diff --git a/data/03/8E/87/038E87839231FFC5DA94FE53FADAFD23.xml b/data/03/8E/87/038E87839231FFC5DA94FE53FADAFD23.xml new file mode 100644 index 00000000000..e90b1841f9a --- /dev/null +++ b/data/03/8E/87/038E87839231FFC5DA94FE53FADAFD23.xml @@ -0,0 +1,167 @@ + + + +The first Palearctic species of Discheramocephalus Johnson (Coleoptera, Ptiliidae, Ptiliinae) from Korea with an annotated catalog of the genus Discheramocephalus + + + +Author + +Jang, Taeyoung +0000-0001-8560-752X +Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea +taeyoung.jang@cbnu.ac.kr + + + +Author + +Park, Jong-Seok +0000-0003-2063-2534 +Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea +jpark16@cbnu.ac.kr + +text + + +Zootaxa + + +2024 + +2024-11-28 + + +5541 + + +4 + + +539 +548 + + + + +http://dx.doi.org/10.11646/zootaxa.5360.3.8 + +journal article +10.11646/zootaxa.5541.4.6 +1175-5326 +14241254 +6ACC4F8A-422B-4E9A-B152-22D649A5980C + + + + + + +Genus + +Discheramocephalus +Johnson, 2007 + + + + + + + +Type +species + +Discheramocephalus semisulcatus +Johnson, 2007 + +. + + + + + + + +Discheramocephalus +Johnson, 2007: 223 + + +; + +Grebennikov 2008: 314 + +(key to the genus + +Discheramocephalus + +); 2009: 117; + +Darby 2013: 240 + +; + +2016: 418 + +(key to the Peruvian + +Discheramocephalus + +); 2019: 21 (key to the genera of the Sarawakian +Ptiliidae +); 2020: 14. + + + + + +Distribution. +Eastern Palearctic ( +South Korea +) and Pantropical (Afrotropical realm: +Cameroon +, +Ecuador +, +Madagascar +, +Tanzania +; Australasian realm: +Solomon Islands +; Indomalayan realm: +Indonesia +, +Malaysia +; Neotropical realm: +Bolivia +, +Peru +) distribution. Additionally, some undescribed + +Discheramocephalus +species + +are known from the +USA +: +Florida +( +Dybas 1980 +, as “undescribed genus”), +Brazil +, +Congo +, +Sri Lanka +( +Johnson 2007 +), and +Australia +: +Queensland +( +Grebennikov 2008 +). + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87839231FFC5DA94FF47FE2BFEA4.xml b/data/03/8E/87/038E87839231FFC5DA94FF47FE2BFEA4.xml index 65380034215..aa018d88876 100644 --- a/data/03/8E/87/038E87839231FFC5DA94FF47FE2BFEA4.xml +++ b/data/03/8E/87/038E87839231FFC5DA94FF47FE2BFEA4.xml @@ -1,56 +1,56 @@ - - - -The first Palearctic species of Discheramocephalus Johnson (Coleoptera, Ptiliidae, Ptiliinae) from Korea with an annotated catalog of the genus Discheramocephalus + + + +The first Palearctic species of Discheramocephalus Johnson (Coleoptera, Ptiliidae, Ptiliinae) from Korea with an annotated catalog of the genus Discheramocephalus - - -Author + + +Author -Jang, Taeyoung -0000-0001-8560-752X -Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea -taeyoung.jang@cbnu.ac.kr +Jang, Taeyoung +0000-0001-8560-752X +Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea +taeyoung.jang@cbnu.ac.kr - - -Author + + +Author -Park, Jong-Seok -0000-0003-2063-2534 -Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea -jpark16@cbnu.ac.kr +Park, Jong-Seok +0000-0003-2063-2534 +Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea +jpark16@cbnu.ac.kr -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-28 + +2024 + +2024-11-28 - -5541 + +5541 - -4 + +4 - -539 -548 + +539 +548 - -http://dx.doi.org/10.11646/zootaxa.5360.3.8 + +http://dx.doi.org/10.11646/zootaxa.5360.3.8 -journal article -10.11646/zootaxa.5541.4.6 -1175-5326 -14241254 -6ACC4F8A-422B-4E9A-B152-22D649A5980C +journal article +10.11646/zootaxa.5541.4.6 +1175-5326 +14241254 +6ACC4F8A-422B-4E9A-B152-22D649A5980C @@ -66,7 +66,7 @@ Discheramocephalini - + Type genus @@ -76,6 +76,8 @@ genus . + + Grebennikov 2009: 115 (phylogenetic tree with morphological data matrix; key to the genera of the tribe diff --git a/data/03/8E/87/038E87839232FFC6DA94F96CFD41F8A6.xml b/data/03/8E/87/038E87839232FFC6DA94F96CFD41F8A6.xml new file mode 100644 index 00000000000..73f01490abf --- /dev/null +++ b/data/03/8E/87/038E87839232FFC6DA94F96CFD41F8A6.xml @@ -0,0 +1,90 @@ + + + +The first Palearctic species of Discheramocephalus Johnson (Coleoptera, Ptiliidae, Ptiliinae) from Korea with an annotated catalog of the genus Discheramocephalus + + + +Author + +Jang, Taeyoung +0000-0001-8560-752X +Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea +taeyoung.jang@cbnu.ac.kr + + + +Author + +Park, Jong-Seok +0000-0003-2063-2534 +Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea +jpark16@cbnu.ac.kr + +text + + +Zootaxa + + +2024 + +2024-11-28 + + +5541 + + +4 + + +539 +548 + + + + +http://dx.doi.org/10.11646/zootaxa.5360.3.8 + +journal article +10.11646/zootaxa.5541.4.6 +1175-5326 +14241254 +6ACC4F8A-422B-4E9A-B152-22D649A5980C + + + + + + + +Discheramocephalus jaeseoki + +sp. nov. + + + + + + + +Discheramocephalus jaeseoki + + +sp. nov. + + + + + +Distribution. +Type +locality: +Korea +( +Gyeongnam Prov. +). + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87839235FFC3DA94FBABFC10F8EF.xml b/data/03/8E/87/038E87839235FFC3DA94FBABFC10F8EF.xml index 7569904fbe9..cac377d2c0e 100644 --- a/data/03/8E/87/038E87839235FFC3DA94FBABFC10F8EF.xml +++ b/data/03/8E/87/038E87839235FFC3DA94FBABFC10F8EF.xml @@ -1,56 +1,56 @@ - - - -The first Palearctic species of Discheramocephalus Johnson (Coleoptera, Ptiliidae, Ptiliinae) from Korea with an annotated catalog of the genus Discheramocephalus + + + +The first Palearctic species of Discheramocephalus Johnson (Coleoptera, Ptiliidae, Ptiliinae) from Korea with an annotated catalog of the genus Discheramocephalus - - -Author + + +Author -Jang, Taeyoung -0000-0001-8560-752X -Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea -taeyoung.jang@cbnu.ac.kr +Jang, Taeyoung +0000-0001-8560-752X +Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea +taeyoung.jang@cbnu.ac.kr - - -Author + + +Author -Park, Jong-Seok -0000-0003-2063-2534 -Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea -jpark16@cbnu.ac.kr +Park, Jong-Seok +0000-0003-2063-2534 +Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju, Chungbuk Province 28644, Republic of Korea +jpark16@cbnu.ac.kr -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-28 + +2024 + +2024-11-28 - -5541 + +5541 - -4 + +4 - -539 -548 + +539 +548 - -http://dx.doi.org/10.11646/zootaxa.5360.3.8 + +http://dx.doi.org/10.11646/zootaxa.5360.3.8 -journal article -10.11646/zootaxa.5541.4.6 -1175-5326 -14241254 -6ACC4F8A-422B-4E9A-B152-22D649A5980C +journal article +10.11646/zootaxa.5541.4.6 +1175-5326 +14241254 +6ACC4F8A-422B-4E9A-B152-22D649A5980C @@ -65,12 +65,14 @@ - + ( Figs. 1–4 ) + + Diagnosis. Adult specimens of @@ -96,6 +98,8 @@ by the pronotal inner depression reaching the middle of the pronotum, two times Fig. 1C ). + + Description. Body length @@ -208,6 +212,8 @@ As in Fig. 3G (chord broken during dissection). + + Materials examined (n=3, @@ -217,7 +223,7 @@ Materials examined (n=3, - + Holotype (deposited in NIBR @@ -227,31 +233,25 @@ Materials examined (n=3, (dry, abdomen dissected and mounted on plastic card), “ SNU] -Col.