEFEF950" box="[193,317,1680,1697]" italics="true" pageId="3" pageNumber="18">Okenia digitata</emphasis>
(MNHN)
</taxonomicName>
, dorsal view of the living animal. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="D5BD92D6DD6EFF86FFB0FC5FFEA0FBC9" ID-DOI="http://doi.org/10.5281/zenodo.5398995" ID-Zenodo-Dep="5398995" httpUri="https://zenodo.org/record/5398995/files/figure.png" pageId="4" pageNumber="19" startId="4.[114,125,1021,1038]" targetBox="[119,1203,204,981]" targetPageId="4">
<paragraph id="817DC25EDD6EFF86FFB0FC5FFEA0FBC9" blockId="4.[114,1211,1021,1080]" pageId="4" pageNumber="19">
FIG. 3. —
<emphasis id="B3B61E4CDD6EFF86FF09FC5FFF1BFBFF" bold="true" box="[203,216,1021,1038]" pageId="4" pageNumber="19">A</emphasis>
,
<taxonomicName id="46C2B9DDDD6EFF86FF24FC5FFDC4FBFF" authority="(MNHN)" baseAuthorityName="MNHN" box="[230,519,1021,1038]" class="Gastropoda" family="Polyceridae" genus="Kaloplocamus" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="species" species="ramosus">
<emphasis id="B3B61E4CDD6EFF86FF24FC5FFE74FBFF" box="[230,439,1021,1038]" italics="true" pageId="4" pageNumber="19">Kaloplocamus ramosus</emphasis>
(MNHN)
</taxonomicName>
, rachis of the radula;
<emphasis id="B3B61E4CDD6EFF86FD12FC5FFD1FFBFF" bold="true" box="[720,732,1021,1038]" pageId="4" pageNumber="19">B</emphasis>
,
<taxonomicName id="46C2B9DDDD6EFF86FD28FC5FFBC8FBFF" authority="(MNHN)" baseAuthorityName="MNHN" box="[746,1035,1021,1038]" class="Gastropoda" family="Polyceridae" genus="Kaloplocamus" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="species" species="ramosus">
<emphasis id="B3B61E4CDD6EFF86FD28FC5FFC79FBFF" box="[746,954,1021,1038]" italics="true" pageId="4" pageNumber="19">Kaloplocamus ramosus</emphasis>
(MNHN)
</taxonomicName>
, inner lateral teeth;
<emphasis id="B3B61E4CDD6EFF86FFB1FBB0FFBCFBD2" bold="true" box="[115,127,1042,1059]" pageId="4" pageNumber="19">C</emphasis>
,
<taxonomicName id="46C2B9DDDD6EFF86FF4EFBB0FE62FBD2" authority="(MNHN)" baseAuthorityName="MNHN" box="[140,417,1042,1059]" class="Gastropoda" family="Polyceridae" genus="Kaloplocamus" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="species" species="ramosus">
<emphasis id="B3B61E4CDD6EFF86FF4EFBB0FE90FBD2" box="[140,339,1042,1059]" italics="true" pageId="4" pageNumber="19">Kaloplocamus ramosus</emphasis>
(MNHN)
</taxonomicName>
, outer lateral teeth;
<emphasis id="B3B61E4CDD6EFF86FD93FBB0FD9EFBD2" bold="true" box="[593,605,1042,1059]" pageId="4" pageNumber="19">D</emphasis>
,
<taxonomicName id="46C2B9DDDD6EFF86FDABFBB0FCF4FBD2" authority="(MNHN)" baseAuthorityName="MNHN" box="[617,823,1042,1059]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="species" species="digitata">
<emphasis id="B3B61E4CDD6EFF86FDABFBB0FD29FBD2" box="[617,746,1042,1059]" italics="true" pageId="4" pageNumber="19">Okenia digitata</emphasis>
(MNHN)
</taxonomicName>
, radular teeth morphology. Scale bars: A, C, 100 µm; B, 20 µm; D, 10 µm.
</paragraph>
</caption>
<subSubSection id="C9D891D5DD6EFF87FFB0FBC6FDDCFE4E" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="discussion">
<paragraph id="817DC25EDD6EFF86FFB0FBC6FF15FB8D" blockId="4.[114,649,1124,1704]" box="[114,214,1124,1150]" pageId="4" pageNumber="19">REMARKS</paragraph>
<paragraph id="817DC25EDD6EFF86FFB0FB27FF0AFA01" blockId="4.[114,649,1124,1704]" pageId="4" pageNumber="19">
The genus
<taxonomicName id="46C2B9DDDD6EFF86FF35FB27FDC1FB6E" authority="Edmunds, 1966" authorityName="Edmunds" authorityYear="1966" box="[247,514,1157,1183]" class="Gastropoda" family="Goniodorididae" genus="Teshia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6EFF86FF35FB27FEFFFB6E" box="[247,316,1157,1183]" italics="true" pageId="4" pageNumber="19">Teshia</emphasis>
<bibRefCitation id="E553BFAFDD6EFF86FE86FB27FDC1FB6E" author="Edmunds M." box="[324,514,1157,1183]" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="4" pageNumber="19" pagination="69 - 72" part="37" refId="ref8352" refString="Edmunds M. 1966. - Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana. Proceedings of the Malacological Society of London 37: 69 - 72." title="Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana" type="journal article" year="1966">Edmunds, 1966</bibRefCitation>
</taxonomicName>
, was introduced by
<bibRefCitation id="E553BFAFDD6EFF86FF27FB01FE68FB4C" author="Edmunds M." box="[229,427,1187,1213]" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="4" pageNumber="19" pagination="69 - 72" part="37" refId="ref8352" refString="Edmunds M. 1966. - Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana. Proceedings of the Malacological Society of London 37: 69 - 72." title="Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana" type="journal article" year="1966">Edmunds (1966)</bibRefCitation>
for
