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Order
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<paragraph pageId="0" pageNumber="165">Diagnosis.</paragraph>
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Ventral transverse musculature absent, less glandular structures on body surface. Grasping spines gently or abruptly curved (
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). Collarette absent, present or small that is almost absent. Intestinal diverticula absent or present. One or two rows of teeth, teeth-rows arranged in comb or fan shape. One paired or two paired lateral fins with or without rayless zone (
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).

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<treatment ID-GBIF-Taxon="191645974" LSID="urn:lsid:plazi:treatment:2E574A6B47265CBBB7E28CD4BEED44DB" httpUri="http://treatment.plazi.org/id/2E574A6B47265CBBB7E28CD4BEED44DB" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="2E574A6B-4726-5CBB-B7E2-8CD4BEED44DB" authority="(Philippi, 1844) s. l." authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">Siciliaria calcarae (Philippi, 1844) s. l.</taxonomicName>
<treatment id="2E574A6B47265CBBB7E28CD4BEED44DB" ID-GBIF-Taxon="191645974" LSID="urn:lsid:plazi:treatment:2E574A6B47265CBBB7E28CD4BEED44DB" httpUri="http://treatment.plazi.org/id/2E574A6B47265CBBB7E28CD4BEED44DB" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Clausiliidae" scope_infraOrder="Clausilioidei" scope_order="Stylommatophora" scope_phylum="Mollusca" scope_superFamily="Clausilioidea">
<subSubSection id="EEA7FE1D8DD3D6F3DD07D36D291A9495" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="6A47D3167E9103629EEC3EC1A02A727A" pageId="0" pageNumber="1">
<taxonomicName id="A1602ACD02AE797C267F2A4AE53FA7EA" ID-CoL="8THDN" LSID="2E574A6B-4726-5CBB-B7E2-8CD4BEED44DB" authority="(Philippi, 1844) s. l." authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">Siciliaria calcarae (Philippi, 1844) s. l.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="106F4C6490AEFFCD82C47084DB6BBE70" pageId="0" pageNumber="1" type="remarks">
<paragraph id="B9EB394D2DE0241A1611D272D279D172" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="0FDF1B16AAC21D8555E29A3461FEB4B1" pageId="0" pageNumber="1">
In the COI tree two subclades of
<taxonomicName authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
<taxonomicName id="2508C016B51DC2E2B3164A868547E5CF" authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis id="8221C6E6C5C744C1938B6816F7D287ED" italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
</taxonomicName>
s. l. appear together with
<taxonomicName lsidName="S. leucophryna" pageId="0" pageNumber="1" rank="species" species="leucophryna">
<emphasis italics="true" pageId="0" pageNumber="1">S. leucophryna</emphasis>
<taxonomicName id="7523DBE2BA3A0C54FC50F3E505F0EA7C" lsidName="S. leucophryna" pageId="0" pageNumber="1" rank="species" species="leucophryna">
<emphasis id="5B20E08B96560F030F36F9EF21A3074E" italics="true" pageId="0" pageNumber="1">S. leucophryna</emphasis>
</taxonomicName>
in an unresolved trichotomy. The subspecies of
<taxonomicName authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
<taxonomicName id="94CC188137EFC0415D0836E31DACE5DC" authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis id="0DE0D679A9F7C97FEFF6E3B045E755BE" italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
</taxonomicName>
are distributed within two haplogroups, one (haplogroup 1) comprising the nominate subspecies as well as
<taxonomicName lsidName="S. c. subsp. belliemii" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="belliemii">
<emphasis italics="true" pageId="0" pageNumber="1">S. c. belliemii</emphasis>
<taxonomicName id="275296F6187935C463073F65F997E669" lsidName="S. c. subsp. belliemii" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="belliemii">
<emphasis id="73C14C14707A232FE3092FE289D24BCE" italics="true" pageId="0" pageNumber="1">S. c. belliemii</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="S. c. subsp. borgettensis" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="borgettensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. c. borgettensis</emphasis>
<taxonomicName id="1B626092F489E505DD3AD12381E77236" lsidName="S. c. subsp. borgettensis" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="borgettensis">
<emphasis id="74E6628B4BDA0A763914C28E6C3231DC" italics="true" pageId="0" pageNumber="1">S. c. borgettensis</emphasis>
</taxonomicName>
ssp. nov.,
<taxonomicName lsidName="S. c. subsp. orlandoi" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="orlandoi">
<emphasis italics="true" pageId="0" pageNumber="1">S. c. orlandoi</emphasis>
<taxonomicName id="FEAC59ED06365A7A4811D9ED1FF1DBD4" lsidName="S. c. subsp. orlandoi" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="orlandoi">
<emphasis id="7963DE3EA5443508B90AABD0DC6959E6" italics="true" pageId="0" pageNumber="1">S. c. orlandoi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="S. c. subsp. parajatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="parajatinensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. c. parajatinensis</emphasis>
