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<document id="42FA6019C6DC966995B709802B2136D8" ID-DOI="10.11646/zootaxa.4027.2.8" ID-GBIF-Dataset="8dcb1038-d556-49c8-8649-50202fa770f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="236450" ID-ZooBank="9096376A-C64F-46D7-8E75-55B56E495A4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461282324405" checkinUser="plazi" docAuthor="Teixeira, Juliana Henrique, Rizzo, Alexandra Elaine &amp; Santos, Cinthya Simone Gomes" docDate="2015" docId="4F7A8122592AFFD4E59FBAF7FAEFFD06" docLanguage="en" docName="zt04027p295.pdf" docOrigin="Zootaxa 4027 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diplocirrus salazarvallejoi Teixeira, Rizzo &amp; Santos, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="292" masterDocId="B343F95A592EFFD1E508BB05FFC8FFF9" masterDocTitle="Three new species of Diplocirrus Haase, 1915 (Polychaeta: Flabelligeridae) from Campos Basin in SE Brazil" masterLastPageNumber="295" masterPageNumber="287" pageNumber="291" updateTime="1698614352147" updateUser="plazi">
<mods:mods id="84D2AA9248B22CE0B53DFCEC7B629B7E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C20ACBCA54BC1AB99E64517BE2551E2">
<mods:title id="23F6B7F4572B43DBF0D0EBF073420E06">Three new species of Diplocirrus Haase, 1915 (Polychaeta: Flabelligeridae) from Campos Basin in SE Brazil</mods:title>
</mods:titleInfo>
<mods:name id="E9893980CA043DBDCF457FB2E21C2220" type="personal">
<mods:role id="F5BD0033C183E2766CFA715B31FB1D7B">
<mods:roleTerm id="F73C3D958AC7904E7BAE2FFC6F1B410A">Author</mods:roleTerm>
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<mods:namePart id="367B70CA28FB079FC22BAF1FFA96D6A2">Teixeira, Juliana Henrique</mods:namePart>
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<mods:namePart id="8CE69BD6C090130FBE298CED3BC966F9">Rizzo, Alexandra Elaine</mods:namePart>
</mods:name>
<mods:name id="18878EA515BD3B5F6165CF346E584390" type="personal">
<mods:role id="2C243DA13DFF0453E771224B5F0832B4">
<mods:roleTerm id="A02F3C6ECB63240353991A061F46263B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="37A1565908F004C3CCF0BED1134E36ED">Santos, Cinthya Simone Gomes</mods:namePart>
</mods:name>
<mods:typeOfResource id="DD7AEFA603118892B3B790CEB4684978">text</mods:typeOfResource>
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<mods:title id="EB4731D1EFCEE22A2A9D9A6CEE375470">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F32907E74F34A33CBFB5A7A372A16DDC">
<mods:date id="441E8715516B824F61EFFE80D8C069DB">2015</mods:date>
<mods:detail id="C33C8E77A1D13098526F7F230C095E6E" type="volume">
<mods:number id="90581B54F643D2BE07904D056C8E5FD8">4027</mods:number>
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<mods:detail id="10D2CF2DB0FA2C68FE8F1FB037D017C4" type="issue">
<mods:number id="019606B1FFF176398D0373F6300D4D20">2</mods:number>
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<mods:start id="EFDF9BF5A6865C3615AF801338AC192B">287</mods:start>
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<mods:classification id="FFA01E6B9DB824CDFF41F5C9DD79C588">journal article</mods:classification>
<mods:identifier id="1A2289B115AB07B5BF4A2F895D8DEA97" type="DOI">10.11646/zootaxa.4027.2.8</mods:identifier>
<mods:identifier id="FD1AF2296EAD9FB382B966B5939AFE79" type="GBIF-Dataset">8dcb1038-d556-49c8-8649-50202fa770f6</mods:identifier>
<mods:identifier id="EE341A664376FA107B97EAB001024A43" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B245A7B913552F28033BB4D111924BD4" type="Zenodo-Dep">236450</mods:identifier>
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</mods:mods>
<treatment id="4F7A8122592AFFD4E59FBAF7FAEFFD06" ID-DOI="http://doi.org/10.5281/zenodo.6098266" ID-GBIF-Taxon="119640878" ID-Zenodo-Dep="6098266" LSID="urn:lsid:plazi:treatment:4F7A8122592AFFD4E59FBAF7FAEFFD06" httpUri="http://treatment.plazi.org/id/4F7A8122592AFFD4E59FBAF7FAEFFD06" lastPageId="5" lastPageNumber="292" pageId="4" pageNumber="291">
<subSubSection id="8FC963BF592AFFD5E59FBAF7FEFCFDD5" pageId="4" pageNumber="291" type="nomenclature">
<paragraph id="C76C3034592AFFD5E59FBAF7FDF3FDF5" blockId="4.[151,571,498,556]" box="[151,571,498,524]" pageId="4" pageNumber="291">
<heading id="9C248758592AFFD5E59FBAF7FDF3FDF5" bold="true" box="[151,571,498,524]" fontSize="11" level="1" pageId="4" pageNumber="291" reason="1">
<emphasis id="F5A7EC26592AFFD5E59FBAF7FDF3FDF5" bold="true" box="[151,571,498,524]" pageId="4" pageNumber="291">
<taxonomicName id="00D34BB7592AFFD5E59FBAF7FE1EFDF5" ID-CoL="36JVZ" box="[151,470,498,524]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="291" phylum="Annelida" rank="species" species="salazarvallejoi" status="sp. nov.">
<emphasis id="F5A7EC26592AFFD5E59FBAF7FE1EFDF5" bold="true" box="[151,470,498,524]" italics="true" pageId="4" pageNumber="291">Diplocirrus salazarvallejoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94515D592AFFD5E4D6BAF7FDF3FDF5" box="[478,571,498,524]" pageId="4" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C76C3034592AFFD5E59FB910FEFCFDD5" blockId="4.[151,571,498,556]" box="[151,308,533,556]" pageId="4" pageNumber="291">
(
<figureCitation id="5FE82CB1592AFFD5E596B910FF31FDD5" box="[158,249,533,556]" captionStart="FIGURE 2" captionStartId="4.[151,244,1815,1837]" captionTargetBox="[232,1322,830,1793]" captionTargetId="figure@4.[217,1369,815,1793]" captionTargetPageId="4" captionText="FIGURE 2. Diplocirrus salazarvallejoi sp. nov. A. Holotype (UERJ 5572), anterior region in oblique lateral view and posterior in dorsal view. B. Same, chaetiger 10, notochaetae. C. Same, neurochaetae. Scale bars: A: 0.5 mm, B: 30 µm, C: 18 µm." httpUri="https://zenodo.org/record/236452/files/figure.png" pageId="4" pageNumber="291">Figure 2</figureCitation>
A–C)
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592AFFD5E59FB95FFCD2FD6D" pageId="4" pageNumber="291" type="materials_examined">
<paragraph id="C76C3034592AFFD5E59FB95FFCD2FD6D" blockId="4.[151,1436,602,764]" pageId="4" pageNumber="291">
<emphasis id="F5A7EC26592AFFD5E59FB95FFEF5FD88" bold="true" box="[151,317,602,625]" pageId="4" pageNumber="291">
<typeStatus id="18688E96592AFFD5E59FB95FFF07FD88" box="[151,207,602,625]" pageId="4" pageNumber="291">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="BFC470A4592AFFD5E44DB95FFE42FD88" box="[325,394,602,625]" name="Brazil" pageId="4" pageNumber="291">Brazil</collectingCountry>
. Atlantic Ocean. Southeastern coast, Campos Basin.
<typeStatus id="18688E96592AFFD5E6CDB95FFBE0FD88" box="[965,1064,602,625]" pageId="4" pageNumber="291" type="holotype">Holotype</typeStatus>
(
<collectionCode id="A1C2A8F1592AFFD5E13FB95FFBB2FD88" box="[1079,1146,602,625]" pageId="4" pageNumber="291">UERJ</collectionCode>
5572-HAB8 E6 R3),
<quantity id="002B9DD1592AFFD5E068B95FFF66FD6D" metricMagnitude="2" metricUnit="m" metricValue="3.871" pageId="4" pageNumber="291" unit="m" value="387.1">387.1 m</quantity>
,
<date id="B36D16F4592AFFD5E5B1B978FEA4FD6D" box="[185,364,637,660]" pageId="4" pageNumber="291" value="2009-01-31">31 January 2009</date>
, (
<geoCoordinate id="A2E756F3592AFFD5E476B978FDF2FD6D" box="[382,570,637,660]" direction="south" orientation="latitude" pageId="4" pageNumber="291" precision="1" value="-22.431934">22º 25' 54,966&quot; S</geoCoordinate>
,
<geoCoordinate id="A2E756F3592AFFD5E74EB978FCC5FD6D" box="[582,781,637,660]" direction="west" orientation="longitude" pageId="4" pageNumber="291" precision="1" value="-40.294315">40º 17' 39,534&quot; W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592AFFD5E5CFB9A5FDF0FD05" pageId="4" pageNumber="291" type="diagnosis">
<paragraph id="C76C3034592AFFD5E5CFB9A5FDF0FD05" blockId="4.[151,1436,602,764]" pageId="4" pageNumber="291">
<emphasis id="F5A7EC26592AFFD5E5CFB9A5FEF4FD4E" bold="true" box="[199,316,672,695]" pageId="4" pageNumber="291">Diagnosis.</emphasis>
Sediment particles concentrated at the base of papillae, all of same size on ventral and dorsal regions, 6– 8 notochaetae and 4–5 neurochaetae per bundle throughout body, ventrolateral gonopores from chaetigers 3–12, notochaetae with small articles basally.
</paragraph>
</subSubSection>
<caption id="93AC60BC592AFFD5E59FBC12FAC8F8B0" httpUri="https://zenodo.org/record/236452/files/figure.png" pageId="4" pageNumber="291" targetBox="[232,1322,830,1793]" targetPageId="4">
<paragraph id="C76C3034592AFFD5E59FBC12FAC8F8B0" blockId="4.[151,1436,1814,1866]" pageId="4" pageNumber="291">
<emphasis id="F5A7EC26592AFFD5E59FBC12FEC6F8D4" bold="true" box="[151,270,1815,1837]" pageId="4" pageNumber="291">FIGURE 2.</emphasis>
<taxonomicName id="00D34BB7592AFFD5E41EBC13FDD5F8D5" box="[278,541,1814,1836]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="291" phylum="Annelida" rank="species" species="salazarvallejoi" status="sp. nov.">
<emphasis id="F5A7EC26592AFFD5E41EBC13FDD5F8D5" box="[278,541,1814,1836]" italics="true" pageId="4" pageNumber="291">Diplocirrus salazarvallejoi</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592AFFD5E72DBC12FDBBF8D4" bold="true" box="[549,627,1815,1837]" pageId="4" pageNumber="291">
<taxonomicNameLabel id="EE94515D592AFFD5E72DBC12FDBBF8D4" box="[549,627,1815,1837]" pageId="4" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Holotype (UERJ5572), anterior region in oblique lateral view and posterior in dorsal view. B. Same, chaetiger 10, notochaetae. C. Same, neurochaetae. Scale bars: A: 0.5 mm, B: 30 µm, C: 18 µm.
</paragraph>
</caption>
<subSubSection id="8FC963BF592AFFD4E5CFBC6AFC5AFEA6" lastPageId="5" lastPageNumber="292" pageId="4" pageNumber="291" type="description">
<paragraph id="C76C3034592AFFD4E5CFBC6AFC5AFEA6" blockId="4.[151,1435,1903,2030]" lastBlockId="5.[151,1436,152,767]" lastPageId="5" lastPageNumber="292" pageId="4" pageNumber="291">
<emphasis id="F5A7EC26592AFFD5E5CFBC6AFE9AF87F" bold="true" box="[199,338,1903,1926]" pageId="4" pageNumber="291">Description.</emphasis>
<typeStatus id="18688E96592AFFD5E456BC6AFE09F87F" box="[350,449,1903,1926]" pageId="4" pageNumber="291" type="holotype">Holotype</typeStatus>
incomplete; body cylindrical swollen along first 8 chaetigers, tapering in median and posterior region;
<quantity id="002B9DD1592AFFD5E459BC97FE55F850" box="[337,413,1938,1961]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="291" unit="mm" value="10.0">10 mm</quantity>
long,
<quantity id="002B9DD1592AFFD5E4EABC97FDFCF850" box="[482,564,1938,1961]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="291" unit="mm" value="1.2">1.2 mm</quantity>
wide, cephalic cage
<quantity id="002B9DD1592AFFD5E61FBC97FC9FF850" box="[791,855,1938,1961]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="291" unit="mm" value="1.0">1 mm</quantity>
long, for 20 chaetigers. Body whitish, tunic thin with papillae, fine sediment adhered to base of capitate papillae (longer than wide), all of same size on ventral and dorsal regions, up to 4 times longer on the lateral. Numerous rows of papillae on chaetiger 10. Longest papillae 1/5 as long as median notochaetae. Cephalic hood exposed, but anterior end features were either eroded or not visible. Cephalic cage chaetae ¼ as long as body width. Only notochaetae of chaetiger 1 form cephalic cage, 6–8 noto-and 4–5 neurochaetae per bundle. Notochaetae slightly longer in median and posterior region, 1/3 as long as body width, neurochaetae of the same length throughout body. From middle region, notochaetae twice as long as neurochaetae. Ventrolateral gonopores in chaetigers 3 to 12. Noto- and neurochaetae multiarticulate, articles of same size, finely tuned falcate tip. From chaetigers 1–13th, chaetae are twice longer than in following chaetigers.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592BFFD4E5CFBA6FFF23FD8D" pageId="5" pageNumber="292" type="discussion">
<paragraph id="C76C3034592BFFD4E5CFBA6FFF23FD8D" blockId="5.[151,1436,152,767]" pageId="5" pageNumber="292">
<emphasis id="F5A7EC26592BFFD4E5CFBA6FFEFDFE78" bold="true" box="[199,309,362,385]" pageId="5" pageNumber="292">Remarks.</emphasis>
<taxonomicName id="00D34BB7592BFFD4E437BA6FFDABFE78" box="[319,611,362,385]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="salazarvallejoi" status="sp. nov.">
<emphasis id="F5A7EC26592BFFD4E437BA6FFDABFE78" box="[319,611,362,385]" italics="true" pageId="5" pageNumber="292">Diplocirrus salazarvallejoi</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592BFFD4E765BA6FFD0DFE78" bold="true" box="[621,709,362,385]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="EE94515D592BFFD4E765BA6FFD0DFE78" box="[621,709,362,385]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="00D34BB7592BFFD4E654BA6FFACDFE78" authority="Malmgren, 1867" authorityName="Malmgren" authorityYear="1867" box="[860,1285,362,385]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="glaucus">
<emphasis id="F5A7EC26592BFFD4E654BA6FFBFDFE78" box="[860,1077,362,385]" italics="true" pageId="5" pageNumber="292">Diplocirrus glaucus</emphasis>
(
<bibRefCitation id="A3424DC5592BFFD4E14EBA6FFB35FE78" author="Malmgren" box="[1094,1277,362,385]" pageId="5" pageNumber="292" refString="Malmgren, A. J. (1867) Annulata Polychaeta: Spetsbergiae, Groenlandiae, Islandiae et Scandinaviae hactenus cognita. Kongelige Vetenskaps-Akademiens Forhandlingar, 4, 127 - 235." type="journal article" year="1867">Malmgren, 1867</bibRefCitation>
)
</taxonomicName>
because both have long papillae on chaetal lobes, about 1/5 as long as median notochaetal length, anterior chaetigers longer than those that follow. Only notochaetae of chaetiger 1 are involved in cephalic cage. In contrast,
<taxonomicName id="00D34BB7592BFFD4E158BAB5FABCFE3E" box="[1104,1396,432,455]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="salazarvallejoi" status="sp. nov.">
<emphasis id="F5A7EC26592BFFD4E158BAB5FABCFE3E" box="[1104,1396,432,455]" italics="true" pageId="5" pageNumber="292">Diplocirrus salazarvallejoi</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592BFFD4E074BAB5FF0EFE10" bold="true" pageId="5" pageNumber="292">
<taxonomicNameLabel id="EE94515D592BFFD4E074BAB5FF0EFE10" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has: a) 6–8 notochaetae and 4–5 neurochaetae per bundle throughout body, b) ventrolateral gonopores from chaetigers 3–12, and c) notochaetae with short articles basally.
<taxonomicName id="00D34BB7592BFFD4E65FBAF0FBF9FDF5" box="[855,1073,501,524]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="glaucus">
<emphasis id="F5A7EC26592BFFD4E65FBAF0FBF9FDF5" box="[855,1073,501,524]" italics="true" pageId="5" pageNumber="292">Diplocirrus glaucus</emphasis>
</taxonomicName>
has a) 2–3 notochaetae and 0–1 neurochaetae in anterior chaetigers, 7–8 notochaetae and 4–5 neurochaetae in median chaetigers, b) gonopores not observed, and c) notochaetae with medium size articles basally. Previous records of
<taxonomicName id="00D34BB7592BFFD4E140B93FFB77FDA8" box="[1096,1215,570,593]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="glaucus">
<emphasis id="F5A7EC26592BFFD4E140B93FFB77FDA8" box="[1096,1215,570,593]" italics="true" pageId="5" pageNumber="292">D. glaucus</emphasis>
</taxonomicName>
may belong to this species.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592BFFD4E5CFB985FEE3FD25" pageId="5" pageNumber="292" type="etymology">
<paragraph id="C76C3034592BFFD4E5CFB985FEE3FD25" blockId="5.[151,1436,152,767]" pageId="5" pageNumber="292">
<emphasis id="F5A7EC26592BFFD4E5CFB985FE8FFD6E" bold="true" box="[199,327,640,663]" pageId="5" pageNumber="292">Etymology.</emphasis>
This species is dedicated to Dr. Sergio I. Salazar-Vallejo for his contribution to the study of polychaetes, especially the family
<taxonomicName id="00D34BB7592BFFD4E488B9A7FDEAFD40" box="[384,546,674,697]" class="Polychaeta" family="Flabelligeridae" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="family">Flabelligeridae</taxonomicName>
, and for his encouragement and guidance to the first author (JHT) on the study of flabelligerids.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592BFFD4E5CFB9EDFAEFFD06" box="[199,1319,744,767]" pageId="5" pageNumber="292" type="distribution">
<paragraph id="C76C3034592BFFD4E5CFB9EDFAEFFD06" blockId="5.[151,1436,152,767]" box="[199,1319,744,767]" pageId="5" pageNumber="292">
<emphasis id="F5A7EC26592BFFD4E5CFB9EDFE91FD06" bold="true" box="[199,345,744,767]" pageId="5" pageNumber="292">Distribution.</emphasis>
Southeastern Atlantic coast, Campos Basin,
<collectingCountry id="BFC470A4592BFFD4E630B9EDFCB4FD06" box="[824,892,744,767]" name="Brazil" pageId="5" pageNumber="292">Brazil</collectingCountry>
, in soft bottom, only from
<quantity id="002B9DD1592BFFD4E192B9EDFB16FD06" box="[1178,1246,744,767]" metricMagnitude="2" metricUnit="m" metricValue="3.87" pageId="5" pageNumber="292" unit="m" value="387.0">387 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="47D1F3A712F25F1D9E3551FBC1B29599" ID-DOI="10.11646/zootaxa.4027.2.8" ID-GBIF-Dataset="8dcb1038-d556-49c8-8649-50202fa770f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="236450" ID-ZooBank="9096376A-C64F-46D7-8E75-55B56E495A4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461282324405" checkinUser="plazi" docAuthor="Teixeira, Juliana Henrique, Rizzo, Alexandra Elaine &amp; Santos, Cinthya Simone Gomes" docDate="2015" docId="4F7A8122592BFFD7E59FB84AFAD1FF0D" docLanguage="en" docName="zt04027p295.pdf" docOrigin="Zootaxa 4027 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diplocirrus acafi Teixeira, Rizzo &amp; Santos, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="293" masterDocId="B343F95A592EFFD1E508BB05FFC8FFF9" masterDocTitle="Three new species of Diplocirrus Haase, 1915 (Polychaeta: Flabelligeridae) from Campos Basin in SE Brazil" masterLastPageNumber="295" masterPageNumber="287" pageNumber="292" updateTime="1698614352147" updateUser="plazi">
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<mods:title id="511E90C85336C1052D99F4D38C2C52FB">Three new species of Diplocirrus Haase, 1915 (Polychaeta: Flabelligeridae) from Campos Basin in SE Brazil</mods:title>
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<mods:namePart id="370E0CC2B0F7F1E481C98A14F8EDECEE">Teixeira, Juliana Henrique</mods:namePart>
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<mods:namePart id="DC2FCB3CE4499BCFD74C31814C321B25">Rizzo, Alexandra Elaine</mods:namePart>
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<mods:namePart id="8AD818EF12F91E66141E25D0041DDC19">Santos, Cinthya Simone Gomes</mods:namePart>
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<mods:date id="07E1B860C6BF4EC3D2FBD41225ED81D7">2015</mods:date>
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<treatment id="4F7A8122592BFFD7E59FB84AFAD1FF0D" ID-DOI="http://doi.org/10.5281/zenodo.6098268" ID-GBIF-Taxon="119640877" ID-Zenodo-Dep="6098268" LSID="urn:lsid:plazi:treatment:4F7A8122592BFFD7E59FB84AFAD1FF0D" httpUri="http://treatment.plazi.org/id/4F7A8122592BFFD7E59FB84AFAD1FF0D" lastPageId="6" lastPageNumber="293" pageId="5" pageNumber="292">
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<paragraph id="C76C3034592BFFD4E59FB84AFE01FC93" blockId="5.[151,457,847,905]" box="[151,457,847,874]" pageId="5" pageNumber="292">
<heading id="9C248758592BFFD4E59FB84AFE01FC93" bold="true" box="[151,457,847,874]" fontSize="11" level="1" pageId="5" pageNumber="292" reason="1">
<emphasis id="F5A7EC26592BFFD4E59FB84AFE01FC93" bold="true" box="[151,457,847,874]" pageId="5" pageNumber="292">
<taxonomicName id="00D34BB7592BFFD4E59FB84AFEACFC90" ID-CoL="36JVH" box="[151,356,847,873]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="acafi" status="sp. nov.">
<emphasis id="F5A7EC26592BFFD4E59FB84AFEACFC90" bold="true" box="[151,356,847,873]" italics="true" pageId="5" pageNumber="292">Diplocirrus acafi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94515D592BFFD4E464B855FE01FC93" box="[364,457,848,874]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C76C3034592BFFD4E59FB877FEFEFC70" blockId="5.[151,457,847,905]" box="[151,310,882,905]" pageId="5" pageNumber="292">
(
<figureCitation id="5FE82CB1592BFFD4E596B877FF3FFC70" box="[158,247,882,905]" captionStart="FIGURE 3" captionStartId="6.[151,244,1685,1707]" captionTargetBox="[230,1356,298,1664]" captionTargetId="figure@6.[230,1356,298,1664]" captionTargetPageId="6" captionText="FIGURE 3. Diplocirrus acafi sp. nov. (A) Holotype (UERJ 5573), oblique lateral view. (B) Same, chaetiger 10, notochaetae. (C) Same, neurochaetae. (D) Papillae capitate. Scale bars: A: 0.6 mm, B: 27 µm, C: 20 µm, D: 25 µm." httpUri="https://zenodo.org/record/236453/files/figure.png" pageId="5" pageNumber="292">Figure 3</figureCitation>
A–D)
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592BFFD4E59FB8BDFF08FB85" pageId="5" pageNumber="292" type="materials_examined">
<paragraph id="C76C3034592BFFD4E59FB8BDFF08FB85" blockId="5.[151,1436,952,2015]" pageId="5" pageNumber="292">
<emphasis id="F5A7EC26592BFFD4E59FB8BDFE59FC36" bold="true" box="[151,401,952,975]" pageId="5" pageNumber="292">
<typeStatus id="18688E96592BFFD4E59FB8BDFF07FC36" box="[151,207,952,975]" pageId="5" pageNumber="292">Type</typeStatus>
material.
<collectingCountry id="BFC470A4592BFFD4E44EB8BDFE44FC36" box="[326,396,952,975]" name="Brazil" pageId="5" pageNumber="292">Brasil</collectingCountry>
.
</emphasis>
Atlantic Ocean. Southeastern coast, Campos Basin.
<typeStatus id="18688E96592BFFD4E6C6B8BDFBFDFC36" box="[974,1077,952,975]" pageId="5" pageNumber="292" type="holotype">Holotype</typeStatus>
: (
<collectionCode id="A1C2A8F1592BFFD4E140B8BDFB43FC36" box="[1096,1163,952,975]" pageId="5" pageNumber="292">UERJ</collectionCode>
5573-HAB16 C3 R3),
<quantity id="002B9DD1592BFFD4E089B8BDFF66FC08" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="5" pageNumber="292" unit="m" value="77.0">77 m</quantity>
, 0
<date id="B36D16F4592BFFD4E5C1B8DFFE8FFC08" box="[201,327,986,1009]" pageId="5" pageNumber="292" value="2009-07-03">3 July 2009</date>
(
<geoCoordinate id="A2E756F3592BFFD4E45FB8DFFDDFFC08" box="[343,535,986,1009]" direction="south" orientation="latitude" pageId="5" pageNumber="292" precision="1" value="-22.780653">22º 46' 50,350&quot; S</geoCoordinate>
<geoCoordinate id="A2E756F3592BFFD4E728B8DFFD17FC08" box="[544,735,986,1009]" direction="west" orientation="longitude" pageId="5" pageNumber="292" precision="1" value="-41.060856">41º 3' 39,080&quot; W</geoCoordinate>
).
<typeStatus id="18688E96592BFFD4E7FBB8DFFC97FC08" box="[755,863,986,1009]" pageId="5" pageNumber="292" type="paratype">Paratypes</typeStatus>
: 1
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(
<collectionCode id="A1C2A8F1592BFFD4E6E3B8DFFBE6FC08" box="[1003,1070,986,1009]" pageId="5" pageNumber="292">UERJ</collectionCode>
5574-HAB8 A6 R2),
<quantity id="002B9DD1592BFFD4E010B8DFFABCFC08" box="[1304,1396,986,1009]" metricMagnitude="2" metricUnit="m" metricValue="4.008" pageId="5" pageNumber="292" unit="m" value="400.8">400.8 m</quantity>
, 0
<date id="B36D16F4592BFFD4E099B8DFFEFAFBED" pageId="5" pageNumber="292" value="2009-02-01">1 February 2009</date>
(
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<geoCoordinate id="A2E756F3592BFFD4E4F7B8F8FD0BFBED" box="[511,707,1021,1044]" direction="west" orientation="longitude" pageId="5" pageNumber="292" precision="1" value="-41.32988">41º 19' 47,574&quot; W</geoCoordinate>
), 1
<typeStatus id="18688E96592BFFD4E7E1B8F8FC8CFBED" box="[745,836,1021,1044]" pageId="5" pageNumber="292" type="paratype">paratype</typeStatus>
(
<collectionCode id="A1C2A8F1592BFFD4E65AB8F8FC5DFBED" box="[850,917,1021,1044]" pageId="5" pageNumber="292">UERJ</collectionCode>
5577-HAB13 FOZ 43 R2),
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,
<date id="B36D16F4592BFFD4E1F2B8F8FA53FBED" box="[1274,1435,1021,1044]" pageId="5" pageNumber="292" value="2009-03-15">15 March 2009</date>
(
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,
<geoCoordinate id="A2E756F3592BFFD4E462BF25FDFAFBCE" box="[362,562,1056,1079]" direction="west" orientation="longitude" pageId="5" pageNumber="292" precision="1" value="-40.537247">40º 32' 14,086&quot; W</geoCoordinate>
) 1
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(
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5576-HAB16 B4 R2),
<quantity id="002B9DD1592BFFD4E6F0BF25FBF7FBCE" box="[1016,1087,1056,1079]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="5" pageNumber="292" unit="m" value="107.0">107 m</quantity>
, 0
<date id="B36D16F4592BFFD4E150BF25FB53FBCE" box="[1112,1179,1056,1079]" pageId="5" pageNumber="292">2 July</date>
. 2009 (
<geoCoordinate id="A2E756F3592BFFD4E1E3BF25FA54FBCE" box="[1259,1436,1056,1079]" direction="south" orientation="latitude" pageId="5" pageNumber="292" precision="1" value="-23.166893">23º 10' 0,820&quot; S</geoCoordinate>
<geoCoordinate id="A2E756F3592BFFD4E59FBF47FE99FBA0" box="[151,337,1090,1113]" direction="west" orientation="longitude" pageId="5" pageNumber="292" precision="1" value="-41.05353">41º 3' 12,720&quot; W</geoCoordinate>
), 1
<typeStatus id="18688E96592BFFD4E472BF47FE1EFBA0" box="[378,470,1090,1113]" pageId="5" pageNumber="292" type="paratype">paratype</typeStatus>
(
<collectionCode id="A1C2A8F1592BFFD4E4ECBF47FDEDFBA0" box="[484,549,1090,1113]" pageId="5" pageNumber="292">UERJ</collectionCode>
5575-HAB17 FOZ 43 R2),
<quantity id="002B9DD1592BFFD4E65DBF47FC46FBA0" box="[853,910,1090,1113]" metricMagnitude="1" metricUnit="m" metricValue="9.7" pageId="5" pageNumber="292" unit="m" value="97.0">97 m</quantity>
,
<date id="B36D16F4592BFFD4E692BF47FBECFBA0" box="[922,1060,1090,1113]" pageId="5" pageNumber="292" value="2009-07-24">24 July 2009</date>
(
<geoCoordinate id="A2E756F3592BFFD4E13BBF47FB38FBA0" box="[1075,1264,1090,1113]" direction="south" orientation="latitude" pageId="5" pageNumber="292" precision="1" value="-22.20744">22º 12' 26,776&quot; S</geoCoordinate>
<geoCoordinate id="A2E756F3592BFFD4E1FFBF47FF7CFB85" direction="west" orientation="longitude" pageId="5" pageNumber="292" precision="1" value="-40.237495">40º 14' 14,985&quot; W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592BFFD4E5CFBF8DFBF2FB38" pageId="5" pageNumber="292" type="diagnosis">
<paragraph id="C76C3034592BFFD4E5CFBF8DFBF2FB38" blockId="5.[151,1436,952,2015]" pageId="5" pageNumber="292">
<emphasis id="F5A7EC26592BFFD4E5CFBF8DFEF4FB66" bold="true" box="[199,316,1160,1183]" pageId="5" pageNumber="292">Diagnosis.</emphasis>
Sediment particles of about the same size and homogeneously distributed throughout body, all papillae of the same length along body, 7 noto- and 5 neurochaetae per bundle throughout body.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592BFFD4E5CFBFC8FCFAF95E" pageId="5" pageNumber="292" type="description">
<paragraph id="C76C3034592BFFD4E5CFBFC8FCFAF95E" blockId="5.[151,1436,952,2015]" pageId="5" pageNumber="292">
<emphasis id="F5A7EC26592BFFD4E5CFBFC8FE9AFB1D" bold="true" box="[199,338,1229,1252]" pageId="5" pageNumber="292">Description.</emphasis>
<typeStatus id="18688E96592BFFD4E455BFC8FE08FB1D" box="[349,448,1229,1252]" pageId="5" pageNumber="292" type="holotype">Holotype</typeStatus>
complete; body cylindrical, anteriorly tapered, light brown;
<quantity id="002B9DD1592BFFD4E166BFC8FB79FB1D" box="[1134,1201,1229,1252]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="292" unit="mm" value="5.0">5 mm</quantity>
long,
<quantity id="002B9DD1592BFFD4E1F7BFC8FA9EFB1D" box="[1279,1366,1229,1252]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="292" unit="mm" value="0.4">0.4 mm</quantity>
wide, cephalic cage 0.7 long, for 29 chaetigers.
<typeStatus id="18688E96592BFFD4E752BFF5FD0AFAFE" box="[602,706,1264,1287]" pageId="5" pageNumber="292" type="paratype">Paratypes</typeStatus>
with 24–32 chaetigers,
<quantity id="002B9DD1592BFFD4E6CDBFF5FBFBFAFE" box="[965,1075,1264,1287]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="5.0" metricValueMin="3.2" pageId="5" pageNumber="292" unit="mm" value="4.1" valueMax="5.0" valueMin="3.2">3.2–5 mm</quantity>
long,
<quantity id="002B9DD1592BFFD4E170BFF5FB31FAFE" box="[1144,1273,1264,1287]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="292" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
wide, cephalic cage
<quantity id="002B9DD1592BFFD4E5DABE17FE9EFAD0" box="[210,342,1298,1321]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="292" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
long. Tunic with thin layer of fine sediment grains, papillate. Papillae ½ as long as neurochaetae, papillae capitate (longer than wide). Fine sediment particles on papillae basis only, papillae are not distributed homogeneously, more numerous on the anterior and posterior region and chaetal lobes, the same-sized throughout. Four rows of papillae per chaetiger. Cephalic hood not exposed. Anterior region not dissected to avoid damage. Cephalic cage chaetae as long as body width. Only notochaetae of chaetiger 1 form cephalic cage, chaetae directed dorsally. Seven notochaetae per bundle throughout, anterior dorsal margin of first chaetiger papillate, papillae similar to those along body. Some notochaetae twice as long as the others in the same bundle, 5 neurochaetae per bundle along body. Chaetigers all of same size. Ventrolateral gonopores absent. Chaetae emerge from body wall. Median notochaetae about ½ of body width. Notochaetae 1/6 longer than neurochaetae. All notochaetae multiarticulated capillaries. Neurochaetae multiarticulated capillaries only in first chaetiger; from chaetiger 2 to end of body, chaetae thicker and falcate. All articles of noto- and neurochaetae of about same length. Neurochaetae thicker than notochaetae and of same thickness along capillary length. Posterior region tapered; terminal anus.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592BFFD4E5CFBDB7FADBF845" pageId="5" pageNumber="292" type="discussion">
<paragraph id="C76C3034592BFFD4E5CFBDB7FADBF845" blockId="5.[151,1436,952,2015]" pageId="5" pageNumber="292">
<emphasis id="F5A7EC26592BFFD4E5CFBDB7FEFDF930" bold="true" box="[199,309,1714,1737]" pageId="5" pageNumber="292">Remarks.</emphasis>
<taxonomicName id="00D34BB7592BFFD4E434BDB7FE3CF930" box="[316,500,1714,1737]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="acafi" status="sp. nov.">
<emphasis id="F5A7EC26592BFFD4E434BDB7FE3CF930" box="[316,500,1714,1737]" italics="true" pageId="5" pageNumber="292">Diplocirrus acafi</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592BFFD4E4F3BDB7FD98F930" bold="true" box="[507,592,1714,1737]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="EE94515D592BFFD4E4F3BDB7FD98F930" box="[507,592,1714,1737]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="00D34BB7592BFFD4E78BBDB7FC25F930" authority="Hartman, 1965" authorityName="Hartman" authorityYear="1965" box="[643,1005,1714,1737]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="octobranchus">
<emphasis id="F5A7EC26592BFFD4E78BBDB7FCFDF930" box="[643,821,1714,1737]" italics="true" pageId="5" pageNumber="292">D. octobranchus</emphasis>
(Hartman, 1965)
</taxonomicName>
have fine sediment particles only on the base of the papillae, absence of ventrolateral gonopores, and only notochaetae of chaetiger 1 make up the cephalic cage. However,
<taxonomicName id="00D34BB7592BFFD4E402BDFDFE0FF8F6" box="[266,455,1784,1807]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="acafi" status="sp. nov.">
<emphasis id="F5A7EC26592BFFD4E402BDFDFE0FF8F6" box="[266,455,1784,1807]" italics="true" pageId="5" pageNumber="292">Diplocirrus acafi</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592BFFD4E4DBBDFDFDE5F8F6" bold="true" box="[467,557,1784,1807]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="EE94515D592BFFD4E4DBBDFDFDE5F8F6" box="[467,557,1784,1807]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has sediment of the same size homogeneously distributed, papillae in greater number in the anterior and posterior regions and in chaetal lobes, all papillae of about the same size along the body, 4 rows per chaetiger, 7 noto- and 5 neurochaetae per bundle throughout the body.
<taxonomicName id="00D34BB7592BFFD4E129BC38FAF2F8AD" box="[1057,1338,1853,1876]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="octobranchus">
<emphasis id="F5A7EC26592BFFD4E129BC38FAF2F8AD" box="[1057,1338,1853,1876]" italics="true" pageId="5" pageNumber="292">Diplocirrus octobranchus</emphasis>
</taxonomicName>
presents sediment particles composed of a few larger sand grains, especially on the dorsum, papillae of varying lengths, longer dorsally and on chaetal lobes, may be as long as chaetae, shorter in the rest of the body, 4–5 rows per chaetiger, 4–5 notochaetae and 2–8 neurochaetae in the first chaetiger, 7 noto- and 2–5 neurochaetae in the median region.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592BFFD7E5CFBCCDFF29FF28" lastPageId="6" lastPageNumber="293" pageId="5" pageNumber="292" type="etymology">
<paragraph id="C76C3034592BFFD7E5CFBCCDFF29FF28" blockId="5.[151,1436,952,2015]" lastBlockId="6.[151,1435,152,244]" lastPageId="6" lastPageNumber="293" pageId="5" pageNumber="292">
<emphasis id="F5A7EC26592BFFD4E5CFBCCDFE8FF826" bold="true" box="[199,327,1992,2015]" pageId="5" pageNumber="292">Etymology.</emphasis>
The species epithet,
<taxonomicName id="00D34BB7592BFFD4E731BCCDFDA6F826" box="[569,622,1992,2015]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="292" phylum="Annelida" rank="species" species="acafi">
<emphasis id="F5A7EC26592BFFD4E731BCCDFDA6F826" box="[569,622,1992,2015]" italics="true" pageId="5" pageNumber="292">acafi</emphasis>
</taxonomicName>
, is a composition from Dr.
<emphasis id="F5A7EC26592BFFD4E6A5BCCDFC08F826" bold="true" box="[941,960,1992,2015]" pageId="5" pageNumber="292">A</emphasis>
ntonio
<emphasis id="F5A7EC26592BFFD4E107BCCDFBF8F826" bold="true" box="[1039,1072,1992,2015]" pageId="5" pageNumber="292">Ca</emphasis>
rlos de
<emphasis id="F5A7EC26592BFFD4E18FBCCDFB5FF826" bold="true" box="[1159,1175,1992,2015]" pageId="5" pageNumber="292">F</emphasis>
re
<emphasis id="F5A7EC26592BFFD4E1A4BCCDFB7BF826" bold="true" box="[1196,1203,1992,2015]" pageId="5" pageNumber="292">i</emphasis>
tas, whose species is named, for his friendship and contribution to the world of nature photography, including the new species described herein.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF5928FFD7E5CFBBD8FAD1FF0D" box="[199,1305,221,244]" pageId="6" pageNumber="293" type="distribution">
<paragraph id="C76C30345928FFD7E5CFBBD8FAD1FF0D" blockId="6.[151,1435,152,244]" box="[199,1305,221,244]" pageId="6" pageNumber="293">
<emphasis id="F5A7EC265928FFD7E5CFBBD8FE91FF0D" bold="true" box="[199,345,221,244]" pageId="6" pageNumber="293">Distribution.</emphasis>
Southeastern Atlantic coast, Campos Basin,
<collectingCountry id="BFC470A45928FFD7E630BBD8FCB4FF0D" box="[824,892,221,244]" name="Brazil" pageId="6" pageNumber="293">Brazil</collectingCountry>
, in soft bottom from
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depth.
