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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.104844" ID-Pensoft-Pub="1313-2970-1168-231" ID-Pensoft-UUID="6928794CB97051F9AAE3B64050E5FAF3" ID-ZooBank="28A21391A4334E808845E5C0996EFAF0" ModsDocID="1313-2970-1168-231" checkinTime="1688439534387" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Kluge, Nikita J. &amp; Gattolliat, Jean-Luc" docDate="2023" docId="267D6366A4C1527686B8B3FE40CBBDDD" docLanguage="en" docName="ZooKeys 1168: 231-266" docOrigin="ZooKeys 1168" docPubDate="2023-07-03" docSource="http://dx.doi.org/10.3897/zookeys.1168.104844" docTitle="Arcobaetis bornensis Kaltenbach &amp; Kluge &amp; Gattolliat 2023, sp. nov." docType="treatment" docUuid="1FD12A10-3730-4CED-AC63-2EFCE4894444" docUuidSource="ZooBank" docVersion="2" id="6928794CB97051F9AAE3B64050E5FAF3" lastPageNumber="231" masterDocId="6928794CB97051F9AAE3B64050E5FAF3" masterDocTitle="A new, widespread genus of Baetidae from South Asia (Insecta, Ephemeroptera)" masterLastPageNumber="266" masterPageNumber="231" pageNumber="231" updateTime="1688439835494" updateUser="ExternalLinkService">
<mods:mods id="67BCC134EAEF756D33668216359C68E1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="980C99C0148478DD6A41382BCC9FCA19">
<mods:title id="4F64CF46669E9DC1709382EE2AEA7F78">A new, widespread genus of Baetidae from South Asia (Insecta, Ephemeroptera)</mods:title>
</mods:titleInfo>
<mods:name id="50B8536147DD1A212A34A8C63B641E7C" type="personal">
<mods:role id="741A5A7C9CFE13A0D1219C4EF8598774">
<mods:roleTerm id="91281AF5CA2652C88A88B89C542993CF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A745C59CC9AE98CA6DAAC1CA66034DC6">Kaltenbach, Thomas</mods:namePart>
<mods:nameIdentifier id="B80CEB6AE2645A03FEB66ACEBB6E15AA" type="ORCID">https://orcid.org/0000-0001-8052-0388</mods:nameIdentifier>
<mods:affiliation id="1DCA50B1FEA5E5A688F0F1FE5A678394">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier id="E4DC4233C4BBD17D14C27DC4D361463C" type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:name id="D449DCCF5868D68FEF97C360BD75BA35" type="personal">
<mods:role id="5EFB5CB71C952A19DC8629C31A7F76D9">
<mods:roleTerm id="AC617567E36FC4410E430AD66C079894">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4468E7C1BA89F450FBD4EDC3B527226E">Kluge, Nikita J.</mods:namePart>
<mods:nameIdentifier id="B00F0815E7BE1872FAFC950E40D16AF4" type="ORCID">https://orcid.org/0000-0001-9741-7790</mods:nameIdentifier>
<mods:affiliation id="8D08A9EFBAEA4D3D4C688390C7272ADB">Department of Entomology, Biological Faculty, Saint-Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
</mods:name>
<mods:name id="2823F3F58FB66F86EEF42BA477C66E18" type="personal">
<mods:role id="AF82F22FBA12D94120B6542682365A28">
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<mods:namePart id="74ACD336EB5DFD588C8305590E68E8A8">Gattolliat, Jean-Luc</mods:namePart>
<mods:nameIdentifier id="C70353E62F2E03A695E2CDEF17A4D6F4" type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
<mods:affiliation id="C692750427DC48034E57F9BCB19108FB">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:title id="C1ED10716938E5F82A708D67F5EB6671">ZooKeys</mods:title>
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<mods:part id="6843C4049ADD161F804673A0E3CCEBA1">
<mods:date id="B2BB4A2FD0BD05701C2314752F71FEEA">2023</mods:date>
<mods:detail id="5B0C18C91D751F34159B28FF13022E56" type="pubDate">
<mods:number id="8E0B1207BC1193399B625DED8811B127">2023-07-03</mods:number>
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<mods:detail id="58769B0BE0B6030DC832CA41406E08CA" type="volume">
<mods:number id="F866396B38B0FD1ED51EF00AE2AE5BA6">1168</mods:number>
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<mods:start id="8CC5D663BBB17095230EE4CDEE5552F6">231</mods:start>
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<mods:identifier id="73015C56E95A64EB19C7CA6805FA61CC" type="DOI">http://dx.doi.org/10.3897/zookeys.1168.104844</mods:identifier>
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<treatment id="267D6366A4C1527686B8B3FE40CBBDDD" LSID="urn:lsid:zoobank.org:act:1FD12A10-3730-4CED-AC63-2EFCE4894444" httpUri="http://treatment.plazi.org/id/267D6366A4C1527686B8B3FE40CBBDDD" lastPageNumber="231" pageId="0" pageNumber="231">
<subSubSection id="1F0B274C9D5DEA8636FA42DC8B427713" pageId="0" pageNumber="231" type="nomenclature">
<paragraph id="EB0272366B3A16EE0D617885E753999C" pageId="0" pageNumber="231">
