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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.392.6328" ID-PMC="PMC3974435" ID-Pensoft-Pub="1313-2970-392-1" ID-Pensoft-UUID="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" ID-PubMed="24715782" ID-Zenodo-Dep="578044" ModsDocID="1313-2970-392-1" checkinTime="1451246138192" checkinUser="pensoft" docAuthor="Breure, Abraham S. H. &amp; Ablett, Jonathan D." docDate="2014" docId="E25E3CB4CB4799ADCD60273D51992A6A" docLanguage="en" docName="ZooKeys 392: 1-367" docOrigin="ZooKeys 392" docPubDate="2014-03-21" docSource="http://dx.doi.org/10.3897/zookeys.392.6328" docTitle="Bulimus chemnitzioides Forbes 1850" docType="treatment" docVersion="5" id="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" lastPageNumber="35" masterDocId="FFCF5A59FFB1FF93FFF10B43FFAFFFF5" masterDocTitle="Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London" masterLastPageNumber="367" masterPageNumber="1" pageNumber="35" updateTime="1668158078298" updateUser="ExternalLinkService">
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<mods:title>Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Breure, Abraham S. H.</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, Leiden, the Netherlands</mods:affiliation>
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<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Natural History Museum, Division of Higher Invertebrates, London, SW 7 5 BD, UK</mods:affiliation>
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<taxonomicName LSID="E25E3CB4-CB47-99AD-CD60-273D51992A6A" authority="Forbes, 1850" authorityName="Forbes" authorityYear="1850" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Bulimus chemnitzioides" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="chemnitzioides">Bulimus chemnitzioides Forbes, 1850</taxonomicName>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-F Bostryx species. A-C Bostryx angrandianus (Pilsbry, 1897), lectotype of Bulimus radiatus Morelet, 1863 NHMUK 1893.2.4.198 (H = 23.7) D Bostryx montagnei (d'Orbigny, 1837), lectotype NHMUK 1854.12.4.194 (H = 23.3) E Bostryx serotinus (Morelet, 1860), lectotype NHMUK 1893.2.4.204 (H = 25.5) F Bostryx nigropileatus (Reeve, 1849), lectotype of Bulimus stenacme Pfeiffer, 1857 (H = 28.0). G-J Naesiotus species. G Naesiotus wolfi (Reibisch, 1892), paralectotype NHMUK 1894.6.8.7 (H = 12.19) H Naesiotus fernandezae Weyrauch, 1958, paratype NHMUK 19831 (H = 15.1) I Naesiotus chemnitzioides (Forbes, 1850), lectotype NHMUK 1855.4.5.23 (H = 18.5) J Naesiotus rugiferus (Sowerby I, 1833), lectotype NHMUK 1975178 (H = 12.5). All enlarged." httpUri="https://binary.pensoft.net/fig/26635" pageId="34" pageNumber="35">Figs 12I</figureCitation>
<figureCitation captionStart="Figure L12" captionStartId="F12L" captionText="Figure L 12. Labels. i Bulimus cercicola Morelet, 1863 ii Bulimus chamaeleon Pfeiffer, 1855 iii Otostomus championi Martens, 1893 iv Bulimus chemnitzioides Forbes, 1850 v Bulimus chimborasensis Reeve, 1848 vi Drymaeus chiriquiensis da Costa, 1901 vii Bulimus cinereus Reeve, 1849." httpUri="https://binary.pensoft.net/fig/26763" pageId="34" pageNumber="35">, L12iv</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="reference_group">
<paragraph pageId="34" pageNumber="35">
<taxonomicName authorityName="Forbes" authorityYear="1850" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="chemnitzioides">Bulimus chemnitzioides</taxonomicName>
<bibRefCitation author="Forbes, E" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="183" pageNumber="184" pagination="53 - 56" refId="B55" refString="Forbes, E, 1850. On the species of Mollusca collected during the surveying voyages of the Herald and Pandora, by Captain Kellett and Lieut. Wood. Proceedings of the Zoological Society of London 1850: 53 - 56" title="On the species of Mollusca collected during the surveying voyages of the Herald and Pandora, by Captain Kellett and Lieut. Wood." volume="1850" year="1850">Forbes 1850</bibRefCitation>
: 55, pl. 9 figs 6a-b; Pfeiffer 1854 in
<bibRefCitation author="Kuester, HC" journalOrPublisher="Systematisches Conchylien-Cabinet von Martini und Chemnitz" pageId="184" pageNumber="185" pagination="1 - 395" refId="B74" refString="Kuester, HC, Pfeiffer, L, 1840-1855 1840-1865. Die Gattungen Bulimus, Partula, Achatinella, Achatina und Azeca. Systematisches Conchylien-Cabinet von Martini und Chemnitz 13 (1): 1 - 395" title="Die Gattungen Bulimus, Partula, Achatinella, Achatina und Azeca." volume="13" year="1840 - 1855">
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
and Pfeiffer 1840-1865
</bibRefCitation>
: 94, pl. 31 figs 21-23.
</paragraph>
<paragraph pageId="34" pageNumber="35">
<taxonomicName class="Gastropoda" family="Bulimulidae" genus="Bulimulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="chemnitzioides">Bulimulus chemnitzioides</taxonomicName>
;
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Manual of Conchology" pageId="188" pageNumber="189" pagination="1 - 399" refId="B150" refString="Pilsbry, HA, 1897-1898. American Bulimulidae: Bulimulus, Neopetraeus, Oxychona and South American Drymaeus. Manual of Conchology 11: 1 - 399" title="American Bulimulidae: Bulimulus, Neopetraeus, Oxychona and South American Drymaeus." volume="11" year="1897 - 1898">Pilsbry 1897 [1897-1898]</bibRefCitation>
: 124, pl. 24 figs 45-46.
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="type locality">
<paragraph pageId="34" pageNumber="35">Type locality.</paragraph>
<paragraph pageId="34" pageNumber="35">
[Ecuador] &quot;Chatham Island, Gelepagos [sic,
<normalizedToken originalValue="Galápagos]”">Galapagos]&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="label">
<paragraph pageId="34" pageNumber="35">Label.</paragraph>
<paragraph pageId="34" pageNumber="35">&quot;Chatham Island / Galapagos / Proc. Zool. Soc. / fig. 6&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="dimensions">
<paragraph pageId="34" pageNumber="35">Dimensions.</paragraph>
<paragraph pageId="34" pageNumber="35">&quot;Long. 19, diam. 4 mil..&quot;; figured specimen herein H 18.5, D 3.64, W 16.0.</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="type material">
<paragraph pageId="34" pageNumber="35">Type material.</paragraph>
<paragraph pageId="34" pageNumber="35">NHMUK 1855.4.5.23, lectotype; 1850.4.23.14-15, two paralectotypes (ex Kellett and Wood).</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="remarks">
<paragraph pageId="34" pageNumber="35">Remarks.</paragraph>
<paragraph pageId="34" pageNumber="35">
The specimen originally figured is here designated lectotype (
<emphasis bold="true" pageId="34" pageNumber="35">design. n.</emphasis>
). The two other specimens also originate from the voyage of Kellett and Wood (
<bibRefCitation author="Forbes, E" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="183" pageNumber="184" pagination="53 - 56" refId="B55" refString="Forbes, E, 1850. On the species of Mollusca collected during the surveying voyages of the Herald and Pandora, by Captain Kellett and Lieut. Wood. Proceedings of the Zoological Society of London 1850: 53 - 56" title="On the species of Mollusca collected during the surveying voyages of the Herald and Pandora, by Captain Kellett and Lieut. Wood." volume="1850" year="1850">Forbes 1850</bibRefCitation>
,
<bibRefCitation author="Seemann, B" journalOrPublisher="Reeve and Co., London" pageId="189" pageNumber="190" refId="B168" refString="Seemann, B, 1853. Narrative of the voyage of H.M.S. Herald during the years 1845-51, under the command of Captain Henry Kellett, R.N., C.B.; during a circumnavigation of the globe, and three cruises to the Arctic regions in search of Sir John Franklin. Reeve and Co., London" title="Narrative of the voyage of H. M. S. Herald during the years 1845 - 51, under the command of Captain Henry Kellett, R. N., C. B.; during a circumnavigation of the globe, and three cruises to the Arctic regions in search of Sir John Franklin." year="1853">Seemann 1853</bibRefCitation>
), and are considered to be part of the original series.
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="current systematic position">
<paragraph pageId="34" pageNumber="35">Current systematic position.</paragraph>
<paragraph pageId="34" pageNumber="35">
<taxonomicName authorityName="Tryon" authorityYear="1867" class="Gastropoda" family="Bulimulidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="34" pageNumber="35" phylum="Mollusca" rank="family">Bulimulidae</taxonomicName>
,
<taxonomicName baseAuthorityName="Forbes" baseAuthorityYear="1850" class="Gastropoda" family="Bulimulidae" genus="Naesiotus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="34" pageNumber="35" phylum="Mollusca" rank="species" species="chemnitzioides">
<emphasis italics="true" pageId="34" pageNumber="35">Naesiotus chemnitzioides</emphasis>
</taxonomicName>
(Forbes, 1850).
</paragraph>
</subSubSection>
</treatment>
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SUBFAMILY
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AMBRYSINAE
<bibRefCitation id="0E664B68FF80FFDEFDD5822A05FDFA8B" author="Usinger RL" box="[529,701,1293,1318]" pageId="13" pageNumber="1258" pagination="5 - 16" refId="ref25189" refString="Usinger RL. 1941. Key to the subfamilies of Naucoridae with a generic synopsis of the new subfamily Ambrysinae (Hemiptera). Annals of the Entomological Society of America 34: 5 - 16." type="journal article" year="1941">USINGER, 1941</bibRefCitation>
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<figureCitation id="F2CC2A1CFF80FFDEFEB6821E068CFAFC" box="[370,460,1337,1362]" captionStart="Figure 3" captionStartId="8.[145,230,1468,1490]" captionTargetBox="[172,733,195,1428]" captionTargetId="figure-532@8.[172,733,195,1428]" captionTargetPageId="8" captionText="Figure 3. Mouthpart positions: recessed rostrum of Cheirochela thailandana (a); lateral views of Laccocoris spurcus (b) and Australambrysus crenulatus (c). White arrows indicate positions of labrum with respect to the functional anterior margin of the head." figureDoi="http://doi.org/10.5281/zenodo.6994613" httpUri="https://zenodo.org/record/6994613/files/figure.png" pageId="13" pageNumber="1258">FIGS 3C</figureCitation>
,
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,
<figureCitation id="F2CC2A1CFF80FFDEFDD5821E057CFAFC" box="[529,572,1337,1362]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="13" pageNumber="1258">11A</figureCitation>
</paragraph>
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species:
</emphasis>
<taxonomicName id="ADF74D1AFF80FFDEFEFB824805D3FA28" ID-CoL="7QM5Z" authority="Stal, 1862" authorityName="Stal" authorityYear="1862" box="[319,659,1391,1413]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="5883EA8BFF80FFDEFEFB8248055AFA28" box="[319,538,1391,1413]" italics="true" pageId="13" pageNumber="1258">Ambrysus signoreti</emphasis>
<bibRefCitation id="0E664B68FF80FFDEFDE5824805D3FA28" author="Stal C" box="[545,659,1391,1413]" pageId="13" pageNumber="1258" pagination="437 - 462" refId="ref25046" refString="Stal C. 1862. Hemiptera Mexicana enumeravit speciesque novas descripsit (continuatio). Entomologische Zeitung Stettiner 23: 437 - 462." type="journal article" year="1862">Stål, 1862</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF80FFDEFF6782830383F9F4" pageId="13" pageNumber="1258" type="discussion">
<paragraph id="6A483699FF80FFDEFF67828304CEFA71" blockId="13.[163,780,1443,1894]" lastBlockId="13.[827,1442,1295,1500]" pageId="13" pageNumber="1258">
<emphasis id="5883EA8BFF80FFDEFF67828306C6FA15" box="[163,390,1443,1465]" italics="true" pageId="13" pageNumber="1258">Taxonomic history:</emphasis>
This subfamily was established by Usinger in 1941, but was downgraded to tribe level and placed in
<taxonomicName id="ADF74D1AFF80FFDEFE5582C60586FA5B" authority="(Popov, 1970)" authorityName="LEACH" authorityYear="1815" baseAuthorityName="Popov" baseAuthorityYear="1970" box="[401,710,1505,1527]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">
Naucorinae (
<bibRefCitation id="0E664B68FF80FFDEFDEB82C605FBFA5B" author="Popov YA" box="[559,699,1505,1526]" pageId="13" pageNumber="1258" pagination="93 - 98" refId="ref23866" refString="Popov YA. 1970. Notes on the classification of the recent Naucoridae (Heteroptera: Nepomorpha). Bulletin of the Polish Academy of Sciences (Biological Sciences) 18: 93 - 98." type="journal article" year="1970">Popov, 1970</bibRefCitation>
)
</taxonomicName>
, then transferred to
<taxonomicName id="ADF74D1AFF80FFDEFEA082D80567F9B8" baseAuthorityName="Reynoso-Velasco &amp; Sites" baseAuthorityYear="2021" box="[356,551,1535,1557]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
(
<bibRefCitation id="0E664B68FF80FFDEFDF882D907A2F99E" author="Stys P &amp; Jansson A" pageId="13" pageNumber="1258" pagination="1 - 44" refId="ref25103" refString="Stys P, Jansson A. 1988. Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica 50: 1 - 44." type="journal article" year="1988">Štys &amp; Jansson, 1988</bibRefCitation>
) and ultimately elevated and restored to subfamily rank (
<bibRefCitation id="0E664B68FF80FFDEFE77811A079EF9DD" author="Reynoso-Velasco D &amp; Sites RW" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFD04811A079EF9DD" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites, 2021</taxonomicName>
</bibRefCitation>
). Significant recent changes in the generic composition of the subfamily include the transfer of
<taxonomicName id="ADF74D1AFF80FFDEFF0481BE066BF903" authorityName="Stal" authorityYear="1876" box="[192,299,1689,1710]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFF0481BE066BF903" box="[192,299,1689,1710]" italics="true" pageId="13" pageNumber="1258">Pelocoris</emphasis>
</taxonomicName>
in from
<taxonomicName id="ADF74D1AFF80FFDEFE5581BE055FF902" authorityName="LEACH" authorityYear="1815" box="[401,543,1689,1711]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
, and the erection of the genera
<emphasis id="5883EA8BFF80FFDEFEEF819006BAF961" box="[299,506,1719,1740]" italics="true" pageId="13" pageNumber="1258">Australambrysus</emphasis>
Reynoso &amp;
<taxonomicName id="ADF74D1AFF80FFDEFD4C8190044BF961" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[648,779,1719,1741]" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF80FFDEFD4C8190044BF961" author="Sites RW" box="[648,779,1719,1741]" pageId="13" pageNumber="1258" pagination="72 - 94" refId="ref24528" refString="Sites RW. 2021. A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species. Zootaxa 4958: 72 - 94." type="journal article" year="2021">Sites, 2021</bibRefCitation>
</taxonomicName>
and
<emphasis id="5883EA8BFF80FFDEFF1A81F106EBF946" box="[222,427,1750,1771]" italics="true" pageId="13" pageNumber="1258">Maculambrysus</emphasis>
(
<bibRefCitation id="0E664B68FF80FFDEFE0581F1079DF8A7" author="Reynoso-Velasco D &amp; Sites RW" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFD0081F1079DF8A7" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites, 2021</taxonomicName>
</bibRefCitation>
). The subfamily currently includes
<taxonomicName id="ADF74D1AFF80FFDEFD5081D20672F885" authority="Stal, 1862" authorityName="Stal" authorityYear="1862" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFD5081D2044BF8A7" box="[660,779,1781,1802]" italics="true" pageId="13" pageNumber="1258">Ambrysus</emphasis>
<bibRefCitation id="0E664B68FF80FFDEFF6780340672F885" author="Stal C" box="[163,306,1811,1833]" pageId="13" pageNumber="1258" pagination="437 - 462" refId="ref25046" refString="Stal C. 1862. Hemiptera Mexicana enumeravit speciesque novas descripsit (continuatio). Entomologische Zeitung Stettiner 23: 437 - 462." type="journal article" year="1862">Stål, 1862</bibRefCitation>
</taxonomicName>
,
<emphasis id="5883EA8BFF80FFDEFE8F80340575F884" box="[331,565,1811,1833]" italics="true" pageId="13" pageNumber="1258">Australambrysus</emphasis>
,
<taxonomicName id="ADF74D1AFF80FFDEFD8A8033044BF884" authorityName="De Carlo" authorityYear="1963" box="[590,779,1811,1833]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFD8A8033044BF884" box="[590,779,1811,1833]" italics="true" pageId="13" pageNumber="1258">Carvalhoiella</emphasis>
</taxonomicName>
D e C a r l o, 1 9 6 3,
<emphasis id="5883EA8BFF80FFDEFEBB8015057FF8E5" box="[383,575,1842,1864]" italics="true" pageId="13" pageNumber="1258">C a t a r a c t o c o r i s</emphasis>
U s i n g e r, 1 9 4 1,
<emphasis id="5883EA8BFF80FFDEFF678076063AF8CB" box="[163,378,1873,1894]" italics="true" pageId="13" pageNumber="1258">H y g r o p e t r o c o r i s</emphasis>
S i t e s, 2 0 1 5,
<emphasis id="5883EA8BFF80FFDEFDF38076044BF8CB" box="[567,779,1872,1894]" italics="true" pageId="13" pageNumber="1258">M a c u l a m b r y s u s</emphasis>
Reynoso &amp;
<taxonomicName id="ADF74D1AFF80FFDEFC1F82280333FA89" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[987,1139,1295,1317]" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF80FFDEFC1F82280333FA89" author="Sites RW" box="[987,1139,1295,1317]" pageId="13" pageNumber="1258" pagination="72 - 94" refId="ref24528" refString="Sites RW. 2021. A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species. Zootaxa 4958: 72 - 94." type="journal article" year="2021">Sites, 2021</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF80FFDEFB4E822802DAFA89" authority="De Carlo" authorityName="De Carlo" authorityYear="1935" box="[1162,1434,1295,1317]" class="Insecta" family="Naucoridae" genus="Melloiella" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFB4E82280252FA89" box="[1162,1298,1295,1316]" italics="true" pageId="13" pageNumber="1258">Melloiella</emphasis>
De Carlo
</taxonomicName>
, 1 9 3 5,
<emphasis id="5883EA8BFF80FFDEFC4B82090346FAEE" box="[911,1030,1325,1347]" italics="true" pageId="13" pageNumber="1258">Pe l o c o r i s</emphasis>
,
<emphasis id="5883EA8BFF80FFDEFBD98209033FFAEE" box="[1053,1151,1326,1347]" italics="true" pageId="13" pageNumber="1258">P i c r o p s</emphasis>
L a R i v e r s, 1 9 5 2 a n d
<taxonomicName id="ADF74D1AFF80FFDEFCFF826B0399FACC" authority="J. Polhemus, 1991" authorityName="J. Polhemus" authorityYear="1991" box="[827,1241,1356,1378]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFCFF826B04BBFACC" box="[827,1019,1356,1377]" italics="true" pageId="13" pageNumber="1258">Procryphocricos</emphasis>
J.
<bibRefCitation id="0E664B68FF80FFDEFBD9826B0399FACC" author="Polhemus JT" box="[1053,1241,1356,1378]" pageId="13" pageNumber="1258" pagination="119 - 123" refId="ref23840" refString="Polhemus JT. 1991. A new and primitive genus of Cryphocricinae (Heteroptera: Naucoridae). Pan-Pacific Entomologist 67: 119 - 123." type="journal article" year="1991">Polhemus, 1991</bibRefCitation>
</taxonomicName>
. Species of all of these genera, except
<taxonomicName id="ADF74D1AFF80FFDEFBFC824C03E3FA2D" authorityName="Stal" authorityYear="1876" box="[1080,1187,1387,1408]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFBFC824C03E3FA2D" box="[1080,1187,1387,1408]" italics="true" pageId="13" pageNumber="1258">Pelocoris</emphasis>
</taxonomicName>
, were included along with
<taxonomicName id="ADF74D1AFF80FFDEFCBD82AE0356FA33" authorityName="Signoret" authorityYear="1850" box="[889,1046,1417,1438]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFCBD82AE0356FA33" box="[889,1046,1417,1438]" italics="true" pageId="13" pageNumber="1258">Cryphocricos</emphasis>
</taxonomicName>
in the subfamily
<taxonomicName id="ADF74D1AFF80FFDEFB2F82AE02E2FA32" baseAuthorityName="Reynoso-Velasco &amp; Sites" baseAuthorityYear="2021" box="[1259,1442,1417,1439]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
prior to the revision by
<bibRefCitation id="0E664B68FF80FFDEFBA9828F04C6FA71" author="Reynoso-Velasco D &amp; Sites RW" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFAA1828F04C6FA71" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
.
