diff --git a/data/C5/2E/90/C52E90E569395E37A5551C76AB307B2C.xml b/data/C5/2E/90/C52E90E569395E37A5551C76AB307B2C.xml
new file mode 100644
index 00000000000..316201e1764
--- /dev/null
+++ b/data/C5/2E/90/C52E90E569395E37A5551C76AB307B2C.xml
@@ -0,0 +1,1077 @@
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+<mods:mods id="3CD73D3E0E8EA7FDDC8FD8099020A2CB" xmlns:mods="http://www.loc.gov/mods/v3">
+<mods:titleInfo id="045F5DF8E8161496FA83483B0A070B3D">
+<mods:title id="5AC12D7D382AAC30B5D2184FC963264C">Integrative delimitation of a new Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae) from the Caucasus with a supplement to the identification guide of Caucasian and Irano-Anatolian species</mods:title>
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+<mods:namePart id="0999C67984F1288B88420DC3BAE1C3CA">Hrivniak, Ľuboš</mods:namePart>
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+<mods:namePart id="5F04F97F91541B0DB2797D8587957BFB">Sroka, Pavel</mods:namePart>
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+<subSubSection id="7DCD2F92A2E1F40FD73FD31BB9539825" type="nomenclature">
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+<taxonomicName id="C63DF23522006A985E1913795E8FDB26" authority="Hrivniak &amp; Sroka" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="1D55D833AF50796DB59F80D25959A60D" italics="true">Epeorus</emphasis>
+(
+<emphasis id="3CB2E33C24BF21169FA9E788ED3A9BEC" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="5672655DAFB72CB8EE24DC764CB0CFCD" italics="true">abditus</emphasis>
+Hrivniak &amp; Sroka
+</taxonomicName>
+<taxonomicNameLabel id="FD7DA0168D895DED92BAF36ED83A4B17" rank="species">sp. nov.</taxonomicNameLabel>
+</paragraph>
+</subSubSection>
+<subSubSection id="6F9BE8C35E20EAA6BB333ED0572D88F3" type="description">
+<paragraph id="304C575EB1F411EED99B46501C1F83DD">
+<figureCitation id="8F20CD50B532CE6A6B2AC73CBB6253FF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">Figs 4</figureCitation>
+, 
+<figureCitation id="4E7E2D5A0118F269489D2CCBFDF884EA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5</figureCitation>
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EPDAE" type="type material">
+<paragraph id="E738E717938D4F7A2A4114EF4014C269">
+<heading id="0263BB900351D391CED24BF6F33175BE" reason="title">Type material.</heading>
+</paragraph>
+<paragraph id="A04CD21EE42B4171E1B892B970BA560C">
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+<emphasis id="39BE1B72E4CEC42F9A05D762CF3035FB" bold="true" italics="true">
+<typeStatus id="43B2640749880FFFA089C98731C9B667" type="holotype">Holotype</typeStatus>
+</emphasis>
+• 
+<specimenCount id="2A38B19DA8C533B7DCBFB2B26170FBC9" count="1" type="larva">female larva</specimenCount>
+(
+<abbrev id="ABBRID0E1DAE" xlink_title="GenBank">
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+, 
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+, 
+<elevation id="44030DE9DA96F7512F9F52609DE2EC11" metricMagnitude="3" metricUnit="m" metricValue="1.599" unit="m" value="1599.0">
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+; code: CAM 6, 
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+, 
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+leg.
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+<typeStatus id="9878F40BC1E3C2FC05D7A655131714E5" type="paratype">Paratypes</typeStatus>
+</emphasis>
+• 
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+(mounted on slide), same data as holotype
+</materialsCitation>
+• 
+<materialsCitation id="601B326BEDC8778E47DF465F0E3BA38D" collectingDate="2013-04-19" collectionCode="GB" collectorName="A. V. Martynov" country="Georgia" county="Kobuleti district" elevation="792" latitude="41.716946" location="Kintrishi River" longLatPrecision="19" longitude="42.044724" municipality="Khino (Didvake) village" specimenCode="PP 987171" specimenCount="1" specimenCount-larva="1" stateProvince="Adjara" typeStatus="paratype">
+<specimenCount id="CD31FD768E1BE8731DDCD0013DAD904E" count="1" type="larva">1 larva</specimenCount>
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+; mounted on slide), 
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+: 
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+, 
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+, vicinity of 
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+, 
+<location id="95DBD8FA3AD4C55B61F529827D6AD03B" LSID="urn:lsid:plazi:treatment:C52E90E569395E37A5551C76AB307B2C:95DBD8FA3AD4C55B61F529827D6AD03B" country="Georgia" county="Kobuleti district" latitude="41.716946" longLatPrecision="19" longitude="42.044724" municipality="Khino (Didvake) village" name="Kintrishi River" stateProvince="Adjara">Kintrishi River</location>
+, 
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+<quantity id="9EFC3DA60BE50142F4E98F385C948F91" metricMagnitude="2" metricUnit="m" metricValue="7.92" unit="m" value="792.0">792 m</quantity>
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+<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5EAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[42.044722,41.716944]}">
+<geoCoordinate id="1199AB1BD52E49529B0069F8ADC707DA" degrees="41" direction="north" minutes="43" orientation="latitude" precision="15" seconds="01" value="41.716946">41°43'01&quot;N</geoCoordinate>
+, 
+<geoCoordinate id="C92B9E6B4554DAF37E0761D145CF2520" degrees="42" direction="east" minutes="02" orientation="longitude" precision="15" seconds="41" value="42.044724">42°02'41&quot;E</geoCoordinate>
+</named-content>
+; code: No 6, 
+<date id="09F6525573AF4D99432DB45684DD41EA" value="2013-04-19">
+<collectingDate id="3B5C0D26B0AA4F3485994AD5A4CAA6CB" value="2013-04-19">19. 4. 2013</collectingDate>
+</date>
+, 
+<collectorName id="312E996734F2154756717DBB50D54BAB">A. V. Martynov</collectorName>
+leg
+</materialsCitation>
+• 
+<materialsCitation id="7259272E05985A628B9D0236D5A31DDA" collectingDate="2019-05-03" collectionCode="GB" collectorName="L. Hrivniak" country="Georgia" elevation="1514" latitude="42.243084" location="Stori River" longLatPrecision="1" longitude="45.495693" municipality="South of Alazani Pass" specimenCode="PP 987168" specimenCount="1" specimenCount-larva="1" stateProvince="Kakheti Province" typeStatus="paratype">
+<specimenCount id="ADE002773B32902861ADFC58C65B2A7D" count="1" type="larva">1 larva</specimenCount>
+(
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+, mouthparts mounted on slide), 
+<collectingCountry id="24D1F23112B3B03807A9BD65C3A6E1BC" name="Georgia">Georgia</collectingCountry>
+: 
+<collectingRegion id="8EB9EF59CCC2DC5E283426C3DB902B36" country="Georgia" name="Kakheti">Kakheti Province</collectingRegion>
+, 
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+, 
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+</named-content>
+; code GEO 60 / 2019, 
+<date id="5C0F18CB5AC0730526C224D01BE8DA9D" value="2019-05-03">
+<collectingDate id="F29F6613629E71DDF56533ACBFA00D16" value="2019-05-03">3. 5. 2019</collectingDate>
+</date>
+; 
+<collectorName id="CB72FE5DF55EAB6605DE348ABA72CFA0">Ľ. Hrivniak</collectorName>
+leg
+</materialsCitation>
+• 
+<materialsCitation id="8B7AA2230AAD8BBE740063F8AFC19C73" collectingDate="2013-05-19" collectionCode="V" collectorName="V. V. Martynov" country="Russia" elevation="2192" latitude="43.241943" location="left tributary of Baksan River" longLatPrecision="19" longitude="42.56361" municipality="Terskol village" specimenCount="1" specimenCount-larva="1" stateProvince="Kabardino-Balkaria" typeStatus="paratype">
+<specimenCount id="B655F7C5D09B25AF3E43556469FC9456" count="1" type="larva">1 larva</specimenCount>
+, 
+<collectingCountry id="B6097FCB742DDB9DCED1F33E344FFE94" name="Russia">Russia</collectingCountry>
+: 
+<collectingRegion id="A2BDA2C45BFACE7087820DED1DF3A618" country="Russia" name="Kabardino-Balkariya">Kabardino-Balkaria</collectingRegion>
+, vicinity of 
+<collectingMunicipality id="6E0C205C73381001CD887407E8FD617D">Terskol village</collectingMunicipality>
+, 
+<location id="FCA1FBD79B4E2E2E3FD7F7AEB17E5BBF" LSID="urn:lsid:plazi:treatment:C52E90E569395E37A5551C76AB307B2C:FCA1FBD79B4E2E2E3FD7F7AEB17E5BBF" country="Russia" latitude="43.241943" longLatPrecision="19" longitude="42.56361" municipality="Terskol village" name="left tributary of Baksan River" stateProvince="Kabardino-Balkaria">left tributary of Baksan River</location>
+, 
+<elevation id="D8311613F68267778B1985CC6A9337BF" metricMagnitude="3" metricUnit="m" metricValue="2.1919999999999997" unit="m" value="2192.0">
+<quantity id="1DE3F22C87FF46704CCD68DE4C657620" metricMagnitude="3" metricUnit="m" metricValue="2.1919999999999997" unit="m" value="2192.0">2192 m</quantity>
+a. s. l.
