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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.999.53130" ID-GBIF-Dataset="a4c2ed66-477c-4416-9942-dd8111a2eb22" ID-PMC="PMC7723882" ID-Pensoft-Pub="1313-2970-999-109" ID-Pensoft-UUID="833665678A4F5EEF87330B7783687C86" ID-PubMed="33343214" ID-ZooBank="B8D6EF7CBFED4758A1EBA58F1711BD30" ModsDocID="1313-2970-999-109" checkinTime="1607107203662" checkinUser="pensoft" docAuthor="Prandi, Massimo, Grossi, Paschoal C. &amp; Vaz-De-Mello, Fernando Z." docDate="2020" docId="E6A84C6B50FC55C299456A179087C179" docLanguage="en" docName="ZooKeys 999: 109-145" docOrigin="ZooKeys 999" docSource="http://dx.doi.org/10.3897/zookeys.999.53130" docTitle="Megasoma (Lycophontes) joergenseni subsp. joergenseni joergenseni (Bruch 1910" docType="treatment" docVersion="4" id="833665678A4F5EEF87330B7783687C86" lastPageNumber="109" masterDocId="833665678A4F5EEF87330B7783687C86" masterDocTitle="Revision of the Megasoma (Megasoma) gyas (Jablonsky in Herbst, 1785) species group (Coleoptera, Scarabaeidae, Dynastinae)" masterLastPageNumber="145" masterPageNumber="109" pageNumber="109" updateTime="1668169988366" updateUser="ExternalLinkService">
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<mods:title>Revision of the Megasoma (Megasoma) gyas (Jablonsky in Herbst, 1785) species group (Coleoptera, Scarabaeidae, Dynastinae)</mods:title>
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<mods:namePart>Prandi, Massimo</mods:namePart>
<mods:affiliation>Via Del Seminario 16, I- 25087, Salo, BS, Italy</mods:affiliation>
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<mods:nameIdentifier type="email">prandi.m@libero.it</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Grossi, Paschoal C.</mods:namePart>
<mods:affiliation>Universidade Federal Rural de Pernambuco, Departamento de Agronomia / Fitossanidade, Laboratorio de Taxonomia de Insetos, Programa de Pos-graduacao em Entomologia Agricola-PPGEA, 52171 - 900, Recife, Pernambuco, Brazil</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vaz-De-Mello, Fernando Z.</mods:namePart>
<mods:affiliation>Universidade Federal de Mato Grosso, Instituto de Biociencias, Departamento de Biologia e Zoologia, Colecao Entomologica, 78060 - 900, Cuiaba, Mato Grosso, Brazil</mods:affiliation>
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<mods:number>999</mods:number>
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<taxonomicName LSID="E6A84C6B-50FC-55C2-9945-6A179087C179" authority="joergenseni (Bruch, 1910)" authorityName="joergenseni (Bruch" authorityYear="1910" baseAuthorityName="Bruch" baseAuthorityYear="1910" class="Insecta" family="Dynastidae" genus="Megasoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megasoma (Lycophontes) joergenseni subsp. joergenseni" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="joergenseni" subGenus="Lycophontes" subSpecies="joergenseni">Megasoma (Lycophontes) joergenseni joergenseni (Bruch, 1910)</taxonomicName>
<figureCitation captionStart="Figure 26" captionStartId="F20" captionText="Figure 26. A M. joergenseni joergenseni ♂ B M. joergenseni joergenseni ♀." figureDoi="10.3897/zookeys.999.53130.figure26" httpUri="https://binary.pensoft.net/fig/481450" pageId="0" pageNumber="109">Figure 26</figureCitation>
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<taxonomicName authorityName="Bruch" authorityYear="1910" class="Insecta" family="Dynastidae" genus="Lycophontes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycophontes" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">Lycophontes</taxonomicName>
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Bruch, 1910: 74.
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<paragraph pageId="0" pageNumber="109">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="109">
The subgenus
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, of the genus
<taxonomicName class="Insecta" family="Dynastidae" genus="Megasoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megasoma" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Megasoma</emphasis>
</taxonomicName>
, is a group comprised of small-sized taxa.
