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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1072.72978" ID-GBIF-Dataset="5abb2e98-44e8-4554-9e15-fa892088595d" ID-PMC="PMC8629907" ID-Pensoft-Pub="1313-2970-1072-129" ID-Pensoft-UUID="9D8AA58A90535BAAB59216914F04E621" ID-PubMed="34899010" ModsDocID="1313-2970-1072-129" checkinTime="1637640646787" checkinUser="pensoft" docAuthor="Ng, Peter K. L. &amp; Yang, Chien-Hui" docDate="2021" docId="4F563B84BA4F5460A96A9606FBC33594" docLanguage="en" docName="ZooKeys 1072: 129-165" docOrigin="ZooKeys 1072" docPubDate="2021-11-22" docSource="http://dx.doi.org/10.3897/zookeys.1072.72978" docTitle="Dicranodromia erinaceus Ng &amp; Yang 2021, sp. nov." docType="treatment" docVersion="4" id="9D8AA58A90535BAAB59216914F04E621" lastPageNumber="129" masterDocId="9D8AA58A90535BAAB59216914F04E621" masterDocTitle="On two new species of deep-sea carrier crabs (Crustacea, Brachyura, Homolodromiidae, Dicranodromia) from Taiwan and the Philippines, with notes on other Indo-West Pacific species" masterLastPageNumber="165" masterPageNumber="129" pageNumber="129" updateTime="1668151158991" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>On two new species of deep-sea carrier crabs (Crustacea, Brachyura, Homolodromiidae, Dicranodromia) from Taiwan and the Philippines, with notes on other Indo-West Pacific species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Peter K. L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5946-0608</mods:nameIdentifier>
<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Chien-Hui</mods:namePart>
<mods:affiliation>National Taiwan Ocean University, 2 Pei-Ning Road, Keelung 20224, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">chyang@ntou.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-11-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1072</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>165</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1072.72978</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1072.72978</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1072-129</mods:identifier>
<mods:identifier type="Pensoft-UUID">9D8AA58A90535BAAB59216914F04E621</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="190549119" LSID="urn:lsid:zoobank.org:act:C1BEA5CC-F350-47D1-8A66-ECBFC3606197" httpUri="http://treatment.plazi.org/id/4F563B84BA4F5460A96A9606FBC33594" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection pageId="0" pageNumber="129" type="nomenclature">
<paragraph pageId="0" pageNumber="129">
<taxonomicName LSID="http://zoobank.org/C1BEA5CC-F350-47D1-8A66-ECBFC3606197" authority="Ng &amp; Yang, 2021" authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus" status="sp. nov.">Dicranodromia erinaceus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="129">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Colour in life. A, B Dicranodromia karubar Guinot, 1993, ♂ (28.7 x 34.7 mm) (ZRC 2020.0348), Java C D. karubar Guinot, 1993, ovigerous ♀ (30.1 x 35.5 mm) (ZRC 2020.0349), Java D D. karubar Guinot, 1993, ovigerous ♀ (24.8 x 31.5 mm) (ZRC 2020.0348), Java E D. erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan F D. erinaceus sp. nov., paratype ♂ (6.9 x 9.5 mm) (ZRC 2021.0084), Taiwan. Photographs: T. - Y. Chan." figureDoi="10.3897/zookeys.1072.72978.figure7" httpUri="https://binary.pensoft.net/fig/612913" pageId="0" pageNumber="129">Figures 7E, F</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A overall view B dorsal view of carapace C front and anterior part of carapace D left orbit and first anterolateral spine." figureDoi="10.3897/zookeys.1072.72978.figure13" httpUri="https://binary.pensoft.net/fig/612919" pageId="0" pageNumber="129">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of right chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure14" httpUri="https://binary.pensoft.net/fig/612920" pageId="0" pageNumber="129">, 14</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Dicranodromia erinaceus sp. nov. A-E holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan F paratype ♀ (13.8 x 18.5 mm) (ZRC 2020.0467), Taiwan A right P 3 B right P 5 C right P 5 propodus and dactylus D right P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca and female gonopores." figureDoi="10.3897/zookeys.1072.72978.figure15" httpUri="https://binary.pensoft.net/fig/612921" pageId="0" pageNumber="129">, 15</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Dicranodromia erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure16" httpUri="https://binary.pensoft.net/fig/612922" pageId="0" pageNumber="129">, 16</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan B, D-F, K-M D. erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan C, G-J, N-P D. robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A-C right anterior part of carapace (setae removed or not drawn) D right P 3 propodus and dactylus E right P 5 propodus and dactylus F right P 4 propodus and dactylus G right P 3 propodus and dactylus H right P 5 propodus and dactylus I left P 5 propodus and dactylus J left P 4 propodus and dactylus K, N left G 1 (ventral view) L, O left G 1 (dorsal view) M, P left G 2. Setae for all structures not figured. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129">, 21A, B, D-F, K-M</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="reference_group">
<paragraph pageId="0" pageNumber="129">
