diff --git a/data/03/D2/38/03D2383B8500C33AD9B2A015A99276B5.xml b/data/03/D2/38/03D2383B8500C33AD9B2A015A99276B5.xml new file mode 100644 index 00000000000..4d78eb876b4 --- /dev/null +++ b/data/03/D2/38/03D2383B8500C33AD9B2A015A99276B5.xml @@ -0,0 +1,457 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips leai +Moulton + + + + + + + +( +Figs 9–11 +, +93–101 +) + + + + + + + +Ecacanthothrips leai +Moulton, 1947: 176 + + +. + + + + +E. leai + +was described based on one damaged female taken from +Kuala Lumpur +, Peninsular +Malaysia +. +Palmer and Mound (1978) +recorded under this name from Java, New +Guinea +, the +Solomon Islands +, and the Ryukyu Islands in addition to Peninsular +Malaysia +. Subsequently, however, +Okajima (1983) +discriminated the specimens from the Ryukyus, southern +Japan +, as a distinct species, + +E. moundi +. + +This species is included in the + +inarmatus + +-group, and exhibits conspicuous variation. A long series of females and males of + +leai + +listed below have been examined from several sites of Peninsular +Malaysia +, at least one site is near the +type +locality, and its intraspecific variation associated with allometric growth has been understood in detail, although geographical variation has not been clarified yet. According to +Palmer and Mound (1978) +, this species shows somewhat confused geographical variation in the length of head, the comparative length of anteromarginal and anteroangular setae in large male, and in the colour of tibiae. However, it is difficult to know whether these local populations contain additional distinct species or not. Furthermore, even within Peninsular +Malaysia +, there are slight variations between local populations in the colour of antennae and tibiae. The specimens from mountainous areas have the antennal segment III somewhat darker, yellowish brown, and the tibiae scarcely shaded with brown, whereas the specimens from lowlands have the antennal segment III paler, brownish yellow, and the tibiae usually clear yellow. +Three females +and +seven males +from +Bali +Is., +Indonesia +, have mid and hind tibiae brownish. +Nine females +and +seven males +from Borneo have the heads rather distinctly reticulate. +Two males +from +Sumatra +and +Sulawesi +, +Indonesia +, listed in the doubtful specimens below have longer head proportions, which are more than 1.5 times as long as wide, and +one female +and +four males +from mountainous area of +Bali +Is. are exceptionally small, +1.8–2.4 mm +. The difference between + +leai + +and + +moundi + +is discussed under + +moundi + +. + + + + +Diagnosis +. Body brown to dark brown, all tibiae yellow, antennal segments IV–VI with base yellowish. +Female +. Body length +2.5–3.6mm +. Head ( +Fig. 93 +) 1.2–1.3 times as long as wide, dorsal surface reticulate, but the reticles weak along midline, cheeks distinctly constricted just behind eyes. Antennae ( +Fig. 95 +) about 2.0 times as long as head; segment VIII distinctly constricted basally, pedicellate; segment V without distinct apical neck, shorter than segment IV. Pronotal pa the longest, aa and am setae subequal in small female, aa and ml elongate in large female. Fore wing with 11–19 duplicated cilia. Pelta ( +Fig. 97 +) bell-shaped. Tube ( +Fig. 100 +) 0.55–0.56 times as long as head. +Male +. Body length +2.2–3.2mm +. Head ( +Fig. 94 +) 1.23–1.40 times as long as wide, dorsal surface scarcely sculptured; cheeks each with two or three stout setae in large male. Pronotal aa, ml and pa setae elongate, but am reduced in large male. Fore tibia with sub-basal inner tubercle in large male. Fore wing with 11–22 duplicated cilia. Tube 0.51–0.54 times as long as head. + + +Specimens examined. + +Peninsular + +Malaysia + +, +Gnung Tengkolok +, +9 females +and +6 males +, on dead leaves and branches, + +27.v.1983 + +, +T +. +Senoh +; + + +Cameron Highland +, +Tanah Rata +, +2 females +, on dead branches, + +24.vii.1976 + +, SO, + + +same locality above, +1 female +and +3 males +, + +8.v.1981 + +, +W. Suzuki +; + + +Cameron Highland +, nr. +Brinchang +, +79 females +and +48 males +, on dead leaves and branches, + +25–27.viii.1990 + +, TN & SO; + + +Cameron Highland +, nr. +Tanah Rata +, +Robinson +water fall, +9 females +and +1 male +, on dead leaves and branches, + +28.viii.1990 + +, TN & SO; + + +Cameron Highland +, nr. +Tanah Rata +, +5 females +and +1 male +, on dead leaves and branches, + +29.viii.1990 + +, TN & SO; + + +Cameron Highland +, foot of +Gnung Jasar +, +17 females +and +2 males +, on dead leaves and branches, + +29.viii.1990 + +, TN & SO; + + +about +20km +N from +Kuala Lumpur +, +Templer Park +, +33 females +and +28 males +, + +12–15.viii.1990 + +, TN & SO; + + +Fraser’s Hill +, +1 female +and +5 males +, on dead leaves and branches, + +13–14.ix.1990 + +, TN & SO. + + + +Borneo + +, +Sabah +, about +3km +N from +Kundasang +, +9 females +and +7 males +, on dead leaves and branches, + +6.ix.1990 + +, TN & SO + +. + + +Indonesia + +, E. +Java +, +Mt Arjuna +, + +1400–1600m + +alt., +1 male +, on dead leaves, + +19.iv.1981 + +, +T +. +Senoh +; + + +Bali +Is. +, +Buleleng +, +Yehketipat +, +3 females +and +7 males +, + +8.iii.2005 + +, SO + +. + + +Doubtful specimens. + + +Indonesia + +, +Sumatra +, nr. +Toba lake +, +1 female +and +1 male +, + +4.v.1990 + +. +H. Matsumoto + +; + +Sulawesi +, +Talaud +, +Karakelong Is. +, +Ponto +, +1 male +, + +30.vii.1979 + +, +N. Kashiwai + +; + +Bali +Is., +Candi Kuning +, alt. about + +1200m + +, +1 female +and +4 males +, + +26.vii.1984 + +, SO + +. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B8501C338D9B2A67BAA7570CB.xml b/data/03/D2/38/03D2383B8501C338D9B2A67BAA7570CB.xml new file mode 100644 index 00000000000..2deb089ca46 --- /dev/null +++ b/data/03/D2/38/03D2383B8501C338D9B2A67BAA7570CB.xml @@ -0,0 +1,119 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips spinipes +(Bagnall) + + + + + + + + + + +Phloeothrips spinipes +Bagnall, 1908: 195–196 + + +. + + + +This species was described based on the unique +holotype +female collected from New +Guinea +, and has three junior synonyms. Subsequently, it was recorded from the Kei Islands ( +Priesner 1930 +: described as synonymous species, + +E. bagnalli + +) and the +Solomon Islands +( +Mound 1970 +). It is presumably included in the + +inarmatus + +-group. According to the literature ( +Mound 1970 +; +Palmer & Mound 1978 +), it may be very similar to + +E. leai + +, and the fore femur bears a series of tuberculate setae at least in medium to large sized individuals, the mid and hind femora bear a series of stout setae with blunt or dilated apices, and the pelta is triangular with lateral wider reticulation. However, even in large individuals, + +leai + +does not have these character states. In + +leai + +, the fore femur has no tuberculate setae, the mid and hind femora bear a series of short and pointed setae and the pelta is bell-shaped. Unfortunately, this species has not been available for this study. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B8501C33BD9B2A1C4AA967582.xml b/data/03/D2/38/03D2383B8501C33BD9B2A1C4AA967582.xml new file mode 100644 index 00000000000..6e52b3d1a51 --- /dev/null +++ b/data/03/D2/38/03D2383B8501C33BD9B2A1C4AA967582.xml @@ -0,0 +1,249 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips moundi +Okajima + + + + + + + +( +Figs 12–13 +, +102–110 +) + + + + + + + +Ecacanthothrips moundi +Okajima, 2006: 245–247 + + +. + + + + +E. moundi + +was described from the Okinawa-Amami group of Islands, the Ryukyu Islands, southern +Japan +, and included in the + +inarmatus + +-group. It is somewhat similar to + +leai + +, but can be distinguished by the following features: body larger, +3.2–3.9mm +in female, +2.5–4.3mm +in male; head ( +Figs 102 & 103 +) longer, 1.3–1.5 times as long as wide in female, 1.50–1.95 times in male (about 1.3 times as long as wide even in large male in + +leai + +); cheeks weakly constricted behind eyes; antennal segment V slender, with a distinct apical neck. Moreover, in large male, pronotal anteromarginal setae are elongated in + +moundi + +, whereas they are not elongate in + +leai + +. The colour of mid and hind tibiae is variable in the +type +series of + +moundi + +, usually brownish, but often yellowish in the male. A series of females and males listed below collected from northern +Vietnam +could probably be identified as this species, but they have some small differences. They have the mid and hind tibiae clear yellow and the antennal segment V longer and more slender, usually longer than segment IV. However, most other character states are indistinguishable from the +type +series. Another species described from +Japan +, + +E. inarmatus + +, is also very similar to this species, but + +moundi + +can be distinguished by the following features: body uniformly dark brown; antennal segment III darker, dark