+</elevation>
+; 
+<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXFAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[42.563611,43.241944]}">
+<geoCoordinate id="DA38D69C36CD5C292305FBC0B8394540" degrees="43" direction="north" minutes="14" orientation="latitude" precision="15" seconds="31" value="43.241943">43°14'31&quot;N</geoCoordinate>
+, 
+<geoCoordinate id="EAEE4FFA4BA57E4E136828A3C853FBEC" degrees="42" direction="east" minutes="33" orientation="longitude" precision="15" seconds="49" value="42.56361">42°33'49&quot;E</geoCoordinate>
+</named-content>
+; 
+<date id="845CBE14044D86E114C1A4A0F3B64ECA" value="2013-05-19">
+<collectingDate id="4360B4FEFF39D17F6215BC921B705388" value="2013-05-19">19. 5. 2013</collectingDate>
+</date>
+, 
+<collectorName id="15FD50208AA000943C80FDFF3C0BE2B8">V. V. Martynov</collectorName>
+leg
+</materialsCitation>
+• 
+<materialsCitation id="C2C0E6749C6AF2675972B047E0C8AC44" collectingDate="2013-05-19" collectionCode="GB" collectorName="V. V. Martynov" country="Russia" elevation="1904" latitude="43.35" location="right tributary of Baksan River" longLatPrecision="1144" longitude="42.866665" municipality="Tyrnyauz village" specimenCode="PP 987169" specimenCount="2" specimenCount-larva="2" stateProvince="Kabardino-Balkaria" typeStatus="paratype">
+<specimenCount id="A0ED058B88B2E22D372298EE9B3D70FB" count="2" type="larva">2 larvae</specimenCount>
+(one barcoded, 
+<abbrev id="ABBRID0E3FAE" xlink_title="GenBank">
+<collectionCode id="824DC5B1B2740AD0E1380DC35F0D63AF">GB</collectionCode>
+</abbrev>
+: 
+<ext-link id="121F9B5FBC364060CCF99408BFA4BAB0" ext-link-type="gen" xlink_href="PP987169" xlink_type="simple">
+<specimenCode id="F8E4B656E8E6587BE0A8BAD9F7104191">PP 987169</specimenCode>
+</ext-link>
+), 
+<collectingCountry id="43F79941219E7A6E7C4E97B718284758" name="Russia">Russia</collectingCountry>
+: 
+<collectingRegion id="D440A87BCD7E7F2BE2A3116FB2EBF69E" country="Russia" name="Kabardino-Balkariya">Kabardino-Balkaria</collectingRegion>
+, vicinity of 
+<collectingMunicipality id="C587C81C3254AE0DB2E363B4A674B402">Tyrnyauz village</collectingMunicipality>
+, 
+<location id="7468657EEEA83DC5CE6F9D42AC4166D6" LSID="urn:lsid:plazi:treatment:C52E90E569395E37A5551C76AB307B2C:7468657EEEA83DC5CE6F9D42AC4166D6" country="Russia" latitude="43.35" longLatPrecision="1144" longitude="42.866665" municipality="Tyrnyauz village" name="right tributary of Baksan River" stateProvince="Kabardino-Balkaria">right tributary of Baksan River</location>
+, 
+<elevation id="FC2FB079B840F22879ECA87FD8BDB94D" metricMagnitude="3" metricUnit="m" metricValue="1.904" unit="m" value="1904.0">
+<quantity id="DFD00A5C070CAA2C673CA60A5F0B3170" metricMagnitude="3" metricUnit="m" metricValue="1.904" unit="m" value="1904.0">1904 m</quantity>
+a. s. l.
+</elevation>
+, 
+<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIGAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[42.866667,43.350000]}">
+<geoCoordinate id="DD4859BEE7F938FB5FFF3FFBA6F1826E" degrees="43" direction="north" minutes="21" orientation="latitude" precision="925" value="43.35">43°21'N</geoCoordinate>
+, 
+<geoCoordinate id="7CB931D04A86B56BB88D0A2FCC80235A" degrees="42" direction="east" minutes="52" orientation="longitude" precision="925" value="42.866665">42°52'E</geoCoordinate>
+</named-content>
+; 
+<date id="7C854934FAA2A5481E483E16DDB7F359" value="2013-05-19">
+<collectingDate id="D28E5F7179D969A38008CAFD8D54A0FC" value="2013-05-19">19. 5. 2013</collectingDate>
+</date>
+, 
+<collectorName id="0C6A086D2089E592F1F0B38747C162CB">V. V. Martynov</collectorName>
+leg.
+</materialsCitation>
+</paragraph>
+<paragraph id="421B58B0CF3DCBC7A936758738DB30CD">
+Type material is deposited in 
+<named-content id="CEB5F24CADE8426BE2144E725273F38B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/biology-centre-academy-sciences-czech-republic-institute-entomology" xlink_title="Biology Centre of the Czech Academy of Sciences, Institute of Entomology">
+<collectionCode id="1FF946C10398F7977E50D72932B7A8E0" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" type="Other institutional type">IECA</collectionCode>
+</named-content>
+.
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0ETGAE" type="etymology">
+<paragraph id="DDEA4693F430FE23336A246DDBCA3428">
+<heading id="278D200BED6C23E401C1ACED25D53A9A" reason="title">Etymology.</heading>
+</paragraph>
+<paragraph id="DB2878DB57AFD64BA2CBF5E8C5F9EF00">
+The species name 
+<taxonomicName id="600BF7919B7542C97EE53E378DD49C49" authority="(Latin)" baseAuthorityName="Latin" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus">
+<emphasis id="9E416166616B828A318A677B4CD3BAC4" italics="true">abditus</emphasis>
+(Latin)
+</taxonomicName>
+means hidden. It refers to rare distribution and morphological similarity with related species.
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EDHAE" type="distribution">
+<paragraph id="F1721BA5D08A8CAD19F9B68E911608DC">
+<heading id="D47B22DDC39480EAB2E019D72A46298B" reason="title">Distribution and habitat preferences of larvae.</heading>
+</paragraph>
+<paragraph id="22021A3A06A79DAD7B1692E2AE5829C1">
+<taxonomicName id="0681791AE7B4F573A95D0268672656D7" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="5AC1EA3CA0AD9E96914422C113938C3A" italics="true">Epeorus</emphasis>
+(
+<emphasis id="21A052392B2CCD9B506117768E25AF76" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="CDCE1D465E7675DC8CCC7B0848E17F7D" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="8B3457F27D82B158575CC6B63AEDBABD" rank="species">sp. nov.</taxonomicNameLabel>
+has relatively wide distribution in the Caucasus region but appears to be relatively rare due to low number of specimens obtained by extensive sampling. They were found in the Pontic Mountains and the Lesser Caucasus (northeast 
+<collectingCountry id="98712350E9ABD47975046F8BF1EFC7B1" name="Turkey">Türkiye</collectingCountry>
+and southwest 
+<collectingCountry id="A91A7F5F13D685BB6C80A9131BE012A2" name="Georgia">Georgia</collectingCountry>
+), and the central (
+<collectingCountry id="B830340EA674244488641920F90E3FF7" name="Russia">Russia</collectingCountry>
+: 
+<collectingRegion id="D8B72AB4420A0901038C9F85CB20731F" country="Russia" name="Kabardino-Balkariya">Kabardino-Balkaria</collectingRegion>
+) and eastern (
+<collectingCountry id="5DE10EAA5E9003D18DDFC19D8E06AA25" name="Georgia">Georgia</collectingCountry>
+: 
+<collectingRegion id="18C734C85A4D87E4801A98C72CF3B8B0" country="Georgia" name="Kakheti">Kakheti</collectingRegion>
+) parts of the Greater Caucasus (Fig. 
+<figureCitation id="7F3C6110E630499D82D490868FF1C701" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Epeorus (Caucasiron) abditus sp. nov.; global localisation of a study area (lower left corner) and our sampling sites investigated between 2008–2019 (lower right corner)." figureDoi="10.3897/zookeys.1214.131266.figure1" httpUri="https://binary.pensoft.net/fig/1150504">1</figureCitation>
+). The larvae were found in low abundance in cold and clear streams and rivers between 792 and 
+<quantity id="0678982A7A7E502F1BBFC9EC2BE4893C" metricMagnitude="3" metricUnit="m" metricValue="2.1919999999999997" unit="m" value="2192.0">2192 m</quantity>
+a. s. l. on stony bed substrate in turbulent flow (Fig. 
+<figureCitation id="BDD6892C4A9EB1858512BB7B93D94625" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The habitat (type locality) of Epeorus (Caucasiron) abditus sp. nov. from northeast Türkiye (photo: G. Türkmen)." figureDoi="10.3897/zookeys.1214.131266.figure3" httpUri="https://binary.pensoft.net/fig/1150506">3</figureCitation>
+). They were not recorded in urban and agricultural areas within the region, where many localities were investigated. Larvae co-occurred with 
+<taxonomicName id="605710B007F1A6D2BC73B766EF84F75F" authorityName="Tshernova" authorityYear="1938" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="znojkoi" subGenus="Caucasiron">
+<emphasis id="E6540C73EC5C5D4101970D36C4BCF584" italics="true">E.</emphasis>
+(
+<emphasis id="A2B1C5AFADAD82FFC02F4EBF49F707F1" italics="true">C.</emphasis>
+) 
+<emphasis id="9C74E037A5E6CADCEC1A0B4E84DF6583" italics="true">znojkoi</emphasis>
+</taxonomicName>
+, 
+<taxonomicName id="7381C801738715E2F63288A5438B6F14" authority="(Braasch, 1979)" baseAuthorityName="Braasch" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="alpestris" subGenus="Caucasiron">
+<emphasis id="53F9913D8365CD8B0FE3C55304A99A0B" italics="true">E.</emphasis>
+(
+<emphasis id="034F6226902435C1D4F24F332BA0CDCF" italics="true">C.</emphasis>
+) 
+<emphasis id="E08B404B1FD8860DAFB2E141D0ED65E8" italics="true">alpestris</emphasis>
+(Braasch, 1979)
+</taxonomicName>
+, 
+<taxonomicName id="C18EFA9CC44C8EB5FBC261BB116DB8BB" baseAuthorityName="Braasch" baseAuthorityYear="1978" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="magnus" subGenus="Caucasiron">
+<emphasis id="F65F5DCED0E01DFBF567ED1B60CC44B3" italics="true">E.</emphasis>
+(
+<emphasis id="1CB76528A5C7ECF310A3B6F4ACF33C23" italics="true">C.</emphasis>
+) 
+<emphasis id="25A769CC08103960C438B287A8027A0B" italics="true">magnus</emphasis>
+</taxonomicName>
+.