<taxonomicName authority="Ng &amp; Yang, 2021" authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia doederleini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="doederleini" status="sp. nov.">Dicranodromia doederleini</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.1163/1568540041958617" author="Ho, P-H" journalOrPublisher="Crustaceana" pageId="0" pageNumber="129" pagination="641 - 668" refId="B15" refString="Ho, P-H, Ng, PKL, Chan, T-Y, Lee, D-A, 2004. New records of 31 species of brachyuran crabs from the joint Taiwan-France Expeditions, &quot;TAIWAN 2000&quot; and &quot;TAIWAN 2001&quot;, off deep waters in Taiwan. Crustaceana 77 (6): 641 - 668, DOI: https://doi.org/10.1163/1568540041958617" title="New records of 31 species of brachyuran crabs from the joint Taiwan-France Expeditions, &quot; TAIWAN 2000 &quot; and &quot; TAIWAN 2001 &quot;, off deep waters in Taiwan." url="https://doi.org/10.1163/1568540041958617" volume="77" year="2004">Ho et al. 2004</bibRefCitation>
: 643, fig. 1B;
<bibRefCitation author="Ahyong, ST" editor="Chan, T-Y" journalOrPublisher="Keelung, Taiwan" pageId="0" pageNumber="129" pagination="27 - 180" refId="B2" refString="Ahyong, ST, Naruse, T, Tan, SH, Ng, PKL, 2009. Part II. Infraorder Brachyura: Sections Dromiacea, Raninoida, Cyclodorippoida. In: Chan, T-Y, Ng, PKL, Ahyong, ST, Tan, SH, Eds., Crustacean Fauna of Taiwan: Brachyuran Crabs, Volume I. Keelung, Taiwan: 27 - 180" title="Part II. Infraorder Brachyura: Sections Dromiacea, Raninoida, Cyclodorippoida." volumeTitle="Crustacean Fauna of Taiwan: Brachyuran Crabs, Volume I." year="2009">Ahyong et al. 2009</bibRefCitation>
: 129, fig. 93 (not fig. 94);
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="129" refId="B30" refString="Ng, PKL, Shih, H-T, Ho, P-H, Wang, C-H, 2017. An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum 70(3 &amp; 4): 1-185." title="An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum 70 (3 &amp; 4): 1 - 185." year="2017">Ng et al. 2017</bibRefCitation>
: 27 (list). (non
<taxonomicName authority="Ng &amp; Yang, 2021" authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia doederleini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="doederleini" status="sp. nov.">Dicranodromia doederleini</taxonomicName>
Ortmann, 1892)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="materials_examined">
<paragraph pageId="0" pageNumber="129">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="129">
<materialsCitation accessionNumber="B00126" collectingDate="2013-05-27" collectorName="N. O. Ocean Researcher" country="TAIWAN" elevation="984" latitude="22.233334" location="Liu-Qiu Island" longLatPrecision="1260" longitude="119.98333" municipality="Taiwan" specimenCount="♀" typeStatus="holotype">
<collectingCountry name="Taiwan">TAIWAN</collectingCountry>
:
<typeStatus>holotype</typeStatus>
<specimenCount>♀</specimenCount>
(14.0
<normalizedToken originalValue="×">x</normalizedToken>
18.0 mm), station CP4091,
<geoCoordinate degrees="22" direction="north" minutes="14" orientation="latitude" precision="925" value="22.233334">22°14'N</geoCoordinate>
,
<geoCoordinate degrees="119" direction="east" minutes="59" orientation="longitude" precision="925" value="119.98333">119°59'E</geoCoordinate>
, among numerous mud tubes, off small
<location LSID="urn:lsid:plazi:treatment:4F563B84BA4F5460A96A9606FBC33594:CB8FC0140FAD0FF30E536AB810D8C64F" country="TAIWAN" latitude="22.233334" longLatPrecision="1260" longitude="119.98333" municipality="Taiwan" name="Liu-Qiu Island">Liu-Qiu Island</location>
, southeast
<collectingMunicipality>Taiwan</collectingMunicipality>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.84" metricValueMax="9.94" metricValueMin="9.74" unit="m" value="984.0" valueMax="994.0" valueMin="974.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.84" metricValueMax="9.94" metricValueMin="9.74" unit="m" value="984.0" valueMax="994.0" valueMin="974.0">974-994 m</elevation>
</quantity>
, coll.
<collectorName>N.O. Ocean Researcher</collectorName>
1,
<collectingDate value="2013-05-27">27 May 2013</collectingDate>
(NTOU
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/B00126">B00126</accessionNumber>
)
</materialsCitation>
;
<materialsCitation accessionNumber="OK351335" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="N. O. Ocean Researcher" country="Taiwan" elevation="984" latitude="22.233334" location="Liu-Qiu Island" longLatPrecision="1260" longitude="119.98333" municipality="Taiwan" specimenCount="2" specimenCount-female="2" typeStatus="paratypes">
<typeStatus>paratypes</typeStatus>
<specimenCount type="female">2 ♀♀</specimenCount>
(13.2
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.76" unit="mm" value="17.6">17.6 mm</quantity>
, 13.8
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.85" unit="mm" value="18.5">18.5 mm</quantity>
), same data as holotype (ZRC 2020.0467, COI sequence:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OK351335">OK351335</accessionNumber>
)
</materialsCitation>
;
<materialsCitation accessionNumber="OK331334" collectingDate="2020-11-15" collectorName="T. - Y. Chan" country="Taiwan" elevation="985" latitude="22.305666" location="southwestern Taiwan" longLatPrecision="12" longitude="119.99184" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
(6.9
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.5" unit="mm" value="9.5">9.5 mm</quantity>
), station CP4212,
<geoCoordinate degrees="22" direction="north" minutes="18.34" orientation="latitude" precision="9" value="22.305666">22°18.34'N</geoCoordinate>
,
<geoCoordinate degrees="119" direction="east" minutes="59.51" orientation="longitude" precision="9" value="119.99184">119°59.51'E</geoCoordinate>
,
<location LSID="urn:lsid:plazi:treatment:4F563B84BA4F5460A96A9606FBC33594:C98FC0FEE1FED55D4BEF033F30827753" country="Taiwan" latitude="22.305666" longLatPrecision="12" longitude="119.99184" name="southwestern Taiwan">southwestern Taiwan</location>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.845" metricValueMax="10.08" metricValueMin="9.61" unit="m" value="984.5" valueMax="1008.0" valueMin="961.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.845" metricValueMax="10.08" metricValueMin="9.61" unit="m" value="984.5" valueMax="1008.0" valueMin="961.0">961-1008 m</elevation>
</quantity>
, coll.
