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data/06/98/1B/06981B0650102FF3C3351718F585F10B.xml
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<mods:namePart>Marshall, Stephen A.</mods:namePart>
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<paragraph pageId="21" pageNumber="50">
<taxonomicName authority="Paykull, 1790" authorityName="Paykull" authorityYear="1790" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus (Paykull, 1790)</taxonomicName>
<treatment id="06981B0650102FF3C3351718F585F10B" ID-GBIF-Taxon="159361056" LSID="urn:lsid:plazi:treatment:06981B0650102FF3C3351718F585F10B" httpUri="http://treatment.plazi.org/id/06981B0650102FF3C3351718F585F10B" lastPageNumber="50" pageId="21" pageNumber="50" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae">
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<taxonomicName id="9E95008E9CADA3A19C630547A20E8E6F" ID-CoL="4R2FJ" authority="Paykull, 1790" authorityName="Paykull" authorityYear="1790" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus (Paykull, 1790)</taxonomicName>
</paragraph>
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<subSubSection pageId="21" pageNumber="50" type="materials">
<paragraph pageId="21" pageNumber="50">Materials.</paragraph>
<paragraph pageId="21" pageNumber="50">
<subSubSection id="C838D9D9A5DF876D7A7B7308BD9EF7E5" pageId="21" pageNumber="50" type="materials">
<paragraph id="5AA2615B233BA9799E7DA0A516C28160" pageId="21" pageNumber="50">Materials.</paragraph>
<paragraph id="D55C8F0C136CE97696BE960EF7ADD46C" pageId="21" pageNumber="50">
CANADA: ON: Wellington Co., Guelph, University Arboretum, rotting
<taxonomicName class="Agaricomycetes" family="Polyporaceae" genus="Polyporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Polyporus squamosus" order="Polyporales" pageId="21" pageNumber="50" phylum="Basidiomycota" rank="species" species="squamosus">Polyporus squamosus</taxonomicName>
<taxonomicName id="9CD1EB75F9DE8FF62C2702A8B780CB0F" class="Agaricomycetes" family="Polyporaceae" genus="Polyporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Polyporus squamosus" order="Polyporales" pageId="21" pageNumber="50" phylum="Basidiomycota" rank="species" species="squamosus">Polyporus squamosus</taxonomicName>
, 29-VI-2008, A. Brunke (2); Guelph, University campus nr. horse pen, grass sweep, 13-VII-2008, C. Ho (1).
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="50" type="diagnosis">
<paragraph pageId="21" pageNumber="50">Diagnosis.</paragraph>
<paragraph pageId="21" pageNumber="50">
<subSubSection id="AD451C13A4781FA887FA9DA8BEAD4064" pageId="21" pageNumber="50" type="diagnosis">
<paragraph id="A3A884AAB1F6A4376AF04E4787432B02" pageId="21" pageNumber="50">Diagnosis.</paragraph>
<paragraph id="D45429AD9168BF85BCDC23A088F16C42" pageId="21" pageNumber="50">
This species may be separated from other northeastern
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
<taxonomicName id="D4C340C2AFF2B23179B7291265B73152" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
by the combination of: elytra with three rows of coarse punctures on the disc; head without a pair of punctures between the ocular punctures; pronotum with three punctures in each dorsal row (Fig. 12).
</paragraph>
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<paragraph id="1D3A1C4EC9B8DC7E4124643FEC50031B" pageId="21" pageNumber="50">
This Palaearctic species was first detected in North America by
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana (1971)</bibRefCitation>
<bibRefCitation id="A7635F1E92FF747B0D678998C54DE205" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana (1971)</bibRefCitation>
based on specimens collected in Massachusetts. It was subsequently recorded from New Jersey, New York and Washington by
<bibRefCitation author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana (1990)</bibRefCitation>
<bibRefCitation id="AEE2A51F2E866E92D6CE3486DFE90A80" author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana (1990)</bibRefCitation>
, and New Hampshire by
<bibRefCitation author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler (2001)</bibRefCitation>
<bibRefCitation id="F8D70E2C260ED5622897B01DF3883F6C" author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler (2001)</bibRefCitation>
. Ithas been present in North America since at least 1942 based on a collection in New Jersey and was introduced to the west coast (Washington) as early as 1979 (
<bibRefCitation author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana 1990</bibRefCitation>
<bibRefCitation id="475A57F4DA1A0DE8EC3C9D6BAAF4B754" author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana 1990</bibRefCitation>
).
<bibRefCitation author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="341 - 345" title="Adventive species of Quedius (Coleoptera, Staphylinidae) in North America: a survey and new Canadian record." volume="22" year="2009">Majka et al. (2009)</bibRefCitation>
<bibRefCitation id="DDE773E79DE0C508F03E04564E460976" author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="341 - 345" title="Adventive species of Quedius (Coleoptera, Staphylinidae) in North America: a survey and new Canadian record." volume="22" year="2009">Majka et al. (2009)</bibRefCitation>
newly recorded it from Canada (New Brunswick) based on specimens collected on carrion. We here newly report
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus</taxonomicName>
<taxonomicName id="3BBEC648F4708362EF609C110087B7AA" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus</taxonomicName>
from Ontario (Map 35). In its native range,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus</taxonomicName>
<taxonomicName id="713D7B49A410247C1D555821E6063210" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus</taxonomicName>
is widespread in the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="B5239F3994C6AE417DE68700C860719E" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
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and Smetana 2004
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). This species frequents disturbed habitats throughout its range and is typically attracted to decaying organic matter; the Ontario specimens were found in a
<taxonomicName genus="Polyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyporus" pageId="21" pageNumber="50" phylum="Platyhelminthes" rank="genus">Polyporus</taxonomicName>
<taxonomicName id="1ED2E6CFAF1D45D48B860F420D3A867B" genus="Polyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyporus" pageId="21" pageNumber="50" phylum="Platyhelminthes" rank="genus">Polyporus</taxonomicName>
fungus in an almost liquid state of decay.
</paragraph>
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<treatment ID-GBIF-Taxon="159361051" LSID="urn:lsid:plazi:treatment:0CA2B084C9044BA620449E75BF6207AD" httpUri="http://treatment.plazi.org/id/0CA2B084C9044BA620449E75BF6207AD" lastPageNumber="44" pageId="15" pageNumber="44">
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<paragraph pageId="15" pageNumber="44">
<taxonomicName authority="Horn, 1884" authorityName="Horn" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus (Horn, 1884)</taxonomicName>
<treatment id="0CA2B084C9044BA620449E75BF6207AD" ID-GBIF-Taxon="159361051" LSID="urn:lsid:plazi:treatment:0CA2B084C9044BA620449E75BF6207AD" httpUri="http://treatment.plazi.org/id/0CA2B084C9044BA620449E75BF6207AD" lastPageNumber="44" pageId="15" pageNumber="44" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae">
<subSubSection id="F443148F04711C63AE2D0AE2C4EE29C2" pageId="15" pageNumber="44" type="nomenclature">
<paragraph id="1B12EA28AC4D4B8FDA47B37A96329A1F" pageId="15" pageNumber="44">
<taxonomicName id="E8C13EC6ACE62128A7D7291514ECA1A0" ID-CoL="6GMQD" authority="Horn, 1884" authorityName="Horn" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus (Horn, 1884)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="44" type="materials">
<paragraph pageId="15" pageNumber="44">Materials.</paragraph>
<paragraph pageId="15" pageNumber="44">
<subSubSection id="5FC23678628A463CD4F6D6B5069B4CA0" pageId="15" pageNumber="44" type="materials">
<paragraph id="CCD9D465A65D009950BF67D5B1E32308" pageId="15" pageNumber="44">Materials.</paragraph>
<paragraph id="BECC909295552ADF198A6FE950CB3B3C" pageId="15" pageNumber="44">
CANADA: ON: Kent Co., Rondeau Prov. Pk., spicebush trail, Carolinian forest, yellow pans, 25-V-2003, M. Buck and S. Paiero (1); Rondeau Prov. Pk., spicebush trail, Carolinian forest, yellow pans, 16 to 17-VI-2003, Buck and Carscadden (1); Wainfleet bog, 8km S of Welland, DIE pt. 1 -north ditch (control), 10 to 24-V-1988, A. Stirling (1); 7 to 13-VI-1988, all collected by A. Stirling: D2H pt. 2 - '1962
<normalizedToken originalValue="zone’">zone'</normalizedToken>
<normalizedToken id="24011B68B021CB01DAC5CFF625922A45" originalValue="zone’">zone'</normalizedToken>
(1), D4H pt. 4 - '1980
<normalizedToken originalValue="zone’">zone'</normalizedToken>
<normalizedToken id="29704EF0909C58E0C8084FA8451FD5B2" originalValue="zone’">zone'</normalizedToken>
(1), D5H pt. 5 - '1985
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<normalizedToken id="A4B0EF71DF7B8C5906E73040C7932509" originalValue="zone’">zone'</normalizedToken>
(1), D5E pt. 5 - '1985
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<normalizedToken id="A3F93B7A74B273657F994B1F945BD5DD" originalValue="zone’">zone'</normalizedToken>
(1).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="44" type="diagnosis">
<paragraph pageId="15" pageNumber="44">Diagnosis.</paragraph>
<paragraph pageId="15" pageNumber="44">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<subSubSection id="6026D33367AB9D5D8CC1A255F17A008A" pageId="15" pageNumber="44" type="diagnosis">
<paragraph id="597D3C470D7F2B4D9839888C19BC1A93" pageId="15" pageNumber="44">Diagnosis.</paragraph>
<paragraph id="BC17A9CED501C4962F42CB35032DD699" pageId="15" pageNumber="44">
<taxonomicName id="3296770F41DECDE6501D42DD7B1EBD9A" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
can be easily distinguished from other northeastern species of the genus with a sparsely punctate forebody, with the exception of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
<taxonomicName id="DC8361F7CAD8EC199EEEDD3CBB21344A" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
, by its small size (&lt;3.6mm from clypeus to abdominal apex).
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank (1975)</bibRefCitation>
<bibRefCitation id="79E0C38E7295BFF7DB1CA414CF5B41DB" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank (1975)</bibRefCitation>
stated that
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="CB9F0879709F4F4EB42E72A25F79DA98" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
could be separated from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
<taxonomicName id="4E1796975F9F184FC311269CEF634B55" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
by the paler coloration, the smaller eyes and the head broader behind the eyes. Head shape was found to be highly variable in both males and females of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="B2678CAAA5C30C51ACD759CFECA0A2BA" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
and this character should not be used to identify this species. However, the length of the pronotum appears to be a reliable character and is subequal to that of the elytra in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="BBD009200C5DEEDAB3F2F1481D703553" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
(Fig. 9) and clearly shorter in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
<taxonomicName id="6605E4E0BBB7D6073CD00F1346949984" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
. It should also be noted that the parameres of the aedeagus in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="79680F3E1019C93E0AFF33921CD4B846" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
possess characteristic short, stout setae (Fig. 19) which were not illustrated by
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank (1975)</bibRefCitation>
<bibRefCitation id="F242F7EEFD02BB3D3BBB9B61D730CFBC" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank (1975)</bibRefCitation>
but mentioned later in
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">Frank (1981a)</bibRefCitation>
<bibRefCitation id="DFEAB41FF3985C7342EC26D4F6C6849F" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">Frank (1981a)</bibRefCitation>
.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
<taxonomicName id="A0E7CB9B814782BB6FA4580719735F77" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
lacks these setae on the parameres.
</paragraph>
<paragraph pageId="15" pageNumber="44">
<paragraph id="A0FCDB041B9C235CBD073480EC2352AD" pageId="15" pageNumber="44">
This species was previously known from Florida (Frank 1981), District of Columbia, Massachusetts, Louisiana, Virginia (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
<bibRefCitation id="FE31781FCE4C1C66F4A9176A91D02670" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
), and New Hampshire (
<bibRefCitation author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
<bibRefCitation id="B9C8053AC0754A626764C15D1441E76F" author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
). Herein we record it as new from Canada (Ontario) based on eight specimens, all collected in the Carolinian region of southern Ontario (Map 23).
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="5A30609DC60CA754C19F32BFB372B564" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
has previously been collected by
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, in
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(
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
<bibRefCitation id="22E49D1772877010C0081DBC28B68E76" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
) and at lights (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">Frank 1981a</bibRefCitation>
<bibRefCitation id="473C309A61B68CE8C9C4FCC73289F46C" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">Frank 1981a</bibRefCitation>
); all Canadian specimens were collected in wet habitats with abundant moss (bog, slough forest). In a survey of southern Ontario peatlands,
<bibRefCitation author="Blades, DCA" editor="Finnamore, AT" journalOrPublisher="Ottawa" pageId="25" pageNumber="54" title="Terrestrial arthropods of Canadian peatlands: Synopsis of pan trap collections at four southern Ontario peatlands" volumeTitle="Terrestrial arthropods of peatlands, with particular reference to Canada. Memoirs of the Entomological Society of Canada No. 169" year="1994">Blades and Marshall (1994)</bibRefCitation>
<bibRefCitation id="BD3E45EFABD4BAA8B5131ABAFE1E88BA" author="Blades, DCA" editor="Finnamore, AT" journalOrPublisher="Ottawa" pageId="25" pageNumber="54" title="Terrestrial arthropods of Canadian peatlands: Synopsis of pan trap collections at four southern Ontario peatlands" volumeTitle="Terrestrial arthropods of peatlands, with particular reference to Canada. Memoirs of the Entomological Society of Canada No. 169" year="1994">Blades and Marshall (1994)</bibRefCitation>
reported
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
<taxonomicName id="95561E79B9C330EC3B6519854394FC57" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
from Wainfleet bog based on a series of specimens found by the first author to be misidentified
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="FC08B112EE88C2C4E1E8C47229ECF075" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
. To our knowledge, no true
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
<taxonomicName id="4A97F2F6DAF5B6AD5B863AFB47AAA502" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius pusio" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="pusio">Erichsonius pusio</taxonomicName>
have been found in southern Ontario peatlands.
</paragraph>
</subSubSection>

136
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<mods:title id="68AEED947A3864BA61C300C5E6D0D5B1">Contributions to the faunistics and bionomics of Staphylinidae (Coleoptera) in northeastern North America: discoveries made through study of the University of Guelph Insect Collection, Ontario, Canada</mods:title>
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<treatment ID-GBIF-Taxon="159361047" LSID="urn:lsid:plazi:treatment:0FC9077C4C27197D87DD93B6E356A98C" httpUri="http://treatment.plazi.org/id/0FC9077C4C27197D87DD93B6E356A98C" lastPageNumber="38" pageId="9" pageNumber="38">
<subSubSection pageId="9" pageNumber="38" type="nomenclature">
<paragraph pageId="9" pageNumber="38">
<taxonomicName authority="Scopoli, 1763" authorityName="Scopoli" authorityYear="1763" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis (Scopoli, 1763)</taxonomicName>
<treatment id="0FC9077C4C27197D87DD93B6E356A98C" ID-GBIF-Taxon="159361047" LSID="urn:lsid:plazi:treatment:0FC9077C4C27197D87DD93B6E356A98C" httpUri="http://treatment.plazi.org/id/0FC9077C4C27197D87DD93B6E356A98C" lastPageNumber="38" pageId="9" pageNumber="38" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Steninae">
<subSubSection id="0F4B2DAF3BF56136EE9A2ED7DD85089E" pageId="9" pageNumber="38" type="nomenclature">
<paragraph id="407217F0754E2FE23A56D6A14E283503" pageId="9" pageNumber="38">
<taxonomicName id="BD41693E32F2902C252BFE4DC5A802E5" ID-CoL="6ZNP8" authority="Scopoli, 1763" authorityName="Scopoli" authorityYear="1763" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis (Scopoli, 1763)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="38" type="materials">
<paragraph pageId="9" pageNumber="38">Materials.</paragraph>
<paragraph pageId="9" pageNumber="38">UNITED STATES: MA:Middlesex Co., Groton, 30-IV-2010, T. Murray, (1).</paragraph>
<paragraph pageId="9" pageNumber="38">CANADA: ON:Halton Reg., Milton, Derry Rd. and 4th line, grass field, yellow pans, 23 to 24-VI-2001, S. Paiero, (1); Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.744, -81.507, soybean field, pitfall, 4-VIII-2010 (1), A. Brunke; Auburn, Limekiln Line, 43.736, -81.506, hedgerow, pitfall, 26-V-2010 (1), A. Brunke; Benmiller, Sharpes Creek Line, 43.691, -81.608, hedgerow nr. creek, canopy trap in buckthorn, 11-V-2009 (2), A. Brunke;Brucefield, London Rd. nr. Centennial Rd., 43.509, -81.516, hedgrerow nr. creek, pitfall, 11-V-2009 (1), A. Brunke; Ottawa, Carleton Place, 12-IX-1992, W. Bennett, (1); Peel Reg., Cooksville, pond margin, 30-V-1993, C. Krupke, (1); Waterloo Reg., Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, hedgerow, pitfall, 18-V-2010 (1), A. Brunke; Wellington Co., Guelph, 10-IX-1980, Y. Deedat; Guelph, 17-IX-1980, Y. Deedat; Guelph, University Arboretum, 1-X-2005, M. Bergeron, (1); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, canopy trap, 23-VI-2009 (1), A. Brunke; York Reg., Stouffville, V-1982, Brian Brown, (1);</paragraph>
<subSubSection id="DF35A7E711DA499AFE5FDB4960327C2E" pageId="9" pageNumber="38" type="materials">
<paragraph id="81DA876406EE6F76756E6CD6EE4BD7BA" pageId="9" pageNumber="38">Materials.</paragraph>
<paragraph id="0E54A5FA20A0276B9DEC4CAED839A64D" pageId="9" pageNumber="38">UNITED STATES: MA:Middlesex Co., Groton, 30-IV-2010, T. Murray, (1).</paragraph>
<paragraph id="CA2CD1CA66734119BEAA8C3AABA4BD48" pageId="9" pageNumber="38">CANADA: ON:Halton Reg., Milton, Derry Rd. and 4th line, grass field, yellow pans, 23 to 24-VI-2001, S. Paiero, (1); Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.744, -81.507, soybean field, pitfall, 4-VIII-2010 (1), A. Brunke; Auburn, Limekiln Line, 43.736, -81.506, hedgerow, pitfall, 26-V-2010 (1), A. Brunke; Benmiller, Sharpes Creek Line, 43.691, -81.608, hedgerow nr. creek, canopy trap in buckthorn, 11-V-2009 (2), A. Brunke;Brucefield, London Rd. nr. Centennial Rd., 43.509, -81.516, hedgrerow nr. creek, pitfall, 11-V-2009 (1), A. Brunke; Ottawa, Carleton Place, 12-IX-1992, W. Bennett, (1); Peel Reg., Cooksville, pond margin, 30-V-1993, C. Krupke, (1); Waterloo Reg., Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, hedgerow, pitfall, 18-V-2010 (1), A. Brunke; Wellington Co., Guelph, 10-IX-1980, Y. Deedat; Guelph, 17-IX-1980, Y. Deedat; Guelph, University Arboretum, 1-X-2005, M. Bergeron, (1); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, canopy trap, 23-VI-2009 (1), A. Brunke; York Reg., Stouffville, V-1982, Brian Brown, (1);</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="38" type="diagnosis">
<paragraph pageId="9" pageNumber="38">Diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="38">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis</taxonomicName>
<subSubSection id="BC770B117E9A15DA4A2522EAB1BBC72E" pageId="9" pageNumber="38" type="diagnosis">
<paragraph id="48B3846D5B12B94E5D3BE6E7A3CB4941" pageId="9" pageNumber="38">Diagnosis.</paragraph>
<paragraph id="7294772DAAF5AB419152484DF68780EF" pageId="9" pageNumber="38">
<taxonomicName id="EFC5F8BD84FF07DEE6F39C06D111B611" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis</taxonomicName>
is, at present, only reliably distinguished from congeners in North America by its characteristic aedeagus (Fig. 14).
</paragraph>
<caption pageId="9" pageNumber="38">
<paragraph pageId="9" pageNumber="38">
<caption id="6C9D3E27004C7340CDD9032B04A6DF04" pageId="9" pageNumber="38">
<paragraph id="A8C9FAA0B0433D478F773C029A0AB389" pageId="9" pageNumber="38">
Maps 13-16. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 13
<taxonomicName class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis</taxonomicName>
<taxonomicName id="50E9C63138BA7AD6A0EAE991C9A68B8D" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis</taxonomicName>
(Scopoli) (
<bibRefCitation author="Puthz, V" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="27" pageNumber="56" title="Stenus clavicornis (Scop.) neu fur Nordamerika" volume="71" year="1975">Puthz 1975</bibRefCitation>
<bibRefCitation id="BA20B9706F4A1150A1691EBE7354A6B1" author="Puthz, V" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="27" pageNumber="56" title="Stenus clavicornis (Scop.) neu fur Nordamerika" volume="71" year="1975">Puthz 1975</bibRefCitation>
,
<bibRefCitation author="Puthz, V" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="27" pageNumber="56" title="Stenus clavicornis (Scop.) nun auch in den U. S. A. (Staphyl.)." volume="90" year="1994">Puthz 1994</bibRefCitation>
<bibRefCitation id="E49CAB4CBEDDF17B0A223A076AE8B9B4" author="Puthz, V" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="27" pageNumber="56" title="Stenus clavicornis (Scop.) nun auch in den U. S. A. (Staphyl.)." volume="90" year="1994">Puthz 1994</bibRefCitation>
,
<bibRefCitation author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
<bibRefCitation id="6A0CBDF2C269FEA8D48F5ED12E63DEA7" author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
,
<bibRefCitation author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
<bibRefCitation id="3D1327C24BE45F3B9C3D1943DAE64501" author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
,
<bibRefCitation author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
<bibRefCitation id="5EA165554C6A337D0103F9BE754B3E78" author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
,
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="26" pageNumber="55" title="Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada" year="2010">Klimaszewski et al. 2010</bibRefCitation>
<bibRefCitation id="EE4F41A15297EFCD4EB934E4DB7F85BB" author="Klimaszewski, J" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="26" pageNumber="55" title="Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada" year="2010">Klimaszewski et al. 2010</bibRefCitation>
) 14
<taxonomicName class="Insecta" family="Staphylinidae" genus="Eustilicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustilicus tristis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="tristis">Eustilicus tristis</taxonomicName>
<taxonomicName id="3AA761259417D718AF559531F388DBF9" class="Insecta" family="Staphylinidae" genus="Eustilicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustilicus tristis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="tristis">Eustilicus tristis</taxonomicName>
(Melsheimer) (
<bibRefCitation author="Melsheimer, FE" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="27" pageNumber="56" pagination="26 - 43" title="Descriptions of new species of North American Coleoptera." volume="2" year="1844">Melsheimer 1884</bibRefCitation>
<bibRefCitation id="B65E3D35B76452B16C864B2FED405A3D" author="Melsheimer, FE" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="27" pageNumber="56" pagination="26 - 43" title="Descriptions of new species of North American Coleoptera." volume="2" year="1844">Melsheimer 1884</bibRefCitation>
,
<bibRefCitation author="Sanderson, MW" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="27 - 30" title="The North American species of Stilicolina Casey (Coleoptera, Staphylinidae)." volume="20" year="1947">Sanderson 1947</bibRefCitation>
<bibRefCitation id="F789BC4DD2C3247505BC6937ABCA13A1" author="Sanderson, MW" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="27 - 30" title="The North American species of Stilicolina Casey (Coleoptera, Staphylinidae)." volume="20" year="1947">Sanderson 1947</bibRefCitation>
,
<bibRefCitation pageId="9" pageNumber="38">Herman 1970</bibRefCitation>
<bibRefCitation id="25C2D4845566B0471FAC17C9ABB0E551" pageId="9" pageNumber="38">Herman 1970</bibRefCitation>
) 15
<taxonomicName class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus</taxonomicName>
<taxonomicName id="7C3FE50ACFD2D9DF923CCF78FAC09165" class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus</taxonomicName>
(Mannerheim) (
<bibRefCitation author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
<bibRefCitation id="C4E0B3C2D06E33EB19F5256B1696818C" author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
) 16
<taxonomicName class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus minutus" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="minutus">Scopaeus minutus</taxonomicName>
<taxonomicName id="F1343BE849497D8871D2F770C6DFCE8F" class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus minutus" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="minutus">Scopaeus minutus</taxonomicName>
Erichson (
<bibRefCitation author="Frisch, J" journalOrPublisher="Organisms Diversity and Evolution" pageId="26" pageNumber="55" pagination="27 - 53" title="Rove beetles of the subtribe Scopaeina Mulsant and Rey (Coleoptera: Staphylinidae) in the West Palaearctic: phylogeny, biogeography and species catalogue." volume="2" year="2002">Frisch et al. 2002</bibRefCitation>
<bibRefCitation id="3668BAD13412AB0DDA9A04A4484869A5" author="Frisch, J" journalOrPublisher="Organisms Diversity and Evolution" pageId="26" pageNumber="55" pagination="27 - 53" title="Rove beetles of the subtribe Scopaeina Mulsant and Rey (Coleoptera: Staphylinidae) in the West Palaearctic: phylogeny, biogeography and species catalogue." volume="2" year="2002">Frisch et al. 2002</bibRefCitation>
).
</paragraph>
</caption>
<paragraph pageId="9" pageNumber="38">
<paragraph id="5135A7BC694EF726549C3FAD5D2C849B" pageId="9" pageNumber="38">
This exotic, Palaearctic species was first recognized in North America by
<bibRefCitation author="Puthz, V" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="27" pageNumber="56" title="Stenus clavicornis (Scop.) neu fur Nordamerika" volume="71" year="1975">Puthz (1975)</bibRefCitation>
<bibRefCitation id="D6BC83B9DFB3B94C10EA678E18EF95D1" author="Puthz, V" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="27" pageNumber="56" title="Stenus clavicornis (Scop.) neu fur Nordamerika" volume="71" year="1975">Puthz (1975)</bibRefCitation>
based on specimens collected in
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="637CC993655BB12081DB45116F5F6716" originalValue="Québec">Quebec</normalizedToken>
as early as 1968 in Orsainville. Since then it has been detected in New Brunswick (
<bibRefCitation author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
<bibRefCitation id="B758AEA0C8BFFE89DF05B4057B0C950E" author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
), Prince Edward Island (
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="26" pageNumber="55" title="Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada" year="2010">Klimaszewski et al. 2010</bibRefCitation>
<bibRefCitation id="5860AE213F8621929BF7A2892ED94E1F" author="Klimaszewski, J" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="26" pageNumber="55" title="Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada" year="2010">Klimaszewski et al. 2010</bibRefCitation>
), New Hampshire, Pennsylvania (
<bibRefCitation author="Puthz, V" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="27" pageNumber="56" title="Stenus clavicornis (Scop.) nun auch in den U. S. A. (Staphyl.)." volume="90" year="1994">Puthz 1994</bibRefCitation>
<bibRefCitation id="E9E693E77EAEDF38E7EE30D10504E654" author="Puthz, V" journalOrPublisher="Entomologische Blatter fur Biologie und Systematik der Kafer" pageId="27" pageNumber="56" title="Stenus clavicornis (Scop.) nun auch in den U. S. A. (Staphyl.)." volume="90" year="1994">Puthz 1994</bibRefCitation>
), Maine (
<bibRefCitation author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
<bibRefCitation id="DD5F6FCBD2A2A429A73C11F6FBBAF1CC" author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
), and Nova Scotia (
<bibRefCitation author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
<bibRefCitation id="A247FB1EBCF7110A13D9AB26EA8CB595" author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
). Herein we report
<taxonomicName class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis</taxonomicName>
<taxonomicName id="0DB6EA0AFE08F1DC644BE4EF26FE73A3" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis</taxonomicName>
from Ontario and Massachusetts on the basis of specimens collected as early as 1980 and 2010, respectively (Map 13). Its native range is very broad and includes most of the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="11EA900179C325C04D101E15B6D5DD94" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
<normalizedToken id="67CE4B6E36E973D62B67E14044BB891B" originalValue="Löbl">Loebl</normalizedToken>
and Smetana 2004
</bibRefCitation>
). In North America, this species has been collected from a variety of habitats including open fields, agricultural land, woodlots, and the margins of ponds and salt marshes.