</paragraph>
</subSubSection>
</treatment>
</document>

243
data/4F/7A/81/4F7A8122592CFFD5E59FBE8FFAC8FE58.xml Normal file
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@ -0,0 +1,243 @@
<document id="D06BF78486DE1409B146F65B21E4F902" ID-DOI="10.11646/zootaxa.4027.2.8" ID-GBIF-Dataset="8dcb1038-d556-49c8-8649-50202fa770f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="236450" ID-ZooBank="9096376A-C64F-46D7-8E75-55B56E495A4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461282324405" checkinUser="plazi" docAuthor="Teixeira, Juliana Henrique, Rizzo, Alexandra Elaine &amp; Santos, Cinthya Simone Gomes" docDate="2015" docId="4F7A8122592CFFD5E59FBE8FFAC8FE58" docLanguage="en" docName="zt04027p295.pdf" docOrigin="Zootaxa 4027 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diplocirrus rugosus Teixeira, Rizzo &amp; Santos, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="291" masterDocId="B343F95A592EFFD1E508BB05FFC8FFF9" masterDocTitle="Three new species of Diplocirrus Haase, 1915 (Polychaeta: Flabelligeridae) from Campos Basin in SE Brazil" masterLastPageNumber="295" masterPageNumber="287" pageNumber="289" updateTime="1698614352147" updateUser="plazi">
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<mods:title id="6C9D5B32347186590E946902466AF78F">Three new species of Diplocirrus Haase, 1915 (Polychaeta: Flabelligeridae) from Campos Basin in SE Brazil</mods:title>
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<mods:namePart id="4E4FBD10A173F70F53497ED3130FF97C">Teixeira, Juliana Henrique</mods:namePart>
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<mods:namePart id="71F6A8CA4289BFA81B6C24AA6CD9FF9C">Rizzo, Alexandra Elaine</mods:namePart>
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<mods:namePart id="54652C9EF29D6B6ADA99A83938B74D9F">Santos, Cinthya Simone Gomes</mods:namePart>
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<subSubSection id="8FC963BF592CFFD3E59FBE8FFEFBFA3D" pageId="2" pageNumber="289" type="nomenclature">
<paragraph id="C76C3034592CFFD3E59FBE8FFE25FA5C" blockId="2.[151,493,1418,1476]" box="[151,493,1418,1445]" pageId="2" pageNumber="289">
<heading id="9C248758592CFFD3E59FBE8FFE25FA5C" bold="true" box="[151,493,1418,1445]" fontSize="11" level="1" pageId="2" pageNumber="289" reason="1">
<emphasis id="F5A7EC26592CFFD3E59FBE8FFE25FA5C" bold="true" box="[151,493,1418,1445]" pageId="2" pageNumber="289">
<taxonomicName id="00D34BB7592CFFD3E59FBE8FFE40FA5D" ID-CoL="36JVY" box="[151,392,1418,1444]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="289" phylum="Annelida" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="F5A7EC26592CFFD3E59FBE8FFE40FA5D" bold="true" box="[151,392,1418,1444]" italics="true" pageId="2" pageNumber="289">Diplocirrus rugosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94515D592CFFD3E498BE8EFE25FA5C" box="[400,493,1419,1445]" pageId="2" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C76C3034592CFFD3E59FBEA8FEFBFA3D" blockId="2.[151,493,1418,1476]" box="[151,307,1453,1476]" pageId="2" pageNumber="289">
(
<figureCitation id="5FE82CB1592CFFD3E596BEA8FF3DFA3D" box="[158,245,1453,1476]" captionStart="FIGURE 1" captionStartId="3.[151,244,1792,1814]" captionTargetBox="[231,1351,402,1768]" captionTargetId="figure@3.[231,1355,402,1768]" captionTargetPageId="3" captionText="FIGURE 1. Diplocirrus rugosus sp. nov. (A) Holotype (UERJ 5570), ventral view. (B) Same, oblique lateral view. (C) Same, chaetiger 10, notochaetae (inset: basal and medial chaetal regions). (D) Same, one neurochaeta. (E) Capitate papillae. Scale bars: A – B: 0.5 mm, C: 45 µm, D-E: 30 µm." httpUri="https://zenodo.org/record/236451/files/figure.png" pageId="2" pageNumber="289">Figure 1</figureCitation>
A–E)
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592CFFD3E59FBEF7FDA7F9B6" pageId="2" pageNumber="289" type="materials_examined">
<paragraph id="C76C3034592CFFD3E59FBEF7FDA7F9B6" blockId="2.[151,1436,1522,2031]" pageId="2" pageNumber="289">
<emphasis id="F5A7EC26592CFFD3E59FBEF7FEF6F9F0" bold="true" box="[151,318,1522,1545]" pageId="2" pageNumber="289">
<typeStatus id="18688E96592CFFD3E59FBEF7FF07F9F0" box="[151,207,1522,1545]" pageId="2" pageNumber="289">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="BFC470A4592CFFD3E44EBEF7FE42F9F0" box="[326,394,1522,1545]" name="Brazil" pageId="2" pageNumber="289">Brazil</collectingCountry>
. Atlantic Ocean. Southeastern coast, Campos Basin.
<typeStatus id="18688E96592CFFD3E6C2BEF7FBF9F9F0" box="[970,1073,1522,1545]" pageId="2" pageNumber="289" type="holotype">Holotype</typeStatus>
: (
<collectionCode id="A1C2A8F1592CFFD3E14CBEF7FB4FF9F0" box="[1092,1159,1522,1545]" pageId="2" pageNumber="289">UERJ</collectionCode>
5570-HAB
<date id="B36D16F4592CFFD3E1F6BEF7FAFDF9F0" box="[1278,1333,1522,1545]" pageId="2" pageNumber="289" value="1917-01-05">17 I5</date>
R3),
<quantity id="002B9DD1592CFFD3E07BBEF7FF66F9D5" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="289" unit="m" value="140.0">140 m</quantity>
,
<date id="B36D16F4592CFFD3E5B2BD10FE8AF9D5" box="[186,322,1557,1580]" pageId="2" pageNumber="289" value="2009-07-21">21 Jul. 2009</date>
, (
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,
<geoCoordinate id="A2E756F3592CFFD3E72ABD10FD24F9D5" box="[546,748,1557,1580]" direction="west" orientation="longitude" pageId="2" pageNumber="289" precision="1" value="-40.254368">40º 15' 15.721&quot; W</geoCoordinate>
).
<typeStatus id="18688E96592CFFD3E608BD10FCA9F9D5" box="[768,865,1557,1580]" pageId="2" pageNumber="289" type="paratype">Paratype</typeStatus>
: (
<collectionCode id="A1C2A8F1592CFFD3E67CBD10FC70F9D5" box="[884,952,1557,1580]" pageId="2" pageNumber="289">UERJ</collectionCode>
5571-HAB
<date id="B36D16F4592CFFD3E126BD10FBACF9D5" box="[1070,1124,1557,1580]" pageId="2" pageNumber="289" value="2013-01-05">13 I5</date>
R2; 43 inds),
<quantity id="002B9DD1592CFFD3E008BD10FA8FF9D5" box="[1280,1351,1557,1580]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="2" pageNumber="289" unit="m" value="147.0">147 m</quantity>
, 0
<date id="B36D16F4592CFFD3E069BD10FF04F9B6" pageId="2" pageNumber="289" value="2009-07-06">6 Jul. 2009</date>
(
<geoCoordinate id="A2E756F3592CFFD3E5D3BD3DFE5DF9B6" box="[219,405,1592,1615]" direction="south" orientation="latitude" pageId="2" pageNumber="289" precision="1" value="-21.382738">21º 22' 57.858&quot; S</geoCoordinate>
<geoCoordinate id="A2E756F3592CFFD3E493BD3DFDABF9B6" box="[411,611,1592,1615]" direction="west" orientation="longitude" pageId="2" pageNumber="289" precision="1" value="-40.254345">40º 15' 15.648&quot; W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592CFFD3E5CFBD5FFD4DF96D" pageId="2" pageNumber="289" type="diagnosis">
<paragraph id="C76C3034592CFFD3E5CFBD5FFD4DF96D" blockId="2.[151,1436,1522,2031]" pageId="2" pageNumber="289">
<emphasis id="F5A7EC26592CFFD3E5CFBD5FFEFCF988" bold="true" box="[199,308,1626,1649]" pageId="2" pageNumber="289">Diagnosis</emphasis>
. Sediment particles dispersed, body papillae abundant, twice as long in chaetal lobes, 4–5 notochaetae per ramus, 3–4 neurochaetae throughout the body.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592CFFD2E5CFBDA5FDC2FEA5" lastPageId="3" lastPageNumber="290" pageId="2" pageNumber="289" type="description">
<paragraph id="C76C3034592CFFD2E5CFBDA5FDC2FEA5" blockId="2.[151,1436,1522,2031]" lastBlockId="3.[151,1436,152,348]" lastPageId="3" lastPageNumber="290" pageId="2" pageNumber="289">
<emphasis id="F5A7EC26592CFFD3E5CFBDA5FE83F94E" bold="true" box="[199,331,1696,1719]" pageId="2" pageNumber="289">Description</emphasis>
.
<typeStatus id="18688E96592CFFD3E45FBDA5FE72F94E" box="[343,442,1696,1719]" pageId="2" pageNumber="289" type="holotype">Holotype</typeStatus>
complete (HAB
<date id="B36D16F4592CFFD3E764BDA5FD69F94E" box="[620,673,1696,1719]" pageId="2" pageNumber="289" value="1917-01-05">17 I5</date>
R3;
<figureCitation id="5FE82CB1592CFFD3E7DCBDA5FCC7F94E" box="[724,783,1696,1719]" captionStart="FIGURE 1" captionStartId="3.[151,244,1792,1814]" captionTargetBox="[231,1351,402,1768]" captionTargetId="figure@3.[231,1355,402,1768]" captionTargetPageId="3" captionText="FIGURE 1. Diplocirrus rugosus sp. nov. (A) Holotype (UERJ 5570), ventral view. (B) Same, oblique lateral view. (C) Same, chaetiger 10, notochaetae (inset: basal and medial chaetal regions). (D) Same, one neurochaeta. (E) Capitate papillae. Scale bars: A – B: 0.5 mm, C: 45 µm, D-E: 30 µm." httpUri="https://zenodo.org/record/236451/files/figure.png" pageId="2" pageNumber="289">Fig. 1</figureCitation>
A),
<quantity id="002B9DD1592CFFD3E633BDA5FC44F94E" box="[827,908,1696,1719]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="289" unit="mm" value="7.3">7.3 mm</quantity>
long,
<quantity id="002B9DD1592CFFD3E6C6BDA5FBC3F94E" box="[974,1035,1696,1719]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="289" unit="mm" value="1.0">1 mm</quantity>
wide, cephalic cage
<quantity id="002B9DD1592CFFD3E1E1BDA5FAF2F94E" box="[1257,1338,1696,1719]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="289" unit="mm" value="2.5">2.5 mm</quantity>
long, for 32 chaetigers. Body cylindrical, swollen anteriorly, yellowish brown in color.
<typeStatus id="18688E96592CFFD3E6C7BDC7FBFFF920" box="[975,1079,1730,1753]" pageId="2" pageNumber="289" type="paratype">Paratypes</typeStatus>
complete,
<quantity id="002B9DD1592CFFD3E1A3BDC7FAE2F920" box="[1195,1322,1730,1753]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="7.3" metricValueMin="1.4" pageId="2" pageNumber="289" unit="mm" value="4.35" valueMax="7.3" valueMin="1.4">1.4–7.3 mm</quantity>
long,
<quantity id="002B9DD1592CFFD3E064BDC7FF23F905" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="2" pageNumber="289" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4– 0.7 mm</quantity>
wide, cephalic cage
<quantity id="002B9DD1592CFFD3E4DDBDE0FD9FF905" box="[469,599,1765,1788]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="2.5" metricValueMin="0.3" pageId="2" pageNumber="289" unit="mm" value="1.4" valueMax="2.5" valueMin="0.3">0.3–2.5 mm</quantity>
long, for 20–35 chaetigers. Tunic with a thin layer of fine sediment grains, papillated. Papillae short relative to size of chaetae, papillae capitate. Fine sediment particles on papillae basis only, papillae ¼ as long as notochaetae, about eight transverse rows in chaetiger 10, papillae twice as long as chaetal lobes, with same size throughout body. Cephalic hood exposed in one specimen; palps and branchiae lost; prostomium low cone; eyes not observed. Caruncle poorly developed, not reaching posterior margin of branchial plate, lateral ridges low, median keel not projected. Palps long, thick; palp keels rounded, reduced. Lateral lips larger, thick, dorsal lip smaller, rhomboid, ventral lip reduced, rounded. Cephalic cage chaetae as long as body width. Only notochaetae of chaetiger 1 present in cephalic cage, chaetae directed dorsally. 4–5 notochaetae per ramus throughout body, anterior dorsal margin of first chaetiger papillated, papillae similar to those along body. Chaetigers all of the same length. 3–4 neurochaetae per bundle throughout body. Ventrolateral gonopores absent. Chaetae emerge from body wall. Noto- and neurochaetae of same size from chaetiger 2, notochaetae as long as ½ body width and neurochaetae ¼ as long as notochaetae. All noto- and neurochaetae multiarticulated. Notochaetae with articles longer basally, medium-sized medially and short distally; neurochaetae with all articles of same-size, tips falcate, smooth. Both chaetae tapering distally. Posterior region tapers into a rounded lobe, terminal anus.
</paragraph>
</subSubSection>
<caption id="93AC60BC592DFFD2E59FBC05FE0FF8A9" httpUri="https://zenodo.org/record/236451/files/figure.png" pageId="3" pageNumber="290" targetBox="[231,1351,402,1768]" targetPageId="3">
<paragraph id="C76C3034592DFFD2E59FBC05FE0FF8A9" blockId="3.[151,1437,1791,1872]" pageId="3" pageNumber="290">
<emphasis id="F5A7EC26592DFFD2E59FBC05FED9F8EF" bold="true" box="[151,273,1792,1814]" pageId="3" pageNumber="290">FIGURE 1.</emphasis>
<taxonomicName id="00D34BB7592DFFD2E414BDFAFE2AF8EC" box="[284,482,1791,1813]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="290" phylum="Annelida" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="F5A7EC26592DFFD2E414BDFAFE2AF8EC" box="[284,482,1791,1813]" italics="true" pageId="3" pageNumber="290">Diplocirrus rugosus</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592DFFD2E4E5BC05FDF6F8EF" bold="true" box="[493,574,1792,1814]" pageId="3" pageNumber="290">
<taxonomicNameLabel id="EE94515D592DFFD2E4E5BC05FDF6F8EF" box="[493,574,1792,1814]" pageId="3" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) Holotype (UERJ5570), ventral view. (B) Same, oblique lateral view. (C) Same, chaetiger 10, notochaetae (inset: basal and medial chaetal regions). (D) Same, one neurochaeta. (E) Capitate papillae. Scale bars: A–B: 0.5 mm,C: 45 µm, D-E: 30 µm.
</paragraph>
</caption>
<subSubSection id="8FC963BF592DFFD5E5CFBC73FC5DFEC0" lastPageId="4" lastPageNumber="291" pageId="3" pageNumber="290" type="discussion">
<paragraph id="C76C3034592DFFD5E5CFBC73FC5DFEC0" blockId="3.[151,1436,1910,2037]" lastBlockId="4.[151,1436,152,417]" lastPageId="4" lastPageNumber="291" pageId="3" pageNumber="290">
<emphasis id="F5A7EC26592DFFD2E5CFBC73FEFDF874" bold="true" box="[199,309,1910,1933]" pageId="3" pageNumber="290">Remarks.</emphasis>
<taxonomicName id="00D34BB7592DFFD2E448BC73FDD3F874" box="[320,539,1910,1933]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="290" phylum="Annelida" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="F5A7EC26592DFFD2E448BC73FDD3F874" box="[320,539,1910,1933]" italics="true" pageId="3" pageNumber="290">Diplocirrus rugosus</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592DFFD2E72FBC73FD48F874" bold="true" box="[551,640,1910,1933]" pageId="3" pageNumber="290">
<taxonomicNameLabel id="EE94515D592DFFD2E72FBC73FD48F874" box="[551,640,1910,1933]" pageId="3" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="00D34BB7592DFFD2E615BC73FC51F874" box="[797,921,1910,1933]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="290" phylum="Annelida" rank="genus">
<emphasis id="F5A7EC26592DFFD2E615BC73FC51F874" box="[797,921,1910,1933]" italics="true" pageId="3" pageNumber="290">Diplocirrus</emphasis>
</taxonomicName>
sp. from
<collectingCountry id="BFC470A4592DFFD2E104BC73FBA4F874" box="[1036,1132,1910,1933]" name="Morocco" pageId="3" pageNumber="290">Morocco</collectingCountry>
because both species have anterior chaetigers of similar size to those present in the posterior ones, body coated with sediment and many short papillae, giving a velvety appearance. However,
<taxonomicName id="00D34BB7592DFFD2E794BCBEFCBBF82B" box="[668,883,1979,2002]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="290" phylum="Annelida" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="F5A7EC26592DFFD2E794BCBEFCBBF82B" box="[668,883,1979,2002]" italics="true" pageId="3" pageNumber="290">Diplocirrus rugosus</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592DFFD2E671BCBEFC07F82B" bold="true" box="[889,975,1979,2002]" pageId="3" pageNumber="290">
<taxonomicNameLabel id="EE94515D592DFFD2E671BCBEFC07F82B" box="[889,975,1979,2002]" pageId="3" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="00D34BB7592DFFD2E152BCBEFB1EF82B" box="[1114,1238,1979,2002]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="290" phylum="Annelida" rank="genus">
<emphasis id="F5A7EC26592DFFD2E152BCBEFB1EF82B" box="[1114,1238,1979,2002]" italics="true" pageId="3" pageNumber="290">Diplocirrus</emphasis>
</taxonomicName>
sp. from
<collectingCountry id="BFC470A4592DFFD2E033BCBEFA53F82B" box="[1339,1435,1979,2002]" name="Morocco" pageId="3" pageNumber="290">Morocco</collectingCountry>
because the first has dispersed sediment particles and papillae twice as long as chaetal lobes, while the second has abundant sediment particles forming a thin layer and short papillae laterally.
<taxonomicName id="00D34BB7592AFFD5E6D3BB9DFB7BFF56" box="[987,1203,152,175]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="291" phylum="Annelida" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="F5A7EC26592AFFD5E6D3BB9DFB7BFF56" box="[987,1203,152,175]" italics="true" pageId="4" pageNumber="291">Diplocirrus rugosus</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592AFFD5E1B4BB9DFADBFF56" bold="true" box="[1212,1299,152,175]" pageId="4" pageNumber="291">
<taxonomicNameLabel id="EE94515D592AFFD5E1B4BB9DFADBFF56" box="[1212,1299,152,175]" pageId="4" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="00D34BB7592AFFD5E59FBBBFFE7FFF28" box="[151,439,186,209]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="291" phylum="Annelida" rank="species" species="salazarvallejoi" status="sp. nov.">
<emphasis id="F5A7EC26592AFFD5E59FBBBFFE7FFF28" box="[151,439,186,209]" italics="true" pageId="4" pageNumber="291">Diplocirrus salazarvallejoi</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592AFFD5E4B6BBBFFDDBFF28" bold="true" box="[446,531,186,209]" pageId="4" pageNumber="291">
<taxonomicNameLabel id="EE94515D592AFFD5E4B6BBBFFDDBFF28" box="[446,531,186,209]" pageId="4" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="00D34BB7592AFFD5E74DBBBFFD34FF28" box="[581,764,186,209]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="291" phylum="Annelida" rank="species" species="acafi" status="sp. nov.">
<emphasis id="F5A7EC26592AFFD5E74DBBBFFD34FF28" box="[581,764,186,209]" italics="true" pageId="4" pageNumber="291">Diplocirrus acafi</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592AFFD5E60BBBBFFC90FF28" bold="true" box="[771,856,186,209]" pageId="4" pageNumber="291">
<taxonomicNameLabel id="EE94515D592AFFD5E60BBBBFFC90FF28" box="[771,856,186,209]" pageId="4" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
because the first has body wall with sand particles and the others not. Furthermore,
<taxonomicName id="00D34BB7592AFFD5E4C5BBD8FD6DFF0D" box="[461,677,221,244]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="291" phylum="Annelida" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="F5A7EC26592AFFD5E4C5BBD8FD6DFF0D" box="[461,677,221,244]" italics="true" pageId="4" pageNumber="291">Diplocirrus rugosus</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592AFFD5E7A5BBD8FCCCFF0D" bold="true" box="[685,772,221,244]" pageId="4" pageNumber="291">
<taxonomicNameLabel id="EE94515D592AFFD5E7A5BBD8FCCCFF0D" box="[685,772,221,244]" pageId="4" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has papillae twice as long as chaetal lobes while
<taxonomicName id="00D34BB7592AFFD5E017BBD8FEFDFEEE" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="291" phylum="Annelida" rank="species" species="salazarvallejoi" status="sp. nov.">
<emphasis id="F5A7EC26592AFFD5E017BBD8FEFDFEEE" italics="true" pageId="4" pageNumber="291">Diplocirrus salazarvallejoi</emphasis>
</taxonomicName>
<emphasis id="F5A7EC26592AFFD5E434BA05FE59FEEE" bold="true" box="[316,401,256,279]" pageId="4" pageNumber="291">
<taxonomicNameLabel id="EE94515D592AFFD5E434BA05FE59FEEE" box="[316,401,256,279]" pageId="4" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
described below has all papillae of same size on ventral and dorsal regions and
<taxonomicName id="00D34BB7592AFFD5E1ECBA05FA53FEEE" box="[1252,1435,256,279]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="291" phylum="Annelida" rank="species" species="acafi" status="sp. nov.">
<emphasis id="F5A7EC26592AFFD5E1ECBA05FA53FEEE" box="[1252,1435,256,279]" italics="true" pageId="4" pageNumber="291">Diplocirrus acafi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94515D592AFFD5E59FBA27FF24FEC0" box="[151,236,290,313]" pageId="4" pageNumber="291" rank="species">
<emphasis id="F5A7EC26592AFFD5E59FBA27FF24FEC0" bold="true" box="[151,236,290,313]" pageId="4" pageNumber="291">sp. nov.</emphasis>
</taxonomicNameLabel>
described below has all papillae of the same length along body.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592AFFD5E5CFBA40FC81FE86" pageId="4" pageNumber="291" type="etymology">
<paragraph id="C76C3034592AFFD5E5CFBA40FC81FE86" blockId="4.[151,1436,152,417]" pageId="4" pageNumber="291">
<emphasis id="F5A7EC26592AFFD5E5CFBA40FE8FFEA5" bold="true" box="[199,327,325,348]" pageId="4" pageNumber="291">Etymology.</emphasis>
The epithet of this new species refers to the presence of short papillae, which, when viewed under a light microscope, give a rugose or rough appearance to the body.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592AFFD5E5CFBA8FFAC8FE58" box="[199,1280,394,417]" pageId="4" pageNumber="291" type="distribution">
<paragraph id="C76C3034592AFFD5E5CFBA8FFAC8FE58" blockId="4.[151,1436,152,417]" box="[199,1280,394,417]" pageId="4" pageNumber="291">
<emphasis id="F5A7EC26592AFFD5E5CFBA8FFE91FE58" bold="true" box="[199,345,394,417]" pageId="4" pageNumber="291">Distribution.</emphasis>
Southeastern Atlantic coast, Campos Basin,
<collectingCountry id="BFC470A4592AFFD5E630BA8FFCB4FE58" box="[824,892,394,417]" name="Brazil" pageId="4" pageNumber="291">Brazil</collectingCountry>
, in soft bottom from
<quantity id="002B9DD1592AFFD5E155BA8FFB5BFE58" box="[1117,1171,394,417]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="291" unit="m" value="52.0">52 m</quantity>
to
<specimenCount id="D1D5FBBD592AFFD5E1BDBA8FFAC8FE58" box="[1205,1280,394,417]" pageId="4" pageNumber="291" type="generic">
<quantity id="002B9DD1592AFFD5E1BDBA8FFB34FE58" box="[1205,1276,394,417]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="4" pageNumber="291" unit="m" value="147.0">147 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E5F1DC256FFEAC51F8C076A3D3A052E0">Three new species of Diplocirrus Haase, 1915 (Polychaeta: Flabelligeridae) from Campos Basin in SE Brazil</mods:title>
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<taxonomicName id="00D34BB7592FFFD0E59FB8B1FE76FC36" ID-CoL="84N8G" authority="Haase, 1915" authorityName="Haase" authorityYear="1915" box="[151,446,948,975]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="288" phylum="Annelida" rank="genus">
<emphasis id="F5A7EC26592FFFD0E59FB8B1FE76FC36" bold="true" box="[151,446,948,975]" pageId="1" pageNumber="288">
<emphasis id="F5A7EC26592FFFD0E59FB8B1FEEAFC37" bold="true" box="[151,290,948,974]" italics="true" pageId="1" pageNumber="288">Diplocirrus</emphasis>
<bibRefCitation id="A3424DC5592FFFD0E420B8B0FE76FC36" author="Haase" box="[296,446,949,975]" pageId="1" pageNumber="288" refString="Haase, P. (1915) Boreale und arktisch Chloraemiden. Wissenschaftliche Meeresuntersuchungen der Kommission zur Wissenschaftlichen Untersuchung der Deutschen Meere, Neue Folge, Kiel, 17, 169 - 228." type="journal article" year="1915">Haase, 1915</bibRefCitation>
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<emphasis id="F5A7EC26592FFFD0E59FB8FFFF34FBE8" box="[151,252,1018,1041]" italics="true" pageId="1" pageNumber="288">Bradiella</emphasis>
<treatmentCitation id="46721625592FFFD0E40AB8FFFE08FBE8" author="Rullier" box="[258,448,1018,1041]" page="188" pageId="1" pageNumber="288" year="1965">
<bibRefCitation id="A3424DC5592FFFD0E40AB8FFFE5AFBE8" author="Rullier" box="[258,402,1018,1041]" pageId="1" pageNumber="288" refString="Rullier, F. (1965) Contribution a la faune des annelides polychetes de l'Australie. University of Queensland Papers, Department of Zoology, 2 (9), 163 - 201." type="journal article" year="1965">Rullier, 1965</bibRefCitation>
:188
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.
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<paragraph id="C76C3034592FFFD0E59FBF18FD9BFBCD" blockId="1.[151,948,1018,1111]" box="[151,595,1053,1076]" pageId="1" pageNumber="288">
<taxonomicName id="00D34BB7592FFFD0E59FBF18FD87FBCD" ID-CoL="152b707b-57bf-480a-af80-bb9bd7b09e0b" authority="Buzhinskaja, 1993:231" authorityName="Buzhinskaja" authorityPageNumber="231" authorityYear="1993" box="[151,591,1053,1076]" class="Polychaeta" family="Flabelligeridae" genus="Diversibranchius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="288" phylum="Annelida" rank="genus">
<emphasis id="F5A7EC26592FFFD0E59FBF18FE98FBCD" box="[151,336,1053,1076]" italics="true" pageId="1" pageNumber="288">Diversibranchius</emphasis>
Buzhinskaja, 1993:231
</taxonomicName>
.
</paragraph>
<paragraph id="C76C3034592FFFD0E59FBF45FC7CFBAE" blockId="1.[151,948,1018,1111]" box="[151,948,1088,1111]" pageId="1" pageNumber="288">
<emphasis id="F5A7EC26592FFFD0E59FBF45FEE5FBAE" bold="true" box="[151,301,1088,1111]" pageId="1" pageNumber="288">Type-species.</emphasis>
<taxonomicName id="00D34BB7592FFFD0E43BBF45FD63FBAE" ID-CoL="7D995" authority="Malmgren, 1867" authorityName="Malmgren" authorityYear="1867" box="[307,683,1088,1111]" class="Polychaeta" family="Flabelligeridae" genus="Trophonia" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="288" phylum="Annelida" rank="species" species="glauca">
<emphasis id="F5A7EC26592FFFD0E43BBF45FE39FBAE" box="[307,497,1088,1111]" italics="true" pageId="1" pageNumber="288">Trophonia glauca</emphasis>
<bibRefCitation id="A3424DC5592FFFD0E4FFBF45FD63FBAE" author="Malmgren" box="[503,683,1088,1111]" pageId="1" pageNumber="288" refString="Malmgren, A. J. (1867) Annulata Polychaeta: Spetsbergiae, Groenlandiae, Islandiae et Scandinaviae hactenus cognita. Kongelige Vetenskaps-Akademiens Forhandlingar, 4, 127 - 235." type="journal article" year="1867">Malmgren, 1867</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592FFFD0E59FBF80FD92FAB0" pageId="1" pageNumber="288" type="diagnosis">
<paragraph id="C76C3034592FFFD0E59FBF80FD92FAB0" blockId="1.[151,1436,1157,1353]" pageId="1" pageNumber="288">
<emphasis id="F5A7EC26592FFFD0E59FBF80FECDFB65" bold="true" box="[151,261,1157,1180]" pageId="1" pageNumber="288">Diagnosis</emphasis>
(from
<bibRefCitation id="A3424DC5592FFFD0E454BF80FCC8FB65" author="Salazar-Vallejo" box="[348,768,1157,1180]" pageId="1" pageNumber="288" refString="Salazar-Vallejo, S. I. &amp; Buzhinskaja, G. (2011) Revision of Diplocirrus Haase, 1915, including Bradiella Rullier, 1965, and Diversibranchius Buzhinskaja, 1993 (Polychaeta, Flabelligeridae). ZooKeys, 106, 1 - 45. http: // dx. doi. org / 10.3897 / zookeys. 106.795" type="journal article" year="2011">Salazar-Vallejo &amp; Buzhinskaja, 2011</bibRefCitation>
). Body clavate or subcylindrical, often anteriorly swollen. Cephalic cage variably developed. Body papillae abundant and short, giving a velvety appearance, or very long, giving a hirsute outlook, sometimes adhering sediment particles. All chaetae multiarticulated capillaries; thick, sometimes falcate neurochaetae. Branchiae sessile, 4 pairs, distal branchiae thicker, often shorter, proximal branchiae thinner, often longer, sometimes basally lamellate. Gonopodial papillae present in chaetiger 4 or 5, or a series of paired ventrolateral gonopores along some anterior chaetigers.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="B342C9C6DDE5EA4EA1E09F36DA8305C1">Three new species of Diplocirrus Haase, 1915 (Polychaeta: Flabelligeridae) from Campos Basin in SE Brazil</mods:title>
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<mods:namePart id="51188EDB55118D465647346E7F80D68D">Santos, Cinthya Simone Gomes</mods:namePart>
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<heading id="9C248758592FFFD0E59FBE9EFA8AFA4C" bold="true" box="[151,1346,1434,1461]" fontSize="11" level="1" pageId="1" pageNumber="288" reason="1">
<emphasis id="F5A7EC26592FFFD0E59FBE9EFA8AFA4C" bold="true" box="[151,1346,1434,1461]" pageId="1" pageNumber="288">
Key to the species
<taxonomicName id="00D34BB7592FFFD0E474BE9FFD6CFA4C" ID-CoL="84N8G" authority="Haase, 1915" authorityName="Haase" authorityYear="1915" box="[380,676,1434,1461]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="288" phylum="Annelida" rank="genus">
<emphasis id="F5A7EC26592FFFD0E474BE9FFDCEFA4D" bold="true" box="[380,518,1434,1460]" italics="true" pageId="1" pageNumber="288">Diplocirrus</emphasis>
<bibRefCitation id="A3424DC5592FFFD0E706BE9EFD6CFA4C" author="Haase" box="[526,676,1435,1461]" pageId="1" pageNumber="288" refString="Haase, P. (1915) Boreale und arktisch Chloraemiden. Wissenschaftliche Meeresuntersuchungen der Kommission zur Wissenschaftlichen Untersuchung der Deutschen Meere, Neue Folge, Kiel, 17, 169 - 228." type="journal article" year="1915">Haase, 1915</bibRefCitation>
</taxonomicName>
(modified from
<bibRefCitation id="A3424DC5592FFFD0E667BE9EFAF3FA4C" author="Salazar-Vallejo" box="[879,1339,1435,1461]" pageId="1" pageNumber="288" refString="Salazar-Vallejo, S. I. &amp; Buzhinskaja, G. (2011) Revision of Diplocirrus Haase, 1915, including Bradiella Rullier, 1965, and Diversibranchius Buzhinskaja, 1993 (Polychaeta, Flabelligeridae). ZooKeys, 106, 1 - 45. http: // dx. doi. org / 10.3897 / zookeys. 106.795" type="journal article" year="2011">Salazar-Vallejo &amp; Buzhinskaja, 2011</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8FC963BF592FFFD0E59FBEE5FA53F820" pageId="1" pageNumber="288" type="key">
<keyStep id="7C272891592FFFD0E59FBEE5FA54F9F5" pageId="1" pageNumber="288">
<paragraph id="C76C3034592FFFD0E59FBEE5FA53FA0E" blockId="1.[151,1436,1504,2009]" box="[151,1435,1504,1527]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBEE5FA53FA0E" box="[151,1435,1504,1527]" pageId="1" pageNumber="288">1. Body papillae abundant............................................................................................................................................. 2</keyLead>
</paragraph>
<paragraph id="C76C3034592FFFD0E59FBD07FA54F9F5" blockId="1.[151,1436,1504,2009]" box="[151,1436,1535,1561]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBD07FA54F9F5" box="[151,1436,1535,1561]" pageId="1" pageNumber="288">
- Body papillae scarce, long, tunic looks bare
<emphasis id="F5A7EC26592FFFD0E7BCBD07FB7DF9E0" box="[692,1205,1538,1561]" italics="true" pageId="1" pageNumber="288">
..........................................................
<taxonomicName id="00D34BB7592FFFD0E131BD07FB7DF9E0" ID-CoL="84N8G" box="[1081,1205,1538,1561]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="288" phylum="Annelida" rank="genus">Diplocirrus</taxonomicName>
</emphasis>
sp. from
<collectingCountry id="BFC470A4592FFFD0E014BD07FA4DF9E0" box="[1308,1413,1538,1561]" name="Sri Lanka" pageId="1" pageNumber="288">Sri Lanka</collectingCountry>
(*)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C272891592FFFD0E59FBD27FA54F9A0" pageId="1" pageNumber="288">
<paragraph id="C76C3034592FFFD0E59FBD27FA53F9C0" blockId="1.[151,1436,1504,2009]" box="[151,1435,1570,1593]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBD27FA53F9C0" box="[151,1435,1570,1593]" pageId="1" pageNumber="288">2(1). Body papillae short, giving a velvety appearance..................................................................................................... 3</keyLead>
</paragraph>
<paragraph id="C76C3034592FFFD0E59FBD47FA54F9A0" blockId="1.[151,1436,1504,2009]" box="[151,1436,1602,1625]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBD47FA54F9A0" box="[151,1436,1602,1625]" pageId="1" pageNumber="288">- Body papillae long, giving a hirsute appearance. ................................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C272891592FFFD0E59FBD67FA53F960" pageId="1" pageNumber="288">
<paragraph id="C76C3034592FFFD0E59FBD67FA54F980" blockId="1.[151,1436,1504,2009]" box="[151,1436,1634,1657]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBD67FA54F980" box="[151,1436,1634,1657]" pageId="1" pageNumber="288">3(2). Body without sand particles ......................................................................................................................................4</keyLead>
</paragraph>
<paragraph id="C76C3034592FFFD0E59FBD87FA53F960" blockId="1.[151,1436,1504,2009]" box="[151,1435,1666,1689]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBD87FA53F960" box="[151,1435,1666,1689]" pageId="1" pageNumber="288">- Body with sand particles ........................................................................................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C272891592FFFD0E59FBDA7FA53F920" pageId="1" pageNumber="288">
<paragraph id="C76C3034592FFFD0E59FBDA7FA53F940" blockId="1.[151,1436,1504,2009]" box="[151,1435,1698,1721]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBDA7FA53F940" box="[151,1435,1698,1721]" pageId="1" pageNumber="288">4(3). Ventrolateral gonopores present in some anterior chaetigers.................................................................................... 5</keyLead>
</paragraph>
<paragraph id="C76C3034592FFFD0E59FBDC7FA53F920" blockId="1.[151,1436,1504,2009]" box="[151,1435,1730,1753]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBDC7FA53F920" box="[151,1435,1730,1753]" pageId="1" pageNumber="288">- Ventrolateral gonopores absent ................................................................................................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C272891592FFFD0E59FBDE7FA53F8C0" pageId="1" pageNumber="288">
<paragraph id="C76C3034592FFFD0E59FBDE7FA54F8E0" blockId="1.[151,1436,1504,2009]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBDE7FA54F8E0" pageId="1" pageNumber="288">
5(4). First chaetiger with long chaetae, about half as long as body width; caruncle posteriorly expanded ........................
<emphasis id="F5A7EC26592FFFD0E5EABC07FC1BF8E0" box="[226,979,1794,1817]" italics="true" pageId="1" pageNumber="288">.................................................................................................................</emphasis>
<taxonomicName id="00D34BB7592FFFD0E6D0BC07FAE1F8E0" ID-CoL="36JVM" authority="Gallardo, 1968" authorityName="Gallardo" authorityYear="1968" box="[984,1321,1794,1817]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="288" phylum="Annelida" rank="species" species="erythroporus">
<emphasis id="F5A7EC26592FFFD0E6D0BC07FB4BF8E0" box="[984,1155,1794,1817]" italics="true" pageId="1" pageNumber="288">D. erythroporus</emphasis>
<bibRefCitation id="A3424DC5592FFFD0E181BC07FAE1F8E0" author="Gallardo" box="[1161,1321,1794,1817]" pageId="1" pageNumber="288" refString="Gallardo, V. A. (1968) Polychaeta from the Bay of Nha Trang, South Viet Nam. Naga Reports, 4 (3), 35 - 279." type="journal article" year="1968">Gallardo, 1968</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="BFC470A4592FFFD0E03FBC07FA5DF8E0" box="[1335,1429,1794,1817]" name="Vietnam" pageId="1" pageNumber="288">Vietnam</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="C76C3034592FFFD0E59FBC27FA53F8C0" blockId="1.[151,1436,1504,2009]" box="[151,1435,1826,1849]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBC27FA53F8C0" box="[151,1435,1826,1849]" pageId="1" pageNumber="288">- Anterior end with short chaetae, 1/3–1/5 as long as body width; caruncle posteriorly tapering .............................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C272891592FFFD0E59FBC47FA53F820" pageId="1" pageNumber="288">
<paragraph id="C76C3034592FFFD0E59FBC47FA53F880" blockId="1.[151,1436,1504,2009]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBC47FA53F880" pageId="1" pageNumber="288">
6(5). Median chaetigers with neurochaetae tapering, 22–25 articles, and tip delicately falcate; cirriform branchiae with lamellae along basal ¼–1/5 of each filament
<emphasis id="F5A7EC26592FFFD0E7B3BC67FC12F880" box="[699,986,1890,1913]" italics="true" pageId="1" pageNumber="288">...........................................</emphasis>
<taxonomicName id="00D34BB7592FFFD0E6D6BC67FAEAF880" ID-CoL="36JVK" authority="(Rullier, 1965)" authorityName="Rullier" authorityYear="1965" baseAuthorityName="Rullier" baseAuthorityYear="1965" box="[990,1314,1890,1913]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="288" phylum="Annelida" rank="species" species="branchiatus">
<emphasis id="F5A7EC26592FFFD0E6D6BC67FBB5F880" box="[990,1149,1890,1913]" italics="true" pageId="1" pageNumber="288">D. branchiatus</emphasis>
(
<bibRefCitation id="A3424DC5592FFFD0E184BC67FAD1F880" author="Rullier" box="[1164,1305,1890,1913]" pageId="1" pageNumber="288" refString="Rullier, F. (1965) Contribution a la faune des annelides polychetes de l'Australie. University of Queensland Papers, Department of Zoology, 2 (9), 163 - 201." type="journal article" year="1965">Rullier, 1965</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="BFC470A4592FFFD0E027BC67FA5CF880" box="[1327,1428,1890,1913]" name="Australia" pageId="1" pageNumber="288">Australia</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="C76C3034592FFFD0E59FBC87FA53F820" blockId="1.[151,1436,1504,2009]" pageId="1" pageNumber="288">
<keyLead id="7C229301592FFFD0E59FBC87FA53F820" pageId="1" pageNumber="288">
- Median chaetigers with neurochaetae barely tapering, 8–11 articles, and tip markedly falcate; cirriform branchiae with lamellae along basal 1/3–1/2 of each filament ................................................................................................... ....................................................................................