brown; hind tibiae variable in colour, yellow to brown, at least not darker than femora; pronotal anteromarginal setae elongate in large male, almost as long as anteroangulars or a little longer ( + +inarmatus + +also has elongate anteromarginal setae in large male, but much shorter than anteroangulars). In contrast to the uniformly dark brown body of + +moundi +, +inarmatus + +is slightly bicoloured with head, thorax and anterior abdominal segments paler, tinged with orange yellow when alive (see p. +53 in +Okajima & Masumoto, 2022 +). These two species have allopatric distribution, + +moundi + +known only from the subtropical Ryukyu Islands, whereas + +inarmatus + +ocurs in the temperate region ( +Hokkaido +, Honshu, Shikoku and Kyushu, and those dependent islands) and +Guangdong +, +China +( +Han 1997 +). One small male listed below from +Thailand +is very similar to the smallest male from +Vietnam +, but it has intermediate antennal segments somewhat shorter. + + +Specimens examined. + + +Japan + +, + +Ryukyu Islands + +, holotype female and paratype +3 females +and +2 males +from + + +Okinawa +Is. +, + + +1 female +and +1 male +from +Amami-ohshima Is. +(data described in +Okajima, 2006 +) + +. + + +Vietnam + +, +Lao Cai Province +, +Sa Pa +, +13 females +and +8 males +, on dead leaves and branches, + +6.viii.2000 + +, SO + +. + + +Thailand + +, +Doi Suthep +, + +1100m + +alt., +1 male +, on dead leaves, + +11.vii.1976 + +, SO + +. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B8501C33BD9B2A442AD017795.xml b/data/03/D2/38/03D2383B8501C33BD9B2A442AD017795.xml new file mode 100644 index 00000000000..64409b06e95 --- /dev/null +++ b/data/03/D2/38/03D2383B8501C33BD9B2A442AD017795.xml @@ -0,0 +1,139 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips nigellus +Okajima + + + + + + + +( +Figs 14 +, +111–119 +) + + + + + + + +Ecacanthothrips nigellus +Okajima, 1983: 59–61 + + +. + + + +This species was described from northern +Thailand +based on +two females +, and is included in the + +inarmatus + +-group. It is very similar to + +leai + +, but has the tibiae and antennal segments darker ( +Fig. 14 +). Moreover, antennal segments III and IV ( +Figs 113–115 +) are heavier than those of + +leai + +, and segment III bears extraordinary dark and thick sense cones. The inner side of antennal segment III is distinctly swollen at basal half and segment IV has thicker basal stem. The basal width of this segment is almost as wide as the apical width in + +nigellus + +, though the stems of these segments are slender in + +leai + +, the basal width being much narrower than apical width. Unfortunately, the intraspecific variation of this species is not yet understood, because only two large females are known so far. + + + + +Specimens examined. + + +Thailand + +, +holotype +female and a + +paratype +female (data described in +Okajima, 1983 +). + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B8502C338D9B2A089AA377469.xml b/data/03/D2/38/03D2383B8502C338D9B2A089AA377469.xml new file mode 100644 index 00000000000..151586e0ef1 --- /dev/null +++ b/data/03/D2/38/03D2383B8502C338D9B2A089AA377469.xml @@ -0,0 +1,267 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips tenuicornis +Okajima + + + + + + + +( +Figs 15–16 +, +120–126 +) + + + + + + + +Ecacanthothrips tenuicornis +Okajima, 1983: 63–65 + + +. + + + + +E. tenuicornis + +was originally described from Luzon Is., the +Philippines +based on +eight females +and +five males +together with non-paratypic +three females +taken from Mindanao Is. This species could well be included in the + +inarmatus + +-group and is somewhat similar to + +leai + +, although it has smaller body, +1.8–2.6mm +in the female, +1.8–2.2mm +in the male. Moreover, the head of + +tenuicornis + +( +Figs 120 & 121 +) is shorter, about 1.1 times as long as wide or shorter, although the head of + +leai + +( +Figs 93 & 94 +) is longer, usually 1.2–1.4 times as long as wide even in small female. In contrast to + +inarmatus + +and + +moundi + +, the pronotal anteromarginal setae in this species are not elongate even in large males. Furthermore, + +tenuicornis + +is apparently most similar to + +brevicornis + +described above, but can be discriminated by the antennal structure mentioned in the key above. However, its intraspecific variation derived from allometric growth is not understood in detail. Females and males listed in the doubtful specimens collected from Peninsular +Malaysia +and +Thailand +are very similar to this species, but have somewhat larger bodies, +2.5–2.9mm +in female and 2.0– +2.6mm +in male. + + + + +Specimens examined. +The + + +Philippines + +, +holotype +female and + +paratype +females and males from Luzon Is.; non-paratypic +3 females +from Mindanao Is. (data described in +Okajima, 1983 +). + + +Doubtful specimens. + +Peninsular +Malaysia + +, about +20km +N from + +Kuala Lumpur +, +Templer Park +, +2 males +, on dead leaves, + +15.viii.1990 + +, TN & SO + +. + + +Thailand + +, +Phuket +Is. +, nr. +Tone Sai +water fall, +2 females +and +5 males +, on dead leaves and branches, 11–14, + +ix.1992 + +, TN & SO + +; + +Phuket +Is. +, +Rang Hill +, +3 females +and +2 males +, on dead leaves and branches, + +14.ix.1992 + +, TN & SO; nr + +, + +Chiang Mai +, foot of +Doi Pui +, on dead leaves and branches, +3 males +, + +1.ix.1991 + +, +1 male +, + +24.viii.1992 + +, TN & SO + +; +Chiang Mai +, farm of + +Chiang Mai +University +, +1 female +and +3 males +, on dead leaves and branches, + +26.viii.1992 + +, TN & SO + +. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B8502C339D9B2A42EACA77003.xml b/data/03/D2/38/03D2383B8502C339D9B2A42EACA77003.xml new file mode 100644 index 00000000000..333e267884e --- /dev/null +++ b/data/03/D2/38/03D2383B8502C339D9B2A42EACA77003.xml @@ -0,0 +1,216 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips tibialis +(Ashmead) + + + + + + + +( +Figs 17–18 +, +127–135 +) + + + + + + + +Idolothrips tibialis +Ashmead, 1905: 20 + + +. + + + +This species was described from Luzon Island, the +Philippines +, based on a single damaged female, and is widely distributed mainly in the tropical and subtropical regions in the Old World. It exhibits highly confusing intraspecific variation which include not only size related variation but also geographical variation, and has more than 10 junior synonyms ( +Palmer & Mound 1978 +). Especially, it has geographical variation between samples from different sites in the colour of tibiae and intermediate antennal segments ( +Okajima 1983 +; +Palmer & Mound 1978 +). However, the structural characteristics of + +tibialis + +are relatively stable despite the size related variation, and it is not difficult to distinguish from two closely related species, + +andrei + +and + +claricornis + +. Although + +andrei + +and + +claricornis + +have antennal segments III and IV yellowish ( +Figs 58 +& +78 +), the postocular cheek setae pointed in male, the mid and hind femora each with only one pale stout seta with blunt or dilated apex ( +Figs 62 +& +82 +) and the pelta bell-shaped ( +Figs 60 +& +80 +), + +tibialis + +has antennal segments III and IV brownish ( +Fig. 129 +), the postocular cheek setae dilated in male, the mid and hind femora each with two or three (often four or more) dark and stout setae with blunt or pointed apices ( +Fig. 133 +) and the pelta triangular ( +Fig. 132 +). Moreover, + +tibialis + +has some differences from both + +andrei + +and + +claricornis + +in the variation associated with allometric growth. In large males, + +tibialis + +has the pronotal anteromarginal setae reduced and a sub-apical inner tubercle on the fore tibia, whereas + +andrei + +and + +claricornis + +have the pronotal anteromarginal setae elongate and no tibial sub-apical tubercle. + + + + +Specimens examined +(detailed data are omitted). Numerous females and males from the following localities: + +Indonesia + +( +Bali +, +Java +, Lombok, +Sulawesi +); + +Peninsular +Malaysia + +; + +Singapore + +; +Borneo +(Kalimantan, Sabah); + +Thailand + +; + +Vietnam + +; + +the +Philippines + +(Mindanao, Luzon); + +Taiwan + +; + +Japan + +(the Ryukyu Islands). + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B8506C33DD9B2A338ADEA73C0.xml b/data/03/D2/38/03D2383B8506C33DD9B2A338ADEA73C0.xml new file mode 100644 index 00000000000..3c17b4bc3f8 --- /dev/null +++ b/data/03/D2/38/03D2383B8506C33DD9B2A338ADEA73C0.xml @@ -0,0 +1,422 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + +Key to + +Hoplandrothrips + +species from Southeast Asia and +Taiwan + + + + + + + +1. Antennal segment III with four or five sense cones........................................................... 2 + + +- Antennal segment III with two or three sense cones.......................................................... 