<taxonomicName id="80F5BF019548D08EC843CE2660E45D79" lsidName="S. c. subsp. parajatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="parajatinensis">
<emphasis id="8FC9C26E12D6D515569E01ABADCE58E9" italics="true" pageId="0" pageNumber="1">S. c. parajatinensis</emphasis>
</taxonomicName>
ssp. nov. and the other one (haplogroup 2) consisting of
<taxonomicName lsidName="S. c. subsp. cruenta" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="cruenta">
<emphasis italics="true" pageId="0" pageNumber="1">S. c. cruenta</emphasis>
<taxonomicName id="98358FDCF510935975D011B39C9649B3" lsidName="S. c. subsp. cruenta" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="cruenta">
<emphasis id="79572FF649D8721BCB91ABE3747688B6" italics="true" pageId="0" pageNumber="1">S. c. cruenta</emphasis>
</taxonomicName>
ssp. nov. and
<taxonomicName lsidName="S. c. subsp. jatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="jatinensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. c. jatinensis</emphasis>
<taxonomicName id="419604FB70474A1E5EFE8A54F507ADA3" lsidName="S. c. subsp. jatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="jatinensis">
<emphasis id="CB640E13F01B61B884E404D0892B8C69" italics="true" pageId="0" pageNumber="1">S. c. jatinensis</emphasis>
</taxonomicName>
ssp. nov. Unfortunately,
<taxonomicName lsidName="S. parajatinensis" pageId="0" pageNumber="1" rank="species" species="parajatinensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. parajatinensis</emphasis>
<taxonomicName id="EA9627E4D2D2FABF12156000E73F0DB1" lsidName="S. parajatinensis" pageId="0" pageNumber="1" rank="species" species="parajatinensis">
<emphasis id="B0059C74EBF75F67B0AF068F0D5BFCA3" italics="true" pageId="0" pageNumber="1">S. parajatinensis</emphasis>
</taxonomicName>
could not be included into the ITS2 analysis. Yet, given the low distances among all representatives of
<taxonomicName lsidName="S. calcarae" pageId="0" pageNumber="1" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae</emphasis>
<taxonomicName id="7D208E341DA9B71E9D77AD647C68ADAB" lsidName="S. calcarae" pageId="0" pageNumber="1" rank="species" species="calcarae">
<emphasis id="062E33A7097B0C9DCECCC4DC229A1B9A" italics="true" pageId="0" pageNumber="1">S. calcarae</emphasis>
</taxonomicName>
in the ITS2 tree (and the resulting bad resolution), this data set does not tell us much concerning these relationships. Thus, presently the close relationship between
<taxonomicName lsidName="S. c. subsp. orlandoi" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="orlandoi">
<emphasis italics="true" pageId="0" pageNumber="1">S. c. orlandoi</emphasis>
<taxonomicName id="9457285584FA01673B78B337A2989755" lsidName="S. c. subsp. orlandoi" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="orlandoi">
<emphasis id="0BA49446253C0153C59A6850CF083EAA" italics="true" pageId="0" pageNumber="1">S. c. orlandoi</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="S. c. subsp. parajatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="parajatinensis">
<emphasis italics="true" pageId="0" pageNumber="1">S. c. parajatinensis</emphasis>
<taxonomicName id="908F1D7633BF4A08D6D7E996DE074B89" lsidName="S. c. subsp. parajatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="c." subSpecies="parajatinensis">
<emphasis id="6E8088A2DDA9D80604002F7C0040A495" italics="true" pageId="0" pageNumber="1">S. c. parajatinensis</emphasis>
</taxonomicName>
hast to rely on mt data only. Concerning the anatomy of the genital organs, shell morphology, distribution, ecology and haplogroup of
<taxonomicName authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
<taxonomicName id="B4D9FDD7D67E9DE08F4479717D9AEF9F" authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis id="1A8B7130D5AC8E46ACFEDD50146F87D2" italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
</taxonomicName>
, two general types can be distinguished for each category.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="genital organs">
<paragraph pageId="0" pageNumber="1">Genital organs.</paragraph>
<paragraph pageId="0" pageNumber="1">Two main anatomical types of male (penial) internal sculpturing are found. The first type presents clear longitudinal pleats that run from the origin of penial pseudopapilla down to (or almost to) the atrium. These longitudinal pleats can be either (almost) completely smooth (Monte Belliemi) or more or less markedly fringed (Calatubo, Romitello, Ficuzza, Jato city park, Monte Kumeta, and Castelluzzo). The second type presents transverse and strong smooth pleats (Bonifato, Gibilmesi, and Visicari).</paragraph>
<subSubSection id="AD64FFF5CCA79A9ED1799354B5138465" pageId="0" pageNumber="1" type="genital organs">
<paragraph id="A2B00AE121A9EF2B4CC689AE04332255" pageId="0" pageNumber="1">Genital organs.</paragraph>
<paragraph id="F9B7C80552599EBF18559D59861B1BAD" pageId="0" pageNumber="1">Two main anatomical types of male (penial) internal sculpturing are found. The first type presents clear longitudinal pleats that run from the origin of penial pseudopapilla down to (or almost to) the atrium. These longitudinal pleats can be either (almost) completely smooth (Monte Belliemi) or more or less markedly fringed (Calatubo, Romitello, Ficuzza, Jato city park, Monte Kumeta, and Castelluzzo). The second type presents transverse and strong smooth pleats (Bonifato, Gibilmesi, and Visicari).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="sculpturing of the surface of the teleoconch">
