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Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil + + + +Author + +Biganzoli-Rangel, Alejandro Jose +0000-0003-4426-3286 +Department of Biology, York University, Toronto, Ontario, Canada + + + +Author + +Carvalho, Gervásio Silva +0000-0002-3835-8644 +Escola de Ciências da Saúde e da Vida, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga, 6681, 90619 - 900, Porto Alegre, Rio Grande do Sul, Brasil (retired) + + + +Author + +Paladini, Andressa +0000-0001-8894-6092 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil & Departamento de Zoologia, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal do Paraná, Av. Cel. Francisco H. dos Santos, 100, Jardim das Américas, 81530 - 000, Curitiba, Paraná, Brasil + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +45 +73 + + + +journal article +10.3897/asp.83.e130043 +2000DF4D-D639-4E57-90EB-590A80CD9A6C + + + + +3.4. 9. + +Mahanarva +( +Ipiranga +) +bahiaensis +Carvalho & Webb, 2004 + + + + + +Figure 6 F + + + + + + + +Mahanarva +( +Ipiranga +) +bahiaensis + +Carvalho and Webb, 2004: 384 + + + + + + + + +Diagnosis. + +Tegmina brownish with two transverse bands yellowish; subgenital plate elongated, paramere with a triangular subapical spine and acutely rounded apex. + + + +Remarks. + + +The male +holotype +was colected in Camacã, +Bahia +, +Brazil +and is currently deposited on British Museum of Natural History ( + +BMNH + +). In this study, we examined +three specimens +from +Bahia +and the pattern of tegmina and male genitalia are very similar to + +M. +( +I. +) +rubicunda + +suggesting that + +M. +( +I. +) +bahiaensis + +can be a synonym of + +M. +( +I. +) +rubicunda + +. Since we didn’t have access to the +holotype +in person and more sampling from +Bahia +, specifically in the type locality, we chose to not synonymize both species. The diagnosis provided here was based on original description made by +Carvalho and Webb (2004) +. + + + + +Material examined. + + +BRASIL +, + +Bahia + +: Encruzilhada, +xi. 1972 +, Alvarenga & Seabra leg., +1 ♂ +, +2 ♀ +( + +MCTP + +). + + + + \ No newline at end of file diff --git a/data/3F/56/7B/3F567BC3D4F75B5A9A14B30FE831B94E.xml b/data/3F/56/7B/3F567BC3D4F75B5A9A14B30FE831B94E.xml new file mode 100644 index 00000000000..9377816d85b --- /dev/null +++ b/data/3F/56/7B/3F567BC3D4F75B5A9A14B30FE831B94E.xml @@ -0,0 +1,174 @@ + + + +Taxonomic review, morphometry, and phylogenetic analysis of Mahanarva (Ipiranga) Fennah, 1968 (Hemiptera: Auchenorrhyncha: Cercopidae) + + + +Author + +Meneghetti, Jéssica +0000-0001-6633-6728 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil + + + +Author + +Biganzoli-Rangel, Alejandro Jose +0000-0003-4426-3286 +Department of Biology, York University, Toronto, Ontario, Canada + + + +Author + +Carvalho, Gervásio Silva +0000-0002-3835-8644 +Escola de Ciências da Saúde e da Vida, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga, 6681, 90619 - 900, Porto Alegre, Rio Grande do Sul, Brasil (retired) + + + +Author + +Paladini, Andressa +0000-0001-8894-6092 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil & Departamento de Zoologia, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal do Paraná, Av. Cel. Francisco H. dos Santos, 100, Jardim das Américas, 81530 - 000, Curitiba, Paraná, Brasil + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +45 +73 + + + +journal article +10.3897/asp.83.e130043 +2000DF4D-D639-4E57-90EB-590A80CD9A6C + + + + +3.4. 10. + +Mahanarva +( +Ipiranga +) +takiyae +Paladini & Cavichioli, 2014 + + + + + +Figure 6 G + + + + + + + +Mahanarva +( +Ipiranga +) +takiyae + +Paladini and Cavichioli, 2014: 481 + + + + + + + + +Diagnosis. + +Tegmina black with four orange maculae; subgenital plate elongated with a spine at apex, paramere with a subapical hook-like spine. + + + +Remarks. + + +This species was recently described by +Paladini and Cavichioli (2014) +, presenting a complete and comprehensive description, as well as habitus illustrations and drawings of the male and female genitalia. The diagnosis presented here was based on the original description. + + + + +Material examined. + + +BRASIL +, + +Brasília + +: +Distrito Federal +, +ii. 1965 +J. A. P. Dutra leg., +paratypes +1 ♂ +, +1 ♀ +( + +DZUP + +); idem: FLONA de Brasília, Gleba 1, + +15 ° 45 ′ 00.0 ″ S +48 ° 04 ′ 00.0 ″ W + +, +1220 m +, +13. xi. 2012 +, D. Luz, +1 ♀ +( + +DZUP + +); idem: Fazenda Água Limpas, Mata de Galeria, +09–23. xi. 2017 +, Malaise, J. R. P Luz leg., +3 ♂ +, +2 ♀ +( + +DZUP + +); idem: Cerrado, +08–22. xii. 2017 +, Malaise, J. R. P Luz leg., +4 ♂ +( + +DZUP + +). + + + + \ No newline at end of file diff --git a/data/61/0A/0A/610A0A3BE5BD5A1680C9DDBCC77A9104.xml b/data/61/0A/0A/610A0A3BE5BD5A1680C9DDBCC77A9104.xml new file mode 100644 index 00000000000..34341f57fd0 --- /dev/null +++ b/data/61/0A/0A/610A0A3BE5BD5A1680C9DDBCC77A9104.xml @@ -0,0 +1,281 @@ + + + +Taxonomic review, morphometry, and phylogenetic analysis of Mahanarva (Ipiranga) Fennah, 1968 (Hemiptera: Auchenorrhyncha: Cercopidae) + + + +Author + +Meneghetti, Jéssica +0000-0001-6633-6728 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil + + + +Author + +Biganzoli-Rangel, Alejandro Jose +0000-0003-4426-3286 +Department of Biology, York University, Toronto, Ontario, Canada + + + +Author + +Carvalho, Gervásio Silva +0000-0002-3835-8644 +Escola de Ciências da Saúde e da Vida, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga, 6681, 90619 - 900, Porto Alegre, Rio Grande do Sul, Brasil (retired) + + + +Author + +Paladini, Andressa +0000-0001-8894-6092 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil & Departamento de Zoologia, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal do Paraná, Av. Cel. Francisco H. dos Santos, 100, Jardim das Américas, 81530 - 000, Curitiba, Paraná, Brasil + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +45 +73 + + + +journal article +10.3897/asp.83.e130043 +2000DF4D-D639-4E57-90EB-590A80CD9A6C + + + + +3.4. 7. + +Mahanarva +( +Ipiranga +) +aguirrei +( +Berg, 1879 +) + + + + + +Figures 6 D +, +10 A – I + + + + + + + +Tomaspis aguirrei + +Berg, 1879 +c: 216. + +Metcalf 1961: 85 + + + + + + + + + + +Mahanarva +( +Ipiranga +) +aguirrei + + +: + +Fennah 1968: 187 + + + + + + + + +Measurements male. + +Head length: 1.24; head width: 2.31; pronotum length: 2, 13; pronotum width: 3.43; tegmen length: 9; tegmen width: 2.7; total length: 11.47 (in mm). + + + +Diagnosis. + +Tegmina black or brownish with one thin transverse band whitish between basal and median third and an incomplete band between median and apical third; subgenital plate with excavation on the basal third, rounded apex, paramere apex subquadrangular, spine long and slender with an acute apex. + + + +Redescription. + + +Head black, vertex subretangular with a slightly marked median carina, ocelli closer to each other than to eyes, separated by about one diameter from each other; tylus subretangular, black, with a slightly marked carina on the basal third, becoming indistinct towards the apex (Fig. +10 A +); antenna black, pedicel sparsely setose; basal body of flagellum subcylindrical with a single and short arista shorter than the pedicel length. Postclypeus black, inflated, with convex profile, longitudinal carina present, lateral grooves slightly marked (Fig. +10 B, C +); rostrum with second segment centrally yellowish and laterally black and third segment black, reaching the base of mesocoxae. Pronotum black, hexagonal, with muscular insertions slightly apparent, median carina slightly marked; anterior and anterolateral margins straight; posterolateral with slight sinuosity and posterior margin medially grooved; scutellum black, with slight central concavity and slightly marked horizontal grooves. Tegmina (Fig. +10 A, B +) black or brownish, narrow, with one transverse band whitish between basal and median third and an incomplete band between median and apical third resembling two rounded spots; veins M and Cu 1 united at base, veins A 1 and A 2 distincts, apical reticulation developed, prominent venation. Hindwings hyaline, with brownish