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<document id="D0544B11B77DEED3B5D92239AD2D6ACD" ID-DOI="10.11646/zootaxa.5519.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="13916293" ID-ZooBank="F46DC8A0-EBF2-431A-B486-75595CEA886A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728587797957" checkinUser="plazi" docAuthor="Sawant, Dattaprasad, Joshi, Shantanu, Pawar, Ujwala, Naik, Pratiksha, Khan, Fahim, Nawge, Viraj &amp; Kunte, Krushnamegh" docDate="2024" docId="03D387630C328E4472B0FC7BFB95572D" docLanguage="en" docName="zootaxa.5519.4.6.pdf" docOrigin="Zootaxa 5519 (4)" docSource="http://zoobank.org/f46dc8a0-ebf2-431a-b486-75595cea886a" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Devadatta adii Joshi, Sawant &amp; Kunte" docType="treatment" docVersion="2" lastPageNumber="581" masterDocId="FFEAFF1B0C338E4E7227FFD5FFFE566C" masterDocTitle="Devadatta adii sp. nov. (Odonata: Zygoptera: Devadattidae) from Upper Siang District, Arunachal Pradesh, India" masterLastPageNumber="589" masterPageNumber="571" pageNumber="572" updateTime="1728588231075" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5001B8E800A4A76D87330BA9A2C5996A">Devadatta adii sp. nov. (Odonata: Zygoptera: Devadattidae) from Upper Siang District, Arunachal Pradesh, India</mods:title>
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<mods:namePart id="294D5D3DC966189DEB7AB9FCB6F83C98">Sawant, Dattaprasad</mods:namePart>
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<mods:affiliation id="CCE968B1D77DA76E83A6831D0B26CBB9">Government Medical College and Hospital, Sindhudurg, 416534, Maharashtra, India &amp; National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, Karnataka, India 560065</mods:affiliation>
<mods:nameIdentifier id="4656AB12C63BB469334CD8C9D64D8765" type="email">dattaprasad.101@gmail.com</mods:nameIdentifier>
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<mods:namePart id="018C037E7BD90F35F55CBEBE104CFFF2">Joshi, Shantanu</mods:namePart>
<mods:affiliation id="FF2AA0E857F23A21F1FE1AA5B1800B53">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, Karnataka, India 560065 &amp; Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, USA 72703</mods:affiliation>
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<mods:namePart id="8017C83ECE6B013816CFC7B71A479887">Pawar, Ujwala</mods:namePart>
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<mods:namePart id="4F812B7C07F3D823F0A395E2B9C7C1EB">Naik, Pratiksha</mods:namePart>
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<mods:affiliation id="01D667307BF9043B15CA262D9FF27B88">Sustainability Department, The Tata Power Company Limited, Carnac Bunder, Mumbai, 400009, Maharashtra, India</mods:affiliation>
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<mods:namePart id="2EB4641F31AF8893B421BC91D4A3EFB5">Khan, Fahim</mods:namePart>
<mods:nameIdentifier id="071A4F05ACA3C9EEC4D623F8C6AF80BA" type="ORCID">0000-0003-3473-0960</mods:nameIdentifier>
<mods:affiliation id="9A7DEB71892014C88633D0375D0A83DF">National Centre for Biological Sciences, GKVK Campus, Bellary Rd, Bengaluru, Karnataka, India 560065</mods:affiliation>
<mods:nameIdentifier id="E97984CB580CE3969E8775783947D35D" type="email">fahimkhanj49@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0B8A962CD233240E3DAC24D9A8C768FC">Kunte, Krushnamegh</mods:namePart>
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<treatment id="03D387630C328E4472B0FC7BFB95572D" LSID="urn:lsid:plazi:treatment:03D387630C328E4472B0FC7BFB95572D" httpUri="http://treatment.plazi.org/id/03D387630C328E4472B0FC7BFB95572D" lastPageId="10" lastPageNumber="581" pageId="1" pageNumber="572">
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<taxonomicName id="4C7A4DF60C328E4F72B0FC7BFD8E55A5" authority="Joshi, Sawant &amp; Kunte" authorityName="Joshi, Sawant &amp; Kunte" box="[151,624,942,969]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="572" phylum="Arthropoda" rank="species" species="adii">
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<emphasis id="B90EEA670C328E4F72B0FC7BFEB955A5" bold="true" box="[151,327,942,969]" italics="true" pageId="1" pageNumber="572">Devadatta adii</emphasis>
Joshi, Sawant &amp; Kunte
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(
<figureCitation id="13412AF00C328E4F72B8FC06FEF75581" box="[159,265,979,1005]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1943,1967]" captionStartId-2="5.[151,250,1943,1967]" captionStartId-3="6.[151,250,1849,1873]" captionStartId-4="7.[151,250,1943,1967]" captionStartId-5="8.[151,250,1650,1674]" captionStartId-6="9.[151,250,1943,1967]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[214,1373,181,1919]" captionTargetBox-2="[214,1373,181,1919]" captionTargetBox-3="[232,1356,151,1837]" captionTargetBox-4="[214,1373,181,1919]" captionTargetBox-5="[151,1436,181,1627]" captionTargetBox-6="[214,1373,181,1919]" captionTargetId-0="figure-18@2.[214,1373,181,1919]" captionTargetId-1="figure-17@3.[214,1373,181,1919]" captionTargetId-2="figure-17@5.[214,1373,181,1919]" captionTargetId-3="figure-18@6.[232,1356,151,1837]" captionTargetId-4="figure-17@7.[214,1373,181,1919]" captionTargetId-5="figure-18@8.[151,1436,181,1627]" captionTargetId-6="figure-17@9.[214,1373,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." captionText-1="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." captionText-2="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." captionText-3="FIGURE 4. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (e–f) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (k–l) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (q–r) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm." captionText-4="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." captionText-5="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." captionText-6="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916295" figureDoi-1="http://doi.org/10.5281/zenodo.13916297" figureDoi-2="http://doi.org/10.5281/zenodo.13916299" figureDoi-3="http://doi.org/10.5281/zenodo.13916301" figureDoi-4="http://doi.org/10.5281/zenodo.13916303" figureDoi-5="http://doi.org/10.5281/zenodo.13916305" figureDoi-6="http://doi.org/10.5281/zenodo.13916307" httpUri-0="https://zenodo.org/record/13916295/files/figure.png" httpUri-1="https://zenodo.org/record/13916297/files/figure.png" httpUri-2="https://zenodo.org/record/13916299/files/figure.png" httpUri-3="https://zenodo.org/record/13916301/files/figure.png" httpUri-4="https://zenodo.org/record/13916303/files/figure.png" httpUri-5="https://zenodo.org/record/13916305/files/figure.png" httpUri-6="https://zenodo.org/record/13916307/files/figure.png" pageId="1" pageNumber="572">Figs. 1–7</figureCitation>
)
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.