4 + + + + + +2. Antennal segment III with five (rarely six) sense cones....................................... + + +quinqueconus +sp. n. + + + + + +- Antennal segment III with four sense cones................................................................. 3 + + + + + +3. Head 1.15–1.30 times as long as wide in both sexes, sculptured entirely with distinct polygonal reticulation; antennal segment III and V subequal in length, segment V without apical neck; pelta with rather slender lateral wings; tibiae yellow; small to medium sized species, +1.8–2.7mm +long............................................................... + +flavipes + + + + + +- Head about 1.4–1.6 times as long as wide in female ( +Fig. 187 +), 1.5–1.9 times in male ( +Fig. 188 +), sculptured with fine reticulation, but reticles weaker behind posterior ocelli; antennal segment V longer than segment III, with distinct apical neck ( +Fig. 189 +); pelta with rather thick lateral wings ( +Fig. 193 +); mid and hind tibiae brownish; large sized species, +2.7–4.4mm +long................................................................................................ + + +formosae +sp. n. + + + + + + + +4. Head elongate, about 1.4 times as long as wide or longer, frequently with constriction just behind eyes; maxillary stylets rather short, retracted to basal 1/3–1/2 of head capsule, not reaching postocular setae..................................... 5 + + +- Head shorter than 1.3 times as long as wide, often as long as wide, without distinct constriction behind eyes; maxillary stylets usually retracted to eyes, at least reaching postocular setae..................................................... 8 + + + + + +5. Prosternal basantra present ( +Figs 167 +& +239 +); body bicoloured (yellow and brown), at least anterior abdominal segment paler than pterothorax...................................................................................... 6 + + + + +- Prosternal basantra absent ( +Fig. 201 +); body uniformly brown to dark brown....................................... 7 + + + + + + +6. Body largely yellow ( +Figs 26 & 27 +); head, prothorax, abdominal segments II–VII and legs yellow; pterothorax brownish, abdominal segments VIII–IX and tube brown................................................. + + +basantratus +sp. n. + + + + + + +- Body largely brown to dark brown ( +Figs 43 & 44 +); abdominal segments II–VI yellow to brown, gradually darkened exteriorly, at least segments II–III yellowish, but these segments sometimes darker in male.................... + + +samirseni +comb. n. + + + + + + + + +7. Antennal segment III yellow, about 2.0 times as long as wide ( +Fig. 171 +); antennal segment VIII distinctly constricted at base, pedicellate; sub-basal wing setae S3 expanded ( +Fig. 175 +)......................................... + + +flavicornis +sp. n. + + + + + + +- Antennal segment III brown with extreme base yellowish, 1.5–1.7 times as long as wide ( +Fig. 199 +); antennal segment VIII scarcely constricted at base; sub-basal wing setae S3 pointed, at least not distinctly expanded ( +Fig. 203 +)... + + +graminicola +sp. n. + + + + + + + +8. Body bicoloured brown and yellow, or largely yellow........................................................ 9 + + +- Body uniformly brown to dark brown.................................................................... 11 + + + + + +9. Antennal segment III with two sense cones; body bicolourous, head and prothorax brown, at least darker than pterothorax and abdomen except for brown tube................................................................... + +coloratus + + + + +- Antennal segment III with three sense cones; body largely yellow, head and thorax largely yellow.................... 10 + + + + + +10. Dorsal surface of head almost smooth ( +Fig. 136 +); posterior ocelli not in contact with eyes; antennal segment VIII not distinctly constricted at base ( +Fig. 140 +); maxillary stylets retracted to eyes; fore tarsus with a small tooth in female ( +Fig. 137 +); fore femur and tibia unarmed even in large male ( +Fig. 139 +); postocular setae about 1/2 length of eyes or shorter, rather close together in female; mesopresternum boat-shaped, not divided............................................. + + +adraneoides +sp. n. + + + + + + +- Dorsal surface of head fully sculptured with polygonal reticulation ( +Fig. 144 +), but very weak; posterior ocelli in contact with eyes; antennal segment VIII constricted at base, pedicellate ( +Fig. 150 +); maxillary stylets retracted to postocular setae, usually not reaching eyes; fore tarsus unarmed in female ( +Fig 145 +); fore femur and tibiae armed with inner apical tubercles at least in large male ( +Fig. 147 +); postocular setae much longer than 1/2 length of eyes, not close together in female; mesopresternum divided into two lateral large triangles and a small median sclerite ( +Fig. 149 +)........................... + + +aseanae +sp. n. + + + + + + + +11. Head almost as long as wide, with bulged cheeks........................................................... 12 + + +- Head longer than 1.1 times as long as wide................................................................ 14 + + + + + +12. Antennal segment III slightly longer than segment IV ( +Fig. 257 +); cheeks finely tuberculate at least in female ( +Fig. 255 +); abdominal sternite VIII without pore plate in male................................................ + + +sulawesi +sp. n. + + + + + +- Antennal segment III as long as segment IV, or shorter; cheeks serrate, not tuberculate; abdominal sternite VIII with a median weak pore plate in male............................................................................... 13 + + + + + +13. Ocellar region almost smooth, without reticulation; eyes slightly larger on ventral surface ( +Figs 177 & 178 +).... + + +floresi +sp. n. + + + + + + +- Ocellar region distinctly sculptured with fine reticulation; eyes not larger on ventral surface ( +Figs 215 & 216 +).... + +obesametae + + + + + + + +14. Mesopresternum boat-shaped, not divided ( +Fig. 209 +); head and pronotum smooth ( +Fig. 205 +); fore femur and tibia unarmed even in large male ( +Fig. 208 +); postocular setae rather close together in female.............................. + + +laurencei +sp. n. + + + + + + +- Mesopresternum divided into three plates, lateral two triangles and median small plate; head and pronotum sculptured with reticulation; fore femur and tibia armed with tubercles in large male (male of + +thailandicus + +is unknown); postocular setae not close together in female............................................................................... 15 + + + + + + +15. Antennal segment VIII distinctly constricted at base, pedicellate ( +Fig. 156 +); all tibiae yellow; postocular setae much shorter than eyes ( +Fig. 154 +)............................................................................. + +asianus + +sp. n +. + + + + + + +- Antennal segment VIII not constricted at base, cone-shaped (cf. +Fig. 251 +); fore tibiae brownish, mid and hind tibiae dark brown, or at least brownish medially; postocular setae a little shorter than eyes.......................................... 16 + + + + + + +16. Head about 1.2 times as long as wide ( +Figs 244 & 245 +); antennal segment III rather slender, longer than 2.0 times as long as wide, longer than segment IV ( +Fig. 250 +); mouth cone not reaching prospinasternum ( +Fig. 252 +); sense cones on segment III stouter, a little shorter than 1/2 length of the segment................................................ + +sapae + +sp. n +. + + + + + + +- Head about 1.1 times as long as wide, or shorter ( +Fig. 264 +); antennal segment III short and stout, shorter than 1.9 times as long as wide, almost as long as segment IV, or a little shorter ( +Fig. 265 +); mouth cone reaching prospinasternum ( +Fig. 268 +); sense cones on segment III stouter, a little longer than 1/2 length of the segment.......................... + + +thailandicus +sp. n. + + + + + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B850BC336D9B2A0C1AABD751B.xml b/data/03/D2/38/03D2383B850BC336D9B2A0C1AABD751B.xml new file mode 100644 index 00000000000..af9d7bebcb0 --- /dev/null +++ b/data/03/D2/38/03D2383B850BC336D9B2A0C1AABD751B.xml @@ -0,0 +1,489 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips +Bagnall + + + + + + + + + + +Ecacanthothrips +Bagnall, 1909: 348 + + +. Type-species: + +Acanthothrips sanguineus +Bagnall + +(synonymised with + +Idolothrips tibialis +Ashmead + +by +Palmer & Mound, 1978 +), by monotypy. + + + +The genus + +Ecacanthothrips + +shares most generic features with + +Hoplandrothrips + +, and seems to be distinguished only by the larger number of sense cones on antennal segment III. + +Ecacanthothrips + +species have more than six (usually more than 10) stout sense cones on the relatively enlarged antennal segment III (cf. +Fig. 113 +), and this is the only autapomorphy mentioned