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<document id="4F164EEDD7DC3C3658015252D9049C31" ID-DOI="10.11646/phytotaxa.642.2.8" ID-ISSN="1179-3163" ID-Zenodo-Dep="13214331" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1722602268773" checkinUser="felipe" docAuthor="Cornejo, Xavier &amp; Weigend, Maximillian" docDate="2024" docId="2C226759EB72FFC4FF56BC416341FB61" docLanguage="en" docName="phytotaxa.642.2.8.pdf" docOrigin="Phytotaxa 642 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.642.2.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Nasa ojedae Cornejo &amp; Weigend, sp. nov." docType="treatment" docVersion="3" lastPageNumber="206" masterDocId="D01B1F21EB73FFC1FFDEB975676AFFAD" masterDocTitle="Nasa ojedae (Loasaceae): A new species from the montane forests of Reserva El Corazón, western Andes of Ecuador" masterLastPageNumber="207" masterPageNumber="201" pageNumber="202" updateTime="1725884228673" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="06D840C9789758436F634258D92E74F6">Nasa ojedae (Loasaceae): A new species from the montane forests of Reserva El Corazón, western Andes of Ecuador</mods:title>
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<mods:namePart id="6B2E7E95435D80C34962FD5F853D9A66">Cornejo, Xavier</mods:namePart>
<mods:affiliation id="D7C82C022626B9A98CE0B2E5C60AC9E9">Herbario GUAY, Departamento de Botánica, Facultad de Ciencias Naturales, Universidad de Guayaquil, Av. Raúl Gómez Lince s. n. y</mods:affiliation>
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<mods:namePart id="58731FEF2C8F3FD1C1A31F0AB54E495D">Weigend, Maximillian</mods:namePart>
<mods:nameIdentifier id="00535798EEDA7D1E62909D2D910505DF" type="ORCID">0000-0003-0813-6650</mods:nameIdentifier>
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<mods:typeOfResource id="02A5DD4B2BF00304DEA3E54AA41B409D">text</mods:typeOfResource>
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<mods:title id="5560CB42502963EE11037A491445FA9E">Phytotaxa</mods:title>
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<mods:part id="217C621E60A4F13CBEF5B30540F790C9">
<mods:date id="91203A5260B694F4DB76B8BD86159E47">2024</mods:date>
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<mods:number id="E57EA72997CD36F574A06FCEE3CF4EC9">2024-03-28</mods:number>
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<mods:number id="69209EDA6DCC386EE3847EA5806A4833">642</mods:number>
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<mods:classification id="12FB40E0B0FC0B787A32AAFE08E70E08">journal article</mods:classification>
<mods:identifier id="2AE7F458D60FBF329C4A0D548DF1D6BD" type="DOI">10.11646/phytotaxa.642.2.8</mods:identifier>
<mods:identifier id="29A69A58E65FB5C3679DF2F1904FFADE" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="32BEF2B0078FA258F6D1EEB142AED709" type="Zenodo-Dep">13214331</mods:identifier>
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<treatment id="2C226759EB72FFC4FF56BC416341FB61" LSID="urn:lsid:plazi:treatment:2C226759EB72FFC4FF56BC416341FB61" httpUri="http://treatment.plazi.org/id/2C226759EB72FFC4FF56BC416341FB61" lastPageId="5" lastPageNumber="206" pageId="1" pageNumber="202">
<subSubSection id="EC9185C4EB72FFC0FF56BC41658AFAE3" box="[136,736,1332,1359]" pageId="1" pageNumber="202" type="nomenclature">
<paragraph id="A434D64FEB72FFC0FF56BC41658AFAE3" blockId="1.[136,736,1332,1359]" box="[136,736,1332,1359]" pageId="1" pageNumber="202">
<heading id="FF7C6123EB72FFC0FF56BC41658AFAE3" box="[136,736,1332,1359]" fontSize="11" level="2" pageId="1" pageNumber="202" reason="3">
<taxonomicName id="638BADCCEB72FFC0FF56BC416695FAE3" authority="Cornejo &amp; Weigend" authorityName="Cornejo &amp; Weigend" box="[136,511,1332,1359]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="1" pageNumber="202" phylum="Tracheophyta" rank="species" species="ojedae" status="sp. nov.">
<emphasis id="96FF0A5DEB72FFC0FF56BC41667FFAE2" bold="true" box="[136,277,1332,1359]" italics="true" pageId="1" pageNumber="202">Nasa ojedae</emphasis>
Cornejo &amp; Weigend
</taxonomicName>
,
<emphasis id="96FF0A5DEB72FFC0FDD4BC406535FAE3" box="[522,607,1333,1358]" italics="true" pageId="1" pageNumber="202">
<taxonomicNameLabel id="8DCCB726EB72FFC0FDD4BC406535FAE3" box="[522,607,1333,1358]" pageId="1" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3CB0CACAEB72FFC0FDB0BC4165BDFAE3" box="[622,727,1332,1358]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[136,234,2063,2085]" captionStartId-1="3.[136,234,1919,1941]" captionStartId-2="4.[136,234,1992,2014]" captionTargetBox-0="[153,1433,531,2034]" captionTargetBox-1="[197,1391,190,1895]" captionTargetBox-2="[170,1417,190,1968]" captionTargetId-0="figure-206@2.[151,1435,530,2036]" captionTargetId-1="figure-23@3.[197,1391,190,1895]" captionTargetId-2="figure-29@4.