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(IBC-BR299),
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,
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,
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,
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(
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, 94.957 0 E;
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a.s.l.
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<collectingDate id="EF80E95D0C328E4F7319FBEAFE515235" box="[318,431,1087,1113]" pageId="1" pageNumber="572" value="2022-05-23">23.v.2022</collectingDate>
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,
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,
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, &amp;
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leg.
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<typeStatus id="54C188D70C328E4F72E0FBB6FEBF5211" box="[199,321,1123,1149]" pageId="1" pageNumber="572" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D7CFDFC0C328E4F736AFBB7FE7352EC" box="[333,397,1122,1152]" count="2" pageId="1" pageNumber="572" type="male">2 ♂♂</specimenCount>
</emphasis>
(IBC-BR297, IBC-BR300), location, date of collection and collector same as
<typeStatus id="54C188D70C328E4F7727FBB6FA985211" box="[1280,1382,1123,1149]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
;
<materialsCitation id="3B123C280C328E4F7755FBB7FC7852CD" collectingDate="2022-05-22" pageId="1" pageNumber="572" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F7755FBB7FA6252EC" bold="true" box="[1394,1436,1122,1152]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F7755FBB7FA6252EC" box="[1394,1436,1122,1152]" count="1" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
</emphasis>
(IBC-BR383) same location and collectors as
<typeStatus id="54C188D70C328E4F7084FB52FCF652CD" box="[675,776,1159,1185]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
,
<date id="FFC410B50C328E4F7133FB52FC7852CD" box="[788,902,1159,1185]" pageId="1" pageNumber="572" value="2022-05-22">
<collectingDate id="EF80E95D0C328E4F7133FB52FC7852CD" box="[788,902,1159,1185]" pageId="1" pageNumber="572" value="2022-05-22">22.v.2022</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B123C280C328E4F71B4FB53FE0E52A9" collectingDate="2022-05-29" collectorName="Shantanu Joshi" country="India" elevation="1090" latitude="28.6337" location="Ramsing" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F71B4FB53FC4052C8" bold="true" box="[915,958,1158,1188]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F71B4FB53FC4052C8" box="[915,958,1158,1188]" count="1" pageId="1" pageNumber="572" type="male">1 ♂</specimenCount>
</emphasis>
(IBC-BR451), same location as holotype,
<date id="FFC410B50C328E4F72B0FB7EFEFB52A9" box="[151,261,1195,1221]" pageId="1" pageNumber="572" value="2022-05-29">
<collectingDate id="EF80E95D0C328E4F72B0FB7EFEFB52A9" box="[151,261,1195,1221]" pageId="1" pageNumber="572" value="2022-05-29">29.v.2022</collectingDate>
</date>
<collectorName id="268F53A30C328E4F7328FB7EFE4552A9" box="[271,443,1195,1221]" pageId="1" pageNumber="572">Shantanu Joshi</collectorName>
leg.
</materialsCitation>
;
<emphasis id="B90EEA670C328E4F73D8FB7FFDD552A4" bold="true" box="[511,555,1194,1224]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F73D8FB7FFDD552A4" box="[511,555,1194,1224]" count="1" pageId="1" pageNumber="572" type="female">1 ♀</specimenCount>
</emphasis>
(IBC-BR314) all details are the same as
<typeStatus id="54C188D70C328E4F762AFB7EFB8D52A9" box="[1037,1139,1195,1221]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
.
<materialsCitation id="3B123C280C328E4F76A7FB79FA925361" collectingDate="2024-05-23" collectorName="Krushnamegh Kunte &amp; Fahim Khan &amp; Nitin Kawthankar &amp; Yangchen Lhamo &amp; Yuktha JG" country="India" elevation="460" latitude="28.6642" location="Sirring Nala" municipality="Upper Siang District" pageId="1" pageNumber="572" specimenCount="3" specimenCount-female="3" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F76A7FB79FB4952A4" bold="true" box="[1152,1207,1196,1224]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F76A7FB79FB4952A4" box="[1152,1207,1196,1224]" count="3" pageId="1" pageNumber="572" type="female">3♀♀</specimenCount>
</emphasis>
(IBC-BX307, IBC-BX309, IBC-BX313)
<location id="8EA560AE0C328E4F73B2FB1AFDDA5285" LSID="urn:lsid:plazi:treatment:03D387630C328E4472B0FC7BFB95572D:8EA560AE0C328E4F73B2FB1AFDDA5285" box="[405,548,1231,1257]" country="India" latitude="28.6642" municipality="Upper Siang District" name="Sirring Nala" pageId="1" pageNumber="572" stateProvince="Arunachal Pradesh">Sirring Nala</location>
,
<collectingMunicipality id="6BA1AC0F0C328E4F7016FB1AFCDF5285" box="[561,801,1231,1257]" pageId="1" pageNumber="572">Upper Siang District</collectingMunicipality>
,
<collectingRegion id="49BEF8970C328E4F7109FB1AFBF65285" box="[814,1032,1231,1257]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="F36D76E50C328E4F7632FB1AFBB15285" box="[1045,1103,1231,1257]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
(
<geoCoordinate id="EE4E50B20C328E4F7647FB1AFB235285" box="[1120,1245,1231,1257]" degrees="28.6642" direction="north" orientation="latitude" pageId="1" pageNumber="572" precision="5" value="28.6642">28.6642 N</geoCoordinate>
, 94.994 3 E;
<elevation id="0057D1460C328E4F7755FB1AFF145361" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="572" unit="m" value="460.0">
<quantity id="4C829B900C328E4F7755FB1AFF535361" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="572" unit="m" value="460.0">460 m</quantity>
a.s.l.
