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<mods:title id="7F889CDB7AA91E70EA693151A5780B82">Revalidation of the genus Sadala Simon, 1880 with the description of a new genus of Neotropical huntsman spiders (Araneae, Sparassidae)</mods:title>
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<taxonomicName id="4C3E4D7AFFB1C91EFF32FF1DFE7DFF72" ID-CoL="99BVD" authority="Simon, 1880" authorityName="Simon IBSP" authorityYear="1880" box="[151,499,151,178]" class="Arachnida" family="Sparassidae" genus="Sadala" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="punicea">
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<taxonomicName id="4C3E4D7AFFB1C91EFF32FF1DFE7DFF72" ID-CoL="99BVD" authority="Simon, 1880" authorityName="Simon " authorityYear="1880" box="[151,499,151,178]" class="Arachnida" family="Sparassidae" genus="Sadala" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="punicea">
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<emphasis id="B94AEAEBFFB1C91EFF32FF1DFEDCFF72" bold="true" box="[151,338,151,178]" italics="true" pageId="16" pageNumber="17">Sadala punicea</emphasis>
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<bibRefCitation id="EFAF4B08FFB1C91EFEFCFF1DFE7DFF72" author="Simon, E." box="[345,499,151,177]" pageId="16" pageNumber="17" pagination="223 - 351" refId="ref51803" refString="Simon, E. (1880) Revision de la famille des Sparassidae (Arachnides). Actes de la Societe Linneenne de Bordeaux, 34 (2 / 3 / 4), 223 - 351." type="journal article" year="1880">Simon, 1880</bibRefCitation>
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</taxonomicName>
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@ -76,7 +76,7 @@ comb. rest.
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<paragraph id="8B8136F9FFB1C91EFF32FE8AFA0BFEDB" blockId="16.[151,1436,255,372]" box="[151,1413,255,280]" pageId="16" pageNumber="17">
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<treatmentCitation id="0A9F10E8FFB1C91EFF32FE8AFE61FEDB" author="Simon, E." box="[151,495,256,280]" page="328" pageId="16" pageNumber="17" year="1880">
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<taxonomicName id="4C3E4D7AFFB1C91EFF32FE8AFE61FEDB" ID-CoL="99BVD" authority="Simon, 1880: 328" authorityName="Simon IBSP" authorityPageNumber="328" authorityYear="1880" box="[151,495,256,280]" class="Arachnida" family="Sparassidae" genus="Sadala" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="punicea">
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<taxonomicName id="4C3E4D7AFFB1C91EFF32FE8AFE61FEDB" ID-CoL="99BVD" authority="Simon, 1880" authorityName="Simon " authorityPageNumber="328" authorityYear="1880" box="[151,495,256,280]" class="Arachnida" family="Sparassidae" genus="Sadala" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="punicea">
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<emphasis id="B94AEAEBFFB1C91EFF32FE8AFEBAFED4" box="[151,308,256,280]" italics="true" pageId="16" pageNumber="17">Sadala punicea</emphasis>
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<materialsCitation id="3B563CA4FFB1C91EFE9FFE8AFE61FEDB" ID-GBIF-Occurrence="3861775533" box="[314,495,256,280]" collectionCode="MNHN" country="Peru" latitude="-3.3167" location="Pebas" longLatPrecision="7" longitude="-71.8167" pageId="16" pageNumber="17" specimenCode="MNHN 3571" specimenCount="1" stateProvince="Loreto" typeStatus="holotype">
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<bibRefCitation id="EFAF4B08FFB1C91EFE9FFE8AFE61FEDB" author="Simon, E." box="[314,495,256,280]" pageId="16" pageNumber="17" pagination="223 - 351" refId="ref51803" refString="Simon, E. (1880) Revision de la famille des Sparassidae (Arachnides). Actes de la Societe Linneenne de Bordeaux, 34 (2 / 3 / 4), 223 - 351." type="journal article" year="1880">Simon, 1880: 328</bibRefCitation>
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@ -103,7 +103,7 @@ from
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<paragraph id="8B8136F9FFB1C91EFF32FE95FD9AFEF5" blockId="16.[151,1436,255,372]" box="[151,532,286,310]" pageId="16" pageNumber="17">
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<treatmentCitation id="0A9F10E8FFB1C91EFF32FE95FD9EFEF5" author="Simon, E." box="[151,528,286,310]" page="35" pageId="16" pageNumber="17" year="1897">
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<taxonomicName id="4C3E4D7AFFB1C91EFF32FE95FD9EFEF5" authority=": Simon 1897: 35" authorityName="Simon" authorityPageNumber="35" authorityYear="1897" box="[151,528,286,310]" class="Arachnida" family="Sparassidae" genus="Sparassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="puniceus">
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<taxonomicName id="4C3E4D7AFFB1C91EFF32FE95FD9EFEF5" authority=": Simon 1897" authorityName="Simon" authorityPageNumber="35" authorityYear="1897" box="[151,528,286,310]" class="Arachnida" family="Sparassidae" genus="Sparassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="puniceus">
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<emphasis id="B94AEAEBFFB1C91EFF32FE95FED0FEF5" box="[151,350,287,310]" italics="true" pageId="16" pageNumber="17">Sparassus puniceus</emphasis>
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:
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<bibRefCitation id="EFAF4B08FFB1C91EFEC9FE94FD9EFEF5" author="Simon, E." box="[364,528,286,310]" pageId="16" pageNumber="17" pagination="1 - 192" refId="ref51872" refString="Simon, E. (1897) Histoire naturelle des araignees. Tome Second. Deuxieme Edition. Roret, Paris, pp. 1 - 192. https: // doi. org / 10.5962 / bhl. title. 51973" type="book chapter" year="1897">Simon 1897: 35</bibRefCitation>
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<document id="FF2896DBD2C5CDA2B39B543F23E4A204" ID-DOI="10.3897/zse.101.143073" ID-ZooBank="32EEBC10-2098-48B5-888F-2429C1A6A326" ID-publisher-id="143073" URI-arpha="4E33F87F-1D0D-5091-BC65-E7DF2AC0602E" URI-zoobank="http://zoobank.org/32EEBC10-2098-48B5-888F-2429C1A6A326" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1739460107938" checkinUser="pensoft" docAuthor="Srikampha, Khanet, Wesener, Thomas, Likhitrakarn, Natdanai, Sutcharit, Chirasak & Srisonchai, Ruttapon" docDate="2025" docId="858B300701D35F8499C8711C5855CF65" docLanguage="en" docName="Zoosystematics and Evolution 101(1): 273-300" docOrigin="Zoosystematics and Evolution 101 (1)" docSource="https://zse.pensoft.net/article/143073/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Sphaerobelum nonghinensis Srikampha & Srisonchai 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="300" masterDocId="4E33F87F1D0D5091BC65E7DF2AC0602E" masterDocTitle="The millipede genus Sphaerobelum Verhoeff, 1924, in Thailand, with descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="300" masterPageNumber="273" pageNumber="273" updateTime="1739469338090" updateUser="admin">
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<mods:mods id="F4CEDF848AC74FBED79D263F1BA7A3E3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="28B0B53D4048AF38D971A792F43B3E63">
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<mods:title id="323A56709A1E70CD62E133D97691A779">The millipede genus Sphaerobelum Verhoeff, 1924, in Thailand, with descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="DDCD2A57D8E5B002837C814911B527FB" type="personal">
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<mods:role id="533519D4A5FAA216653E2D1FC815FFEA">
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<mods:roleTerm id="FEEE232434097D92A84BA066DADFDFCA">Author</mods:roleTerm>
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<mods:namePart id="10FA2AABE7761D9BFE4947493E8F4CFC">Srikampha, Khanet</mods:namePart>
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<mods:nameIdentifier id="5950636DA99AE19421FB3DD0F14BC6AF" type="ORCID">https://orcid.org/0009-0008-9446-2084</mods:nameIdentifier>
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<mods:affiliation id="9CED9C1FAF0D25BA8ED46AF004DDE13B">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="FA0BB7F3867ED5A44B956143CF6FB5FB" type="personal">
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<mods:roleTerm id="936BE830303136255ED05410BFAAA5B0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5BAFD7E2AA62D9B755D9DF02742A09C5">Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier id="2FC2DA7B94EC080F02828886D51691F9" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
|
||||
<mods:affiliation id="7660763C3ECBC3D9E3B83C8ADBD3A374">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Study of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
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</mods:name>
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<mods:name id="76AD4B9235601ACB9ECF66FB0333F8EE" type="personal">
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<mods:roleTerm id="09C5BA8655050F0C4BFCB176B15F8998">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4CBA5BE3B074650744500F2D07D6FC12">Likhitrakarn, Natdanai</mods:namePart>
|
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<mods:nameIdentifier id="BDD26A0D1E030BAF71086F09672EDD36" type="ORCID">0000-0002-1306-317X</mods:nameIdentifier>
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<mods:affiliation id="338595064DEA7DA3CB8185A4C76A6F40">Program of Agriculture, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand & Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Phayathai Road, Patumwan, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="B5659838C5A2A99F66EB8FC06E8953CE" type="personal">
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<mods:roleTerm id="A1EE4D89FE5CDFBCB591BA4088AA6CFB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D69A1511BAE648354E9BFE09846ADC47">Sutcharit, Chirasak</mods:namePart>
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<mods:nameIdentifier id="B0416473447AF7C7BF83F3FF12E73604" type="ORCID">0000-0001-7670-9540</mods:nameIdentifier>
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<mods:affiliation id="37B9FC9483D7C546621F471EBD762F5F">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Phayathai Road, Patumwan, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="F1EC3A3748E117B0C98D393A0D38026B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E048BA7950D98FE0BE257FAFFD23FFA4">Srisonchai, Ruttapon</mods:namePart>
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<mods:nameIdentifier id="BD0174845A80DFF44EBD9CF5B152A119" type="ORCID">0000-0002-7142-0999</mods:nameIdentifier>
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<mods:affiliation id="172B064D6C706F2751681C410A591D88">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand & Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Phayathai Road, Patumwan, Bangkok 10330, Thailand</mods:affiliation>
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<mods:title id="F5D59B2665EBBCB7966DCFEDF6D14CA6">Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part id="CEBE08466305F28AFFB9F6EF45DBCF52">
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<mods:date id="6D12CB9EA38CA9512A12EACB92E466E1">2025</mods:date>
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<mods:number id="1F0BA2361DA4FD8BDA38D1AC265A7155">2025-02-13</mods:number>
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<mods:detail id="C7D02BA1FDF5757BDBEB712C20EE12F3" type="volume">
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<mods:number id="06DF38AAF4B4CA8021BB97B1102E509F">101</mods:number>
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</mods:detail>
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<mods:detail id="A4C2BB506E5A1A558AB8D307ACA8F468" type="issue">
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<mods:number id="DCC344F6D9838AC140BCC77F0A718297">1</mods:number>
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<mods:start id="9CA74D4EE5AD194AC291459D68256512">273</mods:start>
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<mods:end id="CCB9C1A00692FAD0FB7A2AC203A22D8A">300</mods:end>
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</mods:extent>
