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<document id="AF77FEC69E4EB01F28422819061FDD78" ID-DOI="10.11646/zootaxa.5519.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13916275" ID-ZooBank="4FDC3321-611F-413C-B543-290E61712C43" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728587748317" checkinUser="plazi" docAuthor="Jelínek, Josef &amp; Hájek, Jiří" docDate="2024" docId="1523AC3AF8292965BDE987BAFC9EF972" docLanguage="en" docName="zootaxa.5519.4.7.pdf" docOrigin="Zootaxa 5519 (4)" docSource="http://zoobank.org/4fdc3321-611f-413c-b543-290e61712c43" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Viettherchnus helferi Jelínek &amp; Hájek, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="599" masterDocId="E91AD442F82F296CBD7E8205FF88FFCE" masterDocTitle="Three new species of the genus Viettherchnus Kirejtshuk, 1985 (Coleoptera: Nitidulidae: Nitidulinae: Cyllodini)" masterLastPageNumber="600" masterPageNumber="590" pageNumber="596" updateTime="1728588161917" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="21F203DA38DB2416EF76B84142EC1AE5">Three new species of the genus Viettherchnus Kirejtshuk, 1985 (Coleoptera: Nitidulidae: Nitidulinae: Cyllodini)</mods:title>
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<mods:namePart id="E58AAF4F2D19B05E1F58D285722719D6">Jelínek, Josef</mods:namePart>
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<mods:namePart id="D389AA021957969CC0481244A9B39E9A">Hájek, Jiří</mods:namePart>
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<treatment id="1523AC3AF8292965BDE987BAFC9EF972" LSID="urn:lsid:plazi:treatment:1523AC3AF8292965BDE987BAFC9EF972" httpUri="http://treatment.plazi.org/id/1523AC3AF8292965BDE987BAFC9EF972" lastPageId="9" lastPageNumber="599" pageId="6" pageNumber="596">
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<paragraph id="9D351D2CF829296ABDE987BAFE75FA14" blockId="6.[151,509,1471,1535]" box="[151,509,1471,1498]" pageId="6" pageNumber="596">
<heading id="C67DAA40F829296ABDE987BAFE75FA14" bold="true" box="[151,509,1471,1498]" fontSize="11" level="1" pageId="6" pageNumber="596" reason="1">
<emphasis id="AFFEC13EF829296ABDE987BAFE75FA14" bold="true" box="[151,509,1471,1498]" pageId="6" pageNumber="596">
<taxonomicName id="5A8A66AFF829296ABDE987BAFE10FA14" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[151,408,1471,1498]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="596" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF829296ABDE987BAFE10FA14" bold="true" box="[151,408,1471,1498]" italics="true" pageId="6" pageNumber="596">Viettherchnus helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F829296ABCE187C5FE75FA14" box="[415,509,1472,1498]" pageId="6" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="D5904EA7F829296ABDE987E1FE24FA30" box="[151,428,1508,1535]" pageId="6" pageNumber="596" type="description">
<paragraph id="9D351D2CF829296ABDE987E1FE24FA30" blockId="6.[151,509,1471,1535]" box="[151,428,1508,1535]" pageId="6" pageNumber="596">
(
<figureCitation id="05B101A9F829296ABDE187E1FF66FA30" box="[159,238,1508,1534]" captionStart="FIGURES 14" captionStartId="3.[152,270,1922,1946]" captionTargetBox="[168,1421,213,1867]" captionTargetId="figure-32@3.[151,1436,181,1899]" captionTargetPageId="3" captionText="FIGURES 14. Dorsal habitus of Viettherchnus. 1. V. orszuliki sp. nov. (holotype ♂); 2. V. apriculus sp. nov. (holotype ♂); 3. V. helferi sp. nov. (paratype ♀, dark form; Kalimantan); 4. V. helferi sp. nov. (holotype ♂). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916277" httpUri="https://zenodo.org/record/13916277/files/figure.png" pageId="6" pageNumber="596">Figs. 3</figureCitation>
,
<figureCitation id="05B101A9F829296ABD8587E0FE83FA31" box="[251,267,1509,1535]" captionStart="FIGURES 57" captionStartId="4.[152,270,932,956]" captionTargetBox="[176,1411,207,909]" captionTargetId="figure-19@4.[176,1411,181,909]" captionTargetPageId="4" captionText="FIGURES 57. Head of male Viettherchnus. 5. V. orszuliki sp. nov.; 6. V. apriculus sp. nov.; 7. V. helferi sp. nov. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.13916279" httpUri="https://zenodo.org/record/13916279/files/figure.png" pageId="6" pageNumber="596">7</figureCitation>
,
<figureCitation id="05B101A9F829296ABC6987E1FEAFFA30" box="[279,295,1508,1534]" captionStart="FIGURES 89" captionStartId="5.[152,270,1037,1061]" captionTargetBox="[173,1417,205,998]" captionTargetId="figure-32@5.[151,1436,181,1013]" captionTargetPageId="5" captionText="FIGURES 89. Ventral habitus of Viettherchnus. 8. V. orszuliki sp. nov. (paratype ♂); 9. V. helferi sp. nov. (paratype ♀; Kalimantan). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916281" httpUri="https://zenodo.org/record/13916281/files/figure.png" pageId="6" pageNumber="596">9</figureCitation>
,
<figureCitation id="05B101A9F829296ABC4D87E1FE2CFA30" box="[307,420,1508,1535]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="6" pageNumber="596">12, 15, 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F829296ABDE98429FA8DF8D0" pageId="6" pageNumber="596" type="materials_examined">
<paragraph id="9D351D2CF829296ABDE98429FA8DF8D0" blockId="6.[151,1437,1580,2039]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDE98429FEC1F988" bold="true" box="[151,329,1580,1606]" pageId="6" pageNumber="596">Type material.</emphasis>
<materialsCitation id="2DE21771F829296ABC2D8429FE24F918" collectingDate="2011-11-30" collectorName="J. Hajek &amp; J. Schneider &amp; P. Votruba" country="Indonesia" determinerName="Hajek" elevation="66" latitude="-1.2233334" location="KALIMANTAN" longLatPrecision="130" longitude="116.37666" pageId="6" pageNumber="596" specimenCount="2" specimenCount-male="2" stateProvince="Kalimantan Utara" typeStatus="holotype">
<typeStatus id="4231A38EF829296ABC2D8429FE33F988" box="[339,443,1580,1606]" pageId="6" pageNumber="596" type="holotype">Holotype</typeStatus>
<specimenCount id="8B8CD6A5F829296ABCB88428FE53F989" box="[454,475,1581,1607]" count="1" pageId="6" pageNumber="596" type="male"></specimenCount>
(NMPC), labelled: “
<collectingCountry id="E59D5DBCF829296ABFAC8429FCE5F988" box="[722,877,1580,1606]" name="Indonesia" pageId="6" pageNumber="596">INDONESIA</collectingCountry>
, E
<collectingRegion id="5F4ED3CEF829296ABEE68429FBA2F988" box="[920,1066,1580,1606]" country="Indonesia" name="Kalimantan Utara" pageId="6" pageNumber="596">KALIMANTAN</collectingRegion>
/ca.
<quantity id="5A72B0C9F829296AB9198429FB3AF988" box="[1127,1202,1580,1607]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="596" unit="km" value="50.0">50 km</quantity>
<collectingRegion id="5F4ED3CEF829296AB9C28428FA02F988" box="[1212,1418,1580,1607]" country="Indonesia" name="W of BALIKPAPAN" pageId="6" pageNumber="596">W of BALIKPAPAN</collectingRegion>
/ PT Fajar Surya Swadaya [area] /
<geoCoordinate id="F8BE7BEBF829296ABF6C8454FD0AF9A2" box="[530,642,1617,1644]" degrees="01" direction="south" minutes="13.4" orientation="latitude" pageId="6" pageNumber="596" precision="92" value="-1.2233334">01°13.4S</geoCoordinate>
,
<geoCoordinate id="F8BE7BEBF829296ABFF18454FC86F9A2" box="[655,782,1617,1644]" degrees="116" direction="east" minutes="22.6" orientation="longitude" pageId="6" pageNumber="596" precision="92" value="116.37666">116°22.6E</geoCoordinate>
,
<quantity id="5A72B0C9F829296ABE658455FCDDF9A5" box="[795,853,1616,1643]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="596" unit="m" value="66.0">
<elevation id="16A7FA1FF829296ABE658455FCDDF9A5" box="[795,853,1616,1643]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="596" unit="m" value="66.0">66 m</elevation>
</quantity>
/
<collectorName id="307F78FAF829296ABE138454FC45F9A4" box="[877,973,1616,1643]" pageId="6" pageNumber="596">J. Hájek</collectorName>
,
<collectorName id="307F78FAF829296ABEA78454FBEBF9A4" box="[985,1123,1616,1643]" pageId="6" pageNumber="596">J. Schneider</collectorName>
&amp; /
<collectorName id="307F78FAF829296AB9E78454FA85F9A4" box="[1177,1293,1616,1643]" pageId="6" pageNumber="596">P. Votruba</collectorName>
leg., 27+
<date id="E9343BECF829296AB8058455FF63F940" pageId="6" pageNumber="596" value="2011-11-30">
<collectingDate id="F970C204F829296AB8058455FF63F940" pageId="6" pageNumber="596" value="2011-11-30">30. xi.2011</collectingDate>
</date>
[p] // stream in primary rainforest /ca
<quantity id="5A72B0C9F829296ABFD98471FD5CF941" box="[679,724,1652,1679]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="596" unit="m" value="2.0">
<elevation id="16A7FA1FF829296ABFD98471FD5CF941" box="[679,724,1652,1679]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="596" unit="m" value="2.0">2 m</elevation>
</quantity>
wide sandy+mud bottom / collecting in stream and tributaries / on vegetation and dead wood [p] //
<typeStatus id="4231A38EF829296ABF38849DFD64F97C" box="[582,748,1688,1714]" pageId="6" pageNumber="596" type="holotype">HOLOTYPUS</typeStatus>
<specimenCount id="8B8CD6A5F829296ABF88849CFC83F97D" box="[758,779,1689,1715]" count="1" pageId="6" pageNumber="596" type="male"></specimenCount>
/
<taxonomicName id="5A8A66AFF829296ABE5B849DFB8AF97C" authorityName="Kirejtshuk" authorityYear="1985" box="[805,1026,1688,1714]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF829296ABE5B849DFB8AF97C" box="[805,1026,1688,1714]" italics="true" pageId="6" pageNumber="596">VIETTHERCHNUS</emphasis>
</taxonomicName>
/
<taxonomicName id="5A8A66AFF829296AB962849DFBEDF97C" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[1052,1125,1688,1714]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="596" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF829296AB962849DFBEDF97C" box="[1052,1125,1688,1714]" italics="true" pageId="6" pageNumber="596">helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F829296AB911849CFB40F97D" box="[1135,1224,1689,1715]" pageId="6" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
/ JelÍnek &amp;
<determinerName id="68E9F1E8F829296AB827849DFA14F97C" box="[1369,1436,1688,1714]" pageId="6" pageNumber="596">Hájek</determinerName>
det. 2024 [red label, p]”
</materialsCitation>
.
<materialsCitation id="2DE21771F829296ABCC784B8FBD5F918" box="[441,1117,1724,1751]" collectingDate="2011-11-30" collectorName="J. Hajek &amp; J. Schneider &amp; P. Votruba" country="Indonesia" determinerName="Hajek" elevation="66" latitude="-1.2233334" location="KALIMANTAN" longLatPrecision="130" longitude="116.37666" pageId="6" pageNumber="596" specimenCount="39" specimenCount-female="19" specimenCount-male="20" stateProvince="Kalimantan Utara" typeStatus="paratype">
<typeStatus id="4231A38EF829296ABCC784B8FDA2F919" box="[441,554,1725,1751]" pageId="6" pageNumber="596" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="8B8CD6A5F829296ABF4884B9FDFAF919" box="[566,626,1724,1751]" count="20" pageId="6" pageNumber="596" type="male">20 ♂</specimenCount>
,
<specimenCount id="8B8CD6A5F829296ABF0184B9FD31F919" box="[639,697,1724,1751]" count="19" pageId="6" pageNumber="596" type="female">19 ♀</specimenCount>
, the same data as holotype (NMPC)
</materialsCitation>
;
<materialsCitation id="2DE21771F829296AB91484B9FBE4F934" collectingDate="1997-12-03" collectorName="Ivo Jenis" country="Indonesia" county="Banjaran Bintang" location="Taiping" municipality="Bukit Berapit" pageId="6" pageNumber="596" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Utara" typeStatus="paratype">
<specimenCount id="8B8CD6A5F829296AB91484B9FB1FF919" box="[1130,1175,1724,1751]" count="1" pageId="6" pageNumber="596" type="male">1 ♂</specimenCount>
, “MALAYSIA-Perak /
<collectingCounty id="745465A0F829296ABDE984E5FED4F934" box="[151,348,1760,1786]" pageId="6" pageNumber="596">Banjaran Bintang</collectingCounty>
/
<collectingMunicipality id="7D518756F829296ABC0E84E5FD8EF934" box="[368,518,1760,1786]" pageId="6" pageNumber="596">Bukit Berapit</collectingMunicipality>
(
<location id="98554BF7F829296ABF6B84E5FDE6F934" LSID="urn:lsid:plazi:treatment:1523AC3AF8292965BDE987BAFC9EF972:98554BF7F829296ABF6B84E5FDE6F934" box="[533,622,1760,1786]" country="Indonesia" county="Banjaran Bintang" municipality="Bukit Berapit" name="Taiping" pageId="6" pageNumber="596" stateProvince="Kalimantan Utara">Taiping</location>
) / 11.-
<date id="E9343BECF829296ABFD384E5FCADF934" box="[685,805,1760,1786]" pageId="6" pageNumber="596" value="1997-12-03">
<collectingDate id="F970C204F829296ABFD384E5FCADF934" box="[685,805,1760,1786]" pageId="6" pageNumber="596" value="1997-12-03">12.3.1997</collectingDate>
</date>
/
<collectorName id="307F78FAF829296ABE4784E4FC15F934" box="[825,925,1760,1787]" pageId="6" pageNumber="596">Ivo Jeniš</collectorName>
leg. [p]” (NMPC)
</materialsCitation>
;
<materialsCitation id="2DE21771F829296AB90984E5FA89F8D0" country="Indonesia" location="All" pageId="6" pageNumber="596" specimenCount="1" specimenCount-male="1" stateProvince="PRAGENSE" typeStatus="paratype">
<specimenCount id="8B8CD6A5F829296AB90984E5FB2AF935" box="[1143,1186,1760,1787]" count="1" pageId="6" pageNumber="596" type="male">1 ♂</specimenCount>
, “MUS.