<taxonomicName id="40C3DFF48047C4DF2FF3EE8CF0AFF002" LSID="https://zoobank.org/1FD12A10-3730-4CED-AC63-2EFCE4894444" authority="Kaltenbach &amp; Kluge &amp; Gattolliat, 2023" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Arcobaetis bornensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="bornensis" status="sp. nov.">Arcobaetis bornensis</taxonomicName>
<taxonomicNameLabel id="3848080217923AA30B42606CE55BC5D3" pageId="0" pageNumber="231">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="1CB51B31737A70E335C0C0C4EE6715F1" pageId="0" pageNumber="231" type="description">
<paragraph id="76671AAD192128573ACEFC9EE62BEA97" pageId="0" pageNumber="231">
<figureCitation id="D512AF34CB3830AB97FD49A4BEE9568C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">Figs 10</figureCitation>
<figureCitation id="2FDBE2D2863218BB6DBD256949021CB4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">, 11</figureCitation>
<figureCitation id="9013873A21CD51EEC3F542D72FF9254A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Arcobaetis bornensis sp. nov., larva a foreleg b hind leg c habitat (photo Kate Baker, University Exeter, UK)." figureDoi="10.3897/zookeys.1168.104844.figure12" httpUri="https://binary.pensoft.net/fig/872214" pageId="0" pageNumber="231">, 12</figureCitation>
<figureCitation id="FA8AF4E01C57B00F12F192F238B5352E" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Arcobaetis gen. nov., distribution." figureDoi="10.3897/zookeys.1168.104844.figure19" httpUri="https://binary.pensoft.net/fig/872221" pageId="0" pageNumber="231">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="761838AF6EABF57DACA1A703301AF7DF" pageId="0" pageNumber="231" type="type material">
<paragraph id="03422A0C1B4DE63AD89B7DE9380D78CC" pageId="0" pageNumber="231">Type material.</paragraph>
<paragraph id="CBF0C177B009103B57A94B31E65660D1" pageId="0" pageNumber="231">
<emphasis id="D03176D27553501AB0EE258422C0168A" bold="true" pageId="0" pageNumber="231">
<emphasis id="413B85A56CE522771AFDD5753C0C3D94" italics="true" pageId="0" pageNumber="231">Holotype</emphasis>
.
</emphasis>
Brunei • female larva (premature); Temburong District, Ulu Temburong National Park, near Kuala Belalong Field Studies Centre (KBFSC);
<geoCoordinate id="9F9820267D38E6EE39BC4586A9244DF0" degrees="04" direction="north" minutes="32" orientation="latitude" precision="15" seconds="55" value="4.548611">04°32'55&quot;N</geoCoordinate>
,
<geoCoordinate id="2D2E897AE5CFBEA42F6923EFD88401AC" degrees="115" direction="east" minutes="09" orientation="longitude" precision="15" seconds="27" value="115.1575">115°09'27&quot;E</geoCoordinate>
; 103 m; May 2014; leg. Kate Baker; on slide; GBIFCH00465126; MZL.
<emphasis id="B1B8A69B2F1AF8D477AA09592A3D694E" bold="true" italics="true" pageId="0" pageNumber="231">Paratypes.</emphasis>
11 larvae; same data as holotype; 5 on slides; GBIFCH00465127, GBIFCH00465128, GBIFCH00465129, GBIFCH00592613, GBIFCH00592645; MZL; 6 in alcohol; GBIFCH00515212, GBIFCH00975703; MZL.
</paragraph>
</subSubSection>
<subSubSection id="1E219551EA9ABD8AA07E4EC4222DF135" pageId="0" pageNumber="231" type="diagnosis">
<paragraph id="0A959F0132368D73F925B7CEC9F6B479" pageId="0" pageNumber="231">Diagnosis.</paragraph>
<paragraph id="C57D14DEE2C816F9256BF84F3278577E" pageId="0" pageNumber="231">
<emphasis id="8E4F2473CE94C59DCAAFCB7C48AA8FB0" bold="true" pageId="0" pageNumber="231">Larva</emphasis>
(Table
<tableCitation id="88052229DEF9508D7CA70E10C97EB1DF" captionStart="Table 1" captionStartId="T1" captionText="Table 1. States of selected larval characters of Arcobaetis gen. nov. Figure numbers refer to those in this paper, while those of A. gracilentus comb. nov. refer to Kang et al. (1994)." httpUri="http://table.plazi.org/id/2503F8B5E92CE0301DEF57DAFBE238D1" pageId="0" pageNumber="231" tableUuid="2503F8B5E92CE0301DEF57DAFBE238D1">1</tableCitation>
). The following combination of characters distinguish
<taxonomicName id="FC8075D6BF16CAFD629FA1DB3637B0AB" lsidName="A. bornensis" pageId="0" pageNumber="231" rank="species" species="bornensis">
<emphasis id="5748CA551F84AC952B76893DDF4E8464" italics="true" pageId="0" pageNumber="231">A. bornensis</emphasis>
</taxonomicName>
sp. nov. from other species of
<taxonomicName id="D11FA87E0A832C5B5249DF38B16B9068" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="genus">
<emphasis id="5ACBF93C632AF7E35B7354AFF45B7893" italics="true" pageId="0" pageNumber="231">Arcobaetis</emphasis>
</taxonomicName>
gen. nov.: A) distal margins of segments in middle part of flagellum with enlarged spines (Fig.