<paragraph pageId="0" pageNumber="1">Sculpturing of the surface of the teleoconch.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="423EA979EE665573DC7434AA6E533866" pageId="0" pageNumber="1" type="sculpturing of the surface of the teleoconch">
<paragraph id="3D3CB69015BD3B0B537128632C96C3C5" pageId="0" pageNumber="1">Sculpturing of the surface of the teleoconch.</paragraph>
<paragraph id="C1F40D8688EB175D42FBBF288501920D" pageId="0" pageNumber="1">
The first type corresponds to
<taxonomicName authority="calcarae" lsidName="S. calcarae subsp. calcarae" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae calcarae</emphasis>
<taxonomicName id="C602262362298E0F03269FFF489BC0BB" authority="calcarae" lsidName="S. calcarae subsp. calcarae" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="calcarae">
<emphasis id="3D66217F9CA353B770042CBE96424429" italics="true" pageId="0" pageNumber="1">S. calcarae calcarae</emphasis>
</taxonomicName>
(Philippi, 1844) morphotype, which is more or less striated (Bonifato, Ficuzza, Romitello, Visicari, Gibilmesi, and Castelluzzo), whereas the second type presents a markedly ribbed shell (Calatubo, Jato city park, Monte Kumeta, and Monte Belliemi).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="8ED50EC1FC899F0F6429FADA596E9307" pageId="0" pageNumber="1" type="distribution">
<paragraph id="2650398E7F697FA201F32F7317DEDED5" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="26807DE9D79D6185D0DFDFDE6B27519C" pageId="0" pageNumber="1">
Throughout the whole
<taxonomicName lsidName="S. calcarae" pageId="0" pageNumber="1" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae</emphasis>
<taxonomicName id="246EA337248B0DB9571CFD5D92E06531" lsidName="S. calcarae" pageId="0" pageNumber="1" rank="species" species="calcarae">
<emphasis id="DE3E484422F4448BAFDCC7145AF4FF07" italics="true" pageId="0" pageNumber="1">S. calcarae</emphasis>
</taxonomicName>
distribution, the nominate subspecies
<taxonomicName authority="calcarae" lsidName="S. calcarae subsp. calcarae" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae calcarae</emphasis>
<taxonomicName id="6DB7282BDC36651D3EF70E379C3472F3" authority="calcarae" lsidName="S. calcarae subsp. calcarae" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="calcarae">
<emphasis id="480A34CBFDA13F26A250D03D93EE7599" italics="true" pageId="0" pageNumber="1">S. calcarae calcarae</emphasis>
</taxonomicName>
has the widest distribution, whereas all the remaining subspecies have an isolated, punctiform distribution. The populations from Alcamo, Visicari and Castelluzzo belong to the widely distributed nominate subspecies (1)m whereas the remaining populations (Ficuzza, Calatubo, Monte Kumeta, Jato city park, Romitello, Gibilmesi and Monte Belliemi) belong to limited, punctiform populations (2).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">The populations belonging to this subclade were collected in two main ecological niches. Most of them (type 1) were found on limestone rocks and cliffs (Jato city park, Bonifato, Calatubo, Gibilmesi, Monte Kumeta, Romitello, Monte Belliemi and Visicari), whereas the other populations (type 2), from Ficuzza and Castelluzzo, were exclusively found on tree trunks.</paragraph>
<subSubSection id="3BF1B07F25809D1E6551E9BA53F20B84" pageId="0" pageNumber="1" type="ecology">
<paragraph id="7A4D78E9D15935D3A469E9E2FA1508AF" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="5069DB0DC8428BFB48B8626C8B4D4807" pageId="0" pageNumber="1">The populations belonging to this subclade were collected in two main ecological niches. Most of them (type 1) were found on limestone rocks and cliffs (Jato city park, Bonifato, Calatubo, Gibilmesi, Monte Kumeta, Romitello, Monte Belliemi and Visicari), whereas the other populations (type 2), from Ficuzza and Castelluzzo, were exclusively found on tree trunks.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="haplotypes">
<paragraph pageId="0" pageNumber="1">Haplotypes.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="CFC2D4DFE63097420E2A0A12510EA87D" pageId="0" pageNumber="1" type="haplotypes">
<paragraph id="0ED52BCAC841CE799E9FDDF7C2F03C43" pageId="0" pageNumber="1">Haplotypes.</paragraph>
<paragraph id="B18E22BC21E36A16C9ED6C14AD12B775" pageId="0" pageNumber="1">
Concerning the haplogroups of the
<taxonomicName authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
<taxonomicName id="453CABB5797DE8EED1994CEC02F70F5C" authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis id="489748F757680C1D6FD38F53CC9E057B" italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
</taxonomicName>
clade, eight populations fall into subclade 1 (Bonifato, Calatubo, Romitello, Monte Belliemi, Visicari and Catselluzzo, Monte Kumeta, Ficuzza), whereas two populations (Gibilmesi, and Jato city park) fall in to the subclade 2.