</elevation>
),
<date id="FFC410B50C328E4F7325FB26FE8D5361" box="[258,371,1267,1293]" pageId="1" pageNumber="572" value="2024-05-23">
<collectingDate id="EF80E95D0C328E4F7325FB26FE8D5361" box="[258,371,1267,1293]" pageId="1" pageNumber="572" value="2024-05-23">23.v.2024</collectingDate>
</date>
,
<collectorName id="268F53A30C328E4F7358FB26FD925361" box="[383,620,1267,1293]" pageId="1" pageNumber="572">Krushnamegh Kunte</collectorName>
,
<collectorName id="268F53A30C328E4F705EFB26FCF95361" box="[633,775,1267,1293]" pageId="1" pageNumber="572">Fahim Khan</collectorName>
,
<collectorName id="268F53A30C328E4F7133FB26FC1F5361" box="[788,993,1267,1293]" pageId="1" pageNumber="572">Nitin Kawthankar</collectorName>
,
<collectorName id="268F53A30C328E4F71CBFB26FB4B5361" box="[1004,1205,1267,1293]" pageId="1" pageNumber="572">Yangchen Lhamo</collectorName>
,
<collectorName id="268F53A30C328E4F76E6FB26FAC95361" box="[1217,1335,1267,1293]" pageId="1" pageNumber="572">Yuktha JG</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B123C280C328E4F775EFB21FA9C5339" collectingDate="2024-06-02" collectorName="Krushnamegh Kunte &amp; Fahim Khan &amp; Nitin Kawthankar &amp; Yangchen Lhamo &amp; Yuktha JG" country="India" county="Upper Siang District" elevation="360" latitude="28.7639" location="Yinkiong" municipality="Likor Road" pageId="1" pageNumber="572" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<emphasis id="B90EEA670C328E4F775EFB21FA62537C" bold="true" box="[1401,1436,1268,1296]" pageId="1" pageNumber="572">
<specimenCount id="9D7CFDFC0C328E4F775EFB21FA62537C" box="[1401,1436,1268,1296]" count="1" pageId="1" pageNumber="572" type="female">1♀</specimenCount>
</emphasis>
(IBC-BX390)
<location id="8EA560AE0C328E4F731FFAC2FE5E535D" LSID="urn:lsid:plazi:treatment:03D387630C328E4472B0FC7BFB95572D:8EA560AE0C328E4F731FFAC2FE5E535D" box="[312,416,1303,1329]" country="India" county="Upper Siang District" latitude="28.7639" municipality="Likor Road" name="Yinkiong" pageId="1" pageNumber="572" stateProvince="Arunachal Pradesh">Yinkiong</location>
to
<collectingMunicipality id="6BA1AC0F0C328E4F73E4FAC2FDB8535D" box="[451,582,1303,1329]" pageId="1" pageNumber="572">Likor Road</collectingMunicipality>
,
<collectingCounty id="62A44EF90C328E4F7076FAC2FCC4535D" box="[593,826,1303,1329]" pageId="1" pageNumber="572">Upper Siang District</collectingCounty>
,
<collectingRegion id="49BEF8970C328E4F7164FAC2FBE5535D" box="[835,1051,1303,1329]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="F36D76E50C328E4F7602FAC2FBA1535D" box="[1061,1119,1303,1329]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
(
<geoCoordinate id="EE4E50B20C328E4F764BFAC2FB19535D" box="[1132,1255,1303,1329]" degrees="28.7639" direction="north" orientation="latitude" pageId="1" pageNumber="572" precision="5" value="28.7639">28.7639 N</geoCoordinate>
, 94.899 9 E;
<elevation id="0057D1460C328E4F7755FAC2FF175339" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="572" unit="m" value="360.0">
<quantity id="4C829B900C328E4F7755FAC2FF535339" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="572" unit="m" value="360.0">360 m</quantity>
a.s.l.
</elevation>
),
<date id="FFC410B50C328E4F72D8FAEEFE845339" box="[255,378,1339,1365]" pageId="1" pageNumber="572" value="2024-06-02">
<collectingDate id="EF80E95D0C328E4F72D8FAEEFE845339" box="[255,378,1339,1365]" pageId="1" pageNumber="572" value="2024-06-02">02.vi.2024</collectingDate>
</date>
,
<collectorName id="268F53A30C328E4F73A2FAEEFD8E5339" box="[389,624,1339,1365]" pageId="1" pageNumber="572">Krushnamegh Kunte</collectorName>
,
<collectorName id="268F53A30C328E4F705BFAEEFCF75339" box="[636,777,1339,1365]" pageId="1" pageNumber="572">Fahim Khan</collectorName>
,
<collectorName id="268F53A30C328E4F7132FAEEFC1E5339" box="[789,992,1339,1365]" pageId="1" pageNumber="572">Nitin Kawthankar</collectorName>
,
<collectorName id="268F53A30C328E4F71CDFAEEFB4C5339" box="[1002,1202,1339,1365]" pageId="1" pageNumber="572">Yangchen Lhamo</collectorName>
,
<collectorName id="268F53A30C328E4F769BFAEEFACF5339" box="[1212,1329,1339,1365]" pageId="1" pageNumber="572">Yuktha JG</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C328E4F72E0FA8AFE5B5389" pageId="1" pageNumber="572" type="etymology">
<paragraph id="8BC536750C328E4F72E0FA8AFDB553F1" blockId="1.[151,1437,1050,1509]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0FA8AFEB05315" bold="true" box="[199,334,1375,1401]" pageId="1" pageNumber="572">Etymology.</emphasis>
The specific epithet ‘
<emphasis id="B90EEA670C328E4F7018FA8AFD8A5315" box="[575,628,1375,1401]" italics="true" pageId="1" pageNumber="572">adii’</emphasis>
honours the indigenous Adi tribe of the Upper Siang District,
<collectingRegion id="49BEF8970C328E4F770FFA8AFF0D53F1" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="572">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="F36D76E50C328E4F72D9FA56FEC653F1" box="[254,312,1411,1437]" name="India" pageId="1" pageNumber="572">India</collectingCountry>
and neighbouring areas.
</paragraph>
<paragraph id="8BC536750C328E4F72E0FA72FE5B5389" blockId="1.[151,1437,1050,1509]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0FA72FE0C53AD" bold="true" box="[199,498,1446,1473]" pageId="1" pageNumber="572">Proposed English Name.</emphasis>
Golden Grissette: The common English name is given due to the golden yellow pterothorax of the male.
</paragraph>
</subSubSection>
<subSubSection id="C36065FE0C328E4472B0F9C7FB95572D" lastPageId="10" lastPageNumber="581" pageId="1" pageNumber="572" type="description">
<paragraph id="8BC536750C328E4F72B0F9C7FCFD5041" blockId="1.[151,1437,1554,2013]" box="[151,771,1554,1581]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72B0F9C7FD7F5040" bold="true" box="[151,641,1554,1581]" pageId="1" pageNumber="572">
Description of
<typeStatus id="54C188D70C328E4F7360F9C6FE505041" box="[327,430,1555,1581]" pageId="1" pageNumber="572" type="holotype">holotype</typeStatus>
male IBC-BR299
</emphasis>
(
<figureCitation id="13412AF00C328E4F70B7F9C6FD045041" box="[656,762,1555,1581]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1943,1967]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[214,1373,181,1919]" captionTargetId-0="figure-18@2.[214,1373,181,1919]" captionTargetId-1="figure-17@3.[214,1373,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." captionText-1="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916295" figureDoi-1="http://doi.org/10.5281/zenodo.13916297" httpUri-0="https://zenodo.org/record/13916295/files/figure.png" httpUri-1="https://zenodo.org/record/13916297/files/figure.png" pageId="1" pageNumber="572">Figs. 1–2</figureCitation>
)
</paragraph>
<paragraph id="8BC536750C328E4F72B0F9E2FACE50D1" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72B0F9E2FF2A503D" box="[151,212,1591,1617]" italics="true" pageId="1" pageNumber="572">Head</emphasis>
(
<figureCitation id="13412AF00C328E4F72CFF9E2FEC2503D" box="[232,316,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1a</figureCitation>
). Head black, marked as follows: labrum yellowish brown; lower half of mandibles pale blue; labium bluish-yellow except for a round dark area at the center of the upper margin in life; anteclypeus pale blue; postclypeus blackish-brown; frons black. Median ocellus creamy yellow, rest shiny brown; antennae black, upper margin of pedicel pale bluish yellow. Eyes upper half brown, lower half grey in life, darker post-mortem.