<taxonomicName id="00D34BB7592FFFD0E61CBCC7FBADF820" ID-CoL="36JVW" authority="Buzhinskaja, 1994" authorityName="Buzhinskaja" authorityYear="1994" box="[788,1125,1986,2009]" class="Polychaeta" family="Flabelligeridae" genus="Diplocirrus" kingdom="Animalia" order="Terebellida" pageId="1" pageNumber="288" phylum="Annelida" rank="species" species="nicolaji">
<emphasis id="F5A7EC26592FFFD0E61CBCC7FC4EF820" box="[788,902,1986,2009]" italics="true" pageId="1" pageNumber="288">D. nicolaji</emphasis>
(
<bibRefCitation id="A3424DC5592FFFD0E69CBCC7FB96F820" author="Buzhinskaja" box="[916,1118,1986,2009]" pageId="1" pageNumber="288" refString="Buzhinskaja, G. N. (1994 (1993 )) Diversibranchus nicolaji gen. et sp. n. from the Sea of Japan with unique branchial structure (Polychaeta: Flabelligeridae). Zoosystematica Rossica, 2, 229 - 231." type="journal article" year="1994">Buzhinskaja, 1994</bibRefCitation>
)
</taxonomicName>
(Northwestern Sea of
<collectingCountry id="BFC470A4592FFFD0E05EBCC7FA5CF820" box="[1366,1428,1986,2009]" name="Japan" pageId="1" pageNumber="288">Japan</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

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<document id="5E614324B83FE2721B372B0CEC977143" ID-DOI="10.11646/zootaxa.3626.1.5" ID-GBIF-Dataset="135b7553-b75c-4020-b1a6-c689a05b7e66" ID-ISSN="1175-5326" ID-Zenodo-Dep="218625" ID-ZooBank="58AD6857-8CDD-4423-88D0-619CD8D793AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460581053656" checkinUser="plazi" docAuthor="Joshi, Jahnavi &amp; Edgecombe, Gregory D." docDate="2013" docId="4F7A87F2FFC0FFA9FF0BFF28FDB3F871" docLanguage="en" docName="zt03626p145.pdf" docOrigin="Zootaxa 3626 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Digitipes periyarensis Joshi &amp; Edgecombe, 2013, n. sp." docType="treatment" docVersion="7" lastPageNumber="116" masterDocId="B343FF8AFFD1FFB8FF9CFFBFFFE3FFFC" masterDocTitle="Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity" masterLastPageNumber="145" masterPageNumber="99" pageNumber="116" updateTime="1698324300789" updateUser="plazi">
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<mods:title id="3AB7D0888BCEB73DF2B48684F6DEE712">Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity</mods:title>
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<mods:namePart id="7313F4FA5173E644CA51E0CF3E09F95C">Joshi, Jahnavi</mods:namePart>
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<mods:namePart id="A1F1C31D1686DEC97A36DA7B6B476917">Edgecombe, Gregory D.</mods:namePart>
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<treatment id="4F7A87F2FFC0FFA9FF0BFF28FDB3F871" ID-DOI="http://doi.org/10.5281/zenodo.3508271" ID-GBIF-Taxon="119563399" ID-Zenodo-Dep="3508271" LSID="urn:lsid:plazi:treatment:4F7A87F2FFC0FFA9FF0BFF28FDB3F871" httpUri="http://treatment.plazi.org/id/4F7A87F2FFC0FFA9FF0BFF28FDB3F871" lastPageNumber="116" pageId="17" pageNumber="116">
<subSubSection id="8FC9656FFFC0FFA9FF0BFF28FCF5FEE4" pageId="17" pageNumber="116" type="nomenclature">
<paragraph id="C76C36E4FFC0FFA9FF0BFF28FE06FF4D" blockId="17.[151,485,151,210]" box="[151,485,151,177]" pageId="17" pageNumber="116">
<heading id="9C248188FFC0FFA9FF0BFF28FE06FF4D" bold="true" box="[151,485,151,177]" fontSize="11" level="1" pageId="17" pageNumber="116" reason="1">
<emphasis id="F5A7EAF6FFC0FFA9FF0BFF28FE06FF4D" bold="true" box="[151,485,151,177]" pageId="17" pageNumber="116">
<taxonomicName id="00D34D67FFC0FFA9FF0BFF28FE7EFF4D" box="[151,413,151,177]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="species" species="periyarensis" status="sp. nov.">
<emphasis id="F5A7EAF6FFC0FFA9FF0BFF28FE7EFF4D" bold="true" box="[151,413,151,177]" italics="true" pageId="17" pageNumber="116">Digitipes periyarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94578DFFC0FFA9FE3FFF28FE06FF4D" box="[419,485,151,177]" pageId="17" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF0BFF05FEC4FF2E" blockId="17.[151,485,151,210]" box="[151,295,186,210]" pageId="17" pageNumber="116">
(
<figureCitation id="5FE82A61FFC0FFA9FF03FF05FEFCFF2E" box="[159,287,186,210]" captionStart-0="FIGURES 57 – 62" captionStart-1="FIGURES 63 – 67" captionStartId-0="18.[151,264,1777,1800]" captionStartId-1="19.[151,264,1159,1182]" captionTargetBox-0="[151,1435,219,1729]" captionTargetBox-1="[151,1435,193,1137]" captionTargetId-0="figure@18.[151,1436,219,1731]" captionTargetId-1="figure@19.[151,1436,193,1138]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 57 – 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." captionText-1="FIGURES 63 – 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 – 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri-0="https://zenodo.org/record/218635/files/figure.png" httpUri-1="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Figs 57–67</figureCitation>
)
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF0BFEBCFCF5FEE4" blockId="17.[151,790,258,280]" box="[151,790,258,280]" pageId="17" pageNumber="116">
<taxonomicName id="00D34D67FFC0FFA9FF0BFEBCFF17FEE4" box="[151,244,259,280]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC0FFA9FF0BFEBCFF17FEE4" box="[151,244,259,280]" italics="true" pageId="17" pageNumber="116">Digitipes</emphasis>
</taxonomicName>
putative species 4. Joshi and Karanth, 2012: figs 2, 3.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF0BFEFAFBCEFE7D" pageId="17" pageNumber="116" type="materials_examined">
<paragraph id="C76C36E4FFC0FFA9FF0BFEFAFBCEFE7D" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF0BFEFAFEBAFEA2" bold="true" box="[151,345,325,350]" pageId="17" pageNumber="116">
<typeStatus id="18688846FFC0FFA9FF0BFEFAFF31FEA1" box="[151,210,325,349]" pageId="17" pageNumber="116">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="18688846FFC0FFA9FEFCFEFAFE2BFEA1" box="[352,456,325,349]" pageId="17" pageNumber="116" type="holotype">Holotype</typeStatus>
CES
<date id="B36D1024FFC0FFA9FD9FFEFAFDB9FEA1" box="[515,602,325,349]" pageId="17" pageNumber="116" value="37-09-10">091037</date>
, female, from Periyar, Kerala,
<collectingCountry id="BFC47674FFC0FFA9FC28FEFAFC13FEA1" box="[948,1008,325,349]" name="India" pageId="17" pageNumber="116">India</collectingCountry>
, 9°58402 N 77°34902 E, leg. J. Joshi,
<date id="B36D1024FFC0FFA9FF0BFED6FF0CFE7D" box="[151,239,361,385]" pageId="17" pageNumber="116" value="2009-09">ix.2009</date>
.
<typeStatus id="18688846FFC0FFA9FF65FED5FEB8FE7E" box="[249,347,362,386]" pageId="17" pageNumber="116" type="paratype">Paratype</typeStatus>
CES
<date id="B36D1024FFC0FFA9FE09FED6FE0FFE7D" box="[405,492,361,385]" pageId="17" pageNumber="116" value="38-09-10">091038</date>
, female, from
<typeStatus id="18688846FFC0FFA9FD13FED5FD5CFE7E" box="[655,703,362,386]" pageId="17" pageNumber="116">type</typeStatus>
locality, same collection details.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF55FE32FDB7FE59" box="[201,596,397,421]" pageId="17" pageNumber="116" type="etymology">
<paragraph id="C76C36E4FFC0FFA9FF55FE32FDB7FE59" blockId="17.[151,1437,325,1933]" box="[201,596,397,421]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF55FE32FEB3FE59" bold="true" box="[201,336,397,421]" pageId="17" pageNumber="116">Etymology.</emphasis>
For the Periyar region.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF55FE0FFB46FDA5" pageId="17" pageNumber="116" type="diagnosis">
<paragraph id="C76C36E4FFC0FFA9FF55FE0FFB46FDA5" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF55FE0FFEA0FE35" bold="true" box="[201,323,432,457]" pageId="17" pageNumber="116">Diagnosis.</emphasis>
Cephalic plate slightly longer than wide. Basal three antennal articles glabrous dorsally. Forcipular tooth plates longer than wide. Paramedian sutures nearly complete on most sternites. All legs lacking tarsal, tibial and femoral spurs. Femur of ultimate leg with pair of longitudinal grooves on dorsal surface, the more medial of them contiguous with a longitudinal groove on the prefemur and tibia; tarsus of ultimate legs markedly more slender than short, robust tibia and femur. Coxopleural process short; pores relatively sparse.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC0FFA9FF55FDDAFB2FF9D9" pageId="17" pageNumber="116" type="description">
<paragraph id="C76C36E4FFC0FFA9FF55FDDAFE8AFD39" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
<emphasis id="F5A7EAF6FFC0FFA9FF55FDDAFEB8FD82" bold="true" box="[201,347,613,638]" pageId="17" pageNumber="116">Description.</emphasis>
Length to
<quantity id="002B9B01FFC0FFA9FE78FDDAFDDFFD81" box="[484,572,613,637]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="17" pageNumber="116" unit="mm" value="58.0">58 mm</quantity>
. 17 antennal articles; basal three articles glabrous dorsally, two glabrous ventrally. Anterior longitudinal median furrow at most ca 20% length of cephalic plate, considerably shorter in
<typeStatus id="18688846FFC0FFA9FF0BFD12FF1AFD39" box="[151,249,685,709]" pageId="17" pageNumber="116" type="holotype">holotype</typeStatus>
(
<figureCitation id="5FE82A61FFC0FFA9FE94FD12FEBFFD39" box="[264,348,685,709]" captionStart="FIGURES 57 – 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 – 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 57</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FD6EFA92FCF1" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">Cephalic plate and T1 orange; anterior tergites mixed blue and yellow-brown; tergites in posterior half of trunk mostly light brown with blue margins; antennae and legs 1–20 pale yellow apart from pale blue tarsi on leg 20.</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FCA6FF0EFC61" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
Forcipular coxosternal tooth plates longer than wide, markedly so in
<typeStatus id="18688846FFC0FFA9FC46FCA6FBDFFCCD" box="[986,1084,793,817]" pageId="17" pageNumber="116" type="holotype">holotype</typeStatus>
(
<figureCitation id="5FE82A61FFC0FFA9FBD0FCA6FB31FCCD" box="[1100,1234,793,817]" captionStart="FIGURES 57 – 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 – 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Figs 58, 59</figureCitation>
); four main teeth, not arranged in groups, the outer tooth smaller than the inner three; base of tooth plates defined by relative acute oblique sutures diverging at 105°–125°. Trochanteroprefemoral process with indistinctly defined teeth along inner margin.
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FC16FA9DFC3D" blockId="17.[151,1437,325,1933]" box="[201,1406,937,962]" pageId="17" pageNumber="116">Second maxillary claw with slender accessory spurs. Article 2 of telopodite bearing a slender spine distally.</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FC72FDE2FBAD" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
Tergites with paramedian sutures complete from TT4 or 5, short suture confined to anterior edge of TT2–3. Tergites marginate from 5 or 6. Tergites smooth, lacking median ridge or keels (
<figureCitation id="5FE82A61FFC0FFA9FBF8FC4EFB5FFBF5" box="[1124,1212,1009,1033]" captionStart="FIGURES 63 – 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 – 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 – 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 66</figureCitation>
) apart from weak, incomplete median keel on tergite of ultimate leg-bearing segment; lateral part of posterior tergites with low, irregularly anastomosing ridges.
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FBE2FD3DFB65" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">Sternites with paramedian sutures nearly complete in most segments, especially in anterior part of trunk, those in posterior few segments ca 80% length of sternite.</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FB1AFADCFAF9" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
Tergite of ultimate leg-bearing segment with lateral margins gently converging posteriorly, posterolateral margins mostly straight, converging to a broad, evenly rounded posteromedial margin (
<figureCitation id="5FE82A61FFC0FFA9FB31FB76FAE5FB1D" box="[1197,1286,1225,1249]" captionStart="FIGURES 57 – 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 – 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 61</figureCitation>
). Sternite of ultimate leg-bearing segment with sides converging posteriorly, posterior margin gently concave (
<figureCitation id="5FE82A61FFC0FFA9FB42FB52FAD0FAF9" box="[1246,1331,1261,1285]" captionStart="FIGURES 57 – 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 – 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFC0FFA9FF55FAAEFB2FF9D9" blockId="17.[151,1437,325,1933]" pageId="17" pageNumber="116">
Coxopleural process short, barely inflected from posterolateral margin of coxopleuron in ventral view (
<figureCitation id="5FE82A61FFC0FFA9FAF3FAAEFF55FAB1" captionStart="FIGURES 57 – 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 – 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 62</figureCitation>
), with two apical spines, lacking lateral spine(s). Pore field terminating strongly beneath dorsal margin of coxopleuron (
<figureCitation id="5FE82A61FFC0FFA9FEAFFAE6FE6BFA8D" box="[307,392,1369,1393]" captionStart="FIGURES 57 – 62" captionStartId="18.[151,264,1777,1800]" captionTargetBox="[151,1435,219,1729]" captionTargetId="figure@18.[151,1436,219,1731]" captionTargetPageId="18" captionText="FIGURES 57 – 62. Digitipes periyarensis n. sp. Holotype CES 091037 (female). 57. Cephalic plate and T 1, dorsal view, scale = 1 mm. 58. Forcipular segment, ventral view, scale = 2 mm. 59. Tooth plates, ventral view, scale = 0.5 mm. 60. Coxopleuron, lateral view, scale = 1 mm. 61. Tergite of ultimate leg-bearing segment, dorsal view, scale = 2 mm. 62. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218635/files/figure.png" pageId="17" pageNumber="116">Fig. 60</figureCitation>
); pores relative sparse; non-porose area on coxopleural process short. Ultimate leg prefemur with width at distal end nearly 40% its length, of nearly equal width along its length (
<figureCitation id="5FE82A61FFC0FFA9FBFBFAC2FB5EFA69" box="[1127,1213,1405,1429]" captionStart="FIGURES 63 – 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 – 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 – 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 64</figureCitation>
). Prefemoral spines robust, with a short, curved apex (
<figureCitation id="5FE82A61FFC0FFA9FD87FA1EFD93FA45" box="[539,624,1441,1465]" captionStart="FIGURES 63 – 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 – 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 – 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 63</figureCitation>
): VL 3, VM 2(3), DM 2. Femur with width at its distal end 40–50% its length (
<figureCitation id="5FE82A61FFC0FFA9FF70FA7AFEA3FA21" box="[236,320,1477,1501]" captionStart="FIGURES 63 – 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 – 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 – 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 64</figureCitation>
). Dorsal, medial and lateral surfaces of prefemur and femur rugose, with large, shallow pits; similar pits adjacent to longitudinal groove on dorsomedial side of tibia (
<figureCitation id="5FE82A61FFC0FFA9FCEEFA56FC24F9FD" box="[882,967,1513,1537]" captionStart="FIGURES 63 – 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 – 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 – 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="17" pageNumber="116">Fig. 65</figureCitation>
). Ultimate leg tarsus 1 2.5 times length of tarsus 2; tarsus 1 2–3.3 times longer than pretarsus; pretarsus with pair of short accessory claws.
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<emphasis id="F5A7EAF6FFC0FFA9FF55F98EFEADF9B6" bold="true" box="[201,334,1585,1610]" pageId="17" pageNumber="116">Discussion.</emphasis>
This species is the most morphologically distinct among Indian
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<emphasis id="F5A7EAF6FFC0FFA9FBBEF98DFB6AF9B5" box="[1058,1161,1586,1609]" italics="true" pageId="17" pageNumber="116">Digitipes</emphasis>
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, readily identified by its complete absence of tarsal spurs, the long paramedian sutures on the sternites (nearly complete versus not more than 35% the length of the sternites in other Indian
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), and by the robust, dorsally-grooved prefemur, femur and tibia of the ultimate legs. Complete paramedian sutures on the sternites are known elsewhere in
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in the
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species,
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<emphasis id="F5A7EAF6FFC0FFA9FEB1F97DFE29F925" box="[301,458,1729,1753]" italics="true" pageId="17" pageNumber="116">D. verdascens</emphasis>
Attems.
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The two known specimens are both females so no data are at hand with respect to the condition of the femoral process in males. Despite its distinctive character states, its membership in
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<emphasis id="F5A7EAF6FFC0FFA9FC74F8B5FBACF8DD" box="[1000,1103,1802,1825]" italics="true" pageId="17" pageNumber="116">Digitipes</emphasis>
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is indicated by the support at the nodes in the phylogeny that unite it with Group A of Joshi and Karanth (2012) (posterior probability 0.94 for it grouping with
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<emphasis id="F5A7EAF6FFC0FFA9FEA0F8EDFE11F895" box="[316,498,1874,1897]" italics="true" pageId="17" pageNumber="116">D. coonoorensis</emphasis>
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and
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<emphasis id="F5A7EAF6FFC0FFA9FDB5F8EDFD7FF895" box="[553,668,1873,1897]" italics="true" pageId="17" pageNumber="116">D. indicus</emphasis>
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) and that clade as a whole grouping with Groups B and C (posterior probability 1, bootstrap support 100%).
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Jangi and Dass, 1984
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<emphasis id="F5A7EAF6FFC2FFABFF0BFA33FEB9FA5D" box="[151,346,1419,1441]" italics="true" pageId="19" pageNumber="118">Digitipes chhotanii</emphasis>
Jangi and Dass, 1984: 46
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, figs 80–86.
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<emphasis id="F5A7EAF6FFC2FFABFF0BFA71FEE8FA1A" bold="true" box="[151,267,1486,1510]" pageId="19" pageNumber="118">Remarks.</emphasis>
This species was based on a single male from Suranganar Reserve, Tamil Nadu (collected in 1949). Additional records of
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were made by Yadav (1993b) from Maharashtra, and from Kerala by Sureshan
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. (2006). We have not seen any specimens that conform to the original description of
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with respect to a posterior bifurcation on the anterior median furrow on the cephalic plate, 2.5 glabrous antennal articles, or pilose tarsi on the ultimate leg. Its identity as
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<paragraph id="C76C36E4FFC2FFABFF0BF952FD89F8FB" blockId="19.[151,618,1773,1799]" box="[151,618,1773,1799]" pageId="19" pageNumber="118">
<heading id="9C248188FFC2FFABFF0BF952FD89F8FB" bold="true" box="[151,618,1773,1799]" fontSize="11" level="1" pageId="19" pageNumber="118" reason="1">
<taxonomicName id="00D34D67FFC2FFABFF0BF952FD89F8FB" ID-CoL="363VV" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[151,618,1773,1799]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="118" phylum="Arthropoda" rank="species" species="pruthii">
<emphasis id="F5A7EAF6FFC2FFABFF0BF952FD89F8FB" bold="true" box="[151,618,1773,1799]" pageId="19" pageNumber="118">
<emphasis id="F5A7EAF6FFC2FFABFF0BF952FEBEF8FB" bold="true" box="[151,349,1773,1799]" italics="true" pageId="19" pageNumber="118">Digitipes pruthii</emphasis>
Jangi and Dass, 1984
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C76C36E4FFC2FFABFF0BF88AFD21F8B6" blockId="19.[151,706,1844,1866]" box="[151,706,1844,1866]" pageId="19" pageNumber="118">
<taxonomicName id="00D34D67FFC2FFABFF0BF88AFDA6F8B6" authority="Jangi and Dass, 1984: 41" authorityName="Jangi and Dass" authorityPageNumber="41" authorityYear="1984" box="[151,581,1844,1866]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="118" phylum="Arthropoda" rank="species" species="pruthii">
<emphasis id="F5A7EAF6FFC2FFABFF0BF88AFEDCF8B6" box="[151,319,1844,1866]" italics="true" pageId="19" pageNumber="118">Digitipes pruthii</emphasis>
Jangi and Dass, 1984: 41
</taxonomicName>
, figs 51–57.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC2FFACFF0BF8C8FF03FE98" lastPageId="20" lastPageNumber="119" pageId="19" pageNumber="118" type="discussion">
<paragraph id="C76C36E4FFC2FFACFF0BF8C8FF03FE98" blockId="19.[151,1436,1911,2008]" lastBlockId="20.[151,1436,151,356]" lastPageId="20" lastPageNumber="119" pageId="19" pageNumber="118">
<emphasis id="F5A7EAF6FFC2FFABFF0BF8C8FEE8F873" bold="true" box="[151,267,1911,1935]" pageId="19" pageNumber="118">Remarks.</emphasis>
We provisionally accept the validity of
<taxonomicName id="00D34D67FFC2FFABFD42F8C7FCACF873" box="[734,847,1911,1935]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="118" phylum="Arthropoda" rank="species" species="pruthii">
<emphasis id="F5A7EAF6FFC2FFABFD42F8C7FCACF873" box="[734,847,1911,1935]" italics="true" pageId="19" pageNumber="118">D. pruthii</emphasis>
</taxonomicName>
as a species, though it is known from its
<typeStatus id="18688846FFC2FFABFAA6F8C8FA7FF873" box="[1338,1436,1911,1935]" pageId="19" pageNumber="118" type="holotype">holotype</typeStatus>
alone, collected in the Salem District of Tamil Nadu in 1929. Collecting by JJ at the
<typeStatus id="18688846FFC2FFABFBCCF823FB63F848" box="[1104,1152,1948,1972]" pageId="19" pageNumber="118">type</typeStatus>
locality of
<taxonomicName id="00D34D67FFC2FFABFA9EF823FA93F84F" box="[1282,1392,1947,1971]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="118" phylum="Arthropoda" rank="species" species="pruthii">
<emphasis id="F5A7EAF6FFC2FFABFA9EF823FA93F84F" box="[1282,1392,1947,1971]" italics="true" pageId="19" pageNumber="118">D. pruthii</emphasis>
</taxonomicName>
did not yield any specimens of
<taxonomicName id="00D34D67FFC2FFABFE56F87FFDD2F82B" box="[458,561,1984,2007]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC2FFABFE56F87FFDD2F82B" box="[458,561,1984,2007]" italics="true" pageId="19" pageNumber="118">Digitipes</emphasis>
</taxonomicName>
, and none of our collections include specimens that possess certain characters of the
<typeStatus id="18688846FFC5FFACFF7AFF28FEABFF53" box="[230,328,151,175]" pageId="20" pageNumber="119" type="holotype">holotype</typeStatus>
that appear to be distinctive, i.e. a posterior birfurcation on the anterior median furrow on the cephalic plate, a “broad-based and blunt conical” distomedial process on the ultimate leg femur of males, and a short tergite of the ultimate leg-bearing segment. The distomedial process on the femur of the
<typeStatus id="18688846FFC5FFACFB32FF60FAF3FF0B" box="[1198,1296,223,247]" pageId="20" pageNumber="119" type="holotype">holotype</typeStatus>
differs from all definite
<taxonomicName id="00D34D67FFC5FFACFE86FEBAFE62FEE0" box="[282,385,261,284]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFE86FEBAFE62FEE0" box="[282,385,261,284]" italics="true" pageId="20" pageNumber="119">Digitipes</emphasis>
</taxonomicName>
in its substantially lesser projection, and the membership of this species in the genus is not solidly established. It is also geographically anomalous, being the only purported record of
<taxonomicName id="00D34D67FFC5FFACFB09FE96FB1FFEBC" box="[1173,1276,297,320]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFB09FE96FB1FFEBC" box="[1173,1276,297,320]" italics="true" pageId="20" pageNumber="119">Digitipes</emphasis>
</taxonomicName>
in the Eastern Ghats.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="4A6488E475D914C6FC6279C4E55567E6">Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity</mods:title>
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<paragraph id="C76C36E4FFC5FFACFF0BFD15FCFAFD38" blockId="20.[151,793,682,708]" box="[151,793,682,708]" pageId="20" pageNumber="119">
<heading id="9C248188FFC5FFACFF0BFD15FCFAFD38" bold="true" box="[151,793,682,708]" fontSize="11" level="1" pageId="20" pageNumber="119" reason="1">
<emphasis id="F5A7EAF6FFC5FFACFF0BFD15FCFAFD38" bold="true" box="[151,793,682,708]" pageId="20" pageNumber="119">
<taxonomicName id="00D34D67FFC5FFACFF0BFD15FD44FD38" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[151,679,682,708]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="gravelyi" status="comb. nov.">
<emphasis id="F5A7EAF6FFC5FFACFF0BFD15FE64FD38" bold="true" box="[151,391,682,708]" italics="true" pageId="20" pageNumber="119">Otostigmus gravelyi</emphasis>
(Jangi and Dass, 1984)
</taxonomicName>
<taxonomicNameLabel id="EE94578DFFC5FFACFD33FD15FCFAFD38" box="[687,793,682,708]" pageId="20" pageNumber="119">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C76C36E4FFC5FFACFF0BFD4DFD93FCD9" blockId="20.[151,721,753,806]" pageId="20" pageNumber="119">
<taxonomicName id="00D34D67FFC5FFACFF0BFD4DFDB0FCFB" authority="Jangi and Dass, 1984: 41" authorityName="Jangi and Dass" authorityPageNumber="41" authorityYear="1984" box="[151,595,753,775]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="F5A7EAF6FFC5FFACFF0BFD4DFEAEFCFB" box="[151,333,753,775]" italics="true" pageId="20" pageNumber="119">Digitipes gravelyi</emphasis>
Jangi and Dass, 1984: 41
</taxonomicName>
, figs 58–64.
<taxonomicName id="00D34D67FFC5FFACFF0BFCAEFEAEFCDA" box="[151,333,784,806]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="F5A7EAF6FFC5FFACFF0BFCAEFEAEFCDA" box="[151,333,784,806]" italics="true" pageId="20" pageNumber="119">Digitipes gravelyi</emphasis>
</taxonomicName>
. Sureshan
<emphasis id="F5A7EAF6FFC5FFACFE20FCAEFE0AFCDA" box="[444,489,784,806]" italics="true" pageId="20" pageNumber="119">et al</emphasis>
. 2006: 2287.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC5FFACFF0BFCECFCC2F9A2" pageId="20" pageNumber="119" type="discussion">
<paragraph id="C76C36E4FFC5FFACFF0BFCECFD19FB2E" blockId="20.[151,1437,850,1630]" pageId="20" pageNumber="119">
<emphasis id="F5A7EAF6FFC5FFACFF0BFCECFEE8FC97" bold="true" box="[151,267,851,875]" pageId="20" pageNumber="119">Remarks.</emphasis>
This species was described on the basis of its
<typeStatus id="18688846FFC5FFACFCB8FCEDFC65FC96" box="[804,902,850,874]" pageId="20" pageNumber="119" type="holotype">holotype</typeStatus>
alone, a female collected from Parambikulam, Kerala, in 1914. Specimens from the Quilon district in Kerala were subsequently assigned to it by Sureshan
<emphasis id="F5A7EAF6FFC5FFACFAFFFCC7FA76FC72" box="[1379,1429,886,911]" italics="true" pageId="20" pageNumber="119">et al</emphasis>
. (2006). Collections by JJ at the
<typeStatus id="18688846FFC5FFACFD9EFC24FDD1FC4F" box="[514,562,923,947]" pageId="20" pageNumber="119">type</typeStatus>
locality did not produce any specimens that correspond to the
<typeStatus id="18688846FFC5FFACFA9CFC25FA81FC4E" box="[1280,1378,922,946]" pageId="20" pageNumber="119" type="holotype">holotype</typeStatus>
with respect to the presence of four apical spines and two strong lateral spines on the coxopleural process, having as many as four spines in the VM and VL rows on the ultimate leg prefemur, and showing a transition from smooth TT1–2 to keeled, tuberculate T3. We have in fact not identified these character states in specimens of
<taxonomicName id="00D34D67FFC5FFACFAA9FBB8FA7FFBE2" box="[1333,1436,1031,1054]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFAA9FBB8FA7FFBE2" box="[1333,1436,1031,1054]" italics="true" pageId="20" pageNumber="119">Digitipes</emphasis>
</taxonomicName>
from any other locality, apart from the specimens assigned to
<taxonomicName id="00D34D67FFC5FFACFCF5FB93FC08FBBF" box="[873,1003,1067,1091]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="F5A7EAF6FFC5FFACFCF5FB93FC08FBBF" box="[873,1003,1067,1091]" italics="true" pageId="20" pageNumber="119">D. gravelyi</emphasis>
</taxonomicName>
by Sureshan
<emphasis id="F5A7EAF6FFC5FFACFB10FB93FB23FBBF" box="[1164,1216,1067,1091]" italics="true" pageId="20" pageNumber="119">et al</emphasis>
. (2006). The latter (photographs provided by D. Balan) indicate that these distinctive character states of the
<typeStatus id="18688846FFC5FFACFBE7FBF1FB3EFB9A" box="[1147,1245,1102,1126]" pageId="20" pageNumber="119" type="holotype">holotype</typeStatus>
are conserved in other specimens, even from other localities. Accordingly we regard these characters as potentially being diagnostic of this species. The otostigmine collected by JJ at the
<typeStatus id="18688846FFC5FFACFC94FB28FCDBFB53" box="[776,824,1175,1199]" pageId="20" pageNumber="119">type</typeStatus>
locality of
<taxonomicName id="00D34D67FFC5FFACFC22FB27FBA8FB52" box="[958,1099,1174,1199]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="graveleyi">
<emphasis id="F5A7EAF6FFC5FFACFC22FB27FBA8FB52" box="[958,1099,1174,1199]" italics="true" pageId="20" pageNumber="119">D. graveleyi</emphasis>
</taxonomicName>
is clearly a different species, identified below as
<taxonomicName id="00D34D67FFC5FFACFEEFFB04FD14FB2E" authority="Pocock, 1890" authorityName="Pocock" authorityYear="1890" box="[371,759,1210,1235]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC5FFACFEEFFB04FDB0FB2F" box="[371,595,1211,1235]" italics="true" pageId="20" pageNumber="119">Otostigmus ruficeps</emphasis>
Pocock, 1890
</taxonomicName>
.
</paragraph>
<paragraph id="C76C36E4FFC5FFACFF55FB61FCC2F9A2" blockId="20.[151,1437,850,1630]" pageId="20" pageNumber="119">
The male has not been described for “
<emphasis id="F5A7EAF6FFC5FFACFDEFFB5FFD6CFB0B" box="[627,655,1248,1271]" italics="true" pageId="20" pageNumber="119">D.</emphasis>
”
<taxonomicName id="00D34D67FFC5FFACFD3EFB61FD1DFB0A" box="[674,766,1246,1270]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="F5A7EAF6FFC5FFACFD3EFB61FD1DFB0A" box="[674,766,1246,1270]" italics="true" pageId="20" pageNumber="119">gravelyi</emphasis>
</taxonomicName>
so its membership in the genus was not solidly established.
<collectingCountry id="BFC47674FFC5FFACFF0BFABCFF55FAE7" box="[151,182,1283,1307]" name="American Samoa" pageId="20" pageNumber="119">As</collectingCountry>
noted above under discussion of the genus, some of its characters deviate from very stable conditions shared by the other species of
<taxonomicName id="00D34D67FFC5FFACFE1DFA98FE0BFAC2" box="[385,488,1319,1342]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFE1DFA98FE0BFAC2" box="[385,488,1319,1342]" italics="true" pageId="20" pageNumber="119">Digitipes</emphasis>
</taxonomicName>
. It has 19–21 antennal articles, whereas all other Indian species have a conserved number of 17 articles. It has five teeth on each forcipular tooth plate, in contrast to a highly conserved pattern of only four teeth in the other Indian and African species. No
<taxonomicName id="00D34D67FFC5FFACFCEBFAD0FC3DFA7A" box="[887,990,1391,1414]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFCEBFAD0FC3DFA7A" box="[887,990,1391,1414]" italics="true" pageId="20" pageNumber="119">Digitipes</emphasis>
</taxonomicName>
in
<collectingCountry id="BFC47674FFC5FFACFB92FAD1FBABFA7A" box="[1038,1096,1390,1414]" name="India" pageId="20" pageNumber="119">India</collectingCountry>
or Africa has dorsal spines accompanying the two apical spines on the coxopleural process, and none has more than a single lateral spine (versus two very strong lateral spines in “
<emphasis id="F5A7EAF6FFC5FFACFDE2FA07FD71FA33" box="[638,658,1464,1487]" italics="true" pageId="20" pageNumber="119">D</emphasis>
”.
<taxonomicName id="00D34D67FFC5FFACFD33FA09FCE8FA32" box="[687,779,1462,1486]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="F5A7EAF6FFC5FFACFD33FA09FCE8FA32" box="[687,779,1462,1486]" italics="true" pageId="20" pageNumber="119">gravelyi</emphasis>
</taxonomicName>
). These character states are, however, present in various species of
<taxonomicName id="00D34D67FFC5FFACFE92FA64FE6EFA0E" box="[270,397,1499,1522]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFE92FA64FE6EFA0E" box="[270,397,1499,1522]" italics="true" pageId="20" pageNumber="119">Otostigmus</emphasis>
</taxonomicName>
, and indeed there is little to separate
<emphasis id="F5A7EAF6FFC5FFACFCA9FA64FC83FA0E" box="[821,864,1499,1522]" italics="true" pageId="20" pageNumber="119">“D.</emphasis>
”
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<emphasis id="F5A7EAF6FFC5FFACFCE8FA64FC33FA0F" box="[884,976,1499,1523]" italics="true" pageId="20" pageNumber="119">gravelyi</emphasis>
</taxonomicName>
from
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<emphasis id="F5A7EAF6FFC5FFACFB89FA64FAEFFA0F" box="[1045,1292,1499,1523]" italics="true" pageId="20" pageNumber="119">Otostigmus rugulosus</emphasis>
Porat, 1876
</taxonomicName>
. Because we have no new collections of the species, the possibility of synonymy with
<taxonomicName id="00D34D67FFC5FFACFBC7FA40FB08F9EA" box="[1115,1259,1534,1558]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="rugulosus">
<emphasis id="F5A7EAF6FFC5FFACFBC7FA40FB08F9EA" box="[1115,1259,1534,1558]" italics="true" pageId="20" pageNumber="119">
<collectingCountry id="BFC47674FFC5FFACFBC7FA40FB91F9EA" box="[1115,1138,1535,1558]" name="Iceland" pageId="20" pageNumber="119">O</collectingCountry>
. rugulosus
</emphasis>
</taxonomicName>
is not followed up here, but the close similarity warrants the reassignment of the species to
<taxonomicName id="00D34D67FFC5FFACFB9FF99CFB60F9C6" box="[1027,1155,1571,1594]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFB9FF99CFB60F9C6" box="[1027,1155,1571,1594]" italics="true" pageId="20" pageNumber="119">Otostigmus</emphasis>
</taxonomicName>
as the new combination
<taxonomicName id="00D34D67FFC5FFACFF0BF9F8FCFEF9A2" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[151,797,1606,1630]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="gravelyi" subGenus="Otostigmus">
<emphasis id="F5A7EAF6FFC5FFACFF0BF9F8FEF4F9A2" box="[151,279,1607,1630]" italics="true" pageId="20" pageNumber="119">Otostigmus</emphasis>
(
<emphasis id="F5A7EAF6FFC5FFACFEBBF9F8FE45F9A2" box="[295,422,1607,1630]" italics="true" pageId="20" pageNumber="119">Otostigmus</emphasis>
)
<emphasis id="F5A7EAF6FFC5FFACFE2AF9F9FDF1F9A2" box="[438,530,1606,1630]" italics="true" pageId="20" pageNumber="119">gravelyi</emphasis>
(Jangi and Dass, 1984)
</taxonomicName>
.
</paragraph>
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59
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Genus
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Porat, 1876
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Subgenus
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Porat, 1876
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species.
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Porat, 1876
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, by subsequent designation of Pocock (1891).
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<mods:title id="0181A42D07638DDEB6DC6D0753DF25C8">Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity</mods:title>
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<mods:namePart id="789834ECBE72E53B11C8C519A723A2E0">Edgecombe, Gregory D.</mods:namePart>
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<treatment id="4F7A87F2FFC5FFAFFF0BF90DFAA8FAF0" ID-DOI="http://doi.org/10.5281/zenodo.6154132" ID-GBIF-Taxon="119563412" ID-Zenodo-Dep="6154132" LSID="urn:lsid:plazi:treatment:4F7A87F2FFC5FFAFFF0BF90DFAA8FAF0" httpUri="http://treatment.plazi.org/id/4F7A87F2FFC5FFAFFF0BF90DFAA8FAF0" lastPageId="23" lastPageNumber="122" pageId="20" pageNumber="119">
<subSubSection id="8FC9656FFFC5FFACFF0BF90DFD7DF8AD" pageId="20" pageNumber="119" type="nomenclature">
<paragraph id="C76C36E4FFC5FFACFF0BF90DFDD1F930" blockId="20.[151,562,1714,1773]" box="[151,562,1714,1740]" pageId="20" pageNumber="119">
<heading id="9C248188FFC5FFACFF0BF90DFDD1F930" bold="true" box="[151,562,1714,1740]" fontSize="11" level="1" pageId="20" pageNumber="119" reason="1">
<taxonomicName id="00D34D67FFC5FFACFF0BF90DFDD1F930" ID-CoL="4B7ZC" authority="Pocock, 1890" authorityName="Pocock" authorityYear="1890" box="[151,562,1714,1740]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC5FFACFF0BF90DFDD1F930" bold="true" box="[151,562,1714,1740]" pageId="20" pageNumber="119">
<emphasis id="F5A7EAF6FFC5FFACFF0BF90DFE6AF930" bold="true" box="[151,393,1714,1740]" italics="true" pageId="20" pageNumber="119">Otostigmus ruficeps</emphasis>
Pocock, 1890
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C76C36E4FFC5FFACFF0BF96AFEC4F911" blockId="20.[151,562,1714,1773]" box="[151,295,1749,1773]" pageId="20" pageNumber="119">
(
<figureCitation id="5FE82A61FFC5FFACFF03F96AFEFCF911" box="[159,287,1749,1773]" captionStart-0="FIGURES 68 – 73" captionStart-1="FIGURES 74 – 77" captionStartId-0="21.[151,264,1870,1893]" captionStartId-1="22.[151,264,970,993]" captionTargetBox-0="[151,1435,337,1836]" captionTargetBox-1="[395,1198,193,948]" captionTargetId-0="figure@21.[151,1436,337,1849]" captionTargetId-1="figure@22.[388,1198,193,949]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 68 – 73. Otostigmus ruficeps Pocock, 1890. 68. Cephalic plate and T 1, dorsal view, CES 091013, scale = 2 mm. 69. Forcipular segment, ventral view, CES 091048, scale = 1 mm. 70. Tooth plates, ventral view, CES 001013, scale = 1 mm. 71. Coxopleuron, lateral view, CES 091330, scale = 1 mm. 72. Tergite of ultimate leg-bearing segment, dorsal view, CES 091330, scale = 2 mm. 73. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, CES 091330, scale = 1 mm." captionText-1="FIGURES 74 – 77. Otostigmus ruficeps Pocock, 1890. 74. Ultimate leg prefemur, ventral view, CES 091327, scale = 1 mm. 75. Ultimate leg femur, dorsal view, CES 091327, scale = 1 mm. 76. Spiracle on segment 3, CES 091013, scale = 0.5 mm. 77. TT 17 – 20, dorsal view, CES 08960, scale = 1 mm." httpUri-0="https://zenodo.org/record/218637/files/figure.png" httpUri-1="https://zenodo.org/record/218638/files/figure.png" pageId="20" pageNumber="119">Figs 68–77</figureCitation>
)
</paragraph>
<paragraph id="C76C36E4FFC5FFACFF0BF8A1FD7DF8AD" blockId="20.[151,790,1821,1904]" pageId="20" pageNumber="119">
<taxonomicName id="00D34D67FFC5FFACFF0BF8A1FDCBF8CF" authority="Pocock, 1890: 247" authorityName="Pocock" authorityPageNumber="247" authorityYear="1890" box="[151,552,1821,1843]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC5FFACFF0BF8A1FE81F8CF" box="[151,354,1821,1843]" italics="true" pageId="20" pageNumber="119">Otostigmus ruficeps</emphasis>
Pocock, 1890: 247
</taxonomicName>
, pl. 12, figs 2–2b.