92
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<taxonomicName authority="Casey, 1915" authorityName="Casey" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus vulgatus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="vulgatus">Philonthus vulgatus Casey, 1915</taxonomicName>
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<paragraph pageId="21" pageNumber="50">Materials.</paragraph>
<paragraph pageId="21" pageNumber="50">
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<paragraph id="636EF2BF68885269B939A4FB40ED6567" pageId="21" pageNumber="50">Materials.</paragraph>
<paragraph id="A682E084046397744DEB77CC8B0EC85D" pageId="21" pageNumber="50">
CANADA: PEI: Brackley Beach, National Park, milieu
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(=marshy environment), 2-VIII-1979, R. Sexton (13).
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<paragraph pageId="21" pageNumber="50">Diagnosis.</paragraph>
<paragraph pageId="21" pageNumber="50">
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<paragraph id="CA04DB7431B33E22332FB7F507F6771B" pageId="21" pageNumber="50">Diagnosis.</paragraph>
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This species can be easily distinguished from other northeastern
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
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by the combination of: pronotum with punctures widely distributed and not in rows; legs and antennae completely dark; head with hind angles indistinct (Fig. 11).
</paragraph>
<paragraph pageId="21" pageNumber="50">
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<taxonomicName id="E1E49B753A4DA85A74E8918E0B01997E" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus vulgatus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="vulgatus">Philonthus vulgatus</taxonomicName>
was previously known from Alaska, Alberta, British Columbia, Idaho, Illinois, Indiana, Iowa, Kansas, Maine, Manitoba, Massachusetts, Michigan, Minnesota, Montana, Nebraska, New Hampshire, New York, North Dakota, Nova Scotia, Ontario,
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, Saskatchewan, South Dakota, Utah, Washington, and Wisconsin (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
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). Herein we newly report this species from Prince Edward Island (Map 34).
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is a hygrophilous species that has been collected in beaver lodges, at lights, and along water margins in debris or emergent vegetation (
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).
</paragraph>
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104
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<paragraph pageId="4" pageNumber="33">Materials.</paragraph>
<paragraph pageId="4" pageNumber="33">UNITED STATES: VA: Giles Co., Cascades Recreation Area, sifted from leaf litter in hardwood forest, 11 to 25-V-2008, A. Brunke (1).</paragraph>
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<paragraph id="285AD9B777BA006CAC21A9B8D92488EE" pageId="4" pageNumber="33">UNITED STATES: VA: Giles Co., Cascades Recreation Area, sifted from leaf litter in hardwood forest, 11 to 25-V-2008, A. Brunke (1).</paragraph>
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<paragraph pageId="4" pageNumber="33">Diagnosis.</paragraph>
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<paragraph id="4807DC919B8331C4005C49DB4AA6F92A" pageId="4" pageNumber="33">Diagnosis.</paragraph>
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<taxonomicName id="80F68E592CF9FE174DD5C83E11DB2F48" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
is distinguished from other species of the genus in northeastern North America by the combination of: pronotum and elytra without
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and without pale or reddish markings; small size (&lt;1.7mm from the clypeus to the elytral apex); middle-tibia with two apical spines; basal abdominal segments with long lateral bristles.
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When
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell (1976)</bibRefCitation>
<bibRefCitation id="4108E8F7847CCEAF5E1BCE02E1DB0FDB" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell (1976)</bibRefCitation>
described this species (under the homonymic name
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<taxonomicName id="D109658F57C912C18886AF6325E98D63" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus micans" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="micans">Sepedophilus micans</taxonomicName>
), eight specimens were known from scattered localities in Alabama, Maryland, District of Columbia, and North Carolina. Recently, five specimens of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
<taxonomicName id="973726E59B8F6B43D13537E23E08671F" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
were found in Cuivre River State Park, Missouri, at blacklight and under bark (
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
<bibRefCitation id="3956F8E3DBC4567B1C3354001EDEA5AE" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
).
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
<bibRefCitation id="CA0B2C7838DAFC411BF4DC79F09662BA" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
also noted that
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
<taxonomicName id="54069BE9B2BE8C2A735DFE094453172A" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
has been found in Ohio and Florida but without further details. We here report
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<taxonomicName id="D42DB9495DFDB2FFB14659C1C6501729" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
as new for Virginia, contributing to the faunistics of this poorly known species (Map 4).
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
<taxonomicName id="47B553F47824A48F6A6E2AF901BB58FF" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
was recommended for state listing as S3 rank in Missouri based on rarity there and elsewhere (
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
<bibRefCitation id="B32A0226905276CFF6DEFFCA0B43E3BA" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
). Although one specimen has been found on a dead chicken (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="47765BA738FECB5397C8D125275482B0" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
),
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
<taxonomicName id="09D57286F7F204170AD88779C10C9AD8" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
is probably a litter or subcortical species.
</paragraph>
</subSubSection>

114
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<paragraph pageId="13" pageNumber="42">Materials.</paragraph>
<paragraph pageId="13" pageNumber="42">UNITED STATES: VA: Giles Co., Mountain Lake, on dead fox squirrel, 11 to 25-V-2008, A. Brunke (3).</paragraph>
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<subSubSection id="A79E5CC09DAE9A430F062FA0ACB0F482" pageId="13" pageNumber="42" type="materials">
<paragraph id="A757FAB008BF6E2E1ADFA1BD9531E109" pageId="13" pageNumber="42">Materials.</paragraph>
<paragraph id="2260ECDBCA53F3C1713E0806EBD9B3E1" pageId="13" pageNumber="42">UNITED STATES: VA: Giles Co., Mountain Lake, on dead fox squirrel, 11 to 25-V-2008, A. Brunke (3).</paragraph>
<paragraph id="D5C5637F0932039728F1D7566FD12B54" pageId="13" pageNumber="42">
CANADA: PEI: Long Pond, National Park, milieu
<normalizedToken originalValue="marécageux">marecageux</normalizedToken>
<normalizedToken id="E168F24761BB918B04F253B8FB959F23" originalValue="marécageux">marecageux</normalizedToken>
(=marshy environment), 30-VII-1979, R. Sexton (2); West Covehead,
<normalizedToken originalValue="débris">debris</normalizedToken>
<normalizedToken id="D6A1F76A1110F8B8995A5DEBEE3FC481" originalValue="débris">debris</normalizedToken>
sur la plage (=beach debris), 25-VII-1979, R. Sexton, (3).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="42" type="diagnosis">
<paragraph pageId="13" pageNumber="42">Diagnosis.</paragraph>
<paragraph pageId="13" pageNumber="42">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius siegwaldii" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="siegwaldii">Bisnius siegwaldii</taxonomicName>
<subSubSection id="EC6601E06EAD08B2E07B561229876533" pageId="13" pageNumber="42" type="diagnosis">
<paragraph id="6F9033F4C1BFF8AF8706B2B849C47C93" pageId="13" pageNumber="42">Diagnosis.</paragraph>
<paragraph id="59937918CF63FE4891D21E0548835059" pageId="13" pageNumber="42">
<taxonomicName id="AF4210E0551F1335A311F5AA98744FA6" class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius siegwaldii" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="siegwaldii">Bisnius siegwaldii</taxonomicName>
is easily recognized among other species of the genus in the northeast by the combination of: body not bicolored; elytra dark; pronotum with at least five punctures in each dorsal row; head with punctures arranged to form a
<normalizedToken originalValue="‘V’">'V'</normalizedToken>
<normalizedToken id="0A941F70416F907AA97E9B5B9BB9D86A" originalValue="‘V’">'V'</normalizedToken>
(
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="456FB6F9E2C50658E73E4C98842F6909" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
) (Fig. 7).
</paragraph>
<caption pageId="13" pageNumber="42">
<paragraph pageId="13" pageNumber="42">
<caption id="175A5B1CE85EAB3DADE05DE10A27798F" pageId="13" pageNumber="42">
<paragraph id="530F021F65592FF702C9B72CCC48D8A5" pageId="13" pageNumber="42">
Figures 7-12. Dorsal habitus. 7
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius siegwaldii" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="siegwaldii">Bisnius siegwaldii</taxonomicName>
<taxonomicName id="80BE3D07EDDFD6464CC527BC6848DDCB" class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius siegwaldii" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="siegwaldii">Bisnius siegwaldii</taxonomicName>
(Mannerheim)8
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius brachycephalus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="brachycephalus">Erichsonius brachycephalus</taxonomicName>
<taxonomicName id="6A01E347FB658D1736BAC10CCDB67D1C" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius brachycephalus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="brachycephalus">Erichsonius brachycephalus</taxonomicName>
Frank9
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="A61DFCFCBD6C27576843760066B057AD" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
(Horn) 10
<taxonomicName class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis</taxonomicName>
<taxonomicName id="F0310A4CDF772D1AD2B925A62857DCBC" class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis</taxonomicName>
(LeConte)11
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus vulgatus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="vulgatus">Philonthus vulgatus</taxonomicName>
<taxonomicName id="AC5E14CFF939841797B51E90DFF058B6" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus vulgatus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="vulgatus">Philonthus vulgatus</taxonomicName>
Casey12
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus</taxonomicName>
<taxonomicName id="40DC38189DF06C73CDFF67BF51C20F10" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus</taxonomicName>
(Paykull).
</paragraph>
</caption>
<paragraph pageId="13" pageNumber="42">
<paragraph id="280A8E453FF68EE9A9FA5C26FE9CBBF2" pageId="13" pageNumber="42">
This species is transcontinental across northern North America with several collections made further south in both the east and west. It is currently known from Alaska, Alberta, British Columbia, California, Connecticut, Maine, Manitoba, Michigan, Montana, Newfoundland, New Brunswick, New Hampshire, New York, North Carolina, Nova Scotia, Northwest Territories, Ontario, Oregon,
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="F842F3870512E1777528AA6B624E7099" originalValue="Québec">Quebec</normalizedToken>
, Saskatchewan, Tennessee, Vermont, Washington, West Virginia, Wisconsin, and Yukon Territory (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="E071ADAA5D8D193DCE7C65B6A5191ABB" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
). Herein we newly record it from Prince Edward Island and Virginia (Map 20).
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius siegwaldii" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="siegwaldii">Bisnius siegwaldii</taxonomicName>
<taxonomicName id="1FB4248FABD32607CB1EE50FD3934E52" class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius siegwaldii" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="siegwaldii">Bisnius siegwaldii</taxonomicName>
is a common species found in carrion, dung, rotting fungi, decaying plant matter and wood, moss, and in vegetation near water (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="ED7FA00534F4ECD30D0192B61FCD4850" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
).
</paragraph>
</subSubSection>

94
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<subSubSection pageId="14" pageNumber="43" type="nomenclature">
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<taxonomicName authority="Horn, 1884" authorityName="Horn" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus (Horn, 1884)</taxonomicName>
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<subSubSection id="2D95B929D5DB4E6A6D9192B1729DC29B" pageId="14" pageNumber="43" type="nomenclature">
<paragraph id="D40EB90C02E5FA8507CD2AB70EACF9C0" pageId="14" pageNumber="43">
<taxonomicName id="51FA963AFCC33664E8BE5CC286BA90AC" ID-CoL="6GN2H" authority="Horn, 1884" authorityName="Horn" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus (Horn, 1884)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="43" type="materials">
<paragraph pageId="14" pageNumber="43">Materials.</paragraph>
<paragraph pageId="14" pageNumber="43">
<subSubSection id="01AFADE6153D7D9770A50BF9212BA442" pageId="14" pageNumber="43" type="materials">
<paragraph id="5252BE079DBEB7C5E57F38B3EAAB0E82" pageId="14" pageNumber="43">Materials.</paragraph>
<paragraph id="FFAB1A85C37284184B9A883586AF1282" pageId="14" pageNumber="43">
CANADA: PEI: Long Pond National Park, milieu
<normalizedToken originalValue="marécageux">marecageux</normalizedToken>
<normalizedToken id="8263B312B82F10C59C039F7020229C13" originalValue="marécageux">marecageux</normalizedToken>
(=marshy environment), 30-VII-1979, R. Sexton (2).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="44" type="diagnosis">
<paragraph pageId="15" pageNumber="44">
<pageBreakToken pageId="15" pageNumber="44" start="start">Diagnosis</pageBreakToken>
<subSubSection id="A105D6AE7C5CFC00B0CB0543630275E0" pageId="15" pageNumber="44" type="diagnosis">
<paragraph id="84509FCCC31BED8BB9F11DA36749126F" pageId="15" pageNumber="44">
<pageBreakToken id="FBBFB7CDDF0BC893E8E3FDA8211A08FD" pageId="15" pageNumber="44" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="15" pageNumber="44">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus</taxonomicName>
<paragraph id="8D5924FC399E7E458EE52F526FD15CDC" pageId="15" pageNumber="44">
<taxonomicName id="697E83CF75E38FFE70808A10050FD54C" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus</taxonomicName>
can be recognized among the other densely punctate species in northeastern North America by its larger size (&gt;4.4mm, from clypeus to abdominal apex), and the apical portion of the median lobe of the aedeagus, which is distinctly thin and sinuate in lateral view (Fig. 18).
</paragraph>
<paragraph pageId="15" pageNumber="44">
<paragraph id="CD6327C7BF49CA7B5FD9C0024EF641B1" pageId="15" pageNumber="44">
This species is widely distributed and was previously known from Alaska, British Columbia, Illinois, Maine, Massachusetts, New Brunswick, New Hampshire, Newfoundland, New Jersey, New York, Northwest Territories, Nova Scotia, Ontario,
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="E54D284C95FF3628993F83E2BC4080A6" originalValue="Québec">Quebec</normalizedToken>
, Washington, Wisconsin (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
<bibRefCitation id="367B7869848C6A58527615DEA1147DA0" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
),Virginia (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">Frank 1981a</bibRefCitation>
<bibRefCitation id="8BDBB74413E1A6E8B77D1261B104D1C3" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">Frank 1981a</bibRefCitation>
), and Rhode Island (
<bibRefCitation author="Sikes, DS" journalOrPublisher="Zootaxa" pageId="27" pageNumber="56" pagination="1 - 38" title="The beetle fauna of the state of Rhode Island, USA (Coleoptera): 656 new state records." volume="340" year="2003">Sikes 2003</bibRefCitation>
<bibRefCitation id="A28F9C13CD4AE0621360325ABBC38E89" author="Sikes, DS" journalOrPublisher="Zootaxa" pageId="27" pageNumber="56" pagination="1 - 38" title="The beetle fauna of the state of Rhode Island, USA (Coleoptera): 656 new state records." volume="340" year="2003">Sikes 2003</bibRefCitation>
). Herein we newly report this species from Prince Edward Island (Map 22).
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<taxonomicName id="C1E473EBE02AEE5E25FFEE44E8B4FDB3" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus</taxonomicName>
is found in a variety of habitats near water and can be collected in great numbers by sifting or treading the debris present there.
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<paragraph pageId="4" pageNumber="33">Materials.</paragraph>
<paragraph pageId="4" pageNumber="33">CANADA: ON: Bruce Co., Dorcas Bay, dunes, pans under malaise, 5 to 13-VI-1999, S.A. Marshall (1).</paragraph>
<subSubSection id="D649BA4AA21C3A71B4B58C9480002672" pageId="4" pageNumber="33" type="materials">
<paragraph id="348FD32B2659CD36988EC4E34116B431" pageId="4" pageNumber="33">Materials.</paragraph>
<paragraph id="B87B53B0462D2927062B0D1FB417C962" pageId="4" pageNumber="33">CANADA: ON: Bruce Co., Dorcas Bay, dunes, pans under malaise, 5 to 13-VI-1999, S.A. Marshall (1).</paragraph>
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<paragraph pageId="4" pageNumber="33">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="33">
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<paragraph id="DB26420007B4DCD9D746E8209534408A" pageId="4" pageNumber="33">Diagnosis.</paragraph>
<paragraph id="7E27B7FC310D859132798FF9239C4799" pageId="4" pageNumber="33">
<taxonomicName id="2779D018353BA9570133AB5465EE60CC" class="Insecta" family="Staphylinidae" genus="Ischnosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnosoma flavicolle" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="flavicolle">Ischnosoma flavicolle</taxonomicName>
is easily distinguished from others of the genus by the distinctly bicolored elytra that each lack a humeral spot (
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) (Fig. 3).
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This species is primarily southeastern in distribution, with a northward extension along the Atlantic coast, and was known previously from Alabama, Arkansas, District of Columbia, Florida, Georgia, Illinois, Kansas, Kentucky, Louisiana, Maryland, Massachusetts, Mississippi, Missouri, New Hampshire, New Jersey, New York, North Carolina, Oklahoma, South Carolina, Tennessee, Texas, and Virginia (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 169" title="A revision of the genera Mycetoporus Mannerheimand Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) of North and Central America." volume="156" year="1991">Campbell 1991</bibRefCitation>
<bibRefCitation id="A759ACBF0CC67FC0524E8618237C769F" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 169" title="A revision of the genera Mycetoporus Mannerheimand Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) of North and Central America." volume="156" year="1991">Campbell 1991</bibRefCitation>
). Herein we here record this species from Canada (Ontario) for the first time (Map 3). This record represents a significant range expansion and is surprising since the forest of the Bruce Peninsula is known for its dominant boreal elements compared to the more
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Carolinian forests in south-western Ontario (
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). Throughout its range
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is frequently collected from sifted litter in a variety of forest types, and hammocks in the southern extremes; it has also been found in grasslands, carrion, under bark (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 169" title="A revision of the genera Mycetoporus Mannerheimand Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) of North and Central America." volume="156" year="1991">Campbell 1991</bibRefCitation>
<bibRefCitation id="4FB12B51A46C533283B918CA9642A0C5" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 169" title="A revision of the genera Mycetoporus Mannerheimand Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) of North and Central America." volume="156" year="1991">Campbell 1991</bibRefCitation>
), and in stream drift (
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
<bibRefCitation id="3CD6D765E35B55B33A9D1043B764AA64" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
). The record reported here is from an inland dune on the Lake Huron side of the Bruce Peninsula. Further collecting is necessary to delimit the full range of this apparently widespread species.
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<taxonomicName authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus couleensis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="couleensis">Philonthus couleensis Hatch, 1957</taxonomicName>
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<paragraph pageId="18" pageNumber="47">Materials.</paragraph>
<paragraph pageId="18" pageNumber="47">
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<paragraph id="37B6177452D3ED4902F01097DA2E613B" pageId="18" pageNumber="47">Materials.</paragraph>
<paragraph id="214D09EC2CEA2746CBEC80D7A57690C9" pageId="18" pageNumber="47">
CANADA: PEI: Brackley Beach, National Park, milieu
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(=marshy environment), 2-VIII-1979, R. Sexton (1); Long Pond, National Park, milieu
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(=marshy environment), 30-VII-1979, R. Sexton (2).
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<paragraph pageId="18" pageNumber="47">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="47">
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<paragraph id="57C5FA2522029423184020BE5B42C371" pageId="18" pageNumber="47">Diagnosis.</paragraph>
<paragraph id="BB7AD218FF598B42C2E457AFEA72FF43" pageId="18" pageNumber="47">
This species is, at present, best identified by the shape of the median lobe of the aedeagus and branches of the paramere (Fig. 499 in
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="6CA3DBE6C78D7063B679DB100F13F579" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
).
</paragraph>
<paragraph pageId="18" pageNumber="47">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus couleensis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="couleensis">Philonthus couleensis</taxonomicName>
<paragraph id="226AF87D9E79A882F36F2D1D5BE52BA9" pageId="18" pageNumber="47">
<taxonomicName id="3C04BA58F023925BE34E292DE07B3C14" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus couleensis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="couleensis">Philonthus couleensis</taxonomicName>
was previously known from Alberta, British Columbia, Idaho, Illinois, Indiana, Manitoba, Massachusetts, Michigan, New Brunswick, Newfoundland, New Jersey, Northwest Territories, New York, Nova Scotia, Ontario, Saskatchewan, Washington, and Wisconsin (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="B92CE2BD393287B4CC869CD488AD877B" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
). Herein we newly report it from Prince Edward Island (Map 29). This species is hygrophilous and has been collected from a variety of wet microhabitats near water (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
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).
</paragraph>
<caption pageId="18" pageNumber="47">
<paragraph pageId="18" pageNumber="47">
<caption id="AF07802C25A8C248F261F3827A445285" pageId="18" pageNumber="47">
<paragraph id="933531F92CBE9080FF170377D1A3523D" pageId="18" pageNumber="47">
Maps 29-32. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 29
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus couleensis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="couleensis">Philonthus couleensis</taxonomicName>
<taxonomicName id="E77FA0DEE6F97E163B1B3F3B0C640D8D" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus couleensis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="couleensis">Philonthus couleensis</taxonomicName>
Hatch (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
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) 30
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus gavius" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="gavius">Philonthus gavius</taxonomicName>
<taxonomicName id="A3C1961FC3435B7B5B5BC5357A5151AB" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus gavius" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="gavius">Philonthus gavius</taxonomicName>
Smetana 31
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus leechensis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="leechensis">Philonthus leechensis</taxonomicName>
<taxonomicName id="EDBDFB12AADA3FA79A74B970AC654A22" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus leechensis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="leechensis">Philonthus leechensis</taxonomicName>
Hatch (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="290F97F226AA786F2FFCDC7F59235C9F" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
) 32
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus lindrothi" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="lindrothi">Philonthus lindrothi</taxonomicName>
<taxonomicName id="B8F2FCE1F92B8D4618D0EAA5B149A76D" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus lindrothi" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="lindrothi">Philonthus lindrothi</taxonomicName>
Smetana (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="E4246F85536B08A324B9ADBD7F12D233" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
,
<bibRefCitation author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
<bibRefCitation id="57DB0D1CD3CA0FE7483C7AC0C7660DE7" author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
Paquin and
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2001
</bibRefCitation>
).

88
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<taxonomicName authority="Campbell, 1979" authorityName="Campbell" authorityYear="1979" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus ornatus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="ornatus">Tachyporus ornatus Campbell, 1979</taxonomicName>
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<paragraph pageId="8" pageNumber="37">Materials.</paragraph>
<paragraph pageId="8" pageNumber="37">CANADA: ON: Wellington Co., Belwood Lake, lake margin, fallen log overhang, 3-VI-2008, S.A. Marshall (1).</paragraph>
<subSubSection id="7F3D9E39472CD6CD9C45FED2B79695BD" pageId="8" pageNumber="37" type="materials">
<paragraph id="E37519FDBD2A38076A9FA58160AF044A" pageId="8" pageNumber="37">Materials.</paragraph>
<paragraph id="AAAA65915DCB9AA6EE746CE8D8D5A9B8" pageId="8" pageNumber="37">CANADA: ON: Wellington Co., Belwood Lake, lake margin, fallen log overhang, 3-VI-2008, S.A. Marshall (1).</paragraph>
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<paragraph pageId="8" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="37">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus ornatus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="ornatus">Tachyporus ornatus</taxonomicName>
<subSubSection id="8DE80F6EE0230EAA7BABCCEBFBE94390" pageId="8" pageNumber="37" type="diagnosis">
<paragraph id="4F13EE74A6DC884FBF4E6E9FC946A3BE" pageId="8" pageNumber="37">Diagnosis.</paragraph>
<paragraph id="77DE09A5372D73D249DCFFF7C1B8C4E9" pageId="8" pageNumber="37">
<taxonomicName id="00AF2AFCA30A11998C0B80F9EB21DD99" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus ornatus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="ornatus">Tachyporus ornatus</taxonomicName>
can be distinguished from all other large northeastern members of the genus except
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus lecontei" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="lecontei">Tachinus lecontei</taxonomicName>
<taxonomicName id="C4830A4F1B88579CCAFC97954ED9CF37" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus lecontei" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="lecontei">Tachinus lecontei</taxonomicName>
,by the combination of a non-bicolored abdomen and the crisp, dark markings on the elytra. From
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus lecontei" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="lecontei">Tachinus lecontei</taxonomicName>
<taxonomicName id="E6BB3D4820705533D0EEB70207BB8DDB" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus lecontei" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="lecontei">Tachinus lecontei</taxonomicName>
it is most easily identified by the fine microsculpture of the elytra which produces a strong metallic sheen (the former species completely lacks microsculpture).
</paragraph>
<paragraph pageId="8" pageNumber="37">
<paragraph id="8FE45977020C562D5F5627BB34EA5DF5" pageId="8" pageNumber="37">
This species is transcontinental in North America with a disjunct population in the Rocky Mountains of Colorado. It was previously known from the following states and provinces: Alberta, Colorado, Manitoba, Massachusetts, New Hampshire, New Jersey, New York, North Dakota, Pennsylvania,
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="FC1B4094691AC7E3846568FF7D3BA992" originalValue="Québec">Quebec</normalizedToken>
, Saskatchewan, and Vermont. Herein we newly record it from Ontario (Map 11). The only habitat data in
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." volume="109" year="1979">Campbell (1979)</bibRefCitation>
<bibRefCitation id="A66DF572AA254241A7743D08664B04BE" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." volume="109" year="1979">Campbell (1979)</bibRefCitation>
- &quot;treading under Alnus&quot;, and the lakeside habitat of the Ontario specimen recorded here suggest an affinity for decaying organic matter near water, but further collecting is necessary to confirm this.
</paragraph>
</subSubSection>

96
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<taxonomicName authority="Zetterstedt, 1828" authorityName="Zetterstedt" authorityYear="1828" class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites fenestralis" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="fenestralis">Porrhodites fenestralis (Zetterstedt, 1828)</taxonomicName>
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</paragraph>
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<subSubSection pageId="3" pageNumber="32" type="materials">
<paragraph pageId="3" pageNumber="32">Materials.</paragraph>
<paragraph pageId="3" pageNumber="32">CANADA: ON:Thunder Bay Distr. Pukaskwa Natl. Pk. beach trail, dunes, 30-VII-2003, S.M. Paiero (1).</paragraph>
<subSubSection id="0B41E58F6769C8B7710506F8A333BA9F" pageId="3" pageNumber="32" type="materials">
<paragraph id="2517EC9A1499A6521CA5767BE6110B5A" pageId="3" pageNumber="32">Materials.</paragraph>
<paragraph id="5BB3F89F9D48BFAF0583FC46745FFDF9" pageId="3" pageNumber="32">CANADA: ON:Thunder Bay Distr. Pukaskwa Natl. Pk. beach trail, dunes, 30-VII-2003, S.M. Paiero (1).</paragraph>
</subSubSection>
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<paragraph pageId="3" pageNumber="32">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="32">
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<paragraph id="ADF54D6591B89642CC6878B3E71CDEDC" pageId="3" pageNumber="32">Diagnosis.</paragraph>
<paragraph id="12729C253F8F803C56BA88FC263C5D74" pageId="3" pageNumber="32">
This species can be readily distinguished from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites inflatus" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="inflatus">Porrhodites inflatus</taxonomicName>
<taxonomicName id="B2E1F7112FD7DA0A7994F8625D6964B5" class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites inflatus" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="inflatus">Porrhodites inflatus</taxonomicName>
(Hatch), the only other member of the genus in North America, by the combination of: pronotal margins evenly arcuate; metasternum without microsculpture; antennomere two distinctly longer than segment three (Campbell 1984) (Fig. 2).
</paragraph>
<paragraph lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites fenestralis" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="fenestralis">Porrhodites fenestralis</taxonomicName>
<paragraph id="D16E38D30D4180D5EF11E38D710605D3" lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32">
<taxonomicName id="0C2CFCCA974654B384495BF7D5F43571" class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites fenestralis" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="fenestralis">Porrhodites fenestralis</taxonomicName>
is a holarctic, boreal to subarctic species known in the Nearctic region from Alberta, British Columbia, Manitoba, Newfoundland, Northwest Territories, Quebec, Yukon Territory, and Alaska, with a relict population in the Rocky Mountains of Montana and Wyoming (Campbell 1984).In the Palaearctic
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<pageBreakToken id="F3A0376ACABFE717CF4160333525AE8E" pageId="4" pageNumber="33" start="start">it</pageBreakToken>
is known from Austria, Czech Republic, Finland, Germany, Italy, Norway, Russia (North European Territory, Far East, East Siberia, and West Siberia), Sweden, and Switzerland (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="B155E1A9A5B5D9907F51B5CA5CE6BC55" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
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and Smetana 2004
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).
<bibRefCitation author="Ganglbauer, L" journalOrPublisher="Carl Gerold's Sohn, Wien" pageId="26" pageNumber="55" title="Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebeites. 2. Familienreihe Staphylinoidea. Theil I. Staphylinidae, Pselaphidae" year="1895">Ganglbauer (1895)</bibRefCitation>
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reported it from &quot;Lake Superior&quot;, which could be Michigan, Minnesota or Ontario; the above specimen thus represents the first Ontario record of this species (Map 2). The new locality in Ontario confirms that
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<taxonomicName id="118DB1B10D70E83ACE2C3C26A9A9C5DA" class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites fenestralis" order="Coleoptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="fenestralis">Porrhodites fenestralis</taxonomicName>
is transboreal in Canada. This species is typically collected in summer to early fall, and most known specimens with collection data were captured in flight; a long series was found swarming on a pine (
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="genus">Pinus</taxonomicName>
<taxonomicName id="62533618DF4C1557B4129D99456BE461" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="4" pageNumber="33" phylum="Tracheophyta" rank="genus">Pinus</taxonomicName>
)tree (Campbell 1984). Another specimen was found in a deer mouse (
<taxonomicName class="Mammalia" family="Sciuridae" genus="Pteromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteromyscus" order="Rodentia" pageId="4" pageNumber="33" phylum="Chordata" rank="genus">Pteromyscus</taxonomicName>
<taxonomicName id="729F13CA78E1A4293737E08369B81083" class="Mammalia" family="Sciuridae" genus="Pteromyscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteromyscus" order="Rodentia" pageId="4" pageNumber="33" phylum="Chordata" rank="genus">Pteromyscus</taxonomicName>
) nest (Campbell 1984).
</paragraph>
</subSubSection>

