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Kugelann, 1798: 421. Type species:
<taxonomicName authorityName="Herbst" authorityYear="1783" class="Insecta" family="Coccinellidae" genus="Chrysomela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysomela scutellata" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="scutellata">Chrysomela scutellata</taxonomicName>
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Herbst, 1783, by subsequent designation by
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.8975" author="Crotch, GR" journalOrPublisher="Janson, London" pageId="0" pageNumber="61" refId="B5" refString="Crotch, GR, 1874. A revision of the coleopterous family Coccinellidae. E. W. Janson, London, DOI: https://doi.org/10.5962/bhl.title.8975" title="A revision of the coleopterous family Coccinellidae. E. W." url="https://doi.org/10.5962/bhl.title.8975" year="1874">Crotch 1874</bibRefCitation>
<bibRefCitation id="8EEEE093F047427DA022495504A338B9" DOI="https://doi.org/10.5962/bhl.title.8975" author="Crotch, GR" journalOrPublisher="Janson, London" pageId="0" pageNumber="61" refId="B5" refString="Crotch, GR, 1874. A revision of the coleopterous family Coccinellidae. E. W. Janson, London, DOI: https://doi.org/10.5962/bhl.title.8975" title="A revision of the coleopterous family Coccinellidae. E. W." url="https://doi.org/10.5962/bhl.title.8975" year="1874">Crotch 1874</bibRefCitation>
.
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<taxonomicName class="Secernentea" family="Coccinellidae" genus="Strongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Nematoda" rank="genus">Strongylus</taxonomicName>
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Panzer, 1813: 114.
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<taxonomicName authorityName="Dejean" authorityYear="1821" class="Insecta" family="Coccinellidae" genus="Cacidula" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cacidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Cacidula</taxonomicName>
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Dejean, 1821: 132. Type species:
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<taxonomicName id="B48FB2F65A17E840DD1E9061447EAAD7" authorityName="Fabricius" authorityYear="1792" class="Insecta" family="Coccinellidae" genus="Chrysomela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysomela pectoralis" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pectoralis">Chrysomela pectoralis</taxonomicName>
Fabricius, 1792 (=
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Herbst, 1783).
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<taxonomicName id="29FF0D455C808B1067698478A34324A3" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Coccinellidae" genus="Cacicula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacicula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Cacicula</taxonomicName>
Stephens, 1831: 397.
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<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
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<paragraph id="918772CAFCCAB522180D44124876B22E" pageId="0" pageNumber="61">Diagnosis.</paragraph>
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Representatives of the genus
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<emphasis italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
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</taxonomicName>
with its general body shape may resemble
<taxonomicName authorityName="Kapur" authorityYear="1948" class="Insecta" family="Coccinellidae" genus="Tetrabrachys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tetrabrachys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Tetrabrachys</emphasis>
<taxonomicName id="546154551BBF15C1ED24741F51F551C4" authorityName="Kapur" authorityYear="1948" class="Insecta" family="Coccinellidae" genus="Tetrabrachys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tetrabrachys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="06187542FB1D93BF25B2FCAD6C97AA1E" italics="true" pageId="0" pageNumber="61">Tetrabrachys</emphasis>
</taxonomicName>
Kapur, however, it can be separated based on the structure of the tarsi which are tetramerous in both genera but in
<taxonomicName authorityName="Kugelann" authorityYear="1798" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
<taxonomicName id="FA412CB6632235B079CB2218F0B5EE72" authorityName="Kugelann" authorityYear="1798" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="4A97388BFCA350C635F41A1F1DF339D8" italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
</taxonomicName>
the first tarsomere is sub-triangularly broadened apically and the second is elongate and distinctly lobbed, while in
<taxonomicName authorityName="Kapur" authorityYear="1948" class="Insecta" family="Coccinellidae" genus="Tetrabrachys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tetrabrachys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Tetrabrachys</emphasis>
<taxonomicName id="E65BD04DE1AB5FC21C6D5A2872BB117F" authorityName="Kapur" authorityYear="1948" class="Insecta" family="Coccinellidae" genus="Tetrabrachys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tetrabrachys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="393468DF106E4EFD02D4FC5BFA914F12" italics="true" pageId="0" pageNumber="61">Tetrabrachys</emphasis>
</taxonomicName>
both the first and second are narrow, elongate and without lobes. Moreover, in
<taxonomicName authorityName="Kapur" authorityYear="1948" class="Insecta" family="Coccinellidae" genus="Tetrabrachys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tetrabrachys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Tetrabrachys</emphasis>
<taxonomicName id="CB399CDA4782590CD7AFF10B2D3E24A1" authorityName="Kapur" authorityYear="1948" class="Insecta" family="Coccinellidae" genus="Tetrabrachys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tetrabrachys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="46AEDA93EE37B2DFB67159D8E7700D1B" italics="true" pageId="0" pageNumber="61">Tetrabrachys</emphasis>
</taxonomicName>
the apical maxillary palpomere is widely securiform, and beetles are brachypterous, while in
<taxonomicName authorityName="Kugelann" authorityYear="1798" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
<taxonomicName id="A5FCB45703FFA63EAB8430B10A56108E" authorityName="Kugelann" authorityYear="1798" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="E41A9B5852ED565CF282A477245D770D" italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
</taxonomicName>
the apical maxillary palpomere is only slightly widened and the second pair of wings is functional.