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was recently described from a few scattered localities restricted to Bosco Ficuzza (Corleone, Sicily) (
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: 371). The arrangement of the internal penial sculpturing of the holotype seems not matching with what was found by our dissection, where, instead of &quot;two long weak longitudinal furrows&quot; (
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: 372) a set of transverse, interrupted fleshy smooth pleats were found instead. Yet, the authors provided besides pictures and description of the external morphology of the genital organs, only very few details on the internal morphology of the genital organs, namely: &quot;internal walls of penis show two long weak longitudinal furrows; conic penial papilla, with slightly pointed apex and a restriction to the base&quot;. The brief description and the quality of the images (
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<bibRefCitation id="3657D07916FE1CC7EDF966D4B347A45E" author="Liberto, F" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="365 - 384" refId="B42" refString="Liberto, F, Reitano, A, Giglio, S, Colomba, MS, Sparacio, I, 2016. Two new Clausiliidae (Gastropoda Pulmonata) of Sicily (Italy). Biodiversity Journal 7 (3): 365 - 384" title="Two new Clausiliidae (Gastropoda Pulmonata) of Sicily (Italy)." volume="7" year="2016">Liberto et al. 2016</bibRefCitation>
: 375) do not allow accurate comparison of the internal features with those found in the present analysis. Moreover, the pseudopapilla was erroneously considered as a true penial papilla.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="7D8D38C38F9406CECFBA9CD7C0522780" pageId="0" pageNumber="1">
The position of
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<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae orlandoi</emphasis>
<taxonomicName id="48CF861A8D977C443022D3C25D90EF1D" lsidName="S. calcarae subsp. orlandoi" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="orlandoi">
<emphasis id="F9854A099DFE647DB969B66861C3D4FE" italics="true" pageId="0" pageNumber="1">S. calcarae orlandoi</emphasis>
</taxonomicName>
in the COI tree within haplogroup 1 is close to sequences of a population of
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<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
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<emphasis id="BB3B4D2FCE6CAD3BD1EF8193AC984E3F" italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
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from the western slopes of Monte Kumeta. Despite the close phylogentic relationship of this population with
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<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae orlandoi</emphasis>
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<emphasis id="21AECEA2F55A3098C6B57E385D86B5DF" italics="true" pageId="0" pageNumber="1">S. calcarae orlandoi</emphasis>
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, the differences in the arrangement of the internal male genital organs (a set of transverse, interrupted fleshy smooth pleats and epiphallus with two main fringed chords vs. smooth longitudinal pleats and smooth proximal epiphallus) as well as the remarkable differences concerning shell morphology (see also
<bibRefCitation author="Liberto, F" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="365 - 384" refId="B42" refString="Liberto, F, Reitano, A, Giglio, S, Colomba, MS, Sparacio, I, 2016. Two new Clausiliidae (Gastropoda Pulmonata) of Sicily (Italy). Biodiversity Journal 7 (3): 365 - 384" title="Two new Clausiliidae (Gastropoda Pulmonata) of Sicily (Italy)." volume="7" year="2016">Liberto et al. 2016</bibRefCitation>
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: 372 for a comparative discussion with conspecific taxa) we propose the population from Monte Kumeta as
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<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae parajatinensis</emphasis>
<taxonomicName id="61D6FF62E412F9986E65D306FFEAFF2A" lsidName="S. calcarae subsp. parajatinensis" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="parajatinensis">
<emphasis id="2C634F1938675D4C3D0152B65A39C5E7" italics="true" pageId="0" pageNumber="1">S. calcarae parajatinensis</emphasis>
</taxonomicName>
ssp. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<paragraph id="F0E13287E14627587273CB72BF100CFC" pageId="0" pageNumber="1">
Two additional new subspecies of
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<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
<taxonomicName id="257CF66F3353B025D7AC0ECC39EF6292" authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis id="22D115A12B91B85A86666C827620AA8E" italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
</taxonomicName>
will be described in the following sections:
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<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae borgettensis</emphasis>
<taxonomicName id="B035BF0936F47B3F9E826B3758265AF8" lsidName="S. calcarae subsp. borgettensis" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="borgettensis">
<emphasis id="91EDBD2D38FFB6F0A03EF4AA695008F0" italics="true" pageId="0" pageNumber="1">S. calcarae borgettensis</emphasis>
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ssp. nov. and
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<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae cruenta</emphasis>
<taxonomicName id="D20B1A3B58112893A6101B11B6F4EE98" lsidName="S. calcarae subsp. cruenta" pageId="0" pageNumber="1" rank="subSpecies" species="calcarae" subSpecies="cruenta">
<emphasis id="9793D3E26EB5A19D8886317ABD41AA38" italics="true" pageId="0" pageNumber="1">S. calcarae cruenta</emphasis>
</taxonomicName>
ssp. nov. Both taxa, despite showing close phylogenetic relationships with the other
<taxonomicName authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
<taxonomicName id="D1731B588EDEB33D032032011A5D5A49" authorityName="s. l." baseAuthorityName="Philippi" baseAuthorityYear="1844" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria calcarae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="calcarae">
<emphasis id="1D777D3166CC7F8D4207726F05AF7EEC" italics="true" pageId="0" pageNumber="1">Siciliaria calcarae</emphasis>
</taxonomicName>
taxa, present distinctive shell and genital characters strong enough to propose their description as new subspecies.
