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<mods:title id="34119A0DA0A6EB1297179F86A1D33411">Hidden diversity of Freyastera (Asteroidea, Brisingida, Freyellidae) at great depth: Description of new species and remarks on species boundaries</mods:title>
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<bibRefCitation id="61C885A2DD3082E7616D44179B952CBE" author="Sladen WP" refId="B61" refString="Sladen WP (1889) Report on the Asteroidea. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876, Zoology 30 (51): 125 - 174." year="1889">Sladen, 1889</bibRefCitation>
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<figureCitation id="5D53475598F67517C773F37B8E38FF34" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">Fig. 11</figureCitation>
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<taxonomicName id="2C3A638AAE04D41A9819C78437424CEE" ID-CoL="6JMW7" authorityName="Sladen" authorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="C563541356DD66A6AB5F4A7300B7F9D6" italics="true">Freyella benthophila</emphasis>
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:
<treatmentCitation id="0AE7006FF5A1F939773E6EEB86C6C8A8" author="Sladen WP" page="641" year="1889">
<bibRefCitation id="A9CC9A76EEFD653FFA192DCBFCA0E984" author="Sladen WP" firstAuthor="Sladen" page="641" refId="B61" refString="Sladen WP (1889) Report on the Asteroidea. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 18731876, Zoology 30 (51): 125174." year="1889">Sladen 1889: 641</bibRefCitation>
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;
<treatmentCitation id="F340CC9FAC14869354D39591BA6DD108" author="Wood-Mason J &amp; Alcock A" page="440" year="1891">
<bibRefCitation id="D6676395B4E10A539D72BA416AFFB300" DOI="10.1080/00222939109459221" author="Wood-Mason J &amp; Alcock A" firstAuthor="Wood-Mason" page="440" refId="B65" refString="Wood-Mason J, Alcock A (1891) LII. Natural history notes from H. M. Indian marine survey Steamer “ Investigator ”. Series II, No. 1. On the results of deep-sea dredging during the season 189091, Annals and Magazine of Natural History 8 (48): 427452. https://doi.org/10.1080/00222939109459221" year="1891">Wood-Mason and Alcock 1891: 440</bibRefCitation>
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;
<treatmentCitation id="444A56D7367D0B89414A2390B3CC2DE3" author="Alcock A" page="121" year="1893">
<bibRefCitation id="3054972F5F320A2828840F70B44349C1" DOI="10.1080/00222939308677478" author="Alcock A" firstAuthor="Alcock" issue="62" journalOrPublisher="An account of the collection of deep-sea Asteroidea, Annals and Magazine of Natural History" page="121" pagination="73-121" refId="B1" refString="Alcock A (1893) XV. — Natural history notes from H. M. Indian marine survey Steamer Investigator, Commander C. F. Oldham, R. N., commanding. — Series II., No. 7. An account of the collection of deep-sea Asteroidea, Annals and Magazine of Natural History 11 (62): 73121. https://doi.org/10.1080/00222939308677478" title="XV. - Natural history notes from H. M. Indian marine survey Steamer Investigator, Commander C. F. Oldham, R. N., commanding. - Series II., No. 7." volume="11" year="1893">Alcock 1893: 121</bibRefCitation>
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<treatmentCitation id="EC5950DBC0A26EA9013C54C06B9CD168" author="Fisher WK" page="24" year="1928">
<bibRefCitation id="9F15E74BD202E0A4BB6680AEAB9E4761" DOI="10.5479/si.03629236.145.1" author="Fisher WK" firstAuthor="Fisher" journalOrPublisher="Bulletin of the United States National Museum" page="24" pagination="1-347" refId="B18" refString="Fisher WK (1928) Asteroidea of the North Pacific and adjacent waters. Part 2. Forcipulata (part). Bulletin of the United States National Museum 76: 1347. https://doi.org/10.5479/si.03629236.145.1" title="Asteroidea of the North Pacific and adjacent waters. Part 2. Forcipulata (part)." volume="76" year="1928">Fisher 1928: 24</bibRefCitation>
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;
<treatmentCitation id="DC9147646DDE945B0D95734621C3D2D2" author="Madsen FJ" page="84" year="1951">
<bibRefCitation id="4FE23894900B6AB08B838CCF077BCA94" author="Madsen FJ" firstAuthor="Madsen" issue="6" journalOrPublisher="Reports of the Swedish Deep-sea Expedition" page="84" pagination="73-92" refId="B38" refString="Madsen FJ (1951) Asteroidea. Reports of the Swedish Deep-sea Expedition 2 (6): 7392." title="Asteroidea." volume="2" year="1951">Madsen 1951: 84</bibRefCitation>
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;
<treatmentCitation id="557F6D28D88DCB9042C0A59965A90868" author="Cherbonnier G &amp; Sibuet M" page="1356" year="1972">
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;
<treatmentCitation id="32B97CE1E230150FB96ADEC958E9FE5F" author="Sibuet M" page="292" year="1975">
<bibRefCitation id="2B26DF3EDF33F38D2A01EF62FDF68535" author="Sibuet M" firstAuthor="Sibuet" issue="289" journalOrPublisher="Bulletin du Muséum National dHistoire Naturelle, Zoologie" page="292" pagination="281-297" refId="B60" refString="Sibuet M (1975) Astérides abyssales de lAtlantique sud. (Résultats de la campagne Walda, juin-juillet-août 1971). Bulletin du Muséum National dHistoire Naturelle, Zoologie 3 (289): 281297. [in French] https://archimer.ifremer.fr/doc/00000/5426/" title="Astérides abyssales de lAtlantique sud. (Résultats de la campagne Walda, juin-juillet-août 1971)." url="https://archimer.ifremer.fr/doc/00000/5426/" volume="3" year="1975">Sibuet 1975: 292</bibRefCitation>
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;
<treatmentCitation id="28B91F7B7C8931686D0D9A1F440D4377" author="Korovchinsky NM &amp; Galkin SV" page="1215" year="1984">
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(in key);
<treatmentCitation id="1C898A1033EAC913E9046BFF1DAD9529" author="Galkin SV &amp; Korovchinsky NM" page="165" year="1984">
<bibRefCitation id="E0259B4AE579F7FC5789A929D2E96417" author="Galkin SV &amp; Korovchinsky NM" firstAuthor="Galkin" journalOrPublisher="Trudy Instituta Okeanologii Akademii Nauk SSSR" page="165" pagination="164-178" refId="B20" refString="Galkin SV, Korovchinsky NM (1984) Vertical and geographical distribution of the starfishes of the genus Freyella (Brisingidae) with some remarks on their ecology and evolution. Trudy Instituta Okeanologii Akademii Nauk SSSR 119: 164178. [in Russian with English abstract]" title="Vertical and geographical distribution of the starfishes of the genus Freyella (Brisingidae) with some remarks on their ecology and evolution." volume="119" year="1984">Galkin and Korovchinsky 1984: 165</bibRefCitation>
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;
<treatmentCitation id="EEBF65DD7DBC8BFF6A33F2D0EA7E6B29" author="Zhang R &amp; Fau M &amp; Mah C &amp; Eléaume M &amp; Zhang D &amp; Zhou Y &amp; Lu B &amp; Wang C" page="17" year="2024">
<bibRefCitation id="A1FA64723AE658B3B47E13F035EC47D1" DOI="10.1016/j.ympev.2023.107993" author="Zhang R &amp; Fau M &amp; Mah C &amp; Eléaume M &amp; Zhang D &amp; Zhou Y &amp; Lu B &amp; Wang C" etAl="et al." firstAuthor="Zhang" page="17" refId="B69" refString="Zhang R, Fau M, Mah C, Eléaume M, Zhang D, Zhou Y, Lu B, Wang C (2024) Systematics of deep-sea starfish order Brisingida (Echinodermata: Asteroidea), with a revised classification and assessments of morphological characters. Molecular Phylogenetics and Evolution 191: 107993. https://doi.org/10.1016/j.ympev.2023.107993" year="2024">Zhang et al. 2024: 17</bibRefCitation>
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.
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<taxonomicName id="6FF705C0C0387DD9251ADE2BB6E8F3CF" ID-CoL="6JMVB" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyellidea" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="B5C0B7D2805D045CAEE55C95625E2FD2" italics="true">Freyellidea benthophila</emphasis>
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<comment id="DD73B0B8B90C0D0DC398BE869B65EA9C">
:
<treatmentCitation id="021E7FED219A1C214F74504547F9AACD" author="Fisher WK" page="429" year="1917">
<bibRefCitation id="88E8FCB76720670056BC7B8F8DDA6AEA" DOI="10.1080/00222931709487030" author="Fisher WK" firstAuthor="Fisher" issue="120" journalOrPublisher="Annals and Magazine of Natural History" page="429" pagination="418-431" refId="B17" refString="Fisher WK (1917) LI. — New genera and species of Brisingidae. Annals and Magazine of Natural History 20 (120): 418431. https://doi.org/10.1080/00222931709487030" title="LI. - New genera and species of Brisingidae." volume="20" year="1917">Fisher 1917: 429</bibRefCitation>
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<taxonomicName id="C6BE173D5A2B646648A3F1AA8FE466B4" ID-CoL="6JNTB" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="B3C0255F75E80FEC91C49C8317BE48B6" italics="true">Freyastera benthophila</emphasis>
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:
<treatmentCitation id="8FF025D9396C03F6B74F8565E325F7AD" author="Downey ME" page="36" year="1986">
<bibRefCitation id="E558285011D41E3A16F75FF6E3ECC38E" DOI="10.5479/si.00810282.435" author="Downey ME" firstAuthor="Downey" journalOrPublisher="Smithsonian Contributions to Zoology" page="36" pagination="1-57" refId="B15" refString="Downey ME (1986) Revision of the Atlantic Brisingida (Echinodermata: Asteroidea), with description of a new genus and family. Smithsonian Contributions to Zoology 435: 157. https://doi.org/10.5479/si.00810282.435" title="Revision of the Atlantic Brisingida (Echinodermata: Asteroidea), with description of a new genus and family." volume="435" year="1986">Downey 1986: 36</bibRefCitation>
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<treatmentCitation id="B0BA42781DC17AFE6FFCC9142881CF59" author="McKnight DG" page="173" year="1993">
<bibRefCitation id="D232BC0A6B0F231F64CC5950EFEDEC79" DOI="10.1080/03014223.1993.10422858" author="McKnight DG" firstAuthor="McKnight" issue="3" journalOrPublisher="New Zealand Journal of Zoology" page="173" pagination="165-190" refId="B43" refString="McKnight DG (1993) Records of echinoderms (excluding holothurians) from the Norfolk Ridge and Three Kings Rise north of New Zealand. New Zealand Journal of Zoology 20 (3): 165190. https://doi.org/10.1080/03014223.1993.10422858" title="Records of echinoderms (excluding holothurians) from the Norfolk Ridge and Three Kings Rise north of New Zealand." volume="20" year="1993">McKnight 1993: 173</bibRefCitation>
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<treatmentCitation id="F626251ACE7A77685CBCB19A899D7CD4" author="Mah CL" page="78" year="1998">
<bibRefCitation id="908E6BCBA7FE1ABF926FF6B50C4F7518" author="Mah CL" firstAuthor="Mah" page="78" refId="B40" refString="Mah CL (1998) A phylogeny, taxonomic revision, and natural history of the order Brisingida (Asteroidea). Masters Thesis, San Francisco State University, San Francisco, US." year="1998">Mah 1998: 78</bibRefCitation>
</treatmentCitation>
; Mah in
<treatmentCitation id="91059725D4CEF818A62C8C22C3A23EBC" author="Clark AM &amp; Mah C" page="318" year="2001">
<bibRefCitation id="C0B3CFD01B8DAC1CE108B38231167439" author="Clark AM &amp; Mah C" firstAuthor="Clark" journalOrPublisher="Echinoderm studies" page="318" pagination="229-347" refId="B11" refString="Clark AM, Mah C (2001) An index of names of recent Asteroidea — Part 4: Forcipulatida and Brisingida. Echinoderm studies 6: 229347." title="An index of names of recent Asteroidea - Part 4: Forcipulatida and Brisingida." volume="6" year="2001">Clark and Mah 2001: 318</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="D106981D77C582794381B55F16EC76B2" author="Dilman AB" page="38" year="2014">
<bibRefCitation id="874486DBAA4B24A377DE0EB23AE767FE" DOI="10.15298/invertzool.11.1.05" author="Dilman AB" firstAuthor="Dilman" issue="1" journalOrPublisher="Invertebrate Zoology" page="38" pagination="25-42" refId="B13" refString="Dilman AB (2014) Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Asteroidea. Invertebrate Zoology 11 (1): 2542. https://doi.org/10.15298/invertzool.11.1.05" title="Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Asteroidea." volume="11" year="2014">Dilman 2014: 38</bibRefCitation>
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;
<treatmentCitation id="A12F551B364AC4511404CD0A39A22FEC" author="Moreau CV &amp; Aguera A &amp; Jossart Q &amp; Danis B" page="16" year="2015">
<bibRefCitation id="BA0D2E3202564942C3F8CF9D95E12FE5" DOI="10.3897/BDJ.3.e7062" author="Moreau CV &amp; Aguera A &amp; Jossart Q &amp; Danis B" etAl="et al." firstAuthor="Moreau" page="16" refId="B47" refString="Moreau CV, Aguera A, Jossart Q, Danis B (2015) Southern Ocean Asteroidea: a proposed update for the Register of Antarctic Marine Species. Biodiversity Data Journal 3: e 7062. https://doi.org/10.3897/BDJ.3.e7062" year="2015">Moreau et al. 2015: 16</bibRefCitation>
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;
<treatmentCitation id="060B453BE89B35DCEE29F880A5B3E41E" author="Moreau C &amp; Mah C &amp; Agüera A &amp; Améziane N &amp; Barnes D &amp; Crokaert G &amp; Eléaume M &amp; Griffiths H &amp; Guillaumot C &amp; Hemery LG &amp; Jażdżewska A." page="147" year="2018">
<bibRefCitation id="34E9D9E67CF8B3696979C972309CF39B" DOI="10.3897/zookeys.747.22751" author="Moreau C &amp; Mah C &amp; Agüera A &amp; Améziane N &amp; Barnes D &amp; Crokaert G &amp; Eléaume M &amp; Griffiths H &amp; Guillaumot C &amp; Hemery LG &amp; Jażdżewska A." etAl="et al." firstAuthor="Moreau" journalOrPublisher="ZooKeys" page="147" pagination="141-156" refId="B48" refString="Moreau C, Mah C, Agüera A, Améziane N, Barnes D, Crokaert G, Eléaume M, Griffiths H, Guillaumot C, Hemery LG, Jażdżewska A. (2018) Antarctic and sub-Antarctic Asteroidea database. ZooKeys 747: 141156. https://doi.org/10.3897/zookeys.747.22751" title="Antarctic and sub-Antarctic Asteroidea database." volume="747" year="2018">Moreau et al. 2018: 147</bibRefCitation>
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<bibRefCitation id="E4754E98D810631CA988AE5EF29C92BA" DOI="10.1016/j.dsr.2019.103163" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" etAl="et al." firstAuthor="Zhang" page="7" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al. 2019: 7</bibRefCitation>
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(in key).
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</paragraph>
<paragraph id="46B8BA2707E2E4AE1F79CB38503139E2">
<treatmentCitationGroup id="FBBFD9C50C62E2347D64A26A31BD8A4A">
<taxonomicName id="D9A558244EB9E2E379310CE99EFCB29B" ID-CoL="L9YD" authorityName="Ludwig" authorityYear="1903" class="Asteroidea" family="Freyellidae" genus="Belgicella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="racowitzana">
<emphasis id="CD596ABC0642744C387D2D84F7DBA9A4" italics="true">Belgicella racowitzana</emphasis>
</taxonomicName>
<comment id="60E5C6594317C624434C8FD236EB1EAF">
:
<treatmentCitation id="6C628D62D3C0252EC1EC86281085CC26" author="Ludwig H" page="59" year="1903">
<bibRefCitation id="624137B222149B3DD7942865C3C8BA9E" author="Ludwig H" firstAuthor="Ludwig" page="59" refId="B37" refString="Ludwig H (1903) Seesterne. Résultats du voyage du S. Y. Belgica en 1897-1898 - 1899. Rapports scientifiques, 172. [in Germany]" year="1903">Ludwig 1903: 59</bibRefCitation>
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<bibRefCitation id="E0C49E66AD01A19CD2A179F1B670069C" author="Koehler R" firstAuthor="Koehler" issue="6" journalOrPublisher="Zoologischer Anzeiger" page="141" pagination="140-147" refId="B30" refString="Koehler R (1907) Asteries, Ophiures et Echinides recueillis dans les mers australes par la “ Scotia ” (19021904). Zoologischer Anzeiger 32 (6): 140147. [in French]" title="Asteries, Ophiures et Echinides recueillis dans les mers australes par la “ Scotia ” (1902-1904)." volume="32" year="1907">Koehler 1907: 141</bibRefCitation>
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;
<treatmentCitation id="89CFF3E7C1552770151288F0148D0517" author="Koehler R" page="245" year="1908">
<bibRefCitation id="62D9F330C9E35C914655D7671222E526" author="Koehler R" firstAuthor="Koehler" page="245" refId="B31" refString="Koehler R (1908) Astéries, Ophiures et Echinides de lExpedition Antarctique National Ecossaise. Scottish National Antarctic Expedition. Report on the scientific results of the voyage of S. Y. Scotia during the years 1902, 1903, and 1904 5 (13): 193313. [in French]" year="1908">Koehler 1908: 245</bibRefCitation>
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;
<treatmentCitation id="848C9FC22A7EF4A45433FF6083A93CF9" author="Döderlein L" page="293" year="1928">
<bibRefCitation id="F83AF2E2DEB801C176A980EE0D8CE9C7" author="Döderlein L" firstAuthor="Doderlein" page="293" refId="B14" refString="Döderlein L (1928) Die seesterne der Deutschen Sudpolar-Expedition, 19011903. Deutsche Sudpolar Expediiton XIX. Zoologie XI 1901-1903 19 (11): 289301. [in Germany]" year="1928">Döderlein 1928: 293</bibRefCitation>
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;
<treatmentCitation id="B5DEDE0D3E2C31BAEB74DDEE0773D6C2" author="Fisher WK" page="6" year="1928">
<bibRefCitation id="47DFB3FCB42EAA49431958F2D78A90EF" DOI="10.5479/si.03629236.145.1" author="Fisher WK" firstAuthor="Fisher" journalOrPublisher="Bulletin of the United States National Museum" page="6" pagination="1-347" refId="B18" refString="Fisher WK (1928) Asteroidea of the North Pacific and adjacent waters. Part 2. Forcipulata (part). Bulletin of the United States National Museum 76: 1347. https://doi.org/10.5479/si.03629236.145.1" title="Asteroidea of the North Pacific and adjacent waters. Part 2. Forcipulata (part)." volume="76" year="1928">Fisher 1928: 6</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="50BDE28C506959566598A96DB3112BBD" author="Fisher WK" page="75" year="1940">
<bibRefCitation id="8D4F4CFF9FC6ECDED21C912B8A2EA490" author="Fisher WK" firstAuthor="Fisher" journalOrPublisher="Discovery Reports" page="75" pagination="69-306" refId="B19" refString="Fisher WK (1940) Asteroidea. Discovery Reports 20: 69306." title="Asteroidea." volume="20" year="1940">Fisher 1940: 75</bibRefCitation>
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; A. M.
<treatmentCitation id="7D238F844A8903AC3C501880DF51B281" author="Clark AM" page="68" year="1962">
<bibRefCitation id="A71D0A1D9D000C92F8FFF2EBA2D9D211" author="Clark AM" firstAuthor="Clark" journalOrPublisher="Series B, Zoology and botany, Vol. IX" page="68" refId="B9" refString="Clark AM (1962) Asteroidea. Reports (B. A. N. Z. Antarctic Research Expedition, (19291931 )). Series B, Zoology and botany, Vol. IX, 104 pp." title="Asteroidea. Reports (B. A. N. Z. Antarctic Research Expedition, (1929-1931 ))." year="1962">Clark 1962: 68</bibRefCitation>
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;
<treatmentCitation id="FBBEA340972CC46FDF7EAA104DBB4C39" author="Jangoux M &amp; Massin C" page="91" year="1986">
<bibRefCitation id="1CF9A1BF54743560FD4EF3947C162AD1" author="Jangoux M &amp; Massin C" firstAuthor="Jangoux" journalOrPublisher="Bulletin de lInstitut Royal des Sciences Naturelles de Belgique, Biologie" page="91" pagination="83-97" refId="B26" refString="Jangoux M, Massin C (1986) Catalogue commenté des types dEchinodermes actuels conservés dans les collections nationales belges. Bulletin de lInstitut Royal des Sciences Naturelles de Belgique, Biologie 56: 8397." title="Catalogue commenté des types dEchinodermes actuels conservés dans les collections nationales belges." volume="56" year="1986">Jangoux and Massin 1986: 91</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="9B3D865A9D280381CF17E49FE931CB2B" author="Mah CL" page="78" year="1998">
<bibRefCitation id="5DE794913309144CCD09FCD7C86AF74E" author="Mah CL" firstAuthor="Mah" page="78" refId="B40" refString="Mah CL (1998) A phylogeny, taxonomic revision, and natural history of the order Brisingida (Asteroidea). Masters Thesis, San Francisco State University, San Francisco, US." year="1998">Mah 1998: 78</bibRefCitation>
</treatmentCitation>
; Mah in
<treatmentCitation id="879875B6867C6D757942761A0648350A" author="Clark AM &amp; Mah C" page="317" year="2001">
<bibRefCitation id="8B90EE34210408B9A9AC940044C93984" author="Clark AM &amp; Mah C" firstAuthor="Clark" journalOrPublisher="Echinoderm studies" page="317" pagination="229-347" refId="B11" refString="Clark AM, Mah C (2001) An index of names of recent Asteroidea — Part 4: Forcipulatida and Brisingida. Echinoderm studies 6: 229347." title="An index of names of recent Asteroidea - Part 4: Forcipulatida and Brisingida." volume="6" year="2001">Clark and Mah 2001: 317</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="41D2838BE450D35AAE7D9FE3D3728B63" author="Moreau CV &amp; Aguera A &amp; Jossart Q &amp; Danis B" page="5" year="2015">
<bibRefCitation id="747F612768F979701808DFA564B82B23" DOI="10.3897/BDJ.3.e7062" author="Moreau CV &amp; Aguera A &amp; Jossart Q &amp; Danis B" etAl="et al." firstAuthor="Moreau" page="5" refId="B47" refString="Moreau CV, Aguera A, Jossart Q, Danis B (2015) Southern Ocean Asteroidea: a proposed update for the Register of Antarctic Marine Species. Biodiversity Data Journal 3: e 7062. https://doi.org/10.3897/BDJ.3.e7062" year="2015">Moreau et al. 2015: 5</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="486216C44BC9D381FD7AA857AF3ACDF2" author="Moreau C &amp; Mah C &amp; Agüera A &amp; Améziane N &amp; Barnes D &amp; Crokaert G &amp; Eléaume M &amp; Griffiths H &amp; Guillaumot C &amp; Hemery LG &amp; Jażdżewska A." page="147" year="2018">
<bibRefCitation id="FF7E878DD1AF084175B3E1D72943F280" DOI="10.3897/zookeys.747.22751" author="Moreau C &amp; Mah C &amp; Agüera A &amp; Améziane N &amp; Barnes D &amp; Crokaert G &amp; Eléaume M &amp; Griffiths H &amp; Guillaumot C &amp; Hemery LG &amp; Jażdżewska A." etAl="et al." firstAuthor="Moreau" journalOrPublisher="ZooKeys" page="147" pagination="141-156" refId="B48" refString="Moreau C, Mah C, Agüera A, Améziane N, Barnes D, Crokaert G, Eléaume M, Griffiths H, Guillaumot C, Hemery LG, Jażdżewska A. (2018) Antarctic and sub-Antarctic Asteroidea database. ZooKeys 747: 141156. https://doi.org/10.3897/zookeys.747.22751" title="Antarctic and sub-Antarctic Asteroidea database." volume="747" year="2018">Moreau et al. 2018: 147</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3MBI" type="material">
<paragraph id="7A1293C9FCF2FCF55DEF01603B8449F7">
<heading id="AE5FC0CA6F80E3E8AA40F2A8AC1087BF" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="09B70ABB2BBAFF14271262D95B15BE56">
<materialsCitation id="2551CE5310E6EAF69851CEE5BE3E48DA" collectionCode="NHMUK" specimenCode="NHMUK 1890.5.7.1078" specimenCount="1" typeStatus="holotype">
<specimenCode id="1951F6373A8A4A7034AD6EA37E0A0B04">
<named-content id="9521337229E85C25EB15BDFDF01F21D4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum">NHMUK</named-content>
1890.5.7.1078
</specimenCode>
(
<emphasis id="C9F04889517DE1E4B2492120C1BE80C5" bold="true" italics="true">
<typeStatus id="3771B0539E9B47A719E17E8584C84FDE" type="holotype">holotype</typeStatus>
</emphasis>
) (Fig.
<figureCitation id="3BCDC3FE3CDAE5A8BF1C88DE7FD5FE32" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">11 A, B, E, G</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="F16D9023316F0A2A414F513899AC14B6" ID-DOI="10.3897/zse.101.144918.figure11" ID-arpha="19358642-E36E-5ACF-A3DF-400F1352B9A0" httpUri="https://binary.pensoft.net/fig/1304662" startId="F11">
<paragraph id="08A9BE488BFEC8D5356E91FB0F25FD6B">
<label id="C84CC8A76C3D6783FE99AB7C6F13AEE5">Figure 11.</label>
</paragraph>
<paragraph id="DF5679411E216EB422A040D95FDCB1CC">
<taxonomicName id="A0F8AFC81F13CE9E85BDFEBE3BFEA4E8" authority="Sladen, 1889" authorityName="Sladen" authorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="F3046B8565672D98237029A638B24D3D" italics="true">Freyastera benthophila</emphasis>
<bibRefCitation id="AAFAAF0C90A9E55B26254A90C4DD7AC8" author="Sladen WP" refId="B61" refString="Sladen WP (1889) Report on the Asteroidea. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876, Zoology 30 (51): 125 - 174." year="1889">Sladen, 1889</bibRefCitation>
</taxonomicName>
.