[170,1417,190,1968]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Distribution map of Nasa ojedae (*)." captionText-1="FIGURE 2. Nasa ojedae. A. Flower at anthesis, lateral-adaxial view; B. Flower at anthesis, lateral view, inner floral nectary at lower right corner; C. Two detached petals and epipetalous fascicles of stamens attached to base, inner view; D. Fruit nearly to maturity with persistent sepals, lateral view; seeds preserved in alcohol at upper left corner and fruit transversal section exhibiting 5 parietal placentae at lower right corner; E. A mature basal leaf blade, adaxial view; F. Habitat, an Andean forested ravine (A–F, based on X. Cornejo, J. Josse 10058, the type). Photos by Xavier Cornejo." captionText-2="FIGURE 3. Nasa ojedae. A. Whitish indument; B. Arachnoid trichomes; C–D. Stinging setae, close up of glandular base at upper right corner; E. Leaf blade beneath; F. Glochidiate trichomes (A–F, based on X. Cornejo &amp; J. Josse 10058, the type). Photos by Xavier Cornejo." figureDoi-0="http://doi.org/10.5281/zenodo.13214333" figureDoi-1="http://doi.org/10.5281/zenodo.13214337" figureDoi-2="http://doi.org/10.5281/zenodo.13214339" httpUri-0="https://zenodo.org/record/13214333/files/figure.png" httpUri-1="https://zenodo.org/record/13214337/files/figure.png" httpUri-2="https://zenodo.org/record/13214339/files/figure.png" pageId="1" pageNumber="202">Figs. 1–3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EC9185C4EB72FFC0FF56BCF563EEFA70" pageId="1" pageNumber="202" type="diagnosis">
<paragraph id="A434D64FEB72FFC0FF56BCF563EEFA70" blockId="1.[136,1451,1408,1574]" pageId="1" pageNumber="202">
<emphasis id="96FF0A5DEB72FFC0FF56BCF563EEFA70" italics="true" pageId="1" pageNumber="202">
This new species of
<taxonomicName id="638BADCCEB72FFC0FE97BCF56610FA38" authorityName="Weigend" authorityYear="2006" box="[329,378,1408,1429]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="1" pageNumber="202" phylum="Tracheophyta" rank="genus">Nasa</taxonomicName>
is a biennial herb with yellow corollas and white arachnoid indument, similar to
<taxonomicName id="638BADCCEB72FFC0FB50BCF56665FA14" authority="(Werderm.) Weigend" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Werderm." baseAuthorityYear="1935" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="1" pageNumber="202" phylum="Tracheophyta" rank="species" species="profundilobata">N. profundilobata (Werderm.) Weigend</taxonomicName>
but differing from it and from all species of the
<taxonomicName id="638BADCCEB72FFC0FD0ABCD16412FA14" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Kunth" baseAuthorityYear="1808" box="[724,888,1444,1465]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="1" pageNumber="202" phylum="Tracheophyta" rank="species" species="ranunculifolia">N. ranunculifolia</taxonomicName>
group by the ovaries with five (vs. three) placentae and the fruits opening by five (vs. three) apical valves, and the seeds with alveolate (vs. reticulate) sculpture.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="EC9185C4EB72FFC0FF56BC996578F988" pageId="1" pageNumber="202" type="materials_examined">
<paragraph id="A434D64FEB72FFC0FF56BC996578F988" blockId="1.[136,1451,1408,1574]" pageId="1" pageNumber="202">
<materialsCitation id="14E3DC12EB72FFC0FF56BC996612F988" collectingDate="2022-08-20" collectionCode="GUAY" collectorName="X. Cornejo &amp; J. Josse" country="Ecuador" elevation="2600" latitude="-2.05" location="Reserva El Corazon" longLatPrecision="1307" longitude="-78.9" municipality="Type" pageId="1" pageNumber="202" specimenCount="1" stateProvince="Chimborazo" typeStatus="holotype">
<collectingMunicipality id="44504C35EB72FFC0FF56BC9967D0F9AC" box="[136,186,1516,1537]" pageId="1" pageNumber="202">Type</collectingMunicipality>
:—
<collectingCountry id="DC9C96DFEB72FFC0FF08BC996620F9AC" box="[214,330,1516,1537]" name="Ecuador" pageId="1" pageNumber="202">ECUADOR</collectingCountry>
.
<collectingRegion id="664F18ADEB72FFC0FE86BC9966B9F9AF" box="[344,467,1516,1538]" country="Ecuador" name="Chimborazo" pageId="1" pageNumber="202">Chimborazo</collectingRegion>
:
<location id="A1548094EB72FFC0FE02BC9965CBF9AC" LSID="urn:lsid:plazi:treatment:2C226759EB72FFC4FF56BC416341FB61:A1548094EB72FFC0FE02BC9965CBF9AC" box="[476,673,1516,1538]" country="Ecuador" latitude="-2.05" longLatPrecision="1307" longitude="-78.9" municipality="Type" name="Reserva El Corazon" pageId="1" pageNumber="202" stateProvince="Chimborazo">Reserva El Corazón</location>
,
<geoCoordinate id="C1BFB088EB72FFC0FD72BC99659CF9AC" box="[684,758,1516,1537]" degrees="2" direction="south" minutes="03" orientation="latitude" pageId="1" pageNumber="202" precision="925" value="-2.05">2°03’ S</geoCoordinate>
,
<geoCoordinate id="C1BFB088EB72FFC0FCDCBC996435F9AC" box="[770,863,1516,1538]" degrees="78" direction="west" minutes="54" orientation="longitude" pageId="1" pageNumber="202" precision="925" value="-78.9">78°54’ W</geoCoordinate>
,
<quantity id="63737BAAEB72FFC0FCB4BC9964DCF9AC" box="[874,950,1516,1538]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="202" unit="m" value="2600.0">
<elevation id="2FA6317CEB72FFC0FCB4BC9964DCF9AC" box="[874,950,1516,1538]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="202" unit="m" value="2600.0">2600 m</elevation>
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,
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<collectingDate id="C0710967EB72FFC0FC1FBC99632AF9AC" box="[961,1088,1516,1537]" pageId="1" pageNumber="202" value="2022-08-20">20 Aug 2022</collectingDate>
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(fl, fr),
<emphasis id="96FF0A5DEB72FFC0FB4EBC9962C1F9AC" box="[1168,1451,1516,1537]" italics="true" pageId="1" pageNumber="202">
<collectorName id="097EB399EB72FFC0FB4EBC996391F9AC" box="[1168,1275,1516,1537]" pageId="1" pageNumber="202">X. Cornejo</collectorName>
&amp;
<collectorName id="097EB399EB72FFC0FAC2BC996202F9AC" box="[1308,1384,1516,1537]" pageId="1" pageNumber="202">J. Josse</collectorName>
10058
</emphasis>
(
