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<document id="AE5C1EAF81A151329D6BA100158E055F" ID-DOI="10.3897/phytokeys.250.136139" ID-publisher-id="136139" URI-arpha="163F7496-5C12-5455-AB48-30AD78073DF0" article-type="research-article" checkinTime="1734707767354" checkinUser="pensoft" docAuthor="Sukhorukov, Alexander P., Kushunina, Maria, Nilova, Maya V., Baider, Cláudia &amp; Sennikov, Alexander N." docDate="2024" docId="7DA71355D55D5B14B346A5210CE0F8C1" docLanguage="en" docName="PhytoKeys 250: 21-94" docOrigin="PhytoKeys 250" docSource="https://phytokeys.pensoft.net/article/136139/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Achyranthes mauritiana Moq." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="94" masterDocId="163F74965C125455AB4830AD78073DF0" masterDocTitle="Africa and Arabia encompass a much greater species diversity in the Achyranthes aspera aggregate (Amaranthaceae, achyranthoid clade): Evidence from morphological and chorological data" masterLastPageNumber="94" masterPageNumber="21" pageNumber="21" updateTime="1734708299152" updateUser="ExternalLinkService">
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<mods:title id="ECDBB5D2DF92B6648C7D830D870B3BF1">Africa and Arabia encompass a much greater species diversity in the Achyranthes aspera aggregate (Amaranthaceae, achyranthoid clade): Evidence from morphological and chorological data</mods:title>
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<mods:namePart id="CA8E87BC4A6214E9F70D2F4BA2606464">Sukhorukov, Alexander P.</mods:namePart>
<mods:nameIdentifier id="0F4B1876D9A5F201D12FAABE2C40BBA8" type="ORCID">0000-0003-2220-826X</mods:nameIdentifier>
<mods:affiliation id="0407EE30846EC3309F53FC2C0D9B685A">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia &amp; Laboratory Herbarium (TK), Tomsk State University, Lenin Ave. 36, 634050, Tomsk, Russia</mods:affiliation>
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<mods:namePart id="FA6B18AC2BD513ED256AA99DC89B1F3C">Kushunina, Maria</mods:namePart>
<mods:nameIdentifier id="3DEEBB42113B216E519982F79D08AF0D" type="ORCID">0000-0003-2541-6494</mods:nameIdentifier>
<mods:affiliation id="43554BA33C8B5BF97ED3138FA5FB15D7">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
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<mods:namePart id="D664B35A000F60879DB89D6D2141FE72">Nilova, Maya V.</mods:namePart>
<mods:nameIdentifier id="521FAC3E78C9A2089B4D697122F74D0C" type="ORCID">0000-0001-5340-5915</mods:nameIdentifier>
<mods:affiliation id="DF8262058CA360BE718318C92AAC3951">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
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<mods:namePart id="7E8D8BC5309FAF9468777F1FF01FF86B">Baider, Cláudia</mods:namePart>
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<mods:affiliation id="2C3C221DFBD434AC6B59D22FA1E4BF25">Department of Plant Physiology, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
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<mods:namePart id="5239582095FE6EF5FBDCE52A1F7C8EF7">Sennikov, Alexander N.</mods:namePart>
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<mods:affiliation id="83BB30BB50FEA4C2F1A8A1E1E3AE3ED5">The Mauritius Herbarium, RE Vaughan Building, Agricultural Services, Ministry of Agro-Industry and Food Security, Réduit, 80835, Mauritius</mods:affiliation>
</mods:name>
<mods:typeOfResource id="1C92660CDF2B28C66C60C4D1F9DC682D">text</mods:typeOfResource>
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<mods:title id="112DBDAA4445C4B25AD266611318AAF5">PhytoKeys</mods:title>
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<mods:date id="E544F4E9BBA3ADACC02F3F32DFD0CD26">2024</mods:date>
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<mods:number id="F5C28E975568174E5723C68A4CE6E06F">250</mods:number>
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<treatment id="7DA71355D55D5B14B346A5210CE0F8C1" ID-DOI="http://doi.org/10.5281/zenodo.14536758" ID-Zenodo-Dep="14536758" ID-arpha="7DA71355-D55D-5B14-B346-A5210CE0F8C1" LSID="urn:lsid:plazi:treatment:7DA71355D55D5B14B346A5210CE0F8C1" httpUri="http://treatment.plazi.org/id/7DA71355D55D5B14B346A5210CE0F8C1">
<subSubSection id="5A0E0D77BE04DA837A721D4719F6EDBC" type="nomenclature">
<paragraph id="6EB6AAA14F8185407FFC42E1E2EB25CB">
<taxonomicName id="3D0A36261F87EAC4D36A816F7DE07C4A" ID-CoL="999J" authority="Moq." authorityName="Moq." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="mauritiana">
<emphasis id="2B1568C2F0639F808314A34DC80AB2AE" italics="true">Achyranthes mauritiana</emphasis>
Moq.
</taxonomicName>
in DC., Prodr. 13 (2): 313 (1849).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBIAK" type="Lectotype (designated here)">
<paragraph id="B320B40DBB80F181956C5F46B9F901E2">
<heading id="87B3935AE62825409F71C897C5162E47" reason="title">
<typeStatus id="9966C801924B349437DECEC4ECB52F82" type="lectotype">Lectotype</typeStatus>
(designated here).
</heading>
</paragraph>
<paragraph id="A49701B4A00A559BE1FFD37E02EEBC8D">
<materialsCitation id="54935639C295643E7385E885F65E5E9D" collectionCode="P, G" country="Mauritius" location="Mauritius" specimenCode="P 00487020, FI-WEBB 154894, G 00688999" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="93FCC5043C74038048F81FE92393D31C" name="Mauritius">Mauritius</collectingCountry>
, [17681773], [
<emphasis id="16ED7B9C953F24DF19C503A2CE6A3715" italics="true">Commerson</emphasis>
]
<emphasis id="C42FCBFB91DFE1C269FD1E3E762C2735" italics="true">s. n</emphasis>
. (
<specimenCode id="A1BA27FC6CC19257527EDB2462F6F388" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P 00487020</specimenCode>
[image!];
<typeStatus id="15C7DDFBCC6E169E1A135DD5D12FDF9F" type="isolectotype">isolectotypes</typeStatus>
<specimenCode id="F2DE824C619D752E5F18B930472CDB7F" collectionCode="FI-WEBB">FI-WEBB 154894</specimenCode>
!, G-DC
<specimenCode id="BB72E820320CCC0B3464617EDE717070" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G 00688999</specimenCode>
[image!]).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELIAK" type="description">
<paragraph id="1FCA822671708583D421EB3E340B7236">
<heading id="79D07A02CDF7885A475EFEC1902E63AE" reason="title">Description.</heading>
</paragraph>
<paragraph id="141DB7A8D44C70346E28A4090E1E9F8D">
(Fig.
<figureCitation id="520277B205F24410FC3DE97A34AB01E6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A herbarium specimen of Achyranthes mauritiana (Réunion, Rivière des Galets, Jul 1851, Boivin s. n., MAU 0017619)." figureDoi="10.3897/phytokeys.250.136139.figure13" httpUri="https://binary.pensoft.net/fig/1203857">13</figureCitation>
). Annual (?); stem upright, with spreading lateral branches; leaf petioles
<quantity id="E92D7DD663CB94614D28D7C2CB585BE1" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" unit="mm" value="30.0" valueMax="40.0" valueMin="20.0">2040 mm</quantity>
long, blades 30100 ×
<quantity id="D6DD617137E37EA9230B43D7D11049BE" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" unit="mm" value="30.0" valueMax="40.0" valueMin="20.0">2040 mm</quantity>
, rhombic or elliptic, tip attenuate, almost glabrous or sparsely pubescent at the veins, green, sometimes turning black when dry; inflorescence up to
<quantity id="E4ACC9FBFD05BF3B70B95E27EB027480" metricMagnitude="-1" metricUnit="m" metricValue="3.0" unit="cm" value="30.0">30 cm</quantity>
long; bract 2.53.0 mm long, bracteoles 3.0
<quantity id="050C4E85E1A3B31596630A745A60F7D3" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
; perianth 4.05.0 mm long; stamens 5, anthers ~
<quantity id="6979C7C8A95C3176CF4992D22FFD07E4" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.4 mm</quantity>
long; style 1.0
<quantity id="8E80797DB7207E13996A3FBF3E19D53C" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
long; fruit
<quantity id="9113E9F4CFD23EF9D96542B00F9238A6" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.72.1 mm</quantity>
long.