<emphasis id="9FDA37508CDEE9027D7A300E3AC4E3A1" bold="true">A, B, F, G.</emphasis>
Holotype
<specimenCode id="1FB1655D72D36F3463981D9315709BBD">
<named-content id="DD3A442166C6D766D868C3B622FE42D3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum">NHMUK</named-content>
1890.5.7.1078
</specimenCode>
;
<emphasis id="61446F445168BC4E1096BC6819A193AC" bold="true">C, F, H, I.</emphasis>
NHM 9237;
<emphasis id="A5EB48A70AEECE9497AD3EB59EB7B029" bold="true">D.</emphasis>
<specimenCode id="0F08CC2E7F08CBBAFB78C988AF2DA5F3">
<named-content id="A6E9949BE37EBB9AEC17B40670A1CEE1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">MNHN</named-content>
-IE-2022-2192
</specimenCode>
.
<emphasis id="C411172BB14065E89F913A2AA760EF86" bold="true">A D.</emphasis>
Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates;
<emphasis id="6AAF30080648CDE6F807EC4037584E9D" bold="true">E.</emphasis>
Actinal view of the specimen;
<emphasis id="11178B11AB9E9FA362FB829E96E1CD34" bold="true" italics="true">F.</emphasis>
Actinal view of disk;
<emphasis id="D530D8B54A4EF61A1278DD2B429B8714" bold="true">G.</emphasis>
Abactinal view of arm genital area, spines mostly lost;
<emphasis id="1D680E04C17A6119D57131FF722A8CA6" bold="true">H.</emphasis>
Actinal view of arm genital area;
<emphasis id="528C9B7862CA92D696A691975C99A554" bold="true">I.</emphasis>
Distal part of arm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EEPBI" type="material">
<paragraph id="62D488DCD7D7D00AAE956219A35EE9FE">
<heading id="1298A78649A999ADC1E98454355807FC" reason="title">Other material.</heading>
</paragraph>
<paragraph id="F1B71A9D317F46E8CA6C45526F35C493">
IA 3-2168;
<specimenCode id="BB6D81D1422557C16FED92EB5A7DA60C">
<named-content id="3B24C4ED5887D004C787BFBB530BD9AC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="269BC4A3180FD7B93F23B36FB8A20CA2" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2008-1057
</specimenCode>
;
<specimenCode id="9D14AA20386AB655025926DDD76525FC">
<named-content id="3D9A5839C67F8B9D934E4537DC96706C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="6656E9841457F9891D94C8F37C606CAA" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2008-1073
</specimenCode>
;
<specimenCode id="3E6908B7A290473E40D4716077395919">
<named-content id="4792544C95CCCC83C5F071BE26BEF5FF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="8F7024A0564D1C7211EBDEA83F793CDE" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2008-1081
</specimenCode>
;
<specimenCode id="802D5D967459A2A96D2E14DB64556540">
<named-content id="3297990A9214BCD1353412A5F3D32B42" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="081D1CFB13D043F2C86F6150FE8F146E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2017-1829
</specimenCode>
;
<specimenCode id="26BB32CD20B5AFC535BEC82CFE73C264">
<named-content id="8A586A46F28226E378457EBE78503C43" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="D76204BF10D04403B284944BE4F5DFA6" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2017-1948
</specimenCode>
;
<specimenCode id="3D41D52E1B88705388B55AB3B7586690">
<named-content id="03E3FA72C6785FDD97F4A320C9632141" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="6F82A2B03D68B1AB0930B425FE8D88E7" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2019-5752
</specimenCode>
;
<specimenCode id="0E0E4C6AD5064C82C2100281C12E05A0">
<named-content id="50E8CEC73528F948671562399C8995CD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="823A1C5DAA2694C88839E2C19242F65F" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2022-2192
</specimenCode>
(Fig.
<figureCitation id="2BA9CEA31EFDBD7D237E28902FB83704" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">11 D</figureCitation>
);
<specimenCode id="2DFD527B648C074B4DDBC5BB7EEDCB1F">
<named-content id="E1D46D473CD34AF85BDDD3647DBECEE2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="4A3842A7DCDA31DB91EAA365EADE29FA" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2022-2193
</specimenCode>
;
<specimenCode id="7D48D4F700EB4BDD32258A94E362BE07">
<named-content id="398C82EDEB0F3C1A63DB29A5359C82A6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="E12FABEB635CA3D3F579F2B2663DCBA2" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2022-2194
</specimenCode>
;
<specimenCode id="37072A7288167DFB6F5AF19A7490E34E">
<named-content id="C4016B24E7C2C72DA423886BDA5103C8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="29E03BA9E0A144FCB828867663197798" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2022-2195
</specimenCode>
;
<specimenCode id="A6222F24C5CC8B873FA94E586183C9E3">
<named-content id="A34520E041BD70DAA751DCD333F420E0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="D26A454DF014F2151CE12E380EC59A94" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
-IE-2022-2196
</specimenCode>
;
<specimenCode id="43C25846E855BDD3C77F9014303B300F">
<named-content id="9BDE934649994D945312AD2395CEB8E8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum">
<collectionCode id="5F42C0BC9C57FA7953C32597006C30FC" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
9237
</specimenCode>
(Fig.
<figureCitation id="28EEC3D6562BC8B29F372265A06EB3A7" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">11 C, F, H, I</figureCitation>
);
<abbrev id="ABBRID0ECRBI" xlink_title="National Oceanography Centre">NOC</abbrev>
11908 # 44;
<abbrev id="ABBRID0EGRBI" xlink_title="National Oceanography Centre">NOC</abbrev>
12930 # 64;
<abbrev id="ABBRID0EKRBI" xlink_title="National Oceanography Centre">NOC</abbrev>
12930 # 78;
<abbrev id="ABBRID0EORBI" xlink_title="National Oceanography Centre">NOC</abbrev>
13200 # 27;
<abbrev id="ABBRID0ESRBI" xlink_title="National Oceanography Centre">NOC</abbrev>
13200 # 60;
<abbrev id="ABBRID0EWRBI" xlink_title="National Oceanography Centre">NOC</abbrev>
52701 # 42;
<abbrev id="ABBRID0E1RBI" xlink_title="National Oceanography Centre">NOC</abbrev>
54901 # 7;
<abbrev id="ABBRID0E5RBI" xlink_title="National Oceanography Centre">NOC</abbrev>
54902 # 1;
<abbrev id="ABBRID0ECSBI" xlink_title="National Oceanography Centre">NOC</abbrev>
54903 # 1;
<specimenCode id="FDF77845BBCBE61C96D38DDF2BFF4E44">SO 268-1_29_014</specimenCode>
;
<specimenCode id="5D607BE6CB2569CB4792C327589FAA64">SO 268-2_109_189</specimenCode>
; SO 268-02 _ 99 _ 04. (Suppl. material
<xref id="DDE121F241BF507B6EA695D7915A258A" ref-type="supplementary-material" rid="S1">1</xref>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKSBI" type="diagnosis">
<paragraph id="C5167EC0995B7D2656847BC927560879">
<heading id="00DAD739DB4BAF37FF5E63F2568BF46B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="1CB9D294B91F3A0207D02016AF48D62A">
Arms six. Abactinal armature of disk with one primary central plate at center of the disk and a primary interradial plate at each interradius (Fig.
<figureCitation id="1A741F8654B37105324AEEC17BB6E4F1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">11 A D</figureCitation>
). Central and interradial primary plates shield-like, much larger than the rest of abactinal disk plates (primary plates present in most specimens examined, but indistinguishable in some individuals). Abactinal disk and arm plates evenly covered with numerous short spinelets and small pedicellariae (Fig.
<figureCitation id="79A70D31469BB148DC3D266B9EB7901B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">11 G</figureCitation>
). Beyond genital area, abactinal arm with bands of small pedicellariae. Adambulacral plate with one subambulacral spine. Proximal subambulacral spines with modified tips (truncated or flattened bifurcated) (Fig.
<figureCitation id="862CF0A3F6D871F9A42B77FEF3685BBA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">11 E, H</figureCitation>
). Furrow spine absent. Oral plate with 23 actinostomal spines and one suboral spine, each covered with membranous sheath, but the tips are bare (Fig.
<figureCitation id="B5882629CDEED7BA25DA6409B8FBAB58" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">11 E, F</figureCitation>
). Spines covered with small pedicellariae. Large pedicellariae absent. Inferomarginal plates and lateral spines occur at alternate adambulacral plates beyond genital area (Fig.
<figureCitation id="65E83C567E908232CD6553B4C40110FD" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Freyastera benthophila Sladen, 1889. A, B, F, G. Holotype NHMUK 1890.5. 7.1078; C, F, H, I. NHM 9237; D. MNHN - IE- 2022-2192. A D. Abactinal view of disk, red arrows show the primary interradial plates, yellow arrows show the primary central plates; E. Actinal view of the specimen; F. Actinal view of disk; G. Abactinal view of arm genital area, spines mostly lost; H. Actinal view of arm genital area; I. Distal part of arm." figureDoi="10.3897/zse.101.144918.figure11" httpUri="https://binary.pensoft.net/fig/1304662">11 I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EETBI" type="distribution">
<paragraph id="F74E3D8CDD983FC86023DE5DF2119124">
<heading id="4F69239BC36E48A7D14A1443D8EED6D7" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="1C4A3E1C0B3D2FD372671F565F4D89DB">
North Atlantic, Southern Pacific, Eastern Pacific, Southern Ocean.
<quantity id="451CCAFF9B287CB01118269B5AE497BC" metricMagnitude="3" metricUnit="m" metricValue="3.725" metricValueMax="5.0" metricValueMin="2.45" unit="m" value="3725.0" valueMax="5000.0" valueMin="2450.0">24505000 m</quantity>
. Type locality: Southern Pacific,
<quantity id="AA0E4CD7CADB159D9AEE8B76EC3DA1DE" metricMagnitude="3" metricUnit="m" metricValue="4.663" unit="m" value="4663.0">4663 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJTBI" type="remarks">
<paragraph id="8BFE485A644BCB1A7D1146895B11C9AB">
<heading id="3A574FEA6C29994E20B8988BF23B569D" reason="title">
Remarks on
<taxonomicName id="3740412700A1851C5C6FE910844851C0" authorityName="Sladen" authorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="3E64A29691EF081665BB9D8FAF0D98AB" italics="true">F. benthophila</emphasis>
</taxonomicName>
and other six-armed
<taxonomicName id="687CB986AE4A1D4342C591801C0EC092" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="B905055EDCD22C0239E9E7DBF1E9F945" italics="true">Freyella</emphasis>
</taxonomicName>
.
</heading>
</paragraph>
<paragraph id="1FF65BC90078F5105272EDAFC5344A52">
<taxonomicName id="22514342747551C9713095704E11C429" authorityName="Sladen" authorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="1D4610C3FD82814101751E9444EE67CA" italics="true">F. benthophila</emphasis>
</taxonomicName>
stands in a peculiar position by having shared morphological characters with both
<taxonomicName id="9AD4A4C7D8B503E8B6F52D35E6255A07" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="3D82D0FB941211454B06E9AFB9F556EE" italics="true">Freyastera</emphasis>
</taxonomicName>
and
<taxonomicName id="6CB32378D037775E8F2C21EEF76DFFA7" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="256188BAEBEFA3DF31768902976AB9BD" italics="true">Freyella</emphasis>
</taxonomicName>
. As a species with six arms, it is readily distinguished from other
<taxonomicName id="3981FDD4D1DAA7C5E303B3979CABE711" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="FD6171B3F4C88888891A9B8C0A9CAFE5" italics="true">Freyella</emphasis>
</taxonomicName>
species with more than six arms. Compared with other
<taxonomicName id="4AA8550FCAF28E2751214B123A0B5DC7" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="C52C39410B25615FFF5DD25DD125E775" italics="true">Freyastera</emphasis>
</taxonomicName>
, it is differentiated from most species by having many small spinelets on its abactinal disk and arms and distinguished from
<taxonomicName id="12BF3A5AB9E983022A069146D3A51637" authorityName="Madsen" authorityYear="1956" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="mortenseni">
<emphasis id="01D6E2F4AA0B65CFF357474C3884EBAA" italics="true">Freyastera mortenseni</emphasis>
</taxonomicName>
,
<taxonomicName id="A8DB6685E5AD5297F1236091F0427E64" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="delicata">
<emphasis id="86F590C83741930EEE2C6FC09DC13D7E" italics="true">F. delicata</emphasis>
</taxonomicName>
, and
<taxonomicName id="29D4B9EE3A4CE2FB8A64CD322115828B" baseAuthorityName="A. H. Clark" baseAuthorityYear="1939" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="mexicana">
<emphasis id="EA9E11AA7807F5FDC20B9554F6DFB29D" italics="true">F. mexicana</emphasis>
</taxonomicName>
, which also bears multiple spines on abactinal plates, by the absence of furrow spines and more distantly located inferomarginal plates. Phylogenetic analysis put it within the
<taxonomicName id="0FBA9EA49985ECD9527D435851102B58" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="164B0CCDBE12B7F1439C153957ADE818" italics="true">Freyella</emphasis>
</taxonomicName>
clade (Fig.
<figureCitation id="08FB0E6B540B08B5A1F7A0F809767B71" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Phylogenetic tree of Freyastera using COI, 16 S, 12 S, and 28 S concatenated dataset. Tree topology follows that of the ML tree. Numbers at each node indicate UFBS / PP support values." figureDoi="10.3897/zse.101.144918.figure12" httpUri="https://binary.pensoft.net/fig/1304663">12</figureCitation>
;
<bibRefCitation id="90D2F3A7636BE27D03E6BE016600713D" DOI="10.1016/j.ympev.2023.107993" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B69" refString="Zhang R, Fau M, Mah C, Eléaume M, Zhang D, Zhou Y, Lu B, Wang C (2024) Systematics of deep-sea starfish order Brisingida (Echinodermata: Asteroidea), with a revised classification and assessments of morphological characters. Molecular Phylogenetics and Evolution 191: 107993. https://doi.org/10.1016/j.ympev.2023.107993" year="2024">Zhang et al. 2024</bibRefCitation>
), indicating that having six arms is not an autapomorphy of the
<taxonomicName id="68F5310A9E068C650BCF6F9A94FE0564" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="1553E0EB5A82F31E07201B47401B4127" italics="true">Freyastera</emphasis>
</taxonomicName>
clade.
</paragraph>
<caption id="2A321641FDCEBDAA08C3F904F3140433" ID-DOI="10.3897/zse.101.144918.figure12" ID-arpha="035C5477-75E0-5C32-B86E-A2358F1E55FA" httpUri="https://binary.pensoft.net/fig/1304663" startId="F12">
<paragraph id="B63AB63CA12F4B1A93E669B242300DB6">
<label id="C7AD8AE7CF177F734055731FF074C408">Figure 12.</label>
</paragraph>
<paragraph id="9B12F64EC72528034C1B9E71A381AF16">
Phylogenetic tree of
<taxonomicName id="E269FDC86F45E3411AD0B00A0ECB9F47" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="BF6DFE6C680284B273918D516E31B763" italics="true">Freyastera</emphasis>
</taxonomicName>
using
<abbrev id="ABBRID0EPXBI" xlink_title="c oxidase subunit I">COI</abbrev>
,
<abbrev id="ABBRID0ETXBI" xlink_title="mitochondrial 16S ribosomal RNA">16 S</abbrev>
,
<abbrev id="ABBRID0EXXBI" xlink_title="mitochondrial 12S ribosomal RNA">12 S</abbrev>
, and
<abbrev id="ABBRID0E2XBI" xlink_title="nuclear 28S ribosomal RNA">28 S</abbrev>
concatenated dataset. Tree topology follows that of the
<abbrev id="ABBRID0E6XBI" xlink_title="maximum likelihood">ML</abbrev>
tree. Numbers at each node indicate UFBS / PP support values.
</paragraph>
</caption>
<paragraph id="F750F1C5950120E348D1D22339D482B8">
There are three other six-armed
<taxonomicName id="711249B3E2745A34D941FC5BEDBFB14B" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="4A9AD46D98DA97478D2B2D38D605DA9F" italics="true">Freyella</emphasis>
</taxonomicName>
species whose taxonomic position needs to be re-evaluated, including
<taxonomicName id="93BBAB5E2EC6757AF3F86307C62E4B45" authority="H. L. Clark, 1920" authorityName="H. L. Clark" authorityYear="1920" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="oligobrachia">
<emphasis id="E2B460DB23A16A763E9DB716BA1C99EE" italics="true">F. oligobrachia</emphasis>
H. L.
<bibRefCitation id="87363EBCD60CC13B71762020049C4A69" author="Clark HL" issue="3" journalOrPublisher="Memoirs of the Museum of Comparative Zoology" pagination="70-154" refId="B7" refString="Clark HL (1920) Reports on the scientific results of the expedition to the Eastern Tropical Pacific, in charge of Alexander Agassiz, by the US Fish commission Steamer Albatross, from October 1905 to March 1905, Lt. Cmdr. L. M. Garrett, USN, Commanding. XXXII. Asteroidea. Memoirs of the Museum of Comparative Zoology 39 (3): 70-154." title="Reports on the scientific results of the expedition to the Eastern Tropical Pacific, in charge of Alexander Agassiz, by the US Fish commission Steamer Albatross, from October 1905 to March 1905, Lt. Cmdr. L. M. Garrett, USN, Commanding. XXXII. Asteroidea." volume="39" year="1920">Clark, 1920</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="0940BA15EF3607D9B15CA60278108A52" authority="Baranova, 1957" authorityName="Baranova" authorityYear="1957" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="hexactis">
<emphasis id="B4FD21D4F0BD2D0D8D767CAE2D55F40C" italics="true">F. hexactis</emphasis>
<bibRefCitation id="0A566A8F86624B5D0A784D33F77AD3D6" author="Baranova ZI" journalOrPublisher="Investigations of Far-East Seas" pagination="149-266" refId="B3" refString="Baranova ZI (1957) Echinoderms of the Bering Sea. Investigations of Far-East Seas 4: 149-266. [in Russian]" title="Echinoderms of the Bering Sea." volume="4" year="1957">Baranova, 1957</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="B97C00A9ED1258187B02CDFC09E2FE49" authority="Korovchinsky &amp; Galkin, 1984" authorityName="Korovchinsky &amp; Galkin" authorityYear="1984" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="vitjazi">
<emphasis id="C8A78705D697CAF4BC85282CD8585667" italics="true">F. vitjazi</emphasis>
<bibRefCitation id="C97A32C0A66F0168AC0D1FEA4C8B03E3" author="Korovchinsky NM &amp; Galkin SV" issue="8" journalOrPublisher="Zoologicheskii Zhurnal" pagination="1205-1215" refId="B34" refString="Korovchinsky NM, Galkin SV (1984) New data on the fauna of starfishes of the genus Freyella (Brisingidae). Zoologicheskii Zhurnal 63 (8): 1205-1215. [in Russian with English abstract]" title="New data on the fauna of starfishes of the genus Freyella (Brisingidae)." volume="63" year="1984">Korovchinsky &amp; Galkin, 1984</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="F18A02C473108D6DE559CCF1FDFE3CD0" authorityName="H. L. Clark" authorityYear="1920" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="oligobrachia">
<emphasis id="C8C32974810482ADC36D829489D75184" italics="true">F. oligobrachia</emphasis>
</taxonomicName>
and
<taxonomicName id="FD341200F74E466A3E1D17D80173C011" authorityName="Korovchinsky &amp; Galkin" authorityYear="1984" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="vitjazi">
<emphasis id="42153D1A4CCCD45BC370B4FF0353ED8B" italics="true">F. vitjazi</emphasis>
</taxonomicName>
have inferomarginals corresponding to alternate adambulacral plates according to their original descriptions; thus, they should be retained within
<taxonomicName id="03B23590E172AB3B3197266B94FA216A" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="1FD5A06478E80F4D9EE600DD3B29620A" italics="true">Freyella</emphasis>
</taxonomicName>
.
<taxonomicName id="80CD4C8D96401342541F8DA82445389C" authorityName="Baranova" authorityYear="1957" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="hexactis">
<emphasis id="57EB4FF42A90BC35BB823079E8287FFA" italics="true">F. hexactis</emphasis>
</taxonomicName>
was described to have inferomarginal plates to every third adambulacrals, then to alternate adambulacrals, then at the end of the arm, to each adambulacral plate (
<bibRefCitation id="9F6DBC37ED7521AAA2EDC5F2B0F821FA" author="Baranova" firstAuthor="Baranova" journalOrPublisher="Investigations of Far-East Seas" pagination="149-266" refId="B3" refString="Baranova ZI (1957) Echinoderms of the Bering Sea. Investigations of Far-East Seas 4: 149266. [in Russian]" title="Echinoderms of the Bering Sea." volume="4" year="1957">Baranova 1957</bibRefCitation>
). It is difficult to justify it as either
<taxonomicName id="EF0CD400770AC5EB422F6D7F2CF80E1A" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="0C5AD1B5FB7DCB36C9D1FADEB86B127F" italics="true">Freyella</emphasis>
</taxonomicName>
or
<taxonomicName id="53AF29525BD137D3E047584A9138A7CC" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="182C985F3BC8FC0498A8DC685048C7BB" italics="true">Freyastera</emphasis>
</taxonomicName>
simply by this description. However, this species closely resembles
<taxonomicName id="3844AFF8ECD70CAF4F214C1E52EC250A" authorityName="H. L. Clark" authorityYear="1920" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="oligobrachia">
<emphasis id="AC6D0951D18ADD8C7BF72C9A92FD1DCC" italics="true">F. oligobrachia</emphasis>
</taxonomicName>
and
<taxonomicName id="6D742BC0172E16586EAEEA9F15EE9AE5" authorityName="Sladen" authorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="8E57F0971FD5479BB7B34673DAFE82CB" italics="true">F. benthophila</emphasis>
</taxonomicName>
, as was shown in the key in
<bibRefCitation id="167BAB091E1962D040295E4F0BDB133A" DOI="10.1016/j.dsr.2019.103163" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" etAl="et al." firstAuthor="Zhang" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al. (2019)</bibRefCitation>
. It is necessary to examine the types of
<taxonomicName id="EE70FD9CA02FA3C6C1A107F5FB19062C" authorityName="H. L. Clark" authorityYear="1920" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="oligobrachia">
<emphasis id="D61906761D4DC81C79921AC9185B45B1" italics="true">F. oligobrachia</emphasis>
</taxonomicName>
and
<taxonomicName id="B8585FF3E689C2DF6A071E6D25296A08" authorityName="Baranova" authorityYear="1957" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="hexactis">
<emphasis id="CAF08A81A15646D38432AA4A5AADF0B3" italics="true">F. hexactis</emphasis>
</taxonomicName>
to be certain of their generic affinities and their relationship with
<taxonomicName id="754130E9246FCDD13E504E9217A0B0A6" authorityName="Sladen" authorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="A65451DF7CD6BCDD3374D873F107C2D7" italics="true">F. benthophila</emphasis>
</taxonomicName>
.
<taxonomicName id="BABF2D53F8607E1B09CC6C80CF6AE95A" authorityName="Korovchinsky &amp; Galkin" authorityYear="1984" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="vitjazi">
<emphasis id="8B0DDDAE8BF9D1B0209F746C0AB775BC" italics="true">F. vitjazi</emphasis>
</taxonomicName>
, on the other hand, could be differentiated from the other six-armed
<taxonomicName id="28DAE945EC53CC9326E1D02192410B9E" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="41185746966ACE21B4F4BC406301344B" italics="true">Freyella</emphasis>
</taxonomicName>
species by having a diagonal line of three spines on each adambulacral plate. These three species are retained as
<taxonomicName id="75219C8F1DE004183E7FC829FAE0E9E8" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="774B260B56A94943F8FE281717F58DF2" italics="true">Freyella</emphasis>
</taxonomicName>
for the time being, but they should be compared with in any future descriptions of six-armed freyellids.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,118 @@
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<paragraph id="A4FBA132418AAFBF34D77700863F9909">
<taxonomicName id="E64080A7B84997BFD71B4DBA6984CAB4" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-7">
<emphasis id="94AC0995FEDF4DE4D4159213F224BB60" italics="true">Freyastera</emphasis>
sp. 7
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<paragraph id="3B2C075988E3A358DF26B266907B3C46">
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<taxonomicName id="13AF9713634FB6887197433BD0DCE2FE" ID-CoL="6JNSY" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="tuberculata">
<emphasis id="1391C7C3B9A35CCA9C13F03B1D56B8DD" italics="true">Freyastera tuberculata</emphasis>
</taxonomicName>
:
<bibRefCitation id="D6C157B4C677D1D10D1AF255E845039C" DOI="10.3897/BDJ.3.e7062" author="Moreau" etAl="et al." firstAuthor="Moreau" refId="B47" refString="Moreau CV, Aguera A, Jossart Q, Danis B (2015) Southern Ocean Asteroidea: a proposed update for the Register of Antarctic Marine Species. Biodiversity Data Journal 3: e 7062. https://doi.org/10.3897/BDJ.3.e7062" year="2015">Moreau et al. 2015</bibRefCitation>
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;
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<bibRefCitation id="3AD8260CB07C2E5082DF62B23A80EF99" DOI="10.3897/zookeys.747.22751" author="Moreau" etAl="et al." firstAuthor="Moreau" journalOrPublisher="ZooKeys" pagination="141-156" refId="B48" refString="Moreau C, Mah C, Agüera A, Améziane N, Barnes D, Crokaert G, Eléaume M, Griffiths H, Guillaumot C, Hemery LG, Jażdżewska A. (2018) Antarctic and sub-Antarctic Asteroidea database. ZooKeys 747: 141156. https://doi.org/10.3897/zookeys.747.22751" title="Antarctic and sub-Antarctic Asteroidea database." volume="747" year="2018">Moreau et al. 2018</bibRefCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EPDBI" type="material">
<paragraph id="887FB6F4BF860E6E5CF43B3078B0764D">
<heading id="292E119AAECA4810CAF462ED86C77754" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="FFFD2D40E39A2E7DB20B8BE6CCB1F145">
ULB_S 59-MT 2, multiple broken arms from different individuals. (Suppl. material
<xref id="BD93AFF4803E7954615125A0375811DC" ref-type="supplementary-material" rid="S1">1</xref>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZDBI" type="distribution">
<paragraph id="0956893020F2DA7D259BC356AB9BE538">
<heading id="CECF23787FE08A9EB0460AC1B2AA54DE" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="35C7BF14E1EA2293CED4965755689D1E">
Weddle Sea,
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@ -1,73 +1,73 @@
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@ -86,7 +86,6 @@
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<emphasis id="93C38A5F8289175BFC284F67360541BF" italics="true">Freyella mortenseni</emphasis>
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<emphasis id="595F8AB35BC7FAAFCC12E8822E17F03F" italics="true">Freyastera mortenseni</emphasis>
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@ -133,7 +130,6 @@
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@ -142,15 +138,23 @@
<heading id="6FF9DAD62A727352F245887E21522138" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="16A5D12B9F325BC254CBA1603759D6E6">
RSIOAST 0102 (Fig.