<typeStatus id="7B3068EDEB72FFC0FF1ABF656677F98B" box="[196,285,1552,1574]" pageId="1" pageNumber="202" type="holotype">holotype</typeStatus>
:
<collectionCode id="C29A4E8AEB72FFC0FEF8BF65661BF988" box="[294,369,1552,1573]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15759" lsid="urn:lsid:biocol.org:col:15759" name="Universidad de Guayaquil" pageId="1" pageNumber="202" type="Herbarium">GUAY</collectionCode>
!;
</materialsCitation>
<materialsCitation id="14E3DC12EB72FFC0FEA0BF656567F988" box="[382,525,1552,1574]" collectingDate="2022-08-20" collectionCode="QCA" collectorName="X. Cornejo &amp; J. Josse" country="Ecuador" elevation="2600" latitude="-2.05" location="Reserva El Corazon" longLatPrecision="1307" longitude="-78.9" municipality="Type" pageId="1" pageNumber="202" specimenCount="1" stateProvince="Chimborazo" typeStatus="isotype">
<typeStatus id="7B3068EDEB72FFC0FEA0BF6566A9F98B" box="[382,451,1552,1574]" pageId="1" pageNumber="202" type="isotype">isotype</typeStatus>
<collectionCode id="C29A4E8AEB72FFC0FE17BF656696F988" box="[457,508,1552,1573]" collectionName="QCA" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15650" lsid="urn:lsid:biocol.org:col:15650" name="Pontificia Universidad Católica del Ecuador" pageId="1" pageNumber="202" type="Herbarium">QCA</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EC9185C4EB72FFC3FF56BF21637FFE51" lastPageId="2" lastPageNumber="203" pageId="1" pageNumber="202" type="description">
<paragraph id="A434D64FEB72FFC3FF56BF21637FFE51" blockId="1.[136,1452,1620,2080]" lastBlockId="2.[136,1452,158,509]" lastPageId="2" lastPageNumber="203" pageId="1" pageNumber="202">
<emphasis id="96FF0A5DEB72FFC0FF56BF21660BF9C3" box="[136,353,1620,1646]" italics="true" pageId="1" pageNumber="202">Stout biennial herb</emphasis>
c.
<quantity id="63737BAAEB72FFC0FE58BF2166D9F9C2" box="[390,435,1620,1647]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="202" unit="m" value="2.0">2 m</quantity>
tall, stem to c.
<quantity id="63737BAAEB72FFC0FDB8BF2165DEF9C2" box="[614,692,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="202" unit="cm" value="1.5">1.5 cm</quantity>
thick; plant covered with brown stinging setae,
<quantity id="63737BAAEB72FFC0FB05BF216253F9C2" box="[1243,1337,1620,1647]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="202" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, and shorter glochidiate trichomes,
<quantity id="63737BAAEB72FFC0FE3EBF0D650DF93E" box="[480,615,1656,1683]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="202" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
long, and initially flat-filiform, twisted hyaline trichomes, then forming a white arachnoid indument.
<emphasis id="96FF0A5DEB72FFC0FE69BFE965D9F91B" box="[439,691,1692,1718]" italics="true" pageId="1" pageNumber="202">Terminal young shoots</emphasis>
with a thick arachnoid white indument of foamy appearance.
<emphasis id="96FF0A5DEB72FFC0FA80BFE862C6F91B" box="[1374,1452,1693,1718]" italics="true" pageId="1" pageNumber="202">Leaves</emphasis>
alternate; petioles to c.
<quantity id="63737BAAEB72FFC0FE4BBFB566B6F976" box="[405,476,1728,1755]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="202" unit="cm" value="17.0">17 cm</quantity>
long, suberect to ascending, spreading at the lower half of the plant, inserted at base of lamina and sometimes peltate to c.
<quantity id="63737BAAEB72FFC0FDE6BF91651BF952" box="[568,625,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="202" unit="cm" value="1.0">1 cm</quantity>
from leaf margin at base, densely covered with erect hyaline glochidiate trichomes,
<quantity id="63737BAAEB72FFC0FED8BE7D66E5F88E" box="[262,399,1800,1827]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="202" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
long, and translucent light brown setae, c.
<quantity id="63737BAAEB72FFC0FCA3BE7D64D5F88E" box="[893,959,1800,1827]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="202" unit="mm" value="3.0">3 mm</quantity>
long; lamina widely ovate to pentagonous, 14–40 ×
<quantity id="63737BAAEB72FFC0FF32BE596635F8EA" box="[236,351,1836,1863]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" metricValueMax="4.0" metricValueMin="1.2" pageId="1" pageNumber="202" unit="cm" value="26.0" valueMax="40.0" valueMin="12.0">12–40 cm</quantity>
, the base truncate to cordate, the sinus up to c.
<quantity id="63737BAAEB72FFC0FCAFBE5964C3F8EA" box="[881,937,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="202" unit="cm" value="3.0">3 cm</quantity>
deep, margins dentate to subpalmately lobed, the lobes 3 on each side, broadly triangular, 2–5 ×
<quantity id="63737BAAEB72FFC0FD1EBE256441F8C6" box="[704,811,1872,1899]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="7.0" metricValueMin="2.5" pageId="1" pageNumber="202" unit="cm" value="4.75" valueMax="7.0" valueMin="2.5">2.5–7 cm</quantity>
, adaxial and abaxial surfaces covered with setae c.
<quantity id="63737BAAEB72FFC0FAB5BE2562C6F8C6" box="[1387,1452,1872,1899]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="202" unit="mm" value="3.0">3 mm</quantity>
long and glochidiate trichomes,
<quantity id="63737BAAEB72FFC0FE29BE0165EAF822" box="[503,640,1908,1935]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="202" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
long; leaf venation palmate, with 5–7 costae arising near base, tertiary veins sulcate above (fresh), all veins prominent beneath (fresh).