+</paragraph>
+<caption id="09154D057F7F925696C50C8EAAAD852D" ID-DOI="10.3897/zookeys.1214.131266.figure1" ID-arpha="3C0F4EB0-E0CC-53EE-9DDF-07867558A812" httpUri="https://binary.pensoft.net/fig/1150504" startId="F1">
+<paragraph id="1311C9E219077BEBB02F5632FE3C7311">
+<label id="E6BBE125839AED402ADB063CC2AEFBE7">Figure 1.</label>
+</paragraph>
+<paragraph id="1808C5480D67836D805C65E08ADFCF4F">
+Distribution of 
+<taxonomicName id="2EE9F1F411F4B9AE5D4926C211B30026" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="D921AB770483A9AE8EFDB44F056AD766" italics="true">Epeorus</emphasis>
+(
+<emphasis id="73DE571606BE173A94FBBF39D95EF1DF" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="5E73E34968EDAB51C193425058AA4C02" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="87983AE766736480E27A827A0D13C71F" rank="species">sp. nov.</taxonomicNameLabel>
+; global localisation of a study area (lower left corner) and our sampling sites investigated between 2008–2019 (lower right corner).
+</paragraph>
+</caption>
+<caption id="3723BB96E94CD557B3C63B8B9BA6B727" ID-DOI="10.3897/zookeys.1214.131266.figure2" ID-arpha="A3B29615-0758-5503-95B1-456E459226DA" httpUri="https://binary.pensoft.net/fig/1150505" startId="F2">
+<paragraph id="F1F54F3C96640B6A1EDDFF186CF05B53">
+<label id="4067FD39CB219E2D78EBAC37DF3DA425">Figure 2.</label>
+</paragraph>
+<paragraph id="62FC1A6D4846EA1489DD8DB8A01D2478">
+<abbrev id="ABBRID0EMKAE" xlink_title="cytochrome oxidase subunit I">COI</abbrev>
+gene tree with results of molecular species delimitation tools and larval morphology according to 
+<bibRefCitation id="726DA3C0060EF1ED179F84D435433A25" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. (2020 b</bibRefCitation>
+). Black points correspond to posterior probability 1. Delimitation of 
+<taxonomicName id="E91C5D57ABAE3DCAAE1D88FF5842A843" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="422CA510C1442E558F0C0570F823EB39" italics="true">Epeorus</emphasis>
+(
+<emphasis id="4FF11EA2F7767B53BF448E7D2AFA529C" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="2684446D7F627151574E3B901A20F06E" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="7896085B3271BA3D1B68710F5C42AE80" rank="species">sp. nov.</taxonomicNameLabel>
+is highlighted by the dashed frame.
+</paragraph>
+</caption>
+<caption id="D1F24A560084A3EEE9D819DBEF6A405C" ID-DOI="10.3897/zookeys.1214.131266.figure3" ID-arpha="B49225C8-83E3-5375-941A-0142BCEBCE30" httpUri="https://binary.pensoft.net/fig/1150506" startId="F3">
+<paragraph id="E28F876D1F8F0F312A048D98828BB1BA">
+<label id="FAE7B39E08A91695BE84C836225119C7">Figure 3.</label>
+</paragraph>
+<paragraph id="FA9F062B00C1169E72D788AEBF19D331">
+The habitat (type locality) of 
+<taxonomicName id="FC0639064346C079339EC4BBBCC8A138" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="061C1E82FEC53F106F011BCAEB1CD7D4" italics="true">Epeorus</emphasis>
+(
+<emphasis id="2DE98B6986AF39AE076E687DD3E8BE86" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="711B144837A450719D8A4FB5630A75AE" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="457AA17BFAC8BDE5522A6415C60DF8A9" rank="species">sp. nov.</taxonomicNameLabel>
+from northeast Türkiye (photo: G. Türkmen).
+</paragraph>
+</caption>
+</subSubSection>
+<subSubSection id="SECID0ECMAE" type="description">
+<paragraph id="130BFDBFF78572B54E1F6E81A4553767">
+<heading id="0F650BCA52C121208086FEA100B5B6E1" reason="title">Description of larva.</heading>
+</paragraph>
+<paragraph id="EE2BF1931ECD7DAE02434F3937C74479">
+General colouration of larvae yellowish brown with dark brown maculation. Body length of late instar larvae: ca 
+<quantity id="D0B669949B6EFA1D1AD72F0391F4A57F" metricMagnitude="-2" metricUnit="m" metricValue="1.33" unit="mm" value="13.3">13.3 mm</quantity>
+(female), 
+<quantity id="7849D12E8391437AB14E38C58039E1C8" metricMagnitude="-2" metricUnit="m" metricValue="1.1175" metricValueMax="1.125" metricValueMin="1.1099999999999999" unit="mm" value="11.175" valueMax="11.25" valueMin="11.1">11.1–11.25 mm</quantity>
+(male). Length of cerci approximately 1.2 × body length.
+</paragraph>
+<paragraph id="E84CACC83E8ADAFCBC0396FBCE8E1C7F">
+<emphasis id="3C1395F5A8F54868C73559D7EDA79BF2" bold="true">
+<emphasis id="FFDAB513225E83AEE23B92327D4308EE" bold="true" italics="true">Head</emphasis>
+.
+</emphasis>
+Shape oval to trapezoidal. Anterior and lateral margin rounded, posterior margin rounded in female (Fig. 
+<figureCitation id="ACCF05BCF914F0B52784B5636A8C1605" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 E</figureCitation>
+) and slightly rounded in male (Fig. 
+<figureCitation id="1403F9ECB6689FCAD83F707D53BC54B9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 D</figureCitation>
+). Head dimensions of late instar larvae: length ca 
+<quantity id="E60EB0FF18F1E2F11331571B152F216C" metricMagnitude="-3" metricUnit="m" metricValue="4.5" unit="mm" value="4.5">4.5 mm</quantity>
+, width ca 
+<quantity id="33E2E2C750F487BDF4978ACBC61F2157" metricMagnitude="-3" metricUnit="m" metricValue="3.2" unit="mm" value="3.2">3.2 mm</quantity>
+in female; length ca 
+<quantity id="B531B2208679A299C9567A35C590519D" metricMagnitude="-3" metricUnit="m" metricValue="4.05" unit="mm" value="4.05">4.05 mm</quantity>
+, width 
+<quantity id="88BF492A265E0B3E8024C9DAC7E4601D" metricMagnitude="-3" metricUnit="m" metricValue="2.775" metricValueMax="2.8" metricValueMin="2.75" unit="mm" value="2.775" valueMax="2.8" valueMin="2.75">2.75–2.8 mm</quantity>
+in male. Head width / length ratio: 
+<quantity id="EE459858B39CE1B1EDC44DCEDAECBFC9" metricMagnitude="-2" metricUnit="m" metricValue="3.7338" metricValueMax="3.7592" metricValueMin="3.7084" unit="in" value="1.47" valueMax="1.48" valueMin="1.46">1.46–1.48 in</quantity>
+female; 
+<quantity id="7318E3C37B1B906500D53A968A0F5846" metricMagnitude="-2" metricUnit="m" metricValue="3.7718999999999996" metricValueMax="3.8354" metricValueMin="3.7084" unit="in" value="1.4849999999999999" valueMax="1.51" valueMin="1.46">1.46–1.51 in</quantity>
+male.
+</paragraph>
+<caption id="277D1F08D0F008F14468E5B2B88840A9" ID-DOI="10.3897/zookeys.1214.131266.figure4" ID-arpha="98D7E1D9-AF82-51C3-B486-D38DE4422DFF" httpUri="https://binary.pensoft.net/fig/1150507" startId="F4">
+<paragraph id="F388C62665EDF4211DA11E8A63E2FA81">
+<label id="4B3B157C6CD87C3814FBB8B555E06B44">Figure 4.</label>
+</paragraph>
+<paragraph id="53BF4B1FC94511C07F675E7C5629CF68">
+<taxonomicName id="7E9EAF8A1A478EF7EE51172EED49DB9A" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="B86606B4C64F19933A2357558E269728" italics="true">Epeorus</emphasis>
+(
+<emphasis id="9BABB56BFF3C89FC63834412FE1B2B5E" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="7525D93D43192B4992FED37E6051771D" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="C0D01C3B9797DCBD0A851C93D16357AB" rank="species">sp. nov.</taxonomicNameLabel>
+, larva 
+<emphasis id="B81C3BFE113BBC56DA722BA9F0C3DA28" bold="true">A</emphasis>
+habitus in dorsal view 
+<emphasis id="434C5819F50621B1C161445C233787B1" bold="true">B</emphasis>
+habitus in ventral view 
+<emphasis id="ACDE563F2F07F499A56F9AA91B881977" bold="true">C</emphasis>
+habitus in lateral view 
+<emphasis id="F7C7C5001D0548F299A528106305CAC1" bold="true">D</emphasis>
+head of male in dorsal view 
+<emphasis id="E561FBE3D036B7B84B39BCEEA0C6C346" bold="true">E</emphasis>
+head of female in dorsal view 
+<emphasis id="283F615C3DDEFEF782E912CA2DD1F209" bold="true">F</emphasis>
+middle leg in dorsal view 
+<emphasis id="8523BFF4D9C4CB2F0116855750422DCF" bold="true">G</emphasis>
+abdominal terga 
+<emphasis id="24CB8EE11069AB9B2E38456F3E6E42C2" bold="true">H, I</emphasis>
+abdominal sterna II – VI (arrow points on median maculation) 
+<emphasis id="1A1973DEC6BE3E0AFD5DF265A27A36DF" bold="true">J</emphasis>
+gills VII (in natural position from ventral view).