<taxonomicName id="46C2B9DDDD6EFF86FE21FB01FEFDFB2D" authority="Edmunds, 1966" authorityName="Edmunds" authorityYear="1966" class="Gastropoda" family="Goniodorididae" genus="Teshia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="species" species="digitata">
<emphasis id="B3B61E4CDD6EFF86FE21FB01FD4AFB4C" box="[483,649,1187,1213]" italics="true" pageId="4" pageNumber="19">Teshia digitata</emphasis>
<bibRefCitation id="E553BFAFDD6EFF86FFB0FB60FEFDFB2D" author="Edmunds M." box="[114,318,1218,1244]" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="4" pageNumber="19" pagination="69 - 72" part="37" refId="ref8352" refString="Edmunds M. 1966. - Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana. Proceedings of the Malacological Society of London 37: 69 - 72." title="Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana" type="journal article" year="1966">Edmunds, 1966</bibRefCitation>
</taxonomicName>
, as different from
<taxonomicName id="46C2B9DDDD6EFF86FDEDFB60FECEFB0A" authority="Menke, 1830" authorityName="Menke" authorityYear="1830" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6EFF86FDEDFB60FD45FB2D" box="[559,646,1218,1244]" italics="true" pageId="4" pageNumber="19">Okenia</emphasis>
Menke, 1830
</taxonomicName>
. The main characteristic used by Edmunds to separate both genera was that
<taxonomicName id="46C2B9DDDD6EFF86FD84FB5DFD44FAE8" authorityName="Edmunds" authorityYear="1966" box="[582,647,1279,1305]" class="Gastropoda" family="Goniodorididae" genus="Teshia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6EFF86FD84FB5DFD44FAE8" box="[582,647,1279,1305]" italics="true" pageId="4" pageNumber="19">Teshia</emphasis>
</taxonomicName>
had three sets of branchial leaves whereas
<taxonomicName id="46C2B9DDDD6EFF86FDF9FABCFD4BFAC9" authorityName="Menke" authorityYear="1830" box="[571,648,1310,1336]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6EFF86FDF9FABCFD4BFAC9" box="[571,648,1310,1336]" italics="true" pageId="4" pageNumber="19">Okenia</emphasis>
</taxonomicName>
just had one, all other features of
<taxonomicName id="46C2B9DDDD6EFF86FE3FFA9FFD82FAA6" authorityName="Edmunds" authorityYear="1966" box="[509,577,1341,1367]" class="Gastropoda" family="Goniodorididae" genus="Teshia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6EFF86FE3FFA9FFD82FAA6" box="[509,577,1341,1367]" italics="true" pageId="4" pageNumber="19">Teshia</emphasis>
</taxonomicName>
being identical to those of
<taxonomicName id="46C2B9DDDD6EFF86FE96FAF9FE63FA84" authorityName="Menke" authorityYear="1830" box="[340,416,1371,1397]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6EFF86FE96FAF9FE63FA84" box="[340,416,1371,1397]" italics="true" pageId="4" pageNumber="19">Okenia</emphasis>
</taxonomicName>
. In our opinion, this difference is not enough to separate a different genus and therefore both genera are here regarded as synonyms, with
<taxonomicName id="46C2B9DDDD6EFF86FE89FA15FDF3FA20" authority="Menke, 1830" authorityName="Menke" authorityYear="1830" box="[331,560,1463,1489]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6EFF86FE89FA15FE5AFA20" box="[331,409,1463,1489]" italics="true" pageId="4" pageNumber="19">Okenia</emphasis>
Menke,1830
</taxonomicName>
, having priority.
</paragraph>
<paragraph id="817DC25EDD6EFF86FFB0FA57FC38FA20" blockId="4.[114,649,1124,1704]" lastBlockId="4.[677,1210,1125,1704]" pageId="4" pageNumber="19">
The specimens from
<collectingCountry id="F9D582CEDD6EFF86FE98FA57FE61F9FE" box="[346,418,1525,1551]" name="Ghana" pageId="4" pageNumber="19">Ghana</collectingCountry>
and
<collectingCountry id="F9D582CEDD6EFF86FE1EFA57FDEBF9FE" box="[476,552,1525,1551]" name="Angola" pageId="4" pageNumber="19">Angola</collectingCountry>
are very similar in their colour pattern, however they s h ow s o m e d i f f e re n c e s. Ed m u n d s (1 9 6 6) described his single specimen as being grey with dark maroon spots on the dorsal and lateral areas of the body, and pale yellow spots on the flanks. The long rhinophores were grey on the stalk and maroon on the club. Gill and rhinophores were both scattered by maroon spots and small white dots. On the contrary, our specimens are white instead of grey, and have black small spots instead of maroon. The rhinophores of our specimens are white and the gill is yellow. The radula appears to be similar between the material of
<collectingCountry id="F9D582CEDD6EFF86FBF9FA9FFB45FAA6" box="[1083,1158,1341,1367]" name="Ghana" pageId="4" pageNumber="19">Ghana</collectingCountry>
and
<collectingCountry id="F9D582CEDD6EFF86FD67FAF9FD36FA84" box="[677,757,1371,1397]" name="Angola" pageId="4" pageNumber="19">Angola</collectingCountry>
. Both have an outer lateral tooth, folded on the apex, much smaller than the innermost one, which has a strong cusp with eight to ten wide denticles in the inner side.
