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<mods:title id="D0185523E44FD31E3C8D419F84707204">Contributions of a small collection of terrestrial microsnails (Pupilloidea, Hypselostomatidae) from Myanmar with description of three new species</mods:title>
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Genus
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<taxonomicName id="ECB777C2D5D06BEA6CF671A304F632E2" authority=")" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bifidaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bifidaria (Bensonella)" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="subGenus" subGenus="Bensonella">Bifidaria (Bensonella)</taxonomicName>
Pilsbry &amp; Vanatta, 1900: 591.
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<taxonomicName id="FF4364985F3B6A3ADD3029F244790352" authority=")" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia (Bensonella)" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="subGenus" subGenus="Bensonella">Boysidia (Bensonella)</taxonomicName>
.
<bibRefCitation id="80D8D2CEC5977E339FBD184F55431EC4" author="Pilsbry, HA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="157" refId="B74" refString="Pilsbry, HA, 1917 1918. [1916-1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113-256, pls. 14-38 (1917); pp. 257-380, pls 39-49 (1918)]" title="[1916 - 1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113 - 256, pls. 14 - 38 (1917); pp. 257 - 380, pls 39 - 49 (1918)]" year="1917">Pilsbry 1917</bibRefCitation>
: 198.
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<taxonomicName id="424A935F1FC8F7799DD97D9E55AB0707" authority=")" authorityName="Pilsbry" authorityYear="1917" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia (Paraboysidia)" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="subGenus" subGenus="Paraboysidia">Boysidia (Paraboysidia)</taxonomicName>
Pilsbry, 1917: 174, 201.
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<paragraph id="153224D3614380559CD448AD5FC4D23F" pageId="0" pageNumber="157">Type species.</paragraph>
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<emphasis id="FC5DCA5DD334D3CF3920863BE9672A42" italics="true" pageId="0" pageNumber="157">Pupa plicidens</emphasis>
Benson, 1849, by original designation.
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<paragraph id="FFABB11C4FF8BCFCC2F3D19324FBC837" pageId="0" pageNumber="157">Remarks.</paragraph>
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The generic status and diagnostic morphological characters of the type species were recently revised (
<bibRefCitation id="F7E4DEC50FA42015D1FE202753B5B8A8" DOI="https://doi.org/10.1080/00222933.2022.2152750" author="Pall-Gergely, B" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="157" pagination="2011 - 2029" refId="B49" refString="Pall-Gergely, B, White, TS, 2023. Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45-48): 2011 - 2029, DOI: https://doi.org/10.1080/00222933.2022.2152750" title="Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)." url="https://doi.org/10.1080/00222933.2022.2152750" volume="56" year="2023">
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and White 2023
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). The type species was believed to possess
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teeth, i.e., apertural barriers forming hooks that point outside of the aperture (
<bibRefCitation id="54DDC43E5042BD6A6497739D75E1840B" author="Pilsbry, HA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="157" refId="B74" refString="Pilsbry, HA, 1917 1918. [1916-1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113-256, pls. 14-38 (1917); pp. 257-380, pls 39-49 (1918)]" title="[1916 - 1918] Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Series 2, Pulmonata (Vol. 24): Pupillidae (Gastrocoptinae). Academy of Natural Sciences of Philadelphia 380 pp. [pp. 113 - 256, pls. 14 - 38 (1917); pp. 257 - 380, pls 39 - 49 (1918)]" year="1917">Pilsbry 1917</bibRefCitation>
). Later, when specimens without hooked apertural barriers were found in collections, normal and hooked teeth were hypothesised as intraspecific variability (
<bibRefCitation id="E292DF1C9FF3905B9172770F38EC8ED9" author="Budha, PB" editor="Budha, PB" journalOrPublisher="D. dissertation. Evolutionary Ecology Group, University of Antwerp, Antwerp" pageId="0" pageNumber="157" pagination="189 - 201" refId="B12" refString="Budha, PB, Backeljau, T, 2017. First report of Paraboysidia landourensis (Pilsbry, 1915) from Nepal, with description of a putative new species and a note on Bensonella plicidens (Benson, 1849) (Stylommatophora: family unclear). In: Budha, PB, Ed., Taxonomy of terrestrial molluscs of Nepal Ph. D. dissertation. Evolutionary Ecology Group, University of Antwerp, Antwerp: 189 - 201" title="First report of Paraboysidia landourensis (Pilsbry, 1915) from Nepal, with description of a putative new species and a note on Bensonella plicidens (Benson, 1849) (Stylommatophora: family unclear)." volumeTitle="Taxonomy of terrestrial molluscs of Nepal Ph." year="2017">Budha and Backeljau 2017</bibRefCitation>
).