<emphasis id="F5A7EAF6FFC5FFACFF0BF883FE8AF8AE" box="[151,361,1852,1874]" italics="true" pageId="20" pageNumber="119">
<taxonomicName id="00D34D67FFC5FFACFF0BF883FE86F8AE" box="[151,357,1852,1874]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="species" species="ruficeps">Otostigmus ruficeps</taxonomicName>
.
</emphasis>
Lewis, 1996: 828, figs 20–23.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC5FFACFF0BF8E4FCF5F88C" box="[151,790,1882,1904]" pageId="20" pageNumber="119" type="description">
<paragraph id="C76C36E4FFC5FFACFF0BF8E4FCF5F88C" blockId="20.[151,790,1821,1904]" box="[151,790,1882,1904]" pageId="20" pageNumber="119">
<taxonomicName id="00D34D67FFC5FFACFF0BF8E4FF17F88C" box="[151,244,1883,1904]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFF0BF8E4FF17F88C" box="[151,244,1883,1904]" italics="true" pageId="20" pageNumber="119">Digitipes</emphasis>
</taxonomicName>
putative species 9. Joshi and Karanth, 2012: figs 2, 3.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC5FFADFF0BF822FCD9FEE0" lastPageId="21" lastPageNumber="120" pageId="20" pageNumber="119" type="diagnosis">
<paragraph id="C76C36E4FFC5FFADFF0BF822FCD9FEE0" blockId="20.[151,1436,1949,2009]" lastBlockId="21.[151,1436,151,284]" lastPageId="21" lastPageNumber="120" pageId="20" pageNumber="119">
<emphasis id="F5A7EAF6FFC5FFACFF0BF822FEF2F84A" bold="true" box="[151,273,1949,1974]" pageId="20" pageNumber="119">Diagnosis.</emphasis>
<taxonomicName id="00D34D67FFC5FFACFE84F821FE74F849" box="[280,407,1950,1973]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="20" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC5FFACFE84F821FE74F849" box="[280,407,1950,1973]" italics="true" pageId="20" pageNumber="119">Otostigmus</emphasis>
</taxonomicName>
with 21 elongate antennal articles, the basal 2–2.5 glabrous dorsally; forcipular tooth plates with 4+4 large teeth, small (fifth) inner tooth variably present; four teeth along margin of forcipular trochanteroprefemoral process; longitudinal median depression on posterior half of tergite of ultimate leg-bearing segment; two apical spines on coxopleural process, dorsal spine rarely present; lateral spine mostly lacking; two tarsal spurs on legs
<date id="B36D1024FFC4FFADFEE5FF60FDE2FF0B" box="[377,513,223,247]" pageId="21" pageNumber="120" value="1919-01-04" valueMax="1919-01-01" valueMin="1919-01-04">1–4 to 1–19</date>
(usually
<date id="B36D1024FFC4FFADFDF0FF60FCE7FF0B" box="[620,772,223,247]" pageId="21" pageNumber="120" value="1919-01-17" valueMax="1919-01-01" valueMin="1919-01-17">1–17 to 1–19</date>
), one on leg 20; a tibial spur typically on legs 1–6 to 1–8; ultimate leg prefemur with 2 VM spines and 2–3 VL spines.
</paragraph>
</subSubSection>
<caption id="93AC666CFFC4FFADFF0BF8F1FAA5F83C" httpUri="https://zenodo.org/record/218637/files/figure.png" pageId="21" pageNumber="120" targetBox="[151,1435,337,1836]" targetPageId="21">
<paragraph id="C76C36E4FFC4FFADFF0BF8F1FAA5F83C" blockId="21.[151,1436,1870,1984]" pageId="21" pageNumber="120">
<emphasis id="F5A7EAF6FFC4FFADFF0BF8F1FEC4F89F" bold="true" box="[151,295,1870,1893]" pageId="21" pageNumber="120">FIGURES 68</emphasis>
–
<emphasis id="F5A7EAF6FFC4FFADFEA8F8F1FEB7F89F" bold="true" box="[308,340,1870,1891]" pageId="21" pageNumber="120">73.</emphasis>
<taxonomicName id="00D34D67FFC4FFADFEC7F8F0FD5FF898" authority="Pocock, 1890" authorityName="Pocock" authorityYear="1890" box="[347,700,1870,1892]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="120" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC4FFADFEC7F8F0FE2DF898" box="[347,462,1871,1892]" italics="true" pageId="21" pageNumber="120">Otostigmus</emphasis>
<emphasis id="F5A7EAF6FFC4FFADFE49F8F1FDC4F898" box="[469,551,1870,1892]" italics="true" pageId="21" pageNumber="120">ruficeps</emphasis>
Pocock, 1890
</taxonomicName>
. 68. Cephalic plate and T1, dorsal view, CES091013, scale = 2 mm. 69. Forcipular segment, ventral view, CES091048, scale = 1mm. 70. Tooth plates, ventral view, CES001013, scale = 1 mm. 71. Coxopleuron, lateral view, CES091330, scale = 1 mm. 72. Tergite of ultimate leg-bearing segment, dorsal view, CES091330, scale = 2 mm. 73. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, CES091330, scale = 1 mm.
</paragraph>
</caption>
<caption id="93AC666CFFC7FFAEFF0BFC75FD68FBE1" httpUri="https://zenodo.org/record/218638/files/figure.png" pageId="22" pageNumber="121" targetBox="[395,1198,193,948]" targetPageId="22">
<paragraph id="C76C36E4FFC7FFAEFF0BFC75FD68FBE1" blockId="22.[151,1437,970,1053]" pageId="22" pageNumber="121">
<emphasis id="F5A7EAF6FFC7FFAEFF0BFC75FEC5FC1C" bold="true" box="[151,294,970,993]" pageId="22" pageNumber="121">FIGURES 74</emphasis>
–
<emphasis id="F5A7EAF6FFC7FFAEFEA8FC74FEB0FC1C" bold="true" box="[308,339,971,992]" pageId="22" pageNumber="121">77.</emphasis>
<taxonomicName id="00D34D67FFC7FFAEFEC3FC74FD5DFC1C" authority="Pocock, 1890" authorityName="Pocock" authorityYear="1890" box="[351,702,970,992]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="121" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC7FFAEFEC3FC74FE30FC1C" box="[351,467,971,992]" italics="true" pageId="22" pageNumber="121">Otostigmus</emphasis>
<emphasis id="F5A7EAF6FFC7FFAEFE45FC75FDC8FC1C" box="[473,555,970,992]" italics="true" pageId="22" pageNumber="121">ruficeps</emphasis>
Pocock, 1890
</taxonomicName>
. 74. Ultimate leg prefemur, ventral view, CES091327, scale = 1 mm. 75. Ultimate leg femur, dorsal view, CES091327, scale = 1 mm. 76. Spiracle on segment 3, CES091013, scale = 0.5 mm. 77. TT17–20, dorsal view, CES08960, scale = 1 mm.
</paragraph>
</caption>
<subSubSection id="8FC9656FFFC7FFAEFF55FBF5FDDDFB32" pageId="22" pageNumber="121" type="materials_examined">
<paragraph id="C76C36E4FFC7FFAEFF55FBF5FDDDFB32" blockId="22.[151,1437,1098,2022]" pageId="22" pageNumber="121">
<emphasis id="F5A7EAF6FFC7FFAEFF55FBF5FE21FB9F" bold="true" box="[201,450,1098,1123]" pageId="22" pageNumber="121">Material examined.</emphasis>
CES
<date id="B36D1024FFC7FFAEFD90FBF5FD80FB9E" box="[524,611,1098,1122]" pageId="22" pageNumber="121" value="13-09-10">091013</date>
, from Pushpagiri Reserve Forest, Karnataka; CES
<date id="B36D1024FFC7FFAEFB68FBF5FAA8FB9E" box="[1268,1355,1098,1122]" pageId="22" pageNumber="121" value="48-09-10">091048</date>
, from Parambikulam Tiger Reserve, Kerala; CES
<date id="B36D1024FFC7FFAEFD0CFBD1FD04FB7A" box="[656,743,1134,1158]" pageId="22" pageNumber="121" value="27-09-13">091327</date>
, CES
<date id="B36D1024FFC7FFAEFCB7FBD1FC61FB7A" box="[811,898,1134,1158]" pageId="22" pageNumber="121" value="30-09-13">091330</date>
, CES
<date id="B36D1024FFC7FFAEFC5BFBD1FBFEFB7A" box="[967,1053,1134,1158]" pageId="22" pageNumber="121" value="31-09-13">091331</date>
, from Siruvani Reserve Forests, Kerala; CES
<date id="B36D1024FFC7FFAEFEB6FB2DFE63FB56" box="[298,384,1170,1194]" pageId="22" pageNumber="121" value="21-09-10">091021</date>
, from Bisale Ghat Reserve Forests, Karanataka; CES
<date id="B36D1024FFC7FFAEFB98FB2DFBB8FB56" box="[1028,1115,1170,1194]" pageId="22" pageNumber="121" value="42-09-13">091342</date>
, from Achankovil Reserve Forest, Kerala. All leg. J. Joshi, 2009.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC7FFAFFF55FB65FD0FFEE0" lastPageId="23" lastPageNumber="122" pageId="22" pageNumber="121" type="description">
<paragraph id="C76C36E4FFC7FFAEFF55FB65FA7FFAC6" blockId="22.[151,1437,1098,2022]" pageId="22" pageNumber="121">
<emphasis id="F5A7EAF6FFC7FFAEFF55FB65FEB9FB0F" bold="true" box="[201,346,1242,1267]" pageId="22" pageNumber="121">Description.</emphasis>
[Based on CES material from Karnataka and Kerala] Length to
<quantity id="002B9B01FFC7FFAEFBB4FB65FB99FB0E" box="[1064,1146,1242,1266]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="22" pageNumber="121" unit="mm" value="36.0">36 mm</quantity>
. 21 antennal articles apart from one specimen with 17; basal 2–2.5 articles glabrous dorsally, usually 2.2–2.33, one specimen with third article glabrous apart from its distal half on inner side only. Longitudinal median furrow on anterior 25% of cephalic plate.
</paragraph>
<paragraph id="C76C36E4FFC7FFAEFF5BFAF9FAE2FA7E" blockId="22.[151,1437,1098,2022]" pageId="22" pageNumber="121">Cephalic plate and T1 brown or orange-brown except for blue anterior part of cephalic plate; TT2 or 3 to 21 mostly blue or with brown pigment increasing from ca T12; basal glabrous part of antenna pale blue.</paragraph>
<paragraph id="C76C36E4FFC7FFAEFF55FA31FD86F9EE" blockId="22.[151,1437,1098,2022]" pageId="22" pageNumber="121">
Forcipular coxosternal tooth plates with four main teeth (
<figureCitation id="5FE82A61FFC7FFAEFCFEFA31FC09FA5A" box="[866,1002,1422,1446]" captionStart="FIGURES 68 – 73" captionStartId="21.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1836]" captionTargetId="figure@21.[151,1436,337,1849]" captionTargetPageId="21" captionText="FIGURES 68 – 73. Otostigmus ruficeps Pocock, 1890. 68. Cephalic plate and T 1, dorsal view, CES 091013, scale = 2 mm. 69. Forcipular segment, ventral view, CES 091048, scale = 1 mm. 70. Tooth plates, ventral view, CES 001013, scale = 1 mm. 71. Coxopleuron, lateral view, CES 091330, scale = 1 mm. 72. Tergite of ultimate leg-bearing segment, dorsal view, CES 091330, scale = 2 mm. 73. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, CES 091330, scale = 1 mm." httpUri="https://zenodo.org/record/218637/files/figure.png" pageId="22" pageNumber="121">Figs 69, 70</figureCitation>
), inner two usually grouped together, outer tooth smaller than the inner three; tiny fifth tooth at inner margin on one side of two specimens. Trochanteroprefemoral process with three (
<figureCitation id="5FE82A61FFC7FFAEFD1BFA69FD38FA12" box="[647,731,1494,1518]" captionStart="FIGURES 68 – 73" captionStartId="21.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1836]" captionTargetId="figure@21.[151,1436,337,1849]" captionTargetPageId="21" captionText="FIGURES 68 – 73. Otostigmus ruficeps Pocock, 1890. 68. Cephalic plate and T 1, dorsal view, CES 091013, scale = 2 mm. 69. Forcipular segment, ventral view, CES 091048, scale = 1 mm. 70. Tooth plates, ventral view, CES 001013, scale = 1 mm. 71. Coxopleuron, lateral view, CES 091330, scale = 1 mm. 72. Tergite of ultimate leg-bearing segment, dorsal view, CES 091330, scale = 2 mm. 73. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, CES 091330, scale = 1 mm." httpUri="https://zenodo.org/record/218637/files/figure.png" pageId="22" pageNumber="121">Fig. 70</figureCitation>
) or usually four teeth, arranged as distal tooth and two teeth along inner margin or four uniform teeth.
</paragraph>
<paragraph id="C76C36E4FFC7FFAEFF55F9A1FED2F9A6" blockId="22.[151,1437,1098,2022]" pageId="22" pageNumber="121">Second maxillary claw with slender, needle-like accessory spurs. Article 2 of telopodite bearing a slender spine distally.</paragraph>
<paragraph id="C76C36E4FFC7FFAEFF55F9D9FB12F95E" blockId="22.[151,1437,1098,2022]" pageId="22" pageNumber="121">
Tergites with paramedian sutures complete from TT5 to 7; tergites marginate from 6 to 8. Tergites smooth, lacking median ridge or keels (
<figureCitation id="5FE82A61FFC7FFAEFE6DF935FDA5F95E" box="[497,582,1674,1698]" captionStart="FIGURES 74 – 77" captionStartId="22.[151,264,970,993]" captionTargetBox="[395,1198,193,948]" captionTargetId="figure@22.[388,1198,193,949]" captionTargetPageId="22" captionText="FIGURES 74 – 77. Otostigmus ruficeps Pocock, 1890. 74. Ultimate leg prefemur, ventral view, CES 091327, scale = 1 mm. 75. Ultimate leg femur, dorsal view, CES 091327, scale = 1 mm. 76. Spiracle on segment 3, CES 091013, scale = 0.5 mm. 77. TT 17 – 20, dorsal view, CES 08960, scale = 1 mm." httpUri="https://zenodo.org/record/218638/files/figure.png" pageId="22" pageNumber="121">Fig. 77</figureCitation>
). Sternites with paramedian sutures confined to anterior part.
</paragraph>
<paragraph id="C76C36E4FFC7FFAEFF55F910FA64F8F2" blockId="22.[151,1437,1098,2022]" pageId="22" pageNumber="121">
Two tarsal spurs on legs
<date id="B36D1024FFC7FFAEFE7BF911FD9DF93A" box="[487,638,1710,1734]" pageId="22" pageNumber="121" value="1919-01-17" valueMax="1919-01-01" valueMin="1919-01-17">1–17 to 1–19</date>
except for one individual with two spurs on legs 1–4 only; one tarsal spur on leg 20; a tibial spur on legs 1–6, 1–7 or 1–8 except for one individual with spur on leg 1 only (same specimen as with restricted number of legs with two tarsal spurs). A femoral spur variably present on leg 1 only.
</paragraph>
<paragraph id="C76C36E4FFC7FFAEFF55F8A5FAF1F862" blockId="22.[151,1437,1098,2022]" pageId="22" pageNumber="121">
Tergite of ultimate leg-bearing segment with nearly parallel lateral margins, usually with straight posterolateral margins, pointed posteromedially (
<figureCitation id="5FE82A61FFC7FFAEFDACF881FD68F8AA" box="[560,651,1854,1878]" captionStart="FIGURES 68 – 73" captionStartId="21.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1836]" captionTargetId="figure@21.[151,1436,337,1849]" captionTargetPageId="21" captionText="FIGURES 68 – 73. Otostigmus ruficeps Pocock, 1890. 68. Cephalic plate and T 1, dorsal view, CES 091013, scale = 2 mm. 69. Forcipular segment, ventral view, CES 091048, scale = 1 mm. 70. Tooth plates, ventral view, CES 001013, scale = 1 mm. 71. Coxopleuron, lateral view, CES 091330, scale = 1 mm. 72. Tergite of ultimate leg-bearing segment, dorsal view, CES 091330, scale = 2 mm. 73. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, CES 091330, scale = 1 mm." httpUri="https://zenodo.org/record/218637/files/figure.png" pageId="22" pageNumber="121">Fig. 72</figureCitation>
); one specimen with a more transverse posterior margin; shallow longitudinal median depression on posterior half of tergite. Sternite of ultimate leg-bearing segment with sides converging relatively strongly posteriorly, nearly straight, posterior margin variably concave (
<figureCitation id="5FE82A61FFC7FFAEFB2EF839FAE6F862" box="[1202,1285,1926,1950]" captionStart="FIGURES 68 – 73" captionStartId="21.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1836]" captionTargetId="figure@21.[151,1436,337,1849]" captionTargetPageId="21" captionText="FIGURES 68 – 73. Otostigmus ruficeps Pocock, 1890. 68. Cephalic plate and T 1, dorsal view, CES 091013, scale = 2 mm. 69. Forcipular segment, ventral view, CES 091048, scale = 1 mm. 70. Tooth plates, ventral view, CES 001013, scale = 1 mm. 71. Coxopleuron, lateral view, CES 091330, scale = 1 mm. 72. Tergite of ultimate leg-bearing segment, dorsal view, CES 091330, scale = 2 mm. 73. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, CES 091330, scale = 1 mm." httpUri="https://zenodo.org/record/218637/files/figure.png" pageId="22" pageNumber="121">Fig. 73</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFC7FFAFFF55F815FD0FFEE0" blockId="22.[151,1437,1098,2022]" lastBlockId="23.[151,1436,151,1293]" lastPageId="23" lastPageNumber="122" pageId="22" pageNumber="121">
Coxopleural process moderately long, not obviously inflected from posterolateral margin of coxopleuron in ventral view (
<figureCitation id="5FE82A61FFC7FFAEFEADF871FE67F81A" box="[305,388,1998,2022]" captionStart="FIGURES 68 – 73" captionStartId="21.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1836]" captionTargetId="figure@21.[151,1436,337,1849]" captionTargetPageId="21" captionText="FIGURES 68 – 73. Otostigmus ruficeps Pocock, 1890. 68. Cephalic plate and T 1, dorsal view, CES 091013, scale = 2 mm. 69. Forcipular segment, ventral view, CES 091048, scale = 1 mm. 70. Tooth plates, ventral view, CES 001013, scale = 1 mm. 71. Coxopleuron, lateral view, CES 091330, scale = 1 mm. 72. Tergite of ultimate leg-bearing segment, dorsal view, CES 091330, scale = 2 mm. 73. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, CES 091330, scale = 1 mm." httpUri="https://zenodo.org/record/218637/files/figure.png" pageId="22" pageNumber="121">Fig. 73</figureCitation>
), with two apical spines except for one individual with three apical spines on one side; lateral spine lacking except for one individual with one lateral spine on one side. Pores relatively dense; non-porose area on coxopleural process long, narrow, reaching posterior margin of sternite of ultimate leg-bearing segment (
<figureCitation id="5FE82A61FFC6FFAFFAF2FF03FF54FF0B" captionStart="FIGURES 68 – 73" captionStartId="21.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1836]" captionTargetId="figure@21.[151,1436,337,1849]" captionTargetPageId="21" captionText="FIGURES 68 – 73. Otostigmus ruficeps Pocock, 1890. 68. Cephalic plate and T 1, dorsal view, CES 091013, scale = 2 mm. 69. Forcipular segment, ventral view, CES 091048, scale = 1 mm. 70. Tooth plates, ventral view, CES 001013, scale = 1 mm. 71. Coxopleuron, lateral view, CES 091330, scale = 1 mm. 72. Tergite of ultimate leg-bearing segment, dorsal view, CES 091330, scale = 2 mm. 73. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, CES 091330, scale = 1 mm." httpUri="https://zenodo.org/record/218637/files/figure.png" pageId="23" pageNumber="122">Fig. 73</figureCitation>
). Ultimate leg prefemoral spines: VL 2 or 3, VM 2, DM 0 (
<figureCitation id="5FE82A61FFC6FFAFFCC8FF60FC4AFF0B" box="[852,937,223,247]" captionStart="FIGURES 74 – 77" captionStartId="22.[151,264,970,993]" captionTargetBox="[395,1198,193,948]" captionTargetId="figure@22.[388,1198,193,949]" captionTargetPageId="22" captionText="FIGURES 74 – 77. Otostigmus ruficeps Pocock, 1890. 74. Ultimate leg prefemur, ventral view, CES 091327, scale = 1 mm. 75. Ultimate leg femur, dorsal view, CES 091327, scale = 1 mm. 76. Spiracle on segment 3, CES 091013, scale = 0.5 mm. 77. TT 17 – 20, dorsal view, CES 08960, scale = 1 mm." httpUri="https://zenodo.org/record/218638/files/figure.png" pageId="23" pageNumber="122">Fig. 74</figureCitation>
). Ultimate leg tarsus 1 1.7–2 times length of tarsus 2; pretarsus with pair of short accessory claws.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC6FFAFFF55FE98FAA8FAF0" pageId="23" pageNumber="122" type="discussion">
<paragraph id="C76C36E4FFC6FFAFFF55FE98FD18FCA0" blockId="23.[151,1436,151,1293]" pageId="23" pageNumber="122">
<emphasis id="F5A7EAF6FFC6FFAFFF55FE98FEADFEBC" bold="true" box="[201,334,295,320]" pageId="23" pageNumber="122">Discussion.</emphasis>
This species was resolved as sister species to all other Indian
<taxonomicName id="00D34D67FFC6FFAFFC61FE96FB87FEBC" box="[1021,1124,297,320]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC6FFAFFC61FE96FB87FEBC" box="[1021,1124,297,320]" italics="true" pageId="23" pageNumber="122">Digitipes</emphasis>
</taxonomicName>
in the molecular phylogeny (Joshi and Karanth 2012), and the clade that unites the other species had strong node support (posterior probability 1, bootstrap support 100%). No specimens have a distomedial process on the femur of the ultimate leg (
<figureCitation id="5FE82A61FFC6FFAFFAA5FED0FA6DFE7B" box="[1337,1422,367,391]" captionStart="FIGURES 74 – 77" captionStartId="22.[151,264,970,993]" captionTargetBox="[395,1198,193,948]" captionTargetId="figure@22.[388,1198,193,949]" captionTargetPageId="22" captionText="FIGURES 74 – 77. Otostigmus ruficeps Pocock, 1890. 74. Ultimate leg prefemur, ventral view, CES 091327, scale = 1 mm. 75. Ultimate leg femur, dorsal view, CES 091327, scale = 1 mm. 76. Spiracle on segment 3, CES 091013, scale = 0.5 mm. 77. TT 17 – 20, dorsal view, CES 08960, scale = 1 mm." httpUri="https://zenodo.org/record/218638/files/figure.png" pageId="23" pageNumber="122">Fig. 75</figureCitation>
), rendering assignment to
<taxonomicName id="00D34D67FFC6FFAFFE37FE2AFDF1FE50" box="[427,530,405,428]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC6FFAFFE37FE2AFDF1FE50" box="[427,530,405,428]" italics="true" pageId="23" pageNumber="122">Digitipes</emphasis>
</taxonomicName>
unproven. Most specimens of this species differ from Indian
<taxonomicName id="00D34D67FFC6FFAFFB59FE2AFACFFE50" box="[1221,1324,405,428]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC6FFAFFB59FE2AFACFFE50" box="[1221,1324,405,428]" italics="true" pageId="23" pageNumber="122">Digitipes</emphasis>
</taxonomicName>
in having two tarsal spurs and a tibial spur on much more posterior segments than in the other species (i.e., two tarsal spurs to legs 17–19 versus not further than to leg 14, and usually confined to legs 1–3 or 1–4; a tibial spur to legs 6–8 versus confined to leg 1 only). We note, however, that a specimen nested within the molecular phylogeny for the species (CES
<date id="B36D1024FFC6FFAFFF4DFD9BFECBFDC0" box="[209,296,548,572]" pageId="23" pageNumber="122" value="42-09-13">091342</date>
) has two tarsal spurs only as far as leg 4 and a tibial spur on leg 1 only. Other characters that differ from Indian
<taxonomicName id="00D34D67FFC6FFAFFEBFFDF6FE69FD9C" box="[291,394,585,608]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC6FFAFFEBFFDF6FE69FD9C" box="[291,394,585,608]" italics="true" pageId="23" pageNumber="122">Digitipes</emphasis>
</taxonomicName>
are the non-exact transition to glabrous antennal articles (a partial glabrous extent is seen on article 3 versus an abrupt transition between articles 2 and 3, 3 and 4, or 4 and
<quantity id="002B9B01FFC6FFAFFBB3FDD3FBBFFD78" box="[1071,1116,620,644]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="23" pageNumber="122" unit="in" value="5.0">5 in</quantity>
all Indian
<taxonomicName id="00D34D67FFC6FFAFFB47FDD2FAA2FD78" box="[1243,1345,621,644]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC6FFAFFB47FDD2FAA2FD78" box="[1243,1345,621,644]" italics="true" pageId="23" pageNumber="122">Digitipes</emphasis>
</taxonomicName>
studied here), and the longitudinal depression on the tergite of the ultimate leg-bearing segment. These characters are shared by
<taxonomicName id="00D34D67FFC6FFAFFE90FD0AFD71FD30" authority="Pocock, 1890" authorityName="Pocock" authorityYear="1890" box="[268,658,692,716]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC6FFAFFE90FD0AFE0DFD30" box="[268,494,692,716]" italics="true" pageId="23" pageNumber="122">Otostigmus ruficeps</emphasis>
Pocock, 1890
</taxonomicName>
, until now known only from its
<typeStatus id="18688846FFC6FFAFFB9CFD0BFB86FD30" box="[1024,1125,692,716]" pageId="23" pageNumber="122" type="holotype">holotype</typeStatus>
, from Madras. Comparison of the
<typeStatus id="18688846FFC6FFAFFF72FD68FEB3FD13" box="[238,336,727,751]" pageId="23" pageNumber="122" type="holotype">holotype</typeStatus>
(NHM 1890.1.23.22) and our collections strongly indicates conspecificity, and the species diagnosis is revised above based on shared characters that provide a distinctive combination within
<taxonomicName id="00D34D67FFC6FFAFFA80FD42FEC8FCCB" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC6FFAFFA80FD42FA78FCE8" box="[1308,1435,765,788]" italics="true" pageId="23" pageNumber="122">Otostigmus</emphasis>
(
<emphasis id="F5A7EAF6FFC6FFAFFF3CFC9FFEC3FCCB" box="[160,288,800,823]" italics="true" pageId="23" pageNumber="122">Otostigmus</emphasis>
)
</taxonomicName>
. Characters of the ultimate leg are added to the diagnosis based on our material because the
<typeStatus id="18688846FFC6FFAFFAA6FCA0FA7FFCCB" box="[1338,1436,799,823]" pageId="23" pageNumber="122" type="holotype">holotype</typeStatus>
lacked ultimate legs even at the time of its description.
</paragraph>
<paragraph id="C76C36E4FFC6FFAFFF48FCD8FAA8FAF0" blockId="23.[151,1436,151,1293]" pageId="23" pageNumber="122">
Lewis (1996) redescribed the
<typeStatus id="18688846FFC6FFAFFDA8FCD8FD75FC83" box="[564,662,871,895]" pageId="23" pageNumber="122" type="holotype">holotype</typeStatus>
of
<taxonomicName id="00D34D67FFC6FFAFFD58FCD7FCA6FC7C" box="[708,837,872,896]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC6FFAFFD58FCD7FCA6FC7C" box="[708,837,872,896]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFD58FCD7FD38FC83" box="[708,731,872,895]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. ruficeps
</emphasis>
</taxonomicName>
and noted that the species keys out to
<taxonomicName id="00D34D67FFC6FFAFFA80FCD7FE4EFC58" authority="Haase, 1887" authorityName="Haase" authorityYear="1887" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="F5A7EAF6FFC6FFAFFA80FCD7FEE7FC58" italics="true" pageId="23" pageNumber="122">Otostigmus multidens</emphasis>
(Haase, 1887)
</taxonomicName>
using the standard keys (Kraepelin 1903; Attems 1930b). He noted that the number of forcipular teeth (5+5) in the
<typeStatus id="18688846FFC6FFAFFE40FC10FDDDFC3B" box="[476,574,943,967]" pageId="23" pageNumber="122" type="holotype">holotype</typeStatus>
of
<taxonomicName id="00D34D67FFC6FFAFFDFAFC0FFD07FC34" box="[614,740,944,968]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC6FFAFFDFAFC0FFD07FC34" box="[614,740,944,968]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFDFAFC0FFD9EFC3B" box="[614,637,944,967]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. ruficeps
</emphasis>
</taxonomicName>
is at the extremity of the range observed in
<taxonomicName id="00D34D67FFC6FFAFFB43FC0FFA93FC34" box="[1247,1392,944,968]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="F5A7EAF6FFC6FFAFFB43FC0FFA93FC34" box="[1247,1392,944,968]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFB43FC0FFB15FC3B" box="[1247,1270,944,967]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. multidens
</emphasis>
</taxonomicName>
but concluded that no clear distinction could be made between the two species, and proposed the junior synonymy of
<taxonomicName id="00D34D67FFC6FFAFFF0BFC47FEF7FBEC" box="[151,276,1016,1040]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC6FFAFFF0BFC47FEF7FBEC" box="[151,276,1016,1040]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFF0BFC47FF4DFBF3" box="[151,174,1016,1039]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. ruficeps
</emphasis>
</taxonomicName>
. The present collections suggest that the small inner tooth on the tooth plates of the
<typeStatus id="18688846FFC6FFAFFB50FC48FACDFBF3" box="[1228,1326,1015,1039]" pageId="23" pageNumber="122" type="holotype">holotype</typeStatus>
relates to variability within a species that more commonly has 4+4 large teeth. Redescription of
<taxonomicName id="00D34D67FFC6FFAFFB0EFBA2FAC5FBC8" box="[1170,1318,1052,1076]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="F5A7EAF6FFC6FFAFFB0EFBA2FAC5FBC8" box="[1170,1318,1052,1076]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFB0EFBA2FB4AFBC8" box="[1170,1193,1053,1076]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. multidens
</emphasis>
</taxonomicName>
by Lewis (2001) and examination of material of that species from
<collectingCountry id="BFC47674FFC6FFAFFC91FB80FC90FBAB" box="[781,883,1087,1111]" name="Thailand" pageId="23" pageNumber="122">Thailand</collectingCountry>
,
<collectingCountry id="BFC47674FFC6FFAFFCE2FB80FC3EFBAB" box="[894,989,1087,1111]" name="Vietnam" pageId="23" pageNumber="122">Vietnam</collectingCountry>
and New
<collectingCountry id="BFC47674FFC6FFAFFBD3FB80FB43FBAB" box="[1103,1184,1087,1111]" name="Guinea-Bissau" pageId="23" pageNumber="122">Guinea</collectingCountry>
lead us to maintain
<taxonomicName id="00D34D67FFC6FFAFFA1DFBFFFF12FB80" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC6FFAFFA1DFBFFFF12FB80" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFA1DFBFFFA7BFBAB" box="[1409,1432,1088,1111]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. ruficeps
</emphasis>
</taxonomicName>
as a valid species from
<collectingCountry id="BFC47674FFC6FFAFFE62FBDBFDD8FB80" box="[510,571,1124,1148]" name="India" pageId="23" pageNumber="122">India</collectingCountry>
.
<taxonomicName id="00D34D67FFC6FFAFFDDAFBDAFD36FB80" box="[582,725,1124,1148]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="F5A7EAF6FFC6FFAFFDDAFBDAFD36FB80" box="[582,725,1124,1148]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFDDAFBDAFDBEFB80" box="[582,605,1125,1148]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. multidens
</emphasis>
</taxonomicName>
has small teeth interpolated between some of the large teeth on the tooth plates, and has tibial spurs extending as far posteriorly as legs 16–18 (versus legs
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the
<typeStatus id="18688846FFC6FFAFFA87FB37FA9EFB5C" box="[1307,1405,1160,1184]" pageId="23" pageNumber="122" type="holotype">holotype</typeStatus>
of
<taxonomicName id="00D34D67FFC6FFAFFF0BFB12FEF0FB38" box="[151,275,1196,1220]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC6FFAFFF0BFB12FEF0FB38" box="[151,275,1196,1220]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFF0BFB12FF4DFB38" box="[151,174,1197,1220]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. ruficeps
</emphasis>
</taxonomicName>
and legs 1–6, 7 or
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our specimens). These differences are consistent with separate species, although the shared presence of four teeth on the forcipular trochanteroprefemoral process and a longitudinal depression on the tergite of the ultimate leg-bearing segment suggest that
<taxonomicName id="00D34D67FFC6FFAFFCB3FB4AFC48FAF0" box="[815,939,1268,1292]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="ruficeps">
<emphasis id="F5A7EAF6FFC6FFAFFCB3FB4AFC48FAF0" box="[815,939,1268,1292]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFCB3FB4AFCA5FAF0" box="[815,838,1269,1292]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. ruficeps
</emphasis>
</taxonomicName>
and
<taxonomicName id="00D34D67FFC6FFAFFC7EFB4AFB92FAF0" box="[994,1137,1268,1292]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="122" phylum="Arthropoda" rank="species" species="multidens">
<emphasis id="F5A7EAF6FFC6FFAFFC7EFB4AFB92FAF0" box="[994,1137,1268,1292]" italics="true" pageId="23" pageNumber="122">
<collectingCountry id="BFC47674FFC6FFAFFC7EFB4AFC1AFAF0" box="[994,1017,1269,1292]" name="Iceland" pageId="23" pageNumber="122">O</collectingCountry>
. multidens
</emphasis>
</taxonomicName>
are closely related.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="4F7A87F2FFD0FFBBFF0BFA97FADCFEC3" ID-DOI="http://doi.org/10.5281/zenodo.6154112" ID-GBIF-Taxon="119563410" ID-Zenodo-Dep="6154112" LSID="urn:lsid:plazi:treatment:4F7A87F2FFD0FFBBFF0BFA97FADCFEC3" httpUri="http://treatment.plazi.org/id/4F7A87F2FFD0FFBBFF0BFA97FADCFEC3" lastPageId="3" lastPageNumber="101" pageId="1" pageNumber="100">
<subSubSection id="8FC9656FFFD0FFB9FF0BFA97FDE7FABE" box="[151,516,1320,1347]" pageId="1" pageNumber="100" type="nomenclature">
<paragraph id="C76C36E4FFD0FFB9FF0BFA97FDE7FABE" blockId="1.[151,516,1320,1347]" box="[151,516,1320,1347]" pageId="1" pageNumber="100">
<heading id="9C248188FFD0FFB9FF0BFA97FDE7FABE" bold="true" box="[151,516,1320,1347]" fontSize="11" level="1" pageId="1" pageNumber="100" reason="1">
<emphasis id="F5A7EAF6FFD0FFB9FF0BFA97FDE7FABE" bold="true" box="[151,516,1320,1347]" pageId="1" pageNumber="100">
Genus
<taxonomicName id="00D34D67FFD0FFB9FF72FA96FDE7FABE" ID-CoL="45DM" ID-ENA="1096233" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[238,516,1320,1347]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD0FFB9FF72FA96FEB9FABF" bold="true" box="[238,346,1321,1347]" italics="true" pageId="1" pageNumber="100">Digitipes</emphasis>
Attems, 1930
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD0FFB9FF0BFACFFC8BFA74" box="[151,872,1391,1416]" pageId="1" pageNumber="100" type="materials_examined">
<paragraph id="C76C36E4FFD0FFB9FF0BFACFFC8BFA74" blockId="1.[151,1437,1391,2032]" box="[151,872,1391,1416]" pageId="1" pageNumber="100">
<emphasis id="F5A7EAF6FFD0FFB9FF0BFACFFED1FA74" bold="true" box="[151,306,1391,1416]" pageId="1" pageNumber="100">
<typeStatus id="18688846FFD0FFB9FF0BFACFFF31FA74" box="[151,210,1392,1416]" pageId="1" pageNumber="100">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="00D34D67FFD0FFB9FEA5FACEFD20FA7B" authority="Attems, 1930" authorityName="Attems" authorityYear="1930" box="[313,707,1391,1416]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="verdascens">
<emphasis id="F5A7EAF6FFD0FFB9FEA5FACEFDC1FA74" box="[313,546,1392,1416]" italics="true" pageId="1" pageNumber="100">Digitipes verdascens</emphasis>
Attems, 1930
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD0FFB9FF55FA2CFD64F9E4" pageId="1" pageNumber="100" type="diagnosis">
<paragraph id="C76C36E4FFD0FFB9FF55FA2CFD64F9E4" blockId="1.[151,1437,1391,2032]" pageId="1" pageNumber="100">
<emphasis id="F5A7EAF6FFD0FFB9FF55FA2CFEA0FA50" bold="true" box="[201,323,1427,1452]" pageId="1" pageNumber="100">Diagnosis.</emphasis>
Otostigmini lacking spiracle on segment 7. Forcipular tooth plates with four teeth, the inner two grouped. Coxopleural process with two apical spines, no dorsal spines, single lateral spine present or absent. Conical to subcyclindrical distomedial process on femur of ultimate leg in male, longitudinal furrow along medial surface of femur terminating on the process.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD0FFBBFF55F99CFADCFEC3" lastPageId="3" lastPageNumber="102" pageId="1" pageNumber="100" type="discussion">
<paragraph id="C76C36E4FFD0FFB9FF55F99CFAA7F8C4" blockId="1.[151,1437,1391,2032]" pageId="1" pageNumber="100">
<emphasis id="F5A7EAF6FFD0FFB9FF55F99CFEADF9C0" bold="true" box="[201,334,1571,1596]" pageId="1" pageNumber="100">Discussion.</emphasis>
<taxonomicName id="00D34D67FFD0FFB9FEC8F99AFE58F9C0" box="[340,443,1573,1596]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD0FFB9FEC8F99AFE58F9C0" box="[340,443,1573,1596]" italics="true" pageId="1" pageNumber="100">Digitipes</emphasis>
</taxonomicName>
was erected as a monotypic genus to accommodate
<taxonomicName id="00D34D67FFD0FFB9FB9FF99AFB42F9C0" box="[1027,1185,1572,1596]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="verdascens">
<emphasis id="F5A7EAF6FFD0FFB9FB9FF99AFB42F9C0" box="[1027,1185,1572,1596]" italics="true" pageId="1" pageNumber="100">D. verdascens</emphasis>
</taxonomicName>
from the then
<collectingCountry id="BFC47674FFD0FFB9FAD9F99BFF01F9A3" name="Democratic Republic of the Congo" pageId="1" pageNumber="100">Belgian Congo</collectingCountry>
(
<collectingCountry id="BFC47674FFD0FFB9FF6DF9F8FDBAF9A3" box="[241,601,1607,1631]" name="Democratic Republic of the Congo" pageId="1" pageNumber="100">Democratic Republic of Congo</collectingCountry>
). Attems (1930a) noted that it was easily distinguished from other genera of Otostigmini by the process on the femur of the ultimate legs of the male. Two additional species with
<typeStatus id="18688846FFD0FFB9FAB8F9D2FABCF979" box="[1316,1375,1645,1669]" pageId="1" pageNumber="100">types</typeStatus>
from the
<collectingCountry id="BFC47674FFD0FFB9FF5DF930FDC2F95B" box="[193,545,1679,1703]" name="Democratic Republic of the Congo" pageId="1" pageNumber="100">Democratic Republic of Congo</collectingCountry>
were subsequently attributed to the genus:
<taxonomicName id="00D34D67FFD0FFB9FB96F92EFAA6F95B" authority="Kraus, 1957" authorityName="Kraus" authorityYear="1957" box="[1034,1349,1679,1704]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="katangensis">
<emphasis id="F5A7EAF6FFD0FFB9FB96F92EFB52F954" box="[1034,1201,1680,1704]" italics="true" pageId="1" pageNumber="100">D. katangensis</emphasis>
Kraus, 1957
</taxonomicName>
, and
<taxonomicName id="00D34D67FFD0FFB9FA1DF92EFE45F930" authority="Dobroruka, 1968" authorityName="Dobroruka" authorityYear="1968" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="krausi">
<emphasis id="F5A7EAF6FFD0FFB9FA1DF92EFF3EF930" italics="true" pageId="1" pageNumber="100">D. krausi</emphasis>
Dobroruka, 1968
</taxonomicName>
. The former at least demonstrably has the distomedial process on the ultimate leg femur in the male (Kraus, 1957: figs. 8–10), whereas the latter is known only from the female.
