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<document id="69861B1725EE91F336E387A2C6790BDE" ID-DOI="10.5281/zenodo.12520315" ID-ISSN="1407-8619" ID-Zenodo-Dep="12520315" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719250516759" checkinUser="felipe" docAuthor="Vitali, Francesco" docDate="2017" docId="AD3087C2482BFF807C4AFCD43BDF654A" docLanguage="en" docName="BalticJColeopterol.17.2.147-160.pdf" docOrigin="Baltic Journal of Coleopterology 17 (2)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.2:BalticJColeopterol.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeopterol.2012-2021.journal_article" docStyleVersion="2" docTitle="Tillomorphites spinipes Vitali, 2017, n. sp." docType="treatment" docVersion="2" lastPageNumber="151" masterDocId="5109FFBA482FFF847D64FFF4384E6072" masterDocTitle="Two new fossil species of Tillomorphites Vitali (Coleoptera: Cerambycidae) and remarks on the morphological evolution, mimicry, biogeography and phylogeny of the tribe Tillomorphini" masterLastPageNumber="160" masterPageNumber="147" pageNumber="151" updateTime="1719251310870" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="D48BF85036A1748BCB3A3DA9B9252C55">Two new fossil species of Tillomorphites Vitali (Coleoptera: Cerambycidae) and remarks on the morphological evolution, mimicry, biogeography and phylogeny of the tribe Tillomorphini</mods:title>
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<mods:title id="705E516717AE419846CA831FD3A4AC8B">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="C358C9AAE5D63A16E27D11B20FA4553C">2017</mods:date>
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<heading id="7E6E81B8482BFF807C4AFCD43A32634A" bold="true" box="[302,636,800,824]" fontSize="10" level="2" pageId="4" pageNumber="151" reason="4">
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<taxonomicName id="E2994D57482BFF807C4AFCD43A7A634A" authorityName="Vitali" authorityYear="2017" box="[302,564,800,824]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="151" phylum="Arthropoda" rank="species" species="spinipes" status="sp. nov.">
<emphasis id="17EDEAC6482BFF807C4AFCD43A7A634A" bold="true" box="[302,564,800,824]" italics="true" pageId="4" pageNumber="151">Tillomorphites spinipes</emphasis>
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<paragraph id="252636D4482BFF807C4AFCB439F5632A" blockId="4.[302,636,800,856]" box="[302,443,832,856]" pageId="4" pageNumber="151">
(
<figureCitation id="BDA22A51482BFF807C52FCB439FD632A" box="[310,435,832,856]" captionStart="Fig" captionStartId="5.[227,263,620,644]" captionTargetBox="[227,775,189,600]" captionTargetId="figure-9@5.[227,775,189,600]" captionTargetPageId="5" captionText="Fig. 3a Tillomorphites spinipes n. sp., Holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12520339" httpUri="https://zenodo.org/record/12520339/files/figure.png" pageId="4" pageNumber="151">Figs. 3a - b</figureCitation>
)
</paragraph>
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<subSubSection id="6D83655F482BFF807C4AFC743A5364CA" pageId="4" pageNumber="151" type="materials_examined">
<paragraph id="252636D4482BFF807C4AFC7439D263EA" blockId="4.[302,813,896,984]" box="[302,412,896,920]" pageId="4" pageNumber="151">
<emphasis id="17EDEAC6482BFF807C4AFC7439D263EA" bold="true" box="[302,412,896,920]" pageId="4" pageNumber="151">
<typeStatus id="FA228876482BFF807C4AFC7439D263EA" box="[302,412,896,920]" pageId="4" pageNumber="151" type="holotype">Holotype</typeStatus>
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<paragraph id="252636D4482BFF807C4AFC5439E363AA" blockId="4.[302,813,896,984]" pageId="4" pageNumber="151">Baltic amber, ex coll. M. Veta, author’s coll. FS56BS30.</paragraph>
<paragraph id="252636D4482BFF807C4AFBF43A5364CA" blockId="4.[302,813,1024,1208]" pageId="4" pageNumber="151">
The type is included in a 23 x 13 x
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rectangular piece of yellow amber with no observable syninclusions. The beetle is mostly covered by turbidity, especially on the head and along the right side; moreover, it lacks the right antennomeres VII-XI.