146
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<mods:title id="4ABAE63C53EE94A92C71E3D1E2998BC8">Contributions to the faunistics and bionomics of Staphylinidae (Coleoptera) in northeastern North America: discoveries made through study of the University of Guelph Insect Collection, Ontario, Canada</mods:title>
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<treatment ID-GBIF-Taxon="159361024" LSID="urn:lsid:plazi:treatment:79DEC4F84AAD77744BD2601C0C9261DC" httpUri="http://treatment.plazi.org/id/79DEC4F84AAD77744BD2601C0C9261DC" lastPageId="3" lastPageNumber="32" pageId="1" pageNumber="30">
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<paragraph pageId="1" pageNumber="30">
<taxonomicName authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" class="Insecta" family="Staphylinidae" genus="Omalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omalium repandum" order="Coleoptera" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="repandum">Omalium repandum Erichson, 1840</taxonomicName>
<treatment id="79DEC4F84AAD77744BD2601C0C9261DC" ID-GBIF-Taxon="159361024" LSID="urn:lsid:plazi:treatment:79DEC4F84AAD77744BD2601C0C9261DC" httpUri="http://treatment.plazi.org/id/79DEC4F84AAD77744BD2601C0C9261DC" lastPageId="3" lastPageNumber="32" pageId="1" pageNumber="30" scope_family="Staphylinidae" scope_order="Coleoptera">
<subSubSection id="63E08F41D68879025CE13CD87EE1A73C" pageId="1" pageNumber="30" type="nomenclature">
<paragraph id="8B3DC02FCCD3DCC3ACBCFA41B5957D33" pageId="1" pageNumber="30">
<taxonomicName id="66ADFF1371533285B4F63321EF8C0094" ID-CoL="499Y6" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" class="Insecta" family="Staphylinidae" genus="Omalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omalium repandum" order="Coleoptera" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="repandum">Omalium repandum Erichson, 1840</taxonomicName>
</paragraph>
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<subSubSection pageId="1" pageNumber="30" type="materials">
<paragraph pageId="1" pageNumber="30">Materials.</paragraph>
<paragraph pageId="1" pageNumber="30">
<subSubSection id="C54887A1E68CB4B9E63BEDA265B9A556" pageId="1" pageNumber="30" type="materials">
<paragraph id="886760B9D8A186FC68C6459FA3BF5759" pageId="1" pageNumber="30">Materials.</paragraph>
<paragraph id="558095B01CD98C308007A7FF840C87B2" pageId="1" pageNumber="30">
CANADA: ON: Chatam-Kent Co., Tilbury, pitfall trap, 9-VI-1994 (2), T. Sawinksi; Essex Co., Leamington, pitfall trap, 15 to 22-V-1992 (1), 22-V-1992 (1), 9-VI-1993 (1), Palichuck; Point Pelee Natl. Pk., Visitor Centre, malaise and pans, 22 to 29-V-2000, (2), 26-IX to 10-sX-2000 (1) O. Lonsdale. Huron Co., Auburn, Londesboro Rd. nr. Hwy 8, 43.728, -81.529, hedgerow, pitfall, 26-X-2009, A. Brunke (1); Benmiller, Sharpes Creek Line, 43.691, -81.608, hedgerow nr. creek, pitfall, 2-XI-2009, A. Brunke (1); Brucefield, London Rd. nr. Centennial Rd., 43.509, -81.516, hedgrerow nr. creek, pitfall, 27-IX-2009 (1), 12-X-2009 (4) A. Brunke; Waterloo Reg., Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, hedgerow, pitfall, 28-IX-2009 (6), 13-X-2009 (2) A. Brunke; Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, hedgerow, canopy trap in buckthorn, 10-XI-2009, A. Brunke (1); Blair, Fountain St. S. nr Speed River, 43.391, -80.373, hedgerow, pitfall, 24-XI-2009, A. Brunke (1); Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, hedgerow, canopy trap in buckthorn, 18-V-2010, A. Brunke (2); Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, hedgerow, pitfall, 1-VI-2010, A. Brunke (1); Blair, Whistlebare Rd. and Township Rd. 1, 43.367, -80.358, hedgerow, canopy trap in buckthorn, 18-V-2010 (1), 21-IX-2010 (1), A. Brunke; Blair, Whistlebare Rd. and Township Rd. 1, 43.367, -80.358, hedgerow, pitfall, 5-X-2010, A. Brunke (1); St. Jacobs, 'Stuart
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, 3-VI-1993 (1), 14-VI-1994 (1), T. Sawinksi.
</paragraph>
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<subSubSection lastPageId="3" lastPageNumber="32" pageId="1" pageNumber="30" type="diagnosis">
<paragraph pageId="1" pageNumber="30">Diagnosis.</paragraph>
<paragraph lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30">
<subSubSection id="DF1BA37F8F885DE6E62E369C7C169550" lastPageId="3" lastPageNumber="32" pageId="1" pageNumber="30" type="diagnosis">
<paragraph id="5AB4861D0BA1F620BB6B0233FA56CC94" pageId="1" pageNumber="30">Diagnosis.</paragraph>
<paragraph id="80A9F5F63A273C51782F32CCB3B063E6" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30">
As the genus
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<taxonomicName id="A21B2E7462D618B93FED47A8649AF511" class="Insecta" family="Staphylinidae" genus="Omalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omalium" order="Coleoptera" pageId="1" pageNumber="30" phylum="Arthropoda" rank="genus">Omalium</taxonomicName>
currently lacks a rigorous definition and consists of a heterogeneous assemblage of species (
<bibRefCitation author="Newton, A" editor="Arnett, RH" journalOrPublisher="CRC Press, New York" pageId="27" pageNumber="56" title="Staphylinidae Latrielle, 1802" volumeTitle="American beetles: Archostemata, Myxophaga, Adephaga Polyphaga: Staphyliniformia" year="2000">Newton et al. 2000</bibRefCitation>
<bibRefCitation id="39C4AA9CF2FD4B0899447BC95AEB507B" author="Newton, A" editor="Arnett, RH" journalOrPublisher="CRC Press, New York" pageId="27" pageNumber="56" title="Staphylinidae Latrielle, 1802" volumeTitle="American beetles: Archostemata, Myxophaga, Adephaga Polyphaga: Staphyliniformia" year="2000">Newton et al. 2000</bibRefCitation>
), this species is distinguished from all others of the subfamily occurring in the northeast by the combination of: tarsomere five longer than one to four combined; tarsomeres one to four not conspicuously broadened and without dense setae ventrally; maxillary palpomere three not greatly enlarged relative to segment four (
<bibRefCitation author="Newton, A" editor="Arnett, RH" journalOrPublisher="CRC Press, New York" pageId="27" pageNumber="56" title="Staphylinidae Latrielle, 1802" volumeTitle="American beetles: Archostemata, Myxophaga, Adephaga Polyphaga: Staphyliniformia" year="2000">Newton et al. 2000</bibRefCitation>
<bibRefCitation id="9CC284C1AF8B8165D0AA9C0F5D66C72A" author="Newton, A" editor="Arnett, RH" journalOrPublisher="CRC Press, New York" pageId="27" pageNumber="56" title="Staphylinidae Latrielle, 1802" volumeTitle="American beetles: Archostemata, Myxophaga, Adephaga Polyphaga: Staphyliniformia" year="2000">Newton et al. 2000</bibRefCitation>
); antennomeres
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<pageBreakToken id="B04E664AC7506CBB8A79DBEBC5F648A4" pageId="2" pageNumber="31" start="start">eight</pageBreakToken>
to ten elongate; pronotum bright orange with finely crenulate, evenly arcuate margins (Fig. 1).
</paragraph>
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<paragraph pageId="2" pageNumber="31">
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<paragraph id="1BBFE7B44C34C31DEF11230CF915E886" pageId="2" pageNumber="31">
Figures 1-6. Dorsal habitus. 1
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<taxonomicName id="5BB738CFF390B361794DBC3C6D36A4D1" class="Insecta" family="Staphylinidae" genus="Omalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omalium repandum" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="repandum">Omalium repandum</taxonomicName>
Erichson 2
<taxonomicName class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites fenestralis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="fenestralis">Porrhodites fenestralis</taxonomicName>
<taxonomicName id="7A8F13DC8092C8FD8534883F01F1FE30" class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites fenestralis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="fenestralis">Porrhodites fenestralis</taxonomicName>
(Zetterstedt) 3
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnosoma flavicolle" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="flavicolle">Ischnosoma flavicolle</taxonomicName>
<taxonomicName id="9F92FAF7EA79C41358ED0DD3271CA093" class="Insecta" family="Staphylinidae" genus="Ischnosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnosoma flavicolle" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="flavicolle">Ischnosoma flavicolle</taxonomicName>
(LeConte) 4
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
<taxonomicName id="C4A1C7947C38815C7B977087BDE1C1F4" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
Campbell 5
<taxonomicName class="Insecta" family="Staphylinidae" genus="Eustilicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustilicus tristis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="tristis">Eustilicus tristis</taxonomicName>
<taxonomicName id="9A9976D6D426A7D01F8EA775E1E6370B" class="Insecta" family="Staphylinidae" genus="Eustilicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustilicus tristis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="tristis">Eustilicus tristis</taxonomicName>
(Melsheimer) 6
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius cephalicus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="cephalicus">Bisnius cephalicus</taxonomicName>
<taxonomicName id="9AAFEDF06BDF3B8EE4241A948D2E43C1" class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius cephalicus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="cephalicus">Bisnius cephalicus</taxonomicName>
(Casey).
</paragraph>
</caption>
<paragraph pageId="2" pageNumber="31">
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<paragraph id="69C5E3A31D5B132BB3C9739C2BA4C48A" pageId="2" pageNumber="31">
<taxonomicName id="F2AE5BC75A366C4CC3AAAF7D73648D7F" class="Insecta" family="Staphylinidae" genus="Omalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omalium repandum" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="repandum">Omalium repandum</taxonomicName>
was previously known from Missouri, South Carolina, Texas (
<bibRefCitation author="Herman, L" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="26" pageNumber="55" pagination="1 - 4218" title="Catalog of the Staphylinidae (Insecta, Coleoptera): 1758 to the end of the second millennium" volume="265" year="2001">Herman 2001</bibRefCitation>
<bibRefCitation id="DC6561F20A331C8FF9CABD6A16B01C64" author="Herman, L" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="26" pageNumber="55" pagination="1 - 4218" title="Catalog of the Staphylinidae (Insecta, Coleoptera): 1758 to the end of the second millennium" volume="265" year="2001">Herman 2001</bibRefCitation>
), Indiana (
<bibRefCitation author="Blatchley, WS" journalOrPublisher="The Nature Publishing CO., Indianapolis, Indiana" pageId="25" pageNumber="54" title="An illustrated descriptive catalog of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana" year="1910">Blatchley 1910</bibRefCitation>
<bibRefCitation id="413DF02845EB1BD43E813879249BFA67" author="Blatchley, WS" journalOrPublisher="The Nature Publishing CO., Indianapolis, Indiana" pageId="25" pageNumber="54" title="An illustrated descriptive catalog of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana" year="1910">Blatchley 1910</bibRefCitation>
), Minnesota, Massachusetts, Georgia (
<bibRefCitation author="Lundgren, RW" editor="Peck, SB" journalOrPublisher="Arthropods of Florida and Neighboring Land Areas" pageId="27" pageNumber="56" pagination="1 - 180" title="Family Staphylinidae" volume="16" year="1998">Lundgren 1998</bibRefCitation>
<bibRefCitation id="CC032EAB62B264D37E7F08DA2D6F6ACC" author="Lundgren, RW" editor="Peck, SB" journalOrPublisher="Arthropods of Florida and Neighboring Land Areas" pageId="27" pageNumber="56" pagination="1 - 180" title="Family Staphylinidae" volume="16" year="1998">Lundgren 1998</bibRefCitation>
) and Quebec (
<bibRefCitation author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
<bibRefCitation id="A01079D604B669026235F99A7FBC435C" author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
). Herein we provide the first specimen-based records of this species in Canada and newly record it from Ontario based on several, relatively recent collections made in southern Ontario (Map 1). The above specimens were caught by passive traps placed in or near forests and thus no microhabitat data was available but
<bibRefCitation author="Blatchley, WS" journalOrPublisher="The Nature Publishing CO., Indianapolis, Indiana" pageId="25" pageNumber="54" title="An illustrated descriptive catalog of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana" year="1910">Blatchley (1910)</bibRefCitation>
<bibRefCitation id="B155DB0C2C4A94DBFFBF900113240BB0" author="Blatchley, WS" journalOrPublisher="The Nature Publishing CO., Indianapolis, Indiana" pageId="25" pageNumber="54" title="An illustrated descriptive catalog of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana" year="1910">Blatchley (1910)</bibRefCitation>
reported it as 'frequent, under dead
<normalizedToken originalValue="leaves’">leaves'</normalizedToken>
<normalizedToken id="4950EAF9225CCA04E076A70CC96F0C1A" originalValue="leaves’">leaves'</normalizedToken>
.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Omalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omalium repandum" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="repandum">Omalium repandum</taxonomicName>
<taxonomicName id="7155E1C27366A1680C34B184F5D2C04A" class="Insecta" family="Staphylinidae" genus="Omalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omalium repandum" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="repandum">Omalium repandum</taxonomicName>
therefore appears to be forest litter dwelling and is probably widespread in central and eastern North America. The specimens from Huron County, Ontario and the provincial record from Quebec probably represent the northern limit of its range.
</paragraph>
<caption pageId="3" pageNumber="32">
<paragraph pageId="3" pageNumber="32">
<pageBreakToken pageId="3" pageNumber="32" start="start">Maps</pageBreakToken>
<caption id="F61D0998F064B02F5177E54451B1D68C" pageId="3" pageNumber="32">
<paragraph id="00221A8572E46E696D743E2C5E4D5E79" pageId="3" pageNumber="32">
<pageBreakToken id="841D301B0FEDBC941FD13A95D46A5F4D" pageId="3" pageNumber="32" start="start">Maps</pageBreakToken>
1-4. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 1
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<taxonomicName id="9DDC54DA32429754AB1C268EB852CBEC" class="Insecta" family="Staphylinidae" genus="Omalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omalium repandum" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="repandum">Omalium repandum</taxonomicName>
Erichson (
<bibRefCitation author="Blatchley, WS" journalOrPublisher="The Nature Publishing CO., Indianapolis, Indiana" pageId="25" pageNumber="54" title="An illustrated descriptive catalog of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana" year="1910">Blatchley 1910</bibRefCitation>
<bibRefCitation id="10FCDE9EA3E23EA90E5FCDA98FD6AC23" author="Blatchley, WS" journalOrPublisher="The Nature Publishing CO., Indianapolis, Indiana" pageId="25" pageNumber="54" title="An illustrated descriptive catalog of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana" year="1910">Blatchley 1910</bibRefCitation>
,
<bibRefCitation author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
<bibRefCitation id="BCA5F91845C6D87B1C348968590157BB" author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
,
<bibRefCitation author="Lundgren, RW" editor="Peck, SB" journalOrPublisher="Arthropods of Florida and Neighboring Land Areas" pageId="27" pageNumber="56" pagination="1 - 180" title="Family Staphylinidae" volume="16" year="1998">Lundgren 1998</bibRefCitation>
<bibRefCitation id="A7300B636AD10B7BB446F936D681E062" author="Lundgren, RW" editor="Peck, SB" journalOrPublisher="Arthropods of Florida and Neighboring Land Areas" pageId="27" pageNumber="56" pagination="1 - 180" title="Family Staphylinidae" volume="16" year="1998">Lundgren 1998</bibRefCitation>
) 2
<taxonomicName class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites fenestralis" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="fenestralis">Porrhodites fenestralis</taxonomicName>
<taxonomicName id="86C3E5AA0AF04F52A449F0F606C20769" class="Insecta" family="Staphylinidae" genus="Porrhodites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porrhodites fenestralis" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="fenestralis">Porrhodites fenestralis</taxonomicName>
(Zetterstedt) (Campbell 1984) 3
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnosoma flavicolle" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="flavicolle">Ischnosoma flavicolle</taxonomicName>
<taxonomicName id="EBE7E3E0436EEF28F5676B2BDAC05C55" class="Insecta" family="Staphylinidae" genus="Ischnosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnosoma flavicolle" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="flavicolle">Ischnosoma flavicolle</taxonomicName>
(LeConte) (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 169" title="A revision of the genera Mycetoporus Mannerheimand Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) of North and Central America." volume="156" year="1991">Campbell 1991</bibRefCitation>
<bibRefCitation id="730A728ED986FB83AB1A8BA9818B6143" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 169" title="A revision of the genera Mycetoporus Mannerheimand Ischnosoma Stephens (Coleoptera: Staphylinidae: Tachyporinae) of North and Central America." volume="156" year="1991">Campbell 1991</bibRefCitation>
) 4
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
<taxonomicName id="A1F7A2B6B417A8713887FFE9DD7FD21F" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus campbelli" order="Coleoptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="campbelli">Sepedophilus campbelli</taxonomicName>
Herman (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="5646B9C8819ED50D314049E2F82F6BB6" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
,
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
<bibRefCitation id="E9BFEAAEFF5CB958FA80C2818A09CF90" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
).
</paragraph>
</caption>

92
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@ -1,72 +1,72 @@
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<paragraph pageId="18" pageNumber="47">
<taxonomicName authority="LeConte, 1863" authorityName="LeConte" authorityYear="1863" class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis (LeConte, 1863)</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="47" type="materials">
<paragraph pageId="18" pageNumber="47">Materials.</paragraph>
<paragraph pageId="18" pageNumber="47">CANADA: ON: Essex Co.,Middle Is., shore, yellow pans, 11-VI-2003, S.A. Marshall (1); Niagara Reg., Grimsby, J. Pettit (1).</paragraph>
<subSubSection id="AC6F5C3BDB7D61892D9989D1FF86D524" pageId="18" pageNumber="47" type="materials">
<paragraph id="73974AA4374EFD39121894694C971A2F" pageId="18" pageNumber="47">Materials.</paragraph>
<paragraph id="75E509A134F7C2C16B7CEC3523E2CC0B" pageId="18" pageNumber="47">CANADA: ON: Essex Co.,Middle Is., shore, yellow pans, 11-VI-2003, S.A. Marshall (1); Niagara Reg., Grimsby, J. Pettit (1).</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="47" type="diagnosis">
<paragraph pageId="18" pageNumber="47">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="47">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis</taxonomicName>
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<paragraph id="4147F8EFEDEEE8B9711A626CED4EA52F" pageId="18" pageNumber="47">Diagnosis.</paragraph>
<paragraph id="F202A3E3C9347810B9232C67D65E1087" pageId="18" pageNumber="47">
<taxonomicName id="2D37D9A8FC0F07E06BEBEB041ACA5631" class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis</taxonomicName>
is easily recognized among other orange and black species of the genus in northeastern North America by the elytra with a broad, pale apical area (Fig. 10). In other northeastern species, this pale area is restricted to a narrow strip.
</paragraph>
<paragraph pageId="18" pageNumber="47">
<paragraph id="056D8AB704D32C01712383494FFAEC9C" pageId="18" pageNumber="47">
This species was previously known from Arizona, California, Colorado, Iowa, Maine, Maryland, Michigan, New Hampshire, New Mexico, New York, Nova Scotia, Pennsylvania,
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="04001FDE601020C54101F9391F059184" originalValue="Québec">Quebec</normalizedToken>
, Texas, and Virginia (
<bibRefCitation author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
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). Herein we newly record it from Ontario (Map 28). It is also known from Costa Rica, Guatemala, Mexico, and Panama (unverified record) (
<bibRefCitation author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
<bibRefCitation id="248D995AC769855908F1CF005743519D" author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
). Unlike
<taxonomicName class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius occidentoides" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="occidentoides">Neobisnius occidentoides</taxonomicName>
<taxonomicName id="ED147701ECF7E385F738F4B21B25FB5D" class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius occidentoides" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="occidentoides">Neobisnius occidentoides</taxonomicName>
,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis</taxonomicName>
<taxonomicName id="0469FAD7C4C09B0325197F9CFEA98CE1" class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis</taxonomicName>
is strongly associated with the margins of rivers and lakes and is found in litter or under debris. The specimen recorded here from
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Middle Island (in Lake Erie) was taken in pan traps on a gravel lake shore.
</paragraph>
</subSubSection>

100
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leechensis Hatch, 1957
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<paragraph pageId="19" pageNumber="48">Materials.</paragraph>
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<paragraph id="79532B7B6D8ACDBBBD2686D0701FBC91" pageId="19" pageNumber="48">Materials.</paragraph>
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CANADA: PEI: Brackley Beach, National Park, milieu
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(=marshy environment), 2-VIII-1979, R. Sexton, (3).
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<paragraph pageId="19" pageNumber="48">Diagnosis.</paragraph>
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may be recognized among other northeastern
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except for
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by the combination of: pronotum with five punctures in the dorsal row on at least one side; temples without a carina; tergite eight not emarginate in either sex; elytra without distinct markings on the disc; hind tarsus with first segment shorter than last segment; legs completely dark. From
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it can be differentiated by the distinct, obtuse hind angles of the head.
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species was previously known from Alaska, Alberta, Arizona, British Columbia, California, Colorado, Idaho, Manitoba, Minnesota, Montana, Newfoundland, Northwest Territories, Oregon,
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, Saskatchewan, Washington, Wisconsin, and Yukon Territory (
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). Herein we newly record it from Prince Edward Island, representing only the fifth locality known in northeastern North America (Map 31).
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appears to be a hygrophilous species, occurring mostly in northern or montane wetlands. It remains unrecorded in the eastern United States.
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<taxonomicName id="63135889E88E5D777434A9FC8BC298FE" ID-CoL="76ZCV" authority="Smetana, 1995" authorityName="Smetana" authorityYear="1995" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus gavius" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="gavius">Philonthus gavius Smetana, 1995</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="47" type="materials">
<paragraph pageId="18" pageNumber="47">Materials.</paragraph>
<paragraph pageId="18" pageNumber="47">UNITED STATES: MD:Prince George Co., Cedarville St. Pk., at lights, 24 to 29-VII-2008, S. Paiero (1).</paragraph>
<subSubSection id="C69FD884027FE2905A05A6B10D0C229D" pageId="18" pageNumber="47" type="materials">
<paragraph id="545BE1AD65CEB252EB42BDACBFBE4B95" pageId="18" pageNumber="47">Materials.</paragraph>
<paragraph id="AFD00B71219D38FA59E5E1946C6CFB96" pageId="18" pageNumber="47">UNITED STATES: MD:Prince George Co., Cedarville St. Pk., at lights, 24 to 29-VII-2008, S. Paiero (1).</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="47" type="diagnosis">
<paragraph pageId="18" pageNumber="47">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="47">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus gavius" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="gavius">Philonthus gavius</taxonomicName>
<subSubSection id="5B10D2B1FEBF7C50CB73E20883F857D0" pageId="18" pageNumber="47" type="diagnosis">
<paragraph id="664B5D3515B8DD6D97091F0D950C4AA0" pageId="18" pageNumber="47">Diagnosis.</paragraph>
<paragraph id="0B2384CB881AA5ECD80D8C12D86E5D47" pageId="18" pageNumber="47">
<taxonomicName id="07F72578851075634B50C491BB0CA872" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus gavius" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="gavius">Philonthus gavius</taxonomicName>
is recognized among other northeastern
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
<taxonomicName id="A01E9BBFCE224C38FBAABF383AB33896" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
by the combination of: tergite eight distinctly emarginate in both sexes; elytra with most punctures separated by their widths; eyes longer than the temples.
</paragraph>
<paragraph pageId="18" pageNumber="47">
<paragraph id="661C9FFB312D30CA82FEECB52A848323" pageId="18" pageNumber="47">
This species was previously known from Arkansas, Illinois, Louisiana, Oklahoma, Tennessee, and Texas (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="34C3EBCCAC88ABE6E4C0CF07B413C988" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
). It was newly recorded from Missouri by
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
<bibRefCitation id="FC4E56973268999D46E5566DAA45870B" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
based on one specimen at a blacklight. Another specimen collected at a mercury vapour light represents a new record for Maryland (Map 30) and northeastern North America, and extends the known distribution considerably eastward. The habitat preferences of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus gavius" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="gavius">Philonthus gavius</taxonomicName>
<taxonomicName id="4872562140720B620E4990E482DA4675" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus gavius" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="gavius">Philonthus gavius</taxonomicName>
remain unknown but it is probably a species associated with wet areas similar to most other members of the
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
<taxonomicName id="A4581031C07C3882238E793D0BAC7C95" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
'Quadricollis
<normalizedToken originalValue="Group’">Group'</normalizedToken>
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sensu
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana (1995)</bibRefCitation>
<bibRefCitation id="41996A58E78CB9EF656AF2C530A44495" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana (1995)</bibRefCitation>
.
</paragraph>
</subSubSection>