</paragraph>
<paragraph id="817DC25EDD6EFF87FD67FA74FDDCFE4E" blockId="4.[677,1210,1125,1704]" lastBlockId="5.[113,647,206,447]" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<bibRefCitation id="E553BFAFDD6EFF86FD67FA74FCA2FA01" author="Edmunds M." box="[677,865,1494,1520]" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="4" pageNumber="19" pagination="69 - 72" part="37" refId="ref8352" refString="Edmunds M. 1966. - Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana. Proceedings of the Malacological Society of London 37: 69 - 72." title="Teshia digitata gen. and sp. nov., a dorid nudibranch from Ghana" type="journal article" year="1966">Edmunds (1966)</bibRefCitation>
described the gill of
<taxonomicName id="46C2B9DDDD6EFF86FB86FA77FB7AFA01" authorityName="Edmunds" authorityYear="1966" box="[1092,1209,1493,1520]" class="Gastropoda" family="Goniodorididae" genus="Teshia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="19" phylum="Mollusca" rank="species" species="digitata">
<emphasis id="B3B61E4CDD6EFF86FB86FA77FB7AFA01" box="[1092,1209,1493,1520]" italics="true" pageId="4" pageNumber="19">O. digitata</emphasis>
</taxonomicName>
as having three different sets of branchial leaves that emerge from three different points of the body. Instead, we observed that our specimens have three different sets emerging from a single point. The collector observed this character on the living animals, which after preservation cannot be properly observed. On preserved animals the gill seems to be just as Edmunds description except for the presence of a triangular area differently pigmented that joins the three sets of gills. We consi- der that the gills in both animals are similar and that Edmunds description could be based on preserved animals, for this reason the true nature of the gills could have been misinterpreted.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,216 @@
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<subSubSection id="C9D891D5DD6BFF83FD2CFC33FBB3FC5A" box="[750,1136,912,939]" pageId="1" pageNumber="16" type="nomenclature">
<paragraph id="817DC25EDD6BFF83FD2CFC33FBB3FC5A" blockId="1.[750,1136,912,969]" box="[750,1136,912,939]" pageId="1" pageNumber="16">
<heading id="DA357532DD6BFF83FD2CFC33FBB3FC5A" box="[750,1136,912,939]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="16" reason="2">
<taxonomicName id="46C2B9DDDD6BFF83FD2CFC33FBB3FC5A" ID-CoL="9G5LY" authority="(Bergh, 1907)" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[750,1136,912,939]" class="Gastropoda" family="Goniodorididae" genus="Okenia" isUncertain="true" kingdom="Animalia" order="Nudibranchia" pageId="1" pageNumber="16" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD6BFF83FD2CFC33FCFDFC5A" bold="true" box="[750,830,913,939]" italics="true" pageId="1" pageNumber="16">Okenia</emphasis>
cf.
<emphasis id="B3B61E4CDD6BFF83FCA4FC33FC12FC5A" bold="true" box="[870,977,913,939]" italics="true" pageId="1" pageNumber="16">amoenula</emphasis>
(
<bibRefCitation id="E553BFAFDD6BFF83FC1DFC33FBABFC5A" author="Bergh R." bookContentInfo="144 p." box="[991,1128,913,939]" journalOrPublisher="Transactions of the South African Philosophical Society" pageId="1" pageNumber="16" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh, 1907</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FC45FC0DFC14FC38" box="[903,983,943,969]" pageId="1" pageNumber="16" type="description">
<paragraph id="817DC25EDD6BFF83FC45FC0DFC14FC38" blockId="1.[750,1136,912,969]" box="[903,983,943,969]" pageId="1" pageNumber="16">
(
<figureCitation id="19F9DEDBDD6BFF83FC4DFC0DFC0FFC38" box="[911,972,943,969]" captionStart="FIG" captionStartId="2.[114,125,758,775]" captionTargetBox="[120,1207,205,709]" captionTargetId="figure-509@2.[111,1218,196,722]" captionTargetPageId="2" captionText="FIG. 1. — Okenia cf. amoenula (MNHN), dorsal view of the living animal. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5398989" httpUri="https://zenodo.org/record/5398989/files/figure.png" pageId="1" pageNumber="16">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FD67FC43FB79FC08" box="[677,1210,993,1017]" pageId="1" pageNumber="16" type="reference_group">
<paragraph id="817DC25EDD6BFF83FD67FC43FB79FC08" blockId="1.[677,1210,993,1192]" box="[677,1210,993,1017]" pageId="1" pageNumber="16">
<taxonomicName id="46C2B9DDDD6BFF83FD67FC43FBDFFC08" authority="Bergh, 1907: 80 - 82" authorityName="Bergh" authorityPageNumber="80 - 82" authorityYear="1907" box="[677,1052,993,1017]" class="Gastropoda" family="Goniodorididae" genus="Idaliella" kingdom="Animalia" order="Nudibranchia" pageId="1" pageNumber="16" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD6BFF83FD67FC43FC90FC08" box="[677,851,993,1017]" italics="true" pageId="1" pageNumber="16">Idaliella amoenula</emphasis>
<bibRefCitation id="E553BFAFDD6BFF83FC9BFC43FBDFFC08" author="Bergh R." bookContentInfo="144 p." box="[857,1052,993,1017]" journalOrPublisher="Transactions of the South African Philosophical Society" pageId="1" pageNumber="16" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh, 1907: 80-82</bibRefCitation>
</taxonomicName>
, pl. 8, figs 6-11.
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FD67FBA7FC4EFBA1" pageId="1" pageNumber="16" type="materials_examined">
<paragraph id="817DC25EDD6BFF83FD67FBA7FC4EFBA1" blockId="1.[677,1210,993,1192]" pageId="1" pageNumber="16">
MATERIAL EXAMINED. —
<materialsCitation id="31AAC803DD6BFF83FC63FBA6FC4EFBA1" pageId="1" pageNumber="16">
<emphasis id="B3B61E4CDD6BFF83FC63FBA6FB91FBEF" bold="true" box="[929,1106,1028,1054]" pageId="1" pageNumber="16">
<collectingRegion id="43060CBCDD6BFF83FC63FBA6FB8EFBEF" box="[929,1101,1028,1054]" country="Angola" name="Luanda" pageId="1" pageNumber="16">Luanda Province</collectingRegion>
.