</paragraph>
</subSubSection>
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<paragraph id="252636D4482BFF807C4AFB143A7C648A" blockId="4.[302,813,1248,1496]" box="[302,562,1248,1272]" pageId="4" pageNumber="151">
<emphasis id="17EDEAC6482BFF807C4AFB143A7C648A" bold="true" box="[302,562,1248,1272]" pageId="4" pageNumber="151">Differential diagnosis</emphasis>
</paragraph>
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Though scarcely visible, this species is well characterised by the presence of an apical spine on the antennomere V and of strong erect setae on the meso- and metatibiae. Moreover, it differs from
<taxonomicName id="E2994D57482BFF807CDCFA743A7E65EA" authorityName="Vitali" authorityYear="2011" box="[440,560,1408,1432]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="151" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="17EDEAC6482BFF807CDCFA743A7E65EA" box="[440,560,1408,1432]" italics="true" pageId="4" pageNumber="151">T. robustus</emphasis>
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in the larger body size and from
<taxonomicName id="E2994D57482BFF807CF8FA5439B365CA" authorityName="Vitali" authorityYear="2017" box="[412,509,1440,1464]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="151" phylum="Arthropoda" rank="species" species="otiliae" status="n. sp.">
<emphasis id="17EDEAC6482BFF807CF8FA5439B365CA" box="[412,509,1440,1464]" italics="true" pageId="4" pageNumber="151">T. otiliae</emphasis>
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in the elytra and the antennae completely covered with erect setae.
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<emphasis id="17EDEAC6482BFF807C4AF9F439F2666A" bold="true" box="[302,444,1536,1560]" pageId="4" pageNumber="151">Description</emphasis>
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Sex unknown, Body length ~
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.
</paragraph>
<paragraph id="252636D4482BFF807C4AF9B43C5A616A" blockId="4.[302,813,1536,1720]" lastBlockId="4.[850,1361,192,1208]" pageId="4" pageNumber="151">Eyes reniform. Antennae 11-segmented, shorter than body, covered by some short sub-erect setae; antennomeres III - V with a long fine spine at the apex; antennomeres V - X externally dentate at the apex; antennomere proportions according to the formula:?:?:?:?:?: 1.2: 1.1: 0.9: 1.0: 0.8: 0.5.</paragraph>
<paragraph id="252636D4482BFF807E36FED43C0E61CA" blockId="4.[850,1361,192,1208]" pageId="4" pageNumber="151">Pronotum longer than wide, obtusely angulated before the middle and strongly inflated before the base; regularly convergent anteriorly; disc seemingly uneven, smooth and glabrous. Scutellum transverse.</paragraph>
<paragraph id="252636D4482BFF807E36FE343BDF62CA" blockId="4.[850,1361,192,1208]" pageId="4" pageNumber="151">Elytra parallel-sided, feebly depressed on the middle of the disc, about 2.7 times as long as wide at humeri; base feebly convex, concave in correspondence of the posterior angles of the prothorax; humeri rounded; lateral margins furrowed; apex largely rounded; disc smooth, throughout covered with some semi-erect long setae.</paragraph>
<paragraph id="252636D4482BFF807E36FD343CAF63AA" blockId="4.[850,1361,192,1208]" pageId="4" pageNumber="151">Legs relatively short and robust, apparently smooth, femora clavate; tibiae linear, covered with 3 - 4 conspicuous sub-erect black setae, as long as those of antennae or elytra, on the outer side and some short semi-recumbent setae on the inner side; metatarsus about one-half as long as metatibia; metatarsomere I as long as II and III together or as onychium; metatarsomeres II and III transverse.</paragraph>
<paragraph id="252636D4482BFF807E36FC143D5F64CA" blockId="4.[850,1361,192,1208]" pageId="4" pageNumber="151">Prosternum seemingly smooth; intercoxal process fairly narrow, triangular; procoxae globose, procoxal cavities rounded, externally rounded; visible urosternites (I-V) progressively shortened to the apex, very finely and densely punctured, covered with a fine dense recumbent pubescence and some sparse erect setae.</paragraph>
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<subSubSection id="6D83655F482BFF807E36FB143BDF654A" pageId="4" pageNumber="151" type="etymology">
<paragraph id="252636D4482BFF807E36FB143B9A648A" blockId="4.[850,1360,1248,1336]" box="[850,980,1248,1272]" pageId="4" pageNumber="151">
<emphasis id="17EDEAC6482BFF807E36FB143B9A648A" bold="true" box="[850,980,1248,1272]" pageId="4" pageNumber="151">Etymology</emphasis>
</paragraph>
<paragraph id="252636D4482BFF807E36FAF43BDF654A" blockId="4.[850,1360,1248,1336]" pageId="4" pageNumber="151">
From the Latin “
<emphasis id="17EDEAC6482BFF807970FAF43C39656A" box="[1044,1143,1280,1304]" italics="true" pageId="4" pageNumber="151">spini-pes</emphasis>
”, i.e. “with spined legs”.