<taxonomicName authority="jorgenseni" class="Insecta" family="Dynastidae" genus="Megasoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megasoma jorgenseni subsp. jorgenseni" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="subSpecies" species="jorgenseni" subSpecies="jorgenseni">
<emphasis italics="true" pageId="0" pageNumber="109">Megasoma jorgenseni jorgenseni</emphasis>
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is a completely-pubescent taxon, occurring in central-northern Argentina and southeastern Bolivia. The total body length varies from 30 to 40 mm (Fig.
<figureCitation captionStart="Figure 26" captionStartId="F20" captionText="Figure 26. A M. joergenseni joergenseni ♂ B M. joergenseni joergenseni ♀." figureDoi="10.3897/zookeys.999.53130.figure26" httpUri="https://binary.pensoft.net/fig/481450" pageId="0" pageNumber="109">26</figureCitation>
, courtesy of Mushi-sha). The original type of Bruch (San Luis, Mendoza) considered lost, was recently found by the Argentinean entomologist FC Penco in a private collection, and is now deposited in the UNLP collection.
</paragraph>
<caption doi="10.3897/zookeys.999.53130.figure26" httpUri="https://binary.pensoft.net/fig/481450" pageId="0" pageNumber="109" start="Figure 26" startId="F20">
<paragraph pageId="0" pageNumber="109">
<emphasis bold="true" pageId="0" pageNumber="109">Figure 26.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="109">A</emphasis>
<taxonomicName authority="joergenseni" lsidName="M. joergenseni subsp. joergenseni" pageId="0" pageNumber="109" rank="subSpecies" species="joergenseni" subSpecies="joergenseni">
<emphasis italics="true" pageId="0" pageNumber="109">M. joergenseni joergenseni</emphasis>
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♂
<emphasis bold="true" pageId="0" pageNumber="109">B</emphasis>
<taxonomicName authority="joergenseni" lsidName="M. joergenseni subsp. joergenseni" pageId="0" pageNumber="109" rank="subSpecies" species="joergenseni" subSpecies="joergenseni">
<emphasis italics="true" pageId="0" pageNumber="109">M. joergenseni joergenseni</emphasis>
</taxonomicName>
♀.
</paragraph>
</caption>
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</treatment>
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<taxonomicName id="0B6613607816A915EA2BBF1C4297D9DC" ID-CoL="4MW36" authority="Bavay &amp; Dautzenberg, 1909" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" class="Gastropoda" family="Achatinidae" genus="Prosopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopeas excellens" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excellens">Prosopeas excellens Bavay &amp; Dautzenberg, 1909</taxonomicName>
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<paragraph id="CBE690FAAC743BA2725E73C75886F239" pageId="0" pageNumber="1">
<taxonomicName id="52DE609CCD4876470AC48ADE78CE3B4C" class="Gastropoda" family="Achatinidae" genus="Prosopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopeas excellens" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excellens">Prosopeas excellens</taxonomicName>
Bavay &amp; Dautzenberg, 1909d[1908]: 247, 248. Type locality: Phong Tho [Phong Tho District, Lai Chau Province, Vietnam], Muong Bo [probably refers to the Nam Sai Commune, Sa Pa District, Lao Cai Province, Vietnam].
<bibRefCitation id="4B96C8CF597FE58854304163DFF7FE8E" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="279 - 288" title="Description de coquilles nouvelles de ľIndo-Chine (6 e suite)." volume="57" year="1909 c">Bavay and Dautzenberg 1909c</bibRefCitation>
: 282, 283, pl. 10, figs 11, 12.
<bibRefCitation id="4528828D37BF35B5A7BEDC235EC7C006" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko 2011</bibRefCitation>
: 10.