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(Fig.
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@ -161,7 +165,9 @@ RSIOAST 0102 (Fig.
<emphasis id="02ED9B1477F3CF2D37E86FD7E3F9BF85" italics="true">Freyastera mortenseni</emphasis>
<bibRefCitation id="9B2BFAF97CBFB2A3C3AA702D5692B7F4" author="Madsen FJ" journalOrPublisher="Galathea Report" pagination="23-32" refId="B39" refString="Madsen FJ (1956) Echinoidea, Asteroidea, and Ophiuroidea from depths exceeding 6000 meters. Galathea Report 2: 23-32." title="Echinoidea, Asteroidea, and Ophiuroidea from depths exceeding 6000 meters." volume="2" year="1956">Madsen, 1956</bibRefCitation>
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, RSIOAST 0102.
,
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.
<emphasis id="CE3A51109F8A0242456EC2B1BA9F853C" bold="true">A.</emphasis>
Actinal view of disk;
<emphasis id="5361FCA5A595FCC040AA213EE63C93E2" bold="true">B.</emphasis>
@ -196,9 +202,7 @@ in length) cluster in transverse bands on abactinal arm beyond genital area. Pro
<paragraph id="E4FFDFAFFF1BFC147499962429808E3A">
Southwest Pacific: Kermadec Trench; Northwest Pacific: Mariana Trench, Parece Vela basin.
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locality: Kermadec Trench,
. Type locality: Kermadec Trench,
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</paragraph>

View file

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</mods:role>
<mods:namePart id="AF255B9C706133EE55FA852ACE622029">Mao, Jingwen</mods:namePart>
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<mods:affiliation id="A92832A36D3D2143ACA09FBA18FCB3C0">State Key Laboratory of Submarine Geoscience, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, 310012, China</mods:affiliation>
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<mods:namePart id="105BCE5AC072FA5128DAE6FF2DB2F7C4">Wang, Chunsheng</mods:namePart>
<mods:affiliation id="D9D68C5FAB9F50B6740A6608BA76225B">State Key Laboratory of Submarine Geoscience, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, 310012, China &amp; Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, 310012, China &amp; Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519082, China</mods:affiliation>
</mods:name>
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<mods:namePart id="501536255D4AD3918D452588479B1C06">Zhang, Dongsheng</mods:namePart>
<mods:affiliation id="79EF98FDBA24D38A1E029DF317EB2796">State Key Laboratory of Submarine Geoscience, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, 310012, China &amp; Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, 310012, China &amp; Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519082, China</mods:affiliation>
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<mods:title id="50193FB01754BC4932FD29240AC3C233">Zoosystematics and Evolution</mods:title>
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<mods:part id="82AF1CA8C6541E9FA026AC3FBA3075E5">
<mods:date id="9A31B3E22E79DB312D6FC93FD5AB4F00">2025</mods:date>
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<mods:number id="6AEBD8F026C09D38D407D9FEB42439F4">2025-04-10</mods:number>
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<mods:number id="38A1D048841DDAF150F991E3F81F45B3">101</mods:number>
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<mods:number id="2FE954AD728EC24E4BA39080FD6A8EB2">2</mods:number>
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<mods:identifier id="13B3F3C475E77516370717FC883F09EB" type="DOI">10.3897/zse.101.144918</mods:identifier>
<mods:identifier id="7087B568F9CDADCF911A001219C66A4E" type="ZooBank">4611CD60-658B-42C4-B23C-2DA9185A85E3</mods:identifier>
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<treatment id="654269D362F05022BEE1FE432B986D74" ID-arpha="654269D3-62F0-5022-BEE1-FE432B986D74" LSID="urn:lsid:plazi:treatment:654269D362F05022BEE1FE432B986D74" httpUri="http://treatment.plazi.org/id/654269D362F05022BEE1FE432B986D74" scope_class="Asteroidea" scope_family="Freyellidae" scope_order="Brisingida">
<subSubSection id="1A66DF637622041820A2D6AC26EABD13" type="nomenclature">
<paragraph id="DB4E5E6DAB7AAAA9EFEA69BB8C7F1754">
<taxonomicName id="61A4CCEAB45FCC81829AD8DF9897666C" ID-CoL="848TW" authority="Zhang et al., 2019" authorityName="Zhang" authorityYear="2019" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="delicata">
<emphasis id="B8108A19DC563087C07D790FFB81247E" italics="true">Freyastera delicata</emphasis>
<bibRefCitation id="0749FC014F6E0E7E0E37BB77DD72743A" DOI="10.1016/j.dsr.2019.103163" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al., 2019</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="88E63A8FE4DE23E666B8FB6ADA16B526" type="description">
<paragraph id="E639C1F278432866A3127E6A2A7F42FF">
<figureCitation id="5BBD23BF2DE8F8CC7DD65E5AB34D2756" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">Figs 5</figureCitation>
,
<figureCitation id="93CEDFD7236A38308CC2BD4389572915" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. In situ photos of Freyastera species. A. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117; B. Freyastera jiaolongi sp. nov. paratype RSIOAST 0112; C. Freyastera jiaolongi sp. nov. paratype RSIOAST 0113; D. Freyastera delicata RSIOAST 0135; E. Freyastera basketa RSIOAST 0200; F. Freyastera sp. 5 RSIOAST 0116; G. Freyastera sp. 5 RSIOAST 0107; H. Freyastera sp. 6 RSIOAST 0124; I. Freyastera sp. 6 RSIOAST 0125." figureDoi="10.3897/zse.101.144918.figure14" httpUri="https://binary.pensoft.net/fig/1304665">14 D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="29AC42956FA9F24DE88E6C9A6C2C88B9" type="reference_group">
<paragraph id="D6967DFEF565CCD106720FA398DAFC9D">
<treatmentCitationGroup id="CAD9042DA645121EDE1340DF134F7DC9">
<taxonomicName id="75AAF4FDFC18AEC5203B8A73654ED0F1" ID-CoL="848TW" authorityName="Zhang" authorityYear="2019" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="delicata">
<emphasis id="5682746B34C76F32E25F5ACBC5DA007E" italics="true">Freyastera delicata</emphasis>
</taxonomicName>
<comment id="BA26841CE6E972E2EF77333D464A0DA1">
:
<treatmentCitation id="1E54CFE5228225CA63152FECD77DFEAF" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" page="2" year="2019">
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</treatmentCitation>
;
<treatmentCitation id="42D11D2D49EB6E4A2E778FABEEB50527" author="Zhang R &amp; Fau M &amp; Mah C &amp; Eléaume M &amp; Zhang D &amp; Zhou Y &amp; Lu B &amp; Wang C" page="10" year="2024">
<bibRefCitation id="571CB6FE336EDF763D09C1D9AF895A86" DOI="10.1016/j.ympev.2023.107993" author="Zhang R &amp; Fau M &amp; Mah C &amp; Eléaume M &amp; Zhang D &amp; Zhou Y &amp; Lu B &amp; Wang C" etAl="et al." firstAuthor="Zhang" page="10" refId="B69" refString="Zhang R, Fau M, Mah C, Eléaume M, Zhang D, Zhou Y, Lu B, Wang C (2024) Systematics of deep-sea starfish order Brisingida (Echinodermata: Asteroidea), with a revised classification and assessments of morphological characters. Molecular Phylogenetics and Evolution 191: 107993. https://doi.org/10.1016/j.ympev.2023.107993" year="2024">Zhang et al. 2024: 10</bibRefCitation>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EDXAG" type="material">
<paragraph id="440AB4250DA43FE2420A930E05BF5CBE">
<heading id="AD90A2EF8C7BB2D63692182ABE8A63CA" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="005150002CB9423421FFFBA9B36126AA">
<materialsCitation id="7232FE736FFB68EC44F8FFDC73E39BE4" collectionCode="RSIOAST" specimenCode="RSIOAST 0022" specimenCount="1" typeStatus="holotype">
<specimenCode id="54A321222E6698D6AA914E83C6089E57">RSIOAST 0022</specimenCode>
(
<emphasis id="CD43BCBD24131B84D5C5691BA21A0D30" bold="true" italics="true">
<typeStatus id="7F8486339D99ED2C886C511777A5DB5A" type="holotype">holotype</typeStatus>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="2B318D7D52700F275D2DB8B1B759A13E" collectionCode="RSIOAST" specimenCode="RSIOAST 0135" specimenCount="1">
<specimenCode id="DEE55ADF0F3D87A88999A58EF5F7BECA">RSIOAST 0135</specimenCode>
(Figs
<figureCitation id="3AA5135F8D3CF92E6EADE50C422B8F76" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">5</figureCitation>
,
<figureCitation id="D9D4A706057BCD7C869198614A02D50F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. In situ photos of Freyastera species. A. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117; B. Freyastera jiaolongi sp. nov. paratype RSIOAST 0112; C. Freyastera jiaolongi sp. nov. paratype RSIOAST 0113; D. Freyastera delicata RSIOAST 0135; E. Freyastera basketa RSIOAST 0200; F. Freyastera sp. 5 RSIOAST 0116; G. Freyastera sp. 5 RSIOAST 0107; H. Freyastera sp. 6 RSIOAST 0124; I. Freyastera sp. 6 RSIOAST 0125." figureDoi="10.3897/zse.101.144918.figure14" httpUri="https://binary.pensoft.net/fig/1304665">14 D</figureCitation>
). (Suppl. material
<xref id="E8BD7E46AAE534320C1B512EF1A65AD0" ref-type="supplementary-material" rid="S1">1</xref>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYXAG" type="diagnosis">
<paragraph id="D08FEB4E44AB74C8E9DD8008CC963F08">
<heading id="BBC0206E28A1042F5848F7A20158F1E7" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="7813567D0D88E3B4EAD3D67404906A3A">
Arms six. Abactinal disk and arm plate each bears multiple spines (usually 48), most measuring
<quantity id="F4BBE1946BFF73301DA634CB9F073F4E" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
in length, not covered by membranous sheath. Pedicellariae absent on abactinal surface of disk and arm genital area. Pedicellariae cluster in transverse bands on abactinal arm beyond genital area. One long aboral spine and one subambulacral spine on each adambulacral plate. Proximal subambulacral spines with truncate end. Lateral spine corresponds to every adambulacral plate beyond genital area. Oral plate bears two actinostomal spines, one suboral spine, and one aboral furrow spine.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4XAG" type="distribution">
<paragraph id="93E19B667C9575BB6FC0BD406AE869D7">
<heading id="7BD2A977E31E43D0D1431B9B4A16160B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="559CF8C2324523D34DCB5D636DB37911">
Northwest Pacific: Caiwei Seamount, O-Hakucho Guyot,
<quantity id="42FB5E0029A9BF56E9EDFFB53516D786" metricMagnitude="3" metricUnit="m" metricValue="3.7215" metricValueMax="4.322" metricValueMin="3.121" unit="m" value="3721.5" valueMax="4322.0" valueMin="3121.0">31214322 m</quantity>
. Type locality: Caiwei Seamount,
<quantity id="D930C56E7FFFE1475300F77C9966241F" metricMagnitude="3" metricUnit="m" metricValue="4.322" unit="m" value="4322.0">4322 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECYAG" type="remarks">
<paragraph id="E70FA5803099FE9711107630911CBB81">
<heading id="13571B2A56DB2D01E56627FC29D646A7" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="362E0AA8AC57188AE714FD7BF4EF6AD9">
The species was described based on a single specimen (
<typeStatus id="D41E6A5A88152AD1C87C6C6CA9C6A65C" type="holotype">holotype</typeStatus>
<specimenCode id="1CEBD8B94B59A53E581FC82825D45FF7">RSIOAST 0022</specimenCode>
) (
<bibRefCitation id="C96E54E230DE85094A97B5CBE1ADE94F" DOI="10.1016/j.dsr.2019.103163" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al. 2019</bibRefCitation>
). An additional specimen was examined in the present study, providing information on intraspecific variations of the species. The specimen
<specimenCode id="A85AEAEEE60D245881003D6A60747124">RSIOAST 0135</specimenCode>
(Fig.
<figureCitation id="47A895428466564A934EA3538EECE3CA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">5</figureCitation>
) is smaller compared to the
<typeStatus id="0B4E4A421F498793038CEE1BFF96DE8B" type="holotype">holotype</typeStatus>
, with maximum R =
<quantity id="2BFFBD917351D431C17A32560EC8C589" metricMagnitude="-1" metricUnit="m" metricValue="1.35" unit="mm" value="135.0">135 mm</quantity>
, r =
<quantity id="C539ED047E68334BBE9748CCA9B85309" metricMagnitude="-3" metricUnit="m" metricValue="4.1" unit="mm" value="4.1">4.1 mm</quantity>
, R / r ≈ 33. The arm measures
<quantity id="A54715DFA874F37F2C70261261526703" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
wide at the base, whereas the widest part of arm measures
<quantity id="75C78646515203009B28822435AD2CE8" metricMagnitude="-3" metricUnit="m" metricValue="4.6" unit="mm" value="4.6">4.6 mm</quantity>
. The genital area is about
<quantity id="7D975D05066D05821CE528628C66A711" metricMagnitude="-3" metricUnit="m" metricValue="9.5" unit="mm" value="9.5">9.5 mm</quantity>
in length. Gonads well-developed, female (Fig.
<figureCitation id="A20A8AD4988BA69380A49FFCF327D07E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">5 G</figureCitation>
). Abactinal arm spines slightly longer than those in the
<typeStatus id="EA2A6EC7E6F469D99932891749B5F449" type="holotype">holotype</typeStatus>
, but not aligned in transverse rows as in the
<typeStatus id="A23C37A6B31D340C9B38D44C6E24DFDE" type="holotype">holotype</typeStatus>
(Fig.
<figureCitation id="D00501B6BA723D93C2A5BA48440C38A3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">5 D</figureCitation>
). The new specimen shares other characters with the
<typeStatus id="A2DF7464038A0E9DD509EF54BC8E52E8" type="holotype">holotype</typeStatus>
, including having no trace of pedicellariae on abactinal disk and genital area (Fig.
<figureCitation id="C83287F76DD27A218449CB9656D983A1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">5 B, D</figureCitation>
), mouth plate with two actinostomal spines, one suboral spine, and one aboral furrow spine (Fig.
<figureCitation id="54F8994282C74DA1F610B2C77E5BDF8F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">5 A</figureCitation>
), adambulacral plate with one subambulacral spine and one aboral spine pointing upwardly (Fig.
<figureCitation id="11CD5249855C5A88B9747E89BDADC254" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">5 C, E</figureCitation>
), abactinal arm plate with several spines, and arm beyond genital area with transverse bands of pedicellariae (Fig.
<figureCitation id="5F1848FF5E6B063CC4757097C70FF271" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freyastera delicata Zhang et al., 2019, RSIOAST 0135. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Actinal view of middle part of arm; F. Abactinal view of arm beyond genital area; G. Female gonads; H. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure5" httpUri="https://binary.pensoft.net/fig/1304656">5 H</figureCitation>
). The new specimen was found at
<quantity id="8ED8F9BC7D6425BFC1F21E74D2D69D44" metricMagnitude="3" metricUnit="m" metricValue="3.121" unit="m" value="3121.0">3121 m</quantity>
depth from O-Hakucho Guyot, close to the type locality (Fig.
<figureCitation id="F2AAED2E04F6C74FAEBBDB79E312DDC8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Freyastera and six-armed Freyella specimens examined in the present study and type localities of each species." figureDoi="10.3897/zse.101.144918.figure1" httpUri="https://binary.pensoft.net/fig/1304652">1</figureCitation>
), but is
<quantity id="8C1694949CDCA51B57F655A13AD74FF8" metricMagnitude="3" metricUnit="m" metricValue="1.201" unit="m" value="1201.0">1201 m</quantity>
shallower than the depth of the
<typeStatus id="4166D779E30AA05240ABFF1DED395655" type="holotype">holotype</typeStatus>
(
<quantity id="CA80F8F4282750A322923E5EF011A00F" metricMagnitude="3" metricUnit="m" metricValue="4.322" unit="m" value="4322.0">4322 m</quantity>
).
</paragraph>
<caption id="BD88D3D9C5C6FA93B569618ECBB15F25" ID-DOI="10.3897/zse.101.144918.figure5" ID-arpha="140F37E6-E686-50F2-A097-EFEEF395787F" httpUri="https://binary.pensoft.net/fig/1304656" startId="F5">
<paragraph id="D96D11CBAB85E27934F549012B75DE51">
<label id="E12903B6D77EBB2603D36ECBCF7531AA">Figure 5.</label>
</paragraph>
<paragraph id="1A8C6F9A839A11F59C0116AAB64501C0">
<taxonomicName id="61EE40C25C541D4316B45052814A1FC1" authority="Zhang et al., 2019" authorityName="Zhang" authorityYear="2019" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="delicata">
<emphasis id="1D9C332327616DE3896D3BE1909EB263" italics="true">Freyastera delicata</emphasis>
<bibRefCitation id="5AEFB153BB4C52DD0E3325380424B5E0" DOI="10.1016/j.dsr.2019.103163" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al., 2019</bibRefCitation>
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,
<specimenCode id="23EEA6A3694D76F088FD0D20D97F7593">RSIOAST 0135</specimenCode>
.
<emphasis id="8E848FFFE26F1DF946FC40DBAA7F7286" bold="true">A.</emphasis>
Actinal view of disk;
<emphasis id="284952E1FDB582E179B70FBC77B6DE78" bold="true">B.</emphasis>
Abactinal view of disk;
<emphasis id="5F0756FA9476E96F59566A1CDC156376" bold="true">C.</emphasis>
Actinal view of arm genital area;
<emphasis id="EB7E1E710C9ABFA92D9B338DE281D2DE" bold="true">D.</emphasis>
Abactinal view of genital area;
<emphasis id="2FB069CC6AADF4EE9272E66A99BA349B" bold="true">E.</emphasis>
Actinal view of middle part of arm;
<emphasis id="FD686F70BEFCB9D09A58B74C79DBD4D8" bold="true" italics="true">F.</emphasis>
Abactinal view of arm beyond genital area;
<emphasis id="DA06CC0DC94D412F67B0BDDD44B19BCF" bold="true">G.</emphasis>
Female gonads;
<emphasis id="5ECE3C1ABEE69579BCEA94FBB6383F7E" bold="true">H.</emphasis>
abactinal view of middle part of arm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -1,73 +1,73 @@
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<taxonomicName id="4F34FCAEA429D5C6C5F993DB37A74B38" ID-CoL="6JNSY" authority="(Sladen, 1889)" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="tuberculata">
@ -86,7 +86,6 @@
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<emphasis id="BC5ADDF732C2B92099BFC7B9A7588162" italics="true">Freyella tuberculata</emphasis>
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@ -113,12 +112,10 @@
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.
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<emphasis id="23F3041D4ECAAA2F9C19FE0F123F63D7" italics="true">Freyellidea tuberculata</emphasis>
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@ -133,12 +130,10 @@
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.
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<emphasis id="92D30FF890C0721341EB324E8B1245C7" italics="true">Freyastera tuberculata</emphasis>
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@ -173,7 +168,6 @@
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(in key).
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@ -182,24 +176,32 @@
<heading id="788661A378A959764E8FAD07F5A9D10F" reason="title">Material examined.</heading>
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1890.5.7.1076
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(
<emphasis id="819B491064459D414D32A4AE24D01334" bold="true" italics="true">
<typeStatus id="EB126BE9794EFD5261A36BB726051430" type="syntype">syntype</typeStatus>
</emphasis>
) (Fig.
<figureCitation id="A3E08044B0F4FEA476490F80E7C4F212" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A D. Freyastera loricata (Korovchinsky &amp; Galkin, 1984) type ECH 00161 (photos courtesy of Dr. Anna Dilman); E I. Freyastera tuberculata (Sladen, 1889) type NHMUK 1890.5. 7.1076. A, E. abactinal view of disk; B, F. Actinal view of disk; C. G. Abactinal view of arm genital area; D, H. Actinal view of proximal part of arm; I. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure9" httpUri="https://binary.pensoft.net/fig/1304660">9 E I</figureCitation>
);
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1890.5. 7.1077 (
)
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;
<materialsCitation id="94DAAE083692310356605F8E71D245ED" collectionCode="NHMUK" specimenCode="NHMUK 1890.5.7.1077" specimenCount="1" typeStatus="syntype">
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1890.5.7.1077
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(
<emphasis id="4C912E3BA1EB87759ECAF280EE07950E" bold="true" italics="true">
<typeStatus id="0596D2D44EE3EA922CA02D35B3C1917B" type="syntype">syntype</typeStatus>
</emphasis>
).
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELYAI" type="material">
@ -207,26 +209,41 @@
<heading id="DD8D89793170A30489103AE6C9D38EAA" reason="title">Other material.</heading>
</paragraph>
<paragraph id="9868C38C8E71400B82A20F6F2F2D6766">
<specimenCode id="60823CF90B9D0BA85AEFC79F33F4BC7C">
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<collectionCode id="6DD7A909981445ECAFA6BDF0D508E77D" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
- IE- 2008-1363;
-IE-2008-1363
</specimenCode>
;
<specimenCode id="AC4085EA46F40D2729728CAC73277F16">
<named-content id="6170CE83E373CB9EB225D0D94F1F3F07" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="82D4C7AA87E77EE4C3BF8A3EBCF1579A" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
- IE- 2017-1819;
-IE-2017-1819
</specimenCode>
;
<specimenCode id="7F1083A025850DCBD5FB45FA8ED7C877">
<named-content id="A890C9F2570864A44F6B05A6E466DAAC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="46BF349EC0E51869A7CEF0257FC1F1BB" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
- IE- 2019-5479;
-IE-2019-5479
</specimenCode>
;
<specimenCode id="F6EA690550F2668D5EB7773CB878021A">
<named-content id="D01177952F17334E708F20BDA0015C48" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="1E0F77729D4636220D5FFCBEA58EF82E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
- IE- 2019-5743;
-IE-2019-5743
</specimenCode>
;
<specimenCode id="417552912C74F6CA2600C6F4DEF9B226">
<named-content id="D0E7DF8CF5AC9D6309ED03AE6C908864" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="E5F64924CC9CC6AD2B4653FCECE54595" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
- IE- 2019-5745. (Suppl. material
-IE-2019-5745
</specimenCode>
. (Suppl. material
<xref id="F42671A5FF2AEF68F04C02F32A86EF45" ref-type="supplementary-material" rid="S1">1</xref>
).
</paragraph>
@ -238,9 +255,7 @@
<paragraph id="C77FE37689D647FE05A67D976969C406">
Atlantic Ocean; Eastern Pacific; Indian Ocean; Bay of Bengal.