for this genus. There is usually a larger number of sense cones on the ventral surface of the segment than on the dorsal surface. Two + +Ecacanthothrips + +species from +Japan +, + +E. inarmatus + +and + +E. moundi + +, usually have about 10 sense cones (cf. +Fig. 104 +), but infrequently have six or seven sense cones in small individuals. In particular, + +E. coniger + +from Borneo and Peninsular +Malaysia +has more than 60 dark sense cones even in small individuals ( +Fig. 88 +). In contrast, + +Hoplandrothrips + +species usually have three rather slender sense cones on that segment. Even in + +Hoplandrothrips + +, however, a widespread species, + +H. flavipes + +, as well as four species described from +Japan +and one new species described below from +Taiwan +have four rather stout sense cones on that segment. Moreover, + +E. andrei + +from +Indonesia +, Peninsular +Malaysia +and +Thailand +has four or five, but rarely two or three, sense cones on a rather slender segment III ( +Fig. 59 +), although it is included undoubtedly in the + +tibialis + +species-group of + +Ecacanthothrips + +. Members of this species-group include not only the widespread type-species of the genus, + +E. tibialis + +, but also + +E. claricornis + +from the +Philippines +and +Sulawesi +, +Indonesia +, all have a median tubercle on the inner margin of the fore femur in both sexes ( +Okajima 1983 +). Furthermore, + +Hoplandrothrips quinqueconus + + +sp. n. + +from +Taiwan +newly described below, unusually has five (rarely six) sense cones and may be closely related to + +H. ryukyuensis + +from the Ryukyu Islands. Therefore, the difference of sense cone number seems not to satisfactorily distinguish these two genera, and the genus + +Ecacanthothrips + +could well be treated as an Oriental species-group of the worldwide genus + +Hoplandrothrips + +, as suggested by +Okajima (2006) +. + +However, there is an essential morphological difference in these sense cones on antennal segment III between + +E. andrei + +and + +Hoplandrothrips + +species. + +In + +E. andrei + +the sense cones are comparatively small and all arise unusually on the ventral apex of the segment (see + +Fig. +5 + +in + +Palmer +et al. +1978 + +), but those of + +Hoplandrothrips + +species, including species with four or five sense cones on that segment, have the sense cones rather large and with one or two usually arising on the inner apex, and two, rarely one or three, on the outer apex of the segment. Frequently, the outer one is placed ventrally, but such a condition is widely found in the +Phlaeothripidae +. Even in + +H. flavipes + +, the formation of the four sense cones is not so different from ‘usual position’ as +Fig. 104A +in +Okajima (2006) +. It appears that there is a qualitative difference between the sense cones, even if the number is the same or reversed. Thus, it might be interpreted that + +the sense cones on antennal segment III of + +E. andrei + +are not homologous with those of + +Hoplandrothrips + +species, or at least no evidence is found that they are homologous. + +In comparative morphology, it is very important to verify structural homology by evaluating relative positional relationships. This qualitative difference could probably be explained by the fact that the common ancestor of + +E +. +andrei + +and + +E +. +claricornis + +had at least as many sense cones as in + +E +. +claricornis + +which is undoubtedly most closely related to + +E +. +andrei + +, then those sense cones have been reduced in number in + +E +. +andrei + +, but not conspicuously changed in + +E +. +claricornis + +. Therefore, these two genera, + +Ecacanthothrips + +and + +Hoplandrothrips + +, are both retained tentatively in the present study. However, +this is only a temporary treatment and an arbitrary distortion. +Even if the homology of the sense cones in both genera is rejected, it will not immediately lead to a clear separation between these two genera and the relationship between them remains insufficiently clarified. In comparison with some of the synapomorphies shared by these two genera, the autapomorphy involving the sense cones of + +Ecacanthothrips + +presumably has only a limited phylogenetic significance. We consider that + +Hoplandrothrips + +should be a synonym for + +Ecacanthothrips + +, but there are problems with + +Hoplandrothrips + +that need to be resolved, so we will not express that formally at this time. One problem is the relationship between + +Hoplandrothrips + +and + +Malacothrips + +, which might also be considered as a single genus, as discussed below under + +Hoplandrothrips + +. Moreover, + +Hoplandrothrips + +, includes at least 130 species, and repeated nomenclatural changes can cause taxonomic confusion. Incidentally, + +Malacothrips + +is the oldest name among these three genera. In addition, + +Ecacanthothrips + +is considered to be sister group to the + +Hoplandrothrips ryukyuensis + +- group (see below under + +H. flavipes + +), and it is probably not so closely related to the + +flavipes + +-group (see below under + +H. coloratus + +) as was previously considered. + + + + +Among the 11 described species, + +andrei + +, + +claricornis + +and + +tibialis + +, are included in the + +tibialis + +-group, with the head rather longer, a median tubercle on the inner margin of fore femur in both sexes, and the fore coxa strongly extruded postero-externally in large males. Four species, + +inarmatus + +, + +leai + +, + +moundi + +and + +nigellus + +, are included in the + +inarmatus + +-group, with a rather long head and unarmed median inner margin of fore femur. Two species, + +spinipes + +and + +tenuicornis + +, may also be included in the + +inarmatus + +-group, but the relationships of the remaining two species, + +coniger + +and + +kolibaci + +are unclear. It is noticeable that in all species belonging to these two groups, most of the sense cones are on the ventral rather than the dorsal surface of antennal segment III. + +E. coniger + +is peculiar in having antennal segment III strongly distorted ( +Fig. 88 +) with numerous stout sense cones which are situated asymmetrically, biased antero-externally, also the mouth cone is extremely long and pointed ( +Fig. 91 +) and extending beyond the prospinasternum. However, most other character states are similar to those of + +inarmatus + +-group. + +E. kolibaci + +, based on +two females +from +Sichuan +, +China +, has the head short and four stout sense cones on antennal segment III. According to the original description ( +Pelikan 2000 +), most of these four thick sense cones are ventral on the large antennal segment III, and the species is here retained in this genus with some hesitation. The species has not been available for this study and further observations are needed. Especially in the + +inarmatus + +-group, each species shows confusing size related variation as well as geographical variation, and it is very difficult to discriminate each species. + + +In this paper, 12 Oriental + +Ecacanthothrips +species + +are recognised in this genus, including + +E. brevicornis + + +sp. n. + +, here newly described from +Bali +Is., +Indonesia +. + + +Diagnosis. +Small to medium sized species, usually macropterous. Most features very similar to those of + +Hoplandrothrips + +, but with the following differences: antennal segment III usually enlarged (not enlarged in + +andrei + +and + +brevicornis + +), much wider than segment IV, with more than six sense cones (usually more than 10) and if fewer, then mostly on ventral apex of the segment; fore femur often with a median tubercle or tooth on inner surface in addition to a pair of apical tubercles in both sexes. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B850CC337D9B2A5F8ADEA73BA.xml b/data/03/D2/38/03D2383B850CC337D9B2A5F8ADEA73BA.xml new file mode 100644 index 00000000000..f7cce472464 --- /dev/null +++ b/data/03/D2/38/03D2383B850CC337D9B2A5F8ADEA73BA.xml @@ -0,0 +1,304 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + +Key to + +Ecacanthothrips + +species + + + + + + +(*: + +E. spinipes + +and + +E. kolibaci + +are based on literature) + + + + + + +1. Fore femur with a median tubercle on inner margin in both sexes (cf. +Figs 56 & 57 +), this tubercle present but scarcely visible even in very small female; fore coxa strongly extruded postero-externally in large male ( + +tibialis + +-group)................. 