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<document id="AFEB7102B62FF055390ECD0203AC57FD" ID-DOI="10.4081/nhs.2019.394" ID-ISSN="2385-0922" ID-Zenodo-Dep="12523847" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719276420026" checkinUser="felipe" docAuthor="Pasini, Giovanni, Garassino, Alessandro &amp; Sami, Marco" docDate="2019" docId="410487CFFFC03149A327B0B8FEE4FF75" docLanguage="en" docName="NatHistSci.6.1.27-32.pdf" docOrigin="Natural History Sciences 6 (1)" docSource="http://dx.doi.org/10.4081/nhs.2019.394" docStyle="DocumentStyle:926963A4B76E17DFFD817EF231EB8DA9.1:NatHistSci.2024-.journal_article" docStyleId="926963A4B76E17DFFD817EF231EB8DA9" docStyleName="NatHistSci.2024-.journal_article" docStyleVersion="1" docTitle="Monodaeus bortolottii Delle Cave 1988" docType="treatment" docVersion="2" lastPageNumber="30" masterDocId="BD3DFFB7FFC3314DA356B63FFF96FFB5" masterDocTitle="Decapod assemblage from the late Miocene (early-middle Messinian) of the Romagna Apennines nearby Brisighella, Emilia-Romagna (N Italy)" masterLastPageNumber="32" masterPageNumber="27" pageNumber="30" updateTime="1719276794770" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="6B1E0054F13FE1F80B8DFF6493ED69E3">Decapod assemblage from the late Miocene (early-middle Messinian) of the Romagna Apennines nearby Brisighella, Emilia-Romagna (N Italy)</mods:title>
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<mods:namePart id="FDAAF36317827F7CD9B8862E7E181CBE">Pasini, Giovanni</mods:namePart>
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<mods:namePart id="E53E2753E63AB014430065290864A39F">Garassino, Alessandro</mods:namePart>
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<mods:namePart id="B90A9FAD56FE1ED48CCD0A277D5607BC">Sami, Marco</mods:namePart>
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<subSubSection id="81B76552FFC0314EA327B0B8FE12F975" pageId="3" pageNumber="30" type="nomenclature">
<paragraph id="C91236D9FFC0314EA327B0B8FE12F975" blockId="3.[113,756,1671,1728]" pageId="3" pageNumber="30">
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<taxonomicName id="0EAD4D5AFFC0314EA327B0B8FD3DF914" ID-CoL="BTRNJ" authority="Delle Cave, 1988: 123 - 126" authorityName="Delle Cave" authorityPageNumber="123 - 126" authorityYear="1988" box="[113,683,1671,1697]" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA327B0B8FEF8F914" box="[113,366,1671,1697]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
Delle Cave, 1988: 123-126
</taxonomicName>
, Pl. 1, figs. 1, 2, Pl. 2, figs. 1-5.