<collectorName>T.-Y. Chan</collectorName>
,
<collectingDate value="2020-11-15">15 Nov. 2020</collectingDate>
(ZRC 2021.0084, COI sequence:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OK331334">OK331334</accessionNumber>
)
</materialsCitation>
;
<materialsCitation collectingDate="2020-11-15" collectorName="T. - Y. Chan" country="Taiwan" elevation="985" latitude="22.305666" location="southwestern Taiwan" longLatPrecision="12" longitude="119.99184" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
(8.2
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.25" unit="mm" value="12.5">12.5 mm</quantity>
), station CP4212,
<geoCoordinate degrees="22" direction="north" minutes="18.34" orientation="latitude" precision="9" value="22.305666">22°18.34'N</geoCoordinate>
,
<geoCoordinate degrees="119" direction="east" minutes="59.51" orientation="longitude" precision="9" value="119.99184">119°59.51'E</geoCoordinate>
,
<location LSID="urn:lsid:plazi:treatment:4F563B84BA4F5460A96A9606FBC33594:7F216E1F0B74D7DB0D5D490DFE55297E" country="Taiwan" latitude="22.305666" longLatPrecision="12" longitude="119.99184" name="southwestern Taiwan">southwestern Taiwan</location>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.845" metricValueMax="10.08" metricValueMin="9.61" unit="m" value="984.5" valueMax="1008.0" valueMin="961.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.845" metricValueMax="10.08" metricValueMin="9.61" unit="m" value="984.5" valueMax="1008.0" valueMin="961.0">961-1008 m</elevation>
</quantity>
, coll.
<collectorName>T.-Y. Chan</collectorName>
,
<collectingDate value="2020-11-15">15 Nov. 2020</collectingDate>
(ZRC 2021.0085)
</materialsCitation>
.
<materialsCitation accessionNumber="B00127" collectingDate="2000-08-04" country="Taiwan" elevation="731" latitude="24.448334" location="Others" longLatPrecision="125" longitude="122.30167" municipality="Taiwan" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:4F563B84BA4F5460A96A9606FBC33594:5294A940F992DC150FB1156AE60D68A6" country="Taiwan" latitude="24.448334" longLatPrecision="125" longitude="122.30167" municipality="Taiwan" name="Others">Others</location>
:
<specimenCount type="female">1 ♀</specimenCount>
(7.5
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" unit="mm" value="11.1">11.1 mm</quantity>
, carapace badly damaged),
<geoCoordinate degrees="24" direction="north" minutes="26.9" orientation="latitude" precision="92" value="24.448334">24°26.9'N</geoCoordinate>
,
<geoCoordinate degrees="122" direction="east" minutes="18.1" orientation="longitude" precision="92" value="122.30167">122°18.1'E</geoCoordinate>
,
<collectingMunicipality>Taiwan</collectingMunicipality>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.31" metricValueMax="8.24" metricValueMin="6.38" unit="m" value="731.0" valueMax="824.0" valueMin="638.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.31" metricValueMax="8.24" metricValueMin="6.38" unit="m" value="731.0" valueMax="824.0" valueMin="638.0">638-824 m</elevation>
</quantity>
, coll. R/V &quot;Fishery Researcher 1&quot;,
<collectingDate value="2000-08-04">4 August 2000</collectingDate>
(NTOU
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/B00127">B00127</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="129">Carapace longitudinally subovate, widest across intestinal-mesobranchial regions; dorsal surface prominently convex, lateral surfaces covered with numerous spinules, those on median part relatively lower, sometimes granular, with short stiff setae, denser on lateral parts but not obscuring margins; short stiff setae present on pereiopods, thoracic sternum and pleon but not obscuring surface or margins. Branchiocardiac groove distinct, curving medially anteriorly. Each pseudorostral lobe triangular, inner margin straight, outer margin gently convex, directed anteriorly, inner margin with two or three spinules; exorbital tooth spiniform, directed obliquely laterally, anterior margin with two or three spinules; supraorbital margin separated from external orbital tooth by shallow concave cleft, posterior part with three spines; infraorbital margin with prominent triangular lobe, posterior margin with spinules, just visible in dorsal view. Rostrum present as one or two longer spinules in small specimens, barely discernible or just visible as a sharp granule in larger specimens. Epistome covered with spinules on anterior half; posterior half gently upturned, with median fissure, surface not covered with spinules, posterior margin entire. Basal antennal article subquadrate; surfaces covered by spinules and granules; anteroexternal tooth short. Eyes with short peduncle. Third maxilliped relatively narrow; merus subovate with low anterointernal lobe, slightly shorter than ischium; ischium subtrapezoidal, distal half wider than proximal part with inner margin convex; palp (carpus, propodus, dactylus) long, reaching to median part of ischium when folded; exopod with proximal third widest, outer margin with low sharp granules on proximal third. Chelipeds densely covered with stiff setae on most parts; merus and carpus with outer surface and margins lined with spinules and granules; palm relatively short, outer surface and margins covered with numerous sharp granules; fingers thick, wide, occluding surface hollowed; pollex with deep U-shaped depression distally. P2 and P3 relatively long, P3 longer than P2; merus with low tooth on distal extensor margin, length to width ratio of P2 and P3 merus 5.2 and 4.5, respectively; proximal part of extensor margin with low spinules, flexor margin with numerous spinules; propodus almost straight, unarmed, length to width ratio of P2 and P3 propodus 6.7 and 8.0, respectively; dactylus sickle-shaped, flexor margin lined with 15 or 16 spines, terminating in strongly incurved claw, propodus about twice length of dactylus. P4 stouter, shorter than P5; length to width ratio of P4 and P5 merus 3.5 and 5.0, respectively; proximal part of extensor margin of merus with low spinules, flexor margin with numerous spinules; P4 and P5 propodus without median spinules on outer surface, length to width ratio of P4 and P5 propodus 3.5 and 4.7, respectively, distal margin fringed by sharp spines bracketing dactylus; dactylus claw-like, strongly incurved, extensor margin unarmed, flexor margin unarmed or with two weak spines. Thoracic sternite 7 with strong transverse ridge from posterior inner part of female gonopore, lateral part raised, forming triangular tubercle, curving posteriorly to join oblique ridge formed by sternites 7 and 8 with distinct groove between them that leads to spermathecal aperture at centre of triangular tubercle. Male and female pleons with six free somites and telson; male telson distinctly subovate; female telson wide, triangular, with gently sinuous margins. G1 stout, endopod distally covered by dense long setae, subdistal part of outer margin with two lobes, proximal lobe larger, prominent; G2 endopod gradually tapering to sharp tip.</paragraph>