<emphasis id="96FF0A5DEB72FFC0FCB0BEED64A0F81F" box="[878,970,1944,1970]" italics="true" pageId="1" pageNumber="202">Flowers</emphasis>
supraxillary in terminal forked, frondose, erect monochasia
<quantity id="63737BAAEB72FFC0FE8DBEC966A9F87A" box="[339,451,1980,2007]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" metricValueMax="4.0" metricValueMin="1.2" pageId="1" pageNumber="202" unit="cm" value="26.0" valueMax="40.0" valueMin="12.0">12–40 cm</quantity>
long, bearing 3 to 5 suberect flowers, bracts absent. Pedicels c.
<quantity id="63737BAAEB72FFC0FB48BEC963A7F87A" box="[1174,1229,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="202" unit="cm" value="5.0">5 cm</quantity>
long, setose. Calyx cup-shaped, c. 1.3 ×
<quantity id="63737BAAEB72FFC0FEAABE9566A9F856" box="[372,451,2016,2043]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="202" unit="cm" value="1.7">1.7 cm</quantity>
, very densely light brown setose and with erect white glochidiate trichomes, lobes erect, ovate to ovate-lanceolate,
<quantity id="63737BAAEB72FFC0FE6DB1716558F7B2" box="[435,562,2052,2079]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="1" pageNumber="202" unit="cm" value="2.1" valueMax="2.5" valueMin="1.7">1.7–2.5 cm</quantity>
long, fused for ⅓ to ½ of the length, the base setose, abaxially faintly veined and with densely arranged white glochidiate trichomes, adaxially with abundant white glochidiate trichomes, the apex acuminate, the tip glandulose. Corolla bowl-shaped, petals 5, imbricate at anthesis, obovate, 3.5–3.8 ×
<quantity id="63737BAAEB71FFC3FAB2B9B767C7FEAC" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="2" pageNumber="203" unit="cm" value="2.3" valueMax="2.6" valueMin="2.0">2–2.6 cm</quantity>
, the apex rounded, sulfur-yellow, esetulose, abaxially with short (
<quantity id="63737BAAEB71FFC3FC7BB993635AFEAC" box="[933,1072,230,257]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="203" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
long), erect trichomes, adaxially glabrous. Nectar scales with rectangular back, c.
<quantity id="63737BAAEB71FFC3FD69B87F6590FE88" box="[695,762,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="203" unit="mm" value="5.0">5 mm</quantity>
long, with 2 to 4 obovoid to oblong sacs near base on back, 3–4 ×
<quantity id="63737BAAEB71FFC3FF11B85B663DFEE4" box="[207,343,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="2" pageNumber="203" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
, scale neck thickened, 3-lobed, laterally protracted into two stiffly erect, vertical wings 6–8 ×
<quantity id="63737BAAEB71FFC3FAB2B85B67DDFEC0" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="203" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
. Staminodia 2 per scale, linear,
<quantity id="63737BAAEB71FFC3FDC1B82765F3FEC0" box="[543,665,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="203" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
long, densely and shortly pilose. Stamens numerous, in 5 epipetalous fascicles of 10 to 20 each, filaments
<quantity id="63737BAAEB71FFC3FDEFB80365C6FE3C" box="[561,684,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.2" metricValueMin="1.3" pageId="2" pageNumber="203" unit="mm" value="17.5" valueMax="22.0" valueMin="13.0">13–22 mm</quantity>
long, anthers 1.5–2 ×
<quantity id="63737BAAEB71FFC3FC6CB8036342FE3C" box="[946,1064,374,401]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="203" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
, brownish or light brown to grey. Style stout, c.
<quantity id="63737BAAEB71FFC3FEF7B8EF661CFE18" box="[297,374,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="203" unit="cm" value="2.2">2.2 cm</quantity>
long in carpellate phase. Ovary inferior, unilocular, with 5 parietal placentae.
<emphasis id="96FF0A5DEB71FFC3FB36B8EF6225FE19" box="[1256,1359,410,436]" italics="true" pageId="2" pageNumber="203">Capsules</emphasis>
broadly elliptic, c. 3.2 ×
<quantity id="63737BAAEB71FFC3FE9BB8CB66FCFE74" box="[325,406,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="203" unit="cm" value="2.3">2.3 cm</quantity>
, with abundant, white glochidiate trichomes, crowned by a persistent calyx, opening with 5 apical valves, pedicel ca.
<quantity id="63737BAAEB71FFC3FE78B8976698FE50" box="[422,498,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="203" unit="cm" value="5.5">5.5 cm</quantity>
long. Seeds obovate, 1.2–1.5 ×
<quantity id="63737BAAEB71FFC3FC84B89764F7FE50" box="[858,925,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="203" unit="mm" value="1.0">1 mm</quantity>
, alveolate.
</paragraph>
</subSubSection>
<caption id="F0F486C7EB71FFC3FF56B17A6501F788" ID-DOI="http://doi.org/10.5281/zenodo.13214333" ID-Zenodo-Dep="13214333" box="[136,619,2063,2086]" httpUri="https://zenodo.org/record/13214333/files/figure.png" pageId="2" pageNumber="203" startId="2.[136,234,2063,2085]" targetBox="[153,1433,531,2034]" targetPageId="2" targetType="figure">
<paragraph id="A434D64FEB71FFC3FF56B17A6501F788" blockId="2.[136,619,2063,2086]" box="[136,619,2063,2086]" pageId="2" pageNumber="203">
<emphasis id="96FF0A5DEB71FFC3FF56B17A6669F788" bold="true" box="[136,259,2063,2085]" pageId="2" pageNumber="203">FIGURE 1.</emphasis>
Distribution map of
<taxonomicName id="638BADCCEB71FFC3FE12B1656529F788" box="[460,579,2064,2085]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="2" pageNumber="203" phylum="Tracheophyta" rank="species" species="ojedae">
<emphasis id="96FF0A5DEB71FFC3FE12B1656529F788" box="[460,579,2064,2085]" italics="true" pageId="2" pageNumber="203">Nasa ojedae</emphasis>
</taxonomicName>
(*).