</emphasis>
Punta das Palmeirinhas,
<geoCoordinate id="E4F6A499DD6BFF83FCF5FBBDFCBFFBC6" box="[823,892,1055,1079]" degrees="9" direction="south" minutes="09" orientation="latitude" pageId="1" pageNumber="16" precision="925" value="-9.15">9°09S</geoCoordinate>
,
<geoCoordinate id="E4F6A499DD6BFF83FC4BFBBDFC1DFBC4" box="[905,990,1054,1079]" degrees="12" direction="east" minutes="58" orientation="longitude" pageId="1" pageNumber="16" precision="925" value="12.966666">12°58E</geoCoordinate>
,
<date id="F57CE49EDD6BFF83FC29FBBDFBFAFBC6" box="[1003,1081,1055,1079]" pageId="1" pageNumber="16" value="1987-02">II.1987</date>
,
<specimenCount id="97C409D7DD6BFF83FB85FBBDFB79FBC6" box="[1095,1210,1055,1079]" pageId="1" pageNumber="16" type="generic">1 specimen</specimenCount>
<quantity id="463A6FBBDD6BFF83FD67FB9AFD23FBBF" box="[677,736,1079,1104]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="16" unit="mm" value="6.0">6 mm</quantity>
preserved length.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FD67FBFEFB61FB59" pageId="1" pageNumber="16" type="distribution">
<paragraph id="817DC25EDD6BFF83FD67FBFEFB61FB59" blockId="1.[677,1210,993,1192]" pageId="1" pageNumber="16">
DISTRIBUTION. —
<taxonomicName id="46C2B9DDDD6BFF83FCAFFBFFFBE3FB84" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[877,1056,1117,1141]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="1" pageNumber="16" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD6BFF83FCAFFBFFFBE3FB84" box="[877,1056,1117,1141]" italics="true" pageId="1" pageNumber="16">Okenia amoenula</emphasis>
</taxonomicName>
has only been reported from
<collectingCountry id="F9D582CEDD6BFF83FCFDFBD4FC02FB7F" box="[831,961,1142,1166]" name="South Africa" pageId="1" pageNumber="16">South Africa</collectingCountry>
(
<bibRefCitation id="E553BFAFDD6BFF83FC13FBD4FBB4FB7F" author="Gosliner T. M." bookContentInfo="Sea Challengers, Monterey, 136 p." box="[977,1143,1142,1166]" pageId="1" pageNumber="16" refId="ref8424" refString="Gosliner T. M. 1987 a. - Nudibranchs of Southern Africa. A Guide to Opisthobranch Mollusks of Southern Africa. Sea Challengers, Monterey, 136 p." title="Nudibranchs of Southern Africa. A Guide to Opisthobranch Mollusks of Southern Africa" type="book" year="1987">Gosliner 1987a</bibRefCitation>
). This paper is the first reference for this species to
<collectingCountry id="F9D582CEDD6BFF83FB97FB32FB5DFB59" box="[1109,1182,1168,1192]" name="Angola" pageId="1" pageNumber="16">Angola</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD6BFF83FD67FB42FBB9F956" pageId="1" pageNumber="16" type="description">
<paragraph id="817DC25EDD6BFF83FD67FB42FCF6FB09" blockId="1.[677,1210,1248,1704]" box="[677,821,1248,1274]" pageId="1" pageNumber="16">DESCRIPTION</paragraph>
<paragraph id="817DC25EDD6BFF83FD67FB5DFBB9F956" blockId="1.[677,1210,1248,1704]" pageId="1" pageNumber="16">
The background body colour is opaque white in the living animal. In the centre of the dorsum there are a few reddish spots and short lines of the same colour (
<figureCitation id="19F9DEDBDD6BFF83FCF4FAF9FCB6FA84" box="[822,885,1371,1397]" captionStart="FIG" captionStartId="2.[114,125,758,775]" captionTargetBox="[120,1207,205,709]" captionTargetId="figure-509@2.[111,1218,196,722]" captionTargetPageId="2" captionText="FIG. 1. — Okenia cf. amoenula (MNHN), dorsal view of the living animal. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5398989" httpUri="https://zenodo.org/record/5398989/files/figure.png" pageId="1" pageNumber="16">Fig. 1</figureCitation>
). The oral tentacles are white and have four yellow tips each. The dorsum has, on each lateral margin, ten short papillae with yellow pigmentation on the apex. All over the dorsum there is a network of spicules. The rhinophores are white, non-retractile, and ringed with 18 lamellae. There are 11 unipinnate (the largest one is bipinnate) branchial leaves white in colour, having the rachis yellow (in the distal region edged with two red lines). The tail has a yellow line in the centre with few red spots.
</paragraph>
</subSubSection>
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<paragraph id="817DC25EDD68FF80FFB0FD54FC95FCF6" blockId="2.[114,854,758,775]" box="[114,854,758,775]" pageId="2" pageNumber="17">
FIG. 1. —
<taxonomicName id="46C2B9DDDD68FF80FF03FD54FE75FCF6" authority="(MNHN)" baseAuthorityName="MNHN" box="[193,438,758,775]" class="Gastropoda" family="Goniodorididae" genus="Okenia" isUncertain="true" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FF03FD54FF39FCF6" box="[193,250,758,775]" italics="true" pageId="2" pageNumber="17">Okenia</emphasis>
cf.