<quantity id="8CDB02566FC594A452D3A988469F06BE" metricMagnitude="3" metricUnit="m" metricValue="4.2725" metricValueMax="5.18" metricValueMin="3.3649999999999998" unit="m" value="4272.5" valueMax="5180.0" valueMin="3365.0">33655180 m</quantity>
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<typeStatus id="06D0A3A2BFBD81D56EC003EEE76D8721">Type</typeStatus>
locality: Atlantic Ocean: between Canary Islands and
. Type locality: Atlantic Ocean: between Canary Islands and
<collectingCountry id="701729540F51C29984C8952BCEB47482" name="Cape Verde">Cape Verde</collectingCountry>
Islands; between the coast of Africa and the Island of
<collectingCountry id="B26932B0A4526A7448301341259D90AC" name="Saint Helena, Ascension and Tristan da Cunha">Ascension</collectingCountry>

View file

@ -0,0 +1,334 @@
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<treatment id="AC6048DB979B54BEA70F9BD52F1BF82D" ID-arpha="AC6048DB-979B-54BE-A70F-9BD52F1BF82D" LSID="urn:lsid:plazi:treatment:AC6048DB979B54BEA70F9BD52F1BF82D" httpUri="http://treatment.plazi.org/id/AC6048DB979B54BEA70F9BD52F1BF82D" scope_class="Asteroidea" scope_family="Freyellidae" scope_order="Brisingida">
<subSubSection id="BD3B5732AF355EB229503E5C7E16A787" type="nomenclature">
<paragraph id="737C3A94609F6E928E23658CF76B1CD3">
<heading id="DCE14862A4F5EDCD742903E15151B357" reason="label">Genus</heading>
<taxonomicName id="F9A51A93D6C740F96A26DCF4BDF29C7F" ID-CoL="7P2S3" authority="Downey, 1986" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="070DF78FEE80B988DA3CE33E192DFA8F" italics="true">Freyastera</emphasis>
<bibRefCitation id="3D39BB6420F1FCED907BE22EAE2A74C1" DOI="10.5479/si.00810282.435" author="Downey ME" journalOrPublisher="Smithsonian Contributions to Zoology" pagination="1-57" refId="B15" refString="Downey ME (1986) Revision of the Atlantic Brisingida (Echinodermata: Asteroidea), with description of a new genus and family. Smithsonian Contributions to Zoology 435: 1-57. https://doi.org/10.5479/si.00810282.435" title="Revision of the Atlantic Brisingida (Echinodermata: Asteroidea), with description of a new genus and family." volume="435" year="1986">Downey, 1986</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="17C53408F89357A3060C0840509B5E91">
<heading id="F172F51606CA26B7E16CD7F5F736D487" reason="title">Type species.</heading>
</paragraph>
<paragraph id="8734920DD49DE75B3C072CC1801B6677">
<taxonomicName id="E4C46D1EF728EF8BDD95A0C218BBC65B" authority="Perrier, 1885" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="3BC0A839E5307659F1078273455EF96D" italics="true">Freyella sexradiata</emphasis>
<bibRefCitation id="D0F9547712AE383391D6FCF2B66B15A2" author="Perrier E" issue="8" journalOrPublisher="Annales des sciences naturelles, Zoologie" pagination="1-72" refId="B53" refString="Perrier E (1885) Premiere note Preliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman. Annales des sciences naturelles, Zoologie 22 (8): 1-72. [in French]" title="Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman." volume="22" year="1885">Perrier, 1885</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESKAE" type="diagnosis">
<paragraph id="81871548F26FD8A61AB470427D59022E">
<heading id="87D82FBC55C42418288CC96F64A60FA0" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="165667225D2AE1224BB0A67DBFE0DD75">Arms six. Papulae absent. One pair of gonads on each arm. Disk very small, arms extremely long and slender. The first pair of inferomarginal plates appears later on arm, not in contact with the odontophore. Inferomarginal plates and lateral spines generally correspond to every adambulacral plate beyond genital area. Abactinal arm in genital region covered with pavement of spinate plates. Adambulacral plates elongated.</paragraph>
</subSubSection>
<subSubSection id="SECID0EXKAE" type="remarks">
<paragraph id="0B962F4F6647C33F732F92B0AF17E20E">
<heading id="B673CB80D9E14E3950C228357925162D" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="8B02EA46F33D2A5B13652FAA77BD8FDF">
<taxonomicName id="64ECFE15D3E0CDB77876B1E42D98B4E1" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="266A2CAC00609AA45B8F98A70D85B166" italics="true">Freyastera</emphasis>
</taxonomicName>
and
<taxonomicName id="1E543CA90F30B9F0CFF9C09C4E4F5732" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="B08134A7BF2DFFDAFFCD5EDA2AA5666A" italics="true">Freyella</emphasis>
</taxonomicName>
were redefined by
<bibRefCitation id="F5DA71FF27883EDFF4CFD77B51B20EF5" DOI="10.1016/j.ympev.2023.107993" author="Zhang R &amp; Fau M &amp; Mah C &amp; Eléaume M &amp; Zhang D &amp; Zhou Y &amp; Lu B &amp; Wang C" etAl="et al." firstAuthor="Zhang" refId="B69" refString="Zhang R, Fau M, Mah C, Eléaume M, Zhang D, Zhou Y, Lu B, Wang C (2024) Systematics of deep-sea starfish order Brisingida (Echinodermata: Asteroidea), with a revised classification and assessments of morphological characters. Molecular Phylogenetics and Evolution 191: 107993. https://doi.org/10.1016/j.ympev.2023.107993" year="2024">Zhang et al. (2024)</bibRefCitation>
, with the number of arms and the arrangement of inferomarginal plates as key diagnostic characters. The genus
<taxonomicName id="A79A263F568A1DF5D66DFF20A905D6D3" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="52B7B722DB7C6CB55A3C8B488F30A094" italics="true">Freyastera</emphasis>
</taxonomicName>
currently contains seven species (
<bibRefCitation id="E529915227FD48F381DDA71D647480EC" author="Mah" firstAuthor="Mah" refId="B42" refString="Mah CL (2025) . World Asteroidea Database. Freyastera Downey, 1986. [World Register of Marine Species] https://www.marinespecies.org/aphia.php?p=taxdetails&amp;id=123216" url="https://www.marinespecies.org/aphia.php?p=taxdetails&amp;id=123216" year="2025">Mah 2025</bibRefCitation>
):
<taxonomicName id="63F4CD9086AE4C6BD4C55CC22EAB0B5F" authority="Zhang et al., 2019" authorityName="Zhang" authorityYear="2019" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="basketa">
<emphasis id="920FECF06873E302BBF10D7501E4673C" italics="true">Freyastera basketa</emphasis>
<bibRefCitation id="8864B149BF589D7FD5C965ED7324CC74" DOI="10.1016/j.dsr.2019.103163" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al., 2019</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D36A1720114EFCECF4BEF866489842BE" authority="Zhang et al., 2019" authorityName="Zhang" authorityYear="2019" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="delicata">
<emphasis id="22DFA44D6F80F8E341BDA0F8EBB4DB55" italics="true">Freyastera delicata</emphasis>
<bibRefCitation id="DBF608C509B159CC3497F3A97FB0CF0E" DOI="10.1016/j.dsr.2019.103163" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al., 2019</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4CCC108D6B03D77C0AE5BF6CAA5E79B2" authority="McKnight, 2006" authorityName="McKnight" authorityYear="2006" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="digitata">
<emphasis id="8B5D8832EE78355D64B6E8539A5896BC" italics="true">Freyastera digitata</emphasis>
<bibRefCitation id="295428C44027C5DCAA405A8D0436A8AC" author="McKnight DG" journalOrPublisher="NIWA Biodiversity Memoir" pagination="1-187" refId="B44" refString="McKnight DG (2006) The marine fauna of New Zealand, Echinodermata: Asteroidea (Sea-stars). 3. Orders Velatida, Spinulosida, Forcipulatida, Brisingida with addenda to Paxillosida, Valvatida. NIWA Biodiversity Memoir 120: 1-187." title="The marine fauna of New Zealand, Echinodermata: Asteroidea (Sea-stars). 3. Orders Velatida, Spinulosida, Forcipulatida, Brisingida with addenda to Paxillosida, Valvatida." volume="120" year="2006">McKnight, 2006</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="D5DF2DB67AF715C11B8C3AB0CD273FD8" authority="(A. H. Clark 1939)" baseAuthorityName="A. H. Clark" baseAuthorityYear="1939" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="mexicana">
<emphasis id="D309A65E3D4BE7C23C469B7C730A253B" italics="true">Freyastera mexicana</emphasis>
(A. H.
<bibRefCitation id="32DF2EEF717992DAB6F9E1EED83A1075" DOI="10.5479/si.00963801.86-3056.441" author="Clark" firstAuthor="Clark" journalOrPublisher="National Museum" pagination="441-456" refId="B8" refString="Clark AH (1939) Echinoderms of the Smithsonian-Hartford Expedition, 1937, with other West Indian records. Proceedings of the U. S. National Museum 86: 441456. https://doi.org/10.5479/si.00963801.86-3056.441" title="Echinoderms of the Smithsonian-Hartford Expedition, 1937, with other West Indian records. Proceedings of the U. S." volume="86" year="1939">Clark 1939</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B30EC22E21D8D86E3C72207C999B57CC" authority="(Madsen 1956)" baseAuthorityName="Madsen" baseAuthorityYear="1956" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="mortenseni">
<emphasis id="97A1BF654D6F02497336F54F5DB94169" italics="true">Freyastera mortenseni</emphasis>
(
<bibRefCitation id="26F4605AAA787F518ABD7A8DCE1163C6" author="Madsen" firstAuthor="Madsen" journalOrPublisher="Galathea Report" pagination="23-32" refId="B39" refString="Madsen FJ (1956) Echinoidea, Asteroidea, and Ophiuroidea from depths exceeding 6000 meters. Galathea Report 2: 2332." title="Echinoidea, Asteroidea, and Ophiuroidea from depths exceeding 6000 meters." volume="2" year="1956">Madsen 1956</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="3D1593866C808576224F12C51095BB7C" authority="(Perrier 1885)" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="BC683C57C8C6BB93049D34FA3EF9C84A" italics="true">Freyastera sexradiata</emphasis>
(
<bibRefCitation id="13364B749028313DF6A6C9D633CA5361" author="Perrier" firstAuthor="Perrier" issue="8" journalOrPublisher="Annales des sciences naturelles, Zoologie" pagination="1-72" refId="B53" refString="Perrier E (1885) Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman. Annales des sciences naturelles, Zoologie 22 (8): 172. [in French]" title="Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman." volume="22" year="1885">Perrier 1885</bibRefCitation>
)
</taxonomicName>
; and
<taxonomicName id="A198E5F404F8407AF89EF1DFADFD8BBA" authority="(Sladen 1889)" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="tuberculata">
<emphasis id="320A06EFA1A83E7EEE4C0EB1C026166A" italics="true">Freyastera tuberculata</emphasis>
(
<bibRefCitation id="962CBC47AB7F3C87B6816B120ABF9A16" author="Sladen" firstAuthor="Sladen" refId="B61" refString="Sladen WP (1889) Report on the Asteroidea. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 18731876, Zoology 30 (51): 125174." year="1889">Sladen 1889</bibRefCitation>
)
</taxonomicName>
. A new species,
<taxonomicName id="0ED700EB4B899125BF37B5F3155ED7A2" authorityName="Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang" authorityYear="2025" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="jiaolongi" status="sp. nov.">
<emphasis id="F16913B88EB6693751C8C24228CEC43D" italics="true">Freyastera jiaolongi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DDF1BE7EC4925545CE21C101D78250F6" rank="species">sp. nov.</taxonomicNameLabel>
, is described in the present study. Two species previously belonged to
<taxonomicName id="0CCFE4FF6C54CAE87131EB202158602C" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="D0A1B8A9E548CC6CC98650A2F6BF6C74" italics="true">Freyella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C2C37819B50EE246791EB8141BD38" authority="Koehler, 1907" authorityName="Koehler" authorityYear="1907" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="giardi">
<emphasis id="12627D5A2329FFE1A5DE5A0E7792EDE4" italics="true">F. giardi</emphasis>
<bibRefCitation id="8619796BA8595548319BD4F849799BA4" author="Koehler R" issue="6" journalOrPublisher="Zoologischer Anzeiger" pagination="140-147" refId="B30" refString="Koehler R (1907) Asteries, Ophiures et Echinides recueillis dans les mers australes par la &quot; Scotia &quot; (1902 - 1904). Zoologischer Anzeiger 32 (6): 140-147. [in French]" title="Asteries, Ophiures et Echinides recueillis dans les mers australes par la “ Scotia ” (1902-1904)." volume="32" year="1907">Koehler, 1907</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="88D0B85A982179633971876313BCEEA6" authority="Korovchinsky &amp; Galkin, 1984" authorityName="Korovchinsky &amp; Galkin" authorityYear="1984" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="loricata">
<emphasis id="CFD4910DCC4A7E76B4465786F2A43D61" italics="true">F. loricata</emphasis>
<bibRefCitation id="ABFDC735B96D4E515131C4BDA5CC784C" author="Korovchinsky NM &amp; Galkin SV" issue="8" journalOrPublisher="Zoologicheskii Zhurnal" pagination="1205-1215" refId="B34" refString="Korovchinsky NM, Galkin SV (1984) New data on the fauna of starfishes of the genus Freyella (Brisingidae). Zoologicheskii Zhurnal 63 (8): 1205-1215. [in Russian with English abstract]" title="New data on the fauna of starfishes of the genus Freyella (Brisingidae)." volume="63" year="1984">Korovchinsky &amp; Galkin, 1984</bibRefCitation>
</taxonomicName>
, are reassigned here to
<taxonomicName id="D86495AA53154787F60716EF331FE3D5" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="FAB6168AA9FDF501A6900B59AEEECABD" italics="true">Freyastera</emphasis>
</taxonomicName>
as new species combinations. The genus is hence composed of 10 species in total. Furthermore, seven unnamed species are included in the molecular analyses:
<taxonomicName id="B7B0EA3B4BB574C0048FD97E5EC4B314" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-2">
<emphasis id="C89D54C923855A59919DEB925F18DE22" italics="true">Freyastera</emphasis>
sp. 2
</taxonomicName>
,
<taxonomicName id="49035233972B4C776F22C601F653DE1B" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-3">
<emphasis id="795E2739B4F221C00778C74F18B43569" italics="true">Freyastera</emphasis>
sp. 3
</taxonomicName>
,
<taxonomicName id="EF4414920CEA30388ED02A6D11A9FFBC" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-5">
<emphasis id="9D10C97E60E09216E467C4082E4870CC" italics="true">Freyastera</emphasis>
sp. 5
</taxonomicName>
,
<taxonomicName id="C86B74D3DE17B5E2D4B7EBBB859F64EC" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-6">
<emphasis id="C88D4FC115C351683406F969E081202E" italics="true">Freyastera</emphasis>
sp. 6
</taxonomicName>
,
<taxonomicName id="7752B9EF3C67E7B922B0CBFD7B9EE507" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-7">
<emphasis id="9A32F6F1CF97133A2BFC5182F79E2334" italics="true">Freyastera</emphasis>
sp. 7
</taxonomicName>
,
<taxonomicName id="D6098BB1D71C98CAD4FCD1A27A8EEB2D" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="E6DD4B5CD7CA1A7F716D331CE4B26401" italics="true">Freyastera</emphasis>
sp.
</taxonomicName>
Yap, and
<taxonomicName id="4A950E2697B5E51A5698F43186B2D0AB" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" isUncertain="true" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="tuberculata">
<emphasis id="2CA098A2696B5C63A81DA07C462DE221" italics="true">Freyastera</emphasis>
cf.
<emphasis id="3F1E4C6F948650185A2F648AAAA22E20" italics="true">tuberculata</emphasis>
</taxonomicName>
. Among them, precise identification of
<taxonomicName id="7F78F144174FEDAEB7744FF5EB250B2C" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-2">
<emphasis id="1EA4C44BAB8D7D77F5F082F19E6459A6" italics="true">Freyastera</emphasis>
sp. 2
</taxonomicName>
,
<taxonomicName id="70B382F7A88D5E754C53A33D5A0E6476" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-7">
<emphasis id="F40AF17DA2D33913D12BBF2F2265431B" italics="true">Freyastera</emphasis>
sp. 7
</taxonomicName>
, and
<taxonomicName id="311ED4285869288E42D6C4322A20F354" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" isUncertain="true" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="tuberculata">
<emphasis id="A0CA27123B90EA0B709469B7DF5CAD31" italics="true">Freyastera</emphasis>
cf.
<emphasis id="8FC51C2462E24853CDD1C75E0301E4B6" italics="true">tuberculata</emphasis>
</taxonomicName>
was not feasible, as they were morphologically similar to
<taxonomicName id="A4E6DA19BC5C756F52234B57A8B0B4C8" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="tuberculata">
<emphasis id="FF6D3B75C60E57402D30B5DFCABEB774" italics="true">Freyastera tuberculata</emphasis>
</taxonomicName>
,
<taxonomicName id="B051FA22D140458E4AD96EFCA722CD08" authorityName="Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang" authorityYear="2025" baseAuthorityName="Koehler" baseAuthorityYear="1907" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="giardi" status="comb. nov.">
<emphasis id="3F360C710C6ADED271F52C40B61E8EFB" italics="true">Freyastera giardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2102229DA527010C56B415FAD2FD51AA" rank="species">comb. nov.</taxonomicNameLabel>
, and
<taxonomicName id="AF30BB4F72D92C24F809EF78E046997F" authorityName="Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang" authorityYear="2025" baseAuthorityName="Korovchinsky &amp; Galkin" baseAuthorityYear="1984" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="loricata" status="comb. nov.">
<emphasis id="708C6E7D53004A043DBDA49F64690AE8" italics="true">Freyastera loricata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CDC0C23B3033C07F39462781E5791BA" rank="species">comb. nov.</taxonomicNameLabel>
(see
<taxonomicName id="AE423358F65450E96F73FAE49C9F4415" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="tuberculata">
<emphasis id="B4199621F0883A9429C796F99CB86C26" italics="true">Freyastera tuberculata</emphasis>
</taxonomicName>
species complex).
<taxonomicName id="C22738ABEFED9958022E3402E22A3B01" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="539ED44AB2AC1A8C65BD10429E1D8323" italics="true">Freyastera</emphasis>
sp.
</taxonomicName>
Yap (
<specimenCode id="C339872EFAFE8071F308DDC01232B313">RSIOAST 0041</specimenCode>
) was reported based on one broken arm (
<bibRefCitation id="A084388EA89FBAE7B438321E7A0D592C" DOI="10.1016/j.dsr.2019.103163" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al. 2019</bibRefCitation>
), whereas specimens of
<taxonomicName id="7B0C1B0BC3DF289898F209AF5A4707C1" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-3">
<emphasis id="D065891B4F932284D899827996EA9AB2" italics="true">Freyastera</emphasis>
sp. 3
</taxonomicName>
(
<specimenCode id="AD15B378085321464C2ACB50B8C3F4F7">RSIOAST 0057</specimenCode>
),
<taxonomicName id="6F7B99D8B57D1191DEBC4C3A650293FD" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-5">
<emphasis id="3F5786E0A57AF5EBF8B316DC7885E519" italics="true">Freyastera</emphasis>
sp. 5
</taxonomicName>
(
<specimenCode id="685AF11AE81FC6390B74FA4BC4956DDC">RSIOAST 0107</specimenCode>
,
<specimenCode id="23361FF10FECBC1EB1770B1B0C3090EF">RSIOAST 0116</specimenCode>
,
<specimenCode id="653DDE0656F6939432FF20D3462EFAFB">RSIOAST 0138</specimenCode>
) (Fig.
<figureCitation id="1F02F569B4208D3696F32D2DADD90215" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. In situ photos of Freyastera species. A. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117; B. Freyastera jiaolongi sp. nov. paratype RSIOAST 0112; C. Freyastera jiaolongi sp. nov. paratype RSIOAST 0113; D. Freyastera delicata RSIOAST 0135; E. Freyastera basketa RSIOAST 0200; F. Freyastera sp. 5 RSIOAST 0116; G. Freyastera sp. 5 RSIOAST 0107; H. Freyastera sp. 6 RSIOAST 0124; I. Freyastera sp. 6 RSIOAST 0125." figureDoi="10.3897/zse.101.144918.figure14" httpUri="https://binary.pensoft.net/fig/1304665">14 F G</figureCitation>
), and
<taxonomicName id="1B5A815935258344D3B80A1B58537DAD" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-6">
<emphasis id="4BB1AAEEC2C1463541CE736E8B84F437" italics="true">Freyastera</emphasis>
sp. 6
</taxonomicName>
(
<specimenCode id="06B9B4098B645AFC92981E9DD5ED8331">RSIOAST 0124</specimenCode>
,
<specimenCode id="F9667020B84F38380851443898B11EA9">RSIOAST 0125</specimenCode>
) (Fig.
<figureCitation id="F54A03E52A2EB731FFE7564E41BA847F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. In situ photos of Freyastera species. A. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117; B. Freyastera jiaolongi sp. nov. paratype RSIOAST 0112; C. Freyastera jiaolongi sp. nov. paratype RSIOAST 0113; D. Freyastera delicata RSIOAST 0135; E. Freyastera basketa RSIOAST 0200; F. Freyastera sp. 5 RSIOAST 0116; G. Freyastera sp. 5 RSIOAST 0107; H. Freyastera sp. 6 RSIOAST 0124; I. Freyastera sp. 6 RSIOAST 0125." figureDoi="10.3897/zse.101.144918.figure14" httpUri="https://binary.pensoft.net/fig/1304665">14 H, I</figureCitation>
) were severely damaged during collection, with only arm fragments available, or kept frozen after collection, which hinders proper morphological identification and description. However, genetic information from these specimens showed high diversity, and species delimitation tools set them as seven species units, distinguishable from other known species with genetic data available. They were therefore included in the present study to discuss the phylogeny and diversity of
<taxonomicName id="72817AF937F493CAA59AE30CD7B6E72E" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="A8AE2B4F76468C6CEEF2B4276415FAE7" italics="true">Freyastera</emphasis>
</taxonomicName>
. Proper definition and description of these species units need to be done with more specimens in good condition in the future.
</paragraph>
<paragraph id="7F43CFE9BC08CB6BD99710DFD2377A77">
The following systematic accounts include a key, diagnoses, illustrations, and remarks on each known
<taxonomicName id="C31B59759EC0AAB96F23E5160083F079" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="89B45B50457B843B0CCCDFACF2E922E8" italics="true">Freyastera</emphasis>
</taxonomicName>
species or species complex based on specimens or photos examined, except one species,
<taxonomicName id="C37286980E394CB6ABEEF4A7F6FAF781" authority="McKnight, 2006" authorityName="McKnight" authorityYear="2006" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="digitata">
<emphasis id="1FDB2314CFDE92AB4D9F38F7BEE57D3F" italics="true">Freyastera digitata</emphasis>
<bibRefCitation id="534791FF0684DAF3D5C3F6980EA0E3E0" author="McKnight DG" journalOrPublisher="NIWA Biodiversity Memoir" pagination="1-187" refId="B44" refString="McKnight DG (2006) The marine fauna of New Zealand, Echinodermata: Asteroidea (Sea-stars). 3. Orders Velatida, Spinulosida, Forcipulatida, Brisingida with addenda to Paxillosida, Valvatida. NIWA Biodiversity Memoir 120: 1-187." title="The marine fauna of New Zealand, Echinodermata: Asteroidea (Sea-stars). 3. Orders Velatida, Spinulosida, Forcipulatida, Brisingida with addenda to Paxillosida, Valvatida." volume="120" year="2006">McKnight, 2006</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="34AD19B54695EE88A8E2E303140EFA78" authorityName="McKnight" authorityYear="2006" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="digitata">
<emphasis id="DCC379E5D19BC6054384BF7C0CEF8434" italics="true">F. digitata</emphasis>
</taxonomicName>
was described to have an inferomarginal plate corresponding to every 23 adambulacral plates (
<bibRefCitation id="2FEC928355E963F49A75743269CEFFA2" author="McKnight" firstAuthor="McKnight" journalOrPublisher="NIWA Biodiversity Memoir" pagination="1-187" refId="B44" refString="McKnight DG (2006) The marine fauna of New Zealand, Echinodermata: Asteroidea (Sea-stars). 3. Orders Velatida, Spinulosida, Forcipulatida, Brisingida with addenda to Paxillosida, Valvatida. NIWA Biodiversity Memoir 120: 1187." title="The marine fauna of New Zealand, Echinodermata: Asteroidea (Sea-stars). 3. Orders Velatida, Spinulosida, Forcipulatida, Brisingida with addenda to Paxillosida, Valvatida." volume="120" year="2006">McKnight 2006</bibRefCitation>
), but the
<typeStatus id="82C96FE9411238373CA2D318D3C5AD06" type="holotype">holotype</typeStatus>
was broken with only
<quantity id="ED556BA30DBE32C03F489A11BCD06332" metricMagnitude="-2" metricUnit="m" metricValue="2.2" unit="mm" value="22.0">22 mm</quantity>
of arm left, thus the arrangement of inferomarginal plates beyond the genital area is largely unknown. This species greatly resembles
<taxonomicName id="F65FEF1038C4E821E98A4321A91E532F" authorityName="Sladen" authorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="9B08AC6661C7C0F0CD00DC4C03CEECAC" italics="true">Freyella benthophila</emphasis>
</taxonomicName>
in the absence of a furrow spine and abactinal plates with several short spinelets. The taxonomic position of
<taxonomicName id="79F47EC8AF7A7032617099489F6674B2" authorityName="McKnight" authorityYear="2006" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="digitata">
<emphasis id="2A4F1F4CBD1C415FED862302C907A3B6" italics="true">F. digitata</emphasis>
</taxonomicName>
needs to be re-evaluated with further examination of the
<typeStatus id="4756CBA62E8468DDF9955CFB9D9DB558" type="holotype">holotype</typeStatus>
and other complete specimens. It is thus not included in the diagnostic key of the genus for the time being.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,316 @@
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<paragraph id="4D6948F89BBB19190DB94B7F73426939">
<taxonomicName id="661544A32C9DB5C1C4108019E9F96EAC" ID-CoL="848TV" authority="Zhang et al., 2019" authorityName="Zhang" authorityYear="2019" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="basketa">
<emphasis id="ED21CE253949DA16E56F1D357DEE3C94" italics="true">Freyastera basketa</emphasis>
<bibRefCitation id="06C4EDFA0197C14A3E350315271F3962" DOI="10.1016/j.dsr.2019.103163" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al., 2019</bibRefCitation>
</taxonomicName>
</paragraph>
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<paragraph id="0E52014A4FD8AF65C24FBF53E43E9A5D">
<figureCitation id="B96784A849AA7F7100CBFFA10B0FD03D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Freyastera basketa Zhang et al., 2019. A F. RSIOAST 0200; G H. RSIOAST 0201. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Female gonad; F. Abactinal view of distal part of arm; G. Actinal view of arm genital area; H. Distalmost part of arm." figureDoi="10.3897/zse.101.144918.figure6" httpUri="https://binary.pensoft.net/fig/1304657">Figs 6</figureCitation>
,
<figureCitation id="F3BA30D2B317361BEBD18B523BE457F8" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. In situ photos of Freyastera species. A. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117; B. Freyastera jiaolongi sp. nov. paratype RSIOAST 0112; C. Freyastera jiaolongi sp. nov. paratype RSIOAST 0113; D. Freyastera delicata RSIOAST 0135; E. Freyastera basketa RSIOAST 0200; F. Freyastera sp. 5 RSIOAST 0116; G. Freyastera sp. 5 RSIOAST 0107; H. Freyastera sp. 6 RSIOAST 0124; I. Freyastera sp. 6 RSIOAST 0125." figureDoi="10.3897/zse.101.144918.figure14" httpUri="https://binary.pensoft.net/fig/1304665">14E</figureCitation>
</paragraph>
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<paragraph id="2DC5B2157D167F2EF055B3F71E48AF4D">
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<emphasis id="6F6E35D9D69D2B482FBD346AEB76DFAF" italics="true">Freyastera basketa</emphasis>
</taxonomicName>
<comment id="CF571414C3357173EEB9615B0BBBDBE0">
:
<treatmentCitation id="2820491BE97D50418BFDAD43DFD58EBE" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" page="4" year="2019">
<bibRefCitation id="E0672487E933DBBD7258F2CDF042C4A7" DOI="10.1016/j.dsr.2019.103163" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" etAl="et al." firstAuthor="Zhang" page="4" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al. 2019: 4</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="2F5F2E8E572EF01DCC14F0F848DA21F5" author="Zhang R &amp; Fau M &amp; Mah C &amp; Eléaume M &amp; Zhang D &amp; Zhou Y &amp; Lu B &amp; Wang C" page="2" year="2024">
<bibRefCitation id="B8A07C2134F0562248AAF1B76FC89343" DOI="10.1016/j.ympev.2023.107993" author="Zhang R &amp; Fau M &amp; Mah C &amp; Eléaume M &amp; Zhang D &amp; Zhou Y &amp; Lu B &amp; Wang C" etAl="et al." firstAuthor="Zhang" page="2" refId="B69" refString="Zhang R, Fau M, Mah C, Eléaume M, Zhang D, Zhou Y, Lu B, Wang C (2024) Systematics of deep-sea starfish order Brisingida (Echinodermata: Asteroidea), with a revised classification and assessments of morphological characters. Molecular Phylogenetics and Evolution 191: 107993. https://doi.org/10.1016/j.ympev.2023.107993" year="2024">Zhang et al. 2024: 2</bibRefCitation>
</treatmentCitation>
,
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</paragraph>
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<paragraph id="403C64A4632B77DC11F40C0582FCD143">
<heading id="5C9AA99C4F1F48A2C6EA6DC3B1A66386" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="4E123C598C3EA13A5303ADC348A6A83B">
<materialsCitation id="150A56887354395FAA8E65F2D29E65BF" collectionCode="RSIOAST" specimenCode="RSIOAST 0038" specimenCount="1" typeStatus="holotype">
<specimenCode id="3ADAC708A687E5B034F269CCF70A2FD4">RSIOAST 0038</specimenCode>
(
<emphasis id="8342AB311A5ED44187BD51DE399E4326" bold="true" italics="true">
<typeStatus id="96DF4152EBC263D4D7702305E7186277" type="holotype">holotype</typeStatus>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="D4CEDFD01D21039954FC7D31FD0CCC71" collectionCode="RSIOAST" specimenCode="RSIOAST 0006" specimenCount="1" typeStatus="paratype">
<specimenCode id="9AD987E281289DDC5663D15053455C84">RSIOAST 0006</specimenCode>
(
<emphasis id="D71A17E1B3FF472047A6ED7EE6D5ED40" bold="true" italics="true">
<typeStatus id="92462F32A38D728C9A9A81CB870FDB2B" type="paratype">paratype</typeStatus>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="EFD582D21D633919B17E2869129B4C33" collectionCode="RSIOAST" specimenCode="RSIOAST 0008" specimenCount="1" typeStatus="paratype">
<specimenCode id="A64581A08EE795578F1A399979898A4E">RSIOAST 0008</specimenCode>
(
<emphasis id="1939314884B9858E6A485110C8C5166F" bold="true" italics="true">
<typeStatus id="9D6EB2F9CFFACFBF26C9A6E3F05A12D6" type="paratype">paratype</typeStatus>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="10EC5F89D1B236BCB4567E71CDC03A8F" collectionCode="RSIOAST" specimenCode="RSIOAST 0039" specimenCount="1" typeStatus="paratype">
<specimenCode id="EF2EC4E8C413B668478B5BC2365F97AF">RSIOAST 0039</specimenCode>
(
<emphasis id="5B073870FA6F015577143C9515D108FD" bold="true" italics="true">
<typeStatus id="E829DED230ED5DAEB8E9152AD3A36CD6" type="paratype">paratype</typeStatus>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="8B64B5F626ACB922420EFB60A905254A" collectionCode="RSIOAST" specimenCode="RSIOAST 0200" specimenCount="1">
<specimenCode id="A6AD5FCB03047B8D686F5F2F71376B10">RSIOAST 0200</specimenCode>
(Figs
<figureCitation id="5C01771AE9BBD162FAA4087346D9FFE8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Freyastera basketa Zhang et al., 2019. A F. RSIOAST 0200; G H. RSIOAST 0201. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Female gonad; F. Abactinal view of distal part of arm; G. Actinal view of arm genital area; H. Distalmost part of arm." figureDoi="10.3897/zse.101.144918.figure6" httpUri="https://binary.pensoft.net/fig/1304657">6 A F</figureCitation>
,
<figureCitation id="24138959C50CF0AC4A8582B71917F829" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. In situ photos of Freyastera species. A. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117; B. Freyastera jiaolongi sp. nov. paratype RSIOAST 0112; C. Freyastera jiaolongi sp. nov. paratype RSIOAST 0113; D. Freyastera delicata RSIOAST 0135; E. Freyastera basketa RSIOAST 0200; F. Freyastera sp. 5 RSIOAST 0116; G. Freyastera sp. 5 RSIOAST 0107; H. Freyastera sp. 6 RSIOAST 0124; I. Freyastera sp. 6 RSIOAST 0125." figureDoi="10.3897/zse.101.144918.figure14" httpUri="https://binary.pensoft.net/fig/1304665">14 E</figureCitation>
)
</materialsCitation>
;
<materialsCitation id="7413FE1C42C83A3952997E10221184AD" collectionCode="RSIOAST" specimenCode="RSIOAST 0201" specimenCount="1">
<specimenCode id="36A6BA2AEB1E0198B9B053C1867ACB1F">RSIOAST 0201</specimenCode>
(Fig.