</heading>
</paragraph>
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<paragraph id="C91236D9FFC0314EA327B0DEFD47F805" blockId="3.[113,757,1761,1969]" pageId="3" pageNumber="30">
<treatmentCitationGroup id="E9BD11F7FFC0314EA327B0DEFD47F805" pageId="3" pageNumber="30">
<taxonomicName id="0EAD4D5AFFC0314EA327B0DEFEF3F94E" ID-CoL="BTRNJ" authorityName="Delle Cave" authorityYear="1988" box="[113,357,1761,1787]" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA327B0DEFEF3F94E" box="[113,357,1761,1787]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
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– De Angeli &amp; Garassino 2006: 74. — De Angeli
<emphasis id="FBD9EACBFFC0314EA258B13FFED0F8AC" box="[270,326,1791,1817]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2009: 185, 195, fig. 16. —
<treatmentCitation id="480C10C8FFC0314EA12DB0C0FEB6F88D" author="Schweitzer C. E. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." page="126" pageId="3" pageNumber="30" year="2010">
<bibRefCitation id="AD3C4B28FFC0314EA12DB0C0FEB6F88D" author="Schweitzer C. E. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." pageId="3" pageNumber="30" pagination="1 - 222" refId="ref5819" refString="Schweitzer C. E., Feldmann R. M., Garassino A., Karasawa H. &amp; Schweigert G., 2010 - Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10: 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="FBD9EACBFFC0314EA327B121FF3EF88D" box="[113,168,1822,1848]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2010: 126
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="480C10C8FFC0314EA21BB121FDF7F88D" author="Garassino A. &amp; Pasini G. &amp; De Angeli A. &amp; Charbonnier S. &amp; Famiani F. &amp; Baldanza A. &amp; Bizzarri R." box="[333,609,1821,1848]" page="52" pageId="3" pageNumber="30" year="2012">
<bibRefCitation id="AD3C4B28FFC0314EA21BB121FDF7F88D" author="Garassino A. &amp; Pasini G. &amp; De Angeli A. &amp; Charbonnier S. &amp; Famiani F. &amp; Baldanza A. &amp; Bizzarri R." box="[333,609,1821,1848]" pageId="3" pageNumber="30" pagination="1 - 61" refId="ref5024" refString="Garassino A., Pasini G., De Angeli A., Charbonnier S., Famiani F., Baldanza A. &amp; Bizzarri R., 2012 - The decapod community from the Early Pliocene (Zanclean) of &quot; La Serra &quot; quarry (San Miniato, central Italy): sedimentology, systematics, and palaeoenvironmental implications. Annales de Paleontologie, 98: 1 - 61." type="journal article" year="2012">
Garassino
<emphasis id="FBD9EACBFFC0314EA297B121FE6EF88D" box="[449,504,1822,1848]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2012: 52
</bibRefCitation>
</treatmentCitation>
. — Baldanza
<emphasis id="FBD9EACBFFC0314EA327B102FF3EF8E3" box="[113,168,1852,1878]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2013: 347, 348, fig. 13. —
<treatmentCitation id="480C10C8FFC0314EA284B103FF0AF8C0" author="Pasini G. &amp; Garassino A." page="325" pageId="3" pageNumber="30" year="2013">
<bibRefCitation id="AD3C4B28FFC0314EA284B103FF0AF8C0" author="Pasini G. &amp; Garassino A." pageId="3" pageNumber="30" pagination="319 - 328" refId="ref5378" refString="Pasini G. &amp; Garassino A., 2013 a - Records of brachyuran crabs from the Pliocene (Piacenzian) of Reggio Emilia (Emilia Romagna, N Italy). Boletin de la Sociedad Geologica Mexicana, 65 (2): 319 - 328." type="journal article" year="2013">Pasini &amp; Garassino 2013a: 325</bibRefCitation>
</treatmentCitation>
, 326, fig. 5. —
<treatmentCitation id="480C10C8FFC0314EA215B165FD04F8C1" author="Pasini G. &amp; Garassino A." box="[323,658,1882,1908]" page="344" pageId="3" pageNumber="30" year="2013">
<bibRefCitation id="AD3C4B28FFC0314EA215B165FD04F8C1" author="Pasini G. &amp; Garassino A." box="[323,658,1882,1908]" pageId="3" pageNumber="30" pagination="343 - 346" refId="ref5426" refString="Pasini G. &amp; Garassino A., 2013 b - New record of crabs (Decapoda, Brachyura) from the early Pliocene of Cassine (Piedmont, northern Italy). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 270 (3): 343 - 346." type="journal article" year="2013">Pasini &amp; Garassino 2013b: 344</bibRefCitation>
</treatmentCitation>
, 345, fig. 2. —
<treatmentCitation id="480C10C8FFC0314EA3FBB147FE09F826" author="Pasini G. &amp; Garassino A. &amp; Hyzny M. &amp; Baldanza A. &amp; Bizzarri R. &amp; Famiani F." box="[173,415,1912,1939]" page="254" pageId="3" pageNumber="30" year="2014">