</paragraph>

236
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@ -1,192 +1,192 @@
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<taxonomicName LSID="https://zoobank.org/E6927A6E-1532-4746-9679-4391A1F4C4C9" authority="gen. et" authorityName="gen. et" authorityYear="2022" family="Pseudospirobolellidae" genus="Siliquobolellus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Siliquobolellus prasankokae" order="Spirobolida" pageId="0" pageNumber="313" rank="species" species="prasankokae" status="sp. nov.">Siliquobolellus prasankokae gen. et</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="313">sp. nov.</taxonomicNameLabel>
<treatment id="2E9151E1675E59D8840599C81BBF9B62" ID-GBIF-Taxon="199507291" LSID="urn:lsid:zoobank.org:act:E6927A6E-1532-4746-9679-4391A1F4C4C9" httpUri="http://treatment.plazi.org/id/2E9151E1675E59D8840599C81BBF9B62" lastPageNumber="313" pageId="0" pageNumber="313" scope_class="Diplopoda" scope_family="Pseudospirobolellidae" scope_order="Spirobolida" scope_subOrder="Spirobolidea">
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<paragraph id="4E5979E0894CB754450C6260C98E4474" pageId="0" pageNumber="313">
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<taxonomicNameLabel id="5A2BBEE0FD414EE26E3F450D4EB59548" pageId="0" pageNumber="313">sp. nov.</taxonomicNameLabel>
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<paragraph pageId="0" pageNumber="313">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. External morphology of Siliquobolellus gen. nov. A - C. S. amicusdraconis gen. et sp. nov. A. Head, lateral view; B. Head, ventral view; C. Posterior end, lateral view; D. Posterior end, lateral view (S. prasankokae gen. et sp. nov.); E. Posterior end, lateral view (S. constrictus gen. et sp. nov.); F-I. S. prasankokae gen. et sp. nov.; F. Posterior end, ventral view; G, H. Male leg, latero-ventral view; I. Body rings, dorsal colour pattern; J. S. constrictus gen. et sp. nov., body rings, dorsal colour pattern. Av = anal valves; Gst = gnathochilarial stipes; IL = incisura lateralis Me = mentum; Mst = Mandibular stipes; Pre = preanal ring; Sub = subanal scale." figureDoi="10.3897/zse.98.90032.figure2" httpUri="https://binary.pensoft.net/fig/732595" pageId="0" pageNumber="313">Figs 2</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live Siliquobolellus gen. nov. species from Thailand. A, B. S. amicusdraconis gen. et sp. nov. (from Hub Pa Tard), female (paratype, CUMZ-D 00149 - 2); C, D. S. prasankokae gen. et sp. nov. (from Pha Thai), male (holotype, CUMZ-D 00148 - 1)." figureDoi="10.3897/zse.98.90032.figure6" httpUri="https://binary.pensoft.net/fig/732599" pageId="0" pageNumber="313">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Geographic location of the type localities of Siliquobolellus gen. nov. species." figureDoi="10.3897/zse.98.90032.figure7" httpUri="https://binary.pensoft.net/fig/732600" pageId="0" pageNumber="313">, 7</figureCitation>
<subSubSection id="85EE347981F576ADFCCD7BAE04547A9D" pageId="0" pageNumber="313" type="description">
<paragraph id="1855B2AE1D6D17179A233BBED646ED6A" pageId="0" pageNumber="313">
<figureCitation id="758C301B2D578A8D0BDCECB4CCE2C538" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. External morphology of Siliquobolellus gen. nov. A - C. S. amicusdraconis gen. et sp. nov. A. Head, lateral view; B. Head, ventral view; C. Posterior end, lateral view; D. Posterior end, lateral view (S. prasankokae gen. et sp. nov.); E. Posterior end, lateral view (S. constrictus gen. et sp. nov.); F-I. S. prasankokae gen. et sp. nov.; F. Posterior end, ventral view; G, H. Male leg, latero-ventral view; I. Body rings, dorsal colour pattern; J. S. constrictus gen. et sp. nov., body rings, dorsal colour pattern. Av = anal valves; Gst = gnathochilarial stipes; IL = incisura lateralis Me = mentum; Mst = Mandibular stipes; Pre = preanal ring; Sub = subanal scale." figureDoi="10.3897/zse.98.90032.figure2" httpUri="https://binary.pensoft.net/fig/732595" pageId="0" pageNumber="313">Figs 2</figureCitation>
<figureCitation id="3C180A46D9FF531A6C183E50B7982772" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">, 5</figureCitation>
<figureCitation id="BDB1C74EB6C56C55B1D3903DA45C0C07" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live Siliquobolellus gen. nov. species from Thailand. A, B. S. amicusdraconis gen. et sp. nov. (from Hub Pa Tard), female (paratype, CUMZ-D 00149 - 2); C, D. S. prasankokae gen. et sp. nov. (from Pha Thai), male (holotype, CUMZ-D 00148 - 1)." figureDoi="10.3897/zse.98.90032.figure6" httpUri="https://binary.pensoft.net/fig/732599" pageId="0" pageNumber="313">, 6</figureCitation>
<figureCitation id="6461E8EE8BE67FD2E1BA21E0A4ACCBE3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Geographic location of the type localities of Siliquobolellus gen. nov. species." figureDoi="10.3897/zse.98.90032.figure7" httpUri="https://binary.pensoft.net/fig/732600" pageId="0" pageNumber="313">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="313" type="materials_examined">
<paragraph pageId="0" pageNumber="313">Material studied.</paragraph>
<paragraph pageId="0" pageNumber="313">
<materialsCitation collectingDate="2020-07-28" collectorName="P. Pimvichai, P. Prasankok, S. Saratan" country="Thailand" elevation="350" latitude="18.600292" location="Pha Thai" longLatPrecision="1" longitude="99.900276" municipality="Ngao District" specimenCount="1" specimenCount-male="1" stateProvince="Lampang Province" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="313">
<emphasis italics="true" pageId="0" pageNumber="313">
<typeStatus>Holotype</typeStatus>
<subSubSection id="70CA98F9014E81B6250264D1F6765667" pageId="0" pageNumber="313" type="materials_examined">