<figureCitation id="EAFD76F2456C1A2C706B0164441093F1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Freyastera basketa Zhang et al., 2019. A F. RSIOAST 0200; G H. RSIOAST 0201. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Female gonad; F. Abactinal view of distal part of arm; G. Actinal view of arm genital area; H. Distalmost part of arm." figureDoi="10.3897/zse.101.144918.figure6" httpUri="https://binary.pensoft.net/fig/1304657">6 G, H</figureCitation>
)
</materialsCitation>
;
<materialsCitation id="2EEB6B83B656A742032821C1BAA8EEB2" collectionCode="NHMUK" specimenCode="NHMUK 8963" specimenCount="1">
<specimenCode id="6674B6BEE24183BF51579E43E37035D9">
<named-content id="32F2BCDB27A0D93B8A91D0060C2AB06F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum">NHMUK</named-content>
8963
</specimenCode>
</materialsCitation>
;
<materialsCitation id="DDF13969B689794E0BDFED91E23F596A" collectionCode="SO" specimenCode="SO 268-2_174_116" specimenCount="1">
<specimenCode id="D0869B6EE784D5119D4F430D58D4B370">SO 268-2_174_116</specimenCode>
. (Suppl. material
<xref id="B6D3364AEF14D56550018FDA18F19E11" ref-type="supplementary-material" rid="S1">1</xref>
)
</materialsCitation>
.
</paragraph>
<caption id="527B17974613E5300A295534CBD22D17" ID-DOI="10.3897/zse.101.144918.figure6" ID-arpha="2F1CA906-E2B1-531F-BD1E-FC7CD3BF4D0F" httpUri="https://binary.pensoft.net/fig/1304657" startId="F6">
<paragraph id="D53AFA4C0C99C7013A476D07525664B4">
<label id="0E1D485E11FDF8502CC649021AEBF2BA">Figure 6.</label>
</paragraph>
<paragraph id="C9A6621606C36E9E58370D57A46BDF66">
<taxonomicName id="E7623CA0B73CE35C9CFC93F207377D2B" authority="Zhang et al., 2019" authorityName="Zhang" authorityYear="2019" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="basketa">
<emphasis id="9401A4CAE16782CCFF5CB5F63FF0625C" italics="true">Freyastera basketa</emphasis>
<bibRefCitation id="5A054D3B7F89C7B25DB41AC38BC283C5" DOI="10.1016/j.dsr.2019.103163" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al., 2019</bibRefCitation>
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.
<emphasis id="82E583FEA6E98FF56F9843CAF97ED6D5" bold="true">
A
<emphasis id="14892E8164812DDD9AF3B077C22D1D4B" italics="true">F.</emphasis>
</emphasis>
<specimenCode id="78670897B6C3B29B0B1A2AD34919EE31">RSIOAST 0200</specimenCode>
;
<emphasis id="E48210CA5D4C92040AE1DF867F95135B" bold="true">G H.</emphasis>
<specimenCode id="C2615F6FB84058E68FDD2EBD1FDD84F8">RSIOAST 0201</specimenCode>
.
<emphasis id="C58C9DED82F9A173F0C8B52ABD2848F1" bold="true">A.</emphasis>
Actinal view of disk;
<emphasis id="87C2C329B4AE064D0C3D60C5204C2787" bold="true">B.</emphasis>
Abactinal view of disk;
<emphasis id="E185A4B2252F628BF16C76A4CB8D8804" bold="true">C.</emphasis>
Actinal view of arm genital area;
<emphasis id="E3238971788758C3CA4E217DC9FC9125" bold="true">D.</emphasis>
Abactinal view of genital area;
<emphasis id="24BFCA5C82DCE2EA766AB285F1C6C0C8" bold="true">E.</emphasis>
Female gonad;
<emphasis id="52308A33210226C74FF0C777681CEDD0" bold="true" italics="true">F.</emphasis>
Abactinal view of distal part of arm;
<emphasis id="4A5880BB9A58DB96238255107D9C4B1B" bold="true">G.</emphasis>
Actinal view of arm genital area;
<emphasis id="9B746854A6F1571E055C800FCD427259" bold="true">H.</emphasis>
Distalmost part of arm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EBABG" type="diagnosis">
<paragraph id="0970CF1DBCC50001246C6E4B734439BC">
<heading id="B0787D05A74DD8C53296DA84F3DF07C9" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="3AF22BE2805E57E265DDAFD2D322D975">
Arms 6. Abactinal disk and arm plate each bears one long, sharp spine, about
<quantity id="5C1051DA475C8DAF26C7F2B2CB4C5B34" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
in length, not covered by membranous sheath. Enlarged pedicellariae (about
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in length) with curved valves present on oral spines and abactinal surface of disk and arm genital area. Small pedicellariae (less than
<quantity id="2B8B95B0FB2969C4A196B21414F1E103" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
in length) cluster in transverse bands on abactinal arm beyond genital area. One aboral furrow spine and one subambulacral spine on each adambulacral plate. Proximal subambulacral spines with truncate end. One lateral spine corresponding to every adambulacral plate beyond genital area. Oral plate bears one actinostomal spine, one suboral spine, and one furrow spine, all covered by large pedicellariae.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGABG" type="distribution">
<paragraph id="B9978740A2012FBFF10BFE89B14EE326">
<heading id="569DD028A3AD1784FFB62AF29DA1FE51" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="8803662B137CE0067E39C03F066319ED">
Northwest Pacific: Mariana Trench, Yap Trench, Lamont Seamount, Pigafetta Basin; Eastern Pacific: Clarion-Clipperton Zone (CCZ).
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. Type locality: Mariana Trench, Yap Trench, Lamont Seamount,
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.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELABG" type="remarks">
<paragraph id="63D86D99A2E7C9A021A051D83FEBDC66">
<heading id="103A0E13DF576C9B1083505D0FF18697" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="0AE7636ECAA64682EE50E831689F7E7C">
This species is characterized by having enlarged pedicellariae with curved valves on oral spines and abactinal surface of disk and arm genital area (Fig.
<figureCitation id="ABF98EC732081B3DE2B5CBFBC5662DA6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Freyastera basketa Zhang et al., 2019. A F. RSIOAST 0200; G H. RSIOAST 0201. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm genital area; D. Abactinal view of genital area; E. Female gonad; F. Abactinal view of distal part of arm; G. Actinal view of arm genital area; H. Distalmost part of arm." figureDoi="10.3897/zse.101.144918.figure6" httpUri="https://binary.pensoft.net/fig/1304657">6 A, B, G</figureCitation>
), and one long, unsheathed spine on each abactinal arm plate. In some specimens, the proximal adambulacral spines are also equipped with large pedicellariae. The large pedicellariae are found in other
<taxonomicName id="108A9649A319E95E7898D1B3961F8796" authorityName="Fisher" authorityYear="1928" class="Asteroidea" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="order">Brisingida</taxonomicName>
species, such as
<taxonomicName id="9EC927517917EFED6F0127FB39FF1C9E" authority="Korovchinsky &amp; Galkin, 1984" authorityName="Korovchinsky &amp; Galkin" authorityYear="1984" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="macropedicellaria">
<emphasis id="0BF00915166437ADF0C7390FE1F741EE" italics="true">Freyella macropedicellaria</emphasis>
<bibRefCitation id="4BE5A77BEFB082061A365BCC2E1D7D7A" author="Korovchinsky NM &amp; Galkin SV" issue="8" journalOrPublisher="Zoologicheskii Zhurnal" pagination="1205-1215" refId="B34" refString="Korovchinsky NM, Galkin SV (1984) New data on the fauna of starfishes of the genus Freyella (Brisingidae). Zoologicheskii Zhurnal 63 (8): 1205-1215. [in Russian with English abstract]" title="New data on the fauna of starfishes of the genus Freyella (Brisingidae)." volume="63" year="1984">Korovchinsky &amp; Galkin, 1984</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="501B191570212C6CAB2E7101F68EAC5F" authority="Sladen, 1889" authorityName="Sladen" authorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="remex">
<emphasis id="8CAFB4231B3F1DBCD81AAE5D871D6566" italics="true">Freyella remex</emphasis>
<bibRefCitation id="4DC6DAFA3896B506AECD74F458EA7C4E" author="Sladen WP" refId="B61" refString="Sladen WP (1889) Report on the Asteroidea. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876, Zoology 30 (51): 125 - 174." year="1889">Sladen, 1889</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="D993EC5BA133B302F1EF5268C040E7B6" authority="(Korovchinsky 1976)" baseAuthorityName="Korovchinsky" baseAuthorityYear="1976" class="Asteroidea" family="Freyellidae" genus="Astrocles" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="japonicus">
<emphasis id="1CA59B68407F752D8789E400FC34A44D" italics="true">Astrocles japonicus</emphasis>
(
<bibRefCitation id="B7B31FF7E49EC75AF0842CDF60CD1B79" author="Korovchinsky" firstAuthor="Korovchinsky" journalOrPublisher="Trudy Instituta Okeanologii Akademii Nauk SSSR" pagination="165-177" refId="B33" refString="Korovchinsky NM (1976) New data on the deep-sea seastars (Asteroidea, Brisingidae) from the North-Western Pacific. Trudy Instituta Okeanologii Akademii Nauk SSSR 99: 165177. [in Russian with English abstract]" title="New data on the deep-sea seastars (Asteroidea, Brisingidae) from the North-Western Pacific." volume="99" year="1976">Korovchinsky 1976</bibRefCitation>
)
</taxonomicName>
, as well as
<taxonomicName id="DF6DA97DF188A3530C6D42D38B8F85BE" class="Asteroidea" family="Freyellidae" genus="Freyellaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="6F53A4560914FE2B8CA7B2767A95E2F5" italics="true">Freyellaster</emphasis>
</taxonomicName>
species (
<bibRefCitation id="4587FA1D14A206F0946B26424883D864" DOI="10.1016/j.ympev.2023.107993" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B69" refString="Zhang R, Fau M, Mah C, Eléaume M, Zhang D, Zhou Y, Lu B, Wang C (2024) Systematics of deep-sea starfish order Brisingida (Echinodermata: Asteroidea), with a revised classification and assessments of morphological characters. Molecular Phylogenetics and Evolution 191: 107993. https://doi.org/10.1016/j.ympev.2023.107993" year="2024">Zhang et al. 2024</bibRefCitation>
). In
<taxonomicName id="809E1EA5D1E9A762AD75DA8A651C3FE0" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="6275282D5CFBC4B3E9BEA520D3D123E1" italics="true">Freyastera</emphasis>
</taxonomicName>
, only
<taxonomicName id="A38A1C06023B1322E920D943C6BDAD02" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="basketa">
<emphasis id="B9EFF7A1B08487CF935E54428FD5DDA6" italics="true">F. basketa</emphasis>
</taxonomicName>
and
<taxonomicName id="1CD4678EFB84EC70FA7F2EE7C000E9C6" baseAuthorityName="Madsen" baseAuthorityYear="1956" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="mortenseni">
<emphasis id="5B6A66B509348F6986C3D5C521D946B7" italics="true">F. mortenseni</emphasis>
</taxonomicName>
were found to have large pedicellariae.
<taxonomicName id="6CEEAFC51F2E9FB824261BA9C80090A2" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="basketa">
<emphasis id="BDDBEF2C4D4AE97A22260D6ECC664EDD" italics="true">F. basketa</emphasis>
</taxonomicName>
has rather long, sharp spines on abactinal disk and arms, one to each plate, whereas
<taxonomicName id="3DD9952148DC422CE6C777B553CAEBF4" baseAuthorityName="Madsen" baseAuthorityYear="1956" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="mortenseni">
<emphasis id="BCF53B3D30C095BFF17158C5EE6F8AA2" italics="true">F. mortenseni</emphasis>
</taxonomicName>
has several spines on each plate. Furthermore,
<taxonomicName id="866955D0F4A6A45EBC69BAD920EC5577" baseAuthorityName="Madsen" baseAuthorityYear="1956" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="mortenseni">
<emphasis id="44A60EAECDAB11F524654025F9D3AD27" italics="true">F. mortenseni</emphasis>
</taxonomicName>
was reported at greater depth (
<quantity id="C09DC2AB343C9144D586326F4CFFF885" metricMagnitude="3" metricUnit="m" metricValue="6.025" metricValueMax="6.2" metricValueMin="5.85" unit="m" value="6025.0" valueMax="6200.0" valueMin="5850.0">5850 to 6200 m</quantity>
) than
<taxonomicName id="28C3506E285AED6CB8724998246D863B" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="basketa">
<emphasis id="06064A338FEA5EB5BDCCD12FDF1A036B" italics="true">F. basketa</emphasis>
</taxonomicName>
(
<quantity id="A2B7CE7B3ABBCB8E2125BAE81234D494" metricMagnitude="3" metricUnit="m" metricValue="4.564" metricValueMax="4.991" metricValueMin="4.137" unit="m" value="4564.0" valueMax="4991.0" valueMin="4137.0">4137 to 4991 m</quantity>
). The newly examined specimens extend the distribution range of
<taxonomicName id="A663C41A4503720DA19BDD2F4D96978C" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="basketa">
<emphasis id="DCB395C13BD46C055E7C60A6218691A0" italics="true">F. basketa</emphasis>
</taxonomicName>
to CCZ in the Eastern Pacific (Fig.
<figureCitation id="0BBE1E2C735A1F1B3F2B72BCCCAFFC2B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Freyastera and six-armed Freyella specimens examined in the present study and type localities of each species." figureDoi="10.3897/zse.101.144918.figure1" httpUri="https://binary.pensoft.net/fig/1304652">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,483 @@
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<paragraph id="D15628ECC9CA7703B5B2BD3B751F2E95">
<taxonomicName id="FA634A05411376E772BB9CF43D6F48F1" authorityName="Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang" authorityYear="2025" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="jiaolongi" status="sp. nov.">
<emphasis id="59B0F9DFD952810628321747F87BB277" italics="true">Freyastera jiaolongi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C778957FD352CA89A93D2D93619EF106" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="879844D3BD131EB0F2BDCD683A427702" type="description">
<paragraph id="2554B456BEC046C58069B518F2A7DB32">
<figureCitation id="A92B09B4E01FBDD590A045AE4183B492" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm in genital area; D. Abactinal view of arm in genital area; E. Actinal view of middle part of arm; F. Actinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure2" httpUri="https://binary.pensoft.net/fig/1304653">Figs 2</figureCitation>
,
<figureCitation id="12EB0F3488C20D62C670E86EEAC43606" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A D. Abactinal view of arm, from basal part to beyond genital area; E. Abactinal view of middle part of arm; F. Abactinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure3" httpUri="https://binary.pensoft.net/fig/1304654">3</figureCitation>
,
<figureCitation id="FA428D8917FE33C2643CAFBE9D00643C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Freyastera jiaolongi sp. nov. paratypes. A, D. RSIOAST 0112; B, C, E, F. RSIOAST 0 AS 113. A, B. Abactinal view of arm in genital area; C. Actinal view of arm in genital area; D. Female gonads; E. Actinal view of middle part of arm; F. Abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure4" httpUri="https://binary.pensoft.net/fig/1304655">4</figureCitation>
,
<figureCitation id="7B89396E994D14F51B42A0ECB2D9F361" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. In situ photos of Freyastera species. A. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117; B. Freyastera jiaolongi sp. nov. paratype RSIOAST 0112; C. Freyastera jiaolongi sp. nov. paratype RSIOAST 0113; D. Freyastera delicata RSIOAST 0135; E. Freyastera basketa RSIOAST 0200; F. Freyastera sp. 5 RSIOAST 0116; G. Freyastera sp. 5 RSIOAST 0107; H. Freyastera sp. 6 RSIOAST 0124; I. Freyastera sp. 6 RSIOAST 0125." figureDoi="10.3897/zse.101.144918.figure14" httpUri="https://binary.pensoft.net/fig/1304665">14 A C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="92E0FBB574855C1E65502658FA3A148A" type="reference_group">
<paragraph id="B2B38CB006945065BCE278A42749D136">
<treatmentCitationGroup id="3B61BF57D38AEED846555513E7227C0A">
<treatmentCitation id="5B891A7396883F1910C470097C8C1787">
<taxonomicName id="B51B977D1B16351E301EE01995A2BB41" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-4">
<emphasis id="1A7D6CB256347787DB351FE01A85BC5E" italics="true">Freyastera</emphasis>
sp. 4
</taxonomicName>
:
<bibRefCitation id="0D1026597ECD0D0913CA8AF3FB452460" DOI="10.1016/j.ympev.2023.107993" author="Zhang" etAl="et al." firstAuthor="Zhang" refId="B69" refString="Zhang R, Fau M, Mah C, Eléaume M, Zhang D, Zhou Y, Lu B, Wang C (2024) Systematics of deep-sea starfish order Brisingida (Echinodermata: Asteroidea), with a revised classification and assessments of morphological characters. Molecular Phylogenetics and Evolution 191: 107993. https://doi.org/10.1016/j.ympev.2023.107993" year="2024">Zhang et al. 2024</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3BAG" type="material">
<paragraph id="8AC43D13FFF637627864B5331B8983A9">
<heading id="2BEC66F0B7A2D9F7B6B9B5E2E72D5D78" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="5F3F57035220A2D2AC77FFCB482B3BA0">
<materialsCitation id="25EB05FBB967BFC1FC5153BA5FB017DF" collectingDate="2021-11-16" collectionCode="RSIOAST" elevation="3523" latitude="12.330671" location="Kyushu-Palau Ridge" longLatPrecision="1" longitude="134.4721" specimenCode="RSIOAST 0117" specimenCount="1" typeStatus="holotype">
<emphasis id="FA0D0F865F0725B166758CDC273C9772" bold="true">
<emphasis id="A014E0D0C1A32BCA09FA6F3AF58437BE" bold="true" italics="true">
<typeStatus id="008C0D454AE3767A39A3202DCF5BDA25" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCode id="FE2A97CF64B77097E502BF3E6E9343AD">RSIOAST 0117</specimenCode>
(Figs
<figureCitation id="5A48F14BD402740B15DDFD681F8804C9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm in genital area; D. Abactinal view of arm in genital area; E. Actinal view of middle part of arm; F. Actinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure2" httpUri="https://binary.pensoft.net/fig/1304653">2</figureCitation>
,
<figureCitation id="34AD494614980526C9D5F63B759D71BB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A D. Abactinal view of arm, from basal part to beyond genital area; E. Abactinal view of middle part of arm; F. Abactinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure3" httpUri="https://binary.pensoft.net/fig/1304654">3</figureCitation>
),
<location id="C80219E2C392446605435CCDBE18F825" LSID="urn:lsid:plazi:treatment:C5234970A9EB5EAFBD7BBA2B0AC2AC29:C80219E2C392446605435CCDBE18F825" latitude="12.330671" longLatPrecision="1" longitude="134.4721" name="Kyushu-Palau Ridge">Kyushu-Palau Ridge</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESCAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[134.472114,12.330671]}">
<geoCoordinate id="23556FEA962290B60D7C9EF5ED4C0A08" degrees="12.330671" direction="north" orientation="latitude" precision="1" value="12.330671">12.330671°N</geoCoordinate>
,
<geoCoordinate id="914D0E669626C86B14F3BC34E0DDD600" degrees="134.472114" direction="east" orientation="longitude" precision="1" value="134.4721">134.472114°E</geoCoordinate>
</named-content>
,
<quantity id="1BCB4F3A8F8A8D2E56620EE541825F9E" metricMagnitude="3" metricUnit="m" metricValue="3.523" unit="m" value="3523.0">
<elevation id="A1CC2E03A0EC7C29145D745F9A1BA652" metricMagnitude="3" metricUnit="m" metricValue="3.523" unit="m" value="3523.0">3523 m</elevation>
</quantity>
;
<date id="3ECF20CD59D1767B24FF147F6FAAF0C3" value="2021-11-16">
<collectingDate id="A5DF4DBA2CD08072A4B1C384801F79CE" value="2021-11-16">16 Nov. 2021</collectingDate>
</date>
; cruise DY 68, dive JL 199
</materialsCitation>
.
<materialsCitation id="0F5C7DDB49FDC64509CCFB3C3BA24B78" collectingDate="2021-11-13" collectionCode="RSIOAST" elevation="3641" latitude="13.30964" location="Kyushu-Palau Ridge" longLatPrecision="1" longitude="134.59198" specimenCode="RSIOAST 0112" specimenCount="1" typeStatus="paratype">
<emphasis id="6D7552D0E4CE920C6DDAEC7158C84FC9" bold="true">
<emphasis id="C036AF06B3250B0348ACF3BD7380B35D" bold="true" italics="true">
<typeStatus id="6DECDA484CE9DA097975087CC20B0C16" type="paratype">Paratype</typeStatus>
</emphasis>
1.