<taxonomicName id="00D34D67FFD0FFB9FBE8F966FA84F90C" box="[1140,1383,1752,1776]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="katangensis">
<emphasis id="F5A7EAF6FFD0FFB9FBE8F966FA84F90C" box="[1140,1383,1752,1776]" italics="true" pageId="1" pageNumber="100">Digitipes katangensis</emphasis>
</taxonomicName>
was placed in synonymy with
<taxonomicName id="00D34D67FFD0FFB9FE26F942FC8DF8E8" authority="Kraepelin, 1903" authorityName="Kraepelin" authorityYear="1903" box="[442,878,1788,1812]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="reichardti">
<emphasis id="F5A7EAF6FFD0FFB9FE26F942FD4CF8E8" box="[442,687,1788,1812]" italics="true" pageId="1" pageNumber="100">Otostigmus reichardti</emphasis>
Kraepelin, 1903
</taxonomicName>
, by Lewis (2004), extending its geographic range to
<collectingCountry id="BFC47674FFD0FFB9FF28F89FFEFAF8C4" box="[180,281,1824,1848]" name="Tanzania" pageId="1" pageNumber="100">Tanzania</collectingCountry>
and
<collectingCountry id="BFC47674FFD0FFB9FED3F89FFE46F8C4" box="[335,421,1824,1848]" name="Angola" pageId="1" pageNumber="100">Angola</collectingCountry>
. The current combination for this species is
<taxonomicName id="00D34D67FFD0FFB9FC08F89EFAA3F8C4" authority="Kraepelin, 1903" authorityName="Kraepelin" authorityYear="1903" box="[916,1344,1824,1848]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="reichardti">
<emphasis id="F5A7EAF6FFD0FFB9FC08F89EFB8CF8C4" box="[916,1135,1824,1848]" italics="true" pageId="1" pageNumber="100">Digitipes reichardti</emphasis>
(Kraepelin, 1903)
</taxonomicName>
.
</paragraph>
<paragraph id="C76C36E4FFD0FFBAFF55F8FBFA98FF28" blockId="1.[151,1437,1391,2032]" lastBlockId="2.[151,1437,151,2012]" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100">
The first attribution of non-African species to
<taxonomicName id="00D34D67FFD0FFB9FD52F8FAFCD6F8A0" box="[718,821,1861,1884]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD0FFB9FD52F8FAFCD6F8A0" box="[718,821,1861,1884]" italics="true" pageId="1" pageNumber="100">Digitipes</emphasis>
</taxonomicName>
was made by Jangi and Dass (1984), who assigned six new species from peninsular
<collectingCountry id="BFC47674FFD0FFB9FE7DF8D7FDF9F87C" box="[481,538,1896,1920]" name="India" pageId="1" pageNumber="100">India</collectingCountry>
to the genus. Their justification for this assignment was the shared presence of two characters in these species that separated
<taxonomicName id="00D34D67FFD0FFB9FD05F832FD1CF858" box="[665,767,1933,1956]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD0FFB9FD05F832FD1CF858" box="[665,767,1933,1956]" italics="true" pageId="1" pageNumber="100">Digitipes</emphasis>
</taxonomicName>
from
<taxonomicName id="00D34D67FFD0FFB9FCD9F832FC26F858" box="[837,965,1933,1956]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD0FFB9FCD9F832FC26F858" box="[837,965,1933,1956]" italics="true" pageId="1" pageNumber="100">Otostigmus</emphasis>
</taxonomicName>
in their key: the femur of the ultimate leg in the male bears a distomedial process and “claw of 2nd maxilla without spur”. The latter refers to a lack of accessory spurs on the pretarsus, and was also listed by Attems (1930a, b) as diagnostic of
<taxonomicName id="00D34D67FFD0FFB9FB54F867FACCF813" box="[1224,1327,2008,2031]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD0FFB9FB54F867FACCF813" box="[1224,1327,2008,2031]" italics="true" pageId="1" pageNumber="100">Digitipes</emphasis>
</taxonomicName>
. Another second maxillary character was also regarded as diagnostic of the genus by Attems (1930a, b), the lack of a spine on article 2 of the telopodite (presence of a spine in this position being widespread throughout
<taxonomicName id="00D34D67FFD3FFBAFB27FF03FA8CFF28" box="[1211,1391,188,212]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="family">Scolopendridae</taxonomicName>
).
</paragraph>
<paragraph id="C76C36E4FFD3FFBAFF5BFF60FDF6FD30" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="101">
Lewis (2004) cautioned that
<taxonomicName id="00D34D67FFD3FFBAFD96FF5EFD92FF04" box="[522,625,225,248]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFD96FF5EFD92FF04" box="[522,625,225,248]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
could prove to be a junior synonym of
<taxonomicName id="00D34D67FFD3FFBAFBB1FF5FFADBFF0B" authority="Porat, 1876" authorityName="Porat" authorityYear="1876" box="[1069,1336,223,247]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFBB1FF5FFB4EFF0B" box="[1069,1197,224,247]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
Porat, 1876
</taxonomicName>
, because its only reliable diagnostic character (the process on the ultimate leg femur) is confined to males. Lewis (2004) pointed out instances within
<taxonomicName id="00D34D67FFD3FFBAFE78FE97FD87FEC3" box="[484,612,296,319]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFE78FE97FD87FEC3" box="[484,612,296,319]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
in which an absence of a spine on article 2 of the telopodite had been recorded, e.g., in
<taxonomicName id="00D34D67FFD3FFBAFEC0FEF2FD57FE98" authority="Schileyko, 1995" authorityName="Schileyko" authorityYear="1995" box="[348,692,332,356]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="reservatus">
<emphasis id="F5A7EAF6FFD3FFBAFEC0FEF2FE10FE98" box="[348,499,333,356]" italics="true" pageId="2" pageNumber="101">
<collectingCountry id="BFC47674FFD3FFBAFEC0FEF2FE90FE98" box="[348,371,333,356]" name="Iceland" pageId="2" pageNumber="101">O</collectingCountry>
. reservatus
</emphasis>
Schileyko, 1995
</taxonomicName>
, such that this character would not consistently separate
<taxonomicName id="00D34D67FFD3FFBAFAA9FEF2FA7FFE98" box="[1333,1436,333,356]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFAA9FEF2FA7FFE98" box="[1333,1436,333,356]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
from
<taxonomicName id="00D34D67FFD3FFBAFF45FECFFEBAFE7B" box="[217,345,368,391]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFF45FECFFEBAFE7B" box="[217,345,368,391]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
. The absence of this spine has also been recorded in various South American species of
<collectingCountry id="BFC47674FFD3FFBAFA1DFECFFA76FE7B" box="[1409,1429,368,391]" name="Iceland" pageId="2" pageNumber="101">
<emphasis id="F5A7EAF6FFD3FFBAFA1DFECFFA76FE7B" box="[1409,1429,368,391]" italics="true" pageId="2" pageNumber="101">O</emphasis>
</collectingCountry>
. (
<emphasis id="F5A7EAF6FFD3FFBAFF3CFE2AFEA1FE50" box="[160,322,405,428]" italics="true" pageId="2" pageNumber="101">Parotostigmus</emphasis>
) by Chagas-Jr.
<emphasis id="F5A7EAF6FFD3FFBAFE6FFE2AFDC7FE50" box="[499,548,404,428]" italics="true" pageId="2" pageNumber="101">et al</emphasis>
. (2007), who likewise questioned the separation of
<taxonomicName id="00D34D67FFD3FFBAFBF6FE2AFB32FE50" box="[1130,1233,405,428]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFBF6FE2AFB32FE50" box="[1130,1233,405,428]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
from
<taxonomicName id="00D34D67FFD3FFBAFA89FE2AFA77FE50" box="[1301,1428,405,428]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFA89FE2AFA77FE50" box="[1301,1428,405,428]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
. An even more serious defect of this character as regards the assignment of Indian species to
<taxonomicName id="00D34D67FFD3FFBAFB3AFE06FAEEFE2C" box="[1190,1293,441,464]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFB3AFE06FAEEFE2C" box="[1190,1293,441,464]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
, however, is that all of them in fact have a slender, pigmented spine on segment 2, such that its absence may be restricted to African species alone. Lewis (2004) was likewise critical of the utility of the pretarsal accessory spurs on the second maxilla as a diagnostic character of
<taxonomicName id="00D34D67FFD3FFBAFD1CFD9AFD04FDC0" box="[640,743,549,572]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFD1CFD9AFD04FDC0" box="[640,743,549,572]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
because the size of these spurs is so variable between species of
<taxonomicName id="00D34D67FFD3FFBAFF29FDF7FED7FDA3" box="[181,308,584,607]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFF29FDF7FED7FDA3" box="[181,308,584,607]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
, ranging to rudimentary expression in some species. Within
<taxonomicName id="00D34D67FFD3FFBAFC42FDF6FBA6FD9C" box="[990,1093,585,608]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFC42FDF6FBA6FD9C" box="[990,1093,585,608]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
an absence of accessory spurs also appears to be confined to African species (
<taxonomicName id="00D34D67FFD3FFBAFD3BFDD2FCA7FD78" box="[679,836,620,644]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="verdascens">
<emphasis id="F5A7EAF6FFD3FFBAFD3BFDD2FCA7FD78" box="[679,836,620,644]" italics="true" pageId="2" pageNumber="101">D. verdascens</emphasis>
</taxonomicName>
and
<taxonomicName id="00D34D67FFD3FFBAFCE7FDD2FBE8FD78" box="[891,1035,620,644]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="reichardti">
<emphasis id="F5A7EAF6FFD3FFBAFCE7FDD2FBE8FD78" box="[891,1035,620,644]" italics="true" pageId="2" pageNumber="101">D. reichardti</emphasis>
</taxonomicName>
) because we observe their presence in the three Indian species assigned to
<taxonomicName id="00D34D67FFD3FFBAFDD5FD2EFD53FD54" box="[585,688,657,680]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFDD5FD2EFD53FD54" box="[585,688,657,680]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
by Jangi and Dass (1984) that we have examined as well as in the two new species described herein.
</paragraph>
<paragraph id="C76C36E4FFD3FFBAFF5BFD68FB54FACC" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="101">
Thus we agree with Lewis (2004) and Chagas
<emphasis id="F5A7EAF6FFD3FFBAFD56FD66FD18FD0C" box="[714,763,728,752]" italics="true" pageId="2" pageNumber="101">et al</emphasis>
. (2007) that the two peculiarities of the second maxilla cited by Attems (1930a, b) as diagnostic for
<taxonomicName id="00D34D67FFD3FFBAFDD0FD42FD50FCE8" box="[588,691,765,788]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFDD0FD42FD50FCE8" box="[588,691,765,788]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
are of no value for the generic concept if it is meant to encompass Indian species. However, we maintain that the shared presence of a distomedial process on the femur of the ultimate leg in males is a strong argument that most of the Indian species of
<taxonomicName id="00D34D67FFD3FFBAFB94FCFAFB8CFCA0" box="[1032,1135,837,860]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFB94FCFAFB8CFCA0" box="[1032,1135,837,860]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
named by Jangi and Dass (1984) together with new species erected here are most closely related to tropical African
<taxonomicName id="00D34D67FFD3FFBAFB2AFCD6FAFEFC7C" box="[1206,1309,873,896]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFB2AFCD6FAFEFC7C" box="[1206,1309,873,896]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
. Though a similar process is present on the tibia of males in the Brazilian
<taxonomicName id="00D34D67FFD3FFBAFCE5FC32FF31FC3B" authority="Brolemann, 1902" authorityName="Brolemann" authorityYear="1902" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="tibialis" subGenus="Parotostigmus">
<emphasis id="F5A7EAF6FFD3FFBAFCE5FC32FC1AFC58" box="[889,1017,909,932]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
(
<emphasis id="F5A7EAF6FFD3FFBAFB90FC32FB4DFC58" box="[1036,1198,909,932]" italics="true" pageId="2" pageNumber="101">Parotostigmus</emphasis>
)
<emphasis id="F5A7EAF6FFD3FFBAFB5DFC33FAECFC58" box="[1217,1295,908,932]" italics="true" pageId="2" pageNumber="101">tibialis</emphasis>
Brölemann, 1902
</taxonomicName>
, a potentially homologous process on the femur is not known in any other member of Otostigmini. This peculiarity is, in the absence of any other competing hypothesis of relationship (i.e., better support for separate sister groups of African and Indian
<taxonomicName id="00D34D67FFD3FFBAFDD5FC46FD53FBEC" box="[585,688,1017,1040]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFDD5FC46FD53FBEC" box="[585,688,1017,1040]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
within
<taxonomicName id="00D34D67FFD3FFBAFC8DFC47FC72FBF3" box="[785,913,1016,1039]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFC8DFC47FC72FBF3" box="[785,913,1016,1039]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
), indicative of affinity. In Indian species, a longitudinal furrow on the distal half of the medial surface of the femur terminates on the process (
<figureCitation id="5FE82A61FFD3FFBAFB6BFBA3FAA2FBC8" box="[1271,1345,1052,1076]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Digitipes barnabasi Jangi and Dass, 1984. 7. Ultimate leg prefemur, ventral view, CES 091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES 091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES 091033, scale = 1 mm. 10. TT 18 – 20, dorsal view, CES 091033, scale = 1 mm. 11. TT 18 – 20, dorsal view, CES 091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES 091017, scale = 0.5 mm." httpUri="https://zenodo.org/record/218627/files/figure.png" pageId="2" pageNumber="101">Figs 9</figureCitation>
,
<figureCitation id="5FE82A61FFD3FFBAFAD2FBA3FA8FFBC8" box="[1358,1388,1052,1076]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1858,1881]" captionTargetBox="[151,1435,859,1803]" captionTargetId="figure@8.[151,1436,859,1804]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Digitipes coonoorensis Jangi and Dass, 1984. 19. Ultimate leg prefemur, ventral view, CES 08960 (female), scale = 0.5 mm. 20. Ultimate leg femur, dorsal view, CES 091088 (male), scale = 0.5 mm. 21. Ultimate leg femur, medial view, CES 091088, scale = 0.5 mm. 22. TT 18 – 20, dorsal view, CES 08960, scale = 1 mm. 23. Spiracle on segment 3, CES 07125, scale = 0.25 mm." httpUri="https://zenodo.org/record/218629/files/figure.png" pageId="2" pageNumber="101">21</figureCitation>
,
<figureCitation id="5FE82A61FFD3FFBAFAE5FBA3FF55FBAB" captionStart="FIGURES 36 – 45" captionStartId="12.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 36 – 45. Digitipes jonesii (Verhoeff, 1938). 36 – 40, CES 091062 (male); 41 – 45, CES 091090 (male). 36. Ultimate leg prefemur, ventral view, scale = 1 mm. 37. Ultimate leg femur, dorsal view, scale = 1 mm. 38. Ultimate leg femur, medial view, scale = 1 mm. 39. TT 18 – 19, dorsal view, scale = 1 mm. 40. Spiracle on segment 3, lateral view, scale = 0.5 mm. 41. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 42. Ultimate leg femur, medial view, scale = 0.5 mm. 43. Ultimate leg femur, ventral view, scale = 1 mm. 44. TT 18 – 20, dorsal view, scale = 1 mm. 45. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218632/files/figure.png" pageId="2" pageNumber="101">38, 42</figureCitation>
,
<figureCitation id="5FE82A61FFD3FFBAFF5DFB80FF03FBAB" box="[193,224,1087,1111]" captionStart="FIGURES 52 – 56" captionStartId="16.[151,264,1116,1139]" captionTargetBox="[165,1408,185,1098]" captionTargetId="figure@16.[165,1409,185,1099]" captionTargetPageId="16" captionText="FIGURES 52 – 56. Digitipes jangii n. sp. CES 08912 (holotype male) except Fig. 56, CES 08915 (paratype male). 52. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 53. Ultimate leg femur, dorsal view, scale = 0.5 mm. 54. Ultimate leg femur, medial view, scale = 0.5 mm. 55. TT 18 – 20, dorsal view, scale = 0.5 mm. 56. Spiracle on segment 3, scale = 0.25 mm." httpUri="https://zenodo.org/record/218634/files/figure.png" pageId="2" pageNumber="101">54</figureCitation>
), and a corresponding furrow was depicted in the African
<taxonomicName id="00D34D67FFD3FFBAFCEFFBFEFBE0FBA4" box="[883,1027,1088,1112]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="reichardti">
<emphasis id="F5A7EAF6FFD3FFBAFCEFFBFEFBE0FBA4" box="[883,1027,1088,1112]" italics="true" pageId="2" pageNumber="101">D. reichardti</emphasis>
</taxonomicName>
(“
<taxonomicName id="00D34D67FFD3FFBAFBBCFBFEFB24FBA4" box="[1056,1223,1088,1112]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="katangensis">
<emphasis id="F5A7EAF6FFD3FFBAFBBCFBFEFB24FBA4" box="[1056,1223,1088,1112]" italics="true" pageId="2" pageNumber="101">D. katangensis</emphasis>
</taxonomicName>
”) by Kraus (1957: figs 8, 9). The association of the furrow and process strengthens the basis for homology. In Indian species, the flattened medial surface of the femur bears a dense field of fine pores that we infer to be the openings of epidermal glands. The alternative to recognising the Indian species as
<taxonomicName id="00D34D67FFD3FFBAFCDDFB12FC4BFB38" box="[833,936,1197,1220]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFCDDFB12FC4BFB38" box="[833,936,1197,1220]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
would be to relegate them to
<taxonomicName id="00D34D67FFD3FFBAFA9CFB12FA9CFB38" box="[1280,1407,1197,1220]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFA9CFB12FA9CFB38" box="[1280,1407,1197,1220]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
, a genus that already includes some 120 species. We consider that this would create more rather than less taxonomic confusion because
<taxonomicName id="00D34D67FFD3FFBAFEF3FB4AFE0CFAF0" box="[367,495,1269,1292]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFEF3FB4AFE0CFAF0" box="[367,495,1269,1292]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
is demonstrably non-monophyletic (Vahtera
<emphasis id="F5A7EAF6FFD3FFBAFC6CFB4AFBC0FAF0" box="[1008,1059,1268,1292]" italics="true" pageId="2" pageNumber="101">et al</emphasis>
. 2012): molecular data resolve it as polyphyletic, and none of its putatively diagnostic morphological characters is apomorphic.
</paragraph>
<paragraph id="C76C36E4FFD3FFBAFF5BFA82FD40F988" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="101">
We are left with a situation in which
<taxonomicName id="00D34D67FFD3FFBAFDF5FA82FD33FAA8" box="[617,720,1341,1364]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFDF5FA82FD33FAA8" box="[617,720,1341,1364]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
is morphologically diagnosed by a character that is confined to one sex, an uncommon practice in centipede taxonomy but of course one that is commonplace in spiders and millipedes, where the male palps and gonopods, respectively, are often necessary for taxonomic diagnoses. Specimens that possess the femoral process make up a small percentage of our total sample of
<taxonomicName id="00D34D67FFD3FFBAFB50FA16FAD1FA3C" box="[1228,1330,1449,1472]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFB50FA16FAD1FA3C" box="[1228,1330,1449,1472]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
, they are consistently large, and we have not detected incipient stages of its development in small specimens. Accordingly we consider it likely that it is a structure that emerges with sexual maturity. Despite these limitations in applying this character to the diagnosis of the genus, access to molecular tools makes it possible to associate females to males from the same collection, and the monophyly of the Indian radiation of
<taxonomicName id="00D34D67FFD3FFBAFC66F986FB82F9AC" box="[1018,1121,1593,1616]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFC66F986FB82F9AC" box="[1018,1121,1593,1616]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
is robustly supported by the sequence data (Joshi and Karanth 2011, 2012).
</paragraph>
<paragraph id="C76C36E4FFD3FFBBFF5BF93FFADCFEC3" blockId="2.[151,1437,151,2012]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="102" pageId="2" pageNumber="101">
<collectingCountry id="BFC47674FFD3FFBAFF5BF93FFF05F964" box="[199,230,1664,1688]" name="American Samoa" pageId="2" pageNumber="101">As</collectingCountry>
noted under discussion of
<taxonomicName id="00D34D67FFD3FFBAFDBAF93FFCEEF964" box="[550,781,1664,1688]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="gravelyi">
<emphasis id="F5A7EAF6FFD3FFBAFDBAF93FFCEEF964" box="[550,781,1664,1688]" italics="true" pageId="2" pageNumber="101">Otostigmus gravelyi</emphasis>
</taxonomicName>
below, Indian
<taxonomicName id="00D34D67FFD3FFBAFC22F93EFBC6F964" box="[958,1061,1665,1688]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFC22F93EFBC6F964" box="[958,1061,1665,1688]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
to the exclusion of that species (originally assigned to
<taxonomicName id="00D34D67FFD3FFBAFE3FF91AFDE9F940" box="[419,522,1701,1724]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFE3FF91AFDE9F940" box="[419,522,1701,1724]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
but transferred herein to
<taxonomicName id="00D34D67FFD3FFBAFCAAF91AFC55F940" box="[822,950,1701,1724]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFCAAF91AFC55F940" box="[822,950,1701,1724]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
) share a suite of characters that is highly conserved and is shared by African congeners. Forcipular tooth plates with 4+4 teeth are shared by all specimens of Indian and African species apart from one specimen of
<taxonomicName id="00D34D67FFD3FFBAFC87F952FC36F8F8" box="[795,981,1773,1796]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD3FFBAFC87F952FC36F8F8" box="[795,981,1773,1796]" italics="true" pageId="2" pageNumber="101">D. coonoorensis</emphasis>
</taxonomicName>
with five teeth on one tooth plate. The arrangement of the teeth shows a consistent pattern of the inner pair being grouped, the second tooth being the largest and having an asymmetrical outline, its outer margin less steeply sloping than its inner margin, a seta being situated behind the outer edge of the second tooth, and the outer tooth being the smallest and distinctly posterior to the other three. Indian
<taxonomicName id="00D34D67FFD3FFBAFE05F8C2FDE3F868" box="[409,512,1917,1940]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFE05F8C2FDE3F868" box="[409,512,1917,1940]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
have a highly conserved number of 17 antennal articles, fixed in all species apart from a single specimen of
<taxonomicName id="00D34D67FFD3FFBAFE55F81EFD62F844" box="[457,641,1953,1976]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD3FFBAFE55F81EFE3EF844" box="[457,477,1953,1976]" italics="true" pageId="2" pageNumber="101">D</emphasis>
.
<emphasis id="F5A7EAF6FFD3FFBAFE71F81EFD62F844" box="[493,641,1953,1976]" italics="true" pageId="2" pageNumber="101">coonoorensis</emphasis>
</taxonomicName>
with 18 articles on one antenna and two specimens of
<taxonomicName id="00D34D67FFD3FFBAFB67F81EFA8AF844" box="[1275,1385,1953,1976]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFD3FFBAFB67F81EFA8AF844" box="[1275,1385,1953,1976]" italics="true" pageId="2" pageNumber="101">D. jonesii</emphasis>
</taxonomicName>
that have 18 or 18/19 articles. African
<taxonomicName id="00D34D67FFD3FFBAFDBCF87AFD64F820" box="[544,647,1989,2012]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFDBCF87AFD64F820" box="[544,647,1989,2012]" italics="true" pageId="2" pageNumber="101">Digitipes</emphasis>
</taxonomicName>
have 16–17 antennal articles. Unlike most
<taxonomicName id="00D34D67FFD3FFBAFBE4F87AFB14F820" box="[1144,1271,1989,2012]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD3FFBAFBE4F87AFB14F820" box="[1144,1271,1989,2012]" italics="true" pageId="2" pageNumber="101">Otostigmus</emphasis>
</taxonomicName>
, the transition from glabrous to densely setose antennal articles in
<taxonomicName id="00D34D67FFD2FFBBFD7CFF26FCA5FF4C" box="[736,838,153,176]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFD7CFF26FCA5FF4C" box="[736,838,153,176]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
apart from
<taxonomicName id="00D34D67FFD2FFBBFC50FF26FBBAFF4C" box="[972,1113,152,176]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="chhotanii">
<emphasis id="F5A7EAF6FFD2FFBBFC50FF26FBBAFF4C" box="[972,1113,152,176]" italics="true" pageId="3" pageNumber="102">D. chhotanii</emphasis>
</taxonomicName>
corresponds precisely to the boundaries of articles (i.e., two or three glabrous articles rather than 2.25, 2.33, 2.5, etc.). All Indian
<taxonomicName id="00D34D67FFD2FFBBFB66FF02FA82FF28" box="[1274,1377,189,212]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFB66FF02FA82FF28" box="[1274,1377,189,212]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
have two apical spines on the coxopleural process and lack dorsal spines. These character states are shared by all African
<taxonomicName id="00D34D67FFD2FFBBFF0BFEBAFF1DFEE0" box="[151,254,261,284]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFF0BFEBAFF1DFEE0" box="[151,254,261,284]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
as well. The transfer of
<taxonomicName id="00D34D67FFD2FFBBFD88FEBAFCE0FEE0" box="[532,771,260,284]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="graveleyi">
<emphasis id="F5A7EAF6FFD2FFBBFD88FEBAFCE0FEE0" box="[532,771,260,284]" italics="true" pageId="3" pageNumber="102">Otostigmus graveleyi</emphasis>
</taxonomicName>
allows these stable characters to be added to the diagnosis of
<taxonomicName id="00D34D67FFD2FFBBFF29FE96FEFFFEBC" box="[181,284,297,320]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFF29FE96FEFFFEBC" box="[181,284,297,320]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
, though it is recognised that all of them are observed in various species of
<taxonomicName id="00D34D67FFD2FFBBFBFEFE97FB01FEC3" box="[1122,1250,296,319]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFBFEFE97FB01FEC3" box="[1122,1250,296,319]" italics="true" pageId="3" pageNumber="102">Otostigmus</emphasis>
</taxonomicName>
as well.
</paragraph>
</subSubSection>
</treatment>
</document>

248
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<treatment id="4F7A87F2FFD2FFBEFF0BFC48FEF0F9D0" ID-DOI="http://doi.org/10.5281/zenodo.6154114" ID-GBIF-Taxon="119563405" ID-Zenodo-Dep="6154114" LSID="urn:lsid:plazi:treatment:4F7A87F2FFD2FFBEFF0BFC48FEF0F9D0" httpUri="http://treatment.plazi.org/id/4F7A87F2FFD2FFBEFF0BFC48FEF0F9D0" lastPageId="6" lastPageNumber="105" pageId="3" pageNumber="102">
<subSubSection id="8FC9656FFFD2FFBBFF0BFC48FD07FB84" pageId="3" pageNumber="102" type="nomenclature">
<paragraph id="C76C36E4FFD2FFBBFF0BFC48FD6CFBED" blockId="3.[151,655,1015,1074]" box="[151,655,1015,1041]" pageId="3" pageNumber="102">
<heading id="9C248188FFD2FFBBFF0BFC48FD6CFBED" bold="true" box="[151,655,1015,1041]" fontSize="11" level="1" pageId="3" pageNumber="102" reason="1">
<taxonomicName id="00D34D67FFD2FFBBFF0BFC48FD6CFBED" ID-CoL="363VN" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[151,655,1015,1041]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD2FFBBFF0BFC48FD6CFBED" bold="true" box="[151,655,1015,1041]" pageId="3" pageNumber="102">
<emphasis id="F5A7EAF6FFD2FFBBFF0BFC48FE60FBED" bold="true" box="[151,387,1015,1041]" italics="true" pageId="3" pageNumber="102">Digitipes barnabasi</emphasis>
Jangi and Dass, 1984
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C76C36E4FFD2FFBBFF0BFBA5FEFAFBCE" blockId="3.[151,655,1015,1074]" box="[151,281,1050,1074]" pageId="3" pageNumber="102">
(
<figureCitation id="5FE82A61FFD2FFBBFF03FBA5FEF3FBCE" box="[159,272,1050,1074]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 12" captionStartId-0="4.[151,264,1882,1905]" captionStartId-1="5.[151,264,1877,1900]" captionTargetBox-0="[151,1435,195,1857]" captionTargetBox-1="[151,1435,193,1854]" captionTargetId-0="figure@4.[151,1436,195,1858]" captionTargetId-1="figure@5.[151,1436,193,1856]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1 – 6. Digitipes barnabasi Jangi and Dass, 1984. CES 091355 (female) except Figs 1, 4, CES 091353 (female). 1. Cephalic plate and T 1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm." captionText-1="FIGURES 7 – 12. Digitipes barnabasi Jangi and Dass, 1984. 7. Ultimate leg prefemur, ventral view, CES 091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES 091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES 091033, scale = 1 mm. 10. TT 18 – 20, dorsal view, CES 091033, scale = 1 mm. 11. TT 18 – 20, dorsal view, CES 091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES 091017, scale = 0.5 mm." httpUri-0="https://zenodo.org/record/218626/files/figure.png" httpUri-1="https://zenodo.org/record/218627/files/figure.png" pageId="3" pageNumber="102">Figs 1–12</figureCitation>
)
</paragraph>
<paragraph id="C76C36E4FFD2FFBBFF0BFBDCFD07FB84" blockId="3.[151,840,1122,1237]" box="[151,740,1122,1144]" pageId="3" pageNumber="102">
<taxonomicName id="00D34D67FFD2FFBBFF0BFBDCFD84FB84" authority="Jangi and Dass, 1984: 44" authorityName="Jangi and Dass" authorityPageNumber="44" authorityYear="1984" box="[151,615,1122,1144]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD2FFBBFF0BFBDCFE82FB84" box="[151,353,1122,1144]" italics="true" pageId="3" pageNumber="102">Digitipes barnabasi</emphasis>
Jangi and Dass, 1984: 44
</taxonomicName>
, figs 73–79.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD2FFBBFF0BFB3DFCD3FB28" pageId="3" pageNumber="102" type="description">
<paragraph id="C76C36E4FFD2FFBBFF0BFB3DFCD3FB28" blockId="3.[151,840,1122,1237]" pageId="3" pageNumber="102">
<taxonomicName id="00D34D67FFD2FFBBFF0BFB3DFE82FB6B" box="[151,353,1153,1175]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD2FFBBFF0BFB3DFE82FB6B" box="[151,353,1153,1175]" italics="true" pageId="3" pageNumber="102">Digitipes barnabasi</emphasis>
</taxonomicName>
. Yadav 1993: 171; Sureshan
<emphasis id="F5A7EAF6FFD2FFBBFD0CFB3DFD5FFB6B" box="[656,700,1153,1175]" italics="true" pageId="3" pageNumber="102">et al</emphasis>
. 2006: 2287.
<taxonomicName id="00D34D67FFD2FFBBFF0BFB1EFF17FB4A" box="[151,244,1185,1206]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFF0BFB1EFF17FB4A" box="[151,244,1185,1206]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
putative species 7. Joshi and Karanth, 2012: figs 2, 3, 7.
<taxonomicName id="00D34D67FFD2FFBBFF0BFB7FFF17FB29" box="[151,244,1216,1237]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFF0BFB7FFF17FB29" box="[151,244,1216,1237]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
putative species 8. Joshi and Karanth, 2012: figs 2, 3, 7.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD2FFBBFF0BFABEFBCBFA55" pageId="3" pageNumber="102" type="diagnosis">
<paragraph id="C76C36E4FFD2FFBBFF0BFABEFBCBFA55" blockId="3.[151,1437,1281,1989]" pageId="3" pageNumber="102">
<emphasis id="F5A7EAF6FFD2FFBBFF0BFABEFEF2FAE6" bold="true" box="[151,273,1281,1306]" pageId="3" pageNumber="102">Diagnosis.</emphasis>
<taxonomicName id="00D34D67FFD2FFBBFEBEFABDFE6AFAE5" box="[290,393,1282,1305]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFEBEFABDFE6AFAE5" box="[290,393,1282,1305]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
with basal two antennal articles glabrous dorsally and ventrally. Flat median ridge consistently present from T3 to 5–20, irregular rugose ridges at least weakly distinct on lateral parts of tergites in posterior half of trunk; five continuous, tuberculate longitudinal keels from TT6–9 variably present. Coxopleural process relatively short, lateral spine consistently present; pore field extending close to dorsal margin of coxopleuron. Dorsomedial process on ultimate leg femur of male relatively short.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD2FFBBFF5BFA0AFB45F9A1" pageId="3" pageNumber="102" type="materials_examined">
<paragraph id="C76C36E4FFD2FFBBFF5BFA0AFB45F9A1" blockId="3.[151,1437,1281,1989]" pageId="3" pageNumber="102">
<emphasis id="F5A7EAF6FFD2FFBBFF5BFA0AFE52FA32" bold="true" box="[199,433,1461,1486]" pageId="3" pageNumber="102">Material examined.</emphasis>
<specimenCount id="D1D5FD6DFFD2FFBBFE24FA0AFDB2FA31" box="[440,593,1461,1485]" pageId="3" pageNumber="102" type="generic">12 specimens</specimenCount>
of
<taxonomicName id="00D34D67FFD2FFBBFDEBFA09FD3DFA31" box="[631,734,1462,1485]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFDEBFA09FD3DFA31" box="[631,734,1462,1485]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
putative species 8 (
<taxonomicName id="00D34D67FFD2FFBBFC22FA09FBB0FA31" box="[958,1107,1461,1485]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD2FFBBFC22FA09FBB0FA31" box="[958,1107,1461,1485]" italics="true" pageId="3" pageNumber="102">D. barnabasi</emphasis>
</taxonomicName>
of Joshi and Karanth, 2012), including CES
<date id="B36D1024FFD2FFBBFEA7FA66FE71FA0D" box="[315,402,1497,1521]" pageId="3" pageNumber="102" value="48-09-13">091348</date>
, CES
<date id="B36D1024FFD2FFBBFE4CFA66FDC6FA0D" box="[464,549,1497,1521]" pageId="3" pageNumber="102" value="53-09-13">091353</date>
and CES
<date id="B36D1024FFD2FFBBFD12FA66FD00FA0D" box="[654,739,1497,1521]" pageId="3" pageNumber="102" value="55-09-13">091355</date>
from near
<typeStatus id="18688846FFD2FFBBFCC3FA65FC6CFA0E" box="[863,911,1498,1522]" pageId="3" pageNumber="102">type</typeStatus>
locality, and four of
<taxonomicName id="00D34D67FFD2FFBBFBE0FA65FB00FA0D" box="[1148,1251,1498,1521]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD2FFBBFBE0FA65FB00FA0D" box="[1148,1251,1498,1521]" italics="true" pageId="3" pageNumber="102">Digitipes</emphasis>
</taxonomicName>
putative species 7; CES registration numbers in Joshi and Karanth (2012: fig. 2, Table S1). The examined individuals were collected across the evergreen forests of the Western Ghats encompassing 8°–19° N latitude and
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elevation (distribution maps in Joshi and Karanth 2012: fig. 7c); all leg. J. Joshi,
<date id="B36D1024FFD2FFBBFBBCF9FAFB41F9A1" box="[1056,1186,1605,1629]" pageId="3" pageNumber="102" value="2007" valueMax="2010">2007–2010</date>
.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD2FFBEFF55F9D6FDA4FC58" lastPageId="6" lastPageNumber="105" pageId="3" pageNumber="102" type="description">
<paragraph id="C76C36E4FFD2FFBBFF55F9D6FD3BF911" blockId="3.[151,1437,1281,1989]" pageId="3" pageNumber="102">
<emphasis id="F5A7EAF6FFD2FFBBFF55F9D6FEB9F97E" bold="true" box="[201,346,1641,1666]" pageId="3" pageNumber="102">Description.</emphasis>
Length to
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. 17 antennal articles; basal two (three in one individual) articles glabrous both dorsally (
<figureCitation id="5FE82A61FFD2FFBBFE9CF932FEA3F959" box="[256,320,1677,1701]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1882,1905]" captionTargetBox="[151,1435,195,1857]" captionTargetId="figure@4.[151,1436,195,1858]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Digitipes barnabasi Jangi and Dass, 1984. CES 091355 (female) except Figs 1, 4, CES 091353 (female). 1. Cephalic plate and T 1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm." httpUri="https://zenodo.org/record/218626/files/figure.png" pageId="3" pageNumber="102">Fig. 1</figureCitation>
) and ventrally. Cephalic plate and tergites punctate. Longitudinal median furrow on anterior 20% of cephalic plate, occasionally slightly longer; pair of short, medially converging furrows on posterior part of cephalic plate at least faint, in some specimens well defined.
</paragraph>
<paragraph id="C76C36E4FFD2FFBBFF55F946FBB2F8C9" blockId="3.[151,1437,1281,1989]" pageId="3" pageNumber="102">Cephalic plate and T1 red-brown, the following tergites more deeply chestnut brown, variably with some blue pigmentation near the lateral margin; legs pale brown, often with a light purple tinge.</paragraph>
<paragraph id="C76C36E4FFD2FFBBFF55F8FEFBD7F839" blockId="3.[151,1437,1281,1989]" pageId="3" pageNumber="102">
Forcipular coxosternal tooth plates slightly longer than wide (
<figureCitation id="5FE82A61FFD2FFBBFC00F8FEFC06F8A5" box="[924,997,1857,1881]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1882,1905]" captionTargetBox="[151,1435,195,1857]" captionTargetId="figure@4.[151,1436,195,1858]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Digitipes barnabasi Jangi and Dass, 1984. CES 091355 (female) except Figs 1, 4, CES 091353 (female). 1. Cephalic plate and T 1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm." httpUri="https://zenodo.org/record/218626/files/figure.png" pageId="3" pageNumber="102">Fig. 2</figureCitation>
), with four main teeth, the inner two grouped together, sometimes partially fused, outer two acute, the outermost smaller than the other three (
<figureCitation id="5FE82A61FFD2FFBBFADBF8DAFA6FF881" box="[1351,1420,1893,1917]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1882,1905]" captionTargetBox="[151,1435,195,1857]" captionTargetId="figure@4.[151,1436,195,1858]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Digitipes barnabasi Jangi and Dass, 1984. CES 091355 (female) except Figs 1, 4, CES 091353 (female). 1. Cephalic plate and T 1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm." httpUri="https://zenodo.org/record/218626/files/figure.png" pageId="3" pageNumber="102">Fig. 3</figureCitation>
); single strong seta posterior to lateral edge of second tooth; base of tooth plates defined by oblique sutures diverging at 130–140°. Trochanteroprefemoral process with two inner teeth and apical tooth.
</paragraph>
<caption id="93AC666CFFD5FFBCFF0BF8E5FB51F830" httpUri="https://zenodo.org/record/218626/files/figure.png" pageId="4" pageNumber="103" targetBox="[151,1435,195,1857]" targetPageId="4">
<paragraph id="C76C36E4FFD5FFBCFF0BF8E5FB51F830" blockId="4.[151,1437,1882,1996]" pageId="4" pageNumber="103">
<emphasis id="F5A7EAF6FFD5FFBCFF0BF8E5FEFFF88C" bold="true" box="[151,284,1882,1905]" pageId="4" pageNumber="103">FIGURES 1</emphasis>
–
<emphasis id="F5A7EAF6FFD5FFBCFEB4F8E4FED6F88C" bold="true" box="[296,309,1883,1904]" pageId="4" pageNumber="103">6</emphasis>
.