venation, Cu 1 thickened at the base, hamuli with three or four spines. Legs black or brownish, metathoracic femur with an apical spine; tibia with two lateral spines, basal one smaller than the ones located at apical crown, which has 14 spines arranged in two rows; basitarsus with three rows of spines covered by long setae; subungueal process present. — +MALE: Genitalia +: Pygofer (Fig. +10 D, E +) with one finger-like process between anal tube and subgenital plate; subgenital plate long, with rounded apex; basal process short and rounded followed by an excavation, dorsal margin covered with small tooth-like spines. Paramere (Fig. +10 F +) subretangular, dorsal margin rounded and developed, apex subquadrangular, subapical hook-like spine slender bearing small denticles in the inferior margin, with an acute apex directed outwards and forward. Aedeagus (Fig. +10 G – I +) subcylindrical, narrowing towards apex with a pair of straight dorsal processes, shorter than half the length of the shaft and inserted between middle and apical third of aedeagus. — +FEMALE +: specimens unknown. + + + + + + + +Mahanarva +( +Ipiranga +) +aguirrei + +. Habitus: +A +dorsal view; +B +lateral view; +C +ventral view. Male genitalia, pygofer: +D +lateral view; +E +ventral view. Paramere: +F +lateral view. Aedeagus: +G +lateral view; +H +dorsal view; +I +ventral view. Scale bars in mm. + + + + + +Remarks. + + +This species shows similarities to + +M. +( +I. +) +nefasta + + +sp. nov. + +mainly due to the pattern and coloration of the tegmina, but presents some unique characters: a rounded subgenital plate apex, a subquadrangular paramere apex, and a slender, acute paramere spine. This species is registered for the first time in +Brazil +. + + + + +Distribution. + + +Brazil +([new record] +Paraná +), +Paraguay +. + + + + +Material examined. + + +BRASIL +, + +Paraná + +: Umuarama, +5–10. x. 1980 +, A. Yamamoto leg., +1 ♂ +( + +MCTP + +); S. J. dos Pinhais, + +25 ° 36 ′ 18 ″ S +49 ° 11 ′ 37 ″ W + +, +880 m +, +01–31. xii. 2020 +, Malaise, A. C. Domahovski leg., +1 ♂ +( + +DZUP + +); idem: +01–31. i. 2020 +, A. C. Domahovski leg., +1 ♂ +( + +DZUP + +). +PARAGUAY +, + +Canindeyú + +: Reserva Natural del Bosque Mbaracayu, Jejui-mi, Mal. 3, 29. iii – 09. iv. 1996, A. C. F Costa leg., +1 ♂ +( + +DZUP + +). + + + + \ No newline at end of file diff --git a/data/A6/33/32/A633320C43865416AE4804D368C303BE.xml b/data/A6/33/32/A633320C43865416AE4804D368C303BE.xml new file mode 100644 index 00000000000..dc971c5ed8c --- /dev/null +++ b/data/A6/33/32/A633320C43865416AE4804D368C303BE.xml @@ -0,0 +1,340 @@ + + + +Taxonomic review, morphometry, and phylogenetic analysis of Mahanarva (Ipiranga) Fennah, 1968 (Hemiptera: Auchenorrhyncha: Cercopidae) + + + +Author + +Meneghetti, Jéssica +0000-0001-6633-6728 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil + + + +Author + +Biganzoli-Rangel, Alejandro Jose +0000-0003-4426-3286 +Department of Biology, York University, Toronto, Ontario, Canada + + + +Author + +Carvalho, Gervásio Silva +0000-0002-3835-8644 +Escola de Ciências da Saúde e da Vida, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga, 6681, 90619 - 900, Porto Alegre, Rio Grande do Sul, Brasil (retired) + + + +Author + +Paladini, Andressa +0000-0001-8894-6092 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil & Departamento de Zoologia, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal do Paraná, Av. Cel. Francisco H. dos Santos, 100, Jardim das Américas, 81530 - 000, Curitiba, Paraná, Brasil + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +45 +73 + + + +journal article +10.3897/asp.83.e130043 +2000DF4D-D639-4E57-90EB-590A80CD9A6C + + + + +3.4. 11. + +Mahanarva +( +Ipiranga +) +nefasta + +sp. nov. + + + + +Figures 6 I +, +13 A – L + + + + +Measurements male / female. + +Head length: 0.84 / 0.97; head width: 1.87 / 2.02; pronotum length: 1.57 / 1.83; pronotum width: 2.75 / 3.01; tegmen length: 7.67 / 8.35; tegmen width: 2.33 / 2.4; total length: 9.1 / 10.22 (in mm). + + + +Diagnosis. + +Tegmina brownish with one complete, yellowish transverse band between basal and median third, and an incomplete transverse band between median and apical third; subgenital plate not excavated at the basal portion and with an acute apex; paramere apex rounded. + + + +Description. + + +Head black, vertex subretangular with a slightly marked median carina in the apex, but indistinct on its base, ocelli closer to each other than to eyes, separated by about one diameter from each other; tylus quadrangular, black, with its basal third with slightly marked carina, becoming indistinct towards the apex (Fig. +13 A +); antenna brownish, pedicel sparsely setose; basal body of flagellum subcylindrical with a single and short arista shorter than the pedicel length. Postclypeus black, inflated, with convex profile, longitudinal carina present, lateral grooves slightly marked (Fig. +13 B, C +); rostrum with second segment yellowish and third segment brownish, reaching the base of mesocoxae. Pronotum black, hexagonal, with muscular insertions slightly apparent, median carina slightly marked; anterior and anterolateral margins straight; posterolateral with slight sinuosity and posterior margin medially grooved, humeral angles acute; scutellum black, with slight central concavity and slightly marked horizontal grooves. Tegmina (Fig. +13 A, B +) brownish, narrow, with one transverse complete band yellowish between basal and median third and an incomplete yellowish band resembling rounded spots, between median and apical third; veins M and Cu 1 united at base, veins A 1 and A 2 distincts, apical reticulation undeveloped, prominent venation. Hindwings hyaline, with brownish venation, Cu 1 thickened at the base, hamuli with three spines. Legs light brown, metathoracic femur with a small apical spine; tibia with two lateral spines, basal one smaller than the ones located at apical crown, which has 12–14 spines arranged in two rows; basitarsus with three rows of spines covered by long setae; subungueal process present. — +MALE: Genitalia +: Pygofer (Fig. +13 D, E +) with one finger-like processes between anal tube and subgenital plate; subgenital plate long, not excavated at the basal portion, covered with small tooth-like spines, and narrowing towards the apex, which is acute. Paramere (Fig. +13 F +) subretangular, dorsal margin rounded and developed, apex rounded; subapical hook-like spine directed outwards and forward. Aedeagus (Fig. +13 G – I +) subcylindrical, narrowing towards apex with a pair of straight dorsal processes, shorter than half the length of the shaft and inserted between middle and apical third of aedeagus. — +FEMALE +: Morphology similar to that of the male, but slightly larger in relation to total body length. First valvula of ovipositor long (Fig. +13 J, K +), with acute apex, basal process rounded and developed, directed downwards; second valvula (Fig. +13 L +) long with rounded apex and dorsal margin covered by teeth in apical third; third valvula short and wide, with rounded apex, covered ventrally by long setae. + + + + + + + +Mahanarva +( +Ipiranga +) +nefasta + + +sp. nov. + +Habitus: +A +dorsal view; +B +lateral view; +C +ventral view. Male genitalia, pygofer: +D +lateral view; +E +ventral view. Paramere: +F +lateral view. Aedeagus: +G +lateral view; +H +dorsal view; +I +ventral view. Female genitalia, ovipositor: +J +lateral view. First valvulae: +K +lateral view. Second valvulae: +L +lateral view. Scale bars in mm. + + + + + + + + +Mahanarva +( +Ipiranga +) +obliqua + + +sp. nov. + +in its natural habitat. +A +– +C + +Mahanarva +( +Ipiranga +) +obliqua + +: Brazil, Paraná, Piraquara (photo by Raul Czelujinski Varella); +D +Site where the species was collected (photo by Raul Czelujinski Varella). + + + + + +Remarks. + + +This species was being treated as + +M. +( +I. +) +aguirrei + +mainly by the pattern and coloration of the tegmina, and the phylogenetic analysis presented here corroborates a close relationship between these species (Fig. +4 +). Despite this, they differ in several aspects in relation to male genitalia: while + +M. +( +I. +) +aguirrei + +presents a long subgenital plate with rounded apex and a rounded depression on its basal third, + +M. +( +I. +) +nefasta + + +sp. nov. + +has an acute subgenital plate apex. Moreover, + +M. +( +I. +) +aguirrei + +has a subquadrangular paramere apex, which differs from the rounded paramere apex seen in + +M. +( +I. +) +nefasta + + +sp. nov. + +In comparison to the other species within the subgenus, this species presents a more developed basal process of the ovipositor. + + + + +Etymology. + + +The new species name is due to the fact that it had long been mistaken for + +M. +( +I. +) +aguirrei + +in our morphological analyses. + + + + +Distribution. + + +Brazil +( +Minas Gerais +). + + + + +Material examined. + + +BRASIL +, + +Holotype +♂ +: +Minas Gerais + +: Uberaba, +04. i. 1984 +, Koller, W. W. leg. ( + +MCTP + +); + + +Paratypes + + +: same data as +holotype +, +8 ♂ +, +6 ♀ +( + +DZUP + +). + + + + +Adicional material examined. + + +BRASIL +, + +Minas Gerais + +: Uberaba, +04. i. 1984 +, Koller, W. W. leg. +9 ♂ +, +3 ♀ +( + +MCTP + +). + + + + \ No newline at end of file diff --git a/data/AA/B8/3F/AAB83F0BC6B956EC8441953986AC8C91.xml b/data/AA/B8/3F/AAB83F0BC6B956EC8441953986AC8C91.xml new file mode 100644 index 00000000000..6edcb7719b7 --- /dev/null +++ b/data/AA/B8/3F/AAB83F0BC6B956EC8441953986AC8C91.xml @@ -0,0 +1,1354 @@ + + + +Taxonomic review, morphometry, and phylogenetic analysis of Mahanarva (Ipiranga) Fennah, 1968 (Hemiptera: Auchenorrhyncha: Cercopidae) + + + +Author + +Meneghetti, Jéssica +0000-0001-6633-6728 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil + + + +Author + +Biganzoli-Rangel, Alejandro Jose +0000-0003-4426-3286 +Department of Biology, York University, Toronto, Ontario, Canada + + + +Author + +Carvalho, Gervásio Silva +0000-0002-3835-8644 +Escola de Ciências da Saúde e da Vida, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga, 6681, 90619 - 900, Porto Alegre, Rio Grande do Sul, Brasil (retired) + + + +Author + +Paladini, Andressa +0000-0001-8894-6092 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil & Departamento de Zoologia, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal do Paraná, Av. Cel. Francisco H. dos Santos, 100, Jardim das Américas, 81530 - 000, Curitiba, Paraná, Brasil + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +45 +73 + + + +journal article +10.3897/asp.83.e130043 +2000DF4D-D639-4E57-90EB-590A80CD9A6C + + + + +3.4. 6. + +Mahanarva +( +Ipiranga +) +integra +( +Walker, 1858 +) + + + + + +Figures 6 C +, +9 A – L + + + + + + + +Monecphora integra +Walker, 1858 +a: 86 + + + + + + + + +Delassor rubicundus integer + + +[sic]: +Fennah 1953 +c: 350; + +Metcalf 1961: 169 + + + + + + + + + +Mahanarva +( +Ipiranga +) +integra + + +: + +Carvalho and Webb, 2005: 69 + +; +stat. nov. + + + + + + + + +Monecphora moreirae + +Lallemand, 1924: 381 +. + +Metcalf 1961: 228 + +. + + + + + + + + + +Mahanarva +( +Ipiranga +) +moreirae + + +: + +Fennah 1968: 187 + +; + +Carvalho and Webb 2005 + +; +syn. nov. + + + + + + + +Measurements male / female. + +Head length: 1.05 / 1.155; head width: 1.99 / 2.2; pronotum length: 1.85 / 1.98; pronotum width: 2.99 / 3.36; tegmen length: 8.66 / 9.18; tegmen width: 2.96 / 3.14; total length: 10.46 / 11.12 (in mm). + + + +Diagnosis. + +Tegmina black with two wide transverse orange bands, with margins approximately straight, posterior band never fragmented into spots; pygofer with one finger-like process between anal tube and subgenital plate, subgenital plate with an acute apex, basal process short and rounded. + + + +Redescription. + + +Head black, vertex subretangular with a slightly marked median carina, scarce pubescence, ocelli closer to each other than to eyes, separated by about one and a half diameter from each other; tylus quadrangular, black, basal third with slightly marked carina, becoming indistinct towards the apex (Fig. +9 A +); antenna brownish, pedicel sparsely setose; basal body of flagellum subcylindrical with a single and short arista shorter than the pedicel length. Postclypeus black, inflated, with convex profile, longitudinal carina present, lateral grooves slightly marked (Fig. +9 B, C +); rostrum with second segment reddish and third segment black, reaching the base of mesocoxae. Pronotum black, hexagonal, with muscular insertions slightly apparent, median carina slightly marked; anterior and anterolateral margins straight, posterolateral with slight sinuosity and posterior margin medially grooved, humeral angles acute scutellum black, with slight central concavity and slightly marked horizontal grooves. Tegmina (Fig. +9 A, B +) narrow, black, with two transverse wide bands orange: one between basal and medium third and another between medium and apical third; veins M and Cu 1 united at base, veins A 1 and A 2 distincts, apical reticulation developed, prominent venation. Hindwings hyaline, with brownish venation, Cu 1 thickened at the base, hamuli with three or four spines. Legs reddish, metathoracic femur with a small apical spine; tibia with two lateral spines, basal one smaller than the ones located at apical crown, which has 12–14 spines arranged in two rows; basitarsus with three rows of spines covered by long setae; subungueal process present. — +MALE: Genitalia +: Pygofer (Fig. +9 D +) with one finger-like process between anal tube and subgenital plate; subgenital plate narrowing towards apex, which is acute and spine-like directed outward (in ventral view) (Fig. +9 E +), dorsal margin with rounded elevation covered with small tooth-like spines; basal process of subgenital plate short and rounded followed by and excavation. Paramere (Fig. +9 F +) subretangular, dorsal margin rounded and developed, apex subtriangular and slightly acute with long setae; subapical hook-like spine bearing small denticles in the inferior margin, directed outwards and forward. Aedeagus (Fig. +9 G – I +) subcylindrical, narrowing towards apex, with a pair of straight dorsal processes, shorter than half length of shaft and inserted between median and apical third of aedeagus. — +FEMALE +: Morphology similar to that of the male, but slightly larger in relation to total body length. First valvula of ovipositor long (Fig. +9 J, K +), with acute apex, basal process rounded and undeveloped, directed downwards; second valvula (Fig. +9 L +) long with rounded apex and dorsal margin covered by teeth in apical third; third valvula short and wide, with rounded apex, covered ventrally by long setae. + + + + + + + +Mahanarva +( +Ipiranga +) +integra + +. Habitus: +A +dorsal view; +B +lateral view; +C +ventral view. Male genitalia, pygofer: +D +lateral view; +E +ventral view. Paramere: +F +lateral view. Aedeagus: +G +lateral view; +H +dorsal view; +I +ventral view. Female genitalia, ovipositor: +J +lateral view. First valvulae: +K +lateral view. Second valvulae: +L +lateral view. Scale bars in mm. + + + +Remarks. + +M. +( +I. +) +integra + +shares morphological similarities in the male genitalia with + +M. +( +I. +) +rubicunda + +and + +M. +( +I. +) +rubripennis + +. These include a rounded and short basal process on the subgenital plate followed by an excavation and a rounded elevated dorsal margin. + +M. +( +I. +) +integra + +exhibits variation in tegmina proportion with some specimens having wider tegmina comparing to others. While the color and pattern of transverse bands on the tegmina resembles those of + +M. +( +I. +) +rubicunda + +, the bands on + +M. +( +I. +) +integra + +are wider. Furthermore, + +M. +( +I. +) +integra + +has a predominantly black body color with reddish legs. + + + + +Distribution. + + +Brazil +( +Rio de Janeiro +, +São Paulo +[new record], +Paraná +, +Santa Catarina +and +Rio Grande do Sul +), +Argentina +(Misiones). + + + + +Material examined. + + +BRASIL +, + +Rio de Janeiro + +: Guaporé, +03. i. 2008 +, P. C. Grossi leg., +3 ♀ +( + +DZUP + +); + +São Paulo + +: Cássia dos Coqueiros, +x. 1954 +, M. P. Barretto col., +1 ♀ +( +MZSP +); + +Paraná + +: Apucarana, 1975, H. Alvarenga, +1 ♀ +( + +DZUP + +); Araucária, 02. xi. 83, Palú, M. R. leg., +1 ♂ +( + +DZUP + +); Vila Velha, 23. ii. 65, C. Dipterologia, +1 ♀ +( + +DZUP + +); Mariópolis, +06. xii. 1983 +, Exc. Dep. Zool- UFPR, +1 ♂ +( + +DZUP + +); Piraguara, Mananciais da Serra, + +25 ° 29 ′ 46 ″ S +48 ° 58 ′ 46 ″ W + +, +02. i. 2008 +, Grossi, P. C. leg., +4 ♀ +( + +DZUP + +); idem: P. G. Grossi & Paladini leg., +1 ♂ +( + +DZUP + +); idem: +17. xi. 2009 +, Grossi, P. C. leg., +1 ♀ +( + +DZUP + +); Campo Largo, BR 277 Km 115, +21. xi. 2009 +, Dias, F. M. S & Serram leg., E. B. F, +1 ♂ +( + +DZUP + +); +Tijucas do Sul +, Associação dos prof. UFPR Chácara, + +25 ° 50 ′ 14 ″ S +49 ° 02 ′ 57 ″ W + +, A Paladini & R. R. Cavichioli leg., +1 ♂ +, +1 ♀ +, 1 without genitalia ( + +DZUP + +); idem: Vossoroca, +17. i. 2003 +, Excursão Fauna Local- DZOO, (no col. data), +3 ♀ +( + +DZUP + +); São José dos Pinhais, + +25 ° 36 ′ 18 ″ S +49 ° 11 ′ 37 ″ W + +, i-vii. 2014, A. C. Domahovski leg., +18 ♂ +, 1 without genitalia ( + +MCTP + +); idem: +880 m +, +12. x. 2013 +, A. C. Domahovski leg., +1 ♂ +( + +DZUP + +); Jaguariaíva, +28. xii. 1966 +, F. Giacomel leg, +1 ♂ +( + +MCTP + +); Foz do Iguaçu, +16. xii. 1965 +, (no col. data), +1 ♂ +( + +MCTP + +); Curitiba, +15–27. x. 1983 +, E. C. Costa leg., +1 ♂ +( + +DZUP + +); Fênix, +30. iii. 1987 +, Dutra, R. leg., +1 ♂ +( + +DZUP + +); Ponta Grossa, P. E. de Vila Velha, +25–26. xi. 2011 +, Grosso, Santos & Melo leg., +1 ♂ +( + +DZUP + +); + +Santa Catarina + +: Nova Teutonia, + +27 ° 11 ′ 00.0 ″ S +52 ° 23 ′ 00.0 ″ W + +, +xi. 1982 +, Fritz Plaumann leg., +1 ♂ +( + +DZUP + +), +2 ♀ +( + +DZUP + +); idem: +ii. 1981 +, Fritz Plaumann leg., +19 ♂ +( + +DZUP + +), +1 ♀ +( + +DZUP + +); idem: +xi. 1977 +, Fritz Plaumann leg., +2 ♂ +( + +DZUP + +), +2 ♀ +( + +DZUP + +); Lages, +2. x. 1983 +, col. R. L. CHI., +1 ♂ +( + +DZUP + +); Joinville, +10. xi. 1974 +, P. Moure col., +1 ♂ +( + +DZUP + +); Ponte Alta, +12. ii. 1973 +, A. M. Sakakibara, +2 ♀ +( + +DZUP + +); Campo Alto, Sta. Cecília, 12. ii. 73, Sakakibara e Mielke, leg., +1 ♀ +( + +DZUP + +) e 1 without genitalia ( + +DZUP + +); idem: +26. ii. 1968 +, Moure & Mielke, +1 ♀ +( + +DZUP + +); Rio Vermelho, ii. 74, Abílio leg., +3 ♂ +, +8 ♀ +( + +DZUP + +); Rancho Queimado, +15–18. xi. 1995 +, A. Bonaldo leg., +1 ♂ +( + +MCTP + +); idem: +15–18. xi. 1995 +, L. Moura leg., +2 ♂ +( + +MCTP + +); Florianópolis, +27. iv. 1980 +, Butignol C. A. leg., +2 ♂ +( + +MCTP + +); Ponte Alta, +12. ii. 1973 +, A. M. Sakakibara, +1 ♀ +( + +MCTP + +); (no location data), +10. ii. 1973 +, Lopez, P. leg., +1 ♀ +( + +MCTP + +); Campos Novos, +7. i. 1989 +, Mansur, C. & Périco, E., leg., +1 ♂ +( + +MCTP + +); idem: +5. i. 1989 +, Fernandes, J. A. & Penz, C. M. leg., +1 ♂ +, +1 ♀ +( + +MCTP + +); idem: +6. i. 1989 +, Mansur, C. & Pericie., leg., +1 ♂ +, +2 ♀ +, 1 without genitalia ( + +MCTP + +); idem: +6. i. 1989 +, Fernandes, J. A., leg., +1 ♀ +( + +MCTP + +); idem: +6. i. 1989 +, Penz, CM & Fernandes, J. A., leg., +1 ♀ +( + +MCTP + +): idem: +04. i. 1989 +, Equipe do Projeto E. col., +1 ♂ +( + +MCTP + +); Campo Alegre, +15. ii. 2008 +, Pinto, Â. P. col., +3 ♂ +, +5 ♀ +( + +MCTP + +); idem: Rio do Turvo-SC 301, +16. ii. 2008 +, Pinto, Â. P. col., +1 ♀ +( + +MCTP + +); Criciúma, Parque José Milaneze, +13. xii. 2004 +, Martins, F. leg., +1 ♀ +( + +MCTP + +); idem: +27. xii. 2004 +, Martins, F. leg., +1 ♂ +( + +MCTP + +); + +Rio Grande do Sul + +: São Francisco de Paula, Pró-Mata, + +29 ° 28 ′ 49 ″ S +50 ° 10 ′ 28 ″ W + +, +14–17. xii. 2009 +, A. Paladini leg., +2 ♂ +, +4 ♀ +( + +DZUP + +); idem: +14–17. xii. 2009 +, A Paladini & R. R. Cavichioli, +1 ♂ +( + +DZUP + +); idem: +25. ii. 1999 +, A. Köhler leg., +1 ♀ +( + +MCTP + +); idem: +19–21. iii. 1998 +, Carvalho, G. S. leg., +8 ♂ +, +2 ♀ +( + +MCTP + +); idem: +26. iv. 2003 +, (no col. data), +1 ♂ +, +1 ♀ +( + +MCTP + +); idem: +19–20. xi. 1998 +, Carvalho & Pulz, leg., +2 ♂ +, +4 ♀ +( + +MCTP + +); idem: +15–16. v. 2010 +, Carvalho, G. S. leg., +1 ♂ +, +1 ♀ +( + +MCTP + +); idem: +15. iii. 1996 +, Birgit Harter leg., +1 ♀ +( + +MCTP + +); idem: +27. iv. 2003 +, (no col. data), +1 ♀ +( + +MCTP + +); idem: +23. iv. 1999 +, Carvalho, G. S. leg., +1 ♀ +( + +MCTP + +); idem: +2. v. 1997 +, Carvalho, G. S. leg., +4 ♂ +, +1 ♀ +( + +MCTP + +); idem: +11. iv. 1997 +, Carvalho, G. S. leg., +2 ♀ +( + +MCTP + +); idem: +20–21. x. 2001 +, (no col. data), +1 ♀ +( + +MCTP + +); idem: +24. iii. 1998 +, Koehler, leg., +1 ♂ +( + +MCTP + +); idem: +27. iv. 2003 +, Lab. Ent. Sis. leg., +4 ♂ +( + +MCTP + +); idem: +27-29. iii. 2006 +, E. L. C. Silva & A. A. Lise leg., +1 ♀ +( + +MCTP + +); +Montenegro +, + +29 ° 38 ′ 00.0 ″ S +51 ° 28 ′ 00.0 ″ W + +, +12. xi. 2009 +, A. Paladini leg., +3 ♂ +, +1 ♀ +( + +DZUP + +); idem: +29. x. 2009 +, A. Paladini leg., +1 ♂ +( + +DZUP + +); idem: +19. x. 2001 +, Ott, A. P. leg., +1 ♂ +, +2 ♀ +( + +MCTP + +); idem: +30. v. 2011 +, Citros- Pq. Gaúcho leg., +1 ♂ +( + +MCTP + +); idem: +17. xi. 2011 +, Citros- Pq. Gaúcho leg., +4 ♂ +, +4 ♀ +( + +MCTP + +); idem: +29. xii. 2011 +, Citros- Pq. Gaúcho, leg., +1 ♂ +, +3 ♀ +( + +MCTP + +); idem: +12. i. 2012 +, Citros- Pq. Gaúcho leg., +1 ♂ +, +1 ♀ +( + +MCTP + +); idem: +1. xii. 2011 +, Citros- Pq. Gaúcho leg., +8 ♂ +, +3 ♀ +( + +MCTP + +); idem: +15. xii. 2011 +, Citros- Pq. Gaúcho leg., +2 ♂ +, +13 ♀ +( + +MCTP + +); idem: +3. xi. 2011 +, Citros- Pq. Gaúcho leg., +1 ♂ +( + +MCTP + +); idem: +7. i. 2011 +, Citros- Pq. Gaúcho leg., +1 ♀ +( + +MCTP + +); idem: +8. x. 2009 +, Gadelha, Y. E. A. col., +1 ♀ +( + +MCTP + +); idem: +14. xii. 2009 +, Gadelha, Y. E. A. col., +1 ♂ +, +6 ♀ +( + +MCTP + +); idem: +12. xi. 2009 +, Gadelha, Y. E. A. col., +1 ♂ +, +2 ♀ +( + +MCTP + +); idem: +27. xi. 2009 +, Gadelha, Y. E. A. col., +1 ♂ +, +4 ♀ +( + +MCTP + +); idem: +27. xi. 2009 +Carvalho, G. S. leg., +3 ♂ +, +2 ♀ +( + +MCTP + +); idem: +29. xii. 2009 +, Gadelha, Y. E. A. col., +2 ♀ +( + +MCTP + +); Passo Fundo, + +28 ° 13 ′ 40 ″ S +52 ° 24 ′ 19 ″ W + +, +18. xi. 2011 +, S. Lampert leg., +1 ♂ +( + +DZUP + +); Bento Gonçalves, +28. iii. 2007 +, Bertin, A. leg., +2 ♂ +( + +MCTP + +); idem: +8. x. 2008 +, Tognon, R. col., +1 ♂ +( + +MCTP + +); idem: +20. xi. 2008 +, Tognon, R. col., +1 ♂ +( + +MCTP + +); Viamão, +25. xi. 1994 +, Petersen, A. leg., +6 ♂ +, +11 ♀ +( + +MCTP + +); idem: +7. xi. 1995 +, Petersen, A., +2 ♂ +, +3 ♀ +( + +MCTP + +); idem: +7. x. 1994 +, Exc LSE leg., +1 ♂ +( + +MCTP + +); idem: +2. xii. 1994 +, Petersen, A. leg., +1 ♀ +( + +MCTP + +); idem: +14. xi. 1998 +, Prates, P. leg., +1 ♀ +( + +MCTP + +); Guaíba, +24. x. 1995 +, Petersen, A. leg., +1 ♀ +( + +MCTP + +); Torres, +20. v. 1995 +, (no col. data), +1 ♀ +( + +MCTP + +); idem: Col. São Pedro, +8. xi. 1997 +, Carvalho, G. leg., +4 ♂ +, +3 ♀ +( + +MCTP + +); +Salvador do Sul +, +ix – xii. 1994 +, A. Specht, leg., +1 ♀ +( + +MCTP + +); Ijuí, +4. xi. 1978 +, A. Dressler leg., +1 ♂ +( + +MCTP + +); Santa Maria, +8. xi. 1978 +, J. Abreu, +1 ♂ +( + +MCTP + +); idem: +26. x. 1978 +, L. A. Grohe leg., +1 ♂ +( + +MCTP + +); idem: +21. x. 1978 +, L. C. Becker leg., +1 ♂ +( + +MCTP + +); Pelotas, +6. iii. 1979 +, Ortiz leg., 1 without genitalia ( + +MCTP + +); Porto Alegre, +i. 1981 +, Carvalho, G. leg., +1 ♀ +( + +MCTP + +); idem: +28. xii. 1983 +, E. Corseuil leg., +1 ♀ +( + +MCTP + +); idem: +2. iv. 1963 +, C. Trés leg., +1 ♂ +( + +MCTP + +); idem: +6. xii. 1979 +, Butignol leg., 1 without genitalia ( + +MCTP + +); Canela, +23. xii. 1961 +, A. Lise leg., +1 ♂ +( + +MCTP + +); Cotiporã, + +28 ° 59 ′ 00.0 ″ S +51 ° 38 ′ 00.0 ″ W + +, +6–8. i. 2011 +, Pinto, A. P, & Silva, J. G. leg., +2 ♂ +, +2 ♀ +; Barão de Cotegipe, +15. i. 1967 +, F. Giacomel leg., +3 ♂ +( + +DZUP + +), +1 ♀ +( + +MCTP + +); idem: +21. i. 1967 +, F. Giacomel leg., +1 ♂ +( + +DZUP + +), +1 ♀ +( + +DZUP + +); Vila Maria, +26. ii. 1998 +, Specht, A. col., +1 ♂ +( + +MCTP + +); Capão da Canoa, +08. i. 1983 +, Trois, C. leg., +1 ♂ +( + +MCTP + +); Frederico Westphalen, +vii. 2009 +, Silva, E. R; Bussato, D. leg., +1 ♂ +, +1 ♀ +( + +MCTP + +). +ARGENTINA +, + +Misiones + +: Cataratas del Iguazú, +24. xi. 1980 +, Willink-Claps-Dominguez leg., +1 ♂ +( + +MCTP + +); idem: +26. xi. 1980 +, Willink-Claps leg, +1 ♀ +( + +MCTP + +); idem: Bernardo de Irigoyen, +12. xi. 1973 +, Escobar-Claps, +1 ♀ +( + +MCTP + +); idem: +xii. 1961 +, L. N. Alem, A. Martinez leg., +1 ♂ +( +MZSP +). + + + + \ No newline at end of file diff --git a/data/B2/40/54/B24054E42BBF53DEAC5979328D9A0580.xml b/data/B2/40/54/B24054E42BBF53DEAC5979328D9A0580.xml new file mode 100644 index 00000000000..45f79fe10df --- /dev/null +++ b/data/B2/40/54/B24054E42BBF53DEAC5979328D9A0580.xml @@ -0,0 +1,533 @@ + + + +Genetic and multivariate morphometric analyses unveil cryptic diversity in sympatric populations of the Mediterranean wool carder bee, identified as Anthidium undulatum Dours, 1873 (Hymenoptera: Megachilidae: Anthidiini) + + + +Author + +Kasparek, Max +0000-0002-5604-6791 +Mönchhofstr. 