</paragraph>
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<paragraph id="6CF452D2D60F9C19D93359C60A1E69DC" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="1411F15DEF62C2A6BF8A407F5F47B7DF" pageId="0" pageNumber="1">Syntype MNHN-IM-2000-4661 from &quot;Muong-Bo et Phong-Tho&quot; (1 shell; Fig. 21F). Specimens from Tarng Kong, Ban Phak Kard village, Pek District, Xieng Khaung Province (Figs 21G, 55C).</paragraph>
</subSubSection>
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<paragraph id="A01D9DAB46599FE8B37F9846FABA5AA9" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="A9AC8E8A4608BA89FDF955335867275D" pageId="0" pageNumber="1">
Vietnam (
<bibRefCitation id="9A0DF3D6890449ECFDA6A2590E0B0AC7" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko 2011</bibRefCitation>
)
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Figure 21.
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A paralectotypes NHMUK 1856.9.15.68/2-11 B, C CUMZ collections D, E
<taxonomicName id="5A227AAF230B08802E00CB8B466DAD17" class="Gastropoda" family="Achatinidae" genus="Prosopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopeas anceyi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="anceyi">Prosopeas anceyi</taxonomicName>
D syntype of &quot;
<taxonomicName id="B951406AF219931E750C4D4BCB7B58F2" class="Gastropoda" family="Helicinidae" genus="Achatinoidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Achatinoidea macilentum" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="macilentum">macilentum</taxonomicName>
&quot; MNHN-IM-2000-4693 and E CUMZ collection F, G
<taxonomicName id="61DC6ECB85D8C707D90EB70D4C2797D3" class="Gastropoda" family="Achatinidae" genus="Prosopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopeas excellens" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="excellens">Prosopeas excellens</taxonomicName>
F syntype MNHN-IM-2000-4661 and G CUMZ collection H, I
<taxonomicName id="F90798D2CB421A94C1B633212C9C1FFD" class="Gastropoda" family="Achatinidae" genus="Prosopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopeas turricula" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="turricula">Prosopeas turricula</taxonomicName>
H syntype NHMUK 1895.8.1.10 and I CUMZ collection.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

185
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<treatment id="E6A86BEC98C55104D0A9498BD8B07BEF" ID-GBIF-Taxon="154127593" LSID="urn:lsid:plazi:treatment:E6A86BEC98C55104D0A9498BD8B07BEF" httpUri="http://treatment.plazi.org/id/E6A86BEC98C55104D0A9498BD8B07BEF" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
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<paragraph id="4B2E9EC206B047E37D67027685A62482" pageId="30" pageNumber="31">
<taxonomicName id="65E1590AA94D7833D3526F890A2381AF" ID-CoL="QNBR" authority="Boucomont, 1928" authorityName="Boucomont" authorityYear="1928" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium (Neocanthidium) centrale" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="centrale" subGenus="Neocanthidium">Canthidium (Neocanthidium) centrale Boucomont, 1928</taxonomicName>
Plate 10A
</paragraph>
</subSubSection>
<subSubSection id="56D08948CBE3CE7298BFEFD59DD84005" pageId="31" pageNumber="32" type="reference_group">
<paragraph id="753D80356F4B30F21DCD4C94B4A6B9A0" pageId="31" pageNumber="32">
<taxonomicName id="A6C402B7C040D5B06419C7252A75222C" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium centrale" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="centrale">
<pageBreakToken id="E3E8829F9B52799D1C9D597DDD6639FE" pageId="31" pageNumber="32" start="start">Canthidium</pageBreakToken>
centrale
</taxonomicName>
Boucomont, 1928b: 203 (original description. Type locality: Panama, Chiriqui, Guyane
<normalizedToken id="A4A546BF92B19B128FB676BDC9E5DBC9" originalValue="française">francaise</normalizedToken>
: St-Jean du Maroni, St-Laurent du Maroni, Guyane hollandaise, Surinam).