<taxonomicName id="00D34D67FFD5FFBCFED7F8E4FD1EF88C" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[331,765,1882,1904]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD5FFBCFED7F8E4FDF4F88C" box="[331,535,1882,1904]" italics="true" pageId="4" pageNumber="103">Digitipes barnabasi</emphasis>
Jangi and Dass, 1984
</taxonomicName>
. CES091355 (female) except Figs 1, 4, CES091353 (female). 1. Cephalic plate and T1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm.
</paragraph>
</caption>
<caption id="93AC666CFFD4FFBDFF0BF8EAFC6EF83B" httpUri="https://zenodo.org/record/218627/files/figure.png" pageId="5" pageNumber="104" targetBox="[151,1435,193,1854]" targetPageId="5">
<paragraph id="C76C36E4FFD4FFBDFF0BF8EAFC6EF83B" blockId="5.[151,1436,1877,1991]" pageId="5" pageNumber="104">
<emphasis id="F5A7EAF6FFD4FFBDFF0BF8EAFEFFF897" bold="true" box="[151,284,1877,1900]" pageId="5" pageNumber="104">FIGURES 7</emphasis>
–
<emphasis id="F5A7EAF6FFD4FFBDFEB6F8EAFEA0F896" bold="true" box="[298,323,1877,1898]" pageId="5" pageNumber="104">12</emphasis>
.
<taxonomicName id="00D34D67FFD4FFBDFEC7F8E9FCF1F897" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[347,786,1877,1899]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD4FFBDFEC7F8E9FDCBF897" box="[347,552,1877,1899]" italics="true" pageId="5" pageNumber="104">Digitipes barnabasi</emphasis>
Jangi and Dass, 1984
</taxonomicName>
. 7. Ultimate leg prefemur, ventral view, CES091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES091033, scale = 1 mm. 10. TT18–20, dorsal view, CES091033, scale = 1 mm. 11. TT18–20, dorsal view, CES091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES091017, scale = 0.5 mm.
</paragraph>
</caption>
<paragraph id="C76C36E4FFD7FFBEFF55FF28FA9DFF53" blockId="6.[151,1437,151,1580]" box="[201,1406,151,176]" pageId="6" pageNumber="105">Second maxillary claw with slender accessory spurs. Article 2 of telopodite bearing a slender spine distally.</paragraph>
<paragraph id="C76C36E4FFD7FFBEFF55FF03FC69FE7B" blockId="6.[151,1437,151,1580]" pageId="6" pageNumber="105">
Tergites with paramedian sutures complete from TT5 to 7. Tergites fully marginate starting from TT5 to 8, incipient margins as far anteriorly as T2. Tergites mostly smooth apart from longitudinal median ridge beginning from TT3 to 5 (
<figureCitation id="5FE82A61FFD7FFBEFECDFEBBFE40FEE0" box="[337,419,260,284]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Digitipes barnabasi Jangi and Dass, 1984. 7. Ultimate leg prefemur, ventral view, CES 091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES 091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES 091033, scale = 1 mm. 10. TT 18 – 20, dorsal view, CES 091033, scale = 1 mm. 11. TT 18 – 20, dorsal view, CES 091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES 091017, scale = 0.5 mm." httpUri="https://zenodo.org/record/218627/files/figure.png" pageId="6" pageNumber="105">Fig. 11</figureCitation>
) and at least faint (variably strong), irregular rugose ridges on lateral part of tergites in posterior half of trunk; a few individuals with strong ridges and keels/tubercles (
<figureCitation id="5FE82A61FFD7FFBEFBD4FE98FB7CFEC3" box="[1096,1183,295,319]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Digitipes barnabasi Jangi and Dass, 1984. 7. Ultimate leg prefemur, ventral view, CES 091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES 091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES 091033, scale = 1 mm. 10. TT 18 – 20, dorsal view, CES 091033, scale = 1 mm. 11. TT 18 – 20, dorsal view, CES 091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES 091017, scale = 0.5 mm." httpUri="https://zenodo.org/record/218627/files/figure.png" pageId="6" pageNumber="105">Fig. 10</figureCitation>
); three keels between paramedian sutures, one immediately lateral to them, those more laterally poorly-defined, anastomosing. Paramedian sutures 10–20% length of sternites in mid body region.
</paragraph>
<paragraph id="C76C36E4FFD7FFBEFF55FE2BFDEBFE33" blockId="6.[151,1437,151,1580]" pageId="6" pageNumber="105">First four to at least 14 (usually 4–7) pairs of legs with two tarsal spurs, the subsequent to 20 with one. A tibial and a femoral spur on leg 1 only.</paragraph>
<paragraph id="C76C36E4FFD7FFBEFF55FE63FC15FDA3" blockId="6.[151,1437,151,1580]" pageId="6" pageNumber="105">
Tergite of ultimate leg-bearing segment with faintly convex, nearly parallel lateral margins; posterior margin variably sinuous, rounded posteromedially (
<figureCitation id="5FE82A61FFD7FFBEFD0CFE40FD3BFDEB" box="[656,728,511,535]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1882,1905]" captionTargetBox="[151,1435,195,1857]" captionTargetId="figure@4.[151,1436,195,1858]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Digitipes barnabasi Jangi and Dass, 1984. CES 091355 (female) except Figs 1, 4, CES 091353 (female). 1. Cephalic plate and T 1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm." httpUri="https://zenodo.org/record/218626/files/figure.png" pageId="6" pageNumber="105">Fig. 5</figureCitation>
). Sternite of ultimate leg-bearing segment relatively elongate trapezoidal, with sides converging posteriorly, nearly straight, posterior margin gently to moderately concave (
<figureCitation id="5FE82A61FFD7FFBEFAF2FD9BFF44FDA3" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1882,1905]" captionTargetBox="[151,1435,195,1857]" captionTargetId="figure@4.[151,1436,195,1858]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Digitipes barnabasi Jangi and Dass, 1984. CES 091355 (female) except Figs 1, 4, CES 091353 (female). 1. Cephalic plate and T 1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm." httpUri="https://zenodo.org/record/218626/files/figure.png" pageId="6" pageNumber="105">Fig. 6</figureCitation>
); weak trace of an incomplete longitudinal median furrow variably present.
</paragraph>
<paragraph id="C76C36E4FFD7FFBEFF55FDD3FDA4FC58" blockId="6.[151,1437,151,1580]" pageId="6" pageNumber="105">
Coxopleural process relatively short, stoutly conical in ventral view with marked inflection from posterolateral margin of coxopleuron, with two apical spines, one lateral spine (
<figureCitation id="5FE82A61FFD7FFBEFC04FD30FC02FD5B" box="[920,993,655,679]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1882,1905]" captionTargetBox="[151,1435,195,1857]" captionTargetId="figure@4.[151,1436,195,1858]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Digitipes barnabasi Jangi and Dass, 1984. CES 091355 (female) except Figs 1, 4, CES 091353 (female). 1. Cephalic plate and T 1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm." httpUri="https://zenodo.org/record/218626/files/figure.png" pageId="6" pageNumber="105">Fig. 6</figureCitation>
). Pores dense, variable in size, nearly reaching dorsal margin of coxopleuron (
<figureCitation id="5FE82A61FFD7FFBEFDF9FD0BFD53FD30" box="[613,688,692,716]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1882,1905]" captionTargetBox="[151,1435,195,1857]" captionTargetId="figure@4.[151,1436,195,1858]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Digitipes barnabasi Jangi and Dass, 1984. CES 091355 (female) except Figs 1, 4, CES 091353 (female). 1. Cephalic plate and T 1, dorsal view, scale = 0.5 mm. 2. Forcipular segment, ventral view, scale = 1 mm. 3. Tooth plates, ventral view, scale = 0.5 mm. 4. Coxopleuron, lateral view, scale = 0.5 mm. 5. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 6. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 2 mm." httpUri="https://zenodo.org/record/218626/files/figure.png" pageId="6" pageNumber="105">Figs 4</figureCitation>
,
<figureCitation id="5FE82A61FFD7FFBEFD23FD0BFD3DFD30" box="[703,734,692,716]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Digitipes barnabasi Jangi and Dass, 1984. 7. Ultimate leg prefemur, ventral view, CES 091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES 091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES 091033, scale = 1 mm. 10. TT 18 – 20, dorsal view, CES 091033, scale = 1 mm. 11. TT 18 – 20, dorsal view, CES 091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES 091017, scale = 0.5 mm." httpUri="https://zenodo.org/record/218627/files/figure.png" pageId="6" pageNumber="105">12</figureCitation>
); non-porose area on process narrow, not extending as far as posterior margin of sternite of ultimate leg-bearing segment, variably less than half length to that margin. Ultimate leg prefemur with width at distal end about 25% its length; prefemoral spines at most moderately large: VL 3 (
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one individual), VM 2, DM 0 or 1 (
<figureCitation id="5FE82A61FFD7FFBEFDB6FCA0FD8CFCCB" box="[554,623,799,823]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Digitipes barnabasi Jangi and Dass, 1984. 7. Ultimate leg prefemur, ventral view, CES 091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES 091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES 091033, scale = 1 mm. 10. TT 18 – 20, dorsal view, CES 091033, scale = 1 mm. 11. TT 18 – 20, dorsal view, CES 091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES 091017, scale = 0.5 mm." httpUri="https://zenodo.org/record/218627/files/figure.png" pageId="6" pageNumber="105">Fig. 7</figureCitation>
). Distomedial process in males bluntly conical, extending only about as far as distal end of femur on its dorsal side (
<figureCitation id="5FE82A61FFD7FFBEFD08FCFBFD3AFCA0" box="[660,729,836,860]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Digitipes barnabasi Jangi and Dass, 1984. 7. Ultimate leg prefemur, ventral view, CES 091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES 091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES 091033, scale = 1 mm. 10. TT 18 – 20, dorsal view, CES 091033, scale = 1 mm. 11. TT 18 – 20, dorsal view, CES 091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES 091017, scale = 0.5 mm." httpUri="https://zenodo.org/record/218627/files/figure.png" pageId="6" pageNumber="105">Fig. 8</figureCitation>
); shallow groove along most of length of femur on its medial surface (
<figureCitation id="5FE82A61FFD7FFBEFF64FCD8FEDDFC83" box="[248,318,871,895]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@5.[151,1436,193,1856]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Digitipes barnabasi Jangi and Dass, 1984. 7. Ultimate leg prefemur, ventral view, CES 091355 (female), scale = 1 mm, 8. Ultimate leg femur, dorsal view, CES 091033 (male), scale = 1 mm. 9. Ultimate leg femur, medial view, CES 091033, scale = 1 mm. 10. TT 18 – 20, dorsal view, CES 091033, scale = 1 mm. 11. TT 18 – 20, dorsal view, CES 091355, scale = 1 mm. 12. Coxopleuron, lateral view, CES 091017, scale = 0.5 mm." httpUri="https://zenodo.org/record/218627/files/figure.png" pageId="6" pageNumber="105">Fig. 9</figureCitation>
). Ultimate leg tarsus 1 1.8–2.2 times length of tarsus 2; tarsus 1 3–4.8 times longer than pretarsus; pretarsus with pair of accessory claws.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD7FFBEFF55FC10FEF0F9D0" pageId="6" pageNumber="105" type="discussion">
<paragraph id="C76C36E4FFD7FFBEFF55FC10FC09FA84" blockId="6.[151,1437,151,1580]" pageId="6" pageNumber="105">
<emphasis id="F5A7EAF6FFD7FFBEFF55FC10FEADFC34" bold="true" box="[201,334,943,968]" pageId="6" pageNumber="105">Discussion.</emphasis>
PS7 of Joshi and Karanth (2012) was represented by four individuals (three of them immature) that shared all diagnostic characters with
<taxonomicName id="00D34D67FFD7FFBEFDF8FC6AFD1BFC10" box="[612,760,980,1004]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD7FFBEFDF8FC6AFD1BFC10" box="[612,760,980,1004]" italics="true" pageId="6" pageNumber="105">D. barnabasi</emphasis>
</taxonomicName>
, including 17 antennal articles of which two are glabrous on both the dorsal and ventral sides, the coxopleural process with a similar shape and the pore field approximating the dorsal margin of the coxopleuron, leg 20 bearing a tarsal spur, a longitudinal median ridge on the tergites, and irregular ridges on the lateral part of the tergites. In the molecular phylogeny based on mtDNA data, the monophyly of PS7 was not supported. Both PS7 and PS8 have a higher frequency of specimens with two tarsal spurs on more than just the anterior three or four leg pairs than in other Indian
<taxonomicName id="00D34D67FFD7FFBEFB98FB36FB88FB5C" box="[1028,1131,1161,1184]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD7FFBEFB98FB36FB88FB5C" box="[1028,1131,1161,1184]" italics="true" pageId="6" pageNumber="105">Digitipes</emphasis>
</taxonomicName>
, two spurs being present at least as far as leg
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both putative species. Because we have not identified any consistent diagnostic morphological characters to distinguish PS7 from PS8, together with the small sample of the former, we have referred both to a single species,
<taxonomicName id="00D34D67FFD7FFBEFDB3FB4AFD2AFAF0" box="[559,713,1268,1292]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD7FFBEFDB3FB4AFD2AFAF0" box="[559,713,1268,1292]" italics="true" pageId="6" pageNumber="105">D. barnabasi</emphasis>
</taxonomicName>
. Geographically these two putative units show very distinct distributions, with PS7 occurring in the southern and central parts of the Western Ghats and PS8 being restricted to the northern parts of the Western Ghats. Henceforth,
<taxonomicName id="00D34D67FFD7FFBEFC9CFA82FC74FAA8" box="[768,919,1340,1364]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD7FFBEFC9CFA82FC74FAA8" box="[768,919,1340,1364]" italics="true" pageId="6" pageNumber="105">D. barnabasi</emphasis>
</taxonomicName>
becomes one of the most widely distributed
<taxonomicName id="00D34D67FFD7FFBEFF0BFADEFF1DFA84" box="[151,254,1377,1400]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD7FFBEFF0BFADEFF1DFA84" box="[151,254,1377,1400]" italics="true" pageId="6" pageNumber="105">Digitipes</emphasis>
</taxonomicName>
species of the Western Ghats showing strong population structure.
</paragraph>
<paragraph id="C76C36E4FFD7FFBEFF55FA3AFEF0F9D0" blockId="6.[151,1437,151,1580]" pageId="6" pageNumber="105">
In the synonymy above we include a specimen from the
<collectingCountry id="BFC47674FFD7FFBEFCC6FA3AFC4DFA60" box="[858,942,1413,1436]" name="India" pageId="6" pageNumber="105">Kannur</collectingCountry>
district, Kerala, assigned to the species by Surehan
<emphasis id="F5A7EAF6FFD7FFBEFF67FA16FECDFA3C" box="[251,302,1448,1472]" italics="true" pageId="6" pageNumber="105">et al</emphasis>
. (2006); a photograph of the specimen provided by D. Balan shows characters corresponding to
<taxonomicName id="00D34D67FFD7FFBEFA1DFA16FEEAFA18" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD7FFBEFA1DFA16FEEAFA18" italics="true" pageId="6" pageNumber="105">D. barnabasi</emphasis>
</taxonomicName>
as understood here. A record from the Pune district, Maharashtra, by Yadav (1993a) is within the geographic distribution of
<taxonomicName id="00D34D67FFD7FFBEFE5AFA4EFDB8F9F4" box="[454,603,1520,1544]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD7FFBEFE5AFA4EFDB8F9F4" box="[454,603,1520,1544]" italics="true" pageId="6" pageNumber="105">D. barnabasi</emphasis>
</taxonomicName>
and is accordingly listed above, but we have not been able to access the specimens.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="4F7A87F2FFD7FFB1FF0BF9C0FDB2FB14" ID-DOI="http://doi.org/10.5281/zenodo.6154116" ID-GBIF-Taxon="119563400" ID-Zenodo-Dep="6154116" LSID="urn:lsid:plazi:treatment:4F7A87F2FFD7FFB1FF0BF9C0FDB2FB14" httpUri="http://treatment.plazi.org/id/4F7A87F2FFD7FFB1FF0BF9C0FDB2FB14" lastPageId="9" lastPageNumber="108" pageId="6" pageNumber="105">
<subSubSection id="8FC9656FFFD7FFBEFF0BF9C0FD41F8E3" pageId="6" pageNumber="105" type="nomenclature">
<paragraph id="C76C36E4FFD7FFBEFF0BF9C0FD54F965" blockId="6.[151,695,1663,1722]" box="[151,695,1663,1689]" pageId="6" pageNumber="105">
<heading id="9C248188FFD7FFBEFF0BF9C0FD54F965" bold="true" box="[151,695,1663,1689]" fontSize="11" level="1" pageId="6" pageNumber="105" reason="1">
<taxonomicName id="00D34D67FFD7FFBEFF0BF9C0FD54F965" ID-CoL="363VQ" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[151,695,1663,1689]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD7FFBEFF0BF9C0FD54F965" bold="true" box="[151,695,1663,1689]" pageId="6" pageNumber="105">
<emphasis id="F5A7EAF6FFD7FFBEFF0BF9C0FE49F965" bold="true" box="[151,426,1663,1689]" italics="true" pageId="6" pageNumber="105">Digitipes coonoorensis</emphasis>
Jangi and Dass, 1984
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C76C36E4FFD7FFBEFF0BF91DFEC4F946" blockId="6.[151,695,1663,1722]" box="[151,295,1698,1722]" pageId="6" pageNumber="105">
(
<figureCitation id="5FE82A61FFD7FFBEFF03F91DFEFCF946" box="[159,287,1698,1722]" captionStart-0="FIGURES 13 – 18" captionStart-1="FIGURES 19 – 23" captionStartId-0="7.[151,264,1726,1749]" captionStartId-1="8.[151,264,1858,1881]" captionTargetBox-0="[151,1435,193,1703]" captionTargetBox-1="[151,1435,859,1803]" captionTargetId-0="figure@7.[151,1436,193,1705]" captionTargetId-1="figure@8.[151,1436,859,1804]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." captionText-1="FIGURES 19 – 23. Digitipes coonoorensis Jangi and Dass, 1984. 19. Ultimate leg prefemur, ventral view, CES 08960 (female), scale = 0.5 mm. 20. Ultimate leg femur, dorsal view, CES 091088 (male), scale = 0.5 mm. 21. Ultimate leg femur, medial view, CES 091088, scale = 0.5 mm. 22. TT 18 – 20, dorsal view, CES 08960, scale = 1 mm. 23. Spiracle on segment 3, CES 07125, scale = 0.25 mm." httpUri-0="https://zenodo.org/record/218628/files/figure.png" httpUri-1="https://zenodo.org/record/218629/files/figure.png" pageId="6" pageNumber="105">Figs 13–23</figureCitation>
)
</paragraph>
<paragraph id="C76C36E4FFD7FFBEFF0BF954FD41F8E3" blockId="6.[151,816,1770,1854]" pageId="6" pageNumber="105">
<taxonomicName id="00D34D67FFD7FFBEFF0BF954FD64F8FC" authority="Jangi and Dass, 1984: 43" authorityName="Jangi and Dass" authorityPageNumber="43" authorityYear="1984" box="[151,647,1770,1792]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD7FFBEFF0BF954FE63F8FC" box="[151,384,1771,1792]" italics="true" pageId="6" pageNumber="105">Digitipes coonoorensis</emphasis>
Jangi and Dass, 1984: 43
</taxonomicName>
, figs 62–72.
<taxonomicName id="00D34D67FFD7FFBEFF0BF8B5FE63F8E3" box="[151,384,1802,1823]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD7FFBEFF0BF8B5FE63F8E3" box="[151,384,1802,1823]" italics="true" pageId="6" pageNumber="105">Digitipes coonoorensis</emphasis>
</taxonomicName>
. Sureshan
<emphasis id="F5A7EAF6FFD7FFBEFE6CF8B5FDF8F8E3" box="[496,539,1801,1823]" italics="true" pageId="6" pageNumber="105">et al</emphasis>
. 2006: 2287.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD7FFBEFF0BF896FCD3F8C2" box="[151,816,1832,1854]" pageId="6" pageNumber="105" type="description">
<paragraph id="C76C36E4FFD7FFBEFF0BF896FCD3F8C2" blockId="6.[151,816,1770,1854]" box="[151,816,1832,1854]" pageId="6" pageNumber="105">
<taxonomicName id="00D34D67FFD7FFBEFF0BF896FF17F8C2" box="[151,244,1833,1854]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD7FFBEFF0BF896FF17F8C2" box="[151,244,1833,1854]" italics="true" pageId="6" pageNumber="105">Digitipes</emphasis>
</taxonomicName>
putative species 1. Joshi and Karanth, 2012: figs 2, 3, 5.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD7FFBEFF0BF8D5FBB9F812" pageId="6" pageNumber="105" type="diagnosis">
<paragraph id="C76C36E4FFD7FFBEFF0BF8D5FBB9F812" blockId="6.[151,1437,1898,2030]" pageId="6" pageNumber="105">
<emphasis id="F5A7EAF6FFD7FFBEFF0BF8D5FEF2F87F" bold="true" box="[151,273,1898,1923]" pageId="6" pageNumber="105">Diagnosis.</emphasis>
<taxonomicName id="00D34D67FFD7FFBEFE8BF8D3FE9DF87F" box="[279,382,1900,1923]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD7FFBEFE8BF8D3FE9DF87F" box="[279,382,1900,1923]" italics="true" pageId="6" pageNumber="105">Digitipes</emphasis>
</taxonomicName>
with basal three antennal articles glabrous dorsally. Tergite paramedian sutures complete from TT6 to 8; tergites fully marginate starting from 7 to 9; tergites smooth, longitudinal median ridge rarely present. Coxopleural process relatively long, stout, variably inflected outward in ventral view, lateral spine present; pore field terminating far beneath dorsal margin of coxopleuron; leg 20 without tarsal spur.
</paragraph>
</subSubSection>
<caption id="93AC666CFFD6FFBFFF0BF901FEDAF8B3" httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="7" pageNumber="106" targetBox="[151,1435,193,1703]" targetPageId="7">
<paragraph id="C76C36E4FFD6FFBFFF0BF901FEDAF8B3" blockId="7.[151,1437,1726,1871]" pageId="7" pageNumber="106">
<emphasis id="F5A7EAF6FFD6FFBFFF0BF901FEC4F92F" bold="true" box="[151,295,1726,1749]" pageId="7" pageNumber="106">FIGURES 13</emphasis>
–
<emphasis id="F5A7EAF6FFD6FFBFFEA8F901FEB0F92F" bold="true" box="[308,339,1726,1747]" pageId="7" pageNumber="106">18.</emphasis>
<taxonomicName id="00D34D67FFD6FFBFFEC7F900FCC4F928" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[347,807,1726,1748]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="106" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD6FFBFFEC7F900FDA7F928" box="[347,580,1727,1748]" italics="true" pageId="7" pageNumber="106">Digitipes coonoorensis</emphasis>
Jangi and Dass, 1984
</taxonomicName>
. CES08960 (female) except Fig. 15, CES07125 (female) and Fig. 16, CES091088 (male). 13. Cephalic plate and T1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="8FC9656FFFD6FFB0FF55F8C4FD94FF28" lastPageId="8" lastPageNumber="107" pageId="7" pageNumber="106" type="materials_examined">
<paragraph id="C76C36E4FFD6FFB0FF55F8C4FD94FF28" blockId="7.[151,1436,1915,2011]" lastBlockId="8.[151,1437,151,788]" lastPageId="8" lastPageNumber="107" pageId="7" pageNumber="106">
<emphasis id="F5A7EAF6FFD6FFBFFF55F8C4FE55F868" bold="true" box="[201,438,1915,1940]" pageId="7" pageNumber="106">Material examined.</emphasis>
<specimenCount id="D1D5FD6DFFD6FFBFFE5DF8C4FDB9F86F" box="[449,602,1915,1939]" pageId="7" pageNumber="106" type="generic">11 specimens</specimenCount>
of
<taxonomicName id="00D34D67FFD6FFBFFD14F8C2FD0DF868" box="[648,750,1917,1940]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD6FFBFFD14F8C2FD0DF868" box="[648,750,1917,1940]" italics="true" pageId="7" pageNumber="106">Digitipes</emphasis>
</taxonomicName>
putative species 1 (
<taxonomicName id="00D34D67FFD6FFBFFC40F8C2FB75F868" box="[988,1174,1917,1940]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="106" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD6FFBFFC40F8C2FB75F868" box="[988,1174,1917,1940]" italics="true" pageId="7" pageNumber="106">D. coonoorensis</emphasis>
</taxonomicName>
of Joshi and Karanth, 2012), including CES07125, CES07132 and CES07134 from close to
<typeStatus id="18688846FFD6FFBFFC26F81FFC09F844" box="[954,1002,1952,1976]" pageId="7" pageNumber="106">type</typeStatus>
locality; CES registration numbers in Joshi and Karanth (2012: fig. 2, Table S1). These individuals were collected largely in high elevation forests (&lt;1800 msl) of the southern and central parts of the Western Ghats,
<collectingCountry id="BFC47674FFD9FFB0FC06FF28FC37FF53" box="[922,980,151,175]" name="India" pageId="8" pageNumber="107">India</collectingCountry>
(distribution map in Joshi and Karanth 2012: fig. 5c); all leg. J. Joshi,
<date id="B36D1024FFD9FFB0FE6DFF03FD90FF28" box="[497,627,188,212]" pageId="8" pageNumber="107" value="2007" valueMax="2010">2007–2010</date>
.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD9FFB1FF55FF60FB67FCA0" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107" type="description">
<paragraph id="C76C36E4FFD9FFB0FF55FF60FA7EFEC3" blockId="8.[151,1437,151,788]" pageId="8" pageNumber="107">
<emphasis id="F5A7EAF6FFD9FFB0FF55FF60FEB8FF04" bold="true" box="[201,347,223,248]" pageId="8" pageNumber="107">Description.</emphasis>
Length to
<quantity id="002B9B01FFD9FFB0FE40FF60FDD2FF0B" box="[476,561,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="107" unit="mm" value="64.0">64 mm</quantity>
. 17 antennal articles, 18 on one side in one individual in which abnormally short distal articles suggest damage; basal three articles glabrous dorsally (
<figureCitation id="5FE82A61FFD9FFB0FB91FEBBFB87FEE0" box="[1037,1124,260,284]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1726,1749]" captionTargetBox="[151,1435,193,1703]" captionTargetId="figure@7.[151,1436,193,1705]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="8" pageNumber="107">Fig. 13</figureCitation>
), 2.3–2.5 articles glabrous ventrally. Cephalic plate and tergites finely punctate. Longitudinal median furrow 15–20% length of cephalic plate.
</paragraph>
<paragraph id="C76C36E4FFD9FFB0FF55FEF3FA7FFE50" blockId="8.[151,1437,151,788]" pageId="8" pageNumber="107">Colour variable: cephalic plate, basal (glabrous) part of antennae and T1 often blue; trunk may display gradual increase in brown pigment posteriorly, sometimes reverting to blue in posterior tergites; exceptionally cephalic plate and tergites all brown, or most of head and T1 brown with mostly blue tergites from T2; legs pale blue-brown.</paragraph>
<paragraph id="C76C36E4FFD9FFB0FF5BFE08FECDFDC0" blockId="8.[151,1437,151,788]" pageId="8" pageNumber="107">
Forcipular coxosternal tooth plate wider than long (
<figureCitation id="5FE82A61FFD9FFB0FC89FE08FC79FE33" box="[789,922,439,463]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1726,1749]" captionTargetBox="[151,1435,193,1703]" captionTargetId="figure@7.[151,1436,193,1705]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="8" pageNumber="107">Figs 14, 15</figureCitation>
), with four main teeth, the inner two usually grouped, the outer tooth smaller than the inner three (
<figureCitation id="5FE82A61FFD9FFB0FC9AFE63FCBEFE08" box="[774,861,476,500]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1726,1749]" captionTargetBox="[151,1435,193,1703]" captionTargetId="figure@7.[151,1436,193,1705]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="8" pageNumber="107">Fig. 15</figureCitation>
); bases of tooth plates defined by oblique sutures diverging at a relatively obtuse angle of 140–150°. Trochanteroprefemoral process with two distinct teeth along inner margin.
</paragraph>
<paragraph id="C76C36E4FFD9FFB0FF55FDF8FDEFFD78" blockId="8.[151,1437,151,788]" pageId="8" pageNumber="107">Second maxillary claw with slender, needle-like accessory spurs. Article 2 of telopodite bearing a long, slender, pigmented spine distally.</paragraph>
<paragraph id="C76C36E4FFD9FFB0FF55FD30FCC6FCE8" blockId="8.[151,1437,151,788]" pageId="8" pageNumber="107">
Tergites with paramedian sutures complete from TT6 to 8. Tergites marginate from 7, 8 or 9. Tergites smooth (
<figureCitation id="5FE82A61FFD9FFB0FF03FD0BFF1BFD30" box="[159,248,692,716]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1858,1881]" captionTargetBox="[151,1435,859,1803]" captionTargetId="figure@8.[151,1436,859,1804]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Digitipes coonoorensis Jangi and Dass, 1984. 19. Ultimate leg prefemur, ventral view, CES 08960 (female), scale = 0.5 mm. 20. Ultimate leg femur, dorsal view, CES 091088 (male), scale = 0.5 mm. 21. Ultimate leg femur, medial view, CES 091088, scale = 0.5 mm. 22. TT 18 – 20, dorsal view, CES 08960, scale = 1 mm. 23. Spiracle on segment 3, CES 07125, scale = 0.25 mm." httpUri="https://zenodo.org/record/218629/files/figure.png" pageId="8" pageNumber="107">Fig. 22</figureCitation>
), longitudinal median ridge mostly absent, faint from T
<quantity id="002B9B01FFD9FFB0FC0EFD0BFC22FD30" box="[914,961,692,716]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="8" pageNumber="107" unit="in" value="9.0">9 in</quantity>
CES08960. Sternites usually with short paramedian sutures, ca 10% length of sternites in mid body region, at most about 25% length of sternites, sometimes with a short extent on posterior part of sternite.
</paragraph>
<caption id="93AC666CFFD9FFB0FF0BF8FDFEEEF849" httpUri="https://zenodo.org/record/218629/files/figure.png" pageId="8" pageNumber="107" targetBox="[151,1435,859,1803]" targetPageId="8">
<paragraph id="C76C36E4FFD9FFB0FF0BF8FDFEEEF849" blockId="8.[151,1436,1858,1973]" pageId="8" pageNumber="107">
<emphasis id="F5A7EAF6FFD9FFB0FF0BF8FDFEC4F8A4" bold="true" box="[151,295,1858,1881]" pageId="8" pageNumber="107">FIGURES 19</emphasis>
–
<emphasis id="F5A7EAF6FFD9FFB0FEA8F8FCFEB7F8A4" bold="true" box="[308,340,1859,1880]" pageId="8" pageNumber="107">23.</emphasis>
<taxonomicName id="00D34D67FFD9FFB0FEC6F8FBFCC6F8A5" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" box="[346,805,1859,1881]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD9FFB0FEC6F8FBFDA0F8A5" box="[346,579,1860,1881]" italics="true" pageId="8" pageNumber="107">Digitipes coonoorensis</emphasis>
Jangi and Dass, 1984
</taxonomicName>
. 19. Ultimate leg prefemur, ventral view, CES08960 (female), scale = 0.5 mm. 20. Ultimate leg femur, dorsal view, CES091088 (male), scale = 0.5 mm. 21. Ultimate leg femur, medial view, CES091088, scale = 0.5 mm. 22. TT18–20, dorsal view, CES08960, scale = 1 mm. 23. Spiracle on segment 3, CES07125, scale = 0.25 mm.
</paragraph>
</caption>
<paragraph id="C76C36E4FFD8FFB1FF55FF28FD25FEE0" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="108">
Spiracle of segment 3 with narrow elliptical outline (
<figureCitation id="5FE82A61FFD8FFB1FCBDFF28FC96FF53" box="[801,885,151,175]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1858,1881]" captionTargetBox="[151,1435,859,1803]" captionTargetId="figure@8.[151,1436,859,1804]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Digitipes coonoorensis Jangi and Dass, 1984. 19. Ultimate leg prefemur, ventral view, CES 08960 (female), scale = 0.5 mm. 20. Ultimate leg femur, dorsal view, CES 091088 (male), scale = 0.5 mm. 21. Ultimate leg femur, medial view, CES 091088, scale = 0.5 mm. 22. TT 18 – 20, dorsal view, CES 08960, scale = 1 mm. 23. Spiracle on segment 3, CES 07125, scale = 0.25 mm." httpUri="https://zenodo.org/record/218629/files/figure.png" pageId="9" pageNumber="108">Fig. 23</figureCitation>
), long axis oblique to horizontal plane; spiracles from segment 5 with rounded outline, irregular inner margin of peritrema, long axis of spiracular opening vertical. All spiracles with regularly scalloped inner margin of peritrema, folding into a smooth band of trichomes beneath which atrium has strongly humped wall and floor.
</paragraph>
<paragraph id="C76C36E4FFD8FFB1FF55FE98FBA3FE98" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="108">First three (four in one individual) pairs of legs with two tarsal spurs, the subsequent up to 19 with one. Leg 20 without tarsal spur, except one individual. A tibial and a femoral spur on leg 1 only.</paragraph>
<paragraph id="C76C36E4FFD8FFB1FF55FED0FA74FE33" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="108">
Tergite of ultimate leg-bearing segment with gently convex lateral margins that weakly converge posteriorly; posterior margin gently sinuous, bluntly pointed to rounded posteromedially (
<figureCitation id="5FE82A61FFD8FFB1FB86FE2BFB92FE50" box="[1050,1137,404,428]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1726,1749]" captionTargetBox="[151,1435,193,1703]" captionTargetId="figure@7.[151,1436,193,1705]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="9" pageNumber="108">Fig. 17</figureCitation>
). Sternite of ultimate legbearing segment with sides converging posteriorly, nearly straight, posterior margin moderately concave (
<figureCitation id="5FE82A61FFD8FFB1FAABFE08FA69FE33" box="[1335,1418,439,463]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1726,1749]" captionTargetBox="[151,1435,193,1703]" captionTargetId="figure@7.[151,1436,193,1705]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="9" pageNumber="108">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFD8FFB1FF55FE63FB67FCA0" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="108">
Coxopleural process moderately long, robustly conical, set off from course of posterolateral margin of coxopleuron by a pronounced inflection, its apex variably with a slight to distinct lateral orientation (
<figureCitation id="5FE82A61FFD8FFB1FA95FE40FABEFDEB" box="[1289,1373,511,535]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1726,1749]" captionTargetBox="[151,1435,193,1703]" captionTargetId="figure@7.[151,1436,193,1705]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="9" pageNumber="108">Fig. 18</figureCitation>
); two apical spines, one lateral spine (absent on one side in two individuals). Coxopleural pores relatively dense, approximately bimodal size distribution; pore field relatively narrow, terminating moderately to strongly beneath dorsal margin of coxopleuron (
<figureCitation id="5FE82A61FFD8FFB1FE66FDD3FDB3FD78" box="[506,592,620,644]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1726,1749]" captionTargetBox="[151,1435,193,1703]" captionTargetId="figure@7.[151,1436,193,1705]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="9" pageNumber="108">Fig. 16</figureCitation>
); non-porose area on coxopleural process at least 60% length to posterior margin of sternite of ultimate leg-bearing segment, sometimes nearly reaching that margin (
<figureCitation id="5FE82A61FFD8FFB1FB3AFD30FB18FD5B" box="[1190,1275,655,679]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1726,1749]" captionTargetBox="[151,1435,193,1703]" captionTargetId="figure@7.[151,1436,193,1705]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Digitipes coonoorensis Jangi and Dass, 1984. CES 08960 (female) except Fig. 15, CES 07125 (female) and Fig. 16, CES 091088 (male). 13. Cephalic plate and T 1, dorsal view, scale = 1 mm. 14. Forcipular segment, ventral view, scale = 1 mm. 15. Tooth plates, ventral view, scale = 1 mm. 16. Coxopleuron, lateral view, scale = 1 mm. 17. Tergite of ultimate legbearing segment, dorsal view, scale = 0.5 mm. 18. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218628/files/figure.png" pageId="9" pageNumber="108">Fig. 18</figureCitation>
). Ultimate leg prefemur with width at distal end about 30% its length. Prefemoral spines moderately large (
<figureCitation id="5FE82A61FFD8FFB1FB36FD0BFB1DFD30" box="[1194,1278,692,716]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1858,1881]" captionTargetBox="[151,1435,859,1803]" captionTargetId="figure@8.[151,1436,859,1804]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Digitipes coonoorensis Jangi and Dass, 1984. 19. Ultimate leg prefemur, ventral view, CES 08960 (female), scale = 0.5 mm. 20. Ultimate leg femur, dorsal view, CES 091088 (male), scale = 0.5 mm. 21. Ultimate leg femur, medial view, CES 091088, scale = 0.5 mm. 22. TT 18 – 20, dorsal view, CES 08960, scale = 1 mm. 23. Spiracle on segment 3, CES 07125, scale = 0.25 mm." httpUri="https://zenodo.org/record/218629/files/figure.png" pageId="9" pageNumber="108">Fig. 19</figureCitation>
): VL 3 (small fourth spine rarely present), VM 2 (one specimen with 3 on one side), DM 0 or 1. Distomedial process in males bluntly conical, extending about as far as the distal end of the femur on its dorsal side (
<figureCitation id="5FE82A61FFD8FFB1FBF1FD43FB21FCE8" box="[1133,1218,764,788]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1858,1881]" captionTargetBox="[151,1435,859,1803]" captionTargetId="figure@8.[151,1436,859,1804]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Digitipes coonoorensis Jangi and Dass, 1984. 19. Ultimate leg prefemur, ventral view, CES 08960 (female), scale = 0.5 mm. 20. Ultimate leg femur, dorsal view, CES 091088 (male), scale = 0.5 mm. 21. Ultimate leg femur, medial view, CES 091088, scale = 0.5 mm. 22. TT 18 – 20, dorsal view, CES 08960, scale = 1 mm. 23. Spiracle on segment 3, CES 07125, scale = 0.25 mm." httpUri="https://zenodo.org/record/218629/files/figure.png" pageId="9" pageNumber="108">Fig. 20</figureCitation>
); groove on medial surface of femur extending about half length of femur (
<figureCitation id="5FE82A61FFD8FFB1FD62FCA0FCB2FCCB" box="[766,849,799,823]" captionStart="FIGURES 19 – 23" captionStartId="8.[151,264,1858,1881]" captionTargetBox="[151,1435,859,1803]" captionTargetId="figure@8.[151,1436,859,1804]" captionTargetPageId="8" captionText="FIGURES 19 – 23. Digitipes coonoorensis Jangi and Dass, 1984. 19. Ultimate leg prefemur, ventral view, CES 08960 (female), scale = 0.5 mm. 20. Ultimate leg femur, dorsal view, CES 091088 (male), scale = 0.5 mm. 21. Ultimate leg femur, medial view, CES 091088, scale = 0.5 mm. 22. TT 18 – 20, dorsal view, CES 08960, scale = 1 mm. 23. Spiracle on segment 3, CES 07125, scale = 0.25 mm." httpUri="https://zenodo.org/record/218629/files/figure.png" pageId="9" pageNumber="108">Fig. 21</figureCitation>
). Ultimate leg tarsus 1 1.6–2.5 times length of tarsus 2; tarsus 1 2.2–5 times longer than pretarsus; pretarsus with pair of short accessory claws.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD8FFB1FF55FCD8FDB2FB14" pageId="9" pageNumber="108" type="discussion">
<paragraph id="C76C36E4FFD8FFB1FF55FCD8FB83FC10" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="108">
<emphasis id="F5A7EAF6FFD8FFB1FF55FCD8FEADFC7C" bold="true" box="[201,334,871,896]" pageId="9" pageNumber="108">Discussion.</emphasis>
Absence of a leg 20 tarsal spur was used by Jangi and Dass (1984: 41) in the first couplet of their key to Indian species of
<taxonomicName id="00D34D67FFD8FFB1FE32FC32FDF7FC58" box="[430,532,909,932]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD8FFB1FE32FC32FDF7FC58" box="[430,532,909,932]" italics="true" pageId="9" pageNumber="108">Digitipes</emphasis>
</taxonomicName>
, allowing
<taxonomicName id="00D34D67FFD8FFB1FD12FC32FCA5FC58" box="[654,838,909,932]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD8FFB1FD12FC32FCA5FC58" box="[654,838,909,932]" italics="true" pageId="9" pageNumber="108">D. coonoorensis</emphasis>
</taxonomicName>
(spur absent) to be distinguished from
<taxonomicName id="00D34D67FFD8FFB1FA9BFC32FA7FFC58" box="[1287,1436,908,932]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="barnabasi">
<emphasis id="F5A7EAF6FFD8FFB1FA9BFC32FA7FFC58" box="[1287,1436,908,932]" italics="true" pageId="9" pageNumber="108">D. barnabasi</emphasis>
</taxonomicName>
and
<taxonomicName id="00D34D67FFD8FFB1FF5BFC0EFED8FC34" box="[199,315,944,968]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFD8FFB1FF5BFC0EFED8FC34" box="[199,315,944,968]" italics="true" pageId="9" pageNumber="108">D. indicus</emphasis>
</taxonomicName>
(spur present). We observe this character to remain reliable for separation of
<taxonomicName id="00D34D67FFD8FFB1FB39FC0EFABFFC34" box="[1189,1372,945,968]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD8FFB1FB39FC0EFABFFC34" box="[1189,1372,945,968]" italics="true" pageId="9" pageNumber="108">D. coonoorensis</emphasis>
</taxonomicName>
apart from one individual resolved within the species phylogenetically that possesses a spur.