</paragraph>
<paragraph id="8BC536750C328E4F72E0F912FED05145" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0F912FE69508D" box="[199,407,1735,1761]" italics="true" pageId="1" pageNumber="572">Thorax. Prothorax</emphasis>
(
<figureCitation id="13412AF00C328E4F7381F912FDEA508D" box="[422,532,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1b, d</figureCitation>
). Anterior and posterior lobes black, middle lobes brown; propleuron dark brown with downward projection at posterior end: posterior margin pruinosed bluish-white in life, dark brown to black post-mortem.
</paragraph>
<paragraph id="8BC536750C328E4F72E0F8E6FDE851B1" blockId="1.[151,1437,1554,2013]" pageId="1" pageNumber="572">
<emphasis id="B90EEA670C328E4F72E0F8E6FEB35121" box="[199,333,1843,1869]" italics="true" pageId="1" pageNumber="572">Pterothorax</emphasis>
(
<figureCitation id="13412AF00C328E4F7347F8E6FE155121" box="[352,491,1843,1869]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="1" pageNumber="572">Fig. 1c, e–f</figureCitation>
). Mesostigmal plates and mesepisternum matt black with greenish metallic sheen; yellow stripe on mesepisternum above mesopleural suture; pterothorax laterally golden yellow in life, pale brown post-mortem; upper part of mesepimeron pale brown, faint brown markings on the interpleural suture and anterior part of metepisternum; mesinfraepisternum brown, metinfraepisternum yellow. Pterothorax ventrally pale yellow, small black spot at posterior edge.
</paragraph>
<caption id="DF0566FD0C318E4C72B0F842FDE3519B" ID-DOI="http://doi.org/10.5281/zenodo.13916295" ID-Zenodo-Dep="13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="2" pageNumber="573" startId="2.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="2" targetType="figure">
<paragraph id="8BC536750C318E4C72B0F842FDE3519B" blockId="2.[151,1437,1943,2039]" pageId="2" pageNumber="573">
<emphasis id="B90EEA670C318E4C72B0F842FEED51C3" bold="true" box="[151,275,1943,1967]" pageId="2" pageNumber="573">FIGURE 1.</emphasis>
<taxonomicName id="4C7A4DF60C318E4C733FF842FE5151C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[280,431,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="573" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C318E4C733FF842FE5151C3" box="[280,431,1943,1967]" italics="true" pageId="2" pageNumber="573">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C318E4C7392F842FDFB51C3" bold="true" box="[437,517,1943,1967]" pageId="2" pageNumber="573">
<taxonomicNameLabel id="A23D571C0C318E4C7392F842FDFB51C3" box="[437,517,1943,1967]" pageId="2" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c).
</paragraph>
</caption>
<caption id="DF0566FD0C308E4D72B0F842FDB2519B" ID-DOI="http://doi.org/10.5281/zenodo.13916297" ID-Zenodo-Dep="13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="3" pageNumber="574" startId="3.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="3" targetType="figure">
<paragraph id="8BC536750C308E4D72B0F842FDB2519B" blockId="3.[151,1437,1943,2039]" pageId="3" pageNumber="574">
<emphasis id="B90EEA670C308E4D72B0F842FEED51C3" bold="true" box="[151,275,1943,1967]" pageId="3" pageNumber="574">FIGURE 2.</emphasis>
<taxonomicName id="4C7A4DF60C308E4D733DF842FE4C51C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[282,434,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="574" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C308E4D733DF842FE4C51C3" box="[282,434,1943,1967]" italics="true" pageId="3" pageNumber="574">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C308E4D739EF842FDF451C3" bold="true" box="[441,522,1943,1967]" pageId="3" pageNumber="574">
<taxonomicNameLabel id="A23D571C0C308E4D739EF842FDF451C3" box="[441,522,1943,1967]" pageId="3" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BC536750C378E4A72E0FF42FD7D56B9" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FF42FF0256DC" box="[199,252,151,176]" italics="true" pageId="4" pageNumber="575">Legs</emphasis>
. Coxae pale brown with pruinosed blue in life, pale brown post-mortem; femur, tibiae brown. Tibiofemoral joint, tarsus and spines dark brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FF0AFF355709" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FF0AFEF55694" box="[199,267,223,248]" italics="true" pageId="4" pageNumber="575">Wings</emphasis>
(
<figureCitation id="13412AF00C378E4A7339FF0AFE725695" box="[286,396,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Fig. 1c–g</figureCitation>
) very faintly tinted with brown; venation black; Pt trapezoid, upper margin is ½ of lower margin, blackish brown, nervures black, covers almost 3 ½ cells below; a slight bulge in the upper wing margin just proximal to pt on anterior side. Arculus at level of Ax 4–5. Ax
<quantity id="4C829B900C378E4A7147FEF2FC72572D" box="[864,908,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="575" unit="in" value="7.0">7 in</quantity>
right FW,
<quantity id="4C829B900C378E4A7620FEF2FBCD572D" box="[1031,1075,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="4" pageNumber="575" unit="in" value="8.0">8 in</quantity>
rest. Px
<quantity id="4C829B900C378E4A76B0FEF3FB05572D" box="[1175,1275,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="8.763" metricValueMax="8.89" metricValueMin="8.636" pageId="4" pageNumber="575" unit="in" value="34.5" valueMax="35.0" valueMin="34.0">34–35 in</quantity>
FW,
<quantity id="4C829B900C378E4A771EFEF3FA62572D" box="[1337,1436,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="4" pageNumber="575" unit="in" value="25.5" valueMax="26.0" valueMin="25.0">25–26 in</quantity>
HW.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FEBAFB1457BD" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FEBAFECF57E5" box="[199,305,367,393]" italics="true" pageId="4" pageNumber="575">Abdomen</emphasis>
(
<figureCitation id="13412AF00C378E4A7366FEBAFE6D57E5" box="[321,403,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Fig. 1c</figureCitation>
). Pale brown, marked with S1 pale brown, minute small black spots on both ends of lower margin of tergite; S2 with a darker brown lateral patch; S3–6 brownish black lateral stripe, fainter on S4–5, S7–10 black, S6–10 bluish pruinescence, S9–10 almost completely covered in bluish pruinescence in life.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FE0EFB8A540D" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FE0EFECF5799" box="[199,305,475,501]" italics="true" pageId="4" pageNumber="575">Genitalia</emphasis>
(
<figureCitation id="13412AF00C378E4A7318FE0EFE525799" box="[319,428,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@3.[214,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="4" pageNumber="575">Fig. 2a–e</figureCitation>