</emphasis>
<specimenCode id="B957F10C9EFA0FB784F0CB73C5040D0B">RSIOAST 0112</specimenCode>
(Fig.
<figureCitation id="D5787C5135840F99332E436640E7C9ED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A D. Abactinal view of arm, from basal part to beyond genital area; E. Abactinal view of middle part of arm; F. Abactinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure3" httpUri="https://binary.pensoft.net/fig/1304654">3 A, D</figureCitation>
),
<location id="2C1881303B9ADACBF7449C4FCA3A0718" LSID="urn:lsid:plazi:treatment:C5234970A9EB5EAFBD7BBA2B0AC2AC29:2C1881303B9ADACBF7449C4FCA3A0718" latitude="13.30964" longLatPrecision="1" longitude="134.59198" name="Kyushu-Palau Ridge">Kyushu-Palau Ridge</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECDAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[134.591983,13.309640]}">
<geoCoordinate id="24FE93917A5294F016DE7C7B296F6328" degrees="13.309640" direction="north" orientation="latitude" precision="1" value="13.30964">13.309640°N</geoCoordinate>
,
<geoCoordinate id="380E3EB2762A1F6C020BC5BD7E7FBF45" degrees="134.591983" direction="east" orientation="longitude" precision="1" value="134.59198">134.591983°E</geoCoordinate>
</named-content>
,
<quantity id="75AAB7AA3E6FA5341EBBEF75D6DE45F6" metricMagnitude="3" metricUnit="m" metricValue="3.641" unit="m" value="3641.0">
<elevation id="EC0840454BED8D4837B31FAC53CAC355" metricMagnitude="3" metricUnit="m" metricValue="3.641" unit="m" value="3641.0">3641 m</elevation>
</quantity>
;
<date id="8A02A1ECE884F4373F86518F8FF3B057" value="2021-11-13">
<collectingDate id="C6CF001554FF616C0D31C2BDC98A1168" value="2021-11-13">13 Nov. 2021</collectingDate>
</date>
; cruise DY 68, dive JL 197
</materialsCitation>
.
<materialsCitation id="0B6A92FB15F466CFCF64644ACF0A03CE" collectingDate="2021-11-13" collectionCode="RSIOAST" elevation="3541" latitude="13.310851" location="Kyushu-Palau Ridge" longLatPrecision="1" longitude="134.58904" specimenCode="RSIOAST 0113" specimenCount="1" typeStatus="paratype">
<emphasis id="AF876D343FC36AED794C890E875E6657" bold="true">
<emphasis id="8D5F5E62F158D2DC1D96BC6B4AF831DE" bold="true" italics="true">
<typeStatus id="693F7551B357DD8D247AB101EFCEA65E" type="paratype">Paratype</typeStatus>
</emphasis>
2.
</emphasis>
<specimenCode id="C467D4B11D09370241899D482F1536B1">RSIOAST 0113</specimenCode>
(Fig.
<figureCitation id="61388EF6A060F32774A93C711EB72C6D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A D. Abactinal view of arm, from basal part to beyond genital area; E. Abactinal view of middle part of arm; F. Abactinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure3" httpUri="https://binary.pensoft.net/fig/1304654">3 B, C, E, F</figureCitation>
),
<location id="F14382C7CC2D97F0ABAF03E2D75B99C6" LSID="urn:lsid:plazi:treatment:C5234970A9EB5EAFBD7BBA2B0AC2AC29:F14382C7CC2D97F0ABAF03E2D75B99C6" latitude="13.310851" longLatPrecision="1" longitude="134.58904" name="Kyushu-Palau Ridge">Kyushu-Palau Ridge</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESDAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[134.589032,13.310851]}">
<geoCoordinate id="6E3B93C0F73D077E868A9DB279B955D5" degrees="13.310851" direction="north" orientation="latitude" precision="1" value="13.310851">13.310851°N</geoCoordinate>
,
<geoCoordinate id="E7A0BEAC0527BF9E5C3A913DE4D86D05" degrees="134.589032" direction="east" orientation="longitude" precision="1" value="134.58904">134.589032°E</geoCoordinate>
</named-content>
,
<quantity id="6BA53AFE387B595E6AC0699EB5560626" metricMagnitude="3" metricUnit="m" metricValue="3.541" unit="m" value="3541.0">
<elevation id="DA18E9EED038264A178D5568D6CABCA3" metricMagnitude="3" metricUnit="m" metricValue="3.541" unit="m" value="3541.0">3541 m</elevation>
</quantity>
;
<date id="60204F5D6B897FD9836C0742CEF5B064" value="2021-11-13">
<collectingDate id="A344CD9B508E5AA2FD52A3263F595393" value="2021-11-13">13 Nov. 2021</collectingDate>
</date>
; cruise DY 68, dive JL 197
</materialsCitation>
. All type specimens were deposited at the Repository of Second Institute of Oceanography, Ministry of Natural Resources,
<collectingCountry id="B5543D8E07B0569D538EC39D7B0A8558" name="China">China</collectingCountry>
(
<collectionCode id="D628A53C1D1487E387481733776CEE65">SIOMNR</collectionCode>
) (Suppl. material
<xref id="522F0B32393394C836101CC369BFDA6D" ref-type="supplementary-material" rid="S1">1</xref>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2DAG" type="diagnosis">
<paragraph id="5D942F4136717A63FD54B0A370456293">
<heading id="3F4660F617DE23F22F8A459FBDB206B6" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="E62AC5D44B61220799F93BC1FE070D00">
Arms six. Abactinal disk with short hirsute spinelets, about
<quantity id="B112260A30C5FDF18D069C82E102CD52" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
in length, surrounded by small pedicellariae. Abactinal arm plates each bears one spine, covered with a membranous sheath loaded with small pedicellariae. Pedicellariae on abactinal disk and arm less than
<quantity id="63D678756CD85B1C81B89340F900FACD" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
in length. Pedicellariae cluster in pads or transverse bands on abactinal arm beyond genital area. Each adambulacral plate with 13 aboral furrow spines and one subambulacral spine. Lateral spines long, corresponding to each adambulacral plate beyond genital area. Oral plate with 12 actinostomal spines, one suboral spine, and one aboral spine along the furrow margin.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAEAG" type="description">
<paragraph id="7409FB45B042A09960DA1B21BF01436F">
<heading id="EDBBA61C09C98BF2BD06908DEDC7F041" reason="title">Description.</heading>
</paragraph>
<paragraph id="243F0816E61664A286BA9BE784134260">
<typeStatus id="2AD7805C9263BA4C285D31455CFAC202" type="holotype">Holotype</typeStatus>
: a complete specimen, with one arm regenerating. r =
<quantity id="0CCFD2C9FCC933F0C7F04FB263D03791" metricMagnitude="-3" metricUnit="m" metricValue="5.5" unit="mm" value="5.5">5.5 mm</quantity>
, R about
<quantity id="047F60700B81EAEA9600FE307B3DBF42" metricMagnitude="-1" metricUnit="m" metricValue="2.2" unit="mm" value="220.0">220 mm</quantity>
. Height of disk
<quantity id="608C25AE72BC828C3D8385147C3C2CD9" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
. Arm measures
<quantity id="69F88C77E04E908ECD310CA0837AA82F" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
at base,
<quantity id="34806129727DD4AAE85BDB42F6F25F15" metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
at the widest part. Genital inflation extends about
<quantity id="9A488028BC6BE979598C0E8E6C2F7B60" metricMagnitude="-2" metricUnit="m" metricValue="1.35" unit="mm" value="13.5">13.5 mm</quantity>
. Female.
<typeStatus id="696FBC87920978F82A0EDEF0CF45AFD3" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="C43E29149CB1A351FBD10568F6CA407C">RSIOAST 0112</specimenCode>
&amp;
<specimenCode id="2156DC9B977C657A0C2923D25CB65299">RSIOAST 0113</specimenCode>
, disks absent, seven detached arms mixed in one jar. Longest arm measures about
<quantity id="6011ED7656C5FAF5BB1DCCA77740829B" metricMagnitude="-1" metricUnit="m" metricValue="2.3" unit="mm" value="230.0">230 mm</quantity>
. Arm
<quantity id="53BCBDF0058446BD0BACD95B3D3DC4AC" metricMagnitude="-3" metricUnit="m" metricValue="4.1" unit="mm" value="4.1">4.1 mm</quantity>
wide at base,
<quantity id="2C8BC5ABAE77863BDA51C3889B1D7783" metricMagnitude="-3" metricUnit="m" metricValue="4.7" unit="mm" value="4.7">4.7 mm</quantity>
at the widest part. Genital inflation extends about
<quantity id="A4B2001E85C31F99B73E7D553177D043" metricMagnitude="-2" metricUnit="m" metricValue="1.6" unit="mm" value="16.0">16 mm</quantity>
. Female.
</paragraph>
<paragraph id="2D9C803502DF35822A1478EC3B88D51C">
Abactinal surface of disk with small rounded plates, each with one short spinelet measuring about
<quantity id="2E70D9DBE66D271F0B653099BE6AC790" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
in length (Fig.
<figureCitation id="148C12B7D0501ED35063A40E057B760D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm in genital area; D. Abactinal view of arm in genital area; E. Actinal view of middle part of arm; F. Actinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure2" httpUri="https://binary.pensoft.net/fig/1304653">2 B</figureCitation>
). At the base of the spinelets is a circle of 26 small pedicellariae, less than
<quantity id="5F5A2AD0006F02711FC83AF6AAA3736F" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
in length. Spinelets are more crowded and longer at center of the disk, where the anus located but obscured by a cluster of spinelets. Madreporite with a subcentral cleavage,
<quantity id="8A95F23E054864DA0625E003BF805AE8" metricMagnitude="-3" metricUnit="m" metricValue="1.3" unit="mm" value="1.3">1.3 mm</quantity>
in diameter, rather large and protruding, situated near the interradial margin of the disk above the odontophore. The surface of madreporite also covered with spinelets and small pedicellariae.
</paragraph>
<paragraph id="ADD7BAD256FF79CCCFF9026A2705A916">
Basal part of arm with slight genital inflation. A pair of gonads to each arm. Female gonads encapsulated in a long sac, about
<quantity id="8535C98CD6D4DED43C8AD89B96C26D56" metricMagnitude="-2" metricUnit="m" metricValue="1.3" unit="mm" value="13.0">13 mm</quantity>
in length (Fig.
<figureCitation id="7D96119EEDD7D6EC093970CA3B31F45E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Freyastera jiaolongi sp. nov. paratypes. A, D. RSIOAST 0112; B, C, E, F. RSIOAST 0 AS 113. A, B. Abactinal view of arm in genital area; C. Actinal view of arm in genital area; D. Female gonads; E. Actinal view of middle part of arm; F. Abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure4" httpUri="https://binary.pensoft.net/fig/1304655">4 D</figureCitation>
). Abactinal armature of arm in the genital area with irregular polygonal-shaped plates, much larger than those on disk (Fig.
<figureCitation id="9D4F4FA802AC7865F05B49E0E811E8D0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm in genital area; D. Abactinal view of arm in genital area; E. Actinal view of middle part of arm; F. Actinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure2" httpUri="https://binary.pensoft.net/fig/1304653">2 D</figureCitation>
). Each plate carries one short spine, about
<quantity id="EFF83BBF27C2AC8B4DD21F7B0DAB7A6F" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
in length, covered with a tegument loaded with minute pedicellariae (Figs
<figureCitation id="BA3581EB4BB98ECAA6A212203071C0B2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm in genital area; D. Abactinal view of arm in genital area; E. Actinal view of middle part of arm; F. Actinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure2" httpUri="https://binary.pensoft.net/fig/1304653">2 D</figureCitation>
,
<figureCitation id="2B739BADA224E78D3D80ABB56B84ABEE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A D. Abactinal view of arm, from basal part to beyond genital area; E. Abactinal view of middle part of arm; F. Abactinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure3" httpUri="https://binary.pensoft.net/fig/1304654">3 A</figureCitation>
,
<figureCitation id="8AA7C4E20F34AFF8F7DBAF714B4CADDD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Freyastera jiaolongi sp. nov. paratypes. A, D. RSIOAST 0112; B, C, E, F. RSIOAST 0 AS 113. A, B. Abactinal view of arm in genital area; C. Actinal view of arm in genital area; D. Female gonads; E. Actinal view of middle part of arm; F. Abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure4" httpUri="https://binary.pensoft.net/fig/1304655">4 A, B</figureCitation>
). A minority of larger plates carry two or even three spines. The surface of arm plates also scattered with small pedicellariae. The abactinal plates do not seem to go far beyond genital area. The spines become shorter at the end of genital area, sheathed with pedicellariae-covered tegument (Fig.
<figureCitation id="CAA665E818C5EECD498A47EFB4F3E455" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A D. Abactinal view of arm, from basal part to beyond genital area; E. Abactinal view of middle part of arm; F. Abactinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure3" httpUri="https://binary.pensoft.net/fig/1304654">3 B, C</figureCitation>
). Beyond genital area, the pedicellariae cluster in round tegumentary pads (Fig.
<figureCitation id="659542C5ED6C6FE2EF91C1A30EE2A8AE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A D. Abactinal view of arm, from basal part to beyond genital area; E. Abactinal view of middle part of arm; F. Abactinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure3" httpUri="https://binary.pensoft.net/fig/1304654">3 D</figureCitation>
). At the distal part of the arm, pedicellariae pads or transverse bands occur alternately (Figs
<figureCitation id="EC4365ED13019915C962CBEDF6105067" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A D. Abactinal view of arm, from basal part to beyond genital area; E. Abactinal view of middle part of arm; F. Abactinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure3" httpUri="https://binary.pensoft.net/fig/1304654">3 E, F</figureCitation>
,
<figureCitation id="80F10EFF61F1BCDA1C51E2CB6E5979BB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Freyastera jiaolongi sp. nov. paratypes. A, D. RSIOAST 0112; B, C, E, F. RSIOAST 0 AS 113. A, B. Abactinal view of arm in genital area; C. Actinal view of arm in genital area; D. Female gonads; E. Actinal view of middle part of arm; F. Abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure4" httpUri="https://binary.pensoft.net/fig/1304655">4 F</figureCitation>
).
</paragraph>
<paragraph id="A578717ABCBFA853AD0718EBBD65F935">
Adambulacral plate narrow and elongated (Figs
<figureCitation id="B78D55998D2EE75851113D2CB5BDA42E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm in genital area; D. Abactinal view of arm in genital area; E. Actinal view of middle part of arm; F. Actinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure2" httpUri="https://binary.pensoft.net/fig/1304653">2 C, E</figureCitation>
,
<figureCitation id="89FD6F7C20C6037C7BEC28B84D130D46" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Freyastera jiaolongi sp. nov. paratypes. A, D. RSIOAST 0112; B, C, E, F. RSIOAST 0 AS 113. A, B. Abactinal view of arm in genital area; C. Actinal view of arm in genital area; D. Female gonads; E. Actinal view of middle part of arm; F. Abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure4" httpUri="https://binary.pensoft.net/fig/1304655">4 C</figureCitation>
). Each bearing one central subambulacral spine and 13 small aboral furrow spines. Furrow spines at the base of arm measure less than
<quantity id="2AC08D4FD6AA2C6DA44FE4670A2403F4" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
in length, covered with a few scattered small pedicellariae and usually a tuft of pedicellariae at the tip. Furrow spines become fewer and smaller at middle to distal part of arm (Fig.
<figureCitation id="6DD60F184792B83D8F35DD500D515B23" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Freyastera jiaolongi sp. nov. paratypes. A, D. RSIOAST 0112; B, C, E, F. RSIOAST 0 AS 113. A, B. Abactinal view of arm in genital area; C. Actinal view of arm in genital area; D. Female gonads; E. Actinal view of middle part of arm; F. Abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure4" httpUri="https://binary.pensoft.net/fig/1304655">4 E</figureCitation>
), then disappear. Subambulacral spines sheathed, covered with dense small pedicellariae. Subambulacral spines about
<quantity id="582F9CBAE7D76A160FB2A1C8585021B4" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 mm</quantity>
in length at base of arm,
<quantity id="E45A4A3EB7D11B633AB283388F3CAAD8" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
at middle part of arm. The first inferomarginal plate occurs at about the 7
<superScript id="5391069A96C855FB58ED5F7C32785D00">th</superScript>
adambulacral plate, corresponding to alternate or every third adambulacral plate within the genital area and then to every adambulacral plate beyond about 20
<superScript id="FA51A4CBA3E9F070D79859DEE2B1E38B">th</superScript>
adambulacrals. Inferomarginal plates each bear one long sheathed lateral spine, similar in form to subambulacral spines but much longer, about
<quantity id="7C69458C8F465D1526E78C3719FB38E4" metricMagnitude="-2" metricUnit="m" metricValue="2.0" unit="mm" value="20.0">20 mm</quantity>
in length at middle part of arm.
</paragraph>
<paragraph id="DFD76FA6F84A84BB59A2418F493A2F1A">
Oral plate with 34 spines in total (Fig.
<figureCitation id="28E802E16F46CFB47C4DB2BCD081F7E6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm in genital area; D. Abactinal view of arm in genital area; E. Actinal view of middle part of arm; F. Actinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure2" httpUri="https://binary.pensoft.net/fig/1304653">2 A</figureCitation>
). One or two actinostomal spines along the proximal margin of the plate, about
<quantity id="17F748C1B1D51F202E94D2D12D36EFC8" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
in length. One suboral spine, the most robust of all oral spines,
<quantity id="DCA0F91C83D301284D9F5735A672D207" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
in length, situated not at the center of the plate but near the proximal-furrow margin. One aboral spine similar in size to the actinostomal spine, siting at the distal furrow margin of the plate. All mouth spines with pointed or obtuse tips, covered with minute pedicellariae.
</paragraph>
<paragraph id="3F77A7F522E1D0F52ECAD32343696BFC">
A small crustacean skeleton found in the mouth of the
<typeStatus id="20D29EC7EEA35564765DE3CEBEAF9329" type="holotype">holotype</typeStatus>
(Fig.
<figureCitation id="0E16BBB4A57E1AE0F35D2E428C40F2B0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Freyastera jiaolongi sp. nov. holotype RSIOAST 0117. A. Actinal view of disk; B. Abactinal view of disk; C. Actinal view of arm in genital area; D. Abactinal view of arm in genital area; E. Actinal view of middle part of arm; F. Actinal view of distal part of arm." figureDoi="10.3897/zse.101.144918.figure2" httpUri="https://binary.pensoft.net/fig/1304653">2 A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPGAG" type="etymology">
<paragraph id="5437E8F1273E094149DA1FB0253EC68F">
<heading id="06FBC8FF57459B93565C6B4DD11B566F" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="3DB90963649F1396B599EAB04C66DD20">
Named after the Chinese manned submersible
<emphasis id="999EF652D34E227C4C05F661607278CC" italics="true">Jiaolong</emphasis>
that collected all
<specimenCount id="C2DCD2A42EF6B69BF823C67C39C73441" count="3" type="generic">three specimens</specimenCount>
of the new species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXGAG" type="distribution">
<paragraph id="F3FA927767E2B3102BED4670204E8CED">
<heading id="6BFE5D8E44973B6BE66FF427AB197F3A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="08DB309D101C9962AB0BAC9CDFAA8BE2">
Northwest Pacific: Kyushu-Palau Ridge.
<quantity id="285E6632D172D2F47E8205266078D987" metricMagnitude="3" metricUnit="m" metricValue="3.582" metricValueMax="3.641" metricValueMin="3.523" unit="m" value="3582.0" valueMax="3641.0" valueMin="3523.0">35233641 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3GAG" type="remarks">
<paragraph id="2AA69826261FB9B0C880360D6EF824AB">
<heading id="E97D333BC67F0C607AD511C22A5C72BC" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="8B136838DAA039C162FF38FDAA2D480F">
The new species is morphologically most similar to the type species of the genus,
<taxonomicName id="68C80EF7E796DEE1DD7A0807BCED19AD" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="06C36B8D30E267C3FB798AB90575AA0B" italics="true">Freyastera sexradiata</emphasis>
</taxonomicName>
, in the armature of abactinal arm plate, adambulacral plate, and oral plate. It differs from
<taxonomicName id="D20799304B77DED48EB974348C651B36" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="8741519212A587ED6B59AA5188B46AB2" italics="true">F. sexradiata</emphasis>
</taxonomicName>
mainly in the armature of abactinal disk. In
<taxonomicName id="AFDA19ECC17EFA36F2CD833604FAFDC1" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="80D8983644FBFCAE5028E55BF54914A9" italics="true">F. sexradiata</emphasis>
</taxonomicName>
(r =
<quantity id="7ACAEC414683969EF7DD4E45F8FF994C" metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
), the abactinal surface of disk bears “ fairly elongated spines, ” and pedicellariae are absent from the disk (
<bibRefCitation id="F828CD9C34B840A2953D450603E2BF1E" author="Perrier E" journalOrPublisher="Expéditions Scientifiques du Travailleur et du Talisman" pagination="1-431" refId="B54" refString="Perrier E (1894) Stellerides. Expeditions Scientifiques du Travailleur et du Talisman 3: 1-431. [in French]" title="Stellérides." volume="3" year="1894">Perrier, 1894</bibRefCitation>
). In the new species (r =
<quantity id="B5DC921D7E6D5732BEFC12C2AC38881D" metricMagnitude="-3" metricUnit="m" metricValue="5.5" unit="mm" value="5.5">5.5 mm</quantity>
), the abactinal surface of disk is equipped with small hirsute spinelets, about
<quantity id="FAFF8D476CA6C42192E806270E6C57DA" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
in length, which are circled by numerous small pedicellariae. The two species are also geographically distant (Fig.
<figureCitation id="B4C78D21AC073EC4E84EA12FD25E8CDD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Freyastera and six-armed Freyella specimens examined in the present study and type localities of each species." figureDoi="10.3897/zse.101.144918.figure1" httpUri="https://binary.pensoft.net/fig/1304652">1</figureCitation>
).
<taxonomicName id="61F69D1F00573A75C0D4DC446CC239E0" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="4E598B11493A2D689280A20D589B8913" italics="true">F. sexradiata</emphasis>
</taxonomicName>
is known from the North Atlantic, whereas the new species is distributed in the Kyushu-Palau Ridge in the Philippine Sea. Genetically, the new species is close to another undescribed species,
<taxonomicName id="42E487C2B9799C60A0C6031020309057" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-5">
<emphasis id="ECD2150692CC0111EE96C49C747F4DE0" italics="true">Freyastera</emphasis>
sp. 5
</taxonomicName>
(
<abbrev id="ABBRID0EZIAG" xlink_title="c oxidase subunit I">COI</abbrev>
distance 2.64 % 3.52 %). The latter was not described here as only fragmented arms were collected, but in these arm fragments, none of the adambulacral plates bear any furrow spines, which is distinguishable from the new species.
</paragraph>
<caption id="D13D1F57BB2882FB037C78B1DE5E0ED8" ID-DOI="10.3897/zse.101.144918.figure1" ID-arpha="A7F05EEC-915D-5548-9AD3-2D481DA65B18" httpUri="https://binary.pensoft.net/fig/1304652" startId="F1">
<paragraph id="403880F8BC24C503AFF3D3F47230AB8D">
<label id="8F71A0FCC9B71B62C9DA6637BB0440B7">Figure 1.</label>
</paragraph>
<paragraph id="5B2AFD5412215C4B949D892691A72639">
Distribution of
<taxonomicName id="0881E3FD7E9315963C6105EDA291BFBF" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="0C1F9F8934FBE8EBE015169728E3127E" italics="true">Freyastera</emphasis>
</taxonomicName>
and six-armed
<taxonomicName id="FC5A70924F75B391355695AF6209C0EF" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="54D100BDDDA6CD73850D826B7334DFC9" italics="true">Freyella</emphasis>
</taxonomicName>
specimens examined in the present study and type localities of each species.
</paragraph>
</caption>
<caption id="942B69C79BB3BAA7045EEA9D7FF141F2" ID-DOI="10.3897/zse.101.144918.figure2" ID-arpha="3E08CF5D-2650-54D2-9757-A1B2EBE78EF3" httpUri="https://binary.pensoft.net/fig/1304653" startId="F2">
<paragraph id="2A7A289B7C6F8D1090F022C0A54448F5">
<label id="E3836969C4462FEFF0448DACE424AFE5">Figure 2.</label>
</paragraph>
<paragraph id="B62B3EFEC7297874AF5384D25DC80436">
<taxonomicName id="B7F54027811D273553BAB9491CD79E3D" authorityName="Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang" authorityYear="2025" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="jiaolongi" status="sp. nov.">
<emphasis id="ED7D030026D015EAC1F616FAB4B783AB" italics="true">Freyastera jiaolongi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA1DA2657468E395388125BF686635A5" rank="species">sp. nov.</taxonomicNameLabel>
holotype
<specimenCode id="9C4F53AAF271FB91B6B1A50BCB5A412C">RSIOAST 0117</specimenCode>
.
<emphasis id="5686888819E36EE8ABDE9906D7355AE5" bold="true">A.</emphasis>
Actinal view of disk;
<emphasis id="C583AFACF85B6D303FB8F7067CDEB3F7" bold="true">B.</emphasis>
Abactinal view of disk;
<emphasis id="94BE021130B40C9E5E335AE446FEC086" bold="true">C.</emphasis>
Actinal view of arm in genital area;
<emphasis id="E17CC6164AA569E317849A7763CB53E4" bold="true">D.</emphasis>
Abactinal view of arm in genital area;
<emphasis id="D0866352508018F9CBFAFF05E643A56C" bold="true">E.</emphasis>
Actinal view of middle part of arm;
<emphasis id="32E2CD227D965EF3DF12BF9E1045DA13" bold="true" italics="true">F.</emphasis>
Actinal view of distal part of arm.