</paragraph>
<paragraph id="C76C36E4FFD8FFB1FF55FC47FDD2FB80" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="108">
A record of
<taxonomicName id="00D34D67FFD8FFB1FECDFC46FDE4FBEC" box="[337,519,1017,1040]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFD8FFB1FECDFC46FDE4FBEC" box="[337,519,1017,1040]" italics="true" pageId="9" pageNumber="108">D. coonoorensis</emphasis>
</taxonomicName>
was made by Yadav (1993b: 324) from Sindhudurg, Maharashtra, more northerly than any material of the species examined by us. We have not seen the material and have not included the record in the synonymy. All the individuals examined here were found in the shola forests or grasslands of high elevation areas of the southern Western Ghats.
</paragraph>
<paragraph id="C76C36E4FFD8FFB1FF55FB37FDB2FB14" blockId="9.[151,1437,151,1256]" pageId="9" pageNumber="108">
Light-microscopic description of the spiracles is included for this species. For comparative purposes, the spiracle on segment 3 is depicted for other species of the genus as well (
<figureCitation id="5FE82A61FFD8FFB1FC74FB13FB8CFB38" box="[1000,1135,1196,1220]" captionStart="FIGURES 36 – 45" captionStartId="12.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 36 – 45. Digitipes jonesii (Verhoeff, 1938). 36 – 40, CES 091062 (male); 41 – 45, CES 091090 (male). 36. Ultimate leg prefemur, ventral view, scale = 1 mm. 37. Ultimate leg femur, dorsal view, scale = 1 mm. 38. Ultimate leg femur, medial view, scale = 1 mm. 39. TT 18 – 19, dorsal view, scale = 1 mm. 40. Spiracle on segment 3, lateral view, scale = 0.5 mm. 41. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 42. Ultimate leg femur, medial view, scale = 0.5 mm. 43. Ultimate leg femur, ventral view, scale = 1 mm. 44. TT 18 – 20, dorsal view, scale = 1 mm. 45. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218632/files/figure.png" pageId="9" pageNumber="108">Figs 40, 45</figureCitation>
,
<figureCitation id="5FE82A61FFD8FFB1FBE2FB13FB7EFB38" box="[1150,1181,1196,1220]" captionStart="FIGURES 52 – 56" captionStartId="16.[151,264,1116,1139]" captionTargetBox="[165,1408,185,1098]" captionTargetId="figure@16.[165,1409,185,1099]" captionTargetPageId="16" captionText="FIGURES 52 – 56. Digitipes jangii n. sp. CES 08912 (holotype male) except Fig. 56, CES 08915 (paratype male). 52. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 53. Ultimate leg femur, dorsal view, scale = 0.5 mm. 54. Ultimate leg femur, medial view, scale = 0.5 mm. 55. TT 18 – 20, dorsal view, scale = 0.5 mm. 56. Spiracle on segment 3, scale = 0.25 mm." httpUri="https://zenodo.org/record/218634/files/figure.png" pageId="9" pageNumber="108">56</figureCitation>
,
<figureCitation id="5FE82A61FFD8FFB1FB37FB13FB29FB38" box="[1195,1226,1196,1220]" captionStart="FIGURES 63 – 67" captionStartId="19.[151,264,1159,1182]" captionTargetBox="[151,1435,193,1137]" captionTargetId="figure@19.[151,1436,193,1138]" captionTargetPageId="19" captionText="FIGURES 63 – 67. Digitipes periyarensis n. sp. Holotype CES 091037 (female) except Fig. 63, CES 091038 (female). 63. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 64. Ultimate leg, dorsal view, scale = 1 mm. 65. Ultimate leg femur and tibia, dorsal view, scale = 0.5 mm. 66. TT 18 – 20, dorsal view, scale = 1 mm. 67. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218636/files/figure.png" pageId="9" pageNumber="108">67</figureCitation>
), but we have not identified obvious taxonomic variation.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="FEB9B7F52CF601724A132BE606AE627E" ID-DOI="10.11646/zootaxa.3626.1.5" ID-GBIF-Dataset="135b7553-b75c-4020-b1a6-c689a05b7e66" ID-ISSN="1175-5326" ID-Zenodo-Dep="218625" ID-ZooBank="58AD6857-8CDD-4423-88D0-619CD8D793AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460581053656" checkinUser="plazi" docAuthor="Joshi, Jahnavi &amp; Edgecombe, Gregory D." docDate="2013" docId="4F7A87F2FFD8FFB6FF0BFA83FB8EFD78" docLanguage="en" docName="zt03626p145.pdf" docOrigin="Zootaxa 3626 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Digitipes jonesii Verhoeff 1938, n. comb" docType="treatment" docVersion="8" lastPageNumber="112" masterDocId="B343FF8AFFD1FFB8FF9CFFBFFFE3FFFC" masterDocTitle="Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity" masterLastPageNumber="145" masterPageNumber="99" pageNumber="108" updateTime="1698324300789" updateUser="plazi">
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<mods:title id="C7C421AAC7634F3848F2E99B980F74D8">Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity</mods:title>
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<mods:namePart id="803F1F8C88F8A9F43C878F33AB309746">Joshi, Jahnavi</mods:namePart>
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<mods:namePart id="6EA391291444754206E69A32FCEDA3BD">Edgecombe, Gregory D.</mods:namePart>
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<treatment id="4F7A87F2FFD8FFB6FF0BFA83FB8EFD78" ID-DOI="http://doi.org/10.5281/zenodo.6154118" ID-GBIF-Taxon="119563402" ID-Zenodo-Dep="6154118" LSID="urn:lsid:plazi:treatment:4F7A87F2FFD8FFB6FF0BFA83FB8EFD78" httpUri="http://treatment.plazi.org/id/4F7A87F2FFD8FFB6FF0BFA83FB8EFD78" lastPageId="14" lastPageNumber="112" pageId="9" pageNumber="108">
<subSubSection id="8FC9656FFFD8FFB1FF0BFA83FD86FA05" pageId="9" pageNumber="108" type="nomenclature">
<paragraph id="C76C36E4FFD8FFB1FF0BFA83FD70FAA9" blockId="9.[151,659,1339,1398]" box="[151,659,1339,1366]" pageId="9" pageNumber="108">
<heading id="9C248188FFD8FFB1FF0BFA83FD70FAA9" bold="true" box="[151,659,1339,1366]" fontSize="11" level="1" pageId="9" pageNumber="108" reason="1">
<emphasis id="F5A7EAF6FFD8FFB1FF0BFA83FD70FAA9" bold="true" box="[151,659,1339,1366]" pageId="9" pageNumber="108">
<taxonomicName id="00D34D67FFD8FFB1FF0BFA83FDCAFAA9" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" box="[151,553,1339,1366]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="jonesii" status="comb. nov.">
<emphasis id="F5A7EAF6FFD8FFB1FF0BFA83FEB9FAAA" bold="true" box="[151,346,1340,1366]" italics="true" pageId="9" pageNumber="108">Digitipes jonesii</emphasis>
(Verhoeff, 1938)
</taxonomicName>
<taxonomicNameLabel id="EE94578DFFD8FFB1FDACFA84FD70FAA9" box="[560,659,1339,1365]" pageId="9" pageNumber="108">n. comb</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C76C36E4FFD8FFB1FF0BFAE1FEC4FA8A" blockId="9.[151,659,1339,1398]" box="[151,295,1374,1398]" pageId="9" pageNumber="108">
(
<figureCitation id="5FE82A61FFD8FFB1FF03FAE1FEFCFA8A" box="[159,287,1374,1398]" captionStart-0="FIGURES 24 – 29" captionStart-1="FIGURES 30 – 35" captionStart-2="FIGURES 36 – 45" captionStartId-0="10.[151,264,1636,1659]" captionStartId-1="11.[151,264,1877,1900]" captionStartId-2="12.[151,264,1877,1900]" captionTargetBox-0="[151,1435,187,1624]" captionTargetBox-1="[151,1435,232,1854]" captionTargetBox-2="[151,1435,193,1854]" captionTargetId-0="figure@10.[151,1436,187,1627]" captionTargetId-1="figure@11.[151,1436,193,1856]" captionTargetId-2="figure@12.[151,1436,193,1856]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." captionText-1="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." captionText-2="FIGURES 36 – 45. Digitipes jonesii (Verhoeff, 1938). 36 – 40, CES 091062 (male); 41 – 45, CES 091090 (male). 36. Ultimate leg prefemur, ventral view, scale = 1 mm. 37. Ultimate leg femur, dorsal view, scale = 1 mm. 38. Ultimate leg femur, medial view, scale = 1 mm. 39. TT 18 – 19, dorsal view, scale = 1 mm. 40. Spiracle on segment 3, lateral view, scale = 0.5 mm. 41. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 42. Ultimate leg femur, medial view, scale = 0.5 mm. 43. Ultimate leg femur, ventral view, scale = 1 mm. 44. TT 18 – 20, dorsal view, scale = 1 mm. 45. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri-0="https://zenodo.org/record/218630/files/figure.png" httpUri-1="https://zenodo.org/record/218631/files/figure.png" httpUri-2="https://zenodo.org/record/218632/files/figure.png" pageId="9" pageNumber="108">Figs 24–45</figureCitation>
)
</paragraph>
<paragraph id="C76C36E4FFD8FFB1FF0BFA18FD86FA05" blockId="9.[151,838,1446,1591]" pageId="9" pageNumber="108">
<taxonomicName id="00D34D67FFD8FFB1FF0BFA18FCA1FA40" authority="Verhoeff, 1938: 384" authorityName="Verhoeff" authorityPageNumber="384" authorityYear="1938" box="[151,834,1446,1469]" class="Chilopoda" family="Scolopendridae" genus="Arthrorhabdus" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="jonesii" subGenus="Trachycormocephalus">
<emphasis id="F5A7EAF6FFD8FFB1FF0BFA18FECFFA41" box="[151,300,1447,1469]" italics="true" pageId="9" pageNumber="108">Arthrorhabdus</emphasis>
(
<emphasis id="F5A7EAF6FFD8FFB1FEA7FA18FDFFFA41" box="[315,540,1447,1469]" italics="true" pageId="9" pageNumber="108">Trachycormocephalus</emphasis>
)
<emphasis id="F5A7EAF6FFD8FFB1FDB6FA17FD8DFA41" box="[554,622,1448,1469]" italics="true" pageId="9" pageNumber="108">jonesii</emphasis>
Verhoeff, 1938: 384
</taxonomicName>
.
<taxonomicName id="00D34D67FFD8FFB1FF0BFA79FDAAFA27" authority="Jangi and Dass, 1984: 46" authorityName="Jangi and Dass" authorityPageNumber="46" authorityYear="1984" box="[151,585,1477,1499]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFD8FFB1FF0BFA79FEA0FA27" box="[151,323,1477,1499]" italics="true" pageId="9" pageNumber="108">Digitipes indicus</emphasis>
Jangi and Dass, 1984: 46
</taxonomicName>
, figs 87–92.
<emphasis id="F5A7EAF6FFD8FFB1FD50FA7AFCCEFA27" bold="true" box="[716,813,1477,1500]" pageId="9" pageNumber="108">Syn. nov.</emphasis>
<taxonomicName id="00D34D67FFD8FFB1FF0BFA5AFEA0FA06" box="[151,323,1508,1530]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFD8FFB1FF0BFA5AFEA0FA06" box="[151,323,1508,1530]" italics="true" pageId="9" pageNumber="108">Digitipes indicus</emphasis>
</taxonomicName>
. Sureshan
<emphasis id="F5A7EAF6FFD8FFB1FE2FFA5AFE3CFA06" box="[435,479,1508,1530]" italics="true" pageId="9" pageNumber="108">et al</emphasis>
. 2006: 2287.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD8FFB1FF0BF9BCFCD3F9CB" pageId="9" pageNumber="108" type="description">
<paragraph id="C76C36E4FFD8FFB1FF0BF9BCFCD3F9CB" blockId="9.[151,838,1446,1591]" pageId="9" pageNumber="108">
<taxonomicName id="00D34D67FFD8FFB1FF0BF9BCFF17F9E4" box="[151,244,1539,1560]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD8FFB1FF0BF9BCFF17F9E4" box="[151,244,1539,1560]" italics="true" pageId="9" pageNumber="108">Digitipes</emphasis>
</taxonomicName>
putative species 2. Joshi and Karanth, 2012: figs 2, 3, 6.
<taxonomicName id="00D34D67FFD8FFB1FF0BF99DFF17F9CB" box="[151,244,1570,1591]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD8FFB1FF0BF99DFF17F9CB" box="[151,244,1570,1591]" italics="true" pageId="9" pageNumber="108">Digitipes</emphasis>
</taxonomicName>
putative species 3. Joshi and Karanth, 2012: figs 2, 3, 6.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD8FFB1FF0BF9DCFE63F8F0" pageId="9" pageNumber="108" type="diagnosis">
<paragraph id="C76C36E4FFD8FFB1FF0BF9DCFE63F8F0" blockId="9.[151,1436,1635,1984]" pageId="9" pageNumber="108">
<emphasis id="F5A7EAF6FFD8FFB1FF0BF9DCFEF2F980" bold="true" box="[151,273,1635,1660]" pageId="9" pageNumber="108">Diagnosis.</emphasis>
<taxonomicName id="00D34D67FFD8FFB1FE8BF9DAFE9DF980" box="[279,382,1637,1660]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD8FFB1FE8BF9DAFE9DF980" box="[279,382,1637,1660]" italics="true" pageId="9" pageNumber="108">Digitipes</emphasis>
</taxonomicName>
with basal three antennal articles glabrous dorsally. Tergite paramedian sutures complete from TT6 to 8; tergites smooth, longitudinal median ridge usually absent, when present faint, incomplete. Coxopleural process relatively short, lateral spine variably present. Dorsomedial process on ultimate leg femur of male extending further than distal end of the femur on its dorsal side. Legs variably bearing many short, fine setae. Leg 20 with a tarsal spur.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFD8FFB1FF55F8A8FB51F83C" pageId="9" pageNumber="108" type="materials_examined">
<paragraph id="C76C36E4FFD8FFB1FF55F8A8FB51F83C" blockId="9.[151,1436,1635,1984]" pageId="9" pageNumber="108">
<emphasis id="F5A7EAF6FFD8FFB1FF55F8A8FE51F8CC" bold="true" box="[201,434,1815,1840]" pageId="9" pageNumber="108">Material examined.</emphasis>
<specimenCount id="D1D5FD6DFFD8FFB1FE25F8A8FDB2F8D3" box="[441,593,1815,1839]" pageId="9" pageNumber="108" type="generic">20 specimens</specimenCount>
of
<taxonomicName id="00D34D67FFD8FFB1FDEAF8A6FD3EF8CC" box="[630,733,1817,1840]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD8FFB1FDEAF8A6FD3EF8CC" box="[630,733,1817,1840]" italics="true" pageId="9" pageNumber="108">Digitipes</emphasis>
</taxonomicName>
putative species 3 (
<taxonomicName id="00D34D67FFD8FFB1FC27F8A6FBCEF8CC" box="[955,1069,1816,1840]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFD8FFB1FC27F8A6FBCEF8CC" box="[955,1069,1816,1840]" italics="true" pageId="9" pageNumber="108">D. indicus</emphasis>
</taxonomicName>
of Joshi and Karanth, 2012) and 25 of
<taxonomicName id="00D34D67FFD8FFB1FF40F882FEA0F8A8" box="[220,323,1853,1876]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFD8FFB1FF40F882FEA0F8A8" box="[220,323,1853,1876]" italics="true" pageId="9" pageNumber="108">Digitipes</emphasis>
</taxonomicName>
putative species 2 including CES07196 and CES07198 from the
<typeStatus id="18688846FFD8FFB1FBAFF882FB80F8A9" box="[1075,1123,1853,1877]" pageId="9" pageNumber="108">type</typeStatus>
locality of
<taxonomicName id="00D34D67FFD8FFB1FB75F882FABBF8A8" box="[1257,1368,1853,1876]" class="Chilopoda" family="Scolopendridae" genus="Arthrorhabdus" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFD8FFB1FB75F882FABBF8A8" box="[1257,1368,1853,1876]" italics="true" pageId="9" pageNumber="108">D. jonesii</emphasis>
</taxonomicName>
; CES registration numbers in Joshi and Karanth (2012: fig. 2, Table S1). The specimens were collected from low and mid elevation forests (100–1500 msl) of the southern Western Ghats, including four specimens from the
<typeStatus id="18688846FFD8FFB1FB6FF83AFAC0F861" box="[1267,1315,1925,1949]" pageId="9" pageNumber="108">type</typeStatus>
locality of
<taxonomicName id="00D34D67FFD8FFB1FF0BF816FEE9F83C" box="[151,266,1960,1984]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFD8FFB1FF0BF816FEE9F83C" box="[151,266,1960,1984]" italics="true" pageId="9" pageNumber="108">D. indicus</emphasis>
</taxonomicName>
(distribution maps in Joshi and Karanth 2012: fig. 5c); all leg. J. Joshi,
<date id="B36D1024FFD8FFB1FBB1F817FB4DF83C" box="[1069,1198,1960,1984]" pageId="9" pageNumber="108" value="2007" valueMax="2010">2007–2010</date>
.
</paragraph>
</subSubSection>
<caption id="93AC666CFFDBFFB2FF0BF9DBFEDAF909" httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="10" pageNumber="109" targetBox="[151,1435,187,1624]" targetPageId="10">
<paragraph id="C76C36E4FFDBFFB2FF0BF9DBFEDAF909" blockId="10.[151,1436,1636,1781]" pageId="10" pageNumber="109">
<emphasis id="F5A7EAF6FFDBFFB2FF0BF9DBFEC5F985" bold="true" box="[151,294,1636,1659]" pageId="10" pageNumber="109">FIGURES 24</emphasis>
–
<emphasis id="F5A7EAF6FFDBFFB2FEAFF9DBFEB0F985" bold="true" box="[307,339,1636,1657]" pageId="10" pageNumber="109">29.</emphasis>
<taxonomicName id="00D34D67FFDBFFB2FEC5F9DAFD4FF987" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" box="[345,684,1636,1659]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDBFFB2FEC5F9DAFE1CF986" box="[345,511,1637,1658]" italics="true" pageId="10" pageNumber="109">Digitipes jonesii</emphasis>
(Verhoeff, 1938)
</taxonomicName>
(putative species 3 of Joshi and Karanth 2012). CES091062 (male) except Fig. 27, CES091047 (female). 24. Cephalic plate and T1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="8FC9656FFFDBFFB5FF55F89EFBCBFB5C" lastPageId="13" lastPageNumber="112" pageId="10" pageNumber="109" type="description">
<paragraph id="C76C36E4FFDBFFB2FF55F89EFC59F87E" blockId="10.[151,1436,1825,2030]" pageId="10" pageNumber="109">
<emphasis id="F5A7EAF6FFDBFFB2FF55F89EFEB8F8C6" bold="true" box="[201,347,1825,1850]" pageId="10" pageNumber="109">Description.</emphasis>
Length to
<quantity id="002B9B01FFDBFFB2FE4FF89DFDC7F8C6" box="[467,548,1826,1850]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="109" unit="mm" value="55.0">55 mm</quantity>
. 17 antennal articles (apart from two individuals with 18/18 and 18/19 articles); basal three articles glabrous dorsally (
<figureCitation id="5FE82A61FFDBFFB2FDD6F8F9FD47F8A2" box="[586,676,1862,1886]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="10" pageNumber="109">Figs 24</figureCitation>
,
<figureCitation id="5FE82A61FFDBFFB2FD2EF8F9FD32F8A2" box="[690,721,1862,1886]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="10" pageNumber="109">30</figureCitation>
), ca 2.5 articles glabrous ventrally. Cephalic plate and tergites punctate. Longitudinal median furrow 10–20% length of cephalic plate.
</paragraph>
<paragraph id="C76C36E4FFDBFFB2FF55F831FF19F812" blockId="10.[151,1436,1825,2030]" pageId="10" pageNumber="109">Cephalic plate blue-olive green or blue anteriorly and mostly brown posteriorly; most of trunk tergites mixed blue-brown, darker in posterior half of trunk; basal (glabrous) part of antenna light blue, distal part variably purplish.</paragraph>
<caption id="93AC666CFFDAFFB3FF0BF8EAFDF3F81A" httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="11" pageNumber="110" targetBox="[151,1435,232,1854]" targetPageId="11">
<paragraph id="C76C36E4FFDAFFB3FF0BF8EAFDF3F81A" blockId="11.[151,1437,1877,2022]" pageId="11" pageNumber="110">
<emphasis id="F5A7EAF6FFDAFFB3FF0BF8EAFECAF896" bold="true" box="[151,297,1877,1900]" pageId="11" pageNumber="110">FIGURES 30</emphasis>
–
<emphasis id="F5A7EAF6FFDAFFB3FEAAF8EAFEB5F896" bold="true" box="[310,342,1877,1898]" pageId="11" pageNumber="110">35.</emphasis>
<taxonomicName id="00D34D67FFDAFFB3FEC2F8E9FD5AF890" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" box="[350,697,1877,1900]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="11" pageNumber="110" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDAFFB3FEC2F8E9FDE5F897" box="[350,518,1878,1899]" italics="true" pageId="11" pageNumber="110">Digitipes jonesii</emphasis>
(Verhoeff, 1938)
</taxonomicName>
(putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES07166 (female); 31, 34, 35, CES091090 (male). 30. Cephalic plate and TT1–2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm.
</paragraph>
</caption>
<caption id="93AC666CFFDDFFB4FF0BF8EAFA7EF81A" httpUri="https://zenodo.org/record/218632/files/figure.png" pageId="12" pageNumber="111" targetBox="[151,1435,193,1854]" targetPageId="12">
<paragraph id="C76C36E4FFDDFFB4FF0BF8EAFA7EF81A" blockId="12.[151,1437,1877,2022]" pageId="12" pageNumber="111">
<emphasis id="F5A7EAF6FFDDFFB4FF0BF8EAFEC5F896" bold="true" box="[151,294,1877,1900]" pageId="12" pageNumber="111">FIGURES 36</emphasis>
–
<emphasis id="F5A7EAF6FFDDFFB4FEA8F8EAFEB0F896" bold="true" box="[308,339,1877,1898]" pageId="12" pageNumber="111">45.</emphasis>
<taxonomicName id="00D34D67FFDDFFB4FEC5F8E9FD51F890" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" box="[345,690,1877,1900]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="12" pageNumber="111" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDDFFB4FEC5F8E9FDE3F897" box="[345,512,1878,1899]" italics="true" pageId="12" pageNumber="111">Digitipes jonesii</emphasis>
(Verhoeff, 1938)
</taxonomicName>
. 36–40, CES091062 (male); 41–45, CES091090 (male). 36. Ultimate leg prefemur, ventral view, scale = 1 mm. 37. Ultimate leg femur, dorsal view, scale = 1 mm. 38. Ultimate leg femur, medial view, scale = 1 mm. 39. TT18–19, dorsal view, scale = 1 mm. 40. Spiracle on segment 3, lateral view, scale = 0.5 mm. 41. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 42. Ultimate leg femur, medial view, scale = 0.5 mm. 43. Ultimate leg femur, ventral view, scale = 1 mm. 44. TT18–20, dorsal view, scale = 1 mm. 45. Spiracle on segment 3, lateral view, scale = 0.25 mm.
</paragraph>
</caption>
<paragraph id="C76C36E4FFDCFFB5FF55FF28FBA2FEE0" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="112">
Forcipular coxosternal tooth plate at least slightly wider than long, with four main teeth (
<figureCitation id="5FE82A61FFDCFFB5FB39FF28FB1FFF53" box="[1189,1276,151,175]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="13" pageNumber="112">Figs 25</figureCitation>
,
<figureCitation id="5FE82A61FFDCFFB5FA9BFF28FAC6FF53" box="[1287,1317,151,175]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="13" pageNumber="112">31</figureCitation>
), the inner two usually separated from the outer two (
<figureCitation id="5FE82A61FFDCFFB5FD19FF03FD03FF28" box="[645,736,188,212]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="13" pageNumber="112">Figs 26</figureCitation>
,
<figureCitation id="5FE82A61FFDCFFB5FD72FF03FCEEFF28" box="[750,781,188,212]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="13" pageNumber="112">32</figureCitation>
), sometimes partially fused, outer tooth smaller than the inner three; bases of tooth plates defined by oblique sutures diverging at a relatively obtuse angle of 135–150°. Trochanteroprefemoral process with two distinct teeth along inner margin (
<figureCitation id="5FE82A61FFDCFFB5FC7CFEBBFBD7FEE0" box="[992,1076,260,284]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="13" pageNumber="112">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFDCFFB5FF55FE98FA9DFEC3" blockId="13.[151,1437,151,2012]" box="[201,1406,295,320]" pageId="13" pageNumber="112">Second maxillary claw with slender accessory spurs. Article 2 of telopodite bearing a slender spine distally.</paragraph>
<paragraph id="C76C36E4FFDCFFB5FF55FEF3FD2EFE08" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="112">Tergites with paramedian sutures complete from TT6 to 8. Tergites marginate starting from 7 to 13, most specimens starting from 8–10. Tergites smooth apart from faint rugosity on lateral parts in some specimens; longitudinal median ridge mostly absent, a few specimens with a faint, posteriorly incomplete flat-topped ridge from TT3 or 4. Sternites with paramedian sutures mostly 10–15% length of sternites in mid body region, exceptionally confined to anterior edge of sternite.</paragraph>
<paragraph id="C76C36E4FFDCFFB5FF55FE40FE27FDA3" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="112">First three or four pairs of legs with two tarsal spurs, the subsequent up to 20 with one. A tibial and a femoral spur on leg 1 only. All articles of legs in some populations (especially in PS3 of Joshi and Karanth 2012) bear numerous short, fine setae.</paragraph>
<paragraph id="C76C36E4FFDCFFB5FF55FDD3FE73FD13" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="112">
Tergite of ultimate leg-bearing segment with gently convex lateral margins that faintly converge posteriorly; straight to weakly sinuous posterolateral margins, pointed posteromedially (
<figureCitation id="5FE82A61FFDCFFB5FC6FFD30FBA8FD5B" box="[1011,1099,655,679]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="13" pageNumber="112">Figs 28</figureCitation>
,
<figureCitation id="5FE82A61FFDCFFB5FBC5FD30FB9AFD5B" box="[1113,1145,655,679]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="13" pageNumber="112">34</figureCitation>
). Sternite of ultimate legbearing segment with sides converging relatively strongly posteriorly, nearly straight, posterior margin moderately concave (
<figureCitation id="5FE82A61FFDCFFB5FE9DFD68FEBAFD13" box="[257,345,727,751]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="13" pageNumber="112">Figs 29</figureCitation>
,
<figureCitation id="5FE82A61FFDCFFB5FEF8FD68FE60FD13" box="[356,387,727,751]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="13" pageNumber="112">35</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFDCFFB5FF55FD43FBCBFB5C" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="112">
Coxopleural process mostly relatively short (
<figureCitation id="5FE82A61FFDCFFB5FD57FD43FCC3FCE8" box="[715,800,764,788]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="13" pageNumber="112">Fig. 29</figureCitation>
), at most moderately long but stoutly conical in ventral view (
<figureCitation id="5FE82A61FFDCFFB5FF40FCA0FECCFCCB" box="[220,303,799,823]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="13" pageNumber="112">Fig. 35</figureCitation>
), its long axis often oriented slightly laterally; two apical spines; lateral spine(s) either absent on both sides of coxopleural process (most common in PS3 of Joshi and Karanth 2012), present only on one side, or present on both sides (most common in PS2 of Joshi and Karanth 2012), ranging from minute to similar size to apical spines (
<figureCitation id="5FE82A61FFDCFFB5FF72FC33FEA0FC58" box="[238,323,908,932]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="13" pageNumber="112">Fig. 35</figureCitation>
). Pores relatively dense; pore field not reaching dorsal margin of coxopleuron (
<figureCitation id="5FE82A61FFDCFFB5FB51FC33FAC4FC58" box="[1229,1319,908,932]" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="13" pageNumber="112">Figs 27</figureCitation>
,
<figureCitation id="5FE82A61FFDCFFB5FAAFFC33FAB1FC58" box="[1331,1362,908,932]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="13" pageNumber="112">33</figureCitation>
); nonporose area on coxopleural process narrow, from less than half length to posterior margin of sternite of ultimate leg-bearing segment to nearly reaching that margin. Ultimate leg prefemur with width at distal end ca 25% its length; prefemoral spines mostly moderately large (
<figureCitation id="5FE82A61FFDCFFB5FD46FC48FCCCFBF3" box="[730,815,1015,1039]" captionStart="FIGURES 36 – 45" captionStartId="12.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 36 – 45. Digitipes jonesii (Verhoeff, 1938). 36 – 40, CES 091062 (male); 41 – 45, CES 091090 (male). 36. Ultimate leg prefemur, ventral view, scale = 1 mm. 37. Ultimate leg femur, dorsal view, scale = 1 mm. 38. Ultimate leg femur, medial view, scale = 1 mm. 39. TT 18 – 19, dorsal view, scale = 1 mm. 40. Spiracle on segment 3, lateral view, scale = 0.5 mm. 41. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 42. Ultimate leg femur, medial view, scale = 0.5 mm. 43. Ultimate leg femur, ventral view, scale = 1 mm. 44. TT 18 – 20, dorsal view, scale = 1 mm. 45. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218632/files/figure.png" pageId="13" pageNumber="112">Fig. 36</figureCitation>
): VL 2 or more commonly 3, VM 1 or usually 2, rarely 0, DM 0 or more commonly 1. Distomedial process in males bluntly conical (
<figureCitation id="5FE82A61FFDCFFB5FB9EFBA3FBB5FBC8" box="[1026,1110,1052,1076]" captionStart="FIGURES 36 – 45" captionStartId="12.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 36 – 45. Digitipes jonesii (Verhoeff, 1938). 36 – 40, CES 091062 (male); 41 – 45, CES 091090 (male). 36. Ultimate leg prefemur, ventral view, scale = 1 mm. 37. Ultimate leg femur, dorsal view, scale = 1 mm. 38. Ultimate leg femur, medial view, scale = 1 mm. 39. TT 18 – 19, dorsal view, scale = 1 mm. 40. Spiracle on segment 3, lateral view, scale = 0.5 mm. 41. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 42. Ultimate leg femur, medial view, scale = 0.5 mm. 43. Ultimate leg femur, ventral view, scale = 1 mm. 44. TT 18 – 20, dorsal view, scale = 1 mm. 45. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218632/files/figure.png" pageId="13" pageNumber="112">Fig. 43</figureCitation>
), extending distinctly further than distal end of the femur on its dorsal side, with a few fine setae on its apex (
<figureCitation id="5FE82A61FFDCFFB5FBA6FB80FB6CFBAB" box="[1082,1167,1087,1111]" captionStart="FIGURES 36 – 45" captionStartId="12.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 36 – 45. Digitipes jonesii (Verhoeff, 1938). 36 – 40, CES 091062 (male); 41 – 45, CES 091090 (male). 36. Ultimate leg prefemur, ventral view, scale = 1 mm. 37. Ultimate leg femur, dorsal view, scale = 1 mm. 38. Ultimate leg femur, medial view, scale = 1 mm. 39. TT 18 – 19, dorsal view, scale = 1 mm. 40. Spiracle on segment 3, lateral view, scale = 0.5 mm. 41. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 42. Ultimate leg femur, medial view, scale = 0.5 mm. 43. Ultimate leg femur, ventral view, scale = 1 mm. 44. TT 18 – 20, dorsal view, scale = 1 mm. 45. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218632/files/figure.png" pageId="13" pageNumber="112">Fig. 37</figureCitation>
); shallow groove along distal 40–65% of femur on its medial surface (
<figureCitation id="5FE82A61FFDCFFB5FD39FBDBFCCAFB80" box="[677,809,1124,1148]" captionStart="FIGURES 36 – 45" captionStartId="12.[151,264,1877,1900]" captionTargetBox="[151,1435,193,1854]" captionTargetId="figure@12.[151,1436,193,1856]" captionTargetPageId="12" captionText="FIGURES 36 – 45. Digitipes jonesii (Verhoeff, 1938). 36 – 40, CES 091062 (male); 41 – 45, CES 091090 (male). 36. Ultimate leg prefemur, ventral view, scale = 1 mm. 37. Ultimate leg femur, dorsal view, scale = 1 mm. 38. Ultimate leg femur, medial view, scale = 1 mm. 39. TT 18 – 19, dorsal view, scale = 1 mm. 40. Spiracle on segment 3, lateral view, scale = 0.5 mm. 41. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 42. Ultimate leg femur, medial view, scale = 0.5 mm. 43. Ultimate leg femur, ventral view, scale = 1 mm. 44. TT 18 – 20, dorsal view, scale = 1 mm. 45. Spiracle on segment 3, lateral view, scale = 0.25 mm." httpUri="https://zenodo.org/record/218632/files/figure.png" pageId="13" pageNumber="112">Figs 38, 42</figureCitation>
). Ultimate leg tarsus 1 1.8–2.5 times length of tarsus 2; tarsus 1 3.3–4 times longer than pretarsus; pretarsus with pair of accessory claws.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFDCFFB6FF55FB14FB8EFD78" lastPageId="14" lastPageNumber="113" pageId="13" pageNumber="112" type="discussion">
<paragraph id="C76C36E4FFDCFFB5FF55FB14FDC0F8F8" blockId="13.[151,1437,151,2012]" pageId="13" pageNumber="112">
<emphasis id="F5A7EAF6FFDCFFB5FF55FB14FEADFB38" bold="true" box="[201,334,1195,1220]" pageId="13" pageNumber="112">Discussion.</emphasis>
Lewis (2010b) noted that the description of
<taxonomicName id="00D34D67FFDCFFB5FCDCFB13FB04FB38" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" box="[832,1255,1196,1220]" class="Chilopoda" family="Scolopendridae" genus="Arthrorhabdus" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDCFFB5FCDCFB13FBD4FB38" box="[832,1079,1196,1220]" italics="true" pageId="13" pageNumber="112">Arthrorhabdus jonesii</emphasis>
Verhoeff, 1938
</taxonomicName>
, described from the Ponmudi Hills, corresponds closely to that of
<taxonomicName id="00D34D67FFDCFFB5FD22FB6FFBAFFB14" authority="Hasse, 1887" authorityName="Hasse" authorityYear="1887" box="[702,1100,1232,1256]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="F5A7EAF6FFDCFFB5FD22FB6FFC59FB14" box="[702,954,1232,1256]" italics="true" pageId="13" pageNumber="112">Otostigmus ceylonicus</emphasis>
Hasse, 1887
</taxonomicName>
, and that this species could be a misplaced
<taxonomicName id="00D34D67FFDCFFB5FEB8FB4AFE40FAF0" box="[292,419,1269,1292]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDCFFB5FEB8FB4AFE40FAF0" box="[292,419,1269,1292]" italics="true" pageId="13" pageNumber="112">Otostigmus</emphasis>
</taxonomicName>
. Jangi and Dass (1984) did not identify any new material of this species, their description being extracted from Verhoeff (1938), and faunal lists by Sureshan
<emphasis id="F5A7EAF6FFDCFFB5FC09FAA6FC24FACC" box="[917,967,1304,1328]" italics="true" pageId="13" pageNumber="112">et al</emphasis>
. (2003, 2006) likewise attributed no new specimens to the species. Collecting at the
<typeStatus id="18688846FFDCFFB5FD1FFA82FD50FAA9" box="[643,691,1341,1365]" pageId="13" pageNumber="112">type</typeStatus>
locality by JJ found only a single species that corresponds to the description of
<taxonomicName id="00D34D67FFDCFFB5FEA1FADFFE4AFA84" box="[317,425,1376,1400]" class="Chilopoda" family="Scolopendridae" genus="Arthrorhabdus" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDCFFB5FEA1FADFFE4AFA84" box="[317,425,1376,1400]" italics="true" pageId="13" pageNumber="112">A. jonesii</emphasis>
</taxonomicName>
,
<taxonomicName id="00D34D67FFDCFFB5FE25FADEFDC3FA84" box="[441,544,1377,1400]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDCFFB5FE25FADEFDC3FA84" box="[441,544,1377,1400]" italics="true" pageId="13" pageNumber="112">Digitipes</emphasis>
</taxonomicName>
PS2 of Joshi and Karanth (2012). Shared details include 17 antennal articles, three of which are glabrous, four teeth on each forcipular tooth plate, legs 1–4 with two tarsal spurs, a coxopleural process with two apical spines and a lateral spine, and three ventrolateral spines on the ultimate leg prefemur. Lewis (2010b) noted that the original assignment of the species to
<taxonomicName id="00D34D67FFDCFFB5FC5FFA73FB8BFA18" box="[963,1128,1484,1508]" class="Chilopoda" family="Scolopendridae" genus="Arthrorhabdus" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDCFFB5FC5FFA73FB8BFA18" box="[963,1128,1484,1508]" italics="true" pageId="13" pageNumber="112">Arthrorhabdus</emphasis>
</taxonomicName>
would predict that it has scolopendrine-type spiracles (i.e., with a three-flapped valve), but the
<typeStatus id="18688846FFDCFFB5FC7EFA4FFBA7F9F4" box="[994,1092,1520,1544]" pageId="13" pageNumber="112" type="holotype">holotype</typeStatus>
in fact has otostigmine-type spiracles, identical to those of
<taxonomicName id="00D34D67FFDCFFB5FE6FF9AAFD50F9D0" box="[499,691,1556,1580]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFDCFFB5FE6FF9AAFD50F9D0" box="[499,691,1556,1580]" italics="true" pageId="13" pageNumber="112">Digitipes indicus</emphasis>
</taxonomicName>
(i.e., strongly elliptical on segment 3, rounded on the subsequent segments, with a humped atrium). The
<typeStatus id="18688846FFDCFFB5FDF4F987FD29F9AC" box="[616,714,1592,1616]" pageId="13" pageNumber="112" type="holotype">holotype</typeStatus>
of
<taxonomicName id="00D34D67FFDCFFB5FD64F986FADEF9AC" authority="Zoologische Staatssammlung, Munich" authorityName="Zoologische Staatssammlung, Munich" box="[760,1341,1592,1616]" class="Chilopoda" family="Scolopendridae" genus="Arthrorhabdus" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDCFFB5FD64F986FC8AF9AC" box="[760,873,1593,1616]" italics="true" pageId="13" pageNumber="112">D. jonesii</emphasis>
(Zoologische Staatssammlung, Munich)
</taxonomicName>
has the conformation of forcipular teeth (inner two teeth grouped, the outer tooth the smallest) and trochanteroprefemural process (two teeth on the inner margin) shared with Indian
<taxonomicName id="00D34D67FFDCFFB5FCD1F93EFC50F964" box="[845,947,1665,1688]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDCFFB5FCD1F93EFC50F964" box="[845,947,1665,1688]" italics="true" pageId="13" pageNumber="112">Digitipes</emphasis>
</taxonomicName>
. The specimen corresponds precisely to a female of
<taxonomicName id="00D34D67FFDCFFB5FE97F91AFE92F940" box="[267,369,1701,1724]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDCFFB5FE97F91AFE92F940" box="[267,369,1701,1724]" italics="true" pageId="13" pageNumber="112">Digitipes</emphasis>
</taxonomicName>
PS2 of Joshi and Karanth (2012), which we cannot separate from
<taxonomicName id="00D34D67FFDCFFB5FBF0F91AFF31F91C" authority="Jangi and Dass, 1984" authorityName="Jangi and Dass" authorityYear="1984" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFDCFFB5FBF0F91AFB02F940" box="[1132,1249,1700,1724]" italics="true" pageId="13" pageNumber="112">D. indicus</emphasis>
Jangi and Dass, 1984
</taxonomicName>
. Accordingly we place
<taxonomicName id="00D34D67FFDCFFB5FE65F976FD97F91C" box="[505,628,1736,1760]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFDCFFB5FE65F976FD97F91C" box="[505,628,1736,1760]" italics="true" pageId="13" pageNumber="112">D. indicus</emphasis>
</taxonomicName>
in subjective synonymy of
<taxonomicName id="00D34D67FFDCFFB5FC4EF976FBA4F91C" box="[978,1095,1737,1760]" class="Chilopoda" family="Scolopendridae" genus="Arthrorhabdus" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDCFFB5FC4EF976FBA4F91C" box="[978,1095,1737,1760]" italics="true" pageId="13" pageNumber="112">D. jonesii</emphasis>
</taxonomicName>
, and the putative record of
<taxonomicName id="00D34D67FFDCFFB5FF0BF953FEDFF8F8" box="[151,316,1772,1796]" class="Chilopoda" family="Scolopendridae" genus="Arthrorhabdus" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDCFFB5FF0BF953FEDFF8F8" box="[151,316,1772,1796]" italics="true" pageId="13" pageNumber="112">Arthrorhabdus</emphasis>
</taxonomicName>
in
<collectingCountry id="BFC47674FFDCFFB5FEC3F953FE7AF8F8" box="[351,409,1772,1796]" name="India" pageId="13" pageNumber="112">India</collectingCountry>
is removed.