<emphasis id="B3B61E4CDD68FF80FED8FD54FEA8FCF6" box="[282,363,758,775]" italics="true" pageId="2" pageNumber="17">amoenula</emphasis>
(MNHN)
</taxonomicName>
, dorsal view of the living animal. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="C9D891D5DD68FF80FFB0FCD2FCF5FB6E" pageId="2" pageNumber="17" type="discussion">
<paragraph id="817DC25EDD68FF80FFB0FCD2FF15FC79" blockId="2.[114,648,880,1704]" box="[114,214,880,906]" pageId="2" pageNumber="17">REMARKS</paragraph>
<paragraph id="817DC25EDD68FF80FFB0FC2DFEFDFBB2" blockId="2.[114,648,880,1704]" pageId="2" pageNumber="17">The single specimen collected was badly preserved, making dissection and investigation of the internal anatomy impossible. In the process of removing the foregut the radula was lost. For that reason a positive identification of the specimen could not be done.</paragraph>
<paragraph id="817DC25EDD68FF80FFB0FBE5FDAFF959" blockId="2.[114,648,880,1704]" pageId="2" pageNumber="17">
<bibRefCitation id="E553BFAFDD68FF80FFB0FBE5FEC1FB90" author="Bergh R." bookContentInfo="144 p." box="[114,258,1095,1121]" journalOrPublisher="Transactions of the South African Philosophical Society" pageId="2" pageNumber="17" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh (1907)</bibRefCitation>
described
<taxonomicName id="46C2B9DDDD68FF80FEBAFBE5FF70FB71" authority="(Bergh, 1907)" baseAuthorityName="Bergh" baseAuthorityYear="1907" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FEBAFBE5FDF3FB90" box="[376,560,1095,1121]" italics="true" pageId="2" pageNumber="17">Okenia amoenula</emphasis>
(
<bibRefCitation id="E553BFAFDD68FF80FDFDFBE5FF68FB71" author="Bergh R." bookContentInfo="144 p." journalOrPublisher="Transactions of the South African Philosophical Society" pageId="2" pageNumber="17" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh, 1907</bibRefCitation>
)
</taxonomicName>
in the binomen
<taxonomicName id="46C2B9DDDD68FF80FEB1FBC4FF6FFB6E" authority="Bergh, 1907" authorityName="Bergh" authorityYear="1907" class="Gastropoda" family="Goniodorididae" genus="Idaliella" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FEB1FBC4FDFBFB71" box="[371,568,1126,1152]" italics="true" pageId="2" pageNumber="17">Idaliella amoenula</emphasis>
<bibRefCitation id="E553BFAFDD68FF80FD83FBC4FF6FFB6E" author="Bergh R." bookContentInfo="144 p." journalOrPublisher="Transactions of the South African Philosophical Society" pageId="2" pageNumber="17" part="17" refId="ref8210" refString="Bergh R. 1907. - The Opisthobranchiata of South Africa. Transactions of the South African Philosophical Society 17: 144 p., 14 pls." title="The Opisthobranchiata of South Africa" type="book" year="1907">Bergh, 1907</bibRefCitation>
</taxonomicName>
, as being a white animal with three crimson lines on the dorsum, which joined together in the anterior part, and a crimson patch on the anal area. The dorsal papillae were reddish and yellow, the branchial leaves had a crimson colour, and the rhinophores were yellow. Our specimen, as well as Berghs, has a white background colour, red and yellow pigmentation on the lateral papillae, and the branchial leaves are also reddish (with two red lines). However, our specimen shows a few differences in colour pattern from Berghs material (white rhinophores instead of yellow, white branchial leaves instead of crimson), and with that described by
<bibRefCitation id="E553BFAFDD68FF80FEC3F9B1FE7AF9DC" author="Gosliner T. M." bookContentInfo="Sea Challengers, Monterey, 136 p." box="[257,441,1555,1581]" pageId="2" pageNumber="17" refId="ref8424" refString="Gosliner T. M. 1987 a. - Nudibranchs of Southern Africa. A Guide to Opisthobranch Mollusks of Southern Africa. Sea Challengers, Monterey, 136 p." title="Nudibranchs of Southern Africa. A Guide to Opisthobranch Mollusks of Southern Africa" type="book" year="1987">Gosliner (1987a)</bibRefCitation>
from
<collectingCountry id="F9D582CEDD68FF80FE3CF9B1FD44F9DC" box="[510,647,1555,1581]" name="South Africa" pageId="2" pageNumber="17">South Africa</collectingCountry>
(dorsum with more red pigmentation in Gosliners specimen and yellow lateral papillae, but also with red spots), which could be caused by differences in size. Otherwise they are identical.
</paragraph>
<paragraph id="817DC25EDD68FF80FD67FCD2FCF5FB6E" blockId="2.[677,1212,880,1704]" pageId="2" pageNumber="17">
<taxonomicName id="46C2B9DDDD68FF80FD67FCD2FC43FC7A" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[677,896,880,907]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FD67FCD2FC43FC7A" box="[677,896,880,907]" italics="true" pageId="2" pageNumber="17">Okenia amoenula</emphasis>
</taxonomicName>
has been considered by
<bibRefCitation id="E553BFAFDD68FF80FD67FC2DFCBEFC58" author="Schmekel L." box="[677,893,911,937]" journalOrPublisher="Veliger" pageId="2" pageNumber="17" pagination="355 - 360" part="21" refId="ref8798" refString="Schmekel L. 1979. - First record of Okenia impexa Marcus, 1957 from the Western Atlantic in the Mediterranean. Veliger 21: 355 - 360." title="First record of Okenia impexa Marcus, 1957 from the Western Atlantic in the Mediterranean" type="journal article" year="1979">Schmekel (1979)</bibRefCitation>
as a synonym of
<taxonomicName id="46C2B9DDDD68FF80FBA0FC2DFC35FC39" authority="(Ihering, 1886)" baseAuthorityName="Ihering" baseAuthorityYear="1886" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="mediterranea">
<emphasis id="B3B61E4CDD68FF80FBA0FC2DFCF6FC39" italics="true" pageId="2" pageNumber="17">Okenia mediterranea</emphasis>
(Ihering, 1886)
</taxonomicName>
.