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<mods:classification id="49D807F7FC4790FD9BE6B8E7C873E962">journal article</mods:classification>
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<mods:identifier id="DE28EBF5444DD2D022CA160579A21249" type="DOI">10.3897/zse.101.143073</mods:identifier>
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<mods:identifier id="148FA806DF9D3D9E56154A373C2E6DA0" type="ZooBank">32EEBC10-2098-48B5-888F-2429C1A6A326</mods:identifier>
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</mods:mods>
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<treatment id="858B300701D35F8499C8711C5855CF65" ID-arpha="858B3007-01D3-5F84-99C8-711C5855CF65" ID-zoobank="https://zoobank.org/C638CEF4-6641-45E4-9472-B46874557100" LSID="urn:lsid:plazi:treatment:858B300701D35F8499C8711C5855CF65" LSID-ZBK="urn:lsid:zoobank.org:act:C638CEF4-6641-45E4-9472-B46874557100" httpUri="http://treatment.plazi.org/id/858B300701D35F8499C8711C5855CF65" scope_class="Diplopoda" scope_family="Zephroniidae" scope_order="Sphaerotheriida" scope_subFamily="Zephroniinae" scope_tribe="Zephroniini">
|
||||
<subSubSection id="1077E765E72A51C1C24068D7E8B009AF" type="nomenclature">
|
||||
<paragraph id="BC3697CD4C18E72EE367E21F800D8DF4">
|
||||
<taxonomicName id="16850743547793937BC30541FDF222FD" authority="Srikampha & Srisonchai" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="9A4BDFED90EFCC668134CB5058CB20A3" italics="true">Sphaerobelum nonghinensis</emphasis>
|
||||
Srikampha & Srisonchai
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="D6D9332186FFB0703C21A533870EECDF" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="376C8FF20726346FCEA3F0545978DA47" type="description">
|
||||
<paragraph id="A4C88A1B980353C8DCA6EAD901CCC6FE">
|
||||
<figureCitation id="28C6E9A652C907D4EAA0A7030F0DA55B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photograph of live specimens of Sphaerobelum nonghinensis sp. nov. A – F. Paratypes (MZKKU - MYR 0002); G, H. Habitat at the type locality (limestone). Scale bars: 10.0 mm." figureDoi="10.3897/zse.101.143073.figure2" httpUri="https://binary.pensoft.net/fig/1249824">Figs 2</figureCitation>
|
||||
,
|
||||
<figureCitation id="2D2D09E80265362C4CF901E0EE885E2B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A. Head region, lateral view; B. Left antenna, anterior view; C. Antennal disc, anterior view; D. Apical cones on antennal disc, lateral view; E. Organ of Tömösváry, lateral view; F. Epipharynx, ventral view; G. Gnathochilarium, ventral view; H. Sensory cones on inner palpi, anterior view; I. Mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipite. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure3" httpUri="https://binary.pensoft.net/fig/1249825">3</figureCitation>
|
||||
,
|
||||
<figureCitation id="57C71B001CF1105C21E65D6164EAF53C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Sphaerobelum nonghinensis sp. nov. A – E. ♂ paratype (MZKKU - MYR 0002); F, G. ♀ paratype (MZKKU - MYR 0002). A. Leg 11, arrow points to femoral ridge; B. Small spines on femur; C. Stigmatic plate 1; D. Coxae of legs 1; E. Coxae of legs 2 with gonopores; F. Leg 2 with vulvae; G. Subanal plate. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, Sp = apico-mesally spine of leg, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure4" httpUri="https://binary.pensoft.net/fig/1249826">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="82D8836916256FE87003D801E4E1B23C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A – C. Telopods, anterior, posterior, and ventral views, respectively; D. Right posterior telopod, anterior view; E – G. Anterior telopod, anterior, sublateral, and subventral views, respectively. Abbreviations: sp = telopoditomere 4 of posterior telopod’s spine. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure5" httpUri="https://binary.pensoft.net/fig/1249827">5</figureCitation>
|
||||
,
|
||||
<figureCitation id="BA402AD3F4D46EB60C5B9B9857CEC1F2" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM of endoterga of midbody tergites. A, B. Sphaerobelum nonghinensis sp. nov. (♂ paratype, MZKKU - MYR 0002); C, D. Sphaerobelum onyx sp. nov. (♂ paratype, MZKKU - MYR 0004); E, F. Sphaerobelum petramurum sp. nov. (♂ paratype, MZKKU - MYR 0006). Abbreviations: cp = cuticular impression, ia = inner area, ma = middle area, oa = outer area, pm = posterior margin. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure14" httpUri="https://binary.pensoft.net/fig/1249836">14 A, B</figureCitation>
|
||||
,
|
||||
<figureCitation id="25B625C25215CE8E5F083BBDDEB70FB9" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Line drawings of posterior telopods of Sphaerobelum species in Thailand. A, B. Sphaerobelum aesculus; C, D. Sphaerobelum meridionalis, modified from Bhansali & Wesener, 2022; E, F. Sphaerobelum nonghinensis sp. nov.; G, H. Sphaerobelum onyx sp. nov.; I, J. Sphaerobelum petramurum sp. nov.; K, L. Sphaerobelum truncatum; M, N. Sphaerobelum turcosa. A, B, E – N. Right posterior telopods; C, D. Left posterior telopods. A, D, E, G, I, K, M. Posterior view; B, C, F, H, J, L, N. Anterior view. Abbreviations: cr-t = crenulated teeth, ml = membranous lobe, sp = spine on telopoditomere 4 of posterior telopod. Scale bar: 0.5 millimetres." figureDoi="10.3897/zse.101.143073.figure15" httpUri="https://binary.pensoft.net/fig/1249837">15 E, F</figureCitation>
|
||||
,
|
||||
<figureCitation id="3F843E97E07C7EEC814A2D5C2628181F" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution map of Sphaerobelum species in Thailand. Different symbols with numbers represent species: 1 = S. aesculus, 2 = S. meridionalis, 3 = S. nonghinensis sp. nov., 4 = S. onyx sp. nov., 5 = S. petramurum sp. nov., 6 = S. truncatum, 7 = S. turcosa. Black symbol = type locality; white symbol = other locality." figureDoi="10.3897/zse.101.143073.figure16" httpUri="https://binary.pensoft.net/fig/1249838">16</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EDMAG" type="type material">
|
||||
<paragraph id="29B775D4E37AEAF84328DE103DF430BC">
|
||||
<heading id="9F9C864812D86264EB535E91A389964A" reason="title">Type material.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="40F9F95DD3B61BFEE98BDF2CF21F9531">
|
||||
<materialsCitation id="960B209816871FF8E792F26370E7ED2F" collectingDate="2021-10-23" collectionCode="MZKKU" collectorName="K. Srikampha & N. Srimongkol" country="Thailand" elevation="680" latitude="17.050554" location="Suan Hin Pha Ngam" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" specimenCode="MZKKU-MYR 0001" specimenCount="1" specimenCount-male="1" stateProvince="Loei Province" typeStatus="holotype">
|
||||
<emphasis id="A062725333A3A047D69F9728912BB102" bold="true">
|
||||
<emphasis id="EFF97EA9AB9B2BB9075CD9AB42A9CB5F" bold="true" italics="true">
|
||||
<typeStatus id="4D538CF685E37FBBE2DDAD826CDDE0F8" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<collectingCountry id="2F906CA2499C8A4946E502EC803B7728" name="Thailand">Thailand</collectingCountry>
|
||||
•
|
||||
<specimenCount id="434531289947FDA177C92654AAED65BC" count="1" type="male">♂</specimenCount>
|
||||
(
|
||||
<specimenCode id="81C4038696ED050C9F58AD2C81A9D0E8">
|
||||
<named-content id="C2298CED6371416AC286B5BFCC6F66BD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0001
|
||||
</specimenCode>
|
||||
),
|
||||
<collectingRegion id="BB65D6246BEABD46E9465B8B5D2C8703" country="Thailand" name="Loei">Loei Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="6520D9B424B0513EF36E6087C2645A09">Nong Hin District</collectingMunicipality>
|
||||
,
|
||||
<location id="B60D9F74585004EA3CB81043E61C28CF" LSID="urn:lsid:plazi:treatment:858B300701D35F8499C8711C5855CF65:B60D9F74585004EA3CB81043E61C28CF" country="Thailand" latitude="17.050554" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" name="Suan Hin Pha Ngam" stateProvince="Loei Province">Suan Hin Pha Ngam</location>
|
||||
(Hin Pha Ngam Park),
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWMAG" specific-use="{"type":"Point","coordinates":[101.744444,17.050556]}">
|
||||
<geoCoordinate id="52BB953F092CBF8EE1ABD27E0A507F62" degrees="17" direction="north" minutes="03" orientation="latitude" precision="15" seconds="02" value="17.050554">17°03'02"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="4566373984EAA7967B0F4C14133A8760" degrees="101" direction="east" minutes="44" orientation="longitude" precision="15" seconds="40" value="101.74444">101°44'40"E</geoCoordinate>
|
||||
</named-content>
|
||||
, ca.
|
||||
<elevation id="3B6ED29EE53122B7F458030E3B1DA7C6" metricMagnitude="2" metricUnit="m" metricValue="6.8" unit="m" value="680.0">
|
||||
<quantity id="FAE074C2B80D9A7A32C39BABEC621455" metricMagnitude="2" metricUnit="m" metricValue="6.8" unit="m" value="680.0">680 m</quantity>
|
||||
a. s. l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="846D241DA469BC9AB00F8ED250649FA0" value="2021-10-23">
|
||||
<collectingDate id="ABAAD1B096D79647792D9EDFE36557FD" value="2021-10-23">23 October 2021</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="307FB7AA7D44873AAB2B5DDBA337C9FD">K. Srikampha</collectorName>
|
||||
and
|
||||
<collectorName id="FE0F22565B028D3FBCF817AC00ED93FD">N. Srimongkol</collectorName>
|
||||
coll.
|
||||
</materialsCitation>
|
||||
<materialsCitation id="DBD55938A90D989C22C0CB7F0377CCB6" collectingDate="2021-10-23" collectionCode="MZKKU" collectorName="K. Srikampha & N. Srimongkol" country="Thailand" elevation="680" latitude="17.050554" location="Thailand" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" specimenCode="MZKKU-MYR 0002" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Loei" typeStatus="paratype">
|
||||
<emphasis id="6F5CDCE71E71EF00F31949331C07779E" bold="true">
|
||||
<emphasis id="DAE6FB34F823BE9EBA647902C0216E67" bold="true" italics="true">
|
||||
<typeStatus id="B4C08D5F434FD6BB54332B7151BE86D2" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<collectingCountry id="C3D02D3867676FE05F8AF91D3A44DFAA" name="Thailand">Thailand</collectingCountry>
|
||||
• 17 ♂♂, 12 ♀♀ (
|
||||
<specimenCode id="37E856B36A915C02FED6A532B1D2B28A">
|
||||
<named-content id="07517447A9F18760AA27211ED5299689" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0002
|
||||
</specimenCode>
|
||||
),
|
||||
<specimenCount id="31E89748FE1C83ABD67E51F94FA0D8F5" count="2" type="male">2 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="0A82CF15D60B50173279C4A34F93A585" count="1" type="female">♀</specimenCount>
|
||||
for
|
||||
<abbrev id="ABBRID0EENAG" xlink_title="scanning electron microscopy">SEM</abbrev>
|
||||
(
|
||||
<specimenCode id="E7C42453FBF5A4EEC372151CFA31B1AC">
|
||||
<named-content id="F2D40C9E2CEB83296F8D6CBD6B8BF57E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0002
|
||||
</specimenCode>
|
||||
),
|
||||
<backReference id="5C1D24B1D5F29B338D487B5E5201A356">same data as holotype</backReference>
|
||||
</materialsCitation>
|
||||
. •
|
||||
<materialsCitation id="94BDC3CEAA068BC2B497359F32E73BD6" collectingDate="2024-09-27" collectionCode="MZKKU" collectorName="K. Srikampha" country="Thailand" elevation="680" latitude="17.050554" location="Suan Hin Pha Ngam" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" specimenCode="MZKKU-MYR 0002" specimenCount="13" specimenCount-female="9" specimenCount-male="4" stateProvince="Loei" typeStatus="paratype">
|
||||
<specimenCount id="53C9533BA7F100B13277C9C3C2651444" count="4" type="male">4 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="07A1C52A67FFAA46807B28CE16DCD262" count="9" type="female">9 ♀♀</specimenCount>
|
||||
(
|
||||
<specimenCode id="1179FB61C093A021081200CAA7370524">
|
||||
<named-content id="0F4CF07E0740449BEDC5E7F44FEB17B9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0002
|
||||
</specimenCode>
|
||||
),
|
||||
<backReference id="3E0EE435F41CB80EC285E8809DECD765">same data as holotype</backReference>
|
||||
,
|
||||
<date id="3160A846B391780CF9E9CDCD6A5AFB2C" value="2024-09-27">
|
||||
<collectingDate id="7A92A1FD76BF55FD8603A71E4CE5AF5C" value="2024-09-27">27 September 2024</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="3055A71980BF8F3397D045A284791BAB">K. Srikampha</collectorName>
|
||||
coll.