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</treatment>
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<document id="48819933CF376EEE1552A30077CBA2D3" ID-DOI="10.5281/zenodo.12520315" ID-ISSN="1407-8619" ID-Zenodo-Dep="12520315" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719250516759" checkinUser="felipe" docAuthor="Vitali, Francesco" docDate="2017" docId="AD3087C2482EFF807E62FE943AA262AA" docLanguage="en" docName="BalticJColeopterol.17.2.147-160.pdf" docOrigin="Baltic Journal of Coleopterology 17 (2)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.2:BalticJColeopterol.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeopterol.2012-2021.journal_article" docStyleVersion="2" docTitle="Tillomorphites otiliae Vitali, 2017, n. sp." docType="treatment" docVersion="2" lastPageNumber="151" masterDocId="5109FFBA482FFF847D64FFF4384E6072" masterDocTitle="Two new fossil species of Tillomorphites Vitali (Coleoptera: Cerambycidae) and remarks on the morphological evolution, mimicry, biogeography and phylogeny of the tribe Tillomorphini" masterLastPageNumber="160" masterPageNumber="147" pageNumber="148" updateTime="1719251310870" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="7447509279D1B50169D0BA90FE912BA2">Two new fossil species of Tillomorphites Vitali (Coleoptera: Cerambycidae) and remarks on the morphological evolution, mimicry, biogeography and phylogeny of the tribe Tillomorphini</mods:title>
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<emphasis id="17EDEAC6482EFF857E62FE943C70610A" bold="true" box="[774,1086,352,376]" pageId="1" pageNumber="148">
<taxonomicName id="E2994D57482EFF857E62FE943BB9610A" authorityName="Vitali" authorityYear="2017" box="[774,1015,352,376]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="otiliae" status="sp. nov.">
<emphasis id="17EDEAC6482EFF857E62FE943BB9610A" bold="true" box="[774,1015,352,376]" italics="true" pageId="1" pageNumber="148">Tillomorphites otiliae</emphasis>
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<taxonomicNameLabel id="0CDE57BD482EFF857964FE943C70610A" box="[1024,1086,352,376]" pageId="1" pageNumber="148" rank="species">n. sp.</taxonomicNameLabel>
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(
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)
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</emphasis>
</paragraph>
<paragraph id="252636D4482EFF857E62FE143BF3626A" blockId="1.[774,1285,448,536]" pageId="1" pageNumber="148">Female, Baltic amber, ex coll. M. Veta, author’s coll. FS65BS35.</paragraph>
<paragraph id="252636D4482EFF857E62FDB43CC8630A" blockId="1.[774,1286,576,888]" pageId="1" pageNumber="148">
The type is included in a rectangular-trapezoidal, 20 x 15 x
<quantity id="E2619B31482EFF857E19FD943B8E620A" box="[893,960,608,632]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="148" unit="mm" value="5.0">5 mm</quantity>
, oval piece of yellow-orange amber containing the following syninclusions: a Trichoptera, two
<taxonomicName id="E2994D57482EFF857E97FD543C0562CA" box="[1011,1099,672,696]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
<taxonomicName id="E2994D57482EFF857933FD543C8B62CA" authorityName="Billberg" authorityYear="1820" box="[1111,1221,672,696]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
(one present as head), an exuvium of a possible neanic stage of Acridoidea (only four legs present), many stellate hairs and wood dust. The beetle is partially covered by turbidity along the right side and lacks the left antennomeres V- XI; the ovipositor is fully exposed.