<taxonomicName id="46C2B9DDDD68FF80FBC1FC0CFD22FC14" baseAuthorityName="Ihering" baseAuthorityYear="1886" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="mediterranea">
<emphasis id="B3B61E4CDD68FF80FBC1FC0CFD22FC14" italics="true" pageId="2" pageNumber="17">Okenia mediterranea</emphasis>
</taxonomicName>
, like our material of
<taxonomicName id="46C2B9DDDD68FF80FC1CFC6EFBADFC16" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[990,1134,972,999]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FC1CFC6EFBADFC16" box="[990,1134,972,999]" italics="true" pageId="2" pageNumber="17">O. amoenula</emphasis>
</taxonomicName>
, has a yellow line in the centre of the tail with some red spots, and the rhinophores are white. However, our specimen lacks tubercles in the centre of the dorsum, which is a definitive character that separates the two species (
<bibRefCitation id="E553BFAFDD68FF80FB8FFBC4FCEAFB6E" author="Valdes A. &amp; Ortea J." journalOrPublisher="Veliger" pageId="2" pageNumber="17" pagination="223 - 234" part="38" refId="ref8858" refString="Valdes A. &amp; Ortea J. 1995. - Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species. Veliger 38: 223 - 234." title="Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species" type="journal article" year="1995">Valdés &amp; Ortea 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C9D891D5DD68FF80FD67FB01FC9AF959" pageId="2" pageNumber="17" type="description">
<paragraph id="817DC25EDD68FF80FD67FB01FC9AF959" blockId="2.[677,1212,880,1704]" pageId="2" pageNumber="17">
<taxonomicName id="46C2B9DDDD68FF80FD67FB01FCA7FB4C" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[677,868,1187,1213]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FD67FB01FCA7FB4C" box="[677,868,1187,1213]" italics="true" pageId="2" pageNumber="17">Okenia amoenula</emphasis>
</taxonomicName>
is similar in colour to
<taxonomicName id="46C2B9DDDD68FF80FBAFFB01FBDFFB2D" authority="Valdes &amp; Ortea, 1995" authorityName="Valdes &amp; Ortea" authorityYear="1995" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="hispanica">
<emphasis id="B3B61E4CDD68FF80FBAFFB01FCCEFB2D" italics="true" pageId="2" pageNumber="17">Okenia hispanica</emphasis>
<bibRefCitation id="E553BFAFDD68FF80FCD4FB60FBDFFB2D" author="Valdes A. &amp; Ortea J." box="[790,1052,1218,1244]" journalOrPublisher="Veliger" pageId="2" pageNumber="17" pagination="223 - 234" part="38" refId="ref8858" refString="Valdes A. &amp; Ortea J. 1995. - Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species. Veliger 38: 223 - 234." title="Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species" type="journal article" year="1995">Valdés &amp; Ortea, 1995</bibRefCitation>
</taxonomicName>
. Both species have a white background colour, a white tail with a yellow line, and yellow pigmentation on the gill. However, they are clearly distinguishable by several external features. The dorsum of
<taxonomicName id="46C2B9DDDD68FF80FBB2FA9EFD2EFA85" authorityName="Valdes &amp; Ortea" authorityYear="1995" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="hispanica">
<emphasis id="B3B61E4CDD68FF80FBB2FA9EFD2EFA85" italics="true" pageId="2" pageNumber="17">O. hispanica</emphasis>
</taxonomicName>
has two large pink patches, just behind each rhinophore, and another one on the anal area, the rhinophores are pink, and the oral tentacles are white, whereas in
<taxonomicName id="46C2B9DDDD68FF80FC0EFA14FB9BFA20" baseAuthorityName="Bergh" baseAuthorityYear="1907" box="[972,1112,1462,1489]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="amoenula">
<emphasis id="B3B61E4CDD68FF80FC0EFA14FB9BFA20" box="[972,1112,1462,1489]" italics="true" pageId="2" pageNumber="17">O. amoenula</emphasis>
</taxonomicName>
the dorsum is white with red spots or lines (lacking large patches), the rhinophores are white or yellow and the oral tentacles are white with four small yellow spots. Other differences include the absence of tubercles on the dorsum of
<taxonomicName id="46C2B9DDDD68FF80FC18F9F2FB9CF99A" authorityName="Valdes &amp; Ortea" authorityYear="1995" box="[986,1119,1616,1643]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="17" phylum="Mollusca" rank="species" species="hispanica">
<emphasis id="B3B61E4CDD68FF80FC18F9F2FB9CF99A" box="[986,1119,1616,1643]" italics="true" pageId="2" pageNumber="17">O. hispanica</emphasis>
</taxonomicName>
and the radular morphology, already compared by
<bibRefCitation id="E553BFAFDD68FF80FBB7F9CDFC96F959" author="Valdes A. &amp; Ortea J." journalOrPublisher="Veliger" pageId="2" pageNumber="17" pagination="223 - 234" part="38" refId="ref8858" refString="Valdes A. &amp; Ortea J. 1995. - Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species. Veliger 38: 223 - 234." title="Revised taxonomy of some species of the genus Okenia Menke, 1830 (Mollusca: Nudibranchia) from the Atlantic Ocean, with the description of a new species" type="journal article" year="1995">Valdés &amp; Ortea (1995)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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Genus
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Menke, 1830
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Leuckart, 1828
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Genus
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Müller, 1781
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Müller, 1776
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<paragraph id="817DC25EDD6FFF87FFB3FDD8FF16FD63" blockId="5.[113,649,634,1704]" box="[113,213,634,660]" pageId="5" pageNumber="20">REMARKS</paragraph>
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<bibRefCitation id="E553BFAFDD6FFF87FFB3FD38FE88FD45" author="MacFarland F. M." box="[113,331,666,692]" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="5" pageNumber="20" pagination="35 - 34" part="18" refId="ref8483" refString="MacFarland F. M. 1905. - A preliminary account of the Dorididae of Monterey Bay, California. Proceedings of the Biological Society of Washington 18: 35 - 34." title="A preliminary account of the Dorididae of Monterey Bay, California" type="journal volume" year="1905">MacFarland (1905)</bibRefCitation>
introduced the genus
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<emphasis id="B3B61E4CDD6FFF87FD8CFD38FD45FD45" box="[590,646,666,692]" italics="true" pageId="5" pageNumber="20">Laila</emphasis>