</paragraph>
</caption>
<caption id="F0F486C7EB70FFC2FF56BE0A6558F788" ID-DOI="http://doi.org/10.5281/zenodo.13214337" ID-Zenodo-Dep="13214337" httpUri="https://zenodo.org/record/13214337/files/figure.png" pageId="3" pageNumber="204" startId="3.[136,234,1919,1941]" targetBox="[197,1391,190,1895]" targetPageId="3" targetType="figure">
<paragraph id="A434D64FEB70FFC2FF56BE0A6558F788" blockId="3.[136,1452,1919,2085]" pageId="3" pageNumber="204">
<emphasis id="96FF0A5DEB70FFC2FF56BE0A666EF839" bold="true" box="[136,260,1919,1941]" pageId="3" pageNumber="204">FIGURE 2.</emphasis>
<emphasis id="96FF0A5DEB70FFC2FED2BEF566E6F839" box="[268,396,1919,1941]" italics="true" pageId="3" pageNumber="204">
<taxonomicName id="638BADCCEB70FFC2FED2BEF566E2F839" box="[268,392,1919,1941]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="species" species="ojedae">Nasa ojedae</taxonomicName>
.
</emphasis>
A. Flower at anthesis, lateral-adaxial view; B. Flower at anthesis, lateral view, inner floral nectary at lower right corner; C. Two detached petals and epipetalous fascicles of stamens attached to base, inner view; D. Fruit nearly to maturity with persistent sepals, lateral view; seeds preserved in alcohol at upper left corner and fruit transversal section exhibiting 5 parietal placentae at lower right corner; E. A mature basal leaf blade, adaxial view; F. Habitat, an Andean forested ravine (A–F, based on
<emphasis id="96FF0A5DEB70FFC2FB34BE9967A1F789" italics="true" pageId="3" pageNumber="204">X. Cornejo, J. Josse 10058,</emphasis>
the type). Photos by Xavier Cornejo.
</paragraph>
</caption>
<caption id="F0F486C7EB77FFC5FF56BEBD67B7F78B" ID-DOI="http://doi.org/10.5281/zenodo.13214339" ID-Zenodo-Dep="13214339" httpUri="https://zenodo.org/record/13214339/files/figure.png" pageId="4" pageNumber="205" startId="4.[136,234,1992,2014]" targetBox="[170,1417,190,1968]" targetPageId="4" targetType="figure">
<paragraph id="A434D64FEB77FFC5FF56BEBD67B7F78B" blockId="4.[136,1452,1992,2086]" pageId="4" pageNumber="205">
<emphasis id="96FF0A5DEB77FFC5FF56BEBD666CF870" bold="true" box="[136,262,1992,2014]" pageId="4" pageNumber="205">FIGURE 3.</emphasis>
<emphasis id="96FF0A5DEB77FFC5FED1BEBC66FBF870" box="[271,401,1992,2014]" italics="true" pageId="4" pageNumber="205">
<taxonomicName id="638BADCCEB77FFC5FED1BEBC66E7F870" box="[271,397,1992,2014]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="4" pageNumber="205" phylum="Tracheophyta" rank="species" species="ojedae">Nasa ojedae</taxonomicName>
.
</emphasis>
A. Whitish indument; B. Arachnoid trichomes; C–D. Stinging setae, close up of glandular base at upper right corner; E. Leaf blade beneath; F. Glochidiate trichomes (A–F, based on
<emphasis id="96FF0A5DEB77FFC5FCA7BE9863F3F7AC" box="[889,1177,2028,2050]" italics="true" pageId="4" pageNumber="205">X. Cornejo &amp; J. Josse 10058,</emphasis>
the type). Photos by Xavier Cornejo.
</paragraph>
</caption>
<subSubSection id="EC9185C4EB76FFC4FF63B9EB630FFD20" pageId="5" pageNumber="206" type="discussion">
<paragraph id="A434D64FEB76FFC4FF63B9EB630FFD20" blockId="5.[136,1452,158,1228]" pageId="5" pageNumber="206">
Discussion:—
<taxonomicName id="638BADCCEB76FFC4FE82B9EA6682FF15" box="[348,488,158,184]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ojedae">
<emphasis id="96FF0A5DEB76FFC4FE82B9EA6682FF15" box="[348,488,158,184]" italics="true" pageId="5" pageNumber="206">Nasa ojedae</emphasis>
</taxonomicName>
is clearly most similar to
<taxonomicName id="638BADCCEB76FFC4FCD3B9EA649EFF15" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Werderm." baseAuthorityYear="1935" box="[781,1012,158,184]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="profundilobata">
<emphasis id="96FF0A5DEB76FFC4FCD3B9EA649EFF15" box="[781,1012,158,184]" italics="true" pageId="5" pageNumber="206">Nasa profundilobata</emphasis>
</taxonomicName>
and can be assigned to the informal
<taxonomicName id="638BADCCEB76FFC4FA4CB9EA6642FF71" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Kunth" baseAuthorityYear="1808" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ranunculifolia">
<emphasis id="96FF0A5DEB76FFC4FA4CB9EA6642FF71" italics="true" pageId="5" pageNumber="206">N. ranunculifolia</emphasis>
</taxonomicName>
group within
<taxonomicName id="638BADCCEB76FFC4FE14B9B665A5FF71" authorityName="sensu Urban &amp; Gilg" authorityYear="1900" box="[458,719,194,221]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="series" series="Grandiflorae">
<emphasis id="96FF0A5DEB76FFC4FE14B9B6656EFF71" box="[458,516,195,220]" italics="true" pageId="5" pageNumber="206">Nasa</emphasis>
ser.