<collectingRegion id="5F4ED3CEF829296AB98384E5FA06F934" box="[1277,1422,1760,1786]" country="Indonesia" name="PRAGENSE" pageId="6" pageNumber="596">PRAGENSE</collectingRegion>
/
<collectingRegion id="5F4ED3CEF829296ABDE98501FECFF8D0" box="[151,327,1796,1822]" country="Indonesia" name="TENASSERIM" pageId="6" pageNumber="596">TENASSERIM</collectingRegion>
/ COLL.HELFER [p]” (NMPC).
<location id="98554BF7F829296ABFBE8501FD6CF8D0" LSID="urn:lsid:plazi:treatment:1523AC3AF8292965BDE987BAFC9EF972:98554BF7F829296ABFBE8501FD6CF8D0" box="[704,740,1796,1822]" country="Indonesia" name="All" pageId="6" pageNumber="596" stateProvince="PRAGENSE">All</location>
<typeStatus id="4231A38EF829296ABF958500FCDEF8D1" box="[747,854,1797,1823]" pageId="6" pageNumber="596" type="paratype">paratypes</typeStatus>
with the appropriate printed red label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F829296BBDB9852DFD83F865" lastPageId="7" lastPageNumber="597" pageId="6" pageNumber="596" type="description">
<paragraph id="9D351D2CF829296ABDB9852DFB6EF88C" blockId="6.[151,1437,1580,2039]" box="[199,1254,1831,1859]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB9852DFD07F88C" bold="true" box="[199,655,1831,1858]" pageId="6" pageNumber="596">
Description of male
<typeStatus id="4231A38EF829296ABCC9852DFDA8F88C" box="[439,544,1832,1858]" pageId="6" pageNumber="596" type="holotype">holotype</typeStatus>
.
<emphasis id="AFFEC13EF829296ABF528522FD07F88C" bold="true" box="[556,655,1831,1858]" italics="true" pageId="6" pageNumber="596">Habitus.</emphasis>
</emphasis>
Broadly oval, convex, glabrous, smooth and shining.
</paragraph>
<paragraph id="9D351D2CF829296ABDB9854EFB06F860" blockId="6.[151,1437,1580,2039]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB9854EFED2F8A8" bold="true" box="[199,346,1867,1894]" italics="true" pageId="6" pageNumber="596">Colouration.</emphasis>
Black, sides and base of pronotum with narrow reddish border, disc of pronotum with pair of vaguely delimited reddish spots. Disc of elytra orange, humeral bulges, sides and apex of elytra black. Ventral surface red-brown. Clypeus, legs and antennae brown-yellow, antennal club black (
<figureCitation id="05B101A9F829296AB9428591FB09F860" box="[1084,1153,1940,1966]" captionStart="FIGURES 14" captionStartId="3.[152,270,1922,1946]" captionTargetBox="[168,1421,213,1867]" captionTargetId="figure-32@3.[151,1436,181,1899]" captionTargetPageId="3" captionText="FIGURES 14. Dorsal habitus of Viettherchnus. 1. V. orszuliki sp. nov. (holotype ♂); 2. V. apriculus sp. nov. (holotype ♂); 3. V. helferi sp. nov. (paratype ♀, dark form; Kalimantan); 4. V. helferi sp. nov. (holotype ♂). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916277" httpUri="https://zenodo.org/record/13916277/files/figure.png" pageId="6" pageNumber="596">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF829296BBDB985B2FB00FE00" blockId="6.[151,1437,1580,2039]" lastBlockId="7.[151,1437,151,1964]" lastPageId="7" lastPageNumber="597" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB985B2FE8DF81C" bold="true" box="[199,261,1975,2002]" italics="true" pageId="6" pageNumber="596">Head</emphasis>
across eyes narrower than anterior margin of pronotum. Frons almost flat, between antennal insertions with sharp transverse edge projecting over basal portions of clypeus, arcuate in middle and with blunt prominent teeth laterally besides antennal insertions. Punctures of frons fairly equal in size to eye-facets, separated by 1.0 1.5 diameters, interspaces indistinctly alutaceous. Clypeus flat, situated markedly beneath level of frons, truncate anteriorly, alutaceous with a few dispersed shallow punctures. Labrum subtruncate anteriorly with small shallow incision.Mandibles abruptly curved, outer margins arcuate, sharp, raised over shallowly bifid apex, on right mandible ending in short prominent tip (
<figureCitation id="05B101A9F828296BBC958320FDA6FE8E" box="[491,558,293,320]" captionStart="FIGURES 57" captionStartId="4.[152,270,932,956]" captionTargetBox="[176,1411,207,909]" captionTargetId="figure-19@4.[176,1411,181,909]" captionTargetPageId="4" captionText="FIGURES 57. Head of male Viettherchnus. 5. V. orszuliki sp. nov.; 6. V. apriculus sp. nov.; 7. V. helferi sp. nov. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.13916279" httpUri="https://zenodo.org/record/13916279/files/figure.png" pageId="7" pageNumber="597">Fig. 7</figureCitation>
). Antennae slightly longer than width of head across eyes (ratio HEAW/ANLE = 0.96), antennal club occupying ca. one third of antenna length, distinctly shorter than antennomeres IIVIII combined, 1.64× longer than wide, widest at distal end of antennomere IX. Scapus ca. 1.7× longer than wide, almost twice as wide as pedicel. Antennomere III twice as long as wide, IV and V subequal, somewhat longer than wide, VI as long as wide, VII and VIII transverse, VIII wider than preceding ones, adjacent to club.
</paragraph>
<paragraph id="9D351D2CF828296BBDB983DCFB9EFD6B" blockId="7.[151,1437,151,1964]" pageId="7" pageNumber="597">
<emphasis id="AFFEC13EF828296BBDB983DCFEB4FE3D" bold="true" box="[199,316,473,499]" italics="true" pageId="7" pageNumber="597">Pronotum</emphasis>
convex, widest at posterior angles, arcuately narrowed anteriad, ratio WPR1/WPR3 = 1.94, WPR1/ LEPR = 1.83. Anterior margin trapezoidal, not bordered, anterior angles obtusely angulate, blunt. Lateral margins broadly arcuate, finely bordered, not explanate. Basal margin not bordered, in middle projecting into broad short truncate lobe concealing basal portion of scutellar shield, at sides broadly arcuate. Posterior angles obtusely rounded without well-defined tips. Surface of pronotum vaulted both longitudinally and transversely, punctate like frons. Scutellar shield large, subtriangular with rounded apex, punctate like pronotum.
</paragraph>
<paragraph id="9D351D2CF828296BBDB980ABFE6FFCB5" blockId="7.[151,1437,151,1964]" pageId="7" pageNumber="597">
<emphasis id="AFFEC13EF828296BBDB980ABFE87FD07" bold="true" box="[199,271,686,713]" italics="true" pageId="7" pageNumber="597">Elytra</emphasis>
almost as long as their combined width (ratio LELY/WELY = 0.98), widest somewhat behind humeral angles and gradually arcuately narrowed posteriad, simultaneously rounded apically. Lateral margins finely bordered, not explanate, not visible simultaneously in their entirety (dorsal view). Surface strongly convex with distinct traces of rather irregular series of punctures; punctures almost equal in size to those of pronotum, within one series separated by more than one diameter. Interstries with smaller and widely spaced diffuse punctures. Interspaces feebly alutaceous.
</paragraph>
<paragraph id="9D351D2CF828296BBDB98180FE02FACA" blockId="7.[151,1437,151,1964]" pageId="7" pageNumber="597">
<emphasis id="AFFEC13EF828296BBDB98180FED3FC51" bold="true" box="[199,347,901,927]" pageId="7" pageNumber="597">Ventral side.</emphasis>
Antennal furrows converging posteriad. Prosternum very short with sharp mediolongtudinal carina vanishing at distal end of prosternal process, coarsely and densely punctate. Apical margin of prosternal process blunt, subtruncate. Hypomera concave, indistinctly sparsely punctate, moderately shining, with raised sharp ridge nearly parallel to basal margin of pronotum. Mesoventrite roof-shaped, impunctate with fine mediolongitudinal carina. Mesocoxal axillary lines arcuate, running close to posterior margin of mesocoxal cavity. Metaventrite broadly and flatly transversely vaulted; its anterior intercoxal process broad, bordered, anterior margin shallowly concave, overlapping base of mesoventrite with anterior lateral angles somewhat prominent; posterior intercoxal margin truncate. Surface with punctures equal in size to eye-facets, separated by 1.01.5 diameters, interspaces obsoletely alutaceous. Discrimen absent. Metacoxal axillary lines arcuate, running close to posterior margin of metacoxal cavity. Abdominal ventrites punctate like metaventrite, punctures closer, mostly separated by less than one diameter (
<figureCitation id="05B101A9F828296BBC4686EFFEF4FACA" box="[312,380,1258,1284]" captionStart="FIGURES 89" captionStartId="5.[152,270,1037,1061]" captionTargetBox="[173,1417,205,998]" captionTargetId="figure-32@5.[151,1436,181,1013]" captionTargetPageId="5" captionText="FIGURES 89. Ventral habitus of Viettherchnus. 8. V. orszuliki sp. nov. (paratype ♂); 9. V. helferi sp. nov. (paratype ♀; Kalimantan). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916281" httpUri="https://zenodo.org/record/13916281/files/figure.png" pageId="7" pageNumber="597">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF828296BBDB9870BFC81FA30" blockId="7.[151,1437,151,1964]" pageId="7" pageNumber="597">
<emphasis id="AFFEC13EF828296BBDB9870BFE8DFAE6" bold="true" box="[199,261,1294,1320]" italics="true" pageId="7" pageNumber="597">Legs.</emphasis>
Profemur 1.5× longer than wide, its anterior margin rectilinear, proximal anterior angle rectangular; ventral surface with fine longitudinal carina in posterior half. Protibia ca. 2.5× longer than wide, subparallel-sided; outer subapical angle rectangular, outer margin densely denticulate, dorsal surface with dense and coarse punctures bearing semierect bristles, ventral surface with irregular longitudinal series of minute spines. Protarsomeres IIII 0.75× the width of protibia, V as long as IIV combined. Meso- and metafemur broadly oval, 2.00× and 1.85× longer than wide respectively. Meso- and metatibia almost 3× longer than wide, in basal third abruptly dilated distally, then almost parallel-sided. Tarsal claws simple.
</paragraph>
<paragraph id="9D351D2CF828296BBDB9840DFD36F989" blockId="7.[151,1437,151,1964]" pageId="7" pageNumber="597">
<emphasis id="AFFEC13EF828296BBDB9840DFEFEF9ED" bold="true" box="[199,374,1544,1571]" italics="true" pageId="7" pageNumber="597">Male genitalia.</emphasis>
Penis broadly oval, in distal third gradually narrowed towards bluntly pointes apex (
<figureCitation id="05B101A9F828296BB844840DFA07F9EC" box="[1338,1423,1544,1570]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="7" pageNumber="597">Fig. 12</figureCitation>
). Tegmen oval, broadly rounded apically (
<figureCitation id="05B101A9F828296BBF238429FD39F989" box="[605,689,1580,1607]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="7" pageNumber="597">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF828296BBDB98455FF4BF937" blockId="7.[151,1437,151,1964]" pageId="7" pageNumber="597">
<emphasis id="AFFEC13EF828296BBDB98455FEADF9A4" bold="true" box="[199,293,1616,1642]" pageId="7" pageNumber="597">Female.</emphasis>
Habitus as in male, from which it differs in frons and clypeus in the same plane, separated by indistinct transverse depression; frontoclypeal suture absent; punctures on clypeus markedly finer than those of frons. Mandibles and scapus simple, not expanded. Protarsomeres IIII narrower, 0.6× the width of protibia. Exposed posterior margin of tergite VIII absent. Ovipositor serrate in apical third, apex acutely pointed, without styli (
<figureCitation id="05B101A9F828296BB81184BEFF3EF937" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="7" pageNumber="597">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF828296BBDB98507FD2CF8AA" blockId="7.[151,1437,151,1964]" pageId="7" pageNumber="597">
<emphasis id="AFFEC13EF828296BBDB98507FEB7F8D2" bold="true" box="[199,319,1794,1820]" pageId="7" pageNumber="597">Variation.</emphasis>
Colour of pronotum and disc of elytra sometimes dark red to nearly black, extent of red colouration sometimes more or less reduced (
<figureCitation id="05B101A9F828296BBF778523FDC3F88E" box="[521,587,1830,1856]" captionStart="FIGURES 14" captionStartId="3.[152,270,1922,1946]" captionTargetBox="[168,1421,213,1867]" captionTargetId="figure-32@3.[151,1436,181,1899]" captionTargetPageId="3" captionText="FIGURES 14. Dorsal habitus of Viettherchnus. 1. V. orszuliki sp. nov. (holotype ♂); 2. V. apriculus sp. nov. (holotype ♂); 3. V. helferi sp. nov. (paratype ♀, dark form; Kalimantan); 4. V. helferi sp. nov. (holotype ♂). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916277" httpUri="https://zenodo.org/record/13916277/files/figure.png" pageId="7" pageNumber="597">Fig. 3</figureCitation>
); specimens with darker colouration prevail in the
<typeStatus id="4231A38EF828296BB9018522FB27F88F" box="[1151,1199,1831,1857]" pageId="7" pageNumber="597">type</typeStatus>
series. Reddish sides of pronotum sometimes more extended mesad.
</paragraph>
<paragraph id="9D351D2CF828296BBDB9856BFD83F865" blockId="7.[151,1437,151,1964]" pageId="7" pageNumber="597">
<emphasis id="AFFEC13EF828296BBDB9856BFEF6F846" bold="true" box="[199,382,1902,1928]" pageId="7" pageNumber="597">Measurements.</emphasis>
Body length 2.43.0 mm (
<typeStatus id="4231A38EF828296BBFD28568FC98F849" box="[684,784,1901,1927]" pageId="7" pageNumber="597" type="holotype">holotype</typeStatus>
<quantity id="5A72B0C9F828296BBE698568FCF7F846" box="[791,895,1901,1928]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" pageId="7" pageNumber="597" unit="mm" value="2.95">2.95 mm</quantity>
), width 1.72.0 mm (
<typeStatus id="4231A38EF828296BB9118568FB5BF849" box="[1135,1235,1901,1927]" pageId="7" pageNumber="597" type="holotype">holotype</typeStatus>
<quantity id="5A72B0C9F828296BB9A58568FACBF846" box="[1243,1347,1901,1928]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="7" pageNumber="597" unit="mm" value="1.95">1.95 mm</quantity>
), height
<quantity id="5A72B0C9F828296BBDE98594FE96F862" box="[151,286,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="7" pageNumber="597" unit="mm" value="1.3" valueMax="1.5" valueMin="1.1">1.11.5 mm</quantity>
(
<typeStatus id="4231A38EF828296BBC528594FE18F865" box="[300,400,1937,1963]" pageId="7" pageNumber="597" type="holotype">holotype</typeStatus>
<quantity id="5A72B0C9F828296BBCE98594FE77F865" box="[407,511,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="597" unit="mm" value="1.5">1.50 mm</quantity>
).