<taxonomicName authorityName="Kugelann" authorityYear="1798" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
<taxonomicName id="3EF1D1129112DD23C0AA2360C551CD15" authorityName="Kugelann" authorityYear="1798" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="908C0BBB648C83680C8331476D4DFCA1" italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
</taxonomicName>
is also externally similar to European species of
<taxonomicName class="Insecta" family="Coccinellidae" genus="Rhyzobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhyzobius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Rhyzobius</emphasis>
<taxonomicName id="EE68772DD57C34A1EBAA3BBB53FEDBF0" class="Insecta" family="Coccinellidae" genus="Rhyzobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhyzobius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="6086037D0BF9AE729DDA0C0B1DE0A382" italics="true" pageId="0" pageNumber="61">Rhyzobius</emphasis>
</taxonomicName>
but it can be separated based on the following characters: body almost parallel sided, elytra covered with punctures of two sizes, larger punctures arranged in nine rows (in
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. litophiloides" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="litophiloides">
<emphasis italics="true" pageId="0" pageNumber="61">C. litophiloides</emphasis>
<taxonomicName id="DA86469CA5978674739A70AB7B2B0888" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. litophiloides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="litophiloides">
<emphasis id="833CB629F1EF40489B6963B20E71879E" italics="true" pageId="0" pageNumber="61">C. litophiloides</emphasis>
</taxonomicName>
some of them are reduced), base of the pronotum not bordered, while in
<taxonomicName class="Insecta" family="Coccinellidae" genus="Rhyzobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhyzobius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Rhyzobius</emphasis>
<taxonomicName id="A11AB94892F878506AD07A5C7FAB71FD" class="Insecta" family="Coccinellidae" genus="Rhyzobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhyzobius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="F94B2A0A0ABAB14A4310315EF0B5F611" italics="true" pageId="0" pageNumber="61">Rhyzobius</emphasis>
</taxonomicName>
the lateral body outline is broadly rounded, the elytra are covered with single sized, randomly arranged punctures, and base of the pronotum with distinct bordering line.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">Description.</paragraph>
<paragraph pageId="0" pageNumber="61">
<subSubSection id="D001D03CF17B7CFCFBFE1AE8A9C92CAC" pageId="0" pageNumber="61" type="description">
<paragraph id="902888584A071FD88C8A6A7A9ABBA25A" pageId="0" pageNumber="61">Description.</paragraph>
<paragraph id="0B4D13734D5EC9A7CADEB687C14EBE86" pageId="0" pageNumber="61">
Body elongate-oval, with sides parallel (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1C-H</figureCitation>
<figureCitation id="2FC446C82F51CDFBD16D1643084C7582" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1C-H</figureCitation>
), body flattened in lateral view, convex in cross-section; dorsum covered with setiferous punctures of two sizes (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7A</figureCitation>
<figureCitation id="E8D48A2E382C87A318E44B62FDEB5925" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7A</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9A</figureCitation>
<figureCitation id="34911E179DAE583AB0A606F9165024EB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9A</figureCitation>
), hairs directed forwards on pronotum, backwards on elytra.
</paragraph>
<caption doi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 1.</emphasis>
<caption id="4C06A1D1ECABF62E107F4D36EDE56F97" doi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61" start="Figure 1" startId="F1">
<paragraph id="0CE8AE57786818388C4B7A96867C6D03" pageId="0" pageNumber="61">
<emphasis id="BAB4C064AE442F2133E0892ED91DE6DF" bold="true" pageId="0" pageNumber="61">Figure 1.</emphasis>
Immature stages and habitus of adult species of
<taxonomicName authorityName="Kugelann" authorityYear="1798" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
<taxonomicName id="6AB3108AE52CD4894F2AD7B2A844D7DC" authorityName="Kugelann" authorityYear="1798" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="055AD9AFDEA19D20BACCFD25444741B7" italics="true" pageId="0" pageNumber="61">Coccidula</emphasis>
</taxonomicName>
Kugelann
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. rufa" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="rufa">
<emphasis italics="true" pageId="0" pageNumber="61">C. rufa</emphasis>
<emphasis id="38523A3FCB081370BBFF9D4975E49D50" bold="true" pageId="0" pageNumber="61">A</emphasis>
<taxonomicName id="37B48F1B864659912A77F377758EDD49" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. rufa" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B26D8AC1DCB95C07961E651793683934" italics="true" pageId="0" pageNumber="61">C. rufa</emphasis>
</taxonomicName>
larva
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. rufa" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="rufa">
<emphasis italics="true" pageId="0" pageNumber="61">C. rufa</emphasis>
<emphasis id="84D21702BB8B5EA3539DE13018606669" bold="true" pageId="0" pageNumber="61">B</emphasis>
<taxonomicName id="C3DA4AFE9D2A7C82D7856271517766F2" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. rufa" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="1B7883AED4E62163D47B45B5174F749C" italics="true" pageId="0" pageNumber="61">C. rufa</emphasis>
</taxonomicName>
pupa
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. lepida" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="lepida">
<emphasis italics="true" pageId="0" pageNumber="61">C. lepida</emphasis>
<emphasis id="E14E979426EA96D6A4303B23F5B7B08C" bold="true" pageId="0" pageNumber="61">C</emphasis>
<taxonomicName id="2FDA7F803578163897A5DC23E976F7CD" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. lepida" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="lepida">
<emphasis id="A39FD0330B04617C083C548568C6B6C0" italics="true" pageId="0" pageNumber="61">C. lepida</emphasis>
</taxonomicName>
, dorsal
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. lepida" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="lepida">
<emphasis italics="true" pageId="0" pageNumber="61">C. lepida</emphasis>
<emphasis id="7A1BF88E26661D9F6157AF79215F3D7E" bold="true" pageId="0" pageNumber="61">D</emphasis>
<taxonomicName id="D2D845196158FED9B7D15E98048EAFE0" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. lepida" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="lepida">
<emphasis id="A3C912AD9DD3D1F969A90AB30EAD1B3F" italics="true" pageId="0" pageNumber="61">C. lepida</emphasis>
</taxonomicName>
, ventral
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. litophiloides" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="litophiloides">
<emphasis italics="true" pageId="0" pageNumber="61">C. litophiloides</emphasis>
<emphasis id="F2E714E2709975EFD473E88414550003" bold="true" pageId="0" pageNumber="61">E</emphasis>
<taxonomicName id="45504C9AE542AFDF9E78ECC22C8BB227" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. litophiloides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="litophiloides">
<emphasis id="1B13C29F1A313F51ABDCB19CA3DDC8D7" italics="true" pageId="0" pageNumber="61">C. litophiloides</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="61">F</emphasis>
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. reitteri" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="reitteri">
<emphasis italics="true" pageId="0" pageNumber="61">C. reitteri</emphasis>
<emphasis id="3C901AE524560ADB3BA285B7650F1677" bold="true" pageId="0" pageNumber="61">F</emphasis>
<taxonomicName id="AEC1A1A2AB5FA64D624C44719001DE50" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. reitteri" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="DDF690FE591AD9E7A0BD27E30E3087A8" italics="true" pageId="0" pageNumber="61">C. reitteri</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="61">G</emphasis>