</paragraph>
<caption id="EBEBFD10B616B9722A4552B2184AA397" ID-DOI="10.3897/phytokeys.250.136139.figure13" ID-arpha="1C53C6A4-B038-553A-A315-AF10CD5E7308" httpUri="https://binary.pensoft.net/fig/1203857" startId="F13">
<paragraph id="389537740B05F16DAB62F6AA2C827072">
<label id="E300B0AA1DCD0B5A75BA376934562A0F">Figure 13.</label>
</paragraph>
<paragraph id="C3D7879DAA85A1F2410A19EA1872A906">
A herbarium specimen of
<taxonomicName id="7E21FFA87A91176F8314B602FEC50696" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="mauritiana">
<emphasis id="18E0F6AF00CB795E52093996DC9FD900" italics="true">Achyranthes mauritiana</emphasis>
</taxonomicName>
(Réunion, Rivière des Galets, Jul 1851,
<emphasis id="08D73CB8B9E66C433DFD9A8C61DBEC6E" italics="true">Boivin s. n.</emphasis>
, MAU 0017619).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EPJAK" type="Taxonomic notes">
<paragraph id="6DC0A1FCF59859759C506E384B7501E5">
<heading id="80722DB1CB77AA8973E288298029E5A1" reason="title">Taxonomic notes.</heading>
</paragraph>
<paragraph id="C4F8B2400B452E93EB7DF7112AEE05DE">
The specimens seen are represented by upper twigs only, and our present description is based on the scarce material available in the collections
<collectionCode id="B56DE5C04A7B33BFE1BA317B68DED0E1">FI-WEBB</collectionCode>
and
<collectionCode id="9693D8313CB98D28EE438D84E55D4B3A" country="Mauritius" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13560" lsid="urn:lsid:biocol.org:col:13560" name="The Mauritius Herbarium" type="Herbarium">MAU</collectionCode>
.
<collectionCode id="14EA8172BBE476584A6D2384FFBE56D7" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
description provided by
<bibRefCitation id="24685937759E1551545B85635121FCD3" author="Cavaco A" editor="Humbert H" firstAuthor="Cavaco" journalOrPublisher="Typ. Firmin-Didot &amp; Co., Paris" pagination="1-56" refId="B23" refString="Cavaco A (1954) Amaranthaceae. In: Humbert H (Ed.) Flore de Madagascar et des Comores. Typ. Firmin-Didot &amp; Co., Paris, 156." title="Amaranthaceae." volumeTitle="Flore de Madagascar et des Comores." year="1954">Cavaco (1954)</bibRefCitation>
is insufficient for any comparison of the material from the Mascarenes,
<collectingCountry id="D93811CF109D6ED52A999BE508053D36" name="Madagascar">Madagascar</collectingCountry>
and
<collectingCountry id="7741AA16C1A8F71D5B3BC821129B504A" name="Comoros">Comoros</collectingCountry>
, because the morphometry of the reproductive characters was not indicated. So far,
<taxonomicName id="3DB7E69D498A05F4AE51C46B2F58E8C2" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="mauritiana">
<emphasis id="BDBA55A5F4646BF4D606706AE30BC1F9" italics="true">A. mauritiana</emphasis>
</taxonomicName>
is still poorly known.
<bibRefCitation id="CDDE30F4AA24A0F01BF0539B2C19E42B" author="Townsend CC" editor="Bosser J &amp; Cadet Th &amp; Guèho J &amp; Marais W" firstAuthor="Townsend" journalOrPublisher="La Réunion, Maurice, Rodrigues. Kew, ORSTOM, Imprimerie Gouvernement &amp; Centre Universitaire, London - Paris - Port-Louis - Saint-Denis" pagination="1-29" refId="B124" refString="Townsend CC (1994) Amaranthacées. In: Bosser J, Cadet Th, Guèho J, Marais W (Eds) Flore des Mascareignes. La Réunion, Maurice, Rodrigues. Kew, ORSTOM, Imprimerie Gouvernement &amp; Centre Universitaire, London Paris Port-Louis Saint-Denis, 129." title="Amaranthacées." volumeTitle="Flore des Mascareignes." year="1994">Townsend (1994)</bibRefCitation>
synonymized
<taxonomicName id="F8A7CBA82DAAE3A1A99C6F71E97F2BB2" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="mauritiana">
<emphasis id="9FE1C4BA8F7010629EF02DE726A3BF97" italics="true">A. mauritiana</emphasis>
</taxonomicName>
with
<taxonomicName id="41220AE634837B990CB6CEB2354DC53A" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="porphyrostachya">
<emphasis id="9F9E052344059047C47B87B4562D17B2" italics="true">A. aspera var. porphyrostachya</emphasis>
</taxonomicName>
[≡
<taxonomicName id="90C34AE7E063EAC07CE623A2325A192D" baseAuthorityName="Lebrun" baseAuthorityYear="1973" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="porphyrostachya">
<emphasis id="F958F1F520F5641B65886152E0D8E8FC" italics="true">A. porphyrostachya</emphasis>
</taxonomicName>
], but this synonymy cannot be accepted due to their different leaf shape and morphometry of the reproductive characters. Only a few old collections cited below were from the Mascarenes, with a more recent collection from Agaléga (Mauritian territory). Nevertheless,
<taxonomicName id="F054064A16D28480507CF2BB649B4605" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="mauritiana">
<emphasis id="EC552AA62B0C481B12217BF4753BB489" italics="true">A. mauritiana</emphasis>
</taxonomicName>
seems to be a distinct species due to its rhombic or elliptic, almost glabrous leaves as compared with
<taxonomicName id="86A62E93F3EA0F3DB65ECA61D678719E" baseAuthorityName="Ghazanfar" baseAuthorityYear="1992" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="6F4E7869C8AA994020EB9CF20FB689CE" italics="true">A. aspera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="930CA5401321208A70E90861D49C5806" sensu="stricto">s. str.</taxonomicNameLabel>
Its reproductive characters are the same as in
<taxonomicName id="6A0D0726353D4613BF27CE3CA2C93F0F" baseAuthorityName="Ghazanfar" baseAuthorityYear="1992" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="067B8FC713370C1E6CBBA6B3585311CD" italics="true">A. aspera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C5FFB55017E91F37FBC8C968445A33E" sensu="stricto">s. str.</taxonomicNameLabel>
The leaf characters are quite similar to those of
<taxonomicName id="7C85DA6B4C3BC9EAA4C18B62FD70BBA5" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="acuminata">
<emphasis id="AB89FBBE4A58F2BA5947158B5F0D3AEB" italics="true">A. acuminata</emphasis>
</taxonomicName>
, but the morphometry of the reproductive characters is different. Further studies are needed to explore the morphology, phylogenetic relationships and ecology of
<taxonomicName id="AACD7BF170085D91A56C0DF10B32EC76" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="mauritiana">
<emphasis id="4229B06C1F754C94F02737D37F436310" italics="true">A. mauritiana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1F8C5588E4C9726040EDD5445AEFC15F">
<bibRefCitation id="C9D58A07764A6BB132754A7C3539288A" author="Townsend CC" editor="Bosser J &amp; Cadet Th &amp; Guèho J &amp; Marais W" firstAuthor="Townsend" journalOrPublisher="La Réunion, Maurice, Rodrigues. Kew, ORSTOM, Imprimerie Gouvernement &amp; Centre Universitaire, London - Paris - Port-Louis - Saint-Denis" pagination="1-29" refId="B124" refString="Townsend CC (1994) Amaranthacées. In: Bosser J, Cadet Th, Guèho J, Marais W (Eds) Flore des Mascareignes. La Réunion, Maurice, Rodrigues. Kew, ORSTOM, Imprimerie Gouvernement &amp; Centre Universitaire, London Paris Port-Louis Saint-Denis, 129." title="Amaranthacées." volumeTitle="Flore des Mascareignes." year="1994">Townsend (1994)</bibRefCitation>
reported
<taxonomicName id="9220AC2142B031DC7EFF024ADB59F9E8" authority="(Hook. &amp; Arn.) C. C. Towns." authorityName="C. C. Towns." baseAuthorityName="Hook. &amp; Arn." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="velutina">
<emphasis id="3B4E27376DBC263BA63FC180667CF0CD" italics="true">A. aspera var. velutina</emphasis>
(Hook. &amp; Arn.) C. C. Towns.