</paragraph>
</subSubSection>
<caption id="C9F54DA4F8272964BDE68568FB62F867" ID-DOI="http://doi.org/10.5281/zenodo.13916285" ID-Zenodo-Dep="13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="8" pageNumber="598" startId="8.[152,270,1901,1925]" targetBox="[185,1403,212,1865]" targetPageId="8" targetType="figure">
<paragraph id="9D351D2CF8272964BDE68568FB62F867" blockId="8.[151,1437,1901,1961]" pageId="8" pageNumber="598">
<emphasis id="AFFEC13EF8272964BDE68568FED1F84B" bold="true" box="[152,345,1901,1925]" pageId="8" pageNumber="598">FIGURES 1017.</emphasis>
Genitalia of
<taxonomicName id="5A8A66AFF8272964BCA48568FDECF84B" authorityName="Kirejtshuk" authorityYear="1985" box="[474,612,1901,1925]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF8272964BCA48568FDECF84B" box="[474,612,1901,1925]" italics="true" pageId="8" pageNumber="598">Viettherchnus</emphasis>
</taxonomicName>
. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16.
<taxonomicName id="5A8A66AFF8272964BC778597FEF1F867" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[265,377,1937,1961]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="598" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF8272964BC778597FEF1F867" box="[265,377,1937,1961]" italics="true" pageId="8" pageNumber="598">V. orszuliki</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF8272964BCFE8594FE58F867" bold="true" box="[384,464,1937,1961]" pageId="8" pageNumber="598">
<taxonomicNameLabel id="B4CD7C45F8272964BCFE8594FE58F867" box="[384,464,1937,1961]" pageId="8" pageNumber="598" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 11, 14.
<taxonomicName id="5A8A66AFF8272964BF578597FD2AF867" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[553,674,1937,1961]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="598" phylum="Arthropoda" rank="species" species="apriculus" status="sp. nov.">
<emphasis id="AFFEC13EF8272964BF578597FD2AF867" box="[553,674,1937,1961]" italics="true" pageId="8" pageNumber="598">V. apriculus</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF8272964BFD68594FD70F867" bold="true" box="[680,760,1937,1961]" pageId="8" pageNumber="598">
<taxonomicNameLabel id="B4CD7C45F8272964BFD68594FD70F867" box="[680,760,1937,1961]" pageId="8" pageNumber="598" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 12, 15, 17.
<taxonomicName id="5A8A66AFF8272964BE068597FC5BF867" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[888,979,1937,1961]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="598" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF8272964BE068597FC5BF867" box="[888,979,1937,1961]" italics="true" pageId="8" pageNumber="598">V. helferi</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF8272964BEA48594FBA2F867" bold="true" box="[986,1066,1937,1961]" pageId="8" pageNumber="598">
<taxonomicNameLabel id="B4CD7C45F8272964BEA48594FBA2F867" box="[986,1066,1937,1961]" pageId="8" pageNumber="598" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Scale bar 0.5 mm.
</paragraph>
</caption>
<caption id="C9F54DA4F8262965BDE68621FE8AFBAE" ID-DOI="http://doi.org/10.5281/zenodo.13916287" ID-Zenodo-Dep="13916287" httpUri="https://zenodo.org/record/13916287/files/figure.png" pageId="9" pageNumber="599" startId="9.[152,255,1060,1084]" targetBox="[151,1436,181,1036]" targetPageId="9" targetType="figure">
<paragraph id="9D351D2CF8262965BDE68621FE8AFBAE" blockId="9.[151,1436,1060,1120]" pageId="9" pageNumber="599">
<emphasis id="AFFEC13EF8262965BDE68621FEA0FBF2" bold="true" box="[152,296,1060,1084]" pageId="9" pageNumber="599">FIGURE 18.</emphasis>
Habitat of
<taxonomicName id="5A8A66AFF8262965BCDC8621FDFFFBF2" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[418,631,1060,1084]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="599" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF8262965BCDC8621FDFFFBF2" box="[418,631,1060,1084]" italics="true" pageId="9" pageNumber="599">Viettherchnus helferi</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF8262965BFFE8621FD5BFBF2" bold="true" box="[640,723,1060,1084]" pageId="9" pageNumber="599">
<taxonomicNameLabel id="B4CD7C45F8262965BFFE8621FD5BFBF2" box="[640,723,1060,1084]" pageId="9" pageNumber="599" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at the type locality (Indonesia, Eastern Kalimantan, PT Fajar Surya Swadaya).
</paragraph>
</caption>
<subSubSection id="D5904EA7F8262965BDB986AFFF47F9BA" pageId="9" pageNumber="599" type="diagnosis">
<paragraph id="9D351D2CF8262965BDB986AFFE23FAFE" blockId="9.[151,1437,1194,1724]" pageId="9" pageNumber="599">
<emphasis id="AFFEC13EF8262965BDB986AFFE59FB0A" bold="true" box="[199,465,1194,1220]" pageId="9" pageNumber="599">Differential diagnosis.</emphasis>
Antennal club shorter than antennomeres IIVIII combined resembles conditions in
<taxonomicName id="5A8A66AFF8262965BDE986CBFE17FB26" authorityName="Kirejtshuk &amp; Kirk-Spriggs" authorityYear="1996" box="[151,415,1230,1256]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="599" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="AFFEC13EF8262965BDE986CBFE17FB26" box="[151,415,1230,1256]" italics="true" pageId="9" pageNumber="599">Viettherchnus plagiatus</emphasis>
</taxonomicName>
and
<taxonomicName id="5A8A66AFF8262965BCA886CAFDDAFB26" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[470,594,1230,1256]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="599" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF8262965BCA886CAFDDAFB26" box="[470,594,1230,1256]" italics="true" pageId="9" pageNumber="599">V. orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F8262965BF2786CBFD38FB26" box="[601,688,1230,1256]" pageId="9" pageNumber="599" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="5A8A66AFF8262965BFC986CBFC17FB26" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[695,927,1230,1256]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="599" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF8262965BFC986CBFC17FB26" box="[695,927,1230,1256]" italics="true" pageId="9" pageNumber="599">Viettherchnus helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F8262965BED986CBFC75FB26" box="[935,1021,1230,1256]" pageId="9" pageNumber="599" rank="species">sp. nov.</taxonomicNameLabel>
differs from them in different colour pattern (elytra without several ligter spots), and in male sex in anterior margin of frons expanded anteriad to overlap basal portion of clypeus.
</paragraph>
<paragraph id="9D351D2CF8262965BDB9873FFE5AFA52" blockId="9.[151,1437,1194,1724]" pageId="9" pageNumber="599">
<emphasis id="AFFEC13EF8262965BDB9873FFE74FA9A" bold="true" box="[199,508,1338,1364]" pageId="9" pageNumber="599">Collection circumstances.</emphasis>
At the
<typeStatus id="4231A38EF8262965BF25873FFD03FA9A" box="[603,651,1338,1364]" pageId="9" pageNumber="599">type</typeStatus>
locality in
<collectingRegion id="5F4ED3CEF8262965BE68873FFC5FFA9A" box="[790,983,1338,1364]" country="Indonesia" name="Kalimantan Timur" pageId="9" pageNumber="599">East Kalimantan</collectingRegion>
,
<taxonomicName id="5A8A66AFF8262965BE98873EFBC7FA9A" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[998,1103,1338,1364]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="599" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF8262965BE98873EFBC7FA9A" box="[998,1103,1338,1364]" italics="true" pageId="9" pageNumber="599">V. helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F8262965B924873FFB3DFA9A" box="[1114,1205,1338,1364]" pageId="9" pageNumber="599" rank="species">sp. nov.</taxonomicNameLabel>
was collected from unidentified fungi growing on the bottom side of dead tree trunk fallen over small stream in remnant of lowland primary rainforest (
<figureCitation id="05B101A9F8262965BC0F8787FE4DFA52" box="[369,453,1410,1436]" captionStart="FIGURE 18" captionStartId="9.[152,255,1060,1084]" captionTargetBox="[151,1436,181,1036]" captionTargetId="figure-32@9.[151,1436,181,1036]" captionTargetPageId="9" captionText="FIGURE 18. Habitat of Viettherchnus helferi sp. nov. at the type locality (Indonesia, Eastern Kalimantan, PT Fajar Surya Swadaya)." figureDoi="http://doi.org/10.5281/zenodo.13916287" httpUri="https://zenodo.org/record/13916287/files/figure.png" pageId="9" pageNumber="599">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF8262965BDB987A3FF47F9BA" blockId="9.[151,1437,1194,1724]" pageId="9" pageNumber="599">
<emphasis id="AFFEC13EF8262965BDB987A3FE11FA0E" bold="true" box="[199,409,1446,1472]" pageId="9" pageNumber="599">Name derivation.</emphasis>
The new species is dedicated to the Bohemian physician, explorer and naturalist Johann Wilhelm Helfer also known as Jan Vilém Helfer (18101840). He was probably the first, who collected in 1837 1839 plants and animals in Tenasserim (
<collectingCountry id="E59D5DBCF8262965BF1A87EBFD5FF9C6" box="[612,727,1518,1544]" name="Myanmar" pageId="9" pageNumber="599">Myanmar</collectingCountry>
), among them also
<specimenCount id="8B8CD6A5F8262965BEC787EBFBC3F9C6" box="[953,1099,1518,1544]" count="1" pageId="9" pageNumber="599" type="generic" typeStatus="paratype">one paratype</specimenCount>
of the new species. After his sudden death on Andaman Islands (
<date id="E9343BECF8262965BF608417FD66F9E2" box="[542,750,1554,1580]" pageId="9" pageNumber="599" value="1840-01-30">
January 30
<superScript id="6AFFB064F8262965BFE78417FD2EF9EE" attach="none" box="[665,678,1554,1568]" fontSize="6" pageId="9" pageNumber="599">th</superScript>
, 1840
</date>
), his widow, Pauline Helfer donated her husbands collections to the Bohemian Museum, nowadays the National Museum, Prague. The specific epithet is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F8262965BDB9847BFC9EF972" pageId="9" pageNumber="599" type="distribution">
<paragraph id="9D351D2CF8262965BDB9847BFC9EF972" blockId="9.[151,1437,1194,1724]" pageId="9" pageNumber="599">
<emphasis id="AFFEC13EF8262965BDB9847BFEE9F956" bold="true" box="[199,353,1662,1688]" pageId="9" pageNumber="599">Distribution.</emphasis>
<taxonomicName id="5A8A66AFF8262965BC1B847BFDC3F956" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[357,587,1662,1688]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="599" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF8262965BC1B847BFDC3F956" box="[357,587,1662,1688]" italics="true" pageId="9" pageNumber="599">Viettherchnus helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F8262965BF2E847BFD2DF956" box="[592,677,1662,1688]" pageId="9" pageNumber="599" rank="species">sp. nov.</taxonomicNameLabel>
seems to be widespread species, currently it is known from
<collectingCountry id="E59D5DBCF8262965B851847BFA14F956" box="[1327,1436,1662,1688]" name="Indonesia" pageId="9" pageNumber="599">Indonesia</collectingCountry>
(
<collectingRegion id="5F4ED3CEF8262965BDE184A7FEA0F972" box="[159,296,1698,1724]" country="Indonesia" name="Kalimantan Utara" pageId="9" pageNumber="599">Kalimantan</collectingRegion>
),
<collectingCountry id="E59D5DBCF8262965BC4484A7FE29F972" box="[314,417,1698,1724]" name="Malaysia" pageId="9" pageNumber="599">Malaysia</collectingCountry>
(
<collectingRegion id="5F4ED3CEF8262965BCCE84A7FE7EF972" box="[432,502,1698,1724]" country="Malaysia" name="Perak" pageId="9" pageNumber="599">Perak</collectingRegion>
),
<collectingCountry id="E59D5DBCF8262965BF7784A7FDFEF972" box="[521,630,1698,1724]" name="Myanmar" pageId="9" pageNumber="599">Myanmar</collectingCountry>
(Tenasserim).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,230 @@
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<mods:title id="F91035DA864AAA44E0FECD6606921A47">Three new species of the genus Viettherchnus Kirejtshuk, 1985 (Coleoptera: Nitidulidae: Nitidulinae: Cyllodini)</mods:title>
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<paragraph id="9D351D2CF82A2969BDE9869EFD97FB7B" blockId="5.[151,543,1179,1241]" box="[151,543,1179,1206]" pageId="5" pageNumber="595">
<heading id="C67DAA40F82A2969BDE9869EFD97FB7B" bold="true" box="[151,543,1179,1206]" fontSize="11" level="1" pageId="5" pageNumber="595" reason="1">
<emphasis id="AFFEC13EF82A2969BDE9869EFD97FB7B" bold="true" box="[151,543,1179,1206]" pageId="5" pageNumber="595">
<taxonomicName id="5A8A66AFF82A2969BDE9869EFE32FB78" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[151,442,1179,1206]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="595" phylum="Arthropoda" rank="species" species="apriculus" status="sp. nov.">
<emphasis id="AFFEC13EF82A2969BDE9869EFE32FB78" bold="true" box="[151,442,1179,1206]" italics="true" pageId="5" pageNumber="595">Viettherchnus apriculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F82A2969BCBF869EFD97FB7B" box="[449,543,1179,1205]" pageId="5" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="D5904EA7F82A2969BDE986BAFEEDFB17" box="[151,357,1215,1241]" pageId="5" pageNumber="595" type="description">
<paragraph id="9D351D2CF82A2969BDE986BAFEEDFB17" blockId="5.[151,543,1179,1241]" box="[151,357,1215,1241]" pageId="5" pageNumber="595">
(