<emphasis id="96FF0A5DEB76FFC4FDE5B9B765A5FF71" box="[571,719,194,220]" italics="true" pageId="5" pageNumber="206">Grandiflorae</emphasis>
</taxonomicName>
. This is a morphologically rather well-defined group of biennial to perennial herbs or small shrubs, restricted to the higher elevations of the Andean mountain range, generally found well above
<quantity id="63737BAAEB76FFC4FED2B87F6608FE88" box="[268,354,266,293]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="206" unit="m" value="3000.0">3000 m</quantity>
ranging to nearly
<quantity id="63737BAAEB76FFC4FDEBB87F65E1FE88" box="[565,651,266,293]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="206" unit="m" value="5000.0">5000 m</quantity>
in elevation (
<bibRefCitation id="C01AABBEEB76FFC4FCF6B87F6313FE89" author="Weigend, M. &amp; Gottschling, M." box="[808,1145,266,292]" pageId="5" pageNumber="206" pagination="120 - 142" refId="ref3385" refString="Weigend, M. &amp; Gottschling, M. (2006) Evolution of funnel-revolver flowers and ornithophily in Nasa (Loasaceae). Plant Biology 8: 120 - 142. https: // doi. org / 10.1055 / s- 2005 - 873034" type="journal article" year="2006">Weigend &amp; Gottschling 2006</bibRefCitation>
,
<bibRefCitation id="C01AABBEEB76FFC4FB58B87F6241FE89" author="Henning, T. &amp; Rodriguez Rodriguez, E. F. &amp; Weigend, M." box="[1158,1323,266,292]" pageId="5" pageNumber="206" pagination="47 - 93" refId="ref2866" refString="Henning, T., Rodriguez Rodriguez, E. F. &amp; Weigend, M. (2011) A revision of Nasa ranunculifolia group (Nasa ser. Grandiflorae pro parte) Loasaceae. Botanical Journal of the Linnaean Society 167: 47 - 93. https: // doi. org / 10.1111 / j. 1095 - 8339.2011.01164. x" type="journal article" year="2011">Henning 2011</bibRefCitation>
). The
<taxonomicName id="638BADCCEB76FFC4FAACB87E6642FEE5" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Kunth" baseAuthorityYear="1808" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ranunculifolia">
<emphasis id="96FF0A5DEB76FFC4FAACB87E6642FEE5" italics="true" pageId="5" pageNumber="206">Nasa ranunculifolia</emphasis>
</taxonomicName>
group is also characterized by palmate to peltate leaves and stiff erect lateral wings on the nectar scales (
<bibRefCitation id="C01AABBEEB76FFC4FF4EB8276671FEC1" author="Urban, I." box="[144,283,338,364]" pageId="5" pageNumber="206" pagination="364 - 388" refId="ref3157" refString="Urban, I. (1886) Die Bestaubungseinrichtungen der Loasaceen. Jahrbuch des Botanischen Gartens Berlin 4: 364 - 388." type="journal article" year="1886">Urban 1886</bibRefCitation>
,
<bibRefCitation id="C01AABBEEB76FFC4FEF6B8276560FEC1" author="Urban, I. &amp; Gilg, W." box="[296,522,338,364]" pageId="5" pageNumber="206" pagination="1 - 368" refId="ref3180" refString="Urban, I. &amp; Gilg, W. (1900) Monographia Loasacearum. Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosum 76: 1 - 368." type="journal article" year="1900">Urban &amp; Gilg 1900</bibRefCitation>
,
<bibRefCitation id="C01AABBEEB76FFC4FDC8B8276594FEC1" author="Henning, T. &amp; Rodriguez Rodriguez, E. F. &amp; Weigend, M." box="[534,766,338,365]" pageId="5" pageNumber="206" pagination="47 - 93" refId="ref2866" refString="Henning, T., Rodriguez Rodriguez, E. F. &amp; Weigend, M. (2011) A revision of Nasa ranunculifolia group (Nasa ser. Grandiflorae pro parte) Loasaceae. Botanical Journal of the Linnaean Society 167: 47 - 93. https: // doi. org / 10.1111 / j. 1095 - 8339.2011.01164. x" type="journal article" year="2011">
Henning
<emphasis id="96FF0A5DEB76FFC4FDA1B82665D8FEC1" box="[639,690,338,364]" italics="true" pageId="5" pageNumber="206">et al</emphasis>
. 2011
</bibRefCitation>
). However,
<taxonomicName id="638BADCCEB76FFC4FC59B826637EFEC1" box="[903,1044,338,364]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ojedae">
<emphasis id="96FF0A5DEB76FFC4FC59B826637EFEC1" box="[903,1044,338,364]" italics="true" pageId="5" pageNumber="206">Nasa ojedae</emphasis>
</taxonomicName>
is unique and differs from all other species in the
<taxonomicName id="638BADCCEB76FFC4FEF3B802669AFE3D" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Kunth" baseAuthorityYear="1808" box="[301,496,374,400]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ranunculifolia">
<emphasis id="96FF0A5DEB76FFC4FEF3B802669AFE3D" box="[301,496,374,400]" italics="true" pageId="5" pageNumber="206">N. ranunculifolia</emphasis>
</taxonomicName>
group by having ovaries with 5 parietal placentae and fruits apically opening by 5 valves (vs. flowers with 3 parietal placentae and fruits opening by 3 valves). The group comprises several biennial species:
<taxonomicName id="638BADCCEB76FFC4FF39B8CA6514FE75" authority="(Weigend 2006: 77)" baseAuthorityName="Weigend" baseAuthorityPageNumber="77" baseAuthorityYear="2006" box="[231,638,446,472]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis id="96FF0A5DEB76FFC4FF39B8CA66FDFE75" box="[231,407,446,472]" italics="true" pageId="5" pageNumber="206">Nasa magnifica</emphasis>
(Weigend 2006: 77)
</taxonomicName>
, with pale pink flowers and red-and-white nectar scales from the western Andean slope of
<collectingCountry id="DC9C96DFEB76FFC4FE92B89666E8FE50" box="[332,386,483,509]" name="Peru" pageId="5" pageNumber="206">Peru</collectingCountry>
,
<taxonomicName id="638BADCCEB76FFC4FE51B8966499FE51" authority="T. Henning &amp; Weigend (2011: 61)" authorityName="T. Henning &amp; Weigend" authorityPageNumber="61" authorityYear="2011" box="[399,1011,482,509]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="tulipadiaboli">
<emphasis id="96FF0A5DEB76FFC4FE51B8966508FE51" box="[399,610,482,508]" italics="true" pageId="5" pageNumber="206">Nasa tulipadiaboli</emphasis>
T.
<bibRefCitation id="C01AABBEEB76FFC4FD57B8976499FE51" author="Henning, T. &amp; Rodriguez Rodriguez, E. F. &amp; Weigend, M." box="[649,1011,482,508]" pageId="5" pageNumber="206" pagination="47 - 93" refId="ref2866" refString="Henning, T., Rodriguez Rodriguez, E. F. &amp; Weigend, M. (2011) A revision of Nasa ranunculifolia group (Nasa ser. Grandiflorae pro parte) Loasaceae. Botanical Journal of the Linnaean Society 167: 47 - 93. https: // doi. org / 10.1111 / j. 1095 - 8339.2011.01164. x" type="journal article" year="2011">Henning &amp; Weigend (2011: 61)</bibRefCitation>
</taxonomicName>
with yellow flowers with very narrow and pointy petals from the eastern Andes of
<collectingCountry id="DC9C96DFEB76FFC4FDA4BB7265DAFD8C" box="[634,688,519,545]" name="Peru" pageId="5" pageNumber="206">Peru</collectingCountry>
, and
<taxonomicName id="638BADCCEB76FFC4FD32BB7264BEFD8D" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Werderm." baseAuthorityYear="1935" box="[748,980,518,544]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="profundilobata">
<emphasis id="96FF0A5DEB76FFC4FD32BB7264BEFD8D" box="[748,980,518,544]" italics="true" pageId="5" pageNumber="206">Nasa profundilobata</emphasis>
</taxonomicName>
from
<collectingCountry id="DC9C96DFEB76FFC4FBC4BB736310FD8D" box="[1050,1146,518,544]" name="Ecuador" pageId="5" pageNumber="206">Ecuador</collectingCountry>
. Only
<taxonomicName id="638BADCCEB76FFC4FB1DBB7262C1FD8D" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Werderm." baseAuthorityYear="1935" box="[1219,1451,518,544]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="profundilobata">
<emphasis id="96FF0A5DEB76FFC4FB1DBB7262C1FD8D" box="[1219,1451,518,544]" italics="true" pageId="5" pageNumber="206">Nasa profundilobata</emphasis>
</taxonomicName>
shares the presence of rounded, yellow petals and arachnoid pubescence, but differs from
<taxonomicName id="638BADCCEB76FFC4FB52BB5E6393FDE9" box="[1164,1273,554,580]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ojedae">
<emphasis id="96FF0A5DEB76FFC4FB52BB5E6393FDE9" box="[1164,1273,554,580]" italics="true" pageId="5" pageNumber="206">N. ojedae</emphasis>
</taxonomicName>
in ovaries with three placentae and fruits with three valves, like all other species of this group. Also, the leaves of
<taxonomicName id="638BADCCEB76FFC4FB1BBB3A62C1FDC5" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Werderm." baseAuthorityYear="1935" box="[1221,1451,590,616]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="profundilobata">
<emphasis id="96FF0A5DEB76FFC4FB1BBB3A62C1FDC5" box="[1221,1451,590,616]" italics="true" pageId="5" pageNumber="206">Nasa profundilobata</emphasis>
</taxonomicName>
are very deeply lobed with lobes longer than wide, not widely triangular as in
<taxonomicName id="638BADCCEB76FFC4FC2DBB066334FD21" box="[1011,1118,626,652]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ojedae">
<emphasis id="96FF0A5DEB76FFC4FC2DBB066334FD21" box="[1011,1118,626,652]" italics="true" pageId="5" pageNumber="206">N. ojedae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EC9185C4EB76FFC4FF63BBE36475FD79" pageId="5" pageNumber="206" type="etymology">
<paragraph id="A434D64FEB76FFC4FF63BBE36475FD79" blockId="5.[136,1452,158,1228]" pageId="5" pageNumber="206">Etymology:—The epithet honors Mr. Luis Ojeda and his family, by whose initiative and love for nature a patch of Andean forests on the property in El Corazón is conserved.</paragraph>
</subSubSection>
<subSubSection id="EC9185C4EB76FFC4FF63BBAB668DFD55" box="[189,487,734,761]" pageId="5" pageNumber="206" type="vernacular_names">
<paragraph id="A434D64FEB76FFC4FF63BBAB668DFD55" blockId="5.[136,1452,158,1228]" box="[189,487,734,761]" pageId="5" pageNumber="206">Common names:—Ortiga.</paragraph>
</subSubSection>
<subSubSection id="EC9185C4EB76FFC4FF63BA776206FBB4" pageId="5" pageNumber="206" type="biology_ecology">
<paragraph id="A434D64FEB76FFC4FF63BA7765DFFC7D" blockId="5.[136,1452,158,1228]" pageId="5" pageNumber="206">
Habitat and distribution:—
<collectionCode id="C29A4E8AEB76FFC4FE33BA77656BFCB1" box="[493,513,770,796]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="5" pageNumber="206" type="Herbarium">A</collectionCode>
robust, biennial herb in montane Andean wet forests, persisting in forested ravines and secondary forests on well-drained soils, c.
<quantity id="63737BAAEB76FFC4FD7ABA536591FCEC" box="[676,763,806,833]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="206" unit="m" value="2600.0">2600 m</quantity>
elevation at the Reserva El Corazón, at southernmost tip of northern Andean block. In the
<typeStatus id="7B3068EDEB76FFC4FE07BA3E6563FCC8" box="[473,521,843,869]" pageId="5" pageNumber="206">type</typeStatus>
locality,
<taxonomicName id="638BADCCEB76FFC4FDB1BA3E6593FCC9" box="[623,761,842,868]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ojedae">
<emphasis id="96FF0A5DEB76FFC4FDB1BA3E6593FCC9" box="[623,761,842,868]" italics="true" pageId="5" pageNumber="206">Nasa ojedae</emphasis>
</taxonomicName>
is flowering at same time with the sympatric
<taxonomicName id="638BADCCEB76FFC4FB2EBA3E661FFC25" authorityName="Weigend" authorityYear="2006" baseAuthorityName="Urban &amp; Gilg" baseAuthorityYear="1900" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="subSpecies" species="humboldtiana" subSpecies="humboldtiana">
<emphasis id="96FF0A5DEB76FFC4FB2EBA3E62C6FCC9" box="[1264,1452,842,868]" italics="true" pageId="5" pageNumber="206">N. humboldtiana</emphasis>
subsp.