</paragraph>
<paragraph id="D2C584CAE618EEBD5D649F5B637E94DB" pageId="31" pageNumber="32">
<taxonomicName id="A5F9DF7A881A2D910C32625AA04C2015" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium centrale" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="centrale">Canthidium centrale</taxonomicName>
:
<bibRefCitation id="415FD7446D1C1111D91542BB243D7FBF" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
: 205 (list, distribution);
<bibRefCitation id="7F0BED6654543551DB11DB039AF08546" author="Vulcano, MA" journalOrPublisher="Atas do Simposio sobre a Biota Amazonica" pageId="279" pageNumber="280" pagination="533 - 603" title="Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera)." volume="5" year="1967">Vulcano and Pereira 1967</bibRefCitation>
: 593 (characters in key);
<bibRefCitation id="1993C4DC6E64811550BE6B901CC71630" author="Howden, HF" journalOrPublisher="Contributions of the American Entomological Institute" pageId="270" pageNumber="271" pagination="1 - 204" title="Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae)." volume="18" year="1981">Howden and Young 1981</bibRefCitation>
: 74 (characters in key), 75 (redescription);
<bibRefCitation id="1FDC1C3E6B9B50F1370BD5E00DC711AD" author="Medina, CA" journalOrPublisher="Biota Colombiana" pageId="274" pageNumber="275" pagination="131 - 144" title="Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia." volume="2" year="2001">Medina et al. 2001</bibRefCitation>
: 137 (cited for Colombia);
<bibRefCitation id="5EA2A69F62E8207F66DA2C649830D393" author="Ratcliffe, BC" journalOrPublisher="Zootaxa" pageId="276" pageNumber="277" pagination="1 - 48" title="A checklist of the Scarabaeoidea of Panama." url="https://doi.org/10.11646/zootaxa.32.1.1" volume="32" year="2002">Ratcliffe 2002</bibRefCitation>
: 14 (cited for Panama);
<bibRefCitation id="21A49EFB50516AF329AF6D826D23DC2D" author="Solis, A" journalOrPublisher="Giornale Italiano di Entomologia" pageId="277" pageNumber="278" pagination="1 - 73" title="El genero Canthidium (Coleoptera: Scarabaeidae) en Costa Rica." volume="11" year="2004">
<normalizedToken id="463E9775C05518EC59E2F887B7CC3B0C" originalValue="Solís">Solis</normalizedToken>
and Kohlmann 2004
</bibRefCitation>
: 9 (characters in key), 33 (redescription);
<bibRefCitation id="C316440A730E0755932E41884363362D" author="Solis, A" journalOrPublisher="Zootaxa" pageId="278" pageNumber="279" pagination="1 - 32" title="Checklist and Distribution Atlas of the Scarabaeinae of Costa Rica." volume="3482" year="2012">
<normalizedToken id="B508D839D7AF9346F3148A2F704EBC80" originalValue="Solís">Solis</normalizedToken>
and Kohlmann 2012
</bibRefCitation>
: 5 (cited for Costa Rica); Krajcik, 2012: 62 (complete list of species).
</paragraph>
<paragraph id="466D4196982C557B7C706098C1238FE0" pageId="31" pageNumber="32">
<taxonomicName id="7B7E4C3C9ECCD0AE3E2D8581BDF13C8F" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium (Canthidium) centrale" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="centrale" subGenus="Canthidium">Canthidium (Canthidium) centrale</taxonomicName>
:
<bibRefCitation id="B107839C99116E5E498D4287E677D3AD" author="Martinez, A" journalOrPublisher="Acta Zoologica Mexicana" pageId="273" pageNumber="274" pagination="19 - 40" title="Situacion del genero Canthidium." volume="18" year="1986">
<normalizedToken id="3DD817B764F9B4E748CFF4BF0534135C" originalValue="Martínez">Martinez</normalizedToken>
and Halffter 1986
</bibRefCitation>
: 26 (cited for Ecuador, Panama, Guatemala, Mexico);
<bibRefCitation id="3B0C2037BBD9A78508B5F79B07723355" author="Moron, MA" journalOrPublisher="Argania Editio, Barcelona" pageId="274" pageNumber="275" title="Atlas de los escarabajos de Mexico: Coleoptera: Lamellicornia. Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae." year="2003">
<normalizedToken id="BBF6DFE7AA612BF5016BA75DE4BF8DCE" originalValue="Morón">Moron</normalizedToken>
2003
</bibRefCitation>
: 55 (redescription, cited for Mexico);
<bibRefCitation id="6DB712AF06588155DDF5B85F5118A561" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
: 318-319 (cited for Ecuador);
<bibRefCitation id="4C6BFED0DB4B5AB9E3617CB8FA1051C9" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
: 92 (cited for Ecuador).