<bibRefCitation id="E553BFAFDD6FFF87FFB3FD1BFE91FD22" author="MacFarland F. M." box="[113,338,697,723]" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="5" pageNumber="20" pagination="35 - 34" part="18" refId="ref8483" refString="MacFarland F. M. 1905. - A preliminary account of the Dorididae of Monterey Bay, California. Proceedings of the Biological Society of Washington 18: 35 - 34." title="A preliminary account of the Dorididae of Monterey Bay, California" type="journal volume" year="1905">MacFarland, 1905</bibRefCitation>
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, based on
<taxonomicName id="46C2B9DDDD6FFF87FE1AFD1BFE87FD00" authority="MacFarland, 1905" authorityName="MacFarland" authorityYear="1905" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="species" species="cockerelli">
<emphasis id="B3B61E4CDD6FFF87FE1AFD1BFD4AFD22" box="[472,649,697,723]" italics="true" pageId="5" pageNumber="20">Laila cockerelli</emphasis>
<bibRefCitation id="E553BFAFDD6FFF87FFB3FD75FE87FD00" author="MacFarland F. M." box="[113,324,727,753]" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="5" pageNumber="20" pagination="35 - 34" part="18" refId="ref8483" refString="MacFarland F. M. 1905. - A preliminary account of the Dorididae of Monterey Bay, California. Proceedings of the Biological Society of Washington 18: 35 - 34." title="A preliminary account of the Dorididae of Monterey Bay, California" type="journal volume" year="1905">MacFarland, 1905</bibRefCitation>
</taxonomicName>
, as an “allied” to the genera
<taxonomicName id="46C2B9DDDD6FFF87FFB3FD54FE40FCE1" authority="Johnston, 1838" authorityName="Johnston" authorityYear="1838" box="[113,387,758,784]" class="Gastropoda" family="Polyceridae" genus="Triopha" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FFB3FD54FF04FCE1" box="[113,199,758,784]" italics="true" pageId="5" pageNumber="20">Triopha</emphasis>
Johnston, 1838
</taxonomicName>
and
<taxonomicName id="46C2B9DDDD6FFF87FE00FD57FD41FCE1" authority="Bergh, 1880" authorityName="Bergh" authorityYear="1880" box="[450,642,757,784]" class="Gastropoda" family="Polyceridae" genus="Issa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FE00FD57FE2BFCFF" box="[450,488,757,782]" italics="true" pageId="5" pageNumber="20">Issa</emphasis>
Bergh, 1880
</taxonomicName>
, but he did not refer to the genus
<taxonomicName id="46C2B9DDDD6FFF87FE19FCB6FF6DFCBC" authority="Muller, 1781" authorityName="Muller" authorityYear="1781" class="Gastropoda" family="Polyceridae" genus="Limacia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FE19FCB6FDF3FCDC" box="[475,560,788,813]" italics="true" pageId="5" pageNumber="20">Limacia</emphasis>
Müller, 1781
</taxonomicName>
. The description of
<taxonomicName id="46C2B9DDDD6FFF87FE59FC91FDDFFCBC" authorityName="MacFarland" authorityYear="1905" box="[411,540,819,845]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="species" species="cockerelli">
<emphasis id="B3B61E4CDD6FFF87FE59FC91FDDFFCBC" box="[411,540,819,845]" italics="true" pageId="5" pageNumber="20">L. cockerelli</emphasis>
</taxonomicName>
was later completed with drawings of the radula and the animal in the living form (
<bibRefCitation id="E553BFAFDD6FFF87FE6AFCD3FDB9FC7A" author="MacFarland F. M." bookContentInfo="546 p." box="[424,634,881,907]" journalOrPublisher="Memoirs of the California Academy of Sciences" pageId="5" pageNumber="20" part="6" refId="ref8516" refString="MacFarland F. M. 1966. - Studies of opisthobranchiate mollusks of the Pacific coast of North America. Memoirs of the California Academy of Sciences 6: 546 p., 72 pls." title="Studies of opisthobranchiate mollusks of the Pacific coast of North America" type="book" year="1966">MacFarland 1966</bibRefCitation>
).
<taxonomicName id="46C2B9DDDD6FFF87FFB3FC2DFDDDFC58" authority="Bertsch &amp; Ferreira, 1974" authorityName="Bertsch &amp; Ferreira" authorityYear="1974" box="[113,542,911,937]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="species" species="janssi">
<emphasis id="B3B61E4CDD6FFF87FFB3FC2DFF2CFC59" box="[113,239,911,937]" italics="true" pageId="5" pageNumber="20">Laila janssi</emphasis>
<bibRefCitation id="E553BFAFDD6FFF87FF38FC2DFDDDFC58" author="Bertsch H. &amp; Ferreira A. J." box="[250,542,911,937]" journalOrPublisher="Veliger" pageId="5" pageNumber="20" pagination="343 - 353" part="16" refId="ref8238" refString="Bertsch H. &amp; Ferreira A. J. 1974. - Four new species of nudibranchs from tropical west America. Veliger 16: 343 - 353." title="Four new species of nudibranchs from tropical west America" type="journal article" year="1974">Bertsch &amp; Ferreira, 1974</bibRefCitation>
</taxonomicName>
, presents similar morphological characteristics as
<taxonomicName id="46C2B9DDDD6FFF87FDF1FC0FFF60FC16" authorityName="MacFarland" authorityYear="1905" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="species" species="cockerelli">
<emphasis id="B3B61E4CDD6FFF87FDF1FC0FFF60FC16" italics="true" pageId="5" pageNumber="20">L. cockerelli</emphasis>
</taxonomicName>
, differing on the colour pattern and in some radular features such as a smaller number of teeth rows, a rougher rachidian plate texture and some variations on the shape of the outer lateral teeth (
<bibRefCitation id="E553BFAFDD6FFF87FFBAFBE5FE46FB90" author="Bertsch H. &amp; Ferreira A. J." box="[120,389,1095,1121]" journalOrPublisher="Veliger" pageId="5" pageNumber="20" pagination="343 - 353" part="16" refId="ref8238" refString="Bertsch H. &amp; Ferreira A. J. 1974. - Four new species of nudibranchs from tropical west America. Veliger 16: 343 - 353." title="Four new species of nudibranchs from tropical west America" type="journal article" year="1974">Bertsch &amp; Ferreira 1974</bibRefCitation>
).
</paragraph>
<paragraph id="817DC25EDD6FFF87FFB3FBC4FD86F959" blockId="5.[113,649,634,1704]" pageId="5" pageNumber="20">
The genus
<taxonomicName id="46C2B9DDDD6FFF87FF26FBC4FED8FB71" authorityName="MacFarland" authorityYear="1905" box="[228,283,1126,1152]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FF26FBC4FED8FB71" box="[228,283,1126,1152]" italics="true" pageId="5" pageNumber="20">Laila</emphasis>
</taxonomicName>
has the same morphological characteristics as the genus
<taxonomicName id="46C2B9DDDD6FFF87FEB9FB26FE15FB6C" authorityName="Muller" authorityYear="1781" box="[379,470,1156,1181]" class="Gastropoda" family="Polyceridae" genus="Limacia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FEB9FB26FE15FB6C" box="[379,470,1156,1181]" italics="true" pageId="5" pageNumber="20">Limacia</emphasis>
</taxonomicName>
: pallial margin with numerous club-shaped papillae; presence of tubercles on the notum; veil appendages canaliculated; retractile rhinophores; branchial leaves non retractile; two inner lateral teeth hook-shaped, the innermost being thinner and longer than the outermost, and many outer lateral teeth quadrangularshaped.