<figureCitation id="67498785404F9BCA85C77BB605D392EF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">11f</figureCitation>
); B) labial palp segment III sub-quadrangular, at base narrower than distal margin of segment II (Fig.
<figureCitation id="6A781699E61809AEDB767DC78F3E20E3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">10h</figureCitation>
); C) hind protoptera absent; D) claw with single row of denticles, distalmost denticle larger, other denticles minute (Fig.
<figureCitation id="37CCB847494780D64ACCB6BF8230374A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">11b</figureCitation>
); E) tergalii present on abdominal segments I-VII; F) posterior margin of tergum IV with triangular spines, wider than long (Fig.
<figureCitation id="4413D9CD670B09874B9F56CFB1FE51F6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">11c</figureCitation>
).
</paragraph>
<caption id="3E152A91C4528AA651F1938F19B14887" doi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231" start="Figure 10" startId="F10">
<paragraph id="49054DEB80CC288E4E4B6B6CF2A6776E" pageId="0" pageNumber="231">
<emphasis id="F4FBE11AF19D488F58FF56915F287EA2" bold="true" pageId="0" pageNumber="231">Figure 10.</emphasis>
<taxonomicName id="5C60155167800E8ACAB0B92A5D06789B" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis bornensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="bornensis">
<emphasis id="6509A627EFA1C8F843EE396AA1E7D533" italics="true" pageId="0" pageNumber="231">Arcobaetis bornensis</emphasis>
</taxonomicName>
sp. nov., larva
<emphasis id="43549FE655E034C0028EFBF4858658CA" bold="true" pageId="0" pageNumber="231">a</emphasis>
labrum (left: ventral view; right: dorsal view)
<emphasis id="F628095F6A51C10EC96F6B2ADC0D51CF" bold="true" pageId="0" pageNumber="231">b</emphasis>
right mandible
<emphasis id="2D8B60E9E54038DEF87D51BF9DF78521" bold="true" pageId="0" pageNumber="231">c</emphasis>
right prostheca
<emphasis id="3F699337CA69B1B5A3801972883C4906" bold="true" pageId="0" pageNumber="231">d</emphasis>
left mandible
<emphasis id="AFC9459759DE4FC9E0133FAFD8B6356B" bold="true" pageId="0" pageNumber="231">e</emphasis>
left prostheca
<emphasis id="626C09316B245FEC0D7DC8F5D4B12A4F" bold="true" pageId="0" pageNumber="231">f</emphasis>
hypopharynx and superlinguae
<emphasis id="525E36F60380070D1947B9EA706BF174" bold="true" pageId="0" pageNumber="231">g</emphasis>
maxilla
<emphasis id="B923E11CF59D0A2A535FC59F83BE11EB" bold="true" pageId="0" pageNumber="231">h l</emphasis>
abium (left: ventral view; right: dorsal view)
<emphasis id="537D562B56B0F2BD2CE423979A450125" bold="true" pageId="0" pageNumber="231">i</emphasis>
apex of paraglossa (ventral view)
<emphasis id="7DED68E31BA77D0CCCE46B8E17AADA89" bold="true" pageId="0" pageNumber="231">j</emphasis>
apex of paraglossa (dorsal view).