</paragraph>
</caption>
<caption id="EEA88F2EE457EA8AABD7791EE34331E5" ID-DOI="10.3897/zse.101.144918.figure3" ID-arpha="05931D8F-D25F-5E89-80A8-7B2F6B1BDD87" httpUri="https://binary.pensoft.net/fig/1304654" startId="F3">
<paragraph id="C2013BC67BFA8ED8EC3A12D711C81BEE">
<label id="DA2C44827C9BAC493C8270A8DBE2AD4A">Figure 3.</label>
</paragraph>
<paragraph id="32736CCBF4348777CCD6D067FA6D440F">
<taxonomicName id="B6EF85AFFBB84C1FF573C3EF101A06BF" authorityName="Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang" authorityYear="2025" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="jiaolongi" status="sp. nov.">
<emphasis id="C695DA121AA236F5AA908A461EEC3529" italics="true">Freyastera jiaolongi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51517D90BD74C5BA10AB48770D0FB209" rank="species">sp. nov.</taxonomicNameLabel>
holotype
<specimenCode id="6119ED66C52D09C759BDF8E8F9F45F45">RSIOAST 0117</specimenCode>
.
<emphasis id="62A0A81E98ADD4BEEE57B8CD71CCC05D" bold="true">A D.</emphasis>
Abactinal view of arm, from basal part to beyond genital area;
<emphasis id="CD755AC7384A97CB167A53D3426ADA3F" bold="true">E.</emphasis>
Abactinal view of middle part of arm;
<emphasis id="6CC36ADE5D4D3DDBD2731B2AE8490042" bold="true" italics="true">F.</emphasis>
Abactinal view of distal part of arm.
</paragraph>
</caption>
<caption id="E886920ACB9FFF5F5C4EF9076B6EB4A8" ID-DOI="10.3897/zse.101.144918.figure4" ID-arpha="B1DF333B-A5C9-58D2-A9A2-DBB71D2EFC71" httpUri="https://binary.pensoft.net/fig/1304655" startId="F4">
<paragraph id="EBAFB1CA9ED9ADFD82448E1AB154ABC5">
<label id="D1FF1BA2385728E9ABB9C04D7D304916">Figure 4.</label>
</paragraph>
<paragraph id="6597B0A3D87506483474BFB3C2CC1CFB">
<taxonomicName id="17E0A8952E9718109220436FA99E19DB" authorityName="Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang" authorityYear="2025" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="jiaolongi" status="sp. nov.">
<emphasis id="A0BF9AC4DF7EDCE5B79DF164AC6AD0D9" italics="true">Freyastera jiaolongi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="111FA37453F99BB05B32217AF3BB7173" rank="species">sp. nov.</taxonomicNameLabel>
paratypes.
<emphasis id="1145BAA3D5EAA49F2A06E81570079804" bold="true">A, D.</emphasis>
<specimenCode id="2E04902C5B2D1C56788016CC01BB0080">RSIOAST 0112</specimenCode>
;
<emphasis id="79E906D34596959AE85E4421F9415122" bold="true">
B, C, E,
<emphasis id="9BB3AE9E2CDC21A3EB3B5FE08E17F061" italics="true">F.</emphasis>
</emphasis>
RSIOAST 0 AS 113.
<emphasis id="4A998182DA060BD42FBD11D6FE7F4150" bold="true">A, B.</emphasis>
Abactinal view of arm in genital area;
<emphasis id="A08F92035CCF522007B02BABE8B4A487" bold="true">C.</emphasis>
Actinal view of arm in genital area;
<emphasis id="AEB874A805916E2F1C8DF4056609007E" bold="true">D.</emphasis>
Female gonads;
<emphasis id="486291EFA275884900418F66449F4335" bold="true">E.</emphasis>
Actinal view of middle part of arm;
<emphasis id="AAFABE0E3BF3049C301B157CAA98FE16" bold="true" italics="true">F.</emphasis>
Abactinal view of middle part of arm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -1,73 +1,73 @@
<document id="0EFF3E5A4A1D29F16DFA8134479A529C" ID-DOI="10.3897/zse.101.144918" ID-ZooBank="4611CD60-658B-42C4-B23C-2DA9185A85E3" ID-publisher-id="144918" URI-arpha="9ADAAF17-610F-5BDF-A5A2-3B2BB4BFE7D5" URI-zoobank="http://zoobank.org/4611CD60-658B-42C4-B23C-2DA9185A85E3" article-type="research-article" checkinTime="1744298128406" checkinUser="pensoft" docAuthor="Zhang, Ruiyan, Zhou, Yadong, Mao, Jingwen, Wang, Chunsheng &amp; Zhang, Dongsheng" docDate="2025" docId="D1AA056020DD58A392E2DBA685BA52BF" docLanguage="en" docName="Zoosystematics and Evolution 101(2): 735-760" docOrigin="Zoosystematics and Evolution 101 (2)" docSource="https://zse.pensoft.net/article/144918/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Freyastera loricata Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang 2025, comb. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="760" masterDocId="9ADAAF17610F5BDFA5A23B2BB4BFE7D5" masterDocTitle="Hidden diversity of Freyastera (Asteroidea, Brisingida, Freyellidae) at great depth: Description of new species and remarks on species boundaries" masterLastPageNumber="760" masterPageNumber="735" pageNumber="735" updateTime="1744298223404" updateUser="pensoft">
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<mods:title id="802E0E9D29D64444A4F52BDADD35CF66">Hidden diversity of Freyastera (Asteroidea, Brisingida, Freyellidae) at great depth: Description of new species and remarks on species boundaries</mods:title>
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(in key).
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@ -123,7 +121,9 @@
<heading id="709113D0C58E15C2284D5F79753269ED" reason="title">Material examined.</heading>
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<paragraph id="7987E048FF03DB8FF97EAA853539578B">
ECH 00161 (
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<specimenCode id="82DC567F654F0841F435342784935973">ECH 00161</specimenCode>
(
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<typeStatus id="51B1737106BF6D11CF0C82CA834BFE5B" type="paratype">paratype</typeStatus>
</emphasis>
@ -131,7 +131,9 @@ ECH 00161 (
<figureCitation id="4BDB51B685DD44E928641181DB10703C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A D. Freyastera loricata (Korovchinsky &amp; Galkin, 1984) type ECH 00161 (photos courtesy of Dr. Anna Dilman); E I. Freyastera tuberculata (Sladen, 1889) type NHMUK 1890.5. 7.1076. A, E. abactinal view of disk; B, F. Actinal view of disk; C. G. Abactinal view of arm genital area; D, H. Actinal view of proximal part of arm; I. abactinal view of middle part of arm." figureDoi="10.3897/zse.101.144918.figure9" httpUri="https://binary.pensoft.net/fig/1304660">9 A D</figureCitation>
). (Suppl. material
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).
)
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.
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@ -176,9 +178,7 @@ adambulacral plate, appear at alternate adambulacral plates, and then at every a
<heading id="7095BE843684D90F7AED52725F9E0FC4" reason="title">Distribution.</heading>
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Only known at
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locality. East of the Kuril-Kamchatka Trench.
Only known at type locality. East of the Kuril-Kamchatka Trench.
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<treatment id="DA1413D4EE0C5A17932F6B63B444D82F" ID-arpha="DA1413D4-EE0C-5A17-932F-6B63B444D82F" LSID="urn:lsid:plazi:treatment:DA1413D4EE0C5A17932F6B63B444D82F" httpUri="http://treatment.plazi.org/id/DA1413D4EE0C5A17932F6B63B444D82F" scope_class="Asteroidea" scope_family="Freyellidae" scope_order="Brisingida">
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<taxonomicName id="C196414F31B9F06D7354D99045A82BD2" ID-CoL="6JNSY" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" isUncertain="true" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="tuberculata">
<emphasis id="D0291260646C2272232CEC22078C1776" italics="true">Freyastera</emphasis>
cf.
<emphasis id="7BEA05F9B2C03C2BEA5904C2C5AFE6B9" italics="true">tuberculata</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
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<emphasis id="C7C7F42C0E1741AA7C6606EDF9C7818D" italics="true">Freyastera</emphasis>
cf.
<emphasis id="BCB7AA718F3F4B63BCB36582F20D3023" italics="true">tuberculata</emphasis>
</taxonomicName>
<comment id="3219106CCDF3DA01FFA4BFFE127A8DCE">
:
<treatmentCitation id="9D066375A7EF6547771E3B2820A14708" author="Bribiesca-Contreras G &amp; Dahlgren TG &amp; Amon DJ &amp; Cairns S &amp; Drennan R &amp; Durden JM &amp; Eléaume MP &amp; Hosie AM &amp; Kremenetskaia A &amp; McQuaid K &amp; D OHara T" httpUri="http://treatment.plazi.org/id/A7C19C558CF05ABDB05598C74F785B48" page="42" year="2022">
<bibRefCitation id="F859E14B1C0ACBEF342EAC8343EE78C4" DOI="10.3897/zookeys.1113.82172" author="Bribiesca-Contreras G &amp; Dahlgren TG &amp; Amon DJ &amp; Cairns S &amp; Drennan R &amp; Durden JM &amp; Eléaume MP &amp; Hosie AM &amp; Kremenetskaia A &amp; McQuaid K &amp; D OHara T" etAl="et al." firstAuthor="Bribiesca-Contreras" page="42" refId="B4" refString="Bribiesca-Contreras G, Dahlgren TG, Amon DJ, Cairns S, Drennan R, Durden JM, Eléaume MP, Hosie AM, Kremenetskaia A, McQuaid K, D OHara T (2022) Benthic megafauna of the western Clarion-Clipperton Zone, Pacific Ocean. ZooKeys 1113: 1. https://doi.org/10.3897/zookeys.1113.82172" year="2022">Bribiesca-Contreras et al. 2022: 42</bibRefCitation>
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.
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<treatmentCitationGroup id="FF6606F1C0CB1714A1D390110D60EB14">
<taxonomicName id="DC141ED0995FFDE13AF0EC4CC9747754" ID-CoL="6JNTB" baseAuthorityName="Sladen" baseAuthorityYear="1889" class="Asteroidea" family="Freyellidae" genus="Freyastera" isUncertain="true" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="benthophila">
<emphasis id="D5BF8D453637B11A38C4F81A2659789F" italics="true">Freyastera</emphasis>
cf.
<emphasis id="E02600E875D30F154FDE84C9FC3069C7" italics="true">benthophila</emphasis>
</taxonomicName>
<comment id="6A0E94BE308E03BA9B858803E346F99F">
:
<treatmentCitation id="DE2CF0B10FCE3914BB49F2FAE3BA7D55" author="Glover AG &amp; Wiklund H &amp; Rabone M &amp; Amon DJ &amp; Smith CR &amp; OHara T &amp; Mah CL &amp; Dahlgren TG" page="17" year="2016">
<bibRefCitation id="68F871312B5F9461F71FCF4BE619DFBC" DOI="10.3897/BDJ.4.e7251" author="Glover AG &amp; Wiklund H &amp; Rabone M &amp; Amon DJ &amp; Smith CR &amp; OHara T &amp; Mah CL &amp; Dahlgren TG" etAl="et al." firstAuthor="Glover" page="17" refId="B22" refString="Glover AG, Wiklund H, Rabone M, Amon DJ, Smith CR, OHara T, Mah CL, Dahlgren TG (2016) Abyssal fauna of the UK- 1 polymetallic nodule exploration claim, Clarion-Clipperton Zone, central Pacific Ocean: Echinodermata. Biodiversity data journal 4 (4): e 7251. https://doi.org/10.3897/BDJ.4.e7251" year="2016">Glover et al. 2016: 17</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="4389050C0C64FB44ED9C5B43DA5D03E7" author="Amon DJ &amp; Ziegler AF &amp; Kremenetskaia A &amp; Mah CL &amp; Mooi R &amp; OHara T &amp; Pawson DL &amp; Roux M &amp; Smith CR" page="9" year="2017">
<bibRefCitation id="F7E125A23C753A75AEE4326830F9FD6E" DOI="10.3897/BDJ.5.e11794" author="Amon DJ &amp; Ziegler AF &amp; Kremenetskaia A &amp; Mah CL &amp; Mooi R &amp; OHara T &amp; Pawson DL &amp; Roux M &amp; Smith CR" etAl="et al." firstAuthor="Amon" page="9" refId="B2" refString="Amon DJ, Ziegler AF, Kremenetskaia A, Mah CL, Mooi R, OHara T, Pawson DL, Roux M, Smith CR (2017) Megafauna of the UKSRL exploration contract area and eastern Clarion-Clipperton Zone in the Pacific Ocean: Echinodermata. Biodiversity Data Journal 5: e 11794. https://doi.org/10.3897/BDJ.5.e11794" year="2017">Amon et al. 2017: 9</bibRefCitation>
</treatmentCitation>
; GenBank (
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).
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<paragraph id="5FB3D5594695804F6AB20FA350412006">
<heading id="0061346499263E3A24183A906A7BE691" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="A604D26110C9A911CAEFD256C6B72CDE">
<materialsCitation id="C783F151ACCE8D46BBE774BCBB8809A9" collectionCode="NHMUK" specimenCode="NHMUK 2022.79" specimenCount="1">
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2022.79
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(CCZ_087)
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;
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<named-content id="6FDFABD0EE50A6FE6CF5FBC3E4A1B873" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum">NHMUK</named-content>
2022.80
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(CCZ_157)
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;
<materialsCitation id="8BBA2DB5B41A712360754FA88B375111" collectionCode="NHMUK" specimenCode="NHMUK 9120" specimenCount="1">
<specimenCode id="6147C2159178FED4E14446D2BB4C9ECD">
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9120
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; NHM_413
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;
<materialsCitation id="13003D074184896E0FC93C34AB12513F" collectionCode="SO" specimenCode="SO 268-1_64_074" specimenCount="1">
<specimenCode id="CAEBD91361CE143A518C05A6DDB97E92">SO 268-1_64_074</specimenCode>
. (Suppl. material
<xref id="5022356D6C812468278201D474BEC601" ref-type="supplementary-material" rid="S1">1</xref>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EL4AI" type="distribution">
<paragraph id="B7AD0D4AF840566CE16F4ED30A42A9A4">
<heading id="21494A006D6E121DCCEEBF88AB2BE97C" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="25CB5E465D75056D4489502C4857C293">
Eastern Pacific: Clarion-Clipperton Zone.
<quantity id="6EC7DB4D9F89ABF023CB077FF6E9DBEA" metricMagnitude="3" metricUnit="m" metricValue="4.5055" metricValueMax="5.0" metricValueMin="4.011" unit="m" value="4505.5" valueMax="5000.0" valueMin="4011.0">40115000 m</quantity>
.
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</treatment>
</document>

View file

@ -1,73 +1,73 @@
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<paragraph id="C924815922BA03558D1FEE854A944535">
<taxonomicName id="316210BD50627F6C43AA1C23714F4FBC" authority="(Koehler, 1907)" authorityName="Zhang &amp; Zhou &amp; Mao &amp; Wang &amp; Zhang" authorityYear="2025" baseAuthorityName="Koehler" baseAuthorityYear="1907" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="giardi" status="comb. nov.">
@ -87,7 +87,6 @@
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<emphasis id="3A98514B6B135B3E419837C9FAE3A125" italics="true">Freyella giardi</emphasis>
</taxonomicName>
@ -134,7 +133,6 @@
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(in key).
</comment>
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</subSubSection>
@ -143,36 +141,48 @@
<heading id="5E938840B59AEB542D4B946708778D27" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="08EA71832CEAA850DAF7612964C1EF1B">
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<collectionCode id="C1324C77E296B91F7CC811D9E7791815" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1912.11. 11.30 (
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1912.11.11.30
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(
<emphasis id="865A4E29F9B04AEDB666BAA75C71D8FD" bold="true" italics="true">
<typeStatus id="DAA4AFC12A39640EE68E4A3100196255" type="syntype">syntype</typeStatus>
</emphasis>
) (Fig.
<figureCitation id="24903877C6F89B726086C77507B6FCF7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Freyastera giardi (Koehler, 1907) type specimens. A E. NHMUK 1912.11. 11.30; F. MNHN - IE- 2014-463. A. Abactinal view of disk; B. Actinal view of disk; C. Abactinal view of arm genital area; D. Actinal view of disk and proximal part of arm; E. Abactinal view of middle part of arm; F. Actinal view of proximal part of arm. Yellow arrows show subambulacral spines; red arrows show aboral furrow spines." figureDoi="10.3897/zse.101.144918.figure8" httpUri="https://binary.pensoft.net/fig/1304659">8 A E</figureCitation>
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- IE- 2014-463 (
)
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;
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<named-content id="854102804A60F10A145616282BA7393E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">MNHN</named-content>
-IE-2014-463
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(
<emphasis id="0C35E779418682CE5ECC2352A929D8F3" bold="true" italics="true">
<typeStatus id="BDBE76CDEFC3A35DA45C6BBF38096B14" type="syntype">syntype</typeStatus>
</emphasis>
) (Fig.
<figureCitation id="0C63D9678F70B6F7938F3C65C1C11D87" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Freyastera giardi (Koehler, 1907) type specimens. A E. NHMUK 1912.11. 11.30; F. MNHN - IE- 2014-463. A. Abactinal view of disk; B. Actinal view of disk; C. Abactinal view of arm genital area; D. Actinal view of disk and proximal part of arm; E. Abactinal view of middle part of arm; F. Actinal view of proximal part of arm. Yellow arrows show subambulacral spines; red arrows show aboral furrow spines." figureDoi="10.3897/zse.101.144918.figure8" httpUri="https://binary.pensoft.net/fig/1304659">8 F</figureCitation>
);
<named-content id="C54B5EFB401FD9F51E42A56CF46172DC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/musee-oceanographique-monaco" xlink_title="Musée Océanographique de Monaco">
<collectionCode id="7613A0F8F8780347FE0210BF735F4402" country="Monaco" httpUri="http://grbio.org/cool/cx5v-6vpi" name="Musee Oceanographique Monaco" type="">MOM</collectionCode>
</named-content>
- INV- 0021953 (
)
</materialsCitation>
;
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<specimenCode id="69996E41424EB98AD119B4DDA8B1CF9E">
<named-content id="C54B5EFB401FD9F51E42A56CF46172DC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/musee-oceanographique-monaco" xlink_title="Musée Océanographique de Monaco">MOM</named-content>
-INV-0021953
</specimenCode>
(
<emphasis id="125322BF6E63002D6A7C29FC16862875" bold="true" italics="true">
<typeStatus id="8A62A1541AA1CB66BEB5E226C8CFC20E" type="syntype">syntype</typeStatus>
</emphasis>
). (Suppl. material
<xref id="6FC0C248ED5721E0B3BEA669D0476525" ref-type="supplementary-material" rid="S1">1</xref>
).
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFLAI" type="remarks">
@ -219,9 +229,7 @@ as discussed above, we reassign this species as
<heading id="F4EBE09FD95E1429532AA11E4702188A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="EBFC860CE08CD99D0A35CE9ADE6C1F60">
Only known at
<typeStatus id="CEDA2F61046FBA86E470736BEE5F8200">type</typeStatus>
locality. Weddle Sea.
Only known at type locality. Weddle Sea.
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.