+</paragraph>
+</caption>
+<paragraph id="C9FF29C98FB0809CA5A79D79030C646D">
+Colouration of head: dorsal surface with pair of elongated maculae located along epicranial suture; pale stripes extending from lateral ocelli to lateral edges of head; blurred (or rectangular) macula between ocelli; rounded maculae anterolateral of lateral ocelli; blurred (or triangular) maculae near inner edges of compound eyes; pair of stripes (or scattered smaller maculae) located anteriorly from median ocellus (Fig. 
+<figureCitation id="219E6BC471B6FE566F63B202E4F373C6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 D, E</figureCitation>
+). Compound eyes grey to brownish to black. Ocelli blackish. Antennae yellowish brown, scapus and pedicellus darkened. Hair-like setae along anterior margin of head extend to lateral margins. Dorsal surface of head covered with fine hair-like setae and sparsely distributed stick-like setae. Sparse longer and fine hair-like setae located posteriorly to eyes.
+</paragraph>
+<paragraph id="A270A1FF7FCC855872958E92B4F5523E">
+<emphasis id="4622A57307A8BE61F285E43C871A5BA2" bold="true">
+<emphasis id="E90334557D98C862ADF36D26EF0C1596" bold="true" italics="true">Mouthparts</emphasis>
+.
+</emphasis>
+Labrum (Fig. 
+<figureCitation id="0BF12C56E9A0C92A4BF694BE4FB34BBC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 A</figureCitation>
+) widened anteriorly, with anterior margin slightly rounded or nearly straight (in dorsal view), lateral angles rounded. Dorsal surface (Fig. 
+<figureCitation id="9ECBCB403A0C6C54F85F9F4490121AC7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 A</figureCitation>
+, left) sparsely covered with hair-like setae and short bristle-like setae; 5–6 longer bristle-like setae located antero-medially and two antero-laterally. Epipharynx with longer, shortly plumose bristles situated along lateral to anterior margin (Fig. 
+<figureCitation id="339B6765643A74BA80E134AF4F99F19A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 A</figureCitation>
+, right; range of setation figured as large black dots), and brush of fine hair-like setae medially (not figured); ventral surface with group of 10–16 setae of various size located medio-posteriorly. Outer incisors of both mandibles with three apical teeth (Fig. 
+<figureCitation id="07960F0936AAD928E186A6E2449BE17F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 B, C</figureCitation>
+). Inner incisor of left mandible with three apical teeth, right inner incisor bifurcated. Outer edge of both mandibular incisors with numerous setae (range of setae marked with dashed polygons).
+</paragraph>
+<caption id="C942FCA54A0DE10E982D892A4B16ED92" ID-DOI="10.3897/zookeys.1214.131266.figure5" ID-arpha="2F8561FE-7515-511E-9ECE-71C6DDB4D704" httpUri="https://binary.pensoft.net/fig/1150508" startId="F5">
+<paragraph id="9A506B6BAF75445A34134D507B994A38">
+<label id="C66D9EFC0AD643ED0E1845E028CBF80F">Figure 5.</label>
+</paragraph>
+<paragraph id="DB9B7B597264D557940EE98F41C40D05">
+<taxonomicName id="DC25D68F6B1D2047265ED49BF3CD4AAB" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="9BF283CB9B1DC7900F68F72DE5BAD196" italics="true">Epeorus</emphasis>
+(
+<emphasis id="16146DBC22FF4CC6EBBE161D50BA1A8C" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="CD903E8E2A166CA69196025B6B7AFEED" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="857FAD589E510F2686E678672554CAB1" rank="species">sp. nov.</taxonomicNameLabel>
+, larva 
+<emphasis id="D987F34F962B51984E3D2AD841EB55B9" bold="true">A</emphasis>
+labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) 
+<emphasis id="4B70DE6BB982173E62B946FE899426D0" bold="true">B</emphasis>
+incisors of left mandible 
+<emphasis id="3F32BA8987B4B143ACEF33E76BBC2226" bold="true">C</emphasis>
+incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) 
+<emphasis id="8C60C0BFE50650029949650BDAE8719C" bold="true">D</emphasis>
+setae on dorsal surface of femora 
+<emphasis id="832E9172C3DA290E056FB76F7DD464A7" bold="true">E</emphasis>
+surface and posterior margin of abdominal tergum VII 
+<emphasis id="1647556D0C1B9091C5FBF74C5A2DABBC" bold="true">F</emphasis>
+gill I 
+<emphasis id="C1FFABF1BED0E8E0AE82E9D8799A5448" bold="true">G</emphasis>
+gill III (arrow points to projection on costal margin) 
+<emphasis id="A82BAB69B4679E7D5412F687337526A2" bold="true">H</emphasis>
+gill VII (flattened on slide) 
+<emphasis id="82C7B591F807DC0C7B4967816E3B8023" bold="true">I</emphasis>
+gill VII (in natural position from ventral view) 
+<emphasis id="D9D8A13E04B500A42146EA593AF52B62" bold="true">J, K</emphasis>
+abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) 
+<emphasis id="647BE1EC93524D9A7DCBF44FCE62D00B" bold="true">L</emphasis>
+sternum IX of female.
+</paragraph>
+</caption>
+<paragraph id="BA801B8D534B069333444BE25D42BC2D">
+<emphasis id="BAF688F2C29F0D56BD0C2A2E27C38000" bold="true">
+<emphasis id="C3E42232A67F5E508C0D4B08E7ADC449" bold="true" italics="true">Thorax</emphasis>
+.
+</emphasis>
+Pronotum anteriorly narrowed, lateral edges slightly curved. Metanotum with slight postero-medial projection. Dorsal surface with dark brown maculation as on Fig. 
+<figureCitation id="F1FD60994140EA96240AA4EF523FE360" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 A</figureCitation>
+and covered with fine, hair-like setae (as on abdominal terga and head); sparse longer, hair-like setae along pro-, meso- and metanotal suture.
+</paragraph>
+<paragraph id="BB9D367893D96A3449422D739481D060">
+<emphasis id="FFDFEFC1CA9E806EF4D144890C9E44E3" bold="true">
+<emphasis id="1E5D670B79742A83DEDEAD0829879120" bold="true" italics="true">Legs</emphasis>
+.
+</emphasis>
+Colour pattern of femora as on Fig. 
+<specimenCount id="103A093FC75EAD8F4FF6CDAE8656DB18" count="4" type="generic">
+<figureCitation id="7EEB11918CDD812FD42410BDB30BFF68" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 F</figureCitation>
+.
+</specimenCount>
+Femora without medial hypodermal spot. Femora apically slightly darkened; patella-tibial suture darkened; tarsi proximally and distally darkened. Dorsal surface of femora covered by short (sporadically elongated) apically rounded spatulate setae (Fig. 
+<figureCitation id="AB949B62FB7789B26FE8DB1A7A08B30B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 D</figureCitation>
+). Dorsal margin of tibiae and tarsi with row of long setae; ventral margin of both with short distally accumulated spine-like setae. Tarsal claws with 3–4 denticles.
+</paragraph>
+<paragraph id="8B45015B9852F9EB5D6927AB0F8255A9">
+<emphasis id="0EE2EE442DC0D379CD292954C03F592B" bold="true">
+<emphasis id="80FD4E90F25781265501E6B85EBF1696" bold="true" italics="true">Abdominal terga</emphasis>
+.
+</emphasis>
+Colour pattern of abdominal terga consists of transversal stripe along anterior margin of terga I – IX, medially extending to: i) large median triangular macula on terga II – III (IV), and ii) triangular or T-shaped macula on terga 
+<collectionCode id="11D564E817ED0EC4ECCF3A484943A2AF" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
+– IX (medial macula on tergum VIII and IX often widened). Median macula on terga 
+<collectionCode id="65998E19A4199B494F0C04C3240E8583" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
+– VII surrounded by pale area (Fig. 
+<figureCitation id="B720F90416CB1A2BB5AF65B0210A1D0B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 G</figureCitation>
+). Tergum X without distinct maculation. Lateral margin of terga I – IX with oblique macula. Denticles along posterior margin on terga strongly sclerotised, irregular and pointed (Fig. 
+<figureCitation id="EF33DFA92072E4651753A60A78B3F425" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 E</figureCitation>
+). Surface of terga covered with hair-like setae and sparsely with stick-like setae. Tergum X with short postero-lateral projections (Fig. 
+<figureCitation id="321DA54F1E916E989F04ADFAF6A9E510" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 J, K</figureCitation>
+). Longitudinal medial row of hair-like setae along abdominal terga present.
+</paragraph>
+<paragraph id="915C522FA92AAFC5093AFB3527482B33">
+<emphasis id="907136B1A9C67E539E87454FBC6D5876" bold="true">
+<emphasis id="8D8B89B32C92AECE5DC2150700E39F04" bold="true" italics="true">Abdominal sterna</emphasis>
+.