</taxonomicName>
(≡
<taxonomicName id="3716E527D5D4F8884329672FD5A478F6" authority="Hook. &amp; Arn." authorityName="Hook. &amp; Arn." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="velutina">
<emphasis id="91D70A8438A64D212943C7A329E85D24" italics="true">A. velutina</emphasis>
Hook. &amp; Arn.
</taxonomicName>
) from
<collectingCountry id="368104F670DA93F0C9310F7B5B1A83B1" name="Réunion">Réunion</collectingCountry>
. This robust species with villous leaves occurs only in the Pacific. The plants from the Mascarenes resembling
<taxonomicName id="995C7B942984BC4DD028D65C9D4F9B98" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="velutina">
<emphasis id="3E3CE3D87DC525F72FA0919A32AE4951" italics="true">A. velutina</emphasis>
</taxonomicName>
seem to belong to a different species whose name may be
<taxonomicName id="BA771BA11A15F7D1DF663646B864C218" authority="Willd." authorityName="Willd." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="borbonica">
<emphasis id="2F1AC1B7F1488BAF78F3E3B2ECCE3B6F" italics="true">A. borbonica</emphasis>
Willd.
</taxonomicName>
in Roem. &amp; Schult., Syst. Veg., ed. 15 bis 5: 549 (1819) ≡
<treatmentCitation id="BAD0974E72ABA7E776FEFEDC7E48F520" author="C. C. Towns." issue="3" journal="Kew Bull." page="474" score="22.239024375395413" volume="29" year="1974">
<taxonomicName id="00E5FEA55CAB5EE70FDB2ADD67B972A1" ID-CoL="5LYB8" authority="(Willd. ex Schult.) C. C. Towns., Kew Bull." authorityName="C. C. Towns., Kew Bull." baseAuthorityName="Schult." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="borbonica">
<emphasis id="58B9E381A99DA2AFA4178683EC5AF254" italics="true">A. aspera var. borbonica</emphasis>
(Willd. ex Schult.) C. C. Towns., Kew Bull.
</taxonomicName>
29 (3): 474 (1974)
</treatmentCitation>
. The
<typeStatus id="C4B9E4AE373BCD73AB27CA40FF2A5CB6" type="holotype">holotype</typeStatus>
of
<taxonomicName id="34A9E654F49E69CC2B4409C34CC22D5A" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="borbonica">
<emphasis id="8CDAEB76DBB361E3D324A848029A5387" italics="true">A. borbonica</emphasis>
</taxonomicName>
(“ Insula Borbon ” [
<collectingCountry id="4FA94B9BAE4EF27D6C5963755CF30493" name="Réunion">Réunion</collectingCountry>
]) kept at B-W 05-005 - 010 is represented by a small fragment showing villous leaves and an inflorescence collected in a premature stage. This species is also very poorly known, and no other material except for the type of
<taxonomicName id="05FAF960D12BFE86E87CEEF5EB1CA1AB" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="borbonica">
<emphasis id="9904E7FB0210F9027D01AED9C6187B4A" italics="true">A. borbonica</emphasis>
</taxonomicName>
is seen from the Mascarenes.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFQAK" type="Nomenclatural notes">
<paragraph id="0D801C7FFAD7617DF660A732DA73A95F">
<heading id="E81E3EC2DD57A21A95BDC3924792E05F" reason="title">Nomenclatural notes.</heading>
</paragraph>
<paragraph id="7FEA4950F5679442F06C9318D5048650">
<bibRefCitation id="43053C1CEFD271E14762542DBC782B79" author="Moquin-Tandon CHBA" editor="de Candolle AP" firstAuthor="Moquin-Tandon" journalOrPublisher="13, part 2. Treuttel &amp; Würtz, Paris" pagination="231-424" refId="B84" refString="Moquin-Tandon CHBA (1849) Amaranthaceae. In: de Candolle AP (Ed.) Prodromus systematis naturalis regni vegetabilis, vol. 13, part 2. Treuttel &amp; Würtz, Paris, 231424." title="Amaranthaceae." volumeTitle="Prodromus systematis naturalis regni vegetabilis, vol." year="1849">Moquin-Tandon (1849)</bibRefCitation>
based his
<taxonomicName id="42F4452C88D55E8A0611D524136CFFF0" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="mauritiana">
<emphasis id="C4506E5BF6A1D52F4DC9535B1B5E886F" italics="true">A. mauritiana</emphasis>
</taxonomicName>
on
<specimenCount id="40BAA675B4BE70665FA86AA16BAA25CF" count="2" type="generic">two specimens</specimenCount>
from
<collectingCountry id="2E35D8BD0810058931B2DF79DD30A89A" name="Mauritius">Mauritius</collectingCountry>
without exact label information (seemingly collected by
<collectionCode id="6443CD15E7D43E490F71FFE7761C5224" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P</collectionCode>
. Commerson during his residence on
<collectingCountry id="5CCCF19DF01702013F19C6D8E3B62486" name="Mauritius">Mauritius</collectingCountry>
in 17681773:
<bibRefCitation id="993E29D3C6D08DD435C3A739103F6293" DOI="10.1017/CBO9780511541803" author="Frodin" firstAuthor="Frodin" refId="B40" refString="Frodin DG (2001) Guide to standard floras of the world: An annotated, geographically arranged systematic bibliography of the principal floras, enumerations, checklists and chorological atlases of different areas. 2 nd ed. Cambridge University Press, Cambridge. https://doi.org/10.1017/CBO9780511541803" year="2001">Frodin 2001</bibRefCitation>
), which were kept in the Paris Museum of Natural History and the personal herbarium of Labillardière.
<bibRefCitation id="F2C735A1D37E666D0C74CAFA2D22FB4D" author="Townsend CC" editor="Bosser J &amp; Cadet Th &amp; Guèho J &amp; Marais W" firstAuthor="Townsend" journalOrPublisher="La Réunion, Maurice, Rodrigues. Kew, ORSTOM, Imprimerie Gouvernement &amp; Centre Universitaire, London - Paris - Port-Louis - Saint-Denis" pagination="1-29" refId="B124" refString="Townsend CC (1994) Amaranthacées. In: Bosser J, Cadet Th, Guèho J, Marais W (Eds) Flore des Mascareignes. La Réunion, Maurice, Rodrigues. Kew, ORSTOM, Imprimerie Gouvernement &amp; Centre Universitaire, London Paris Port-Louis Saint-Denis, 129." title="Amaranthacées." volumeTitle="Flore des Mascareignes." year="1994">Townsend (1994)</bibRefCitation>
designated a specimen at G-DC as
<typeStatus id="E47E3FE756AC555DFDE89E13943F35AE" type="lectotype">lectotype</typeStatus>
. This specimen belongs to the collection from
<collectingCountry id="974AD11DE54D6C9E2CB7662D3A93CC6C" name="Mauritius">Mauritius</collectingCountry>
, which was previously kept in Paris but obtained by
<collectionCode id="DFCCC49EEF25E24BE0C74CCFEFD9D763" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
.
<collectionCode id="0B95ED1F9F996788A96D257EA339E508" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P</collectionCode>
. de Candolle for his private herbarium by exchange in 1821 (cf.
<bibRefCitation id="434828E8E5508591345614556631E1E0" author="de Candolle" firstAuthor="de Candolle" journalOrPublisher="Cherbuliez, Genève &amp; Paris" refId="B27" refString="de Candolle A (1862) Mémoires et souvenirs de Augustin-Pyramus de Candolle, écrits par lui-même et publiés par son fils. J. Cherbuliez, Genève &amp; Paris, 618 pp." title="Mémoires et souvenirs de Augustin-Pyramus de Candolle, écrits par lui-même et publiés par son fils. J." year="1862">de Candolle 1862</bibRefCitation>
). Although this specimen is part (a duplicate) of the collection which was used for the original description, it was not cited in the protologue and therefore has no precedence over cited
<typeStatus id="E636DDF73D4CE547E602C8678C83A36A" type="syntype">syntypes</typeStatus>
in
<typeStatus id="2561D6B17DDD96F2993E7C2383370F65" type="lectotype">lectotype</typeStatus>
designation.