</paragraph>
</caption>
<caption id="B537F45767656F201D370CA646053170" doi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231" start="Figure 11" startId="F11">
<paragraph id="37760F873F29CEFABEA3681C1AF52BC8" pageId="0" pageNumber="231">
<emphasis id="15F2404120E6555788931E4A508F1464" bold="true" pageId="0" pageNumber="231">Figure 11.</emphasis>
<taxonomicName id="ED3183E278D062FF38C41A8AB249A025" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis bornensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="bornensis">
<emphasis id="F091401A6A0EB70EE052AA9F1852564B" italics="true" pageId="0" pageNumber="231">Arcobaetis bornensis</emphasis>
</taxonomicName>
sp. nov., larva
<emphasis id="88D3C58D76DAA8DF23AAE69D3C85BC00" bold="true" pageId="0" pageNumber="231">a</emphasis>
foreleg
<emphasis id="A5883844E52B7DF131E0484BD47FD5C2" bold="true" pageId="0" pageNumber="231">b</emphasis>
fore claw
<emphasis id="7FE5CC3CB37FFC90054843ED3AF43C05" bold="true" pageId="0" pageNumber="231">c</emphasis>
tergum IV
<emphasis id="B98BE0936559BEFB79842F26C572FFBC" bold="true" pageId="0" pageNumber="231">d</emphasis>
tergalius IV
<emphasis id="383637919394AE5387992FF6C58788E7" bold="true" pageId="0" pageNumber="231">e</emphasis>
paraproct
<emphasis id="0CC18BB56212C0FEA004C3BF0C353082" bold="true" pageId="0" pageNumber="231">f</emphasis>
antenna (right: dorsal view; left: section in lateral view).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E6E460F130D8E91B637F6C142E9CFDE2" pageId="0" pageNumber="231" type="description">
<paragraph id="47A7C21178CACB16D991E7EDFE4F473B" pageId="0" pageNumber="231">Description.</paragraph>
<paragraph id="30BAD1A0CA95FDC15427CB3CD1E5724E" pageId="0" pageNumber="231">
<emphasis id="E321F4CE7487A68307E2D9FE6CB2F76F" bold="true" pageId="0" pageNumber="231">Larva</emphasis>
(Figs
<figureCitation id="5BEAE6276A553D7A00CE47388ECD166D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">10</figureCitation>
-
<figureCitation id="A1DA2D27A80E26AB83B7DB8B78B861FF" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Arcobaetis bornensis sp. nov., larva a foreleg b hind leg c habitat (photo Kate Baker, University Exeter, UK)." figureDoi="10.3897/zookeys.1168.104844.figure12" httpUri="https://binary.pensoft.net/fig/872214" pageId="0" pageNumber="231">12</figureCitation>
). Body length ca. 3.0 mm.
</paragraph>
<caption id="579D8B826FB2E9E890A56F3BE7082033" doi="10.3897/zookeys.1168.104844.figure12" httpUri="https://binary.pensoft.net/fig/872214" pageId="0" pageNumber="231" start="Figure 12" startId="F12">
<paragraph id="A3494EE76FC6969C1C48124CF85363CE" pageId="0" pageNumber="231">
<emphasis id="141222E714BE30B8B346AA5B5B372E47" bold="true" pageId="0" pageNumber="231">Figure 12.</emphasis>
<taxonomicName id="9F2B79CE3A61850DE26D3278FB952281" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis bornensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="bornensis">
<emphasis id="87C3EB320BD2302B941ED27D315F6DDE" italics="true" pageId="0" pageNumber="231">Arcobaetis bornensis</emphasis>
</taxonomicName>
sp. nov., larva
<emphasis id="F19E81FB8A5036356701895BB55651E6" bold="true" pageId="0" pageNumber="231">a</emphasis>
foreleg
<emphasis id="927A488167B4FA8CEE1A47B4175B1541" bold="true" pageId="0" pageNumber="231">b</emphasis>
hind leg
<emphasis id="675F3E167CA65F42FBD630985048FEE0" bold="true" pageId="0" pageNumber="231">c</emphasis>
habitat (photo Kate Baker, University Exeter, UK).
</paragraph>
</caption>
<paragraph id="AF7CA591C09BD3069ACB796D75A6DE32" pageId="0" pageNumber="231">
<emphasis id="FD771DFD14CE73F6E701B5C675AEE656" bold="true" italics="true" pageId="0" pageNumber="231">Colouration</emphasis>
. Head, thorax, and abdomen dorsally brown, ventrally ochre. Legs ecru. Caudalii pale brown.
</paragraph>
<paragraph id="4845FEC0D89C59E6795AEE74927CF684" pageId="0" pageNumber="231">
<emphasis id="6E1286132B88E182F207762236656C70" bold="true" italics="true" pageId="0" pageNumber="231">Hypodermal colouration.</emphasis>
Unknown.
</paragraph>
<paragraph id="5B7E0B0F8DAE61C825D49845E0230F56" pageId="0" pageNumber="231">
<emphasis id="4EEC85311EA7467B2C94729B4E63CD6A" bold="true" italics="true" pageId="0" pageNumber="231">Antenna.</emphasis>
Flagellum in middle part with enlarged spines at distal margin of segments (Fig.
<figureCitation id="C678D1E59A35FF22ADC4EB6CA8904A39" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">11f</figureCitation>
).