<caption doi="10.3897/zookeys.1072.72978.figure13" httpUri="https://binary.pensoft.net/fig/612919" pageId="0" pageNumber="129" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 13.</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia erinaceus</emphasis>
</taxonomicName>
sp. nov., holotype ♀ (14.0
<normalizedToken originalValue="×">x</normalizedToken>
18.0 mm) (NTOU B00126), Taiwan
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
overall view
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
dorsal view of carapace
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
front and anterior part of carapace
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
left orbit and first anterolateral spine.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1072.72978.figure14" httpUri="https://binary.pensoft.net/fig/612920" pageId="0" pageNumber="129" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 14.</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia erinaceus</emphasis>
</taxonomicName>
sp. nov., holotype ♀ (14.0
<normalizedToken originalValue="×">x</normalizedToken>
18.0 mm) (NTOU B00126), Taiwan
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
ventral view of cephalothorax
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
left third maxilliped
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
frontal view of cephalothorax
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
epistome, antennules, antennae and orbits
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
dorsal view of right chela
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
left chela.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1072.72978.figure15" httpUri="https://binary.pensoft.net/fig/612921" pageId="0" pageNumber="129" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 15.</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia erinaceus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="129">A-E</emphasis>
holotype ♀ (14.0
<normalizedToken originalValue="×">x</normalizedToken>
18.0 mm) (NTOU B00126), Taiwan
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
paratype ♀ (13.8
<normalizedToken originalValue="×">x</normalizedToken>
18.5 mm) (ZRC 2020.0467), Taiwan
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
right P3
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
right P5
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
right P5 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
right P4 propodus and dactylus
<emphasis bold="true" pageId="0" pageNumber="129">E</emphasis>
anterior thoracic sternum and spermatheca
<emphasis bold="true" pageId="0" pageNumber="129">F</emphasis>
posterior thoracic sternum showing spermatheca and female gonopores.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1072.72978.figure16" httpUri="https://binary.pensoft.net/fig/612922" pageId="0" pageNumber="129" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Figure 16.</emphasis>
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia erinaceus</emphasis>
</taxonomicName>
sp. nov., paratype ♂ (8.2
<normalizedToken originalValue="×">x</normalizedToken>
12.5 mm) (ZRC 2021.0085), Taiwan
<emphasis bold="true" pageId="0" pageNumber="129">A</emphasis>
overall view
<emphasis bold="true" pageId="0" pageNumber="129">B</emphasis>
dorsal view of carapace
<emphasis bold="true" pageId="0" pageNumber="129">C</emphasis>
pleon
<emphasis bold="true" pageId="0" pageNumber="129">D</emphasis>
anterior thoracic sternum and G1s in situ.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="variation">
<paragraph pageId="0" pageNumber="129">Variation.</paragraph>
<paragraph pageId="0" pageNumber="129">
None of the specimens examined had a spine or spinule on the extensor margin of the P5 dactylus; and outer surface of the P5 propodus was also unarmed (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan B, D-F, K-M D. erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan C, G-J, N-P D. robusta sp. nov., paratype ♂ (15.2 x 21.0 mm) (ZRC 2018.0095), Philippines A-C right anterior part of carapace (setae removed or not drawn) D right P 3 propodus and dactylus E right P 5 propodus and dactylus F right P 4 propodus and dactylus G right P 3 propodus and dactylus H right P 5 propodus and dactylus I left P 5 propodus and dactylus J left P 4 propodus and dactylus K, N left G 1 (ventral view) L, O left G 1 (dorsal view) M, P left G 2. Setae for all structures not figured. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1072.72978.figure21" httpUri="https://binary.pensoft.net/fig/612927" pageId="0" pageNumber="129">21E</figureCitation>
). Most of the flexor margins of the dactylus were not armed with obvious spines or spinules, although two or three stout setae may be present.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="etymology">
<paragraph pageId="0" pageNumber="129">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="129">
The species is named after the hedgehog,
<taxonomicName authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Erinaceidae" genus="Erinaceus" kingdom="Animalia" lsidName="Erinaceus" order="Erinaceomorpha" pageId="0" pageNumber="129" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Erinaceus</emphasis>
</taxonomicName>
, alluding to the spiny appearance of the carapace and legs. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="remarks">
<paragraph pageId="0" pageNumber="129">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia erinaceus</emphasis>
</taxonomicName>
sp. nov. belongs to the same group of species as
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. spinulosa" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis italics="true" pageId="0" pageNumber="129">D. spinulosa</emphasis>
</taxonomicName>
and
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. delli" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="delli">
<emphasis italics="true" pageId="0" pageNumber="129">D. delli</emphasis>
</taxonomicName>
in its spinose carapace surface and pereiopods, slender and spiniform exorbital tooth, and an acutely triangular suborbital tooth.