</paragraph>
<paragraph id="6A483699FF80FFDEFCFF81220383F9F4" blockId="13.[827,1442,1541,1625]" pageId="13" pageNumber="1258">
<emphasis id="5883EA8BFF80FFDEFCFF8122036AF9B6" box="[827,1066,1541,1563]" italics="true" pageId="13" pageNumber="1258">Revised taxonomy:</emphasis>
The subfamily was recently revised by
<bibRefCitation id="0E664B68FF80FFDEFC728103025CF994" author="Reynoso-Velasco D &amp; Sites RW" box="[950,1308,1572,1594]" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFB508103025CF994" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1172,1308,1572,1594]" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
. No further taxonomic action is proposed here.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF80FFDDFCFF81A5057EFE42" lastPageId="14" lastPageNumber="1259" pageId="13" pageNumber="1258" type="diagnosis">
<paragraph id="6A483699FF80FFDDFCFF81A5057EFE42" blockId="13.[827,1443,1665,1902]" lastBlockId="14.[145,762,197,495]" lastPageId="14" lastPageNumber="1259" pageId="13" pageNumber="1258">
<emphasis id="5883EA8BFF80FFDEFCFF81A504FDF93A" box="[827,957,1666,1687]" italics="true" pageId="13" pageNumber="1258">Diagnosis:</emphasis>
Diagnostic features were provided by
<bibRefCitation id="0E664B68FF80FFDEFCFF818703E5F918" author="Reynoso-Velasco D &amp; Sites RW" box="[827,1189,1696,1718]" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFBD8818703E5F918" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1052,1189,1696,1718]" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
as follows: (1) labium inserted near the anterior margin of the head; (2) labrum well developed; (3) maxillary plates only slightly produced anteriorly; (4) anterior margin of the pronotum generally with a concavity of different depths, but straight in
<taxonomicName id="ADF74D1AFF80FFDEFC73801E0359F8E3" authorityName="Stal" authorityYear="1876" box="[951,1049,1849,1870]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFC73801E0359F8E3" box="[951,1049,1849,1870]" italics="true" pageId="13" pageNumber="1258">Pelocoris</emphasis>
</taxonomicName>
; (5) pronotum with lateral margins smooth or finely crenate; (6) prosternellum concealed by the mesally directed propleural projections, but prosternellum exposed in
<taxonomicName id="ADF74D1AFF83FFDDFE6987C3054FFF54" authorityName="Stal" authorityYear="1876" box="[429,527,228,249]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFE6987C3054FFF54" box="[429,527,228,249]" italics="true" pageId="14" pageNumber="1259">Pelocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF83FFDDFD8787C305B3FF54" authorityName="J. Polhemus" authorityYear="1991" box="[579,755,228,249]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFD8787C305B3FF54" box="[579,755,228,249]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
; (7) forewing macropterous, but brachyptery also present in
<taxonomicName id="ADF74D1AFF83FFDDFED586060689FE9B" authorityName="J. Polhemus" authorityYear="1991" box="[273,457,289,310]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFED586060689FE9B" box="[273,457,289,310]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
; (8) no sternal abdominal sense organs (hydrostats), except in
<taxonomicName id="ADF74D1AFF83FFDDFDFF866705B2FEF8" authorityName="J. Polhemus" authorityYear="1991" box="[571,754,320,341]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFDFF866705B2FEF8" box="[571,754,320,341]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
; (9) compressible air-bubble respiration, but a plastron could be used by brachypterous
<taxonomicName id="ADF74D1AFF83FFDDFDE9865A05B1FE3F" authorityName="J. Polhemus" authorityYear="1991" box="[557,753,381,402]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFDE9865A05B1FE3F" box="[557,753,381,402]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
; (10) pubescent abdomen (scattered long setae in
<taxonomicName id="ADF74D1AFF83FFDDFF55869D0615FE62" authorityName="J. Polhemus" authorityYear="1991" box="[145,341,442,463]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFF55869D0615FE62" box="[145,341,442,463]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
); and (11) males with vesica not modified into a long tubular structure.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF83FFDDFF55853F0529FDE1" pageId="14" pageNumber="1259" type="discussion">
<paragraph id="6A483699FF83FFDDFF55853F0529FDE1" blockId="14.[145,758,536,588]" pageId="14" pageNumber="1259">
<emphasis id="5883EA8BFF83FFDDFF55853F0661FD80" box="[145,289,536,557]" italics="true" pageId="14" pageNumber="1259">C o m m e n t s:</emphasis>
A l l s p e c i e s i n t h i s s u b f a m i l y a r e distributed exclusively in the New World.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D86B7E2FFABE4CF3D79A7ABA834219E6">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:title id="D4143ECFEF14C0D7D461DE18AE5776DB">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="319F8E13E6E36D8126BFC9E3235CF8E8">
<mods:date id="37A9A645DBC376A3F74A04CFF5F1B9D0">2022</mods:date>
<mods:detail id="630E9EC615CE37737ABA9F6B367D8C88" type="pubDate">
<mods:number id="9523F4A7CF6F31DD2E7843F66F6FBA37">2022-02-07</mods:number>
</mods:detail>
<mods:detail id="91EEB9534C64392DF8C52C489CDD4B16" type="volume">
<mods:number id="12596D2080F33C859617D6E4E6AB8879">195</mods:number>
</mods:detail>
<mods:extent id="97657BA7D32F0B910ED4B4B632470EFD" unit="page">
<mods:start id="099B70F3284B932772B9FE318D1ADF2C">1245</mods:start>
<mods:end id="963F07EEE06A5AAAA1AE3BCDA52A9A15">1286</mods:end>
</mods:extent>
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<mods:classification id="2A0F239B9B637E757BC8D315019D7EB7">journal article</mods:classification>
<mods:identifier id="5F7E9DFCA15AFE9D6CB416CC710BCC33" type="CLB-Dataset">121760</mods:identifier>
<mods:identifier id="B4BD98BE923BFCCCE8E0EDD72FD7E41C" type="DOI">10.1093/zoolinnean/zlab105</mods:identifier>
<mods:identifier id="6412A5088F6E6F0C1D0621F19CFBE770" type="GBIF-Dataset">83d946c2-fa80-4948-9160-16625c240886</mods:identifier>
<mods:identifier id="4734A8F96B47E0B6C305CC57F0038FF9" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="C4BAE2C68A2015EFC91423EE4716B72A" type="Zenodo-Dep">6994599</mods:identifier>
<mods:identifier id="E4DBCA7A090EC9573AC925A2905752C1" type="ZooBank">C06A1F94-AF08-4A21-B1F3-A0865FB1A8DF</mods:identifier>
</mods:mods>
<treatment id="E25E878FFF83FFC3FCF887E3060DFB5D" ID-DOI="http://doi.org/10.5281/zenodo.6994697" ID-GBIF-Taxon="199315273" ID-Zenodo-Dep="6994697" LSID="urn:lsid:plazi:treatment:E25E878FFF83FFC3FCF887E3060DFB5D" httpUri="http://treatment.plazi.org/id/E25E878FFF83FFC3FCF887E3060DFB5D" lastPageId="16" lastPageNumber="1261" pageId="14" pageNumber="1259">
<subSubSection id="22ED6512FF83FFDDFCF887E3023EFF76" box="[828,1406,196,221]" pageId="14" pageNumber="1259" type="nomenclature">
<paragraph id="6A483699FF83FFDDFCF887E3023EFF76" blockId="14.[828,1406,196,221]" box="[828,1406,196,221]" pageId="14" pageNumber="1259">
SUBFAMILY
<taxonomicName id="ADF74D1AFF83FFDDFC0787E3022EFF70" ID-CoL="JBP" authority="MONTANDON, 1897" authorityName="MONTANDON" authorityYear="1897" box="[963,1390,196,221]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">CHEIROCHELINAE MONTANDON, 1897</taxonomicName>
E
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF83FFDDFC3C87D70382FEAA" box="[1016,1218,240,265]" pageId="14" pageNumber="1259" type="description">
<paragraph id="6A483699FF83FFDDFC3C87D70382FEAA" blockId="14.[1016,1218,240,265]" box="[1016,1218,240,265]" pageId="14" pageNumber="1259">
<figureCitation id="F2CC2A1CFF83FFDDFC3C87D70312FEAA" box="[1016,1106,240,265]" captionStart="Figure 3" captionStartId="8.[145,230,1468,1490]" captionTargetBox="[172,733,195,1428]" captionTargetId="figure-532@8.[172,733,195,1428]" captionTargetPageId="8" captionText="Figure 3. Mouthpart positions: recessed rostrum of Cheirochela thailandana (a); lateral views of Laccocoris spurcus (b) and Australambrysus crenulatus (c). White arrows indicate positions of labrum with respect to the functional anterior margin of the head." figureDoi="http://doi.org/10.5281/zenodo.6994613" httpUri="https://zenodo.org/record/6994613/files/figure.png" pageId="14" pageNumber="1259">FIGS 3A</figureCitation>
,
<figureCitation id="F2CC2A1CFF83FFDDFB9B87D703CAFEAA" box="[1119,1162,240,265]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="14" pageNumber="1259">10B</figureCitation>
,
<figureCitation id="F2CC2A1CFF83FFDDFB5387D70382FEAA" box="[1175,1218,240,265]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="14" pageNumber="1259">11B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF83FFDDFCED8601021FFE96" box="[809,1375,294,315]" pageId="14" pageNumber="1259" type="type_taxon">
<paragraph id="6A483699FF83FFDDFCED8601021FFE96" blockId="14.[809,1375,294,315]" box="[809,1375,294,315]" pageId="14" pageNumber="1259">
<emphasis id="5883EA8BFF83FFDDFCED860104F9FE96" box="[809,953,294,315]" italics="true" pageId="14" pageNumber="1259">
<typeStatus id="B54C883BFF83FFDDFCED8601041EFE96" box="[809,862,294,315]" pageId="14" pageNumber="1259">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF83FFDDFC0186010218FE96" ID-CoL="TTYV" authority="Hope, 1840" authorityName="Hope" authorityYear="1840" box="[965,1368,294,315]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="5883EA8BFF83FFDDFC0186010392FE96" box="[965,1234,294,315]" italics="true" pageId="14" pageNumber="1259">Cheirochela assamensis</emphasis>
<bibRefCitation id="0E664B68FF83FFDDFB1D86010218FE96" author="Hope FW" box="[1241,1368,294,315]" pageId="14" pageNumber="1259" pagination="435 - 447" refId="ref22124" refString="Hope FW. 1840. Descriptions of some new insects, collected in Assam by William Griffith, Esq., assistant-surgeon in the Madras Medical Service, and attached to the late scientific mission to Assam. Transactions of the Linnean Society of London 18: 435 - 447." type="journal article" year="1840">Hope, 1840</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF83FFC3FCED864205BAFE9A" lastPageId="16" lastPageNumber="1261" pageId="14" pageNumber="1259" type="discussion">
<paragraph id="6A483699FF83FFDCFCED86420604F8C1" blockId="14.[809,1425,356,593]" lastBlockId="15.[163,779,1726,1901]" lastPageId="15" lastPageNumber="1260" pageId="14" pageNumber="1259">
<emphasis id="5883EA8BFF83FFDDFCED86420358FED4" box="[809,1048,356,378]" italics="true" pageId="14" pageNumber="1259">Taxonomic history:</emphasis>
This subfamily has included three tribes:
<taxonomicName id="ADF74D1AFF83FFDDFC0D86A40245FE35" authority="Popov, 1970" authorityName="Popov" authorityYear="1970" box="[969,1285,387,409]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1284" phylum="Arthropoda" rank="tribe" tribe="Cheirochelini">
Cheirochelini
<bibRefCitation id="0E664B68FF83FFDDFBB286A40245FE35" author="Popov YA" box="[1142,1285,387,408]" pageId="14" pageNumber="1259" pagination="93 - 98" refId="ref23866" refString="Popov YA. 1970. Notes on the classification of the recent Naucoridae (Heteroptera: Nepomorpha). Bulletin of the Polish Academy of Sciences (Biological Sciences) 18: 93 - 98." type="journal article" year="1970">Popov, 1970</bibRefCitation>
</taxonomicName>
(Indochina to
<collectingCountry id="12E07609FF83FFDDFC8E868504CBFE15" box="[842,907,418,440]" name="India" pageId="14" pageNumber="1259">India</collectingCountry>
and Borneo), Sagocorini
<bibRefCitation id="0E664B68FF83FFDDFB08868502D0FE1A" author="La Rivers I" box="[1228,1424,418,440]" pageId="14" pageNumber="1259" pagination="1 - 120" refId="ref22462" refString="La Rivers I. 1971. Studies of Naucoridae (Hemiptera). Biological Society of Nevada Memoirs 2: 1 - 120." type="journal article" year="1971">La Rivers, 1971</bibRefCitation>
(New
<collectingCountry id="12E07609FF83FFDDFCB486E70487FE7B" box="[880,967,448,470]" name="Guinea-Bissau" pageId="14" pageNumber="1259">Guinea</collectingCountry>
and the
<collectingCountry id="12E07609FF83FFDDFBF386E70382FE7B" box="[1079,1218,448,470]" name="Philippines" pageId="14" pageNumber="1259">Philippines</collectingCountry>
) and Tanycricini
<bibRefCitation id="0E664B68FF83FFDDFCED86F804B9FE59" author="La Rivers I" box="[809,1017,479,501]" pageId="14" pageNumber="1259" pagination="1 - 120" refId="ref22462" refString="La Rivers I. 1971. Studies of Naucoridae (Hemiptera). Biological Society of Nevada Memoirs 2: 1 - 120." type="journal article" year="1971">La Rivers, 1971</bibRefCitation>
(New
<collectingCountry id="12E07609FF83FFDDFB9286F803F8FE58" box="[1110,1208,479,501]" name="Guinea-Bissau" pageId="14" pageNumber="1259">Guinea</collectingCountry>
). The taxonomic composition of Sagocorini has undergone recent changes because of considerations of evolutionary relationships based on characters related to male genitalia and head structure. The Philippine endemic genera
<taxonomicName id="ADF74D1AFF82FFDCFF3981FA0505F95F" authority="Usinger, 1937" authorityName="Usinger" authorityYear="1937" box="[253,581,1757,1779]" class="Insecta" family="Naucoridae" genus="Asthenocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF82FFDCFF3981FA06D5F95F" box="[253,405,1757,1778]" italics="true" pageId="15" pageNumber="1260">Asthenocoris</emphasis>
Usinger, 1937
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF82FFDCFD9781FA056CF8BD" authority="D. Polhemus &amp; J. Polhemus, 1987" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="1987" class="Insecta" family="Naucoridae" genus="Philippinocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF82FFDCFD9781FA044BF95F" box="[595,779,1757,1778]" italics="true" pageId="15" pageNumber="1260">Philippinocoris</emphasis>
D. Polhemus &amp; J.
<bibRefCitation id="0E664B68FF82FFDCFEB081DC056CF8BD" author="Polhemus DA &amp; Polhemus JT" box="[372,556,1787,1809]" pageId="15" pageNumber="1260" pagination="265 - 269" refId="ref23665" refString="Polhemus DA, Polhemus JT. 1987. A new genus of Naucoridae (Hemiptera) from the Philippines, with comments on zoogeography. Pan-Pacific Entomologist 63: 265 - 269." type="journal article" year="1987">Polhemus, 1987</bibRefCitation>
</taxonomicName>
(as
<taxonomicName id="ADF74D1AFF82FFDCFD9A81DB058FF8BC" authorityName="Montandon" authorityYear="1911" box="[606,719,1788,1809]" class="Insecta" family="Naucoridae" genus="Sagocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF82FFDCFD9A81DB058FF8BC" box="[606,719,1788,1809]" italics="true" pageId="15" pageNumber="1260">Sagocoris</emphasis>
</taxonomicName>
) and
<taxonomicName id="ADF74D1AFF82FFDCFF67803D06ACF89D" authority="La Rivers, 1969" authorityName="La Rivers" authorityYear="1969" box="[163,492,1818,1840]" class="Insecta" family="Naucoridae" genus="Stalocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF82FFDCFF67803D065EF882" box="[163,286,1818,1839]" italics="true" pageId="15" pageNumber="1260">Stalocoris</emphasis>
<bibRefCitation id="0E664B68FF82FFDCFEEC803D06ACF89D" author="La Rivers I" box="[296,492,1818,1840]" pageId="15" pageNumber="1260" pagination="1 - 12" refId="ref22401" refString="La Rivers I. 1969. New naucorid taxa (Hemiptera). Biological Society of Nevada Occasional Papers 20: 1 - 12." type="journal article" year="1969">La Rivers, 1969</bibRefCitation>
</taxonomicName>
were originally placed in Sagocorini, but later transferred to
<taxonomicName id="ADF74D1AFF82FFDCFDBB801E067FF8C1" authority="(Zettel, 2001)" authorityName="LEACH" authorityYear="1815" baseAuthorityName="Zettel" baseAuthorityYear="2001" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">
Naucorinae (
<bibRefCitation id="0E664B68FF82FFDCFF6F80700675F8C1" author="Zettel H" box="[171,309,1879,1901]" pageId="15" pageNumber="1260" pagination="1085 - 1095" refId="ref25492" refString="Zettel H. 2001. Die indische Zwergschwimmwanze Nanonaucoris gen. n., eine neue Gattung der Naucorini (Heteroptera: Naucoridae) aus Sudindien. Linzer biologische Beitrage 33: 1085 - 1095." type="journal article" year="2001">Zettel, 2001</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="3E886611FF83FFDDFF5481C206BFF8FE" ID-DOI="http://doi.org/10.5281/zenodo.6994631" ID-Zenodo-Dep="6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="14" pageNumber="1259" startId="14.[144,226,1765,1787]" targetBox="[144,1424,676,1726]" targetPageId="14">
<paragraph id="6A483699FF83FFDDFF5481C206BFF8FE" blockId="14.[144,1424,1765,1875]" pageId="14" pageNumber="1259">
<emphasis id="5883EA8BFF83FFDDFF5481C2064CF957" bold="true" box="[144,268,1765,1787]" pageId="14" pageNumber="1259">Figure 10.</emphasis>
Habitus examples of each subfamily:
<taxonomicName id="ADF74D1AFF83FFDDFD7F81C20403F956" authorityName="Stal" authorityYear="1862" box="[699,835,1765,1787]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF83FFDDFC8B81C20364F956" authorityName="Stal" authorityYear="1862" box="[847,1060,1765,1787]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="5883EA8BFF83FFDDFC8B81C20364F956" box="[847,1060,1765,1787]" italics="true" pageId="14" pageNumber="1259">Ambrysus signoreti</emphasis>
</taxonomicName>
(a),
<taxonomicName id="ADF74D1AFF83FFDDFB9C81C20241F956" authorityName="MONTANDON" authorityYear="1897" box="[1112,1281,1765,1787]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1260" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF83FFDDFAC981C20653F8B4" authorityName="D. Polhemus, J. Polhemus &amp; Sites" authorityYear="2008" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="thailandana">
<emphasis id="5883EA8BFF83FFDDFAC981C20653F8B4" italics="true" pageId="14" pageNumber="1259">Cheirochela thailandana</emphasis>
</taxonomicName>
(b),
<taxonomicName id="ADF74D1AFF83FFDDFEFA802406A0F8B4" baseAuthorityName="Reynoso-Velasco &amp; Sites" baseAuthorityYear="2021" box="[318,480,1795,1817]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF83FFDDFE2C802405B1F8B4" authorityName="Usinger" authorityYear="1947" box="[488,753,1795,1817]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="hungerfordi">
<emphasis id="5883EA8BFF83FFDDFE2C802405B1F8B4" box="[488,753,1795,1817]" italics="true" pageId="14" pageNumber="1259">Cryphocricos hungerfordi</emphasis>
</taxonomicName>
(c),
<taxonomicName id="ADF74D1AFF83FFDDFCDF802404D2F8B4" authorityName="Sites" authorityYear="2022" box="[795,914,1795,1817]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1261" phylum="Arthropoda" rank="subFamily" status="stat. nov." subFamily="Ilyocorinae">Ilyocorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF83FFDDFC5E80240324F8B4" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[922,1124,1795,1817]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="cimicoides">
<emphasis id="5883EA8BFF83FFDDFC5E80240324F8B4" box="[922,1124,1795,1817]" italics="true" pageId="14" pageNumber="1259">Ilyocoris cimicoides</emphasis>
</taxonomicName>
(d),
<taxonomicName id="ADF74D1AFF83FFDDFB558024025BF8B4" authorityName="STAL" authorityYear="1876" box="[1169,1307,1795,1817]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF83FFDDFAE7802407A7F89B" baseAuthorityName="Stal" baseAuthorityYear="1855" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="limicola">
<emphasis id="5883EA8BFF83FFDDFAE7802407A7F89B" italics="true" pageId="14" pageNumber="1259">Laccocoris limicola</emphasis>
</taxonomicName>
(e),
<taxonomicName id="ADF74D1AFF83FFDDFED3800706EDF89B" authorityName="STAL" authorityYear="1876" box="[279,429,1824,1846]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1267" phylum="Arthropoda" rank="subFamily" subFamily="Limnocorinae">Limnocorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF83FFDDFE73800705DFF89B" authorityName="Montandon" authorityYear="1898" box="[439,671,1824,1846]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis id="5883EA8BFF83FFDDFE73800705DFF89B" box="[439,671,1824,1846]" italics="true" pageId="14" pageNumber="1259">Limnocoris inornatus</emphasis>
</taxonomicName>
(f),
<taxonomicName id="ADF74D1AFF83FFDDFD0F8007041CF89B" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[715,860,1824,1846]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Macrocorinae">Macrocorinae</taxonomicName>
<taxonomicNameLabel id="43B057F0FF83FFDDFCA0800704AAF89B" box="[868,1002,1824,1846]" pageId="14" pageNumber="1259" rank="subFamily">subfam. nov.</taxonomicNameLabel>
:
<taxonomicName id="ADF74D1AFF83FFDDFC338007039DF89B" authorityName="Signoret" authorityYear="1861" box="[1015,1245,1824,1846]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="flavicollis">
<emphasis id="5883EA8BFF83FFDDFC338007039DF89B" box="[1015,1245,1824,1846]" italics="true" pageId="14" pageNumber="1259">Macrocoris flavicollis</emphasis>
</taxonomicName>
(g),
<taxonomicName id="ADF74D1AFF83FFDDFACA800702CDF89B" authorityName="LEACH" authorityYear="1815" box="[1294,1421,1824,1846]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF83FFDDFF54801A0695F8FE" authority="(Thailand)" authorityName=", Zettel" authorityYear="2001" baseAuthorityName="Thailand" box="[144,469,1853,1875]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF83FFDDFF54801A0620F8FE" box="[144,352,1853,1875]" italics="true" pageId="14" pageNumber="1259">Naucoris scutellaris</emphasis>
(Thailand)
</taxonomicName>
(h).
</paragraph>
</caption>
<caption id="3E886611FF82FFDCFF66812302D1F9DF" ID-DOI="http://doi.org/10.5281/zenodo.6994635" ID-Zenodo-Dep="6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="15" pageNumber="1260" startId="15.[162,240,1540,1562]" targetBox="[163,1443,194,1500]" targetPageId="15">
<paragraph id="6A483699FF82FFDCFF66812302D1F9DF" blockId="15.[162,1443,1540,1650]" pageId="15" pageNumber="1260">
<emphasis id="5883EA8BFF82FFDCFF6681230657F9B4" bold="true" box="[162,279,1540,1562]" pageId="15" pageNumber="1260">Figure 11.</emphasis>
Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily:
<taxonomicName id="ADF74D1AFF82FFDCFEDF810606E0F99A" authorityName="Stal" authorityYear="1862" box="[283,416,1569,1591]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF82FFDCFE6F81060530F99A" authorityName="Usinger" authorityYear="1946" box="[427,624,1569,1591]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="5883EA8BFF82FFDCFE6F81060530F99A" box="[427,624,1569,1591]" italics="true" pageId="15" pageNumber="1260">Ambrysus lunatus</emphasis>
</taxonomicName>
(a),
<taxonomicName id="ADF74D1AFF82FFDCFD6781060408F99A" authorityName="MONTANDON" authorityYear="1897" box="[675,840,1569,1591]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF82FFDCFC9081060373F99A" authorityName="D. Polhemus" authorityYear="1986" box="[852,1075,1569,1591]" class="Insecta" family="Naucoridae" genus="Coptocatus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="species" species="kinabalu">
<emphasis id="5883EA8BFF82FFDCFC9081060373F99A" box="[852,1075,1569,1591]" italics="true" pageId="15" pageNumber="1260">Coptocatus kinabalu</emphasis>
</taxonomicName>
(b),
<taxonomicName id="ADF74D1AFF82FFDCFBA28106024DF99A" baseAuthorityName="Reynoso-Velasco &amp; Sites" baseAuthorityYear="2021" box="[1126,1293,1569,1591]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF82FFDCFADC8106065CF9F9" authorityName="Usinger" authorityYear="1947" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="species" species="hungerfordi">
<emphasis id="5883EA8BFF82FFDCFADC8106065CF9F9" italics="true" pageId="15" pageNumber="1260">Cryphocricos hungerfordi</emphasis>
</taxonomicName>
(c),
<taxonomicName id="ADF74D1AFF82FFDCFE87811906F6F9F9" authorityName="Sites" authorityYear="2022" box="[323,438,1598,1620]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1261" phylum="Arthropoda" rank="subFamily" status="stat. nov." subFamily="Ilyocorinae">Ilyocorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF82FFDCFE79811905C1F9F9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[445,641,1598,1620]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="species" species="cimicoides">
<emphasis id="5883EA8BFF82FFDCFE79811905C1F9F9" box="[445,641,1598,1620]" italics="true" pageId="15" pageNumber="1260">Ilyocoris cimicoides</emphasis>
</taxonomicName>
(d),
<taxonomicName id="ADF74D1AFF82FFDCFD6F81190471F9F9" authorityName="STAL" authorityYear="1876" box="[683,817,1598,1620]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF82FFDCFCFD811904B6F9F9" authorityName="Stal" authorityYear="1856" box="[825,1014,1598,1620]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="species" species="spurcus">
<emphasis id="5883EA8BFF82FFDCFCFD811904B6F9F9" box="[825,1014,1598,1620]" italics="true" pageId="15" pageNumber="1260">Laccocoris spurcus</emphasis>
</taxonomicName>
(e),
<taxonomicName id="ADF74D1AFF82FFDCFBDB811903EEF9F9" authorityName="STAL" authorityYear="1876" box="[1055,1198,1598,1620]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1267" phylum="Arthropoda" rank="subFamily" subFamily="Limnocorinae">Limnocorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF82FFDCFB71811902C3F9F9" authorityName="Montandon" authorityYear="1897" box="[1205,1411,1598,1620]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="species" species="ovatulus">
<emphasis id="5883EA8BFF82FFDCFB71811902C3F9F9" box="[1205,1411,1598,1620]" italics="true" pageId="15" pageNumber="1260">Limnocoris ovatulus</emphasis>
</taxonomicName>
(f),
<taxonomicName id="ADF74D1AFF82FFDCFF67817B066DF9DF" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[163,301,1628,1650]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Macrocorinae">Macrocorinae</taxonomicName>
<taxonomicNameLabel id="43B057F0FF82FFDCFEF6817B06F1F9DF" box="[306,433,1628,1650]" pageId="15" pageNumber="1260" rank="subFamily">subfam. nov.</taxonomicNameLabel>
:
<taxonomicName id="ADF74D1AFF82FFDCFE7F817B05D3F9DF" authorityName="Signoret" authorityYear="1861" box="[443,659,1628,1650]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="species" species="flavicollis">
<emphasis id="5883EA8BFF82FFDCFE7F817B05D3F9DF" box="[443,659,1628,1650]" italics="true" pageId="15" pageNumber="1260">Macrocoris flavicollis</emphasis>
</taxonomicName>
(g),
<taxonomicName id="ADF74D1AFF82FFDCFD7A817B0476F9DF" authorityName="LEACH" authorityYear="1815" box="[702,822,1628,1650]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF82FFDCFCFB817B0482F9DF" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[831,962,1628,1650]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF82FFDCFCFB817B0482F9DF" box="[831,962,1628,1650]" italics="true" pageId="15" pageNumber="1260">Afronaucoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF82FFDCFC03817B0362F9DF" box="[967,1058,1628,1650]" pageId="15" pageNumber="1260" rank="genus">gen. nov.</taxonomicNameLabel>
:
<taxonomicName id="ADF74D1AFF82FFDCFBED817B0228F9DF" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1899" box="[1065,1384,1628,1650]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="5883EA8BFF82FFDCFBED817B0228F9DF" box="[1065,1384,1628,1650]" italics="true" pageId="15" pageNumber="1260">Afronaucoris madagascariensis</emphasis>
</taxonomicName>
(h).