). First and second segments ventrally and dorsally dark brown. Third segment pale yellow. Apical segment translucent medially raised on dorsal side ending with a hood, lamina curved anteriorly and extended laterally, segmented in two parts on both sides, upper part forming a shoulder and ending in a thin filamentous arm slightly thicker at apex. Lower part canine-shaped, rounded, curved outward &amp; laterally.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FDBEFE84549D" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FDBEFE5854E9" box="[199,422,619,645]" italics="true" pageId="4" pageNumber="575">Caudal appendages</emphasis>
(
<figureCitation id="13412AF00C378E4A7392FDBEFDD254E9" box="[437,556,619,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@3.[214,1373,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) lateral view of Segment 2; (b) ventral view of Segment 2; (c–e) genital ligula: (c) ventral view; (d) dorsal view; (e) lateral view; (f–h) caudal appendages: (f) dorsal view; (g) ventral view; (h) lateral view. All scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13916297" httpUri="https://zenodo.org/record/13916297/files/figure.png" pageId="4" pageNumber="575">Figs. 2f–h</figureCitation>
). Cerci black, bent inwards at midpoint making ~90° angle with proximal part; cerci apex rounded and expanded; inner margin of proximal part has bulging and numerous small black spines; cerci length more than S10; paraproct dark brown to black, 1⁄2 of the cerci length, broad at base, apex sharply pointed and curved inwards.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FD2EFB7B5579" blockId="4.[151,1437,151,789]" box="[199,1157,762,789]" pageId="4" pageNumber="575">Measurements. Abdomen without appendages 35–36; HW 29–30; total length 45–46.</paragraph>
<paragraph id="8BC536750C378E4A72B0FC97FE015531" blockId="4.[151,1436,834,1293]" box="[151,511,834,861]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FC97FE8B5531" bold="true" box="[151,373,834,861]" pageId="4" pageNumber="575">Variation in Males</emphasis>
(
<figureCitation id="13412AF00C378E4A73A3FC96FE115531" box="[388,495,835,861]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1943,1967]" captionStartId-1="6.[151,250,1849,1873]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[232,1356,151,1837]" captionTargetId-0="figure-17@5.[214,1373,181,1919]" captionTargetId-1="figure-18@6.[232,1356,151,1837]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." captionText-1="FIGURE 4. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (e–f) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (k–l) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (q–r) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916299" figureDoi-1="http://doi.org/10.5281/zenodo.13916301" httpUri-0="https://zenodo.org/record/13916299/files/figure.png" httpUri-1="https://zenodo.org/record/13916301/files/figure.png" pageId="4" pageNumber="575">Figs. 3–4</figureCitation>
):
</paragraph>
<paragraph id="8BC536750C378E4A72B0FCB2FEFB5235" blockId="4.[151,1436,834,1293]" pageId="4" pageNumber="575">
Though in life all males have similar golden yellow lateral pterothorax, post-mortem it changes to dull brown in some individuals (
<figureCitation id="13412AF00C378E4A7340FC5EFE3D55C9" box="[359,451,907,933]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-18@2.[214,1373,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. Devadatta adii sp. nov. Holotype male (IBC-BR299): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) dorsal view of pterothorax; (f) lateral view of pterothorax; (g) left forewing and hindwing. Scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916295" httpUri="https://zenodo.org/record/13916295/files/figure.png" pageId="4" pageNumber="575">Figs. 1c</figureCitation>
,
<figureCitation id="13412AF00C378E4A73F6FC5EFE0E55C9" box="[465,496,907,933]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">3b</figureCitation>
). IBC-BR297, IBC-BR
<specimenCount id="9D7CFDFC0C378E4A70DDFC5EFC9255C9" box="[762,876,907,933]" count="383" pageId="4" pageNumber="575" type="male">383 males</specimenCount>
have brown upper part of mesepimeron, whereas IBC-BR300 and IBC-BR451 have clear golden yellow thorax without any post-mortem browning (
<figureCitation id="13412AF00C378E4A7738FC7AFA6E55A5" box="[1311,1424,943,969]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1943,1967]" captionStartId-1="6.[151,250,1849,1873]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[232,1356,151,1837]" captionTargetId-0="figure-17@5.[214,1373,181,1919]" captionTargetId-1="figure-18@6.[232,1356,151,1837]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." captionText-1="FIGURE 4. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (e–f) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (k–l) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (q–r) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916299" figureDoi-1="http://doi.org/10.5281/zenodo.13916301" httpUri-0="https://zenodo.org/record/13916299/files/figure.png" httpUri-1="https://zenodo.org/record/13916301/files/figure.png" pageId="4" pageNumber="575">Figs. 3–4</figureCitation>
). IBC-BR
<specimenCount id="9D7CFDFC0C378E4A72D2FC06FEA55581" box="[245,347,979,1005]" count="297" pageId="4" pageNumber="575" type="male">297 male</specimenCount>
has 9–10 Ax in FW and HW and 34 Px in FW, 27 Px in HW. IBC-BR
<specimenCount id="9D7CFDFC0C378E4A7641FC06FB325581" box="[1126,1228,979,1005]" count="300" pageId="4" pageNumber="575" type="male">300 male</specimenCount>
has 7–8 Ax in FW and HW and 31–32 Px in FW and 26–27 Px in HW (
<figureCitation id="13412AF00C378E4A70CCFC22FCCE527D" box="[747,816,1015,1041]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">Fig. 3</figureCitation>
). Similarly, there are 37 Px in FW and 28 Px in HW in IBC-BR
<specimenCount id="9D7CFDFC0C378E4A72D1FBCEFEA25259" box="[246,348,1051,1077]" count="383" pageId="4" pageNumber="575" type="male">383 male</specimenCount>
and 36 Px in FW and 27 Px in HW in IBC-BR
<specimenCount id="9D7CFDFC0C378E4A7154FBCFFC275259" box="[883,985,1050,1077]" count="451" pageId="4" pageNumber="575" type="male">451 male</specimenCount>
(
<figureCitation id="13412AF00C378E4A71CEFBCEFBD05259" box="[1001,1070,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@5.[214,1373,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Devadatta adii sp. nov. Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="4" pageNumber="575">Fig. 3</figureCitation>
). Both these males have 8 Ax in all wings.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FBB6FE8C5211" blockId="4.[151,1436,834,1293]" box="[199,370,1123,1149]" pageId="4" pageNumber="575">Measurements.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB52FB9752CD" blockId="4.[151,1436,834,1293]" box="[199,1129,1159,1185]" pageId="4" pageNumber="575">IBC-BR297: Abdomen without appendages 36–37; HW 32–33; total length 46–47.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB7EFBC152A9" blockId="4.[151,1436,834,1293]" box="[199,1087,1195,1221]" pageId="4" pageNumber="575">IBC-BR300: Abdomen without appendages 31–32; HW 28; total length 41–42.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB1AFB975285" blockId="4.[151,1436,834,1293]" box="[199,1129,1231,1257]" pageId="4" pageNumber="575">IBC-BR383: Abdomen without appendages 31–32; HW 29–30; total length 43–44.</paragraph>