<bibRefCitation id="AD3C4B28FFC0314EA3FBB147FE09F826" author="Pasini G. &amp; Garassino A. &amp; Hyzny M. &amp; Baldanza A. &amp; Bizzarri R. &amp; Famiani F." box="[173,415,1912,1939]" pageId="3" pageNumber="30" refId="ref5477" refString="Pasini G., Garassino A., Hyzny M., Baldanza A., Bizzarri R. &amp; Famiani F., 2014 - The bathyal decapod crustacean community from the early Pleistocene of Volterra (Pisa, Tuscany, central Italy). Neues" type="book" year="2014">
Pasini
<emphasis id="FBD9EACBFFC0314EA3A3B146FEBCF827" box="[245,298,1912,1938]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2014: 254
</bibRefCitation>
</treatmentCitation>
, 255, fig. 9B. — Baldanza
<emphasis id="FBD9EACBFFC0314EA1E9B146FD62F827" box="[703,756,1912,1938]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2017: 60, 61, fig. 15C. —
<treatmentCitation id="480C10C8FFC0314EA2DBB1A9FDE5F805" author="Pasini G. &amp; Garassino A. &amp; Sami M." box="[397,627,1942,1968]" page="31" pageId="3" pageNumber="30" year="2018">
<bibRefCitation id="AD3C4B28FFC0314EA2DBB1A9FDE5F805" author="Pasini G. &amp; Garassino A. &amp; Sami M." box="[397,627,1942,1968]" pageId="3" pageNumber="30" refId="ref5541" refString="Pasini G., Garassino A. &amp; Sami M., 2018 - Decapod crustaceans from the late Pliocene (Placentian) nearby Faenza (Emilia-Romagna, N Italy). Natural History Sciences. Atti della Societa italiana di" type="book" year="2018">
Pasini
<emphasis id="FBD9EACBFFC0314EA283B1A8FD9AF805" box="[469,524,1942,1968]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2018: 31
</bibRefCitation>
</treatmentCitation>
, fig. 3A.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="81B76552FFC0314EA033B4DFFB6BFB9C" pageId="3" pageNumber="30" type="materials_examined">
<paragraph id="C91236D9FFC0314EA033B4DFFB6BFB9C" blockId="3.[831,1475,736,1969]" pageId="3" pageNumber="30">
<emphasis id="FBD9EACBFFC0314EA033B4DFFB21FD4E" bold="true" box="[869,1207,736,763]" pageId="3" pageNumber="30">Material and measurements</emphasis>
: 13 carapaces in dorsal view (
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2300 – lcxp:
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, wcxp:
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);
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2307
<emphasis id="FBD9EACBFFC0314EA0D7B522FC19FC82" bold="true" box="[897,911,797,823]" pageId="3" pageNumber="30">–</emphasis>
lcxp:
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, wcxp:
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;
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1154
<emphasis id="FBD9EACBFFC0314EA624B522FA16FC82" bold="true" box="[1394,1408,797,823]" pageId="3" pageNumber="30">–</emphasis>
lcxp:
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, wcxp:
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2309
<emphasis id="FBD9EACBFFC0314EA79FB504FB41FCE0" bold="true" box="[1225,1239,827,853]" pageId="3" pageNumber="30">–</emphasis>
lcxp:
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, wcxp:
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;
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2315
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lcxp:
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, wcxp:
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;
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, wcxp:
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;
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2325 – lcxp:
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, wcxp:
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, wcxp:
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;
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2327 – lcxp:
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, wcxp:
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;
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2328 – lcxp:
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, wcxp:
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;
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2329 – lcxp:
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, wcxp:
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;
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2330 – lcxp:
<quantity id="0E559B3CFFC0314EA784B5CFFA8EFBBF" box="[1234,1304,1008,1034]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="30" unit="mm" value="7.0">7 mm</quantity>
, wcxp:
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;
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2331 – lcxp:
<quantity id="0E559B3CFFC0314EA747B231FBC2FB9C" box="[1041,1108,1038,1065]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="30" unit="mm" value="6.0">6 mm</quantity>
, wcxp:
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).