</paragraph>

View file

@ -0,0 +1,274 @@
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<mods:affiliation id="E013ACF5F677EE3823BC2780FAEDF12A">State Key Laboratory of Submarine Geoscience, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, 310012, China &amp; Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, 310012, China &amp; Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519082, China</mods:affiliation>
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<mods:title id="63C5D3D4AB7B1BC9AF23FEBD53281869">Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part id="5574F2503AE23D16D76F3D240C7728A0">
<mods:date id="18EC8F6990656275280F623A4A8566A7">2025</mods:date>
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<mods:number id="55B71DE8ED0E9D4B9E36D3C84CB80583">2025-04-10</mods:number>
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<mods:number id="F6E76E28C284A97C06148132C2E54707">101</mods:number>
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<mods:number id="6125443C892C38EFCA9BB5D84B882A12">2</mods:number>
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<mods:start id="46AC99C987D2CC2E5A12DE042B91E667">735</mods:start>
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<mods:identifier id="4A5ECC9B0B23D7EB9672E9A241511E8F" type="DOI">10.3897/zse.101.144918</mods:identifier>
<mods:identifier id="F3CB83AE8F8BDA75435560353FDCBBE8" type="ZooBank">4611CD60-658B-42C4-B23C-2DA9185A85E3</mods:identifier>
</mods:mods>
<treatment id="EEA6264E2D09540E9D3219BCBEB5409C" ID-arpha="EEA6264E-2D09-540E-9D32-19BCBEB5409C" LSID="urn:lsid:plazi:treatment:EEA6264E2D09540E9D3219BCBEB5409C" httpUri="http://treatment.plazi.org/id/EEA6264E2D09540E9D3219BCBEB5409C" scope_class="Asteroidea" scope_family="Freyellidae" scope_order="Brisingida">
<subSubSection id="9A667DF8EBDB2B0D312755537508059D" type="nomenclature">
<paragraph id="5FD5C05359F9633A0B38F9ADD646CFB1">
<taxonomicName id="28D23A84DB1330FF8D27CD8F7192D978" ID-CoL="6JNSZ" authority="(Perrier, 1885)" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="BD277B187B63EA8833415D50DC25EDB4" italics="true">Freyastera sexradiata</emphasis>
(
<bibRefCitation id="E224F94C72B30A3BAB061346BD9BB67C" author="Perrier E" issue="8" journalOrPublisher="Annales des sciences naturelles, Zoologie" pagination="1-72" refId="B53" refString="Perrier E (1885) Premiere note Preliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman. Annales des sciences naturelles, Zoologie 22 (8): 1-72. [in French]" title="Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman." volume="22" year="1885">Perrier, 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A44891221037594A10DDAA8A0CC96AA6" type="reference_group">
<paragraph id="0795C04DB1E914E58A64034A2A27F529">
<treatmentCitationGroup id="E0D9A338526D10979C9C78F46D26EDE9">
<taxonomicName id="C8AE4CAD255F5E4EAF8E943445C0934A" ID-CoL="6JN7F" authorityName="Perrier" authorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyella" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="8530740AAD60AC48985172EEAFD1B5B4" italics="true">Freyella sexradiata</emphasis>
</taxonomicName>
<comment id="791565E0C48017B0EA9FCEBCC2822BA6">
:
<treatmentCitation id="E7F5F1EB061EBCAE9A14193AA8C7808F" author="Perrier E" page="6" year="1885">
<bibRefCitation id="784FDD70CF3A95EF4BA6F573E8E0BD53" author="Perrier E" firstAuthor="Perrier" issue="8" journalOrPublisher="Annales des sciences naturelles, Zoologie" page="6" pagination="1-72" refId="B53" refString="Perrier E (1885) Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman. Annales des sciences naturelles, Zoologie 22 (8): 172. [in French]" title="Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman." volume="22" year="1885">Perrier 1885: 6</bibRefCitation>
</treatmentCitation>
; 1894: 89;
<treatmentCitation id="1B37FD6C7B7416E9B873F02324E68432" author="Koehler R" page="129" year="1909">
<bibRefCitation id="C0D090BD89BF2F1F099C70372787E291" author="Koehler R" firstAuthor="Koehler" journalOrPublisher="Résultats des Campagnes Scientifiques Accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco" page="129" pagination="25-28" refId="B32" refString="Koehler R (1909) Echinodermes provenant des campagnes du yacht Princesse-Alise (Astéries, Ophiures, Echinides et Crinoides). Résultats des Campagnes Scientifiques Accomplies sur son Yacht par Albert Ier Prince Souverain de Monaco 34: 2528. [in French]" title="Echinodermes provenant des campagnes du yacht Princesse-Alise (Astéries, Ophiures, Echinides et Crinoides)." volume="34" year="1909">Koehler 1909: 129</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6E142597026565FD3BB64B43CF9F4FE8" author="Grieg JA" page="30" year="1921">
<bibRefCitation id="09A2CDA3F1014A56B387A153F17E545F" author="Grieg JA" firstAuthor="Grieg" page="30" refId="B23" refString="Grieg JA (1921) Echinodermata. Report on the Scientific Results of the “ Michael Sars ” North Atlantic Deep-sea Expedition 1910. Trustees of the Bergen Museum, 144." year="1921">Grieg 1921: 30</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="4BB8D6A95D68D9ABC914989BE94D4ACC" author="Mortensen T" page="129" year="1927">
<bibRefCitation id="611F521322ABD1B18CB52E0355CC310C" DOI="10.5962/bhl.title.6841" author="Mortensen T" firstAuthor="Mortensen" page="129" refId="B50" refString="Mortensen T (1927) Handbook of the Echinoderms of the British Isles. Oxford University Press, 471 pp. https://doi.org/10.5962/bhl.title.6841" year="1927">Mortensen 1927: 129</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="DDC1809CD705B38B984CDC4F818297AF" author="Fisher WK" page="24" year="1928">
<bibRefCitation id="D868C0C49BFBDBE5D83D9EC7E91FEB08" DOI="10.5479/si.03629236.145.1" author="Fisher WK" firstAuthor="Fisher" journalOrPublisher="Bulletin of the United States National Museum" page="24" pagination="1-347" refId="B18" refString="Fisher WK (1928) Asteroidea of the North Pacific and adjacent waters. Part 2. Forcipulata (part). Bulletin of the United States National Museum 76: 1347. https://doi.org/10.5479/si.03629236.145.1" title="Asteroidea of the North Pacific and adjacent waters. Part 2. Forcipulata (part)." volume="76" year="1928">Fisher 1928: 24</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AB60A3323021D81A80EA887FDDD8F13E" author="Madsen FJ" page="84" year="1951">
<bibRefCitation id="CF2BC9BD07F10AC2FD83129A3E51E7F4" author="Madsen FJ" firstAuthor="Madsen" issue="6" journalOrPublisher="Reports of the Swedish Deep-sea Expedition" page="84" pagination="73-92" refId="B38" refString="Madsen FJ (1951) Asteroidea. Reports of the Swedish Deep-sea Expedition 2 (6): 7392." title="Asteroidea." volume="2" year="1951">Madsen 1951: 84</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F3645C8FA0BFF1D21C00077CB97DFE4D" author="Cherbonnier G &amp; Sibuet M" page="1356" year="1972">
<bibRefCitation id="197AF2B5512FED0ED969709B1256F9E0" DOI="10.5962/p.272616" author="Cherbonnier G &amp; Sibuet M" firstAuthor="Cherbonnier" journalOrPublisher="Bulletin du Museum national dhistoire naturelle" page="1356" pagination="1333-1394" refId="B6" refString="Cherbonnier G, Sibuet M (1972) Résultats scientifiques de la campagne Noratlante: Astérides et Ophiurides. Bulletin du Museum national dhistoire naturelle 102: 13331394. [in French] https://doi.org/10.5962/p.272616" title="Résultats scientifiques de la campagne Noratlante: Astérides et Ophiurides." volume="102" year="1972">Cherbonnier and Sibuet 1972: 1356</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="2CB8B3869606A43E70EC8C5E254F3020" author="Korovchinsky NM &amp; Galkin SV" page="1213" year="1984">
<bibRefCitation id="822AEF4F79C6ED45D3DD12F0A8B3BCCE" author="Korovchinsky NM &amp; Galkin SV" firstAuthor="Korovchinsky" issue="8" journalOrPublisher="Zoologicheskii Zhurnal" page="1213" pagination="1205-1215" refId="B34" refString="Korovchinsky NM, Galkin SV (1984) New data on the fauna of starfishes of the genus Freyella (Brisingidae). Zoologicheskii Zhurnal 63 (8): 12051215. [in Russian with English abstract]" title="New data on the fauna of starfishes of the genus Freyella (Brisingidae)." volume="63" year="1984">Korovchinsky and Galkin 1984: 1213</bibRefCitation>
</treatmentCitation>
(in key);
<treatmentCitation id="A577A3BD22842908C98D014A900C24DC" author="Galkin SV &amp; Korovchinsky NM" page="167" year="1984">
<bibRefCitation id="E724CA67662B34ABB3B05CFD9E987A14" author="Galkin SV &amp; Korovchinsky NM" firstAuthor="Galkin" journalOrPublisher="Trudy Instituta Okeanologii Akademii Nauk SSSR" page="167" pagination="164-178" refId="B20" refString="Galkin SV, Korovchinsky NM (1984) Vertical and geographical distribution of the starfishes of the genus Freyella (Brisingidae) with some remarks on their ecology and evolution. Trudy Instituta Okeanologii Akademii Nauk SSSR 119: 164178. [in Russian with English abstract]" title="Vertical and geographical distribution of the starfishes of the genus Freyella (Brisingidae) with some remarks on their ecology and evolution." volume="119" year="1984">Galkin and Korovchinsky 1984: 167</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="1A445156731CA4592D5A675C9BC3CAA7">
<treatmentCitationGroup id="FE71EC24140D2F1E5330FAA0906506FB">
<taxonomicName id="5523FBD2B5F40EA9901892E5F730F74D" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyellidea" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="8AE182A9B4D437BCA1D0FAB15DCEB3DD" italics="true">Freyellidea sexradiata</emphasis>
</taxonomicName>
<comment id="30B8BA14439EAD4ED58EC70D34EB8A84">
:
<treatmentCitation id="3789CAD6AD4736505D6CFF7B504D7914" author="Fisher WK" page="429" year="1917">
<bibRefCitation id="5362A9A7E8010AB34634B7AADA111E96" DOI="10.1080/00222931709487030" author="Fisher WK" firstAuthor="Fisher" issue="120" journalOrPublisher="Annals and Magazine of Natural History" page="429" pagination="418-431" refId="B17" refString="Fisher WK (1917) LI. — New genera and species of Brisingidae. Annals and Magazine of Natural History 20 (120): 418431. https://doi.org/10.1080/00222931709487030" title="LI. - New genera and species of Brisingidae." volume="20" year="1917">Fisher 1917: 429</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6734C83630FA2D9F5BC19B89A01537BD">
<treatmentCitationGroup id="83F9746AE8B00BA4A746AEBE649EB9F0">
<taxonomicName id="513E7D0512C05456B62EB55B228387BD" ID-CoL="6JNSZ" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="87F7CB57F22F636F8E5849CD227DB30C" italics="true">Freyastera sexradiata</emphasis>
</taxonomicName>
<comment id="21715EA6DC9F83AA4C49A460BECA51B5">
:
<treatmentCitation id="6C5BD50E121AFF7E43B808771EA0AD17" author="Downey ME" page="40" year="1986">
<bibRefCitation id="C5D067B7223E2F753A7B18AFEA913795" DOI="10.5479/si.00810282.435" author="Downey ME" firstAuthor="Downey" journalOrPublisher="Smithsonian Contributions to Zoology" page="40" pagination="1-57" refId="B15" refString="Downey ME (1986) Revision of the Atlantic Brisingida (Echinodermata: Asteroidea), with description of a new genus and family. Smithsonian Contributions to Zoology 435: 157. https://doi.org/10.5479/si.00810282.435" title="Revision of the Atlantic Brisingida (Echinodermata: Asteroidea), with description of a new genus and family." volume="435" year="1986">Downey 1986: 40</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="30C87D9D7D9EF58F644222C58864DD01" author="Clark AM &amp; Downey ME" page="481" year="1992">
<bibRefCitation id="BFD193532B8ED287564E8A7053B181C2" author="Clark AM &amp; Downey ME" firstAuthor="Clark" page="481" refId="B10" refString="Clark AM, Downey ME (1992) Starfishes of the Atlantic. London: Chapman and Hall, 794 pp." year="1992">Clark and Downey 1992: 481</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="D47BE8088E87C8707EBDA96CD61BB5ED" author="Mah CL" page="87" year="1998">
<bibRefCitation id="D50E62C553F4A5CDE41563BE5B28320D" author="Mah CL" firstAuthor="Mah" page="87" refId="B40" refString="Mah CL (1998) A phylogeny, taxonomic revision, and natural history of the order Brisingida (Asteroidea). Masters Thesis, San Francisco State University, San Francisco, US." year="1998">Mah 1998: 87</bibRefCitation>
</treatmentCitation>
; Mah in
<treatmentCitation id="A7FB8DFDECFDDFCCABF14DA0FD4736FB" author="Clark AM &amp; Mah C" page="318" year="2001">
<bibRefCitation id="1DD6C372AC57932D0F12921E3F169DA8" author="Clark AM &amp; Mah C" firstAuthor="Clark" journalOrPublisher="Echinoderm studies" page="318" pagination="229-347" refId="B11" refString="Clark AM, Mah C (2001) An index of names of recent Asteroidea — Part 4: Forcipulatida and Brisingida. Echinoderm studies 6: 229347." title="An index of names of recent Asteroidea - Part 4: Forcipulatida and Brisingida." volume="6" year="2001">Clark and Mah 2001: 318</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="16E06A00CF8A22EB00611E5990359F99" author="Dilman AB" page="38" year="2014">
<bibRefCitation id="26758813DFB27511A479D6D69D53CEC0" DOI="10.15298/invertzool.11.1.05" author="Dilman AB" firstAuthor="Dilman" issue="1" journalOrPublisher="Invertebrate Zoology" page="38" pagination="25-42" refId="B13" refString="Dilman AB (2014) Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Asteroidea. Invertebrate Zoology 11 (1): 2542. https://doi.org/10.15298/invertzool.11.1.05" title="Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Asteroidea." volume="11" year="2014">Dilman 2014: 38</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="1BF06CDCBF7BD3FC6C1AB1AA4A204061" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" page="6" year="2019">
<bibRefCitation id="C9B2B2B521207772E4DC26643807B53C" DOI="10.1016/j.dsr.2019.103163" author="Zhang R &amp; Wang C &amp; Zhou Y &amp; Zhang H" etAl="et al." firstAuthor="Zhang" page="6" refId="B68" refString="Zhang R, Wang C, Zhou Y, Zhang H (2019) Morphology and molecular phylogeny of two new species in genus Freyastera (Asteroidea: Brisingida: Freyellidae), with a revised key to close species and ecological remarks. Deep Sea Research Part I: Oceanographic Research Papers 154: 103163. https://doi.org/10.1016/j.dsr.2019.103163" year="2019">Zhang et al. 2019: 6</bibRefCitation>
</treatmentCitation>
(in key).
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBSAG" type="material">
<paragraph id="9727E8F8695E0BF6DD0446D56B307497">
<heading id="7F1D4978CAF04B137CD6256EAB1915DD" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="B04F1D298A05DFFE85ED79768EA151B9">
<materialsCitation id="35F22864B92EEE082D8F6DDFE5C1A510" collectionCode="MNHN" specimenCode="MNHN-IE-2014-165" specimenCount="1" typeStatus="holotype">
<specimenCode id="BE72C6456F120EF88E629910DAF0323B">
<named-content id="ADB8F8C5BC2109027A70F5A1BBB584E2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">MNHN</named-content>
-IE-2014-165
</specimenCode>
(
<emphasis id="8E593437417B171B4D53C15BFD8DA7FE" bold="true" italics="true">
<typeStatus id="EC7938AD02218293F991B0881F889BEA" type="holotype">holotype</typeStatus>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="14E1EAECABCFBDF181321A4911FC3995" collectionCode="MOM" specimenCode="MOM-81 0428" specimenCount="1">
<specimenCode id="1E768D1F46B737252BC42A06904CCBE9">
<named-content id="60692B8407AF8F62F4C8B661D588777F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/musee-oceanographique-monaco" xlink_title="Musée Océanographique de Monaco">MOM</named-content>
-81 0428
</specimenCode>
. (Suppl. material
<xref id="16A06155AA0167185CBAC7B02EA8AA28" ref-type="supplementary-material" rid="S1">1</xref>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWSAG" type="diagnosis">
<paragraph id="99FF69F69AA002AB3C20BAA0C090116F">
<heading id="B52CFB2796647DEA5714BD4E46AD363D" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="6382BD6F79F06098FA10FAD786E00AE9">Arms six. Abactinal disk plate, each with one fairly elongated spine. Pedicellariae absent on abactinal disk. Abactinal arm plates each bears one spine, covered with a membranous sheath loaded with small pedicellariae. Each adambulacral plate with one small aboral furrow spine and one subambulacral spine. Lateral spines corresponding to each adambulacral plate beyond genital area. Oral plate with two actinostomal spines, one suboral spine, and one aboral spine along the furrow margin.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2SAG" type="distribution">
<paragraph id="DEEF445BF9B12B837AEFD5166EAE3422">
<heading id="5BD31C500BBA74188A7CE444DC082D3F" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="9BFA6B8949F18EBC816EDB8DFA650183">
North Atlantic,
<quantity id="1C5F775DE0329443324E1B99987FB7B7" metricMagnitude="3" metricUnit="m" metricValue="4.5649999999999995" metricValueMax="5.11" metricValueMin="4.02" unit="m" value="4565.0" valueMax="5110.0" valueMin="4020.0">40205110 m</quantity>
. Type locality: Between the Azores and
<collectingCountry id="26A3F8B6E4F7F6BE7A09C1D3641BC051" name="France">France</collectingCountry>
,
<quantity id="8BC999D7D4E19D71FF19387247AA379A" metricMagnitude="3" metricUnit="m" metricValue="4.06" unit="m" value="4060.0">4060 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EATAG" type="remarks">
<paragraph id="9C452AE1E7B1A6458E3E30F51E2AA3AE">
<heading id="1F24439D887CAA1ED185DF847DD2F4EE" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="02C2410E28F19F72D9E02A960E162402">
<taxonomicName id="E5413D0D68E15C899A6AF05E811CFB19" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="E55F655FB8BBBF058055C0AC0AA9E173" italics="true">F. sexradiata</emphasis>
</taxonomicName>
is the type species of the genus. The
<typeStatus id="E8B45C8B9D13BAA0A2F7632576D45676" type="holotype">holotype</typeStatus>
was examined in the present study, which demonstrates the key characters of the species as described by
<bibRefCitation id="4FF6FA5CABE2A4BD5C71CD4EA8A67D37" author="Perrier" firstAuthor="Perrier" issue="8" journalOrPublisher="Annales des sciences naturelles, Zoologie" pagination="1-72" refId="B53" refString="Perrier E (1885) Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman. Annales des sciences naturelles, Zoologie 22 (8): 172. [in French]" title="Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman." volume="22" year="1885">Perrier (1885</bibRefCitation>
,
<bibRefCitation id="2D5E0740485AE7E730C851B8DB9D6093" author="Perrier" firstAuthor="Perrier" journalOrPublisher="Expéditions Scientifiques du Travailleur et du Talisman" pagination="1-431" refId="B54" refString="Perrier E (1894) Stellérides. Expéditions Scientifiques du Travailleur et du Talisman 3: 1431. [in French]" title="Stellérides." volume="3" year="1894">1894</bibRefCitation>
), including having abactinal arm plate with one long spine, adambulacral plate with one subambulacral spine and one small aboral furrow spine, oral plate with two actinostomal spines, one suboral spine, and one aboral furrow spine. The arrangement of inferomarginal plates was not described in
<taxonomicName id="418315EE473EB11C10B38109874E3AA6" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="D79893A0185A7C3624FFC9858CBDC249" italics="true">F. sexradiata</emphasis>
</taxonomicName>
(
<bibRefCitation id="7C9371BE270D179E4D46A01609F6F460" author="Perrier E" issue="8" journalOrPublisher="Annales des sciences naturelles, Zoologie" pagination="1-72" refId="B53" refString="Perrier E (1885) Premiere note Preliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman. Annales des sciences naturelles, Zoologie 22 (8): 1-72. [in French]" title="Premiére note Préliminaire des les Echinodermes, recueillis durant les campagnes de dragages sous-marines du Travailleur et du Talisman." volume="22" year="1885">Perrier, 1885</bibRefCitation>
,
<bibRefCitation id="6195AC60C4EF84E2365EA25BBC051B04" author="Perrier E" journalOrPublisher="Expéditions Scientifiques du Travailleur et du Talisman" pagination="1-431" refId="B54" refString="Perrier E (1894) Stellerides. Expeditions Scientifiques du Travailleur et du Talisman 3: 1-431. [in French]" title="Stellérides." volume="3" year="1894">1894</bibRefCitation>
), and the
<typeStatus id="C5E8D7A5BAB825D78EC8FF960946122A" type="holotype">holotype</typeStatus>
was too broken to be examined for such a character. Based on the drawings in
<bibRefCitation id="1064BDB6FF19635054DBE976884BA61C" author="Perrier E" firstAuthor="Perrier" journalOrPublisher="Expéditions Scientifiques du Travailleur et du Talisman" pagination="1-431" refId="B54" refString="Perrier E (1894) Stellérides. Expéditions Scientifiques du Travailleur et du Talisman 3: 1431. [in French]" title="Stellérides." volume="3" year="1894">Perrier (1894)</bibRefCitation>
, the inferomarginal plates appear at each adambulacral plate beyond genital area. Another specimen examined (
<named-content id="1832782A40D996A6B535F99C50D5784F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/musee-oceanographique-monaco" xlink_title="Musée Océanographique de Monaco">
<collectionCode id="43BB63EEBEE58019C95F7B7C75388685" country="Monaco" httpUri="http://grbio.org/cool/cx5v-6vpi" name="Musee Oceanographique Monaco" type="">MOM</collectionCode>
</named-content>
- 81 0428) also has continuous inferomarginal plates beyond genital area. The photos of the
<typeStatus id="39BC209DE3614D6981A2C298C7242EC7" type="holotype">holotype</typeStatus>
of
<taxonomicName id="24FA6FB34547E9FFBAF1BFC6CAF605A9" baseAuthorityName="Perrier" baseAuthorityYear="1885" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="sexradiata">
<emphasis id="51A4D1EC4AAA60D6D09FC15210887E59" italics="true">F. sexradiata</emphasis>
</taxonomicName>
are available on the
<named-content id="C5B125F0167E265BA03FD63964532524" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national dHistoire naturelle">
<collectionCode id="256EBCA9D1CE5920646135F9D238C46F" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
collection website (
<typeStatus id="158C627F03A7DCFF4C32A04E16508897" type="holotype">holotype</typeStatus>
:
<ext-link id="A34244E984656F28DC30AB6E0848B789" ext-link-type="uri" xlink_href="https://science.mnhn.fr/institution/mnhn/collection/ie/item/2014-165" xlink_type="simple">https://science.mnhn.fr/institution/mnhn/collection/ie/item/2014-165</ext-link>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,177 @@
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<mods:title id="29D8DFF7DEA983B96D452046A36F6BCB">Hidden diversity of Freyastera (Asteroidea, Brisingida, Freyellidae) at great depth: Description of new species and remarks on species boundaries</mods:title>
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<mods:identifier id="EBB90A70800019D9936B404F008F7190" type="ZooBank">4611CD60-658B-42C4-B23C-2DA9185A85E3</mods:identifier>
</mods:mods>
<treatment id="FC59587043135738A06C5D58D8DA679B" ID-arpha="FC595870-4313-5738-A06C-5D58D8DA679B" LSID="urn:lsid:plazi:treatment:FC59587043135738A06C5D58D8DA679B" httpUri="http://treatment.plazi.org/id/FC59587043135738A06C5D58D8DA679B" scope_class="Asteroidea" scope_family="Freyellidae" scope_order="Brisingida">
<subSubSection id="E1EF1AEA6A719911C5BC5E9B03630FDD" type="nomenclature">
<paragraph id="B8A8252D4E17D072353731D6A489D1D1">
<taxonomicName id="87A33F4CA83D96ED05EEC47CA346510F" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-2">
<emphasis id="8FB00D262307D5230DE69DB0090129D5" italics="true">Freyastera</emphasis>
sp. 2
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="25221B5B679A1964BD294661888C2179" type="description">
<paragraph id="B5AB8124D7FA3ABCC0CF0FF27DD30B63">
<figureCitation id="515D5B1936FD473E8829E89BC95F9BC3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Freyastera sp. 2. A, B, F. NHMUK 2022.81; C E. RSIOAST 0103. A. Abactinal view of disk; B. Actinal view of disk; C. Abactinal view of arm genital area; D. Zoom-in view of oral plates; E. Abactinal view of middle part of arm; F. Zoom-in view of middle part of arm, showing pedicellariae clustered in small pads." figureDoi="10.3897/zse.101.144918.figure10" httpUri="https://binary.pensoft.net/fig/1304661">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCCD459AA8025B362AE45FD4A5B5CC0F" type="reference_group">
<paragraph id="5E055323C87B655600E9600C61FC861A">
<treatmentCitationGroup id="386C119536510162C458CE9C01E9DF5C">
<taxonomicName id="62C4D1F5AB2FD28F8753CFC82D038374" ID-CoL="7P2S3" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="A7FF0DF864AAB49EEC55EBFB907BFC1C" italics="true">Freyastera</emphasis>
</taxonomicName>
<comment id="43FB28E3CE13075DC43504D0135F8650">
stet. CCZ_201:
<treatmentCitation id="68F1C0694051CF407BD213E03ED337EF" author="Bribiesca-Contreras G &amp; Dahlgren TG &amp; Amon DJ &amp; Cairns S &amp; Drennan R &amp; Durden JM &amp; Eléaume MP &amp; Hosie AM &amp; Kremenetskaia A &amp; McQuaid K &amp; D OHara T" httpUri="http://treatment.plazi.org/id/7C871719323051E2B0B9D8B017C0B3E7" page="44" year="2022">
<bibRefCitation id="BA432CD6F93D383A4C56AF460D634A9F" DOI="10.3897/zookeys.1113.82172" author="Bribiesca-Contreras G &amp; Dahlgren TG &amp; Amon DJ &amp; Cairns S &amp; Drennan R &amp; Durden JM &amp; Eléaume MP &amp; Hosie AM &amp; Kremenetskaia A &amp; McQuaid K &amp; D OHara T" etAl="et al." firstAuthor="Bribiesca-Contreras" page="44" refId="B4" refString="Bribiesca-Contreras G, Dahlgren TG, Amon DJ, Cairns S, Drennan R, Durden JM, Eléaume MP, Hosie AM, Kremenetskaia A, McQuaid K, D OHara T (2022) Benthic megafauna of the western Clarion-Clipperton Zone, Pacific Ocean. ZooKeys 1113: 1. https://doi.org/10.3897/zookeys.1113.82172" year="2022">Bribiesca-Contreras et al. 2022: 44</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE6AI" type="material">
<paragraph id="35141DC0B0CB773B6949310FE0245BC7">
<heading id="BB44BA561C54E917830A7A31E6D9145E" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="28BBDFD1964AB6D21D6F76DB6BA56650">
<materialsCitation id="EC35D3B26DC73B9730506FF1120E36BB" collectionCode="NHMUK" specimenCode="NHMUK 2022.81" specimenCount="1">
<specimenCode id="B1822447F219978213F5303D132893D9">
<named-content id="1534533DBC787067C48CB2351D4A157C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum">NHMUK</named-content>
2022.81
</specimenCode>
(Fig.
<figureCitation id="70ACA76C1E3BC310FE6F6E5821CDB821" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Freyastera sp. 2. A, B, F. NHMUK 2022.81; C E. RSIOAST 0103. A. Abactinal view of disk; B. Actinal view of disk; C. Abactinal view of arm genital area; D. Zoom-in view of oral plates; E. Abactinal view of middle part of arm; F. Zoom-in view of middle part of arm, showing pedicellariae clustered in small pads." figureDoi="10.3897/zse.101.144918.figure10" httpUri="https://binary.pensoft.net/fig/1304661">10 A, B, F</figureCitation>
) (
<taxonomicName id="C2C7C9A84DB8E2927CD9959ABEACD295" authorityName="Downey" authorityYear="1986" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="genus">
<emphasis id="A09F7B6911F4AE1AF4D26F912394096C" italics="true">Freyastera</emphasis>
</taxonomicName>
stet. CCZ_201)
</materialsCitation>
;
<materialsCitation id="D9C6371746A61F87EF6DF95DCD3805E1" collectionCode="RSIOAST" specimenCode="RSIOAST 0103" specimenCount="1">
<specimenCode id="4857030B3F91A64045E8DAA2E3C41D00">RSIOAST 0103</specimenCode>
(Fig.
<figureCitation id="671915C21E419672F3865018EFEE5D5D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Freyastera sp. 2. A, B, F. NHMUK 2022.81; C E. RSIOAST 0103. A. Abactinal view of disk; B. Actinal view of disk; C. Abactinal view of arm genital area; D. Zoom-in view of oral plates; E. Abactinal view of middle part of arm; F. Zoom-in view of middle part of arm, showing pedicellariae clustered in small pads." figureDoi="10.3897/zse.101.144918.figure10" httpUri="https://binary.pensoft.net/fig/1304661">10 C E</figureCitation>
). (Suppl. material
<xref id="95A2A57B71B3DFCB610158D4A8BE7A3C" ref-type="supplementary-material" rid="S1">1</xref>
)
</materialsCitation>
.
</paragraph>
<caption id="747B751DCBA5011411A7E5D462573C17" ID-DOI="10.3897/zse.101.144918.figure10" ID-arpha="0A16E5E1-1BCC-5B87-B7A3-932BF085AD4D" httpUri="https://binary.pensoft.net/fig/1304661" startId="F10">
<paragraph id="18A52A219AC7B975CAD4633900F6270C">
<label id="64339FD474BE03EACC4C2374A3099A95">Figure 10.</label>
</paragraph>
<paragraph id="7A12FB7E45BB2ED1EAA23DFAAF65191C">
<taxonomicName id="937DC55D0D09C80CE46C5ABB7F8EE328" class="Asteroidea" family="Freyellidae" genus="Freyastera" kingdom="Animalia" order="Brisingida" phylum="Echinodermata" rank="species" species="undefined-2">
<emphasis id="1FBEF64E8B697F6C92DC4A409FA4D747" italics="true">Freyastera</emphasis>
sp. 2
</taxonomicName>
.
<emphasis id="B98E01BF42F015F5690DF7BBE4036DF4" bold="true">
A, B,
<emphasis id="59745DC1719B324ED85BBFBEF68604DB" italics="true">F.</emphasis>
</emphasis>
<specimenCode id="AD2E4A293AAF76AA97549CC3318C1A15">
<named-content id="7018C244C77BCE737B057BA1C95108DE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum">NHMUK</named-content>
2022.81
</specimenCode>
;
<emphasis id="BB3665784E968EC316470F1FA083F284" bold="true">C E.</emphasis>
<specimenCode id="AEBC85DBED665EB3877901AC00E9288F">RSIOAST 0103</specimenCode>
.
<emphasis id="1FD329252CBF61B9483D349A5648BC11" bold="true">A.</emphasis>
Abactinal view of disk;
<emphasis id="1B36F0CD0D95D7C52E2A5DC07C7D78A0" bold="true">B.</emphasis>
Actinal view of disk;
<emphasis id="1BAAFD4F82601D1999559DBDFA632ACE" bold="true">C.</emphasis>
Abactinal view of arm genital area;
<emphasis id="9BD23969B443656966355889678F12A0" bold="true">D.</emphasis>
Zoom-in view of oral plates;
<emphasis id="13668BF418DD36C17C710D4B5A625BE6" bold="true">E.</emphasis>
Abactinal view of middle part of arm;
<emphasis id="FBB5D20347CE2A846D07AD253F83A0A4" bold="true" italics="true">F.</emphasis>
Zoom-in view of middle part of arm, showing pedicellariae clustered in small pads.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EUBBI" type="distribution">
<paragraph id="2D16550193455AC61AEE1C6B16533102">
<heading id="828356C5A32FE80C1C95525B922812EA" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="3B0B4F0E88A3A786EDB84BE581D4AA78">
Eastern Pacific: Clarion-Clipperton Zone; Northwest Pacific: Western Philippine Basin.
<quantity id="8556A7B2BCCF55E70B3D1D35FF2AF8CF" metricMagnitude="3" metricUnit="m" metricValue="5.552" metricValueMax="5.9" metricValueMin="5.204" unit="m" value="5552.0" valueMax="5900.0" valueMin="5204.0">52045900 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>