<collectionCode id="079AD64767FA23141CC317CDA3CD7F85" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
specimen at
<collectionCode id="38DEC2FDA51E957CB857A48858B3DDB4" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P</collectionCode>
, which was examined by Moquin-Tandon and is accompanied with his analytical drawing, is designated here as a new
<typeStatus id="73B6172B3E0891234270AE9C98CE6D29" type="lectotype">lectotype</typeStatus>
in agreement with Art. 9.12.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMRAK" type="distribution">
<paragraph id="5F02312897D82079360BF69D4758CC59">
<heading id="49F08282EE0AAE21E91340EAE6EE210F" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="4751E8BDDB22934F9CD0F7F5DA255D73">
<emphasis id="BF13A2106974492E001664AD035B1C9D" bold="true">
<collectingCountry id="429C6595D9AB31962F10995CCE423428" name="Mauritius">Mauritius</collectingCountry>
</emphasis>
: [without exact location and date]
<emphasis id="7BE4C76F9D3CFCFAA3EA49403498640D" italics="true">herb</emphasis>
.
<emphasis id="01D90311A754900E8722A89CC580901C" italics="true">Moquin-Tandon s. n</emphasis>
. (
<collectionCode id="A004457989DF1424258D41FB21DDDE38">FI-WEBB</collectionCode>
154894);
<collectingRegion id="8C1CA90C624E771EDA1FAA632D303606" country="Mauritius" name="Agalega Islands">Agaléga</collectingRegion>
, North Island,
<date id="18628061670EAA6637E2CE8C8009588B" value="1976-07-19">19 Jul 1976</date>
,
<emphasis id="D94F89C16D9701211F4647F7C91E4183" italics="true">
D.
<collectionCode id="75412195225DEB01A95A02CC32012F11" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Stoddart 7237
</emphasis>
(
<collectionCode id="4A14CAAF755499534CFC4689FB248770" country="South Africa" name="University of Stellenbosch">US</collectionCode>
03544411);
</paragraph>
<paragraph id="0DC8DA50249E1BDEB1505F6086384327">
<emphasis id="B0A641F6F10FDCBB4914E84F9D782A9B" bold="true">
<collectingCountry id="EC91BA70B7057A270CF9DAC6DADC6335" name="Réunion">Réunion</collectingCountry>
</emphasis>
(French Overseas Department): Rivière des Galets,
<date id="74139DF3FCD5CD94F2591CED3731CFF4" value="1851-07">Jul 1851</date>
,
<emphasis id="66974BC7602DC55352A2EFCA2BC538B2" italics="true">Boivin s. n.</emphasis>
(
<collectionCode id="6AB156BC992BA51B394842A157FF0435" country="Mauritius" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13560" lsid="urn:lsid:biocol.org:col:13560" name="The Mauritius Herbarium" type="Herbarium">MAU</collectionCode>
0017619).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EASAK" type="distribution">
<paragraph id="6BDAE8B0D21B1BFB3263615B7CE48283">
<heading id="CE273547AF4047AD65BB3FF4A85BE522" reason="title">General distribution.</heading>
</paragraph>
<paragraph id="36DAC234C621F9B62742C457B237704E">
The species seems to be endemic to the Mascarenes (
<collectingCountry id="3E5EE0169C7C0CD78196FFDE85AB65F8" name="Mauritius">Mauritius</collectingCountry>
and
<collectingCountry id="69FC5D681B19B36D749685A4CAB41C82" name="Réunion">Réunion</collectingCountry>
), but recently collected in Agaléga.
<bibRefCitation id="4DE6FC4A3715D37E28ACC0B9CB5D4418" author="Cavaco A" editor="Humbert H" firstAuthor="Cavaco" journalOrPublisher="Typ. Firmin-Didot &amp; Co., Paris" pagination="1-56" refId="B23" refString="Cavaco A (1954) Amaranthaceae. In: Humbert H (Ed.) Flore de Madagascar et des Comores. Typ. Firmin-Didot &amp; Co., Paris, 156." title="Amaranthaceae." volumeTitle="Flore de Madagascar et des Comores." year="1954">Cavaco (1954)</bibRefCitation>
cited some records from
<collectingCountry id="D39AC8E52F35CBF217475092E144EA01" name="Madagascar">Madagascar</collectingCountry>
, but all material seen belongs to other species.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,278 @@
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<mods:title id="D20FC51F8A388F30302B0F17363CE68C">Africa and Arabia encompass a much greater species diversity in the Achyranthes aspera aggregate (Amaranthaceae, achyranthoid clade): Evidence from morphological and chorological data</mods:title>
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<treatment id="8B77D03B00BB56F5B085321BAF8D7103" ID-DOI="http://doi.org/10.5281/zenodo.14536765" ID-Zenodo-Dep="14536765" ID-arpha="8B77D03B-00BB-56F5-B085-321BAF8D7103" LSID="urn:lsid:plazi:treatment:8B77D03B00BB56F5B085321BAF8D7103" httpUri="http://treatment.plazi.org/id/8B77D03B00BB56F5B085321BAF8D7103">
<subSubSection id="7696C5D39B73FCF92771670E8ED60C61" type="nomenclature">
<paragraph id="78E200CF0FB820A6BD448988024E3436">
<treatmentCitation id="DFB5D8D949FFB3931D7B39CD88BB8D3F" author="C. C. Towns." issue="3" journal="Kew Bull." page="424" score="22.239024375395413" volume="34" year="1980">
<taxonomicName id="0D83F4634E68625932AF7CA96CD7F1C9" ID-CoL="997Q" authority="(Suess.) C. C. Towns., Kew Bull." authorityName="C. C. Towns., Kew Bull." baseAuthorityName="Suess." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="fasciculata">
<emphasis id="C907D348F40DC20F8F32E9BBE4CA6F9E" italics="true">Achyranthes fasciculata</emphasis>
(Suess.) C. C. Towns., Kew Bull.
</taxonomicName>
34 (3): 424 (1980)
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9036EDAE0439F85BF2B4EB3DACF8C74" type="reference_group">
<paragraph id="7B9B9755C7908D54973C8AADEDA16EFA">
<treatmentCitationGroup id="5AC951CC792EC5D39EF596EA1FA007E7">
<treatmentCitation id="1C0CA806F32AA6D6B60ADDAE30501B21">
<treatmentCitation id="D4919D695ADA4969433A26CDE53AFAD3" author="Suess." journal="Kew Bull." page="477" score="22.239024375395413" volume="4" year="1949">
<taxonomicName id="86C869DB8D664E53F4525315D3368E25" ID-CoL="4CBRF" authority="Suess., Kew Bull." authorityName="Suess., Kew Bull." class="Magnoliopsida" family="Amaranthaceae" genus="Pandiaka" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="fasciculata">
<emphasis id="865E6C546A9C6207163A3B0CE1AAC8A2" italics="true">Pandiaka fasciculata</emphasis>
Suess., Kew Bull.
</taxonomicName>
4: 477 (1949)
</treatmentCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHMCK" type="Holotype">
<paragraph id="16DA0D422B0ADAC8BD736C5CCE482E2C">
<heading id="671309EB8BEDF65B3BA1FF76CABE4943" reason="title">
<typeStatus id="39BFDA19148E702A1F2FA77FA2AC7E35" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="48D78E844CDF9A3615254932362742F6">
<materialsCitation id="8AE96AD5579BE431178DBD0EC805BA47" collectingDate="1946-02-06" collectionCode="P, J, BM" country="Tanzania" county="Mbulu distr." elevation="2438" location="Nangwa" municipality="Mt. Hanang" specimenCode="K 000243725" specimenCount="1" stateProvince="Manyara" typeStatus="holotype">
Tanganyika Territory [
<collectingCountry id="750408FB822221CF7B6FFC2B5F602B4C" name="Tanzania">Tanzania</collectingCountry>
,
<collectingRegion id="66CAE81DDDB5A797DB29FCB806A5EC90" country="Tanzania" name="Manyara">Manyara Region</collectingRegion>
],
<collectingCounty id="D35816E5A288D205E7BA59D1EF3E5E87">Mbulu distr.</collectingCounty>
, SE slopes of
<collectingMunicipality id="8B8EBE718E3E161346054FADB37DEC7E">Mt. Hanang</collectingMunicipality>
,
<location id="778AE676CE157AF8FB66F70C8434CC75" LSID="urn:lsid:plazi:treatment:8B77D03B00BB56F5B085321BAF8D7103:778AE676CE157AF8FB66F70C8434CC75" country="Tanzania" county="Mbulu distr." municipality="Mt. Hanang" name="Nangwa" stateProvince="Manyara">Nangwa</location>
,
<quantity id="D3B45D4AFF058D6D909BDC9C9B841722" metricMagnitude="3" metricUnit="m" metricValue="2.4384" unit="ft" value="8000.0">
<elevation id="E8FD5C3DE0DD9DA112752818B30D4C36" metricMagnitude="3" metricUnit="m" metricValue="2.4384" unit="ft" value="8000.0">8000 ft</elevation>
</quantity>
,
<date id="3A2599EF55B634860D489B0B29F6BEE0" value="1946-02-06">
<collectingDate id="5F7FF4383DDB972035E89E28B622D2D1" value="1946-02-06">6 February 1946</collectingDate>
</date>
,
<emphasis id="30B32B1EA4FD94EA56CAE986ADB0D55C" italics="true">
<collectionCode id="BBFCD3D0655A375968C9D1E153C37AB9" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P</collectionCode>
.