</paragraph>
</caption>
<paragraph id="6A483699FF82FFC3FCFF819905BAFE9A" blockId="15.[827,1443,1726,1901]" lastBlockId="16.[145,762,197,311]" lastPageId="16" lastPageNumber="1261" pageId="15" pageNumber="1260">
<emphasis id="5883EA8BFF82FFDCFCFF81990367F979" box="[827,1063,1726,1748]" italics="true" pageId="15" pageNumber="1260">Revised taxonomy:</emphasis>
With the taxonomic changes proposed herein, the subfamily
<taxonomicName id="ADF74D1AFF82FFDCFB0E81FA02C3F95E" authorityName="MONTANDON" authorityYear="1897" box="[1226,1411,1757,1779]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
is here restricted to only the previously recognized tribe
<taxonomicName id="ADF74D1AFF82FFDCFC47803D0370F89D" box="[899,1072,1818,1840]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="tribe" tribe="Cheirochelini">Cheirochelini</taxonomicName>
, comprising the Indochinese genera
<taxonomicName id="ADF74D1AFF82FFDCFC56801E035CF8E3" authorityName="Hope" authorityYear="1840" box="[914,1052,1849,1870]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF82FFDCFC56801E035CF8E3" box="[914,1052,1849,1870]" italics="true" pageId="15" pageNumber="1260">Cheirochela</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF82FFDCFB9C801E0396F8E3" authorityName="Montandon" authorityYear="1897" box="[1112,1238,1849,1870]" class="Insecta" family="Naucoridae" genus="Gestroiella" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF82FFDCFB9C801E0396F8E3" box="[1112,1238,1849,1870]" italics="true" pageId="15" pageNumber="1260">Gestroiella</emphasis>
</taxonomicName>
and the Bornean
<taxonomicName id="ADF74D1AFF82FFDCFCFF807F04F8F8C0" authorityName="Montandon" authorityYear="1909" box="[827,952,1880,1901]" class="Insecta" family="Naucoridae" genus="Coptocatus" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="1260" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF82FFDCFCFF807F04F8F8C0" box="[827,952,1880,1901]" italics="true" pageId="15" pageNumber="1260">Coptocatus</emphasis>
</taxonomicName>
. The remaining seven genera in the former tribes Sagocorini and Tanycricini were recovered in a distantly related clade and are here transferred to
<taxonomicName id="ADF74D1AFF9DFFC3FF558625065AFEB5" authorityName="LEACH" authorityYear="1815" box="[145,282,258,280]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
. As such,
<taxonomicName id="ADF74D1AFF9DFFC3FE498625057CFEB5" authorityName="MONTANDON" authorityYear="1897" box="[397,572,258,280]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1260" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
is retained as a valid subfamily but with only three constituent genera.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FF5586470543FBB5" pageId="16" pageNumber="1261" type="diagnosis">
<paragraph id="6A483699FF9DFFC3FF5586470543FBB5" blockId="16.[145,761,352,1048]" pageId="16" pageNumber="1261">
<emphasis id="5883EA8BFF9DFFC3FF5586470647FED8" box="[145,263,352,373]" italics="true" pageId="16" pageNumber="1261">Diagnosis:</emphasis>
Diagnostic features for the subfamily were provided by
<bibRefCitation id="0E664B68FF9DFFC3FEEC86590577FE39" author="Polhemus DA &amp; Polhemus JT &amp; Sites RW" box="[296,567,382,404]" pageId="16" pageNumber="1261" pagination="255 - 279" refId="ref23766" refString="Polhemus DA, Polhemus JT, Sites RW. 2008. A revision of the Indochinese genera Cheirochela and Gestroiella (Heteroptera: Naucoridae), and a review of the tribe Cheirochelini. Raffles Bulletin of Zoology 56: 255 - 279." type="journal article" year="2008">
Polhemus
<emphasis id="5883EA8BFF9DFFC3FE62865806A3FE3E" box="[422,483,382,404]" italics="true" pageId="16" pageNumber="1261">et al.</emphasis>
(2008)
</bibRefCitation>
in a revision of the tribe
<taxonomicName id="ADF74D1AFF9DFFC3FF3986BA06D8FE1E" box="[253,408,413,435]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="tribe" tribe="Cheirochelini">Cheirochelini</taxonomicName>
as the following: the vertex is strongly produced posteriorly; the antennae are stout, with segments 1–3 subequal in length and about as wide as long, and segment 4 slightly longer; the antennae are set in elongate, concave sockets beneath the eyes; the rostrum and labrum are recessed into a deep cavity under the head; the labrum is reduced and barely visible at the base of the rostrum; the maxillary plates adjacent to the rostral cavity lie in a horizontal orientation; the head is strongly prolonged forward of the eyes and rostral cavity, forming an anteclypeal shelf; the fore tarsus and claw are fused to form a single segment; fore and hind tibiae bear pads of short, dense setae ventrally at their apices; abdominal medio- and laterosternites are fused to form undivided sterna on segments IV– VI, lateral sections of these plates adjacent to spiracles bear groups of small, ovate, glabrous depressions; the parameres are symmetrical, overlapping and extending to approximately half the length of the aedeagus; the aedeagus is symmetrical with the central portion sclerotized and apex lyre shaped.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FF558366060DFB5D" pageId="16" pageNumber="1261" type="discussion">
<paragraph id="6A483699FF9DFFC3FF558366060DFB5D" blockId="16.[145,761,1089,1264]" pageId="16" pageNumber="1261">
<emphasis id="5883EA8BFF9DFFC3FF5583660653FBFB" box="[145,275,1089,1110]" italics="true" pageId="16" pageNumber="1261">Comments:</emphasis>
The genera
<taxonomicName id="ADF74D1AFF9DFFC3FE6B8366057BFBFB" authorityName="Hope" authorityYear="1840" box="[431,571,1089,1110]" class="Insecta" family="Naucoridae" genus="Cheirochela" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FE6B8366057BFBFB" box="[431,571,1089,1110]" italics="true" pageId="16" pageNumber="1261">Cheirochela</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9DFFC3FDBD836605B9FBFB" authorityName="Montandon" authorityYear="1897" box="[633,761,1089,1110]" class="Insecta" family="Naucoridae" genus="Gestroiella" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FDBD836605B9FBFB" box="[633,761,1089,1110]" italics="true" pageId="16" pageNumber="1261">Gestroiella</emphasis>
</taxonomicName>
occur from southern
<collectingCountry id="12E07609FF9DFFC3FE528347069FFBDB" box="[406,479,1120,1142]" name="China" pageId="16" pageNumber="1261">China</collectingCountry>
throughout Indochina west to
<collectingCountry id="12E07609FF9DFFC3FF2C83590666FB39" box="[232,294,1150,1172]" name="India" pageId="16" pageNumber="1261">India</collectingCountry>
;
<taxonomicName id="ADF74D1AFF9DFFC3FEF4835806EDFB39" authorityName="Montandon" authorityYear="1909" box="[304,429,1151,1172]" class="Insecta" family="Naucoridae" genus="Coptocatus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FEF4835806EDFB39" box="[304,429,1151,1172]" italics="true" pageId="16" pageNumber="1261">Coptocatus</emphasis>
</taxonomicName>
is restricted to Borneo and is found in all three countries represented on the island. These are large, impressive insects occurring in fastflowing streams.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="E25E878FFF87FFDFFC5384410333F8F0" ID-DOI="http://doi.org/10.5281/zenodo.6994605" ID-GBIF-Taxon="199315250" ID-Zenodo-Dep="6994605" LSID="urn:lsid:plazi:treatment:E25E878FFF87FFDFFC5384410333F8F0" httpUri="http://treatment.plazi.org/id/E25E878FFF87FFDFFC5384410333F8F0" lastPageId="12" lastPageNumber="1256" pageId="10" pageNumber="1255">
<subSubSection id="22ED6512FF87FFD9FC5384410262FCD2" box="[919,1314,870,895]" pageId="10" pageNumber="1255" type="nomenclature">
<paragraph id="6A483699FF87FFD9FC5384410262FCD2" blockId="10.[919,1314,825,895]" box="[919,1314,870,895]" pageId="10" pageNumber="1255">
FAMILY
<taxonomicName id="ADF74D1AFF87FFD9FC3784410262FCD2" ID-CoL="624PT" authority="LEACH, 1815" authorityName="LEACH" authorityYear="1815" box="[1011,1314,870,895]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="1255" phylum="Arthropoda" rank="family">
NAUCORIDAE
<bibRefCitation id="0E664B68FF87FFD9FB5484410262FCD2" author="Leach WE" box="[1168,1314,870,895]" pageId="10" pageNumber="1255" pagination="57 - 192" refId="ref22574" refString="Leach WE. 1815. Entomology. In: Brewster D, ed. The Edinburgh encyclopedia, Vol. 9. Edinburgh: William Blackwood, 57 - 192." type="book chapter" year="1815">LEACH, 1815</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF87FFD9FCED84B0038FFC01" box="[809,1231,918,940]" pageId="10" pageNumber="1255" type="type_taxon">
<paragraph id="6A483699FF87FFD9FCED84B0038FFC01" blockId="10.[809,1231,918,940]" box="[809,1231,918,940]" pageId="10" pageNumber="1255">
<emphasis id="5883EA8BFF87FFD9FCED84B004EDFC01" box="[809,941,919,940]" italics="true" pageId="10" pageNumber="1255">
<typeStatus id="B54C883BFF87FFD9FCED84B0041EFC01" box="[809,862,919,940]" pageId="10" pageNumber="1255">Type</typeStatus>
genus:
</emphasis>
<taxonomicName id="ADF74D1AFF87FFD9FC7D84B00388FC01" ID-CoL="5XV9" authority="Geoffroy, 1762" authorityName="Geoffroy" authorityYear="1762" box="[953,1224,918,940]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="1255" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF87FFD9FC7D84B00360FC01" box="[953,1056,919,940]" italics="true" pageId="10" pageNumber="1255">Naucoris</emphasis>
<bibRefCitation id="0E664B68FF87FFD9FBE384B10388FC01" author="Geoffroy EL" box="[1063,1224,918,940]" pageId="10" pageNumber="1255" refId="ref21927" refString="Geoffroy EL. 1762. Histoire abregee des insectes qui se trouvent aux environs de Paris. Vol. 1. Paris: Durand." type="book" year="1762">Geoffroy, 1762</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF87FFD9FCED84F10207F903" pageId="10" pageNumber="1255" type="discussion">
<paragraph id="6A483699FF87FFD9FCED84F102D1F9BF" blockId="10.[809,1426,981,1555]" pageId="10" pageNumber="1255">
<emphasis id="5883EA8BFF87FFD9FCED84F10346FC47" box="[809,1030,981,1003]" italics="true" pageId="10" pageNumber="1255">Taxonomic history:</emphasis>
The group was first proposed by
<bibRefCitation id="0E664B68FF87FFD9FCED84D304FAFBA7" author="Leach WE" box="[809,954,1012,1034]" pageId="10" pageNumber="1255" pagination="57 - 192" refId="ref22574" refString="Leach WE. 1815. Entomology. In: Brewster D, ed. The Edinburgh encyclopedia, Vol. 9. Edinburgh: William Blackwood, 57 - 192." type="book chapter" year="1815">Leach (1815)</bibRefCitation>
as Naucorida, an incorrect spelling of the family name. This and other family names placed on the Official Index of Rejected and Invalid Family-Group Names by Opinion 681 (1963) include Naucorides (
<bibRefCitation id="0E664B68FF87FFD9FA8283770420FB2E" author="Fallen CF" pageId="10" pageNumber="1255" refId="ref21882" refString="Fallen CF. 1814. Specimen novam Hemiptera disponendi methodum exhibens. Lund: Litteris Berlingianus." type="book" year="1814">Fallén, 1814</bibRefCitation>
; see
<bibRefCitation id="0E664B68FF87FFD9FC58834A03D6FB2E" author="Stys P &amp; Jansson A" box="[924,1174,1133,1157]" pageId="10" pageNumber="1255" pagination="1 - 44" refId="ref25103" refString="Stys P, Jansson A. 1988. Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica 50: 1 - 44." type="journal article" year="1988">Štys &amp; Jansson, 1988</bibRefCitation>
), Naucoridea (
<bibRefCitation id="0E664B68FF87FFD9FA8783490421FB0E" author="Fieber FX" pageId="10" pageNumber="1255" refId="ref21899" refString="Fieber FX. 1851. Genera Hydrocoridum secundum ordinem naturalem in familias disposita. Actis Regiae Bohemicae Societatis Scientiarum. Prague: J. G. Calve." type="book" year="1851">Fieber, 1851</bibRefCitation>
), Naucoridi (
<bibRefCitation id="0E664B68FF87FFD9FBC583AA03E6FB0F" author="Acloque A" box="[1025,1190,1165,1187]" pageId="10" pageNumber="1255" refId="ref21394" refString="Acloque A. 1897. Faune de France contenant la description des especes indigenes disposees en tableaux analytiques: orthopteres, neuropteres, hymenopteres, lepidopteres, hemipteres, dipteres, aphanipteres, thysanopteres, rhipipteres. Paris: J. B. Bailliere &amp; fils." type="book" year="1897">Acloque, 1897</bibRefCitation>
),
<taxonomicName id="ADF74D1AFF87FFD9FB7B83AA0428FB6C" authority="(Costa, 1852)" authorityName="POPOV" authorityYear="1970" baseAuthorityName="Costa" baseAuthorityYear="1852" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">
Naucorini (
<bibRefCitation id="0E664B68FF87FFD9FA8383AA0420FB6C" author="Costa A" pageId="10" pageNumber="1255" pagination="6 - 76" refId="ref21577" refString="Costa A. 1852. Cimicum Regni Neapolitani Centuria 7: 6 - 76 + 1 plate. Royal Academy of Naples." type="book chapter" year="1852">
<collectingRegion id="A833F87BFF87FFD9FA8383AA02CCFB0F" box="[1351,1420,1165,1186]" country="Tanzania" name="Pwani" pageId="10" pageNumber="1255">Costa</collectingRegion>
, 1852
</bibRefCitation>
)
</taxonomicName>
and Naucoriseae (
<bibRefCitation id="0E664B68FF87FFD9FB85838B038EFB6C" author="Fieber FX" box="[1089,1230,1195,1218]" pageId="10" pageNumber="1255" refId="ref21899" refString="Fieber FX. 1851. Genera Hydrocoridum secundum ordinem naturalem in familias disposita. Actis Regiae Bohemicae Societatis Scientiarum. Prague: J. G. Calve." type="book" year="1851">Fieber, 1851</bibRefCitation>
). The family has included Aphelocheirinae (e.g.
<bibRefCitation id="0E664B68FF87FFD9FB4C83ED041EFB53" author="Polhemus DA &amp; Polhemus JT" pageId="10" pageNumber="1255" pagination="167 - 300" refId="ref23698" refString="Polhemus DA, Polhemus JT. 1988. The Aphelocheirinae of tropical Asia (Heteroptera: Naucoridae). Raffles Bulletin of Zoology 36: 167 - 300." type="journal article" year="1988">Polhemus &amp; Polhemus, 1988</bibRefCitation>
) and Potamocorinae (
<bibRefCitation id="0E664B68FF87FFD9FB8883CE03A1FB53" author="Usinger RL" box="[1100,1249,1257,1279]" pageId="10" pageNumber="1255" pagination="5 - 16" refId="ref25189" refString="Usinger RL. 1941. Key to the subfamilies of Naucoridae with a generic synopsis of the new subfamily Ambrysinae (Hemiptera). Annals of the Entomological Society of America 34: 5 - 16." type="journal article" year="1941">Usinger, 1941</bibRefCitation>
) as subfamilies, although most recent authors have considered these to be distinct families related to
<taxonomicName id="ADF74D1AFF87FFD9FBA9820103ACFA91" authorityName="LEACH" authorityYear="1815" box="[1133,1260,1318,1340]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="1255" phylum="Arthropoda" rank="family">Naucoridae</taxonomicName>
in superfamily
<taxonomicName id="ADF74D1AFF87FFD9FCED826204F6FAF6" authorityName="Leach" authorityYear="1815" box="[809,950,1349,1371]" pageId="10" pageNumber="1255" rank="superFamily" superFamily="Naucoroidea">Naucoroidea</taxonomicName>
. A largely unaccepted concept was proposed by
<bibRefCitation id="0E664B68FF87FFD9FC8882430341FAD4" author="De Carlo JA" box="[844,1025,1380,1402]" pageId="10" pageNumber="1255" pagination="159 - 166" refId="ref21730" refString="De Carlo JA. 1971. Valor sistematico del estudio del aparato genital macho en los hemipteros acuaticos y semicuaticos. Division de la familia Naucoridae en dos familias. Caracteristica externa de la capsula genital. Revista de la Sociedad Entomologica Argentina 33: 159 - 166." type="journal article" year="1971">De Carlo (1971)</bibRefCitation>
in which he split the family based on male and female reproductive structures into separate Old and New World families as
<taxonomicName id="ADF74D1AFF87FFD9FB2782860222FA1A" authorityName="LEACH" authorityYear="1815" box="[1251,1378,1441,1463]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="1255" phylum="Arthropoda" rank="family">Naucoridae</taxonomicName>
and Pelocoridae, respectively; although the New World family name Limnocoridae was adopted in place of Pelocoridae because of the Principle of Priority (e.g. López Ruf, 1987).
</paragraph>
<paragraph id="6A483699FF87FFD9FCED811B0207F903" blockId="10.[809,1424,1596,1710]" pageId="10" pageNumber="1255">
<emphasis id="5883EA8BFF87FFD9FCED811B0358F9FC" box="[809,1048,1596,1617]" italics="true" pageId="10" pageNumber="1255">Revised taxonomy:</emphasis>
The family was found to be monophyletic here, thus no changes to the family level taxonomy are proposed. However, eight subfamilies (
<figureCitation id="F2CC2A1CFF87FFD9FCF581BF043DF900" box="[817,893,1688,1710]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="10" pageNumber="1255">Fig. 10</figureCitation>
) are now recognized as described below.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF87FFD8FCED81F00352F9C0" lastPageId="11" lastPageNumber="1256" pageId="10" pageNumber="1255" type="diagnosis">
<paragraph id="6A483699FF87FFD8FCED81F00352F9C0" blockId="10.[809,1426,1750,1895]" lastBlockId="11.[827,1442,1593,1645]" lastPageId="11" lastPageNumber="1256" pageId="10" pageNumber="1255">
<emphasis id="5883EA8BFF87FFD9FCED81F004DFF941" box="[809,927,1751,1772]" italics="true" pageId="10" pageNumber="1255">Diagnosis:</emphasis>
Members of
<taxonomicName id="ADF74D1AFF87FFD9FBF081F103F6F941" authorityName="LEACH" authorityYear="1815" box="[1076,1206,1750,1772]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="1255" phylum="Arthropoda" rank="family">Naucoridae</taxonomicName>
are mostly aquatic, generally flattened, ovate or slightly elongate and with retentorial forelegs (
<bibRefCitation id="0E664B68FF87FFD9FBB5803302C0F884" author="Sites RW &amp; Nichols BJ" box="[1137,1408,1812,1834]" pageId="10" pageNumber="1255" pagination="800 - 808" refId="ref24619" refString="Sites RW, Nichols BJ. 1990. Life history and descriptions of immature stages of Ambrysus lunatus lunatus (Hemiptera: Naucoridae). Annals of the Entomological Society of America 83: 800 - 808." type="journal article" year="1990">
<taxonomicName id="ADF74D1AFF87FFD9FBB5803303EEF887" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1137,1198,1812,1834]" genus="Sites" pageId="10" pageNumber="1270" rank="genus" status="gen. nov.">Sites</taxonomicName>
&amp; Nichols, 1990
</bibRefCitation>
). They can be similar in appearance to
<taxonomicName id="ADF74D1AFF87FFD9FB1E801502CDF8E5" box="[1242,1421,1842,1864]" class="Insecta" family="Belostomatidae" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="1255" phylum="Arthropoda" rank="family">Belostomatidae</taxonomicName>
, although substantially smaller and without the caudal respiratory tubes. More specifically, antennae usually are four-segmented and short; the labium is short, reaching only to near the fore coxae; the membrane of the hemelytra is without venation; the forefemur is usually greatly enlarged; the foretarsus is usually fixed and immobile with a single fixed pretarsal claw; the meso- and metatibiae and meso- and metatarsi usually are fringed with natatorial setae; metathoracic scent glands are ventral in adults and between abdominal terga 3 and
<quantity id="AD0F9B7CFF86FFD8FEF0806A0622F8CE" box="[308,354,1869,1891]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="11" pageNumber="1256" unit="in" value="4.0">4 in</quantity>
nymphs; and the genital capsule is reflexed 180 ° so the aedeagus and parameres are directed anteriorly.
</paragraph>
</subSubSection>
<caption id="3E886611FF86FFD8FF67859F03F9FD63" ID-DOI="http://doi.org/10.5281/zenodo.6994623" ID-Zenodo-Dep="6994623" box="[163,1209,696,718]" httpUri="https://zenodo.org/record/6994623/files/figure.png" pageId="11" pageNumber="1256" startId="11.[163,241,696,718]" targetBox="[399,1207,195,656]" targetPageId="11">
<paragraph id="6A483699FF86FFD8FF67859F03F9FD63" blockId="11.[163,1209,696,718]" box="[163,1209,696,718]" pageId="11" pageNumber="1256">
<emphasis id="5883EA8BFF86FFD8FF67859F064BFD63" bold="true" box="[163,267,696,718]" pageId="11" pageNumber="1256">Figure 7.</emphasis>
Valvulae of female
<taxonomicName id="ADF74D1AFF86FFD8FE19859F05E8FD63" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[477,680,696,718]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="1256" phylum="Arthropoda" rank="species" species="cimicoides">
<emphasis id="5883EA8BFF86FFD8FE19859F05E8FD63" box="[477,680,696,718]" italics="true" pageId="11" pageNumber="1256">Ilyocoris cimicoides</emphasis>
</taxonomicName>
with dentate margins. vlf = valvifer, vlv = valvula.
</paragraph>
</caption>
<caption id="3E886611FF86FFD8FF6782E90507F9AF" ID-DOI="http://doi.org/10.5281/zenodo.6994627" ID-Zenodo-Dep="6994627" httpUri="https://zenodo.org/record/6994627/files/figure.png" pageId="11" pageNumber="1256" startId="11.[163,242,1486,1508]" targetBox="[163,1443,800,1447]" targetPageId="11">
<paragraph id="6A483699FF86FFD8FF6782E90507F9AF" blockId="11.[163,1444,1486,1538]" pageId="11" pageNumber="1256">
<emphasis id="5883EA8BFF86FFD8FF6782E9064CFA4E" bold="true" box="[163,268,1486,1508]" pageId="11" pageNumber="1256">Figure 8.</emphasis>
Mesofemora of
<taxonomicName id="ADF74D1AFF86FFD8FE7D82E805C6FA48" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[441,646,1487,1509]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="1256" phylum="Arthropoda" rank="species" species="cimicoides">
<emphasis id="5883EA8BFF86FFD8FE7D82E805C6FA48" box="[441,646,1487,1509]" italics="true" pageId="11" pageNumber="1256">Ilyocoris cimicoides</emphasis>
</taxonomicName>
(a) and
<taxonomicName id="ADF74D1AFF86FFD8FD1A82E8043DFA48" box="[734,893,1487,1509]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="1256" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="5883EA8BFF86FFD8FD1A82E80418FA48" box="[734,856,1487,1509]" italics="true" pageId="11" pageNumber="1256">Placomerus</emphasis>
sp.
</taxonomicName>
(b) showing the posterodorsal margin flattened and fringed with a dense brush-line of hairs.