</paragraph>
<paragraph id="C76C36E4FFDCFFB6FF5BF8AEFB8EFD78" blockId="13.[151,1437,151,2012]" lastBlockId="14.[151,1436,151,645]" lastPageId="14" lastPageNumber="113" pageId="13" pageNumber="112">
<taxonomicName id="00D34D67FFDCFFB5FF5BF8AEFE65F8D4" box="[199,390,1809,1832]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDCFFB5FF5BF8AEFE65F8D4" box="[199,390,1809,1832]" italics="true" pageId="13" pageNumber="112">Digitipes jonesii</emphasis>
</taxonomicName>
groups putative species 2 and 3 of Joshi and Karanth (2012) because they cannot be distinguished by consistent morphological differences. In the mtDNA phylogeny, these two taxa were resolved as sister species. Both putative species are distinguished from several other Western Ghats
<taxonomicName id="00D34D67FFDCFFB5FB13F8E6FB15F88C" box="[1167,1270,1881,1904]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDCFFB5FB13F8E6FB15F88C" box="[1167,1270,1881,1904]" italics="true" pageId="13" pageNumber="112">Digitipes</emphasis>
</taxonomicName>
by having the basal three antennal articles being glabrous (shared with
<taxonomicName id="00D34D67FFDCFFB5FCB5F8C2FC00F868" box="[809,995,1917,1940]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="112" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="F5A7EAF6FFDCFFB5FCB5F8C2FC00F868" box="[809,995,1917,1940]" italics="true" pageId="13" pageNumber="112">D. coonoorensis</emphasis>
</taxonomicName>
), including individuals with relatively setose legs, and having a long ultimate leg femoral process in the male. Specimens of putative species 2 differ from those of putative species
<quantity id="002B9B01FFDCFFB5FE27F87BFE0AF820" box="[443,489,1988,2012]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="112" unit="in" value="3.0">3 in</quantity>
the frequency of a lateral spine on the coxopleural process, a spine on both sides being the most common state in PS2 (
<figureCitation id="5FE82A61FFDFFFB6FDDBFF28FD2AFF53" box="[583,713,151,175]" captionStart="FIGURES 30 – 35" captionStartId="11.[151,264,1877,1900]" captionTargetBox="[151,1435,232,1854]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURES 30 – 35. Digitipes jonesii (Verhoeff, 1938) (putative species 2 of Joshi and Karanth 2012). 30, 32, 33, CES 07166 (female); 31, 34, 35, CES 091090 (male). 30. Cephalic plate and TT 1 – 2, dorsal view, scale = 0.5 mm. 31. Forcipular segment, ventral view, scale = 1 mm. 32. Tooth plates, ventral view, scale = 0.5 mm. 33. Coxopleuron, lateral view, scale = 1 mm. 34. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 35. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 1 mm." httpUri="https://zenodo.org/record/218631/files/figure.png" pageId="14" pageNumber="113">Figs 33, 35</figureCitation>
) and shared by the
<typeStatus id="18688846FFDFFFB6FC35FF28FBE8FF53" box="[937,1035,151,175]" pageId="14" pageNumber="113" type="holotype">holotype</typeStatus>
of
<taxonomicName id="00D34D67FFDFFFB6FBAFFF26FB42FF4C" box="[1075,1185,153,176]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="jonesii">
<emphasis id="F5A7EAF6FFDFFFB6FBAFFF26FB42FF4C" box="[1075,1185,153,176]" italics="true" pageId="14" pageNumber="113">D. jonesii</emphasis>
</taxonomicName>
, but rare in PS3 (
<figureCitation id="5FE82A61FFDFFFB6FAF6FF28FF00FF28" captionStart="FIGURES 24 – 29" captionStartId="10.[151,264,1636,1659]" captionTargetBox="[151,1435,187,1624]" captionTargetId="figure@10.[151,1436,187,1627]" captionTargetPageId="10" captionText="FIGURES 24 – 29. Digitipes jonesii (Verhoeff, 1938) (putative species 3 of Joshi and Karanth 2012). CES 091062 (male) except Fig. 27, CES 091047 (female). 24. Cephalic plate and T 1, dorsal view, scale = 1 mm. 25. Forcipular segment, ventral view, scale = 1 mm. 26. Tooth plates, ventral view, scale = 0.5 mm. 27. Coxopleuron, lateral view, scale = 1 mm. 28. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 29. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218630/files/figure.png" pageId="14" pageNumber="113">Figs 27, 29</figureCitation>
). Both putative species include specimens with a spine on one side only, and while complete absence of lateral spines is the most frequent state in PS3 (as in the
<typeStatus id="18688846FFDFFFB6FCBEFF60FC67FF0B" box="[802,900,223,247]" pageId="14" pageNumber="113" type="holotype">holotype</typeStatus>
of
<taxonomicName id="00D34D67FFDFFFB6FC31FF5EFBC1FF04" box="[941,1058,224,248]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="F5A7EAF6FFDFFFB6FC31FF5EFBC1FF04" box="[941,1058,224,248]" italics="true" pageId="14" pageNumber="113">D. indicus</emphasis>
</taxonomicName>
), PS2 also includes several large specimens that lack lateral spines.
<collectingCountry id="BFC47674FFDFFFB6FD81FEBBFDDFFEE0" box="[541,572,260,284]" name="American Samoa" pageId="14" pageNumber="113">As</collectingCountry>
such, diagnostic value cannot be ascribed to this character, which appears to vary at the population level. Specimens of PS2 have a more consistent formula of the ultimate leg prefemoral spines (VL 3, VM 2, DM 1), but within the variability exhibited by putative species 3 are individuals that correspond exactly to the (more stable) state of putative species 2. The relatively pronounced setae on the legs are unique to this clade when compared to other Indian
<taxonomicName id="00D34D67FFDFFFB6FD46FE2AFCA2FE50" box="[730,833,405,428]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDFFFB6FD46FE2AFCA2FE50" box="[730,833,405,428]" italics="true" pageId="14" pageNumber="113">Digitipes</emphasis>
</taxonomicName>
. Within PS2, one of the two main clades identified by molecular phylogenetics (Joshi and Karanth 2012) has setae on the legs comparable to those more consistently shown by PS3, whereas the other clade within PS2 has less setose legs. The two clades within PS2 occur in different mountain ranges, geographically close to each other. This feature thus displays some geographic variation between populations of what appears to represent a single species, and prohibits making a clear-cut distinction between PS2 and PS3 on the basis of leg setation. The two putative species are sympatric in their distributions in some parts of their ranges, largely in the southernmost hill ranges of the Western Ghats.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="9BAEB8F6E1CDCC7632AFCA9B6D0AD19F">Revision of the scolopendrid centipede Digitipes Attems, 1930, from India (Chilopoda: Scolopendromorpha): reconciling molecular and morphological estimates of species diversity</mods:title>
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<mods:namePart id="96CDA01E09BAF7FF1DB948880C18002A">Edgecombe, Gregory D.</mods:namePart>
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<mods:typeOfResource id="D937D7C0478419A5BDFF804E169D7681">text</mods:typeOfResource>
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<mods:title id="1922D169E574DE5C0489920D54470C65">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="ACE775EDC9F2C0FB63DD96166817D73B">
<mods:date id="910DF21D961D043C91F63050BFA6334C">2013</mods:date>
<mods:detail id="8DB32807FD151E6CECBD0868E6117B10" type="volume">
<mods:number id="B745A1FA44C34356C69273FBB0D8A22F">3626</mods:number>
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<mods:start id="5D880AB318E3A40CCB8777C7EF39EB75">99</mods:start>
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<mods:identifier id="F17E800C9B46ED6F290C69240DFEFCA5" type="DOI">10.11646/zootaxa.3626.1.5</mods:identifier>
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<treatment id="4F7A87F2FFDFFFA8FF0BFD68FC2CF815" ID-DOI="http://doi.org/10.5281/zenodo.6154120" ID-GBIF-Taxon="119563406" ID-Zenodo-Dep="6154120" LSID="urn:lsid:plazi:treatment:4F7A87F2FFDFFFA8FF0BFD68FC2CF815" httpUri="http://treatment.plazi.org/id/4F7A87F2FFDFFFA8FF0BFD68FC2CF815" lastPageId="16" lastPageNumber="115" pageId="14" pageNumber="113">
<subSubSection id="8FC9656FFFDFFFB6FF0BFD68FCD3FC8B" pageId="14" pageNumber="113" type="nomenclature">
<paragraph id="C76C36E4FFDFFFB6FF0BFD68FE7AFD0D" blockId="14.[151,409,727,786]" box="[151,409,727,753]" pageId="14" pageNumber="113">
<heading id="9C248188FFDFFFB6FF0BFD68FE7AFD0D" bold="true" box="[151,409,727,753]" fontSize="11" level="1" pageId="14" pageNumber="113" reason="1">
<emphasis id="F5A7EAF6FFDFFFB6FF0BFD68FE7AFD0D" bold="true" box="[151,409,727,753]" pageId="14" pageNumber="113">
<taxonomicName id="00D34D67FFDFFFB6FF0BFD68FEB3FD0D" box="[151,336,727,753]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="species" species="jangii" status="sp. nov.">
<emphasis id="F5A7EAF6FFDFFFB6FF0BFD68FEB3FD0D" bold="true" box="[151,336,727,753]" italics="true" pageId="14" pageNumber="113">Digitipes jangii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94578DFFDFFFB6FECBFD68FE7AFD0D" box="[343,409,727,753]" pageId="14" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C76C36E4FFDFFFB6FF0BFD45FEC4FCEE" blockId="14.[151,409,727,786]" box="[151,295,762,786]" pageId="14" pageNumber="113">
(
<figureCitation id="5FE82A61FFDFFFB6FF03FD45FEFCFCEE" box="[159,287,762,786]" captionStart-0="FIGURES 46 – 51" captionStart-1="FIGURES 52 – 56" captionStartId-0="15.[151,264,1870,1893]" captionStartId-1="16.[151,264,1116,1139]" captionTargetBox-0="[151,1435,337,1847]" captionTargetBox-1="[165,1408,185,1098]" captionTargetId-0="figure@15.[151,1436,337,1849]" captionTargetId-1="figure@16.[165,1409,185,1099]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURES 46 – 51. Digitipes jangii n. sp. 46, 50, 51, CES 08912 (holotype male); 47 – 49, CES 08915 (paratype male). 46. Cephalic plate and T 1, dorsal view, scale = 2 mm. 47. Forcipular segment, ventral view, scale = 1 mm. 48. Tooth plates, ventral view, scale = 0.5 mm. 49. Coxopleuron, lateral view, scale = 1 mm. 50. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 51. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." captionText-1="FIGURES 52 – 56. Digitipes jangii n. sp. CES 08912 (holotype male) except Fig. 56, CES 08915 (paratype male). 52. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 53. Ultimate leg femur, dorsal view, scale = 0.5 mm. 54. Ultimate leg femur, medial view, scale = 0.5 mm. 55. TT 18 – 20, dorsal view, scale = 0.5 mm. 56. Spiracle on segment 3, scale = 0.25 mm." httpUri-0="https://zenodo.org/record/218633/files/figure.png" httpUri-1="https://zenodo.org/record/218634/files/figure.png" pageId="14" pageNumber="113">Figs 46–56</figureCitation>
)
</paragraph>
<paragraph id="C76C36E4FFDFFFB6FF0BFCFCFCD3FC8B" blockId="14.[151,816,834,887]" pageId="14" pageNumber="113">
<taxonomicName id="00D34D67FFDFFFB6FF0BFCFCFF17FCA4" box="[151,244,835,856]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDFFFB6FF0BFCFCFF17FCA4" box="[151,244,835,856]" italics="true" pageId="14" pageNumber="113">Digitipes</emphasis>
</taxonomicName>
putative species 5. Joshi and Karanth, 2012: figs 2, 3, 6.
<taxonomicName id="00D34D67FFDFFFB6FF0BFCDDFF17FC8B" box="[151,244,866,887]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDFFFB6FF0BFCDDFF17FC8B" box="[151,244,866,887]" italics="true" pageId="14" pageNumber="113">Digitipes</emphasis>
</taxonomicName>
putative species 6. Joshi and Karanth, 2012: figs 2, 3, 6.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFDFFFB6FF0BFC1BFC46FB93" pageId="14" pageNumber="113" type="materials_examined">
<paragraph id="C76C36E4FFDFFFB6FF0BFC1BFC46FB93" blockId="14.[151,1436,931,2036]" pageId="14" pageNumber="113">
<emphasis id="F5A7EAF6FFDFFFB6FF0BFC1BFEBDFC40" bold="true" box="[151,350,931,956]" pageId="14" pageNumber="113">
<typeStatus id="18688846FFDFFFB6FF0BFC1BFF31FC40" box="[151,210,932,956]" pageId="14" pageNumber="113">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="18688846FFDFFFB6FEF6FC1BFE31FC40" box="[362,466,932,956]" pageId="14" pageNumber="113" type="holotype">Holotype</typeStatus>
CES08912, male, from Kudremukh National Park, Karnataka,
<collectingCountry id="BFC47674FFDFFFB6FB5EFC1BFB1DFC40" box="[1218,1278,932,956]" name="India" pageId="14" pageNumber="113">India</collectingCountry>
, 13°20052 N 75°19289 E, leg. J. Joshi,
<date id="B36D1024FFDFFFB6FE45FC78FDDAFC23" box="[473,569,967,991]" pageId="14" pageNumber="113" value="2008-07">vii.2008</date>
.
<typeStatus id="18688846FFDFFFB6FDD5FC77FD59FC1C" box="[585,698,968,992]" pageId="14" pageNumber="113" type="paratype">Paratypes</typeStatus>
: CES08915, male, CES08907, CES08922, females, from
<typeStatus id="18688846FFDFFFB6FAF0FC77FA7FFC1C" box="[1388,1436,968,992]" pageId="14" pageNumber="113">type</typeStatus>
locality, same collection; CES07219, CES07223, CES07226, CES07230, from Anshi-Dandeli Tiger Reserve, Karnataka; CES07230, from Dodamanne Ghat, Karnataka; CES08288, from Talacauvery Reserve Forest, Karnataka; CES08930, from
<collectingCountry id="BFC47674FFDFFFB6FE6CFB8BFDACFBB0" box="[496,591,1076,1100]" name="India" pageId="14" pageNumber="113">Bombay</collectingCountry>
point, Mahabaleshwar Reserve Forest, Maharashtra; CES
<date id="B36D1024FFDFFFB6FB61FB8BFAB7FBB0" box="[1277,1364,1076,1100]" pageId="14" pageNumber="113" value="20-09-10">091020</date>
, from Bisale Ghat Reserve Forest, Karanataka. All leg. J. Joshi,
<date id="B36D1024FFDFFFB6FC83FBE8FC42FB93" box="[799,929,1111,1135]" pageId="14" pageNumber="113" value="2007" valueMax="2010">2007–2010</date>
.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFDFFFB6FF55FBC3FE5CFB4B" pageId="14" pageNumber="113" type="etymology">
<paragraph id="C76C36E4FFDFFFB6FF55FBC3FE5CFB4B" blockId="14.[151,1436,931,2036]" pageId="14" pageNumber="113">
<emphasis id="F5A7EAF6FFDFFFB6FF55FBC3FEB3FB68" bold="true" box="[201,336,1148,1172]" pageId="14" pageNumber="113">Etymology.</emphasis>
For B. S. Jangi, in recognition of his work on the
<taxonomicName id="00D34D67FFDFFFB6FC4FFBC3FB62FB68" box="[979,1153,1148,1172]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="family">Scolopendridae</taxonomicName>
of
<collectingCountry id="BFC47674FFDFFFB6FB28FBC3FB13FB68" box="[1204,1264,1148,1172]" name="India" pageId="14" pageNumber="113">India</collectingCountry>
, including the identification of
<taxonomicName id="00D34D67FFDFFFB6FECDFB1EFE5BFB44" box="[337,440,1185,1208]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDFFFB6FECDFB1EFE5BFB44" box="[337,440,1185,1208]" italics="true" pageId="14" pageNumber="113">Digitipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFDFFFB6FF55FB7CFEFDFA90" pageId="14" pageNumber="113" type="diagnosis">
<paragraph id="C76C36E4FFDFFFB6FF55FB7CFEFDFA90" blockId="14.[151,1436,931,2036]" pageId="14" pageNumber="113">
<emphasis id="F5A7EAF6FFDFFFB6FF55FB7CFEA0FB20" bold="true" box="[201,323,1219,1244]" pageId="14" pageNumber="113">Diagnosis.</emphasis>
<taxonomicName id="00D34D67FFDFFFB6FED6FB7AFE52FB20" box="[330,433,1221,1244]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFDFFFB6FED6FB7AFE52FB20" box="[330,433,1221,1244]" italics="true" pageId="14" pageNumber="113">Digitipes</emphasis>
</taxonomicName>
with basal three or four antennal articles glabrous both dorsally and ventrally. Forcipular tooth plates wider than long. Tergite paramedian sutures complete from TT4 or 5; tergites smooth, lacking longitudinal median ridge or keels. Coxopleural process long, slender, lateral spine usually present. Leg 20 without tarsal spur. Spines on ultimate leg prefemur moderate to strong. Distomedial process on ultimate leg femur of male short, blunt.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFDFFFA8FF55FAC8FAD4F97D" lastPageId="16" lastPageNumber="115" pageId="14" pageNumber="113" type="description">
<paragraph id="C76C36E4FFDFFFB6FF55FAC8FC59FA24" blockId="14.[151,1436,931,2036]" pageId="14" pageNumber="113">
<emphasis id="F5A7EAF6FFDFFFB6FF55FAC8FEB8FA6C" bold="true" box="[201,347,1399,1424]" pageId="14" pageNumber="113">Description.</emphasis>
Length to
<quantity id="002B9B01FFDFFFB6FE42FAC8FDD0FA6C" box="[478,563,1399,1424]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="14" pageNumber="113" unit="mm" value="47.0">47 mm</quantity>
. 17 antennal articles; basal three (five individuals) or four (six individuals) articles glabrous both dorsally and ventrally. Longitudinal median furrow on anterior 20% of cephalic plate. Short paired longitudinal furrows on posterior part of cephalic plate (
<figureCitation id="5FE82A61FFDFFFB6FCC5FA7FFC4EFA24" box="[857,941,1472,1496]" captionStart="FIGURES 46 – 51" captionStartId="15.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1847]" captionTargetId="figure@15.[151,1436,337,1849]" captionTargetPageId="15" captionText="FIGURES 46 – 51. Digitipes jangii n. sp. 46, 50, 51, CES 08912 (holotype male); 47 – 49, CES 08915 (paratype male). 46. Cephalic plate and T 1, dorsal view, scale = 2 mm. 47. Forcipular segment, ventral view, scale = 1 mm. 48. Tooth plates, ventral view, scale = 0.5 mm. 49. Coxopleuron, lateral view, scale = 1 mm. 50. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 51. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218633/files/figure.png" pageId="14" pageNumber="113">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="C76C36E4FFDFFFB6FF55FA5BFE60F994" blockId="14.[151,1436,931,2036]" pageId="14" pageNumber="113">Anterior part of cephalic plate and basal (glabrous) part of antenna blue, remainder of cephalic plate usually brown; T1 variably light brown or mostly blue; tergites often mostly blue from T2 to middle of body and increasingly brown in posterior segments but may be dark brown from T2; legs usually light blue, in some specimens dark blue.</paragraph>
<paragraph id="C76C36E4FFDFFFB6FF55F9CBFF0EF904" blockId="14.[151,1436,931,2036]" pageId="14" pageNumber="113">
Forcipular coxosternal tooth plates wider than long (
<figureCitation id="5FE82A61FFDFFFB6FCB6F9CBFC51F970" box="[810,946,1652,1676]" captionStart="FIGURES 46 – 51" captionStartId="15.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1847]" captionTargetId="figure@15.[151,1436,337,1849]" captionTargetPageId="15" captionText="FIGURES 46 – 51. Digitipes jangii n. sp. 46, 50, 51, CES 08912 (holotype male); 47 – 49, CES 08915 (paratype male). 46. Cephalic plate and T 1, dorsal view, scale = 2 mm. 47. Forcipular segment, ventral view, scale = 1 mm. 48. Tooth plates, ventral view, scale = 0.5 mm. 49. Coxopleuron, lateral view, scale = 1 mm. 50. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 51. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218633/files/figure.png" pageId="14" pageNumber="113">Figs 47, 48</figureCitation>
), with four main teeth, inner two grouped together, the outer tooth smaller than the inner three; base of tooth plates defined by relatively obtuse oblique sutures diverging at 140–145°. Trochanteroprefemoral process with two distinctly defined teeth along inner margin.
</paragraph>
<paragraph id="C76C36E4FFDFFFB6FF55F8BBFCF4F8E0" blockId="14.[151,1436,931,2036]" box="[201,791,1796,1821]" pageId="14" pageNumber="113">Second maxillary claw with slender accessory spurs.</paragraph>
<paragraph id="C76C36E4FFDFFFB6FF55F897FDA7F874" blockId="14.[151,1436,931,2036]" pageId="14" pageNumber="113">
Tergites with paramedian sutures complete from TT4 or (usually) 5. Tergites marginate starting from 7 to 13, most commonly from 8 or 9. Tergites smooth (
<figureCitation id="5FE82A61FFDFFFB6FD03F8F3FD11F898" box="[671,754,1868,1892]" captionStart="FIGURES 52 – 56" captionStartId="16.[151,264,1116,1139]" captionTargetBox="[165,1408,185,1098]" captionTargetId="figure@16.[165,1409,185,1099]" captionTargetPageId="16" captionText="FIGURES 52 – 56. Digitipes jangii n. sp. CES 08912 (holotype male) except Fig. 56, CES 08915 (paratype male). 52. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 53. Ultimate leg femur, dorsal view, scale = 0.5 mm. 54. Ultimate leg femur, medial view, scale = 0.5 mm. 55. TT 18 – 20, dorsal view, scale = 0.5 mm. 56. Spiracle on segment 3, scale = 0.25 mm." httpUri="https://zenodo.org/record/218634/files/figure.png" pageId="14" pageNumber="113">Fig. 55</figureCitation>
), lacking median ridge or keels. Paramedian sutures 20–35% length of sternites in mid body region.
</paragraph>
<paragraph id="C76C36E4FFDFFFB6FF55F82BFE1AF808" blockId="14.[151,1436,931,2036]" pageId="14" pageNumber="113">First three or four pairs of legs (two pairs in a single specimen) with two tarsal spurs, the subsequent up to 19 with one, 20 without tarsal spur (except one individual). Tibial spur on leg 1 only. Femoral spur usually lacking on leg 1, present in one individual.</paragraph>
<paragraph id="C76C36E4FFDEFFB7FF55FF28FF20FEE0" blockId="15.[151,1436,151,284]" pageId="15" pageNumber="114">
Tergite of ultimate leg-bearing segment with gently convex lateral margins that converge posteriorly; posterolateral margins straight, variably pointed posteromedially (
<figureCitation id="5FE82A61FFDEFFB7FC31FF03FBEAFF28" box="[941,1033,188,212]" captionStart="FIGURES 46 – 51" captionStartId="15.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1847]" captionTargetId="figure@15.[151,1436,337,1849]" captionTargetPageId="15" captionText="FIGURES 46 – 51. Digitipes jangii n. sp. 46, 50, 51, CES 08912 (holotype male); 47 – 49, CES 08915 (paratype male). 46. Cephalic plate and T 1, dorsal view, scale = 2 mm. 47. Forcipular segment, ventral view, scale = 1 mm. 48. Tooth plates, ventral view, scale = 0.5 mm. 49. Coxopleuron, lateral view, scale = 1 mm. 50. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 51. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218633/files/figure.png" pageId="15" pageNumber="114">Fig. 50</figureCitation>
). Sternite of ultimate leg-bearing segment with sides converging posteriorly, weakly convex outwards, posterior margin moderately concave (
<figureCitation id="5FE82A61FFDEFFB7FAF3FF60FF55FEE0" captionStart="FIGURES 46 – 51" captionStartId="15.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1847]" captionTargetId="figure@15.[151,1436,337,1849]" captionTargetPageId="15" captionText="FIGURES 46 – 51. Digitipes jangii n. sp. 46, 50, 51, CES 08912 (holotype male); 47 – 49, CES 08915 (paratype male). 46. Cephalic plate and T 1, dorsal view, scale = 2 mm. 47. Forcipular segment, ventral view, scale = 1 mm. 48. Tooth plates, ventral view, scale = 0.5 mm. 49. Coxopleuron, lateral view, scale = 1 mm. 50. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 51. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218633/files/figure.png" pageId="15" pageNumber="114">Fig. 51</figureCitation>
).
</paragraph>
<caption id="93AC666CFFDEFFB7FF0BF8F1FB31F83C" httpUri="https://zenodo.org/record/218633/files/figure.png" pageId="15" pageNumber="114" targetBox="[151,1435,337,1847]" targetPageId="15">
<paragraph id="C76C36E4FFDEFFB7FF0BF8F1FB31F83C" blockId="15.[151,1437,1870,1984]" pageId="15" pageNumber="114">
<emphasis id="F5A7EAF6FFDEFFB7FF0BF8F1FEC8F89F" bold="true" box="[151,299,1870,1893]" pageId="15" pageNumber="114">FIGURES 46</emphasis>
–
<emphasis id="F5A7EAF6FFDEFFB7FEABF8F1FEB4F89F" bold="true" box="[311,343,1870,1891]" pageId="15" pageNumber="114">51.</emphasis>
<taxonomicName id="00D34D67FFDEFFB7FEFDF8F0FDE0F898" box="[353,515,1871,1892]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="15" pageNumber="114" phylum="Arthropoda" rank="species" species="jangii" status="sp. nov.">
<emphasis id="F5A7EAF6FFDEFFB7FEFDF8F0FDE0F898" box="[353,515,1871,1892]" italics="true" pageId="15" pageNumber="114">Digitipes jangii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94578DFFDEFFB7FD92F8F0FDA8F898" box="[526,587,1871,1892]" pageId="15" pageNumber="114" rank="species">
<emphasis id="F5A7EAF6FFDEFFB7FD92F8F0FDA8F898" bold="true" box="[526,587,1871,1892]" pageId="15" pageNumber="114">n. sp.</emphasis>
</taxonomicNameLabel>
46, 50, 51, CES08912 (holotype male); 47–49, CES08915 (paratype male). 46. Cephalic plate and T1, dorsal view, scale = 2 mm. 47. Forcipular segment, ventral view, scale = 1 mm. 48. Tooth plates, ventral view, scale = 0.5 mm. 49. Coxopleuron, lateral view, scale = 1 mm. 50. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 51. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm.
</paragraph>
</caption>
<caption id="93AC666CFFC1FFA8FF0BFBE3FAA4FB4C" httpUri="https://zenodo.org/record/218634/files/figure.png" pageId="16" pageNumber="115" targetBox="[165,1408,185,1098]" targetPageId="16">
<paragraph id="C76C36E4FFC1FFA8FF0BFBE3FAA4FB4C" blockId="16.[151,1437,1116,1200]" pageId="16" pageNumber="115">
<emphasis id="F5A7EAF6FFC1FFA8FF0BFBE3FEC4FB8E" bold="true" box="[151,295,1116,1139]" pageId="16" pageNumber="115">FIGURES 52</emphasis>
–
<emphasis id="F5A7EAF6FFC1FFA8FEA8FBE2FEB7FB8E" bold="true" box="[308,340,1117,1138]" pageId="16" pageNumber="115">56.</emphasis>
<taxonomicName id="00D34D67FFC1FFA8FEC7FBE1FE1AFB8F" box="[347,505,1118,1139]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="115" phylum="Arthropoda" rank="species" species="jangii" status="sp. nov.">
<emphasis id="F5A7EAF6FFC1FFA8FEC7FBE1FE1AFB8F" box="[347,505,1118,1139]" italics="true" pageId="16" pageNumber="115">Digitipes jangii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE94578DFFC1FFA8FD9CFBE2FDD9FB8E" box="[512,570,1117,1138]" pageId="16" pageNumber="115" rank="species">
<emphasis id="F5A7EAF6FFC1FFA8FD9CFBE2FDD9FB8E" bold="true" box="[512,570,1117,1138]" pageId="16" pageNumber="115">n. sp.</emphasis>
</taxonomicNameLabel>
CES08912 (holotype male) except Fig. 56, CES08915 (paratype male). 52. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 53. Ultimate leg femur, dorsal view, scale = 0.5 mm. 54. Ultimate leg femur, medial view, scale = 0.5 mm. 55. TT18–20, dorsal view, scale = 0.5 mm. 56. Spiracle on segment 3, scale = 0.25 mm.
</paragraph>
</caption>
<paragraph id="C76C36E4FFC1FFA8FF55FB62FAD4F97D" blockId="16.[151,1436,1245,2025]" pageId="16" pageNumber="115">
Coxopleural process long, slender, with two apical spines, mostly with a lateral spine on each side (
<figureCitation id="5FE82A61FFC1FFA8FAA2FB62FF55FAE5" captionStart="FIGURES 46 – 51" captionStartId="15.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1847]" captionTargetId="figure@15.[151,1436,337,1849]" captionTargetPageId="15" captionText="FIGURES 46 – 51. Digitipes jangii n. sp. 46, 50, 51, CES 08912 (holotype male); 47 – 49, CES 08915 (paratype male). 46. Cephalic plate and T 1, dorsal view, scale = 2 mm. 47. Forcipular segment, ventral view, scale = 1 mm. 48. Tooth plates, ventral view, scale = 0.5 mm. 49. Coxopleuron, lateral view, scale = 1 mm. 50. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 51. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218633/files/figure.png" pageId="16" pageNumber="115">Figs 49, 51</figureCitation>
), a few individuals with lateral spine on one side only, one lacking spines on both sides. Pores relatively dense, approximately bimodal mix of large and small pores; pore field extending close to but not reaching dorsal margin of coxopleuron (
<figureCitation id="5FE82A61FFC1FFA8FECEFAF6FE44FA9D" box="[338,423,1353,1377]" captionStart="FIGURES 46 – 51" captionStartId="15.[151,264,1870,1893]" captionTargetBox="[151,1435,337,1847]" captionTargetId="figure@15.[151,1436,337,1849]" captionTargetPageId="15" captionText="FIGURES 46 – 51. Digitipes jangii n. sp. 46, 50, 51, CES 08912 (holotype male); 47 – 49, CES 08915 (paratype male). 46. Cephalic plate and T 1, dorsal view, scale = 2 mm. 47. Forcipular segment, ventral view, scale = 1 mm. 48. Tooth plates, ventral view, scale = 0.5 mm. 49. Coxopleuron, lateral view, scale = 1 mm. 50. Tergite of ultimate leg-bearing segment, dorsal view, scale = 1 mm. 51. Coxopleuron and sternite of ultimate leg-bearing segment, ventral view, scale = 0.5 mm." httpUri="https://zenodo.org/record/218633/files/figure.png" pageId="16" pageNumber="115">Fig. 49</figureCitation>
); non-porose area on process narrow, at least half length to posterior margin of sternite of ultimate leg-bearing segment, sometimes extending to that margin. Ultimate leg prefemur with width at distal end about one-third its length; prefemoral spines usually large, distally curved (
<figureCitation id="5FE82A61FFC1FFA8FC7CFA2EFBD7FA55" box="[992,1076,1425,1449]" captionStart="FIGURES 52 – 56" captionStartId="16.[151,264,1116,1139]" captionTargetBox="[165,1408,185,1098]" captionTargetId="figure@16.[165,1409,185,1099]" captionTargetPageId="16" captionText="FIGURES 52 – 56. Digitipes jangii n. sp. CES 08912 (holotype male) except Fig. 56, CES 08915 (paratype male). 52. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 53. Ultimate leg femur, dorsal view, scale = 0.5 mm. 54. Ultimate leg femur, medial view, scale = 0.5 mm. 55. TT 18 – 20, dorsal view, scale = 0.5 mm. 56. Spiracle on segment 3, scale = 0.25 mm." httpUri="https://zenodo.org/record/218634/files/figure.png" pageId="16" pageNumber="115">Fig. 52</figureCitation>
): VL 3, VM 2, DM 1 or 2, apart from one individual with VL 1/2, VM 1, DM 0. Distomedial process in males blunt, not extending to distal end of femur on its dorsal side (
<figureCitation id="5FE82A61FFC1FFA8FE31FA66FDE2FA0D" box="[429,513,1497,1521]" captionStart="FIGURES 52 – 56" captionStartId="16.[151,264,1116,1139]" captionTargetBox="[165,1408,185,1098]" captionTargetId="figure@16.[165,1409,185,1099]" captionTargetPageId="16" captionText="FIGURES 52 – 56. Digitipes jangii n. sp. CES 08912 (holotype male) except Fig. 56, CES 08915 (paratype male). 52. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 53. Ultimate leg femur, dorsal view, scale = 0.5 mm. 54. Ultimate leg femur, medial view, scale = 0.5 mm. 55. TT 18 – 20, dorsal view, scale = 0.5 mm. 56. Spiracle on segment 3, scale = 0.25 mm." httpUri="https://zenodo.org/record/218634/files/figure.png" pageId="16" pageNumber="115">Fig. 53</figureCitation>
); groove on medial surface of femur confined to distal end of femur or as much as half length of femur (
<figureCitation id="5FE82A61FFC1FFA8FE1BFA42FE38F9E9" box="[391,475,1533,1557]" captionStart="FIGURES 52 – 56" captionStartId="16.[151,264,1116,1139]" captionTargetBox="[165,1408,185,1098]" captionTargetId="figure@16.[165,1409,185,1099]" captionTargetPageId="16" captionText="FIGURES 52 – 56. Digitipes jangii n. sp. CES 08912 (holotype male) except Fig. 56, CES 08915 (paratype male). 52. Ultimate leg prefemur, ventral view, scale = 0.5 mm. 53. Ultimate leg femur, dorsal view, scale = 0.5 mm. 54. Ultimate leg femur, medial view, scale = 0.5 mm. 55. TT 18 – 20, dorsal view, scale = 0.5 mm. 56. Spiracle on segment 3, scale = 0.25 mm." httpUri="https://zenodo.org/record/218634/files/figure.png" pageId="16" pageNumber="115">Fig. 54</figureCitation>
); slightly flattened and variably depigmented area on medial surface of femur bearing dense field of fine pores. Distomedial end of ultimate leg tibia in male either bears (CES08912) or lacks (CES08915, CES
<date id="B36D1024FFC1FFA8FEC1F9FAFE57F9A1" box="[349,436,1605,1629]" pageId="16" pageNumber="115" value="20-09-10">091020</date>
) a weak rounded process similar to that on femur. Ultimate leg tarsus 1 1.6–1.8 times length of tarsus 2; tarsus 1 2.4–6 times longer than pretarsus; pretarsus with pair of short accessory claws.
</paragraph>
</subSubSection>
<subSubSection id="8FC9656FFFC1FFA8FF55F932FC2CF815" pageId="16" pageNumber="115" type="discussion">
<paragraph id="C76C36E4FFC1FFA8FF55F932FC2CF815" blockId="16.[151,1436,1245,2025]" pageId="16" pageNumber="115">
<emphasis id="F5A7EAF6FFC1FFA8FF55F932FEADF95A" bold="true" box="[201,334,1677,1702]" pageId="16" pageNumber="115">Discussion.</emphasis>
Putative species 5 and 6 had previously been observed not to be differentiated on the environmental and morphological axis (Joshi and Karanth 2012). In the current revision we treat them as one species but recognise it as distinct from all previously named congeners. Its distinctive features include the presence of four glabrous antennal articles in some specimens, relatively short forcipular tooth plates, a slender coxopleural process, robust prefemoral spines, and a short femoral process on the male ultimate leg. It is also the only species in our sample apart from
<taxonomicName id="00D34D67FFC1FFA8FDC2F8FDFCEDF8A5" box="[606,782,1858,1881]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="115" phylum="Arthropoda" rank="species" species="periyarensis">
<emphasis id="F5A7EAF6FFC1FFA8FDC2F8FDFCEDF8A5" box="[606,782,1858,1881]" italics="true" pageId="16" pageNumber="115">D. periyarensis</emphasis>
</taxonomicName>
with individuals (4 of
<specimenCount id="D1D5FD6DFFC1FFA8FBBDF8FEFB5CF8A5" box="[1057,1215,1857,1881]" pageId="16" pageNumber="115" type="generic">11 specimens</specimenCount>
, all from the same population) having two DM spines on the ultimate leg prefemur; other Indian
<taxonomicName id="00D34D67FFC1FFA8FBB7F8D9FB71F881" box="[1067,1170,1894,1917]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="F5A7EAF6FFC1FFA8FBB7F8D9FB71F881" box="[1067,1170,1894,1917]" italics="true" pageId="16" pageNumber="115">Digitipes</emphasis>
</taxonomicName>
have at most one DM spine. A character of potential taxonomic importance is a small distomedial process on the ultimate leg tibia in the sole mature male of PS5 (the
<typeStatus id="18688846FFC1FFA8FE43F812FDA2F839" box="[479,577,1965,1989]" pageId="16" pageNumber="115" type="holotype">holotype</typeStatus>
of
<taxonomicName id="00D34D67FFC1FFA8FDF9F811FD2BF839" box="[613,712,1966,1989]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="115" phylum="Arthropoda" rank="species" species="jangii">
<emphasis id="F5A7EAF6FFC1FFA8FDF9F811FD2BF839" box="[613,712,1966,1989]" italics="true" pageId="16" pageNumber="115">D. jangii</emphasis>
</taxonomicName>
) versus its absence in two mature males of PS6. We are reluctant to attribute diagnostic value to this difference without additional samples.
</paragraph>
</subSubSection>
</treatment>
</document>