<taxonomicName id="46C2B9DDDD6FFF87FF0FFAF9FEC5FA84" authorityName="MacFarland" authorityYear="1905" box="[205,262,1371,1397]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FF0FFAF9FEC5FA84" box="[205,262,1371,1397]" italics="true" pageId="5" pageNumber="20">Laila</emphasis>
</taxonomicName>
and
<taxonomicName id="46C2B9DDDD6FFF87FEFFFAF9FE56FA85" authorityName="Muller" authorityYear="1781" box="[317,405,1371,1396]" class="Gastropoda" family="Polyceridae" genus="Limacia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FEFFFAF9FE56FA85" box="[317,405,1371,1396]" italics="true" pageId="5" pageNumber="20">Limacia</emphasis>
</taxonomicName>
only differ in the fact that
<taxonomicName id="46C2B9DDDD6FFF87FF6EFADBFDE9FA65" authority="Muller, 1776" authorityName="Muller" authorityYear="1776" box="[172,554,1401,1428]" class="Gastropoda" family="Polyceridae" genus="Limacia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="species" species="clavigera">
<emphasis id="B3B61E4CDD6FFF87FF6EFADBFEBDFA65" box="[172,382,1401,1428]" italics="true" pageId="5" pageNumber="20">Limacia clavigera</emphasis>
Müller, 1776
</taxonomicName>
, lacks a rachidian plate in the radula, which is present in both
<taxonomicName id="46C2B9DDDD6FFF87FF6FFA15FEE8FA20" authorityName="MacFarland" authorityYear="1905" box="[173,299,1463,1489]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="species" species="cockerelli">
<emphasis id="B3B61E4CDD6FFF87FF6FFA15FEE8FA20" box="[173,299,1463,1489]" italics="true" pageId="5" pageNumber="20">L. cockerelli</emphasis>
</taxonomicName>
and
<taxonomicName id="46C2B9DDDD6FFF87FEA3FA15FE7AFA21" authorityName="Bertsch &amp; Ferreira" authorityYear="1974" box="[353,441,1463,1488]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="species" species="janssi">
<emphasis id="B3B61E4CDD6FFF87FEA3FA15FE7AFA21" box="[353,441,1463,1488]" italics="true" pageId="5" pageNumber="20">L. janssi</emphasis>
</taxonomicName>
. Since most of the morphological features of
<taxonomicName id="46C2B9DDDD6FFF87FE54FA74FE0CFA01" authorityName="MacFarland" authorityYear="1905" box="[406,463,1494,1520]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FE54FA74FE0CFA01" box="[406,463,1494,1520]" italics="true" pageId="5" pageNumber="20">Laila</emphasis>
</taxonomicName>
and
<taxonomicName id="46C2B9DDDD6FFF87FDC5FA77FDA3FA1F" authorityName="Muller" authorityYear="1781" box="[519,608,1493,1518]" class="Gastropoda" family="Polyceridae" genus="Limacia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FDC5FA77FDA3FA1F" box="[519,608,1493,1518]" italics="true" pageId="5" pageNumber="20">Limacia</emphasis>
</taxonomicName>
are very similar, just the presence of rachidian plate in the radula of
<taxonomicName id="46C2B9DDDD6FFF87FEC2F9B1FEF4F9DC" authorityName="MacFarland" authorityYear="1905" box="[256,311,1555,1581]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FEC2F9B1FEF4F9DC" box="[256,311,1555,1581]" italics="true" pageId="5" pageNumber="20">Laila</emphasis>
</taxonomicName>
does not seem to be enough to consider
<taxonomicName id="46C2B9DDDD6FFF87FF13F990FECBF9BD" authorityName="MacFarland" authorityYear="1905" box="[209,264,1586,1612]" class="Gastropoda" family="Polyceridae" genus="Laila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FF13F990FECBF9BD" box="[209,264,1586,1612]" italics="true" pageId="5" pageNumber="20">Laila</emphasis>
</taxonomicName>
as a different genus. Therefore both genera are here regarded as synonyms, with
<taxonomicName id="46C2B9DDDD6FFF87FFB3F9CDFEA6F978" authority="Muller, 1781" authorityName="Muller" authorityYear="1781" box="[113,357,1647,1673]" class="Gastropoda" family="Polyceridae" genus="Limacia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="B3B61E4CDD6FFF87FFB3F9CDFF09F979" box="[113,202,1647,1672]" italics="true" pageId="5" pageNumber="20">Limacia</emphasis>
Müller, 1781
</taxonomicName>
having priority. This idea was already proposed by
<bibRefCitation id="E553BFAFDD6FFF87FEBAF92CFD83F959" author="Ortea J. &amp; Quero A. &amp; Rodriguez G. &amp; Valdes A." box="[376,576,1677,1704]" journalOrPublisher="Revista de Biologia de la Universidad de Oviedo" pageId="5" pageNumber="20" pagination="99 - 107" part="7" refId="ref8690" refString="Ortea J., Quero A., Rodriguez G. &amp; Valdes A. 1989. - Estudio de Limacia clavigera (Muller, 1776) (Mollusca: Nudibranchia), con nota sobre su distribuccion geografica y la validez del genero Laila MacFarland, 1905. Revista de Biologia de la Universidad de Oviedo 7: 99 - 107." title="Estudio de Limacia clavigera (Muller, 1776) (Mollusca: Nudibranchia), con nota sobre su distribuccion geografica y la validez del genero Laila MacFarland, 1905" type="journal article" year="1989">
Ortea
<emphasis id="B3B61E4CDD6FFF87FE79F92FFE2EF959" box="[443,493,1677,1704]" italics="true" pageId="5" pageNumber="20">et al.</emphasis>
(1989)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>