<collectionCode id="2304BD7EB337B73F4FE5E92CF59048AC" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" type="Herbarium">J</collectionCode>
. Greenway 7620
</emphasis>
(
<specimenCode id="EDE78E49D12C48EE1720E523F6A3AB2F" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K 000243725</specimenCode>
!,
<typeStatus id="51418A3AD76659E3B7A57C11D74428B4" type="isotype">isotype</typeStatus>
<collectionCode id="14533610847CB95CD9E4946257D3ECAD" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
!).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPMCK" type="habitat">
<paragraph id="883E70B8EEFD6C81A5D14F91157557D4">
<heading id="AC8324FD7F4CAEF6092696535F93E668" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="4DA0BCAA0E44D312D3CC3AC0889EF0B7">
Upland grassland and mist forests at elevatios
<quantity id="66E73815D2B4A66AB72409D612236459" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.8" metricValueMin="1.8" unit="m" value="2300.0" valueMax="2800.0" valueMin="1800.0">18002800 m</quantity>
a. s. l.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUMCK" type="distribution">
<paragraph id="3E56E1A498F5D264E193D70FA900C8DC">
<heading id="96B0AE57C71651EB8513DDA3DA3B7811" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="7125FF6DF065E52EBF7BFD034C31FAD1">
(Fig.
<figureCitation id="5DCF992C76A1529FCFEED5DF70057206" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution map of Achyranthes fasciculata." figureDoi="10.3897/phytokeys.250.136139.figure19" httpUri="https://binary.pensoft.net/fig/1203863">19</figureCitation>
).
<emphasis id="8A6103641A3EDC3C69BB7E8B31325F4A" bold="true">
<collectingCountry id="8D7F606D1AE48337390E79D624744799" name="Tanzania">Tanzania</collectingCountry>
</emphasis>
(selected): Ngorongoro crater,
<quantity id="DB8A9241026CE1D6DF4C579D4DDABEAC" metricMagnitude="3" metricUnit="m" metricValue="2.4384" unit="ft" value="8000.0">8000 ft</quantity>
,
<date id="E0EE4FC20C72198F9BB9A775C59026F9" value="1932-09-12">12 Sep 1932</date>
,
<emphasis id="AB5D9ECE6BC9CFD251EEEA3ADB49F6AA" italics="true">
<collectionCode id="630CCB80545A37AF90299F0F3DBFF633" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" type="Herbarium">B</collectionCode>
. D. Burtt 4308
</emphasis>
(
<collectionCode id="4F7BFE6913B1303CE11F6D624F2A91B4" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Ngorongoro rest camp,
<quantity id="B536C856A7BE50F4F88E4D4FB5A000F7" metricMagnitude="3" metricUnit="m" metricValue="1.8288" unit="ft" value="6000.0">6000 ft</quantity>
,
<date id="8E8C61C3FFD65ECF5FF8E2947E6667F4" value="1941-04-03">3 Apr 1941</date>
,
<emphasis id="56FCC97F705430E87EDCE91020C1305D" italics="true">
<collectionCode id="846FD2CE27DCA3338DBD8B4F7E4CFE8B" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P</collectionCode>
.
<collectionCode id="07A2FCF8208C01FA338FD312FE21AE50" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectionCode id="12E7FC859F406A63B53151AE48C48D93" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" type="Herbarium">O</collectionCode>
. Balley 2232
</emphasis>
(
<collectionCode id="C185E6F55F33238A1EB0AF973041C0F4" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Ngorongoro crater,
<quantity id="F0148485BEDD8663E3DE6EA44409A12C" metricMagnitude="3" metricUnit="m" metricValue="2.2555199999999997" unit="ft" value="7400.0">7400 ft</quantity>
,
<date id="89EBFB99042FFE7C56160D8A1B8A56E9" value="1966-07-13">13 Jul 1966</date>
,
<emphasis id="ECDA302C3C72975906D0526BFB015738" italics="true">
<collectionCode id="C1B803B02B39A7153A84A302505DB1C0" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P</collectionCode>
.
<collectionCode id="418377103A316EFB8A8035A8D13F3807" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" type="Herbarium">J</collectionCode>
. Greenway &amp;
<collectionCode id="F8E25BEC1B3B02359DECF12CFA8ED0D7" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
. Kanuri 1256662
</emphasis>
(
<collectionCode id="1F07BB273C5E0D8542C207B00CAC4134" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
0000013708887,
<collectionCode id="8C69D0B0E1A0E507E64EE620AFE87826" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Empakaai crater,
<quantity id="F703D45F7285243371C108FA44FB5B0B" metricMagnitude="3" metricUnit="m" metricValue="2.15" unit="m" value="2150.0">2150 m</quantity>
a. s. l.,
<date id="EB6898A950488C91627EF1200805DA94" value="1972-08-10">10 Aug 1972</date>
,
<emphasis id="91238602A3C5194ED58D7607EBC5D6A0" italics="true">
<collectionCode id="F90FA85DB896D1B7A3D5CB53390A889F" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
.
<collectionCode id="144EFB1FA8845F842AB6C0530E7D28C3" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" type="Herbarium">W</collectionCode>
. Frame 27
</emphasis>
(
<collectionCode id="E4D40C88F1E996456B98A6366380EA3B" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
0000013708870,
<collectionCode id="7C6817D410E47B1125F1E71FB870DEB3" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Masai distr., Ngorongoro crater,
<quantity id="4ADBE85B9132045F746760053DF98050" metricMagnitude="3" metricUnit="m" metricValue="2.35" unit="m" value="2350.0">2350 m</quantity>
a. s. l.,
<date id="C70CFFF6C2E9A27793920E2E43DD49C4" value="1977-09-15">15 Sep 1977</date>
,
<emphasis id="034BF3C6DF629AEA542CCC6A975E0AD2" italics="true">
<collectionCode id="8843D564A0F894B5ACD7335BDB9A1453" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" type="Herbarium">J</collectionCode>
. Raynal 19052
</emphasis>
(
<collectionCode id="A18F5E48279D42FE4510B1BD170DC5D2" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
,
<collectionCode id="2620C6B90EAB1F9CB9934F398B50E48D" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
01404450);
<collectingRegion id="4E1F7888641E1DDECC45FACC36AD846F" country="Tanzania" name="Arusha">Arusha Region</collectingRegion>
, Monduli distr., Ketumbeine Forest Reserve,
<quantity id="03CA61D553C93F5FAF7DAAB0FA743675" metricMagnitude="3" metricUnit="m" metricValue="2.33" unit="m" value="2330.0">2330 m</quantity>
a. s. l.,
<date id="A5B71A4EB7540F9D4CADBAEC41CF8D51" value="2000-04-02">2 Apr 2000</date>
,
<emphasis id="0B0DE1D68F7AB79E5E08DD492A05F43A" italics="true">
<collectionCode id="1679A60B61AED26F19F8B7E91C3E48F2" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectionCode id="2EAE5B402E07615D17144501064E6E0C" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
. Gereau &amp; al. 6428
</emphasis>
(
<collectionCode id="562AA7C33300AB75B9AD160EFE9B92FB" country="China" name="Chongqing Museum">CM</collectionCode>
,
<collectionCode id="AE3DD6660C97F4504549E5B8B0F7A15E" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" type="Herbarium">MO</collectionCode>
).