</paragraph>
</caption>
<subSubSection id="22ED6512FF86FFDFFCFF81B10333F8F0" lastPageId="12" lastPageNumber="1257" pageId="11" pageNumber="1256" type="discussion">
<paragraph id="6A483699FF86FFDFFCFF81B10333F8F0" blockId="11.[827,1443,1686,1892]" lastBlockId="12.[809,1425,1803,1886]" lastPageId="12" lastPageNumber="1257" pageId="11" pageNumber="1256">
<emphasis id="5883EA8BFF86FFD8FCFF81B10481F906" box="[827,961,1686,1707]" italics="true" pageId="11" pageNumber="1256">Comments:</emphasis>
The family has been recognized since
<bibRefCitation id="0E664B68FF86FFD8FCFF81940379F967" author="Stys P &amp; Jansson A" box="[827,1081,1715,1739]" pageId="11" pageNumber="1256" pagination="1 - 44" refId="ref25103" refString="Stys P, Jansson A. 1988. Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica 50: 1 - 44." type="journal article" year="1988">Štys &amp; Jansson (1988)</bibRefCitation>
to have five subfamilies loosely defined by a few morphological features with uncertain reliability as being apomorphic and not homoplasious. A concurrent project to determine the evolutionary history of the most species-rich subfamily,
<taxonomicName id="ADF74D1AFF86FFD8FB22800802DDF8E8" baseAuthorityName="Reynoso-Velasco &amp; Sites" baseAuthorityYear="2021" box="[1254,1437,1839,1861]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
, has resulted in the elevation of Ambrysini back to the subfamily rank as originally assigned by
<bibRefCitation id="0E664B68FF81FFDFFD8B802C05B3F88D" author="Usinger RL" box="[591,755,1803,1825]" pageId="12" pageNumber="1257" pagination="5 - 16" refId="ref25189" refString="Usinger RL. 1941. Key to the subfamilies of Naucoridae with a generic synopsis of the new subfamily Ambrysinae (Hemiptera). Annals of the Entomological Society of America 34: 5 - 16." type="journal article" year="1941">Usinger (1941)</bibRefCitation>
. As such, six subfamilies are currently recognized (
<tableCitation id="27750322FF81FFDFFD7F800E07DFF8F0" captionStart="Table 1" captionStartId="2.[146,210,196,217]" captionTargetBox="[145,1381,277,1085]" captionText="Table 1. Current taxonomy of subfamilies, tribes and genera of Naucoridae. Recent taxonomic changes proposed by Reynoso-Velasco &amp; Sites (2021) are indicated with asterisks" httpUri="http://table.plazi.org/id/3E886611FF8FFFD1FF5687E30442FF5A" pageId="12" pageNumber="1257" tableUuid="3E886611FF8FFFD1FF5687E30442FF5A">Table 1</tableCitation>
) as was the case following
<bibRefCitation id="0E664B68FF81FFDFFE18806F05C8F8F0" author="Usinger RL" box="[476,648,1864,1886]" pageId="12" pageNumber="1257" pagination="5 - 16" refId="ref25189" refString="Usinger RL. 1941. Key to the subfamilies of Naucoridae with a generic synopsis of the new subfamily Ambrysinae (Hemiptera). Annals of the Entomological Society of America 34: 5 - 16." type="journal article" year="1941">Usinger (1941)</bibRefCitation>
,
<bibRefCitation id="0E664B68FF81FFDFFD52806F04F0F88C" author="China WE &amp; Miller NCE" box="[662,944,1803,1886]" pageId="12" pageNumber="1257" pagination="257 - 267" refId="ref21513" refString="China WE, Miller NCE. 1955. Check-list of family and subfamily names in Hemiptera-Heteroptera. Annals and Magazine of Natural History 8: 257 - 267." type="journal article" year="1955">
<collectingCountry id="12E07609FF81FFDFFD52806F059CF8F3" box="[662,732,1864,1886]" name="China" pageId="12" pageNumber="1257">China</collectingCountry>
&amp; Miller (1955
</bibRefCitation>
,
<bibRefCitation id="0E664B68FF81FFDFFC7F802C04B1F88C" author="China WE &amp; Miller NCE" box="[955,1009,1803,1825]" pageId="12" pageNumber="1257" pagination="1 - 45" refId="ref21542" refString="China WE, Miller NCE. 1959. Check-list and keys to the families and subfamilies of the Hemiptera-Heteroptera. Bulletin of the British Museum (Natural History) 8: 1 - 45." type="journal article" year="1959">1959</bibRefCitation>
) and
<bibRefCitation id="0E664B68FF81FFDFFBEA802C03A4F88D" author="La Rivers I" box="[1070,1252,1803,1825]" pageId="12" pageNumber="1257" pagination="1 - 120" refId="ref22462" refString="La Rivers I. 1971. Studies of Naucoridae (Hemiptera). Biological Society of Nevada Memoirs 2: 1 - 120." type="journal article" year="1971">La Rivers (1971)</bibRefCitation>
. The taxonomic changes proposed here will increase the number of subfamilies to eight (
<tableCitation id="27750322FF81FFDFFBD7806F0324F8F0" box="[1043,1124,1864,1886]" captionStart="Table 2" captionStartId="13.[164,228,196,217]" captionTargetBox="[163,1434,253,1177]" captionText="Table 2. Revised taxonomy of subfamilies, tribes and genera of Naucoridae" httpUri="http://table.plazi.org/id/3E886611FF80FFDEFF6087E304F9FF77" pageId="12" pageNumber="1257" tableUuid="3E886611FF80FFDEFF6087E304F9FF77">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="9BC40B4E1382E3119B64F4C8B2B2241C" ID-CLB-Dataset="121760" ID-DOI="10.1093/zoolinnean/zlab105" ID-GBIF-Dataset="83d946c2-fa80-4948-9160-16625c240886" ID-ISSN="0024-4082" ID-Zenodo-Dep="6994599" ID-ZooBank="C06A1F94-AF08-4A21-B1F3-A0865FB1A8DF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660324512590" checkinUser="plazi" docAuthor="Sites, Robert W." docDate="2022" docId="E25E878FFF91FFCDFED6817F0416FE9B" docLanguage="en" docName="zlab105.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Heleolaccocoris Sites 2022, GEN. NOV." docType="treatment" docVersion="6" lastPageNumber="1275" masterDocId="1E67FFF7FF8DFFD3FFC487270740FFAD" masterDocTitle="Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)" masterLastPageNumber="1286" masterPageNumber="1245" pageNumber="1273" updateTime="1699366499196" updateUser="ExternalLinkService">
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<mods:title id="FF1457906B6C58C1D51A59F23D519C0A">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:namePart id="7F569CFA19953044354D830B97F13C13">Sites, Robert W.</mods:namePart>
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<mods:date id="21DAC2D3DC02E2AFE6DE84326D41EE41">2022</mods:date>
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<treatment id="E25E878FFF91FFCDFED6817F0416FE9B" ID-DOI="http://doi.org/10.5281/zenodo.6994678" ID-GBIF-Taxon="199315272" ID-Zenodo-Dep="6994678" LSID="urn:lsid:plazi:treatment:E25E878FFF91FFCDFED6817F0416FE9B" httpUri="http://treatment.plazi.org/id/E25E878FFF91FFCDFED6817F0416FE9B" lastPageId="30" lastPageNumber="1275" pageId="28" pageNumber="1273">
<subSubSection id="22ED6512FF91FFCFFED6817F0538F9DD" box="[274,632,1624,1648]" pageId="28" pageNumber="1273" type="nomenclature">
<paragraph id="6A483699FF91FFCFFED6817F0538F9DD" blockId="28.[274,632,1624,1648]" box="[274,632,1624,1648]" pageId="28" pageNumber="1273">
<heading id="310081F5FF91FFCFFED6817F0538F9DD" box="[274,632,1624,1648]" centered="true" fontSize="9" level="2" pageId="28" pageNumber="1273" reason="2">
<taxonomicName id="ADF74D1AFF91FFCFFED6817F06B9F9C3" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[274,505,1624,1647]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="genus" status="gen. nov.">HELEOLACCOCORIS</taxonomicName>
<emphasis id="5883EA8BFF91FFCFFDC4817C0538F9DD" bold="true" box="[512,632,1624,1648]" pageId="28" pageNumber="1273">
<taxonomicNameLabel id="43B057F0FF91FFCFFDC4817C0538F9DD" box="[512,632,1624,1648]" pageId="28" pageNumber="1273" rank="genus">GEN. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="22ED6512FF91FFCDFE8181A40670FC72" lastPageId="30" lastPageNumber="1275" pageId="28" pageNumber="1273" type="description">
<paragraph id="6A483699FF91FFCFFE8181A40505F931" blockId="28.[325,581,1667,1692]" box="[325,581,1667,1692]" pageId="28" pageNumber="1273">
<figureCitation id="F2CC2A1CFF91FFCFFE8181A406DEF936" box="[325,414,1667,1692]" captionStart="Figure 4" captionStartId="9.[163,249,1379,1401]" captionTargetBox="[163,781,195,1339]" captionTargetId="figure-575@9.[163,781,195,1339]" captionTargetPageId="9" captionText="Figure 4. Foretibiae and foretarsi of Laccocorinae: Ctenipocoris asiaticus (a), Heleolaccocoris ovatus (b), Namtokocoris dalanta (c)." figureDoi="http://doi.org/10.5281/zenodo.6994615" httpUri="https://zenodo.org/record/6994615/files/figure.png" pageId="28" pageNumber="1273">FIGS 4B</figureCitation>
,
<figureCitation id="F2CC2A1CFF91FFCFFE6F81A406A4F936" box="[427,484,1667,1692]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="28" pageNumber="1273">5A, B</figureCitation>
,
<figureCitation id="F2CC2A1CFF91FFCFFE3581A4055CF936" box="[497,540,1667,1692]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="28" pageNumber="1273">11E</figureCitation>
,
<figureCitation id="F2CC2A1CFF91FFCFFDED81A40505F931" box="[553,581,1667,1692]" captionStart="Figure 21" captionStartId="29.[164,242,1876,1898]" captionTargetBox="[163,1443,921,1837]" captionTargetId="figure-364@29.[163,1443,921,1837]" captionTargetPageId="29" captionText="Figure 21. Heleolaccocoris horvathi male: dorsal habitus (a), genital capsule with terga IX and X removed (b)." figureDoi="http://doi.org/10.5281/zenodo.6994664" httpUri="https://zenodo.org/record/6994664/files/figure.png" pageId="28" pageNumber="1273">21</figureCitation>
</paragraph>
<paragraph id="6A483699FF91FFCFFF5581E805B4F8AE" blockId="28.[145,761,1743,1795]" pageId="28" pageNumber="1273">
<emphasis id="5883EA8BFF91FFCFFF5581E8069DF949" box="[145,477,1743,1764]" italics="true" pageId="28" pageNumber="1273">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF91FFCFFF1A81CA05B4F8AE" box="[222,756,1773,1795]" pageId="28" pageNumber="1273">E84AF6F5-D470-436E-9D0C-F26EB532CCF4</uuid>
</paragraph>
<paragraph id="6A483699FF91FFCFFF55800A07A5F8CD" blockId="28.[145,760,1836,1889]" pageId="28" pageNumber="1273">
<emphasis id="5883EA8BFF91FFCFFF55800A0668F8EF" box="[145,296,1837,1858]" italics="true" pageId="28" pageNumber="1273">
<typeStatus id="B54C883BFF91FFCFFF55800A0788F8EF" box="[145,200,1837,1858]" pageId="28" pageNumber="1273">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF91FFCFFEF2800B079FF8CD" authority="(Montandon, 1897 b)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="species" species="horvathi">
<emphasis id="5883EA8BFF91FFCFFEF2800B0519F8EC" box="[310,601,1836,1857]" italics="true" pageId="28" pageNumber="1273">Heleolaccocoris horvathi</emphasis>
(
<bibRefCitation id="0E664B68FF91FFCFFDAF800B0795F8CC" author="Montandon AL" pageId="28" pageNumber="1273" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="6A483699FF91FFCFFCED83E10303FB57" blockId="28.[809,1425,1221,1887]" pageId="28" pageNumber="1273">
<emphasis id="5883EA8BFF91FFCFFCED83E104F9FB76" box="[809,953,1222,1243]" italics="true" pageId="28" pageNumber="1273">Description:</emphasis>
Form broadly oval; widest at embolia; dorsoventrally flattened.
</paragraph>
<paragraph id="6A483699FF91FFCFFC8582240391F8AE" blockId="28.[809,1425,1221,1887]" pageId="28" pageNumber="1273">Head broad, less than twice as long as wide, eyes and anterior margin continuously and evenly convex to anterolateral corners of pronotum. Eyes slightly convergent anteriorly; mesal margin straight, anteroventral margin concave; narrow band of cuticle posterior to eye widening lateral to eye. Front of head rolled posteroventrally so labrum set back from functional anterior margin, giving a bull-nosed appearance. Labrum wider than long, apex rounded or pointed, without transverse basal sulcus. Maxillary plate not flap like, but integral with head capsule, immediately posterolateral to labrum. Rostrum with three visible segments, with segments 1 and part of 2 concealed behind labrum. Antennae four segmented with segment 1 short and inconspicuous, 2 and 4 longer, 3 longest; segment 2 enlarged, 2–4 setose. Head and pronotum flattened transversely.</paragraph>
<paragraph id="6A483699FF91FFCEFC85802B06C2FE7D" blockId="28.[809,1425,1221,1887]" lastBlockId="29.[163,779,197,863]" lastPageId="29" lastPageNumber="1274" pageId="28" pageNumber="1273">Pronotum with anterior margin straight or slightly concave between eyes, then concavity becoming more pronounced as margin contours around eye; lateral margins strongly convergent anteriorly and with slight convexity; posterior margin straight except at posterolateral corners. Scutellum triangular, lateral margins sinuate, ~twice as wide as long, heavily punctate. Hemelytra punctate throughout, extending to tip of abdomen. Embolar suture well-defined mesally, but poorly defined or absent posteriorly in some species; clavus well defined with claval and intraclaval sutures.</paragraph>
<paragraph id="6A483699FF90FFCEFF7F86FE05BEFC8F" blockId="29.[163,779,197,863]" pageId="29" pageNumber="1274">Propleuron immediately lateral to coxal cavity with brush of long setae; mesal part irregularly shaped and pruinose; lateral part glabrous. Prosternum sharply carinate, carina widening anteriorly to form a Y shape. Mesosternum tumescent medially with thick brush of long hairs. Metaxyphus well developed and with median carina. Abdominal venter covered with dense pile of appressed hairs; brush of long hairs on midline of III–VII in females, III–VIII in males; glabrous band along lateral margin; laterosternites each with 2–4 oval to elongate, glabrous depressions near spiracles.</paragraph>
<paragraph id="6A483699FF90FFCEFF7F840D03AFFD68" blockId="29.[163,779,197,863]" lastBlockId="29.[827,1444,197,863]" pageId="29" pageNumber="1274">
Profemur elongate for family, not noticeably inflated, anterior margin with dorsal and ventral rows of golden hairs sandwiching one or two rows of short, dark spines. Protarsus single segmented in females, two segmented in males, each with paired movable pretarsal claws. Males with thick pad of hairs on mesotibia and tarsomeres 2 and 3, and to a lesser extent on protibia and both tarsomeres; females with pads of hairs greatly reduced. Mesofemur with posteroventral row of short, spinose setae tightly spaced such that bases of adjacent setae are in contact or nearly so (
<figureCitation id="F2CC2A1CFF90FFCEFC0D869D0353FE7D" box="[969,1043,442,464]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="29" pageNumber="1274">Fig. 5a</figureCitation>
); posterior margin with dense brush of light-coloured setae. Metafemur with posteroventral row of short spinose setae spaced approximately one seta width apart for most of length; with anteroventral seta row noticeably arcuate or with a posteromesal direction change in basal third as it extends basally (
<figureCitation id="F2CC2A1CFF90FFCEFA80857302D4FDC4" box="[1348,1428,595,618]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="29" pageNumber="1274">Fig. 5b</figureCitation>
). Meso- and metatibiae with rows of long, dark-red spines. Long, golden swimming hairs sparse on mesotibia and tarsus, profuse on metatibia and tarsus.
</paragraph>
<paragraph id="6A483699FF90FFCDFC9785E90538FF57" blockId="29.[827,1444,197,863]" lastBlockId="30.[145,761,197,250]" lastPageId="30" lastPageNumber="1275" pageId="29" pageNumber="1274">Proctiger triangular; aedeagus with lateral margins convex or angled, widest near middle; parameres small, symmetrical, touching or nearly touching lateral margins of aedeagus. Female mediosternite VII (subgenital plate) broad basally, strongly convex transversely, lateral margins convergent posteriorly; posterior margin interspecifically variable.</paragraph>
<caption id="3E886611FF90FFCEFF6080730268F8C7" ID-DOI="http://doi.org/10.5281/zenodo.6994664" ID-Zenodo-Dep="6994664" box="[164,1320,1876,1898]" httpUri="https://zenodo.org/record/6994664/files/figure.png" pageId="29" pageNumber="1274" startId="29.[164,242,1876,1898]" targetBox="[163,1443,921,1837]" targetPageId="29">
<paragraph id="6A483699FF90FFCEFF6080730268F8C7" blockId="29.[164,1320,1876,1898]" box="[164,1320,1876,1898]" pageId="29" pageNumber="1274">
<emphasis id="5883EA8BFF90FFCEFF6080730659F8C4" bold="true" box="[164,281,1876,1898]" pageId="29" pageNumber="1274">Figure 21.</emphasis>
<taxonomicName id="ADF74D1AFF90FFCEFEE780730561F8C7" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[291,545,1876,1898]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="1274" phylum="Arthropoda" rank="species" species="horvathi">
<emphasis id="5883EA8BFF90FFCEFEE780730561F8C7" box="[291,545,1876,1898]" italics="true" pageId="29" pageNumber="1274">Heleolaccocoris horvathi</emphasis>
</taxonomicName>
male: dorsal habitus (a), genital capsule with terga IX and X removed (b).
</paragraph>
</caption>
<paragraph id="6A483699FF93FFCDFF55860F06D9FE15" blockId="30.[145,761,296,440]" pageId="30" pageNumber="1275">
<emphasis id="5883EA8BFF93FFCDFF55860F0648FE90" box="[145,264,296,317]" italics="true" pageId="30" pageNumber="1275">Diagnosis:</emphasis>
Beyond attributes unique to the subfamily and tribe (see diagnoses above), additional nonmolecular features to distinguish
<taxonomicName id="ADF74D1AFF93FFCDFDF9864205B9FED7" authorityName="Sites" authorityYear="2022" box="[573,761,357,378]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFDF9864205B9FED7" box="[573,761,357,378]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris</emphasis>
</taxonomicName>
from
<taxonomicName id="ADF74D1AFF93FFCDFF0B86A30606FE34" authorityName="Stal" authorityYear="1876" box="[207,326,388,409]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFF0B86A30606FE34" box="[207,326,388,409]" italics="true" pageId="30" pageNumber="1275">Heleocoris</emphasis>
</taxonomicName>
in
<collectingCountry id="12E07609FF93FFCDFEA886A306EAFE37" box="[364,426,388,410]" name="India" pageId="30" pageNumber="1275">India</collectingCountry>
are not apparent other than geographic association.
</paragraph>
<paragraph id="6A483699FF93FFCDFF5586CB06D8FDD1" blockId="30.[145,761,492,637]" pageId="30" pageNumber="1275">
<emphasis id="5883EA8BFF93FFCDFF5586CB0652FDAC" box="[145,274,492,513]" italics="true" pageId="30" pageNumber="1275">Etymology:</emphasis>
The genus name recognizes the previous generic associations of species in this clade. Thus,
<taxonomicName id="ADF74D1AFF93FFCDFF55850E0600FD93" authorityName="Sites" authorityYear="2022" box="[145,320,553,574]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFF55850E0600FD93" box="[145,320,553,574]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris</emphasis>
</taxonomicName>
is an amalgamation of
<taxonomicName id="ADF74D1AFF93FFCDFD94850E0586FD93" authorityName="Stal" authorityYear="1876" box="[592,710,553,574]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFD94850E0586FD93" box="[592,710,553,574]" italics="true" pageId="30" pageNumber="1275">Heleocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF93FFCDFF55856F064EFDF0" authorityName="Stal" authorityYear="1856" box="[145,270,584,605]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFF55856F064EFDF0" box="[145,270,584,605]" italics="true" pageId="30" pageNumber="1275">Laccocoris</emphasis>
</taxonomicName>
, with the prefix derived from the Greek ἕλειος, marsh dwelling.
</paragraph>
<paragraph id="6A483699FF93FFCDFF5585920670FC72" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<emphasis id="5883EA8BFF93FFCDFF5585920653FD67" box="[145,275,693,714]" italics="true" pageId="30" pageNumber="1275">Comments:</emphasis>
All species of
<taxonomicName id="ADF74D1AFF93FFCDFE058591057CFD66" authorityName="Stal" authorityYear="1856" box="[449,572,694,715]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFE058591057CFD66" box="[449,572,694,715]" italics="true" pageId="30" pageNumber="1275">Laccocoris</emphasis>
</taxonomicName>
from Southeast Asia, the Sunda Islands and
<collectingCountry id="12E07609FF93FFCDFDE885F30588FD44" box="[556,712,724,746]" name="Philippines" pageId="30" pageNumber="1275">Philippines</collectingCountry>
, as well as all species of
<taxonomicName id="ADF74D1AFF93FFCDFE6C85D5056AFCAA" authorityName="Stal" authorityYear="1876" box="[424,554,754,775]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFE6C85D5056AFCAA" box="[424,554,754,775]" italics="true" pageId="30" pageNumber="1275">Heleocoris</emphasis>
</taxonomicName>
from Southeast Asia, including Yunnan, are here transferred to
<taxonomicName id="ADF74D1AFF93FFCDFF558417060BFCE8" authorityName="Sites" authorityYear="2022" box="[145,331,816,837]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFF558417060BFCE8" box="[145,331,816,837]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris</emphasis>
</taxonomicName>
. Although I have not examined all species, because of the clear geographic association for species of
<taxonomicName id="ADF74D1AFF93FFCDFEF2844906F3FC2E" authorityName="Stal" authorityYear="1856" box="[310,435,878,899]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFEF2844906F3FC2E" box="[310,435,878,899]" italics="true" pageId="30" pageNumber="1275">Laccocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF93FFCDFE36844A052DFC2F" authorityName="Stal" authorityYear="1876" box="[498,621,877,898]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF93FFCDFE36844A052DFC2F" box="[498,621,877,898]" italics="true" pageId="30" pageNumber="1275">Heleocoris</emphasis>
</taxonomicName>
, all species from this region are here transferred for taxonomic stability. As such, the following new combinations are proposed:
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF93FFCDFF6D84CF0416FE9B" pageId="30" pageNumber="1275" type="reference_group">
<paragraph id="6A483699FF93FFCDFF6D84CF07FEFBB1" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D84CF05F1FC50" authority="(Montandon, 1910 a)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1910" box="[169,689,1000,1022]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="dissidens" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D84CF0686FC50" box="[169,454,1000,1021]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris dissidens</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE1184CF05E9FC50" author="Montandon AL" box="[469,681,1000,1022]" pageId="30" pageNumber="1275" pagination="1268 - 1271" refId="ref23246" refString="Montandon AL. 1910 a. Deux especes nouvelles de la fam. Naucoridae. Bulletin de la Societe des Sciences de Bucarest, Roumanie 19: 1268 - 1271." type="journal article" year="1910">Montandon, 1910a</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD7C84CF07FEFBB1" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D83020645FBF4" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D830205B8FB97" authority="(Nieser &amp; Chen, 1992)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Nieser &amp; Chen" baseAuthorityYear="1992" box="[169,760,1061,1083]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="floresensis" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D830206A4FB97" box="[169,484,1061,1082]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris floresensis</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE31830205B0FB97" author="Nieser N &amp; Chen PP" box="[501,752,1061,1083]" pageId="30" pageNumber="1275" pagination="30 - 40" refId="ref23429" refString="Nieser N, Chen PP. 1992. Notes on Indonesian waterbugs (Nepomorpha &amp; Gerromorpha). Storkia 1: 30 - 40." type="journal article" year="1992">Nieser &amp; Chen, 1992</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFF5583630645FBF4" box="[145,261,1092,1114]" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D834507FEFB3B" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D834505EDFBD5" authority="(Montandon, 1909)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1909" box="[169,685,1122,1144]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="grandis" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D83450683FBDA" box="[169,451,1122,1143]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris grandis</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE11834505E5FBD5" author="Montandon AL" box="[469,677,1122,1144]" pageId="30" pageNumber="1275" pagination="43 - 61" refId="ref23219" refString="Montandon AL. 1909. Naucoridae. Description d'especes nouvelles. Bulletin de la Societe des Sciences de Bucarest, Roumanie 18: 43 - 61." type="journal article" year="1909">Montandon, 1909</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD72834507FEFB3B" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D838707FEFB79" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D838705F0FB18" authority="(La Rivers, 1970)" authorityName="Sites" authorityYear="2022" baseAuthorityName="La Rivers" baseAuthorityYear="1970" box="[169,688,1184,1206]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="hoogstraali" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D838706A3FB18" box="[169,483,1184,1205]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris hoogstraali</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE37838705E7FB18" author="La Rivers I" box="[499,679,1184,1206]" pageId="30" pageNumber="1275" pagination="496 - 499" refId="ref22426" refString="La Rivers I. 1970. Two new species of Laccocoris from British North Borneo (Hemiptera: Naucoridae: Laccocorinae). Proceedings of the Entomological Society of Washington 72: 496 - 499." type="journal article" year="1970">La Rivers, 1970</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD7C838707FEFB79" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D83FA07FEFABC" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D83FA05F0FB5F" authority="(Montandon, 1897 b)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[169,688,1245,1267]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="horvathi" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D83FA0682FB5F" box="[169,450,1245,1266]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris horvathi</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE1683FA05E8FB5E" author="Montandon AL" box="[466,680,1245,1267]" pageId="30" pageNumber="1275" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD7C83FA07FEFABC" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D823D05F9FA9D" blockId="30.[145,762,693,1880]" box="[169,697,1306,1328]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D823D057DFA82" authority="(Zettel, 2012)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Zettel" baseAuthorityYear="2012" box="[169,573,1306,1328]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="jaechi" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D823D06E0FA82" box="[169,416,1306,1327]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris jaechi</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE6B823D0576FA82" author="Zettel H" box="[431,566,1306,1328]" pageId="30" pageNumber="1275" pagination="27 - 36" refId="ref25608" refString="Zettel H. 2012. New creeping water bugs (Insecta: Heteroptera: Naucoridae) from China and the Philippines. Annalen des Naturhistorischen Museums in Wien 113 B: 27 - 36." type="journal article" year="2012">Zettel, 2012</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD80823D05F9FA9D" box="[580,697,1306,1328]" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D821E07FEFAC0" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D821E05F1FAE3" authority="(Montandon, 1897 b)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[169,689,1337,1359]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="laeviceps" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D821E0684FAE3" box="[169,452,1337,1358]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris laeviceps</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE10821E05E9FAE2" author="Montandon AL" box="[468,681,1337,1359]" pageId="30" pageNumber="1275" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD7C821E07FEFAC0" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D82510687FA07" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D8251060BFA07" authority="(D. Polhemus &amp; J. Polhemus, 2013)" authorityName="Sites" authorityYear="2022" baseAuthorityName="D. Polhemus &amp; J. Polhemus" baseAuthorityYear="2013" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="malayensis" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D82510543FA26" box="[169,515,1398,1419]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris malayensis</emphasis>
(D. Polhemus &amp; J.