128
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@ -1,107 +1,107 @@
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<paragraph pageId="13" pageNumber="42">
<taxonomicName authority="Frank, 1975" authorityName="Frank" authorityYear="1975" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius brachycephalus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="brachycephalus">Erichsonius brachycephalus Frank, 1975</taxonomicName>
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<taxonomicName id="A3217EAD3A5A9DD8DE7B9BA3431EE6F7" ID-CoL="6GMN9" authority="Frank, 1975" authorityName="Frank" authorityYear="1975" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius brachycephalus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="brachycephalus">Erichsonius brachycephalus Frank, 1975</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="42" type="materials">
<paragraph pageId="13" pageNumber="42">Materials.</paragraph>
<paragraph pageId="13" pageNumber="42">
<subSubSection id="9537CD950C11E21865F71D8EA28884D5" pageId="13" pageNumber="42" type="materials">
<paragraph id="08FCF3B5FF03D8AF2386BBA454D6AC77" pageId="13" pageNumber="42">Materials.</paragraph>
<paragraph id="8863D4B9445F943B6D632986B221DE17" pageId="13" pageNumber="42">
CANADA: ON: Huron Co., Brucefield, London Rd. nr. Centennial Rd., 43.509, -81.516, hedgerow nr. creek, canopy trap in buckthorn, 11-V-2009, A. Brunke (1); QC:La
<normalizedToken originalValue="Valleé-de-la-Gatinaeu">Vallee-de-la-Gatinaeu</normalizedToken>
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,Martindale, hutte
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<normalizedToken id="99C008BC42B8FE4B01BA98CE52058534" originalValue="à">a</normalizedToken>
castor (=beaver lodge), 19-IX-1976, R. Sexton (26).
</paragraph>
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<paragraph pageId="13" pageNumber="42">Diagnosis.</paragraph>
<paragraph pageId="13" pageNumber="42">This species is easily recognized among others of the genus with a sparsely punctate forebody by its large size (&gt;4.7mm from clypeus to abdominal apex) and transverse head with slightly converging temples (Fig. 8).</paragraph>
<paragraph lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius brachycephalus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="brachycephalus">Erichsonius brachycephalus</taxonomicName>
<subSubSection id="A11EC7A4C6E9458CB26BA067771E5B5D" lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42" type="diagnosis">
<paragraph id="FE30702DE676816FC38207439CA32B21" pageId="13" pageNumber="42">Diagnosis.</paragraph>
<paragraph id="B133D494C6BECC11DB3B9FA4B078805E" pageId="13" pageNumber="42">This species is easily recognized among others of the genus with a sparsely punctate forebody by its large size (&gt;4.7mm from clypeus to abdominal apex) and transverse head with slightly converging temples (Fig. 8).</paragraph>
<paragraph id="A9A4D48C376302B81DEFA1F339D4F5F0" lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42">
<taxonomicName id="BF0C6E3BEA2AE0C89754D6627E907C2C" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius brachycephalus" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="brachycephalus">Erichsonius brachycephalus</taxonomicName>
was previously known from Illinois, Maine, Massachusetts, New Jersey, Texas (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
<bibRefCitation id="F3357E49EC6B1D2BFA7677653E171545" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
),Virginia (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">Frank 1981a</bibRefCitation>
<bibRefCitation id="D13DE80798B1DCF9A93AEAC536F9BD8B" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">Frank 1981a</bibRefCitation>
), and New Hampshire (
<bibRefCitation author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
<bibRefCitation id="0EB152AA75937888C83F1E418C3F1C7A" author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
).
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
<bibRefCitation id="D38AE897B6BDCFC3644494C85F74DBA3" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
reported it from Missouri based on specimens collected in stream drift and leaf litter. Herein we newly record it from Canada (Ontario
<pageBreakToken pageId="14" pageNumber="43" start="start">and</pageBreakToken>
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<pageBreakToken id="4E2CC5C4552CAAD7EDF871433453C05E" pageId="14" pageNumber="43" start="start">and</pageBreakToken>
<normalizedToken id="50C45C728ECBCE267E2B06DCEC387C3A" originalValue="Québec">Quebec</normalizedToken>
) and suggest that this species is broadly distributed in northeastern North America, reaching its northern limit in southernmost Canada (Map 21). All available habitat data suggest that this species is strongly associated with decaying vegetative debris along the edges of ponds, lakes, streams, and rivers. Despite the long series from Quebec beaver lodges, we have not found this species during our searches of beaver lodges in Ontario.
</paragraph>
<caption pageId="14" pageNumber="43">
<paragraph pageId="14" pageNumber="43">
<caption id="784F0230F3A6E43B91420BF6F33FE55E" pageId="14" pageNumber="43">
<paragraph id="31E5856E045B59002CFFDF739F59B10D" pageId="14" pageNumber="43">
Maps 21-24. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 21
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius brachycephalus" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="brachycephalus">Erichsonius brachycephalus</taxonomicName>
<taxonomicName id="16EC103FE4C1DC81F02BDA23155B65C3" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius brachycephalus" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="brachycephalus">Erichsonius brachycephalus</taxonomicName>
Frank (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
<bibRefCitation id="B5362946E4A3AC62E18C6162681F68CE" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
,
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">1981a</bibRefCitation>
<bibRefCitation id="072BD9CC6D6521DFD6C4932AAFD99D4E" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">1981a</bibRefCitation>
,
<bibRefCitation author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
<bibRefCitation id="5877679FBE749D749FDE9360610A6FE8" author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
) 22
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus</taxonomicName>
<taxonomicName id="F9B078F8F3F7E9DC6C4DF1A38022704A" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus</taxonomicName>
(Horn) (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
<bibRefCitation id="67EF4F854CAC9359AE5F74BB46DA9582" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
,
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">1981a</bibRefCitation>
<bibRefCitation id="CD5D0776E3F6F05D0226D89C10408A2D" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="97 - 106" title="A New Erichsonius Species from Arizona with Discussion on Phylogeny within the Genus (Coleoptera: Staphylinidae)." volume="35" year="1981 a">1981a</bibRefCitation>
,
<bibRefCitation author="Sikes, DS" journalOrPublisher="Zootaxa" pageId="27" pageNumber="56" pagination="1 - 38" title="The beetle fauna of the state of Rhode Island, USA (Coleoptera): 656 new state records." volume="340" year="2003">Sikes 2003</bibRefCitation>
<bibRefCitation id="FFB807B05529D87DDCD14BF5A0769D76" author="Sikes, DS" journalOrPublisher="Zootaxa" pageId="27" pageNumber="56" pagination="1 - 38" title="The beetle fauna of the state of Rhode Island, USA (Coleoptera): 656 new state records." volume="340" year="2003">Sikes 2003</bibRefCitation>
,
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimaszewski et al. 2005</bibRefCitation>
<bibRefCitation id="61C11004FD0146BC097FBA591D2A35EE" author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimaszewski et al. 2005</bibRefCitation>
) 23
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="4CAD9284CE4F642C387B2FFE4BB9E331" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
(Horn) (
<bibRefCitation author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
<bibRefCitation id="E67E44E6EEE2B7E025FEE409EA0F6B64" author="Frank, JH" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="177 - 203" title="A Revision of the New World Species of the Genus Erichsonius Fauvel (Coleoptera: Staphylinidae)." volume="29" year="1975">Frank 1975</bibRefCitation>
,
<bibRefCitation author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
<bibRefCitation id="8A541704919479311D4FB67337377188" author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
) 24
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius amulius" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="amulius">Gabrius amulius</taxonomicName>
<taxonomicName id="27B5FB61C29B415BDD6B2E80009D919D" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius amulius" order="Coleoptera" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="amulius">Gabrius amulius</taxonomicName>
Smetana (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="3DC52ED31B06996B241346D0C90A0F99" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
).
</paragraph>
</caption>

96
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@ -1,78 +1,78 @@
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<paragraph pageId="20" pageNumber="49">
<taxonomicName authority="Smetana, 1965" authorityName="Smetana" authorityYear="1965" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus lindrothi" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="lindrothi">Philonthus lindrothi Smetana, 1965</taxonomicName>
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<paragraph id="E37DDD43C4B077939BE3DB9726FD1FB7" pageId="20" pageNumber="49">
<taxonomicName id="2F284FB439B67C8BD50715D2FA485CB0" ID-CoL="4GB6B" authority="Smetana, 1965" authorityName="Smetana" authorityYear="1965" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus lindrothi" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="lindrothi">Philonthus lindrothi Smetana, 1965</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="49" type="materials">
<paragraph pageId="20" pageNumber="49">Materials.</paragraph>
<paragraph pageId="20" pageNumber="49">
<subSubSection id="5C0B8F4CA7591AD755FA7EF48F5C8D36" pageId="20" pageNumber="49" type="materials">
<paragraph id="9F83CA532DD6EEA4CCC5D78DAD1C40D6" pageId="20" pageNumber="49">Materials.</paragraph>
<paragraph id="C307BAD748409CCB692967F1B1A7008C" pageId="20" pageNumber="49">
CANADA: PEI: Brackley Beach, National Park, milieu
<normalizedToken originalValue="marécageux">marecageux</normalizedToken>
<normalizedToken id="0677DC1375D3580110C4CAE96693E166" originalValue="marécageux">marecageux</normalizedToken>
(=marshy environment), 2-VIII-1979, R. Sexton (6).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="49" type="diagnosis">
<paragraph pageId="20" pageNumber="49">Diagnosis.</paragraph>
<paragraph pageId="20" pageNumber="49">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus lindrothi" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="lindrothi">Philonthus lindrothi</taxonomicName>
<subSubSection id="CD2BF376AAEB6A578F158392FB181E5F" pageId="20" pageNumber="49" type="diagnosis">
<paragraph id="44F544BDC78BBD6B96E0D488D2FA83FF" pageId="20" pageNumber="49">Diagnosis.</paragraph>
<paragraph id="051D19B54D7CA2DBA5E06E05AF8922EE" pageId="20" pageNumber="49">
<taxonomicName id="AB4953899145301E2D83AC638EABF774" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus lindrothi" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="lindrothi">Philonthus lindrothi</taxonomicName>
can be distinguished fromnortheastern congeners other than
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus pseudolodes" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="pseudolodes">Philonthus pseudolodes</taxonomicName>
<taxonomicName id="883B913710457E8E95D36FC43F9FFB6E" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus pseudolodes" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="pseudolodes">Philonthus pseudolodes</taxonomicName>
Smetana 1996 by the combination of: pronotum with five punctures in the dorsal row on at least one side; head with hind angles present but rounded; tergite eight not emarginate in either sex; elytra without distinct markings on the disc; hind tarsus with first segment shorter than last segment; antennae with basal segments not distinctly paler than others and with segments seven and eight elongate. Males can be readily separated from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus pseudolodes" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="pseudolodes">Philonthus pseudolodes</taxonomicName>
<taxonomicName id="35F2105E859D3CFFD8737E46B05BA5EF" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus pseudolodes" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="pseudolodes">Philonthus pseudolodes</taxonomicName>
by the notch in sternite eight not continuing as a grove towards its base (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="38BD9418454A14AC110E1AC993A2D4B3" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
).
</paragraph>
<paragraph pageId="20" pageNumber="49">
<paragraph id="5048FEF0D2715E73483C1D4EF4CA3C6C" pageId="20" pageNumber="49">
This widespread species was previously known from Alaska, Alberta, Arizona, British Columbia, California, Colorado, Idaho, Illinois, Indiana, Iowa, Kansas, Maine, Manitoba, Massachusetts, Michigan, Minnesota, Missouri, Montana, Nebraska, Nevada, New Brunswick, Newfoundland, New Hampshire, New Jersey, New York, North Carolina, North Dakota, Northwest Territories, Nova Scotia, Ohio, Ontario, Oregon, Pennsylvania,
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="18DED04ED083C27EC76C33A88ED6DCBA" originalValue="Québec">Quebec</normalizedToken>
, Rhode Island, Saskatchewan, South Dakota, Vermont, Washington, and Wisconsin (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="5D968CE2E5E5E568FDD419E38E39BCD9" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
). Herein we newly record it from Prince Edward Island (Map 32).
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<taxonomicName id="02F4DA7C59D8B3C037BF6329F79735BA" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus lindrothi" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="lindrothi">Philonthus lindrothi</taxonomicName>
is an extremely common hygrophilous species that occasionally visits lights (
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).
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<paragraph pageId="18" pageNumber="47">Materials.</paragraph>
<paragraph pageId="18" pageNumber="47">CANADA: ON: Essex Co., East Sister Is. Prov. Nature Res., dry pond bed, yellow pans, 30-VII-2003, S.A. Marshall (1); Leamington, pitfall trap, 17-VIII-1993 (1); Huron Co., Centralia, Dev 1A, pitfall, 16-VIII-1992 (1); Middlesex Co., London, southern crop protection research centre, corn pitfalls 3, 19-VII-1993 (1); London, southern crop protection research centre, pitfall/Masner trap, 2-VIII-1995, T. Sawinski (1).</paragraph>
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<paragraph id="EE859E9A180C7CFC5AD13E91D0B256A7" pageId="18" pageNumber="47">Materials.</paragraph>
<paragraph id="D84AECFEC258C17528EBC9747AC1AC8A" pageId="18" pageNumber="47">CANADA: ON: Essex Co., East Sister Is. Prov. Nature Res., dry pond bed, yellow pans, 30-VII-2003, S.A. Marshall (1); Leamington, pitfall trap, 17-VIII-1993 (1); Huron Co., Centralia, Dev 1A, pitfall, 16-VIII-1992 (1); Middlesex Co., London, southern crop protection research centre, corn pitfalls 3, 19-VII-1993 (1); London, southern crop protection research centre, pitfall/Masner trap, 2-VIII-1995, T. Sawinski (1).</paragraph>
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<paragraph pageId="18" pageNumber="47">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="47">
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<paragraph id="3D008050D3E12AD9B519364DB1C12783" pageId="18" pageNumber="47">Diagnosis.</paragraph>
<paragraph id="05C28D7FA7E9F4ED502C0EBE4FA8444D" pageId="18" pageNumber="47">
This species can be distinguished from other orange and black
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in northeastern North America by the combination of head completely lacking microsculpture, elytra with apically paler area limited to a narrow strip, and the maxillary palpi with at least one segment darkened.
</paragraph>
<paragraph pageId="18" pageNumber="47">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius occidentoides" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="occidentoides">Neobisnius occidentoides</taxonomicName>
<paragraph id="BB2EAF41B89460422DEFF152AB825BD3" pageId="18" pageNumber="47">
<taxonomicName id="974B6F4FA62C9590183647D25A0B9BEA" class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius occidentoides" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="occidentoides">Neobisnius occidentoides</taxonomicName>
is a widespread species and was previously known from Alberta, Alabama, Arkansas, Arizona, California, Colorado, Idaho, Illinois, Kansas, Kentucky, Louisiana, Manitoba, Minnesota, Mississippi, Missouri, Montana, Nebraska, Nevada, New Mexico, New York, North Carolina, North Dakota, Ohio, Oklahoma, South Dakota, Tennessee, Texas, Utah, Vermont, Virginia, Washington, and Wyoming (
<bibRefCitation author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
<bibRefCitation id="830F8531758BEC2324E425DD270FBBAD" author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
). This species is also known from Mexico. Herein we newly record it from eastern Canada based on several collections made in the Carolinian region of southern Ontario (Map 27).
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is less strongly associated with water margins than are other bicolored species of the genus, and is frequently collected in agricultural fields with moist soil (
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).
</paragraph>
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<taxonomicName authority="Fabricius, 1792" authorityName="Fabricius" authorityYear="1792" class="Insecta" family="Staphylinidae" genus="Sunius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sunius melanocephalus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="melanocephalus">
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<paragraph id="F5B7B2E458334AAF6CDB764EF1AFC7AA" pageId="12" pageNumber="41">
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<pageBreakToken id="91D5C4B3C5695E6EA849607C0C8D6873" pageId="12" pageNumber="41" start="start">Sunius</pageBreakToken>
melanocephalus (Fabricius, 1792)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="41" type="materials">
<paragraph pageId="12" pageNumber="41">Materials.</paragraph>
<paragraph pageId="12" pageNumber="41">CANADA: ON: Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.742, -81.514, hedgerow, canopy trap in buckthorn, 26-V-2010 (1); Waterloo Reg., Blair, Whistlebare Rd. and Township Rd. 1, 43.367, -80.358, hedgerow, pitfall, 4-V-2010 (1); Wellington Co., Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, hedgerow, pitfall, 4-V-2010 (1).</paragraph>
<subSubSection id="3530889BE09B04CBA85A8EBE2E8817A6" pageId="12" pageNumber="41" type="materials">
<paragraph id="440EFC5F957966D213ACD062F235B96D" pageId="12" pageNumber="41">Materials.</paragraph>
<paragraph id="0413389E8F43DF67FB63C4EF17261768" pageId="12" pageNumber="41">CANADA: ON: Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.742, -81.514, hedgerow, canopy trap in buckthorn, 26-V-2010 (1); Waterloo Reg., Blair, Whistlebare Rd. and Township Rd. 1, 43.367, -80.358, hedgerow, pitfall, 4-V-2010 (1); Wellington Co., Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, hedgerow, pitfall, 4-V-2010 (1).</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="41" type="diagnosis">
<paragraph pageId="12" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="41">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sunius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sunius melanocephalus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="melanocephalus">Sunius melanocephalus</taxonomicName>
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<paragraph id="455E2831391CB53AE7FD23AD20AB20BC" pageId="12" pageNumber="41">Diagnosis.</paragraph>
<paragraph id="C98ECB0D9674095C14FF788B9458C3D5" pageId="12" pageNumber="41">
<taxonomicName id="F0B27FDC44096717C6CB896613B1A277" class="Insecta" family="Staphylinidae" genus="Sunius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sunius melanocephalus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="melanocephalus">Sunius melanocephalus</taxonomicName>
may be easily recognized among other northeastern members of the genus by the combination of the non-serrate lateral margins of the pronotum and the bicolored body.
</paragraph>
<paragraph pageId="12" pageNumber="41">
<paragraph id="8D5D3026C992F4E7D07A8F354D2687B7" pageId="12" pageNumber="41">
This species was accidentally introduced from the Palaearctic region to North America and was first recognized on the continent by
<bibRefCitation author="Hoebeke, ER" journalOrPublisher="Entomological News" pageId="26" pageNumber="55" pagination="19 - 24" title="Sunius melanocephalus (Coleoptera: Staphylinidae), a Palearctic rove beetle new to North America." volume="102" year="1991">Hoebeke (1991)</bibRefCitation>
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from specimens collected in New York as early as 1924. Since then, it has been detected in Pennsylvania, Vermont (
<bibRefCitation author="Byers, RA" journalOrPublisher="The Great Lakes Entomologist" pageId="26" pageNumber="55" pagination="81 - 106" title="Richness and abundance of Carabidae and Staphylinidae (Coleoptera) in northeastern dairy pastures under intensive grazing." volume="33" year="2000">Byers et al. 2000</bibRefCitation>
<bibRefCitation id="859B1076038A70FC614FA79169A91E57" author="Byers, RA" journalOrPublisher="The Great Lakes Entomologist" pageId="26" pageNumber="55" pagination="81 - 106" title="Richness and abundance of Carabidae and Staphylinidae (Coleoptera) in northeastern dairy pastures under intensive grazing." volume="33" year="2000">Byers et al. 2000</bibRefCitation>
), and
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="3199D79C4BA4331FC3484932CBD26874" originalValue="Québec">Quebec</normalizedToken>
(
<bibRefCitation author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
<bibRefCitation id="9C4BD23AC20D7AA6B507D259BB87DC9E" author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
). As specimen data were not provided for the
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="5539AD05222DE91E832C4FB2D8042F97" originalValue="Québec">Quebec</normalizedToken>
record, this
<normalizedToken originalValue="species’">species'</normalizedToken>
<normalizedToken id="D91BB632AFF071F8FD6F250C747EF782" originalValue="species’">species'</normalizedToken>
presence in Canada was uncertain. Herein we verify its occurrence in Canada and newly record it from Ontario based on collections made in 2010 (Map 17); no earlier collections of this species in Ontario were present in DEBU. In the Palaearctic region
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sunius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sunius melanocephalus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="melanocephalus">Sunius melanocephalus</taxonomicName>
<taxonomicName id="0536AF28B871D6B7F5785909255DB26B" class="Insecta" family="Staphylinidae" genus="Sunius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sunius melanocephalus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="melanocephalus">Sunius melanocephalus</taxonomicName>
is widely distributed (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="6661DE05766C76B4D9369872524340EC" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
<normalizedToken id="BA6CA512D55A5455CD3B2B2EF5FECE59" originalValue="Löbl">Loebl</normalizedToken>
and Smetana 2004
</bibRefCitation>
) and inhabits a wide variety of habitats including grasslands, swamps, riverbanks, gardens, parks, arable land and mammal burrows (
<bibRefCitation author="Assing, V" journalOrPublisher="Linzer biologische Beitrage" pageId="25" pageNumber="54" pagination="5 - 135" title="A revision of the Sunius species of the Western Palaearctic region and Middle Asia (Coleoptera: Staphylinidae: Paederinae)." volume="40" year="2008">Assing 2008</bibRefCitation>
<bibRefCitation id="0615368088FDC063D871D6A334909648" author="Assing, V" journalOrPublisher="Linzer biologische Beitrage" pageId="25" pageNumber="54" pagination="5 - 135" title="A revision of the Sunius species of the Western Palaearctic region and Middle Asia (Coleoptera: Staphylinidae: Paederinae)." volume="40" year="2008">Assing 2008</bibRefCitation>
). The Ontario specimens were captured in passive traps at the edges of woodlots.
</paragraph>
<caption pageId="12" pageNumber="41">
<paragraph pageId="12" pageNumber="41">
<caption id="3DF73A0DC78434B8CF6C3BBBDDEF991B" pageId="12" pageNumber="41">
<paragraph id="1E0B4FD45C15E7145B5DCC3DC5287ABA" pageId="12" pageNumber="41">
Maps 17-20. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 17
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sunius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sunius melanocephalus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="melanocephalus">Sunius melanocephalus</taxonomicName>
<taxonomicName id="24B820BCBBF781D676C4F2044E9A8D56" class="Insecta" family="Staphylinidae" genus="Sunius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sunius melanocephalus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="melanocephalus">Sunius melanocephalus</taxonomicName>
Fabricius (
<bibRefCitation author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
<bibRefCitation id="30B84081055AD5441A3760C5AC6E1AE3" author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies 1991</bibRefCitation>
,
<bibRefCitation author="Hoebeke, ER" journalOrPublisher="Entomological News" pageId="26" pageNumber="55" pagination="19 - 24" title="Sunius melanocephalus (Coleoptera: Staphylinidae), a Palearctic rove beetle new to North America." volume="102" year="1991">Hoebeke 1991</bibRefCitation>
<bibRefCitation id="11F990E477452DC3E6A1632A6B7A19E8" author="Hoebeke, ER" journalOrPublisher="Entomological News" pageId="26" pageNumber="55" pagination="19 - 24" title="Sunius melanocephalus (Coleoptera: Staphylinidae), a Palearctic rove beetle new to North America." volume="102" year="1991">Hoebeke 1991</bibRefCitation>
,
<bibRefCitation author="Byers, RA" journalOrPublisher="The Great Lakes Entomologist" pageId="26" pageNumber="55" pagination="81 - 106" title="Richness and abundance of Carabidae and Staphylinidae (Coleoptera) in northeastern dairy pastures under intensive grazing." volume="33" year="2000">Byers et al. 2000</bibRefCitation>
<bibRefCitation id="5E7B0BEDF8AFEE691DF7DD7741DADD0F" author="Byers, RA" journalOrPublisher="The Great Lakes Entomologist" pageId="26" pageNumber="55" pagination="81 - 106" title="Richness and abundance of Carabidae and Staphylinidae (Coleoptera) in northeastern dairy pastures under intensive grazing." volume="33" year="2000">Byers et al. 2000</bibRefCitation>
) 18
<taxonomicName class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acylophorus agilis" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="agilis">Acylophorus agilis</taxonomicName>
<taxonomicName id="701CE23BE3D62CDD6F5A92D2F43A1A96" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acylophorus agilis" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="agilis">Acylophorus agilis</taxonomicName>
Smetana (
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
<bibRefCitation id="B0591A798AC34B925BBC7A1166DAE890" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
,
<bibRefCitation author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="815 - 840" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 4." volume="110" year="1978">1978</bibRefCitation>
<bibRefCitation id="97011ED12A04CF4B37E32F252F3F8AF8" author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="815 - 840" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 4." volume="110" year="1978">1978</bibRefCitation>
) 19
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius cephalicus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="cephalicus">Bisnius cephalicus</taxonomicName>
<taxonomicName id="00E4E5833E9CED0110F8CE133B9E309B" class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius cephalicus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="cephalicus">Bisnius cephalicus</taxonomicName>
(Casey) (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="C36BA18A74258D10F876C48764F6FBD5" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
,
<bibRefCitation author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
<bibRefCitation id="A22A2857A024C0C37C9A66F600E2F41E" author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
Paquin and
<normalizedToken originalValue="Dupérré">Duperre</normalizedToken>
<normalizedToken id="E96A520135B97B8A61E0975009EFFD93" originalValue="Dupérré">Duperre</normalizedToken>
2001
</bibRefCitation>
) 20
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius siegwaldii" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="siegwaldii">Bisnius siegwaldii</taxonomicName>
<taxonomicName id="0B6A799D7F95E72A523F5654C81CBD78" class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius siegwaldii" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="siegwaldii">Bisnius siegwaldii</taxonomicName>
(Mannerheim) (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="409D16332BABFE4B75360364A6B7329D" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
,
<bibRefCitation author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
<bibRefCitation id="CFCB130A7AC0DDA74407DC954B093F12" author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
Paquin and
<normalizedToken originalValue="Dupérré">Duperre</normalizedToken>
<normalizedToken id="86ABE596378AD5B3E731D47C15483340" originalValue="Dupérré">Duperre</normalizedToken>
2001
</bibRefCitation>
).

120
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<taxonomicName authority="Smetana, 1965" authorityName="Smetana" authorityYear="1965" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus neonatus" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="neonatus">Philonthus neonatus Smetana, 1965</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="49" type="materials">
<paragraph pageId="20" pageNumber="49">Materials.</paragraph>
<paragraph pageId="20" pageNumber="49">UNITED STATES: VA: Giles Co., Ripplemead, rte 460 at bridge, flood debris, berlese, 11 to 25-V-2008, A. Brunke (4).</paragraph>
<subSubSection id="DDF4E255BDBF799911EC1E5BA8AF0FA3" pageId="20" pageNumber="49" type="materials">
<paragraph id="BC12AD6CC014E703959507E259B7FFB5" pageId="20" pageNumber="49">Materials.</paragraph>
<paragraph id="D24187ED23756D07C5C810D4C966D6B7" pageId="20" pageNumber="49">UNITED STATES: VA: Giles Co., Ripplemead, rte 460 at bridge, flood debris, berlese, 11 to 25-V-2008, A. Brunke (4).</paragraph>
</subSubSection>
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<paragraph pageId="20" pageNumber="49">Diagnosis.</paragraph>
<paragraph pageId="20" pageNumber="49">
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<paragraph id="D8F7633C80A83A04E428FD46F3D41E8A" pageId="20" pageNumber="49">Diagnosis.</paragraph>
<paragraph id="340E9E3474FDD94D852F6C395362E7DC" pageId="20" pageNumber="49">
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is separated from other northeastern
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by the combination of: pronotum with six punctures in both dorsal rows; pronotum no more than vaguely narrowed anteriorly; elytra distinctly red and without dark markings; elytra with micropunctures between the regular punctures; abdominal segments paler apically.
</paragraph>
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This species was previously known from Arkansas, District of Columbia, Indiana, Iowa, Kansas, Kentucky, Maine, Maryland, Massachusetts, Michigan, Mississippi,
<pageBreakToken pageId="21" pageNumber="50" start="start">Missouri</pageBreakToken>
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, New Hampshire, New Jersey, New York, Ontario, Pennsylvania, and
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="CC02F47BD06C6935520F0BBFAB23AA40" originalValue="Québec">Quebec</normalizedToken>
(
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="2CDD59C64061C6902BE300F6B79ECF51" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
). Herein we newly record it from Virginia (Map 33).
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is a hygrophilous species collected from debris along the margins of creeks, rivers and lakes (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="8F7B13C3EF4BA1D3CEFCF8911E1F098E" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
).
</paragraph>
<caption pageId="21" pageNumber="50">
<paragraph pageId="21" pageNumber="50">
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Maps 33-36. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 33
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<taxonomicName id="939EDEC9023CC9C03B49BBB293DF82FB" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus neonatus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="neonatus">Philonthus neonatus</taxonomicName>
Smetana (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="D9D0111777F8541034E80331AEE5EB83" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
) 34
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus vulgatus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="vulgatus">Philonthus vulgatus</taxonomicName>
<taxonomicName id="0ABCE7486B8C4BE62347B7C5FDF2D79E" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus vulgatus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="vulgatus">Philonthus vulgatus</taxonomicName>
Casey (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="277DDD7F7CF1F46558B047A046141DAE" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
) 35
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus</taxonomicName>
<taxonomicName id="65D7C0F981EB4642E7AFCEAB9A585AA4" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cinctus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cinctus">Quedius cinctus</taxonomicName>
(Paykull) (
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
<bibRefCitation id="7FC8296AF7A38F3132030FCD6862E5F5" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
,
<bibRefCitation author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">1990</bibRefCitation>
<bibRefCitation id="5313456E4F7601A752008FC612B4EC62" author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">1990</bibRefCitation>
,
<bibRefCitation author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
<bibRefCitation id="B5BFE7104BF7F1E8E3665E9DEDAC0E99" author="Chandler, DS" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" title="University of New Hampshire Insect and Arachnid Collections - Coleoptera" url="http://colsa1.unh.edu:591/unhinsects.htm" year="2001">Chandler 2001</bibRefCitation>
,
<bibRefCitation author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="341 - 345" title="Adventive species of Quedius (Coleoptera, Staphylinidae) in North America: a survey and new Canadian record." volume="22" year="2009">Majka et al. 2009</bibRefCitation>
<bibRefCitation id="A485FD62E324BD70F0AE5CA9B43020A0" author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="341 - 345" title="Adventive species of Quedius (Coleoptera, Staphylinidae) in North America: a survey and new Canadian record." volume="22" year="2009">Majka et al. 2009</bibRefCitation>
) 36
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cruentus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cruentus">Quedius cruentus</taxonomicName>
<taxonomicName id="3038E82F5AD399AE09C60822C918E71F" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cruentus" order="Coleoptera" pageId="21" pageNumber="50" phylum="Arthropoda" rank="species" species="cruentus">Quedius cruentus</taxonomicName>
(Olivier) (
<bibRefCitation author="Gusarov, VI" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="374 - 377" title="Quedius cruentus (Olivier) (Coleoptera, Staphylinidae), a Palaearctic Species New to North America." volume="55" year="2001">Gusarov 2001</bibRefCitation>
<bibRefCitation id="CF1D52B6944EC714C94A2052DA41EF74" author="Gusarov, VI" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="374 - 377" title="Quedius cruentus (Olivier) (Coleoptera, Staphylinidae), a Palaearctic Species New to North America." volume="55" year="2001">Gusarov 2001</bibRefCitation>
,
<bibRefCitation author="Hoebeke, ER" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="26" pageNumber="55" pagination="391 - 396" title="New records of Quedius cruentus (Olivier), a Palearctic rove beetle (Coleoptera: Staphylinidae), in northeastern North America." volume="110" year="2008">Hoebeke 2008</bibRefCitation>
<bibRefCitation id="F7398288BC26E5754D25AB55BA5DDFA3" author="Hoebeke, ER" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="26" pageNumber="55" pagination="391 - 396" title="New records of Quedius cruentus (Olivier), a Palearctic rove beetle (Coleoptera: Staphylinidae), in northeastern North America." volume="110" year="2008">Hoebeke 2008</bibRefCitation>
).
</paragraph>
</caption>