</paragraph>
<paragraph id="252636D4482EFF857E62FC543B3F63CA" blockId="1.[774,1285,928,1016]" box="[774,881,928,952]" pageId="1" pageNumber="148">
<emphasis id="17EDEAC6482EFF857E62FC543B3F63CA" bold="true" box="[774,881,928,952]" pageId="1" pageNumber="148">
<typeStatus id="FA228876482EFF857E62FC543B3F63CA" box="[774,881,928,952]" pageId="1" pageNumber="148" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="252636D4482EFF857E62FC343BF3638A" blockId="1.[774,1285,928,1016]" pageId="1" pageNumber="148">Female, Baltic amber, ex coll. M. Veta, author’s coll. FS70BS39.</paragraph>
<paragraph id="252636D4482EFF857E62FBD43B89648A" blockId="1.[774,1286,1056,1272]" pageId="1" pageNumber="148">
The type is included in a 25 x 11 x
<quantity id="E2619B31482EFF8579A5FBD43D4A644A" box="[1217,1284,1056,1080]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="148" unit="mm" value="4.0">4 mm</quantity>
parallelogram-shaped piece of yellow-orange amber with many stellate hairs as syninclusions. The beetle is partially covered by turbidity on the pronotum and the mouth pieces; the abdomen looks inflated due to the possible presence of eggs.
</paragraph>
</subSubSection>
<subSubSection id="6D83655F482EFF857E62FAD43D4B666A" pageId="1" pageNumber="148" type="diagnosis">
<paragraph id="252636D4482EFF857E62FAD43C45654A" blockId="1.[774,1286,1312,1560]" box="[774,1035,1312,1336]" pageId="1" pageNumber="148">
<emphasis id="17EDEAC6482EFF857E62FAD43C45654A" bold="true" box="[774,1035,1312,1336]" pageId="1" pageNumber="148">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="252636D4482EFF857E62FAB43D4B666A" blockId="1.[774,1286,1312,1560]" pageId="1" pageNumber="148">
This species differs from
<taxonomicName id="E2994D57482EFF857950FAB43CFC652A" authorityName="Vitali" authorityYear="2011" box="[1076,1202,1344,1368]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="17EDEAC6482EFF857950FAB43CFC652A" box="[1076,1202,1344,1368]" italics="true" pageId="1" pageNumber="148">T. robustus</emphasis>
</taxonomicName>
and
<taxonomicName id="E2994D57482EFF857995FAB43B2C650A" authorityName="Vitali" authorityYear="2017" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="spinipes" status="n. sp.">
<emphasis id="17EDEAC6482EFF857995FAB43B2C650A" italics="true" pageId="1" pageNumber="148">T. spinipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CDE57BD482EFF857E0AFA943BE3650A" box="[878,941,1376,1400]" pageId="1" pageNumber="148" rank="species">n. sp.</taxonomicNameLabel>
in the absence of short erect setae on the half of each antennomere and in the more elongated and nearly smooth elytra. Moreover, it differs from
<taxonomicName id="E2994D57482EFF857953FA343CF965AA" authorityName="Vitali" authorityYear="2011" box="[1079,1207,1472,1496]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="17EDEAC6482EFF857953FA343CF965AA" box="[1079,1207,1472,1496]" italics="true" pageId="1" pageNumber="148">T. robustus</emphasis>
</taxonomicName>
in the different pronotal shape and from
<taxonomicName id="E2994D57482EFF8579E9FA143D4B658A" authorityName="Vitali" authorityYear="2017" box="[1165,1285,1504,1528]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="17EDEAC6482EFF8579E9FA143D4B658A" box="[1165,1285,1504,1528]" italics="true" pageId="1" pageNumber="148">T. spinipes</emphasis>
</taxonomicName>
in the simply recumbent pubescence of the legs.