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="D7B5B47830C17CDD04AC4324F0AEFCAF" collectingDate="2021-10-23" collectionCode="CUMZ" collectorName="K. Srikampha & N. Srimongkol" country="Thailand" elevation="680" latitude="17.050554" location="Suan Hin Pha Ngam" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" specimenCode="CUMZ - MYR 0031" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loei" typeStatus="paratype">
|
||||
<specimenCount id="A773EA7C471AACA889C0677CED407F4F" count="1" type="male">♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="24691B2C13A5F237ABBC47B4E99C127F" count="1" type="female">♀</specimenCount>
|
||||
(
|
||||
<specimenCode id="DEDBEC0077966832FB182318CAD0DD22">
|
||||
<named-content id="B0ED9D3A32155D829E9AD6D8331513E4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Museum of Zoology, Chulalongkorn University, Thailand">CUMZ</named-content>
|
||||
- MYR 0031
|
||||
</specimenCode>
|
||||
),
|
||||
<backReference id="64E2F515E2F44E29C017346684514DAD">same data as holotype</backReference>
|
||||
</materialsCitation>
|
||||
. •
|
||||
<materialsCitation id="36B7DBF3E3BE7A4581CAF6BF0E1518F4" collectingDate="2021-10-23" collectionCode="NHM" collectorName="K. Srikampha & N. Srimongkol" country="Thailand" elevation="680" latitude="17.050554" location="Suan Hin Pha Ngam" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loei" typeStatus="paratype">
|
||||
<specimenCount id="1702A8D2410716B8387912C420455164" count="1" type="male">♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="46F0A9B1C75808EF65CEF2B1CFABEF71" count="1" type="female">♀</specimenCount>
|
||||
(
|
||||
<abbrev content-type="institution" id="ABBRID0EXNAG" xlink_title="Natural History Museum, England">
|
||||
<collectionCode id="A5CD212EE54B7FB671BE8C1D9D6CF9B1">NHM</collectionCode>
|
||||
</abbrev>
|
||||
),
|
||||
<backReference id="DAAC68156818CBC04727A7C0B27A5ACE">same data as holotype</backReference>
|
||||
</materialsCitation>
|
||||
. •
|
||||
<materialsCitation id="282E1812BD16CB6C19E351082F4F35FE" collectingDate="2021-10-23" collectionCode="NHMD" collectorName="K. Srikampha & N. Srimongkol" country="Thailand" elevation="680" latitude="17.050554" location="Suan Hin Pha Ngam" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loei" typeStatus="paratype">
|
||||
<specimenCount id="F8D030A1A645377CC2EA2AA778A4CEA2" count="1" type="male">♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="1F59C76C0F73003E587FE315BDB521EB" count="1" type="female">♀</specimenCount>
|
||||
(
|
||||
<abbrev content-type="institution" id="ABBRID0E3NAG" xlink_title="Natural History Museum of Denmark, Denmark">
|
||||
<collectionCode id="28B2CE11B05DF3FF84695CDA07434078">NHMD</collectionCode>
|
||||
</abbrev>
|
||||
),
|
||||
<backReference id="1CC7A654B813470C683A596641318DC3">same data as holotype</backReference>
|
||||
</materialsCitation>
|
||||
. •
|
||||
<materialsCitation id="3AE83DA649D4B18C6DE94BCBBF24C972" collectingDate="2021-10-23" collectionCode="NHMW" collectorName="K. Srikampha & N. Srimongkol" country="Thailand" elevation="680" latitude="17.050554" location="Suan Hin Pha Ngam" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loei" typeStatus="paratype">
|
||||
<specimenCount id="D305C5ED895A15E4D70E1984FF208A04" count="1" type="male">♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="EB7B87A5195FC4C4B16EF5F71E02D056" count="1" type="female">♀</specimenCount>
|
||||
(
|
||||
<named-content id="9B77D9216ECE3A23B3C169F93F378318" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-wien-0" xlink_title="Naturhistorisches Museum Wien, Vienna, Austria">
|
||||
<collectionCode id="F67878E14FC6404C0AA5140239F7C7DC" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
|
||||
</named-content>
|
||||
),
|
||||
<backReference id="34D554FF973A9200581CCC09BF2C948A">same data as holotype</backReference>
|
||||
</materialsCitation>
|
||||
. •
|
||||
<materialsCitation id="411D5AB2F5AF396470719CF36935F137" collectingDate="2021-10-23" collectionCode="ZFMK" collectorName="K. Srikampha & N. Srimongkol" country="Thailand" elevation="680" latitude="17.050554" location="Suan Hin Pha Ngam" longLatPrecision="21" longitude="101.74444" municipality="Nong Hin District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loei" typeStatus="paratype">
|
||||
<specimenCount id="173893EC841CD0583563E0766E75A513" count="1" type="male">♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="091330774CBDD54F8575A2169ED964A6" count="1" type="female">♀</specimenCount>
|
||||
(
|
||||
<named-content id="7E7427B7693219EC6F84943FA380FFF6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoological Research Museum Koenig, Bonn, Germany">
|
||||
<collectionCode id="87F881C46BA1995492A6CEB1145265A6" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
|
||||
</named-content>
|
||||
),
|
||||
<backReference id="EAEE9BE4112B7410EEC6C64A96E054A0">same data as holotype</backReference>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ELOAG" type="type material">
|
||||
<paragraph id="0E08CBF8CC82F5DD63D3752FC98902FE">
|
||||
<heading id="D8796006D750CE907128827B0E6108FC" reason="title">Additional material examined — non-type.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="06536A449F49BD31E3D3C77BC0E42184">
|
||||
<materialsCitation id="2CC1D44693916D4E30F74DF8B31905CD" collectingDate="2021-09-12" collectionCode="MZKKU" collectorName="K. Srikampha" country="Thailand" elevation="718" latitude="17.043612" location="Wat Tham Pha Fai" longLatPrecision="21" longitude="101.73611" municipality="Nong Hin District" specimenCount="1" specimenCount-female="1" stateProvince="Loei Province">
|
||||
<collectingCountry id="71F0B8691D590C973EDA8A017C8B8F76" name="Thailand">Thailand</collectingCountry>
|
||||
•
|
||||
<specimenCount id="0750CFBA93B0D4316E69945FC145A750" count="1" type="female">♀</specimenCount>
|
||||
(
|
||||
<named-content id="CB77CBEF0EBE95AA5A41FDBC65611183" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">
|
||||
<collectionCode id="948AEC95F5274E490A5C555D502FC0B2">MZKKU</collectionCode>
|
||||
</named-content>
|
||||
),
|
||||
<collectingRegion id="8082EE5E4FFABFC11C3BAB0BC58A2828" country="Thailand" name="Loei">Loei Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="B8BC56438B0292B38EAF33C4E8C7C676">Nong Hin District</collectingMunicipality>
|
||||
,
|
||||
<location id="2ED6C67231E08D1EC6B8C68223645DB6" LSID="urn:lsid:plazi:treatment:858B300701D35F8499C8711C5855CF65:2ED6C67231E08D1EC6B8C68223645DB6" country="Thailand" latitude="17.043612" longLatPrecision="21" longitude="101.73611" municipality="Nong Hin District" name="Wat Tham Pha Fai" stateProvince="Loei Province">Wat Tham Pha Fai</location>
|
||||
,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZOAG" specific-use="{"type":"Point","coordinates":[101.736111,17.043611]}">
|
||||
<geoCoordinate id="F314E4359FD87293F622E1CF20B5B778" degrees="17" direction="north" minutes="02" orientation="latitude" precision="15" seconds="37" value="17.043612">17°02'37"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="2EBEBA67DE58B124F8BFFAF46375580E" degrees="101" direction="east" minutes="44" orientation="longitude" precision="15" seconds="10" value="101.73611">101°44'10"E</geoCoordinate>
|
||||
</named-content>
|
||||
, ca.
|
||||
<elevation id="6B240B83D4D2A7A52760390E9D35768B" metricMagnitude="2" metricUnit="m" metricValue="7.18" unit="m" value="718.0">
|
||||
<quantity id="849C5A504519D31AEC57BB9D0564629C" metricMagnitude="2" metricUnit="m" metricValue="7.18" unit="m" value="718.0">718 m</quantity>
|
||||
a. s. l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="3C5168A5C2EEFC23A68322335D7A93AC" value="2021-09-12">
|
||||
<collectingDate id="D85A895E5FC5F2CBECC7908DF3CC744E" value="2021-09-12">12 September 2021</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="A82D95D4E4DB629250EC3F7C54044768">K. Srikampha</collectorName>
|
||||
coll.