</paragraph>
</subSubSection>
<subSubSection id="81B76552FFC03149A033B213FEE4FF75" lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30" type="discussion">
<paragraph id="C91236D9FFC0314EA033B213FB9FFA05" blockId="3.[831,1475,736,1969]" pageId="3" pageNumber="30">
<emphasis id="FBD9EACBFFC0314EA033B213FC75FBF3" bold="true" box="[869,995,1068,1094]" pageId="3" pageNumber="30">Discussion</emphasis>
. The studied specimens have been assigned to
<taxonomicName id="0EAD4D5AFFC0314EA0DAB275FAC4FBD1" authority="Delle Cave, 1988" authorityName="Delle Cave" authorityYear="1988" box="[908,1362,1098,1124]" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA0DAB275FB1EFBD1" box="[908,1160,1098,1124]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
Delle Cave,
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having a slightly convex suboctagonal carapace, wider than long; straight front margin, with a weak median incision; short convex anterolateral margin; long convergent posterolateral margin; posterior margin straight medially; dorsal region well marked by grooves, with wide raised epigastric lobes; well-marked suboval protogastric regions; subpentagonal mesogastric regions with narrow, elongate anterior process between protogastric regions; cardiac region well marked by branchiocardiac grooves; small poorly marked hepatic regions; and wide, well-marked, branchial regions.
</paragraph>
<paragraph id="C91236D9FFC03149A033B38BFEE4FF75" blockId="3.[831,1475,736,1969]" lastBlockId="4.[113,755,135,192]" lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
<taxonomicName id="0EAD4D5AFFC0314EA033B38BFBFEFA7B" authorityName="Delle Cave" authorityYear="1988" box="[869,1128,1460,1486]" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA033B38BFBFEFA7B" box="[869,1128,1460,1486]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
</taxonomicName>
has been previously reported from the Pliocene of Volterra (Delle Cave, 1988) and Grosseto (De Angeli
<emphasis id="FBD9EACBFFC0314EA767B3CEFBFCF9BF" box="[1073,1130,1520,1546]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
, 2009). Later
<bibRefCitation id="AD3C4B28FFC0314EA65AB3CFFC5CF99D" author="Pasini G. &amp; Garassino A." pageId="3" pageNumber="30" pagination="319 - 328" refId="ref5378" refString="Pasini G. &amp; Garassino A., 2013 a - Records of brachyuran crabs from the Pliocene (Piacenzian) of Reggio Emilia (Emilia Romagna, N Italy). Boletin de la Sociedad Geologica Mexicana, 65 (2): 319 - 328." type="journal article" year="2013">Pasini &amp; Garassino (2013a</bibRefCitation>
, b) and Garassino, Hyžný &amp; Pasini
<emphasis id="FBD9EACBFFC0314EA624B030FA1EF99D" box="[1394,1416,1551,1576]" italics="true" pageId="3" pageNumber="30">in</emphasis>
Baldanza
<emphasis id="FBD9EACBFFC0314EA0DFB012FC57F9F3" box="[905,961,1580,1606]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
(2013) reported this species from the Pliocene (Piacenzian) of Castellarano and Monticelli di Quattro Castella (Reggio Emilia,
<collectingRegion id="0B69F83BFFC0314EA7D0B056FAD8F931" box="[1158,1358,1641,1668]" country="Italy" name="Emilia-Romagna" pageId="3" pageNumber="30">Emilia-Romagna</collectingRegion>
), from the early Pliocene of Cassine (Alessandria,
<collectingRegion id="0B69F83BFFC0314EA644B0B8FA13F917" box="[1298,1413,1671,1698]" country="Italy" name="Piemonte" pageId="3" pageNumber="30">Piedmont</collectingRegion>
), and from the early Pleistocene of the Poggio i Sodi (Siena,
<collectingRegion id="0B69F83BFFC0314EA069B0FBFC32F96B" box="[831,932,1732,1758]" country="Italy" name="Toscana" pageId="3" pageNumber="30">Tuscany</collectingRegion>