<paragraph id="8BC536750C378E4A72E0FB27FB975361" blockId="4.[151,1436,834,1293]" box="[199,1129,1266,1293]" pageId="4" pageNumber="575">IBC-BR451: Abdomen without appendages 32–33; HW 28–29; total length 41–42.</paragraph>
<paragraph id="8BC536750C378E4A72B0FAEFFCE35339" blockId="4.[151,1437,1338,1941]" box="[151,797,1338,1365]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FAEFFD625338" bold="true" box="[151,668,1338,1365]" pageId="4" pageNumber="575">
Description of
<typeStatus id="54C188D70C378E4A7360FAEEFE4C5339" box="[327,434,1339,1365]" pageId="4" pageNumber="575" type="paratype">paratype</typeStatus>
female IBC-BX390
</emphasis>
(
<figureCitation id="13412AF00C378E4A708CFAEEFCEB5339" box="[683,789,1338,1365]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[151,250,1943,1967]" captionStartId-1="8.[151,250,1650,1674]" captionTargetBox-0="[214,1373,181,1919]" captionTargetBox-1="[151,1436,181,1627]" captionTargetId-0="figure-17@7.[214,1373,181,1919]" captionTargetId-1="figure-18@8.[151,1436,181,1627]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." captionText-1="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.13916303" figureDoi-1="http://doi.org/10.5281/zenodo.13916305" httpUri-0="https://zenodo.org/record/13916303/files/figure.png" httpUri-1="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Figs. 5–6</figureCitation>
)
</paragraph>
<paragraph id="8BC536750C378E4A72B0FA8AFAA253AD" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72B0FA8AFF2A5315" box="[151,212,1375,1401]" italics="true" pageId="4" pageNumber="575">Head</emphasis>
(
<figureCitation id="13412AF00C378E4A72C2FA8AFEA85315" box="[229,342,1374,1401]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5a–b</figureCitation>
). Labrum, labium and mandible pale blue in life, brown post-mortem, labrum dark brown post-mortem; anteclypeus, postclypeus and frons black with blue shine. Rest of head black. Ocelli brown, antennae black. Eyes upper half brown, lower half grey in life, darker post-mortem. Postocular area and occiput black.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FA1EFBFC5389" blockId="4.[151,1437,1338,1941]" box="[199,1026,1482,1509]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FA1EFE665389" box="[199,408,1483,1509]" italics="true" pageId="4" pageNumber="575">Thorax. Prothorax</emphasis>
(
<figureCitation id="13412AF00C378E4A7380FA1EFDEA5389" box="[423,532,1482,1509]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5c–e</figureCitation>
). Similar to male. Slightly pruinosed in life.
</paragraph>
<paragraph id="8BC536750C378E4A72E0FA3AFA98503D" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0FA3AFEB35065" box="[199,333,1519,1545]" italics="true" pageId="4" pageNumber="575">Pterothorax</emphasis>
(
<figureCitation id="13412AF00C378E4A7347FA3AFE305065" box="[352,462,1518,1545]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5f–g</figureCitation>
). Similar to male, except browner laterally. Mesepisternum black with bronze tinge; brownish yellow stripe on mesepisternum just above the mesopleural suture which is greyish blue in life; upper part of mesepimeron brown; faint brown marking on interpleural suture; rest of the lateral pterothorax pale brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F98EFD5150F5" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F98EFF025018" box="[199,252,1627,1652]" italics="true" pageId="4" pageNumber="575">Legs</emphasis>
. Coxae brown. Femur, tibiae brown with bluish pruinescence in life, dark brown post-mortem. Tibiofemoral joint, tarsus and spines blackish brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F976FA8950D1" blockId="4.[151,1437,1338,1941]" box="[199,1399,1698,1725]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F976FEF550D0" box="[199,267,1699,1724]" italics="true" pageId="4" pageNumber="575">Wings</emphasis>
(
<figureCitation id="13412AF00C378E4A733CF976FE9350D1" box="[283,365,1699,1725]" captionStart="FIGURE 6" captionStartId="8.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1627]" captionTargetId="figure-18@8.[151,1436,181,1627]" captionTargetPageId="8" captionText="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi="http://doi.org/10.5281/zenodo.13916305" httpUri="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Fig. 6a</figureCitation>
). Hyaline, similar to male; Pt pale brown. Ax
<quantity id="4C829B900C378E4A7157F976FC6550D1" box="[880,923,1699,1725]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="575" unit="in" value="7.0">7 in</quantity>
all wings. Px
<quantity id="4C829B900C378E4A761DF977FB6350D1" box="[1082,1181,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="8.763" metricValueMax="8.89" metricValueMin="8.636" pageId="4" pageNumber="575" unit="in" value="34.5" valueMax="35.0" valueMin="34.0">34–35 in</quantity>
FW,
<quantity id="4C829B900C378E4A76FEF977FAC250D1" box="[1241,1340,1698,1725]" metricMagnitude="-1" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="4" pageNumber="575" unit="in" value="25.5" valueMax="26.0" valueMin="25.0">25–26 in</quantity>
HW.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F912FE0A5145" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F912FECF508D" box="[199,305,1735,1761]" italics="true" pageId="4" pageNumber="575">Abdomen</emphasis>
(
<figureCitation id="13412AF00C378E4A7364F912FE69508D" box="[323,407,1734,1761]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@7.[214,1373,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Devadatta adii sp. nov. Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c)." figureDoi="http://doi.org/10.5281/zenodo.13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="4" pageNumber="575">Fig. 5c</figureCitation>
). S1–2 dorsally yellowish brown, laterally brown, S3–7 brown with faint, pale yellowish basal rings; dorsum of S6 with a large yellow, rounded, dorsal marking towards posterior margin; S9–10 black; abdomen ventrally pale brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F8E6FD94511D" blockId="4.[151,1437,1338,1941]" pageId="4" pageNumber="575">
<emphasis id="B90EEA670C378E4A72E0F8E6FE575121" box="[199,425,1843,1869]" italics="true" pageId="4" pageNumber="575">Caudal appendages</emphasis>
(
<figureCitation id="13412AF00C378E4A739DF8E6FDD25121" box="[442,556,1843,1869]" captionStart="FIGURE 6" captionStartId="8.[151,250,1650,1674]" captionTargetBox="[151,1436,181,1627]" captionTargetId="figure-18@8.[151,1436,181,1627]" captionTargetPageId="8" captionText="FIGURE 6. Devadatta adii sp. nov. Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view." figureDoi="http://doi.org/10.5281/zenodo.13916305" httpUri="https://zenodo.org/record/13916305/files/figure.png" pageId="4" pageNumber="575">Fig. 6b–d</figureCitation>
). Cerci short, conical, pointed, black, less than length of S10. Paraproct very small, globular, brown; ovipositor brown.