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="DA0CD35D3C035E5D87D9917830185F8D" collectingDate="2021-09-12" collectionCode="MZKKU" collectorName="K. Srikampha" country="Thailand" elevation="344" latitude="17.07528" location="Wat Tham Sung" longLatPrecision="21" longitude="101.78555" municipality="Nong Hin District" specimenCount="1" specimenCount-male="1" stateProvince="Loei Province">
|
||||
<specimenCount id="E75833D55AEBDD78F13E52A8F13EA478" count="1" type="male">♂</specimenCount>
|
||||
(
|
||||
<named-content id="D269A33A894882CA1AAC8312D787C0F8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">
|
||||
<collectionCode id="2E06B90EE04B0B14D96CFF27CC6F9F82">MZKKU</collectionCode>
|
||||
</named-content>
|
||||
),
|
||||
<collectingRegion id="F05582F5E0876228BEFDF0B58D6CD830" country="Thailand" name="Loei">Loei Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="9C5800B26C8A9F85A451A20BC5CAAD8C">Nong Hin District</collectingMunicipality>
|
||||
,
|
||||
<location id="2DC150EFF5A283812B3FFF2893064F2D" LSID="urn:lsid:plazi:treatment:858B300701D35F8499C8711C5855CF65:2DC150EFF5A283812B3FFF2893064F2D" country="Thailand" latitude="17.07528" longLatPrecision="21" longitude="101.78555" municipality="Nong Hin District" name="Wat Tham Sung" stateProvince="Loei Province">Wat Tham Sung</location>
|
||||
(Wat Tham Pha Nang Khoi),
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGPAG" specific-use="{"type":"Point","coordinates":[101.785556,17.075278]}">
|
||||
<geoCoordinate id="E6B62996D6E9D4C557DA441C9EF887D7" degrees="17" direction="north" minutes="04" orientation="latitude" precision="15" seconds="31" value="17.07528">17°04'31"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="1021F8A362CB374993ECC9213DC2E2FE" degrees="101" direction="east" minutes="47" orientation="longitude" precision="15" seconds="08" value="101.78555">101°47'08"E</geoCoordinate>
|
||||
</named-content>
|
||||
, ca.
|
||||
<elevation id="62EB3D208D06F8519C8606302A3A23C6" metricMagnitude="2" metricUnit="m" metricValue="3.44" unit="m" value="344.0">
|
||||
<quantity id="E208434307039AB8A4DA3F39789939BB" metricMagnitude="2" metricUnit="m" metricValue="3.44" unit="m" value="344.0">344 m</quantity>
|
||||
a. s. l.
|
||||
</elevation>
|
||||
,
|
||||
<date id="F87F3B17662DE42864BA607099722878" value="2021-09-12">
|
||||
<collectingDate id="E4C8C071302F7FF258DC0CCDF5A0F442" value="2021-09-12">12 September 2021</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="55F6544CF5489E13ABBDA47D1D17A407">K. Srikampha</collectorName>
|
||||
coll.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ELPAG" type="diagnosis">
|
||||
<paragraph id="59AD886794246920DAE7BE34E59290FF">
|
||||
<heading id="5465C39A77E83C82B22711579FE14F3F" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8321073D386F2FEABC0C7E30A1087E98">
|
||||
<taxonomicName id="87BCAF7873F5444903D74D98151F8877" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="EBEBD13DBA1EF272BCECE30A14F0AB9C" italics="true">S. nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="E78E6E720E6D2BE5BAFB97BA931434FD" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs in the femur of the legs, in which only the apical margin carries a few teeth, a character only shared with
|
||||
<taxonomicName id="7C445112802F8DA291443740246A022C" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="bicorne">
|
||||
<emphasis id="B112A20E90FBDB7CB3CD3F92D66732D5" italics="true">S. bicorne</emphasis>
|
||||
<bibRefCitation id="9DC68EDEE90EC28416D119454AC64314" author="Attems C" issue="2" journalOrPublisher="Mémoires du Muséum National d’Histoire Naturelle" pagination="187-353" refId="B2" refString="Attems C (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum National d'Histoire Naturelle 6 (2): 187-353." title="Die von Dr. C. Dawydoff in französisch Indochina gesammelten Myriopoden." volume="6" year="1938">Attems, 1938</bibRefCitation>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="61205F58397A545C2DE4CDB519243B26" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="bolavensis">
|
||||
<emphasis id="39400D2D1A546B649394468C9494252B" italics="true">S. bolavensis</emphasis>
|
||||
<bibRefCitation id="1AE30F174D91F3BB2429704C623B151A" DOI="10.11646/zootaxa.4563.2.1" author="Wesener T" issue="2" journalOrPublisher="Zootaxa" pagination="201-248" refId="B46" refString="Wesener T (2019) First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae). Zootaxa 4563 (2): 201-248. https://doi.org/10.11646/zootaxa.4563.2.1" title="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)." volume="4563" year="2019">Wesener, 2019</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, as well as
|
||||
<taxonomicName id="D83A08369B06C1D078F1604843977F87" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
|
||||
<emphasis id="8831D1680F823E699B6FADB322161F21" italics="true">S. onyx</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="CCA747396D0C90E4CBE39D896818F94C" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
and
|
||||
<taxonomicName id="57DF1D03D189CF44C867549D7BC35752" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="petramurum" status="sp. nov.">
|
||||
<emphasis id="675C4E04DC4ABC0329DD537C72EA65F2" italics="true">S. petramurum</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="E8D53BFA6E1DE6B416C1B749A2E4C8AC" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
described below.
|
||||
<taxonomicName id="B673235AE13F0E17C3146A056A5B01BB" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="DCCFB9A2E67F390F3545F0C9428CF41A" italics="true">S. nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="3DA757351B3A204A5529B5BD5B716D04" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from both
|
||||
<taxonomicName id="82DB148C83DFDF2F9EDC6B79D9B15B8E" authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="bicorne">
|
||||
<emphasis id="8B26A0002EEB9060042075C8F93DCAF4" italics="true">S. bicorne</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="DC4773B0C642CC3E3EECA0D029E8CB15" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="bolavensis">
|
||||
<emphasis id="E305CEF5F4B4184155DF1D86C204625E" italics="true">S. bolavensis</emphasis>
|
||||
</taxonomicName>
|
||||
in the presence of a “ hook ” on telopoditomere 4 of the posterior telopods. Differs in the unique colouration pattern, black, with orange markings only on the thoracic shield and anal shield from other known
|
||||
<taxonomicName id="469C2214FBD6680C7C335A5F2396E008">
|
||||
<emphasis id="7D0F62EB3AC471D1EB0B72DCDFFFA868" italics="true">Sphaerobelum</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
, e. g., from
|
||||
<taxonomicName id="449438042173DD44E9BB324000C036C8" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="petramurum" status="sp. nov.">
|
||||
<emphasis id="119D1E0BCE295F99E9B335E79780655C" italics="true">S. petramurum</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="92A16AC4B26A67BD3DA43673ED4D65F5" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, which is greenish-bluish brown, and from
|
||||
<taxonomicName id="B104DBA5E272931C271BFCB88CBCBFC7" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
|
||||
<emphasis id="268828DD2B5B7F2762B9FB8EC13E0D50" italics="true">S. onyx</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="8858AE101FBEC3B1A58043E41DD872B5" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, which is entirely black.
|
||||
<taxonomicName id="58615CB2ACC50FA2349A9F6C071A5C6A" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="FBD70CE434EDFA32642C8F252CDB6287" italics="true">S. nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="EBBA804AD5EA446D35D3F52690C5A95D" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
has a “ hook ” on telopoditomere 4 of the posterior telopods that resembles those of
|
||||
<taxonomicName id="6C932CCE50BDC0B21DFE6543940791F6" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="clavigerum">
|
||||
<emphasis id="A388FD480E57EB19D8DB35311C7982F5" italics="true">S. clavigerum</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="8028193EF0F062B8EC03671B25DB3852" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="hirsutum">
|
||||
<emphasis id="3D7D2A93232D89E08D725EA56452EA88" italics="true">S. hirsutum</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="78C551E9A2F9154BED496ABD1E187539" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="laoticum">
|
||||
<emphasis id="EC37FF17C189B9D4EA0133EBFEBE8E0E" italics="true">S. laoticum</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="6F5F19A9A4CD891190E3ECD21F170B60" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
|
||||
<emphasis id="80F24224194288FB014BEE84A0BDC54D" italics="true">S. onyx</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="AF9A7AA371E437362E81C93F30D456C1" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
,
|
||||
<taxonomicName id="8217177EE0FD8465E782731751F5FBAF" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="petramurum" status="sp. nov.">
|
||||
<emphasis id="09310FD415997B0405E4BF6AC60C0FFC" italics="true">S. petramurum</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="93F4D14020FBD8D669F7FAF85D9890B8" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
,
|
||||
<taxonomicName id="1D23B688BD8D6780367966FF258C7FD7" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="schwendingeri">
|
||||
<emphasis id="362958C365C0B4C78240F2D8F413CE60" italics="true">S. schwendingeri</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="1B894BB0AC48295EBF06ABDF3A0D5932" authorityName="Attems" authorityYear="1953" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="separatum">
|
||||
<emphasis id="57E23ED95081E50976BB482682FBB799" italics="true">S. separatum</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="1AEA74754E9E06318BDC417BAD9A36CF" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="splendidum">
|
||||
<emphasis id="2ED0055702A9E1292299492CB219C089" italics="true">S. splendidum</emphasis>
|
||||
</taxonomicName>
|
||||
. The new species differs from
|
||||
<taxonomicName id="4203BA479632B7ADF28CA7CADEB6FEA3" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="clavigerum">
|
||||
<emphasis id="CF0F96DDF8E1FC79F9328AD1E655B12A" italics="true">S. clavigerum</emphasis>
|
||||
</taxonomicName>
|
||||
in the presence of a single short or dot-like locking carina on the anal shield (vs. 3 × long), from
|
||||
<taxonomicName id="15CB2DC3C1B83F04B69C76585185A33D" authorityName="Verhoeff" authorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="hirsutum">
|
||||
<emphasis id="8B84DF1229E5565A198B58E52E01378B" italics="true">S. hirsutum</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="49EE9FBD62FA66927E4333D8D7007EA6" authorityName="Attems" authorityYear="1953" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="separatum">
|
||||
<emphasis id="07DDD1CDFBD8BECDD57A38BA1D95F9F3" italics="true">S. separatum</emphasis>
|
||||
</taxonomicName>
|
||||
in the presence of glabrous tergites (vs. hairy). It differs from
|
||||
<taxonomicName id="8F4EDDE97C99E0FC1B01684CDBA946A5" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="laoticum">
|
||||
<emphasis id="93510AD225C6AE6F209C6B9AEEC928C5" italics="true">S. laoticum</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="1C98062F27CB5303A3C24168FE357EBE" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="schwendingeri">
|
||||
<emphasis id="FBEBA8BA071B6C3EF31988F860694C70" italics="true">S. schwendingeri</emphasis>
|
||||
</taxonomicName>
|
||||
by having <60 apical cones on antenna (vs.