<bibRefCitation id="0E664B68FF93FFCDFF5582B20603FA07" author="Polhemus DA &amp; Polhemus JT" box="[145,323,1429,1451]" pageId="30" pageNumber="1275" pagination="665 - 686" refId="ref23727" refString="Polhemus DA, Polhemus JT. 2013. Guide to the aquatic Heteroptera of Singapore and Peninsular Malaysia. XI. Infraorder Nepomorpha - families Naucoridae and Aphelocheiridae. Raffles Bulletin of Zoology 61: 665 - 686." type="journal article" year="2013">Polhemus, 2013</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFE9682B20687FA07" box="[338,455,1429,1451]" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D82930645FA45" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D829305B9FA64" authority="(Montandon, 1897 d)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[169,761,1460,1482]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="marginatus" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D829306B9FA64" box="[169,505,1460,1481]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris marginatus</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFDC8829305B0FA67" author="Montandon AL" box="[524,752,1460,1482]" pageId="30" pageNumber="1275" pagination="50 - 66" refId="ref23079" refString="Montandon AL. 1897 d. Hemipteres-Heteropteres exotiques. Notes et descriptions, Annales de la Societe Entomologique de Belgique 41: 50 - 66." type="journal article" year="1897">Montandon, 1897d</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFF5582F50645FA45" box="[145,261,1490,1512]" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D82D6060BF988" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D82D60790F988" authority="(Sites &amp; Vitheepradit, 2011)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Sites &amp; Vitheepradit" baseAuthorityYear="2011" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="mcphersoni" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D82D606ADF9AB" box="[169,493,1521,1542]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris mcphersoni</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE3982D60788F988" author="Sites RW &amp; Vitheepradit A" pageId="30" pageNumber="1275" pagination="1 - 16" refId="ref24735" refString="Sites RW, Vitheepradit A. 2011. The Heleocoris (Heteroptera: Naucoridae: Laccocorinae) of Thailand, with the description of a new species. Zootaxa 2736: 1 - 16." type="journal article" year="2011">Sites &amp; Vitheepradit, 2011</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFF138137060BF988" box="[215,331,1552,1574]" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D810907FEF9CF" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D810905F1F9EE" authority="(Lundblad, 1933)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Lundblad" baseAuthorityYear="1933" box="[169,689,1582,1604]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="montandoni" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D810906A8F9EE" box="[169,488,1582,1603]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris montandoni</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE33810905E9F9EE" author="Lundblad O" box="[503,681,1582,1604]" pageId="30" pageNumber="1275" pagination="263 - 489" refId="ref22883" refString="Lundblad O. 1933. Zur Kenntnis der aquatilen und semiaquatilen Hemipteren von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement 12: 1 - 195, 263 - 489, 21 plates." type="book chapter" year="1933">Lundblad, 1933</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD7C810907FEF9CF" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D814B07FEF90D" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D814B05F0F92C" authority="(Montandon, 1909)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1909" box="[169,688,1644,1666]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="nebulosus" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D814B0690F92C" box="[169,464,1644,1665]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris nebulosus</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE24814B05E8F92C" author="Montandon AL" box="[480,680,1644,1666]" pageId="30" pageNumber="1275" pagination="43 - 61" refId="ref23219" refString="Montandon AL. 1909. Naucoridae. Description d'especes nouvelles. Bulletin de la Societe des Sciences de Bucarest, Roumanie 18: 43 - 61." type="journal article" year="1909">Montandon, 1909</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD7C814B07FEF90D" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D818E07FEF970" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D818E05F0F913" authority="(Montandon, 1897 d)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[169,688,1705,1727]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="nervicus" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D818E0680F912" box="[169,448,1705,1727]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris nervicus</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE14818E05E8F912" author="Montandon AL" box="[464,680,1705,1727]" pageId="30" pageNumber="1275" pagination="50 - 66" refId="ref23079" refString="Montandon AL. 1897 d. Hemipteres-Heteropteres exotiques. Notes et descriptions, Annales de la Societe Entomologique de Belgique 41: 50 - 66." type="journal article" year="1897">Montandon, 1897d</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD7C818E07FEF970" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D81C10645F8B7" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D81C105B9F956" authority="(Montandon, 1897 d)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[169,761,1766,1788]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="obscuratus" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D81C106B5F956" box="[169,501,1766,1787]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris obscuratus</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFDCD81C105B0F951" author="Montandon AL" box="[521,752,1766,1788]" pageId="30" pageNumber="1275" pagination="50 - 66" refId="ref23079" refString="Montandon AL. 1897 d. Hemipteres-Heteropteres exotiques. Notes et descriptions, Annales de la Societe Entomologique de Belgique 41: 50 - 66." type="journal article" year="1897">Montandon, 1897d</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFF5580220645F8B7" box="[145,261,1797,1819]" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF93FFCDFF6D800307FEF8F5" blockId="30.[145,762,693,1880]" pageId="30" pageNumber="1275">
<taxonomicName id="ADF74D1AFF93FFCDFF6D800305EDF894" authority="(Montandon, 1897 b)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[169,685,1828,1850]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="ovatus" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFF6D800306F5F894" box="[169,437,1828,1849]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris ovatus</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFE02800305E4F897" author="Montandon AL" box="[454,676,1828,1850]" pageId="30" pageNumber="1275" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF93FFCDFD72800307FEF8F5" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
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<taxonomicName id="ADF74D1AFF93FFCDFC8587E202D1FF77" authority="(Montandon, 1897 b)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[833,1425,197,219]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="staudingeri" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFC8587E203D1FF77" box="[833,1169,197,218]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris staudingeri</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFB6087E202C8FF76" author="Montandon AL" box="[1188,1416,197,219]" pageId="30" pageNumber="1275" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
)
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<taxonomicNameLabel id="43B057F0FF93FFCDFCED87C304DDFF54" box="[809,925,228,250]" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
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<taxonomicName id="ADF74D1AFF93FFCDFC8586250206FEBA" authority="(Montandon, 1897 b)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[833,1350,258,280]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="strabus" status="comb. nov.">
<emphasis id="5883EA8BFF93FFCDFC8586250313FEBA" box="[833,1107,258,279]" italics="true" pageId="30" pageNumber="1275">Heleolaccocoris strabus</emphasis>
(
<bibRefCitation id="0E664B68FF93FFCDFBA08625027EFEB5" author="Montandon AL" box="[1124,1342,258,280]" pageId="30" pageNumber="1275" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
)
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<taxonomicNameLabel id="43B057F0FF93FFCDFA8B86250416FE9B" pageId="30" pageNumber="1275" rank="species">comb. nov.</taxonomicNameLabel>
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59
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Species in the following three new genera were, or would have been in the case of
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, previously recognized within
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. Thus, although molecular evidence reveals strong distinctions among them, generic level morphological differences are subtle at best in these morphologically conserved taxa. As such, I provide a generalized description of each new genus, but with few evident morphological differences.
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242
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<figureCitation id="F2CC2A1CFF93FFCDFBD08420033CFCB2" box="[1044,1148,775,800]" captionStart="Figure 19" captionStartId="25.[164,245,774,796]" captionTargetBox="[163,1444,195,734]" captionTargetId="figure-573@25.[163,1444,195,734]" captionTargetPageId="25" captionText="Figure 19. Revised generic representation of the former genus Naucoris. Dorsal habitus of Naucoris maculatus (a), Australonaucoris congrex (b), Afronaucoris parvulus (c), Indonaucoris minutus sp. nov. (d)." figureDoi="http://doi.org/10.5281/zenodo.6994658" httpUri="https://zenodo.org/record/6994658/files/figure.png" pageId="30" pageNumber="1275">FIGS 19C</figureCitation>
,
<figureCitation id="F2CC2A1CFF93FFCDFB4D842003E5FC8D" box="[1161,1189,775,800]" captionStart="Figure 22" captionStartId="31.[164,242,1851,1873]" captionTargetBox="[163,1444,895,1813]" captionTargetId="figure-311@31.[163,1444,895,1813]" captionTargetPageId="31" captionText="Figure 22. Afronaucoris obscuratus male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c), distal part of aedeagus (d). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994668" httpUri="https://zenodo.org/record/6994668/files/figure.png" pageId="30" pageNumber="1275">22</figureCitation>
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<emphasis id="5883EA8BFF93FFCDFCED84760335FCCB" box="[809,1141,848,870]" italics="true" pageId="30" pageNumber="1275">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
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<typeStatus id="B54C883BFF93FFCDFCED8489041FFC6E" box="[809,863,942,963]" pageId="30" pageNumber="1275">Type</typeStatus>
species:
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<taxonomicName id="ADF74D1AFF93FFCDFC10848A042BFC4C" authority="(Montandon, 1913)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1913" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1275" phylum="Arthropoda" rank="species" species="obscuratus">
<emphasis id="5883EA8BFF93FFCDFC10848A03B3FC6F" box="[980,1267,941,962]" italics="true" pageId="30" pageNumber="1275">Afronaucoris obscuratus</emphasis>
(
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)
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.
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<emphasis id="5883EA8BFF93FFCDFCED832104FEFBB6" box="[809,958,1030,1051]" italics="true" pageId="30" pageNumber="1275">Description:</emphasis>
Form ovate; widest just posterior to middle, at embolia (
<figureCitation id="F2CC2A1CFF93FFCDFBDF8303033CFB94" box="[1051,1148,1060,1082]" captionStart="Figure 22" captionStartId="31.[164,242,1851,1873]" captionTargetBox="[163,1444,895,1813]" captionTargetId="figure-311@31.[163,1444,895,1813]" captionTargetPageId="31" captionText="Figure 22. Afronaucoris obscuratus male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c), distal part of aedeagus (d). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994668" httpUri="https://zenodo.org/record/6994668/files/figure.png" pageId="30" pageNumber="1275">Fig. 22a</figureCitation>
); moderately flattened for family. Head broad, eyes and anterior margin continuously and evenly convex to anterolateral corners of pronotum. Eyes convergent anteriorly; each eye narrowing anteriorly when viewed from above, mesal margin straight or nearly so, anteroventral margin concave; narrow band of cuticle lateral to eye. Labrum arising at front of head between inner margins of eyes; apex broadly rounded. Maxillary plates angled ventromedially, tips rounded, not exceeding apex of labrum. Rostrum with three visible segments, with segments one and part of two concealed behind labrum. Antennae four segmented with segment 1 short and inconspicuous, 2 and 4 longer, 3 longest, segments 2–4 enlarged and setose. Midventer of head sharply carinate. Head and pronotum moderately convex transversely.
</paragraph>
<paragraph id="6A483699FF93FFCCFC8581090596FE9A" blockId="30.[809,1425,1029,1879]" lastBlockId="31.[163,779,197,832]" lastPageId="31" lastPageNumber="1276" pageId="30" pageNumber="1275">Pronotum with anterior margin straight or slightly concave between eyes, then concavity becoming more pronounced as margin wraps around eyes; lateral margins slightly convex and convergent anteriorly; posterior margin mostly straight, with slight concavity medially and convexity laterally to posterolateral corners. Scutellum triangular, slightly wider than long, evenly domed, heavily punctate. Hemelytra extending to tip of abdomen. Embolium well defined with embolar suture, yellowish anteriorly, dark posteriorly; clavus well defined with claval and intraclaval sutures; corium, clavus and membrane punctate and dark with subtle mottling, sparse long hairs and a single light spot at the middle of corium/membrane interface.</paragraph>
<paragraph id="6A483699FF92FFCCFF7F8667044AFD0A" blockId="31.[163,779,197,832]" pageId="31" pageNumber="1276">Propleuron lateral to coxal cavity infuscated and setose; posterolaterally light coloured and glabrous; mesal margin triangular and touching prosternellum such that prosternellum is dorsad of propleuron tip. Probasisternum sharply carinate, carina continuous with midventral carina of head. Mesosternum broadly carinate medially with a brush of long hairs and several isolated short, dark spines, which can be concealed by hairs. Metaxyphus with sharp median carina. Abdominal venter thickly covered with long hairs except narrow glabrous lateral band; laterosternites with paired, oval, glabrous depressions near spiracles.</paragraph>
<paragraph id="6A483699FF92FFCCFF7F85970242FE3F" blockId="31.[163,779,197,832]" lastBlockId="31.[827,1443,197,740]" pageId="31" pageNumber="1276">Profemur inflated, anterior margin with dorsal and ventral rows of golden hairs sandwiching two rows of short, dark spines. Protarsus single segmented with single, fixed, minute pretarsal claw. Meso- and metafemora with posteroventral row of short spines; those on mesofemur densely packed such that bases of adjacent spines are touching or nearly touching; those on metafemur less densely packed, spaced approximately one spine width apart. Meso- and metatibiae with numerous rows of long dark red spines. Long golden swimming hairs sparse on mesotibia and tarsus, profuse on metatibia and tarsus.</paragraph>
<paragraph id="6A483699FF92FFCCFC9786BB032BFD49" blockId="31.[827,1443,197,740]" pageId="31" pageNumber="1276">Male genitalia with aedeagus widest apically, with pronounced flap-like structure on right side with membranous window below; parameres strongly asymmetrical, criss-crossing in basal third; left paramere overlapping right, arcuate, with pronounced lobe on left margin; right paramere slenderer, more arcuate, with elongate sulcus in distal third to embrace aedeagus. Female mediosternite VII (subgenital plate) broad basally, strongly convex transversely, lateral margins convergent posteriorly toward broadly rounded to spatulate apex.</paragraph>
<paragraph id="6A483699FF92FFF3FCFF8429062FFEB5" blockId="31.[827,1442,781,834]" lastBlockId="32.[145,761,197,280]" lastPageId="32" lastPageNumber="1277" pageId="31" pageNumber="1276">
<emphasis id="5883EA8BFF92FFCCFCFF842904FDFC8E" box="[827,957,782,803]" italics="true" pageId="31" pageNumber="1276">Diagnosis:</emphasis>
Morphological features diagnostic for this clade are not evident at this time; however, the distinction of this genus as an independent lineage is supported by its geographic association with the Afrotropical region.
</paragraph>
<caption id="3E886611FF92FFCCFF60801C034BF8C2" ID-DOI="http://doi.org/10.5281/zenodo.6994668" ID-Zenodo-Dep="6994668" httpUri="https://zenodo.org/record/6994668/files/figure.png" pageId="31" pageNumber="1276" startId="31.[164,242,1851,1873]" targetBox="[163,1444,895,1813]" targetPageId="31">
<paragraph id="6A483699FF92FFCCFF60801C034BF8C2" blockId="31.[163,1442,1851,1903]" pageId="31" pageNumber="1276">
<emphasis id="5883EA8BFF92FFCCFF60801C0658F8FC" bold="true" box="[164,280,1851,1873]" pageId="31" pageNumber="1276">Figure 22.</emphasis>
<taxonomicName id="ADF74D1AFF92FFCCFEE6801B055FF8FF" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1913" box="[290,543,1852,1874]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="1276" phylum="Arthropoda" rank="species" species="obscuratus">
<emphasis id="5883EA8BFF92FFCCFEE6801B055FF8FF" box="[290,543,1852,1874]" italics="true" pageId="31" pageNumber="1276">Afronaucoris obscuratus</emphasis>
</taxonomicName>
male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c), distal part of aedeagus (d). Sizes are not proportional among figures.
</paragraph>
</caption>
<paragraph id="6A483699FFADFFF3FF55866605D5FE1E" blockId="32.[145,761,321,435]" pageId="32" pageNumber="1277">
<emphasis id="5883EA8BFFADFFF3FF5586660651FEFB" box="[145,273,321,342]" italics="true" pageId="32" pageNumber="1277">Etymology:</emphasis>
The genus name recognizes the region in which these former members of
<taxonomicName id="ADF74D1AFFADFFF3FDC78647052BFED8" authorityName="Geoffroy" authorityYear="1762" box="[515,619,352,373]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFADFFF3FDC78647052BFED8" box="[515,619,352,373]" italics="true" pageId="32" pageNumber="1277">Naucoris</emphasis>
</taxonomicName>
occur, while still referencing the previous generic association. The prefix is derived from the Latin
<taxonomicName id="ADF74D1AFFADFFF3FE3A86BA056DFE1F" authorityName="Conrad" authorityYear="1858" box="[510,557,413,434]" class="Gastropoda" family="Buccinidae" genus="Afer" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="1277" phylum="Mollusca" rank="genus">
<emphasis id="5883EA8BFFADFFF3FE3A86BA056DFE1F" box="[510,557,413,434]" italics="true" pageId="32" pageNumber="1277">Afer</emphasis>
</taxonomicName>
, African.
</paragraph>
<paragraph id="6A483699FFADFFF3FF5586FB0642FB7F" blockId="32.[145,762,476,1571]" pageId="32" pageNumber="1277">
<emphasis id="5883EA8BFFADFFF3FF5586FB0653FE5C" box="[145,275,476,497]" italics="true" pageId="32" pageNumber="1277">Comments:</emphasis>
<bibRefCitation id="0E664B68FFADFFF3FEE586FB068AFE5C" author="Poisson R" box="[289,458,476,498]" pageId="32" pageNumber="1277" pagination="3 - 94" refId="ref23595" refString="Poisson R. 1949. Exploration du Parc National Albert, I. Mission G. F. de Witte 1933 - 1935. Hemipteres aquatiques. Institut des Parcs Nationaux Congo Belge 58: 3 - 94." type="journal article" year="1949">Poisson (1949)</bibRefCitation>
established the subgenus
<taxonomicName id="ADF74D1AFFADFFF3FF5586DD0667FDA2" authorityName="Poisson" authorityYear="1949" box="[145,295,506,527]" class="Insecta" family="Naucoridae" genus="Naucorisella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFADFFF3FF5586DD0667FDA2" box="[145,295,506,527]" italics="true" pageId="32" pageNumber="1277">Naucorisella</emphasis>
</taxonomicName>
for the Madagascan
<taxonomicName id="ADF74D1AFFADFFF3FDDA86DC067EFD82" authority="Signoret, 1860" authorityName="Signoret" authorityYear="1860" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="5883EA8BFFADFFF3FDDA86DC05B9FDA2" box="[542,761,506,528]" italics="true" pageId="32" pageNumber="1277">Naucoris parvulus</emphasis>
<bibRefCitation id="0E664B68FFADFFF3FF55853E067EFD82" author="Signoret V" box="[145,318,537,559]" pageId="32" pageNumber="1277" pagination="917 - 972" refId="ref24302" refString="Signoret V. 1860. Faune des Hemipteres de Madagascar. (Suite et fin.). Heteropteres. Annales de la Societe Entomologique de France 8: 917 - 972." type="journal article" year="1860">Signoret, 1860</bibRefCitation>
</taxonomicName>
, but the subgenus was considered to be invalid and was synonymized by
<bibRefCitation id="0E664B68FFADFFF3FD858511079CFDC1" author="Stys P &amp; Jansson A" pageId="32" pageNumber="1277" pagination="1 - 44" refId="ref25103" refString="Stys P, Jansson A. 1988. Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica 50: 1 - 44." type="journal article" year="1988">Štys &amp; Jansson (1988)</bibRefCitation>
. The Afrotropical species of
<taxonomicName id="ADF74D1AFFADFFF3FDF2857005E2FDC1" authorityName="Geoffroy" authorityYear="1762" box="[566,674,599,620]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFADFFF3FDF2857005E2FDC1" box="[566,674,599,620]" italics="true" pageId="32" pageNumber="1277">Naucoris</emphasis>
</taxonomicName>
, which excludes the Palaearctic
<taxonomicName id="ADF74D1AFFADFFF3FE2F855105B9FD26" authorityName="Fabricius" authorityYear="1798" box="[491,761,629,651]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="5883EA8BFFADFFF3FE2F855105B9FD26" box="[491,761,629,651]" italics="true" pageId="32" pageNumber="1277">Naucoris maculatus</emphasis>
</taxonomicName>
occurring in northern Africa, are here transferred to
<taxonomicName id="ADF74D1AFFADFFF3FF7785940614FD65" authorityName="Sites" authorityYear="2022" box="[179,340,691,712]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFADFFF3FF7785940614FD65" box="[179,340,691,712]" italics="true" pageId="32" pageNumber="1277">Afronaucoris</emphasis>
</taxonomicName>
. Although only two species were included in the molecular analysis, the phylogenetic framework suggests a geographic occurrence by which to preliminarily associate species with this new genus. I have examined five of the eight described Afrotropical species and these are exceedingly similar in appearance. In fact,
<taxonomicName id="ADF74D1AFFADFFF3FD4A844C07B6FC33" authorityName="Poisson" authorityYear="1949" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="kenyalis">
<emphasis id="5883EA8BFFADFFF3FD4A844C07B6FC33" italics="true" pageId="32" pageNumber="1277">Naucoris kenyalis</emphasis>
</taxonomicName>
was originally described as a subspecies of
<taxonomicName id="ADF74D1AFFADFFF3FF74848F0557FC10" authority="(Poisson, 1949)" baseAuthorityName="Poisson" baseAuthorityYear="1949" box="[176,535,936,958]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="obscuratus">
<emphasis id="5883EA8BFFADFFF3FF74848F0615FC10" box="[176,341,936,957]" italics="true" pageId="32" pageNumber="1277">N. obscuratus</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FEAC848F054DFC10" author="Poisson R" box="[360,525,936,958]" pageId="32" pageNumber="1277" pagination="3 - 94" refId="ref23595" refString="Poisson R. 1949. Exploration du Parc National Albert, I. Mission G. F. de Witte 1933 - 1935. Hemipteres aquatiques. Institut des Parcs Nationaux Congo Belge 58: 3 - 94." type="journal article" year="1949">Poisson, 1949</bibRefCitation>
)
</taxonomicName>
. Of the remaining three species, one is known from only the
<typeStatus id="B54C883BFFADFFF3FD5684E105B8FC71" box="[658,760,966,988]" pageId="32" pageNumber="1277" type="holotype">holotype</typeStatus>
(
<taxonomicName id="ADF74D1AFFADFFF3FF5D84C10600FC57" authorityName="Schaum" authorityYear="1853" box="[153,320,997,1019]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="5883EA8BFFADFFF3FF5D84C10600FC57" box="[153,320,997,1019]" italics="true" pageId="32" pageNumber="1277">N. fuscipennis</emphasis>
</taxonomicName>
) and the other two (
<taxonomicName id="ADF74D1AFFADFFF3FDF084C105E3FC56" authorityName="Bolivar" authorityYear="1879" box="[564,675,998,1019]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="5883EA8BFFADFFF3FDF084C105E3FC56" box="[564,675,998,1019]" italics="true" pageId="32" pageNumber="1277">N. perezii</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFFADFFF3FD2484C107A1FBB4" authorityName="Germar" authorityYear="1838" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="5883EA8BFFADFFF3FD2484C107A1FBB4" italics="true" pageId="32" pageNumber="1277">N. planus</emphasis>
</taxonomicName>
) are known from only the inadequately brief original descriptions of more than 140 years ago (only 20 words in length for
<taxonomicName id="ADF74D1AFFADFFF3FE188365050CFBFB" authorityName="Germar" authorityYear="1838" box="[476,588,1089,1111]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="5883EA8BFFADFFF3FE188365050CFBFB" box="[476,588,1089,1111]" italics="true" pageId="32" pageNumber="1277">N. planus</emphasis>
</taxonomicName>
). Nonetheless, because of their geographic association and to establish taxonomic stability, they are included here. As such, the following new combinations are proposed:
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FFADFFF3FF6D83FD05FDF98F" pageId="32" pageNumber="1277" type="reference_group">
<paragraph id="6A483699FFADFFF3FF6D83FD07FEFAA3" blockId="32.[145,762,476,1571]" pageId="32" pageNumber="1277">