<figureCitation id="05B101A9F82A2969BDE186BAFF67FB17" box="[159,239,1215,1241]" captionStart="FIGURES 14" captionStartId="3.[152,270,1922,1946]" captionTargetBox="[168,1421,213,1867]" captionTargetId="figure-32@3.[151,1436,181,1899]" captionTargetPageId="3" captionText="FIGURES 14. Dorsal habitus of Viettherchnus. 1. V. orszuliki sp. nov. (holotype ♂); 2. V. apriculus sp. nov. (holotype ♂); 3. V. helferi sp. nov. (paratype ♀, dark form; Kalimantan); 4. V. helferi sp. nov. (holotype ♂). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916277" httpUri="https://zenodo.org/record/13916277/files/figure.png" pageId="5" pageNumber="595">Figs. 2</figureCitation>
,
<figureCitation id="05B101A9F82A2969BD8586BAFE83FB17" box="[251,267,1215,1241]" captionStart="FIGURES 57" captionStartId="4.[152,270,932,956]" captionTargetBox="[176,1411,207,909]" captionTargetId="figure-19@4.[176,1411,181,909]" captionTargetPageId="4" captionText="FIGURES 57. Head of male Viettherchnus. 5. V. orszuliki sp. nov.; 6. V. apriculus sp. nov.; 7. V. helferi sp. nov. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.13916279" httpUri="https://zenodo.org/record/13916279/files/figure.png" pageId="5" pageNumber="595">6</figureCitation>
,
<figureCitation id="05B101A9F82A2969BC6986BAFED5FB17" box="[279,349,1215,1241]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="5" pageNumber="595">11, 14</figureCitation>
)
</paragraph>
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<subSubSection id="D5904EA7F82A2969BDE98702FCDAFAA7" pageId="5" pageNumber="595" type="materials_examined">
<paragraph id="9D351D2CF82A2969BDE98702FCDAFAA7" blockId="5.[151,1437,1287,2034]" pageId="5" pageNumber="595">
<emphasis id="AFFEC13EF82A2969BDE98702FECEFAEF" bold="true" box="[151,326,1287,1313]" pageId="5" pageNumber="595">Type material.</emphasis>
<materialsCitation id="2DE21771F82A2969BC318702FCDAFAA7" collectingDate="2009-05-15" collectorName="Hajek" country="Malaysia" determinerName="Hajek" elevation="590" latitude="4.37" location="Cameron Highlands" longLatPrecision="130" longitude="101.333336" pageId="5" pageNumber="595" specimenCount="2" specimenCount-male="2" stateProvince="Perak" typeStatus="holotype">
<typeStatus id="4231A38EF82A2969BC318702FE3FFAEF" box="[335,439,1287,1313]" pageId="5" pageNumber="595" type="holotype">Holotype</typeStatus>
<specimenCount id="8B8CD6A5F82A2969BCC1870DFE5CFAEC" box="[447,468,1288,1314]" count="1" pageId="5" pageNumber="595" type="male"></specimenCount>
(NMPC), labelled: “
<collectingCountry id="E59D5DBCF82A2969BFBF8702FCD0FAEF" box="[705,856,1287,1313]" name="Malaysia" pageId="5" pageNumber="595">MALAYSIA</collectingCountry>
,
<collectingRegion id="5F4ED3CEF82A2969BE1A8702FC2CFAEF" box="[868,932,1287,1313]" country="Malaysia" name="Perak" pageId="5" pageNumber="595">Perak</collectingRegion>
/
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/ BATU [= MILE] 19 vill. env. /
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,
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,
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<elevation id="16A7FA1FF82A2969BF66872EFDD6FA88" box="[536,606,1323,1350]" metricMagnitude="2" metricUnit="m" metricValue="5.9" pageId="5" pageNumber="595" unit="m" value="590.0">590 m</elevation>
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/ JiřÍ
<collectorName id="307F78FAF82A2969BFE3872EFD68FA8B" box="[669,736,1323,1349]" pageId="5" pageNumber="595">Hájek</collectorName>
leg. 5.-
<date id="E9343BECF82A2969BE50872EFC2DFA88" box="[814,933,1323,1350]" pageId="5" pageNumber="595" value="2009-05-15">
<collectingDate id="F970C204F82A2969BE50872EFC2DFA88" box="[814,933,1323,1350]" pageId="5" pageNumber="595" value="2009-05-15">15.5.2009</collectingDate>
</date>
[p] //
<typeStatus id="4231A38EF82A2969BE96872EFB06FA8B" box="[1000,1166,1323,1349]" pageId="5" pageNumber="595" type="holotype">HOLOTYPUS</typeStatus>
<specimenCount id="8B8CD6A5F82A2969B9EB8729FB22FA88" box="[1173,1194,1324,1350]" count="1" pageId="5" pageNumber="595" type="male"></specimenCount>
/
<taxonomicName id="5A8A66AFF82A2969B9C18729FA14FA88" authorityName="Kirejtshuk" authorityYear="1985" box="[1215,1436,1324,1350]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82A2969B9C18729FA14FA88" box="[1215,1436,1324,1350]" italics="true" pageId="5" pageNumber="595">VIETTHERCHNUS</emphasis>
</taxonomicName>
/
<taxonomicName id="5A8A66AFF82A2969BDD8874AFE98FAA7" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[166,272,1359,1385]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="595" phylum="Arthropoda" rank="species" species="apriculus" status="sp. nov.">
<emphasis id="AFFEC13EF82A2969BDD8874AFE98FAA7" box="[166,272,1359,1385]" italics="true" pageId="5" pageNumber="595">apriculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F82A2969BC698755FEE5FAA4" box="[279,365,1360,1386]" pageId="5" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
/ JelÍnek &amp;
<determinerName id="68E9F1E8F82A2969BC88874AFDB1FAA7" box="[502,569,1359,1385]" pageId="5" pageNumber="595">Hájek</determinerName>
det. 2024 [red label, p]”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F82A296ABDB98776FC7FFBDF" lastPageId="6" lastPageNumber="596" pageId="5" pageNumber="595" type="description">
<paragraph id="9D351D2CF82A2969BDB98776FB91FA43" blockId="5.[151,1437,1287,2034]" box="[199,1049,1395,1422]" pageId="5" pageNumber="595">
<emphasis id="AFFEC13EF82A2969BDB98776FE4AFA40" bold="true" box="[199,450,1395,1422]" pageId="5" pageNumber="595">
Description.
<emphasis id="AFFEC13EF82A2969BC1E8776FE4AFA40" bold="true" box="[352,450,1395,1422]" italics="true" pageId="5" pageNumber="595">Habitus.</emphasis>
</emphasis>
Broadly oval, convex, glabrous, smooth and shining.
</paragraph>
<paragraph id="9D351D2CF82A2969BDB98792FA96F9D3" blockId="5.[151,1437,1287,2034]" pageId="5" pageNumber="595">
<emphasis id="AFFEC13EF82A2969BDB98792FED2FA7C" bold="true" box="[199,346,1431,1458]" italics="true" pageId="5" pageNumber="595">Colouration.</emphasis>
Head black with minute orange spots at insertions of antennae, pronotum black with narrow orange borders, somewhat wider at basal margin and vaguely delimited reddish-brown mediolongitudinal area. Scutellar shield and elytra orange, wider border along lateral half of base and lateral margins of elytra, as well as apical portion of elytra brown-black. Legs and antennae brown-yellow, antennal club infuscate (
<figureCitation id="05B101A9F82A2969B9B28406FA99F9D3" box="[1228,1297,1539,1565]" captionStart="FIGURES 14" captionStartId="3.[152,270,1922,1946]" captionTargetBox="[168,1421,213,1867]" captionTargetId="figure-32@3.[151,1436,181,1899]" captionTargetPageId="3" captionText="FIGURES 14. Dorsal habitus of Viettherchnus. 1. V. orszuliki sp. nov. (holotype ♂); 2. V. apriculus sp. nov. (holotype ♂); 3. V. helferi sp. nov. (paratype ♀, dark form; Kalimantan); 4. V. helferi sp. nov. (holotype ♂). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916277" httpUri="https://zenodo.org/record/13916277/files/figure.png" pageId="5" pageNumber="595">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF82A2969BDB98422FB91F848" blockId="5.[151,1437,1287,2034]" pageId="5" pageNumber="595">
<emphasis id="AFFEC13EF82A2969BDB98422FE8DF98C" bold="true" box="[199,261,1575,1602]" italics="true" pageId="5" pageNumber="595">Head</emphasis>
transverse, temples rectilinear, converging posteriad. Frons almost flat, diffusely punctate, its anterior portion between antennal insertions sloping down, almost impunctate. Punctures almost equal in size to eye-facets, separated by 23 diameters, interspaces smooth, strongly shining. Clypeus separately vaulted, at base separated from antennal insertions by deep perpendicular furrows, anterior margin subtrapezoidal, surface with very small and widely separated punctures, interspaces smooth and shining. Labrum almost semicircular with narrow median incision. Mandibles with strong pointed protuberances curved outwards (
<figureCitation id="05B101A9F82A2969BEAF84DEFB9DF93B" box="[977,1045,1755,1781]" captionStart="FIGURES 57" captionStartId="4.[152,270,932,956]" captionTargetBox="[176,1411,207,909]" captionTargetId="figure-19@4.[176,1411,181,909]" captionTargetPageId="4" captionText="FIGURES 57. Head of male Viettherchnus. 5. V. orszuliki sp. nov.; 6. V. apriculus sp. nov.; 7. V. helferi sp. nov. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.13916279" httpUri="https://zenodo.org/record/13916279/files/figure.png" pageId="5" pageNumber="595">Fig. 6</figureCitation>
). Antennae somewhat shorter than width of head across eyes, antennal club oblong, obovate, occupying ca 0.4 antennal length, 1.5× longer than wide, widest at base of antennomere XI. Scape almost semicircular, its anterior (dorsal) side with two sharp parallel longitudinal edges, between them canaliculate. Pedicel twice, antennomere III 1.66× as long as wide, antennomeres IV, VI and VII nearly as long as wide, V slightly longer than neighbouring ones.
</paragraph>
<paragraph id="9D351D2CF82A296ABDB98595FD57FE8F" blockId="5.[151,1437,1287,2034]" lastBlockId="6.[151,1437,151,1401]" lastPageId="6" lastPageNumber="596" pageId="5" pageNumber="595">
<emphasis id="AFFEC13EF82A2969BDB98595FEB4F864" bold="true" box="[199,316,1936,1962]" italics="true" pageId="5" pageNumber="595">Pronotum</emphasis>
transverse, convex both transversely and (less so) longitudinally, widest at posterior angles and strongly curved anteriad. Ratio WPR1/WPR3 =1.91, WPR2/LEPR = 1.82. Anterior margin with trapezoidal emargination, arcuate, not bordered; anterior angles obtusely rounded. Lateral margins broadly arcuate, in posterior half subrectilinear (vertical view), finely bordered, not explanate. Basal margin not bordered, with broad truncate lobe opposing scutellar shield, besides it broadly arcuate. Posterior angles obtuse. Punctures equal in size to those of frons, separated by more than one diameter; interspaces smooth and shining, with sparsely intermixed microscopic punctures bearing short indistinct rudimentary setae. Scutellar shield subtriangular, partly concealed by basal lobe of pronotum, punctate more densely than pronotum.
</paragraph>
<paragraph id="9D351D2CF829296ABDB9834FFB62FE3B" blockId="6.[151,1437,151,1401]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB9834FFE87FEAB" bold="true" box="[199,271,330,357]" italics="true" pageId="6" pageNumber="596">Elytra</emphasis>
widest near base, arcuately narrowed posteriad and simultaneously rounded apically.Disc strongly convex both longitudinally and transversely, lateral margins finely bordered, not explanate, not visible simultaneously from above in their entirety. Surface of elytra seriate punctate, punctures in series nearly equal in size to those of pronotum, neighbouring punctures in one series separated by ca. one diameter, punctures on interstries diffuse, of somewhat variable size, but generally smaller and sparser than those in series. Sutural lines absent.
</paragraph>
<paragraph id="9D351D2CF829296ABDB983FBFB3EFD03" blockId="6.[151,1437,151,1401]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB983FBFE08FDD6" bold="true" box="[199,384,510,537]" italics="true" pageId="6" pageNumber="596">Ventral surface.</emphasis>
Prosternum mediolongitudinally carinate, apical margin flatly arcuate. Mesoventrite situated beneath metaventrite (ventral view), mediolongitudinally carinate. Metaventrite flatly convex, in posterior half with punctures nearly equal to eye-facets and separated by 1.01.5 diameters, becoming larger and closer laterally and finer in anterior half; interspaces smooth and shining. Diverticulum absent. Mesocoxal axillary lines reaching metasternopleural suture at ca. one third of its length. Abdominal ventrites with larger punctures separated by one diameter or less. Metacoxal axillary lines closely bordering posterior margin of coxal cavities.
</paragraph>
<paragraph id="9D351D2CF829296ABDB980D2FEB8FC4F" blockId="6.[151,1437,151,1401]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB980D2FE8DFD3F" bold="true" box="[199,261,727,753]" italics="true" pageId="6" pageNumber="596">Legs.</emphasis>
Profemur 1.5×longer than wide, its anterior margin rectilinear, proximal anterior angle rectangular. Protibia ca. 2.5× longer than wide, subparallel-sided; outer subapical angle rectangular, outer margin densely denticulate. Protarsomeres IIII 0.75× the width of protibia, V as long as IIV combined. Mesofemur subtrapezoidal, 1.74× longer than wide. Metafemur almost oval, posterior margin in basal half rectilinear, 1.93× longer than wide. Tarsal claws simple.
</paragraph>
<paragraph id="9D351D2CF829296ABDB9818FFE88FC07" blockId="6.[151,1437,151,1401]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB9818FFEFEFC6B" bold="true" box="[199,374,906,933]" italics="true" pageId="6" pageNumber="596">Male genitalia.</emphasis>
Penis subparallel-sided, broadly rounded apically (
<figureCitation id="05B101A9F829296ABEC4818EFB84FC6B" box="[954,1036,907,933]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="6" pageNumber="596">Fig. 11</figureCitation>
). Tegmen broadly rounded apically (
<figureCitation id="05B101A9F829296ABDE181AAFF7BFC07" box="[159,243,943,969]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="6" pageNumber="596">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF829296ABDB981D6FE1DFC23" blockId="6.[151,1437,151,1401]" box="[199,405,979,1005]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB981D6FE96FC23" bold="true" box="[199,286,979,1005]" pageId="6" pageNumber="596">Female</emphasis>
unknown.