148
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<treatment ID-GBIF-Taxon="159361037" LSID="urn:lsid:plazi:treatment:A3BC36EC697A80A0CFF138F046E83EED" httpUri="http://treatment.plazi.org/id/A3BC36EC697A80A0CFF138F046E83EED" lastPageId="7" lastPageNumber="36" pageId="5" pageNumber="34">
<subSubSection pageId="5" pageNumber="34" type="nomenclature">
<paragraph pageId="5" pageNumber="34">
<taxonomicName authority="Stephens, 1832" authorityName="Stephens" authorityYear="1832" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami (Stephens, 1832)</taxonomicName>
<treatment id="A3BC36EC697A80A0CFF138F046E83EED" ID-GBIF-Taxon="159361037" LSID="urn:lsid:plazi:treatment:A3BC36EC697A80A0CFF138F046E83EED" httpUri="http://treatment.plazi.org/id/A3BC36EC697A80A0CFF138F046E83EED" lastPageId="7" lastPageNumber="36" pageId="5" pageNumber="34" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Tachyporinae">
<subSubSection id="3FB9B305CBC5A06CA66AAE3EC21BDF6F" pageId="5" pageNumber="34" type="nomenclature">
<paragraph id="98424BF3677A87BD06B8FB053DA03372" pageId="5" pageNumber="34">
<taxonomicName id="D9226F3E353BB8F465E0CA590F273BB2" ID-CoL="4WQ7F" authority="Stephens, 1832" authorityName="Stephens" authorityYear="1832" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami (Stephens, 1832)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="34" type="materials">
<paragraph pageId="5" pageNumber="34">Materials.</paragraph>
<paragraph pageId="5" pageNumber="34">UNITED STATES: NH: Coos Co., Jefferson, under bark, 20-IV-2010, T. Murray (1).</paragraph>
<paragraph pageId="5" pageNumber="34">CANADA: ON: Essex Co., Kingsville, 14-V-1973, R. Roughly (1); Waterloo Reg., Blair, RARE, Cruickston Creek, yellow pan traps, 15 to 20-VI-2006, S.A. Marshall and M. Bergeron (1); Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, hedgerow, pitfall, 13-X-2009 (1), 10-XI-2009 (1), A. Brunke;Blair, Fountain St. S. nr Speed River, 43.391, -80.373, hedgerow, pitfall, 13-X-2009 (1), 10-XI-2009 (1);Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, hedgerow near soybean field, pitfall trap, 2-XI-2010 (2); Wellington Co., Arkell, 27-IX-1986, L. Work (1); Belwood Lake, lake margin, fallen log overhang, 3-VI-2008, S.A. Marshall (1); Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, hedgerow, pitfall, 4-V-2010, A. Brunke (1); Guelph, 19-V-1981, G.M. Grant (1); Guelph, in rotten wood, 20-IV-2007, S.P.L. Luk (1); Guelph, Preservation Park, under bark, 21-IX-2010, S.P.L. Luk (1).</paragraph>
<subSubSection id="EE6194A51C33A37BBD1A59E896F806FD" pageId="5" pageNumber="34" type="materials">
<paragraph id="B6303DC9B001F602A2DBF0C77420E28C" pageId="5" pageNumber="34">Materials.</paragraph>
<paragraph id="AE246D6E8B1E231B263320C8E96983CA" pageId="5" pageNumber="34">UNITED STATES: NH: Coos Co., Jefferson, under bark, 20-IV-2010, T. Murray (1).</paragraph>
<paragraph id="41CEE70881835A65ECFA3266FCBD992B" pageId="5" pageNumber="34">CANADA: ON: Essex Co., Kingsville, 14-V-1973, R. Roughly (1); Waterloo Reg., Blair, RARE, Cruickston Creek, yellow pan traps, 15 to 20-VI-2006, S.A. Marshall and M. Bergeron (1); Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, hedgerow, pitfall, 13-X-2009 (1), 10-XI-2009 (1), A. Brunke;Blair, Fountain St. S. nr Speed River, 43.391, -80.373, hedgerow, pitfall, 13-X-2009 (1), 10-XI-2009 (1);Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, hedgerow near soybean field, pitfall trap, 2-XI-2010 (2); Wellington Co., Arkell, 27-IX-1986, L. Work (1); Belwood Lake, lake margin, fallen log overhang, 3-VI-2008, S.A. Marshall (1); Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, hedgerow, pitfall, 4-V-2010, A. Brunke (1); Guelph, 19-V-1981, G.M. Grant (1); Guelph, in rotten wood, 20-IV-2007, S.P.L. Luk (1); Guelph, Preservation Park, under bark, 21-IX-2010, S.P.L. Luk (1).</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="36" pageId="5" pageNumber="34" type="diagnosis">
<paragraph pageId="5" pageNumber="34">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="34">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
<subSubSection id="52E6B7512A0F7504B68B669421053553" lastPageId="7" lastPageNumber="36" pageId="5" pageNumber="34" type="diagnosis">
<paragraph id="2EA853090E26ACAE05015DE43D41AB2B" pageId="5" pageNumber="34">Diagnosis.</paragraph>
<paragraph id="CC1E29CF3D84740AB9D4E249EA55E1D6" pageId="5" pageNumber="34">
<taxonomicName id="7864DA0BBB426D6D6AFDE069FC8ABD6A" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
may be distinguished from all other members of the genus in eastern North America except
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus testaceus" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="testaceus">Sepedophilus testaceus</taxonomicName>
<taxonomicName id="55DE72E87910BCE2757E9D85ADBF7988" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus testaceus" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="testaceus">Sepedophilus testaceus</taxonomicName>
by the following combination of characters: body size large (&gt;2.3mm from clypeus to elytral apex); tergite seven with a white, apical, palisade fringe and at least one pair of bristles; elytra reddish but without distinct, reddish basal markings; middle-tibia with two apical spines (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="535D7169D5A63287126C76F9FD5812C4" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
). It differs from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus testaceus" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="testaceus">Sepedophilus testaceus</taxonomicName>
<taxonomicName id="803F658ED3620A44F360B8431AD21BE6" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus testaceus" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="testaceus">Sepedophilus testaceus</taxonomicName>
, another exotic species in North America, by the distinctly elongate seventh antennomere, which is subquadrate to weakly transverse in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus testaceus" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="testaceus">Sepedophilus testaceus</taxonomicName>
<taxonomicName id="E2826C8F7C223DCC2B6954B2F0189BF4" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus testaceus" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="testaceus">Sepedophilus testaceus</taxonomicName>
. Specimens of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus testaceus" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="testaceus">Sepedophilus testaceus</taxonomicName>
<taxonomicName id="33C6ED27274EA5FE696391942089D21A" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus testaceus" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="testaceus">Sepedophilus testaceus</taxonomicName>
with rather reddish elytra do exist but these individuals are uniformly pale, while in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
<taxonomicName id="2A4942F897126890C8D3EE0811ABFC65" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
the pronotum is distinctly darker than the elytra.
</paragraph>
<paragraph lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
<paragraph id="32D2460021D76F13BE2AD559D29C7E4D" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
This exotic, Palaearctic species was first collected in North America in Quebec in 1959 (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="C095DB235CE5E449E37DE8F8F7300414" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
) and has since been detected in Nova Scotia (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="9805FACAFD3DF4D2AABDEEFEBBC88002" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
)
<pageBreakToken pageId="6" pageNumber="35" start="start">and</pageBreakToken>
<pageBreakToken id="B9A1DA92E3A950273A52B24A06D41E65" pageId="6" pageNumber="35" start="start">and</pageBreakToken>
New Brunswick (
<bibRefCitation author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
<bibRefCitation id="D1AD2715890026F2F93561321D178D75" author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
). It was listed as questionably present in Ontario (
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="26" pageNumber="55" title="Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada" year="2010">Klimaszewski et al. 2010</bibRefCitation>
<bibRefCitation id="51E97AC13F97A3AE8F8037C51D9235FF" author="Klimaszewski, J" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="26" pageNumber="55" title="Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada" year="2010">Klimaszewski et al. 2010</bibRefCitation>
) but herein we confirm its widespread occurrence in the province as early as 1973 (Map 5). We also newly record
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
<taxonomicName id="9A307EC9FEF11AAA606482A49A2C41EF" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
for the United States (New Hampshire). This species is widespread in the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="D5156322AE9A6E50C2A5476DE2068DA5" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
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<normalizedToken id="17BC4D80C331D08A3A3A54361D281CB8" originalValue="Löbl">Loebl</normalizedToken>
and Smetana 2004
</bibRefCitation>
). In the Nearctic region,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
<taxonomicName id="6D719A9E253964CE7C7D1FBCD28CE124" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
is typically collected from leaf litter and under loose, fungusy bark in disturbed woodland fragments although it also inhabits debris along freshwater and marine shorelines (
<bibRefCitation author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="115 - 150" title="The coastal rove beetles (Coleoptera, Staphylinidae) of Atlantic Canada: a survey and new records." volume="2" year="2008">Majka et al. 2008</bibRefCitation>
<bibRefCitation id="41E3081350D8EC0747F713BB44330E12" author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="115 - 150" title="The coastal rove beetles (Coleoptera, Staphylinidae) of Atlantic Canada: a survey and new records." volume="2" year="2008">Majka et al. 2008</bibRefCitation>
) and open areas including raspberry fields and woodland edges (
<bibRefCitation author="Levesque, C" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="355 - 370" title="Abundance, Diversity and Dispersal Power of Rove Beetles (Coleoptera: Staphylinidae) in a Raspberry Plantation and Adjacent Sites in Eastern Canada." volume="68" year="1995">Levesque and Levesque 1995</bibRefCitation>
<bibRefCitation id="FB4309993D6088E464C2815067EBCF8B" author="Levesque, C" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="355 - 370" title="Abundance, Diversity and Dispersal Power of Rove Beetles (Coleoptera: Staphylinidae) in a Raspberry Plantation and Adjacent Sites in Eastern Canada." volume="68" year="1995">Levesque and Levesque 1995</bibRefCitation>
).
</paragraph>
<caption pageId="7" pageNumber="36">
<paragraph pageId="7" pageNumber="36">
<pageBreakToken pageId="7" pageNumber="36" start="start">Maps</pageBreakToken>
<caption id="2B100EDDEE416EEE7C6E5CD5AA4C88CF" pageId="7" pageNumber="36">
<paragraph id="16C9A786405CFD445E2B5868F16F3FB9" pageId="7" pageNumber="36">
<pageBreakToken id="554336AD49149DDF56EEBECE4A817013" pageId="7" pageNumber="36" start="start">Maps</pageBreakToken>
5-8. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 5
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
<taxonomicName id="A2E9B86EE79A442E2B140EEE5C0294BA" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus marshami" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="marshami">Sepedophilus marshami</taxonomicName>
(Stephens) (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="91F846897D6C0806F672A4DA31C68397" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
,
<bibRefCitation author="Levesque, C" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="355 - 370" title="Abundance, Diversity and Dispersal Power of Rove Beetles (Coleoptera: Staphylinidae) in a Raspberry Plantation and Adjacent Sites in Eastern Canada." volume="68" year="1995">Levesque and Levesque 1995</bibRefCitation>
<bibRefCitation id="FBAE4917030699E477E0FD6F3798F96C" author="Levesque, C" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="355 - 370" title="Abundance, Diversity and Dispersal Power of Rove Beetles (Coleoptera: Staphylinidae) in a Raspberry Plantation and Adjacent Sites in Eastern Canada." volume="68" year="1995">Levesque and Levesque 1995</bibRefCitation>
,
<bibRefCitation author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
<bibRefCitation id="57678052EDC88D838D33618B77A0F273" author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
) 6
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus occultus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="occultus">Sepedophilus occultus</taxonomicName>
<taxonomicName id="5CB754789C14B7C7E4AB05D17177ACD7" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus occultus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="occultus">Sepedophilus occultus</taxonomicName>
(Casey) (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="957018B73C971CFD28D8EC091EED71F9" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
,
<bibRefCitation author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
<bibRefCitation id="C4A02E3765CE2C6E2BEE130EFB87838C" author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
Paquin and
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2001
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,
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) 7
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(Say) (
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) 8
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus versicolor" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="versicolor">Sepedophilus versicolor</taxonomicName>
<taxonomicName id="31EFB8AC6290A2136A9B56C645A353C5" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus versicolor" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="versicolor">Sepedophilus versicolor</taxonomicName>
(Casey) (
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).
</paragraph>
</caption>

106
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<taxonomicName authority="Smetana, 1995" authorityName="Smetana" authorityYear="1995" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius vindex" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="vindex">Gabrius vindex Smetana, 1995</taxonomicName>
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<paragraph pageId="17" pageNumber="46">Materials.</paragraph>
<paragraph pageId="17" pageNumber="46">CANADA: ON: Bruce Co.,Bruce Pen. Natl. Pk., Upper Andrew Lake, beaver lodge, 26-VII-2008, S. A. Marshall (1); Simcoe Co., Noisy River Prov. Pk., Res., beaver lodge, 28-IX-2008, S.A. Marshall (1).</paragraph>
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<paragraph id="2FBA5979C48CC2ECC5120993A721FEC9" pageId="17" pageNumber="46">Materials.</paragraph>
<paragraph id="B5465CAE67714D574EAD82D592E32127" pageId="17" pageNumber="46">CANADA: ON: Bruce Co.,Bruce Pen. Natl. Pk., Upper Andrew Lake, beaver lodge, 26-VII-2008, S. A. Marshall (1); Simcoe Co., Noisy River Prov. Pk., Res., beaver lodge, 28-IX-2008, S.A. Marshall (1).</paragraph>
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<paragraph pageId="17" pageNumber="46">Diagnosis.</paragraph>
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<paragraph id="BDFDD3551DDCD19968050F45EF7BD8D6" pageId="17" pageNumber="46">Diagnosis.</paragraph>
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<taxonomicName id="71AC94903896C11BE24592E215EDFCB6" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius vindex" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="vindex">Gabrius vindex</taxonomicName>
is easily distinguished from all other species of the genus in northeastern North America except
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by the combin
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of: punctures of the elytra dense, separated by their widths or less; size large (at least 6.0mm from clypeus to abdominal apex); the area between the basal lines of tergites two and three punctate. From
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it is most easily separated by the shape of the pronotum which is narrowed anteriorly in
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and parallel in
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.
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This species is transcontinental in northern North America and was previously known from Alaska, Manitoba, Minnesota, and
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(
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). Herein we newly record
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from Ontario (Map 26), representing the only known localities in northeastern North America other than the Gatineau area of
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. All known specimens of
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have been collected in debris adjacent to the
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edge, and
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suggested that beaver lodges provide ideal conditions for this species. The Ontario specimens and the 92 DEBU specimens from beaver lodges in the Gatineau area of
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strongly support this.
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<taxonomicName authority="Fabricius, 1793" authorityName="Fabricius" authorityYear="1793" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus (Fabricius, 1793)</taxonomicName>
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</paragraph>
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<paragraph pageId="23" pageNumber="52">Materials.</paragraph>
<paragraph pageId="23" pageNumber="52">CANADA: ON: Huron Co., Seaforth, 7-VII-1955, D. Keys (1).</paragraph>
<subSubSection id="E9B355CD7BFD445CE6528FD12F00110D" pageId="23" pageNumber="52" type="materials">
<paragraph id="19D8864FBEF91D05C340370F6AC06770" pageId="23" pageNumber="52">Materials.</paragraph>
<paragraph id="47FFEC394134043C5250C45C96A57303" pageId="23" pageNumber="52">CANADA: ON: Huron Co., Seaforth, 7-VII-1955, D. Keys (1).</paragraph>
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<paragraph pageId="23" pageNumber="52">Diagnosis.</paragraph>
<paragraph pageId="23" pageNumber="52">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
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<paragraph id="A82282D589AC3BA9D59F46DEF107EA77" pageId="23" pageNumber="52">Diagnosis.</paragraph>
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may be distinguished from other northeastern
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<taxonomicName id="041FBD75BF535552A98CC2DC1FB91A3B" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
by the combination of: elytra evenly punctate; labrum distinctly bilobed; eyes distinctly shorter than temples; antennomeres one to three not distinctly paler than others; distal antennomeres only slightly transverse; pronotum with sublateral row of punctures longer than dorsal row.
</paragraph>
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This Palaearctic species was first correctly recognized in North America by
<bibRefCitation author="Horn, GH" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" pagination="149 - 167" title="Synopsis of the Quediini of the United States." volume="7" year="1878">Horn (1878)</bibRefCitation>
<bibRefCitation id="590A32E35D031CE7E116BD342C60B843" author="Horn, GH" journalOrPublisher="Transactions of the American Entomological Society" pageId="26" pageNumber="55" pagination="149 - 167" title="Synopsis of the Quediini of the United States." volume="7" year="1878">Horn (1878)</bibRefCitation>
who synonymised
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<taxonomicName id="FC523F44FD8F5286C4D5DF83BBEAF267" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius iracundus" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="iracundus">Quedius iracundus</taxonomicName>
(
<bibRefCitation author="Say, T" journalOrPublisher="Transactions of the American Philosophical Society" pageId="27" pageNumber="56" pagination="409 - 471" title="Descriptions of new North American insects and observations on some already described." volume="4" year="1834">Say 1834</bibRefCitation>
<bibRefCitation id="EED8273496F39EA7C695A48941583138" author="Say, T" journalOrPublisher="Transactions of the American Philosophical Society" pageId="27" pageNumber="56" pagination="409 - 471" title="Descriptions of new North American insects and observations on some already described." volume="4" year="1834">Say 1834</bibRefCitation>
), a species described from North America, with
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<taxonomicName id="9C2394016DFE08EF11323FF9139FFE5E" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
; this synonymy was later confirmed by
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana (1971)</bibRefCitation>
<bibRefCitation id="8FE0EAAAC6D139B72D0AA40FE963A8C6" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana (1971)</bibRefCitation>
. The first verifiable specimens from North
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<pageBreakToken id="5BBD626F8585DD6422BC11816A150DFF" pageId="24" pageNumber="53" start="start">America</pageBreakToken>
were collected in 1874 from Iowa (Smetana) but as
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<taxonomicName id="F84011698506CB0B0357CC0687C9BD75" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius iracundus" order="Coleoptera" pageId="24" pageNumber="53" phylum="Arthropoda" rank="species" species="iracundus">Quedius iracundus</taxonomicName>
(=Q. fulgidus)was described in 1834 from Indiana,
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<taxonomicName id="45959C42991F6B781AFA407436C09271" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="24" pageNumber="53" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
has surely been present long before 1874. Currently,
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<taxonomicName id="914275F470BF1D7C52F0296F57A02201" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="24" pageNumber="53" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
is known from Arizona, British Columbia, California, Colorado, District of Columbia, Georgia, Idaho, Illinois, Indiana, Kansas, Kentucky, Manitoba, Maryland, Massachusetts, Michigan, Minnesota, Mississippi, Missouri, New Jersey, New Mexico, New York, Oregon, Pennsylvania, Texas, Virginia, Washington, West Virginia, Wisconsin (
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
<bibRefCitation id="45E2DCA9CD6E949388367FADE968999F" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
), and Tennessee (
<bibRefCitation author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="815 - 840" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 4." volume="110" year="1978">Smetana 1978</bibRefCitation>
<bibRefCitation id="29C400A4C4F86EE3DB8142CCD814EBB9" author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="815 - 840" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 4." volume="110" year="1978">Smetana 1978</bibRefCitation>
). Herein we newly record this species from eastern Canada (Ontario) (Map 38). This species is strongly synanthropic in North America (
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
<bibRefCitation id="D188024802497F883FA967C2FCF4670D" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
) but occurs regularly in caves in the more southern portion of its range (
<bibRefCitation author="Peck, SB" journalOrPublisher="Journal of Cave and Karst Studies" pageId="27" pageNumber="56" pagination="3 - 8" title="The cave-inhabiting rove beetles of the United States (Coleoptera; Staphylinidae; excluding Aleocharinae and Pselaphinae): diversity and distributions." volume="65" year="2003">Peck and Thayer 2003</bibRefCitation>
<bibRefCitation id="2BA8B4273A838A38CB9879D825B715D2" author="Peck, SB" journalOrPublisher="Journal of Cave and Karst Studies" pageId="27" pageNumber="56" pagination="3 - 8" title="The cave-inhabiting rove beetles of the United States (Coleoptera; Staphylinidae; excluding Aleocharinae and Pselaphinae): diversity and distributions." volume="65" year="2003">Peck and Thayer 2003</bibRefCitation>
). In its native range
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="24" pageNumber="53" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
<taxonomicName id="49FF4E314B416D3694A3BB40373181BA" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="24" pageNumber="53" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
is widely distributed in the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="CE397EF3B715F6A41654D23DC0C7B9C7" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
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and Smetana 2004
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).
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<taxonomicName id="9E4087E17531E21B6CB64FE0959EB6F3" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="24" pageNumber="53" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
appears to be uncommon in the northeastern extreme of its range compared to its close relative
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cruentus" order="Coleoptera" pageId="24" pageNumber="53" phylum="Arthropoda" rank="species" species="cruentus">Quedius cruentus</taxonomicName>
<taxonomicName id="C59751C6494FBBA2E6079D5540757CAF" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius cruentus" order="Coleoptera" pageId="24" pageNumber="53" phylum="Arthropoda" rank="species" species="cruentus">Quedius cruentus</taxonomicName>
, another exotic species that was recently established on the continent (see above) and has since become extremely common.
</paragraph>
</subSubSection>