</paragraph>
</subSubSection>
<subSubSection id="6D83655F482EFF807E62F9B43ACC626A" lastPageId="4" lastPageNumber="151" pageId="1" pageNumber="148" type="description">
<paragraph id="252636D4482EFF857E62F9B43BDA662A" blockId="1.[774,1285,1600,1720]" box="[774,916,1600,1624]" pageId="1" pageNumber="148">
<emphasis id="17EDEAC6482EFF857E62F9B43BDA662A" bold="true" box="[774,916,1600,1624]" pageId="1" pageNumber="148">Description</emphasis>
</paragraph>
<paragraph id="252636D4482EFF867E62F9943C4F608A" blockId="1.[774,1285,1600,1720]" lastBlockId="2.[850,1360,192,248]" lastPageId="2" lastPageNumber="149" pageId="1" pageNumber="148">
Female, Body length 4.3 (
<typeStatus id="FA228876482EFF857947F9943CCB660A" box="[1059,1157,1632,1656]" pageId="1" pageNumber="148" type="holotype">holotype</typeStatus>
) to
<quantity id="E2619B31482EFF8579D5F9943D4B660A" box="[1201,1285,1632,1656]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="148" unit="mm" value="4.9">4.9 mm</quantity>
(
<typeStatus id="FA228876482EFF857E74F9743B2066EA" box="[784,878,1664,1688]" pageId="1" pageNumber="148" type="paratype">paratype</typeStatus>
). Body throughout black except for the elytra, showing a pre-median transversal reddish band (
<typeStatus id="FA228876482DFF867E8CFF343C0460AA" box="[1000,1098,192,216]" pageId="2" pageNumber="149" type="holotype">holotype</typeStatus>
) and a similar pre-apical band (
<typeStatus id="FA228876482DFF867EF7FF143BB8608A" box="[915,1014,224,248]" pageId="2" pageNumber="149" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="252636D4482DFF867E36FED43BA4638A" blockId="2.[850,1361,288,1016]" pageId="2" pageNumber="149">Head short; forehead oblique, smooth; antennal tubercles widely separated and scarcely elevated; interantennal furrow fine; eyes large, finely faceted, very strongly reniform, scarcely prominent; upper eye-lobes well developed; under eye-lobes nearly occupying all space of genae. Palpomeres sub-equal; last palpomere obovate, longer than the previous one. Antennae 11-segmented, shorter than body, glabrous; antennomeres III - IV with a very fine spine at the apex; antennomeres V - X externally dentate at the apex; scape short, bowed; pedicel transverse one-ninth as long as scape; antennomere III club-shaped, one-half as long as scape; antennomeres IV club-shaped, about one-third longer than III; antennomere V even longer; antennomeres VI and VII equal, hardly longer than V; antennomeres VIII and IX equal, hardly shorter than V, antennomere X even shorter, antennomere XI one-third as long as scape (antennomere proportions according to the formula: 1.8: 0.2: 9.0: 1.3 1.6: 1.7: 1.7: 1.5: 1.5: 1.4: 0.6).</paragraph>
<caption id="71E6665C482DFF867C4AFD4939D36287" ID-DOI="http://doi.org/10.5281/zenodo.12520318" ID-Zenodo-Dep="12520318" httpUri="https://zenodo.org/record/12520318/files/figure.png" pageId="2" pageNumber="149" startId="2.[302,338,701,725]" targetBox="[302,812,189,669]" targetPageId="2" targetType="figure">
<paragraph id="252636D4482DFF867C4AFD4939D36287" blockId="2.[302,812,701,757]" pageId="2" pageNumber="149">
Fig. 1a.
<taxonomicName id="E2994D57482DFF867CEBFD493B6662A7" authority="Vitali, 2011" authorityName="Vitali" authorityYear="2011" box="[399,808,701,725]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="17EDEAC6482DFF867CEBFD493AD962A7" box="[399,663,701,725]" italics="true" pageId="2" pageNumber="149">Tillomorphites robustus</emphasis>
<bibRefCitation id="41084B25482DFF867FC5FD493B6662A7" author="Vitali F." box="[673,808,701,725]" pageId="2" pageNumber="149" pagination="1 - 34" refId="ref6734" refString="Vitali F. 2011. Six new fossil Cerambycids included in Baltic and Saxon amber (Coleoptera Cerambycidae). Entomapeiron (P. S.) 4 (1): 1 - 34." type="journal article" year="2011">Vitali, 2011</bibRefCitation>
</taxonomicName>
, Holotype.