</paragraph>
<caption id="0E3D16BB00C784AEC00882018E300717" ID-DOI="10.3897/phytokeys.250.136139.figure19" ID-arpha="F52B75FC-8BE5-5EA0-A42A-D194B9A4A78A" httpUri="https://binary.pensoft.net/fig/1203863" startId="F19">
<paragraph id="59933571DF34156AF42935EA0ACF0BF1">
<label id="186B99D3290F9A1E0FB1E007E68D2A1F">Figure 19.</label>
</paragraph>
<paragraph id="4F0500A640D5E91E952D8B86B596558C">
Distribution map of
<taxonomicName id="CE50FE65F90132C4DB7272DD3E850D56" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="fasciculata">
<emphasis id="CC9C58B78A6DE5B17CFDBA7EEEE11DDB" italics="true">Achyranthes fasciculata</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EEOCK" type="distribution">
<paragraph id="0F1DA2013612C03902C526D2DBA20A30">
<heading id="B4B6DD66CC695609225BB6AA5DA4E768" reason="title">General distribution.</heading>
</paragraph>
<paragraph id="ED14E6C6C482EC342D1B0E718AAE9D6B">
Only known from
<collectingCountry id="659E523070C2E95713CEA1FF21E10EC0" name="Tanzania">Tanzania</collectingCountry>
. Records from
<collectingCountry id="6931DD5FB09CD90AADCE23D5C1F6CEDE" name="Kenya">Kenya</collectingCountry>
are also possible.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,437 @@
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<mods:title id="E1F8DB9D45345A1703A5C81F40E1BE71">Africa and Arabia encompass a much greater species diversity in the Achyranthes aspera aggregate (Amaranthaceae, achyranthoid clade): Evidence from morphological and chorological data</mods:title>
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<mods:affiliation id="F2242A6E3A5E3A44C1DCF898BAACAE9A">The Mauritius Herbarium, RE Vaughan Building, Agricultural Services, Ministry of Agro-Industry and Food Security, Réduit, 80835, Mauritius</mods:affiliation>
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<mods:date id="820C9705150A7465DB95FA90C4FA8A8B">2024</mods:date>
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<subSubSection id="4913E2B6892B18D34C68DD555E6E734B" type="nomenclature">
<paragraph id="3A74DE2A6FD5184E5EC4DE0A8D34DEBC">
<taxonomicName id="68153C49A0801D914A1A9FF5B11388C2" authority="Sukhor." authorityName="Sukhorukov &amp; Kushunina &amp; Nilova &amp; Baider &amp; Sennikov" authorityYear="2024" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="seychellensis" status="sp. nov.">
<emphasis id="E4E946BBBAAD5F99CE495836C0B7719F" italics="true">Achyranthes seychellensis</emphasis>
Sukhor.
</taxonomicName>
<taxonomicNameLabel id="14107B57AACC037A1236CB241CD27299" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3PBK" type="Holotype">
<paragraph id="25CCB6B6E9889C2DBE84AF9EB99516E7">
<heading id="8F4736D3E48848AC5A4B015A8CDB0F28" reason="title">
<typeStatus id="07DF6412861FE2E953C6DF5EB52E2F6A" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="099DBAF0BE95A8CF122156BC6C805F0E">
<materialsCitation id="B2384AD1CC843A1DB12A895E95533D6C" collectingDate="1984-08-18" collectionCode="G" country="Seychelles" location="Farquhar Atoll" specimenCode="MW 0586306, LE 00019370" specimenCount="1" typeStatus="holotype">
<collectingCountry id="4704C609837A3976D67015C747283E13" name="Seychelles">Seychelles</collectingCountry>
,
<location id="CB1C96CA7B287379A474EB362889741E" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:CB1C96CA7B287379A474EB362889741E" country="Seychelles" name="Farquhar Atoll">Farquhar Atoll</location>
, coconut plantation,
<date id="502EA263305D2F2293A197EA8A49753C" value="1984-08-18">
<collectingDate id="6A1E58D055B503CF15408692CE633FD6" value="1984-08-18">18 August 1984</collectingDate>
</date>
,
<emphasis id="5DA5E6C2A89C64976778FC9797DF9275" italics="true">
<collectionCode id="46D6F5A8A7FB2BFC20A2AFFF7DF583F0" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. Ogureeva 608
</emphasis>
(
<specimenCode id="8837A63B11625A6E8ADC1229FFD37858" collectionCode="MW" country="Netherlands" name="Museum Wasmann">MW 0586306</specimenCode>
!,
<typeStatus id="BC38DC80F44121A747AAD885CFDBC6DE" type="isotype">isotype</typeStatus>
<specimenCode id="ABAB45E2EE4529E4D3D416A58A476FD1" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia">LE 00019370</specimenCode>
!).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEQBK" type="description">
<paragraph id="69AA8F28C9C53A4EDB72F4C7C3349F93">
<heading id="45A6630D5C4DD1AFD54AF60EB1B931CB" reason="title">Description.</heading>
</paragraph>
<paragraph id="4D3C27BFB2626FDDBF47368E32430225">
(Fig.
<figureCitation id="AE98D3F036232AC704B239E1A56E5002" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Holotype of Achyranthes seychellensis Sukhor., sp. nov. (Seychelles, Farquhar Atoll, coconut plantation, 18 Aug 1984, G. Ogureeva 608, MW 0586306)." figureDoi="10.3897/phytokeys.250.136139.figure16" httpUri="https://binary.pensoft.net/fig/1203860">16</figureCitation>
). Annual herb up to
<quantity id="691D281E923A91D4B7DCB0F1AF6BFC2A" metricMagnitude="0" metricUnit="m" metricValue="1.5" unit="m" value="1.5">1.5 m</quantity>
tall; stem stout, basally ± roundish, four-angled in the inflorescence; leaf petioles
<quantity id="C0F128FD474C03BCF9110C67BB95E1C5" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" unit="mm" value="17.5" valueMax="25.0" valueMin="10.0">1025 mm</quantity>
long, blades 80120 ×
<quantity id="89EE217DDC4528817BFED13FCCADDE74" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="mm" value="45.0" valueMax="60.0" valueMin="30.0">3060 mm</quantity>
, cuneate, tip shortly acuminate, green above and pale green below, slightly pubescent on veins; inflorescence up to
<quantity id="094849CAF4B4B15D727CEA83D37F7FD2" metricMagnitude="-1" metricUnit="m" metricValue="2.5" unit="cm" value="25.0">25 cm</quantity>
long with hirsute axis; bract 3.54.0 mm long; bracteoles 3.24.0 mm long; perianth
<quantity id="4216898A3B3DE3854CE956917389294D" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.55.5 mm</quantity>
long, ± equal, deep rose or purple inside; stamens 5, anthers 0.751.0 mm long; pseudostaminodes 1.252.0 mm long, brownish or white when dry, fimbriate; style with stigma
<quantity id="7F283E16F6151C7104A6CFA9CFA146D6" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.22.7 mm</quantity>
long; fruit (without style) ± 2.0 mm long.
</paragraph>
<caption id="D9EAC5C3ADD9CB23DD28AAB2A373CCDB" ID-DOI="10.3897/phytokeys.250.136139.figure16" ID-arpha="346ED6EA-EE5F-5B23-896C-A1DFCEFC177B" httpUri="https://binary.pensoft.net/fig/1203860" startId="F16">
<paragraph id="FE95B2F081221B02B0EACBD0D1C15CBA">
<label id="E5CAFAA9A977E0BBF0F23D22A21985F7">Figure 16.</label>
</paragraph>
<paragraph id="B5FD4EAE9B1BA8E3BFFE12F7366D71E1">
Holotype of
<taxonomicName id="125E0999A689E771677C46DB9A43E91F" authority="Sukhor." authorityName="Sukhorukov &amp; Kushunina &amp; Nilova &amp; Baider &amp; Sennikov" authorityYear="2024" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="seychellensis" status="sp. nov.">
<emphasis id="81F248477E6E18EC75C567AAE9848620" italics="true">Achyranthes seychellensis</emphasis>
Sukhor.