</paragraph>
<paragraph id="9D351D2CF829296ABDB981F2FC7FFBDF" blockId="6.[151,1437,151,1401]" box="[199,1015,1014,1041]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB981F2FEF6FBDF" bold="true" box="[199,382,1015,1041]" pageId="6" pageNumber="596">Measurements.</emphasis>
Body length
<quantity id="5A72B0C9F829296ABF6B81F3FDF3FBDF" box="[533,635,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="6" pageNumber="596" unit="mm" value="2.45">2.45 mm</quantity>
, width
<quantity id="5A72B0C9F829296ABFB381F2FCBBFBDF" box="[717,819,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="596" unit="mm" value="1.7">1.70 mm</quantity>
, height
<quantity id="5A72B0C9F829296ABEF281F2FC7BFBDF" box="[908,1011,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="596" unit="mm" value="1.2">1.20 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F829296ABDB9861FFDD1FAFF" pageId="6" pageNumber="596" type="diagnosis">
<paragraph id="9D351D2CF829296ABDB9861FFBDEFB0B" blockId="6.[151,1437,151,1401]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB9861FFE44FBFA" bold="true" box="[199,460,1050,1076]" pageId="6" pageNumber="596">Differential diagnosis</emphasis>
. Large antennal club, shape of scapus and seriate punctation of elytra suggest its close relation to
<taxonomicName id="5A8A66AFF829296ABC2B863AFB85FB97" authority="Kirejtshuk &amp; Kirk-Spriggs, 1996" authorityName="Kirejtshuk &amp; Kirk-Spriggs" authorityYear="1996" box="[341,1037,1087,1113]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="596" phylum="Arthropoda" rank="species" species="sulawesiensis">
<emphasis id="AFFEC13EF829296ABC2B863AFD06FB97" box="[341,654,1087,1113]" italics="true" pageId="6" pageNumber="596">Viettherchnus sulawesiensis</emphasis>
<bibRefCitation id="F91B60DDF829296ABFE9863AFB85FB97" author="Kirejtshuk, A. G. &amp; Kirk-Spriggs, A. H." box="[663,1037,1087,1113]" pageId="6" pageNumber="596" pagination="131 - 138" refId="ref7251" refString="Kirejtshuk, A. G. &amp; Kirk-Spriggs, A. H. (1996) Viettherchnus Kirejtshuk, 1985 and Ceramphosia gen. n. from the Indo-Malayan Region (Coleoptera: Nitidulidae). Zoosystematica Rossica, 4, 131 - 138. [1995]" type="journal article" year="1996">Kirejtshuk &amp; Kirk-Spriggs, 1996</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="E59D5DBCF829296AB901863AFB64FB97" box="[1151,1260,1087,1113]" name="Indonesia" pageId="6" pageNumber="596">Indonesia</collectingCountry>
(
<collectingRegion id="5F4ED3CEF829296AB985863AFAEBFB97" box="[1275,1379,1087,1113]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="596">Sulawesi</collectingRegion>
) and
<collectingCountry id="E59D5DBCF829296ABDE98666FE91FBB3" box="[151,281,1123,1149]" name="Philippines" pageId="6" pageNumber="596">Philippines</collectingCountry>
. The new species differs from the latter in its brightly orange elytra with black apex and sides, and in striking acute lateral projections on mandibles in male. Large mandibular projections occur also in the sympatric
<taxonomicName id="5A8A66AFF829296AB8F68682FF77FB0B" authorityName="Kirejtshuk &amp; Kirk-Spriggs" authorityYear="1996" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="596" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="AFFEC13EF829296AB8F68682FF77FB0B" italics="true" pageId="6" pageNumber="596">V. plagiatus</emphasis>
</taxonomicName>
, in which they are directed anteriad, subparallel-sided and rounded apically.
</paragraph>
<paragraph id="9D351D2CF829296ABDB986CAFDD1FAFF" blockId="6.[151,1437,151,1401]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB986CAFE15FB26" bold="true" box="[199,413,1230,1257]" pageId="6" pageNumber="596">Name derivation.</emphasis>
Latin noun
<taxonomicName id="5A8A66AFF829296ABF4586CAFD2DFB27" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[571,677,1231,1257]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="596" phylum="Arthropoda" rank="species" species="apriculus">
<emphasis id="AFFEC13EF829296ABF4586CAFD2DFB27" box="[571,677,1231,1257]" italics="true" pageId="6" pageNumber="596">apriculus</emphasis>
</taxonomicName>
,
<emphasis id="AFFEC13EF829296ABFC786CAFD42FB26" box="[697,714,1231,1256]" italics="true" pageId="6" pageNumber="596">-i</emphasis>
, m, is diminutive of
<emphasis id="AFFEC13EF829296ABEA686CAFB83FB26" box="[984,1035,1231,1256]" italics="true" pageId="6" pageNumber="596">aper</emphasis>
(boar), referring to characteristic modification of male mandibles resembling the projecting teeth of a wild boar. The specific epithet is a noun in the nominative case, standing in apposition.
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F829296ABDB9873FFE58FAB7" pageId="6" pageNumber="596" type="distribution">
<paragraph id="9D351D2CF829296ABDB9873FFE58FAB7" blockId="6.[151,1437,151,1401]" pageId="6" pageNumber="596">
<emphasis id="AFFEC13EF829296ABDB9873FFEE9FA9A" bold="true" box="[199,353,1338,1364]" pageId="6" pageNumber="596">Distribution.</emphasis>
<taxonomicName id="5A8A66AFF829296ABC14873EFDFEFA9B" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[362,630,1339,1365]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="596" phylum="Arthropoda" rank="species" species="apriculus" status="sp. nov.">
<emphasis id="AFFEC13EF829296ABC14873EFDFEFA9B" box="[362,630,1339,1365]" italics="true" pageId="6" pageNumber="596">Viettherchnus apriculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F829296ABF01873EFD51FA9B" box="[639,729,1339,1365]" pageId="6" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
is only known from the
<typeStatus id="4231A38EF829296ABE85873EFBA3FA9B" box="[1019,1067,1339,1365]" pageId="6" pageNumber="596">type</typeStatus>
locality in Cameron Highlands,
<collectingRegion id="5F4ED3CEF829296ABDE9875AFF52FAB7" box="[151,218,1375,1401]" country="Malaysia" name="Perak" pageId="6" pageNumber="596">Perak</collectingRegion>
, Peninsular
<collectingCountry id="E59D5DBCF829296ABC1C875AFE44FAB7" box="[354,460,1375,1401]" name="Malaysia" pageId="6" pageNumber="596">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="553AB0B04DE1D015F2EDF92D8DCB9543">Three new species of the genus Viettherchnus Kirejtshuk, 1985 (Coleoptera: Nitidulidae: Nitidulinae: Cyllodini)</mods:title>
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<treatment id="1523AC3AF82E2968BDE986E7FAF0F83B" LSID="urn:lsid:plazi:treatment:1523AC3AF82E2968BDE986E7FAF0F83B" httpUri="http://treatment.plazi.org/id/1523AC3AF82E2968BDE986E7FAF0F83B" lastPageId="4" lastPageNumber="594" pageId="1" pageNumber="591">
<subSubSection id="D5904EA7F82E296DBDE986E7FD9DFB33" box="[151,533,1250,1277]" pageId="1" pageNumber="591" type="nomenclature">
<paragraph id="9D351D2CF82E296DBDE986E7FD9DFB33" blockId="1.[151,533,1250,1313]" box="[151,533,1250,1277]" pageId="1" pageNumber="591">
<heading id="C67DAA40F82E296DBDE986E7FD9DFB33" bold="true" box="[151,533,1250,1277]" fontSize="11" level="1" pageId="1" pageNumber="591" reason="1">
<emphasis id="AFFEC13EF82E296DBDE986E7FD9DFB33" bold="true" box="[151,533,1250,1277]" pageId="1" pageNumber="591">
<taxonomicName id="5A8A66AFF82E296DBDE986E7FE38FB33" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[151,432,1250,1277]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF82E296DBDE986E7FE38FB33" bold="true" box="[151,432,1250,1277]" italics="true" pageId="1" pageNumber="591">Viettherchnus orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F82E296DBCC686E6FD9DFB33" box="[440,533,1251,1277]" pageId="1" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F82E296DBDE98702FE24FAEF" box="[151,428,1286,1313]" pageId="1" pageNumber="591" type="description">
<paragraph id="9D351D2CF82E296DBDE98702FE24FAEF" blockId="1.[151,533,1250,1313]" box="[151,428,1286,1313]" pageId="1" pageNumber="591">
(
<figureCitation id="05B101A9F82E296DBDE18702FF65FAEF" box="[159,237,1287,1313]" captionStart="FIGURES 14" captionStartId="3.[152,270,1922,1946]" captionTargetBox="[168,1421,213,1867]" captionTargetId="figure-32@3.[151,1436,181,1899]" captionTargetPageId="3" captionText="FIGURES 14. Dorsal habitus of Viettherchnus. 1. V. orszuliki sp. nov. (holotype ♂); 2. V. apriculus sp. nov. (holotype ♂); 3. V. helferi sp. nov. (paratype ♀, dark form; Kalimantan); 4. V. helferi sp. nov. (holotype ♂). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916277" httpUri="https://zenodo.org/record/13916277/files/figure.png" pageId="1" pageNumber="591">Figs. 1</figureCitation>
,
<figureCitation id="05B101A9F82E296DBD858703FE83FAEF" box="[251,267,1286,1313]" captionStart="FIGURES 57" captionStartId="4.[152,270,932,956]" captionTargetBox="[176,1411,207,909]" captionTargetId="figure-19@4.[176,1411,181,909]" captionTargetPageId="4" captionText="FIGURES 57. Head of male Viettherchnus. 5. V. orszuliki sp. nov.; 6. V. apriculus sp. nov.; 7. V. helferi sp. nov. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.13916279" httpUri="https://zenodo.org/record/13916279/files/figure.png" pageId="1" pageNumber="591">5</figureCitation>
,
<figureCitation id="05B101A9F82E296DBC698702FEAEFAEF" box="[279,294,1287,1313]" captionStart="FIGURES 89" captionStartId="5.[152,270,1037,1061]" captionTargetBox="[173,1417,205,998]" captionTargetId="figure-32@5.[151,1436,181,1013]" captionTargetPageId="5" captionText="FIGURES 89. Ventral habitus of Viettherchnus. 8. V. orszuliki sp. nov. (paratype ♂); 9. V. helferi sp. nov. (paratype ♀; Kalimantan). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916281" httpUri="https://zenodo.org/record/13916281/files/figure.png" pageId="1" pageNumber="591">8</figureCitation>
,
<figureCitation id="05B101A9F82E296DBC4D8702FE2CFAEF" box="[307,420,1287,1313]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="1" pageNumber="591">10, 13, 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F82E296DBDE9874AFB92F9D3" pageId="1" pageNumber="591" type="materials_examined">
<paragraph id="9D351D2CF82E296DBDE9874AFB92F9D3" blockId="1.[151,1437,1358,2033]" pageId="1" pageNumber="591">
<emphasis id="AFFEC13EF82E296DBDE9874AFECCFAA6" bold="true" box="[151,324,1358,1385]" pageId="1" pageNumber="591">Type material.</emphasis>
<materialsCitation id="2DE21771F82E296DBC37874AFA96FA43" collectingDate="2015-04-23" country="Nepal" determinerName="Hajek" elevation="800" location="Manaslu" municipality="Besisahar" pageId="1" pageNumber="591" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="4231A38EF82E296DBC37874AFE39FAA7" box="[329,433,1359,1385]" pageId="1" pageNumber="591" type="holotype">Holotype</typeStatus>
<specimenCount id="8B8CD6A5F82E296DBCC9874AFE44FAA7" box="[439,460,1359,1385]" count="1" pageId="1" pageNumber="591" type="male"></specimenCount>
(NMPC), labelled: “
<collectingCountry id="E59D5DBCF82E296DBFCE874AFC86FAA7" box="[688,782,1359,1385]" name="Nepal" pageId="1" pageNumber="591">NEPAL</collectingCountry>
.
<location id="98554BF7F82E296DBE69874AFCFFFAA7" LSID="urn:lsid:plazi:treatment:1523AC3AF82E2968BDE986E7FAF0F83B:98554BF7F82E296DBE69874AFCFFFAA7" box="[791,887,1359,1385]" country="Nepal" municipality="Besisahar" name="Manaslu" pageId="1" pageNumber="591">Manaslu</location>
env. /
<collectingMunicipality id="7D518756F82E296DBEC3874AFBA2FAA7" box="[957,1066,1359,1385]" pageId="1" pageNumber="591">Besisahar</collectingMunicipality>
/
<quantity id="5A72B0C9F82E296DB943874AFB0CFAA7" box="[1085,1156,1359,1385]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="591" unit="m" value="800.0">
<elevation id="16A7FA1FF82E296DB943874AFB0CFAA7" box="[1085,1156,1359,1385]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="591" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="E9343BECF82E296DB9F1874BFB77FAA7" box="[1167,1279,1358,1385]" pageId="1" pageNumber="591" value="2015-04-23">
<collectingDate id="F970C204F82E296DB9F1874BFB77FAA7" box="[1167,1279,1358,1385]" pageId="1" pageNumber="591" value="2015-04-23">23.4.2015</collectingDate>
</date>
/ lgt. Orszulik [p] //
<typeStatus id="4231A38EF82E296DBDAC8776FEF0FA43" box="[210,376,1395,1421]" pageId="1" pageNumber="591" type="holotype">HOLOTYPUS</typeStatus>
<specimenCount id="8B8CD6A5F82E296DBC038776FE1AFA43" box="[381,402,1395,1421]" count="1" pageId="1" pageNumber="591" type="male"></specimenCount>
/
<taxonomicName id="5A8A66AFF82E296DBCDB8776FD0AFA43" authorityName="Kirejtshuk" authorityYear="1985" box="[421,642,1395,1421]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82E296DBCDB8776FD0AFA43" box="[421,642,1395,1421]" italics="true" pageId="1" pageNumber="591">VIETTHERCHNUS</emphasis>
</taxonomicName>
/
<taxonomicName id="5A8A66AFF82E296DBFEA8776FD7CFA43" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[660,756,1395,1421]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF82E296DBFEA8776FD7CFA43" box="[660,756,1395,1421]" italics="true" pageId="1" pageNumber="591">orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F82E296DBF878776FCC6FA43" box="[761,846,1395,1421]" pageId="1" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
/ JelÍnek &amp;
<determinerName id="68E9F1E8F82E296DBEB18776FB9AFA43" box="[975,1042,1395,1421]" pageId="1" pageNumber="591">Hájek</determinerName>
det. 2024 [red label, p]”
</materialsCitation>
.