</paragraph>
</caption>
<paragraph id="252636D4482DFF867E36FBD43C7A654A" blockId="2.[850,1361,1056,1336]" pageId="2" pageNumber="149">Pronotum longer than wide, ob-ovate, constricted before the base; apex one-fifth wider than base, anteriorly feebly convex, very finely furrowed along its margin; base bilobed, posteriorly convex in the middle, two-fifth narrower than elytral base, finely furrowed along its margin; disc convex, smooth and glabrous. Scutellum small, not guessable since partially covered by turbidity.</paragraph>
<paragraph id="252636D4482DFF867E36FA943BAD664A" blockId="2.[850,1361,1376,1592]" pageId="2" pageNumber="149">Elytra apically inflated, moderately convex, 2.7 times as long as wide at humeri; base feebly convex, concave in correspondence of the basal angles of the pronotum; humeri rounded; lateral margins furrowed; apex largely rounded; disc smooth, covered with some long semi-erect setae at base.</paragraph>
<paragraph id="252636D4482DFF867E36F9943D1E66AA" blockId="2.[850,1361,1632,1752]" pageId="2" pageNumber="149">Legs relatively short and robust, pubescent, femora clavate; tibiae linear; tarsi short; metatarsus one-half as long as metatibia; metatarsomere I as long as II and III together or</paragraph>
<caption id="71E6665C482DFF867C4AF9693A2A66A7" ID-DOI="http://doi.org/10.5281/zenodo.12520321" ID-Zenodo-Dep="12520321" httpUri="https://zenodo.org/record/12520321/files/figure.png" pageId="2" pageNumber="149" startId="2.[302,339,1693,1717]" targetBox="[324,825,870,1611]" targetPageId="2" targetType="figure">
<paragraph id="252636D4482DFF867C4AF9693A2A66A7" blockId="2.[302,813,1693,1749]" pageId="2" pageNumber="149">
Fig1b.
<taxonomicName id="E2994D57482DFF867CE6F9693B6B66C7" authority="Vitali, 2011" authorityName="Vitali" authorityYear="2011" box="[386,805,1693,1717]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="17EDEAC6482DFF867CE6F9693ADC66C7" box="[386,658,1693,1717]" italics="true" pageId="2" pageNumber="149">Tillomorphites robustus</emphasis>
<bibRefCitation id="41084B25482DFF867FF9F9693B6B66C7" author="Vitali F." box="[669,805,1693,1717]" pageId="2" pageNumber="149" pagination="1 - 34" refId="ref6734" refString="Vitali F. 2011. Six new fossil Cerambycids included in Baltic and Saxon amber (Coleoptera Cerambycidae). Entomapeiron (P. S.) 4 (1): 1 - 34." type="journal article" year="2011">Vitali, 2011</bibRefCitation>
</taxonomicName>
, habitus, new reconstruction.
</paragraph>
</caption>
<paragraph id="252636D4482CFF877FB5FF663B5860D7" blockId="3.[721,790,146,165]" box="[721,790,146,165]" pageId="3" pageNumber="150">
<emphasis id="17EDEAC6482CFF877FB5FF663B5860D7" box="[721,790,146,165]" italics="true" pageId="3" pageNumber="150">Vitali F.</emphasis>
</paragraph>
<caption id="71E6665C482CFF877D87FDB03AAF622E" ID-DOI="http://doi.org/10.5281/zenodo.12520323" ID-Zenodo-Dep="12520323" box="[227,737,580,604]" httpUri="https://zenodo.org/record/12520323/files/figure.png" pageId="3" pageNumber="150" startId="3.[227,266,580,604]" targetBox="[227,775,189,558]" targetPageId="3" targetType="figure">
<paragraph id="252636D4482CFF877D87FDB03AAF622E" blockId="3.[227,737,580,604]" box="[227,737,580,604]" pageId="3" pageNumber="150">
Fig. 2a.