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. rufa" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="rufa">
<emphasis italics="true" pageId="0" pageNumber="61">C. rufa</emphasis>
<emphasis id="F5CCF0F716047B926C395CF705EB185A" bold="true" pageId="0" pageNumber="61">G</emphasis>
<taxonomicName id="4B1691C58AE6172ABBB1B8DC5086C9DD" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. rufa" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="0A9A7D0E7FCA8C55EDF57E33F3CE3403" italics="true" pageId="0" pageNumber="61">C. rufa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="61">H</emphasis>
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. scutellata" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="scutellata">
<emphasis italics="true" pageId="0" pageNumber="61">C. scutellata</emphasis>
<emphasis id="FCCC5EF1C416C4CF666CC25A8F6DF35D" bold="true" pageId="0" pageNumber="61">H</emphasis>
<taxonomicName id="886B360806A42B43E109B65DA90D59B1" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. scutellata" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="scutellata">
<emphasis id="51BB7F1E71E19D65BDC014439346988C" italics="true" pageId="0" pageNumber="61">C. scutellata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="61">A, B</emphasis>
<emphasis id="7628F28F7F998375CA329FB3C7D36BA1" bold="true" pageId="0" pageNumber="61">A, B</emphasis>
Gilles San Martin, Wikimedia Commons
<emphasis bold="true" pageId="0" pageNumber="61">C, D</emphasis>
<emphasis id="C99C77BC35B5EBB2A0E7FBAA1743B9B4" bold="true" pageId="0" pageNumber="61">C, D</emphasis>
Danny Haelewaters
<emphasis bold="true" pageId="0" pageNumber="61">G, H</emphasis>
<emphasis id="CC00CD739E192A3C304DF958944C8455" bold="true" pageId="0" pageNumber="61">G, H</emphasis>
Udo Schmidt. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="61">C-H</emphasis>
<emphasis id="A860A5004298001861BFE495453AA621" bold="true" pageId="0" pageNumber="61">C-H</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<paragraph id="0A91F16FF3F00CFD41776B528DFD6525" pageId="0" pageNumber="61">
Head partially withdrawn into prothorax (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1C-H</figureCitation>
<figureCitation id="D3BEFE7FA5F33735878AE2AEADC97238" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1C-H</figureCitation>
); ventral antennal grooves shallow and moderately long, extending to posterior border of an eye (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3E</figureCitation>
<figureCitation id="CF0B6329E343E40B49588B93DADF63C8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3E</figureCitation>
). Eyes prominent, coarsely facetted (7-8 ommatidia per eye width), ocular canthus distinct, about as long as 4-5 ommatidium diameters; interocular distance about 3
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as eye diameter; interfacetal setae present only in basal part; temple behind eye distinctly longer than eye (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3E</figureCitation>
<figureCitation id="437C50D62CC4F86802BC0F7E3F913465" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3E</figureCitation>
). Antennal insertion placed laterally, invisible from above, distance between antennal insertions about same as between eyes; frons around antennal insertions slightly expanded, covering antennal insertions, anterior tentorial pits placed ventrally below antennal insertions. Antennae (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5A, E</figureCitation>
<figureCitation id="E6C0099D49002495F0AF0B77900F4CA3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5A, E</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7A, D</figureCitation>
<figureCitation id="E27F791D0020B1478930846AACA24709" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7A, D</figureCitation>
) longer than maximum head width including eyes, composed of 11 antennomeres (AN); scape simple, without projections, slightly curved; pedicel distinctly narrower than scape, elongate (1.5
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longer than wide); AN 3-8 elongate (AN3 ≈ 3.5
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; AN8 ≈ 1.3
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longer than wide); AN 9-11 forming a loose, asymmetric club, ultimate AN truncate apically. Frontoclypeus short, transverse, anterior margin straight. Labrum entirely exposed, transverse, anterior margin straight. Mandibles asymmetric, bifid apically (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10G</figureCitation>
<figureCitation id="544B04DA0149DF5DB6A68D62C84AEC29" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10G</figureCitation>
), molar part with basal tooth; prostheca distinct. Maxillary stipes (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coccidula lepida LeConte SEM illustrations A pronotum B head and prothorax, ventral C meso and metaventrite D elytra dorsal E ventrite 1. Scale bars: 500 µm (A-C); 1 mm (E, D)." figureDoi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61">2B</figureCitation>
<figureCitation id="EE65AA2AB661EEFC5F0A3D68D7E9DC1C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coccidula lepida LeConte SEM illustrations A pronotum B head and prothorax, ventral C meso and metaventrite D elytra dorsal E ventrite 1. Scale bars: 500 µm (A-C); 1 mm (E, D)." figureDoi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61">2B</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5C</figureCitation>
<figureCitation id="25D869129BA50DF3A98D4FE6A27EF9F5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5C</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9C</figureCitation>
<figureCitation id="A8BCCCAE4892C090678B6FE2A710F4FA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9C</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10H</figureCitation>
<figureCitation id="AD8C1AFBB7F2D356FFF93B85314B3A96" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10H</figureCitation>
) with distinct groove for reception of maxillary palp in repose; palpomere 2 shorter than terminal (4th) one, slightly broadened apically; palpomere 3 about 2.3
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<normalizedToken id="DEC4134E707DF4B3E8332A425DAB8046" originalValue="×">x</normalizedToken>
shorter than terminal one, subtriangular; terminal palpomere slightly securiform; lacinia with stiff setae on outer margin in apical half, with several additional spurs on surface (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10H</figureCitation>
<figureCitation id="0AE3571E852B872E3EB4829761F0336F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10H</figureCitation>
). Labial palps (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3E</figureCitation>
<figureCitation id="ED19CA84C5187514733E2005A49CED0D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3E</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9C</figureCitation>
<figureCitation id="6D893A73D90299565CE51399088438ED" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9C</figureCitation>
) with 3 palpomeres, inserted ventrally on prementum; palpomere 1 very small, apical palpomere as long as and about as broad as penultimate; distance between palp insertions about 1.5-2
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as its width. Prementum subquadrate, transverse apically. Mentum trapezoidal, broadest in anterior part, with horseshoe impression at base (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7C</figureCitation>
<figureCitation id="457348E51A8811D87C5D2E5A3634D905" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7C</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9C</figureCitation>
<figureCitation id="54D5C0C5878D5E09854BB965485C88FF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9C</figureCitation>
). Submentum broad, transverse, with suture invisible.