#: 210417 ChangW-01 / -Loc -: -Gobdol -mine, 118 / -Dongeup-ro +Col.#: +210417 +ChangW-01 / Loc: Gobdol mine, 118 / +Dongeup-ro , -Changwon-si +Changwon-si , / GN, Korea -. / -Date -: +. / Date: 17. iv. 2021 -/ -Leg -: -Jae Seok Oh +/ Leg: +Jae Seok Oh ”. - + Paratypes (deposited in CBNUIC @@ -259,63 +259,57 @@ mine, 118 / 1♂ (slide mounted), “ SNU] -Col.#: 210417 ChangW-01 / -Loc -: -Gobdol -mine, 118 / -Dongeup-ro +Col.#: +210417 +ChangW-01 / Loc: Gobdol mine, 118 / +Dongeup-ro , -Changwon-si +Changwon-si , / GN, Korea -. / -Date -: +. / Date: 17. iv. 2021 -/ -Leg -: -Jae Seok Oh -” (same data as in -holotype -); +/ Leg: +Jae Seok Oh +” (same data as in holotype); + + 1♂ (slide mounted), “ [SNU] -Col.#: 221025 OJS-01 / -Loc -: -Gobdol -mine, 193-118 / 1055beon-gil, -Dongeup-ro -, / Changwon-si, GN, +Col.#: +221025 OJS-01 +/ Loc: Gobdol mine, 193-118 / 1055beon-gil, +Dongeup-ro +, / +Changwon-si +, GN, Korea . / 35° 19' 30"N 128° 39' 14"E -/ -Date -: +/ Date: 25. x. 2022 -/ -Leg -: -Jaeseok Oh +/ Leg: +Jaeseok Oh ” . + + Etymology . Species name dedicated to the arachnologist, Mr. Jaeseok Oh, collector of the types of this species. + + Habitat. The term “Gobdol” on the label refers to the mineral talc. The collection site is an abandoned talc mine, and the entrance was narrowed with piled soil ( @@ -341,6 +335,8 @@ was collected by sifting bat guano and soil near the entrance (J. Oh & J. Se 25 October 2023 ). + + Distribution. Korea diff --git a/data/03/CC/87/03CC87A3FFEAFFF8FF6584D808CDF81F.xml b/data/03/CC/87/03CC87A3FFEAFFF8FF6584D808CDF81F.xml index c74435f1ea6..63932b3d4da 100644 --- a/data/03/CC/87/03CC87A3FFEAFFF8FF6584D808CDF81F.xml +++ b/data/03/CC/87/03CC87A3FFEAFFF8FF6584D808CDF81F.xml @@ -1,90 +1,93 @@ - - - -New Cretaceous neosuchians (Crocodylomorpha) from Thailand bridge the evolutionary history of atoposaurids and paralligatorids + + + +New Cretaceous neosuchians (Crocodylomorpha) from Thailand bridge the evolutionary history of atoposaurids and paralligatorids - - -Author + + +Author -Pochat-Cottilloux, Yohan -Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France -yohan.pochat@orange.fr +Pochat-Cottilloux, Yohan +Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France +yohan.pochat@orange.fr - - -Author + + +Author -Lauprasert, Komsorn -Department of Biology, Faculty of Science, Mahasarakham University, Maha Sarakham, Thailand & Vertebrate Palaeontology and Evolution Research Unit, Excellence Center in Basin Studies and Applied Paleontology, Mahasarakham University, Khamriang, Maha Sarakham, Thailand +Lauprasert, Komsorn +Department of Biology, Faculty of Science, Mahasarakham University, Maha Sarakham, Thailand & Vertebrate Palaeontology and Evolution Research Unit, Excellence Center in Basin Studies and Applied Paleontology, Mahasarakham University, Khamriang, Maha Sarakham, Thailand - - -Author + + +Author -Chanthasit, Phornphen -Sirindhorn Museum, Department of Mineral Resources, Kalasin, Thailand +Chanthasit, Phornphen +Sirindhorn Museum, Department of Mineral Resources, Kalasin, Thailand - - -Author + + +Author -Manitkoon, Sita -Vertebrate Palaeontology and Evolution Research Unit, Excellence Center in Basin Studies and Applied Paleontology, Mahasarakham University, Khamriang, Maha Sarakham, Thailand & Palaeontological Research and Education Centre, Mahasarakham University, Maha Sarakham, Thailand +Manitkoon, Sita +Vertebrate Palaeontology and Evolution Research Unit, Excellence Center in Basin Studies and Applied Paleontology, Mahasarakham University, Khamriang, Maha Sarakham, Thailand & Palaeontological Research and Education Centre, Mahasarakham University, Maha Sarakham, Thailand - - -Author + + +Author -Adrien, Jérôme -Laboratoire Matériaux, Ingénierie et Science, Institut National des Sciences Appliquées de Lyon, Villeurbanne, France +Adrien, Jérôme +Laboratoire Matériaux, Ingénierie et Science, Institut National des Sciences Appliquées de Lyon, Villeurbanne, France - - -Author + + +Author -Lachambre, Joël -Laboratoire Matériaux, Ingénierie et Science, Institut National des Sciences Appliquées de Lyon, Villeurbanne, France +Lachambre, Joël +Laboratoire Matériaux, Ingénierie et Science, Institut National des Sciences Appliquées de Lyon, Villeurbanne, France - - -Author + + +Author -Amiot, Romain -Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France +Amiot, Romain +Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France - - -Author + + +Author -Martin, Jeremy E. -Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France +Martin, Jeremy E. +Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France -text - - -Zoological Journal of the Linnean Society +text + + +Zoological Journal of the Linnean Society - -2024 - -2024-01-09 + +2024 + +2024-01-09 - -202 + +202 - -1 -27 + +1 +27 -journal article -10.1093/zoolinnean/zlad195 -0024-4082 -30554A2-79AF-4C42-B85A-377B8DB832AE +journal article +306265 +10.1093/zoolinnean/zlad195 +3f2ed02b-39d5-4f58-b764-68f8a5f7e20e +0024-4082 +14335145 +30554A2-79AF-4C42-B85A-377B8DB832AE - + @@ -100,7 +103,7 @@ ( -Figs 2–15 +Figs 2–15 ) @@ -187,7 +190,7 @@ a three-dimensionally preserved nearly complete skull. a crocodylomorph characterized by the following unique combination of features: the dorsal part of the postorbital has an anterolaterally facing edge; the quadrate has no fenestrae; there are two waves of enlarged maxillary teeth; the quadratojugal has no ornamentation; the outer surface of the squamosal is dorsolaterally oriented, reduced, and sculpted; and there is a depression on the posterolateral surface of the maxilla. - + Figure 5. SM-2021-1-98 @@ -218,9 +221,9 @@ and SM- The infratemporal fenestra is preserved on the left side in the two most complete specimens: it is not complete but appears triangular; it is bordered