<paragraph id="B337A6C3D53981664141BC7C1075C930" pageId="0" pageNumber="313">Material studied.</paragraph>
<paragraph id="B2FEB11DED8719161D69CFD1A74E529D" pageId="0" pageNumber="313">
<materialsCitation id="E122EB8CF1241A3DAE25F2A59B494F10" collectingDate="2020-07-28" collectorName="P. Pimvichai, P. Prasankok, S. Saratan" country="Thailand" elevation="350" latitude="18.600292" location="Pha Thai" longLatPrecision="1" longitude="99.900276" municipality="Ngao District" specimenCount="1" specimenCount-male="1" stateProvince="Lampang Province" typeStatus="Holotype">
<emphasis id="6B2E7EDB517D098792B03E25A98D2097" bold="true" pageId="0" pageNumber="313">
<emphasis id="CD03F85764D219AA2551BCCDF02D0B5F" italics="true" pageId="0" pageNumber="313">
<typeStatus id="D87E0E6DA3E1D7DA1DCB8AA07A83B442">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount type="male">1 male</specimenCount>
<specimenCount id="C39530C074D672BD560379D900EE2DD3" type="male">1 male</specimenCount>
(CUMZ-D00148-1),
<collectingCountry name="Thailand">Thailand</collectingCountry>
<collectingCountry id="BDFF4F08BAB3D17B3F0A21671E553840" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion country="Thailand" name="Lampang">Lampang Province</collectingRegion>
<collectingRegion id="F8B571F60833856D6644ECD46C7C1052" country="Thailand" name="Lampang">Lampang Province</collectingRegion>
,
<collectingMunicipality>Ngao District</collectingMunicipality>
<collectingMunicipality id="3EDDCF7A1619DDBAD5846616171A6BA6">Ngao District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:2E9151E1675E59D8840599C81BBF9B62:008835CEE2A742248CD81EE181122117" country="Thailand" latitude="18.600292" longLatPrecision="1" longitude="99.900276" municipality="Ngao District" name="Pha Thai" stateProvince="Lampang Province">Pha Thai</location>
<location id="008835CEE2A742248CD81EE181122117" LSID="urn:lsid:plazi:treatment:2E9151E1675E59D8840599C81BBF9B62:008835CEE2A742248CD81EE181122117" country="Thailand" latitude="18.600292" longLatPrecision="1" longitude="99.900276" municipality="Ngao District" name="Pha Thai" stateProvince="Lampang Province">Pha Thai</location>
;
<geoCoordinate degrees="18" direction="north" minutes="36" orientation="latitude" precision="1" seconds="01.05" value="18.600292">18°36'01.05&quot;N</geoCoordinate>
<geoCoordinate id="CED07B468C6727605F02E7B620E66245" degrees="18" direction="north" minutes="36" orientation="latitude" precision="1" seconds="01.05" value="18.600292">18°36'01.05&quot;N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="54" orientation="longitude" precision="1" seconds="01.00" value="99.900276">99°54'01.00&quot;E</geoCoordinate>
<geoCoordinate id="39B976D8B2BEA458A83E1D9B88E71543" degrees="99" direction="east" minutes="54" orientation="longitude" precision="1" seconds="01.00" value="99.900276">99°54'01.00&quot;E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
<elevation id="38D0218951969D3A25506D445CA9ECCC" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
<quantity id="030CE437FC94E50B0FFD0C979343B954" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
a.s.l.
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;
<collectingDate value="2020-07-28">28 July 2020</collectingDate>
<collectingDate id="A150842D8CEBABF494E5624963ABFBC2" value="2020-07-28">28 July 2020</collectingDate>
;
<collectorName>P. Pimvichai</collectorName>
<collectorName id="D215BE08472665860FA2AF022EA0E9FC">P. Pimvichai</collectorName>
,
<collectorName>P. Prasankok</collectorName>
<collectorName id="18FEE56D07F375BF73315B1B31943227">P. Prasankok</collectorName>
and
<collectorName>S. Saratan</collectorName>
<collectorName id="60D7321DAFF3629806C0D88057511F0C">S. Saratan</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="313">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="P. Pimvichai &amp; P. Prasankok &amp; S. Saratan" country="Thailand" elevation="350" latitude="18.600292" location="Pha Thai" longLatPrecision="1" longitude="99.900276" municipality="Ngao District" specimenCount="6" specimenCount-female="6" stateProvince="Lampang" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="313">
<emphasis italics="true" pageId="0" pageNumber="313">
<typeStatus>Paratypes</typeStatus>
<paragraph id="7E0BCBBB1ECCD5B8ED218E758A9D3F5D" pageId="0" pageNumber="313">
<materialsCitation id="89EE7B17AAD2D9AF89E89CC1394FB9EA" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="P. Pimvichai &amp; P. Prasankok &amp; S. Saratan" country="Thailand" elevation="350" latitude="18.600292" location="Pha Thai" longLatPrecision="1" longitude="99.900276" municipality="Ngao District" specimenCount="6" specimenCount-female="6" stateProvince="Lampang" typeStatus="Paratypes">
<emphasis id="5D374D655B05466CEBDB45822A6E4A31" bold="true" pageId="0" pageNumber="313">
<emphasis id="DDFC806BE132384260AD738E092FE66A" italics="true" pageId="0" pageNumber="313">
<typeStatus id="1A077891924DA8CE084ED1CAEAB9248C">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount type="female">6 females</specimenCount>
<specimenCount id="7DE2A3C1FF31D9E4A1C33DAFEC144CBD" type="female">6 females</specimenCount>
(CUMZ-D00148-2); same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="313" type="etymology">
<paragraph pageId="0" pageNumber="313">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="313">The species honors Associate Professor Dr. Pongpun Prasankok, biologist and devoted millipede collector.</paragraph>
<subSubSection id="830A255770850432890C8C1F5DE5D93A" pageId="0" pageNumber="313" type="etymology">
<paragraph id="8D0A4C04BF37EF110B2F28F432FA9549" pageId="0" pageNumber="313">Etymology.</paragraph>