</paragraph>
<paragraph id="03B05FA58230E8A471C6807574570FE4" pageId="31" pageNumber="32">
<taxonomicName id="10464A4B7AB6CD0AE941ABC3B0DC229C" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium (Neocanthidium) centrale" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="centrale" subGenus="Neocanthidium">Canthidium (Neocanthidium) centrale</taxonomicName>
:
<bibRefCitation id="925D97641676501C60558D6152F3A420" author="Cupello, M" journalOrPublisher="Zootaxa" pageId="265" pageNumber="266" pagination="451 - 486" title="On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species." url="https://doi.org/10.11646/zootaxa.4388.4.1" volume="4388" year="2018">Cupello 2018</bibRefCitation>
: 467 (transferred to the subgenus
<taxonomicName id="19255CABB87CE1278E5A58A769587F59" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium (Neocanthidium)" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subGenus" subGenus="Neocanthidium">Canthidium (Neocanthidium)</taxonomicName>
<normalizedToken id="4EA8B043BDFB2EF1C4B35C8B37ADB50E" originalValue="Martínez">Martinez</normalizedToken>
, Halffter &amp; Pereira, 1964. Distribution, cited for Ecuador);
<bibRefCitation id="99C3ED3664908C69BF83771FCCB019E9" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
: 92 (cited for Ecuador).
</paragraph>
<paragraph id="869CBB422345B57DDAAC61003CF88C4E" pageId="31" pageNumber="32">
<taxonomicName id="5C5CE4BDF8655BB2D486A3B487728E36" class="Insecta" family="Scarabaeidae" genus="Neocanthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocanthidium martinezi" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="martinezi">Neocanthidium martinezi</taxonomicName>
Edmonds &amp; Halffter, 1978: 319 (original description);
<bibRefCitation id="96D44D204469FEB3FF08A8D9265E0D86" author="Solis, A" journalOrPublisher="Giornale Italiano di Entomologia" pageId="277" pageNumber="278" pagination="1 - 73" title="El genero Canthidium (Coleoptera: Scarabaeidae) en Costa Rica." volume="11" year="2004">
<normalizedToken id="986A294CA653583F496CB7CBCC7D495A" originalValue="Solís">Solis</normalizedToken>
and Kohlmann 2004
</bibRefCitation>
: 33 (synonym of
<taxonomicName id="582E7FA9665C56A3621B484AFCB8A071" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium centrale" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="centrale">Canthidium centrale</taxonomicName>
Boucomont, 1928);
<bibRefCitation id="09AC4D60ED0C1CE143BB899D7A1A8E6D" author="Solis, A" journalOrPublisher="Zootaxa" pageId="278" pageNumber="279" pagination="1 - 32" title="Checklist and Distribution Atlas of the Scarabaeinae of Costa Rica." volume="3482" year="2012">
<normalizedToken id="406726BBED1D4CC088452F27FAAE6B2F" originalValue="Solís">Solis</normalizedToken>
and Kohlmann 2012
</bibRefCitation>
: 5 (cited as synonym of
<taxonomicName id="35C470CCE2EEF33DF60714312912B284" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium centrale" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="centrale">Canthidium centrale</taxonomicName>
Boucomont, 1928); Cupello, 2018: 467 (comment).