by the jugal laterally, the postorbital anteriorly, and the quadratojugal posteromedially. No spikelike projection inside the fenestra can be seen. In SM- 2023-1- 16 , the antorbital fenestra is present near the triple junction between the maxilla, the lacrimal, and the jugal in lateral view ( -Figs 11D +Figs 11D , -13D +13D ). @@ -237,7 +240,7 @@ is triangular in shape and ~ 1 cm at its widest part. - + Figure 6. SM-2021-1-100 @@ -247,9 +250,9 @@ at its widest part. Cranium ( -Figs 2–4 +Figs 2–4 , -11–14 +11–14 ) @@ -257,15 +260,15 @@ Cranium ( Premasilla ( -Figs 2A, C–E +Figs 2A, C–E , -4A–E, G +4A–E, G , -11A–E, G +11A–E, G , -12A +12A , -13A–D +13A–D ): The premaxilla, together with the anterior part of the nasal, forms the anterior part of the snout. It is straight posteriorly, forming an oblique suture with the maxilla in lateral view. In dorsal view, the opening for the nares is mostly composed of the premaxilla, with participation of the nasal in its posterior part. On the ventral surface, the foramen @@ -279,15 +282,15 @@ The alveoli are circular, and the premaxillary toothrow is curved anteriorly. Masilla ( -Figs 2A, C–E +Figs 2A, C–E , -4A–E, G +4A–E, G , -11A–E, G +11A–E, G , -12A +12A , -13A–D +13A–D ): The two maxillae contact ventrally in an anteroposteriorly straight suture, whereas they are separated dorsally by the nasals, also in an anteroposteriorly straight suture. The triple junction between the maxilla, the premaxilla, and the nasal is situated at the level of the first maxillary alveolus. The contact with the jugal is curved anteriorly to oblique in lateral view. Medially, there is a suture with the ectopterygoid, which is also curved anteriorly. In lateral view, a depression for the insertion of a dentary tooth can be seen at the level of the sixth–seventh maxillary alveoli, where the maxilla curves medially. Dorsally, the maxilla has two planes: one directed dorsomedially to ventrolaterally, and one oriented anteroposteriorly and flat dorsoventrally: the specimens have an altirostral skull. The contact between these two planes is directed posteromedially until the most developed caniniform maxillary tooth (fourth alveolus in @@ -300,7 +303,7 @@ The two maxillae contact ventrally in an anteroposteriorly straight suture, wher 2023-1-16 , it can be assessed that the maxilla contains at least 10 teeth. The largest anterior alveoli, although separated, are the third and the fourth ones, but the fifth one is almost the same size as the third one. A diastema is present between alveoli five and six, probably to accommodate the enlarged dentary tooth. Then the alveoli globally decrease in size again posteriorly; they are mediolaterally compressed and closely spaced. In ventral view, the alveolar rows globally diverge posterolaterally, with a concavity at the level of the fifth–sixth alveolus. The largest tooth is preserved on each side, in addition to some other fragments of other teeth. Those are caniniform teeth, with no particular carinae or ridges, and they are not compressed mediolaterally. - + Figure 7. SM-2021-1-99 @@ -309,7 +312,7 @@ and : caudal vertebrae and limb bones in dorsal (A, C) and ventral (B, D) views. Abbreviations: CD, cervical vertebra; fib, fibula; met, metatarsal; pha, phalanx; r, rib; t, tibia. Scale bars: 1 cm. - + Figure 8. Three-dimensional reconstruction of @@ -317,7 +320,7 @@ Three-dimensional reconstruction of vertebrae in anterior (A) and lateral (B) views. Blue vertebrae are cervical, green are dorsal, red are sacral, and orange are caudal. Abbreviations: c, centrum; dia, diapophysis; ns, neural spine; par, parapophysis; poz, postzygapophysis; prz, prezygapophysis; tp, transverse process. Scale bars: 3 cm. - + Figure 9. Three-dimensional reconstruction of @@ -328,13 +331,13 @@ limb bones and girdles in dorsal (A), lateral (B), and ventral (D) views. C, art Nasal ( -Figs 2A, E +Figs 2A, E , -4A, C, E, G +4A, C, E, G , -11A, E +11A, E , -13A, D +13A, D ): The nasals are paired and elongated anteroposteriorly (no more than @@ -354,13 +357,13 @@ The bone is not visible in ventral view. Lacrimal ( -Figs 2A, C, D +Figs 2A, C, D , -4A–E, G +4A–E, G , -11A, D +11A, D , -13A, C, D +13A, C, D ): The lacrimal is triangular in shape, with one tip directed anteromedially. The posterior curved margin forms a part of the anterior margin of the orbit. The medial side is bordered by the prefrontal, whereas the lateral side contacts the posterior part of the maxilla. The lacrimal is shorter anteroposteriorly than the prefrontal in @@ -374,22 +377,22 @@ The ventral surface is too damaged to be described. In SM- Jugal ( -Figs 2A, C, E, F +Figs 2A, C, E, F , -3A–C, G +3A–C, G , -4 +4 , -11A, C–E, G +11A, C–E, G , -13A, C, D +13A, C, D ): The jugal is elongate and plate-like (taller than wide). Anteriorly, it is directed straight anteroposteriorly, whereas posteriorly it curves mediolaterally (near the contact with the quadratojugal). The jugal sends a convex process to connect with the maxilla on the lateral side, at the same level as the lacrimal, extending to the penultimate alveolus in SM- 2023-1-16. In both specimens, a crest followed by a depression can be seen, with both being directed anteroposteriorly. Posteriorly, the bone connects medially with the quadratojugal almost all the way to the posterior extremity of the skull and almost participates in the articulation with the mandible. The jugal also forms most of the lateral edge of the orbit; however, it does not form a dorsal bulge in this region. Behind the orbit, the jugal has a dorsomedially angled process, which sutures with the postorbital in an anterolaterally curved suture. The posterior margin of this contact, together with the preserved posterior part of the jugal, forms the lateral and anterolateral margins of the infratemporal fenestra. The contact with the quadratojugal is straight, and no foramina or special ornamentation can be seen in dorsal view. - + Figure 10. Three-dimensional reconstruction of @@ -400,13 +403,13 @@ osteoderms. A, dorsal osteoderm of the cervical region. B, C, dorsal (B) and ven Prefrontal ( -Figs 2A, C, D +Figs 2A, C, D , -4A–E, G +4A–E, G , -11A, C–E, G +11A, C–E, G , -13A, C, D +13A, C, D ): The prefrontal is squeezed between the lacrimal laterally and the frontal and the nasal medially, both with straight sutures. Its anteromedialmost part