<materialsCitation id="2DE21771F82E296DB8568776FD60FA7F" collectingDate="2015-04-23" country="Nepal" determinerName="Hajek" elevation="800" location="Manaslu" municipality="Besisahar" pageId="1" pageNumber="591" specimenCount="10" specimenCount-female="4" specimenCount-male="6" typeStatus="paratype">
<typeStatus id="4231A38EF82E296DB8568776FA11FA43" box="[1320,1433,1395,1421]" pageId="1" pageNumber="591" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="8B8CD6A5F82E296DBDE98792FF4AFA7F" box="[151,194,1431,1457]" count="6" pageId="1" pageNumber="591" type="male">6 ♂</specimenCount>
,
<specimenCount id="8B8CD6A5F82E296DBDB38792FF7EFA7F" box="[205,246,1431,1457]" count="4" pageId="1" pageNumber="591" type="female">4 ♀</specimenCount>
, the same data as holotype (KOFM, NMPC)
</materialsCitation>
;
<materialsCitation id="2DE21771F82E296DBF8F8792FD43FA37" country="Nepal" elevation="1260" latitude="27.29" location="Kangchenjunga Himal Mts" longLatPrecision="742" longitude="87.45" pageId="1" pageNumber="591" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="8B8CD6A5F82E296DBF8F8792FC94FA7F" box="[753,796,1431,1457]" count="1" pageId="1" pageNumber="591" type="male">1 ♂</specimenCount>
,
<specimenCount id="8B8CD6A5F82E296DBE598792FCD8FA7F" box="[807,848,1431,1457]" count="1" pageId="1" pageNumber="591" type="female">1 ♀</specimenCount>
, “E-NEPAL, 30/6-1/7.2000, /
<location id="98554BF7F82E296DB9DF8792FF4DFA1B" LSID="urn:lsid:plazi:treatment:1523AC3AF82E2968BDE986E7FAF0F83B:98554BF7F82E296DB9DF8792FF4DFA1B" country="Nepal" latitude="27.29" longLatPrecision="742" longitude="87.45" name="Kangchenjunga Himal Mts" pageId="1" pageNumber="591">Kangchenjunga Himal Mts</location>
, / CHIRUWA vill.
<quantity id="5A72B0C9F82E296DBCE487BEFE6FFA1B" box="[410,487,1467,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="591" unit="m" value="1260.0">
<elevation id="16A7FA1FF82E296DBCE487BEFE6FFA1B" box="[410,487,1467,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="591" unit="m" value="1260.0">1260m</elevation>
</quantity>
/
<geoCoordinate id="F8BE7BEBF82E296DBC8387BEFDD8FA1B" box="[509,592,1467,1493]" degrees="27.29" direction="north" orientation="latitude" pageId="1" pageNumber="591" precision="555" value="27.29">27.29N</geoCoordinate>
<geoCoordinate id="F8BE7BEBF82E296DBF2887BFFD2EFA1B" box="[598,678,1466,1493]" degrees="87.45" direction="east" orientation="longitude" pageId="1" pageNumber="591" precision="555" value="87.45">87.45E</geoCoordinate>
[GPS] env. / David Král lgt. [p] //
<collectingCountry id="E59D5DBCF82E296DB95787BEFB08FA1B" box="[1065,1152,1467,1493]" name="Nepal" pageId="1" pageNumber="591">NEPAL</collectingCountry>
Expedition, / Jan Farkač, David Král / &amp; Jan Schneider, 2000 [p]” (NMPC)
</materialsCitation>
;
<materialsCitation id="2DE21771F82E296DBFA887DAFB9EF9D3" collectingDate="2017-05-04" collectorName="E. Kucera" country="Nepal" location="Beni" municipality="Pokhara" pageId="1" pageNumber="591" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="8B8CD6A5F82E296DBFA887DAFC8AFA37" box="[726,770,1503,1529]" count="1" pageId="1" pageNumber="591" type="male">1 ♂</specimenCount>
, “
<collectingCountry id="E59D5DBCF82E296DBE6687DAFCFAFA37" box="[792,882,1503,1529]" name="Nepal" pageId="1" pageNumber="591">NEPAL</collectingCountry>
central /
<location id="98554BF7F82E296DBEA587DAFB98FA37" LSID="urn:lsid:plazi:treatment:1523AC3AF82E2968BDE986E7FAF0F83B:98554BF7F82E296DBEA587DAFB98FA37" box="[987,1040,1503,1529]" country="Nepal" municipality="Pokhara" name="Beni" pageId="1" pageNumber="591">Beni</location>
west of
<collectingMunicipality id="7D518756F82E296DB9F887DAFB6AFA37" box="[1158,1250,1503,1529]" pageId="1" pageNumber="591">Pokhara</collectingMunicipality>
/ 3.5.-
<date id="E9343BECF82E296DB85B87DBFA05FA37" box="[1317,1421,1502,1529]" pageId="1" pageNumber="591" value="2017-05-04">
<collectingDate id="F970C204F82E296DB85B87DBFA05FA37" box="[1317,1421,1502,1529]" pageId="1" pageNumber="591" value="2017-05-04">4.5.2017</collectingDate>
</date>
/ lgt.
<collectorName id="307F78FAF82E296DBDBD8406FEA3F9D3" box="[195,299,1539,1565]" pageId="1" pageNumber="591">E.Kučera</collectorName>
[p]” (NMPC). All
<typeStatus id="4231A38EF82E296DBF7E8406FDE3F9D3" box="[512,619,1539,1565]" pageId="1" pageNumber="591" type="paratype">paratypes</typeStatus>
with the appropriate printed red label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F82E2968BDB98423FD83F98B" lastPageId="4" lastPageNumber="594" pageId="1" pageNumber="591" type="description">
<paragraph id="9D351D2CF82E296DBDB98423FB52F98F" blockId="1.[151,1437,1358,2033]" box="[199,1242,1574,1601]" pageId="1" pageNumber="591">
<emphasis id="AFFEC13EF82E296DBDB98423FD00F98F" bold="true" box="[199,648,1574,1601]" pageId="1" pageNumber="591">
Description of male
<typeStatus id="4231A38EF82E296DBCC98422FDA8F98F" box="[439,544,1575,1601]" pageId="1" pageNumber="591" type="holotype">holotype</typeStatus>
.
<emphasis id="AFFEC13EF82E296DBF528423FD00F98F" bold="true" box="[556,648,1574,1601]" italics="true" pageId="1" pageNumber="591">Habitus</emphasis>
</emphasis>
broadly oval, convex, glabrous, smooth and shining.
</paragraph>
<paragraph id="9D351D2CF82E296DBDB9844FFDACF93B" blockId="1.[151,1437,1358,2033]" pageId="1" pageNumber="591">
<emphasis id="AFFEC13EF82E296DBDB9844FFED2F9AB" bold="true" box="[199,346,1610,1637]" italics="true" pageId="1" pageNumber="591">Colouration.</emphasis>
Orange, pronotum with four round black spots, two smaller at anterior margin, two larger at basal margin besides scutellum. Each elytron with following black spots: one at humeral bulge, one behind it near mid-length, one round spot at median axis at ca. one third of elytra length; further there are two common spots at suture: one narrow oblong just behind scutellum, and one broad subtriangular one near apex. Legs and antennae yellow, antennal club black-brown (
<figureCitation id="05B101A9F82E296DBCAF84DEFD9AF93B" box="[465,530,1755,1781]" captionStart="FIGURES 14" captionStartId="3.[152,270,1922,1946]" captionTargetBox="[168,1421,213,1867]" captionTargetId="figure-32@3.[151,1436,181,1899]" captionTargetPageId="3" captionText="FIGURES 14. Dorsal habitus of Viettherchnus. 1. V. orszuliki sp. nov. (holotype ♂); 2. V. apriculus sp. nov. (holotype ♂); 3. V. helferi sp. nov. (paratype ♀, dark form; Kalimantan); 4. V. helferi sp. nov. (holotype ♂). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916277" httpUri="https://zenodo.org/record/13916277/files/figure.png" pageId="1" pageNumber="591">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF82E296EBDB984FBFD2AFE63" blockId="1.[151,1437,1358,2033]" lastBlockId="2.[151,1437,151,2039]" lastPageId="2" lastPageNumber="592" pageId="1" pageNumber="591">
<emphasis id="AFFEC13EF82E296DBDB984FBFE8DF8D7" bold="true" box="[199,261,1790,1817]" italics="true" pageId="1" pageNumber="591">Head</emphasis>
transverse, eyes on small conical protuberances, with sharp transverse edges between eye and antenna insertion, temples rectilinear, converging posteriad. Vertex and lateral portions of frons feebly convex, median portion of frons between antennae markedly concave, anterior portion as well as clypeus almost horizontal and moderately transversely convex. Sides of frons besides eyes distinctly punctate, punctures almost equal in size to eye-facets, separated by one diameter or more, interspaces smooth and shining. Median portion as well as clypeus smooth and shining with very fine and widely spaced punctures. Anterior margin of clypeus subtruncate, not bordered. Labrum transverse, anterior margin arcuate laterally, subtruncate in middle without distinct incision. Mandibles arcuately dilated laterally with sharp, raised angulate dorsal edge and large, shallowly concave face against eye (
<figureCitation id="05B101A9F82D296EBC5282BEFEFAFF1B" box="[300,370,186,213]" captionStart="FIGURES 57" captionStartId="4.[152,270,932,956]" captionTargetBox="[176,1411,207,909]" captionTargetId="figure-19@4.[176,1411,181,909]" captionTargetPageId="4" captionText="FIGURES 57. Head of male Viettherchnus. 5. V. orszuliki sp. nov.; 6. V. apriculus sp. nov.; 7. V. helferi sp. nov. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.13916279" httpUri="https://zenodo.org/record/13916279/files/figure.png" pageId="2" pageNumber="592">Fig. 5</figureCitation>
). Antennae as long as width of head across eyes (ratio HEAW/ANLE = 1.00); club oblong oval. Scapus almost 3× wider than pedicel, 1.375× longer than wide, widest at base and subrectangular, gradually narrowed distad, with arcuate sharp anterior margin and well defined shallow anteocular concavity on posterior face. Pedicel length/width = 4/3, hardly wider than following antennomeres. Antennomere III ca. twice as long as wide, IV ca. as long as wide, its anterior side angulate, V 1.33× longer than wide, VI and VII as long as wide, VIII transverse. Ratio ANCL/ANCW = 1.78, antennomere XI nearly as long as IX + X combined, X lunular, shorter than neighbouring ones; ratio ANLE/ANCL = 2.60.
</paragraph>
<paragraph id="9D351D2CF82D296EBDB983B2FEB5FD67" blockId="2.[151,1437,151,2039]" pageId="2" pageNumber="592">
<emphasis id="AFFEC13EF82D296EBDB983B2FEB4FE1F" bold="true" box="[199,316,439,465]" italics="true" pageId="2" pageNumber="592">Pronotum</emphasis>
transverse, strongly transversely convex, widest at posterior angles and strongly arcuately narrowed anteriad. Anterior margin deeply trapezoidally emarginate, indistinctly bordered. Anterior angles broad, prominent, obtuse with more or less rounded tips. Lateral margins broadly arcuate, not explanate, finely bordered, visible simultaneously from above in their entirety. Basal margin not bordered, in front of scutellum with broad truncate projection, besides it broadly arcuate. Posterior angles obtusely angulate. Punctures almost equal in size to eye-facets, separated by several diameters, interspaces smooth and shining. Scutellar shield large, subtriangular, punctate like pronotum.
</paragraph>
<paragraph id="9D351D2CF82D296EBDB980B7FB15FC4F" blockId="2.[151,1437,151,2039]" pageId="2" pageNumber="592">
<emphasis id="AFFEC13EF82D296EBDB980B7FE87FD03" bold="true" box="[199,271,690,717]" italics="true" pageId="2" pageNumber="592">Elytra</emphasis>
widest near base, gradually arcuately narrowed posteriad and simultaneously rounded apically, strongly convex both longitudinally and transversely, without distinct humeral bulges; lateral margins not explanate, not visible simultaneously from above. Surface of each elytron with eight series of punctures nearly equal to those of pronotum and within one row separated by ca. one diameter. Analogous, but diffuse, punctures occur also along apical portion of suture. Sutural lines indistinct. Interstries finely diffusely punctulate with widely spaced punctures corresponding to those of concave median portion of frons. Interspaces smooth and shining.
</paragraph>
<paragraph id="9D351D2CF82D296EBDB9818FFBF7FC07" blockId="2.[151,1437,151,2039]" pageId="2" pageNumber="592">
<emphasis id="AFFEC13EF82D296EBDB9818FFEBDFC6B" bold="true" box="[199,309,906,933]" italics="true" pageId="2" pageNumber="592">Pygidium</emphasis>
subtriangular, broadly rounded apically. Punctures larger than eye-facets and bearing rudimentary setae, separated by one diameter or less, interspaces obsoletely reticulate, somewhat dull.
</paragraph>
<paragraph id="9D351D2CF82D296EBDB981D7FF3DF9A5" blockId="2.[151,1437,151,2039]" pageId="2" pageNumber="592">
<emphasis id="AFFEC13EF82D296EBDB981D7FE09FC22" bold="true" box="[199,385,978,1005]" italics="true" pageId="2" pageNumber="592">Ventral surface.</emphasis>
Postmentum ca. 2.5× wider than long, its anterior margin feebly concave, punctures larger than eye-facets, shallow, separated by one diameter or less, interspaces smooth and shining. Antennal furrows well defined with raised margins, converging posteriorly. Prosternum short with sharp mediolongitudinal carina gradually vanishing on prosternal process; punctures fairly twice as large as eye-facets, flat, separated by less than one diameter, interspaces shining. Prosternal process with blunt arcuate apical margin behind procoxae, its apical wall convex, obliquely sloping dorsad, with subtruncate apical margin. Hypomera broad, concave, impunctate, smooth and shining, postcoxal pieces punctate like prosternum. Mesoventrite situated markedly lower than metaventrite (ventral view), transversely convex, mediolongitudinally carinate. Anterior portion of mesepisternum (prepectus) flat, indistinctly punctate, alutaceous, dull; posterior portion concave, punctate like mesoventrite, shining. Mesocoxal axillary lines in median half transverse, running near posterior margins of mesocoxal cavities, their outer recurrent portion arcuate, meeting metasternopleural suture before its mid-length. Metaventrite almost flat except for large concavities in front of metacoxae, its anterior intercoxal margin concave, extending over intercoxal piece of mesoventrite, posterior intercoxal margin truncate. Discrimen absent. Punctures in median portion indistinct, very fine and widely spaced, becoming larger laterally, at sides fairly equal in size to eye-facets and separated by nearly one diameter. Metacoxal axillary lines running close to posterior margins of metacoxal cavity all along its length, axillary space not developed. Abdominal ventrite I as long as IIIV combined, punctures of all ventrites equal, nearly equal in size to eye-facets, but very shallow, separated by nearly one diameter and bearing long semirecumbent yellow setae; interspaces smooth and shining. Hypopygium transverse, flatly rounded apically (
<figureCitation id="05B101A9F82D296EB80E8428FF2FF9A5" captionStart="FIGURES 89" captionStartId="5.[152,270,1037,1061]" captionTargetBox="[173,1417,205,998]" captionTargetId="figure-32@5.[151,1436,181,1013]" captionTargetPageId="5" captionText="FIGURES 89. Ventral habitus of Viettherchnus. 8. V. orszuliki sp. nov. (paratype ♂); 9. V. helferi sp. nov. (paratype ♀; Kalimantan). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916281" httpUri="https://zenodo.org/record/13916281/files/figure.png" pageId="2" pageNumber="592">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF82D296EBDB98470FF61F839" blockId="2.[151,1437,151,2039]" pageId="2" pageNumber="592">
<emphasis id="AFFEC13EF82D296EBDB98470FE8DF941" bold="true" box="[199,261,1653,1679]" italics="true" pageId="2" pageNumber="592">Legs.</emphasis>
Distances between pro-, meso- and metacoxae as 7: 16: 16. Profemur twice as long as wide, dorsal anterior margin almost rectilinear, ventral one in distal half arcuately emarginate. Posterior margin subrectilinear, in apical portion arcuate. Protibia shorter than femur (33/45), 3.3× longer than wide, subparallel-sided, only in basal fifth arcuately narrowed proximad; outer margin densely crenulate, outer subapical angle subrectangular. Mesofemur broad (length/width= 45/25), its posterior margin almost rectilinear, in ca. 2/3 of its length abruptly curved and then again as subrectilinear running towards distal end of femur. Mesotibia 2.69× longer than wide, mesofemur 1.34× longer than mesotibia, subparallel-sided in two distal thirds, in basal third arcuately narrowed towards base. Metafemur 1.83× longer than wide, widest at two thirds of its length, its posterior margin in two basal thirds subrectilinear, in distal third arcuate. Metatibia as long as femur, subparallel-sided, in basal third arcuately narrowed proximad. Meso- and metatarsomeres IIII bilobed, tarsomere V as long as IIV combined. Tarsal claws simple.