</paragraph>
<paragraph id="8BC536750C378E4A72E0F8AEFB7B51F9" blockId="4.[151,1437,1338,1941]" box="[199,1157,1915,1941]" pageId="4" pageNumber="575">Measurements. Abdomen without appendages 30–31; HW 29–30; total length 39–40.</paragraph>
<caption id="DF0566FD0C368E4B72B0F842FC5451BF" ID-DOI="http://doi.org/10.5281/zenodo.13916299" ID-Zenodo-Dep="13916299" httpUri="https://zenodo.org/record/13916299/files/figure.png" pageId="5" pageNumber="576" startId="5.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="5" targetType="figure">
<paragraph id="8BC536750C368E4B72B0F842FC5451BF" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="576">
<emphasis id="B90EEA670C368E4B72B0F842FEEA51C3" bold="true" box="[151,276,1943,1967]" pageId="5" pageNumber="576">FIGURE 3.</emphasis>
<taxonomicName id="4C7A4DF60C368E4B733CF842FE4D51C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[283,435,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C368E4B733CF842FE4D51C3" box="[283,435,1943,1967]" italics="true" pageId="5" pageNumber="576">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C368E4B739CF842FDF251C3" bold="true" box="[443,524,1943,1967]" pageId="5" pageNumber="576">
<taxonomicNameLabel id="A23D571C0C368E4B739CF842FDF251C3" box="[443,524,1943,1967]" pageId="5" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype male (IBC-BR300): (a) lateral habitus; paratype male (IBC-BR383): (b) lateral habitus; paratype male (IBC-BR451): (c) lateral habitus. All scale bars 5 mm.
</paragraph>
</caption>
<caption id="DF0566FD0C358E4872B0F8ECFD72519D" ID-DOI="http://doi.org/10.5281/zenodo.13916301" ID-Zenodo-Dep="13916301" httpUri="https://zenodo.org/record/13916301/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,250,1849,1873]" targetBox="[232,1356,151,1837]" targetPageId="6" targetType="figure">
<paragraph id="8BC536750C358E4872B0F8ECFD72519D" blockId="6.[151,1436,1849,2034]" pageId="6" pageNumber="577">
<emphasis id="B90EEA670C358E4872B0F8ECFEED513D" bold="true" box="[151,275,1849,1873]" pageId="6" pageNumber="577">FIGURE 4.</emphasis>
<taxonomicName id="4C7A4DF60C358E48733EF8EFFE4E513E" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[281,432,1850,1874]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C358E48733EF8EFFE4E513E" box="[281,432,1850,1874]" italics="true" pageId="6" pageNumber="577">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C358E487391F8EFFDF9513E" bold="true" box="[438,519,1850,1874]" pageId="6" pageNumber="577">
<taxonomicNameLabel id="A23D571C0C358E487391F8EFFDF9513E" box="[438,519,1850,1874]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype male (IBC-BR300): (a) anterior view of head; (b) dorsal view of pterothorax; (c) lateral view of pterothorax; (d) ventral view of segment 2; (e–f) caudal appendages: (e) dorsal view; (f) lateral view; paratype male (IBC-BR383): (g) anterior view of head; (h) dorsal view of pterothorax; (i) lateral view of pterothorax; (j) ventral view of segment 2; (k–l) caudal appendages: (k) dorsal view; (l) lateral view; paratype male (IBC-BR451): (m) anterior view of head; (n) dorsal view of pterothorax; (o) lateral view of pterothorax; (p) ventral view of Segment 2; (q–r) caudal appendages: (q) dorsal view; (r) lateral view. All scale bars 1 mm.
</paragraph>
</caption>
<caption id="DF0566FD0C348E4972B0F842FE2D519B" ID-DOI="http://doi.org/10.5281/zenodo.13916303" ID-Zenodo-Dep="13916303" httpUri="https://zenodo.org/record/13916303/files/figure.png" pageId="7" pageNumber="578" startId="7.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="7" targetType="figure">
<paragraph id="8BC536750C348E4972B0F842FE2D519B" blockId="7.[151,1437,1943,2039]" pageId="7" pageNumber="578">
<emphasis id="B90EEA670C348E4972B0F842FEEC51C3" bold="true" box="[151,274,1943,1967]" pageId="7" pageNumber="578">FIGURE 5.</emphasis>
<taxonomicName id="4C7A4DF60C348E497330F842FE5351C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[279,429,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C348E497330F842FE5351C3" box="[279,429,1943,1967]" italics="true" pageId="7" pageNumber="578">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C348E497395F842FDFC51C3" bold="true" box="[434,514,1943,1967]" pageId="7" pageNumber="578">
<taxonomicNameLabel id="A23D571C0C348E497395F842FDFC51C3" box="[434,514,1943,1967]" pageId="7" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC-BR390): (a) anterior view of head; (b) dorsal view of head; (c) lateral habitus; (d) dorsal view of prothorax; (e) lateral view of prothorax; (f) dorsal view of pterothorax; (g) lateral view of pterothorax. All scale bars 1 mm except (c).