2 + + + +- Fore femur with inner margin almost smooth or with a series of short stout setae on small tubercles; fore coxa not strongly extruded, at most weakly angulate in large male............................................................. 4 + + + + + +2. Antennal segments III and IV brown ( +Fig. 129 +); mid and hind femora anterior margins each with 2–4 (or more) dark stout setae with blunt or pointed apices ( +Fig. 133 +); pronotal am setae reduced, much shorter than aa in large male; fore tibia with a sub-apical inner tubercle in large male; pelta triangular ( +Fig. 132 +).............................................. + +tibialis + + + + + +- Antennal segments III and IV largely yellowish; mid and hind femora anterior margins each with one pale stout seta with blunt or dilated apex (cf. +Fig. 62 +); pronotal am setae elongate, longer than aa in large male; fore tibia without inner sub-apical tubercle even in large male; pelta bell-shaped (cf. +Fig. 60 +)............................................................ 3 + + + + + + +3. Antennal segment III slender, almost as wide as segment IV, usually with 4 or 5, sometimes 3, rarely 2, sense cones on ventral apex ( +Fig. 59 +).................................................................................... + +andrei + + + + + +- Antennal segment III heavy, much wider than segment IV, with more than 10 sense cones ( +Fig. 79 +)............ + +claricornis + + + + + + + +4. Fore femur with a row of short stout setae on small tubercles; mid and hind femora anterior margins with 5 or more stout setae with blunt or dilated apices; pelta widely triangular................................................... + +spinipes + +* + + + +- Fore femur with inner margin almost smooth; mid and hind femora with 8 or more fine short setae with pointed apices; pelta usually bell-shaped.................................................................................... 5 + + + + + +5. Antennal segment III with more than 60 dark stout sense cones distributed asymmetrically on the segment, biased antero-externally ( +Fig. 88 +); antennal segments IV and V with distinct apical neck; mouth cone rather long, extending beyond prospinasternum ( +Fig. 91 +)......................................................................... + +coniger + + + + + +- Antennal segment III with less than 20 sense cones; antennal segments IV and V without distinct apical neck; mouth cone shorter, scarcely reaching prospinasternum (cf. +Fig. 99 +)....................................................... 6 + + + + + + +6. Antennal segment III with four thick sense cones, most of which are on ventral apex of segment................ + +kolibaci + +* + + + + +- Antennal segment III with more than 6 sense cones ( + +inarmatus + +-group)........................................... 7 + + + + + + +7. Head about 1.1 times as long as wide in both sexes (cf. +Figs 65 +& +120 +)........................................... 8 + + + + +- Head longer, 1.2–1.5 times as long as wide in female, 1.2–1.8 times in male (cf. +Figs 93 +& +103 +)....................... 9 + + + + + + +8. Antenna short, shorter than 1.7 times as long as head in female, 1.9 times in male; segment VIII short, 2.1–2.3 times as long as wide, slightly constricted at base; segment III uniformly brown, swollen laterally, shorter than 1.4 times as long as wide, a little shorter than IV ( +Figs 67 & 68 +); mouth cone not reaching prosternal ferna in female; mid and hind tibiae shaded with brown medially; fore wing paler, scarcely shaded medially............................................. + + +brevicornis + +sp. n. + + + + + +- Antenna longer, about 2.0 times as long as head in female, 2.1 times in male; segment VIII slender, 2.8–3.3 times as long as wide, distinctly constricted at base, pedicellate; segment III brown with yellowish base, straight-sided, longer than 1.6 times as long as wide, a little longer than IV ( +Fig. 122 +); mouth cone extending beyond prosternal ferna; mid and hind tibiae yellow; fore wing shaded with brown........................................................................ + +tenuicornis + + + + + + + +9. Mid and hind tibiae usually yellow, but sometimes brownish; head 1.2–1.3 times as long as wide in female, 1.2–1.4 times in male; cheeks distinctly constricted just behind eyes ( +Figs 93 & 94 +)............................................ + +leai + + + + + +- Mid and hind tibiae darker, if yellow, head longer than 1.3 times as long as wide in female, longer than 1.5 times in male; cheeks constricted behind eyes, but not distinct (cf. +Fig. 102 +)....................................................... 10 + + + + + + +10. Sense cones on antennal segment III dark; inner margin of antennal segment III distinctly swollen at basal half; base of segment IV wider, almost as wide as apical width ( +Figs 114 & 115 +)................................................ + +nigellus + + + + + +- Sense cones on antennal segment III pale; inner margin of antennal segment III not swollen at basal half; base of segment IV narrower than apical width (cf. +Fig. 104 +).................................................................. 11 + + + + + + +11. Head, prothorax and abdominal segments II to VI usually yellowish, a little paler than pterothorax; antennal segment III yellowish, much paler than segment II; hind tibiae dark brown, usually darker than femora.................... + +inarmatus + + + + + +- Body uniformly dark brown; antennal segment III dark brown, scarcely paler than segment II; hind tibiae brown to dark brown, almost concolourous with femora, or a little paler....................................................... + +moundi + + + + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B850DC337D9B2A219A9F5769D.xml b/data/03/D2/38/03D2383B850DC337D9B2A219A9F5769D.xml new file mode 100644 index 00000000000..09bb4198672 --- /dev/null +++ b/data/03/D2/38/03D2383B850DC337D9B2A219A9F5769D.xml @@ -0,0 +1,377 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips andrei +Palmer & Mound + + + + + + + +( +Figs 1–2 +, +56–64 +) + + + + + + + +Ecacanthothrips andrei +Palmer & Mound, 1978: 158–160 + + +. + + + +This species was described from Peninsular +Malaysia +and +Singapore +, and is included in the + +tibialis + +-group. +Pelikan (2000) +examined +one female +and +one male +of this species collected from +Sumatra +, +Indonesia +, but did not indicate detailed data. It is now recorded from +Thailand +and +Bali +Is., +Indonesia +, for the first time based on a good number of females and males. Although, two other members of the species-group, + +claricornis + +and + +tibialis + +, have more than 10 sense cones (about +40 in +maximum) on rather enlarged antennal segment III, + +andrei + +have usually four or five (often three, rarely two) sense cones on comparatively slender segment III ( +Figs 58 & 59 +). Apparently, this sense cone number of + +andrei + +appears to recover a distinction between the genera + +Ecacanthothrips + +and + +Hoplandrothrips + +. However, the sense cones of + +andrei + +seem not to be homologous with those of + +Hoplandrothrips + +species as discussed above, judging from their form and position. This condition of the slender antennal segment III with small number of sense cones in + +andrei + +( +Fig. 59 +) within + +Ecacanthothrips + +could because it evolved from a species that had enlarged segment III with larger number of sense cones, such as + +claricornis + +( +Fig. 79 +), but is now the result of secondary reduction during evolution. In essence, amongst the genus + +Ecacanthothrips + +the condition of antennal segment and sense cones of + +andrei + +is an autapomorphy, not a plesiomorphy inherited from + +Hoplandrothrips + +. + + + + +Structural variation of this species is very similar to that of + +tibialis + +. However, it is mainly size related variation, and there is no conspicuous geographical variation. Similar to + +claricornis + +, + +andrei + +has the pelta bell-shaped, only one stout seta with blunt or dilated apex on each mid and hind femur in both sexes ( +Fig. 62 +), and has the anteromarginal pronotal setae elongate, much longer than anteroangular setae in large male. In consequence, + +andrei + +is undoubtedly more closely related to + +claricornis + +than to + +tibialis + +. Furthermore, + +andrei + +and + +claricornis + +have allopatric distribution, although both are sympatric with + +tibialis + +. + + +Specimens examined. + +Peninsular + +Malaysia + +, +Tapah +, +1 male +, on dead leaves, + +26.vii.1976 + +, SO; + + +about + +20km + +N from Kuala Lumpur, +Templer Park +, +37 females +and +27 males +, on dead leaves, + +11–15.viii.1990 + +, TN & SO; + + +Cameron Highland +, nr. +Tanah Rata +, +Robinson +water fall, +1 female +, on dead leaves and branches, + +28.viii.1990 + +, TN & SO + +. + + +Singapore + +, +Macritchie Res. +, +1 female +, on dead +Palmae +fronds, +1 female +, on dead branches, + +7.viii.1990 + +, TN & SO + +. + + +Indonesia + +, +Bali +Is. +, +Tabanan +, +Bengkel +, +Pura Batu Selahan +, +4 females +and +3 males +, + +3.ix.2005 + +, SO + +. + + +Thailand + +, +Phuket +, +Rang Hill +, +12 females +and +9 males +, on dead leaves and branches, + +14.ix.1992 + +, TN & SO; + + +Phuket +, nr. +Tonesai +water fall, +1 female +and +1 male +, on bamboo, + +23.viii.1991 + +, TN & SO, + + +same locality above, +15 females +and +8 males +, on dead leaves and branches, + +11–12.ix.1992 + +, TN & SO; + + +nr. +Chiang Mai +, +Mae Sa +, +3 females +and +6 males +, on dead branches, + +6.ix.1992 + +, TN & SO + +. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B850EC335D9B2A647AB6673C7.xml b/data/03/D2/38/03D2383B850EC335D9B2A647AB6673C7.xml new file mode 100644 index 00000000000..59b08d18ee2 --- /dev/null +++ b/data/03/D2/38/03D2383B850EC335D9B2A647AB6673C7.xml @@ -0,0 +1,223 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Ecacanthothrips claricornis +Okajima + + + + + + + +( +Figs 5–6 +, +75–83 +) + + + + + + + +Ecacanthothrips claricornis +Okajima, 1983: 57–59 + + +. + + + + +E. claricornis + +was described based on +17 females +and +six males +collected from Mindanao Is., the +Philippines +, together with non-paratypic +three females +and +five males +from Luzon Is., and is included in the + +tibialis + +-group. It is newly recorded here from Central and +South Sulawesi +, +Indonesia +, based on a long series of females and males listed below. This species also shows remarkable size related structural variations associated with allometric growth as in + +tibialis + +. Although the specimens from Mindanao, the +type +series, have the femora fully brown, the specimens from both Luzon and +Sulawesi +have the femora largely brown but with apices clear yellow. Moreover, the size related variation of + +claricornis + +is more similar to + +andrei + +rather than + +tibialis + +, and the prothoracic anteromarginal setae elongate, the fore tibia with no sub-apical inner tubercle even in large males, and + +claricornis + +has only one stout seta with blunt or dilated apex on each mid and hind femur in both sexes ( +Fig. 82 +). In contrast to + +andrei + +, this species has somewhat enlarged antennal segment III ( +Figs 78 & 79 +), that is distinctly wider than segment IV, with usually 10–15 sense cones. + + + + +Specimens examined. +The + + +Philippines + +, holotype female and paratype females and males from +Mindanao Is +.; + + +3 females +and +5 males +, from +Luzon Is. +(data described in +Okajima, 1983 +) + +. + + +Indonesia + +, +South Celebes +(= Sulawesi), +Malino +, alt. about + +900m + +, +4 females +and +5 males +, on dead leaves and branches, + +31.vii.1984 + +, +2 females +and +3 males +, on dead Palmae, + +3.viii.1984 + +, SO; + + +Central Celebes +, +near Rantepao +, +Pedamaran +, alt. about + +1000m + +, +62 females +and +45 males +, on dead leaves and branches, + +8–14.viii.1984 + +, SO + +. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B8513C329D9B2A56DAC837695.xml b/data/03/D2/38/03D2383B8513C329D9B2A56DAC837695.xml new file mode 100644 index 00000000000..c529a5607d7 --- /dev/null +++ b/data/03/D2/38/03D2383B8513C329D9B2A56DAC837695.xml @@ -0,0 +1,224 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Hoplandrothrips obesametae +Chen + + + + + + + +( +Figs 38–39 +, +215–224 +) + + + + + + + +Hoplandrothrips obesametae +Chen, 1980: 177–178 + + +. + + + + +H. obesametae + +was described from +Taiwan +based on strongly crushed specimens and was recorded subsequently from +China +( +Tong & Zhang 1989 +) and +Japan +( +Okajima 2006 +). Unfortunately, the +type +specimens have not been available for this study, and it is very difficult to discriminate not only this species but also most of the species described by L. S. Chen that are known only from the original descriptions that were based on strongly crushed +type +specimens. However, +one female +collected from Nanshanchi, +Nantou Hsien +, +Taiwan +, listed below could well be identified as + +H. obesametae +. + +This species may be closely related to + +H. floresi + +newly described above, in having antennal segment VIII short and not distinctly constricted basally, the terminal setae much longer than the tube and abdominal sternite VIII of males with a median small pore plate. +Okajima (2006) +identified as this species several females and males collected from the Ryukyu Islands (Yonakuni Is., Ishigaki Is. and Miyako Is.), and fully described these. They are indistinguishable from the female from +Taiwan +, but are identified as + +obesametae + +with some slight hesitation, due to some small differences from the original description, and further study is required. +Five females +and +one male +collected from Mindanao Is., the +Philippines +, listed below in the doubtful specimens are very similar to this species, in having the head shorter, cheeks swollen, antennal segment III shorter than segment IV, antennal segment VIII short and conical, and abdominal sternite VIII of male with a median small pore plate. However, they have shorter antennal segments III and IV, and shorter mouth-cone, but these differences are probably related to the smaller body size. The bodies of these females are +1.8–2.3mm +in length, though those of + +obesametae + +from the Ryukyu Is. and +Taiwan +are +2.2–2.8mm +. + + +Specimens examined +. + + +Japan + +, +Ryukyu Islands +, +10 females +and +7 males +(data described in +Okajima, 2006 +) + +. + + +Taiwan + +, +Nantou Hsien +, +Nanshanchi +, +1 female +, on dead leaves, + +30.iii.1984 + +, SO + +. + + + + +Doubtful specimens +. +The + + +Philippines + +, +Mindanao Is. +, +North Cotabato +, Ilomavis, +2 females +and +1 male +, on dead leaves, + +26.vii.1979 + +, +1 female +, on dead fern, + +27.vii.1979 + +, +2 females +, on dead leaves, + +28.vii.1979 + +, SO + +. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B8514C32FD9B2A1C4AC507027.xml b/data/03/D2/38/03D2383B8514C32FD9B2A1C4AC507027.xml index e076ab08395..1e6fabb6fab 100644 --- a/data/03/D2/38/03D2383B8514C32FD9B2A1C4AC507027.xml +++ b/data/03/D2/38/03D2383B8514C32FD9B2A1C4AC507027.xml @@ -1,67 +1,67 @@ - - - -The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan - - -Author + + +Author -Okajima, Shûji -0000-0001-7249-671X -Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. -7okajimas2@gmail.com +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com - - -Author + + +Author -Masumoto, Masami -0000-0001-9049-2448 -Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. -masumotoms@gmail.com +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-07-31 + +2024 + +2024-07-31 - -5489 + +5489 - -1 + +1 - -22 -91 + +22 +91 - -http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 -journal article -10.11646/zootaxa.5489.1.4 -1175-5326 -13211341 -373DBA20-A1A7-4A2D-856C-67BF13D83C41 +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 - + + - Hoplandrothrips quinqueconus - sp. n. + @@ -75,9 +75,7 @@ Female (macroptera) -. - -Distended body length: +. Distended body length: 2.8–3.7mm . Body uniformly dark brown. All legs dark brown, but tibiae often scarcely paler distally, all tarsi concolourous with apex of tibia. Antennal segments I and II @@ -126,9 +124,7 @@ subequal in length. Mouth cone scarcely reaching prospinasternum; maxillary styl Fig. 232 ) dilated, S1 and S2 subequal, S3 scarcely longer than S2. Pelta ( Fig. 229 -) bell-shaped, with rather large lateral wings. tergite IX setae S1 and S2 subequal, shorter than tube, slightly dilated. Tube 0.54–0.59 times as long as head, 2.30-2.44 times as long as wide, almost tapering; terminal setae much longer than tube - -. +) bell-shaped, with rather large lateral wings. tergite IX setae S1 and S2 subequal, shorter than tube, slightly dilated. Tube 0.54–0.59 times as long as head, 2.30-2.44 times as long as wide, almost tapering; terminal setae much longer than tube. Measurements @@ -250,12 +246,14 @@ and , on dead leaves and branches , + 5 females and 8 males , 19.viii.1993 , TN & SO. + diff --git a/data/03/D2/38/03D2383B8515C32CD9B2A0E5AAD274DF.xml b/data/03/D2/38/03D2383B8515C32CD9B2A0E5AAD274DF.xml new file mode 100644 index 00000000000..4a878ea0505 --- /dev/null +++ b/data/03/D2/38/03D2383B8515C32CD9B2A0E5AAD274DF.xml @@ -0,0 +1,349 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Hoplandrothrips samirseni +(Varatharajan, Singh & Bala) comb. n. + + + + + + + +( +Figs 43–44 +, +233–243 +) + + + + + + + +Tylothrips samirseni +Varatharajan, Singh & Bala, 2015: 7160-7162 + + +. + + + +This species was originally described from +Manipur +, Northeastern +India +based on a unique +holotype +male collected from the fallen leaves of +Ureno lobata +( +Malvaceae +). The maxillary stylets are relatively long but not reaching the middle of the head capsule and close together medially, and the fore wings are slightly constricted medially with a few duplicated cilia. These character states indicate that this species may not be related to the genus + +Tylothrips + +( +Mound 1977 +, + +Uzun Yiğit +et al. +2021 + +). The original description of + +samirseni + +provides only a few important structural details to recognise its taxonomic position, and the measurements also lack accuracy. The