</paragraph>
</subSubSection>
<subSubSection id="A44AAB637E665D56653A9E229239BF50" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="type specimens">
<paragraph id="3192CF6A077A100F5E0CB5AE9E51E026" pageId="31" pageNumber="32">Type specimens.</paragraph>
<paragraph id="5D8919D03928AD8668431ED616F3491B" pageId="31" pageNumber="32">
<taxonomicName id="27759E11E3DC99BB228E6B4BD4F6A0C3" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium centrale" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="centrale">Canthidium centrale</taxonomicName>
Boucomont, 1928. The holotype (sex unknown) is deposited at the MNHN (see
<bibRefCitation id="9F2DE2A7FBC4B1837B1B5B665CAB4A60" author="Solis, A" journalOrPublisher="Giornale Italiano di Entomologia" pageId="277" pageNumber="278" pagination="1 - 73" title="El genero Canthidium (Coleoptera: Scarabaeidae) en Costa Rica." volume="11" year="2004">
<normalizedToken id="563A27661A7696BC78629E353309AEC7" originalValue="Solís">Solis</normalizedToken>
and Kohlmann 2004
</bibRefCitation>
: 33). Locality: Panama, Chiriqui. Not examined.
</paragraph>
<paragraph id="97BAEEBFF750EB2645372DBA346F2839" pageId="32" pageNumber="33">
<taxonomicName id="532382FC3C0B53DD38C244620ECB5211" class="Insecta" family="Scarabaeidae" genus="Neocanthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocanthidium martinezi" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="martinezi">
<pageBreakToken id="413DA7497E95B50E796DEA77BC807662" pageId="32" pageNumber="33" start="start">Neocanthidium</pageBreakToken>
martinezi
</taxonomicName>
Edmonds &amp; Halffter, 1978. Type material not examined.
</paragraph>
</subSubSection>
<subSubSection id="A92B5FE32C71F9E2ADC3FA225C0BA824" pageId="32" pageNumber="33" type="distribution">
<paragraph id="B429609879653A8DF324397B9B41F57E" pageId="32" pageNumber="33">Distribution.</paragraph>
<paragraph id="00D9E95AC2373C92B6D7FA3115CF3A52" pageId="32" pageNumber="33">Colombia, Ecuador, Guatemala, Panama, and Mexico.</paragraph>
</subSubSection>
<subSubSection id="0E040FE3B269BD38AD872BB11AA53C3E" pageId="32" pageNumber="33" type="records examined">
<paragraph id="710FCBAA3C45348E9C03662D3DCAF919" pageId="32" pageNumber="33">Records examined.</paragraph>
<paragraph id="C887DCF6F73F9C1CE25BC64D0DEED1E2" pageId="32" pageNumber="33">
ESMERALDAS: Majua (1 specimen CEMT); Playa de Oro (1 specimen CEMT); Pote (1 specimen CEMT). LOS
<normalizedToken id="5BA79B602A8D1CE9DB2E16B2CE73BC3A" originalValue="RÍOS">RIOS</normalizedToken>
:
<normalizedToken id="7489E1061BB0DB5DCD550EADC39AF135" originalValue="Río">Rio</normalizedToken>
Palenque Station (1 specimen CEMT).
</paragraph>
</subSubSection>
<subSubSection id="A702D65E27F98B4B35C145C492118DE1" pageId="32" pageNumber="33" type="temporal data">
<paragraph id="334ECDC6FECA7E86E64CB5E8BC3AAB02" pageId="32" pageNumber="33">Temporal data.</paragraph>
<paragraph id="7AF879C1D9E95E054D86C1333E47B059" pageId="32" pageNumber="33">Collected in February and March.</paragraph>
</subSubSection>
<subSubSection id="B42AC07A9822995642CF98BD6DD177E5" pageId="32" pageNumber="33" type="remarks">
<paragraph id="EBD9FD0149206C91B29B32AD241CA47F" pageId="32" pageNumber="33">Remarks.</paragraph>
<paragraph id="DD458014C8512446B5793E96C725B903" pageId="32" pageNumber="33">Inhabits coastal lowland evergreen forests. Collected with pitfall traps baited with human feces.</paragraph>
</subSubSection>
</treatment>
</document>