102
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<paragraph pageId="21" pageNumber="50">Materials.</paragraph>
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<paragraph id="77C0FD08D4B983840B6F69A534AF250F" pageId="21" pageNumber="50">Materials.</paragraph>
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CANADA: ON: Essex Co., Windsor, ~1.5km S Ojibway Prairie, forest-prairie edge, malaise trap, 15-V to 1-VI-2001, S. Paiero (1); Windsor, ~1.5 km S Ojibway Prairie, private prairie, malaise, 5 to 12-VI-2001, S. Paiero (3); Windsor, ~1.5km S Ojibway Prairie, private prairie, malaise, 19 to 30-VI-2001, P. Pratt (2); Hald. -Norfolk Reg., Charlotte 2 Rd., ~480m E of Charlotteville, West Quarterline Rd., 'C.C.S.N. -5', purple prism trap, 13 to 19-VI-2009, S.M. Paiero (1); Halton
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., Milton, Derry Rd. and 4th Line, under composter, 16-X-2008, S. M. Paiero (1); Oxford Co., Woodstock, trails nr. river, 14-VI-2008, S.A. Marshall (1). Simcoe Co., Midhurst, forest nr. Neretva St., 28-IX-2008, A. Brunke and K. Brunke (1); Wellington Co., Guelph, University Campus, dairy bush, dry
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, 22-IX-2008, A. Brunke (1).
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<paragraph pageId="22" pageNumber="51">Diagnosis.</paragraph>
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<paragraph id="2022C4FA478B060057B09592F614D26D" pageId="22" pageNumber="51">Diagnosis.</paragraph>
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may be distinguished from other northeastern
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by the combination of: elytra evenly punctate; labrum distinctly bilobed; eyes distinctly shorter than temples; antennomeres one to three distinctly paler than others; distal antennomeres strongly transverse; pronotum with sublateral row of punctures longer than dorsal row.
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This Palaearctic species was first detected by
<bibRefCitation author="Gusarov, VI" journalOrPublisher="The Coleopterists Bulletin" pageId="26" pageNumber="55" pagination="374 - 377" title="Quedius cruentus (Olivier) (Coleoptera, Staphylinidae), a Palaearctic Species New to North America." volume="55" year="2001">Gusarov (2001)</bibRefCitation>
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based on a specimen collected in New York.
<bibRefCitation author="Hoebeke, ER" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="26" pageNumber="55" pagination="391 - 396" title="New records of Quedius cruentus (Olivier), a Palearctic rove beetle (Coleoptera: Staphylinidae), in northeastern North America." volume="110" year="2008">Hoebeke (2008)</bibRefCitation>
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newly reported
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from Maine, Massachusetts, New Jersey, Ohio, Pennsylvania, and
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, and established its presence in North America as early as 1983 in New York. Herein we newly report this species from Ontario (Map 36). In its native range,
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is widely distributed in the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="DCA58EFB5B225CB915B82E48B660CBC6" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
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and Smetana 2004
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).
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has been found in a variety of habitats in Ontario including forests, prairies, urban greenspace under loose bark, under objects and in decayed fungi.
</paragraph>
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<treatment ID-GBIF-Taxon="159361013" LSID="urn:lsid:plazi:treatment:AF9A90238E2A4242F99EC3257C891487" httpUri="http://treatment.plazi.org/id/AF9A90238E2A4242F99EC3257C891487" lastPageNumber="37" pageId="8" pageNumber="37">
<subSubSection pageId="8" pageNumber="37" type="nomenclature">
<paragraph pageId="8" pageNumber="37">
<taxonomicName authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus (Gravenhorst, 1802)</taxonomicName>
<treatment id="AF9A90238E2A4242F99EC3257C891487" ID-GBIF-Taxon="159361013" LSID="urn:lsid:plazi:treatment:AF9A90238E2A4242F99EC3257C891487" httpUri="http://treatment.plazi.org/id/AF9A90238E2A4242F99EC3257C891487" lastPageNumber="37" pageId="8" pageNumber="37" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Tachyporinae">
<subSubSection id="78999AA62071F84CACB2050B0C87A8F4" pageId="8" pageNumber="37" type="nomenclature">
<paragraph id="3072333BC51B0F53DF883B8B4AD2D65F" pageId="8" pageNumber="37">
<taxonomicName id="27AACB493EF8F6BE52854E822DDFAF25" ID-CoL="54BJF" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus (Gravenhorst, 1802)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="37" type="materials">
<paragraph pageId="8" pageNumber="37">Materials.</paragraph>
<paragraph pageId="8" pageNumber="37">UNITED STATES: MA: Middlesex Co., Groton, 22-IV-2010, T. Murray (1).</paragraph>
<subSubSection id="EF91EE8420C597C5A9A3126D1A72E136" pageId="8" pageNumber="37" type="materials">
<paragraph id="6D46F354950DDAF86C27AC135D8C998E" pageId="8" pageNumber="37">Materials.</paragraph>
<paragraph id="28F123F043DB682BE21B35A05F29F800" pageId="8" pageNumber="37">UNITED STATES: MA: Middlesex Co., Groton, 22-IV-2010, T. Murray (1).</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="37" type="diagnosis">
<paragraph pageId="8" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="37">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus</taxonomicName>
<subSubSection id="0AD0FA69A512EDB3BB4219E0A9ACFB38" pageId="8" pageNumber="37" type="diagnosis">
<paragraph id="051AD40855B99D5343F2B8C6FF936566" pageId="8" pageNumber="37">Diagnosis.</paragraph>
<paragraph id="D504A4DB9A93715E1C71A9F2A00354E6" pageId="8" pageNumber="37">
<taxonomicName id="61CCFC65EBE4402BD8D9DEC50709A1CC" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus</taxonomicName>
is easily distinguished from congeners in northeastern North America by the combination of: pronotum and elytra lacking microsculpture; pronotum with at least borders paler than head; female tergite eight with all lobes of similar size; male sternite seven without apical lobes; small size (3.00-3.75 mm from clypeus to apex of elytra).
</paragraph>
<paragraph pageId="8" pageNumber="37">
<paragraph id="1EA4C8573F68DAE38E5FB1037E8BDD2B" pageId="8" pageNumber="37">
This exotic species was first collected in North America in St. Cyrville,
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="943401CCCBF45CDE62823BDD86A1B81B" originalValue="Québec">Quebec</normalizedToken>
in 1967 and was first recognized in North America by
<bibRefCitation author="Campbell, JM" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="87 - 94" title="New species and record of Tachinus (Coleoptera: Staphylinidae) from North America." volume="107" year="1975">Campbell (1975)</bibRefCitation>
<bibRefCitation id="14A682670588B4A193202B1514F50F49" author="Campbell, JM" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="87 - 94" title="New species and record of Tachinus (Coleoptera: Staphylinidae) from North America." volume="107" year="1975">Campbell (1975)</bibRefCitation>
. Since then it has been detected in Vermont, Pennsylvania (
<bibRefCitation author="Byers, RA" journalOrPublisher="The Great Lakes Entomologist" pageId="26" pageNumber="55" pagination="81 - 106" title="Richness and abundance of Carabidae and Staphylinidae (Coleoptera) in northeastern dairy pastures under intensive grazing." volume="33" year="2000">Byers et al. 2000</bibRefCitation>
<bibRefCitation id="79A7B6300EB4CEDEF55016D7CE919196" author="Byers, RA" journalOrPublisher="The Great Lakes Entomologist" pageId="26" pageNumber="55" pagination="81 - 106" title="Richness and abundance of Carabidae and Staphylinidae (Coleoptera) in northeastern dairy pastures under intensive grazing." volume="33" year="2000">Byers et al. 2000</bibRefCitation>
), Nova Scotia (
<bibRefCitation author="Schuelke, M" journalOrPublisher="Entomologische Blaetter" pageId="27" pageNumber="56" title="Tachinus corticinus Gravenhorst - in Nova Scotia (Canada) (Coleoptera, Staphylinidae)" volume="101" year="2006">
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
<bibRefCitation id="F9110C31CE02E8426EE79FF9269BA17E" author="Schuelke, M" journalOrPublisher="Entomologische Blaetter" pageId="27" pageNumber="56" title="Tachinus corticinus Gravenhorst - in Nova Scotia (Canada) (Coleoptera, Staphylinidae)" volume="101" year="2006">
<normalizedToken id="D6AA10BAB9C7A3A085C43AA0C935A90F" originalValue="Schülke">Schuelke</normalizedToken>
2006
</bibRefCitation>
), New Brunswick, Prince Edward Island (
<bibRefCitation author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
<bibRefCitation id="13A246D7ADF12AB3DAC696F93504890F" author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
) and Ontario (
<bibRefCitation author="Brunke, AJ" journalOrPublisher="Environmental Entomology" pageId="26" pageNumber="55" pagination="1106 - 1116" title="Generalist predators (Coleoptera: Carabidae, Staphylinidae) associated with millipede populations in sweet potato and carrot fields and implications for millipede management." volume="38" year="2009">Brunke et al. 2009</bibRefCitation>
<bibRefCitation id="3B2ADEA36C6CD5CF7041A1B20DE77A71" author="Brunke, AJ" journalOrPublisher="Environmental Entomology" pageId="26" pageNumber="55" pagination="1106 - 1116" title="Generalist predators (Coleoptera: Carabidae, Staphylinidae) associated with millipede populations in sweet potato and carrot fields and implications for millipede management." volume="38" year="2009">Brunke et al. 2009</bibRefCitation>
). Herein we record it as new for Massachusetts (Map 9).
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus</taxonomicName>
<taxonomicName id="8F342A59B2F9A2A84A30E6762FBC7B31" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus</taxonomicName>
is widespread in the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="5FAB40536022DDDBFE63FEC41532FC65" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
<normalizedToken id="D26A7313FA1DD5CBC08D83CC4BE2D559" originalValue="Löbl">Loebl</normalizedToken>
and Smetana 2004
</bibRefCitation>
) and has been collected in a variety of open and forested habitats. Although most individuals captured in Hannover, Germany were brachypterous (
<bibRefCitation author="Assing, V" journalOrPublisher="Bericht der Naturhistorischen Gesellschaft Hannover" pageId="25" pageNumber="54" pagination="173 - 187" title="Die Kurzflugelkafer (Coleoptera: Staphylinidae) eines urbanen Inselbiotopkomplexes in Hannover. 3. Beitrag zur Faunistik und Okologie der Staphyliniden von Hannover." volume="134" year="1992">Assing 1992</bibRefCitation>
<bibRefCitation id="DA6DFDD7370E75004A943E933FC8FCA0" author="Assing, V" journalOrPublisher="Bericht der Naturhistorischen Gesellschaft Hannover" pageId="25" pageNumber="54" pagination="173 - 187" title="Die Kurzflugelkafer (Coleoptera: Staphylinidae) eines urbanen Inselbiotopkomplexes in Hannover. 3. Beitrag zur Faunistik und Okologie der Staphyliniden von Hannover." volume="134" year="1992">Assing 1992</bibRefCitation>
),
<bibRefCitation author="Levesque, C" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="355 - 370" title="Abundance, Diversity and Dispersal Power of Rove Beetles (Coleoptera: Staphylinidae) in a Raspberry Plantation and Adjacent Sites in Eastern Canada." volume="68" year="1995">Levesque and Levesque (1995)</bibRefCitation>
<bibRefCitation id="88D5E18BEA5074621BCB9D285010D0B8" author="Levesque, C" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="355 - 370" title="Abundance, Diversity and Dispersal Power of Rove Beetles (Coleoptera: Staphylinidae) in a Raspberry Plantation and Adjacent Sites in Eastern Canada." volume="68" year="1995">Levesque and Levesque (1995)</bibRefCitation>
found that nearly all individuals captured in
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="E299EC5B1B8F5B006A5685F198ED21CA" originalValue="Québec">Quebec</normalizedToken>
raspberry fields were fully winged. All specimens deposited in DEBU were found to be brachypterous but fully winged individuals do exist in Ontario as
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus</taxonomicName>
<taxonomicName id="823BA5FDB6E9B2177534CC8A00B3F067" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus</taxonomicName>
was captured in small numbers in raised pan traps (A. Brunke unpublished data).
</paragraph>
<caption pageId="8" pageNumber="37">
<paragraph pageId="8" pageNumber="37">
<caption id="785EC93B047CA364FB6B4C87191863B8" pageId="8" pageNumber="37">
<paragraph id="02D26E81BFBE997A293C8645ACD1AE50" pageId="8" pageNumber="37">
Maps 9-12. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 9
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus</taxonomicName>
<taxonomicName id="CE5446FC640EE28072D53D773CAE5582" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus corticinus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="corticinus">Tachinus corticinus</taxonomicName>
(Gravenhorst) (
<bibRefCitation author="Campbell, JM" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="87 - 94" title="New species and record of Tachinus (Coleoptera: Staphylinidae) from North America." volume="107" year="1975">Campbell 1975</bibRefCitation>
<bibRefCitation id="95176E9CE94B8BFDDC6DE4158E8A5E40" author="Campbell, JM" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="87 - 94" title="New species and record of Tachinus (Coleoptera: Staphylinidae) from North America." volume="107" year="1975">Campbell 1975</bibRefCitation>
,
<bibRefCitation author="Levesque, C" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="355 - 370" title="Abundance, Diversity and Dispersal Power of Rove Beetles (Coleoptera: Staphylinidae) in a Raspberry Plantation and Adjacent Sites in Eastern Canada." volume="68" year="1995">Levesque and Levesque 1995</bibRefCitation>
<bibRefCitation id="050B6A030ADAC38BC9629DB350CE5FD8" author="Levesque, C" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="355 - 370" title="Abundance, Diversity and Dispersal Power of Rove Beetles (Coleoptera: Staphylinidae) in a Raspberry Plantation and Adjacent Sites in Eastern Canada." volume="68" year="1995">Levesque and Levesque 1995</bibRefCitation>
,
<bibRefCitation author="Byers, RA" journalOrPublisher="The Great Lakes Entomologist" pageId="26" pageNumber="55" pagination="81 - 106" title="Richness and abundance of Carabidae and Staphylinidae (Coleoptera) in northeastern dairy pastures under intensive grazing." volume="33" year="2000">Byers et al. 2000</bibRefCitation>
<bibRefCitation author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
<bibRefCitation id="F90BB2809CE012DF08C4FF6BDEA2088D" author="Byers, RA" journalOrPublisher="The Great Lakes Entomologist" pageId="26" pageNumber="55" pagination="81 - 106" title="Richness and abundance of Carabidae and Staphylinidae (Coleoptera) in northeastern dairy pastures under intensive grazing." volume="33" year="2000">Byers et al. 2000</bibRefCitation>
<bibRefCitation id="AA0A7D20E4ADF42E645CD956B9D630FC" author="Majka, CG" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="48 - 72" title="Introduced Staphylinidae (Coleoptera) in the Maritime provinces of Canada." volume="140" year="2008 a">Majka and Klimaszewski 2008a</bibRefCitation>
,
<bibRefCitation author="Brunke, AJ" journalOrPublisher="Environmental Entomology" pageId="26" pageNumber="55" pagination="1106 - 1116" title="Generalist predators (Coleoptera: Carabidae, Staphylinidae) associated with millipede populations in sweet potato and carrot fields and implications for millipede management." volume="38" year="2009">Brunke et al. 2009</bibRefCitation>
<bibRefCitation id="BD31A9D629667BDA3724969A8F80DB5D" author="Brunke, AJ" journalOrPublisher="Environmental Entomology" pageId="26" pageNumber="55" pagination="1106 - 1116" title="Generalist predators (Coleoptera: Carabidae, Staphylinidae) associated with millipede populations in sweet potato and carrot fields and implications for millipede management." volume="38" year="2009">Brunke et al. 2009</bibRefCitation>
) 10
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
<taxonomicName id="53EF07B4F1D4147ED02D0112DC4DBB8E" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
Campbell (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." volume="109" year="1979">Campbell 1979</bibRefCitation>
<bibRefCitation id="82FD5B38A4718A43A7037585858124FC" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." volume="109" year="1979">Campbell 1979</bibRefCitation>
,
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimaszewski et al. 2005</bibRefCitation>
<bibRefCitation id="D55CD3AF7EA6E295A680D570FBB37B5D" author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimaszewski et al. 2005</bibRefCitation>
) 11
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus ornatus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="ornatus">Tachyporus ornatus</taxonomicName>
<taxonomicName id="C03BDCA81840BD201312C365BC388339" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus ornatus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="ornatus">Tachyporus ornatus</taxonomicName>
Campbell (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." volume="109" year="1979">Campbell 1979</bibRefCitation>
<bibRefCitation id="2CE07884E8113B2DD175F70246452F4D" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." volume="109" year="1979">Campbell 1979</bibRefCitation>
) 12
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bledius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bledius neglectus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="neglectus">Bledius neglectus</taxonomicName>
<taxonomicName id="7D78910D7ECFAA4712E6B578D8BFC7BE" class="Insecta" family="Staphylinidae" genus="Bledius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bledius neglectus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="neglectus">Bledius neglectus</taxonomicName>
Casey (
<bibRefCitation author="Herman, L" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="26" pageNumber="55" pagination="71 - 172" title="Revision of Bledius and related genera. Part II. The armatus, basalis, and mandibularis (Coleoptera, Staphylinidae, Oxytelinae)." volume="157" year="1976">Herman 1976</bibRefCitation>
<bibRefCitation id="B1E4EB2C6D206F3CABC3BBA58DC98825" author="Herman, L" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="26" pageNumber="55" pagination="71 - 172" title="Revision of Bledius and related genera. Part II. The armatus, basalis, and mandibularis (Coleoptera, Staphylinidae, Oxytelinae)." volume="157" year="1976">Herman 1976</bibRefCitation>
,
<bibRefCitation author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="115 - 150" title="The coastal rove beetles (Coleoptera, Staphylinidae) of Atlantic Canada: a survey and new records." volume="2" year="2008">Majka et al. 2008</bibRefCitation>
<bibRefCitation id="9171AAAC09DDEE7299A526C5A8FD1169" author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="115 - 150" title="The coastal rove beetles (Coleoptera, Staphylinidae) of Atlantic Canada: a survey and new records." volume="2" year="2008">Majka et al. 2008</bibRefCitation>
).
</paragraph>
</caption>

102
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<paragraph pageId="8" pageNumber="37">Materials.</paragraph>
<paragraph pageId="8" pageNumber="37">CANADA: ON: Kent Co., Rondeau Prov. Pk., spicebush trail, Carolinian forest, malaise, 3 to 16-VII-2003 (1), 15-VIII to 7-IX-2003 (1), Marshall et al., 16 to 29-VII-2003, S.A. Marshall (1).</paragraph>
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<paragraph id="25EEC4C55959513FFFFCE40ECA601C3E" pageId="8" pageNumber="37">Materials.</paragraph>
<paragraph id="DE386FD5E03EBE42D7C37202AA1469EB" pageId="8" pageNumber="37">CANADA: ON: Kent Co., Rondeau Prov. Pk., spicebush trail, Carolinian forest, malaise, 3 to 16-VII-2003 (1), 15-VIII to 7-IX-2003 (1), Marshall et al., 16 to 29-VII-2003, S.A. Marshall (1).</paragraph>
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<paragraph pageId="8" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="37">
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<paragraph id="315F9111733BFC7B215D5F63272A1B71" pageId="8" pageNumber="37">Diagnosis.</paragraph>
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can be easily separated from others of the genus except
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and
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by the apical ctenidium of the mesotibia, which extends upwards along the lateral edge (
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). It is best distinguished from
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and
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by the combination of: abdominal sternites four to six with three lateral bristles; abdominal sternites five and six with only one bristle at each side of the midline; smaller size (2.0-2.5mm from clypeus to elytral apex).
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This species is broadly distributed in eastern North America and was previously known from District of Columbia, Florida, Georgia, Illinois, Iowa, Kansas, Kentucky, Maryland, Massachusetts, Michigan, Minnesota, Missouri, New Hampshire, New Jersey, New York, North Carolina, Ohio, Pennsylvania, South Carolina, and Virginia (
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). Herein we newly record it from Canada (Ontario) (Map 8). This species apparently reaches its northern limit in
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Carolinian forests. Little is known about its bionomics although it has been found on mushrooms (
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) like the related
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and in a 'rotten stump with a small
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(
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
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).
</paragraph>
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98
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</paragraph>
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<paragraph pageId="11" pageNumber="40">Materials.</paragraph>
<paragraph pageId="11" pageNumber="40">CANADA: ON: Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.742, -81.514, soybean field, pitfall, 23-VI-2010 (6), 7-VII-2010 (6), 21-VII-2010 (4), 1-IX-2010 (1), A. Brunke; Benmiller, Sharpes Creek Line, 43.691, -81.608, hedgerow nr. creek, canopy trap in buckthorn, 22-VI-2009 (1) A. Brunke; Brucefield, London Rd. nr. Centennial Rd., 43.509, -81.516, soybean field, canopy trap, 22-VI-2009 (1) A. Brunke; Waterloo Reg., Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, soybean field, pitfall, 23-VI-2009 (1), 7-VII-2009 (2), 21-VII-2009 (1), A. Brunke; Blair, Fountain St. S. nr Speed River, 43.391, -80.373, soybean field, pitfall, 23-VI-2009 (3),7-VII-2009 (1) A. Brunke; Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, soybean field, pitfall, 13-VII-2010 (4), A. Brunke; Blair, Whistlebare Rd. and Township Rd. 1, 43.367, -80.358, soybean field, pitfall trap, 15-VI-2010 (3), 27-VII-2010 (1), A. Brunke. Wellington Co., Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, soybean field, pitfall, 13-VII-2010 (2); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, pitfall trap, 23-VI-2009 (23), 7-VII-2009 (4), 21-VII-2009 (11), 4-VIII-2009 (1), 18-VIII-2009 (2), 1-IX-2009 (1), 15-IX-2009 (4); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, vacuum sampled from soybean foliage at 8pm, 16-VII-2009.</paragraph>
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<paragraph id="BB2AFBB6E4DBD5A685C4C142EB64F2A7" pageId="11" pageNumber="40">Materials.</paragraph>
<paragraph id="30E32E59DF1AFBA9D5E2062F1A2A0F5F" pageId="11" pageNumber="40">CANADA: ON: Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.742, -81.514, soybean field, pitfall, 23-VI-2010 (6), 7-VII-2010 (6), 21-VII-2010 (4), 1-IX-2010 (1), A. Brunke; Benmiller, Sharpes Creek Line, 43.691, -81.608, hedgerow nr. creek, canopy trap in buckthorn, 22-VI-2009 (1) A. Brunke; Brucefield, London Rd. nr. Centennial Rd., 43.509, -81.516, soybean field, canopy trap, 22-VI-2009 (1) A. Brunke; Waterloo Reg., Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, soybean field, pitfall, 23-VI-2009 (1), 7-VII-2009 (2), 21-VII-2009 (1), A. Brunke; Blair, Fountain St. S. nr Speed River, 43.391, -80.373, soybean field, pitfall, 23-VI-2009 (3),7-VII-2009 (1) A. Brunke; Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, soybean field, pitfall, 13-VII-2010 (4), A. Brunke; Blair, Whistlebare Rd. and Township Rd. 1, 43.367, -80.358, soybean field, pitfall trap, 15-VI-2010 (3), 27-VII-2010 (1), A. Brunke. Wellington Co., Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, soybean field, pitfall, 13-VII-2010 (2); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, pitfall trap, 23-VI-2009 (23), 7-VII-2009 (4), 21-VII-2009 (11), 4-VIII-2009 (1), 18-VIII-2009 (2), 1-IX-2009 (1), 15-IX-2009 (4); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, vacuum sampled from soybean foliage at 8pm, 16-VII-2009.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="40" type="diagnosis">
<paragraph pageId="11" pageNumber="40">Diagnosis.</paragraph>
<paragraph pageId="11" pageNumber="40">
<subSubSection id="359ADFC1FD8B1103E3A06BE8E0F7A0B4" pageId="11" pageNumber="40" type="diagnosis">
<paragraph id="D184DB07D6A3B46D716F40C246D2DDCA" pageId="11" pageNumber="40">Diagnosis.</paragraph>
<paragraph id="6AD069831E7CF1F1C70592C2AB995A26" pageId="11" pageNumber="40">
The diverse genus
<taxonomicName class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="genus">Scopaeus</taxonomicName>
<taxonomicName id="8F21DC4E48DECD000F3C1609A9F49148" class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="genus">Scopaeus</taxonomicName>
is greatly in need of revision in North America and thus,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus minutus" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="minutus">Scopaeus minutus</taxonomicName>
<taxonomicName id="5703C2DEF06EB4D71A59C6DAB10DD323" class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus minutus" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="minutus">Scopaeus minutus</taxonomicName>
can only be recognized currently by the form of the aedeagus (Fig. 17).
</paragraph>
<paragraph pageId="11" pageNumber="40">
<paragraph id="C84F3BB3B81C203989CA542EAAEF6E5A" pageId="11" pageNumber="40">
This exotic Palaearctic species was first reported from North America by
<bibRefCitation author="Frisch, J" journalOrPublisher="Organisms Diversity and Evolution" pageId="26" pageNumber="55" pagination="27 - 53" title="Rove beetles of the subtribe Scopaeina Mulsant and Rey (Coleoptera: Staphylinidae) in the West Palaearctic: phylogeny, biogeography and species catalogue." volume="2" year="2002">Frisch et al. (2002)</bibRefCitation>
<bibRefCitation id="DC850C1A9BC3CD7CAA5F9FE544A2355E" author="Frisch, J" journalOrPublisher="Organisms Diversity and Evolution" pageId="26" pageNumber="55" pagination="27 - 53" title="Rove beetles of the subtribe Scopaeina Mulsant and Rey (Coleoptera: Staphylinidae) in the West Palaearctic: phylogeny, biogeography and species catalogue." volume="2" year="2002">Frisch et al. (2002)</bibRefCitation>
from Montreal,
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="DAEBD34D639D790D09CE6E8C599C6578" originalValue="Québec">Quebec</normalizedToken>
, Canada; however, no specimen data were provided. Herein we confirm its presence in North America and newly report it from Ontario based on numerous voucher specimens collected from 2009-2010 (Map 16). It is widely distributed in the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="10186BFEE4F763F481ECE180ACF6BB5A" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
<normalizedToken id="6FD68111FFF4D31121AF233701558C8A" originalValue="Löbl">Loebl</normalizedToken>
and Smetana 2004
</bibRefCitation>
). The North American material was collected in passive traps in soybean fields and woodlot edges.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus minutus" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="minutus">Scopaeus minutus</taxonomicName>
<taxonomicName id="9FB0B2DB390E739ED566387DDF01E5F2" class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus minutus" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="minutus">Scopaeus minutus</taxonomicName>
is less hygrophilus than others of the genus (
<bibRefCitation author="Frisch, J" journalOrPublisher="Organisms Diversity and Evolution" pageId="26" pageNumber="55" pagination="27 - 53" title="Rove beetles of the subtribe Scopaeina Mulsant and Rey (Coleoptera: Staphylinidae) in the West Palaearctic: phylogeny, biogeography and species catalogue." volume="2" year="2002">Frisch et al. 2002</bibRefCitation>
<bibRefCitation id="D86052B2755910352C72BB33CD57B0D9" author="Frisch, J" journalOrPublisher="Organisms Diversity and Evolution" pageId="26" pageNumber="55" pagination="27 - 53" title="Rove beetles of the subtribe Scopaeina Mulsant and Rey (Coleoptera: Staphylinidae) in the West Palaearctic: phylogeny, biogeography and species catalogue." volume="2" year="2002">Frisch et al. 2002</bibRefCitation>
) and is typically found in habitats undergoing early stages of succession (
<bibRefCitation author="Bohac, J" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="25" pageNumber="54" pagination="360 - 385" title="Review of the subfamily Paederinae (Coleoptera, Staphylinidae) in Czechoslavakia." volume="82" year="1985">
<normalizedToken originalValue="Boháč">Bohac</normalizedToken>
<bibRefCitation id="1FCCCE214E95848B176B4F5B02F14039" author="Bohac, J" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="25" pageNumber="54" pagination="360 - 385" title="Review of the subfamily Paederinae (Coleoptera, Staphylinidae) in Czechoslavakia." volume="82" year="1985">
<normalizedToken id="67F4D9B1FD805EBF3E6CA226DDAC7900" originalValue="Boháč">Bohac</normalizedToken>
1985
</bibRefCitation>
).