</paragraph>
<paragraph id="39797CF55626F4716A5BBF1C731F1D1A" pageId="0" pageNumber="231">
<emphasis id="872B49FC8CD93A65255CD6121201BD81" bold="true" italics="true" pageId="0" pageNumber="231">Labrum</emphasis>
(Fig.
<figureCitation id="E17BB6E3968026CA40AABF4AFFED2DE7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">10a</figureCitation>
). Length 0.7
<normalizedToken id="641C68CE4DACE4513F82CD649710F6DF" originalValue="×">x</normalizedToken>
maximum width. Dorsally with sub-median seta, sub-marginal arc of ca. three simple setae, and several setae in between.
</paragraph>
<paragraph id="874FA2D2A57F5436A782735AB355FF3A" pageId="0" pageNumber="231">
<emphasis id="60B652CC7677942DB1EE347774F39D33" bold="true" italics="true" pageId="0" pageNumber="231">Right mandible</emphasis>
(Fig.
<figureCitation id="3B950AE2FB9426552C9552CED3945D67" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">10b, c</figureCitation>
). Incisor with five denticles; kinetodontium with three denticles.
</paragraph>
<paragraph id="16D840BF9BC8FD2634ADC889A5590FA7" pageId="0" pageNumber="231">
<emphasis id="50D1F72E89FC965C29BBB36E7C8C0DE0" bold="true" italics="true" pageId="0" pageNumber="231">Left mandible</emphasis>
(Fig.
<figureCitation id="B8C2DD3AE7975FEF0DF1D976FA126D93" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">10d, e</figureCitation>
). Incisor with four denticles; kinetodontium with three denticles.
</paragraph>
<paragraph id="506A82DA6CB29D3C34BB04D020FB046C" pageId="0" pageNumber="231">
<emphasis id="34FDD25A401D775889418708B8FB6F59" bold="true" italics="true" pageId="0" pageNumber="231">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitation id="06F952B591CCE56B4A7F9E4390C13D51" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">10f</figureCitation>
). Lingua slightly shorter than superlinguae, longer than broad. Superlinguae distally straight; lateral margins rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph id="1A47F4C9C14DFFFB429A471C4D26BAE7" pageId="0" pageNumber="231">
<emphasis id="F6B361FCF3C766395ED673319B6C1857" bold="true" italics="true" pageId="0" pageNumber="231">Maxilla</emphasis>
(Fig.
<figureCitation id="925D8AD6670E67A0D6A0B3A51E66408C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">10g</figureCitation>
). Galea-lacinia ventrally with two simple setae just proximad of canines. Medially with one pectinate, spine-like seta and ca. four medium to long, simple setae. Maxillary palp approx. 1.4
<normalizedToken id="C6A9CFCBEE85A56E4AE315C782021B8D" originalValue="×">x</normalizedToken>
as long as galea-lacinia; palp segment II approx. 1.2
<normalizedToken id="DEEEFEB2EB4B40DC51579BE5F2E680AE" originalValue="×">x</normalizedToken>
as long as segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II.
</paragraph>
<paragraph id="6111064822D0CF04CEDA8A4AB4C32803" pageId="0" pageNumber="231">
<emphasis id="F00CD200232504A8887E4A16852A53F7" bold="true" italics="true" pageId="0" pageNumber="231">Labium</emphasis>
(Fig.
<figureCitation id="7A6DF2258F57DABD0BAFE062AEDAD032" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">10h-j</figureCitation>
). Inner margin of glossa with ca. nine spine-like setae; apex with three long robust setae; outer margin with ca. nine spine-like setae; paraglossa ventrally with ca. four medium, simple setae in anteromedial area; dorsally with arc of ca. seven long, spine-like setae in distal area. Labial palp with segment I 0.8
<normalizedToken id="FA91C03674B957FEEA06A4E0540BA749" originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment II dorsally with row of ca. four spine-like setae. Segment III sub-quadrangular; at base narrower than distal margin of segment II; ventral surface with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<paragraph id="C1409CB1EE55A7B2AE3C8550749CED73" pageId="0" pageNumber="231">
<emphasis id="917A755F59A1D5B442BBE7910FBEA435" bold="true" italics="true" pageId="0" pageNumber="231">Hind protoptera</emphasis>
absent.
</paragraph>
<paragraph id="083A81795C404FA90F4D498267A2FF2C" pageId="0" pageNumber="231">
<emphasis id="D18B318951A005B65945E86A023C13EA" bold="true" italics="true" pageId="0" pageNumber="231">Legs</emphasis>
(Figs
<figureCitation id="56F8F07DA65784033BFDB292E07DF968" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">11a, b</figureCitation>
,
<figureCitation id="25946A412083825A4BB4593A159B7455" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Arcobaetis bornensis sp. nov., larva a foreleg b hind leg c habitat (photo Kate Baker, University Exeter, UK)." figureDoi="10.3897/zookeys.1168.104844.figure12" httpUri="https://binary.pensoft.net/fig/872214" pageId="0" pageNumber="231">12a, b</figureCitation>
). Long and slender, hind legs slenderer than fore leg, middle legs unknown. Ratio of leg segments: fore leg 1.5:1.0:0.9:0.3, hind leg 1.4:1.0:0.7:0.2.