</paragraph>
<caption id="C9F54DA4F82C296FBDE68587FB52F870" ID-DOI="http://doi.org/10.5281/zenodo.13916277" ID-Zenodo-Dep="13916277" httpUri="https://zenodo.org/record/13916277/files/figure.png" pageId="3" pageNumber="593" startId="3.[152,270,1922,1946]" targetBox="[168,1421,213,1867]" targetPageId="3" targetType="figure">
<paragraph id="9D351D2CF82C296FBDE68587FB52F870" blockId="3.[151,1436,1922,1983]" pageId="3" pageNumber="593">
<emphasis id="AFFEC13EF82C296FBDE68587FEC9F855" bold="true" box="[152,321,1922,1947]" pageId="3" pageNumber="593">FIGURES 14.</emphasis>
Dorsal habitus of
<taxonomicName id="5A8A66AFF82C296FBC828586FD0EF855" authorityName="Kirejtshuk" authorityYear="1985" box="[508,646,1923,1947]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82C296FBC828586FD0EF855" box="[508,646,1923,1947]" italics="true" pageId="3" pageNumber="593">Viettherchnus</emphasis>
</taxonomicName>
. 1.
<taxonomicName id="5A8A66AFF82C296FBFD28586FC93F855" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[684,795,1923,1947]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF82C296FBFD28586FC93F855" box="[684,795,1923,1947]" italics="true" pageId="3" pageNumber="593">V. orszuliki</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF82C296FBE5C8586FCFBF855" bold="true" box="[802,883,1923,1947]" pageId="3" pageNumber="593">
<taxonomicNameLabel id="B4CD7C45F82C296FBE5C8586FCFBF855" box="[802,883,1923,1947]" pageId="3" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype ♂); 2.
<taxonomicName id="5A8A66AFF82C296FB95D8586FB14F855" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[1059,1180,1923,1947]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="apriculus" status="sp. nov.">
<emphasis id="AFFEC13EF82C296FB95D8586FB14F855" box="[1059,1180,1923,1947]" italics="true" pageId="3" pageNumber="593">V. apriculus</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF82C296FB9DC8586FB7AF855" bold="true" box="[1186,1266,1923,1947]" pageId="3" pageNumber="593">
<taxonomicNameLabel id="B4CD7C45F82C296FB9DC8586FB7AF855" box="[1186,1266,1923,1947]" pageId="3" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype ♂); 3.
<taxonomicName id="5A8A66AFF82C296FBDE985A2FF7AF871" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[151,242,1959,1983]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF82C296FBDE985A2FF7AF871" box="[151,242,1959,1983]" italics="true" pageId="3" pageNumber="593">V. helferi</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF82C296FBD8785A2FEC1F871" bold="true" box="[249,329,1959,1983]" pageId="3" pageNumber="593">
<taxonomicNameLabel id="B4CD7C45F82C296FBD8785A2FEC1F871" box="[249,329,1959,1983]" pageId="3" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(paratype ♀, dark form; Kalimantan); 4.
<taxonomicName id="5A8A66AFF82C296FBF9385A2FCC0F871" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[749,840,1959,1983]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="593" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF82C296FBF9385A2FCC0F871" box="[749,840,1959,1983]" italics="true" pageId="3" pageNumber="593">V. helferi</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF82C296FBE3085A2FC17F871" bold="true" box="[846,927,1959,1983]" pageId="3" pageNumber="593">
<taxonomicNameLabel id="B4CD7C45F82C296FBE3085A2FC17F871" box="[846,927,1959,1983]" pageId="3" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype ♂). Scale bar 1 mm.
</paragraph>
</caption>
<caption id="C9F54DA4F82B2968BDE681A1FF59FC2F" ID-DOI="http://doi.org/10.5281/zenodo.13916279" ID-Zenodo-Dep="13916279" httpUri="https://zenodo.org/record/13916279/files/figure.png" pageId="4" pageNumber="594" startId="4.[152,270,932,956]" targetBox="[176,1411,207,909]" targetPageId="4" targetType="figure">
<paragraph id="9D351D2CF82B2968BDE681A1FF59FC2F" blockId="4.[151,1436,932,993]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BDE681A1FECCFC73" bold="true" box="[152,324,932,957]" pageId="4" pageNumber="594">FIGURES 57.</emphasis>
Head of male
<taxonomicName id="5A8A66AFF82B2968BC9D81A0FDE5FC73" authorityName="Kirejtshuk" authorityYear="1985" box="[483,621,933,957]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82B2968BC9D81A0FDE5FC73" box="[483,621,933,957]" italics="true" pageId="4" pageNumber="594">Viettherchnus</emphasis>
</taxonomicName>
. 5.
<taxonomicName id="5A8A66AFF82B2968BFE681A0FC82FC73" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[664,778,933,957]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF82B2968BFE681A0FC82FC73" box="[664,778,933,957]" italics="true" pageId="4" pageNumber="594">V. orszuliki</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF82B2968BE6A81A0FCEFFC73" bold="true" box="[788,871,933,957]" pageId="4" pageNumber="594">
<taxonomicNameLabel id="B4CD7C45F82B2968BE6A81A0FCEFFC73" box="[788,871,933,957]" pageId="4" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 6.
<taxonomicName id="5A8A66AFF82B2968BEED81A0FB86FC73" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[915,1038,933,957]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="apriculus" status="sp. nov.">
<emphasis id="AFFEC13EF82B2968BEED81A0FB86FC73" box="[915,1038,933,957]" italics="true" pageId="4" pageNumber="594">V. apriculus</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF82B2968B96981A0FBE2FC73" bold="true" box="[1047,1130,933,957]" pageId="4" pageNumber="594">
<taxonomicNameLabel id="B4CD7C45F82B2968B96981A0FBE2FC73" box="[1047,1130,933,957]" pageId="4" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 7.
<taxonomicName id="5A8A66AFF82B2968B9E981A0FB7CFC73" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[1175,1268,933,957]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="AFFEC13EF82B2968B9E981A0FB7CFC73" box="[1175,1268,933,957]" italics="true" pageId="4" pageNumber="594">V. helferi</emphasis>
</taxonomicName>
<emphasis id="AFFEC13EF82B2968B98381A0FAD8FC73" bold="true" box="[1277,1360,933,957]" pageId="4" pageNumber="594">
<taxonomicNameLabel id="B4CD7C45F82B2968B98381A0FAD8FC73" box="[1277,1360,933,957]" pageId="4" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Not in scale.
</paragraph>
</caption>
<paragraph id="9D351D2CF82B2968BDB98637FB89FBBF" blockId="4.[151,1437,1074,2037]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BDB98637FEFEFB83" bold="true" box="[199,374,1074,1101]" italics="true" pageId="4" pageNumber="594">Male genitalia.</emphasis>
Penis broad and short, apical margin broadly rounded with short rounded apical protuberance (
<figureCitation id="05B101A9F82B2968BDE18652FF7BFBBF" box="[159,243,1111,1137]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="4" pageNumber="594">Fig. 10</figureCitation>
). Tegmen 1.25× longer than wide, broadly rounded apically (
<figureCitation id="05B101A9F82B2968BEDF8652FC7CFBBF" box="[929,1012,1111,1137]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="4" pageNumber="594">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF82B2968BDB9867EFBF5FAA3" blockId="4.[151,1437,1074,2037]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BDB9867EFEADFB5B" bold="true" box="[199,293,1147,1173]" pageId="4" pageNumber="594">Female.</emphasis>
Externally similar to male except for absence of secondary sexual characters. Frons almost flat, situated approximately at the same level as clypeus and separated from it by shallow transverse impression between antennal insertions. Entire frons punctate with punctures almost equal in size to eye-facets, separated by one diameter or more, interspaces smooth and shining. Clypeus smooth and shining with very fine and widely spaced indistinct punctures. Mandibles dorsoventrally flattened, not extended laterally, outer margin sharp, but not raised. Scapus ca. twice as long as wide, base not angulate, anterior face convex, posterior one with well defined shallow anteocular concavity. Ovipositor serrate in apical third, apex acutely pointed, without styli (
<figureCitation id="05B101A9F82B2968B9628756FBE7FAA3" box="[1052,1135,1363,1389]" captionStart="FIGURES 1017" captionStartId="8.[152,270,1901,1925]" captionTargetBox="[185,1403,212,1865]" captionTargetId="figure-19@8.[172,1416,199,1878]" captionTargetPageId="8" captionText="FIGURES 1017. Genitalia of Viettherchnus. 1012. penis with internal sac in dorsal view; 1315. Tegmen; 1617. Ovipositor. 10, 13, 16. V. orszuliki sp. nov.; 11, 14. V. apriculus sp. nov.; 12, 15, 17. V. helferi sp. nov. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916285" httpUri="https://zenodo.org/record/13916285/files/figure.png" pageId="4" pageNumber="594">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="9D351D2CF82B2968BDB98772FB03FA33" blockId="4.[151,1437,1074,2037]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BDB98772FEB7FA5F" bold="true" box="[199,319,1399,1425]" pageId="4" pageNumber="594">Variation.</emphasis>
Colour pattern variable, in specimens with most extended black pigmentation pronotum mostly black with orange borders at sides and lateral portions of basal margin, as well as with more or less interrupted zigzag transverse bar on disc; elytra black, each with large orange ellipse in sutural half, reaching from basal margin to ca. two thirds of elytra length and projecting shortly laterad; also humeral angle orange.
</paragraph>
<paragraph id="9D351D2CF82B2968BDB98402FD83F98B" blockId="4.[151,1437,1074,2037]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BDB98402FEF6F9EF" bold="true" box="[199,382,1543,1569]" pageId="4" pageNumber="594">Measurements.</emphasis>
Body length 2.43.0 mm (
<typeStatus id="4231A38EF82B2968BFD28402FC98F9EF" box="[684,784,1543,1569]" pageId="4" pageNumber="594" type="holotype">holotype</typeStatus>
<quantity id="5A72B0C9F82B2968BE698402FCF7F9EF" box="[791,895,1543,1570]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" pageId="4" pageNumber="594" unit="mm" value="2.95">2.95 mm</quantity>
), width 1.72.0 mm (
<typeStatus id="4231A38EF82B2968B9118402FB5BF9EF" box="[1135,1235,1543,1569]" pageId="4" pageNumber="594" type="holotype">holotype</typeStatus>
<quantity id="5A72B0C9F82B2968B9A58402FACBF9EF" box="[1243,1347,1543,1570]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="4" pageNumber="594" unit="mm" value="1.95">1.95 mm</quantity>
), height 1.0
<quantity id="5A72B0C9F82B2968BDB7842EFE96F98B" box="[201,286,1579,1605]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="594" unit="mm" value="1.4">1.4 mm</quantity>
(
<typeStatus id="4231A38EF82B2968BC52842EFE18F98B" box="[300,400,1579,1605]" pageId="4" pageNumber="594" type="holotype">holotype</typeStatus>
<quantity id="5A72B0C9F82B2968BCE9842EFE77F98B" box="[407,511,1579,1605]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="594" unit="mm" value="1.4">1.40 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F82B2968BDB9844AFE3DF81F" pageId="4" pageNumber="594" type="diagnosis">
<paragraph id="9D351D2CF82B2968BDB9844AFB96F97F" blockId="4.[151,1437,1074,2037]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BDB9844AFE58F9A7" bold="true" box="[199,464,1615,1641]" pageId="4" pageNumber="594">Differential diagnosis.</emphasis>
The combination of a marked orange and black colour pattern, antennal club shorter than antennomeres IIVIII combined and distinct serial punctation on elytra suggest close relation of
<taxonomicName id="5A8A66AFF82B2968B9818471FAFEF943" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[1279,1398,1651,1677]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF82B2968B9818471FAFEF943" box="[1279,1398,1651,1677]" italics="true" pageId="4" pageNumber="594">V orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F82B2968B8028476FF4FF97F" pageId="4" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
to
<taxonomicName id="5A8A66AFF82B2968BD948492FCE3F97F" authority="Kirejtshuk &amp; Kirk-Spriggs, 1996" authorityName="Kirejtshuk &amp; Kirk-Spriggs" authorityYear="1996" box="[234,875,1687,1713]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="AFFEC13EF82B2968BD948492FE7AF97F" box="[234,498,1687,1713]" italics="true" pageId="4" pageNumber="594">Viettherchnus plagiatus</emphasis>
<bibRefCitation id="F91B60DDF82B2968BC878492FCE3F97F" author="Kirejtshuk, A. G. &amp; Kirk-Spriggs, A. H." box="[505,875,1687,1713]" pageId="4" pageNumber="594" pagination="131 - 138" refId="ref7251" refString="Kirejtshuk, A. G. &amp; Kirk-Spriggs, A. H. (1996) Viettherchnus Kirejtshuk, 1985 and Ceramphosia gen. n. from the Indo-Malayan Region (Coleoptera: Nitidulidae). Zoosystematica Rossica, 4, 131 - 138. [1995]" type="journal article" year="1996">Kirejtshuk &amp; Kirk-Spriggs, 1996</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="E59D5DBCF82B2968BED18492FB91F97F" box="[943,1049,1687,1713]" name="Malaysia" pageId="4" pageNumber="594">Malaysia</collectingCountry>
.