<bibRefCitation id="777EF0755A2FCA413B862CA875B6C5DB" DOI="https://doi.org/10.1080/00222933.2022.2152750" author="Pall-Gergely, B" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="157" pagination="2011 - 2029" refId="B49" refString="Pall-Gergely, B, White, TS, 2023. Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45-48): 2011 - 2029, DOI: https://doi.org/10.1080/00222933.2022.2152750" title="Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)." url="https://doi.org/10.1080/00222933.2022.2152750" volume="56" year="2023">
<normalizedToken id="3768F947E8BAC1D49AEBA3243BD74412" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
and White (2023)
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showed that the
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<taxonomicName id="1A8719670EA31B26DEAA8563E1C3C227" baseAuthorityName="Benson" baseAuthorityYear="1849" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella plicidens" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="plicidens">
<emphasis id="F33AE3C5C48E6D76034E6F33AA4F24A9" italics="true" pageId="0" pageNumber="157">Bensonella plicidens</emphasis>
</taxonomicName>
was a species with normal (not hooked) barriers, while the Himalayan species with hooked teeth was an undescribed species (
<taxonomicName id="CE8034C2CDEB1ACFF617680EEC3ACB44" authorityName="Páll-Gergely" authorityYear="2023" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella hooki" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="hooki">
<emphasis id="27B52B1ACE36768943081AD391463228" italics="true" pageId="0" pageNumber="157">Bensonella hooki</emphasis>
</taxonomicName>
<normalizedToken id="77413A24F356F9E6026732B112434D04" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
, 2023). Since the
<normalizedToken id="DD67FC225E4B01FAC63134C923171104" originalValue="hooked">'hooked'</normalizedToken>
and 'not
<normalizedToken id="10865AAC5417AF2529C5C3F2CD60DB18" originalValue="hooked">hooked'</normalizedToken>
species are very similar in other shell characters, this trait could not be used as a diagnostic character for recognising
<taxonomicName id="B7E08D783372E7D6CFC06B2968CF131A" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="3E40691553DC9D240DCD65367094BDA7" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
</taxonomicName>
. Rather, this genus is diagnosed by a brownish shell with the last whorl attached to the penultimate whorl, and the presence of separate angular and parietal lamellae.
</paragraph>
<paragraph id="AF66B0CC494BDBAD78BA0F1CC91A7C78" pageId="0" pageNumber="157">
Since the type species of
<taxonomicName id="6AE6F366618B786CF4BE227EF439EF29" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="A9FDA42AF4E2E13F111E41EF09E5FF48" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
</taxonomicName>
and
<taxonomicName id="57DD5584D2FBF0D7FA99ADAFBB6676D1" authorityName="Pilsbry" authorityYear="1917" class="Gastropoda" family="Hypselostomatidae" genus="Paraboysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraboysidia" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="040886A064C71B9166A3DD293C471EB3" italics="true" pageId="0" pageNumber="157">Paraboysidia</emphasis>
</taxonomicName>
(
<taxonomicName id="55251131082199BDA4183039CA98E1A7" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" class="Gastropoda" family="Hypselostomatidae" genus="Boysidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boysidia paviei" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="paviei">
<emphasis id="B6A64069C24110024F7794F5D8769E14" italics="true" pageId="0" pageNumber="157">Boysidia paviei</emphasis>
</taxonomicName>
Bavay &amp; Dautzenberg, 1912) are very similar, the latter has been treated as a junior synonym of the former (
<bibRefCitation id="CB8B77784550F6DBC774DE1A42AEF565" DOI="https://doi.org/10.12657/folmal.029.009" author="Gittenberger, E" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="157" pagination="69 - 90" refId="B22" refString="Gittenberger, E, Gyeltshen, C, Leda, P, Sherub, S, 2021. The superfamilies Pupilloidea and Enoidea (Gastropoda, Eupulmonata) in Bhutan. Folia Malacologica 29 (2): 69 - 90, DOI: https://doi.org/10.12657/folmal.029.009" title="The superfamilies Pupilloidea and Enoidea (Gastropoda, Eupulmonata) in Bhutan." url="https://doi.org/10.12657/folmal.029.009" volume="29" year="2021">Gittenberger et al. 2021</bibRefCitation>
;
<bibRefCitation id="9D7B09D92233DF4FB344D020A612AADC" DOI="https://doi.org/10.1080/00222933.2022.2152750" author="Pall-Gergely, B" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="157" pagination="2011 - 2029" refId="B49" refString="Pall-Gergely, B, White, TS, 2023. Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45-48): 2011 - 2029, DOI: https://doi.org/10.1080/00222933.2022.2152750" title="Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)." url="https://doi.org/10.1080/00222933.2022.2152750" volume="56" year="2023">