connects with the posterolateralmost part of the nasal. It also connects anteriorly with the maxilla in a mediolaterally straight suture. It is wider anteriorly than posteriorly in @@ -418,15 +421,15 @@ It makes most of the medial margin of the orbit. Ventrally, it is damaged but co Frontal ( -Figs 2A, D +Figs 2A, D , -4A, C–E, G +4A, C–E, G , -11A, C–E +11A, C–E , -13A, C, D +13A, C, D , -14A, F +14A, F ): The frontal forms a bridge between the rostrum and the postorbital region of the skull and makes part of the posteromedial margin of both orbits. Those margins are dorsally raised. Anteriorly, it connects with the nasals in a V-shaped suture, with an anteriorly projected process of the frontal. Posteriorly, it sutures with the parietal and the postorbitals in zigzagged sutural surfaces that are straight overall. Laterally, it does not contact with the lacrimal but with the prefrontal anteromedially. It forms the anteromedial margin of the supratemporal fenestra. The dorsal surface of the frontal is slightly concave and bears a sagittal crest. It is not thick dorsoventrally. In @@ -436,23 +439,23 @@ The frontal forms a bridge between the rostrum and the postorbital region of the , which delimit the olfactory tract ( Iordansky 1973 ; -Fig. 2B +Fig. 2B ). Postorbital ( -Figs 2A, C, D +Figs 2A, C, D , -4A–E, G +4A–E, G , -11A, C–E +11A, C–E , -12B, C +12B, C , -13A, C, D +13A, C, D , -14A, C, D, F +14A, C, D, F ): The postorbital forms the posterior margin of the orbit, the anterolateral margin of the supratemporal fenestra, and the anterior margin of the infratemporal fenestra. It has a T-shape, with one branch directed medially, another one posteriorly, and the final one ventrally. Medially, it contacts only the frontal. Posteriorly, the suture with the squamosal is straight (curved posteriorly in SM- @@ -462,15 +465,15 @@ The postorbital forms the posterior margin of the orbit, the anterolateral margi Parietal ( -Figs 2A, F +Figs 2A, F , -4A, F, G +4A, F, G , -11A, E +11A, E , -13A, E +13A, E , -14A, E +14A, E ): This single bone is part of the skull roof. It has a rectangular shape with depressed sides, like an hourglass. It is less wide mediolaterally than the posterior part of the frontal in @@ -498,17 +501,17 @@ The parietal contacts the frontal anteriorly, the supraoccipital posteriorly, an Squamosal ( -Figs 2A, C, D, F +Figs 2A, C, D, F , -4A, C, D, G +4A, C, D, G , -11A, D, F +11A, D, F , -12B, C +12B, C , -13A, D, E +13A, D, E , -14A, C–E +14A, C–E ): The squamosal is T-shaped and forms the posterolateral corner of the skull roof. It contacts the quadrate anteriorly to the external auditory meatus and the otoccipital posteriorly in @@ -523,7 +526,7 @@ or ventrally in SM- 2023-1-16 , but this is probably a taphonomic artefact. The squamosal has a longitudinal groove for the muscles of the external ear flap located dorsally to the external auditory meatus in lateral view. - + Figure 11. SM-2023-1-16 in dorsal (A), ventral (B), lateral (C, D), anterior (E), posterior (F), and ventrolateral (G) views. Abbreviations: IX– XI, foramen for cranial nerves IX–XI; af, antorbital fenestra; bo, basioccipital; cb, crest B; cd, crest/depression on the jugal; cq, cranioquadrate passage; di, depression for the insertion of a dentary tooth; dpc, deltopectoral crest; ec, ectopterygoid; ef, eustachian foramen; f, frontal; fm, foramen @@ -536,15 +539,15 @@ SM-2023-1-16 in dorsal (A), ventral (B), lateral (C, D), anterior (E), posterior Quadratojugal ( -Figs 2A, C, D +Figs 2A, C, D , -4A, C–E, G +4A, C–E, G , -11A +11A , -12B, C +12B, C , -13A, D +13A, D ): This bone extends posteriorly from the dorsomedial corner of the infratemporal fenestra. The posterior part is thin and squeezed between the quadrate medially and the jugal laterally. It extends all the way to the posterolateralmost part of the skull and participates in the articulation with the mandible. The suture with the jugal is straight in SM- @@ -556,19 +559,19 @@ and curved medially in Quadrate ( -Figs 2A–D, F, G +Figs 2A–D, F, G , -3G +3G , -4 +4 , -11A, D, F +11A, D, F , -12B, C +12B, C , -13 +13 , -14B +14B ): The quadrate has a complex shape. Anteriorly, it does not reach the infratemporal fenestra, but it reaches the ventral margin of the supratemporal fenestra and contacts the parietal. Dorsally, it contacts the squamosal anteriorly to the external auditory meatus in @@ -595,7 +598,7 @@ can be seen on the left quadrate of 1–1.5 cm tall in cross-section and curved dorsally. - + Figure 12. Details of SM-2023-1-16: tooth (A), braincase region (B), and outlines of the bones in the braincase region (C). Abbreviations: V, foramen for cranial nerve V; ec, ectopterygoid; j, jugal; ls, laterosphenoid; pbs, parabasisphenoid; pbsr, basisphenoid rostrum; pro, prootic; pt, pterygoid; s, squamosal. Scale bars: 1 cm. @@ -604,15 +607,15 @@ Details of SM-2023-1-16: tooth (A), braincase region (B), and outlines of the bo Supraoccipital ( -Figs 2A, F +Figs 2A, F , -4A, F +4A, F , -11A, F +11A, F , -13A, E +13A, E , -14E +14E ): This bone is triangular in shape in posterior view.It is exposed in dorsal view. Laterally, it connects with the otoccipital. It also does not participate in the formation of the foramen @@ -628,15 +631,15 @@ because those areas are covered by other fragments. Otoccipital ( -Figs 2F +Figs 2F , -4A, C, D, F, G +4A, C, D, F, G , -11F +11F , -13E +13E , -14E +14E ): This bone connects laterally with the quadrate, medially with the supraoccipital, and dorsally with the squamosal in a straight to curved contact. In posterior view, it extends laterally to the lateral edge of the skull (not in SM- @@ -650,13 +653,13 @@ there are three foramina: the lateralmost one is for the internal carotid artery Basioccipital ( -Figs 2B, D, F, G +Figs 2B, D, F, G , -4A, B, F +4A, B, F , -11B, F +11B, F , -13B, E +13B, E ): The basioccipital sutures laterally with the otoccipital (in posterior view), the quadrate (in lateral view), and maybe the pterygoid posteroventrally (in @@ -666,15 +669,15 @@ The basioccipital sutures laterally with the otoccipital (in posterior view), th Palatine ( -Figs 2B, G +Figs 2B, G , -3D, E +3D, E , -4B, G +4B, G , -11B +11B , -13B +13B ): The palatine connects with the maxilla in an anteriorly convex suture in @@ -684,7 +687,7 @@ and SM- The posterior region is heavily damaged and remodelled, but