<taxonomicName authorityName="Ng &amp; Yang" authorityYear="2021" class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia erinaceus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. delli" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="delli">
<emphasis italics="true" pageId="0" pageNumber="129">D. delli</emphasis>
</taxonomicName>
from New Zealand but can be distinguished by the ischium of the third maxilliped being relatively shorter and wider especially at the distal half (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of right chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure14" httpUri="https://binary.pensoft.net/fig/612920" pageId="0" pageNumber="129">14B</figureCitation>
) (versus more slender and rectangular in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. delli" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="delli">
<emphasis italics="true" pageId="0" pageNumber="129">D. delli</emphasis>
</taxonomicName>
, cf.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1708.1.1" author="Ahyong, ST" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 72" refId="B1" refString="Ahyong, ST, 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72, DOI: https://doi.org/10.11646/zootaxa.1708.1.1" title="Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1708.1.1" volume="1708" year="2008">Ahyong 2008</bibRefCitation>
: fig. 4C); proportionately shorter P2 and P3 (e.g., P3 merus 4.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, propodus 8.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A overall view B dorsal view of carapace C front and anterior part of carapace D left orbit and first anterolateral spine." figureDoi="10.3897/zookeys.1072.72978.figure13" httpUri="https://binary.pensoft.net/fig/612919" pageId="0" pageNumber="129">13A</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Dicranodromia erinaceus sp. nov. A-E holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan F paratype ♀ (13.8 x 18.5 mm) (ZRC 2020.0467), Taiwan A right P 3 B right P 5 C right P 5 propodus and dactylus D right P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca and female gonopores." figureDoi="10.3897/zookeys.1072.72978.figure15" httpUri="https://binary.pensoft.net/fig/612921" pageId="0" pageNumber="129">15A</figureCitation>
) (versus P3 merus 6.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, propodus 11.1
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. delli" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="delli">
<emphasis italics="true" pageId="0" pageNumber="129">D. delli</emphasis>
</taxonomicName>
, cf.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1708.1.1" author="Ahyong, ST" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 72" refId="B1" refString="Ahyong, ST, 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72, DOI: https://doi.org/10.11646/zootaxa.1708.1.1" title="Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1708.1.1" volume="1708" year="2008">Ahyong 2008</bibRefCitation>
: fig. 2A, 3D); the proportionately shorter and stouter P4 and P5 (e.g., P5 merus just reaches the branchiocardiac groove in dorsal position, Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A overall view B dorsal view of carapace C front and anterior part of carapace D left orbit and first anterolateral spine." figureDoi="10.3897/zookeys.1072.72978.figure13" httpUri="https://binary.pensoft.net/fig/612919" pageId="0" pageNumber="129">13A</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Dicranodromia erinaceus sp. nov. A-E holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan F paratype ♀ (13.8 x 18.5 mm) (ZRC 2020.0467), Taiwan A right P 3 B right P 5 C right P 5 propodus and dactylus D right P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca and female gonopores." figureDoi="10.3897/zookeys.1072.72978.figure15" httpUri="https://binary.pensoft.net/fig/612921" pageId="0" pageNumber="129">15B</figureCitation>
) (versus longer and more slender, extending beyond branchiocardiac groove in dorsal position in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. delli" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="delli">
<emphasis italics="true" pageId="0" pageNumber="129">D. delli</emphasis>
</taxonomicName>
, cf.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1708.1.1" author="Ahyong, ST" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 72" refId="B1" refString="Ahyong, ST, 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72, DOI: https://doi.org/10.11646/zootaxa.1708.1.1" title="Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1708.1.1" volume="1708" year="2008">Ahyong 2008</bibRefCitation>
: fig. 2A, B); the relatively stouter palm (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of right chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure14" httpUri="https://binary.pensoft.net/fig/612920" pageId="0" pageNumber="129">14F</figureCitation>
) (versus more slender in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. delli" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="delli">
<emphasis italics="true" pageId="0" pageNumber="129">D. delli</emphasis>
</taxonomicName>
, cf.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1708.1.1" author="Ahyong, ST" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 72" refId="B1" refString="Ahyong, ST, 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72, DOI: https://doi.org/10.11646/zootaxa.1708.1.1" title="Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1708.1.1" volume="1708" year="2008">Ahyong 2008</bibRefCitation>
: fig. 3B); and the proportionately wider female telson (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of right chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure14" httpUri="https://binary.pensoft.net/fig/612920" pageId="0" pageNumber="129">14A</figureCitation>
) (versus less wide in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. delli" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="delli">
<emphasis italics="true" pageId="0" pageNumber="129">D. delli</emphasis>
</taxonomicName>
, cf.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1708.1.1" author="Ahyong, ST" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 72" refId="B1" refString="Ahyong, ST, 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72, DOI: https://doi.org/10.11646/zootaxa.1708.1.1" title="Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1708.1.1" volume="1708" year="2008">Ahyong 2008</bibRefCitation>
: fig. 3C). The holotype and only known specimen of
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. delli" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="delli">
<emphasis italics="true" pageId="0" pageNumber="129">D. delli</emphasis>
</taxonomicName>
, an ovigerous female 15.5
<normalizedToken originalValue="×">x</normalizedToken>
19.0 mm from Nukuliau Seamount in New Zealand, is comparable in size to the ovigerous female holotype of
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">D. erinaceus</emphasis>
</taxonomicName>
(14.0
<normalizedToken originalValue="×">x</normalizedToken>
18.0 mm) so the differences are not size-related. The characters of P2-P5 and third maxilliped are also obvious in the smaller female paratype of
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">D. erinaceus</emphasis>
</taxonomicName>
as well as in the smaller male paratypes.