|
||||
<quantity id="A3F654AD5D2D347D44616490EFE97DCE" metricMagnitude="0" metricUnit="m" metricValue="2.1209" metricValueMax="2.1844" metricValueMin="2.0574" unit="in" value="83.5" valueMax="86.0" valueMin="81.0">81–86 in</quantity>
|
||||
<taxonomicName id="716880FCDE51ED8EB5C0CDE82F1C1422" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="laoticum">
|
||||
<emphasis id="5F0D2C647A5834745DEDF770104C1446" italics="true">S. laoticum</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<quantity id="8AD00B5B4C0525552FEC2FADC4CD209E" metricMagnitude="0" metricUnit="m" metricValue="1.3843" metricValueMax="1.8542" metricValueMin="0.9144" unit="in" value="54.5" valueMax="73.0" valueMin="36.0">36–73 in</quantity>
|
||||
<taxonomicName id="731B4D5987A207351F21005174D2EFFD" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="schwendingeri">
|
||||
<emphasis id="8A212B4178C8BA092172E7153EB90D56" italics="true">S. schwendingeri</emphasis>
|
||||
</taxonomicName>
|
||||
) and by having a greenish-brown antenna (vs. red). It differs from
|
||||
<taxonomicName id="CC239319F3BBBC5785020758F5AF2C9C" authorityName="Wesener" authorityYear="2019" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="splendidum">
|
||||
<emphasis id="70211B32A72590CF8B83619658A6BA13" italics="true">S. splendidum</emphasis>
|
||||
</taxonomicName>
|
||||
by having an elongated telopoditomere 2 of the anterior telopod (vs. short, as long as telopoditomere 3).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EF1AG" type="description">
|
||||
<paragraph id="730F978A2F1E002927007B0B91C865F5">
|
||||
<heading id="D37462579639CCE9BC071E3CBD74E9D1" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="059E285E8C118824D3791A7F1FF3BFB7">
|
||||
<emphasis id="700C448624DB2C304DE77651C2D70936" bold="true" italics="true">Measurements</emphasis>
|
||||
: Male
|
||||
<typeStatus id="04F2050B48B202264AC1D70C94908B4A" type="holotype">holotype</typeStatus>
|
||||
; body length
|
||||
<quantity id="F9733C937D2BCD32248BF57815E893AF" metricMagnitude="-2" metricUnit="m" metricValue="2.61" unit="mm" value="26.1">26.1 mm</quantity>
|
||||
; width (broadest) of thoracic shield 13.0 mm, of tergite 7 =
|
||||
<quantity id="0618FA943D1AB234E95A2EC317C2CBA5" metricMagnitude="-2" metricUnit="m" metricValue="1.31" unit="mm" value="13.1">13.1 mm</quantity>
|
||||
; height of tergite 7 =
|
||||
<quantity id="BAD17EF071A2A313A579BDA69DFE270D" metricMagnitude="-3" metricUnit="m" metricValue="7.7" unit="mm" value="7.7">7.7 mm</quantity>
|
||||
. Males: body length =
|
||||
<quantity id="E4A964DA5649E11F00581F4E2AF5D207" metricMagnitude="-2" metricUnit="m" metricValue="2.6149999999999998" metricValueMax="2.79" metricValueMin="2.44" unit="mm" value="26.15" valueMax="27.9" valueMin="24.4">24.4–27.9 mm</quantity>
|
||||
; width (broadest) of thoracic shield =
|
||||
<quantity id="CA7774BD6F43B00A9840F9E14BF55FD8" metricMagnitude="-2" metricUnit="m" metricValue="1.215" metricValueMax="1.25" metricValueMin="1.18" unit="mm" value="12.15" valueMax="12.5" valueMin="11.8">11.8–12.5 mm</quantity>
|
||||
, of tergite 7 =
|
||||
<quantity id="0C54B8BF551850494A54F240F98CB27F" metricMagnitude="-2" metricUnit="m" metricValue="1.265" metricValueMax="1.31" metricValueMin="1.22" unit="mm" value="12.649999999999999" valueMax="13.1" valueMin="12.2">12.2–13.1 mm</quantity>
|
||||
; height of tergite 7 = 7.9–8.0 mm. Females: body length =
|
||||
<quantity id="B603888CA75F7D1C0BC611795CB2F3C2" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" metricValueMax="2.87" metricValueMin="2.57" unit="mm" value="27.2" valueMax="28.7" valueMin="25.7">25.7–28.7 mm</quantity>
|
||||
; width (broadest) of thoracic shield =
|
||||
<quantity id="27CD0023AF0BF2C6DA5AC5C867BAA687" metricMagnitude="-2" metricUnit="m" metricValue="1.385" metricValueMax="1.46" metricValueMin="1.31" unit="mm" value="13.85" valueMax="14.6" valueMin="13.1">13.1–14.6 mm</quantity>
|
||||
, of tergite 7 = 14.0–
|
||||
<quantity id="BB66750650C82CACE57A504C9D27D5AE" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" unit="mm" value="15.7">15.7 mm</quantity>
|
||||
; height of tergite 7 =
|
||||
<quantity id="727D0CEBF208B027A8186B5E76CC6A51" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.1" metricValueMin="8.4" unit="mm" value="9.25" valueMax="10.1" valueMin="8.4">8.4–10.1 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="9B44B4330B33F37EA133273B94CFC042">
|
||||
<emphasis id="7FD317452C67C21F3BBCD84B253F7052" bold="true" italics="true">Colouration</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="DC99F922EA601EC966275C89F598DC09" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photograph of live specimens of Sphaerobelum nonghinensis sp. nov. A – F. Paratypes (MZKKU - MYR 0002); G, H. Habitat at the type locality (limestone). Scale bars: 10.0 mm." figureDoi="10.3897/zse.101.143073.figure2" httpUri="https://binary.pensoft.net/fig/1249824">2 A – F</figureCitation>
|
||||
): Live specimens with dark body colour, shiny. Head, collum, and tergites shiny black. Lateral margins of both thoracic and anal shields contrastingly orangish brown. Antennae brown. Legs and venter greenish brown. Specimens in 70 % alcohol after 4 years changed to brownish black.
|
||||
</paragraph>
|
||||
<caption id="8229D7BD408EAAF26985B447F6DA5658" ID-DOI="10.3897/zse.101.143073.figure2" ID-arpha="9F26BB31-3726-50FC-A455-F246710A90FE" httpUri="https://binary.pensoft.net/fig/1249824" startId="F2">
|
||||
<paragraph id="03D74ADCD555E5B4DD450AF06DAAF6A4">
|
||||
<label id="B8747A64DAF50E990A02D4EF53757C48">Figure 2.</label>
|
||||
</paragraph>
|
||||
<paragraph id="825EACA2AEAC14F84C44ABCB04A6D0D8">
|
||||
Photograph of live specimens of
|
||||
<taxonomicName id="278810F4D4CF3C56958BFA5E23E82B2B" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="E0D2BDC3EC87CE13A05C8872A09F0392" italics="true">Sphaerobelum nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="991BB073A1C6AAE0A343D467952EF755" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="A5C70C706045C3BFCAE6D8C5F0FFE1A3" bold="true">A – F.</emphasis>
|
||||
Paratypes (
|
||||
<specimenCode id="D1706F25F5528B45CE757536DB5CB88F">
|
||||
<named-content id="9039827CA71DA096819B12E200106BF6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0002
|
||||
</specimenCode>
|
||||
);
|
||||
<emphasis id="5C33F69DA9AC5389AA7B1F76CB6E0EB9" bold="true">G, H.</emphasis>
|
||||
Habitat at the type locality (limestone). Scale bars: 10.0 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="B88A6672A390CFC1A2D019E7ED8DDEE5" ID-DOI="10.3897/zse.101.143073.figure3" ID-arpha="A1D4B228-5DAE-5F71-B98D-43B6F9084FCC" httpUri="https://binary.pensoft.net/fig/1249825" startId="F3">
|
||||
<paragraph id="69FF9E79590E70255E0645F81327247F">
|
||||
<label id="7D32EAB167110581C83A5250A5EC3640">Figure 3.</label>
|
||||
</paragraph>
|
||||
<paragraph id="5C9FEE3F029C9B9ECCC715E5BACA26E9">
|
||||
<abbrev id="ABBRID0E54AG" xlink_title="scanning electron microscopy">SEM</abbrev>
|
||||
of
|
||||
<taxonomicName id="E383F2525BAB18CAE4DAD3D8B441EAB2" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="6B01F9351143D3E1C7A185CFDF6AFACC" italics="true">Sphaerobelum nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="FBC5C85FDDA70D799132660597B1DB9B" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, ♂ paratype (
|
||||
<specimenCode id="1176D1E31AD028A951F91D2A4F2F7D31">
|
||||
<named-content id="FC43DCE2841777C34E26312C33A729D4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0002
|
||||
</specimenCode>
|
||||
).
|
||||
<emphasis id="BAC430959ED78E41356B9FE858ED80BE" bold="true">A.</emphasis>
|
||||
Head region, lateral view;
|
||||
<emphasis id="A09506034953CC0CC5C63A73C36B64D6" bold="true">B.</emphasis>
|
||||
Left antenna, anterior view;
|
||||
<emphasis id="5EBE29D10D71B7AC6E23880EDBD13A0C" bold="true">C.</emphasis>
|
||||
Antennal disc, anterior view;
|
||||
<emphasis id="0172EDF5B75B2D199FB76D079DF774C7" bold="true">D.</emphasis>
|
||||
Apical cones on antennal disc, lateral view;
|
||||
<emphasis id="44ECA84F1D89AE4771CD4DEA8A8CC2CC" bold="true">E.</emphasis>
|
||||
Organ of Tömösváry, lateral view;
|
||||
<emphasis id="2208BC43B9F1F15A97E22E08FA2CEEDE" bold="true">F.</emphasis>
|
||||
Epipharynx, ventral view;
|
||||
<emphasis id="C7DE2A2199A873828FCBEC9ABD906824" bold="true">G.</emphasis>
|
||||
Gnathochilarium, ventral view;
|
||||
<emphasis id="3E645F35F3A5AE8878836316A9B31A49" bold="true">H.</emphasis>
|
||||
Sensory cones on inner palpi, anterior view;
|
||||
<emphasis id="66751F6ED335F31FD9E237A37486B312" bold="true">I.</emphasis>
|
||||
Mandible, mesal view. Abbreviations:
|
||||
<abbrev id="ABBRID0EI6AG" xlink_title="3-combed inner tooth of mandible">3 it</abbrev>
|
||||
= 3 - combed inner tooth,
|
||||
<abbrev id="ABBRID0EM6AG" xlink_title="condyles of mandible">co</abbrev>
|
||||
= condylus,
|
||||
<abbrev id="ABBRID0EQ6AG" xlink_title="central tooth">ct</abbrev>
|
||||
= central tooth,
|
||||
<abbrev id="ABBRID0EU6AG" xlink_title="external tooth of epipharynx">et</abbrev>
|
||||
= external tooth,
|
||||
<abbrev id="ABBRID0EY6AG" xlink_title="external tooth of mandibular gnathal lobe">Et</abbrev>
|
||||
= external tooth,
|
||||
<abbrev id="ABBRID0E36AG" xlink_title="inner area of mandible">Ia</abbrev>
|
||||
= inner area,
|
||||
<abbrev id="ABBRID0EAABG" xlink_title="inner palpi of gnathochilarium">ip</abbrev>
|
||||
= inner palpi,
|
||||
<abbrev id="ABBRID0EEABG" xlink_title="lamellae lingulales of gnathochilarium">ll</abbrev>
|
||||
= lamellae linguales,
|
||||
<abbrev id="ABBRID0EIABG" xlink_title="mentum">me</abbrev>
|
||||
= mentum,
|
||||
<abbrev id="ABBRID0EMABG" xlink_title="molar plate of mandible">mp</abbrev>
|
||||
= molar plate,
|
||||
<abbrev id="ABBRID0EQABG" xlink_title="pectinate lamellae of mandible">pl</abbrev>
|
||||
= pectinate lamellae,
|
||||
<abbrev id="ABBRID0EUABG" xlink_title="row of spine on epipharynx">rsp</abbrev>
|
||||
= row of spines,
|
||||
<abbrev id="ABBRID0EYABG" xlink_title="sensory cone">sc</abbrev>
|
||||
= sensory cone,
|
||||
<abbrev id="ABBRID0E3ABG" xlink_title="stipites">st</abbrev>
|
||||
= stipite. Scale bars in millimetres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="3FB6FED4804128C03D1AA9D86FBCC5EC">
|
||||
<emphasis id="6F908B2722A031416DBE40DB6E79F6FC" bold="true" italics="true">Head</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="3FD96CF5CFB14CADCA7D73B098F2B438" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A. Head region, lateral view; B. Left antenna, anterior view; C. Antennal disc, anterior view; D. Apical cones on antennal disc, lateral view; E. Organ of Tömösváry, lateral view; F. Epipharynx, ventral view; G. Gnathochilarium, ventral view; H. Sensory cones on inner palpi, anterior view; I. Mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipite. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure3" httpUri="https://binary.pensoft.net/fig/1249825">3 A, E</figureCitation>
|
||||
): Trapeziform, sparsely setose; setae on anterior part longer than posterior one; each seta located inside small pit. With 75–87 ommatidia (ocelli) in males and
|
||||
<quantity id="1656E3EB07F70AF20620517957AB58CD" metricMagnitude="0" metricUnit="m" metricValue="1.905" metricValueMax="1.9812" metricValueMin="1.8288" unit="in" value="75.0" valueMax="78.0" valueMin="72.0">72–78 in</quantity>
|
||||
females. Aberrant ommatidium present at brim of antennal groove. Organ of Tömösváry situated at brim near ocelli, separated from eye field. No sclerotised crest / ridge between antennal socket and eye field.