<taxonomicName id="E2994D57482CFF877C58FDB03A68622E" authorityName="Vitali" authorityYear="2017" box="[316,550,580,604]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="species" species="otiliae" status="sp. nov.">
<emphasis id="17EDEAC6482CFF877C58FDB03A68622E" box="[316,550,580,604]" italics="true" pageId="3" pageNumber="150">Tillomorphites otiliae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CDE57BD482CFF877F49FDB03A27622E" box="[557,617,580,604]" pageId="3" pageNumber="150" rank="species">n. sp.</taxonomicNameLabel>
, Holotype.
</paragraph>
</caption>
<caption id="71E6665C482CFF877E7DFC5D3CC86393" ID-DOI="http://doi.org/10.5281/zenodo.12520327" ID-Zenodo-Dep="12520327" httpUri="https://zenodo.org/record/12520327/files/figure.png" pageId="3" pageNumber="150" startId="3.[793,832,937,961]" targetBox="[853,1272,208,904]" targetPageId="3" targetType="figure">
<paragraph id="252636D4482CFF877E7DFC5D3CC86393" blockId="3.[793,1285,937,993]" pageId="3" pageNumber="150">
Fig. 2 d.
<taxonomicName id="E2994D57482CFF877EEBFC5D3CED63B3" authorityName="Vitali" authorityYear="2017" box="[911,1187,937,961]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="species" species="otiliae" status="sp. nov.">
<emphasis id="17EDEAC6482CFF877EEBFC5D3CED63B3" box="[911,1187,937,961]" italics="true" pageId="3" pageNumber="150">Tillomorphites otiliae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CDE57BD482CFF8779D0FC5D3CB363B3" box="[1204,1277,937,961]" pageId="3" pageNumber="150" rank="species">n. sp.</taxonomicNameLabel>
, Holotype, colour reconstruction.
</paragraph>
</caption>
<caption id="71E6665C482CFF877D87FC0C3AAC6462" ID-DOI="http://doi.org/10.5281/zenodo.12520331" ID-Zenodo-Dep="12520331" box="[227,738,1016,1040]" httpUri="https://zenodo.org/record/12520331/files/figure.png" pageId="3" pageNumber="150" startId="3.[227,266,1016,1040]" targetBox="[227,775,636,996]" targetPageId="3" targetType="figure">
<paragraph id="252636D4482CFF877D87FC0C3AAC6462" blockId="3.[227,738,1016,1040]" box="[227,738,1016,1040]" pageId="3" pageNumber="150">
Fig. 2b.
<taxonomicName id="E2994D57482CFF877C5BFC0C3A646462" authorityName="Vitali" authorityYear="2017" box="[319,554,1016,1040]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="species" species="otiliae" status="sp. nov.">
<emphasis id="17EDEAC6482CFF877C5BFC0C3A646462" box="[319,554,1016,1040]" italics="true" pageId="3" pageNumber="150">Tillomorphites otiliae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CDE57BD482CFF877F56FC0C3A216462" box="[562,623,1016,1040]" pageId="3" pageNumber="150" rank="species">n. sp.</taxonomicNameLabel>
, Paratype.
</paragraph>
</caption>
<caption id="71E6665C482CFF877D87F9593C316696" ID-DOI="http://doi.org/10.5281/zenodo.12520335" ID-Zenodo-Dep="12520335" httpUri="https://zenodo.org/record/12520335/files/figure.png" pageId="3" pageNumber="150" startId="3.[227,265,1709,1733]" targetBox="[298,1252,1029,1698]" targetPageId="3" targetType="figure">
<paragraph id="252636D4482CFF877D87F9593C316696" blockId="3.[227,1285,1708,1765]" pageId="3" pageNumber="150">
Fig. 2c.
<taxonomicName id="E2994D57482CFF877C2EF9593A0566B7" authorityName="Vitali" authorityYear="2017" box="[330,587,1709,1733]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="species" species="otiliae" status="sp. nov.">
<emphasis id="17EDEAC6482CFF877C2EF9593A0566B7" box="[330,587,1709,1733]" italics="true" pageId="3" pageNumber="150">Tillomorphites otiliae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CDE57BD482CFF877F32F9593AD666B7" box="[598,664,1709,1733]" pageId="3" pageNumber="150" rank="species">n. sp.</taxonomicNameLabel>
, habitus, Fig. 2e.