16, 69120 Heidelberg, Germany + + + +Author + +Boustani, Mira +Laboratory of Zoology, Institute for Biosciences, University of Mons, Place du Parc 20, 7000 Mons, Belgium + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +75 +92 + + + +journal article +10.3897/asp.83.e137570 +05CB5D16-02E4-4323-9294-45A28971CCC2 + + + + + +Anthidium wahrmani +Mavromoustakis, 1948 + +stat. nov. + + + + +Figures 6 +, +7 + + + + + + + +Proanthidium undulatum wahrmani +Mavromoustakis, 1948 + +. Male, female ( +Israel +/ +Palestine +). – Mavromoustakis (1948), stated that the +holotype +is deposited in his own collection. However, it has not been listed in the collection inventory (MoA, 1989). The material presented here was attributed to this taxon based on Mavromoustakis’ description. + + + + + + +Material examined. + + + +ISRAEL +/ +PALESTINE + +: +2 ♀ +, +2 ♂ +; +R +70, +10 km +NNE Nahariya ( + +33.05 N +35.15 E + +), +60 m +; +28. iv. 2018 +; M. Halada leg. ( +CMK +: mk 1788, +ABABX +548-22; +CMK +: mk 1789, +ABABX +450-22; +CMK +: mk 1791, +ABABX +539-22; +CMK +: mk 1790, +ABABX +538-22). – +1 ♀ +; +15 km +E Qiryat Shemona Hermon Foothill ( + +33.25 N +36.73 E + +); +16. v. 1996 +; M. Hauser leg. ( +CMK +- +MH +, mh 067). – +1 ♂ +; Mt. Carmel, old quarry ( + +32.73 N +35.03 E + +); +14. v. 2000 +; M. Török leg. ( + +ZMH + +: zmh 009). – +2 ♂ +; Mt. Carmel Wadi Denia ( + +32.75 N +35.00 E + +); +04. vii. 2000 +; M. Török leg. ( + +ZMH + +: zmh 010-11). — + +JORDAN + +: +1 ♀ +, +1 ♂ +; N. Shuna ( + +32.61 N +35.60 E + +); +20. – 22. iv. 1996 +; Ma. Halada leg. ( +CMK +: ms 0940, 1430). – +1 ♂ +; Jordan valley, Mubalath (Tal al Maqlub) ( + +32.40 N +35.68 E + +); +27. iv. 1996 +; Ma. Halada leg. ( +CMK +: ms 0820). – +1 ♀ +; Petra ( + +30.32 N +35.44 E + +); +14. v. 1995 +; K. Deneš sen. leg. ( +OLL +: oll 0935). — + +LEBANON + +: +1 ♂ +; North Lebanon: Harissa, Al Jawar ( + +34.18 N +35.92 E + +), +1765 m +; +18. vii. 2017 +; M. Boustani leg. ( +CMK +: MBou 245; +MGPCC +093-21). – +1 ♂ +; North Lebanon: Tannourine Reserve, Trail 4 ( + +34.20 N +35.92 E + +), +1781 m +; +27. vii. 2019 +; M. Boustani leg. ( +CMK +: MBou 134; +ABABX +059-20). – +2 ♀ +, +1 ♂ +; Mount Lebanon: Anjar, Guest House ( + +33.72 N +35.93 E + +), +973 m +; +23. vii. 2019 +; M. Boustani & X. van Achter leg. ( +CMK +: mbou 136, +ABABX +057-20; +CMK +: mbou 138:, +ABABX +449-22; coll. M. Boustani: mbou 139). — + +PALESTINE + +: +1 ♂ +; +10 km +E Jerusalem ( + +31.82 N +35.33 E + +); +05. v. 1996 +; M. Hauser leg. ( +CMK +: mh 068). + + + + +Barcode Index Number. + + + +BOLD +: AAO 4344 + +. + + + + +Diagnosis. + + +Maculation of integument dark yellow (as in + +A. undulatum + +, but pale yellow in + +A. libanicum + +). Female: Clypeus yellow without black maculation (as in + +A. libanicum + +, but present in most + +A. undulatum + +). Broad yellow preoccipital band, also covering genae entirely; yellow tergal bands broad, uninterrupted; +T +6 yellow with a black wedge-shaped apical maculation (as in + +A. libanicum + +; in + +A. undulatum + +mostly black with a large yellow spot on each side). Male similar in coloration, but preoccipital band reaching only middle of genae. See also the diagnosis of + +A. undulatum + +. + + + + + + +Face and apical terga of typical females of + +Anthidium + +: +A +, +D + +A. undulatum + +; +B +, +E + +A. libanicum + + +sp. n. + +; +C +, +F + +A. wahrmani + + +stat. nov. + +Note the color of the antennal scape (black in + +A. undulatum + +and + +A. libanicum + +), the color pattern of T 6 (black in + +A. undulatum + +, yellow with black apicomedial wedge-shaped incision in + +A. libanicum + + +sp. n. + +and + +A. wahrmani + + +stat. nov. + +), and overall tone of yellow (pale yellow in + +A. libanicum + + +sp. n. + +, dark bright yellow in + +A. undulatum + +and especially in + +A. wahrmani + + +stat. nov. + +). + + + + + +Habitat. + + +See under + +A. libanicum + +and Figs +10 +, +11 +. + + + + + + + +Anthidium libanicum + + +sp. n. + +Male metasoma, left with black ground color, right a rare color morph with red-brown ground color. The belonging of these two specimens to the same species ( + +A. libanicum + + +sp. n. + +) has been confirmed by genetic barcoding. + + + + + + + +Habitat of + +Anthidium libanicum + + +sp. n. + +in Jord Aarsal in the Anti-Lebanon Mountains, Lebanon, at 1.975 m a. s. l. + + + + + +Distribution. + + +Distribution restricted to the southern Levant with records from +Israel +/ +Palestine +, +Jordan +, +Lebanon +, and southern +Syria +. The distribution overlaps with + +A. libanicum + + +sp. n. + +in +Lebanon +. Taxonomic status in south-eastern +Türkiye +unclear, where it possibly occurs and overlaps with + +A. undulatum + +(Fig. +2 +). + + + + + + +Habitat of + +Anthidium wahrmani + + +stat. nov. + +in the Tannourine Nature Reserve, Lebanon, in the Lebanon Mountains at 1.780 m a. s. l. + + + + + + + +Mountainous habitat in Harissa al Jawar, North Lebanon, in which + +Anthidium wahrmani + +and + +A. libanicum + + +sp. n. + +were found to occur sympatrically. + + + + + \ No newline at end of file diff --git a/data/BA/89/7E/BA897ED245D65E0CB3BA6EA7EB2467E8.xml b/data/BA/89/7E/BA897ED245D65E0CB3BA6EA7EB2467E8.xml new file mode 100644 index 00000000000..71bec985064 --- /dev/null +++ b/data/BA/89/7E/BA897ED245D65E0CB3BA6EA7EB2467E8.xml @@ -0,0 +1,844 @@ + + + +Taxonomic review, morphometry, and phylogenetic analysis of Mahanarva (Ipiranga) Fennah, 1968 (Hemiptera: Auchenorrhyncha: Cercopidae) + + + +Author + +Meneghetti, Jéssica +0000-0001-6633-6728 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil + + + +Author + +Biganzoli-Rangel, Alejandro Jose +0000-0003-4426-3286 +Department of Biology, York University, Toronto, Ontario, Canada + + + +Author + +Carvalho, Gervásio Silva +0000-0002-3835-8644 +Escola de Ciências da Saúde e da Vida, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga, 6681, 90619 - 900, Porto Alegre, Rio Grande do Sul, Brasil (retired) + + + +Author + +Paladini, Andressa +0000-0001-8894-6092 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil & Departamento de Zoologia, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal do Paraná, Av. Cel. Francisco H. dos Santos, 100, Jardim das Américas, 81530 - 000, Curitiba, Paraná, Brasil + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +45 +73 + + + +journal article +10.3897/asp.83.e130043 +2000DF4D-D639-4E57-90EB-590A80CD9A6C + + + + +3.4. 