88
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<paragraph pageId="15" pageNumber="44">Materials.</paragraph>
<paragraph pageId="15" pageNumber="44">CANADA: ON: Simcoe Co. Midhurst, forest nr. Neretva St., under bark of large beech trunk, 4-IX-2009, A. Brunke and K. Brunke (1).</paragraph>
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<paragraph id="413B28C6823557800ACA00A6C0AB7EB1" pageId="15" pageNumber="44">Materials.</paragraph>
<paragraph id="4DF05A51DB2BFAE9C3B2108247FCA784" pageId="15" pageNumber="44">CANADA: ON: Simcoe Co. Midhurst, forest nr. Neretva St., under bark of large beech trunk, 4-IX-2009, A. Brunke and K. Brunke (1).</paragraph>
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<paragraph pageId="15" pageNumber="44">Diagnosis.</paragraph>
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<paragraph id="134BAB20D74C434F467047AACEFCD199" pageId="15" pageNumber="44">Diagnosis.</paragraph>
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may be recognized by the combination of: large size (at least 5.0mm long from clypeus to abdominal apex); eyes large, with temple that is distinctly less than twice as long as the eye; forebody without a greenish metallic lustre; elytra with sparsely distributed punctures that are separated by two to three times their diameter; area between basal lines on tergites two and three punctate
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This apparently rare species was known from only five specimens at the time of its description (
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) from localities in New York and Ohio. It was collected
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<pageBreakToken id="EA444912A23049575EEABEB6E89D47C3" pageId="16" pageNumber="45" start="start">in</pageBreakToken>
Missouri by
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
<bibRefCitation id="7BA04AF642A6DF97184E18E93AFEC373" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
in a flight intercept trap. Herein we newly report it from Canada (Ontario) (Map 24). All known specimens with microhabitat data were collected in deciduous forests in litter, or in the proximity of decaying wood.
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is almost certainly an uncommon specialist of deciduous or mixed forests.
</paragraph>
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90
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<taxonomicName authority="Say, 1834" authorityName="Say" authorityYear="1834" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus opicus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="opicus">Sepedophilus opicus (Say, 1834)</taxonomicName>
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</paragraph>
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<subSubSection pageId="7" pageNumber="36" type="materials">
<paragraph pageId="7" pageNumber="36">Materials.</paragraph>
<paragraph pageId="7" pageNumber="36">CANADA: ON: Hald.-Norfolk Reg., Backus Tract Woods, sifting leaf litter under mushrooms, 7-VI-2009, A. Brunke and L. DesMarteaux (5); Backus Tract Woods, sifted litter in sugar maple-dominated, mesic forest, 2-IV-2010, A. Brunke (1); Kent Co., Rondeau Prov. Pk., spicebush trail, carolinian forest, malaise, 16 to 29-VII-2003, S. Marshall et al. (1); Rondeau Prov. Pk., south point trail, slough forest, sifting leaf litter, 27-IX-2009, A. Brunke and D.K.B. Cheung (1). Lambton Co., Pinery Prov. Pk., Carolinian Trail, hardwood forest, litter around white pines, 17-IV-2010, A. Brunke (3).</paragraph>
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<paragraph id="8DED5692A18E1A87E07A9F00B8E7BFF7" pageId="7" pageNumber="36">Materials.</paragraph>
<paragraph id="49CE4762BC01C1B13832C669A1C824B1" pageId="7" pageNumber="36">CANADA: ON: Hald.-Norfolk Reg., Backus Tract Woods, sifting leaf litter under mushrooms, 7-VI-2009, A. Brunke and L. DesMarteaux (5); Backus Tract Woods, sifted litter in sugar maple-dominated, mesic forest, 2-IV-2010, A. Brunke (1); Kent Co., Rondeau Prov. Pk., spicebush trail, carolinian forest, malaise, 16 to 29-VII-2003, S. Marshall et al. (1); Rondeau Prov. Pk., south point trail, slough forest, sifting leaf litter, 27-IX-2009, A. Brunke and D.K.B. Cheung (1). Lambton Co., Pinery Prov. Pk., Carolinian Trail, hardwood forest, litter around white pines, 17-IV-2010, A. Brunke (3).</paragraph>
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<paragraph pageId="7" pageNumber="36">Diagnosis.</paragraph>
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<paragraph id="2ECB7E6A2112A84B7793C75E904C7A0A" pageId="7" pageNumber="36">Diagnosis.</paragraph>
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<taxonomicName id="A7C609499A4A22A1DD5C2AD478CA118E" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus opicus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="opicus">Sepedophilus opicus</taxonomicName>
can be distinguished from other members of genus in northeastern North America by a combination of: base of elytra with reddish mark
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extending laterally to the margin; basal abdominal segments with long bristles; elytral epipleuron uniformly setose; pronotal microsculpture distinct (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
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).
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This widely distributed species is known from Alabama, Florida, Illinois, Indiana, Iowa, Maryland, Massachusetts, Michigan, Missouri, New Jersey, New York, North Carolina, Pennsylvania,
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, Texas, and Virginia (
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). Herein we newly record it from Ontario (Map 7).
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appears to be confined to deciduous forests in litter, under bark and on mushrooms, and reaches its northern distributional limit in southern Canada. A large number of individuals were found on fresh mushrooms (~35, 5 taken as vouchers) in Backus Tract Woods, Ontario and this may be a preferred microhabitat.
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<taxonomicName authority="Casey, 1884" authorityName="Casey" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus occultus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="occultus">Sepedophilus occultus (Casey, 1884)</taxonomicName>
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</paragraph>
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<paragraph pageId="7" pageNumber="36">Materials.</paragraph>
<paragraph pageId="7" pageNumber="36">CANADA: ON:Huron Co., Benmiller, Sharpes Creek Line, 43.691, -81.608, hedgerow nr. creek, canopy trap in buckthorn, 22-VI-2009, A. Brunke (1); Elgin Co., Orwell, 15-VI-1978, D. Morris (1); Essex Co., Point Pelee Natl. Pk., wood area by W beach, malaise/pan traps, 10 to 23-IX-1999, O. Lonsdale (1); Hald.-Norfolk Reg.,Turkey Point Provincial Park, malaise trap, 3 to 28-VIII-2009, S. Paiero (1); Kent Co., Rondeau Prov. Park, spicebush trail, 42°18'9N, 81°51'6W, Carolinian forest, WPT, 16to17-VI-2003, Paiero and Carscadden (1); Wellington Co., Guelph, University arboretum nature reserve, ex. Beech litter, 3-V-2009, Brunke and Cheung (1); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, hedgerow, pitfall, 2-VI-2009, A. Brunke (1).</paragraph>
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<paragraph id="70471A11355F561B48B1924E04496168" pageId="7" pageNumber="36">Materials.</paragraph>
<paragraph id="56A28EEEAFE5F5D997EDA6C5DBA2BF44" pageId="7" pageNumber="36">CANADA: ON:Huron Co., Benmiller, Sharpes Creek Line, 43.691, -81.608, hedgerow nr. creek, canopy trap in buckthorn, 22-VI-2009, A. Brunke (1); Elgin Co., Orwell, 15-VI-1978, D. Morris (1); Essex Co., Point Pelee Natl. Pk., wood area by W beach, malaise/pan traps, 10 to 23-IX-1999, O. Lonsdale (1); Hald.-Norfolk Reg.,Turkey Point Provincial Park, malaise trap, 3 to 28-VIII-2009, S. Paiero (1); Kent Co., Rondeau Prov. Park, spicebush trail, 42°18'9N, 81°51'6W, Carolinian forest, WPT, 16to17-VI-2003, Paiero and Carscadden (1); Wellington Co., Guelph, University arboretum nature reserve, ex. Beech litter, 3-V-2009, Brunke and Cheung (1); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, hedgerow, pitfall, 2-VI-2009, A. Brunke (1).</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="36" type="diagnosis">
<paragraph pageId="7" pageNumber="36">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="36">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus occultus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="occultus">Sepedophilus occultus</taxonomicName>
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<paragraph id="410FDDBBEFFEA19CC529DB6F032E98F6" pageId="7" pageNumber="36">Diagnosis.</paragraph>
<paragraph id="DB3FB114574AF55752814DB16A53E3AA" pageId="7" pageNumber="36">
<taxonomicName id="7DF9D6BAE4C820F2E6A156DD95A758F9" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus occultus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="occultus">Sepedophilus occultus</taxonomicName>
can be distinguished from other northeastern
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">Sepedophilus</taxonomicName>
<taxonomicName id="FFA14555CFE976112333EB4E2C93851B" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">Sepedophilus</taxonomicName>
with a reddish area at the base of each elytron by the combination of: basal abdominal segments with short bristles only; elytral epipleuron with few or no setae on its basal half; pronotum uniformly colored; elytra without coarse bristles laterally; apical ctenidium of mesotibia restricted to the apex (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="C1C922074B8B5A28785DC1AE6CD49343" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
). The individual from Kent County has a uniformly reddish body and may be slightly teneral. Similar specimens can be recognized by the unique combination of the elytral epipleuron with few or no setae on its basal half and the impunctate elytral apex.
</paragraph>
<paragraph pageId="7" pageNumber="36">
<paragraph id="B6285E34A2F1F04BB50F9C95EE6E37F6" pageId="7" pageNumber="36">
This species is widely distributed in eastern North America and was previously known from Connecticut, District of Columbia, Georgia, Illinois, Iowa, Kentucky, Maryland, Massachusetts, Michigan, Mississippi, New Jersey, New York, Ohio, Pennsylvania (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
<bibRefCitation id="CB868048C9F40B4A89C98DCE8D531A36" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 89" title="A revision of the genus Sepedophilus Gistel (Coleoptera: Staphylinidae) of America north of Mexico." volume="99" year="1976">Campbell 1976</bibRefCitation>
), Missouri (
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
<bibRefCitation id="A79584A99565252082D0C0214DC3187A" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
) Rhode Island (
<bibRefCitation author="Sikes, DS" journalOrPublisher="Zootaxa" pageId="27" pageNumber="56" pagination="1 - 38" title="The beetle fauna of the state of Rhode Island, USA (Coleoptera): 656 new state records." volume="340" year="2003">Sikes 2003</bibRefCitation>
<bibRefCitation id="D2C503A610A258C13EDC94B650590681" author="Sikes, DS" journalOrPublisher="Zootaxa" pageId="27" pageNumber="56" pagination="1 - 38" title="The beetle fauna of the state of Rhode Island, USA (Coleoptera): 656 new state records." volume="340" year="2003">Sikes 2003</bibRefCitation>
), and Quebec (
<bibRefCitation author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
<bibRefCitation id="BB07098CA5619B2FE73421112AD2C0FA" author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
Paquin and
<normalizedToken originalValue="Dupérré">Duperre</normalizedToken>
<normalizedToken id="50BBF812BFCB15F87A490B96F310FFF7" originalValue="Dupérré">Duperre</normalizedToken>
2001
</bibRefCitation>
). Herein we report it as new for Ontario (Map 6).
<taxonomicName class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus occultus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="occultus">Sepedophilus occultus</taxonomicName>
<taxonomicName id="328A4DC50AAFB0ED2E445A9D3A83B3F8" class="Insecta" family="Staphylinidae" genus="Sepedophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sepedophilus occultus" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="occultus">Sepedophilus occultus</taxonomicName>
is a forest-dwelling species that has been collected mainly from leaf litter and under bark. Its presence in the Boreal Forest Region of
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="4C0D03594DDEAD6D3E6D473DB2EAA74E" originalValue="Québec">Quebec</normalizedToken>
(
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<bibRefCitation id="4F64B60F30209CBE59A27CC8C5863140" author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
Paquin and
<normalizedToken originalValue="Dupérré">Duperre</normalizedToken>
<normalizedToken id="35074950CAB0EC866E19823B0916616D" originalValue="Dupérré">Duperre</normalizedToken>
2001
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) is surprising considering its more southern distribution elsewhere but this probably reflects the inadequate knowledge of
<normalizedToken originalValue="Canada’s">Canada's</normalizedToken>
<normalizedToken id="0566534A6359F6ED241D5E9DBC6B6786" originalValue="Canada’s">Canada's</normalizedToken>
boreal insect fauna rather than a disjunct population.
</paragraph>
</subSubSection>

98
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<paragraph pageId="12" pageNumber="41">Materials.</paragraph>
<paragraph pageId="12" pageNumber="41">CANADA: ON: Middlesex Co. London, Southern Crop Protection Research Centre, pitfall trap/Masner trap, 21-VI-1995, T. Sawinski (1); Simcoe Co., Noisy River, Prov. Nature Res., beaver lodge, 28-IX-2008, S.A. Marshall (1).</paragraph>
<subSubSection id="6609148CCB6FEFCF1503F8588A2E9A82" pageId="12" pageNumber="41" type="materials">
<paragraph id="CB9731405D602F5561DACA03F3D50FC2" pageId="12" pageNumber="41">Materials.</paragraph>
<paragraph id="AFB6F0E05E4FC23227B008CB759159C3" pageId="12" pageNumber="41">CANADA: ON: Middlesex Co. London, Southern Crop Protection Research Centre, pitfall trap/Masner trap, 21-VI-1995, T. Sawinski (1); Simcoe Co., Noisy River, Prov. Nature Res., beaver lodge, 28-IX-2008, S.A. Marshall (1).</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="41" type="diagnosis">
<paragraph pageId="12" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="41">
<subSubSection id="B05710602DE4E7EA19C0D31D6220FA13" pageId="12" pageNumber="41" type="diagnosis">
<paragraph id="0FCCAE93177F8E82150B728E5AC03D2A" pageId="12" pageNumber="41">Diagnosis.</paragraph>
<paragraph id="E57508679E708A449B621B00F15579BA" pageId="12" pageNumber="41">
At present,
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<taxonomicName id="1460EE277B5683D85EE1F14F6E50FCC7" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acylophorus agilis" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="agilis">Acylophorus agilis</taxonomicName>
is reliably separated from others in the diverse
<taxonomicName class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acylophorus pronus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="pronus">Acylophorus pronus</taxonomicName>
<taxonomicName id="6CDE5EE0FFC1D9AFE2DE669D0F3C7FA6" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acylophorus pronus" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="pronus">Acylophorus pronus</taxonomicName>
-group only by the characteristically shaped paramere of the aedeagus (Fig. 169 in
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
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).
</paragraph>
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<paragraph id="DC581DE24B09B89113918BD8291B382D" pageId="12" pageNumber="41">
This species is widely distributed in eastern North America, and was previously known from Indiana, Maryland, Massachusetts, Michigan, New York, North Carolina, Pennsylvania, Rhode Island, Missouri, (
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
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), Illinois, and Kentucky (
<bibRefCitation author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="815 - 840" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 4." volume="110" year="1978">Smetana 1978</bibRefCitation>
<bibRefCitation id="2DF2B38E4EF805802BC2C643AFFFB164" author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="815 - 840" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 4." volume="110" year="1978">Smetana 1978</bibRefCitation>
). Herein we newly record it from Canada (Ontario) (Map 18).
<taxonomicName class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acylophorus agilis" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="agilis">Acylophorus agilis</taxonomicName>
<taxonomicName id="A84854674C8A7BF6E9AD2589E226BEF6" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acylophorus agilis" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="agilis">Acylophorus agilis</taxonomicName>
has been collected in a variety of periaquatic habitats including 'floating grass
<normalizedToken originalValue="patches’">patches'</normalizedToken>
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in a eutrophic pond (
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
<bibRefCitation id="0D6BD68429C50D5CD54BF0D90A0AB676" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
), in sediment-laden debris at the edge of forest creeks (
<bibRefCitation author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="815 - 840" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 4." volume="110" year="1978">Smetana 1978</bibRefCitation>
<bibRefCitation id="C8EDE6FCCEA619D468C395A8C65431A6" author="Smetana, A" journalOrPublisher="The Canadian Entomologist" pageId="27" pageNumber="56" pagination="815 - 840" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 4." volume="110" year="1978">Smetana 1978</bibRefCitation>
), at the edge of a sinkhole pond (
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
<bibRefCitation id="229D2F639039126DD34224B9F0BA6007" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous 2008</bibRefCitation>
), and from debris in a beaver lodge.
</paragraph>
</subSubSection>

73
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<paragraph pageId="16" pageNumber="45">Materials.</paragraph>
<paragraph pageId="16" pageNumber="45">CANADA: ON: Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.744, -81.507, soybean field, pitfall, 23-VI-2010 (1), 7-VII-2010 (2), A. Brunke; Auburn, Limekiln Line, 43.736, -81.506, soybean field, pitfall, 23-VI-2010 (1), 7-VII-2010 (1), 4-VIII-2010 (1), 18-VIII-2010 (1), A. Brunke; Waterloo Reg., Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, soybean field, pitfall, 23-VI-2009 (1), 7-VII-2009 (1), 21-VII-2009 (6), 4-VIII-2009 (1), A. Brunke; Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, hedgerow, canopy trap in buckthorn, 27-X-2009, A. Brunke (1); Blair, Fountain St. S. nr Speed River, 43.391 -80.373, soybean field, canopy trap, 15-IX-2009, A. Brunke (1);Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, hedgerow, pitfall, 4-V-2010, A. Brunke (1); Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, soybean field, pitfall, 29-VI-2010, A. Brunke (1); Blair, Whistlebare Rd. and Township Rd. 1, 43.367, -80.358, soybean field, pitfall trap, 13-VII-2010 (1), 25-VIII-2010 (1), A. Brunke; Wellington Co., Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, hedgerow, pitfall, 4-V-2010, A. Brunke (2); Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, soybean, pitfall, 15-VI-2010 (3), 29-VI-2010 (3), A. Brunke; Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, pitfall, 21-VII-2009 (1), 4-VIII-2009 (6), 1-IX-2009 (4), A. Brunke; Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, canopy trap, 4-VIII-2009 (1), 18-VIII-2009 (1), A. Brunke</paragraph>
<subSubSection id="6CF1105DF2E61ED49862234BA1D7CED4" pageId="16" pageNumber="45" type="materials">
<paragraph id="DC84F1EA5040A1105ABC59D0C4BAC7BB" pageId="16" pageNumber="45">Materials.</paragraph>
<paragraph id="8876F607F97B8CE073CCEDCACB09EAD1" pageId="16" pageNumber="45">CANADA: ON: Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.744, -81.507, soybean field, pitfall, 23-VI-2010 (1), 7-VII-2010 (2), A. Brunke; Auburn, Limekiln Line, 43.736, -81.506, soybean field, pitfall, 23-VI-2010 (1), 7-VII-2010 (1), 4-VIII-2010 (1), 18-VIII-2010 (1), A. Brunke; Waterloo Reg., Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, soybean field, pitfall, 23-VI-2009 (1), 7-VII-2009 (1), 21-VII-2009 (6), 4-VIII-2009 (1), A. Brunke; Blair, Dickie Settlement Rd. nr. WhistleBear golf course, 43.373, -80.400, hedgerow, canopy trap in buckthorn, 27-X-2009, A. Brunke (1); Blair, Fountain St. S. nr Speed River, 43.391 -80.373, soybean field, canopy trap, 15-IX-2009, A. Brunke (1);Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, hedgerow, pitfall, 4-V-2010, A. Brunke (1); Blair, Whistlebare Rd. and Township Rd.1, 43.372, -80.362, soybean field, pitfall, 29-VI-2010, A. Brunke (1); Blair, Whistlebare Rd. and Township Rd. 1, 43.367, -80.358, soybean field, pitfall trap, 13-VII-2010 (1), 25-VIII-2010 (1), A. Brunke; Wellington Co., Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, hedgerow, pitfall, 4-V-2010, A. Brunke (2); Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, soybean, pitfall, 15-VI-2010 (3), 29-VI-2010 (3), A. Brunke; Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, pitfall, 21-VII-2009 (1), 4-VIII-2009 (6), 1-IX-2009 (4), A. Brunke; Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, soybean field, canopy trap, 4-VIII-2009 (1), 18-VIII-2009 (1), A. Brunke</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="46" pageId="16" pageNumber="45" type="diagnosis">
<paragraph pageId="16" pageNumber="45">Diagnosis.</paragraph>
<paragraph pageId="16" pageNumber="45">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius appendiculatus" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="appendiculatus">Gabrius appendiculatus</taxonomicName>
<subSubSection id="341795041D1D71EF52C9DA025EBB74DC" lastPageId="17" lastPageNumber="46" pageId="16" pageNumber="45" type="diagnosis">
<paragraph id="5C38B120ADD5BD7553524C56D9F03E32" pageId="16" pageNumber="45">Diagnosis.</paragraph>
<paragraph id="7651F1F82CA94CB69DECB30E2E15F3C2" pageId="16" pageNumber="45">
<taxonomicName id="839D17099252D3CD66F70FCB1CC1DAE9" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius appendiculatus" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="appendiculatus">Gabrius appendiculatus</taxonomicName>
can be distinguished from congenersin the northeast by the following combination of characters: area between basal lines on tergites two and three impunctate; basal antennomeres not distinctly paler than rest of antenna; males with sternite eight broadly notched; females with tergite 10 pointed (as opposed to truncate) apically.
</paragraph>
<paragraph lastPageId="17" lastPageNumber="46" pageId="16" pageNumber="45">
<paragraph id="DD252E428CCFD5CF11121D04973A7D20" lastPageId="17" lastPageNumber="46" pageId="16" pageNumber="45">
This Palaearctic species was first recognized in North America by
<bibRefCitation author="Smetana, A" journalOrPublisher="Le Naturaliste Canadien" pageId="27" pageNumber="56" pagination="175 - 178" title="Gabrius subnigritulus (Reitter), a Palearctic species recently introduced into North America (Coleoptera: Staphylinidae)." volume="116" year="1989">Smetana (1989)</bibRefCitation>
<bibRefCitation id="5DC3DBB3862D2B45F7F8131C59BAD87C" author="Smetana, A" journalOrPublisher="Le Naturaliste Canadien" pageId="27" pageNumber="56" pagination="175 - 178" title="Gabrius subnigritulus (Reitter), a Palearctic species recently introduced into North America (Coleoptera: Staphylinidae)." volume="116" year="1989">Smetana (1989)</bibRefCitation>
(as
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius subnigritulus" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="subnigritulus">Gabrius subnigritulus</taxonomicName>
<taxonomicName id="4F141F48297D53F4E8D21A071765F208" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius subnigritulus" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="subnigritulus">Gabrius subnigritulus</taxonomicName>
(Reitter)) based on specimens collected as early as 1978 in Ormstown,
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="5DFD830B536F5FEA463B1C676C92B809" originalValue="Québec">Quebec</normalizedToken>
and from other localities in British Columbia and Newfoundland; the earliest records from the west coast of North America were from 1979.
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimaszewski et al. (2005)</bibRefCitation>
<bibRefCitation id="BE6645995B0710FF8CF6B489CBEBE6E5" author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimaszewski et al. (2005)</bibRefCitation>
newly recorded it from red spruce-dominated forests in New Brunswick. Herein we report the dispersal of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius appendiculatus" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="appendiculatus">Gabrius appendiculatus</taxonomicName>
<taxonomicName id="1E92F0D5D8AB04544C1FB20407EBD94D" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius appendiculatus" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="appendiculatus">Gabrius appendiculatus</taxonomicName>
into the southern portion of Ontario from specimens collected in 2009-2010 (Map 25). No other specimens before this period are known to have been collected in Ontario. In its native range,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius appendiculatus" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="appendiculatus">Gabrius appendiculatus</taxonomicName>
<taxonomicName id="5BCE66B7F77552786B1B73DC4CFE5664" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius appendiculatus" order="Coleoptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="appendiculatus">Gabrius appendiculatus</taxonomicName>
is widespread in distribution (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="C6729096336983A26BE89370CE8011D9" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
<normalizedToken id="3EA01F29CB70E30253B3F5571E8200E6" originalValue="Löbl">Loebl</normalizedToken>
and Smetana 2004
</bibRefCitation>
). Numerous specimens collected from soybean fields in Ontario further support the
<pageBreakToken pageId="17" pageNumber="46" start="start">conclusion</pageBreakToken>
<pageBreakToken id="79B35A8786832125F44835A42C041461" pageId="17" pageNumber="46" start="start">conclusion</pageBreakToken>
of
<bibRefCitation author="Smetana, A" journalOrPublisher="Le Naturaliste Canadien" pageId="27" pageNumber="56" pagination="175 - 178" title="Gabrius subnigritulus (Reitter), a Palearctic species recently introduced into North America (Coleoptera: Staphylinidae)." volume="116" year="1989">Smetana (1989)</bibRefCitation>
<bibRefCitation id="234F5E90BBE5CC37BFCA9902FA82BA06" author="Smetana, A" journalOrPublisher="Le Naturaliste Canadien" pageId="27" pageNumber="56" pagination="175 - 178" title="Gabrius subnigritulus (Reitter), a Palearctic species recently introduced into North America (Coleoptera: Staphylinidae)." volume="116" year="1989">Smetana (1989)</bibRefCitation>
that this species is well-adapted to agricultural habitat in North America.
</paragraph>
<caption pageId="17" pageNumber="46">
<paragraph pageId="17" pageNumber="46">
<caption id="512D08F83444CB076240AB6C313A1D42" pageId="17" pageNumber="46">
<paragraph id="A9F7A29DD0794928DFB6B468CF45CBFB" pageId="17" pageNumber="46">
Maps 25-28. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 25
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius appendiculatus" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="appendiculatus">Gabrius appendiculatus</taxonomicName>
<taxonomicName id="B59DEE6CF75AC84E5ECF60F64EDCE345" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius appendiculatus" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="appendiculatus">Gabrius appendiculatus</taxonomicName>
(Sharp) (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="1AD746E4EBB27E627A9FBE06BB59EF04" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
,
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimazewski et al. 2005</bibRefCitation>
<bibRefCitation id="769468AFE77F9DEAF8F5EE767EE52A5B" author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimazewski et al. 2005</bibRefCitation>
) 26
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius vindex" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="vindex">Gabrius vindex</taxonomicName>
<taxonomicName id="32605754B15AC5C3EA1DA9AE3CD9FDEE" class="Insecta" family="Staphylinidae" genus="Gabrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gabrius vindex" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="vindex">Gabrius vindex</taxonomicName>
Smetana (
<bibRefCitation author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
<bibRefCitation id="FEBE08F3A112152C7710CFCB4DDED900" author="Smetana, A" journalOrPublisher="Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International" pageId="27" pageNumber="56" title="Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae)" volume="3" year="1995">Smetana 1995</bibRefCitation>
) 27
<taxonomicName class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius occidentoides" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="occidentoides">Neobisnius occidentoides</taxonomicName>
<taxonomicName id="D674C61E4884A132E31DA05CB2B2C25D" class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius occidentoides" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="occidentoides">Neobisnius occidentoides</taxonomicName>
Frank (
<bibRefCitation author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
<bibRefCitation id="AFCE6AC5359C485B860660B992AA3E1A" author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
) 28
<taxonomicName class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis</taxonomicName>
<taxonomicName id="40D1F530829421BDC38C7EEB9AEB652B" class="Insecta" family="Staphylinidae" genus="Neobisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisnius terminalis" order="Coleoptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="terminalis">Neobisnius terminalis</taxonomicName>
(LeConte) (
<bibRefCitation author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
<bibRefCitation id="7B661010295F045245727E25D34C4BCC" author="Frank, JH" journalOrPublisher="Occasional Papers of the Florida State Collection of Arthropods" pageId="26" pageNumber="55" pagination="1 - 160" title="A revision of the New World species of the genus Neobisnius Ganglbauer (Coleoptera: Staphylinidae: Staphylininae)." volume="1" year="1981 b">Frank 1981b</bibRefCitation>
).
</paragraph>
</caption>

86
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<paragraph pageId="12" pageNumber="41">Materials.</paragraph>
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CANADA: ON:Cochrane Dist., N. Moosonee, sandy beach, ridge along coastal marsh,
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,
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,
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and herbs, pitfall trap, 23-VI-1990, J. Pilny, (1).
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Diagnosis.
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is readily distinguished from others of the genus in the northeast by the combination of: body bicolored with orange elytra; pronotum with five punctures in each dorsal row; eyes small, with the space behind them about three times longer (Fig. 6).
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At the time of the most recent revision of the genus, this species was known from only two specimens, from Alberta and Manitoba (
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). Later, one specimen
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collected in the northern Boreal Forest Region of
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="3AB226E17E64681431A6324D7BFAA9FB" originalValue="Québec">Quebec</normalizedToken>
(
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<bibRefCitation id="1A3C5E22822B44B9C7A4122A4E84ADFA" author="Paquin, P" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="27" pageNumber="56" pagination="57 - 98" title="Beetles of the boreal: a faunistic survey carried out in western Quebec." volume="132" year="2001">
Paquin and
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2001
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). Herein we report the fourth specimen known and newly record
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from boreal Ontario (Map 19). This species is apparently transboreal in distribution and its poor representation in collections may be due to a cryptic microhabitat. Its relatively small eyes suggest a subterranean existence in the burrows of mammals, similar to that of certain other
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">Bisnius</taxonomicName>
<taxonomicName id="DB9F3C2F4B609C2458FFF058B5B78753" class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bisnius" order="Coleoptera" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">Bisnius</taxonomicName>
species.
</paragraph>
</subSubSection>

102
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<taxonomicName authority="Mannerheim, 1830" authorityName="Mannerheim" authorityYear="1830" class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus (Mannerheim, 1830)</taxonomicName>
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<paragraph pageId="9" pageNumber="38">Materials.</paragraph>
<paragraph lastPageId="10" lastPageNumber="39" pageId="9" pageNumber="38">
<subSubSection id="40E2636422C4A963055A3C6952D2F9EA" lastPageId="10" lastPageNumber="39" pageId="9" pageNumber="38" type="materials">
<paragraph id="E8B4B1B8236C0913C164094BD7544275" pageId="9" pageNumber="38">Materials.</paragraph>
<paragraph id="E20D95F7CF2E5DE0D9DD8ABE0173DAB4" lastPageId="10" lastPageNumber="39" pageId="9" pageNumber="38">
CANADA: ON: Huron Co., Auburn, Hullett-McKillop Rd. nr. Limekiln Line, 43.742, -81.514, hedgerow, pitfall, 26-V-2010 (1) A. Brunke; Auburn, Limekiln Line, 43.736, -81.506, hedgerow, canopy trap in buckthorn, 26-V-2010 (2) A. Brunke; Benmiller, Sharpes Creek Line, 43.691, -81.608, hedgerow nr. creek, pitfall, 11-V-2009 (1) A. Brunke; Muskoka Reg., S. Waseosa Rd., 8-VII-1996 (1)W. J. Crins; Wellington Co., Guelph, 26-V-1978 (1) Ron O. Kreazer; Guelph, under rock, 16
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III-1983 (1) Brian Brown; Guelph, University Arboretum nature reserve, sifting beech litter, 3-V-2009 (4) A. Brunke and D.K.B. Cheung, sifting litter, 6-VI-2009 (1) A. Brunke; York Co., Toronto, 2-V-1959 (2) R. J. Pilfrey.
</paragraph>
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<paragraph pageId="10" pageNumber="39">Diagnosis.</paragraph>
<paragraph pageId="10" pageNumber="39">
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<paragraph id="D0376CFF3C1515843B522575203BC534" pageId="10" pageNumber="39">Diagnosis.</paragraph>
<paragraph id="78B94DDAA26ADBC4001E5DE1F85E124F" pageId="10" pageNumber="39">
The genus
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is in need of revision in North America, and
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<taxonomicName id="2564A152277057E826F62CD95FB4604E" class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus</taxonomicName>
is currently recognizable in North America only from the characteristic modifications of the male seventh sternite and aedeagus (Fig. 15-16).
</paragraph>
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<paragraph id="195893C2AF291962C5AD3033C53A5AF8" lastPageId="11" lastPageNumber="40" pageId="10" pageNumber="39">
This exotic, Palaearctic species was first recognized in North America by
<bibRefCitation author="Campbell, JM" editor="Bousquet, Y" journalOrPublisher="Agriculture Canada, Ottawa" pageId="26" pageNumber="55" pagination="86 - 129" title="Family Staphylinidae" volumeTitle="Checklist of beetles of Canada and Alaska" year="1991">Campbell and Davies (1991)</bibRefCitation>
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from
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but specimen data were not given and the Palearctic species had not yet been revised at that time. Herein we confirm its presence in North
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based on comparisons with illustrations in
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and newly report it from Ontario based upon specimens collected across southern Ontario as early as 1959 (Map 15).
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is widely distributed in the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="C18EFE09D070DCD01378A9913FF4FD69" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
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and Smetana 2004
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). In North America, specimens have been sifted from deciduous litter in a small fragment of mature forest and found under a rock.
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<taxonomicName id="85682DDFFF74C8177095E7041B19531F" class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus</taxonomicName>
is a common species in its native range and typically inhabits leaf litter and compost (
<bibRefCitation author="Assing, V" journalOrPublisher="Bonner Zoologische Beitrage" pageId="25" pageNumber="54" pagination="33 - 82" title="A revision of the Medon species of the eastern Mediterranean and adjacent regions." volume="52" year="2004">Assing 2004</bibRefCitation>
<bibRefCitation id="D24A2EB1639A7267D97BD7EBE703C4C3" author="Assing, V" journalOrPublisher="Bonner Zoologische Beitrage" pageId="25" pageNumber="54" pagination="33 - 82" title="A revision of the Medon species of the eastern Mediterranean and adjacent regions." volume="52" year="2004">Assing 2004</bibRefCitation>
).
</paragraph>
</subSubSection>