<taxonomicName id="E2994D57482CFF877EEBF9583CEC66B6" authorityName="Vitali" authorityYear="2017" box="[911,1186,1708,1732]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="species" species="otiliae" status="sp. nov.">
<emphasis id="17EDEAC6482CFF877EEBF9583CEC66B6" box="[911,1186,1708,1732]" italics="true" pageId="3" pageNumber="150">Tillomorphites otiliae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CDE57BD482CFF8779D7F9583CB366B6" box="[1203,1277,1708,1732]" pageId="3" pageNumber="150" rank="species">n. sp.</taxonomicNameLabel>
, reconstruction. Paratype, colour reconstruction.
</paragraph>
</caption>
<paragraph id="252636D4482BFF807C4AFF3439EB608A" blockId="4.[302,812,192,248]" pageId="4" pageNumber="151">as onychium; metatarsomeres II and III transverse.</paragraph>
<paragraph id="252636D4482BFF807C4AFED43ACC626A" blockId="4.[302,813,288,536]" pageId="4" pageNumber="151">Prosternum with numerous fine transversal ridges; intercoxal process fairly narrow; procoxae globose, procoxal cavities externally rounded, posteriorly open; mesocoxal cavities laterally closed; visible urosternites (I-V) progressively shortened to the apex, very finely and densely punctured, seemingly granulated, covered with some erect setae.</paragraph>
</subSubSection>
<subSubSection id="6D83655F482BFF807C4AFDB43AA262AA" pageId="4" pageNumber="151" type="etymology">
<paragraph id="252636D4482BFF807C4AFDB439FE622A" blockId="4.[302,813,576,728]" box="[302,432,576,600]" pageId="4" pageNumber="151">
<emphasis id="17EDEAC6482BFF807C4AFDB439FE622A" bold="true" box="[302,432,576,600]" pageId="4" pageNumber="151">Etymology</emphasis>
</paragraph>
<paragraph id="252636D4482BFF807C4AFD943AA262AA" blockId="4.[302,813,576,728]" pageId="4" pageNumber="151">I am honoured to dedicate this new species to my wife Otilia, who gave me as a present both typical specimens and selflessly supports my research providing invaluable assistance.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="8561183CD83861133FF7A81BD25EDCA8">Two new fossil species of Tillomorphites Vitali (Coleoptera: Cerambycidae) and remarks on the morphological evolution, mimicry, biogeography and phylogeny of the tribe Tillomorphini</mods:title>
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<mods:namePart id="4410755B8117E35E6DE0D92C3C8CB184">Vitali, Francesco</mods:namePart>
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<mods:title id="805E2B483039608A0122FD4CBAEDAACC">Baltic Journal of Coleopterology</mods:title>
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<mods:part id="9DA7BA7E5F8E34D8C7DCC37DE2E412DB">
<mods:date id="2095049B11398E19A558E4B22E4C8F81">2017</mods:date>
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<mods:number id="0331D9934290BA8F73C6D8036B5F2BD1">2017-12-31</mods:number>
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<mods:number id="5785C50DDE23B66341680ABE4FAD689B">17</mods:number>
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<mods:number id="D04FD19582E5325D417F6B6A34395374">2</mods:number>
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<mods:identifier id="304903BA0CB40A0C02BDCA11BC1EC440" type="DOI">10.5281/zenodo.12520315</mods:identifier>
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<taxonomicName id="E2994D57482EFF857D87FCF43ACD636A" authority="Vitali, 2011" authorityName="Vitali" authorityYear="2011" box="[227,643,768,792]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="robustus">
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<emphasis id="17EDEAC6482EFF857D87FCF439BC636A" bold="true" box="[227,498,768,792]" italics="true" pageId="1" pageNumber="148">Tillomorphites robustus</emphasis>
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(
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<taxonomicName id="E2994D57482EFF857D87FC743AAF63EA" authority="Vitali, 2011 Baltic" authorityName="Vitali" authorityYear="2011" box="[227,737,896,920]" class="Insecta" family="Cerambycidae" genus="Tillomorphites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="17EDEAC6482EFF857D87FC7439B763EA" box="[227,505,896,920]" italics="true" pageId="1" pageNumber="148">Tillomorphites robustus</emphasis>
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Baltic
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amber, ex coll. M. Veta, author’s coll. FS47BS28.
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