8. + +Mahanarva +( +Ipiranga +) +rubripennis +( +Schmidt, 1922 +) + + + + + +Figures 6 E +, +11 A – L + + + + + + + +Luederwaldtia rubripennis + +Schmidt, 1922: 263 +. + +Metcalf 1961: 539 + + + + + + + + + + +Mahanarva rubripennis + + +: + +Fennah 1968: 186 + + + + + + + + + +Mahanarva +( +Ipiranga +) +rubripennis + + +: + +Paladini and Cavichioli 2014: 483 + +comb. nov. + + + + + + + +Measurements male / female. + +Head length: 0.90 / 1.02; head width: 1.82 / 1.98; pronotum length: 1.65 / 1.82; pronotum width: 2.76 / 3.04; tegmen length: 7.34 / 8.06; tegmen width: 2.26 / 2.56; total length: 8.79 / 10.11 (in mm). + + + +Diagnosis. + +Tegmina red with black, rounded spots on apical plexus of veins; subgenital plate with an acute apex, basal process short and rounded followed by an excavation, paramere with a finger-like apex. + + + +Redescription. + + +Head black, vertex subretangular, with a slightly marked median carina in the apical third, ocelli closer to each other than to eyes, separated by about one and a half diameter from each other; tylus quadrangular, black, with its basal third with slightly marked carina, becoming indistinct towards the apex (Fig. +11 A +); antenna black, pedicel sparsely setose, basal body of flagellum subcylindrical with a single and short arista shorter than the pedicel length. Postclypeus black, inflated, with convex profile, longitudinal carina present, lateral grooves slightly marked (Fig. +11 B, C +); rostrum with second segment reddish and third segment black, reaching the base of mesocoxae. Pronotum black, hexagonal, with muscular insertions slightly apparent, median carina slightly marked; anterior and anterolateral margins straight; posterolateral with slight sinuosity and posterior margin medially grooved, humeral angles acute; scutellum black, with slight central concavity and slightly marked horizontal grooves. Tegmina (Figs +11 A, B +) narrow, red, with black, rounded spots on apical plexus of veins; veins M and Cu 1 united at base, veins A 1 and A 2 distincts, apical reticulation developed, prominent venation. Hindwings hyaline, with brownish venation, Cu 1 thickened at the base, hamuli with three spines. Legs brownish, metathoracic femur with a small apical spine; tibia with two lateral spines, basal one smaller than the ones located at apical crown, which has 10–12 spines arranged in two rows; basitarsus with three rows of spines covered by long setae; subungueal process present. — +MALE: Genitalia +: Pygofer (Fig. +11 D +) with one finger-like process between anal tube and subgenital plate; subgenital plate narrowing towards apex, which is acute and spine-like slightly directed outward (in ventral view) (Fig. +11 E +); dorsal margin with rounded elevation covered with small tooth-like spines; basal process of subgenital plate short and rounded followed by an excavation. Paramere (Fig. +11 F +) subretangular, dorsal margin rounded and developed; with an elongated and finger-like apex; subapical hook-like spine directed outwards and forward. Aedeagus (Fig. +11 G – I +) subcylindrical, narrowing towards apex, with a pair of straight dorsal processes, shorter than half length of shaft and inserted between median and apical thirds of aedeagus. — +FEMALE +: Morphology similar to that of the male, but slightly larger in relation to total body length. First valvula of ovipositor long (Fig. +11 J, K +), with acute apex, basal process rounded and undeveloped, directed downwards and backwards; second valvula long (Fig. +11 L +), with rounded apex and dorsal margin covered by teeth in apical third; third valvula short and wide, with rounded apex, covered ventrally by long setae. + + + + + + + +Mahanarva +( +Ipiranga +) +rubripennis + +. Habitus: +A +dorsal view; +B +lateral view; +C +ventral view. Male genitalia, pygofer: +D +lateral view; +E +ventral view. Paramere: +F +lateral view. Aedeagus: +G +lateral view; +H +dorsal view; +I +ventral view. Female genitalia, ovipositor: +J +lateral view. First valvulae: +K +lateral view. Second valvulae: +L +lateral view. Scale bars in mm. + + + + + +Remarks. + + +The male genitalia of + +M. +( +I. +) +rubripennis + +differs from + +M. +( +I. +) +integra + +due to the elongated paramere apex and the slightly longer subgenital plate. Additionally, the pattern and coloration of the tegmina are quite distinct between these species and can be used as a diagnostic character. + +M. +( +I. +) +rubripennis + +is the only species within the genus that exhibits red tegmina with a pattern of black maculae covering the apical reticulum, making it readily distinguishable from the other species. + + + + +Distribution. + + +Brazil +( +Mato Grosso +[new record], +Minas Gerais +, +Espírito Santo +, +Rio de Janeiro +, +São Paulo +, +Paraná +, +Santa Catarina +, +Rio Grande do Sul +). + + + + +Material examined. + + +BRASIL +, + +Mato Grosso + +: P. N. Chapada dos Guimarães, +i. 2013 +, Sweep M. Savaris & S. Lampert leg., +2 ♂ +( + +DZUP + +); + +Minas Gerais + +: São Gonçalo do Rio Abaixo, +14. ii. 2003 +, A. F. Kumagai col., +2 ♂ +( + +DZUP + +); idem: +4. iv. 2003 +, A. F. Kumagai col., +1 ♂ +, +1 ♀ +( + +DZUP + +); idem: +28. iii. 2003 +, A. F. Kumagai col., +1 ♀ +( + +DZUP + +); + +Espírito Santo + +: Linhares, +vii. 1981 +, C. Elias leg. +1 ♂ +( + +DZUP + +); Santa Teresa, +15. xi. 1967 +, C. T. & C. Elias leg., +2 ♂ +( + +MCTP + +, + +DZUP + +); idem: +12. v. 1967 +, C. T. & C. Elias leg., +2 ♂ +, +1 ♀ +( + +DZUP + +); idem: +07. xii. 1964 +, C. Elias leg., +1 ♂ +, +1 ♀ +( + +DZUP + +); idem: +16–22. v. 1967 +, C. & C. T. Elias leg., +2 ♂ +( + +DZUP + +); idem: +21. ii. 1966 +, C. Tadeu Elias leg. +2 ♀ +( + +DZUP + +); idem: +28. xi. 1966 +, C. T. & C. Elias leg., +1 ♂ +( + +DZUP + +); idem: +26. xi. 1967 +, C. T. & C. Elias leg. +1 ♂ +( + +DZUP + +); idem: 26. x. 64, C. Elias leg., +1 ♂ +( + +DZUP + +); + +Rio de Janeiro + +: Itatiaia, P. N. Itatiaia, +26. xii. 1963 +, Alvarenga leg., +1 ♀ +( + +MCTP + +); + +São Paulo + +: Santos, 10. i. 93, Bittencourt, A. col., +1 ♀ +( + +DZUP + +); São José do Barreiro (Serra Bocaina), +iii. 1973 +, F. M. Oliveira leg., +1 ♀ +( + +DZUP + +); Salesópolis, Estação Biológica de Boracéia, + +23 ° 39 ′ 15.5 ″ S +45 ° 53 ′ 23 ″ W + +, +23–29. xi. 2008 +, A. Paladini, D. R Parizotto, P. C. Grossi, +19 ♂ +, +13 ♀ +( + +DZUP + +); idem: +13. iv. 2001 +, A. M. A. Lima leg., +5 ♂ +; +7 ♀ +( + +DZUP + +); Cananéia, Pq. Est. Ilha do Cardoso, +18–23. x. 2011 +, Exp. Botu Cinza leg., +5 ♂ +, +2 ♀ +( + +MCTP + +); idem: + +25 ° 04 ′ 00.0 ″ S +47 ° 55 ′ 00.0 ″ W + +, +1 ♂ +, +1 ♀ +( + +MCTP + +); Caraguatatuba, +09. xii. 1983 +, Netto leg., +3 ♂ +, +1 ♀ +( + +DZUP + +); Cotia, 11. iii. 73, J. R. Araújo leg., +1 ♂ +, +1 ♀ +( + +DZUP + +); Barueri, +viii. 1958 +, K. Lenco leg., +1 ♂ +( + +DZUP + +); + +Paraná + +: Curitiba, +2. x. 1983 +, Crepaldi S. leg., +1 ♂ +( + +DZUP + +); idem, 9. iv. 81, Juarez C. Oliveira leg., +1 ♂ +( + +DZUP + +); idem: +10. xi. 1985 +, Ferreira leg., +1 ♂ +( + +DZUP + +); idem: 12–13. xi. 83, Luís leg., +1 ♀ +( + +DZUP + +); Piraquara, Mananciais da Serra, + +25 ° 29 ′ 46 ″ S +48 ° 58 ′ 54 ″ W + +, 18. xi. 09, R. R Cavichioli & P. G Grossi, +1 ♀ +( + +DZUP + +); idem: Parque Estadual do Marumbi, + +25 ° 29 ′ 13 ″ S +48 ° 58 ′ 30 ″ W + +, +24. i. 2012 +, Grossi, Cavichioli & Silva leg., +2 ♂ +( + +DZUP + +), +2 ♀ +( + +DZUP + +); idem: +22. xi. 2005 +, R. Gonçalves leg., +1 ♀ +( + +DZUP + +); Antonina, Reserva Rio Cachoeira, + +25 ° 18 ′ 58 ″ S +48 ° 41 ′ 46 ″ W + +, +20–25. xi. 2014 +, Altitude +50 m +, +1 ♂ +, +1 ♀ +( + +DZUP + +); idem: +05–10. xi. 2015 +, Entomologia UFPR, +1 ♀ +( + +DZUP + +); idem: RPPN Guaricica, + +25 ° 18 ′ 58 ″ S +48 ° 41 ′ 46 ″ W + +, +23–27. x. 2017 +, Sweep A. C. Domahovski leg., +2 ♂ +( + +DZUP + +); idem: Rio dos Nunes, +15. vi. 2001 +, A. C. F Da Costa, +1 ♀ +( + +DZUP + +); idem: +16–20. v. 2018 +, Luminosa, R. R. Cavichioli & A. P. Pinto leg. +1 ♂ +( + +DZUP + +); idem, +23–27. i. 2017 +, Luz solo, A. C. Domahovski leg., +1 ♀ +; idem: Reserva Sapitanduva, +20. x. 1986 +, Lev. Ent. PROFAUPAU, Malaise, +1 ♀ +( + +DZUP + +); Guaratuba, Estrada dos Castelhanos, + +25 ° 49 ′ 50 ″ S +48 ° 55 ′ 38 ″ W + +, (no col. data), +1 ♂ +( + +DZUP + +); idem: + +25 ° 50 ′ 00.0 ″ S +48 ° 56 ′ 00.0 ″ W + +, +13. xii. 2004 +, Carvalho, Mauro & Cavichioli, +1 ♂ +, +1 ♀ +( + +DZUP + +); +Pontal do Sul +, +12. x. 2004 +, Paladini, A leg., +4 ♂ +( + +MCTP + +); idem: +11. x. 2004 +, Paladini, A leg., +1 ♂ +, +1 ♀ +( + +MCTP + +); São João de Petrópolis, +6–12. vi. 1967 +, C. & C. T. Elias leg, +1 ♀ +( + +MCTP + +); Paranaguá, Ilha do Mel, +7. x. 1988 +, Zanella, F. C. V, leg., +1 ♀ +( + +MCTP + +); Morretes, +i. 1982 +, A. M. Sakakibara leg., +3 ♂ +, +3 ♀ +( + +DZUP + +); idem: +i. 1984 +, Sakakibara leg., +1 ♀ +( + +DZUP + +); idem: +18. ii. 1985 +, CIIF- luminosa, (no col. data), +1 ♂ +( + +DZUP + +); idem: Viad. Caruru, + +15. xii. +1968, 700 m + +, Pe. J. S. Moure leg., +1 ♀ +( + +DZUP + +); Campo Largo, Est. Faxina km 4, +06. xii. 2002 +, Excursão Fauna Local- DZOO, +2 ♂ +( + +DZUP + +); Alexandra, +10. x. 1970 +, Moure & Mielke, +2 ♀ +( + +DZUP + +); São José dos Pinhais, Rep. Guaricana, + +25 ° 42 ′ 54 ″ S +48 ° 58 ′ 16 ″ W + +, +12–14. xii. 2017 +, Cavichioli & Domahovski, +1 ♀ +( + +DZUP + +); Guaraqueçaba, +13. x. 1983 +, Rupp J. leg., +1 ♀ +( + +DZUP + +); idem: +14. x. 1983 +, Rupp J. leg., +1 ♂ +( + +DZUP + +); idem: +18. ix. 1983 +, Crepaldi S. leg., +1 ♀ +( + +DZUP + +); Serra Negra, +10. iii. 1981 +, Isaias leg., +2 ♀ +( + +DZUP + +); + +Santa Catarina + +: +São Bento do Sul +, Rio Vermelho, + +9. iii. +1974, 850 m + +, Rank leg., +1 ♂ +( + +MCTP + +), +1 ♀ +( + +DZUP + +); + +Rio Grande do Sul + +: São Francisco de Paula, Pró-Mata, +19–21. iii. 1998 +, Carvalho, G. S. leg, +1 ♀ +( + +MCTP + +). + + + + \ No newline at end of file diff --git a/data/BC/F7/46/BCF746E3FC6356F6B3411127F37E1769.xml b/data/BC/F7/46/BCF746E3FC6356F6B3411127F37E1769.xml new file mode 100644 index 00000000000..56059e9bf82 --- /dev/null +++ b/data/BC/F7/46/BCF746E3FC6356F6B3411127F37E1769.xml @@ -0,0 +1,481 @@ + + + +Genetic and multivariate morphometric analyses unveil cryptic diversity in sympatric populations of the Mediterranean wool carder bee, identified as Anthidium undulatum Dours, 1873 (Hymenoptera: Megachilidae: Anthidiini) + + + +Author + +Kasparek, Max +0000-0002-5604-6791 +Mönchhofstr. 16, 69120 Heidelberg, Germany + + + +Author + +Boustani, Mira +Laboratory of Zoology, Institute for Biosciences, University of Mons, Place du Parc 20, 7000 Mons, Belgium + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +75 +92 + + + +journal article +10.3897/asp.83.e137570 +05CB5D16-02E4-4323-9294-45A28971CCC2 + + + + + +Anthidium libanicum +Kasparek + +sp. nov. + + + + +Figures 6 +, +7 + + + + +Type material. + + + +HOLOTYPE +: + +LEBANON + +: male, +North +Lebanon: +Hadath el Jebbe +, +Road +to +Wadi +al +Fouar +( + +34 ° 12 ’ 47.9 ’’ N +35 ° 55 ’ 30.9 ’’ E + +), + +1553 m + +; + +22. viii. 2018 + +; +M. Boustani +leg. ( +CMK +: mbou 249; +ABABX 247-21 +) + +. — +PARATYPES +: + +LEBANON + +: +1 ♂ +; +Beqaa +: Hadath ( + +34 ° 00 ’ 38.1 ’’ N +35 ° 59 ’ 01.1 ’’ E + +), +1441 m +; +7. vii. 2019 +; M. Boustani leg. ( +CMK +: mbou 135; +ABABX +250-21). – +1 ♀ +, North +Lebanon +: Harissa, Al Jawar ( + +34 ° 11 ’ 36.0 ’’ N +35 ° 55 ’ 25, 9 ’’ E + +), +1765 m +; +28. vi. 2017 +; M. Boustani leg. ( +CMK +: mbou 254; +ABABX +058-20). – +4 ♂ +; North +Lebanon +: Tannourine, Harissa El Jawar ( + +34 ° 11 ’ 36.2 ’’ N +35 ° 55 ’ 26.1 ’’ E + +), +1736 m +; +11. vii. 2018 +; M. Boustani & J. Jabbour ( +CMK +: mbou 250-253; +ABABX +421-22, +ABABX +453-22, +ABABX +452-22, +ABABX +451-22). – +1 ♂ +; North +Lebanon +: Tannourine el Tahta, Mar Boutros Church ( + +34 ° 12 ’ 34.8 ’’ N +35 ° 54 ’ 01.3 ’’ E + +), +1207 m +; +27. vi. 2019 +; M. Boustani leg. ( +OLL +: mbou 133: +ABABZ +093-23). – +1 ♂ +; North +Lebanon +: Tannourine El Tahta, Saint Peter Church ( + +34 ° 12 ’ 34.70 ’’ N +35 ° 54 ’ 01.20 ’’ E + +), +1155 m +; +27. vi. 2019 +; X. van Achter leg. (coll. M. Boustani: mbou 132; +ABABX +422-22). – +1 ♂ +; North +Lebanon +: Tannourine, Wadi Ain el Raha ( + +34 ° 12 ’ 34, 5 ’’ N +35 ° 54 ’ 01, 1 ’’ E + +), +1618 m +; +29. vi. 2017 +; M. Boustani leg. ( +CMK +: mbou 247: +ABABZ +092-23). + + + + +Non-type material (examined). + + + +LEBANON + +: +1 ♀ +; +Beqaa +: Jord Aarsal ( + +34 ° 06 ’ 27.0 ’’ N +36 ° 25 ’ 26.4 ’’ E + +), +1975 m +; +11. vii. 2019 +; M. Boustani leg. ( +CMK +: mbou 137; +ABABZ +091-23, DNA not retrieved). – +1 ♂ +; North Lebanon: Tannourine Cedars ( + +34.20 ° N +35.92 ° E + +); +25. vi. 2006 +; N. Nemer leg. ( +CMK +: mbou 248). – +1 ♂ +; North Lebanon: Harissa, Al Jawar ( + +34 ° 11 ’ 36.0 ’’ N +35 ° 55 ’ 25, 9 ’’ E + +), +1765 m +; +28. vi. 2017 +; M. Boustani leg. (coll. M. Boustani: mbou 246). Note: Specimens with barcode sequence have been assigned as +type +material. Specimens not barcoded are classified as non-type material. + + + + +Barcode Index Number. + + + +BOLD +: AEH 0696 + +. + + + + +Diagnosis. + + +The species is characterized in both sexes by a combination of several characters. The body size is on average smaller than in + +A. undulatum + +and + +A. wahrmani + +(average length of the marginal cell 1.67 [1.53–2.10] mm in +11 males +, compared to 2.14 [1.67–2.16] mm in +7 males +of + +A. wahrmani + +and 3.05 [1.46–2.15] mm in +38 males +of + +A. undulatum + +), the color of the body maculations is pale yellow (bright yellow in the other two species), the transition between the black ground color and yellow band on +T +1 and +T +2 is rusty (sometimes additionally with some inconspicuous, blurred rusty maculation within the yellow band) (rusty transition zone absent in the other two species), relatively narrow anterolateral yellow band on scutum (broader in the other two species), and a black antennal scape (red-brown in + +A. wahrmani + +; black or yellow in + +A. undulatum + +). + + +Genetically, + +A. libanicum + +differs from + +A. undulatum + +and + +A. wahrmani + +at seven different nucleotide sites in the DNA sequence of the +COI +gene: 124, 184, 328, 433, 451, 515, 565, and 625 (see Table +3 +for details). + + + + +Description. + + +Female +. +8 mm +. – +Head +: Black with pale yellow clypeus and lower paraocular area (reaching beyond the antennal socket), preoccipital band extending across vertex and malar area and reaching mandibular area; preoccipital band in contact with the eye only in the lower malar area; clypeus slightly convex, with the highest point in the center; densely punctate laterally, less dense and coarser punctures along the midline; anterior margin almost straight, posterior rim straight and smooth; mandible yellow with dark brown teeth; three large teeth and in between them approximately 10 minute teeth (number slightly variable); antennal scape and pedicel black, flagellum mostly reddish brown (darker on the upper side than on the lower side). – +Mesosoma +: Black; pale yellow anterolateral band on scutum; scutellum as seen from above with margin more or less transverse, angled forward toward axillar margin; posterolateral margin transparent lamellate; scutellum posteriorly with pale yellow band; axilla pale yellow. – +Metasoma +: +T +1 – +T +5 with narrow, pale yellow transversal band; broader laterally than medially; +T +6 pale yellow with black outer margin and a black wedge-shaped incision; transition between black ground color and yellow band rusty on +T +1 and +T +2, sometimes additionally with some inconspicuous, blurred rusty maculation on the yellow band; femora and tibiae yellow with some black on the inner face; basitarsi covered with dense felt-like pubescence. + + +Male +. 8–9 (10) mm. – +Head +: Yellow maculation similar to female; yellow maculation of paraocular area reaches middle of upper half of eye; preoccipital band variable, sometimes reaching only upper one third or upper half of eye. – +Mesosoma +: As female. – +Metasoma +: Coloration of +T +1 – +T +5 as in female; apical margin of +T +6 medially protruding; +T +6 with lateral hooked extension; outer margin of +T +7 as seen from above widely rounded with transparent sides and a deep, almost semi-circular emargination. + + +Variation +. The material from +Lebanon +includes +one male +of a light morph. While the general characters agree with those of + +A. libanicum + +, the ground color of the metasomal sterna and terga is dark reddish brown (not black), the color of the anterior vertical side of +T +1 light reddish brown (not dark reddish brown). The body size is significantly larger than the other males examined. It has, e. g., a marginal cell length of +2.2 mm +, as compared to an average length of +1.60 mm +for the remaining specimens (N = 8, only barcoded material). The affiliation of this specimen with + +A. libanicum + +was confirmed by the genetic barcoding sequence. + + + + +Habitat and floral relationships. + + + +Anthidium libanicum + +was collected visiting flowers of + +Ononis natrix + +, + +O. spinosa + +( +Fabaceae +) and + +Stachys distans + +( +Lamiaceae +). The species was collected in both humid Mediterranean and steppic semi-arid habitats (Figs +11 +, +13 +). The localities included clearings of