<emphasis id="8E63FE06D0DF82AD5CD377420BD36A58" bold="true" pageId="0" pageNumber="231">
<emphasis id="A73DF4EA66D9CF0DC369658FC6A50EAB" italics="true" pageId="0" pageNumber="231">Femur</emphasis>
.
</emphasis>
Length of fore femur ca. 3.6
<normalizedToken id="90403DE2F62525C49E65BEA681CC1400" originalValue="×">x</normalizedToken>
maximum width, outer and inner margins almost parallel; length of hind femur ca. 5
<normalizedToken id="E97D013554089DE729B431178A7244F7" originalValue="×">x</normalizedToken>
maximum width, outer margin slightly concave. Outer margin with row of ca. 7 short, curved, spine-like setae, on fore leg larger than on hind leg. On ventral side of fore leg with short, spine-like, pectinate, pointed setae; nearly absent on hind leg.
<emphasis id="9FEAC5A4584B5418C4B295A5E0532A63" bold="true" pageId="0" pageNumber="231">
<emphasis id="5543AFCFF07F9D30790B62D33396C97B" italics="true" pageId="0" pageNumber="231">Tibia</emphasis>
.
</emphasis>
Few short, stout, pointed, pectinate setae near inner margin. Inner margin with row of short, curved, spine-like, pectinate setae, on apex two longer, curved, spine-like, pectinate setae. Outer margin with one apical, spine-like seta; on fore leg without other spine-like setae, on hind leg with few medium, spine-like setae. Patella-tibial suture on foreleg on proximal ¹/2 (difficult to see), on hind leg on proximal ¹/3.
<emphasis id="6661B39BD569EABE7BAB8019F1170A22" bold="true" italics="true" pageId="0" pageNumber="231">Tarsus</emphasis>
. Inner margin with row of short, curved, pectinate, spine-like setae; outer margin without spine-like setae.
<emphasis id="F986536D90BA207BDAAEE5FCD7602647" bold="true" italics="true" pageId="0" pageNumber="231">Claw</emphasis>
with one row of denticles, distalmost denticle larger and directed distad, other denticles minute.
</paragraph>
<paragraph id="67B74184C586A630010B2FF35526805A" pageId="0" pageNumber="231">
<emphasis id="11BB0609D2E5C64E4910A237AD938A05" bold="true" italics="true" pageId="0" pageNumber="231">Abdominal terga</emphasis>
(Fig.
<figureCitation id="4B9F8AAAAF5563F33BB240B8876E166F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">11c</figureCitation>
). Posterior margin of terga: I and II smooth, without spines; III-IX with triangular spines, slightly increasing in length toward IX; wider than long on tergum IV; row of spines on tergum IX interrupted by smaller spines in middle part behind bases of submedian setae (similar to Fig.
<figureCitation id="0EA2274D252DF807C44143C149681BFF" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure15" httpUri="https://binary.pensoft.net/fig/872217" pageId="0" pageNumber="231">15j</figureCitation>
). Posterior margin of tergum X with median concavity with smaller spines (similar to Fig.
<figureCitation id="ECC80A185231F94B38AF9A05D39E82C6" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure15" httpUri="https://binary.pensoft.net/fig/872217" pageId="0" pageNumber="231">15k</figureCitation>
).
</paragraph>
<paragraph id="49CC9C65FC577E1EC9BD81EC2996E178" pageId="0" pageNumber="231">
<emphasis id="DC5C999F11AD50D908CD623EF26C9D49" bold="true" pageId="0" pageNumber="231">
<emphasis id="34E2D45A3FC7919E5D3596889171E131" italics="true" pageId="0" pageNumber="231">Abdominal sternites</emphasis>
.
</emphasis>
Posterior margins of sterna: I-V smooth, without spines; VI-IX with triangular spines.
</paragraph>
<paragraph id="24A3F93B91A0C48C0050CFE1A67F9CEC" pageId="0" pageNumber="231">
<emphasis id="C60B47831107D2395A0A745C81205515" bold="true" italics="true" pageId="0" pageNumber="231">Tergalii</emphasis>
(Fig.
<figureCitation id="A2229479DAEFFB3F0A050C907FE5A16E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">11d</figureCitation>
). Present on abdominal segments I-VII. Tracheation limited to main trunk. Margin with minute denticles intercalating short, simple setae.
</paragraph>
<paragraph id="399631C3858758D4A4E7E6F7E13DC3DC" pageId="0" pageNumber="231">
<emphasis id="9F060437F4BA198BD1359813A266CCAF" bold="true" italics="true" pageId="0" pageNumber="231">Paraproct</emphasis>
(Fig.