<taxonomicName id="ADF74D1AFFADFFF3FF6D83FD05EFFB42" authority="(Linnavuori, 1971)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Linnavuori" baseAuthorityYear="1971" box="[169,687,1242,1264]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="ciliatistylus" status="comb. nov.">
<emphasis id="5883EA8BFFADFFF3FF6D83FD068FFB42" box="[169,463,1242,1263]" italics="true" pageId="32" pageNumber="1277">Afronaucoris ciliatistylus</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FE1B83FD05E7FB42" author="Linnavuori R" box="[479,679,1242,1264]" pageId="32" pageNumber="1277" pagination="340 - 366" refId="ref22692" refString="Linnavuori R. 1971. Hemiptera of the Sudan, with remarks on some species of the adjacent countries. 1. The aquatic and subaquatic families. Annales Zoologici Fennici 8: 340 - 366." type="journal article" year="1971">Linnavuori, 1971</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FD7383FD07FEFAA3" pageId="32" pageNumber="1277" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFADFFF3FF6D823F05B5FA80" blockId="32.[145,762,476,1571]" box="[169,757,1304,1326]" pageId="32" pageNumber="1277">
<taxonomicName id="ADF74D1AFFADFFF3FF6D823F053AFA80" authority="(Schaum, 1853)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Schaum" baseAuthorityYear="1853" box="[169,634,1304,1326]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="fuscipennis" status="comb. nov.">
<emphasis id="5883EA8BFFADFFF3FF6D823F0683FA80" box="[169,451,1304,1325]" italics="true" pageId="32" pageNumber="1277">Afronaucoris fuscipennis</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FE16823F0532FA83" author="Schaum H" box="[466,626,1304,1326]" pageId="32" pageNumber="1277" pagination="356 - 359" refId="ref24196" refString="Schaum H. 1853. Bericht uber die zur Bekanntmachung geeigneten Verhandlungen der Koniglich Preussischen. Akademie Wissenschaften zu Berlin 2: 356 - 359." type="journal article" year="1853">Schaum, 1853</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FD45823F05B5FA80" box="[641,757,1304,1326]" pageId="32" pageNumber="1277" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFADFFF3FF6D8211058DFAE1" blockId="32.[145,762,476,1571]" box="[169,717,1334,1356]" pageId="32" pageNumber="1277">
<taxonomicName id="ADF74D1AFFADFFF3FF6D82110511FAE6" authority="(Poisson, 1949)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Poisson" baseAuthorityYear="1949" box="[169,593,1334,1356]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="kenyalis" status="comb. nov.">
<emphasis id="5883EA8BFFADFFF3FF6D821106E1FAE6" box="[169,417,1334,1355]" italics="true" pageId="32" pageNumber="1277">Afronaucoris kenyalis</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FE7482110509FAE6" author="Poisson R" box="[432,585,1334,1356]" pageId="32" pageNumber="1277" pagination="3 - 94" refId="ref23595" refString="Poisson R. 1949. Exploration du Parc National Albert, I. Mission G. F. de Witte 1933 - 1935. Hemipteres aquatiques. Institut des Parcs Nationaux Congo Belge 58: 3 - 94." type="journal article" year="1949">Poisson, 1949</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FD9C8211058DFAE1" box="[600,717,1334,1356]" pageId="32" pageNumber="1277" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFADFFF3FF6D82720645FA24" blockId="32.[145,762,476,1571]" pageId="32" pageNumber="1277">
<taxonomicName id="ADF74D1AFFADFFF3FF6D827205B9FAC7" authority="(Montandon, 1899)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1899" box="[169,761,1365,1387]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="madagascariensis" status="comb. nov.">
<emphasis id="5883EA8BFFADFFF3FF6D82720556FAC7" box="[169,534,1365,1386]" italics="true" pageId="32" pageNumber="1277">Afronaucoris madagascariensis</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FDE2827205B1FAC7" author="Montandon AL" box="[550,753,1365,1387]" pageId="32" pageNumber="1277" pagination="79 - 81" refId="ref23144" refString="Montandon AL. 1899. Deux especes nouvelles d'Hemipteres Heteropteres des collections du Museum de Paris. Bulletin du Museum d'Histoire Naturelle 2: 79 - 81." type="journal article" year="1899">Montandon, 1899</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FF5582530645FA24" box="[145,261,1396,1418]" pageId="32" pageNumber="1277" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFADFFF3FF6D82B507FEFA6B" blockId="32.[145,762,476,1571]" pageId="32" pageNumber="1277">
<taxonomicName id="ADF74D1AFFADFFF3FF6D82B505EEFA0A" authority="(Montandon, 1913)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1913" box="[169,686,1426,1448]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="obscuratus" status="comb. nov.">
<emphasis id="5883EA8BFFADFFF3FF6D82B50688FA0A" box="[169,456,1426,1447]" italics="true" pageId="32" pageNumber="1277">Afronaucoris obscuratus</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FE1D82B505E5FA0A" author="Montandon AL" box="[473,677,1426,1448]" pageId="32" pageNumber="1277" pagination="219 - 224" refId="ref23328" refString="Montandon AL. 1913. Hemipteres aquatiques. Notes et descriptions de deux especes nouvelles. Bulletin de la Section Scientifique de l'Academie Roumaine 1: 219 - 224." type="journal article" year="1913">Montandon, 1913</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FD7382B507FEFA6B" pageId="32" pageNumber="1277" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFADFFF3FF6D82F7059FFA48" blockId="32.[145,762,476,1571]" box="[169,735,1488,1510]" pageId="32" pageNumber="1277">
<taxonomicName id="ADF74D1AFFADFFF3FF6D82F70524FA48" authority="(Signoret, 1860)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Signoret" baseAuthorityYear="1860" box="[169,612,1488,1510]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="parvulus" status="comb. nov.">
<emphasis id="5883EA8BFFADFFF3FF6D82F706E8FA48" box="[169,424,1488,1509]" italics="true" pageId="32" pageNumber="1277">Afronaucoris parvulus</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FE7382F7051CFA4B" author="Signoret V" box="[439,604,1488,1510]" pageId="32" pageNumber="1277" pagination="917 - 972" refId="ref24302" refString="Signoret V. 1860. Faune des Hemipteres de Madagascar. (Suite et fin.). Heteropteres. Annales de la Societe Entomologique de France 8: 917 - 972." type="journal article" year="1860">Signoret, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FDAF82F7059FFA48" box="[619,735,1488,1510]" pageId="32" pageNumber="1277" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFADFFF3FF6D82C905F5F9A9" blockId="32.[145,762,476,1571]" box="[169,693,1518,1540]" pageId="32" pageNumber="1277">
<taxonomicName id="ADF74D1AFFADFFF3FF6D82C9057AF9AE" authority="(Bolivar, 1879)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Bolivar" baseAuthorityYear="1879" box="[169,570,1518,1540]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="perezii" status="comb. nov.">
<emphasis id="5883EA8BFFADFFF3FF6D82C906CEF9A9" box="[169,398,1518,1540]" italics="true" pageId="32" pageNumber="1277">Afronaucoris perezii</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FE5982C90572F9AE" author="Bolivar I" box="[413,562,1518,1540]" pageId="32" pageNumber="1277" pagination="133 - 146" refId="ref21458" refString="Bolivar I. 1879. Hemipteros nuevos del Museo de Madrid. Anales de la Sociedad Espanola de Historia Natural 8: 133 - 146." type="journal article" year="1879">Bolívar, 1879</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FD8582C905F5F9A9" box="[577,693,1518,1540]" pageId="32" pageNumber="1277" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFADFFF3FF6D812A05FDF98F" blockId="32.[145,762,476,1571]" box="[169,701,1549,1571]" pageId="32" pageNumber="1277">
<taxonomicName id="ADF74D1AFFADFFF3FF6D812A0502F98F" authority="(Germar, 1838)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Germar" baseAuthorityYear="1838" box="[169,578,1549,1570]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="planus" status="comb. nov.">
<emphasis id="5883EA8BFFADFFF3FF6D812A06D1F98F" box="[169,401,1549,1570]" italics="true" pageId="32" pageNumber="1277">Afronaucoris planus</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FE64812A057AF98F" author="Germar EF" box="[416,570,1549,1570]" pageId="32" pageNumber="1277" pagination="121 - 192" refId="ref21952" refString="Germar EF. 1838. Hemiptera Heteroptera promontorii bonae spei nundum descripta, quae collegit C. F. Drege. Revue Entomologique, Publiee par Silbermann 5: 121 - 192." type="journal article" year="1838">Germar, 1838</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FD8D812A05FDF98F" box="[585,701,1549,1571]" pageId="32" pageNumber="1277" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="22ED6512FF94FFCAFC0682F603D8F9BB" pageId="25" pageNumber="1270" type="nomenclature">
<paragraph id="6A483699FF94FFCAFC0682F6025CFA44" blockId="25.[962,1308,1489,1514]" box="[962,1308,1489,1514]" pageId="25" pageNumber="1270">
<heading id="310081F5FF94FFCAFC0682F6025CFA44" box="[962,1308,1489,1514]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="1270" reason="3">
TRIBE
<taxonomicName id="ADF74D1AFF94FFCAFBCA82F603D6FA45" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1038,1174,1489,1513]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Idiocarini">IDIOCARINI</taxonomicName>
<emphasis id="5883EA8BFF94FFCAFB5A82F2025CFA44" bold="true" box="[1182,1308,1489,1513]" pageId="25" pageNumber="1270">
<taxonomicNameLabel id="43B057F0FF94FFCAFB5A82F2025CFA44" box="[1182,1308,1489,1513]" pageId="25" pageNumber="1270" rank="tribe">TRIB. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A483699FF94FFCAFB8282DA03D8F9BB" blockId="25.[1094,1176,1533,1558]" box="[1094,1176,1533,1558]" pageId="25" pageNumber="1270">
<figureCitation id="F2CC2A1CFF94FFCAFB8282DA03D8F9BB" box="[1094,1176,1533,1558]" captionStart="Figure 23" captionStartId="33.[163,241,986,1008]" captionTargetBox="[163,1444,195,946]" captionTargetId="figure-408@33.[163,1444,195,946]" captionTargetPageId="33" captionText="Figure 23. Australonaucoris congrex male: dorsal habitus (a), genital capsule with parameres and terga IX and X removed (b), left paramere (c), right paramere (d). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994670" httpUri="https://zenodo.org/record/6994670/files/figure.png" pageId="25" pageNumber="1270">FIG. 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF94FFC9FCFF8113066CFBDD" lastPageId="26" lastPageNumber="1271" pageId="25" pageNumber="1270" type="description">
<paragraph id="6A483699FF94FFCAFCFF811302CCF9C5" blockId="25.[827,1443,1587,1640]" pageId="25" pageNumber="1270">
<emphasis id="5883EA8BFF94FFCAFCFF811303C7F9E4" box="[827,1159,1587,1609]" italics="true" pageId="25" pageNumber="1270">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF94FFCAFC4C817502CCF9C5" box="[904,1420,1618,1640]" pageId="25" pageNumber="1270">3F13F62A-0CCB-4452-9224-7CE361467AF7</uuid>
</paragraph>
<paragraph id="6A483699FF94FFC9FCFF81B6079BFED9" blockId="25.[827,1444,1680,1886]" lastBlockId="26.[145,761,197,372]" lastPageId="26" lastPageNumber="1271" pageId="25" pageNumber="1270">
<emphasis id="5883EA8BFF94FFCAFCFF81B604FBF90B" box="[827,955,1681,1702]" italics="true" pageId="25" pageNumber="1270">Diagnosis:</emphasis>
Despite the diversity of genera in this tribe, a synapomorphy that unites them is the shape of the left paramere of the males (
<figureCitation id="F2CC2A1CFF94FFCAFAF081E902D3F94E" box="[1332,1427,1742,1764]" captionStart="Figure 23" captionStartId="33.[163,241,986,1008]" captionTargetBox="[163,1444,195,946]" captionTargetId="figure-408@33.[163,1444,195,946]" captionTargetPageId="33" captionText="Figure 23. Australonaucoris congrex male: dorsal habitus (a), genital capsule with parameres and terga IX and X removed (b), left paramere (c), right paramere (d). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994670" httpUri="https://zenodo.org/record/6994670/files/figure.png" pageId="25" pageNumber="1270">Fig. 23c</figureCitation>
). Specifically, they tend to be relatively broad and lack the pronounced lobe of the left side that is present in the other tribes of
<taxonomicName id="ADF74D1AFF94FFCAFBDA800D03E8F8ED" authorityName="LEACH" authorityYear="1815" box="[1054,1192,1834,1856]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
(
<figureCitation id="F2CC2A1CFF94FFCAFB7E800D025DF892" box="[1210,1309,1834,1856]" captionStart="Figure 22" captionStartId="31.[164,242,1851,1873]" captionTargetBox="[163,1444,895,1813]" captionTargetId="figure-311@31.[163,1444,895,1813]" captionTargetPageId="31" captionText="Figure 22. Afronaucoris obscuratus male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c), distal part of aedeagus (d). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994668" httpUri="https://zenodo.org/record/6994668/files/figure.png" pageId="25" pageNumber="1270">Figs 22c</figureCitation>
,
<figureCitation id="F2CC2A1CFF94FFCAFAE8800D0215F892" box="[1324,1365,1834,1855]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="25" pageNumber="1270">24e</figureCitation>
), thus rendering the structure as appearing less modified, whereas this is probably the derived character state given its phylogenetic position in the dendrogram and that the lobe is present in the other tribes. Species of
<taxonomicName id="ADF74D1AFF97FFC9FF5586060641FE9B" authorityName="Montandon" authorityYear="1911" box="[145,257,289,310]" class="Insecta" family="Naucoridae" genus="Sagocoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FF5586060641FE9B" box="[145,257,289,310]" italics="true" pageId="26" pageNumber="1271">Sagocoris</emphasis>
</taxonomicName>
may have secondarily acquired an incipient lobe unrelated to the pronounced lobe of the other tribes.
</paragraph>
<paragraph id="6A483699FF97FFC9FF5586BF053AFCF6" blockId="26.[145,762,408,859]" pageId="26" pageNumber="1271">
<emphasis id="5883EA8BFF97FFC9FF5586BF0651FE00" box="[145,273,408,429]" italics="true" pageId="26" pageNumber="1271">Comments:</emphasis>
The seven New Guinean endemic genera of the former tribes Sagocorini and Tanycricini are here transferred from
<taxonomicName id="ADF74D1AFF97FFC9FE5386F20506FE46" authorityName="MONTANDON" authorityYear="1897" box="[407,582,469,491]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1260" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
to
<taxonomicName id="ADF74D1AFF97FFC9FDA886F205B5FE46" authorityName="LEACH" authorityYear="1815" box="[620,757,469,491]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
. They were recovered with an Australian species of
<taxonomicName id="ADF74D1AFF97FFC9FF55853407BFFD85" authorityName="Geoffroy" authorityYear="1762" box="[145,255,531,552]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FF55853407BFFD85" box="[145,255,531,552]" italics="true" pageId="26" pageNumber="1271">Naucoris</emphasis>
</taxonomicName>
at the root. For this strongly supported clade (pp = 1.0, bs = 100), I propose the new tribe
<taxonomicName id="ADF74D1AFF97FFC9FF5585770645FDCB" authorityName="Sites" authorityYear="2022" box="[145,261,592,614]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Idiocarini">Idiocarini</taxonomicName>
and
<taxonomicName id="ADF74D1AFF97FFC9FE878577054FFDC8" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[323,527,592,613]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF97FFC9FE878577054FFDC8" box="[323,527,592,613]" italics="true" pageId="26" pageNumber="1271">Australonaucoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF97FFC9FDDC857705C0FDC8" box="[536,640,592,613]" pageId="26" pageNumber="1271" rank="genus">gen. nov.</taxonomicNameLabel>
described below, in which to transfer
<taxonomicName id="ADF74D1AFF97FFC9FE378548058CFD29" authorityName="Stal" authorityYear="1876" box="[499,716,623,644]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="congrex">
<emphasis id="5883EA8BFF97FFC9FE378548058CFD29" box="[499,716,623,644]" italics="true" pageId="26" pageNumber="1271">Naucoris congrex</emphasis>
</taxonomicName>
. In addition to
<taxonomicName id="ADF74D1AFF97FFC9FEDC85A906D1FD0E" authorityName="Stal" authorityYear="1876" box="[280,401,654,675]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="congrex">
<emphasis id="5883EA8BFF97FFC9FEDC85A906D1FD0E" box="[280,401,654,675]" italics="true" pageId="26" pageNumber="1271">N. congrex</emphasis>
</taxonomicName>
, five other species of
<taxonomicName id="ADF74D1AFF97FFC9FD5485A905B9FD0E" authorityName="Geoffroy" authorityYear="1762" box="[656,761,654,675]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FD5485A905B9FD0E" box="[656,761,654,675]" italics="true" pageId="26" pageNumber="1271">Naucoris</emphasis>
</taxonomicName>
are known from
<collectingCountry id="12E07609FF97FFC9FE8B858B06FFFD6F" box="[335,447,684,706]" name="Australia" pageId="26" pageNumber="1271">Australia</collectingCountry>
:
<taxonomicName id="ADF74D1AFF97FFC9FE0E858B078BFD4D" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="australicus">
<emphasis id="5883EA8BFF97FFC9FE0E858B05FBFD6C" box="[458,699,684,705]" italics="true" pageId="26" pageNumber="1271">Naucoris australicus</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FD07858C078BFD4D" author="Stal C" pageId="26" pageNumber="1271" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">Stål, 1876</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FF1985EC0538FD4D" authority="Lansbury, 1991" authorityName="Lansbury" authorityYear="1991" box="[221,632,714,736]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="magela">
<emphasis id="5883EA8BFF97FFC9FF1985EC06EEFD72" box="[221,430,714,736]" italics="true" pageId="26" pageNumber="1271">Naucoris magela</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FE7E85ED0538FD4D" author="Lansbury I" box="[442,632,714,736]" pageId="26" pageNumber="1271" pagination="103 - 114" refId="ref22277" refString="Lansbury I. 1991. Naucoridae and Notonectidae (HemipteraHeteroptera) of the Northern Territory, Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences 8: 103 - 114." type="journal article" year="1991">Lansbury, 1991</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FD4E85EC0647FD53" authorityName=", Zettel" authorityYear="2001" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FD4E85EC0647FD53" italics="true" pageId="26" pageNumber="1271">Naucoris scutellaris</emphasis>
</taxonomicName>
(=
<taxonomicName id="ADF74D1AFF97FFC9FEE985CD05B2FD53" authority="J. Polhemus, 1984" authorityName="J. Polhemus" authorityYear="1984" box="[301,754,745,767]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="rhizomatus">
<emphasis id="5883EA8BFF97FFC9FEE985CD055EFD53" box="[301,542,745,767]" italics="true" pageId="26" pageNumber="1271">Naucoris rhizomatus</emphasis>
J.
<bibRefCitation id="0E664B68FF97FFC9FDFA85CE05B2FD53" author="Polhemus JT" box="[574,754,745,767]" pageId="26" pageNumber="1271" pagination="157 - 160" refId="ref23810" refString="Polhemus JT. 1984. A review of the Naucorinae of Australia (Heteroptera: Naucoridae). Journal of the Australian Entomological Society 23: 157 - 160." type="journal article" year="1984">Polhemus, 1984</bibRefCitation>
</taxonomicName>
; some authorities placed this species in
<taxonomicName id="ADF74D1AFF97FFC9FDAE842F0673FC91" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" class="Insecta" family="Naucoridae" genus="Thurselinus" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FDAE842F05B8FCB0" box="[618,760,776,797]" italics="true" pageId="26" pageNumber="1271">Thurselinus</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FF5584010673FC91" author="Distant WL" box="[145,307,806,828]" pageId="26" pageNumber="1271" pagination="258 - 259" refId="ref21779" refString="Distant WL. 1904. New Rhynchota-Cryptocerata. The Entomologist 37: 258 - 259." type="journal article" year="1904">Distant, 1904</bibRefCitation>
</taxonomicName>
),
<taxonomicName id="ADF74D1AFF97FFC9FE8F840005B9FC91" authority="Lansbury, 1985" authorityName="Lansbury" authorityYear="1985" box="[331,761,806,828]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="subaureus">
<emphasis id="5883EA8BFF97FFC9FE8F84000579FC96" box="[331,569,806,828]" italics="true" pageId="26" pageNumber="1271">Naucoris subaureus</emphasis>
Lansbury, 1985
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF97FFC9FF0784610533FCF7" authority="Montandon, 1913" authorityName="Montandon" authorityYear="1913" box="[195,627,837,859]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="5883EA8BFF97FFC9FF07846106E5FCF7" box="[195,421,837,859]" italics="true" pageId="26" pageNumber="1271">Naucoris subopacus</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FE6884620533FCF7" author="Montandon AL" box="[428,627,837,859]" pageId="26" pageNumber="1271" pagination="219 - 224" refId="ref23328" refString="Montandon AL. 1913. Hemipteres aquatiques. Notes et descriptions de deux especes nouvelles. Bulletin de la Section Scientifique de l'Academie Roumaine 1: 219 - 224." type="journal article" year="1913">Montandon, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="6A483699FF97FFC9FF5584A3066CFBDD" blockId="26.[145,762,900,1136]" pageId="26" pageNumber="1271">
<emphasis id="5883EA8BFF97FFC9FF5584A30608FC34" box="[145,328,900,921]" italics="true" pageId="26" pageNumber="1271">Included taxa:</emphasis>
This tribe contains seven genera endemic to New
<collectingCountry id="12E07609FF97FFC9FEB48485068FFC15" box="[368,463,930,952]" name="Guinea-Bissau" pageId="26" pageNumber="1271">Guinea</collectingCountry>
, including
<taxonomicName id="ADF74D1AFF97FFC9FDA28484062CFC7A" authority="Montandon, 1897 d" authorityName="Montandon" authorityYear="1897" class="Insecta" family="Naucoridae" genus="Aptinocoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FDA2848405B8FC15" box="[614,760,931,952]" italics="true" pageId="26" pageNumber="1271">Aptinocoris</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FF5584E6062CFC7A" author="Montandon AL" box="[145,364,961,983]" pageId="26" pageNumber="1271" pagination="50 - 66" refId="ref23079" refString="Montandon AL. 1897 d. Hemipteres-Heteropteres exotiques. Notes et descriptions, Annales de la Societe Entomologique de Belgique 41: 50 - 66." type="journal article" year="1897">Montandon, 1897d</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FEBE84E606ACFC7B" authorityName="La Rivers" authorityYear="1971" box="[378,492,961,982]" class="Insecta" family="Naucoridae" genus="Cavocoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FEBE84E606ACFC7B" box="[378,492,961,982]" italics="true" pageId="26" pageNumber="1271">Cavocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FE3F84E6052AFC7B" authorityName="Montandon" authorityYear="1897" box="[507,618,961,982]" class="Insecta" family="Naucoridae" genus="Idiocarus" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FE3F84E6052AFC7B" box="[507,618,961,982]" italics="true" pageId="26" pageNumber="1271">Idiocarus</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FDBC84E505B2FC7A" authorityName="La Rivers" authorityYear="1971" box="[632,754,962,983]" class="Insecta" family="Naucoridae" genus="Nesocricos" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FDBC84E505B2FC7A" box="[632,754,962,983]" italics="true" pageId="26" pageNumber="1271">Nesocricos</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FF5584C707BFFC58" authorityName="Montandon" authorityYear="1911" box="[145,255,992,1013]" class="Insecta" family="Naucoridae" genus="Sagocoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FF5584C707BFFC58" box="[145,255,992,1013]" italics="true" pageId="26" pageNumber="1271">Sagocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FECD84C706C2FC58" authorityName="La Rivers" authorityYear="1971" box="[265,386,992,1013]" class="Insecta" family="Naucoridae" genus="Tanycricos" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FECD84C706C2FC58" box="[265,386,992,1013]" italics="true" pageId="26" pageNumber="1271">Tanycricos</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF97FFC9FE7184C705FEFC58" authority="La Rivers, 1971" authorityName="La Rivers" authorityYear="1971" box="[437,702,992,1014]" class="Insecta" family="Naucoridae" genus="Warisia" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FE7184C7054DFC58" box="[437,525,992,1013]" italics="true" pageId="26" pageNumber="1271">Warisia</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FDD684C705FEFC58" author="La Rivers I" box="[530,702,992,1014]" pageId="26" pageNumber="1271" pagination="1 - 120" refId="ref22462" refString="La Rivers I. 1971. Studies of Naucoridae (Hemiptera). Biological Society of Nevada Memoirs 2: 1 - 120." type="journal article" year="1971">La Rivers, 1971</bibRefCitation>
</taxonomicName>
. Also included is
<taxonomicName id="ADF74D1AFF97FFC9FED184D9069BFBBE" authorityName="Sites" authorityYear="2022" box="[277,475,1022,1043]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FED184D9069BFBBE" box="[277,475,1022,1043]" italics="true" pageId="26" pageNumber="1271">Australonaucoris</emphasis>
</taxonomicName>
from
<collectingCountry id="12E07609FF97FFC9FDDB84D905CFFBB9" box="[543,655,1022,1044]" name="Australia" pageId="26" pageNumber="1271">Australia</collectingCountry>
. Three of the other species of
<taxonomicName id="ADF74D1AFF97FFC9FEBE833906A3FB9E" authorityName="Geoffroy" authorityYear="1762" box="[378,483,1054,1075]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FEBE833906A3FB9E" box="[378,483,1054,1075]" italics="true" pageId="26" pageNumber="1271">Naucoris</emphasis>
</taxonomicName>
from
<collectingCountry id="12E07609FF97FFC9FDEC833A05D6FB9E" box="[552,662,1053,1075]" name="Australia" pageId="26" pageNumber="1271">Australia</collectingCountry>
also are transferred to
<taxonomicName id="ADF74D1AFF97FFC9FEF2831B06B9FBFC" authorityName="Sites" authorityYear="2022" box="[310,505,1084,1105]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FEF2831B06B9FBFC" box="[310,505,1084,1105]" italics="true" pageId="26" pageNumber="1271">Australonaucoris</emphasis>
</taxonomicName>
(see description of the genus below).