+</emphasis>
+Yellowish, with colouration pattern on sterna I – VIII consisting of rounded median macula (Fig. 
+<figureCitation id="864C4DC0499DAB97502F176202A1CC0E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 H</figureCitation>
+, arrow). In more pigmented specimens, median macula with paired pale spots located medio-posteriorly. Rounded median macula often poorly expressed and only medio-posterior edge of sterna is slightly pigmented (Fig. 
+<figureCitation id="B32EDF095510716638F8D62DF94958EE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 I</figureCitation>
+, arrow). Colouration pattern sporadically restricted to sterna I and II or absent. Sternum IX of female with V-shaped median emargination and surface covered by setae centrally (Fig. 
+<figureCitation id="216300B7319DE7E4E177DCEB7232E312" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 L</figureCitation>
+).
+</paragraph>
+<paragraph id="1137EDD3FBC9983813271B620D91063F">
+<emphasis id="885D560C5F8181A647EB59429F2337CC" bold="true">
+<emphasis id="E689608D4A4B5F43AED5D476C410CDEF" bold="true" italics="true">Gills</emphasis>
+.
+</emphasis>
+Dorsal surface of gill plate I yellowish; of gill plates II – VII greyish on anterior half, brownish on posterior half. Ventral margin of all gill plates yellowish. Costal projection on gill plate III well-developed (Fig. 
+<figureCitation id="12F0E847330385EA14096B4E0EF1EF38" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 G</figureCitation>
+, arrow). Gill plate VII wide (in natural position of ventral view; Figs 
+<figureCitation id="976FBD5D91B055E9EC8B1D3863E32629" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 J</figureCitation>
+, 
+<figureCitation id="47F24B4A68D20016ADA7FCC5BF229055" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 H, I</figureCitation>
+).
+</paragraph>
+<paragraph id="541EAA6DC54F44F2C6C5E5160F104506">
+<emphasis id="EC2210FC660651478105670BA308C974" bold="true">
+<emphasis id="0C3710DF88DEA5E7EBDB93150085F7F7" bold="true" italics="true">Cerci</emphasis>
+.
+</emphasis>
+Yellowish brown, basally darkened.
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EQTAE" type="Subimagoes, imagoes, and eggs">
+<paragraph id="8E9CDA3724739BE5B9F179D54F941EDF">
+<heading id="4B3C5143F4B42A342C4E717D8A8B39D1" reason="title">Subimagoes, imagoes, and eggs.</heading>
+</paragraph>
+<paragraph id="5689DA73DDEB1AC86A99D3CC8E165CCB">Unknown.</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EVTAE" type="Morphological diagnostics of larvae">
+<paragraph id="FDED629BCF4768003E9BEB10941E8A51">
+<heading id="DCD03E3235703147E0BEA32F3B1558B7" reason="title">Morphological diagnostics of larvae.</heading>
+</paragraph>
+<paragraph id="519F861DD8871F4907A3FC960C891D7A">
+<taxonomicName id="D7F6F46868DCC65F9AFA4DA14E1FC208" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="0EBB759FDA1934A2E1E26893D8DA4852" italics="true">Epeorus</emphasis>
+(
+<emphasis id="CC2291BAE7D7D8A5356B77F7113A8E27" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="58FCDCD90958FFE6E823AD27837BF553" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="260BAB76C8D2D40E62EA2C9212F501C4" rank="species">sp. nov.</taxonomicNameLabel>
+can be distinguished by the combination of the following morphological characters: i) femora without median spot (Fig. 
+<figureCitation id="CA0FA1B4DE4593452D075B7F32BBF59D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 F</figureCitation>
+); ii) abdominal sterna with circular median macula as on Fig. 
+<figureCitation id="F8A5354ED4E514D04088BBEACD674091" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 H</figureCitation>
+(colouration may be restricted to medio-posterior part of sterna as on Fig. 
+<figureCitation id="2A877EE9C8AF00290728F24B6FF12F36" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 I</figureCitation>
+); iii) abdominal terga 
+<collectionCode id="43A8C8122D4864F4E370856FB79367ED" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
+– VII with triangular or T-shaped macula surrounded by pale area (Fig. 
+<figureCitation id="CB89B3A24FD5A75EEDC29FBC5E9149AB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 G</figureCitation>
+); iv) tergum X with short postero-lateral projection (Fig. 
+<figureCitation id="2533E9C5BA09BC8CF018252743FCB383" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 J, K</figureCitation>
+, arrow); v) surface of abdominal terga with hair-like setae (Fig. 
+<figureCitation id="F229B981C05941BD5E4C1143E5D6874B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 E</figureCitation>
+); vi) shape of gill plates VII wide (in natural position from ventral view, Figs 
+<figureCitation id="88B45D2EA9042AD2381C978FA3E5CFEC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 J</figureCitation>
+; 
+<figureCitation id="942E54C6397A81E536BDE5A9ED53054A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 H, I</figureCitation>
+).
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EJVAE" type="Morphological affinities">
+<paragraph id="CD7EE124C29EFB39A486727641A23071">
+<heading id="BFE71BB705F883F43147C738BFADE187" reason="title">Morphological affinities.</heading>
+</paragraph>
+<paragraph id="FA531894CFC85E3231FB33183FB7170D">
+<taxonomicName id="1626D7B0B4929465D94EC9A8F68A2ED8" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="FB02DF3255BB2C7E63967648BC3AE1AC" italics="true">Epeorus</emphasis>
+(
+<emphasis id="D63363C440514D2EB71D0544D11A6390" italics="true">Caucasiron</emphasis>
+) 
+<emphasis id="A9BA108033598B38CD417DC4CF0409B1" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="413927101B4B00329B8EEF69ECF48B49" rank="species">sp. nov.</taxonomicNameLabel>
+is similar to several species from the Caucasus and neighbouring Mediterranean and Irano-Anatolian ranges, namely 
+<taxonomicName id="5F0954255B569FA6E9F319D04E4920CD" baseAuthorityName="Braasch" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="alpestris" subGenus="Caucasiron">
+<emphasis id="2E508CA2D54F51089F80DA1DC2F6466A" italics="true">E.</emphasis>
+(
+<emphasis id="B47E0112EB95B15BDC207362DD3B8FA0" italics="true">C.</emphasis>
+) 
+<emphasis id="ED7E1C7F63505F79ACA92AE128745DC9" italics="true">alpestris</emphasis>
+</taxonomicName>
+(distributed in the Greater Caucasus), 
+<taxonomicName id="2CD882649650B38356B628591455CFF0" authority="Hrivniak &amp; Sroka, 2020" authorityName="Hrivniak &amp; Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="alborzicus" subGenus="Caucasiron">
+<emphasis id="D0A90955B3C1C05EB7FFF5933B4AA079" italics="true">E.</emphasis>
+(
+<emphasis id="8680E76EBA0E2AA2008AC0059045F196" italics="true">C.</emphasis>
+) 
+<emphasis id="B874CD55F1C64C897F09B21CB5E13212" italics="true">alborzicus</emphasis>
+Hrivniak &amp; Sroka, 2020
+</taxonomicName>
+(
+<collectingRegion id="5BBE7FDC39E2B88CDC176FEEDFD5D652" country="Iran" name="Alborz">Alborz Mountains</collectingRegion>
+), and 
+<taxonomicName id="36D45E2AE684B95CBA24744401B5EDBF" authority="Hrivniak, 2017" authorityName="Hrivniak" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="bicolliculatus" subGenus="Caucasiron">
+<emphasis id="FB7379F847B21254DE9A84EBA6241047" italics="true">E.</emphasis>
+(
+<emphasis id="5149DD589903B3DC9EB3CD6AEB0C6F7D" italics="true">C.</emphasis>
+) 
+<emphasis id="7E4FDFE6FECAAEDB98F59CC8E8D4530E" italics="true">bicolliculatus</emphasis>
+<bibRefCitation id="D84EA9608967D097972118031599D9F1" DOI="10.11646/zootaxa.4341.3.2" author="Hrivniak Ľ &amp; Sroka P &amp; Godunko RJ &amp; Žurovcová M" journalOrPublisher="Zootaxa" pagination="353-374" refId="B12" refString="Hrivniak L, Sroka P, Godunko RJ, Zurovcova M (2017) Mayflies of the genus Epeorus Eaton, 1881 s. l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Turkiye. Zootaxa 4341: 353 - 374. https: // doi. org / 10.11646 / zootaxa. 4341.3. 2" title="Mayflies of the genus Epeorus Eaton, 1881 s. l. (Ephemeroptera: Heptageniidae) from the Caucasus Mountains: a new species of Caucasiron Kluge, 1997 from Georgia and Türkiye." volume="4341" year="2017">Hrivniak, 2017</bibRefCitation>
+</taxonomicName>
+(Pontic Mountains, Lesser and Greater Caucasus). All of them possess abdominal sterna with a rounded median macula and femora without median spot (
+<bibRefCitation id="B09DDF900FAF83B162DCDD397A9E7E9B" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+).