). Later
<bibRefCitation id="AD3C4B28FFC0314EA0AEB0FCFB5BF968" author="Pasini G. &amp; Garassino A. &amp; Hyzny M. &amp; Baldanza A. &amp; Bizzarri R. &amp; Famiani F." box="[1016,1229,1731,1757]" pageId="3" pageNumber="30" refId="ref5477" refString="Pasini G., Garassino A., Hyzny M., Baldanza A., Bizzarri R. &amp; Famiani F., 2014 - The bathyal decapod crustacean community from the early Pleistocene of Volterra (Pisa, Tuscany, central Italy). Neues" type="book" year="2014">
Pasini
<emphasis id="FBD9EACBFFC0314EA715B0FBFBEDF968" box="[1091,1147,1731,1757]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
(2014)
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reported
<taxonomicName id="0EAD4D5AFFC0314EA66EB0FBFCC1F94E" authorityName="Delle Cave" authorityYear="1988" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA66EB0FBFCC1F94E" italics="true" pageId="3" pageNumber="30">M. bortolottii</emphasis>
</taxonomicName>
from the early Pleistocene of Volterra (Pisa,
<collectingRegion id="0B69F83BFFC0314EA604B0DDFA20F949" box="[1362,1462,1762,1788]" country="Italy" name="Toscana" pageId="3" pageNumber="30">Tuscany</collectingRegion>
), close to the
<typeStatus id="1616887BFFC0314EA081B13FFB9EF8AF" box="[983,1032,1792,1818]" pageId="3" pageNumber="30">type</typeStatus>
locality originally reported by Delle Cave (1988). Recently this species has been reported also from the early Pleistocene of Poggi Gialli (Sinalunga,
<collectingRegion id="0B69F83BFFC0314EA069B165FC37F8C1" box="[831,929,1882,1908]" country="Italy" name="Toscana" pageId="3" pageNumber="30">Tuscany</collectingRegion>
) by De Angeli, Garassino &amp; Pasini
<emphasis id="FBD9EACBFFC0314EA66CB165FAC6F8C6" box="[1338,1360,1882,1907]" italics="true" pageId="3" pageNumber="30">in</emphasis>
Baldanza
<emphasis id="FBD9EACBFFC0314EA069B146FCEEF827" box="[831,888,1912,1938]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
(2017), and from the Pliocene of Faenza (Ravenna,
<collectingRegion id="0B69F83BFFC0314EA069B1A9FB92F804" box="[831,1028,1942,1969]" country="Italy" name="Emilia-Romagna" pageId="3" pageNumber="30">Emilia-Romagna</collectingRegion>
) (
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Pasini
<emphasis id="FBD9EACBFFC0314EA739B1A8FB24F805" box="[1135,1202,1942,1968]" italics="true" pageId="3" pageNumber="30">et al.,</emphasis>
2018
</bibRefCitation>
). This is the first record of this species in the Miocene, notably enlarging its stratigraphic range.
</paragraph>
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71
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<paragraph id="C91236D9FFC0314EA327B012FDA1F9F2" blockId="3.[113,567,1581,1637]" box="[113,567,1581,1607]" pageId="3" pageNumber="30">
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<taxonomicName id="0EAD4D5AFFC0314EA327B012FDA1F9F2" ID-CoL="BTRNJ" authority="Delle Cave, 1988" authorityName="Delle Cave" authorityYear="1988" box="[113,567,1581,1607]" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA327B012FEFBF9F2" box="[113,365,1581,1607]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
Delle Cave, 1988
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Genus
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Linnaeus, 1761
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Genus
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Genus
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Lamarck, 1801
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[=
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