|
||||
</paragraph>
|
||||
<paragraph id="1D27F4ADD9997905A7CFB2EFC3A69866">
|
||||
<emphasis id="D6463C1F55A80A95EA9E3315B65752D1" bold="true" italics="true">Antennae</emphasis>
|
||||
(Figs
|
||||
<figureCitation id="4A5E58A1C8C2D8C05AB17E82CF347B61" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photograph of live specimens of Sphaerobelum nonghinensis sp. nov. A – F. Paratypes (MZKKU - MYR 0002); G, H. Habitat at the type locality (limestone). Scale bars: 10.0 mm." figureDoi="10.3897/zse.101.143073.figure2" httpUri="https://binary.pensoft.net/fig/1249824">2 B, C</figureCitation>
|
||||
,
|
||||
<figureCitation id="3AE7FD1BC29CFC11F0B2CF7D75F409E7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A. Head region, lateral view; B. Left antenna, anterior view; C. Antennal disc, anterior view; D. Apical cones on antennal disc, lateral view; E. Organ of Tömösváry, lateral view; F. Epipharynx, ventral view; G. Gnathochilarium, ventral view; H. Sensory cones on inner palpi, anterior view; I. Mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipite. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure3" httpUri="https://binary.pensoft.net/fig/1249825">3 A – D</figureCitation>
|
||||
): Short and stout, covered by long and dense setae; last antennomere reaching back to leg pair 2 or 3. Lengths of antennomeres 6> 5 = 4> 3> 2 = 1. Antennomere 6 apically rounded, oval; with sensilla basiconica apically. Apical disc flat, with 14–50 apical cones in male or
|
||||
<quantity id="DB4308BDA2808D1108BE8BFF4D73FFAE" metricMagnitude="-1" metricUnit="m" metricValue="8.001" metricValueMax="8.636" metricValueMin="7.366" unit="in" value="31.5" valueMax="34.0" valueMin="29.0">29–34 in</quantity>
|
||||
female.
|
||||
</paragraph>
|
||||
<paragraph id="409503E9188F9E93722AD70DC759579D">
|
||||
<emphasis id="928C8CD8324D8306E866DBFB134BC54F" bold="true" italics="true">Epipharynx</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="F8FCA3270CB405C28D65E6A416BAEE88" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A. Head region, lateral view; B. Left antenna, anterior view; C. Antennal disc, anterior view; D. Apical cones on antennal disc, lateral view; E. Organ of Tömösváry, lateral view; F. Epipharynx, ventral view; G. Gnathochilarium, ventral view; H. Sensory cones on inner palpi, anterior view; I. Mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipite. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure3" httpUri="https://binary.pensoft.net/fig/1249825">3 F</figureCitation>
|
||||
): With a regular central tooth (
|
||||
<abbrev id="ABBRID0EKCBG" xlink_title="central tooth">ct</abbrev>
|
||||
); laterally with numerous external teeth (
|
||||
<abbrev id="ABBRID0EOCBG" xlink_title="external tooth of epipharynx">et</abbrev>
|
||||
), short; inner area of each side with a row of crenate spines (
|
||||
<abbrev id="ABBRID0ESCBG" xlink_title="row of spine on epipharynx">rsp</abbrev>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="50588B6DB310AA380EED5C1A505C7A81">
|
||||
<emphasis id="FE14101F9278582311C6A1E094436E36" bold="true" italics="true">Gnathochilarium</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="4CB095D0DD626F42428A5FBEDC731139" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A. Head region, lateral view; B. Left antenna, anterior view; C. Antennal disc, anterior view; D. Apical cones on antennal disc, lateral view; E. Organ of Tömösváry, lateral view; F. Epipharynx, ventral view; G. Gnathochilarium, ventral view; H. Sensory cones on inner palpi, anterior view; I. Mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipite. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure3" httpUri="https://binary.pensoft.net/fig/1249825">3 G, H</figureCitation>
|
||||
): With long and dense setae. Lamellae linguales (
|
||||
<abbrev id="ABBRID0EBDBG" xlink_title="lamellae lingulales of gnathochilarium">ll</abbrev>
|
||||
) rectangular, flat apically. Central pads (
|
||||
<abbrev id="ABBRID0EFDBG" xlink_title="central pad of gnathochilarium">Cp</abbrev>
|
||||
) modified, with numerous sensory cones. Stipites (
|
||||
<abbrev id="ABBRID0EJDBG" xlink_title="stipites">st</abbrev>
|
||||
) large; located laterally to lamellae linguales. Mentum (
|
||||
<abbrev id="ABBRID0ENDBG" xlink_title="mentum">me</abbrev>
|
||||
) large, fused. Lateral palpi inconspicuous. Inner palpi (
|
||||
<abbrev id="ABBRID0ERDBG" xlink_title="inner palpi of gnathochilarium">ip</abbrev>
|
||||
) with sensory cones (
|
||||
<abbrev id="ABBRID0EVDBG" xlink_title="sensory cone">sc</abbrev>
|
||||
) arranged in two fields.
|
||||
</paragraph>
|
||||
<paragraph id="C0D8E01C90277A4E3BC2968AF982CE20">
|
||||
<emphasis id="5632E905896904DE29B9F7D440E09E4A" bold="true">
|
||||
<emphasis id="B270C00CE7851BCF7FDD725F8173B9EE" bold="true" italics="true">Mandibles</emphasis>
|
||||
(gnathal lobe)
|
||||
</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="5221E375C5C5B2540F00B8D513D0CBAF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A. Head region, lateral view; B. Left antenna, anterior view; C. Antennal disc, anterior view; D. Apical cones on antennal disc, lateral view; E. Organ of Tömösváry, lateral view; F. Epipharynx, ventral view; G. Gnathochilarium, ventral view; H. Sensory cones on inner palpi, anterior view; I. Mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipite. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure3" httpUri="https://binary.pensoft.net/fig/1249825">3 I</figureCitation>
|
||||
): With undivided external tooth (
|
||||
<abbrev id="ABBRID0EEEBG" xlink_title="external tooth of mandibular gnathal lobe">Et</abbrev>
|
||||
) and with conspicuous 3 - combed inner tooth (
|
||||
<abbrev id="ABBRID0EIEBG" xlink_title="3-combed inner tooth of mandible">3 it</abbrev>
|
||||
). With 4–5 rows of pectinate lamellae (
|
||||
<abbrev id="ABBRID0EMEBG" xlink_title="pectinate lamellae of mandible">pl</abbrev>
|
||||
). Inner area (
|
||||
<abbrev id="ABBRID0EQEBG" xlink_title="inner area of mandible">Ia</abbrev>
|
||||
) with group of tiny teeth. Molar plate (
|
||||
<abbrev id="ABBRID0EUEBG" xlink_title="molar plate of mandible">mp</abbrev>
|
||||
) flat, velvet-like; lacking a membranous fringe. Condylus (
|
||||
<abbrev id="ABBRID0EYEBG" xlink_title="condyles of mandible">co</abbrev>
|
||||
) conspicuous, apically with two distinct ridges.
|
||||
</paragraph>
|
||||
<paragraph id="A3F8C0EE7BE1BB5127E88B61DE2BDFC7">
|
||||
<emphasis id="13E2AF207A3160712FE62BFF67642F24" bold="true" italics="true">Tegument</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="ED56FEAD4DB71FE68056E6102A0680EB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photograph of live specimens of Sphaerobelum nonghinensis sp. nov. A – F. Paratypes (MZKKU - MYR 0002); G, H. Habitat at the type locality (limestone). Scale bars: 10.0 mm." figureDoi="10.3897/zse.101.143073.figure2" httpUri="https://binary.pensoft.net/fig/1249824">2 A – F</figureCitation>
|
||||
): Shiny; collum, thoracic shield, tergites, and anal shield sparsely microscopic setose; each seta located in an inconspicuous pit. Anterior margins of midbody tergite and anal shield with a few rows of long setae.
|
||||
</paragraph>
|
||||
<paragraph id="FD35E9FBEC088D457B366CFC7521489A">
|
||||
<emphasis id="81951EA5A1D8D86300DC028F7A20FCBA" bold="true" italics="true">Collum</emphasis>
|
||||
(Figs
|
||||
<figureCitation id="051B21E0D2F15CB1EC798C2363940E80" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photograph of live specimens of Sphaerobelum nonghinensis sp. nov. A – F. Paratypes (MZKKU - MYR 0002); G, H. Habitat at the type locality (limestone). Scale bars: 10.0 mm." figureDoi="10.3897/zse.101.143073.figure2" httpUri="https://binary.pensoft.net/fig/1249824">2 B</figureCitation>
|
||||
,
|
||||
<figureCitation id="17D2DB5DA8AF7263A07A2A583A71AB55" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A. Head region, lateral view; B. Left antenna, anterior view; C. Antennal disc, anterior view; D. Apical cones on antennal disc, lateral view; E. Organ of Tömösváry, lateral view; F. Epipharynx, ventral view; G. Gnathochilarium, ventral view; H. Sensory cones on inner palpi, anterior view; I. Mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipite. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure3" httpUri="https://binary.pensoft.net/fig/1249825">3 A</figureCitation>
|
||||
): Subsemicircular; tip of lateral margin obtuse.
|
||||
</paragraph>
|
||||
<paragraph id="3BFF0BE6A64704D30E895650839EE2CB">
|
||||
<emphasis id="F9BCDA81057EAFD1D0F49EC438CAFDA1" bold="true" italics="true">Thoracic shield</emphasis>
|
||||
: Anteriorly with shallow and large grooves separated by a long ridge. Groove and ridge smooth, without setae. Sloped towards groove laterally with 4 inconspicuous keels.
|
||||
</paragraph>
|
||||
<paragraph id="0E206D46125EDD57F084DE5F1703CCB0">
|
||||
<emphasis id="34A39C23FB15EB0C88B7E7708E19EFBB" bold="true" italics="true">Midbody tergite</emphasis>
|
||||
: With a row of oval impressions at anterior margin. Tips of midbody paratergites projecting posteroventrad.
|
||||
</paragraph>
|
||||
<paragraph id="5C4D4C7A2F9C5E94BB63265894508229">
|
||||
<emphasis id="1206C9E51ACF257E21D9D66C7819CE0F" bold="true" italics="true">Paratergites</emphasis>
|
||||
: Paratergites 3 and 4 slender, slightly attenuate; tip sharp. Paratergites 5–12 wide; tip obtuse.
|
||||
</paragraph>
|
||||
<paragraph id="F595FA80F6311634B3BED59479708484">
|
||||
<emphasis id="2918D12692932023CB42F305FC8A7C39" bold="true" italics="true">Anal shield</emphasis>
|
||||
: Large and well-rounded in female, slenderer in male. Underside with a single, short, black locking carina, as long as those of tergites, sometimes dotlike.