+</paragraph>
+<paragraph id="04E52F9E0BC785626DB5BA33D457FC72">
+<taxonomicName id="D8D9DF000C699022AE1A8F26D54CC90E" baseAuthorityName="Braasch" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="alpestris" subGenus="Caucasiron">
+<emphasis id="7D7D1A16033B60616B5A1F0205EA31E5" italics="true">Epeorus</emphasis>
+(
+<emphasis id="6421B0E75124444B6B98F3073D5693B8" italics="true">C.</emphasis>
+) 
+<emphasis id="BA6B8B3C0FB61FF59D761E11A9B55ACD" italics="true">alpestris</emphasis>
+</taxonomicName>
+can be distinguished from 
+<taxonomicName id="A9333000F3B61121B464A3E00BD679FE" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="E794332C4D98E3BB51068243E3812FC7" italics="true">E.</emphasis>
+(
+<emphasis id="E45F8E65069DBEABEDFC4B92510938B2" italics="true">C.</emphasis>
+) 
+<emphasis id="CBB2869E6DCDD60ECBCECE1C88372A11" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="C3D5CB3BFC98EC2E062D65A4BA117942" rank="species">sp. nov.</taxonomicNameLabel>
+by the absence of postero-lateral projections on the tergum X (
+<bibRefCitation id="280F0989C94C131437832D8E8916FA21" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 17 L) present in 
+<taxonomicName id="4767390B8D9605B9112C4E34B835401B" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="0B06BF15DA0D3D523A212CBD1DAD37C9" italics="true">E.</emphasis>
+(
+<emphasis id="F1B51E3566C605FA06D4E6440C74B4A2" italics="true">C.</emphasis>
+) 
+<emphasis id="29727CF68A5E880A7BBE122CA7E98D14" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="E5D6B99F80B358F7FB759F4F3109E3E7" rank="species">sp. nov.</taxonomicNameLabel>
+(Fig. 
+<figureCitation id="93B54E64223AE4933D81F6B8155D78DA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 J, K</figureCitation>
+, arrow). Additionally, the rounded maculae on abdominal sterna are always present in 
+<taxonomicName id="5AD5A9AC0D7812ABD489BF25D591E93D" baseAuthorityName="Braasch" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="alpestris" subGenus="Caucasiron">
+<emphasis id="4BBC591686A161661DF9E9A2FDB59B37" italics="true">E.</emphasis>
+(
+<emphasis id="012B6DD43BC6053AA3749F9ADC259960" italics="true">C.</emphasis>
+) 
+<emphasis id="DC4858447D830E0962B53542C219E2E2" italics="true">alpestris</emphasis>
+</taxonomicName>
+(
+<bibRefCitation id="AEAFC17CF9CB40BC8BB046696B3B1370" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 16 I), whereas the colouration pattern of abdominal sterna varies from a well-defined pattern (Fig. 
+<figureCitation id="7EA360E5430AC7B801749DE7C47733BF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 H</figureCitation>
+) to an indistinct (Fig. 
+<figureCitation id="24BE36B367E45C4D862515053E68E3BD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 I</figureCitation>
+) or no pattern in 
+<taxonomicName id="07C36C8FE050110BE73E8054065EE759" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="E06916EA3BFC91D1994554E77725B296" italics="true">E.</emphasis>
+(
+<emphasis id="B203DA683E01DD75E4066C0538560E15" italics="true">C.</emphasis>
+) 
+<emphasis id="0B35750E11D050541C5EE59738A13123" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="EA266FC5FC8A5EB56C72699C38D882D3" rank="species">sp. nov.</taxonomicNameLabel>
+Moreover, 
+<taxonomicName id="51EF78ADB9C8D96F30F161865FA16867" baseAuthorityName="Braasch" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="alpestris" subGenus="Caucasiron">
+<emphasis id="006A5985E5AE523DE01DA408195BA079" italics="true">Epeorus</emphasis>
+(
+<emphasis id="03166BE720A78EB1D8DE275B89C2324F" italics="true">C.</emphasis>
+) 
+<emphasis id="F1D46D2F065F6017657E405C366E72E1" italics="true">alpestris</emphasis>
+</taxonomicName>
+is characterised by typical maculation of abdominal terga (
+<bibRefCitation id="DF91B526E5B41982C87C14C6FC168886" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 16 G, H).
+</paragraph>
+<paragraph id="3EC0746CCAA6646552CDEF86DE018D5B">
+<taxonomicName id="68B670BF04BEBF759EBFAABD86E22687" authorityName="Hrivniak &amp; Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="alborzicus" subGenus="Caucasiron">
+<emphasis id="5E0CD262BF38105E8345C5AD9D00AE20" italics="true">Epeorus</emphasis>
+(
+<emphasis id="4BA03725A9831E4CCD6BE1196F0753F0" italics="true">C.</emphasis>
+) 
+<emphasis id="E7CAE8617B37B3844EA6665617630AA1" italics="true">alborzicus</emphasis>
+</taxonomicName>
+possesses abdominal sterna with a large circular medial macula (
+<bibRefCitation id="7DF3FCBB7DC568F1C74AE4F5C4939587" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 40 L – N) and blurred macula (or a pair of rounded maculae) on tergum II and III (
+<bibRefCitation id="F2CC5652149D0E7872BC7EC3BAFE630F" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 40 H, I), in contrast to 
+<taxonomicName id="50783DD95B10330EF8AE0A41E7478C34" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="926A3F90D83F4ABACA2D97D22588FC18" italics="true">E.</emphasis>
+(
+<emphasis id="8BEF57D57C238672246BF1356CDBF970" italics="true">C.</emphasis>
+) 
+<emphasis id="1FDEB1F8501AB88A894856AFDF5FB0A7" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="6277A2845A11C2B81DD989186398486E" rank="species">sp. nov.</taxonomicNameLabel>
+with relatively small rounded medial macula on abdominal sterna (Fig. 
+<figureCitation id="2179C338F04D2E5D8B94E8FF2531CE90" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 H, I</figureCitation>
+) and triangular macula on tergum II and III (Fig. 
+<figureCitation id="4908B94105ECB47F3E922A0DBB6AE4CA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Epeorus (Caucasiron) abditus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G abdominal terga H, I abdominal sterna II – VI (arrow points on median maculation) J gills VII (in natural position from ventral view)." figureDoi="10.3897/zookeys.1214.131266.figure4" httpUri="https://binary.pensoft.net/fig/1150507">4 G</figureCitation>
+).
+</paragraph>
+<paragraph id="24D14FB1A9D2ADE7BDE67979297BAFFC">
+<taxonomicName id="1F084DA6585A55CE7C742C0817C27105" authorityName="Hrivniak" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="bicolliculatus" subGenus="Caucasiron">
+<emphasis id="682C01A48585D5099A33B3CCAD69E9F2" italics="true">Epeorus</emphasis>
+(
+<emphasis id="452A48708568BA973B8D2D406196DB37" italics="true">C.</emphasis>
+) 
+<emphasis id="8386BFB3DD72960EC0052F41D88670F1" italics="true">bicolliculatus</emphasis>
+</taxonomicName>
+differs from 
+<taxonomicName id="D8C4BF05A1EB72D2529A8770C1A9524E" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="0B0DA38690E9F11536F37705D00FAE3E" italics="true">E.</emphasis>
+(
+<emphasis id="FA7CBA5F589A47E47ED2BC493D0ACA86" italics="true">C.</emphasis>
+) 
+<emphasis id="1D869BC88642476A51453D6A290CD6B5" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="35BD54A255F030BA650FDD4F58AB4313" rank="species">sp. nov.</taxonomicNameLabel>
+by the presence of paired postero-medial protuberances on abdominal terga II – IX (
+<bibRefCitation id="A935E01F4AFBFC31D807B33A2A7E8FC8" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 34 H) and basally widened setae on the surface of terga (
+<bibRefCitation id="0178195DFA2A9681F4BE0ACAEA0D90CE" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 35 E) in contrast to basally narrow setae in 
+<taxonomicName id="2FE0FB0D32B941659C650FC0D7D7F65E" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="7BDEC632B5A9FDA6193E5B45BBEBAC79" italics="true">E.</emphasis>
+(
+<emphasis id="522F245BBD9F54F2D57E2D3C11CF8F8F" italics="true">C.</emphasis>
+) 
+<emphasis id="5EFC0AE2D2647EDE382815D0753C4AC2" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="DC8D41D4B04CACDC10A82E8220EDB73B" rank="species">sp. nov.</taxonomicNameLabel>
+(Fig. 
+<figureCitation id="4361B47B213E1C7906B1D34CE2F3B10E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 E</figureCitation>
+).
+</paragraph>
+<paragraph id="90D15EEF37CF61FC843065702C7CB47C">
+The larvae of 
+<taxonomicName id="1DFBFAEDCBE1FF31AEAD779F0561BF04" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="4AC35359D43F3934D5D88FA7FB3021DA" italics="true">E.</emphasis>
+(
+<emphasis id="D0464D7DB90139B28F53FA23F6A9E3D3" italics="true">C.</emphasis>
+) 
+<emphasis id="F35163FA859BB8321028B3DBD444C9FC" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="41AEDC5AE6B10DD238EEBED725AC9D13" rank="species">sp. nov.</taxonomicNameLabel>
+with weakly pigmented abdominal sterna may be erroneously assigned to 
+<taxonomicName id="751DBF41DE16161652760C561CCDF4CD" baseAuthorityName="Braasch" baseAuthorityYear="1978" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="magnus" subGenus="Caucasiron">
+<emphasis id="234D91B93437A4CD147EA4693295F6F8" italics="true">E.</emphasis>
+(
+<emphasis id="7DBE806E822E99EF0C0A0022BF1F62BC" italics="true">C.</emphasis>
+) 
+<emphasis id="C50A95623F9897D4D2B2A8CC67573AC4" italics="true">magnus</emphasis>
+</taxonomicName>
+(distributed in the Greater and Lesser Caucasus, Pontic and Taurus Mountains). This species differs from 
+<taxonomicName id="2B5EBF04E56F4E7C8A0850F58BF12C6D" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="6881CEABAF6F1F651F97EE462802316D" italics="true">E.</emphasis>
+(
+<emphasis id="FCEC16BD18DE962B3B005C0E1F9F3E68" italics="true">C.</emphasis>
+) 
+<emphasis id="EF3C585894973ED6016237291165FDCA" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="E54B7CC89517FB849B11E5ECE8C2BCE3" rank="species">sp. nov.</taxonomicNameLabel>
+by the presence of dense bristle-like setae on the dorsal surface of the labrum (
+<bibRefCitation id="A29ADDB42BAEB4BA41A9D30841780B8D" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 11 A) in contrast to sparse and hair-like setae in 
+<taxonomicName id="37A3144FACACABC0BD00698E14035306" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="A4D558D1BAB2F3685EB60B85D6DDC986" italics="true">E.</emphasis>
+(
+<emphasis id="D3CB8BF24E73B7108D570621037E94A6" italics="true">C.</emphasis>
+) 
+<emphasis id="DB32F01ABF1F8CEC62A667BBCB987FA1" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="9206AA5C15C3B445BB452A4FFF55431E" rank="species">sp. nov.</taxonomicNameLabel>
+(Fig. 