it does not include the choanae in SM-2021-1-97/99 ( -Fig. 3G +Fig. 3G ). However, this could be the case in SM- 2023-1-16 , where the palatine would form its anterior margin, but because of the poor state of preservation it is difficult to assess. The suture with the pterygoid is also anteroposteriorly straight immediately anteriorly to the internal choanae. The paired palatines are flat when they meet at the midline. In ventral view, the lateral margins of the palatines are parallel and straight in @@ -695,7 +698,7 @@ The posterior region is heavily damaged and remodelled, but it does not include 2023-1-16. The vomer cannot be seen. - + Figure 13. Outlines of the bones of SM-2023-1-16 in dorsal (A), ventral (B), lateral (C, D), and posterior (E) views. Abbreviations: IX–XI, foramen for cranial nerve IX–XI; af, antorbital fenestra; bo, basioccipital; cb, crest B; cd, crest/depression on the jugal; cq, cranioquadrate passage; dpc, deltopectoral crest; ec, ectopterygoid; ef, eustachian foramen; f, frontal; fm, foramen magnum; h, humerus; ic, internal choanae; if, infratemporal fenestra; j, jugal; l, lacrimal; m, maxilla; n, nasal; o, orbit; oc, occipital condyle; ot, otoccipital; p, parietal; pal, palatine; pbs, parabasisphenoid; pf, prefrontal; pfp, prefrontal pillar; pm, premaxilla; po, postorbital; ps, proximal surface; pt, pterygoid; q, quadrate; qj, quadratojugal; s, squamosal; sf, supratemporal fenestra; sh, squamosal ‘horn’; sm, maxillary sulcus; so, supraoccipital; sub, suborbital fenestra; v, vertebrae remains. Scale bars: 1 cm. @@ -704,17 +707,17 @@ Outlines of the bones of SM-2023-1-16 in dorsal (A), ventral (B), lateral (C, D) Pterygoid ( -Figs 2B, D, F, G +Figs 2B, D, F, G , -3C–F +3C–F , -4B–D, F, G +4B–D, F, G , -11C +11C , -12B, C +12B, C , -13B, C +13B, C ): The pterygoid is sutured to the palatine anteriorly, the ectopterygoid laterally, and the basioccipital posteriorly. Each pterygoid makes a straight connection with the corresponding ectopterygoid and tends to be more curved ventrally at that point. The internal choanae are ovoid, bordered by prominent anterior margins in ventral view, and have no midline process. They are totally enclosed by the pterygoids in @@ -723,7 +726,7 @@ The pterygoid is sutured to the palatine anteriorly, the ectopterygoid laterally 2023-1-16 ) and are situated anteriorly to the posterior margin of the suborbital fenestra. Anteriorly, a median process of the pterygoid extends to contact the palatine and forms the ventral edge of the interorbital septum. The pterygoid is smooth on all surfaces except on the lateralmost sides. - + Figure 14. SM-2023-1-17 in dorsal (A), ventral (B), lateral (C, D), posterior (E), and anterior (F) views. Abbreviations: cb, crest B; f, frontal; ot, otoccipital; p, parietal; po, postorbital; q, quadrate; s, squamosal; sf, supratemporal fenestra; so, supraoccipital. Scale bars: 1 cm. @@ -732,17 +735,17 @@ SM-2023-1-17 in dorsal (A), ventral (B), lateral (C, D), posterior (E), and ante Ectopterygoid ( -Figs 2B, D +Figs 2B, D , -3C, D–F +3C, D–F , -4B, C, F, G +4B, C, F, G , -11D +11D , -12B, C +12B, C , -13B, D +13B, D ): This bone contacts the jugal and the maxilla laterally and the pterygoid medially (forming the pterygoid flange). The triple junction between the maxilla, the jugal, and the ectopterygoid is situated at the middle of the anterior process of the ectopterygoid in @@ -754,7 +757,7 @@ This bone contacts the jugal and the maxilla laterally and the pterygoid mediall Laterosphenoid ( -Fig. 12B, C +Fig. 12B, C ): The dorsoposterior ridge of the laterosphenoid connects with the parietal. The ventralmost part of the bone contacts the quadrate, forming the foramen for cranial nerve V. @@ -762,11 +765,11 @@ The dorsoposterior ridge of the laterosphenoid connects with the parietal. The v Parabasisphenoid ( -Figs 11C, D +Figs 11C, D , -12B, C +12B, C , -13C, D +13C, D ): The parabasisphenoid is complex. Anteriorly, the parabasisphenoid rostrum is short and not dorsoventrally elevated. It does not contact the laterosphenoid; however, it does contact the prootic and the pterygoid. Posteriorly, it has a plate-like shape directed anterolaterally to posteromedially. It contacts the pterygoid ventrally, the quadrate dorsally, and the basioccipital posteriorly, encircling the pharyngeal foramen. This part also bears a crest directed anteroventrally to posterodorsally. @@ -774,7 +777,7 @@ The parabasisphenoid is complex. Anteriorly, the parabasisphenoid rostrum is sho Prootic ( -Fig. 12B, C +Fig. 12B, C ): This small bone sutures with the laterosphenoid posteriorly on the posterior margin of the foramen for cranial nerve V and the parabasisphenoid rostrum anteriorly. @@ -782,11 +785,11 @@ This small bone sutures with the laterosphenoid posteriorly on the posterior mar Mandible ( -Figs 2B, G +Figs 2B, G , -3 +3 , -4 +4 ) @@ -794,16 +797,16 @@ Mandible ( Dentary ( -Figs 2B–E, G +Figs 2B–E, G , -3B, C +3B, C , -4B–G +4B–G ): This bone is the only tooth-bearing element of the mandible, with at least 13 alveoli. The two rami separate at the level of the fifth dentary tooth and are firmly sutured anteriorly. The symphyseal region remains wide anteriorly and is U-shaped at its anteriormost point. In medial view, the dentary is thin dorsoventrally and forms an acute angle anteriorly. Ventrally and dorsally, the medial suture with the splenial is oblique, directed posterolaterally to anteromedially. The dentary remains uniform in width up to the point of divergence, where it begins to taper off. Posteriorly, the dentary connects dorsally with the surangular, from the end of the toothrow, and ventrally with the angular at its posteriormost part. The ventral surface is smooth, and anteriorly it is curved dorsally. The lateral side shows no groove ventrally to the tooth row. In lateral view, the dorsal margin is sinusoidal, marked by two sets of waves that culminate at the level of the 3rd/4th and 11th/12fth dentary alveoli. There is also a medial depression to accommodate the largest caniniform maxillary tooth at the level of the 9th dentary alveolus, especially visible on the left side. In terms of size, alveoli 4–10 and 13 are relatively small, whereas the 3rd one is the largest, and the 11th and 12th are also large. Alveoli 4–7 and 8–10 are closely spaced, whereas there is a larger space between alveoli 7 and 