<figureCitation id="BD728963723951BEC58C5B88EC55B8CB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">11e</figureCitation>
). Posterior margin with ca. 13 stout spines. Surface scattered with scale bases and micropores.
</paragraph>
<paragraph id="13D0535A438BB3226478FB6EBC2FE1A8" pageId="0" pageNumber="231">
<emphasis id="83576FD6B236F3D7AB436A96AE79AB2C" bold="true" pageId="0" pageNumber="231">
<emphasis id="F6988F768774D8A5CA4C7D0A79BFE23E" italics="true" pageId="0" pageNumber="231">Caudalii</emphasis>
.
</emphasis>
Spines of cerci and paracercus unknown.
</paragraph>
</subSubSection>
<subSubSection id="DFB6314C7BEFD9150EFE2FB7C226F02F" pageId="0" pageNumber="231" type="etymology">
<paragraph id="72194CCC5DA028841DE682D6B6200E89" pageId="0" pageNumber="231">Etymology.</paragraph>
<paragraph id="01072454BCBC493276B8BD75C30CA741" pageId="0" pageNumber="231">The specific epithet refers to the type locality in Borneo (Brunei).</paragraph>
</subSubSection>
<subSubSection id="8B280E57D6000F6597377EEFFA26CF02" pageId="0" pageNumber="231" type="distribution">
<paragraph id="C719FA6A36D5E3F8E2722E1FA872048B" pageId="0" pageNumber="231">Distribution.</paragraph>
<paragraph id="998616D4E60DE6F609C9A98D9670055B" pageId="0" pageNumber="231">
Brunei (Fig.
<figureCitation id="B97E424DB2444C1B15A696F15ED5952D" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Arcobaetis gen. nov., distribution." figureDoi="10.3897/zookeys.1168.104844.figure19" httpUri="https://binary.pensoft.net/fig/872221" pageId="0" pageNumber="231">19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6BEFE511A51EEC4F53ED53C14D948AE7" pageId="0" pageNumber="231" type="biological aspects">
<paragraph id="AB7B638180EC9077627A53AC42E75316" pageId="0" pageNumber="231">Biological aspects</paragraph>
<paragraph id="C6269BE59C6EEB3B443CAD2D28345D17" pageId="0" pageNumber="231">
<emphasis id="7676C80E66C59097EFC1AFA99EF4956D" bold="true" pageId="0" pageNumber="231">
(Fig.
<figureCitation id="C20DAAEE5BC37B4A0E4BA1AEB3F5250F" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Arcobaetis bornensis sp. nov., larva a foreleg b hind leg c habitat (photo Kate Baker, University Exeter, UK)." figureDoi="10.3897/zookeys.1168.104844.figure12" httpUri="https://binary.pensoft.net/fig/872214" pageId="0" pageNumber="231">12c</figureCitation>
).
</emphasis>
The species was found at an altitude of 1380 m in slow current of a small and shallow stream.
</paragraph>
</subSubSection>
</treatment>
</document>

72
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<taxonomicName id="4E66AB13C1C9D7A07435976256C21C67" ID-CoL="3GHBF" authority="Desvignes, 1856" authorityName="Desvignes" authorityYear="1856" class="Insecta" family="Ichneumonidae" genus="Glypta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glypta nigrina" order="Hymenoptera" pageId="0" pageNumber="9042" phylum="Arthropoda" rank="species" species="nigrina">Glypta nigrina Desvignes, 1856</taxonomicName>
</paragraph>
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<paragraph id="7F79512AEA0960D3C7B8F93063314DB8" pageId="0" pageNumber="9042">
<taxonomicName id="23C060810607900598E5205A8064A4A2" genus="Ichneumonoidea" lsidName="Ichneumonoidea flavipes" pageId="0" pageNumber="9042" rank="species" species="flavipes">flavipes</taxonomicName>
Desvignes, 1856
</paragraph>
<paragraph id="8C0C6F88F9E3320550B31D7C96CC4565" pageId="0" pageNumber="9042">
<taxonomicName id="820F87949838701AF8B8C9617ADDB0B0" genus="Ichneumonoidea" lsidName="Ichneumonoidea ruficeps" pageId="0" pageNumber="9042" rank="species" species="ruficeps">ruficeps</taxonomicName>
Desvignes, 1856
</paragraph>
<paragraph id="44A416CAA4591D7171F3DC52725933FE" pageId="0" pageNumber="9042">
<taxonomicName id="E53A25C715EA4D3C87981A5CE3D9BD11" genus="Ichneumonoidea" lsidName="Ichneumonoidea fractigena" pageId="0" pageNumber="9042" rank="species" species="fractigena">fractigena</taxonomicName>
Thomson, 1889
</paragraph>
<paragraph id="2AC8A64C7F15C123C52823FF04008599" pageId="0" pageNumber="9042">
<taxonomicName id="398EC431A10E69B822064F914E38007C" genus="Ichneumonoidea" lsidName="Ichneumonoidea obscura" pageId="0" pageNumber="9042" rank="species" species="obscura">obscura</taxonomicName>
Pfankuch, 1924 unavailable
</paragraph>