</paragraph>
<paragraph id="9D351D2CF82B2968BDB984BEFB81F8AB" blockId="4.[151,1437,1074,2037]" pageId="4" pageNumber="594">
The peculiar colour pattern of the new species resembles that of some
<taxonomicName id="5A8A66AFF82B2968BE9C84BEFB14F91B" box="[994,1180,1723,1749]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BE9C84BEFBCCF91B" box="[994,1092,1723,1749]" italics="true" pageId="4" pageNumber="594">Cyllodes</emphasis>
species
</taxonomicName>
like
<taxonomicName id="5A8A66AFF82B2968B9AB84BEFEB1F937" authority="(Reitter, 1878)" baseAuthorityName="Reitter" baseAuthorityYear="1878" class="Insecta" family="Nitidulidae" genus="Cyllodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="literatus">
<emphasis id="AFFEC13EF82B2968B9AB84BEFA14F91B" box="[1237,1436,1723,1749]" italics="true" pageId="4" pageNumber="594">Cyllodes literatus</emphasis>
(
<bibRefCitation id="F91B60DDF82B2968BDE184DAFEB8F937" author="Reitter, E." box="[159,304,1759,1785]" pageId="4" pageNumber="594" pagination="89 - 90" refId="ref7299" refString="Reitter, E. (1878) [Neue Japanische Kafer]. Deutsche Entomologische Zeitschrift, 22, 89 - 90." type="book chapter" year="1878">Reitter, 1878</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="5A8A66AFF82B2968BC3B84DAFD40F937" authority="Grouvelle, 1914" authorityName="Grouvelle" authorityYear="1914" box="[325,712,1759,1785]" class="Insecta" family="Nitidulidae" genus="Cyllodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="multimaculatus">
<emphasis id="AFFEC13EF82B2968BC3B84DAFD98F937" box="[325,528,1759,1785]" italics="true" pageId="4" pageNumber="594">C. multimaculatus</emphasis>
<bibRefCitation id="F91B60DDF82B2968BF6B84DAFD40F937" author="Grouvelle, A." box="[533,712,1759,1785]" pageId="4" pageNumber="594" pagination="33 - 76" refId="ref6995" refString="Grouvelle, A. (1914) H. Sauter's Formosa-Ausbeute. Rhysodidae, Nitidulidae, Ostomidae, Colydiidae, Passandridae, Cucujidae, Cryptophagidae, Diphyllidae, Lathridiidae, Mycetophagidae, Dermestidae. Archiv fur Naturgeschichte, Abteilung A, 79 (11), 33 - 76. [1913]" type="journal article" year="1914">Grouvelle, 1914</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="5A8A66AFF82B2968BF9784DAFBAAF937" authority="Hisamatsu, 1961" authorityName="Hisamatsu" authorityYear="1961" box="[745,1058,1759,1785]" class="Insecta" family="Nitidulidae" genus="Cyllodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="nakanei">
<emphasis id="AFFEC13EF82B2968BF9784DAFCE9F937" box="[745,865,1759,1785]" italics="true" pageId="4" pageNumber="594">C. nakanei</emphasis>
<bibRefCitation id="F91B60DDF82B2968BE1B84DAFBAAF937" author="Hisamatsu, S." box="[869,1058,1759,1785]" pageId="4" pageNumber="594" pagination="26 - 32" refId="ref7048" refString="Hisamatsu, S. (1961) Four new species of Nitidulidae from Japan (Coleoptera). Transactions of the Shikoku Entomological Society, 7, 26 - 32." type="journal article" year="1961">Hisamatsu, 1961</bibRefCitation>
</taxonomicName>
. Female of
<taxonomicName id="5A8A66AFF82B2968B9E184DAFA14F937" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[1183,1436,1759,1785]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF82B2968B9E184DAFA14F937" box="[1183,1436,1759,1785]" italics="true" pageId="4" pageNumber="594">Viettherchnus orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F82B2968BDE98506FF66F8D3" box="[151,238,1795,1821]" pageId="4" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
differs from them in the abrupt boundary between distinctly punctate frons and indistinctly, finely punctate clypeus (in
<taxonomicName id="5A8A66AFF82B2968BC658522FE5EF88F" box="[283,470,1831,1857]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BC658522FEF5F88F" box="[283,381,1831,1857]" italics="true" pageId="4" pageNumber="594">Cyllodes</emphasis>
species
</taxonomicName>
punctation of frons becomes gradually finer towards clypeus) and blackish pygidium (in mentioned
<emphasis id="AFFEC13EF82B2968BC44854EFE19F8AB" box="[314,401,1867,1893]" italics="true" pageId="4" pageNumber="594">Cyllode</emphasis>
s species pygidium as well as entire ventral surface red).
</paragraph>
<paragraph id="9D351D2CF82B2968BDB9856AFE3DF81F" blockId="4.[151,1437,1074,2037]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BDB9856AFE13F847" bold="true" box="[199,411,1903,1929]" pageId="4" pageNumber="594">Name derivation.</emphasis>
The new species is dedicated to Kamil Orszulik (Frýdek-Místek,
<collectingCountry id="E59D5DBCF82B2968B9D1856AFAE2F847" box="[1199,1386,1903,1929]" name="Czech Republic" pageId="4" pageNumber="594">Czech Republic</collectingCountry>
), an enthusiastic amateur entomologist and collector of the most specimens of the new species. The specific epithet is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F82B2968BDB985DEFAF0F83B" box="[199,1400,2011,2037]" pageId="4" pageNumber="594" type="distribution">
<paragraph id="9D351D2CF82B2968BDB985DEFAF0F83B" blockId="4.[151,1437,1074,2037]" box="[199,1400,2011,2037]" pageId="4" pageNumber="594">
<emphasis id="AFFEC13EF82B2968BDB985DEFEE9F83B" bold="true" box="[199,353,2011,2037]" pageId="4" pageNumber="594">Distribution.</emphasis>
<taxonomicName id="5A8A66AFF82B2968BC1985DEFDEFF83B" authorityName="Jelínek &amp; Hájek" authorityYear="2024" box="[359,615,2011,2037]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="orszuliki" status="sp. nov.">
<emphasis id="AFFEC13EF82B2968BC1985DEFDEFF83B" box="[359,615,2011,2037]" italics="true" pageId="4" pageNumber="594">Viettherchnus orszuliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4CD7C45F82B2968BF1085DEFD4DF83B" box="[622,709,2011,2037]" pageId="4" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
is known from several localities in central and eastern
<collectingCountry id="E59D5DBCF82B2968B85085DEFAFDF83B" box="[1326,1397,2011,2037]" name="Nepal" pageId="4" pageNumber="594">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,108 @@
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<subSubSection id="D5904EA7F82E296DBDE9815EFD92FCB8" box="[151,538,859,886]" pageId="1" pageNumber="591" type="nomenclature">
<paragraph id="9D351D2CF82E296DBDE9815EFD92FCB8" blockId="1.[151,538,859,886]" box="[151,538,859,886]" pageId="1" pageNumber="591">
<heading id="C67DAA40F82E296DBDE9815EFD92FCB8" bold="true" box="[151,538,859,886]" fontSize="11" level="1" pageId="1" pageNumber="591" reason="1">
<taxonomicName id="5A8A66AFF82E296DBDE9815EFD92FCB8" authority="Kirejtshuk, 1985" authorityName="Kirejtshuk" authorityYear="1985" box="[151,538,859,886]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82E296DBDE9815EFD92FCB8" bold="true" box="[151,538,859,886]" pageId="1" pageNumber="591">
<emphasis id="AFFEC13EF82E296DBDE9815EFEC9FCB8" bold="true" box="[151,321,859,886]" italics="true" pageId="1" pageNumber="591">Viettherchnus</emphasis>
<bibRefCitation id="F91B60DDF82E296DBC368159FD92FCB8" author="Kirejtshuk, A. G." box="[328,538,860,886]" pageId="1" pageNumber="591" pagination="157 - 164" refId="ref7079" refString="Kirejtshuk, A. G. (1985) Novye vidy Cyllodes Erichson i Viettherchnus gen. n. (Coleoptera, Nitidulidae) fauny Vietnama i sopredel'nykh territorii [New species of Cyllodes Erichson and Viettherchnus gen. n. (Coleoptera, Nitidulidae) of the fauna of Vietnam and adjoining territories]. In: Medvedev, L. N. (Ed.), Nasekomye Vietnama [Insects of Vietnam]. Nauka, Moskva, pp. 157 - 164. [in Russian]" type="book chapter" year="1985">Kirejtshuk, 1985</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D5904EA7F82E296DBDE981A5FD84FC76" box="[151,524,928,952]" pageId="1" pageNumber="591" type="reference_group">
<paragraph id="9D351D2CF82E296DBDE981A5FD84FC76" blockId="1.[151,524,928,952]" box="[151,524,928,952]" pageId="1" pageNumber="591">
<treatmentCitationGroup id="BD9A3A02F82E296DBDE981A5FD84FC76" box="[151,524,928,952]" pageId="1" pageNumber="591">
<treatmentCitation id="1C2B3B3DF82E296DBDE981A5FD80FC76" author="Kirejtshuk, A. G." box="[151,520,928,952]" page="163" pageId="1" pageNumber="591" year="1985">
<taxonomicName id="5A8A66AFF82E296DBDE981A5FD80FC76" authority="Kirejtshuk, 1985: 163" authorityName="Kirejtshuk" authorityPageNumber="163" authorityYear="1985" box="[151,520,928,952]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82E296DBDE981A5FEA9FC76" box="[151,289,928,952]" italics="true" pageId="1" pageNumber="591">Viettherchnus</emphasis>
<bibRefCitation id="F91B60DDF82E296DBC5981A5FD80FC76" author="Kirejtshuk, A. G." box="[295,520,928,952]" pageId="1" pageNumber="591" pagination="157 - 164" refId="ref7079" refString="Kirejtshuk, A. G. (1985) Novye vidy Cyllodes Erichson i Viettherchnus gen. n. (Coleoptera, Nitidulidae) fauny Vietnama i sopredel'nykh territorii [New species of Cyllodes Erichson and Viettherchnus gen. n. (Coleoptera, Nitidulidae) of the fauna of Vietnam and adjoining territories]. In: Medvedev, L. N. (Ed.), Nasekomye Vietnama [Insects of Vietnam]. Nauka, Moskva, pp. 157 - 164. [in Russian]" type="book chapter" year="1985">Kirejtshuk, 1985: 163</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<subSubSection id="D5904EA7F82E296DBDE981E2FE46FB5F" pageId="1" pageNumber="591" type="materials_examined">
<paragraph id="9D351D2CF82E296DBDE981E2FB96FBCF" blockId="1.[151,1436,998,1169]" box="[151,1054,998,1025]" pageId="1" pageNumber="591">
<typeStatus id="4231A38EF82E296DBDE981E2FF47FBCF" box="[151,207,999,1025]" pageId="1" pageNumber="591">Type</typeStatus>
species:
<taxonomicName id="5A8A66AFF82E296DBC4B81E2FD72FBCF" authority="Kirejtshuk, 1985" authorityName="Kirejtshuk" authorityYear="1985" box="[309,762,998,1025]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="species" species="finitimus">
<emphasis id="AFFEC13EF82E296DBC4B81E2FDBFFBCF" box="[309,567,999,1025]" italics="true" pageId="1" pageNumber="591">Viettherchnus finitimus</emphasis>
<bibRefCitation id="F91B60DDF82E296DBF4081E2FD72FBCF" author="Kirejtshuk, A. G." box="[574,762,998,1025]" pageId="1" pageNumber="591" pagination="157 - 164" refId="ref7079" refString="Kirejtshuk, A. G. (1985) Novye vidy Cyllodes Erichson i Viettherchnus gen. n. (Coleoptera, Nitidulidae) fauny Vietnama i sopredel'nykh territorii [New species of Cyllodes Erichson and Viettherchnus gen. n. (Coleoptera, Nitidulidae) of the fauna of Vietnam and adjoining territories]. In: Medvedev, L. N. (Ed.), Nasekomye Vietnama [Insects of Vietnam]. Nauka, Moskva, pp. 157 - 164. [in Russian]" type="book chapter" year="1985">Kirejtshuk, 1985</bibRefCitation>
</taxonomicName>
(by original designation).
</paragraph>
<paragraph id="9D351D2CF82E296DBDB9860EFE46FB5F" blockId="1.[151,1436,998,1169]" pageId="1" pageNumber="591">
The genus was redescribed by
<bibRefCitation id="F91B60DDF82E296DBF65860EFC10FBEB" author="Kirejtshuk, A. G. &amp; Kirk-Spriggs, A. H." box="[539,920,1035,1061]" pageId="1" pageNumber="591" pagination="131 - 138" refId="ref7251" refString="Kirejtshuk, A. G. &amp; Kirk-Spriggs, A. H. (1996) Viettherchnus Kirejtshuk, 1985 and Ceramphosia gen. n. from the Indo-Malayan Region (Coleoptera: Nitidulidae). Zoosystematica Rossica, 4, 131 - 138. [1995]" type="journal article" year="1996">Kirejtshuk &amp; Kirk-Spriggs (1996)</bibRefCitation>
. It corresponds in most characters to
<taxonomicName id="5A8A66AFF82E296DB84C860EFA1CFBEB" authorityName="Erichson" authorityYear="1843" box="[1330,1428,1035,1061]" class="Insecta" family="Nitidulidae" genus="Cyllodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82E296DB84C860EFA1CFBEB" box="[1330,1428,1035,1061]" italics="true" pageId="1" pageNumber="591">Cyllodes</emphasis>
</taxonomicName>
; both genera shares, contrary to other Cyllodini, lobed metatarsomeres IIII.
<taxonomicName id="5A8A66AFF82E296DBE8E862AFB00FB87" authorityName="Kirejtshuk" authorityYear="1985" box="[1008,1160,1071,1097]" class="Insecta" family="Nitidulidae" genus="Viettherchnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82E296DBE8E862AFB00FB87" box="[1008,1160,1071,1097]" italics="true" pageId="1" pageNumber="591">Viettherchnus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="5A8A66AFF82E296DB863862AFAF7FB87" authorityName="Erichson" authorityYear="1843" box="[1309,1407,1071,1097]" class="Insecta" family="Nitidulidae" genus="Cyllodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="AFFEC13EF82E296DB863862AFAF7FB87" box="[1309,1407,1071,1097]" italics="true" pageId="1" pageNumber="591">Cyllodes</emphasis>
</taxonomicName>
in smaller size (body length as a rule not exceeding 3.0 mm), and by pronounced sexual dimorphism in the shape of frons, mandibles and scape.
</paragraph>
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