<emphasis id="96FF0A5DEB76FFC4FF07BA1B661FFC25" box="[217,373,878,904]" italics="true" pageId="5" pageNumber="206">humboldtiana</emphasis>
</taxonomicName>
, a species recently rediscovered by the senior author 162 years after the
<typeStatus id="7B3068EDEB76FFC4FB13BA1A6397FC24" box="[1229,1277,879,905]" pageId="5" pageNumber="206">type</typeStatus>
collection was gathered by Richard Spruce, in
<date id="D035F08FEB76FFC4FE34BAE76530FC00" box="[490,602,914,941]" pageId="5" pageNumber="206" value="1859-08">Aug 1859</date>
(
<bibRefCitation id="C01AABBEEB76FFC4FDB7BAE76425FC01" author="Henning, T. &amp; Acuna-Castillo, R. &amp; Cornejo, X. &amp; Gonzales, P. &amp; Segovia, E. &amp; Wong Sato, A. &amp; Weigend, M." box="[617,847,914,941]" pageId="5" pageNumber="206" pagination="1 - 19" refId="ref2779" refString="Henning, T., Acuna-Castillo, R., Cornejo, X., Gonzales, P., Segovia, E., Wong Sato, A. &amp; Weigend, M. (2023) When the absence of evidence is not the evidence of absence: Nasa (Loasaceae) rediscoveries from Peru and Ecuador, and the contribution of community science networks. Phytokeys 229: 1 - 19. https: // doi. org / 10.3897 / phytokeys. 229.100082" type="journal article" year="2023">
Henning
<emphasis id="96FF0A5DEB76FFC4FD0CBAE6646EFC01" box="[722,772,914,940]" italics="true" pageId="5" pageNumber="206">et al</emphasis>
. 2023
</bibRefCitation>
). At present, both endangered taxa are only known by less than 5 individuals at the Reserva El Corazón.
</paragraph>
<paragraph id="A434D64FEB76FFC4FF63BAAF6206FBB4" blockId="5.[136,1452,158,1228]" pageId="5" pageNumber="206">
Phenology:
<emphasis id="96FF0A5DEB76FFC4FEE3BAAE663CFC59" box="[317,342,987,1012]" italics="true" pageId="5" pageNumber="206">—</emphasis>
Based on field work carried out from
<date id="D035F08FEB76FFC4FCD5BAAF64A8FC59" box="[779,962,986,1012]" pageId="5" pageNumber="206" value="2020-11">November 2020</date>
to
<date id="D035F08FEB76FFC4FC34BAAF63CEFC59" box="[1002,1188,986,1012]" pageId="5" pageNumber="206" value="2023-11">November 2023</date>
, flowers and immature fruits were observed on the same individual from August to November, the mature fruits were found in January.
</paragraph>
</subSubSection>
<subSubSection id="EC9185C4EB76FFC4FF63BD576341FB61" pageId="5" pageNumber="206" type="conservation">
<paragraph id="A434D64FEB76FFC4FF63BD576341FB61" blockId="5.[136,1452,158,1228]" pageId="5" pageNumber="206">
Preliminary conservation status:—
<taxonomicName id="638BADCCEB76FFC4FD9DBD5665A4FB91" box="[579,718,1058,1084]" class="Magnoliopsida" family="Loasaceae" genus="Nasa" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="ojedae">
<emphasis id="96FF0A5DEB76FFC4FD9DBD5665A4FB91" box="[579,718,1058,1084]" italics="true" pageId="5" pageNumber="206">Nasa ojedae</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="7B3068EDEB76FFC4FC3CBD566378FB90" box="[994,1042,1059,1085]" pageId="5" pageNumber="206">type</typeStatus>
, which was collected in the Reserva El Corazón, in the Andean province of
<collectingRegion id="664F18ADEB76FFC4FD91BD3365B7FBCD" box="[591,733,1094,1120]" country="Ecuador" name="Chimborazo" pageId="5" pageNumber="206">Chimborazo</collectingRegion>
. As the area of occupancy of the species is (dramatically) less than
<quantity id="63737BAAEB76FFC4FF1FBD1F667DFB29" box="[193,279,1130,1156]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="206" unit="km" value="100.0">100 km</quantity>
<superScript id="53FE7B07EB76FFC4FEC9BD1F6675FBD5" attach="left" box="[279,287,1130,1144]" fontSize="6" pageId="5" pageNumber="206">2</superScript>
and due to the steady deforestation and fragmentation in the region, with only small patches of native vegetation remaining, surrounded by crops (pers. obs.), it is suggested that this new species should be assigned as Critically Endangered (
<collectionCode id="C29A4E8AEB76FFC4FE53BDC766DEFB61" box="[397,436,1202,1228]" country="Costa Rica" lsid="urn:lsid:biocol.org:col:15652" name="Museo Nacional de Costa Rica" pageId="5" pageNumber="206" type="Herbarium">CR</collectionCode>
) following the
<collectionCode id="C29A4E8AEB76FFC4FDBCBDC765CCFB61" box="[610,678,1202,1228]" pageId="5" pageNumber="206">IUCN</collectionCode>
criterion
<collectionCode id="C29A4E8AEB76FFC4FCCDBDC76443FB61" box="[787,809,1202,1228]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="5" pageNumber="206" type="Herbarium">B</collectionCode>
1 ab(iii) (
<bibRefCitation id="C01AABBEEB76FFC4FC50BDC76375FB61" author="IUCN Standards &amp; Petitions Committee" box="[910,1055,1202,1228]" pageId="5" pageNumber="206" refId="ref2990" refString="IUCN Standards and Petitions Committee (2022) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15. Prepared by the Standards and Petitions Committee. Downloadable from http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 1 July 2022)" type="book" year="2022">IUCN, 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>