</paragraph>
<caption doi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
<taxonomicName authorityName="LeConte" authorityYear="1852" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula lepida" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="lepida">
<emphasis italics="true" pageId="0" pageNumber="61">Coccidula lepida</emphasis>
<caption id="D22F7AACAB61C4E200406C03FA9723E5" doi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61" start="Figure 2" startId="F2">
<paragraph id="BBA2B6F7A5795A6BD445FC1DA24992BB" pageId="0" pageNumber="61">
<emphasis id="63C648EFA2E9D22DD8859D9F3520C997" bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
<taxonomicName id="C3309C5D1F665C01B55DA38DB6A8B67F" authorityName="LeConte" authorityYear="1852" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula lepida" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="lepida">
<emphasis id="2CB0988297F3402E628AC34253604E6E" italics="true" pageId="0" pageNumber="61">Coccidula lepida</emphasis>
</taxonomicName>
LeConte SEM illustrations
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
<emphasis id="4B04125E311A9CC77BF8CEA9101B8FEA" bold="true" pageId="0" pageNumber="61">A</emphasis>
pronotum
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
<emphasis id="C9E3392D448CDBAC5D60600A2FCDA14A" bold="true" pageId="0" pageNumber="61">B</emphasis>
head and prothorax, ventral
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
<emphasis id="064BFA9516AD7A05646CEC73F849EAA6" bold="true" pageId="0" pageNumber="61">C</emphasis>
meso and metaventrite
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
<emphasis id="BCED575E96CEBB72248AC6940281C118" bold="true" pageId="0" pageNumber="61">D</emphasis>
elytra dorsal
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
<emphasis id="5CFA5464B362B5688F8B221ED911F8B4" bold="true" pageId="0" pageNumber="61">E</emphasis>
ventrite 1. Scale bars: 500
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<emphasis bold="true" pageId="0" pageNumber="61">(A-C)</emphasis>
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<emphasis id="14FEDCFF3338822D062675D573706551" bold="true" pageId="0" pageNumber="61">(A-C)</emphasis>
; 1 mm (
<emphasis bold="true" pageId="0" pageNumber="61">E, D</emphasis>
<emphasis id="0F765574D3F15801460DBB7C7147B757" bold="true" pageId="0" pageNumber="61">E, D</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<paragraph id="BD383059D09003F2B18FF41553D4809F" pageId="0" pageNumber="61">
Anterior margin of pronotum weakly, broadly emarginate (Figs
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<figureCitation id="09DB9AFCBB240AFD0F45EC23FF93E4AC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3B</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5B</figureCitation>
<figureCitation id="DCA9FE2424E89F618A6738F624A2D6CE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5B</figureCitation>
) with anterior corners broadly rounded; lateral margins with moderately (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7B</figureCitation>
<figureCitation id="56E8A1D3E876AB5F6E98CD34C885CFEB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7B</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9B</figureCitation>
<figureCitation id="D6623BD509BE6DBAD46A501C104B950B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9B</figureCitation>
) to distinctly expanded lateral beads (Fig.
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<figureCitation id="E18D924207FD87B535DE98379639FEDD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3B</figureCitation>
), distinctly margined; hind corners sharply pointed; hind margin not bordered. Prothoracic hypomeron smooth, without delimited foveae (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3C</figureCitation>
<figureCitation id="C8B8A4442DE2E76BAC9B47770A7AB1D2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3C</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7C</figureCitation>
<figureCitation id="F619FCBCE48DACB76DA78C049A765D86" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7C</figureCitation>
). Prosternum in front of coxae about as long as longitudinal length of procoxal cavity; anterior margin straight or slightly emarginate with distinct border. Prosternal process about 0.4 times of coxal diameter, surface smooth (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3C</figureCitation>
<figureCitation id="9CA9EBAA4E24020CF0DB76199ED66992" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3C</figureCitation>
) or with lateral carinae (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7C</figureCitation>
<figureCitation id="B41864617527E5E9145F28CB5C52ED04" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7C</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9E</figureCitation>
<figureCitation id="F66427F02DBF0CD976E0689F37CC6D09" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9E</figureCitation>
). Procoxal cavity oval, distinctly bordered anteriorly.