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<paragraph pageId="9" pageNumber="38">Materials.</paragraph>
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CANADA: PEI: Stanhope Beach, National Park,
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. sur plage (=beach debris), 23-VII-1979 (2) R. Sexton.
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<paragraph pageId="9" pageNumber="38">Diagnosis.</paragraph>
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can be identified to the
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group of species by the combination of the complete elytral epipleuron, the undivided labrum and the lack of a suture on the epipleuron (
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). Within this group in northeastern North America, it is best recognized by the combination of: pronotal pubescence directed toward the midline; dark elytral maculation reaching lateral margin of scutellum; basal angle of pronotum strongly sinuate; pronotum with coarse punctures, separated by their widths.
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This species is widely distributed along the coast of eastern North America and was previously known from Georgia, Maine, Maryland, Massachusetts, Newfoundland, New Jersey, New York, North Carolina, Nova Scotia, Rhode Island (
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), and New Brunswick (
<bibRefCitation author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="115 - 150" title="The coastal rove beetles (Coleoptera, Staphylinidae) of Atlantic Canada: a survey and new records." volume="2" year="2008">Majka et al. 2008</bibRefCitation>
<bibRefCitation id="5ADD33176566442A37FEDADD396A3EE7" author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="115 - 150" title="The coastal rove beetles (Coleoptera, Staphylinidae) of Atlantic Canada: a survey and new records." volume="2" year="2008">Majka et al. 2008</bibRefCitation>
). Herein we newly report it from Prince Edward Island (Map 12). It occurs along marine coastline on 'moist, un-vegetated
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, intertidal zones and often away from shore on the leeward sides of islands and peninsulas (
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<bibRefCitation id="9C9BED74E3E3397735EE5E1FCAA065EC" author="Herman, L" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="26" pageNumber="55" pagination="71 - 172" title="Revision of Bledius and related genera. Part II. The armatus, basalis, and mandibularis (Coleoptera, Staphylinidae, Oxytelinae)." volume="157" year="1976">Herman 1976</bibRefCitation>
). The specimen from vegetative beach debris in Prince Edward Island may have been a dispersing individual seeking refuge from desiccation.
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<taxonomicName id="B93A25B97C6D5DB36CACE7E2497883D9" ID-CoL="78PXW" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis Bernhauer, 1908</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="52" pageId="22" pageNumber="51" type="materials">
<paragraph pageId="22" pageNumber="51">Materials.</paragraph>
<paragraph pageId="22" pageNumber="51">UNITED STATES: VT: Bennington Co., Woodford, sifting leaf litter near stream, 1-IV-2010, T. Murray (1).</paragraph>
<paragraph pageId="22" pageNumber="51">
<subSubSection id="57143AC94C71CF7483D7F2D74210E3B8" lastPageId="23" lastPageNumber="52" pageId="22" pageNumber="51" type="materials">
<paragraph id="BA95DABA53F2F75E569E2DA52A1DA26A" pageId="22" pageNumber="51">Materials.</paragraph>
<paragraph id="D32F1336556360CA866B47CCD8D8817B" pageId="22" pageNumber="51">UNITED STATES: VT: Bennington Co., Woodford, sifting leaf litter near stream, 1-IV-2010, T. Murray (1).</paragraph>
<paragraph id="6A0184A9EAF396D7B94DCAF30AE680B2" pageId="22" pageNumber="51">
CANADA: ON:
<normalizedToken originalValue="‘Ont.’">'Ont.'</normalizedToken>
<normalizedToken id="ECC7D3BCACF79B881F85DEFB474469C3" originalValue="‘Ont.’">'Ont.'</normalizedToken>
, 30-IX-1982, G. Abayo (1); Halton Reg., Oakville, nr hwy 25 and Burhamthorpe Rd., meadow, yellow pans, 12 to 14-IX-2003, S.M. Paiero (1); Hamilton Reg., Hamilton, 3-VIII-1984, M.T. Kasserra (1); Waterloo Reg., Blair, Fountain St. S. nr Speed River, 43.391, -80.373, hedgerow, pitfall, 28-IX-2009, A. Brunke (1); Wellington Co., Eramosa, Wellington County Rds. 124 and 29, 43.615, -80.215, hedgerow, pitfall, 4-V-2010 (1), 18-V-2010 (1), 2-XI-2010 (1), A. Brunke; Guelph, 17-VIII-1976, David Levin (1); Guelph, 7-VI-1983, C.F. Langlois (1); Guelph, University Arboretum, hand collected, 16-III-1983, L.B. Carlson (1); Guelph, 30-IX-1983, A. Harris (1); Guelph, 5-VII-1984,
<normalizedToken originalValue="‘maple’">'maple'</normalizedToken>
<normalizedToken id="91607F802A9F08FB96CAE7BBB57B74A3" originalValue="‘maple’">'maple'</normalizedToken>
, T. Young (1); Guelph, 3-IV-1991, M. Kovacevick (1); Guelph, 14-X-1998, T. Phillips (1); Guelph, Victoria Rd. and Conservation Line, 43.580, -80.275, hedgerow, pitfall, 19-V-2009 (1), 17-XI-2009 (1), A. Brunke.
</paragraph>
<paragraph pageId="22" pageNumber="51">
<paragraph id="803359BDCDF2E20EB10F249EB7BF759E" pageId="22" pageNumber="51">
Diagnosis.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<taxonomicName id="FB74158408153D50689141C9A762FA95" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
can be distinguished from other northeastern
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
<taxonomicName id="B5757FB942F3B6287E99770D38D45F88" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
by the combination of: elytra with even punctation; labrum not bilobed; scutellum impunctate; basal antennomeres not distinctly darker than the other segments (Fig. 13).
</paragraph>
<caption pageId="22" pageNumber="51">
<paragraph pageId="22" pageNumber="51">
<caption id="934F5C0D2AB3B5653315025AEE41B9AF" pageId="22" pageNumber="51">
<paragraph id="89DFD99018EBED107BD5B34893DD6EFF" pageId="22" pageNumber="51">
Figure 13. Dorsal habitus of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<taxonomicName id="B41DE7287CB9E8B0F5B286FD1670DA85" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
Bernhauer.
</paragraph>
</caption>
<caption pageId="22" pageNumber="51">
<paragraph pageId="22" pageNumber="51">
<caption id="E05ADA22B5107CE6B73244A25E9EE680" pageId="22" pageNumber="51">
<paragraph id="397E13D4522F935D49986770C5DEB962" pageId="22" pageNumber="51">
Figures 14-19. 14
<taxonomicName class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis</taxonomicName>
<taxonomicName id="F135934EF35B100D4D6BE350FA27D298" class="Insecta" family="Staphylinidae" genus="Stenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenus clavicornis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="clavicornis">Stenus clavicornis</taxonomicName>
(Scopoli), aedeagus in parameral view 15
<taxonomicName class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus</taxonomicName>
<taxonomicName id="100FA02A35C0CCCE8F74910E654E342F" class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus</taxonomicName>
(Mannerheim), 7th sternite 16
<taxonomicName class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus</taxonomicName>
<taxonomicName id="D096FB65698FE846932D4F216D1D7642" class="Insecta" family="Staphylinidae" genus="Medon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Medon fusculus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="fusculus">Medon fusculus</taxonomicName>
, aedeagus in lateral view 17
<taxonomicName class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus minutus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="minutus">Scopaeus minutus</taxonomicName>
<taxonomicName id="2A5543530248E529D7A224082119E14C" class="Insecta" family="Staphylinidae" genus="Scopaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopaeus minutus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="minutus">Scopaeus minutus</taxonomicName>
Erichson, aedeagus in lateral view 18
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus</taxonomicName>
<taxonomicName id="71DB77A4F74156C95BC8EF382B02EC3D" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius nanus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="nanus">Erichsonius nanus</taxonomicName>
(Horn), aedeagus in lateral view 19
<taxonomicName class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
<taxonomicName id="32A6CA81895A17FD274D75A116FDBB6F" class="Insecta" family="Staphylinidae" genus="Erichsonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichsonius parcus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="parcus">Erichsonius parcus</taxonomicName>
(Horn), aedeagus in parameral view.
</paragraph>
</caption>
<paragraph lastPageId="23" lastPageNumber="52" pageId="22" pageNumber="51">
<paragraph id="A80D2BB322E6495EDCD2D6C548817D68" lastPageId="23" lastPageNumber="52" pageId="22" pageNumber="51">
This exotic Palaearctic species was first correctly reported in North America by
<bibRefCitation author="Korge, H" journalOrPublisher="Deutsche Entomologische Zeitschrift (Neue Folge)" pageId="27" pageNumber="56" pagination="332 - 335" title="Beitrage zur Kenntnis der Untergattung Quedius der Gattung Quedius Steph. (Col. Staphylinidae)." volume="9" year="1962">Korge (1962)</bibRefCitation>
<bibRefCitation id="7A153AEC46CC93D49AA1B76730E6C277" author="Korge, H" journalOrPublisher="Deutsche Entomologische Zeitschrift (Neue Folge)" pageId="27" pageNumber="56" pagination="332 - 335" title="Beitrage zur Kenntnis der Untergattung Quedius der Gattung Quedius Steph. (Col. Staphylinidae)." volume="9" year="1962">Korge (1962)</bibRefCitation>
who recognized that
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius parallelus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="parallelus">Quedius parallelus</taxonomicName>
<taxonomicName id="A3438E19C3D96C7AAED6AEC49CB82C1C" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius parallelus" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="parallelus">Quedius parallelus</taxonomicName>
Hatch 1957, a species described from western North America, was synonymous with
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<taxonomicName id="A45978CCA14456D2E3F3C36F2CB5114D" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
Bernhauer 1908.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<pageBreakToken pageId="23" pageNumber="52" start="start">has</pageBreakToken>
<taxonomicName id="6B0FC1BB796A62546AFB04B14F82F485" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<pageBreakToken id="90BAF35037BB604BA7E98D8A5110E8B6" pageId="23" pageNumber="52" start="start">has</pageBreakToken>
been present in North America as early as 1934 based on specimens from Washington and also occurs in British Columbia and Oregon (
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
<bibRefCitation id="D5E934526EE08F0665B17E4254BE41FD" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
). A separate introduction to the east was detected first by
<bibRefCitation author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana (1990)</bibRefCitation>
<bibRefCitation id="FFF9D59BBC0EB76136A5B9874F242F37" author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana (1990)</bibRefCitation>
based on a New Hampshire specimen collected in 1983. Since then
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<taxonomicName id="7D9701E8EC8EF9A006577228549D1965" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
has been detected in New Brunswick and Nova Scotia (
<bibRefCitation author="Majka, CG" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="27" pageNumber="56" pagination="427 - 434" title="New records of introduced species of Quedius Stephens, 1829 (Coleoptera: Staphylinidae) from the Maritime provinces of the Canada." volume="109" year="2007">Majka and Smetana 2007</bibRefCitation>
<bibRefCitation id="773B3CBCDA82C69B21EED663D178E5E6" author="Majka, CG" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="27" pageNumber="56" pagination="427 - 434" title="New records of introduced species of Quedius Stephens, 1829 (Coleoptera: Staphylinidae) from the Maritime provinces of the Canada." volume="109" year="2007">Majka and Smetana 2007</bibRefCitation>
). Herein we newly report it from Ontario and Vermont (Map 37) from specimens collected as early as 1976 and 2010, respectively. The record from Guelph, Ontario represents the earliest known collection in eastern North America. In its native range,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<taxonomicName id="30969D5235D2C2453BEA65C280EC7880" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
is widely distributed in the Palaearctic region (
<bibRefCitation author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
<bibRefCitation id="DFF3F2AAAE548C81305FBE8D37B01248" author="Smetana, A" editor="Lobl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="56" title="Staphylininae" volumeTitle="Catalogue of Palearctic Coleoptera" year="2004">
Smetana in
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
<normalizedToken id="9202B83D03D897B4ED51F5FFE6E3BB43" originalValue="Löbl">Loebl</normalizedToken>
and Smetana 2004
</bibRefCitation>
).
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<taxonomicName id="608A3DA53426AD498EA9EFD8D7596A4D" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
has been collected in Ontario mainly in disturbed habitat such as regenerating woodland, fields and agricultural crops.
</paragraph>
<caption pageId="23" pageNumber="52">
<paragraph pageId="23" pageNumber="52">
<caption id="64CB2E3A433E21332D8A9D18FF5A9D2B" pageId="23" pageNumber="52">
<paragraph id="0D9E90B86F2943424EC4805950C45060" pageId="23" pageNumber="52">
Maps 37-38. Distribution in northeastern North America, sources of data other than DEBU are quoted in parentheses. 37
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
<taxonomicName id="0B90D821A4041108172517FB59B6A680" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius curtipennis" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="curtipennis">Quedius curtipennis</taxonomicName>
Bernhauer (
<bibRefCitation author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana 1990</bibRefCitation>
<bibRefCitation id="AEF93297D1A5062A60058D107D83E13B" author="Smetana, A" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="56" pagination="95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana 1990</bibRefCitation>
,
<bibRefCitation author="Majka, CG" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="27" pageNumber="56" pagination="427 - 434" title="New records of introduced species of Quedius Stephens, 1829 (Coleoptera: Staphylinidae) from the Maritime provinces of the Canada." volume="109" year="2007">Majka and Smetana 2007</bibRefCitation>
<bibRefCitation id="8C18D4E3F5FC0C2427371F456E84CE35" author="Majka, CG" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="27" pageNumber="56" pagination="427 - 434" title="New records of introduced species of Quedius Stephens, 1829 (Coleoptera: Staphylinidae) from the Maritime provinces of the Canada." volume="109" year="2007">Majka and Smetana 2007</bibRefCitation>
,
<bibRefCitation author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="151 - 174" title="Adventive Staphylinidae (Coleoptera) of the Maritime Provinces of Canada: further contributions." volume="2" year="2008 b">Majka and Klimaszewski 2008b</bibRefCitation>
<bibRefCitation id="981F6A02BB9F3B752276A3620EAF5BAE" author="Majka, CG" journalOrPublisher="ZooKeys" pageId="27" pageNumber="56" pagination="151 - 174" title="Adventive Staphylinidae (Coleoptera) of the Maritime Provinces of Canada: further contributions." volume="2" year="2008 b">Majka and Klimaszewski 2008b</bibRefCitation>
) 38
<taxonomicName class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
<taxonomicName id="E433D3D721996B113B3CE15ECBCBF20C" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius fulgidus" order="Coleoptera" pageId="23" pageNumber="52" phylum="Arthropoda" rank="species" species="fulgidus">Quedius fulgidus</taxonomicName>
(Fabricius) (
<bibRefCitation author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
<bibRefCitation id="5B61A9E3E15CCB2DF813DF06CC6E6F28" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="27" pageNumber="56" pagination="1 - 303" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." volume="79" year="1971">Smetana 1971</bibRefCitation>
,
<bibRefCitation author="Peck, SB" journalOrPublisher="Journal of Cave and Karst Studies" pageId="27" pageNumber="56" pagination="3 - 8" title="The cave-inhabiting rove beetles of the United States (Coleoptera; Staphylinidae; excluding Aleocharinae and Pselaphinae): diversity and distributions." volume="65" year="2003">Peck and Thayer 2003</bibRefCitation>
<bibRefCitation id="288C231B87F6085E1855643B82A0E07B" author="Peck, SB" journalOrPublisher="Journal of Cave and Karst Studies" pageId="27" pageNumber="56" pagination="3 - 8" title="The cave-inhabiting rove beetles of the United States (Coleoptera; Staphylinidae; excluding Aleocharinae and Pselaphinae): diversity and distributions." volume="65" year="2003">Peck and Thayer 2003</bibRefCitation>
).
</paragraph>
</caption>

100
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<paragraph pageId="9" pageNumber="38">Materials.</paragraph>
<paragraph pageId="9" pageNumber="38">United States: VA:Giles Co., Ripplemead, Rte. 460 at bridge, flood debris, 11 to 25-V-2008 (6) A. Brunke.</paragraph>
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<paragraph id="888CC5E118FE2BCF75125A0B430F651D" pageId="9" pageNumber="38">Materials.</paragraph>
<paragraph id="BD83EF905C01161AFF88468FE325B64C" pageId="9" pageNumber="38">United States: VA:Giles Co., Ripplemead, Rte. 460 at bridge, flood debris, 11 to 25-V-2008 (6) A. Brunke.</paragraph>
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<paragraph pageId="9" pageNumber="38">Diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="38">
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<paragraph id="4EBDF3078654AB072A6068D8F472BB4E" pageId="9" pageNumber="38">Diagnosis.</paragraph>
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is the only northeastern member of the genus and its distinct habitus will readily identify it as a
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(Fig. 5).
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This species is rarely collected and at the time of the most recent revision, it was only known from scattered localities in District of Columbia, Kentucky, New Jersey, Missouri, Oklahoma, Texas (
<bibRefCitation pageId="9" pageNumber="38">Herman 1970</bibRefCitation>
<bibRefCitation id="8D490988DDF445FD1AE17E79665FAB91" pageId="9" pageNumber="38">Herman 1970</bibRefCitation>
), Arkansas, and South Carolina (
<bibRefCitation author="Sanderson, MW" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="27 - 30" title="The North American species of Stilicolina Casey (Coleoptera, Staphylinidae)." volume="20" year="1947">Sanderson 1947</bibRefCitation>
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).
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was described from
<normalizedToken originalValue="‘Pennsylvania’">'Pennsylvania'</normalizedToken>
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by
<bibRefCitation author="Melsheimer, FE" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="27" pageNumber="56" pagination="26 - 43" title="Descriptions of new species of North American Coleoptera." volume="2" year="1844">Melsheimer (1844)</bibRefCitation>
<bibRefCitation id="03998C9245044482326771215D6F20A6" author="Melsheimer, FE" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="27" pageNumber="56" pagination="26 - 43" title="Descriptions of new species of North American Coleoptera." volume="2" year="1844">Melsheimer (1844)</bibRefCitation>
but no locality was given.
<bibRefCitation author="Sanderson, MW" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="27 - 30" title="The North American species of Stilicolina Casey (Coleoptera, Staphylinidae)." volume="20" year="1947">Sanderson (1947)</bibRefCitation>
<bibRefCitation id="3F252375D55CF73090496CE6C45040B2" author="Sanderson, MW" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="27" pageNumber="56" pagination="27 - 30" title="The North American species of Stilicolina Casey (Coleoptera, Staphylinidae)." volume="20" year="1947">Sanderson (1947)</bibRefCitation>
stated that it had been recorded from Ohio but we could find no mention of this in the literature. We here newly record it from Virginia (Map 14). This species appears to be a specialist in stream and river drift/flood litter, although it is occasionally found in caves (
<bibRefCitation author="Peck, SB" journalOrPublisher="Journal of Cave and Karst Studies" pageId="27" pageNumber="56" pagination="3 - 8" title="The cave-inhabiting rove beetles of the United States (Coleoptera; Staphylinidae; excluding Aleocharinae and Pselaphinae): diversity and distributions." volume="65" year="2003">Peck and Thayer 2003</bibRefCitation>
<bibRefCitation id="86AA7EC34D3B111B55903861AC62A160" author="Peck, SB" journalOrPublisher="Journal of Cave and Karst Studies" pageId="27" pageNumber="56" pagination="3 - 8" title="The cave-inhabiting rove beetles of the United States (Coleoptera; Staphylinidae; excluding Aleocharinae and Pselaphinae): diversity and distributions." volume="65" year="2003">Peck and Thayer 2003</bibRefCitation>
).
<bibRefCitation author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
<bibRefCitation id="50DFB64CF4DD3F06103C3C75A151992C" author="Watrous, L" journalOrPublisher="Lincoln Co., MO. Missouri Department of Natural Resources, Ballwin, Mo." pageId="27" pageNumber="56" title="Contribution to the Taxonomic Inventory of Coleoptera at Cuivre River State Park" year="2008">Watrous (2008)</bibRefCitation>
recently recommended
<taxonomicName class="Insecta" family="Staphylinidae" genus="Eustilicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustilicus tristis" order="Coleoptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="tristis">Eustilicus tristis</taxonomicName>
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for S3 ranking in Missouri based on its specialized, sensitive habitat and general rarity over its known distribution.
</paragraph>
</subSubSection>

102
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.75.767</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-75-29</mods:identifier>
<mods:classification id="3132E991C6A4B3B81738CED2D24A4685">journal article</mods:classification>
<mods:identifier id="F7927C6305927C27247307C11664357C" type="DOI">http://dx.doi.org/10.3897/zookeys.75.767</mods:identifier>
<mods:identifier id="175F9365D80B466602710BBEEE5604AB" type="Pensoft-Pub">1313-2970-75-29</mods:identifier>
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<treatment ID-GBIF-Taxon="159361028" LSID="urn:lsid:plazi:treatment:FE9093178D886E8B7DE728B62D5C4C71" httpUri="http://treatment.plazi.org/id/FE9093178D886E8B7DE728B62D5C4C71" lastPageNumber="37" pageId="8" pageNumber="37">
<subSubSection pageId="8" pageNumber="37" type="nomenclature">
<paragraph pageId="8" pageNumber="37">
<taxonomicName authority="Campbell, 1979" authorityName="Campbell" authorityYear="1979" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni Campbell, 1979</taxonomicName>
<treatment id="FE9093178D886E8B7DE728B62D5C4C71" ID-GBIF-Taxon="159361028" LSID="urn:lsid:plazi:treatment:FE9093178D886E8B7DE728B62D5C4C71" httpUri="http://treatment.plazi.org/id/FE9093178D886E8B7DE728B62D5C4C71" lastPageNumber="37" pageId="8" pageNumber="37" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Tachyporinae">
<subSubSection id="D97772A037F7325F11AC94CF0FF39AA9" pageId="8" pageNumber="37" type="nomenclature">
<paragraph id="16BB642A7531474FD84C924C9C34715B" pageId="8" pageNumber="37">
<taxonomicName id="50A14C83757199E8B462EFA058449E18" ID-CoL="8VSLS" authority="Campbell, 1979" authorityName="Campbell" authorityYear="1979" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni Campbell, 1979</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="37" type="materials">
<paragraph pageId="8" pageNumber="37">Materials.</paragraph>
<paragraph pageId="8" pageNumber="37">CANADA: ON:Huron Co., Benmiller, Sharpes Creek Line, 43.691, -81.608, soybean field, pitfall, 18-IX-2009, A. Brunke (1).</paragraph>
<paragraph pageId="8" pageNumber="37">UNITED STATES: NH: Coos Co., Dixville, leaf litter, 6-IV-2010, T. Murray (1); Jefferson, leaf litter, grassy area near stream, 20-IV-2010, T. Murray (1); Dixville, 4-V-2010, T. Murray (4). MA: Middlesex Co., Groton, sifting hay, flood debris in farm field nr. drainage ditch, 30-IV-2010, T. Murray (1). VT: Orange Co., Topsham, sweeping low vegetation, 22-VI-2010, T. Murray (1).</paragraph>
<subSubSection id="3E937B412BF0FF3EB66FFF2235A1530E" pageId="8" pageNumber="37" type="materials">
<paragraph id="58C13DB0E40A049D858ABBE108DA4869" pageId="8" pageNumber="37">Materials.</paragraph>
<paragraph id="8E457E31903014328BFD9E8CA17B1880" pageId="8" pageNumber="37">CANADA: ON:Huron Co., Benmiller, Sharpes Creek Line, 43.691, -81.608, soybean field, pitfall, 18-IX-2009, A. Brunke (1).</paragraph>
<paragraph id="B09ED8B19F9C3C07EF73641278E42565" pageId="8" pageNumber="37">UNITED STATES: NH: Coos Co., Dixville, leaf litter, 6-IV-2010, T. Murray (1); Jefferson, leaf litter, grassy area near stream, 20-IV-2010, T. Murray (1); Dixville, 4-V-2010, T. Murray (4). MA: Middlesex Co., Groton, sifting hay, flood debris in farm field nr. drainage ditch, 30-IV-2010, T. Murray (1). VT: Orange Co., Topsham, sweeping low vegetation, 22-VI-2010, T. Murray (1).</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="37" type="diagnosis">
<paragraph pageId="8" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="37">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
<subSubSection id="ABF197587EAD993419425F186B9A936B" pageId="8" pageNumber="37" type="diagnosis">
<paragraph id="182F865F56CF0C7BAC632D2911A34BC4" pageId="8" pageNumber="37">Diagnosis.</paragraph>
<paragraph id="95238DE3D498822B30E660812FA128FB" pageId="8" pageNumber="37">
<taxonomicName id="9682BEDFE9104A66B9CE6B66D5936A3A" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
can be easily recognized amongst other northeastern
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">Tachyporus</taxonomicName>
<taxonomicName id="84E3C02DBB383EB10A6F5A180055CA66" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">Tachyporus</taxonomicName>
by the combination of a bicolored abdomen and elytra without black discal markings (Fig. 4). Rarely, specimens occur with a small black marking on the scutellum but it does not extend half the length of the elytra as in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus elegans" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="elegans">Tachinus elegans</taxonomicName>
<taxonomicName id="58B6C8A13EC1B1DA00382857D2D1D811" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus elegans" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="elegans">Tachinus elegans</taxonomicName>
Horn. Additionally,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus elegans" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="elegans">Tachinus elegans</taxonomicName>
<taxonomicName id="1422933824F3673445EEBF79BBE26B3F" class="Insecta" family="Staphylinidae" genus="Tachinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachinus elegans" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="elegans">Tachinus elegans</taxonomicName>
lacks dark markings on the pronotum.
</paragraph>
<paragraph pageId="8" pageNumber="37">
<paragraph id="C0E5FA591CCC0E259566EA171DEA55FD" pageId="8" pageNumber="37">
This distinctive species was known from 12 specimens at the time of its description, all collected from September to November in southern
<normalizedToken originalValue="Québec">Quebec</normalizedToken>
<normalizedToken id="3014726B8787E19CCC3623D0AD4CED1F" originalValue="Québec">Quebec</normalizedToken>
and Connecticut (
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." volume="109" year="1979">Campbell 1979</bibRefCitation>
<bibRefCitation id="D71F272B9754FF0C0CA6CA4C4D24ACD2" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 95" title="A revision of the genus Tachyporus Gravenhorst (Coleoptera: Staphylinidae) of North America." volume="109" year="1979">Campbell 1979</bibRefCitation>
).
<bibRefCitation author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimaszewski et al. (2005)</bibRefCitation>
<bibRefCitation id="321181FC852336715B849F87F99F43FB" author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="26" pageNumber="55" pagination="1 - 48" title="Rove beetles (Coleoptera: Staphylinidae) in red spruce stands, eastern Canada: diversity, abundance, and descriptions of new species." volume="137" year="2005">Klimaszewski et al. (2005)</bibRefCitation>
newly recorded it from red spruce-dominated forest in New Brunswick. Herein we newly report
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
<taxonomicName id="F7D5D0C4C21BCB97CEB6E8F859F73E82" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
from Ontario, New Hampshire, Massachusetts, and Vermont (Map 10). Habitat data suggests that
<taxonomicName class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
<taxonomicName id="190830B424535F567D513187ABB7CC27" class="Insecta" family="Staphylinidae" genus="Tachyporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachyporus browni" order="Coleoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="browni">Tachyporus browni</taxonomicName>
inhabits moist or wet litter/debris near water. Most of the specimens known are from the cooler months of the year and this seasonality is probably responsible for its rarity in collections. This phenomenon is common for many staphylinid groups (e.g., winter-active
<taxonomicName lsidName="" pageId="8" pageNumber="37" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
<taxonomicName id="4D65D7DB634140511DC61264DE7B731C" lsidName="" pageId="8" pageNumber="37" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
in
<bibRefCitation author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 87" title="A revision of the North American Omaliinae (Coleoptera: Staphylinidae). I. The genera Haida Keen, Pseudohaida Hatch, and Eudectoides new genus. 2. The tribe Coryphiini." volume="106" year="1978">Campbell 1978</bibRefCitation>
<bibRefCitation id="B256CBE14C08417B4748392E7EC08297" author="Campbell, JM" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="26" pageNumber="55" pagination="1 - 87" title="A revision of the North American Omaliinae (Coleoptera: Staphylinidae). I. The genera Haida Keen, Pseudohaida Hatch, and Eudectoides new genus. 2. The tribe Coryphiini." volume="106" year="1978">Campbell 1978</bibRefCitation>
) and suggests that increased sampling during September to April will yield further discoveries.
</paragraph>
</subSubSection>