published images indicate that the tube is clearly shorter than the head, although it is much longer than the head in the measurements. This is probably because the magnification of the objective lens was incorrect when using the eyepiece micrometer. However, the images and figures in the description are available for this study and suggest that it is very similar to grass-inhabiting + +Hoplandrothrips + +species from Southeast Asia. Based on that description it cannot be distinguished satisfactorily from specimens collected in +Indonesia +, +Thailand +and +Vietnam +listed below, especially the smaller males. Therefore, these specimens are here identified as + +samirseni + +, but with slight hesitation, because there are some small differences between local populations in addition to the insufficient original description. It undoubtedly belongs to the + +nobilis + +- group, and is related to + +H. basantratus + +described above. Both of them share some character states as follows: head elongate with a constriction just behind compound eyes, rather short maxillary stylets retracted to basal 1/3 of head, pronotal basantra present at least in specimens from Southeast Asia, and the body bicoloured yellow and brown. However, + +samirseni + +differs clearly from + +basantratus + +in having the body largely brown mentioned in the key above. Another grass-inhabiting species, + +H. graminicola + +, may also be related, but has the body uniformly brown and no prosternal basantra. + + + + +The full description of this species is given below based on the specimens from +Java +, +Indonesia +, because the original description is insufficient for the present study. + + +Female (macroptera). +Distended body length: 2.0– +2.9mm +. The following description is based on a female with +2.3mm +body in length. Body largely brown, but partly yellowish; head, thorax, abdominal segments VII–IX and tube brown, abdominal segments II–III yellow, very weakly shaded with brown, segments IV–VI brownish yellow to brown, gradually darkened posteriorly. Antennal segments I–II brown, a little darker than head, segment III yellowish, weakly shaded with brown, segments IV–VIII largely pale brown, bases of IV–VI slightly paler. Femora brown; tibiae and tarsi yellow, tibiae often weakly shaded. Fore wings almost clear. All prominent setae clear. Head ( +Fig. 233 +) elongate, 1.45–1.50 times as long as wide, 1.48 times, dorsal surface almost smooth, but weakly sculptured with reticulation at extreme base, postero-lateral portion, just behind eyes and ocellar region. Cheeks weakly rounded, incised just behind eyes, weakly serrated, with some minute setae. Postocular setae shorter than eyes, dilated. Eyes bulged, about 0.3 times as long as head; posterior ocelli close to eyes, distance between posterior ocelli longer than diameter of an ocellus, 25μm apart from one another. Antennae ( +Fig. 237 +) 1.8–1.9 times as long as head; segment VIII rather elongate, constricted at base, pedicellate; segment III 1.76 times as long as wide, almost as long as segment IV, with three sense cones. Mouth cone short, maxillary stylets retracted to basal 1/3 of head, not reaching postocular setae, 15–20μm apart from each other, 16μm. Pronotum smooth, but weakly reticulate posteriorly, about 0.5–0.6 times as long as head; five pairs of prominent setae elongate, slender and expanded; am much shorter than aa, pa and epim subequal in length. Prosternal basantra ( +Fig. 239 +) present, but small; mesopresternum divided into two lateral large triangle plates and a median circular plate. Metanotum ( +Fig. 240 +) weakly sculptured with longitudinal reticulation; median pair of setae acute, about 40–45μm apart from anterior margin, about 70μm apart from one another. Fore tarsal tooth ( +Fig. 234 +) minute. Fore wings with 3–5 duplicated cilia; three sub-basal setae ( +Fig. 242 +) expanded. Pelta ( +Fig. 241 +) hat-shaped, with rather broad lateral wings, distinctly reticulate. Tergite IX S1 setae almost pointed or blunt, shorter than tube, S2 acute, almost as long as tube or a little shorter. Tube ( +Fig. 243 +) 0.56 times as long as head, about 2.0 times as long as wide. Terminal setae much longer than tube. + + +Measurements +(female in μm). Body length about 2290 (distended). Head length 242, from anterior margin of eyes 212, width across eyes 150, maximum width across cheeks 163; eyes length 70, width 48; diameter of posterior ocelli 13–17; postocular setae 50-52. Antenna total length 450, segments I–VIII length (width) as follows: 50 (40), 50 (33), 60 (34), 62 (33), 62 (28), 52 (24), 50 (21), 41 (13). Pronotum length 135, width 225. Setae on prothorax: am 15–20, aa 35–40, ml 40–45, pa 52–53, epim 52–55, cox about 40. Fore wing length 800. Sub-basal wing setae: +S1 45 +–47, +S2 52 +–55, +S3 50 +–52. Tergite IX setae: S1 110–112, S2 130–132. Tube length 135, maximum width 67; terminal setae 200. + + +Male (macroptera). +Distended body length: +1.7–2.4mm +. Abdominal segments usually darker than female, segments I and II yellowish brown, remaining segments gradually darkened towards tube, pale brown to dark brown; mid femora yellowish, apical 1/5–1/3 of fore femora and basal 1/3 of hind femora often yellowish, mid and hind tibiae often weakly shaded with brown. Pronotal am setae reduced, less than 15μm. Large male: pronotal aa setae elongate, much longer than pa; fore coxa with some stout setae on posterior margin; fore femur ( +Fig. 236 +) swollen, with a ventro-apical tubercle, fore tibia with a sub-basal inner tubercle, but without apical tubercle, fore tarsal tooth stout, wide-based; fore wing with 6 duplicated cilia. Small male: pronotal aa setae almost as long as pa, or a little shorter; fore femur ( +Fig. 235 +) without apical tubercle, fore tibia without sub-basal tubercle, fore tarsal tooth small as in female. Abdominal sternite VIII without pore plate; tergite IX S2 setae about 1/2 length of S1, S1 almost as long as tube. Tube 0.47–0.52 times as long as head. + + +Measurements +(large/small males in μm). Body length about 2340/1720 (distended). Head length 245/222, from anterior margin of eyes 220/195, width across eyes 150/130, maximum width across cheeks 160/140; eyes length 80/63, width 50/40; postocular setae 65–68/40. Antenna total length 405/385, segments I–VIII length as follows: 50/43, 50/45, 63/50, 63/52, 65/52, 53/45, 47/42, 42/40. Pronotum length 180/125, width 260/195. Setae on prothorax: am about 10/about 10, aa 85–90/35–40, ml 75–80/38–40, pa 60/40–45, epim 68–75/40–45, cox 55–60/ about 30. Fore wing length 870/680. Sub-basal wing setae: +S1 45 +/33, S2 65–68/43, +S3 60 +–65/40. Tergite IX setae: S1 120–123/about 100, +S2 58 +–60/50–52. Tube length 125/105, maximum width 71/55; terminal setae 195/155. + + +Specimens examined +. + + +Indonesia + +, +Java +, +Malang +, +Wono-koyo +, ca + +1150m + +alt., +34 females +and +15 males +, on grass, + +23.viii.2005 + +, SO + +. + +Thailand + +, nr. + +Chiang Rai +, +Pha Yao +, +4 females +and +1 male +, on grass, + +5.ix.1992 + +, SO; +2 females +, data very similar to above, but on dead leaves and branches; foot of Doi Inthanon, +1 female +, on grass, + +20.viii.1992 + +, TN & SO; +Pha Hean +, +1 female +and +2 males +, + +3.ix.1992 + +, TN & SO; +Doi Pui +, +1 female +, on grass, + +1.ix.1992 + +, SO + +; + +Saraburi +, +Farm of Kasetsart University +, +2 males +, on grass, + +20.viii.1991 + +, TN & SO + +; + +Chiang Mai +, +1 female +, in paddy field, + +7.v.1978 + +, +K. Yasumatsu + +; +Phuket +Is., + +Phuket +Hill +, +1 male +, + +9.ix.1992 + +, SO + +. + + +Vietnam + +, +Lam Dong Province +, +Bao Loc +, +Dam Bri +, +1 male +, on bamboo, + +27.xii.2001 + +, SO + +. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B851DC327D9B2A019A9A87549.xml b/data/03/D2/38/03D2383B851DC327D9B2A019A9A87549.xml new file mode 100644 index 00000000000..4ac6201af45 --- /dev/null +++ b/data/03/D2/38/03D2383B851DC327D9B2A019A9A87549.xml @@ -0,0 +1,171 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Hoplandrothrips coloratus +Okajima + + + + + + + + +Hoplandrothrips coloratus +Okajima, 2006 + +, 311–312. + + +Described from the Ryukyu Islands (Ishigaki Is. and Iriomote Is.), +Japan +, this species was recorded subsequently from +Taiwan +( +Dang & Qiao 2014 +). Most specimens of the +type +series were collected from subtropical forest litter. It has the head entirely reticulate dorsally, the sub-basal cheek setae small, antennal segment VIII pedicellate with the base distinctly constricted, the fore tarsal tooth minute in female, the pelta with rather slender lateral wings, the intermediate seta between S1 and S2 on the tergite IX rather long, and sternite VIII of males with no pore plate. Because these character states are shared, this species may be closely related to + +H. flavipes + +together with + +H. brunneicinctus + +from +Australia +and + +H. aseanae + +newly described above from Southeast Asia (they are named tentatively + +flavipes + +-group here). However, + +flavipes + +has four stout sense cones on antennal segment III, whereas + +coloratus + +has unusually only two sense cones on that segment. However, + +brunneicinctus + +and + +aseanae + +have morphological structures something intermediate between + +coloratus + +and + +flavipes + +with three sense cones on the antennal segment III. In addition to this species, at least three species, such as + +H. elongatus + +from +Japan +, + +H. hemiflavus + +from +Australia +and + +H. tumiceps + +from New York, North America, also have only two sense cones on antennal segment III, but these species may be not closely related to each other. Unfortunately, specimens from +Taiwan +have not been available for this study. + + + + +Specimens examined. + + +Japan + +, +holotype +female and + + +paratype +females and males from the +Ryukyu Islands +(data described in +Okajima, 2006 +) + +. + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B851EC324D9B2A0ADACBA77BD.xml b/data/03/D2/38/03D2383B851EC324D9B2A0ADACBA77BD.xml new file mode 100644 index 00000000000..6d7df94bf7b --- /dev/null +++ b/data/03/D2/38/03D2383B851EC324D9B2A0ADACBA77BD.xml @@ -0,0 +1,212 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + + +Hoplandrothrips flavipes +Bagnall + + + + + + + + +Hoplandrothrips flavipes +Bagnall, 1923 + +, 628. + + + +H. flavipes + +was originally described from +Kenya +and has six junior synonyms. It is widely distributed mainly in the tropics and subtropics around the world, extending to the temperate region in +Japan +. Sometimes, it makes large colonies on dead leafy branches in Southeast Asia and +Japan +. In spite of the wide distribution of this species, the morphological characteristics are relatively stable, and there is no significant difference depending on the locality. However, colouration of the median abdominal segments is often variable. In most individuals from Southeast Asia to +Japan +, abdominal segment III is usually yellowish, paler than segment II, and segments IV to VII are gradually darkened posteriorly in the female, but this is rather indistinct in the male, and often these segments are uniformly brown in both sexes. Moreover, according to +Ananthakrishnan (1964) +, this species from +India +(= + +Phlaeothrips indicus +Ramakrishna & Margabandhu + +) has the metathorax and abdominal segment I yellowish at least in females. It is unusual in having four stout sense cones on antennal segment III, but this character state is shared with four congeners from +Japan +( +Okajima 2006 +) and + +H. formosae + +newly described below from +Taiwan +. These sense cones are placed in an almost usual position for +Phlaeothripinae +, two sense cones on each inner and outer apex of the segment, but the outer one frequently situated ventrally. However, although these six species share the same sense cone formula on antennal segment III, they are not necessarily closely related to each other. Some large sized species, which are named temporarily + +ryukyuensis + +-group here, such as + +formosae + +, + +jennyae +, +quinqueconus + +and + +ryukyuensis + +, are undoubtedly not so closely related to this species. Moreover, even in the + +flavipes + +-group, + +H. aseanae + +described above and + +H. brunneicinctus + +from +Australia +are probably closely related to + +flavipes + +, though they have three sense cones on that antennal segment. Similarly, + +H. asianus + +described above and + +H. ochraceus + +from +Japan +appear to be included in the + +flavipes + +-group and also have three sense cones on segment III, despite the maxillary stylets scarcely longer and closer together. Finally, + +H. coloratus + +may also be included in this group, although it has only two sense cones on antennal segment III. These species share the following character states: head a little longer than wide, almost entirely sculptured with polygonal reticulation, but reticles often very weak; sub-basal cheek setae rather small; antennal segment VIII distinctly constricted basally, pedicellate; maxillary stylets close together medially, but narrowly apart, retracted to postocular setae, usually not reaching eyes; fore tarsal tooth absent or weakly developed in female; pelta hat-shaped with slender lateral wings; abdominal sternite VIII with no pore plate in male. Considering these relationships, and the character state of sense cones on antennal segment III, there is a possibility that this species is not as closely related to + +Ecacanthothrips + +species as considered previously. +Okajima (2006) +redescribed and illustrated this species based on a long series of females and males collected mainly from both subtropical and temperate regions of +Japan +. + + +Specimens examined +(detailed data are omitted). Numerous females and males from the following localities: + +Indonesia + +( +Bali +, Lombok and +Sulawesi +); + +Peninsular +Malaysia + +; + +Singapore + +; +Borneo +(Kalimantan and Sabah); + +Thailand + +; + +Taiwan + +; + +Japan + +(Honshu, Ogasawara and Ryukyu, data described in +Okajima, 2006 +). + + + + \ No newline at end of file diff --git a/data/03/D2/38/03D2383B851FC32BD9B2A65DAC8871BB.xml b/data/03/D2/38/03D2383B851FC32BD9B2A65DAC8871BB.xml index e5e19e1613b..7b1a26c9183 100644 --- a/data/03/D2/38/03D2383B851FC32BD9B2A65DAC8871BB.xml +++ b/data/03/D2/38/03D2383B851FC32BD9B2A65DAC8871BB.xml @@ -1,67 +1,67 @@ - - - -The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan - - -Author + + +Author -Okajima, Shûji -0000-0001-7249-671X -Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. -7okajimas2@gmail.com +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com - - -Author + + +Author -Masumoto, Masami -0000-0001-9049-2448 -Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. -masumotoms@gmail.com +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-07-31 + +2024 + +2024-07-31 - -5489 + +5489 - -1 + +1 - -22 -91 + +22 +91 - -http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 -journal article -10.11646/zootaxa.5489.1.4 -1175-5326 -13211341 -373DBA20-A1A7-4A2D-856C-67BF13D83C41 +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 - + + - Hoplandrothrips formosae - sp. n. + @@ -75,9 +75,7 @@ Female (macroptera) -. - -Distended body length: +. Distended body length: 3.5–4.4mm . Body uniformly dark brown. Fore tibiae brownish yellow, mid and hind tibiae brownish, slightly paler distally, tarsi brownish yellow. Antennal segments I and II @@ -136,9 +134,7 @@ longer than segment Fig. 196 ) 0.52 times as long as head, 1.93 times as long as wide in holotype -, almost tapering; terminal setae longer than tube - -. +, almost tapering; terminal setae longer than tube. Measurements diff --git a/data/03/D2/38/03D2383B852AC310D9B2A530ADEA74E5.xml b/data/03/D2/38/03D2383B852AC310D9B2A530ADEA74E5.xml new file mode 100644 index 00000000000..82d2b200430 --- /dev/null +++ b/data/03/D2/38/03D2383B852AC310D9B2A530ADEA74E5.xml @@ -0,0 +1,111 @@ + + + +The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan + + + +Author + +Okajima, Shûji +0000-0001-7249-671X +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +7okajimas2@gmail.com + + + +Author + +Masumoto, Masami +0000-0001-9049-2448 +Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. +masumotoms@gmail.com + +text + + +Zootaxa + + +2024 + +2024-07-31 + + +5489 + + +1 + + +22 +91 + + + + +http://dx.doi.org/10.11646/zootaxa.5489.1.4 + +journal article +10.11646/zootaxa.5489.1.4 +1175-5326 +13211341 +373DBA20-A1A7-4A2D-856C-67BF13D83C41 + + + + + + +Key to + +Phlaeothrips + +species from +Vietnam + + + + + + + + +1. Antennal segments III–VIII largely brown ( +Fig. 273 +); antennal segment VIII constricted basally, pedicellate, segments III–VI with short apical neck; mouth cone short and rather pointed, not reaching prosternal ferna ( +Fig. 276 +); fore wing with more than 45 duplicated cilia; pelta trapezoidal ( +Fig. 274 +); abdominal tergites II–VI each with two pairs of sigmoid wing-retaining setae; sternite VIII of male with arched narrow pore plate............................................ + + +annamensis +sp. n. + + + + + + +- Antennal segment III yellow on basal 1/2, brown on distal 1/2 ( +Fig. 285 +), segments IV–VI brown with pedicels yellowish; antennal segment VIII not constricted basally ( +Fig. 286 +), cone-shaped, segments III–VI truncate at apex; mouth cone long, reaching prosternal ferna ( +Fig. 287 +); fore wing with less than 30 duplicated cilia; pelta bell-shaped ( +Figs 282 & 283 +); abdominal tergites II–VII each with two pairs of sigmoid wing-retaining setae; sternite VIII of male without pore plate.................................................................................................... + + +tonkinensis +sp. n. + + + + + + + + \ No newline at end of file