<paragraph id="87E6198DB95A66E5FE2FD1D15241FFE2" pageId="0" pageNumber="313">The species honors Associate Professor Dr. Pongpun Prasankok, biologist and devoted millipede collector.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="313" type="diagnosis">
<paragraph pageId="0" pageNumber="313">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="313">Differing from other species in the genus by having the posterior gonopod apically with a short, sharp protrusion; externally differing by having a row of rectangular dark brown spots middorsally on the body rings.</paragraph>
<subSubSection id="2D9A4ABC1D51722996C9719276C8B123" pageId="0" pageNumber="313" type="diagnosis">
<paragraph id="EA27FF807C0AACE2A416FC26C33FA756" pageId="0" pageNumber="313">Diagnosis.</paragraph>
<paragraph id="657D9C517E75099A7498CD6DCAFDA396" pageId="0" pageNumber="313">Differing from other species in the genus by having the posterior gonopod apically with a short, sharp protrusion; externally differing by having a row of rectangular dark brown spots middorsally on the body rings.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="313" type="description">
<paragraph pageId="0" pageNumber="313">Description.</paragraph>
<paragraph pageId="0" pageNumber="313">Adult male with 41 podous rings, 1 apodous ring. Length ~3 cm, diameter ~3 mm. Adult females with 40 or 41 podous rings, 1 apodous ring. Length ~3 cm, diameter 3.2-3.6 mm.</paragraph>
<paragraph pageId="0" pageNumber="313">
<subSubSection id="7983967C54D46EB89165953372FA2C54" pageId="0" pageNumber="313" type="description">
<paragraph id="E381D0B85FD032BEFA6638A48F308076" pageId="0" pageNumber="313">Description.</paragraph>
<paragraph id="D1A3C59476846C6D33E5EE0268429C18" pageId="0" pageNumber="313">Adult male with 41 podous rings, 1 apodous ring. Length ~3 cm, diameter ~3 mm. Adult females with 40 or 41 podous rings, 1 apodous ring. Length ~3 cm, diameter 3.2-3.6 mm.</paragraph>
<paragraph id="26521D8CA7E605D182E5FB814472CE29" pageId="0" pageNumber="313">
Color of living animal: collum and telson beige; antenna and leg light brown; dorsal part of body rings yellowish brown, with a row of rectangular dark brown spots middorsally on the body rings; head and lateral part of body rings dark brown (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. External morphology of Siliquobolellus gen. nov. A - C. S. amicusdraconis gen. et sp. nov. A. Head, lateral view; B. Head, ventral view; C. Posterior end, lateral view; D. Posterior end, lateral view (S. prasankokae gen. et sp. nov.); E. Posterior end, lateral view (S. constrictus gen. et sp. nov.); F-I. S. prasankokae gen. et sp. nov.; F. Posterior end, ventral view; G, H. Male leg, latero-ventral view; I. Body rings, dorsal colour pattern; J. S. constrictus gen. et sp. nov., body rings, dorsal colour pattern. Av = anal valves; Gst = gnathochilarial stipes; IL = incisura lateralis Me = mentum; Mst = Mandibular stipes; Pre = preanal ring; Sub = subanal scale." figureDoi="10.3897/zse.98.90032.figure2" httpUri="https://binary.pensoft.net/fig/732595" pageId="0" pageNumber="313">2I</figureCitation>
<figureCitation id="EC9076D7F93083B12AF5EE7EA61B446D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. External morphology of Siliquobolellus gen. nov. A - C. S. amicusdraconis gen. et sp. nov. A. Head, lateral view; B. Head, ventral view; C. Posterior end, lateral view; D. Posterior end, lateral view (S. prasankokae gen. et sp. nov.); E. Posterior end, lateral view (S. constrictus gen. et sp. nov.); F-I. S. prasankokae gen. et sp. nov.; F. Posterior end, ventral view; G, H. Male leg, latero-ventral view; I. Body rings, dorsal colour pattern; J. S. constrictus gen. et sp. nov., body rings, dorsal colour pattern. Av = anal valves; Gst = gnathochilarial stipes; IL = incisura lateralis Me = mentum; Mst = Mandibular stipes; Pre = preanal ring; Sub = subanal scale." figureDoi="10.3897/zse.98.90032.figure2" httpUri="https://binary.pensoft.net/fig/732595" pageId="0" pageNumber="313">2I</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live Siliquobolellus gen. nov. species from Thailand. A, B. S. amicusdraconis gen. et sp. nov. (from Hub Pa Tard), female (paratype, CUMZ-D 00149 - 2); C, D. S. prasankokae gen. et sp. nov. (from Pha Thai), male (holotype, CUMZ-D 00148 - 1)." figureDoi="10.3897/zse.98.90032.figure6" httpUri="https://binary.pensoft.net/fig/732599" pageId="0" pageNumber="313">6C, D</figureCitation>
<figureCitation id="01AF15476E387CF65D88D488315661A6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Live Siliquobolellus gen. nov. species from Thailand. A, B. S. amicusdraconis gen. et sp. nov. (from Hub Pa Tard), female (paratype, CUMZ-D 00149 - 2); C, D. S. prasankokae gen. et sp. nov. (from Pha Thai), male (holotype, CUMZ-D 00148 - 1)." figureDoi="10.3897/zse.98.90032.figure6" httpUri="https://binary.pensoft.net/fig/732599" pageId="0" pageNumber="313">6C, D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="313">
<paragraph id="69F2E8C59BECCFEFD000FBE4117CA7DE" pageId="0" pageNumber="313">
Anterior gonopods (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5A, B, D, E</figureCitation>
<figureCitation id="1EFAD93D2048BD9A9AE5058846A07F1F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5A, B, D, E</figureCitation>
) with high coxae, apically narrow, curving mesad, mesal margin sigmoid, posterior surface folding to cover more than half of telopodite, with strong ridge laterally. Telopodite with thick process, with strong ridge distomesally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5B</figureCitation>
<figureCitation id="68248F119E7A35451A129EB77A3C2711" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5B</figureCitation>
, arrow).