</taxonomicName>
,
<taxonomicNameLabel id="950AFEEED0BCFF8ACAEA5F3AD05CC91B" rank="species">sp. nov.</taxonomicNameLabel>
(Seychelles, Farquhar Atoll, coconut plantation, 18 Aug 1984,
<emphasis id="05F4FE55C3080F8D958E2BEFFF013B11" italics="true">G. Ogureeva 608</emphasis>
, MW 0586306).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EIRBK" type="habitat">
<paragraph id="168F9E889FA65C487EB4FB791A416786">
<heading id="FA1822655EB157A901506EEA36ABEDCD" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="F0B6BC85AE1AE1597754D6DB7E8A7BD7">
Calcareous and rocky areas at elevations of
<quantity id="86DE0EE66522F4B24D8B87AA06F57D64" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" unit="m" value="100.0" valueMax="200.0" valueMin="0.0">0200 m</quantity>
a. s. l. It can be a common weed in coconut plantations.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENRBK" type="Flowering at fruiting">
<paragraph id="F0BF92C7A87047481EC37B9BD4B92484">
<heading id="E69EC410E83E1C138008E72F4086DFC6" reason="title">Flowering at fruiting.</heading>
</paragraph>
<paragraph id="D49F8B6E4FB7FD3804FE5496DD3A14F5">Around the year.</paragraph>
</subSubSection>
<subSubSection id="SECID0ESRBK" type="Additional specimens seen">
<paragraph id="5F224384B3F5C2DCEA2C873A873F7353">
<heading id="3DF7EBC8BB2D603254A650AAFCC44C94" reason="title">Additional specimens seen.</heading>
</paragraph>
<paragraph id="1E5EB93B3E1FC8CCE842508FE9548E98">
<emphasis id="287B51931F9F5BD1B0A041F1F99FE53C" bold="true">
<collectingCountry id="CE35A23419E111818F57F73DD7CDAA7B" name="Seychelles">Seychelles</collectingCountry>
</emphasis>
: Sèche Island,
<date id="26FCF5F7774A73FF089657ACDE5E24C4" value="1938-01-23">23 Jan 1938</date>
,
<emphasis id="E3A8F8ABC3C64E8EE7E847E0DAC6D50A" italics="true">Ves 6149</emphasis>
(
<collectionCode id="79DE9FB3C0A6E4056ECF534D5F5CDDF5" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); St. Pierre Island,
<date id="D81F1C79AEE28543BF54EB69561161BC" value="1960-10">Oct 1960</date>
,
<emphasis id="0FA6F75460C4C41AD91577B019C51F6B" italics="true">
<collectionCode id="37E2BA190C9DC4CE3A962C013386E3BC" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
.
<collectionCode id="423155CA09013498C0CCB37730306A02" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" type="Herbarium">J</collectionCode>
. Pigeott s. n.
</emphasis>
(
<collectionCode id="474DD91B0AD753479A9FA178C559FA3B" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Aride Island,
<date id="7D55139286B607A37AA0AF8E47E77C97" value="1983-02">Feb 1983</date>
,
<emphasis id="421805658A37E7CFDB2B777FD5CC29F7" italics="true">
<collectionCode id="C45CAA98C31BDF0AB4DA02CBDA380F6E" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" type="Herbarium">F</collectionCode>
. Friedmann 4469
</emphasis>
(
<collectionCode id="67EA0A9F0C25E821AA34BBF1AAD59B87" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P</collectionCode>
00804851) &amp;
<date id="290FF7CC5FCB45E498913D8524B56966" value="1987-04">Apr 1987</date>
,
<emphasis id="4C8A71726A9699A8AE2246D2502FAB30" italics="true">
<collectionCode id="EA4F2C50A6284345FAC1A5EE851939F2" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" type="Herbarium">F</collectionCode>
. Friedmann 5455
</emphasis>
(
<collectionCode id="7F6F5C7EFDB2EB8045BDE9042E2F471A" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" type="Herbarium">P</collectionCode>
00804847); Bird Island,
<date id="C3C8325810109959F5D47E8423137005" value="1952-09-28">28 Sep 1952</date>
,
<emphasis id="E516CD4633B0A7B16FCCAC8124FBECD9" italics="true">
<collectionCode id="E2C9E397D0304B2F7021DFE75221E4A5" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" type="Herbarium">E</collectionCode>
.
<collectionCode id="00ABC66D0D777418FA5CCFD371D625B5" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" type="Herbarium">S</collectionCode>
. Brown s. n
</emphasis>
. (
<collectionCode id="3CF306DA2293451715BA214109A83174" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
); Cousin Island,
<date id="A306E1BC97ED4A003DA1DEE04FFE03E6" value="1970-01-25">25 Jan 1970</date>
,
<emphasis id="6E5265681FF8061C22045C290338B00F" italics="true">
<collectionCode id="20CE1E4D33A932E9D3B096BCE28DC341" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" type="Herbarium">F</collectionCode>
.
<collectionCode id="BD9188B53226E49F5517177E16CDBC86" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Fosberg 52173
</emphasis>
(
<collectionCode id="0873A87AD4B5D0DE8A4D1D2BE4A65529" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Fouquet Island,
<date id="0AFA9C79B6202DCEF5393F99E9F1C526" value="1976-04-05">5 Apr 1976</date>
,
<emphasis id="995ABBE1E93A04986F7765B954B41967" italics="true">
D.
<collectionCode id="64D65CE984DD78DB8A9CDE622EEF6F4F" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Stoddart 7148
</emphasis>
(
<collectionCode id="2D01376CD597C3092DD9A61BA8D48ED0" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Farquhar Atoll (South),
<date id="2D96CA6BDFD388485E672F9326EFDE1E" value="1967-10-02">2 Oct 1967</date>
,
<emphasis id="598BFE9F0E0DFF06B8CF6219F2E92237" italics="true">Gwynne &amp; Wood 1189</emphasis>
(
<collectionCode id="CD07863A7B7E88497AF400153E1064E5" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Farquhar Atoll (North),
<date id="DDCA129869A458439C4C8A6D385628E4" value="1968-09-19">19 Sep 1968</date>
,
<emphasis id="50D61F8CE9AEF12AEFE4EBC7C6E70CE6" italics="true">
D.
<collectionCode id="B464E51DFDEE172166D83C257A82C1C2" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Stoddart 1376
</emphasis>
(
<collectionCode id="9A410399A6A9D69EA0B57B8532D0BF38" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Aride Island,
<date id="E0CD5E9F70B3D01B1610D8B364A4E6F0" value="1975-07">Jul 1975</date>
,
<emphasis id="69D40FF191B4CE217C75744178219E11" italics="true">Warman 23</emphasis>
(
<collectionCode id="ECF486189FCC6E4F18E33D87C96D7650" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Rémire [Eagle] Island,
<date id="66C99F18A25F950D19E05675BBEF3240" value="1976-10-25">25 Oct 1976</date>
,
<emphasis id="9007D1737BFD1B61922230C167E40CEB" italics="true">
<collectionCode id="3217EED90CCF2DA50C94C77F550B5B34" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" type="Herbarium">S</collectionCode>
.
<collectionCode id="06942F714A6660240D97F322A3DFFC34" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
. Robertson 2319
</emphasis>
(
<collectionCode id="F069D5A2A52D8700D90694D119908B32" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Bird Island,
<date id="3A1C635C7B19716093E9ACA69CE0D4D2" value="1976-03">Mar 1976</date>
,
<emphasis id="61D787F10DB9C18E68D408301CA56F91" italics="true">
D.
<collectionCode id="B1BFD6E312D796ADEF135EAC0E446EA8" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Stoddart 7079
</emphasis>
(
<collectionCode id="4CAB3FB4CEB4AA88D89EBC22F6C141BB" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Denis Island,
<date id="9CD87486904059DCBB12F5097739545A" value="1977-12-10">10 Dec 1977</date>
,
<emphasis id="961818278C9B36A0DFF52E83E8856595" italics="true">
D.
<collectionCode id="3021D5D216844B8C620293D1F4FC5157" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Stoddart 8086
</emphasis>
(
<collectionCode id="6581D37D29D2C2B2C672945ED1B5EDDA" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Platte Island,
<date id="ECE74F6A2588114A691F5F0F84CB111D" value="1980-02-26">26 Feb 1980</date>
,
<emphasis id="CEE76B43C8E35DC5D807999AA8E93137" italics="true">
<collectionCode id="39ED05FF0631F19DDF85197621214133" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" type="Herbarium">S</collectionCode>
.