+<figureCitation id="E49C821893AA3AE4AE9E7273EE4AE1CD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 A</figureCitation>
+). In addition, 
+<taxonomicName id="78A18E349F5125266B69726AF8CF79A0" baseAuthorityName="Braasch" baseAuthorityYear="1978" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="magnus" subGenus="Caucasiron">
+<emphasis id="778BF706E01A321602AA3DC53621D521" italics="true">E.</emphasis>
+(
+<emphasis id="DD45A8D964259E200F1485513A7D1276" italics="true">C.</emphasis>
+) 
+<emphasis id="466D25181292F757FCB509919B1E1CA8" italics="true">magnus</emphasis>
+</taxonomicName>
+often has clearly developed lateral projections on the tergum X (
+<bibRefCitation id="4A1BFC6CFAC077E8796D8306ADEED098" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 11 K – M) in contrast to only short projections in 
+<taxonomicName id="44A054CDC6FB78CFF6DCAADF7CDF7F99" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="0CCDEEF4571940434D9154D0C4D5C25A" italics="true">E.</emphasis>
+(
+<emphasis id="6CC4553EA5AAA0961A54F026FAEE736B" italics="true">C.</emphasis>
+) 
+<emphasis id="F81A998DCD53B07A5CDC3F85559D24B4" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="E018C0A456910E89A6B60A3AF180EE16" rank="species">sp. nov.</taxonomicNameLabel>
+(Fig. 
+<figureCitation id="B82B23E2F3BF0FA0A855BA43CF5E9523" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 J, K</figureCitation>
+).
+</paragraph>
+<paragraph id="9B3B5B1E200ADDA36032F05945CBC571">
+Two species distributed in the western and central Greater Caucasus, namely 
+<taxonomicName id="A2BE127B3796489697BA6E883B5B80BB" authority="(Braasch, 1979)" baseAuthorityName="Braasch" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="soldani" subGenus="Caucasiron">
+<emphasis id="BEA76F816AFAD4BECF2D13130437F277" italics="true">E.</emphasis>
+(
+<emphasis id="98E89777C62468C91FB15CF63D91F732" italics="true">C.</emphasis>
+) 
+<emphasis id="52B0EE25DF04349B2D35188234B2901A" italics="true">soldani</emphasis>
+(Braasch, 1979)
+</taxonomicName>
+and 
+<taxonomicName id="DB3ACD77A369CD4912A930474CFD880D" authority="(Braasch &amp; Zimmerman, 1979)" baseAuthorityName="Braasch &amp; Zimmerman" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="sinitshenkovae" subGenus="Caucasiron">
+<emphasis id="DD19C865E0B7005BA78BACC042A952A6" italics="true">E.</emphasis>
+(
+<emphasis id="C2290BF849D9A4DE769CC3FE94A2BF75" italics="true">C.</emphasis>
+) 
+<emphasis id="B61E818B3AEB62361AD6F35DA68EECD2" italics="true">sinitshenkovae</emphasis>
+(Braasch &amp; Zimmerman, 1979)
+</taxonomicName>
+, have abdominal sterna without or with weakly developed colouration pattern and no femoral spot. 
+<taxonomicName id="D99D2D8E48964CF3DF719CB7A6D2A32F" baseAuthorityName="Braasch" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="soldani" subGenus="Caucasiron">
+<emphasis id="D26E184EB146FDD9574F0662F0AF571F" italics="true">Epeorus</emphasis>
+(
+<emphasis id="96E9369E1ADEF5B650AE5E006DB84314" italics="true">C.</emphasis>
+) 
+<emphasis id="06D5B60211D15551DF17C883B73D2011" italics="true">soldani</emphasis>
+</taxonomicName>
+can also be easily distinguished from 
+<taxonomicName id="739A31B80997F2AA8043756D5C70A476" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="62DB23A1CC0B12DD0219F37288DD1916" italics="true">E.</emphasis>
+(
+<emphasis id="4015BC1E823CC9DE4BDAE67C571CF7E3" italics="true">C.</emphasis>
+) 
+<emphasis id="652350E1F8D293E8EE6DBB0456E1336F" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="21C392D5D6ED8657DFB1C306CE95F885" rank="species">sp. nov.</taxonomicNameLabel>
+by setae on abdominal terga that are basally widened in the former species (
+<bibRefCitation id="AA72FE852E4C56B6243D9C9124DDD209" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 20 E) and narrow in the latter (Fig. 
+<figureCitation id="3E3EFD0426C9AA5595751865BBED5B1E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 E</figureCitation>
+). 
+<taxonomicName id="656F74910F321882F0169780420CA7D1" baseAuthorityName="Braasch &amp; Zimmerman" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="sinitshenkovae" subGenus="Caucasiron">
+<emphasis id="936FD9686EFD2CEBCF7CFF19EB711DE8" italics="true">Epeorus</emphasis>
+(
+<emphasis id="8E573D43E4DE2A1C1E17E1C589C932CB" italics="true">C.</emphasis>
+) 
+<emphasis id="3B71F907DB5FC32C202E57B6677170B2" italics="true">sinitshenkovae</emphasis>
+</taxonomicName>
+can be separated by a poorly developed projection on costal margin of gill plates (
+<bibRefCitation id="85DFC2FF0EAE194D403DD713A4894101" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 26 G) from 
+<taxonomicName id="BC1BA250ACE5F7E67A81560047FAA12D" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="30AA3DFD07CF4A7DB6A71F922CCA8DBA" italics="true">E.</emphasis>
+(
+<emphasis id="B97E4E5E72537099BBE925147FA65700" italics="true">C.</emphasis>
+) 
+<emphasis id="4A6F48862BC79BA5570071C42717439F" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="12DFFF9232B4B95579ADF824AFFB8B4E" rank="species">sp. nov.</taxonomicNameLabel>
+bearing a well-developed projection (Fig. 
+<figureCitation id="8F6FD986A39096B4D258F3B266520A88" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epeorus (Caucasiron) abditus sp. nov., larva A labrum (left half in dorsal view, right half in ventral view; black dots refer to range of setae) B incisors of left mandible C incisors of right mandible (dashed polygons on outer edge of both mandibles refer to range of setae) D setae on dorsal surface of femora E surface and posterior margin of abdominal tergum VII F gill I G gill III (arrow points to projection on costal margin) H gill VII (flattened on slide) I gill VII (in natural position from ventral view) J, K abdominal segments VIII – X in lateral view (arrow points on postero-lateral projection) L sternum IX of female." figureDoi="10.3897/zookeys.1214.131266.figure5" httpUri="https://binary.pensoft.net/fig/1150508">5 G</figureCitation>
+). Additionally, 
+<taxonomicName id="842DF879BB73744C768BF9D9A0083BEF" baseAuthorityName="Braasch &amp; Zimmerman" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="sinitshenkovae" subGenus="Caucasiron">
+<emphasis id="B62F1FD65F02DFFCAFB557758CA9F322" italics="true">E.</emphasis>
+(
+<emphasis id="BE694C5B3264F5CEF7152878F9D0F273" italics="true">C.</emphasis>
+) 
+<emphasis id="F608C733FF96B2604E6F36CC704F1A63" italics="true">sinitshenkovae</emphasis>
+</taxonomicName>
+is characterised by a specific colouration of abdominal terga and femora (
+<bibRefCitation id="014AA89FD160BA45BD4D998CA334E987" DOI="10.3897/zookeys.986.56276" author="Hrivniak Ľ &amp; Sroka P &amp; Bojková J &amp; Godunko RJ" journalOrPublisher="ZooKeys" pagination="1-53" refId="B16" refString="Hrivniak Ľ, Sroka P, Bojková J, Godunko RJ (2020 b) Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae). ZooKeys 986: 1–53. https://doi.org/10.3897/zookeys.986.56276" title="Identification guide to larvae of Caucasian Epeorus (Caucasiron) (Ephemeroptera, Heptageniidae)." volume="986">Hrivniak et al. 2020 b</bibRefCitation>
+, fig. 25 H, F).