oak and cedar woods, scrublands with karstic outcrops and +Lamiaceae +patches, and degraded roadsides lined with + +Ononis +spp. + +bushes. + + + + +Distribution. + + +Endemic to +Lebanon +. It was found in clearings of oak ( + +1.000 +–1.400 +m + +a. s. l.) and cedar woods ( + +1.400 +–1.800 +m + +a. s. l.) on the western slopes of the Mount +Lebanon +chain, and semi-arid steppes on the eastern slopes of the same mountain chain ( +1.400 m +a. s. l.). It was also found on the peak of the Anti-Lebanon mountain chain near the Syrian border in the Oro-Mediterranean steppe ( +1.900 m +a. s. l.). The lower altitudes were not heavily sampled, and the species could well have a larger distribution than reported (Fig. +2 +). + + + + \ No newline at end of file diff --git a/data/BD/EB/90/BDEB90BF607C533C86B04A8ACF62292D.xml b/data/BD/EB/90/BDEB90BF607C533C86B04A8ACF62292D.xml new file mode 100644 index 00000000000..834b586749e --- /dev/null +++ b/data/BD/EB/90/BDEB90BF607C533C86B04A8ACF62292D.xml @@ -0,0 +1,363 @@ + + + +Taxonomic review, morphometry, and phylogenetic analysis of Mahanarva (Ipiranga) Fennah, 1968 (Hemiptera: Auchenorrhyncha: Cercopidae) + + + +Author + +Meneghetti, Jéssica +0000-0001-6633-6728 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil + + + +Author + +Biganzoli-Rangel, Alejandro Jose +0000-0003-4426-3286 +Department of Biology, York University, Toronto, Ontario, Canada + + + +Author + +Carvalho, Gervásio Silva +0000-0002-3835-8644 +Escola de Ciências da Saúde e da Vida, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga, 6681, 90619 - 900, Porto Alegre, Rio Grande do Sul, Brasil (retired) + + + +Author + +Paladini, Andressa +0000-0001-8894-6092 +Departamento de Ecologia e Evolução, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal de Santa Maria, Av. Roraima, 1000, Cidade Universitária, Bairro Camobi, 97105 - 900, Santa Maria, Rio Grande do Sul, Brasil & Departamento de Zoologia, Laboratório de Sistemática e Biogeografia de Hemiptera, Universidade Federal do Paraná, Av. Cel. Francisco H. dos Santos, 100, Jardim das Américas, 81530 - 000, Curitiba, Paraná, Brasil + +text + + +Arthropod Systematics & Phylogeny + + +2025 + +2025-05-09 + + +83 + + +45 +73 + + + +journal article +10.3897/asp.83.e130043 +2000DF4D-D639-4E57-90EB-590A80CD9A6C + + + + +3.4. 11. + +Mahanarva +( +Ipiranga +) +obliqua + +sp. nov. + + + + +Figures 6 H +, +12 A – L +, +14 A – D + + + + +Measurements male / female. + +Head length: 1.25 / 1.32; head width: 2.25 / 2.33; pronotum length: 2.31 / 2.45; pronotum width: 3.72 / 4.05; tegmen length: 10.3 / 10.9; tegmen width: 3.3 / 3.6; total length: 12.2 / 13.25 (in mm). + + + +Diagnosis. + +Tegmina black with an orange spot near base and two wide, orange, transversal bands, zigzag-like; subgenital plates with a rounded apex, dorsal margin without elevation, paramere with subtriangular apex. + + + +Description. + + +Holotype +. Head black, with a slight pubescence, vertex subretangular with a slightly marked median carina, ocelli closer to each other than to eyes, separated by about one and a half diameter from each other; tylus quadrangular, black, with its basal third with a slightly marked carina, becoming indistinct towards the apex (Fig. +12 A +); antenna black, pedicel sparsely setose; basal body of flagellum subcylindrical with a single and short arista shorter than the pedicel length. Postclypeus black, inflated, with convex profile, longitudinal carina strongly marked and lateral grooves slightly marked (Fig. +12 B, C +); rostrum reddish, barely reaching mesocoxae. Pronotum black, hexagonal, with muscular insertions slightly apparent, median carina slightly marked, anterior and anterolateral margins straight; posterolateral with slight sinuosity and posterior margin medially grooved, humeral angles acute; scutellum black, with slight central concavity and slightly marked horizontal grooves. Tegmina (Fig. +12 A, B +) black, narrow, with an orange spot near base and two wide, orange, transversal bands, zigzag-like, one located between basal and medium third and another between medium and apical third; vein R thickened at base, M and Cu 1 united at base and A 1 and A 2 distincts, apical reticulation developed, prominent venation. Hindwings hyaline, with brown venation, Cu 1 thickened at the base, hamuli with three to four spines. Legs reddish, metathoracic femur with a small apical spine; tibia with two lateral spines, basal one smaller than the ones located at apical crown, which has 12–14 spines arranged in two rows; basitarsus with three irregular rows of spines covered by long and dense setae; subungueal process present. — +MALE: Genitalia +: Pygofer (Fig. +12 D, E +) with one subtriangular process between anal tube and subgenital plate; subgenital plate narrowing toward its extension with apex rounded with denticles on inner surface; basal process of dorsal margin short and rounded. Paramere (Fig. +12 F +) subretangular, with a subtriangular apex, dorsal margin rounded and developed, subapical hook-like spine directed outwards and forward. Aedeagus (Fig. +12 G – I +) subcylindrical, narrowing towards apex, with a pair of straight dorsal processes, shorter than half length of shaft and inserted medially. — +FEMALE +: Morphology similar to that of the male, but slightly larger in relation to total body length. First valvula of ovipositor long (Fig. +12 J, K +), with slightly acute apex, basal process rounded and undeveloped, directed downwards; second valvula long (Fig. +12 L +), with rounded apex and dorsal margin covered by teeth in apical third; third valvula short and wide, with rounded apex, covered ventrally by long setae. + + + + + + + +Mahanarva +( +Ipiranga +) +obliqua + + +sp. nov. + +Habitus: +A +dorsal view; +B +lateral view; +C +ventral view. Male genitalia, pygofer: +D +lateral view; +E +ventral view. Paramere: +F +lateral view. Aedeagus: +G +lateral view; +H +dorsal view; +I +ventral view. Female genitalia, ovipositor: +J +lateral view. First valvulae: +K +lateral view. Second valvulae: +L +lateral view. Scale bars in mm. + + + + + +Remarks. + + +This species was initially considered as a variation of + +M. +( +I. +) +rubicunda + +, however upon closer examination, it became evident that significant differences exist in the morphological characters such as the tegmina pattern and the male genitalia. In fact, + +M. +( +I. +) +obliqua + + +sp. nov. + +presents several morphological similarities regarding other + +Ipiranga + +species: a rounded and developed dorsal margin of paramere, a subapical hook-like paramere spine, and a subcylindrical aedeagus with a pair of straight dorsal processes. + + + + +Etymology. + +In Latin, ‘ obliquum’ means zigzag shape, thus this species name refers to the tegmina pattern with transversal bands. + + + +Distribution. + + +Brazil +( +Paraná +and +Santa Catarina +). + + + + +Material examined. + + +BRASIL +, + +Holotype +♂ +: +Paraná + +: Piraquara, riacho próximo à estrada, ativa, + +25 ° 31 ′ 01 ″ S +49 ° 00 ′ 32 ″ W + +, +909 m +a. s. l, +20. iii. 2024 +, AP Pinto, L. Polizelli, & RC Varella leg. ( + +DZUP + +); + + +Paratypes + + +: same data as +holotype +, +3 ♂ +, +2 ♀ +; idem: +07. xii. 2023 +, AP Pinto, J Ehlert, MUM de Almeida & RC Varella leg., +7 ♂ +, +1 ♀ +( + +DZUP + +); Curitiba, +28. vii. 1983 +, L. L. Leite leg. +1 ♂ +, +1 ♀ +( + +DZUP + +449663); idem, +3. ix. 1983 +, R. Amador leg., +2 ♀ +(1 + +MCTP + +, 1 + +DZUP + +449664); idem, Guajuvira, +8. vi. 1980 +, A. Claret, leg., +1 ♂ +( + +MCTP + +); idem, Quatro Barras, +22. ii. 1969 +, Becker & Laroca, +1 ♀ +( + +DZUP + +449660); idem, Jaguariaíva, (no collect data), M. Linsing leg., +2 ♀ +( + +DZUP + +449667, + +DZUP + +449668); + +Santa Catarina + +: Três Barras, +12. iii. 1995 +, Jorge Cheren leg., +1 ♀ +( + +MCTP + +); +1 ♀ +no data ( + +MCTP + +). + + + + +Adicional material examined. + + +BRASIL +, + +Paraná + +: Piraquara, riacho próximo à estrada, ativa, +25 ° 31 ′ 01 ″ S +49 ° 00 ′ 32 ″ W +, +909 m +a. s. l, +20. iii. 2024 +, AP Pinto, L. Polizelli, & RC Varella leg. +6 ♂ +, +2 ♀ +( + +DZUP + +). + + + + \ No newline at end of file