<collectionCode id="87D2A869C70020BF10769FCE4B138E32" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
. Robertson 3021
</emphasis>
(
<collectionCode id="D18143D135AE62D2ABA18B90F140A169" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Poivre Island,
<date id="C16E7424CA1D9A3D0C2C4FF356448C08" value="1984-08-06">6 Aug 1984</date>
,
<emphasis id="E08CCA84490CF327CCEE84707720E5E9" italics="true">
<collectionCode id="BFC174D4C6C80EAA835123212D4490C1" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" type="Herbarium">L</collectionCode>
. Averyanov &amp; al. s. n
</emphasis>
. (
<collectionCode id="CC1A3BE298250AF456B04794D6236D4F" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
,
<collectionCode id="7FB4490264006532635189F8E12DC509" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15585" lsid="urn:lsid:biocol.org:col:15585" name="Main Botanical Garden of the Russian Academy of Sciences" type="Herbarium">MHA</collectionCode>
); Farquhar Atoll,
<date id="23F11D58C0CF1895A3226055D5804BBD" value="1984-08-16">16 Aug 1984</date>
,
<emphasis id="C0315158D1C0AF6C798B3DC23E33B968" italics="true">
<collectionCode id="DA7DEE166CBB98D938997C37EB2C8617" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" type="Herbarium">L</collectionCode>
. Averyanov &amp; al. 580
</emphasis>
(
<collectionCode id="56F9F594D35C1D484D5B386C52450C03" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
).
</paragraph>
<paragraph id="A8F965BBF7D745185F835584BA5286B7">
<emphasis id="9B48A90C34646A77335BFCFC6EBBA5F9" bold="true">
<collectingCountry id="7B02A71DBAB3304F2AD2115BBEEE08E0" name="Mauritius">Mauritius</collectingCountry>
</emphasis>
: Chagos Archipelago, Diego Garcia Atoll, East Island,
<date id="669005FA70BFA44E55077CB155F160F1" value="1967-07-26">26 Jul 1967</date>
,
<emphasis id="4F91350D81343539B68F2E1CDB4FF794" italics="true">
D.
<collectionCode id="208451675A22CA2F2BFF944F58ED5C4E" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Stoddart 875
</emphasis>
(
<collectionCode id="E05C634DF2A69D0C2C14D2A63E8C507E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
); Diego Garcia, near north coast,
<date id="D16C2A0845A5B4C5D11B164C83F74541" value="1971-03-30">30 Mar 1971</date>
,
<emphasis id="665956768160769D06F9FEDFDA693100" italics="true">
<collectionCode id="E7C3D453E9C2955023C6262732E66D38" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" type="Herbarium">A</collectionCode>
.
<collectionCode id="7E97F876B2A40387B83D8B615F606D4A" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Hutson 15
</emphasis>
(
<collectionCode id="135F036EC1A4DBEFD7DBE5B094D9A214" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
); Sea Cow Island,
<date id="ECA317503F0B0D4B9D6860AA4138C187" value="1975-02-21">21 Feb 1975</date>
,
<emphasis id="9DB7917D1718440FAA1EA6B27B235D44" italics="true">
<collectionCode id="430D01917441A290D18B04B7FC401B95" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
.
<collectionCode id="ED6ED83669566D9CA5F40C2E2F132881" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" type="Herbarium">J</collectionCode>
. D. Hirons 50
</emphasis>
(
<collectionCode id="D710A91FECFE519190F12EE6FFA87817" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYTBK" type="Relationship">
<paragraph id="9B9D0FB3E350984C1756EE17F50D57B5">
<heading id="7AFE676D11EC7B622A5EA3F2422DA258" reason="title">Relationship.</heading>
</paragraph>
<paragraph id="A460EF5AFBDE6D2B018483384724920C">
<bibRefCitation id="D562B5A20832826B192CF4D6FC1EF058" author="Friedmann F" firstAuthor="Friedmann" journalOrPublisher="Orstom, Paris" refId="B39" refString="Friedmann F (1994) Flore de Seychelles. Dicotylédones. Orstom, Paris, 663 pp." title="Flore de Seychelles. Dicotylédones." year="1994">Friedmann (1994)</bibRefCitation>
reported three varieties of
<taxonomicName id="ED30B0EE91823DD14498F380786703F3" baseAuthorityName="Ghazanfar" baseAuthorityYear="1992" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="0D8E41F7AF3053822CD6DD617E94F8E9" italics="true">A. aspera</emphasis>
</taxonomicName>
:
<emphasis id="BDB8DCFC83F1B2A090E1E5F1B208D164" italics="true">
var.
<taxonomicName id="DF1A97B4663075CFCD214F997EE9FEF8" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="velutina">
aspera,
<emphasis id="686961EE422DE759583A0DF1EDDAA7FA" italics="true">var. velutina</emphasis>
</taxonomicName>
</emphasis>
, and
<taxonomicName id="0F12BEAAF74D5E635141BBC25381AB2D" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="fruticosa">
<emphasis id="BD9774F45F8878FC018B7B1CF44F1E02" italics="true">var. fruticosa</emphasis>
</taxonomicName>
from the
<collectingCountry id="3C5B136748A9C94DAD60DD97753DD8A2" name="Seychelles">Seychelles</collectingCountry>
. The specimens seen were identified as
<taxonomicName id="97212000222A8EE6AFF5AD9C9F49BAD7" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="aspera">
<emphasis id="CABAF1A262E08263275B65BF6C76EC00" italics="true">var. aspera</emphasis>
</taxonomicName>
or
<taxonomicName id="29D67E98B1DFA05354B1E88B9DC91843" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="velutina">
<emphasis id="492563F63563B503AD38EBAF204DD594" italics="true">var. velutina</emphasis>
</taxonomicName>
. From
<taxonomicName id="E72BE7FAEFCF7C0DE326DC96F336B409" baseAuthorityName="Ghazanfar" baseAuthorityYear="1992" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="5551F6940ECA04A021C0457DCE3D56D1" italics="true">A. aspera</emphasis>
</taxonomicName>
, the new species differs by longer leaf blades (80120 ×
<quantity id="3777862ABD6606978D1B29402A9F8FF9" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="mm" value="45.0" valueMax="60.0" valueMin="30.0">3060 mm</quantity>
vs. 15110 ×
<quantity id="1661D946024A86DC3505FDA7763431A1" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" unit="mm" value="50.0" valueMax="80.0" valueMin="20.0">2080 mm</quantity>
), and longer anthers (0.751.0 mm vs.
<quantity id="734F7DA3514CDD7BB113F377658A8199" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.40.6 mm</quantity>
) and styles (
<quantity id="CC780B98ACF4D45D8EAB6C9E171EC546" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.22.7 mm</quantity>
vs. 1.0
<quantity id="06550D7FEA3B6485F8DE22B4BB525269" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
). The true
<taxonomicName id="E6B4C72EE0C61B10141127E960745181" authority="Hook. &amp; Arn." authorityName="Hook. &amp; Arn." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="velutina">
<emphasis id="FD5B273737557B3C3310A2E267F55AE4" italics="true">A. velutina</emphasis>
Hook. &amp; Arn.
</taxonomicName>
from the Pacific (
<collectingCountry id="B18EEE9703029CE6B00934CE68E90AED" name="Samoa">Samoa</collectingCountry>
, Tahiti, Tuamotu etc.) is a hirsute shrub with longer (ca.
<quantity id="25E72F9A051DD5639E81CDE726E3ABA0" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
long) perianths, but some other characters, i. e. length of anthers and styles are similar. Several specimens from the Aldabra Islands identified as
<taxonomicName id="70D16B229BB2805705EADF2360FEE0D9" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="fruticosa">
<emphasis id="78C07917C575B8F496E37B6916CD62F9" italics="true">var. fruticosa</emphasis>
</taxonomicName>
represent plant fragments with a woody base and need further evalution.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4XBK" type="distribution">
<paragraph id="6FDCA95CD19B108D0AFFC19D8ED281AE">
<heading id="DDD55EE73ECE391FD48189DD0275708F" reason="title">General distribution.</heading>
</paragraph>
<paragraph id="8F0E06AC05864D250A501234B05A11D0">
Endemic to the
<collectingCountry id="83B08BDFD4D5CE8DECC1448D01751D9A" name="Seychelles">Seychelles</collectingCountry>
and Chagos Archipelago.
</paragraph>
</subSubSection>
</treatment>
</document>