</paragraph>
<paragraph pageId="0" pageNumber="129">
Compared to
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. spinulosa" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis italics="true" pageId="0" pageNumber="129">D. spinulosa</emphasis>
</taxonomicName>
,
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">D. erinaceus</emphasis>
</taxonomicName>
can be separated by the carapace being proportionately wider (Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A overall view B dorsal view of carapace C front and anterior part of carapace D left orbit and first anterolateral spine." figureDoi="10.3897/zookeys.1072.72978.figure13" httpUri="https://binary.pensoft.net/fig/612919" pageId="0" pageNumber="129">13B</figureCitation>
,
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Dicranodromia erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure16" httpUri="https://binary.pensoft.net/fig/612922" pageId="0" pageNumber="129">16B</figureCitation>
) (versus carapace transversely narrower in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. spinulosa" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis italics="true" pageId="0" pageNumber="129">D. spinulosa</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 21a;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1708.1.1" author="Ahyong, ST" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 72" refId="B1" refString="Ahyong, ST, 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72, DOI: https://doi.org/10.11646/zootaxa.1708.1.1" title="Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1708.1.1" volume="1708" year="2008">Ahyong 2008</bibRefCitation>
: fig. 1C); the median dorsal surface of the carapace covered with low sharp granules (Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A overall view B dorsal view of carapace C front and anterior part of carapace D left orbit and first anterolateral spine." figureDoi="10.3897/zookeys.1072.72978.figure13" httpUri="https://binary.pensoft.net/fig/612919" pageId="0" pageNumber="129">13B</figureCitation>
,
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Dicranodromia erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure16" httpUri="https://binary.pensoft.net/fig/612922" pageId="0" pageNumber="129">16B</figureCitation>
) (versus covered with spinules in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. spinulosa" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis italics="true" pageId="0" pageNumber="129">D. spinulosa</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 21a;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1708.1.1" author="Ahyong, ST" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 72" refId="B1" refString="Ahyong, ST, 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72, DOI: https://doi.org/10.11646/zootaxa.1708.1.1" title="Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1708.1.1" volume="1708" year="2008">Ahyong 2008</bibRefCitation>
: fig. 1C); the submarginal surface of the posterior margin of the epistome is unarmed (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of right chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure14" httpUri="https://binary.pensoft.net/fig/612920" pageId="0" pageNumber="129">14C, D</figureCitation>
) (versus area armed with short spines in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. spinulosa" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis italics="true" pageId="0" pageNumber="129">D. spinulosa</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 22B); the ischium of the third maxilliped relatively shorter and wider especially at the distal half (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A ventral view of cephalothorax B left third maxilliped C frontal view of cephalothorax D epistome, antennules, antennae and orbits E dorsal view of right chela F left chela." figureDoi="10.3897/zookeys.1072.72978.figure14" httpUri="https://binary.pensoft.net/fig/612920" pageId="0" pageNumber="129">14B</figureCitation>
) (versus more slender and rectangular in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. spinulata" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulata">
<emphasis italics="true" pageId="0" pageNumber="129">D. spinulata</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 21c); the relatively longer P2 and P3 (e.g., P3 propodus 8.0
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dicranodromia erinaceus sp. nov., holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan A overall view B dorsal view of carapace C front and anterior part of carapace D left orbit and first anterolateral spine." figureDoi="10.3897/zookeys.1072.72978.figure13" httpUri="https://binary.pensoft.net/fig/612919" pageId="0" pageNumber="129">13A</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Dicranodromia erinaceus sp. nov. A-E holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan F paratype ♀ (13.8 x 18.5 mm) (ZRC 2020.0467), Taiwan A right P 3 B right P 5 C right P 5 propodus and dactylus D right P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca and female gonopores." figureDoi="10.3897/zookeys.1072.72978.figure15" httpUri="https://binary.pensoft.net/fig/612921" pageId="0" pageNumber="129">15A</figureCitation>
) (versus P3 propodus less than 7
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide), the P2 and P3 propodus twice the length of the dactylus (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Dicranodromia erinaceus sp. nov. A-E holotype ♀ (14.0 x 18.0 mm) (NTOU B 00126), Taiwan F paratype ♀ (13.8 x 18.5 mm) (ZRC 2020.0467), Taiwan A right P 3 B right P 5 C right P 5 propodus and dactylus D right P 4 propodus and dactylus E anterior thoracic sternum and spermatheca F posterior thoracic sternum showing spermatheca and female gonopores." figureDoi="10.3897/zookeys.1072.72978.figure15" httpUri="https://binary.pensoft.net/fig/612921" pageId="0" pageNumber="129">15A</figureCitation>
) (versus 1.7
<normalizedToken originalValue="×">x</normalizedToken>
in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. spinulosa" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis italics="true" pageId="0" pageNumber="129">D. spinulosa</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: fig. 21a;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1708.1.1" author="Ahyong, ST" journalOrPublisher="Zootaxa" pageId="0" pageNumber="129" pagination="1 - 72" refId="B1" refString="Ahyong, ST, 2008. Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura). Zootaxa 1708: 1 - 72, DOI: https://doi.org/10.11646/zootaxa.1708.1.1" title="Deepwater crabs from seamounts and chemosynthetic habitats off eastern New Zealand (Crustacea: Decapoda: Brachyura)." url="https://doi.org/10.11646/zootaxa.1708.1.1" volume="1708" year="2008">Ahyong 2008</bibRefCitation>
: fig. 1C); and the male telson subovate in shape (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Dicranodromia erinaceus sp. nov., paratype ♂ (8.2 x 12.5 mm) (ZRC 2021.0085), Taiwan A overall view B dorsal view of carapace C pleon D anterior thoracic sternum and G 1 s in situ." figureDoi="10.3897/zookeys.1072.72978.figure16" httpUri="https://binary.pensoft.net/fig/612922" pageId="0" pageNumber="129">16C</figureCitation>
) (versus triangular in
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. spinulosa" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis italics="true" pageId="0" pageNumber="129">D. spinulosa</emphasis>
</taxonomicName>
, cf.
<bibRefCitation author="Guinot, D" editor="Crosnier, A" journalOrPublisher="Volume 13, Memoires du Museum national d'Histoire naturelle 163" pageId="0" pageNumber="129" pagination="155 - 282" refId="B12" refString="Guinot, D, 1995. Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900. In: Crosnier, A, Ed., Resultats des Campagnes MUSORSTOM. Volume 13, Memoires du Museum national d'Histoire naturelle 163: 155 - 282" title="Crustacea Decapoda Brachyura: Revision de la famille des Homolodromiidae Alcock, 1900." volumeTitle="Resultats des Campagnes MUSORSTOM." year="1995">Guinot 1995</bibRefCitation>
: figs 21c, 25D).