</paragraph>
<caption doi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
<taxonomicName authorityName="Reitter" authorityYear="1890" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula litophiloides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="litophiloides">
<emphasis italics="true" pageId="0" pageNumber="61">Coccidula litophiloides</emphasis>
<caption id="97F52A4EC78FD08F33960034E659964C" doi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
<paragraph id="E1C4B4555EAB5721355AB0279A5F26A0" pageId="0" pageNumber="61">
<emphasis id="D229FCEF82C8D40A18C616444A548819" bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
<taxonomicName id="4EE615B9753A5A9E8D79D1231F44FE17" authorityName="Reitter" authorityYear="1890" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula litophiloides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="litophiloides">
<emphasis id="0715C7467103B7D333DB9126F3D93988" italics="true" pageId="0" pageNumber="61">Coccidula litophiloides</emphasis>
</taxonomicName>
Reitter SEM illustrations, paratype HNHM
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
<emphasis id="B8208F8383AB5A4380988C77E3FEAF96" bold="true" pageId="0" pageNumber="61">A</emphasis>
body, dorsal
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
<emphasis id="C0C52FD6A16B8B86BF923C4611DE7F31" bold="true" pageId="0" pageNumber="61">B</emphasis>
pronotum
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
<emphasis id="6C258879040F19ADE4D5B05F13C14BDB" bold="true" pageId="0" pageNumber="61">C</emphasis>
head and prothorax, ventral
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
<emphasis id="98C9FDDAE789E0BA92DF13EB4F5F5764" bold="true" pageId="0" pageNumber="61">D</emphasis>
meso and metaventrite
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
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head ventral. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
<emphasis id="3FEC0B415B084EEEFC20901CBB8A96AC" bold="true" pageId="0" pageNumber="61">A</emphasis>
); 500
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(
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<emphasis id="E182A8DAE05CA77E76FDB9E38EF3BEE7" bold="true" pageId="0" pageNumber="61">B-D</emphasis>
); 400
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="A8B5973B7DA816FB2DF6E2ED66422E10" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
<emphasis id="6873489C3972B9CF0F0C7F513321F098" bold="true" pageId="0" pageNumber="61">E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<paragraph id="C9C8A7F3CAED41F5F3DAD4E88DD6C525" pageId="0" pageNumber="61">
Mesoventrite 1.3
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<normalizedToken id="7AE253774B0DA05266AFEB2AEEF2C707" originalValue="×">x</normalizedToken>
longer than its width at the level of mid coxae (Figs
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<figureCitation id="3A35D7180D468416F8AA9CECF63D58C7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1D</figureCitation>
,
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,
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); mesal surface with deep emargination for receiving tip of prosternal process (Fig.
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); anterior margin with completely raised border. Meso-metaventral process narrow (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1D</figureCitation>
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,
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<figureCitation id="DFD45CC8F59D5284C993BE09AAC28E69" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coccidula lepida LeConte SEM illustrations A pronotum B head and prothorax, ventral C meso and metaventrite D elytra dorsal E ventrite 1. Scale bars: 500 µm (A-C); 1 mm (E, D)." figureDoi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61">2C</figureCitation>
,
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,
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), about 0.5 times of mesocoxal diameter, junction slightly arcuate (Figs
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,
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<figureCitation id="953A968FFDAC1C3126BA04105AA9D632" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3D</figureCitation>
,
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,
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,
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), with suture visible. Metendosternite with stalk sub-quadrate, tendons long, separated by a distance of about width of stalk and situated closer to center (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10I</figureCitation>
<figureCitation id="DD5CC3D41331A23D3014AC8354D03DDF" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10I</figureCitation>
). Scutellar shield pentagonal (Figs
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,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9B</figureCitation>
<figureCitation id="D05DCDDF8A475DB1BA3E3F7FA409C303" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9B</figureCitation>
). Elytra at base wider than pronotum, lateral margins clearly visible from above throughout (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coccidula lepida LeConte SEM illustrations A pronotum B head and prothorax, ventral C meso and metaventrite D elytra dorsal E ventrite 1. Scale bars: 500 µm (A-C); 1 mm (E, D)." figureDoi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61">2D</figureCitation>
<figureCitation id="D51090A0563D01CC154FDA84A8141CEB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coccidula lepida LeConte SEM illustrations A pronotum B head and prothorax, ventral C meso and metaventrite D elytra dorsal E ventrite 1. Scale bars: 500 µm (A-C); 1 mm (E, D)." figureDoi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61">2D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3A</figureCitation>
<figureCitation id="46C2178C4E7EB7500FC65D75DEC26B02" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3A</figureCitation>
,
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<figureCitation id="6ADE0B9CF3CDEAA68DED5F1C311782EB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5A</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7A</figureCitation>
<figureCitation id="7E5824BC965C4E20F62547077B5A5386" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7A</figureCitation>
) (except
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<emphasis italics="true" pageId="0" pageNumber="61">C. scutellata</emphasis>
<taxonomicName id="F4F2993144840B6FDD5DAC3D24BE3CAB" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. scutellata" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="scutellata">
<emphasis id="EE54EEEC42C890E5F0ACDC181C1F36FA" italics="true" pageId="0" pageNumber="61">C. scutellata</emphasis>
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where it is obscured in basal part, Fig.