<paragraph id="6EA36C54343EA42BC53747E2FEBEE98C" pageId="0" pageNumber="9042">
<taxonomicName id="C8039F0462F0EF181D2DAEF2BA8E213F" genus="Ichneumonoidea" lsidName="Ichneumonoidea clypeodentata" pageId="0" pageNumber="9042" rank="species" species="clypeodentata">clypeodentata</taxonomicName>
Bauer, 1959
</paragraph>
<paragraph id="2858AA2BC1FD46A7EF4F7D5DDF7BBC4D" pageId="0" pageNumber="9042">
<taxonomicName id="38D48B2F6CAFDD8C9CB5FE3900BD8DF7" genus="Ichneumonoidea" lsidName="Ichneumonoidea habermani" pageId="0" pageNumber="9042" rank="species" species="habermani">habermani</taxonomicName>
Ozols, 1959
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<subSubSection id="0217D9F901F33CEA37398132E4A64ADE" pageId="0" pageNumber="9042" type="distribution">
<paragraph id="9BC63AE2C4ADF36D70C7DE5091003CDC" pageId="0" pageNumber="9042">Distribution</paragraph>
<paragraph id="ADA2322C0C3CE070EEBEC261C4660360" pageId="0" pageNumber="9042">England, Scotland, Ireland</paragraph>
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[[ worker ]]. - L. 2,6 a 2,9 mill. - Comme la precedente, mais il y a une apparence d' oe il (une ou deux facettes obsoletes au tiers anterieur des cotes de la tete). Fossettes antennaires superficielles. La forme du thorax est la meme, mais la face declive, moins abrupte, du metanotum est distinctement bordee et haut et de cote, subconcave, lisse et luisante. Ecaille plus haute et plus mince, trois fois plus haute qu'epaisse (a peine deux fois plus haute qu'epaisse chez la
<taxonomicName id="2ACCEAF6149F5765A265334E02ED6074" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:231002" class="Insecta" family="Formicidae" genus="Ponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ponera jugata Forel" lsidName-HNS="Ponera jugata Forel" order="Hymenoptera" pageNumber="245" phylum="Arthropoda" rank="variety" species="jugata" variety="glabrata">P. jugata v. glabrata</taxonomicName>
), presque aussi epaisse au sommet qu'a la base. Une pilosite dressee, tres courte, plus abondante, surtout sur l'abdomen; pubescence plus longue et plus forte, luisante. Sculpture un peu plus forte que chez la precedente; couleur un peu plus foncee; antennes d'un brun rougeatre.
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[[ queen ]]. - L. 3,0 mill. - Semblable a l'ouvriere. Ecaille un peu plus epaisse a la base qu'au sommet. Ailes manquent. Les yeux sont grands, ovales, situes on peu en arriere du tiers anterieur de la tete, bien plus grands que chez la
<taxonomicName id="15ECC43F6892519B7558D808CA0741B9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138892" class="Insecta" family="Formicidae" genus="Ponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ponera punctatissima Roger" lsidName-HNS="Ponera punctatissima Roger" order="Hymenoptera" pageNumber="246" phylum="Arthropoda" rank="species" species="punctatissima">P. punctatissima</taxonomicName>
typique. Leur bord posterieur atteint presque le milieu des cotes de la tete et leur bord anterieur en depasse a peine le quart anterieur. La tete est beaucoup plus etroite que chez la
<taxonomicName id="EB95E84BFECDF9477A14AA00B6AE7D89" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:186862" class="Insecta" family="Formicidae" genus="Ponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ponera jugata Forel" lsidName-HNS="Ponera jugata Forel" order="Hymenoptera" pageNumber="246" phylum="Arthropoda" rank="species" species="jugata">P. jugata</taxonomicName>
, beaucoup plus longue que large (seulement un peu plus longue que large chez la
<taxonomicName id="52A2FF01E8C9F2D6B1D5EAEDC8C1683E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:186862" class="Insecta" family="Formicidae" genus="Ponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ponera jugata Forel" lsidName-HNS="Ponera jugata Forel" order="Hymenoptera" pageNumber="246" phylum="Arthropoda" rank="species" species="jugata">P. jugata</taxonomicName>
). La position des yeux, leur taille et la forme de la tete distinguent principalement cette race de la
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[[ queen ]].
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<paragraph id="7EAA052568543563BD48CBC8FB746AD9" pageNumber="246">Moramanga (M. Sikora).</paragraph>
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