|
||||
</paragraph>
|
||||
<paragraph id="8AA4818CF58EE16B9F23A1612A20100A">
|
||||
<emphasis id="8EC57800288016F588CDBBE1069D885D" bold="true" italics="true">Endotergum of midbody tergite</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="899A22EAD78640A1CE101BF5D89F8872" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. SEM of endoterga of midbody tergites. A, B. Sphaerobelum nonghinensis sp. nov. (♂ paratype, MZKKU - MYR 0002); C, D. Sphaerobelum onyx sp. nov. (♂ paratype, MZKKU - MYR 0004); E, F. Sphaerobelum petramurum sp. nov. (♂ paratype, MZKKU - MYR 0006). Abbreviations: cp = cuticular impression, ia = inner area, ma = middle area, oa = outer area, pm = posterior margin. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure14" httpUri="https://binary.pensoft.net/fig/1249836">14 A, B</figureCitation>
|
||||
): Posterior margin (
|
||||
<abbrev id="ABBRID0EJIBG" xlink_title="posterior margin of endotergum">pm</abbrev>
|
||||
) flat, regular. Outer area (oa) wide; without setae. Marginal bristles arranged in 3–4 rows; tips of the longest bristles not protruding above posterior margin. Middle area (
|
||||
<abbrev id="ABBRID0ENIBG" xlink_title="middle area of endotergum">ma</abbrev>
|
||||
) with a single row of conspicuous, circular cuticular impressions (
|
||||
<abbrev id="ABBRID0ERIBG" xlink_title="cuticular impression of endotergum">cp</abbrev>
|
||||
); distance between impressions twice as long as individual diameter. Inner area (
|
||||
<abbrev id="ABBRID0EVIBG" xlink_title="inner area of endotergum">ia</abbrev>
|
||||
) without tubercles or setae.
|
||||
</paragraph>
|
||||
<paragraph id="A41030F491C83D00A1678F60BC344544">
|
||||
<emphasis id="0768419B09EB425DFA6D2C69C700FB85" bold="true">
|
||||
<emphasis id="E5CDD17189CF15EF176578A1B714C9A6" bold="true" italics="true">Pleurites</emphasis>
|
||||
(laterotergites)
|
||||
</emphasis>
|
||||
: First pleurite slender, boomeranglike; apical margin attenuated, strongly projecting into a wide tip. Pleurite 2 wider than the first one, projecting into obtuse tip. All remaining pleurites flat and broad, apical margin slightly extended.
|
||||
</paragraph>
|
||||
<paragraph id="D1E2B97DB1AC4345C1E0B56820B4CF3B">
|
||||
<emphasis id="873AD510829D41E164897CE18256B5C3" bold="true" italics="true">Subanal plate</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="378CFAF2C46147A78E9E0102A4DFF068" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Sphaerobelum nonghinensis sp. nov. A – E. ♂ paratype (MZKKU - MYR 0002); F, G. ♀ paratype (MZKKU - MYR 0002). A. Leg 11, arrow points to femoral ridge; B. Small spines on femur; C. Stigmatic plate 1; D. Coxae of legs 1; E. Coxae of legs 2 with gonopores; F. Leg 2 with vulvae; G. Subanal plate. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, Sp = apico-mesally spine of leg, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure4" httpUri="https://binary.pensoft.net/fig/1249826">4 G</figureCitation>
|
||||
): Densely setose; large and wide; well-rounded; apical margin slightly concave.
|
||||
</paragraph>
|
||||
<caption id="36858A272D2EA1DB935A78D6209056F0" ID-DOI="10.3897/zse.101.143073.figure4" ID-arpha="B3608A06-2531-5948-9CDF-D76BEB52E3AB" httpUri="https://binary.pensoft.net/fig/1249826" startId="F4">
|
||||
<paragraph id="C87C374EA704210EA4783A901EB2D400">
|
||||
<label id="45EA09551E3C57B2D43ED7BA38C7FB70">Figure 4.</label>
|
||||
</paragraph>
|
||||
<paragraph id="5AE2A17D2332B1AEF08589DC33502B0D">
|
||||
<abbrev id="ABBRID0EYJBG" xlink_title="scanning electron microscopy">SEM</abbrev>
|
||||
of
|
||||
<taxonomicName id="05BF92582EC6CA04ED57E930AA82F00F" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="090502642C4A97E7D75F9D8A9E48B1C2" italics="true">Sphaerobelum nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="E76E96DC154709094FE672E8FB7B7648" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="BC8A761AE3832FCEDE4F2444D0519E0F" bold="true">A – E.</emphasis>
|
||||
♂ paratype (
|
||||
<specimenCode id="CF6105307C712CD663AC0DCEB00DD04E">
|
||||
<named-content id="4B78A9907728F7093F4977B1A764B60B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0002
|
||||
</specimenCode>
|
||||
);
|
||||
<emphasis id="47713E8B9F99055966E2BA65DCB3AA3E" bold="true">F, G.</emphasis>
|
||||
♀ paratype (
|
||||
<specimenCode id="283E683DEC63E4DC03C29226FB9F72A5">
|
||||
<named-content id="CEB354C98231045FC8E924D3D3008838" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0002
|
||||
</specimenCode>
|
||||
).
|
||||
<emphasis id="06BF0A6EF6E7745A0000CC0306BDED5F" bold="true">A.</emphasis>
|
||||
Leg 11, arrow points to femoral ridge;
|
||||
<emphasis id="4593581BA957859FC932B059F439DF28" bold="true">B.</emphasis>
|
||||
Small spines on femur;
|
||||
<emphasis id="0E6C5C1301232F5236DB8ABA48AC23DE" bold="true">C.</emphasis>
|
||||
Stigmatic plate 1;
|
||||
<emphasis id="27E3CFC41DBFF9D814777BCE5F7840A9" bold="true">D.</emphasis>
|
||||
Coxae of legs 1;
|
||||
<emphasis id="E60FF86C5FB7F2EFE004D539B743B279" bold="true">E.</emphasis>
|
||||
Coxae of legs 2 with gonopores;
|
||||
<emphasis id="B4F17613AFF50370BFCC742FA8F23961" bold="true">F.</emphasis>
|
||||
Leg 2 with vulvae;
|
||||
<emphasis id="66042EAF434581454C4FF0B459D66F8A" bold="true">G.</emphasis>
|
||||
Subanal plate. Abbreviation:
|
||||
<abbrev id="ABBRID0EHLBG" xlink_title="bursa of vulva">bu</abbrev>
|
||||
= bursa,
|
||||
<abbrev id="ABBRID0ELLBG" xlink_title="coxa">cx</abbrev>
|
||||
= coxa,
|
||||
<abbrev id="ABBRID0EPLBG" xlink_title="femur">fe</abbrev>
|
||||
= femur,
|
||||
<abbrev id="ABBRID0ETLBG" xlink_title="operculum of vulva">op</abbrev>
|
||||
= operculum,
|
||||
<abbrev id="ABBRID0EXLBG" xlink_title="postfemur">po</abbrev>
|
||||
= postfemur,
|
||||
<abbrev id="ABBRID0E2LBG" xlink_title="prefemur">pre</abbrev>
|
||||
= prefemur,
|
||||
<abbrev id="ABBRID0E6LBG" xlink_title="apico-mesal spine of leg (tooth)">Sp</abbrev>
|
||||
= apico-mesally spine of leg,
|
||||
<abbrev id="ABBRID0EDMBG" xlink_title="tarsus">ta</abbrev>
|
||||
= tarsus,
|
||||
<abbrev id="ABBRID0EHMBG" xlink_title="tibia">ti</abbrev>
|
||||
= tibia. Scale bars in millimetres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="DBCF747E67DD995455AE62B995A45FCD">
|
||||
<emphasis id="D0C72C091A1DEFE7EB2E3E476577516F" bold="true" italics="true">Stigmatic plates</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="D84CE856D4699DF301D7B74905A7DF94" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Sphaerobelum nonghinensis sp. nov. A – E. ♂ paratype (MZKKU - MYR 0002); F, G. ♀ paratype (MZKKU - MYR 0002). A. Leg 11, arrow points to femoral ridge; B. Small spines on femur; C. Stigmatic plate 1; D. Coxae of legs 1; E. Coxae of legs 2 with gonopores; F. Leg 2 with vulvae; G. Subanal plate. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, Sp = apico-mesally spine of leg, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure4" httpUri="https://binary.pensoft.net/fig/1249826">4 C</figureCitation>
|
||||
): First stigmatic plate slender, apex well-rounded. Second stigmatic plate in both sexes not curved.
|
||||
</paragraph>
|
||||
<paragraph id="1BDDF4F8CEDC0D22DDD69AB0A5BDE390">
|
||||
<emphasis id="B2C0A09547A70FABB6D860731753AC17" bold="true" italics="true">Legs</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="AD01794D045610CC57DDBE544F3339EA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Sphaerobelum nonghinensis sp. nov. A – E. ♂ paratype (MZKKU - MYR 0002); F, G. ♀ paratype (MZKKU - MYR 0002). A. Leg 11, arrow points to femoral ridge; B. Small spines on femur; C. Stigmatic plate 1; D. Coxae of legs 1; E. Coxae of legs 2 with gonopores; F. Leg 2 with vulvae; G. Subanal plate. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, Sp = apico-mesally spine of leg, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure4" httpUri="https://binary.pensoft.net/fig/1249826">4 A – F</figureCitation>
|
||||
): All podomeres sparsely setose. Coxa (
|
||||
<abbrev id="ABBRID0EGNBG" xlink_title="coxa">cx</abbrev>
|
||||
) large; coxal process absent in first legs; legs 2–21 laterally with conspicuous coxal ridge (legs 2 of female slightly smaller than in male). Prefemur (
|
||||
<abbrev id="ABBRID0EKNBG" xlink_title="prefemur">pre</abbrev>
|
||||
) short; apico-mesally with inconspicuous projection. Femur (
|
||||
<abbrev id="ABBRID0EONBG" xlink_title="femur">fe</abbrev>
|
||||
) 1.5 times longer than wide, with long femoral ridge in all leg pairs; apico-mesally with dentate margin, carrying inconspicuous teeth (
|
||||
<abbrev id="ABBRID0ESNBG" xlink_title="apico-mesal spine of leg (tooth)">Sp</abbrev>
|
||||
). Postfemur (
|
||||
<abbrev id="ABBRID0EWNBG" xlink_title="postfemur">po</abbrev>
|
||||
) and tibia (
|
||||
<abbrev id="ABBRID0E1NBG" xlink_title="tibia">ti</abbrev>
|
||||
) short. Tarsus (
|
||||
<abbrev id="ABBRID0E5NBG" xlink_title="tarsus">ta</abbrev>
|
||||
) of midbody legs quite long, 3.4 times longer than wide, as long as length of femur + postfemur. First two leg pairs without apical spine; leg-pair 1 with 2 ventral spines; leg-pair 2 with 5 ventral spines; leg-pair 3 with 5 or 6 ventral spines and 1 apical spine; leg pairs 4–21 with 6 or 7 ventral spines and 1 apical spine. Claw normal, with a small notch at base.
|
||||
</paragraph>
|
||||
<paragraph id="46CE10B30C4C814616C4626F33239722">
|
||||
<emphasis id="615B3789CFFA83012BD38E3236EF003D" bold="true" italics="true">Male gonopore</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="3692E9EA32CD2AC3C1E5B8DEF8834D60" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Sphaerobelum nonghinensis sp. nov. A – E. ♂ paratype (MZKKU - MYR 0002); F, G. ♀ paratype (MZKKU - MYR 0002). A. Leg 11, arrow points to femoral ridge; B. Small spines on femur; C. Stigmatic plate 1; D. Coxae of legs 1; E. Coxae of legs 2 with gonopores; F. Leg 2 with vulvae; G. Subanal plate. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, Sp = apico-mesally spine of leg, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure4" httpUri="https://binary.pensoft.net/fig/1249826">4 E</figureCitation>
|
||||
): Small, triangular; covered by long setae.