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<figureCitation id="9350E6D90EE967B95CA59D3B54486ACA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9A</figureCitation>
), surface covered with punctures of double size, smaller irregularly distributed, larger punctures arranged in nine irregular longitudinal rows. Sutural stria absent. Elytral epipleuron narrow, incomplete, reaching base of ventrite 4 (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1D</figureCitation>
<figureCitation id="F4AE50DD29E946A7B430B12B3BEBA6B0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1D</figureCitation>
), with complete bordering line, epipleural foveae absent. Hind wings fully developed or missing (in
<taxonomicName family="Coccinellidae" kingdom="Animalia" lsidName="C. litophiloides" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="litophiloides">
<emphasis italics="true" pageId="0" pageNumber="61">C. litophiloides</emphasis>
<taxonomicName id="AD4AD2707F6DA86DB28971AFAA33DC34" class="Insecta" family="Coccinellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. litophiloides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="litophiloides">
<emphasis id="A23D218903E3A2C716AC27ED52EC2048" italics="true" pageId="0" pageNumber="61">C. litophiloides</emphasis>
</taxonomicName>
). Metaventral postcoxal lines roundly joined medially, complete laterally, straight or descending (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coccidula lepida LeConte SEM illustrations A pronotum B head and prothorax, ventral C meso and metaventrite D elytra dorsal E ventrite 1. Scale bars: 500 µm (A-C); 1 mm (E, D)." figureDoi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61">2C</figureCitation>
<figureCitation id="F13723461C7DE61CC4A9E3B9A6236AB3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coccidula lepida LeConte SEM illustrations A pronotum B head and prothorax, ventral C meso and metaventrite D elytra dorsal E ventrite 1. Scale bars: 500 µm (A-C); 1 mm (E, D)." figureDoi="10.3897/zookeys.1043.65829.figure2" httpUri="https://binary.pensoft.net/fig/554377" pageId="0" pageNumber="61">2C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3D</figureCitation>
<figureCitation id="0C5AB8731C832CF0201F5A4F534A5980" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coccidula litophiloides Reitter SEM illustrations, paratype HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E head ventral. Scale bars: 1 mm (A); 500 µm (B-D); 400 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure3" httpUri="https://binary.pensoft.net/fig/554378" pageId="0" pageNumber="61">3D</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5D</figureCitation>
<figureCitation id="462BC73FFC514DDFC2E7BE83475622D2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Coccidula reitteri Dodge SEM illustrations, HNHM A body, dorsal B pronotum C head and prothorax, ventral D meso and metaventrite E antenna. Scale bars: 1 mm (A); 500 µm (B-D); 300 µm (E)." figureDoi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61">5D</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7E</figureCitation>
<figureCitation id="22A3A1679D5455B3F317870F532619C3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7E</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9D</figureCitation>
<figureCitation id="9B6641977CCE16EFAEB2C43A582A5F8C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9D</figureCitation>
). Metaventrite with discrimen visible in posterior 2/3.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<paragraph id="1D7A96DE42943DF55751E845CD57A4C7" pageId="0" pageNumber="61">
Trochanters simple, subtriangular, without projection (figs 7E, 9D). Tibiae slightly expanded apically with one apical spur on forelegs, and two in mid and hind legs. Tarsi consisting of four tarsomeres, second tarsomere truncate apically; tarsal claws cleft apically (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9G</figureCitation>
<figureCitation id="6F4B0EB64190BBEBFBD66427F8C3612F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9G</figureCitation>
) with single empodial seta present.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<paragraph id="ECF0007D1CC9D7BE4C39CF202BCC1DF7" pageId="0" pageNumber="61">
Abdomen in both sexes with 6 ventrites (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1D</figureCitation>
<figureCitation id="86F75E6D8C1BF7F8E5D7E4D25C0825F1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1D</figureCitation>
); ventrite 1 about as long as ventrites 2-4 combined, ventrite 2 longer than ventrite 3, ventrites 3-5 subequal in length. Abdominal postcoxal lines (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7E</figureCitation>
<figureCitation id="7643B972CBD7234B50E7A2DFC7674E2C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7E</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9D</figureCitation>
<figureCitation id="A5AB36829AF748FDB9F23449EE4090AA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9D</figureCitation>
) separate medially, recurved and complete, reaching anterior margin of ventrite, posteriorly reaching about half length of ventrite 1. Ventrite 5 in female posteriorly rounded (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7F</figureCitation>
<figureCitation id="68768D3FCB9F9DA32C7026021235E267" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Coccidula rufa (Reitter) SEM illustrations A body, dorsal B pronotum C head and prothorax, ventral D antenna E mesoventrite, metaventrite and ventrite 1 F ventrites 4 - 6, female. Scale bars: 1 mm (A); 500 µm (B, C, E, F); 400 µm (D)." figureDoi="10.3897/zookeys.1043.65829.figure7" httpUri="https://binary.pensoft.net/fig/554383" pageId="0" pageNumber="61">7F</figureCitation>
), in male truncate (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9F</figureCitation>
<figureCitation id="32D9890C5215BBD2D5F8957A952A04CE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Coccidula scutellata (Reitter) SEM illustrations A body, dorsal B pronotum C head, ventral D mesoventrite, metaventrite and ventrite 1 E prosternum F ventrites 4 - 6, male G pro-tarsal claw. Scale bars: 1 mm (A); 500 µm (B, D, E); 400 µm (C, F); 100 µm (G)." figureDoi="10.3897/zookeys.1043.65829.figure9" httpUri="https://binary.pensoft.net/fig/554385" pageId="0" pageNumber="61">9F</figureCitation>
). Ventrite 6 rounded in both sexes.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">
<emphasis italics="true" pageId="0" pageNumber="61">Male terminalia</emphasis>
<paragraph id="B013CEE76E69C3A7A38FA70D07AF1246" pageId="0" pageNumber="61">
<emphasis id="BFD7DBA5057867675D1197A072A52C95" bold="true" pageId="0" pageNumber="61">
<emphasis id="9B1047677E0BAE8A3CC029DD6BFD0321" italics="true" pageId="0" pageNumber="61">Male terminalia</emphasis>
.