</paragraph>
</caption>
<caption id="DF0566FD0C3B8E4672B0F9A7FCCD50C3" ID-DOI="http://doi.org/10.5281/zenodo.13916305" ID-Zenodo-Dep="13916305" httpUri="https://zenodo.org/record/13916305/files/figure.png" pageId="8" pageNumber="579" startId="8.[151,250,1650,1674]" targetBox="[151,1436,181,1627]" targetPageId="8" targetType="figure">
<paragraph id="8BC536750C3B8E4672B0F9A7FCCD50C3" blockId="8.[151,1437,1650,1711]" pageId="8" pageNumber="579">
<emphasis id="B90EEA670C3B8E4672B0F9A7FEED50E7" bold="true" box="[151,275,1650,1675]" pageId="8" pageNumber="579">FIGURE 6.</emphasis>
<taxonomicName id="4C7A4DF60C3B8E46733EF9A6FE4E50E7" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[281,432,1651,1675]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3B8E46733EF9A6FE4E50E7" box="[281,432,1651,1675]" italics="true" pageId="8" pageNumber="579">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3B8E467391F9A6FDF950E7" bold="true" box="[438,519,1651,1675]" pageId="8" pageNumber="579">
<taxonomicNameLabel id="A23D571C0C3B8E467391F9A6FDF950E7" box="[438,519,1651,1675]" pageId="8" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC- BX390): (a) left forewing and hindwing; (b–d) caudal appendages, scale bar 1 mm: (b) dorsal view; (c) ventral view; (d) lateral view.
</paragraph>
</caption>
<paragraph id="8BC536750C3B8E4672B0F908FE0D509B" blockId="8.[151,1436,1757,1999]" box="[151,499,1757,1783]" pageId="8" pageNumber="579">
<emphasis id="B90EEA670C3B8E4672B0F908FE71509B" bold="true" box="[151,399,1757,1783]" pageId="8" pageNumber="579">Variation in Females</emphasis>
(
<figureCitation id="13412AF00C3B8E4673B9F908FE1C509B" box="[414,482,1757,1783]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
):
</paragraph>
<paragraph id="8BC536750C3B8E4672B0F8D4FF4251A3" blockId="8.[151,1436,1757,1999]" pageId="8" pageNumber="579">
Stripe on mesepisternum is usually greyish blue in life, it has variable post-mortem colours (
<figureCitation id="13412AF00C3B8E467690F8D4FB035177" box="[1207,1277,1793,1819]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
). IBC-BX307 and IBC-BX
<specimenCount id="9D7CFDFC0C3B8E467300F8F0FE4E5153" box="[295,432,1829,1855]" count="309" pageId="8" pageNumber="579" type="female">309 females</specimenCount>
have pale bluish yellow stripe on mesepisternum, whereas IBC-BR314 and IBC-BX313 have brownish yellow stripe on mesepisternum (
<figureCitation id="13412AF00C3B8E467091F89CFD04510F" box="[694,762,1865,1891]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-17@9.[214,1373,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Devadatta adii sp. nov. Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="8" pageNumber="579">Fig. 7</figureCitation>
). IBC-BR
<specimenCount id="9D7CFDFC0C3B8E46714AF89CFC17510F" box="[877,1001,1865,1891]" count="314" pageId="8" pageNumber="579" type="female">314 female</specimenCount>
has 7–8 Ax in all the wings and 31–33 Px in FW and 23–27 Px in HW. IBC-BX
<specimenCount id="9D7CFDFC0C3B8E46707AF8B8FD2751EB" box="[605,729,1901,1927]" count="307" pageId="8" pageNumber="579" type="female">307 female</specimenCount>
has clearer wings and 7–8 Ax and 33–36, whereas IBC-BX
<specimenCount id="9D7CFDFC0C3B8E467754F8B8FF1D51C7" count="309" pageId="8" pageNumber="579" type="female">309 female</specimenCount>
has faintly brown enfumed wings and 7–10 Ax and 34–36 Px. IBC-BX
<specimenCount id="9D7CFDFC0C3B8E46762AF844FB7451C7" box="[1037,1162,1937,1963]" count="313" pageId="8" pageNumber="579" type="female">313 female</specimenCount>
has 8–10 Ax and 34–37 Px.
</paragraph>
<caption id="DF0566FD0C3A8E4772B0F842FBF551BF" ID-DOI="http://doi.org/10.5281/zenodo.13916307" ID-Zenodo-Dep="13916307" httpUri="https://zenodo.org/record/13916307/files/figure.png" pageId="9" pageNumber="580" startId="9.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="9" targetType="figure">
<paragraph id="8BC536750C3A8E4772B0F842FBF551BF" blockId="9.[151,1436,1943,2004]" pageId="9" pageNumber="580">
<emphasis id="B90EEA670C3A8E4772B0F842FEEB51C3" bold="true" box="[151,277,1943,1967]" pageId="9" pageNumber="580">FIGURE 7.</emphasis>
<taxonomicName id="4C7A4DF60C3A8E477339F842FE4951C3" authorityName="Sawant &amp; Joshi &amp; Pawar &amp; Naik &amp; Khan &amp; Nawge &amp; Kunte" authorityYear="2024" box="[286,439,1943,1967]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="580" phylum="Arthropoda" rank="species" species="adii" status="sp. nov.">
<emphasis id="B90EEA670C3A8E477339F842FE4951C3" box="[286,439,1943,1967]" italics="true" pageId="9" pageNumber="580">Devadatta adii</emphasis>
</taxonomicName>
<emphasis id="B90EEA670C3A8E4773E7F842FDED51C3" bold="true" box="[448,531,1943,1967]" pageId="9" pageNumber="580">
<taxonomicNameLabel id="A23D571C0C3A8E4773E7F842FDED51C3" box="[448,531,1943,1967]" pageId="9" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype female (IBC-BX307): (a) lateral habitus; paratype female (IBC- BX309): (b) lateral habitus; paratype female (IBC- BX313): (c) lateral habitus. All scale bars 5 mm.
</paragraph>
</caption>
<paragraph id="8BC536750C398E4472E0FF42FE8C56DD" blockId="10.[199,1131,151,321]" box="[199,370,151,177]" pageId="10" pageNumber="581">Measurements.</paragraph>
<paragraph id="8BC536750C398E4472E0FF6EFB9756B9" blockId="10.[199,1131,151,321]" box="[199,1129,186,213]" pageId="10" pageNumber="581">IBC-BR314: Abdomen without appendages 28–29; HW 24–25; total length 34–35.</paragraph>
<paragraph id="8BC536750C398E4472E0FF0AFB955695" blockId="10.[199,1131,151,321]" box="[199,1131,223,249]" pageId="10" pageNumber="581">IBC-BX307: Abdomen without appendages 31–32; HW 31–32; total length 40–41.</paragraph>
<paragraph id="8BC536750C398E4472E0FED6FB955771" blockId="10.[199,1131,151,321]" box="[199,1131,259,285]" pageId="10" pageNumber="581">IBC-BX309: Abdomen without appendages 32–32; HW 30–31; total length 40–41.</paragraph>
<paragraph id="8BC536750C398E4472E0FEF2FB95572D" blockId="10.[199,1131,151,321]" box="[199,1131,295,321]" pageId="10" pageNumber="581">IBC-BX313: Abdomen without appendages 31–32; HW 31–22; total length 41–42.</paragraph>
</subSubSection>
</treatment>
</document>