<taxonomicName class="Malacostraca" family="Homolodromiidae" genus="Dicranodromia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicranodromia spinulosa" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis italics="true" pageId="0" pageNumber="129">Dicranodromia spinulosa</emphasis>
</taxonomicName>
was described from three males and one female from New Caledonia, the holotype female being 7.5
<normalizedToken originalValue="×">x</normalizedToken>
11.0 mm; a size comparable to that of the male specimens of
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">D. erinaceus</emphasis>
</taxonomicName>
we examined.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<bibRefCitation DOI="https://doi.org/10.1163/1568540041958617" author="Ho, P-H" journalOrPublisher="Crustaceana" pageId="0" pageNumber="129" pagination="641 - 668" refId="B15" refString="Ho, P-H, Ng, PKL, Chan, T-Y, Lee, D-A, 2004. New records of 31 species of brachyuran crabs from the joint Taiwan-France Expeditions, &quot;TAIWAN 2000&quot; and &quot;TAIWAN 2001&quot;, off deep waters in Taiwan. Crustaceana 77 (6): 641 - 668, DOI: https://doi.org/10.1163/1568540041958617" title="New records of 31 species of brachyuran crabs from the joint Taiwan-France Expeditions, &quot; TAIWAN 2000 &quot; and &quot; TAIWAN 2001 &quot;, off deep waters in Taiwan." url="https://doi.org/10.1163/1568540041958617" volume="77" year="2004">Ho et al. (2004)</bibRefCitation>
recorded
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. doederleini" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="doederleini">
<emphasis italics="true" pageId="0" pageNumber="129">D. doederleini</emphasis>
</taxonomicName>
from Taiwan from one badly damaged female specimen from northeastern Taiwan (see also
<bibRefCitation author="Ahyong, ST" editor="Chan, T-Y" journalOrPublisher="Keelung, Taiwan" pageId="0" pageNumber="129" pagination="27 - 180" refId="B2" refString="Ahyong, ST, Naruse, T, Tan, SH, Ng, PKL, 2009. Part II. Infraorder Brachyura: Sections Dromiacea, Raninoida, Cyclodorippoida. In: Chan, T-Y, Ng, PKL, Ahyong, ST, Tan, SH, Eds., Crustacean Fauna of Taiwan: Brachyuran Crabs, Volume I. Keelung, Taiwan: 27 - 180" title="Part II. Infraorder Brachyura: Sections Dromiacea, Raninoida, Cyclodorippoida." volumeTitle="Crustacean Fauna of Taiwan: Brachyuran Crabs, Volume I." year="2009">Ahyong et al. 2009</bibRefCitation>
). The specimen is now referred to
<taxonomicName class="Malacostraca" kingdom="Animalia" lsidName="D. erinaceus" order="Decapoda" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="erinaceus">
<emphasis italics="true" pageId="0" pageNumber="129">D. erinaceus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

50
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Northwest territories, Canada (
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).
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<subSubSection pageId="566" pageNumber="567" type="reference_group">
<paragraph pageId="566" pageNumber="567">
<taxonomicName authorityName="LeConte" authorityYear="1855" class="Hexapoda" family="Carabidae" genus="Amara" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="566" pageNumber="567" phylum="Arthropoda" rank="species" species="jacobina">Amara jacobina</taxonomicName>
LeConte, 1855: 346. Type locality: &quot;San Diego [San Diego County], California&quot; (original citation). Holotype [by monotypy; designated lectotype by Hieke (1990: 285)] (♂) in MCZ [# 5665].
</paragraph>
<paragraph pageId="566" pageNumber="567">
<taxonomicName authorityName="LeConte" authorityYear="1855" class="Hexapoda" family="Carabidae" genus="Amara" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="566" pageNumber="567" phylum="Arthropoda" rank="species" species="stupida">Amara stupida</taxonomicName>
LeConte, 1855: 347. Type locality: &quot;Sacramento [Sacramento County], California&quot; (original citation). Holotype [by monotypy; designated lectotype by Hieke (1990: 286)] (♀) in MCZ [# 5666]. Synonymy established by Hieke (1990: 283).
</paragraph>
</subSubSection>
<subSubSection pageId="566" pageNumber="567" type="distribution">
<paragraph pageId="566" pageNumber="567">Distribution.</paragraph>
<paragraph pageId="566" pageNumber="567">
The range of this species extends from
<normalizedToken originalValue="“Washington”">&quot;Washington&quot;</normalizedToken>
(Hayward 1908: 23) to the Baja California Peninsula (Horn 1894: 309) and Arizona (Griffith 1900: 565; Hayward 1908: 23). The records from north-central Utah (Horn 1894: 309),
<normalizedToken originalValue="“Colorado”">&quot;Colorado&quot;</normalizedToken>
(Wickham 1902: 236), and New Mexico (Fall and Cockerell 1907: 159) need confirmation.