</paragraph>
</subSubSection>
</treatment>
</document>

133
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<treatment id="E25E878FFF94FFCAFECF81E80363FAD0" ID-DOI="http://doi.org/10.5281/zenodo.6994666" ID-GBIF-Taxon="199315268" ID-Zenodo-Dep="6994666" LSID="urn:lsid:plazi:treatment:E25E878FFF94FFCAFECF81E80363FAD0" httpUri="http://treatment.plazi.org/id/E25E878FFF94FFCAFECF81E80363FAD0" lastPageNumber="1270" pageId="25" pageNumber="1270">
<subSubSection id="22ED6512FF94FFCAFECF81E805E2F94A" box="[267,674,1742,1768]" pageId="25" pageNumber="1270" type="nomenclature">
<paragraph id="6A483699FF94FFCAFECF81E805E2F94A" blockId="25.[267,674,1742,1768]" box="[267,674,1742,1768]" pageId="25" pageNumber="1270">
<heading id="310081F5FF94FFCAFECF81E805E2F94A" box="[267,674,1742,1768]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="1270" reason="3">
TRIBE
<taxonomicName id="ADF74D1AFF94FFCAFE9381E9055DF948" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[343,541,1742,1766]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Asthenocorini">ASTHENOCORINI</taxonomicName>
<emphasis id="5883EA8BFF94FFCAFDE081F505E2F94A" bold="true" box="[548,674,1743,1767]" pageId="25" pageNumber="1270">
<taxonomicNameLabel id="43B057F0FF94FFCAFDE081F505E2F94A" box="[548,674,1743,1767]" pageId="25" pageNumber="1270" rank="tribe">TRIB. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF94FFCAFEBD81DC0363FAD0" pageId="25" pageNumber="1270" type="description">
<paragraph id="6A483699FF94FFCAFEBD81DC0574F8B9" blockId="25.[377,564,1787,1812]" box="[377,564,1787,1812]" pageId="25" pageNumber="1270">
<figureCitation id="F2CC2A1CFF94FFCAFEBD81DC0692F8B9" box="[377,466,1787,1812]" captionStart="Figure 17" captionStartId="23.[827,914,1816,1838]" captionTargetBox="[929,1337,1400,1776]" captionTargetId="figure-735@23.[929,1337,1400,1776]" captionTargetPageId="23" captionText="Figure 17. Lateral view of head and prothorax of Philippinocoris usingeri showing anteromedian tooth of prosternum (white arrow)." figureDoi="http://doi.org/10.5281/zenodo.6994654" httpUri="https://zenodo.org/record/6994654/files/figure.png" pageId="25" pageNumber="1270">FIGS 17</figureCitation>
,
<figureCitation id="F2CC2A1CFF94FFCAFE1B81DC054BF8BF" box="[479,523,1787,1812]" captionStart="Figure 19" captionStartId="25.[164,245,774,796]" captionTargetBox="[163,1444,195,734]" captionTargetId="figure-573@25.[163,1444,195,734]" captionTargetPageId="25" captionText="Figure 19. Revised generic representation of the former genus Naucoris. Dorsal habitus of Naucoris maculatus (a), Australonaucoris congrex (b), Afronaucoris parvulus (c), Indonaucoris minutus sp. nov. (d)." figureDoi="http://doi.org/10.5281/zenodo.6994658" httpUri="https://zenodo.org/record/6994658/files/figure.png" pageId="25" pageNumber="1270">19D</figureCitation>
,
<figureCitation id="F2CC2A1CFF94FFCAFDDC81DC0574F8B9" box="[536,564,1787,1812]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="25" pageNumber="1270">24</figureCitation>
</paragraph>
<paragraph id="6A483699FF94FFCAFF67801605B4F8C8" blockId="25.[163,779,1840,1893]" pageId="25" pageNumber="1270">
<emphasis id="5883EA8BFF94FFCAFF67801606AFF8EB" box="[163,495,1840,1862]" italics="true" pageId="25" pageNumber="1270">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF94FFCAFF34806805B4F8C8" box="[240,756,1871,1893]" pageId="25" pageNumber="1270">195C2481-7E1F-4692-A007-4AA952D643AE</uuid>
</paragraph>
<paragraph id="6A483699FF94FFCAFCFF844B037AFC13" blockId="25.[827,1443,875,958]" pageId="25" pageNumber="1270">
<emphasis id="5883EA8BFF94FFCAFCFF844B04FBFC2C" box="[827,955,876,897]" italics="true" pageId="25" pageNumber="1270">Diagnosis:</emphasis>
All members of
<taxonomicName id="ADF74D1AFF94FFCAFB49844C0277FC2C" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1165,1335,875,897]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1258" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Asthenocorini">Asthenocorini</taxonomicName>
have an anteromedial tooth of the prosternum (
<figureCitation id="F2CC2A1CFF94FFCAFB3C84AD0204FC32" box="[1272,1348,906,928]" captionStart="Figure 17" captionStartId="23.[827,914,1816,1838]" captionTargetBox="[929,1337,1400,1776]" captionTargetId="figure-735@23.[929,1337,1400,1776]" captionTargetPageId="23" captionText="Figure 17. Lateral view of head and prothorax of Philippinocoris usingeri showing anteromedian tooth of prosternum (white arrow)." figureDoi="http://doi.org/10.5281/zenodo.6994654" httpUri="https://zenodo.org/record/6994654/files/figure.png" pageId="25" pageNumber="1270">Fig. 17</figureCitation>
), except
<taxonomicName id="ADF74D1AFF94FFCAFCFF848F0491FC10" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[827,977,936,957]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF94FFCAFCFF848F0491FC10" box="[827,977,936,957]" italics="true" pageId="25" pageNumber="1270">Indonaucoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF94FFCAFC1C848E037AFC13" box="[984,1082,937,958]" pageId="25" pageNumber="1270" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF94FFCAFCFF84CF0351FB5F" blockId="25.[827,1443,999,1266]" pageId="25" pageNumber="1270">
<emphasis id="5883EA8BFF94FFCAFCFF84CF04FEFC50" box="[827,958,1000,1021]" italics="true" pageId="25" pageNumber="1270">Comments:</emphasis>
This predominantly Philippine clade is represented by the three former members of Sagocorini that were transferred to
<taxonomicName id="ADF74D1AFF94FFCAFB99830303A3FB97" authorityName="LEACH" authorityYear="1815" box="[1117,1251,1060,1082]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
by
<taxonomicName id="ADF74D1AFF94FFCAFACA830302E3FB97" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1294,1443,1060,1082]" genus="Zettel" pageId="25" pageNumber="1285" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF94FFCAFACA830302E3FB97" author="Zettel H" box="[1294,1443,1060,1082]" pageId="25" pageNumber="1270" pagination="89 - 94" refId="ref25528" refString="Zettel H. 2007. Halmaheria skalei gen. n. et sp. n., eine neue Schwimmwanze (Insecta: Heteroptera: Naucoridae) von der Insel Halmahera, Indonesien. Annalen des Naturhistorischen Museums in Wien 108 B: 89 - 94." type="journal article" year="2007">Zettel (2007)</bibRefCitation>
</taxonomicName>
and a tiny (
<quantity id="AD0F9B7CFF94FFCAFC0083640346FBF4" box="[964,1030,1091,1113]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="1270" unit="mm" value="5.0">5 mm</quantity>
length)
<taxonomicName id="ADF74D1AFF94FFCAFBAF83630394FBF4" authorityName="Geoffroy" authorityYear="1762" box="[1131,1236,1092,1113]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFBAF83630394FBF4" box="[1131,1236,1092,1113]" italics="true" pageId="25" pageNumber="1270">Naucoris</emphasis>
</taxonomicName>
-like species from
<collectingCountry id="12E07609FF94FFCAFCFF83450438FBD5" box="[827,888,1122,1144]" name="India" pageId="25" pageNumber="1270">India</collectingCountry>
at its root. For this strongly supported group (pp = 1.0, bs = 100), I propose the new tribe
<taxonomicName id="ADF74D1AFF94FFCAFAC083A702E2FB3B" authorityName="Sites" authorityYear="2022" box="[1284,1442,1152,1174]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="tribe" tribe="Asthenocorini">Asthenocorini</taxonomicName>
and the small
<taxonomicName id="ADF74D1AFF94FFCAFC348387031DFB18" authorityName="Geoffroy" authorityYear="1762" box="[1008,1117,1184,1205]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFC348387031DFB18" box="[1008,1117,1184,1205]" italics="true" pageId="25" pageNumber="1270">Naucoris</emphasis>
</taxonomicName>
-like species at the root is described here as
<taxonomicName id="ADF74D1AFF94FFCAFBCA83990251FB7E" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1038,1297,1214,1235]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">
<emphasis id="5883EA8BFF94FFCAFBCA83990251FB7E" box="[1038,1297,1214,1235]" italics="true" pageId="25" pageNumber="1270">Indonaucoris minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF94FFCAFADD8399022EFB7E" box="[1305,1390,1214,1235]" pageId="25" pageNumber="1270" rank="species">sp. nov.</taxonomicNameLabel>
(see description below).
</paragraph>
<paragraph id="6A483699FF94FFCAFCFF822C0363FAD0" blockId="25.[827,1443,1291,1405]" pageId="25" pageNumber="1270">
<emphasis id="5883EA8BFF94FFCAFCFF822C0499FA8C" box="[827,985,1291,1313]" italics="true" pageId="25" pageNumber="1270">Included taxa:</emphasis>
This tribe contains three genera endemic to the
<collectingCountry id="12E07609FF94FFCAFC55820D0363FAED" box="[913,1059,1322,1344]" name="Philippines" pageId="25" pageNumber="1270">Philippines</collectingCountry>
:
<taxonomicName id="ADF74D1AFF94FFCAFBF6820D0390FA92" authorityName="Usinger" authorityYear="1937" box="[1074,1232,1322,1343]" class="Insecta" family="Naucoridae" genus="Asthenocoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFBF6820D0390FA92" box="[1074,1232,1322,1343]" italics="true" pageId="25" pageNumber="1270">Asthenocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF94FFCAFB27820D02E2FA92" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="1987" box="[1251,1442,1322,1343]" class="Insecta" family="Naucoridae" genus="Philippinocoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFB27820D02E2FA92" box="[1251,1442,1322,1343]" italics="true" pageId="25" pageNumber="1270">Philippinocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF94FFCAFCB5826F04A9FAF0" authorityName="La Rivers" authorityYear="1969" box="[881,1001,1352,1373]" class="Insecta" family="Naucoridae" genus="Stalocoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFCB5826F04A9FAF0" box="[881,1001,1352,1373]" italics="true" pageId="25" pageNumber="1270">Stalocoris</emphasis>
</taxonomicName>
. Also included is
<taxonomicName id="ADF74D1AFF94FFCAFB07826F0221FAF0" authorityName="Sites" authorityYear="2022" box="[1219,1377,1352,1373]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFB07826F0221FAF0" box="[1219,1377,1352,1373]" italics="true" pageId="25" pageNumber="1270">Indonaucoris</emphasis>
</taxonomicName>
from
<collectingRegion id="A833F87BFF94FFCAFCFF82400494FAD0" box="[827,980,1383,1405]" country="India" name="Maharashtra" pageId="25" pageNumber="1270">Maharashtra</collectingRegion>
,
<collectingCountry id="12E07609FF94FFCAFC1B8240035FFAD0" box="[991,1055,1383,1405]" name="India" pageId="25" pageNumber="1270">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

217
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<mods:title id="60B645E64E4C06909B9AA0639C1A3B4E">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<subSubSection id="22ED6512FF95FFCBFC5483CD0269FAAF" box="[912,1321,1258,1283]" pageId="24" pageNumber="1269" type="nomenclature">
<paragraph id="6A483699FF95FFCBFC5483CD0269FAAF" blockId="24.[912,1321,1258,1283]" box="[912,1321,1258,1283]" pageId="24" pageNumber="1269">
<heading id="310081F5FF95FFCBFC5483CD0269FAAF" box="[912,1321,1258,1283]" centered="true" fontSize="9" level="2" pageId="24" pageNumber="1269" reason="3">
TRIBE
<taxonomicName id="ADF74D1AFF95FFCBFC1883CD03E4FAAD" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[988,1188,1258,1282]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Afronaucorini">AFRONAUCORINI</taxonomicName>
<emphasis id="5883EA8BFF95FFCBFB6F83C90269FAAF" bold="true" box="[1195,1321,1258,1282]" pageId="24" pageNumber="1269">
<taxonomicNameLabel id="43B057F0FF95FFCBFB6F83C90269FAAF" box="[1195,1321,1258,1282]" pageId="24" pageNumber="1269" rank="tribe">TRIB. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF95FFCAFC33823107ACF932" lastPageId="25" lastPageNumber="1270" pageId="24" pageNumber="1269" type="description">
<paragraph id="6A483699FF95FFCBFC3382310382FA82" blockId="24.[1015,1218,1302,1327]" box="[1015,1218,1302,1327]" pageId="24" pageNumber="1269">
<figureCitation id="F2CC2A1CFF95FFCBFC3382310321FA80" box="[1015,1121,1302,1327]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="24" pageNumber="1269">FIGS 11H</figureCitation>
,
<figureCitation id="F2CC2A1CFF95FFCBFBAA823103D9FA80" box="[1134,1177,1302,1327]" captionStart="Figure 19" captionStartId="25.[164,245,774,796]" captionTargetBox="[163,1444,195,734]" captionTargetId="figure-573@25.[163,1444,195,734]" captionTargetPageId="25" captionText="Figure 19. Revised generic representation of the former genus Naucoris. Dorsal habitus of Naucoris maculatus (a), Australonaucoris congrex (b), Afronaucoris parvulus (c), Indonaucoris minutus sp. nov. (d)." figureDoi="http://doi.org/10.5281/zenodo.6994658" httpUri="https://zenodo.org/record/6994658/files/figure.png" pageId="24" pageNumber="1269">19C</figureCitation>
,
<figureCitation id="F2CC2A1CFF95FFCBFB6282310382FA82" box="[1190,1218,1302,1327]" captionStart="Figure 22" captionStartId="31.[164,242,1851,1873]" captionTargetBox="[163,1444,895,1813]" captionTargetId="figure-311@31.[163,1444,895,1813]" captionTargetPageId="31" captionText="Figure 22. Afronaucoris obscuratus male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c), distal part of aedeagus (d). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994668" httpUri="https://zenodo.org/record/6994668/files/figure.png" pageId="24" pageNumber="1269">22</figureCitation>
</paragraph>
<paragraph id="6A483699FF95FFCBFCED826B0230FA2D" blockId="24.[809,1425,1356,1408]" pageId="24" pageNumber="1269">
<emphasis id="5883EA8BFF95FFCBFCED826B0335FACC" box="[809,1141,1356,1377]" italics="true" pageId="24" pageNumber="1269">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF95FFCBFCBC824D0230FA2D" box="[888,1392,1386,1408]" pageId="24" pageNumber="1269">92431AE1-96E6-4568-8873-7D5D38B34307</uuid>
</paragraph>
<paragraph id="6A483699FF95FFCBFCED828D03B2F9B6" blockId="24.[809,1424,1449,1563]" pageId="24" pageNumber="1269">
<emphasis id="5883EA8BFF95FFCBFCED828D04F2FA12" box="[809,946,1450,1471]" italics="true" pageId="24" pageNumber="1269">Diagnosis:</emphasis>
Autapomorphies for this clade are not evident at this time, but the distinction as an independent lineage is supported by the geographic association with the Afrotropical region.
</paragraph>
<paragraph id="6A483699FF95FFCBFCED816302D1F8C0" blockId="24.[809,1425,1604,1902]" pageId="24" pageNumber="1269">
<emphasis id="5883EA8BFF95FFCBFCED816304E9F9F4" box="[809,937,1604,1625]" italics="true" pageId="24" pageNumber="1269">Comments:</emphasis>
Three specimens of
<taxonomicName id="ADF74D1AFF95FFCBFB5981630245F9F4" authorityName="Geoffroy" authorityYear="1762" box="[1181,1285,1604,1625]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFB5981630245F9F4" box="[1181,1285,1604,1625]" italics="true" pageId="24" pageNumber="1269">Naucoris</emphasis>
</taxonomicName>
from Africa were recovered as a clade distinct from the other tribe-level clades in
<taxonomicName id="ADF74D1AFF95FFCBFBDD81A603E1F93A" authorityName="LEACH" authorityYear="1815" box="[1049,1185,1665,1687]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
(pp = 1.0, bs = 100). This group was represented in the analysis by two populations of
<taxonomicName id="ADF74D1AFF95FFCBFC2E81980421F95F" authority="Montandon, 1913" authorityName="Montandon" authorityYear="1913" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="species" species="obscuratus">
<emphasis id="5883EA8BFF95FFCBFC2E819803AFF97E" box="[1002,1263,1726,1748]" italics="true" pageId="24" pageNumber="1269">Naucoris obscuratus</emphasis>
<bibRefCitation id="0E664B68FF95FFCBFB3F81990421F95F" author="Montandon AL" pageId="24" pageNumber="1269" pagination="219 - 224" refId="ref23328" refString="Montandon AL. 1913. Hemipteres aquatiques. Notes et descriptions de deux especes nouvelles. Bulletin de la Section Scientifique de l'Academie Roumaine 1: 219 - 224." type="journal article" year="1913">Montandon, 1913</bibRefCitation>
</taxonomicName>
from the
<collectingRegion id="A833F87BFF95FFCBFC1381FA0361F95E" box="[983,1057,1757,1779]" country="Tanzania" name="Pwani" pageId="24" pageNumber="1269">Pwani</collectingRegion>
and
<collectingRegion id="A833F87BFF95FFCBFB9A81FA03DDF95F" box="[1118,1181,1757,1778]" country="Tanzania" name="Mara" pageId="24" pageNumber="1269">Mara</collectingRegion>
regions in
<collectingCountry id="12E07609FF95FFCBFAE781FA02D0F95E" box="[1315,1424,1757,1779]" name="Tanzania" pageId="24" pageNumber="1269">Tanzania</collectingCountry>
with little evolutionary distance from each other.
<taxonomicName id="ADF74D1AFF95FFCBFCED803C03F9F89D" authority="Signoret, 1860" authorityName="Signoret" authorityYear="1860" box="[809,1209,1818,1840]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="5883EA8BFF95FFCBFCED803C0345F882" box="[809,1029,1818,1840]" italics="true" pageId="24" pageNumber="1269">Naucoris parvulus</emphasis>
<bibRefCitation id="0E664B68FF95FFCBFBC9803D03F9F89D" author="Signoret V" box="[1037,1209,1818,1840]" pageId="24" pageNumber="1269" pagination="917 - 972" refId="ref24302" refString="Signoret V. 1860. Faune des Hemipteres de Madagascar. (Suite et fin.). Heteropteres. Annales de la Societe Entomologique de France 8: 917 - 972." type="journal article" year="1860">Signoret, 1860</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="12E07609FF95FFCBFAC5803D02D0F89D" box="[1281,1424,1818,1840]" name="Madagascar" pageId="24" pageNumber="1269">Madagascar</collectingCountry>
was recovered as their sister. As such, for these African species of
<taxonomicName id="ADF74D1AFF95FFCBFBC1807F0330F8C0" authorityName="Geoffroy" authorityYear="1762" box="[1029,1136,1880,1901]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFBC1807F0330F8C0" box="[1029,1136,1880,1901]" italics="true" pageId="24" pageNumber="1269">Naucoris</emphasis>
</taxonomicName>
, I propose
<taxonomicName id="ADF74D1AFF95FFCBFB33807F02D1F8C0" authorityName="Sites" authorityYear="2022" box="[1271,1425,1880,1901]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFB33807F02D1F8C0" box="[1271,1425,1880,1901]" italics="true" pageId="24" pageNumber="1269">Afronaucoris</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="6A483699FF94FFCAFF6787A706A5FA04" blockId="25.[163,392,128,155]" lastBlockId="25.[163,780,875,1449]" pageId="25" pageNumber="1270">
1270 R.W.
<taxonomicName id="ADF74D1AFF94FFCAFEF987A306C8FF34" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[317,392,132,153]" genus="Sites" pageId="25" pageNumber="1270" rank="genus" status="gen. nov.">SITES</taxonomicName>
<taxonomicNameLabel id="43B057F0FF94FFCAFF67844B064AFC2C" box="[163,266,876,897]" pageId="25" pageNumber="1270" rank="genus">gen. nov.</taxonomicNameLabel>
and the containing group in the new tribe
<taxonomicName id="ADF74D1AFF94FFCAFF6784AD061CFC0D" authorityName="Sites" authorityYear="2022" box="[163,348,906,928]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="tribe" tribe="Afronaucorini">Afronaucorini</taxonomicName>
. Also examined morphologically were
<taxonomicName id="ADF74D1AFF94FFCAFF2A848E0443FC13" authority="Linnavuori, 1971" authorityName="Linnavuori" authorityYear="1971" box="[238,771,936,958]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="ciliatistylus">
<emphasis id="5883EA8BFF94FFCAFF2A848E054FFC13" box="[238,527,936,958]" italics="true" pageId="25" pageNumber="1270">Naucoris ciliatistylus</emphasis>
<bibRefCitation id="0E664B68FF94FFCAFDD9848E0443FC13" author="Linnavuori R" box="[541,771,936,958]" pageId="25" pageNumber="1270" pagination="340 - 366" refId="ref22692" refString="Linnavuori R. 1971. Hemiptera of the Sudan, with remarks on some species of the adjacent countries. 1. The aquatic and subaquatic families. Annales Zoologici Fennici 8: 340 - 366." type="journal article" year="1971">Linnavuori, 1971</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF94FFCAFF6784EF050DFC71" authority="Poisson, 1949" authorityName="Poisson" authorityYear="1949" box="[163,589,967,989]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="kenyalis">
<emphasis id="5883EA8BFF94FFCAFF6784EF06CFFC71" box="[163,399,967,989]" italics="true" pageId="25" pageNumber="1270">Naucoris kenyalis</emphasis>
<bibRefCitation id="0E664B68FF94FFCAFE5F84E0050DFC71" author="Poisson R" box="[411,589,967,989]" pageId="25" pageNumber="1270" pagination="3 - 94" refId="ref23595" refString="Poisson R. 1949. Exploration du Parc National Albert, I. Mission G. F. de Witte 1933 - 1935. Hemipteres aquatiques. Institut des Parcs Nationaux Congo Belge 58: 3 - 94." type="journal article" year="1949">Poisson, 1949</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF94FFCAFD5384EF0516FC56" authority="Montandon, 1899" authorityName="Montandon" authorityYear="1899" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="5883EA8BFF94FFCAFD5384EF063BFC56" italics="true" pageId="25" pageNumber="1270">Naucoris madagascariensis</emphasis>
<bibRefCitation id="0E664B68FF94FFCAFE4084C10516FC56" author="Montandon AL" box="[388,598,998,1020]" pageId="25" pageNumber="1270" pagination="79 - 81" refId="ref23144" refString="Montandon AL. 1899. Deux especes nouvelles d'Hemipteres Heteropteres des collections du Museum de Paris. Bulletin du Museum d'Histoire Naturelle 2: 79 - 81." type="journal article" year="1899">Montandon, 1899</bibRefCitation>
</taxonomicName>
. These species are remarkably similar in appearance and are also transferred to
<taxonomicName id="ADF74D1AFF94FFCAFE568304056FFB95" authorityName="Sites" authorityYear="2022" box="[402,559,1059,1080]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFE568304056FFB95" box="[402,559,1059,1080]" italics="true" pageId="25" pageNumber="1270">Afronaucoris</emphasis>
</taxonomicName>
. Three additional species of
<taxonomicName id="ADF74D1AFF94FFCAFEDE836506C3FBFA" authorityName="Geoffroy" authorityYear="1762" box="[282,387,1090,1111]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFEDE836506C3FBFA" box="[282,387,1090,1111]" italics="true" pageId="25" pageNumber="1270">Naucoris</emphasis>
</taxonomicName>
are known from Africa (
<taxonomicName id="ADF74D1AFF94FFCAFD668365054EFBDB" authority="Schaum, 1853" authorityName="Schaum" authorityYear="1853" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="5883EA8BFF94FFCAFD668365067DFBDB" italics="true" pageId="25" pageNumber="1270">Naucoris fuscipennis</emphasis>
<bibRefCitation id="0E664B68FF94FFCAFE898346054EFBDB" author="Schaum H" box="[333,526,1120,1142]" pageId="25" pageNumber="1270" pagination="356 - 359" refId="ref24196" refString="Schaum H. 1853. Bericht uber die zur Bekanntmachung geeigneten Verhandlungen der Koniglich Preussischen. Akademie Wissenschaften zu Berlin 2: 356 - 359." type="journal article" year="1853">Schaum, 1853</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF94FFCAFDE383460604FB39" authority="Bolivar, 1879" authorityName="Bolivar" authorityYear="1879" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="perezii">
<emphasis id="5883EA8BFF94FFCAFDE38346044AFBDB" box="[551,778,1121,1142]" italics="true" pageId="25" pageNumber="1270">Naucoris perezii</emphasis>
<bibRefCitation id="0E664B68FF94FFCAFF6783580604FB39" author="Bolivar I" box="[163,324,1151,1173]" pageId="25" pageNumber="1270" pagination="133 - 146" refId="ref21458" refString="Bolivar I. 1879. Hemipteros nuevos del Museo de Madrid. Anales de la Sociedad Espanola de Historia Natural 8: 133 - 146." type="journal article" year="1879">Bolívar, 1879</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF94FFCAFE4783A705BBFB39" authority="Germar, 1838" authorityName="Germar" authorityYear="1838" box="[387,763,1151,1173]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="5883EA8BFF94FFCAFE4783A7050CFB39" box="[387,588,1151,1173]" italics="true" pageId="25" pageNumber="1270">Naucoris planus</emphasis>
<bibRefCitation id="0E664B68FF94FFCAFD92835805BBFB39" author="Germar EF" box="[598,763,1151,1172]" pageId="25" pageNumber="1270" pagination="121 - 192" refId="ref21952" refString="Germar EF. 1838. Hemiptera Heteroptera promontorii bonae spei nundum descripta, quae collegit C. F. Drege. Revue Entomologique, Publiee par Silbermann 5: 121 - 192." type="journal article" year="1838">Germar, 1838</bibRefCitation>
</taxonomicName>
). Although I know of no museum specimens, because of their African geographic records, these species are provisionally transferred to
<taxonomicName id="ADF74D1AFF94FFCAFDFA83FC059EFB5D" authorityName="Sites" authorityYear="2022" box="[574,734,1243,1264]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFDFA83FC059EFB5D" box="[574,734,1243,1264]" italics="true" pageId="25" pageNumber="1270">Afronaucoris</emphasis>
</taxonomicName>
for taxonomic stability. Further study of these species is needed to confirm their proper generic assignment. The Palaearctic
<taxonomicName id="ADF74D1AFF94FFCAFEA7821F044AFAE1" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[355,778,1335,1357]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="5883EA8BFF94FFCAFEA7821F050FFAE1" box="[355,591,1335,1357]" italics="true" pageId="25" pageNumber="1270">Naucoris maculatus</emphasis>
<bibRefCitation id="0E664B68FF94FFCAFD938210044AFAE1" author="Fabricius JC" box="[599,778,1335,1357]" pageId="25" pageNumber="1270" pagination="511 - 572" refId="ref21860" refString="Fabricius JC. 1798. Ryngota. Os rostro: vagina articulata. Supplementum Entomologiae Systematicae 12: 511 - 572." type="journal article" year="1798">Fabricius, 1798</bibRefCitation>
</taxonomicName>
occurs in northern Africa and was recovered in a different clade with other species of
<taxonomicName id="ADF74D1AFF94FFCAFD9A8252058AFA27" authorityName="Geoffroy" authorityYear="1762" box="[606,714,1397,1418]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFD9A8252058AFA27" box="[606,714,1397,1418]" italics="true" pageId="25" pageNumber="1270">Naucoris</emphasis>
</taxonomicName>
from Southeast Asia (
<taxonomicName id="ADF74D1AFF94FFCAFEA482B40699FA04" authorityName="POPOV" authorityYear="1970" box="[352,473,1427,1449]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini</taxonomicName>
).