8. The ventral margin of the dentary is straight anteroposteriorly. - + Figure 15. Different postcranial parts (SM-2023-1-17): three linked vertebral centra (A), two vertebral centra (B, C), upper part of a vertebra in dorsal (D) and ventral (E) views, and proximal part of limb bone in anterior (F) and posterior (G) views. Abbreviations: tp, tranverse process; V1, vertebra 1; V2, vertebra 2; V3, vertebra 3; zp, zygapophysis. Scale bars: 1 cm. @@ -812,11 +815,11 @@ Different postcranial parts (SM-2023-1-17): three linked vertebral centra (A), t Splenial ( -Figs 2B, G +Figs 2B, G , -3C +3C , -4 +4 ): The splenial is most exposed medially; it forms a vertical plate that sutures with the dentary and with the other splenial at the midline anteriorly. It reaches up to the fifth dentary tooth and is not exposed ventrally. Posteriorly, it becomes thin and plate-like along the medial surface of the dentary (even becoming the medial wall for the last posterior dentary alveoli, from the 11th one) until it disappears posteriorly without meeting the angular or the surangular. In posterior view, the circular foramen @@ -826,11 +829,11 @@ is present at the point where the two splenials diverge (not on the medial sides Angular ( -Figs 2B, D, F, G +Figs 2B, D, F, G , -3A, B, H +3A, B, H , -4B–D, 4F, G +4B–D, 4F, G ): The angular is the ventralmost mandibular element (approximately half of the total length of the mandible). In lateral view, it is elongated, and posteriorly it is curved dorsally. Medially, the angular sutures with the dentary, the surangular, and the articular to form a huge medial depression (adductor chamber; @@ -842,11 +845,11 @@ is not very developed and is visible only in the posteroventral margin of the an Surangular ( -Figs 2C, F +Figs 2C, F , -3A–C, G, H +3A–C, G, H , -4 +4 ): This bone is robust and elongated. Anteriorly, its dorsal process might extend between the dentary and the splenial, but the preserved parts of the specimen show only a contact between the surangular and the dentary anteriorly. Posteriorly, it tends to curve dorsally and sutures with the angular for the rest of its length and medially with the articular. It also becomes more plate-like. This suture is linear anteriorly, and posteriorly it curves dorsally. The bone also curves medially and forms the lateral margin of the articular fossa. Laterally to this, there is a short ridge oriented anteroposteriorly, forming a depression. The dorsal surface is convex, not ornamented, and flattens before the glenoid surface. On the lateral surface, there is a ridge directed posterodorsally to anteroventrally. @@ -854,11 +857,11 @@ This bone is robust and elongated. Anteriorly, its dorsal process might extend b Articular ( -Figs 2A, B, D, F, G +Figs 2A, B, D, F, G , -3G, H +3G, H , -4A, B, D–F +4A, B, D–F ): The articular is the posteriormost element of the mandible. It has two dorsal surfaces separated by a ridge oriented mediolaterally. The anterior surface articulates with the quadrate; the ridge helping to stabilize this articulation as its posterior wall is tall and dorsally edged. The articular fossa is divided into a lateral and a medial portion of equal size by a small ridge oriented anteroposteriorly. The posterior surface (retroarticular process) is concave overall and paddle shaped. It seems to taper posteriorly, but the posteriormost part is broken on each side. Ventromedially, the articular forms the posteromedial wall of the adductor chamber. It sutures with the surangular at its lateral margin and with the angular at its ventral margin. @@ -866,11 +869,11 @@ The articular is the posteriormost element of the mandible. It has two dorsal su Dentition ( -Figs 2C–E, H, I +Figs 2C–E, H, I , -4C–E, G +4C–E, G , -12A +12A ): A large maxillary caniniform tooth is preserved on each side of the skull. It is conical and slightly curved lingually. The base of the tooth crown is ovoid in cross-section, and the apex is pointed. There are no carinae or crenulations visible in @@ -910,11 +913,11 @@ Tennant Axial skeleton ( -Figs 5–8 +Figs 5–8 , -10 +10 , -15A–E +15A–E ) @@ -930,11 +933,11 @@ Most crocodylians possess eight cervical vertebrae, 16 dorsals (lumbar included) , there are at least 32 vertebrae. Given that the areas of transition between the cervicals and the anterior dorsals ( SM-2021-1-98 ; -Fig. 5 +Fig. 5 ) and between the posterior dorsals and the sacrals ( SM-2021-1-100 ; -Fig. 6 +Fig. 6 ) are preserved, we can establish the following: SM-2021-1-97/101 preserves 4 cervicals (5 @@ -948,9 +951,9 @@ preserves 4 cervicals (5 Cervicals ( -Figs 5A–C, E +Figs 5A–C, E , -8 +8 ): The neural arches and spines are tall, narrow, and pointed dorsally. The diapophyseal processes are longer than the parapophyseal processes. The zygapophyses are large in comparison to the centra, with the prezygapophyses being larger than the postzygapophyses. These structures become horizontal posteriorly. All the centra are amphicoelous, and the lateral sides of the centra (between the diapophyseal and parapophyseal processes) are notably depressed. The last cervical vertebra has a tall neural spine (more than two times the height of the centrum). The hypapophyses are broken, but their areas of insertion on the centrum are still visible on all centra. The diapophyseal processes gradually increase in size posteriorly, and they also migrate from the lateral side of the centrum to the lateral side of the neural arch. The parapophyseal processes do not seem to increase in size, but they also migrate dorsally, going from the ventrolateral margin to the lateral side of the centrum posteriorly. @@ -958,11 +961,11 @@ The neural arches and spines are tall, narrow, and pointed dorsally. The diapoph Dorsals ( -Figs 5B–D +Figs 5B–D , -6B, C +6B, C , -8 +8 ): The neural spines are anteroposteriorly longer than those of the cervicals. The first dorsal vertebra has the tallest neural spine (more than two times the height of the centrum). The diapophyseal and parapophyseal processes are too damaged to be described. On the centrum of the first dorsal vertebra, the proximal part of the hypapophysis is preserved; it is half the size of the centrum. The ventral hypapophyses are either absent or too damaged to be seen from the second or third centra. The articular facets of the pre- and postzygapophyses are more horizontally oriented. @@ -970,9 +973,9 @@ The neural spines are anteroposteriorly longer than those of the cervicals. The Sacrals ( -Figs 6 +Figs 6 , -8 +8 ): There are two sacral vertebrae of the same size. The base of their neural spine is long anteroposteriorly for both. On the second vertebra, the transverse processes and their articular surfaces are more developed, probably because they are less damaged than on the first one. The @@ -982,11 +985,11 @@ is directed ventrally on the first sacral. The contact between the two centra is Caudals ( -Figs 6 +Figs 6 , -7B–D +7B–D , -8 +8 ): The first caudal vertebra is large, and its articular surfaces are concave to flat. The other ones are damaged. The second has a neural spine that is preserved; it is high dorsoventrally and long anteroposteriorly. Some of them have two horizontal ridges on the centrum, perhaps for the insertion of the chevron. A chevron is preserved; it is triangular in shape and open in the middle. @@ -994,9 +997,9 @@ The first caudal vertebra is large, and its articular surfaces are concave to fl Ribs ( -Figs 5A +Figs 5A , -7B +7B ): Some ribs are preserved, but it is difficult to identify to which vertebra they were attached. When those structures are preserved, the tuberculum is less developed than the capitulum. The ribs are slightly curved and are rod-like. @@ -1004,14 +1007,14 @@ Some ribs are preserved, but it is difficult to identify to which vertebra they Pectoral girdle ( -Fig.9 +Fig.9 ) Coracoid ( -Fig. 9 +Fig. 9 ): The left coracoid is preserved. This bone is convex overall. The shaft is triangular, and both ends of the bone are extended. The coracoid foramen is visible in the most proximal part. The articular surface with the scapula is flat, and the ventral part of the glenoid fossa is saddle shaped and directed posteriorly. The distal end is more developed anteriorly than posteriorly. @@ -1019,18 +1022,18 @@ The left coracoid is preserved. This bone is convex overall. The shaft is triang Pelvic girdle ( -Figs 6B, C +Figs 6B, C , -9A, B, D +9A, B, D ) Pubis ( -Figs 6B, C +Figs 6B, C , -9A, B, D +9A, B, D ): The two bones are flat. The most proximal part articulating with the rest of the pelvic girdle is missing in both. Overall, it has a round shape, with a convex and narrow dorsal margin. @@ -1038,18 +1041,18 @@ The two bones are flat. The most proximal part articulating with the rest of the Forelimb ( -Figs 11B, C, G +Figs 11B, C, G , -13B, C +13B, C ) Humerus ( -Figs 11B, C, G +Figs 11B, C, G , -13B, C +13B, C ): The humerus is squeezed between the anterior margin of the suborbital fenestra and the right side of the parabasisphenoid rostrum. Its deltopectoral crest is slender and placed anteroproximally. On the proximomedial side, there is a ridge that connects the deltopectoral crest and the triangular-shaped head of the humerus. The humerus is straight and not very curved or twisted. The head of the humerus is very well developed and unusual among crocodyliforms, extending posterolaterally. The articular surface is flat and decreases anteromedially to articulate with the glenoid fossa. There is also a lateral projection where the humeral head reaches the shaft. The area of insertion for the @@ -1059,18 +1062,18 @@ is visible on the posterior side. Distally, on the bicondylar articulation with Hindlimb ( -Figs 7A, B +Figs 7A, B , -9A +9A , BD) Tibia( -Figs 7A, B +Figs 7A, B , -9A, B, D +9A, B, D ): The shaft of the left tibia, although deformed taphonomically, appears to be straight anteroposteriorly and curved laterally. However, both ends of the bone are on the same plane mediolaterally. The proximal articulation surface is flat, with a posterior concavity. The distal articulation surface is ovoid. @@ -1078,7 +1081,7 @@ The shaft of the left tibia, although deformed taphonomically, appears to be str Tibia ( -Fig 9A, B, D +Fig 9A, B, D ): The fibula is straight and very slender. The proximal end is compressed mediolaterally. The distal articular surface comprises two articular surfaces that are flat to convex: one is posteromedial and would have articulated with the calcaneum, and the other one is distal and would have articulated with the astragalus. @@ -1086,7 +1089,7 @@ The fibula is straight and very slender. The proximal end is compressed mediolat Digits ( -Fig 9A, B, D +Fig 9A, B, D ): The preserved metatarsal is concave ventrally. The distal end is separated into two condyles, with a depression on each side laterally and medially to the hemicondyles. The preserved ungual is curved ventrally and pike shaped. @@ -1094,9 +1097,9 @@ The preserved metatarsal is concave ventrally. The distal end is separated into Osteoderms ( -Figs 5–7 +Figs 5–7 , -10 +10 ) @@ -1114,29 +1117,29 @@ either had at least four rows of paravertebral osteoderms or at least two rows o Dorsal shield ( -Figs 5 +Figs 5 , -6A, B +6A, B , -7A, C +7A, C , -10A, B, D +10A, B, D ): The osteoderms are more expanded mediolaterally anteriorly than posteriorly, but their anteroposterior size remains the same. All osteoderms are flat or slightly arched dorsally and ornamented with circular pits, as is seen on the cranial table and the posterolateral side of the mandible. There are some spine- or peg-like processes for articulation with the more anteriorly situated osteoderm that are preserved ( -Fig. 10A +Fig. 10A ). The articulation system is as follows: the anterior osteoderm overlaps the posterior osteoderm, and the left osteoderm tends to overlap the right one. The margins are straight. Ventral shield ( -Figs 5 +Figs 5 , -6C +6C , -7B, D +7B, D , -10C, E +10C, E ): The ventral shield is more damaged; however, it consists of square osteoderms. There are at least two rows of osteoderms. The ornamentation is the same as the dorsal shield; it consists of flat circular pits. The ventral shield is missing in the sacral region. @@ -1160,7 +1163,7 @@ as a monophyletic group (including Varanosuchus sakonnakhonensis ; -Fig. 16 +Fig. 16 , node 4) based on the following combination of characters: a broad altirostral skull (character 3); little participation of the premaxilla in the internarial bar (character 4); the quadrate, squamosal, and otoccipital do not meet to enclose the cranioquadrate passage (character 49); the antorbital fenestra is much smaller than the orbit (character 67); one wave of enlarged maxillary teeth (character 79); dorsal osteoderms