</paragraph>
</subSubSection>
<subSubSection pageId="566" pageNumber="567" type="records">
<paragraph pageId="566" pageNumber="567">Records.</paragraph>
<paragraph pageId="566" pageNumber="567">
<emphasis bold="true" pageId="566" pageNumber="567">USA</emphasis>
: AZ, CA, OR, WA [CO, NM, UT] - Mexico
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="006F8AF6DC73EC7CD4EB3A2D4F3D00A6" authority="Roemer, 1870" authorityName="Roemer" authorityYear="1870" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python euboicus" pageId="22" pageNumber="51" phylum="Chordata" rank="species" species="euboicus">
Python euboicus
<normalizedToken id="6455A0151032C039959404B44BA02181" originalValue="Römer">Roemer</normalizedToken>
, 1870
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<taxonomicNameLabel id="83D50D8CE8FC1D65C37EF54165E698F6" pageId="22" pageNumber="51">[extinct species, considered nomen dubium by Rage 1984]</taxonomicNameLabel>
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<paragraph id="7F487D7D856962E21E196486EBC84610" pageId="22" pageNumber="51">Synonyms:</paragraph>
<paragraph id="A6C2D6CFEC38DF9EAE214291F4933E22" pageId="22" pageNumber="51">
<taxonomicName id="D5872F24040CA15627571B4FCBE18918" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python (Euboicus)" pageId="22" pageNumber="51" phylum="Chordata" rank="subGenus" subGenus="Euboicus">Python Euboicus</taxonomicName>
<normalizedToken id="5E0AB6DE264FFEC8A0461851F2995BDE" originalValue="Römer">Roemer</normalizedToken>
, 1870
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<paragraph id="6857FE01C48653C96D22DFC8936B903A" pageId="22" pageNumber="51">
<taxonomicName id="467198E2D49ED0740212BCC9BACB5BFC" class="Reptilia" genus="Heteropython" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Heteropython euboicus" pageId="22" pageNumber="51" phylum="Chordata" rank="species" species="euboicus">Heteropython euboicus</taxonomicName>
(
<normalizedToken id="10550C39F3414271426F9ABC449F007C" originalValue="Römer">Roemer</normalizedToken>
) -
<bibRefCitation id="D1CA2181FF6D9966A47C3E1C4C3E685B" author="Rochebrune de, AT" journalOrPublisher="Nouvelles Archives du Museum d'Histoire naturelle" pageId="25" pageNumber="54" pagination="271 - 296" title="Revision des ophidiens fossiles du Museum d'Histoire naturelle." volume="3" year="1880">de Rochebrune 1880</bibRefCitation>
</paragraph>
<paragraph id="A146D1319E1EEE627F84DD4221138A0D" pageId="22" pageNumber="51">
<taxonomicName id="69D50E46AC17B95FF59C08D352CEC189" class="Reptilia" genus="Heteropython" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Heteropython euboicus" pageId="22" pageNumber="51" phylum="Chordata" rank="species" species="euboicus">Heteropython euboicus</taxonomicName>
(
<normalizedToken id="1554C47E258F1518B61AD7C9D77A54D5" originalValue="Römer">Roemer</normalizedToken>
) - Kuhn 1939, 1963
</paragraph>
<paragraph id="BB93F8A07A5ECF544977F1637E0418DA" pageId="22" pageNumber="51">
<taxonomicName id="77D25A373724425AAA72A909ABE2E951" class="Squamata" family="Pythonidae" genus="Python" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Python euboicus" pageId="22" pageNumber="51" phylum="Chordata" rank="species" species="euboicus">Python euboicus</taxonomicName>
<normalizedToken id="D1620F436ABCA38802694AB51E3F368E" originalValue="Römer">Roemer</normalizedToken>
-
<bibRefCitation id="251B28CDBE90C226696D626CF7722605" author="Rage, J-C" editor="Wellenhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart-New York" pageId="25" pageNumber="54" title="Serpentes." volumeTitle="Handbuch der Palioherpetologie" year="1984">Rage 1984</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="862505A769B647D05B1CDC6E438C8826" pageId="22" pageNumber="51" type="holotype">
<paragraph id="18BE6860D0C989F6ACCA476A9681BA26" pageId="22" pageNumber="51">Holotype:</paragraph>
<paragraph id="61A761CFB57AF62BB09030FEF074DEB6" pageId="22" pageNumber="51">
Fragment of the trunk portion of the vertebral column (25 vertebrae and ribs), left dentary. No accession number. According to
<bibRefCitation id="2A6824018A912CBF66004374B16F1099" author="Szyndlar, Z" journalOrPublisher="Estudios geologica" pageId="25" pageNumber="54" pagination="103 - 126" title="A Review of Neogene and Quarternary Snakes of Central and Eastern Europe. Part 1: Scolecophidia, Boidae, Colubrinae." volume="47" year="1991">Szyndlar (1991)</bibRefCitation>
the holotype is probably lost.
</paragraph>
</subSubSection>
<subSubSection id="1611BC474DB53BDCC443E5BAE424B847" pageId="22" pageNumber="51" type="type locality">
<paragraph id="31F8AC58A86C16F20AB308E70263D465" pageId="22" pageNumber="51">Type locality:</paragraph>
<paragraph id="67634F838D4C077A9CD8B064084E9212" pageId="22" pageNumber="51">Kimi (Euboea, Greece), early Miocene (MN?3).</paragraph>
</subSubSection>
<subSubSection id="F32B3901C661140E2EEB0199EB31D20B" pageId="22" pageNumber="51" type="remarks">
<paragraph id="4850F84929CB9AD547C4E839C91B779B" pageId="22" pageNumber="51">Remarks:</paragraph>
<paragraph id="0285F08B69879C3F5D3A9C1A66269E67" pageId="22" pageNumber="51">
See
<bibRefCitation id="CDA9302EB37573180FFE95A9CC72140A" author="Szyndlar, Z" journalOrPublisher="Estudios geologica" pageId="25" pageNumber="54" pagination="103 - 126" title="A Review of Neogene and Quarternary Snakes of Central and Eastern Europe. Part 1: Scolecophidia, Boidae, Colubrinae." volume="47" year="1991">Szyndlar (1991)</bibRefCitation>
and
<bibRefCitation id="673D756F968DF6F34BE4BDEB23D5B286" author="Szyndlar, Z" journalOrPublisher="Institute of Systematics and Evolution of Animals. Polish Academy of Sciences, Poland" pageId="25" pageNumber="54" title="Non-erycine Booidea from the Oligocene and Miocene of Europe." year="2003">Szyndlar and Rage (2003: 67-68)</bibRefCitation>
for further information.
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