<normalizedToken id="6AD70EF93D05A42C26F29C9F007E6653" originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
and White 2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,113 @@
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.129.34988" ID-GBIF-Dataset="7a3be85e-9a8f-4749-9984-1e60c69e67d0" ID-PMC="PMC6717120" ID-Pensoft-Pub="1314-2003-129-1" ID-Pensoft-UUID="BF57C6B3C3065AEE9B4B3D47189C908F" ID-PubMed="31523157" ID-Zenodo-Dep="3382366" ModsDocID="1314-2003-129-1" checkinTime="1566623483646" checkinUser="pensoft" docAuthor="Paton, Alan J., Mwanyambo, Montfort, Govaerts, Rafael H. A., Smitha, Kokkaraniyil, Suddee, Somran, Phillipson, Peter B., Wilson, Trevor C., Forster, Paul I. &amp; Culham, Alastair" docDate="2019" docId="F3D6F235C07A5022A211C89BEE1DDAC5" docLanguage="en" docName="PhytoKeys 129: 1-158" docOrigin="PhytoKeys 129" docSource="http://dx.doi.org/10.3897/phytokeys.129.34988" docTitle="Equilabium flaccidum Paton &amp; Mwanyambo &amp; Govaerts &amp; Smitha &amp; Suddee &amp; Phillipson &amp; Wilson &amp; Forster &amp; Culham 2019, comb. nov." docType="treatment" docVersion="6" id="BF57C6B3C3065AEE9B4B3D47189C908F" lastPageNumber="99" masterDocId="BF57C6B3C3065AEE9B4B3D47189C908F" masterDocTitle="Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera" masterLastPageNumber="158" masterPageNumber="1" pageNumber="99" updateTime="1668139430914" updateUser="ExternalLinkService">
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<mods:title>Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paton, Alan J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mwanyambo, Montfort</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Govaerts, Rafael H. A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smitha, Kokkaraniyil</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Suddee, Somran</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Phillipson, Peter B.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wilson, Trevor C.</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Forster, Paul I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Culham, Alastair</mods:namePart>
</mods:name>
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<treatment ID-GBIF-Taxon="159244288" LSID="urn:lsid:plazi:treatment:F3D6F235C07A5022A211C89BEE1DDAC5" httpUri="http://treatment.plazi.org/id/F3D6F235C07A5022A211C89BEE1DDAC5" lastPageNumber="99" pageId="98" pageNumber="99">
<subSubSection pageId="98" pageNumber="99" type="nomenclature">
<paragraph pageId="98" pageNumber="99">
<taxonomicName LSID="F3D6F235-C07A-5022-A211-C89BEE1DDAC5" authority="(Vatke) Mwany. &amp; A. J. Paton" authorityName="Paton &amp; Mwanyambo &amp; Govaerts &amp; Smitha &amp; Suddee &amp; Phillipson &amp; Wilson &amp; Forster &amp; Culham" authorityYear="2019" baseAuthorityName="Vatke" class="Magnoliopsida" family="Lamiaceae" genus="Equilabium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Equilabium flaccidum" order="Lamiales" pageId="98" pageNumber="99" phylum="Tracheophyta" rank="species" species="flaccidum" status="comb. nov.">Equilabium flaccidum (Vatke) Mwany. &amp; A.J.Paton</taxonomicName>
<taxonomicNameLabel pageId="98" pageNumber="99">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="98" pageNumber="99" type="reference_group">
<paragraph pageId="98" pageNumber="99">
<taxonomicName authorityName="Vatke" authorityYear="1881" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coleus flaccidus" order="Lamiales" pageId="98" pageNumber="99" phylum="Tracheophyta" rank="species" species="flaccidus">Coleus flaccidus</taxonomicName>
Vatke in Linnaea 43: 9. 1881.
</paragraph>
<paragraph pageId="98" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Plectranthus flaccidus" order="Lamiales" pageId="98" pageNumber="99" phylum="Tracheophyta" rank="species" species="flaccidus">Plectranthus flaccidus</taxonomicName>
(Vatke)
<normalizedToken originalValue="Gürke">Guerke</normalizedToken>
, Bot. Jahrb. Syst. 19: 206. 1894. Type: Tanzania, Bagamoyo District: Ruvu [Kingani] R., Hildebrandt 1264 (holotype: B, destroyed; isotype: K).
</paragraph>
<paragraph pageId="98" pageNumber="99">
<taxonomicName authorityName="Baker" authorityYear="1900" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Plectranthus fragilis" order="Lamiales" pageId="98" pageNumber="99" phylum="Tracheophyta" rank="species" species="fragilis">Plectranthus fragilis</taxonomicName>
Baker in Fl. Trop. Afr. 5: 407. 1900. Type: Zanzibar, Kirk s.n. (holotype: K).
</paragraph>
</subSubSection>
<subSubSection pageId="98" pageNumber="99" type="distribution">
<paragraph pageId="98" pageNumber="99">Distribution.</paragraph>
<paragraph pageId="98" pageNumber="99">S. Somalia to Mozambique, Comoros.</paragraph>
</subSubSection>
</treatment>
</document>