+</paragraph>
+<paragraph id="46DAAE84A6603C8B343CCA4A5FF9C35B">
+All other species of 
+<emphasis id="140DD2E42D52AEBCE416FDEE442FAA44" italics="true">E.</emphasis>
+(
+<taxonomicName id="85A071B3E8FEF62CBEF714236456AB58" authorityName="Kluge" authorityYear="1997" rank="subGenus" subGenus="Caucasiron">
+<emphasis id="3CBB53EAC7AC61C48C10B18F3042DAE4" italics="true">Caucasiron</emphasis>
+</taxonomicName>
+) from the Caucasus, Mediterranean, and Irano-Anatolian ranges can be easily distinguished from 
+<taxonomicName id="B45E2A85179395CD83AA1299CD8508EE" authorityName="Hrivniak &amp; Sroka &amp; Türkmen &amp; Martynov &amp; Bojková" authorityYear="2024" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="abditus" status="sp. nov." subGenus="Caucasiron">
+<emphasis id="643DBF0E71EC2448E4207EC1F46799EC" italics="true">E.</emphasis>
+(
+<emphasis id="450E638DAC59A481340EABADB14EB47F" italics="true">C.</emphasis>
+) 
+<emphasis id="954D130A64F03469244213A7B65535F4" italics="true">abditus</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="29C6052421EAFFF1059BD7B9AA6C77CC" rank="species">sp. nov.</taxonomicNameLabel>
+by the presence of specific colouration pattern of abdominal sterna and / or presence of femoral spot. These include 
+<taxonomicName id="9651B18CF01D7E3C2051A32F058C5756" authority="(Tshernova, 1938)" baseAuthorityName="Tshernova" baseAuthorityYear="1938" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="caucasicus" subGenus="Caucasiron">
+<emphasis id="07D8E23D2D77B0F6ACF7F354BF223D82" italics="true">E.</emphasis>
+(
+<emphasis id="69F885831BA960770FF774C4C5B12161" italics="true">C.</emphasis>
+) 
+<emphasis id="925DD0B279C99307309F315B59587BF4" italics="true">caucasicus</emphasis>
+(
+<bibRefCitation id="284A32F4DBA03D15E594AE928F841FBE" author="Tshernova OA" issue="42" journalOrPublisher="Trudy Azerbajdshanskogo Filiala AN SSSR, Baku" pagination="55-64" refId="B31" refString="Tshernova OA (1938) Zur Kenntnis der Ephemeropteren Ost-Transkaukasien. Trudy Azerbajdshanskogo Filiala AN SSSR, Baku 7 (42): 55 - 64." title="Zur Kenntnis der Ephemeropteren Ost-Transkaukasien." volume="7" year="1938">Tshernova, 1938</bibRefCitation>
+)
+</taxonomicName>
+, 
+<taxonomicName id="C430643F0E5931F50B909D37CCCB1A3D" authority="(Sinitshenkova, 1976)" baseAuthorityName="Sinitshenkova" baseAuthorityYear="1976" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="nigripilosus" subGenus="Caucasiron">
+<emphasis id="AA48A1170E430BB13BAB98FB32B0FCD4" italics="true">E.</emphasis>
+(
+<emphasis id="FE9F6A8B79FB41F675C9070F51743950" italics="true">C.</emphasis>
+) 
+<emphasis id="6703DAE2312A23913C1D0055C85F69AB" italics="true">nigripilosus</emphasis>
+(
+<bibRefCitation id="C2CA3E056373AFD6779ACEC3C18F20B4" author="Sinitshenkova ND" issue="4" journalOrPublisher="Entomological Review" pagination="85-92" refId="B30" refString="Sinitshenkova ND (1976) Mayflies of the genus Iron Eaton (Ephemeroptera, Heptageniidae) in the fauna of the Caucasus. Entomological Review 55 (4): 85 - 92." title="Mayflies of the genus Iron Eaton (Ephemeroptera, Heptageniidae) in the fauna of the Caucasus." volume="55" year="1976">Sinitshenkova, 1976</bibRefCitation>
+)
+</taxonomicName>
+, 
+<taxonomicName id="ACFE4C063C741752BBEBC4C4E6D5E978" authority="Hrivniak &amp; Sroka, 2020" authorityName="Hrivniak &amp; Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="zagrosicus" subGenus="Caucasiron">
+<emphasis id="EE1FBE4448238A51F79B8BFABD9BA224" italics="true">E.</emphasis>
+(
+<emphasis id="4F623925642A08736DF332A2C207A6E7" italics="true">C.</emphasis>
+) 
+<emphasis id="E582F6F6E619024AFC5F71F17C26C831" italics="true">zagrosicus</emphasis>
+Hrivniak &amp; Sroka, 2020
+</taxonomicName>
+, 
+<taxonomicName id="F8C6605A8BE87F98178C3BE0D17403DF" authority="(Braasch &amp; Soldan, 1979)" baseAuthorityName="Braasch &amp; Soldan" baseAuthorityYear="1979" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Caucasiron">
+<emphasis id="5E3574C12C0E5EF011E3C437FB7B4138" italics="true">E.</emphasis>
+(
+<emphasis id="4B8BA2BC383A3362E0B5EBB3223A7B29" italics="true">C.</emphasis>
+) 
+<emphasis id="95B746E450F6F086DD59E0AB3B92F018" italics="true">iranicus</emphasis>
+(Braasch &amp; Soldán, 1979)
+</taxonomicName>
+, 
+<taxonomicName id="A67F632406C83C914B300D5127B407BE" authority="(Braasch, 1980)" baseAuthorityName="Braasch" baseAuthorityYear="1980" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="longimaculatus" subGenus="Caucasiron">
+<emphasis id="DF2119D5E5F1D591321E6D6F25DAB232" italics="true">E.</emphasis>
+(
+<emphasis id="6422B97F164D796F1C3D468D77E33B37" italics="true">C.</emphasis>
+) 
+<emphasis id="ADEC4D19265E361F93B9EFECD0D21618" italics="true">longimaculatus</emphasis>
+(
+<bibRefCitation id="C08DC190A064C6913EB064842B3F3B0C" author="Braasch D" issue="10-11" journalOrPublisher="Entomologische Nachrichten" pagination="166-173" refId="B5" refString="Braasch D (1980) Beitrag zur Kenntnis der Gattung Iron Eaton (Heptageniidae, Ephemeroptera) im Kaukasus (UdSSR), 2. Entomologische Nachrichten 24 (10 - 11): 166 - 173." title="Beitrag zur Kenntnis der Gattung Iron Eaton (Heptageniidae, Ephemeroptera) im Kaukasus (UdSSR), 2." volume="24" year="1980">Braasch, 1980</bibRefCitation>
+)
+</taxonomicName>
+, 
+<taxonomicName id="D08A20C1E7DA85F95A6638E571C13F7F" authority="Hrivniak, Turkmen &amp; Kazanci, 2019" authorityName="Hrivniak, Turkmen &amp; Kazanci" authorityYear="2019" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="turcicus" subGenus="Caucasiron">
+<emphasis id="D30FDA24FDF09ACFD3CFEA6550444B2A" italics="true">E.</emphasis>
+(
+<emphasis id="0BFFC000653BD0BB0EA421F81E70925E" italics="true">C.</emphasis>
+) 
+<emphasis id="1DD5A113A6035E6283FB203C4D2A0AD4" italics="true">turcicus</emphasis>
+<bibRefCitation id="36E532455C6E3D30A35C6EB1B152A002" DOI="10.11646/zootaxa.4550.1.2" author="Hrivniak Ľ &amp; Sroka P &amp; Türkmen G &amp; Godunko RJ &amp; Kazancı N" journalOrPublisher="Zootaxa" pagination="58-70" refId="B14" refString="Hrivniak L, Sroka P, Turkmen G, Godunko RJ, Kazanci N (2019) A new Epeorus (Caucasiron) (Ephemeroptera: Heptageniidae) species from Turkey based on molecular and morphological evidence. Zootaxa 4550: 58 - 70. https: // doi. org / 10.11646 / zootaxa. 4550.1. 2" title="A new Epeorus (Caucasiron) (Ephemeroptera: Heptageniidae) species from Turkey based on molecular and morphological evidence." volume="4550" year="2019">Hrivniak, Türkmen &amp; Kazancı, 2019</bibRefCitation>
+</taxonomicName>
+, 
+<taxonomicName id="2B2C1B928231AE90E4B5AE197D8D78B8" authorityName="Hrivniak &amp; Sroka" authorityYear="2020" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="shargi" subGenus="Caucasiron">
+<emphasis id="DFB6749F9C79FF1CC9876F64ECBE376C" italics="true">E.</emphasis>
+(
+<emphasis id="4629CCB3D596B8D5DC0851D881BA95CA" italics="true">C.</emphasis>
+) 
+<emphasis id="09E172D3AE063DFC103E633D224CD057" italics="true">shargi</emphasis>
+</taxonomicName>
+, 
+<taxonomicName id="C82DE781041A2BA35FB40D49301081A8" authorityName="Hrivniak &amp; Sroka" authorityYear="2021" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="hyrcanicus" subGenus="Caucasiron">
+<emphasis id="3E65AE9082D5425087377B2844CC1341" italics="true">E.</emphasis>
+(
+<emphasis id="D10C7FF3E27D091E8FDFDB8DD75CCAA7" italics="true">C.</emphasis>
+) 
+<emphasis id="828350E2A9A232DA43917DC307AD726A" italics="true">hyrcanicus</emphasis>
+</taxonomicName>
+, and 
+<taxonomicName id="5928AE2E2DC996DCB945F83FF4CF25E2" authorityName="Hrivniak &amp; Sroka" authorityYear="2022" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tripertitus" subGenus="Caucasiron">
+<emphasis id="E59EE0A2BEFB2758CED7E56085BAF712" italics="true">E.</emphasis>
+(
+<emphasis id="028B3E5A6A787138C9EB7FE67946C2C4" italics="true">C.</emphasis>
+) 
+<emphasis id="85D43524AE2E5DD3907418AD61A81327" italics="true">tripertitus</emphasis>
+</taxonomicName>
+.
+</paragraph>
+</subSubSection>
+</treatment>
+</document>
\ No newline at end of file