</emphasis>
Tegmen (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coccidula litophiloides Reitter A tegmen, lateral B tegmen, inner C penis, lateral D spermatheca. Scale bar: 500 µm (A-D)." figureDoi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61">4A, B</figureCitation>
<figureCitation id="9E385C6B7F243BE0E36804C4D2D1DE5C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coccidula litophiloides Reitter A tegmen, lateral B tegmen, inner C penis, lateral D spermatheca. Scale bar: 500 µm (A-D)." figureDoi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61">4A, B</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8A, B</figureCitation>
<figureCitation id="FDD4F883BCDC0ECB73B7D698A33E7CE9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8A, B</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10A, B</figureCitation>
<figureCitation id="F7C181DD994A0A9D91AEB4542EBA5638" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10A, B</figureCitation>
) symmetrical; parameres articulated with penis guide. Penis (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coccidula litophiloides Reitter A tegmen, lateral B tegmen, inner C penis, lateral D spermatheca. Scale bar: 500 µm (A-D)." figureDoi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61">4C</figureCitation>
<figureCitation id="EA6670EBBF610A9BFD3F3B2C29D3C5B2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coccidula litophiloides Reitter A tegmen, lateral B tegmen, inner C penis, lateral D spermatheca. Scale bar: 500 µm (A-D)." figureDoi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61">4C</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8C</figureCitation>
<figureCitation id="1909928705BFEC3F81906ECE6C8738B7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8C</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10C</figureCitation>
<figureCitation id="CABCAEB2D7CBC4B4FC762C61F6B0B8F2" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10C</figureCitation>
) slender, pointed apically; penis capsule asymmetrical with outer arm reduced, inner arm well developed. Apodeme of male sternum IX simple, not broadened apically (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8D</figureCitation>
<figureCitation id="DFBCB1811BB6699D54094F7AC62913DF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8D</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10D</figureCitation>
<figureCitation id="EF2DCD160C2990AD4A7A538250AD6F76" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10D</figureCitation>
). Tergite X broadly rounded, semicircular (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8D</figureCitation>
<figureCitation id="7D814CC4C4B90CAA42935141FF93535C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8D</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10D</figureCitation>
<figureCitation id="E20DFAA4506D19BD316BB597DFE10D0D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 4.</emphasis>
<taxonomicName authorityName="Reitter" authorityYear="1890" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula litophiloides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="litophiloides">
<emphasis italics="true" pageId="0" pageNumber="61">Coccidula litophiloides</emphasis>
<caption id="116674A37D2B00EF3BDAC13903F988DF" doi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61" start="Figure 4" startId="F4">
<paragraph id="00840A55E1ED3E572C4CF38CD56FDCE0" pageId="0" pageNumber="61">
<emphasis id="52A19140C29496635FEB2446CD63C9BD" bold="true" pageId="0" pageNumber="61">Figure 4.</emphasis>
<taxonomicName id="A03BCB6A84D9021E23ABDBE3D0CDF2AA" authorityName="Reitter" authorityYear="1890" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula litophiloides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="litophiloides">
<emphasis id="9FDB84E4BC23B7D1F99DAFF2B89C8583" italics="true" pageId="0" pageNumber="61">Coccidula litophiloides</emphasis>
</taxonomicName>
Reitter
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
<emphasis id="AACAF8F3790E9C8E83624CEE92514B02" bold="true" pageId="0" pageNumber="61">A</emphasis>
tegmen, lateral
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
<emphasis id="32704FF6E5F1A70BF2872534443C06C9" bold="true" pageId="0" pageNumber="61">B</emphasis>
tegmen, inner
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
<emphasis id="8354BF9055C4E26A342D279BFA1C8A91" bold="true" pageId="0" pageNumber="61">C</emphasis>
penis, lateral
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
<emphasis id="F8C2AC4AB65C29025D0F5A1FF1A3AD68" bold="true" pageId="0" pageNumber="61">D</emphasis>
spermatheca. Scale bar: 500
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="6D6D624EC1A41A50A315A8A26E9CD8CE" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="61">A-D</emphasis>
<emphasis id="B6D33DE423A05476313B9A0532C12667" bold="true" pageId="0" pageNumber="61">A-D</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 5.</emphasis>
<taxonomicName authorityName="Dodge" authorityYear="1938" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula reitteri" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis italics="true" pageId="0" pageNumber="61">Coccidula reitteri</emphasis>
<caption id="2BEADA7A2DE2E8AC3F9D848685E626C8" doi="10.3897/zookeys.1043.65829.figure5" httpUri="https://binary.pensoft.net/fig/554380" pageId="0" pageNumber="61" start="Figure 5" startId="F5">
<paragraph id="F11D012C05054E6AE790C305F45FADF3" pageId="0" pageNumber="61">
<emphasis id="2C3854F483A2775B604358A01FD38A22" bold="true" pageId="0" pageNumber="61">Figure 5.</emphasis>
<taxonomicName id="1EAB3D1827177C2507E6BA2C60742015" authorityName="Dodge" authorityYear="1938" class="Insecta" family="Coccinellidae" genus="Coccidula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccidula reitteri" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="D10603F13D2D13119FBD0DE651B9B18A" italics="true" pageId="0" pageNumber="61">Coccidula reitteri</emphasis>
</taxonomicName>
Dodge SEM illustrations, HNHM
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
<emphasis id="B448271B837C281E446B3AB397BAB505" bold="true" pageId="0" pageNumber="61">A</emphasis>
body, dorsal
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
<emphasis id="70F1B279CE2B472CE83C20472544C556" bold="true" pageId="0" pageNumber="61">B</emphasis>
pronotum
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
<emphasis id="28B381B3D48EE8D1518C0CA3C2611AA9" bold="true" pageId="0" pageNumber="61">C</emphasis>
head and prothorax, ventral
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
<emphasis id="E7638FF1499665D77B9C6EA7A8B387E4" bold="true" pageId="0" pageNumber="61">D</emphasis>
meso and metaventrite
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
<emphasis id="A71A66BF795D51B9259F253ADDB23300" bold="true" pageId="0" pageNumber="61">E</emphasis>
antenna. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
<emphasis id="253CB19A021406DEBEF96034C11C1027" bold="true" pageId="0" pageNumber="61">A</emphasis>
); 500
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="E53A54CDA262CDC28A3FA46F5A650EAB" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="61">B-D</emphasis>
<emphasis id="A198B5E4C51EC0B6F6F6467E53181A95" bold="true" pageId="0" pageNumber="61">B-D</emphasis>
); 300
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="B1C768FAFA9CB8281588B27EBCAE32B9" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
<emphasis id="96F07A52E6649555715884491734CD90" bold="true" pageId="0" pageNumber="61">E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">
<emphasis italics="true" pageId="0" pageNumber="61">Female terminalia</emphasis>
<paragraph id="6259F234A0366B872B69C12D2DAB9EFA" pageId="0" pageNumber="61">
<emphasis id="3D4BF22E6A4DFD782F5C438D38B5244C" bold="true" pageId="0" pageNumber="61">
<emphasis id="E44D0D3B72B7135E4E3DB3A5E7BF19FE" italics="true" pageId="0" pageNumber="61">Female terminalia</emphasis>
.