with a well-developed process located anterolaterally in dorsal parasagittal osteoderms (character 96); two parallel rows of dorsal osteoderms (character 97); a symmetrically developed lateral compression on the maxillary teeth (character 140); and a lacrimal that tapers posteroventrally and does not contact or only slightly contacts the jugal (character 229). @@ -1175,7 +1178,7 @@ Rummy forms a monophyletic group ( -Fig. 16 +Fig. 16 , node 5) defined by the following synapomorphies: no vascular opening on the dorsal surface of the postorbital bar (character 27); the medial quadrate condyle expands ventrally, being separated from the lateral condyle by a deep intercondylar groove (character 170); a sharp ridge along the lateral surface of the angular (character 219); the ulna has a wide and rounded olecranon process (character 260); and a foramen located in palatal view on the premaxilla–maxilla suture near the alveolar border (character 320). @@ -1192,12 +1195,12 @@ Eusuchians (including , and Crocodylia ; -Fig. 16 +Fig. 16 , node 3) are also retrieved as a monophyletic group, with the following synapomorphies: the choanal groove is undivided (character 69); the cervical vertebrae are procoelous (character 92); the dorsal osteoderms have a discrete convexity on the anterior margin (character 96); there are more than two rows of dorsal primary osteoderms (character 97); and the supraoccipital is exposed in the skull roof (character 171). Finally, the two most complete specimens from Phu Sung are retrieved as a monophyletic group ( -Fig. 16 +Fig. 16 , node 8), with the following synapomorphies: the dorsal part of the postorbital has an anterolaterally facing edge (character 29); the quadrate has no fenestrae (character 45); two waves of enlarged maxillary teeth (festooned; character 79); a quadratojugal with no ornamentation (character 145); the outer surface of the squamosal dorsolaterally oriented is reduced and sculpted (character 168); and there is a depression on the posterolateral surface of the maxilla (character 207). diff --git a/data/03/CC/87/03CC87A3FFFFFFFAFC3E84D50A8CFAED.xml b/data/03/CC/87/03CC87A3FFFFFFFAFC3E84D50A8CFAED.xml index 0eec57b3b6b..03f31ce034a 100644 --- a/data/03/CC/87/03CC87A3FFFFFFFAFC3E84D50A8CFAED.xml +++ b/data/03/CC/87/03CC87A3FFFFFFFAFC3E84D50A8CFAED.xml @@ -1,90 +1,93 @@ - - - -New Cretaceous neosuchians (Crocodylomorpha) from Thailand bridge the evolutionary history of atoposaurids and paralligatorids + + + +New Cretaceous neosuchians (Crocodylomorpha) from Thailand bridge the evolutionary history of atoposaurids and paralligatorids - - -Author + + +Author -Pochat-Cottilloux, Yohan -Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France -yohan.pochat@orange.fr +Pochat-Cottilloux, Yohan +Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France +yohan.pochat@orange.fr - - -Author + + +Author -Lauprasert, Komsorn -Department of Biology, Faculty of Science, Mahasarakham University, Maha Sarakham, Thailand & Vertebrate Palaeontology and Evolution Research Unit, Excellence Center in Basin Studies and Applied Paleontology, Mahasarakham University, Khamriang, Maha Sarakham, Thailand +Lauprasert, Komsorn +Department of Biology, Faculty of Science, Mahasarakham University, Maha Sarakham, Thailand & Vertebrate Palaeontology and Evolution Research Unit, Excellence Center in Basin Studies and Applied Paleontology, Mahasarakham University, Khamriang, Maha Sarakham, Thailand - - -Author + + +Author -Chanthasit, Phornphen -Sirindhorn Museum, Department of Mineral Resources, Kalasin, Thailand +Chanthasit, Phornphen +Sirindhorn Museum, Department of Mineral Resources, Kalasin, Thailand - - -Author + + +Author -Manitkoon, Sita -Vertebrate Palaeontology and Evolution Research Unit, Excellence Center in Basin Studies and Applied Paleontology, Mahasarakham University, Khamriang, Maha Sarakham, Thailand & Palaeontological Research and Education Centre, Mahasarakham University, Maha Sarakham, Thailand +Manitkoon, Sita +Vertebrate Palaeontology and Evolution Research Unit, Excellence Center in Basin Studies and Applied Paleontology, Mahasarakham University, Khamriang, Maha Sarakham, Thailand & Palaeontological Research and Education Centre, Mahasarakham University, Maha Sarakham, Thailand - - -Author + + +Author -Adrien, Jérôme -Laboratoire Matériaux, Ingénierie et Science, Institut National des Sciences Appliquées de Lyon, Villeurbanne, France +Adrien, Jérôme +Laboratoire Matériaux, Ingénierie et Science, Institut National des Sciences Appliquées de Lyon, Villeurbanne, France - - -Author + + +Author -Lachambre, Joël -Laboratoire Matériaux, Ingénierie et Science, Institut National des Sciences Appliquées de Lyon, Villeurbanne, France +Lachambre, Joël +Laboratoire Matériaux, Ingénierie et Science, Institut National des Sciences Appliquées de Lyon, Villeurbanne, France - - -Author + + +Author -Amiot, Romain -Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France +Amiot, Romain +Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France - - -Author + + +Author -Martin, Jeremy E. -Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France +Martin, Jeremy E. +Universite Claude Bernard Lyon 1, LGL-TPE, UMR 5276, CNRS, ENS Lyon, Villeurbanne, France -text - - -Zoological Journal of the Linnean Society +text + + +Zoological Journal of the Linnean Society - -2024 - -2024-01-09 + +2024 + +2024-01-09 - -202 + +202 - -1 -27 + +1 +27 -journal article -10.1093/zoolinnean/zlad195 -0024-4082 -30554A2-79AF-4C42-B85A-377B8DB832AE +journal article +306265 +10.1093/zoolinnean/zlad195 +3f2ed02b-39d5-4f58-b764-68f8a5f7e20e +0024-4082 +14335145 +30554A2-79AF-4C42-B85A-377B8DB832AE - + @@ -110,7 +113,7 @@ The status of ‘ Buffetaut & Ingavat, 1983 consists of an incomplete dentary ( -Fig. 17 +Fig. 17 ) and is considered by some as belonging to the genus Sunosuchus @@ -141,7 +144,7 @@ Puértolas-Pascual 2015 ). This taxon is diagnosed on a fragment of dentary, by the following characters: dentary symphysis reaching the level of the sixth alveoli (i.e. five alveoli completely included); no marked angulation of the lower tooth row in its anterior part; no strong outward projection of the lateral rims of the third and fourth dentary alveoli; and an alveolar edge strongly undulated in lateral view ( -Fig. 17 +Fig. 17 ). Although the specimen diagnosing ‘ Goniopholis