|
||||
</paragraph>
|
||||
<paragraph id="AF9414F90E0523A32E0AEEA7A4E2E9AC">
|
||||
<emphasis id="7EEDBCB118151D873EBC7607B0552254" bold="true" italics="true">Anterior telopods</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="9F2FC27B3DACD37CA7A9325A1C5C1592" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A – C. Telopods, anterior, posterior, and ventral views, respectively; D. Right posterior telopod, anterior view; E – G. Anterior telopod, anterior, sublateral, and subventral views, respectively. Abbreviations: sp = telopoditomere 4 of posterior telopod’s spine. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure5" httpUri="https://binary.pensoft.net/fig/1249827">5 A, C, E – G</figureCitation>
|
||||
): First telopoditomere large, stout, apically widening. Telopoditomere 2 large, with projection as long as telopoditomere 3. Immovable finger (process of telopoditomere 2) short; clearly seen in posterior and ventral views; almost as long as telopoditomere 3; strongly curved; attenuate near tip; tip obtuse. Telopoditomere 3 indistinctly demarcated from telopoditomere 4 (with inconspicuous suture), both of equal length and cylindrical; telopoditomere 3 apically with a single spine and telopoditomere 4 with two spines or five spines on some specimens.
|
||||
</paragraph>
|
||||
<caption id="C5C96646AD21A067196F09FDC4DD4D7A" ID-DOI="10.3897/zse.101.143073.figure5" ID-arpha="8807C690-1C69-5C19-A5F5-1DB46AFE3A66" httpUri="https://binary.pensoft.net/fig/1249827" startId="F5">
|
||||
<paragraph id="533A222ADDBBAF3C5D77729A2C5127FA">
|
||||
<label id="CFA589A51EFBABA7DEB54DC1EEABC2B5">Figure 5.</label>
|
||||
</paragraph>
|
||||
<paragraph id="6BEC8685A86CA27A6FA3BC9E7F7AA7F8">
|
||||
<abbrev id="ABBRID0EAPBG" xlink_title="scanning electron microscopy">SEM</abbrev>
|
||||
of
|
||||
<taxonomicName id="3DC993BE0E2EF2A20C2E0A153BE2EF14" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="423EBFD08A461AE64F7AF2BC8F0FD915" italics="true">Sphaerobelum nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="13226CF018818F40157D92FCEB81F096" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, ♂ paratype (
|
||||
<specimenCode id="326E2CD7C04C010DABC0E7B096E9B4EE">
|
||||
<named-content id="40DA7D89ACF0B6EEA9BA3B9B051CCDB9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology, Thailand">MZKKU</named-content>
|
||||
-MYR 0002
|
||||
</specimenCode>
|
||||
).
|
||||
<emphasis id="0DED5AB4DCC4B8CDAEC2BE1634093378" bold="true">A – C.</emphasis>
|
||||
Telopods, anterior, posterior, and ventral views, respectively;
|
||||
<emphasis id="D5CA9942782C8932D60965215AE093BB" bold="true">D.</emphasis>
|
||||
Right posterior telopod, anterior view;
|
||||
<emphasis id="04F18EBB92E457165DDC2672C6FD4C8B" bold="true">E – G.</emphasis>
|
||||
Anterior telopod, anterior, sublateral, and subventral views, respectively. Abbreviations:
|
||||
<abbrev id="ABBRID0E5PBG" xlink_title="spine on telopoditomere 4 of posterior telopod">sp</abbrev>
|
||||
= telopoditomere 4 of posterior telopod’s spine. Scale bars in millimetres.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="AC5483EA0DE8A636CAFF120DEA056421">
|
||||
<emphasis id="614CC254034F9D0C7F93C307690D5B46" bold="true" italics="true">Posterior telopods</emphasis>
|
||||
(Figs
|
||||
<figureCitation id="65D9D9FDF67C4370DCA6249D9B7DD6B7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM of Sphaerobelum nonghinensis sp. nov., ♂ paratype (MZKKU - MYR 0002). A – C. Telopods, anterior, posterior, and ventral views, respectively; D. Right posterior telopod, anterior view; E – G. Anterior telopod, anterior, sublateral, and subventral views, respectively. Abbreviations: sp = telopoditomere 4 of posterior telopod’s spine. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure5" httpUri="https://binary.pensoft.net/fig/1249827">5 A – D</figureCitation>
|
||||
,
|
||||
<figureCitation id="2D330F73D810F720D6DBB15A961CAAC7" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Line drawings of posterior telopods of Sphaerobelum species in Thailand. A, B. Sphaerobelum aesculus; C, D. Sphaerobelum meridionalis, modified from Bhansali & Wesener, 2022; E, F. Sphaerobelum nonghinensis sp. nov.; G, H. Sphaerobelum onyx sp. nov.; I, J. Sphaerobelum petramurum sp. nov.; K, L. Sphaerobelum truncatum; M, N. Sphaerobelum turcosa. A, B, E – N. Right posterior telopods; C, D. Left posterior telopods. A, D, E, G, I, K, M. Posterior view; B, C, F, H, J, L, N. Anterior view. Abbreviations: cr-t = crenulated teeth, ml = membranous lobe, sp = spine on telopoditomere 4 of posterior telopod. Scale bar: 0.5 millimetres." figureDoi="10.3897/zse.101.143073.figure15" httpUri="https://binary.pensoft.net/fig/1249837">15 E, F</figureCitation>
|
||||
): With 4 telopoditomeres. First telopoditomere rectangular, longer than wide. Telopoditomere 2 large. Immovable finger (process of telopoditomere 2) long, massive, hammerlike, 2 times longer than wide, with a characteristic, apically extremely swollen, round, inner margin. With single spine; with thin dark spot near apex; tip
|
||||
<emphasis id="573A5C65EF6D090A666B25A0EC11AEDC" italics="true">in situ</emphasis>
|
||||
, curving ventrad; antero-apical margin a bit collapsed to accommodate tip of telopoditomere 4. Telopoditomere 3 stout; 2 times longer than wide, with single spine at excavate inner margin. Telopoditomere 4 relatively short, half as long as telopoditomere 3, strongly tapering toward apex; with two spines; immediately curved midway by forming a digitiform tip. Telopoditomeres 1 and
|
||||
<quantity id="E2A4A1C5ABD3E50933A0E436715E72BC" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
|
||||
anterior view covered by sparse setae, in posterior view glabrous. Telopoditomeres 3 and 4 glabrous. Inner horns with sharp tips, directed posteriad.
|
||||
</paragraph>
|
||||
<paragraph id="0B04FF17DB51F9119529D6441658A1AB">
|
||||
<emphasis id="8FC634947E790A717598E1FDA09D12D8" bold="true" italics="true">Female sexual characters</emphasis>
|
||||
(Fig.
|
||||
<figureCitation id="19279C9B74FBCC1CD5BC19DFB25D0BBF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. SEM of Sphaerobelum nonghinensis sp. nov. A – E. ♂ paratype (MZKKU - MYR 0002); F, G. ♀ paratype (MZKKU - MYR 0002). A. Leg 11, arrow points to femoral ridge; B. Small spines on femur; C. Stigmatic plate 1; D. Coxae of legs 1; E. Coxae of legs 2 with gonopores; F. Leg 2 with vulvae; G. Subanal plate. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, Sp = apico-mesally spine of leg, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zse.101.143073.figure4" httpUri="https://binary.pensoft.net/fig/1249826">4 F</figureCitation>
|
||||
): Vulva slightly short and slender; sparsely setose; covering almost 1 / 3 of coxa; extending mesally near base of prefemur. Operculum (
|
||||
<abbrev id="ABBRID0EDRBG" xlink_title="operculum of vulva">op</abbrev>
|
||||
) round, slender, mesal margin protruding into a short, well-rounded lobe. Bursa (
|
||||
<abbrev id="ABBRID0EHRBG" xlink_title="bursa of vulva">bu</abbrev>
|
||||
) relatively long, demarcated from operculum by a triangular groove.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ELRBG" type="etymology">
|
||||
<paragraph id="4A061C5E11FF73B07A8945161FA056D4">
|
||||
<heading id="3089DA789282199A487BCB432493D491" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="CFC4220088F2278BFE6677864AB8025A">Named after Nong Hin District, where all specimens were collected. Adjective.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EQRBG" type="distribution">
|
||||
<paragraph id="EE37A2B8D6E209F9E259057B0B4ACFA3">
|
||||
<heading id="96BD17DAB92B44D6EC399821D097737B" reason="title">Distribution and habitat</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BA5C775DBA1AE2EDA91B186E57727C9F">
|
||||
(Fig.
|
||||
<figureCitation id="F1CD421343131B05BB30BC8B44FF23CD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photograph of live specimens of Sphaerobelum nonghinensis sp. nov. A – F. Paratypes (MZKKU - MYR 0002); G, H. Habitat at the type locality (limestone). Scale bars: 10.0 mm." figureDoi="10.3897/zse.101.143073.figure2" httpUri="https://binary.pensoft.net/fig/1249824">2 G, H</figureCitation>
|
||||
).
|
||||
<taxonomicName id="230BC9CEB618ACB07BC765780FF327C8" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="4B2CC6BE2A4F4F9AF91D9A9041C803CC" italics="true">Sphaerobelum nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="7A83573F8D1DFA6CAFB22227352A2492" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is currently known only from the type locality in Nong Hin District,
|
||||
<collectingRegion id="419337EB3C5775700A2E0A1CB869594D" country="Thailand" name="Loei">Loei Province</collectingRegion>
|
||||
. This new species was observed during the daytime in an evergreen forest within limestone habitat. Individuals typically conceal themselves beneath thick layers of leaf litter and decayed logs on rocky surfaces or in crevices, highlighting their adaptation to this unique environment.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EJSBG" type="remarks">
|
||||
<paragraph id="A5BACFD7220C3457EA41B328BBBE8956">
|
||||
<heading id="CCD2A53911ED33E583328FF46EDBE199" reason="title">Remarks.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="530DDDA919B2ED53AC723DD5CAABB34A">
|
||||
Females of
|
||||
<taxonomicName id="53C6B49D8122E9E81E89B1CFE48FF4D3" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="nonghinensis" status="sp. nov.">
|
||||
<emphasis id="25D325A42A69FFA907D5E3B8EC7900E1" italics="true">S. nonghinensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="03B301AF8555D80760278222D19C414A" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
are generally similar in size to males. This new species features a striking black body colour (but with contrasting orange markings on the thoracic shield and anal shield) and a telopod shape comparable to that of
|
||||
<taxonomicName id="9B33C0808A4E47D3A35D41D19C03960C" authorityName="Srikampha & Srisonchai" authorityYear="2025" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
|
||||
<emphasis id="6B29C542E8206B824A15DF242140C5C0" italics="true">S. onyx</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="5C900427986E98F817F17293654C033F" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
However, genetic analysis reveals a notable difference, with a
|
||||
<abbrev id="ABBRID0ENTBG" xlink_title="mitochondrial cytochrome c oxidase subunit 1">COI</abbrev>
|
||||
genetic divergence of 13.24 % to 14.59 %, indicating a significant level of separation between the two species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue