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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.63775" ID-GBIF-Dataset="e6957b3b-f228-4d6a-b430-35d41cdb2caa" ID-PMC="PMC8222308" ID-Pensoft-Pub="1313-2970-1044-375" ID-Pensoft-UUID="EBD9E81E5EC55AA491952682787793A4" ID-PubMed="34183881" ID-ZooBank="D50CC77C2E6941FDA9BD395B025C43AE" ModsDocID="1313-2970-1044-375" checkinTime="1623875406767" checkinUser="pensoft" docAuthor="Will, Kipling &amp; Gueorguiev, Borislav" docDate="2021" docId="F0F208486E535F98AA2CCD26AAD48868" docLanguage="en" docName="ZooKeys 1044: 375-425" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.63775" docTitle="Adelopomorpha tethys Will &amp; Guéorguiev 2021, sp. nov." docType="treatment" docUuid="C4A275B5-DC5E-468D-84A4-23115734DF28" docUuidSource="ZooBank" docVersion="4" id="EBD9E81E5EC55AA491952682787793A4" lastPageNumber="375" masterDocId="EBD9E81E5EC55AA491952682787793A4" masterDocTitle="Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)" masterLastPageNumber="425" masterPageNumber="375" pageNumber="375" updateTime="1668150454977" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Will, Kipling</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7056-9011</mods:nameIdentifier>
<mods:affiliation>University of California Berkeley, Essig Museum of Entomology. Berkeley, California, USA</mods:affiliation>
<mods:nameIdentifier type="email">kipwill@berkeley.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:affiliation>National Museum of Natural History, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1044</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1044.63775</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.63775</mods:identifier>
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<treatment ID-GBIF-Taxon="182754021" LSID="urn:lsid:zoobank.org:act:C4A275B5-DC5E-468D-84A4-23115734DF28" httpUri="http://treatment.plazi.org/id/F0F208486E535F98AA2CCD26AAD48868" lastPageNumber="375" pageId="0" pageNumber="375">
<subSubSection pageId="0" pageNumber="375" type="nomenclature">
<paragraph pageId="0" pageNumber="375">
<taxonomicName LSID="http://zoobank.org/C4A275B5-DC5E-468D-84A4-23115734DF28" authority="Will &amp; Guéorguiev, 2021" authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha tethys" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="tethys" status="sp. nov.">Adelopomorpha tethys</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="375">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">Figs 27</figureCitation>
<figureCitation captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">, 37</figureCitation>
<figureCitation captionStart="Figures 52, 53" captionStartId="F15" captionText="Figures 52, 53. Maps of localities of New Caledonian Oodini." figureDoi="10.3897/zookeys.1044.63775.figures52-53" httpUri="https://binary.pensoft.net/fig/554062" pageId="0" pageNumber="375">, 52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="375" type="materials_examined">
<paragraph pageId="0" pageNumber="375">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="375">
<materialsCitation collectingDate="1984-05-11" collectingDateMax="1984-05-13" collectingDateMin="1984-05-11" collectorName="G. Monteith, D. Cook" country="NEW CALEDONIA" elevation="750" location="New Caledonia" specimenCount="♂" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="375">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:F0F208486E535F98AA2CCD26AAD48868:E5D47BE24DF82D4C95AD7F9C0DC1CDF5" country="NEW CALEDONIA" name="New Caledonia">New Caledonia</location>
•
<specimenCount>♂</specimenCount>
; &quot;
<collectingCountry name="New Caledonia">NEW CALEDONIA</collectingCountry>
<normalizedToken originalValue="Mandjélia">Mandjelia</normalizedToken>
, above
<normalizedToken originalValue="Pouébo">Pouebo</normalizedToken>
<collectingDate value="1984-05-11" valueMax="1984-05-13" valueMin="1984-05-11">11-13 May 1984</collectingDate>
, 6-
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.5" unit="m" value="750.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.5" unit="m" value="750.0">750 m</elevation>
</quantity>
<collectorName>G. Monteith</collectorName>
&amp;
<collectorName>D. Cook</collectorName>
&quot;
</materialsCitation>
<typeStatus>Holotype</typeStatus>
pinned, with genitalia in a separate microvial. Source collection QM, deposited MNHN. Type locality as given on label.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="375" type="materials_examined">
<paragraph pageId="0" pageNumber="375">Other material.</paragraph>
<paragraph pageId="0" pageNumber="375">
<materialsCitation collectingDate="2000-11-22" collectorName="G. B. Monteith. Pyrethrum" country="New Caledonia" elevation="500" location="Sx" longitude="166.01666" municipality="Mt. Koghis" specimenCount="♂">
<collectingCountry name="New Caledonia">New Caledonia</collectingCountry>
•
<specimenCount>♂</specimenCount>
; &quot;21°11'
<location LSID="urn:lsid:plazi:treatment:F0F208486E535F98AA2CCD26AAD48868:769C6D97ACCB812E310C76F15D1E7FB7" country="New Caledonia" longitude="166.01666" municipality="Mt. Koghis" name="Sx">Sx</location>
<geoCoordinate degrees="166" direction="east" minutes="01" orientation="longitude" precision="925" value="166.01666">166°01'E</geoCoordinate>
:
<collectingMunicipality>Mt. Koghis</collectingMunicipality>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<collectingDate value="2000-11-22">22 Nov 2000</collectingDate>
<collectorName>G.B. Monteith. Pyrethrum</collectorName>
trunks &amp; logs. 9931&quot; [ZSM]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="375" type="diagnosis">
<paragraph pageId="0" pageNumber="375">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="375">
Similar to
<taxonomicName lsidName="C. microps" pageId="0" pageNumber="375" rank="species" species="microps">
<emphasis italics="true" pageId="0" pageNumber="375">C. microps</emphasis>
</taxonomicName>
in size and the lack of impressed striae 1-5, but with much larger, prominent eyes and male having protarsomeres 1-3 not expanded. Both
<taxonomicName authorityName="Heller" authorityYear="1916" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha glabra" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="glabra">
<emphasis italics="true" pageId="0" pageNumber="375">Adelopomorpha glabra</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="A. tuberculata" pageId="0" pageNumber="375" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="375">A. tuberculata</emphasis>
</taxonomicName>
also lack impressed striae, but are smaller, and in
<taxonomicName lsidName="A. tethys" pageId="0" pageNumber="375" rank="species" species="tethys">
<emphasis italics="true" pageId="0" pageNumber="375">A. tethys</emphasis>
</taxonomicName>
the male has a pair of setae on ventrite 6 that is lacking in
<taxonomicName lsidName="A. glabra" pageId="0" pageNumber="375" rank="species" species="glabra">
<emphasis italics="true" pageId="0" pageNumber="375">A. glabra</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="A. tuberculata" pageId="0" pageNumber="375" rank="species" species="tuberculata">
<emphasis italics="true" pageId="0" pageNumber="375">A. tuberculata</emphasis>
</taxonomicName>
. The aedeagus is also decisively different (Fig.
<figureCitation captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">27</figureCitation>
), with a relatively short and slightly acute tip.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="375" type="description">
<paragraph pageId="0" pageNumber="375">Description</paragraph>
<paragraph pageId="0" pageNumber="375">
<emphasis bold="true" pageId="0" pageNumber="375">(based on holotype)</emphasis>
.
<emphasis bold="true" pageId="0" pageNumber="375">
<emphasis italics="true" pageId="0" pageNumber="375">Habitus</emphasis>
.
</emphasis>
Small sized, BL: 6.75 mm and BW: 3.25 mm.
<emphasis bold="true" pageId="0" pageNumber="375">
<emphasis italics="true" pageId="0" pageNumber="375">Color and luster</emphasis>
.
</emphasis>
Head dark reddish black; pronotum, elytra and ventral surface of body black, pronotum marginally diaphanous reddish black, extreme apical edge of ventrite 6 deep reddish black medially; legs deep reddish brown; antennae and palpi brown. Integument moderately dull, without spectral iridescence except ventrally.
<emphasis bold="true" pageId="0" pageNumber="375">
<emphasis italics="true" pageId="0" pageNumber="375">Microsculpture and punctation</emphasis>
.
</emphasis>
Dorsal surface of head, pronotum, and elytra with prominent isodiametric meshes; ventral surface with scarcely-visible sculpticells or sculpticells mostly not apparent. Head lacking macropunctation or wrinkles on vertex; head, pronotum and elytra with scattered micropunctation, only on elytra do micropunctures and sculpticells form some irregular rosettes (sensu
<bibRefCitation author="Spence, JR" journalOrPublisher="The Coleopterists' Bulletin" pageId="0" pageNumber="375" pagination="567 - 580" publicationUrl="https://www.jstor.org/stable/4007993" refId="B39" refString="Spence, JR, 1983. Taxonomic status, relationships, and biogeography of Anatrichis LeConte and Oodinus Motschulsky (Carabidae: Oodini). The Coleopterists' Bulletin 36 (4): 567 - 580, https://www.jstor.org/stable/4007993" title="Taxonomic status, relationships, and biogeography of Anatrichis LeConte and Oodinus Motschulsky (Carabidae: Oodini)." url="https://www.jstor.org/stable/4007993" volume="36" year="1983">Spence 1983</bibRefCitation>
); all abdominal ventrites very shallowly wrinkled to nearly smooth.
<emphasis bold="true" pageId="0" pageNumber="375">
<emphasis italics="true" pageId="0" pageNumber="375">Head</emphasis>
.
</emphasis>
Approximately 1/3 as wide as pronotum (Suppl. material 2: Table S1). Eye size moderate, eye somewhat protruded, EyW/HW: 1.35. Frontoclypeal sutures not evident. Mentum without paramedial border.
<emphasis bold="true" pageId="0" pageNumber="375">
<emphasis italics="true" pageId="0" pageNumber="375">Thorax</emphasis>
.
</emphasis>
Pronotum 1 2/3 wider than long (PW/PL: 1.67), with sides evenly, very shallowly rounded from anterior angles to posterior ones; maximal width in posterior 1/5; width at apex 2 2/3
<normalizedToken originalValue="×">x</normalizedToken>
less than at widest point (PW/PA: 2.32). Posterior angles obtuse, broadly rounded. Mesosternum without medial tubercle. Metepisternum 1 1/3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, with lateral margin slightly convex, coadunation with epipleuron along entire length.
<emphasis bold="true" pageId="0" pageNumber="375">
<emphasis italics="true" pageId="0" pageNumber="375">Elytra</emphasis>
.
</emphasis>
Broad, nearly as wide as long (EL/EW: 1.06). Basal margin distinct, forming small, rounded callosity at shoulder, ended medially at level of parascutellar punctures, joined to parascutellar puncture fovea by a short, shallow striole. Apical sinuation scarcely evident, very shallow.
<emphasis bold="true" pageId="0" pageNumber="375">
<emphasis italics="true" pageId="0" pageNumber="375">Female genitalia</emphasis>
.
</emphasis>
Female specimens unknown.
<emphasis bold="true" pageId="0" pageNumber="375">
<emphasis italics="true" pageId="0" pageNumber="375">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view long, almost straight (Fig.
<figureCitation captionStart="Figures 32–40" captionStartId="F8" captionText="Figures 32 - 40. Aedeagus, left lateral view of 32 Coptocarpus microps sp. nov. 33 C. erwini sp. nov. 34 C. cyllodinus (Fauvel) 35 C. amieuensis sp. nov. 36 C. magnus sp. nov. 37 Adelopomorpha tethys sp. nov. 38 A. tuberculata sp. nov. 39 A. glabra Heller 40 right lateral view of tip with arrow indicating the dorsal tooth of C. magnus." figureDoi="10.3897/zookeys.1044.63775.figures32-40" httpUri="https://binary.pensoft.net/fig/554055" pageId="0" pageNumber="375">37</figureCitation>
); lobe in dorsal view with apical lamella moderately long, rounded, triangular and slightly oriented to right (Fig.
<figureCitation captionStart="Figures 22–31" captionStartId="F7" captionText="Figures 22 - 31. Aedeagus, dorsal view of 22 Coptocarpus microps sp. nov. 23 C. erwini sp. nov. 24 C. cyllodinus (Fauvel) 25 C. amieuensis sp. nov. 26 C. magnus sp. nov. 27 Adelopomorpha tethys sp. nov. 28 A. tuberculata sp. nov. 29 A. glabra Heller 30 Adelopomorpha glabra group male from Mt. Koghis 31 Adelopomorpha glabra group male from Gelima." figureDoi="10.3897/zookeys.1044.63775.figures22-31" httpUri="https://binary.pensoft.net/fig/554054" pageId="0" pageNumber="375">27</figureCitation>
); sclerotized portion of endophallus visible in repose, with two sclerites.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="375" type="etymology">
<paragraph pageId="0" pageNumber="375">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="375">
The specific epithet
<taxonomicName authorityName="Will &amp; Guéorguiev" authorityYear="2021" class="Insecta" family="Carabidae" genus="Adelopomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Adelopomorpha tethys" order="Coleoptera" pageId="0" pageNumber="375" phylum="Arthropoda" rank="species" species="tethys">
<emphasis italics="true" pageId="0" pageNumber="375">Adelopomorpha tethys</emphasis>
</taxonomicName>
is treated as a noun in the genitive case and is from the mythological Greek Titan goddess Tethys, said to be the primal source of water to nourish Earth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="375" type="notes">
<paragraph pageId="0" pageNumber="375">Notes.</paragraph>
<paragraph pageId="0" pageNumber="375">The male specimen from Mt. Koghis (&quot;Additional specimen examined&quot; above) differs from the holotype in somewhat larger eyes (EyW/HW&gt; 1.40), mentum with distinct paramedial border, pronotum with anterior submarginal sulcus present laterally each side and absent in medial 1/4, mesosternum with transversely elongate medial tubercle, and somewhat different structure of the median lobe of aedeagus in lateral view. Given the significant distance (~ 280 km) between the collecting localities and the several morphological differences between them, it is possible that the population from Mt. Koghis represents a separate species, and we exclude this specimen from being a paratype. However, given that these are slight and subtle differences in otherwise very similar looking individuals, it is also possible that this specimen is a conspecific variant. Additional specimens and further study are needed to settle this question.</paragraph>
<caption doi="10.3897/zookeys.1044.63775.figures52-53" httpUri="https://binary.pensoft.net/fig/554062" pageId="0" pageNumber="375" start="Figures 52, 53" startId="F15">
<paragraph pageId="0" pageNumber="375">
<emphasis bold="true" pageId="0" pageNumber="375">Figures 52, 53.</emphasis>
Maps of localities of New Caledonian
<taxonomicName authorityName="LaFerte-Senectere" authorityYear="1851" lsidName="" pageId="0" pageNumber="375" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<treatment id="F0F2AC32CDAE2C8A0BAB2609BA5F1D0D" ID-DOI="http://doi.org/10.5281/zenodo.6290511" ID-GBIF-Taxon="100129037" ID-Zenodo-Dep="6290511" LSID="urn:lsid:plazi:treatment:F0F2AC32CDAE2C8A0BAB2609BA5F1D0D" httpUri="http://treatment.plazi.org/id/F0F2AC32CDAE2C8A0BAB2609BA5F1D0D" lastPageNumber="4" pageNumber="4">
<subSubSection id="4A86955457B8AEA03B6BFD66510B19DA" type="nomenclature">
<paragraph id="ED00F7FC59E852B525B0FED6A0E45FD4" pageNumber="4">
<taxonomicName id="9DC9276D84D0958E670E47D3392F60E9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:231981" authority="Forel (Atopogyne)" authorityName="Forel (Atopogyne)" class="Insecta" family="Formicidae" genus="Cremastogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Uploaded - Cremastogaster auberti subsp. jehovae" lsidName-HNS="Cremastogaster auberti subsp. jehovae" order="Hymenoptera" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="auberti" subSpecies="jehovae">Cremastogaster Auberti Emery subsp. Jehovae</taxonomicName>
Forel
</paragraph>
</subSubSection>
<subSubSection id="F89067C02B7BB8803520ADCE52383C35" type="materials_examined">
<paragraph id="D9087969B7075D8A2B1FA1FDFBAC3312" pageNumber="4">[[ worker ]]. - Jerusalem (Schmitz).</paragraph>
</subSubSection>
</treatment>
</document>