</emphasis>
Coxites (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10F</figureCitation>
<figureCitation id="C293BA15AFD564D9BABA06018ED1BCB1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10F</figureCitation>
) distinctly elongate, subtriangular; styli small but visible, bearing several short setae; infundibulum absent (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coccidula litophiloides Reitter A tegmen, lateral B tegmen, inner C penis, lateral D spermatheca. Scale bar: 500 µm (A-D)." figureDoi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61">4D</figureCitation>
<figureCitation id="80FF4A60E29BFADF896F125E0480C294" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coccidula litophiloides Reitter A tegmen, lateral B tegmen, inner C penis, lateral D spermatheca. Scale bar: 500 µm (A-D)." figureDoi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61">4D</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8E</figureCitation>
<figureCitation id="65544E26B8C77F794D9B41251D2FCAE3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8E</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10E</figureCitation>
<figureCitation id="0493F793DEC8A39A435B06FD7E894871" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10E</figureCitation>
); sperm duct simple. Spermatheca (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coccidula litophiloides Reitter A tegmen, lateral B tegmen, inner C penis, lateral D spermatheca. Scale bar: 500 µm (A-D)." figureDoi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61">4D</figureCitation>
<figureCitation id="69CEF3F23974A6DDBAD29B363265C3A7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coccidula litophiloides Reitter A tegmen, lateral B tegmen, inner C penis, lateral D spermatheca. Scale bar: 500 µm (A-D)." figureDoi="10.3897/zookeys.1043.65829.figure4" httpUri="https://binary.pensoft.net/fig/554379" pageId="0" pageNumber="61">4D</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8E</figureCitation>
<figureCitation id="F8E6F858AFD917F4C1D372F75489CF5B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Coccidula rufa (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca. Scale bar: 500 µm (A-E)." figureDoi="10.3897/zookeys.1043.65829.figure8" httpUri="https://binary.pensoft.net/fig/554384" pageId="0" pageNumber="61">8E</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10E</figureCitation>
<figureCitation id="0306F190E5BE18105C07907A72FD3370" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10E</figureCitation>
) worm-like, without clear ramus or nodulus; spermathecal accessory gland small, elongate. Proctiger elongate, rounded apically (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10F</figureCitation>
<figureCitation id="38CE601F07A4C68AF392A2C6BCE29F2C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Coccidula scutellata (Reitter) A tegmen, inner B tegmen, lateral C penis, lateral D male genital segment, dorsal E spermatheca F female genitalia G left and right mandibles H maxilla I metendosternite. Scale bars: 500 µm (A-F); 200 µm (G-I)." figureDoi="10.3897/zookeys.1043.65829.figure10" httpUri="https://binary.pensoft.net/fig/554386" pageId="0" pageNumber="61">10F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="immature stages">
<paragraph pageId="0" pageNumber="61">Immature stages.</paragraph>
<paragraph pageId="0" pageNumber="61">
<subSubSection id="E47CEA825160FD9E0EBA47A0E7F9AC9D" pageId="0" pageNumber="61" type="immature stages">
<paragraph id="1C8998F6488D33FFC1147EB8C9541D8F" pageId="0" pageNumber="61">Immature stages.</paragraph>
<paragraph id="ED897F6DBC852A83C12F8238F9DB01E9" pageId="0" pageNumber="61">
Larva as in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1A</figureCitation>
<figureCitation id="D044D3877D96084A85D3C64D83A4E667" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1A</figureCitation>
, pupa as in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1B</figureCitation>
<figureCitation id="9F568DA56599AB09503CD63C81374214" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Immature stages and habitus of adult species of Coccidula Kugelann A C. rufa larva B C. rufa pupa C C. lepida, dorsal D C. lepida, ventral E C. litophiloides F C. reitteri G C. rufa H C. scutellata A, B Gilles San Martin, Wikimedia Commons C, D Danny Haelewaters G, H Udo Schmidt. Scale bars: 1 mm (C-H)." figureDoi="10.3897/zookeys.1043.65829.figure1" httpUri="https://binary.pensoft.net/fig/554376" pageId="0" pageNumber="61">1B</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="distribution">
<paragraph pageId="0" pageNumber="61">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="61">Holarctic: Asia, Europe, Africa (North), North America.</paragraph>
<subSubSection id="A5CABD74BC353C6A92EC232BCC5117F3" pageId="0" pageNumber="61" type="distribution">
<paragraph id="4E83AB8E3960909EB6FF9DAD55BE29BB" pageId="0" pageNumber="61">Distribution.</paragraph>
<paragraph id="725FBAFEE8F16D9BDC1E20A98445279F" pageId="0" pageNumber="61">Holarctic: Asia, Europe, Africa (North), North America.</paragraph>
</subSubSection>
</treatment>
</document>