</paragraph>
<paragraph pageId="0" pageNumber="313">
<paragraph id="F612FF5A4925C53E02B50CB4D9ADB04A" pageId="0" pageNumber="313">
Posterior gonopods (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5C, F-J</figureCitation>
<figureCitation id="0C22DB9C97DFD75EC980D13DFC34F043" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5C, F-J</figureCitation>
) simple, rounded, with short, smooth coxal part (pcx); telopodital part (pt) twice the length of the pcx, lateral margin folding mesad, forming a deep concavity, the inner lateral margin expanded into a rounded lamella, apically forming a canopy (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5I</figureCitation>
<figureCitation id="FB88A4A4805E1AA3C06212C00994014C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5I</figureCitation>
, arrow), apically with a short, sharp protrusion (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5C, G</figureCitation>
<figureCitation id="154188BE6EFA3D31089E1F7B2D9605E5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5C, G</figureCitation>
, arrows).
</paragraph>
<paragraph pageId="0" pageNumber="313">
<paragraph id="01256580456F645D932A56FEE3F29552" pageId="0" pageNumber="313">
Female vulvae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5K</figureCitation>
<figureCitation id="19C13DCBDA0822A7EE9101B6CF776D64" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siliquobolellus prasankokae gen. et sp. nov., holotype, gonopods (specimen from Pha Thai, CUMZ-D 00148 - 1). A. Anterior gonopod, anterior view; B. Anterior gonopod, posterior view, arrow indicates a a thumb-like process with strong ridge distomesally; C. Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; D. Anterior gonopod, anterior view; E. Anterior gonopod, posterior view; F. Right posterior gonopod, lateral view; G. SEM, Right posterior gonopod, lateral view, arrow indicates a short, sharp protrusion; H. SEM, Mesal part of right posterior gonopod, lateral view; I. SEM, Right posterior gonopod, mesal view, arrow indicates a canopy; J. SEM, Tip of right posterior gonopod, mesal view; K. SEM, left female vulva, posterior mesal view. at = anterior gonopod telopodite; cx = coxa; oeg = opening of efferent groove; op = operculum of vulva; pcx = coxal part of the posterior gonopod telopodite; pt = telopodital part of the posterior gonopod telopodite; st = sternal process." figureDoi="10.3897/zse.98.90032.figure5" httpUri="https://binary.pensoft.net/fig/732598" pageId="0" pageNumber="313">5K</figureCitation>
): simple, valves prominent, the right valve slightly larger than the left valve.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="313" type="dna barcode">
<paragraph pageId="0" pageNumber="313">DNA barcode.</paragraph>
<paragraph pageId="0" pageNumber="313">The GenBank accession number of the COI barcode of the paratype is OP174623 (voucher code CUMZ-D00148).</paragraph>
<subSubSection id="05C634C2CC9F05C68EDB22839E6FC3BE" pageId="0" pageNumber="313" type="dna barcode">
<paragraph id="4932A6E20A1B391F912C1A79F58DD70E" pageId="0" pageNumber="313">DNA barcode.</paragraph>
<paragraph id="F1894A7502542357BA190B4E79F72F08" pageId="0" pageNumber="313">The GenBank accession number of the COI barcode of the paratype is OP174623 (voucher code CUMZ-D00148).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="313" type="habitat">
<paragraph pageId="0" pageNumber="313">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="313">Found under leaf litter and inside a rotten log.</paragraph>
<subSubSection id="D1AE58FE678A57D892E9411EDA225020" pageId="0" pageNumber="313" type="habitat">
<paragraph id="FB51A37249DAAF681742907524A7863C" pageId="0" pageNumber="313">Habitat.</paragraph>
<paragraph id="44E41B488453DD4559F281DDD33ACFF5" pageId="0" pageNumber="313">Found under leaf litter and inside a rotten log.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="313" type="distribution">
<paragraph pageId="0" pageNumber="313">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="313">
<subSubSection id="B49DCAB6BCA1604EE4DF5852AF6DE215" pageId="0" pageNumber="313" type="distribution">
<paragraph id="99FA6EB1A96E423A5D407C5C1DEEB15C" pageId="0" pageNumber="313">Distribution.</paragraph>
<paragraph id="F8B4E57AEEE7B0E022B099B86C5BEECD" pageId="0" pageNumber="313">
Known only from the type locality in Lampang Province, Thailand (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Geographic location of the type localities of Siliquobolellus gen. nov. species." figureDoi="10.3897/zse.98.90032.figure7" httpUri="https://binary.pensoft.net/fig/732600" pageId="0" pageNumber="313">7</figureCitation>
<figureCitation id="A93F6CCFA415200934CAC9C774F6690B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Geographic location of the type localities of Siliquobolellus gen. nov. species." figureDoi="10.3897/zse.98.90032.figure7" httpUri="https://binary.pensoft.net/fig/732600" pageId="0" pageNumber="313">7</figureCitation>
).
</paragraph>
</subSubSection>

708
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