</paragraph>
<caption id="3E886611FF94FFCAFF6084210310FC97" ID-DOI="http://doi.org/10.5281/zenodo.6994658" ID-Zenodo-Dep="6994658" httpUri="https://zenodo.org/record/6994658/files/figure.png" pageId="25" pageNumber="1270" startId="25.[164,245,774,796]" targetBox="[163,1444,195,734]" targetPageId="25">
<paragraph id="6A483699FF94FFCAFF6084210310FC97" blockId="25.[163,1441,774,826]" pageId="25" pageNumber="1270">
<emphasis id="5883EA8BFF94FFCAFF608421065EFCB6" bold="true" box="[164,286,774,796]" pageId="25" pageNumber="1270">Figure 19.</emphasis>
Revised generic representation of the former genus
<taxonomicName id="ADF74D1AFF94FFCAFCA284210488FCB1" authorityName="Geoffroy" authorityYear="1762" box="[870,968,774,796]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF94FFCAFCA284210488FCB1" box="[870,968,774,796]" italics="true" pageId="25" pageNumber="1270">Naucoris</emphasis>
</taxonomicName>
. Dorsal habitus of
<taxonomicName id="ADF74D1AFF94FFCAFB5884210238FCB1" authorityName="Fabricius" authorityYear="1798" box="[1180,1400,774,796]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="5883EA8BFF94FFCAFB5884210238FCB1" box="[1180,1400,774,796]" italics="true" pageId="25" pageNumber="1270">Naucoris maculatus</emphasis>
</taxonomicName>
(a),
<taxonomicName id="ADF74D1AFF94FFCAFF67840306EBFC94" authorityName="Sites" authorityYear="2022" baseAuthorityName="Stal" baseAuthorityYear="1876" box="[163,427,804,826]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="congrex">
<emphasis id="5883EA8BFF94FFCAFF67840306EBFC94" box="[163,427,804,826]" italics="true" pageId="25" pageNumber="1270">Australonaucoris congrex</emphasis>
</taxonomicName>
(b),
<taxonomicName id="ADF74D1AFF94FFCAFE1D84030583FC97" authorityName="Sites" authorityYear="2022" baseAuthorityName="Signoret" baseAuthorityYear="1860" box="[473,707,804,826]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="5883EA8BFF94FFCAFE1D84030583FC97" box="[473,707,804,826]" italics="true" pageId="25" pageNumber="1270">Afronaucoris parvulus</emphasis>
</taxonomicName>
(c),
<taxonomicName id="ADF74D1AFF94FFCAFD2A84030496FC97" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[750,982,804,826]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">
<emphasis id="5883EA8BFF94FFCAFD2A84030496FC97" box="[750,982,804,826]" italics="true" pageId="25" pageNumber="1270">Indonaucoris minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF94FFCAFC1884030368FC97" box="[988,1064,804,826]" pageId="25" pageNumber="1270" rank="species">sp. nov.</taxonomicNameLabel>
(d).
</paragraph>
</caption>
<paragraph id="6A483699FF94FFCAFF6782F507ACF932" blockId="25.[163,779,1490,1695]" pageId="25" pageNumber="1270">
<emphasis id="5883EA8BFF94FFCAFF6782F5061CFA4A" box="[163,348,1490,1511]" italics="true" pageId="25" pageNumber="1270">Included taxa:</emphasis>
The two species included in the molecular analysis were
<taxonomicName id="ADF74D1AFF94FFCAFE2E82D7061AF988" authority="(Signoret, 1860)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Signoret" baseAuthorityYear="1860" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="parvulus" status="comb. nov.">
<emphasis id="5883EA8BFF94FFCAFE2E82D7044AF9A8" box="[490,778,1520,1541]" italics="true" pageId="25" pageNumber="1270">Afronaucoris parvulus</emphasis>
(
<bibRefCitation id="0E664B68FF94FFCAFF6E81280612F988" author="Signoret V" box="[170,338,1551,1573]" pageId="25" pageNumber="1270" pagination="917 - 972" refId="ref24302" refString="Signoret V. 1860. Faune des Hemipteres de Madagascar. (Suite et fin.). Heteropteres. Annales de la Societe Entomologique de France 8: 917 - 972." type="journal article" year="1860">Signoret, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF94FFCAFEA581280697F988" box="[353,471,1551,1573]" pageId="25" pageNumber="1270" rank="species">comb. nov.</taxonomicNameLabel>
from
<collectingCountry id="12E07609FF94FFCAFDDF812805E7F988" box="[539,679,1551,1573]" name="Madagascar" pageId="25" pageNumber="1270">Madagascar</collectingCountry>
and two populations of
<taxonomicName id="ADF74D1AFF94FFCAFE95810907A2F9CF" authority="(Montandon, 1913)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1913" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="1270" phylum="Arthropoda" rank="species" species="obscuratus" status="comb. nov.">
<emphasis id="5883EA8BFF94FFCAFE958109052EF9EE" box="[337,622,1582,1603]" italics="true" pageId="25" pageNumber="1270">Afronaucoris obscuratus</emphasis>
(
<bibRefCitation id="0E664B68FF94FFCAFDBB8109079AF9CF" author="Montandon AL" pageId="25" pageNumber="1270" pagination="219 - 224" refId="ref23328" refString="Montandon AL. 1913. Hemipteres aquatiques. Notes et descriptions de deux especes nouvelles. Bulletin de la Section Scientifique de l'Academie Roumaine 1: 219 - 224." type="journal article" year="1913">Montandon, 1913</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF94FFCAFF2D816B061DF9CF" box="[233,349,1612,1634]" pageId="25" pageNumber="1270" rank="species">comb. nov.</taxonomicNameLabel>
from
<collectingCountry id="12E07609FF94FFCAFE64816B054DF9CF" box="[416,525,1612,1634]" name="Tanzania" pageId="25" pageNumber="1270">Tanzania</collectingCountry>
. Additional taxonomic actions are given below in the formal description of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>

170
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<taxonomicName id="ADF74D1AFF96FFC8FE9885C50693FD54" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[348,467,738,761]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="1272" phylum="Arthropoda" rank="genus" status="gen. nov.">TSINGALA</taxonomicName>
<emphasis id="5883EA8BFF96FFC8FE1F85C20512FD57" bold="true" box="[475,594,738,762]" pageId="27" pageNumber="1272">
<taxonomicNameLabel id="43B057F0FF96FFC8FE1F85C20512FD57" box="[475,594,738,762]" pageId="27" pageNumber="1272" rank="genus">GEN. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF96FFCFFEAE84290689FA9A" lastPageId="28" lastPageNumber="1273" pageId="27" pageNumber="1272" type="description">
<paragraph id="6A483699FF96FFC8FEAE84290503FC8A" blockId="27.[362,579,782,807]" box="[362,579,782,807]" pageId="27" pageNumber="1272">
<figureCitation id="F2CC2A1CFF96FFC8FEAE842906A0FC8B" box="[362,480,782,807]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="27" pageNumber="1272">FIGS 5C, D</figureCitation>
,
<figureCitation id="F2CC2A1CFF96FFC8FE298429055AFC8B" box="[493,538,782,807]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="27" pageNumber="1272">11H</figureCitation>
,
<figureCitation id="F2CC2A1CFF96FFC8FDE384290503FC8A" box="[551,579,782,807]" captionStart="Figure 20" captionStartId="28.[146,226,1107,1129]" captionTargetBox="[145,1425,195,1067]" captionTargetId="figure-347@28.[145,1425,195,1067]" captionTargetPageId="28" captionText="Figure 20. Tsingala humeralis male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994662" httpUri="https://zenodo.org/record/6994662/files/figure.png" pageId="27" pageNumber="1272">20</figureCitation>
</paragraph>
<paragraph id="6A483699FF96FFC8FF67844205BEFC35" blockId="27.[163,779,868,920]" pageId="27" pageNumber="1272">
<emphasis id="5883EA8BFF96FFC8FF67844206AFFCD7" box="[163,495,868,890]" italics="true" pageId="27" pageNumber="1272">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF96FFC8FF2B84A505BEFC35" box="[239,766,898,920]" pageId="27" pageNumber="1272">92EEA10A-6208-404D-8024-C2DBAF79590D</uuid>
</paragraph>
<paragraph id="6A483699FF96FFC8FF6784E505A1FC7B" blockId="27.[163,737,961,983]" box="[163,737,961,983]" pageId="27" pageNumber="1272">
<emphasis id="5883EA8BFF96FFC8FF6784E50673FC7A" box="[163,307,962,983]" italics="true" pageId="27" pageNumber="1272">
<typeStatus id="B54C883BFF96FFC8FF6784E50798FC7A" box="[163,216,962,983]" pageId="27" pageNumber="1272">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF96FFC8FEFB84E6059DFC7B" authority="(Signoret, 1860)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Signoret" baseAuthorityYear="1860" box="[319,733,961,983]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="1272" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="5883EA8BFF96FFC8FEFB84E6055FFC7B" box="[319,543,961,982]" italics="true" pageId="27" pageNumber="1272">Tsingala humeralis</emphasis>
(
<bibRefCitation id="0E664B68FF96FFC8FDE984E60592FC7A" author="Signoret V" box="[557,722,961,983]" pageId="27" pageNumber="1272" pagination="917 - 972" refId="ref24302" refString="Signoret V. 1860. Faune des Hemipteres de Madagascar. (Suite et fin.). Heteropteres. Annales de la Societe Entomologique de France 8: 917 - 972." type="journal article" year="1860">Signoret, 1860</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="6A483699FF96FFC8FF67832706C3FB99" blockId="27.[163,779,1024,1904]" pageId="27" pageNumber="1272">
<emphasis id="5883EA8BFF96FFC8FF678327066BFBB8" box="[163,299,1024,1045]" italics="true" pageId="27" pageNumber="1272">Description:</emphasis>
Form ovate; widest at embolia; moderately flattened for family.
</paragraph>
<paragraph id="6A483699FF96FFC8FF7F831A05BCF9B2" blockId="27.[163,779,1024,1904]" pageId="27" pageNumber="1272">Head broad, eyes and anterior margin continuously and evenly convex to anterolateral corners of pronotum. Eyes slightly divergent anteriorly; mesal margin straight or nearly so, anteroventral margin concave; narrow band of cuticle posterior to eye widening lateral to eye. Front of head rolled posteroventrally so labrum set back from functional anterior margin, giving a bull-nosed appearance. Labrum broadly, evenly rounded with transverse basal sulcus. Maxillary plate not flap like, but integral with head capsule, immediately posterolateral to labrum. Rostrum with three visible segments, with segments 1 and part of 2 concealed behind labrum. Antenna four segmented with segment 1 short and inconspicuous, 2 and 4 longer, 3 longest; segments 2–3 enlarged, 2–4 setose. Head and pronotum moderately convex transversely.</paragraph>
<paragraph id="6A483699FF96FFC8FF7F810F05D5F8DD" blockId="27.[163,779,1024,1904]" pageId="27" pageNumber="1272">Pronotum with anterior margin straight between eyes, then becoming concave as margin contours around eye; lateral margins convergent anteriorly and with slight convexity; posterior margin straight with rounded posterolateral corners. Scutellum triangular, lateral margins sinuate, twice as wide as long, evenly domed, heavily punctate. Hemelytra punctate, extending to tip of abdomen. Embolium well defined with embolar suture, yellowish anteriorly, dark posteriorly; clavus well defined with claval and intraclaval sutures; corium, clavus and embolium with sparse, fine hairs.</paragraph>
<paragraph id="6A483699FF96FFC8FC9787E203B3FD0A" blockId="27.[827,1443,197,1445]" pageId="27" pageNumber="1272">Propleuron immediately lateral to coxal cavity infuscated and covered with long clavate setae; laterally light coloured and glabrous. Prosternum sharply carinate, anterior surface of carina expressed as an elongate triangle to appress rostrum. Mesosternum roundedly carinate medially with a brush of long hairs and several isolated short, dark spines; carina terminates posteriorly as posteroventrally-directed tubercle. Metaxyphus broad with sharp median carina. Abdominal venter mostly glabrous along midline of III–V, covered with recumbent hairs and long, erect hairs laterally; thin, glabrous band along lateral margin; laterosternites each with 2–5 oval, glabrous depressions near spiracles; laterosternite V posterior margin modified as a variously-shaped lobe in females.</paragraph>
<paragraph id="6A483699FF96FFC8FC97859704C9FB63" blockId="27.[827,1443,197,1445]" pageId="27" pageNumber="1272">
Profemur elongate, inflated, anterior margin with dorsal and ventral rows of golden hairs sandwiching one row of short, dark spines. Protarsus single segmented in females, two segmented in males, each with paired movable pretarsal claws. Males with thick pad of setae on mesotibia and tarsomeres 2 and 3, and to a lesser extent on protibia and both tarsomeres; female with pads of setae greatly reduced. Metafemur with anteroventral oblique, straight row of long setae (
<figureCitation id="F2CC2A1CFF96FFC8FC8784E304D3FC74" box="[835,915,963,986]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="27" pageNumber="1272">Fig. 5d</figureCitation>
). Meso- and metafemora with posteroventral rows of short, spinose setae spaced approximately one spine width apart for most of length, tightly spaced in distal fifth of mesofemur (
<figureCitation id="F2CC2A1CFF96FFC8FBAA833803FBFB98" box="[1134,1211,1055,1077]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="27" pageNumber="1272">Fig. 5c</figureCitation>
); posterior margins with dense brush of light-coloured hair; hairs longer on mesofemur. Meso- and metatibiae with rows of long dark-red spines. Long golden swimming hairs sparse on mesotibia and tarsus, profuse on metatibia and tarsus.
</paragraph>
<paragraph id="6A483699FF96FFC8FC9783F004B3FA08" blockId="27.[827,1443,197,1445]" pageId="27" pageNumber="1272">Males with aedeagus stout, narrowing apically; parameres asymmetrical, short, generally hatchet shaped; medial lobes of tergum VIII (pseudoparameres) well-developed. Female mediosternite VII (subgenital plate) broad basally, convex transversely, lateral margins convergent posteriorly, shape at apex differs interspecifically.</paragraph>
<paragraph id="6A483699FF96FFC8FCFF82FB0327F904" blockId="27.[827,1443,1499,1705]" pageId="27" pageNumber="1272">
<emphasis id="5883EA8BFF96FFC8FCFF82FB04FDFA5C" box="[827,957,1500,1521]" italics="true" pageId="27" pageNumber="1272">Diagnosis:</emphasis>
In addition to features unique to the subfamily and tribe (see diagnoses above), the labral apex is rounded, the hemelytra lack a fracture from the distal ends of the embolia through the tip of the claval commissure, the anterior margin of the head is rounded rather than sharply margined, and the genus is endemic to
<collectingCountry id="12E07609FF96FFC8FC1381B40323F904" box="[983,1123,1683,1705]" name="Madagascar" pageId="27" pageNumber="1272">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="6A483699FF96FFC8FCFF81F803C0F8DD" blockId="27.[827,1442,1759,1904]" pageId="27" pageNumber="1272">
<emphasis id="5883EA8BFF96FFC8FCFF81F80485F959" box="[827,965,1759,1780]" italics="true" pageId="27" pageNumber="1272">Etymology:</emphasis>
In Malagasy, the native language of
<collectingCountry id="12E07609FF96FFC8FCFF81D90492F8B9" box="[827,978,1790,1812]" name="Madagascar" pageId="27" pageNumber="1272">Madagascar</collectingCountry>
,
<emphasis id="5883EA8BFF96FFC8FC2681D90308F8BE" box="[994,1096,1790,1811]" italics="true" pageId="27" pageNumber="1272">tsingala</emphasis>
is the word for a naucorid bug. As such, I honour the people of
<collectingCountry id="12E07609FF96FFC8FB2B803A023FF89E" box="[1263,1407,1821,1843]" name="Madagascar" pageId="27" pageNumber="1272">Madagascar</collectingCountry>
by elevating their common name for these insects with formal scientific recognition.
</paragraph>
<caption id="3E886611FF91FFCFFF568374059CFB2B" ID-DOI="http://doi.org/10.5281/zenodo.6994662" ID-Zenodo-Dep="6994662" httpUri="https://zenodo.org/record/6994662/files/figure.png" pageId="28" pageNumber="1273" startId="28.[146,226,1107,1129]" targetBox="[145,1425,195,1067]" targetPageId="28">
<paragraph id="6A483699FF91FFCFFF568374059CFB2B" blockId="28.[145,1425,1107,1158]" pageId="28" pageNumber="1273">
<emphasis id="5883EA8BFF91FFCFFF568374064AFBC5" bold="true" box="[146,266,1107,1129]" pageId="28" pageNumber="1273">Figure 20.</emphasis>
<taxonomicName id="ADF74D1AFF91FFCFFED0837406A5FBC4" authorityName="Sites" authorityYear="2022" baseAuthorityName="Signoret" baseAuthorityYear="1860" box="[276,485,1107,1129]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="5883EA8BFF91FFCFFED0837406A5FBC4" box="[276,485,1107,1129]" italics="true" pageId="28" pageNumber="1273">Tsingala humeralis</emphasis>
</taxonomicName>
male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c). Sizes are not proportional among figures.
</paragraph>
</caption>
<paragraph id="6A483699FF91FFCFFF5583E10689FA9A" blockId="28.[145,760,1221,1519]" pageId="28" pageNumber="1273">
<emphasis id="5883EA8BFF91FFCFFF5583E1065DFB76" box="[145,285,1222,1243]" italics="true" pageId="28" pageNumber="1273">Comments:</emphasis>
The three species formerly held in
<taxonomicName id="ADF74D1AFF91FFCFFF5583C20659FB57" authorityName="Stal" authorityYear="1876" box="[145,281,1252,1274]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF91FFCFFF5583C20659FB57" box="[145,281,1252,1274]" italics="true" pageId="28" pageNumber="1273">Heleocoris</emphasis>
</taxonomicName>
known from
<collectingCountry id="12E07609FF91FFCFFE1C83C30538FB54" box="[472,632,1252,1274]" name="Madagascar" pageId="28" pageNumber="1273">Madagascar</collectingCountry>
are here transferred to
<taxonomicName id="ADF74D1AFF91FFCFFE85822406EAFAB5" authorityName="Sites" authorityYear="2022" box="[321,426,1283,1304]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF91FFCFFE85822406EAFAB5" box="[321,426,1283,1304]" italics="true" pageId="28" pageNumber="1273">Tsingala</emphasis>
</taxonomicName>
. As such, the following new combinations are proposed:
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF91FFCFFF6D8267059BFA42" pageId="28" pageNumber="1273" type="reference_group">
<paragraph id="6A483699FF91FFCFFF6D82670580FAF8" blockId="28.[145,760,1221,1519]" box="[169,704,1344,1366]" pageId="28" pageNumber="1273">
<taxonomicName id="ADF74D1AFF91FFCFFF6D82670504FAF8" authority="(Signoret, 1860)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Signoret" baseAuthorityYear="1860" box="[169,580,1344,1366]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="species" species="humeralis" status="comb. nov.">
<emphasis id="5883EA8BFF91FFCFFF6D826706C9FAF8" box="[169,393,1344,1365]" italics="true" pageId="28" pageNumber="1273">Tsingala humeralis</emphasis>
(
<bibRefCitation id="0E664B68FF91FFCFFE5C8267057CFAFB" author="Signoret V" box="[408,572,1344,1366]" pageId="28" pageNumber="1273" pagination="917 - 972" refId="ref24302" refString="Signoret V. 1860. Faune des Hemipteres de Madagascar. (Suite et fin.). Heteropteres. Annales de la Societe Entomologique de France 8: 917 - 972." type="journal article" year="1860">Signoret, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF91FFCFFD8F82670580FAF8" box="[587,704,1344,1366]" pageId="28" pageNumber="1273" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF91FFCFFF6D82780619FA3E" blockId="28.[145,760,1221,1519]" pageId="28" pageNumber="1273">
<taxonomicName id="ADF74D1AFF91FFCFFF6D8278079EFA3F" authority="(Montandon, 1897 d)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="subSpecies" species="naucoroides" status="comb. nov." subSpecies="naucoroides">
<emphasis id="5883EA8BFF91FFCFFF6D82780512FAD9" box="[169,594,1375,1396]" italics="true" pageId="28" pageNumber="1273">Tsingala naucoroides naucoroides</emphasis>
(
<bibRefCitation id="0E664B68FF91FFCFFDA282780796FA3E" author="Montandon AL" pageId="28" pageNumber="1273" pagination="50 - 66" refId="ref23079" refString="Montandon AL. 1897 d. Hemipteres-Heteropteres exotiques. Notes et descriptions, Annales de la Societe Entomologique de Belgique 41: 50 - 66." type="journal article" year="1897">Montandon, 1897d</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF91FFCFFF21825A0619FA3E" box="[229,345,1405,1427]" pageId="28" pageNumber="1273" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FF91FFCFFF6D82BB0645FA7D" blockId="28.[145,760,1221,1519]" pageId="28" pageNumber="1273">
<taxonomicName id="ADF74D1AFF91FFCFFF6D82BB05B8FA1C" authority="(Poisson, 1962)" baseAuthorityName="Poisson" baseAuthorityYear="1962" box="[169,760,1436,1458]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="subSpecies" species="naucoroides" subSpecies="ambiguus">
<emphasis id="5883EA8BFF91FFCFFF6D82BB0574FA1C" box="[169,564,1436,1457]" italics="true" pageId="28" pageNumber="1273">Tsingala naucoroides ambiguus</emphasis>
(
<bibRefCitation id="0E664B68FF91FFCFFD8C82BB05B0FA1F" author="Poisson R" box="[584,752,1436,1458]" pageId="28" pageNumber="1273" pagination="33 - 44" refId="ref23634" refString="Poisson R. 1962. Les Heleocoris Stal de la region ethiopienne (Heteropteres Naucoridae Laccocorinae). Bulletin de la Societe Scientifique de Bretagne 37: 33 - 44." type="journal article" year="1962">Poisson, 1962</bibRefCitation>
)
</taxonomicName>
comb. nov.
</paragraph>
<paragraph id="6A483699FF91FFCFFF6D82FE059BFA42" blockId="28.[145,760,1221,1519]" box="[169,731,1497,1519]" pageId="28" pageNumber="1273">
<taxonomicName id="ADF74D1AFF91FFCFFF6D82FE0520FA43" authority="(Bergroth, 1893)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Bergroth" baseAuthorityYear="1893" box="[169,608,1497,1519]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="1273" phylum="Arthropoda" rank="species" species="nossibeanus" status="comb. nov.">
<emphasis id="5883EA8BFF91FFCFFF6D82FE06DEFA43" box="[169,414,1497,1518]" italics="true" pageId="28" pageNumber="1273">Tsingala nossibeanus</emphasis>
(
<bibRefCitation id="0E664B68FF91FFCFFE6A82FE0518FA43" author="Bergroth E" box="[430,600,1497,1519]" pageId="28" pageNumber="1273" pagination="210 - 213" refId="ref21441" refString="Bergroth E. 1893. Rhynchota aquatica Madagascariensia. Revue d'Entomologie 12: 210 - 213." type="journal article" year="1893">Bergroth, 1893</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF91FFCFFDA382FE059BFA42" box="[615,731,1497,1519]" pageId="28" pageNumber="1273" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E30C6E51E3C2FEA3ECCF6C8FA30D2F6E">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:name id="979AF9A61BD8612D24CA95FA2A2C454E" type="personal">
<mods:role id="46AF7933E06D19CF538A6C8F9738A809">
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<mods:namePart id="B34B7483FEB415536CA2527D88E075A5">Sites, Robert W.</mods:namePart>
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<mods:typeOfResource id="B0E3D4345C485013AED306FD6C2CD9B4">text</mods:typeOfResource>
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<mods:titleInfo id="76F23950A522A1C4A3FD7461827F8464">
<mods:title id="04B3972B98BAC639D80624F73E0713E2">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="D9C2751E40DDA166899CAED30B0E036C">
<mods:date id="B574E09F3BE7DA3C46578D46B80EB474">2022</mods:date>
<mods:detail id="3C1E8CD722967C8F680064FC95BC7325" type="pubDate">
<mods:number id="B190609DEC124B4704EDA9AAE8548DA8">2022-02-07</mods:number>
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<mods:detail id="DB9F81B27B2C3DFB2D0CCC277ED49AF2" type="volume">
<mods:number id="668460E4B9732803677AD4284BF767E7">195</mods:number>
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<mods:classification id="9FB01C90AEDB440F3EAC5F98AFD6C9ED">journal article</mods:classification>
<mods:identifier id="36735933614C7BC7B5A03F386B8D4FC7" type="CLB-Dataset">121760</mods:identifier>
<mods:identifier id="8BC2639644D2152AB11816F4056312B1" type="DOI">10.1093/zoolinnean/zlab105</mods:identifier>
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<mods:identifier id="463EDF563E0BB58412CF67AA84CB233D" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="7085A8F66BDC3BF6ECD872DDF9E1C0DB" type="Zenodo-Dep">6994599</mods:identifier>
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<treatment id="E25E878FFF97FFC8FED683B8058EFE9A" ID-DOI="http://doi.org/10.5281/zenodo.6994707" ID-GBIF-Taxon="199315265" ID-Zenodo-Dep="6994707" LSID="urn:lsid:plazi:treatment:E25E878FFF97FFC8FED683B8058EFE9A" httpUri="http://treatment.plazi.org/id/E25E878FFF97FFC8FED683B8058EFE9A" lastPageId="27" lastPageNumber="1271" pageId="26" pageNumber="1271">
<subSubSection id="22ED6512FF97FFC9FED683B80537FB15" box="[274,631,1183,1208]" pageId="26" pageNumber="1271" type="nomenclature">
<paragraph id="6A483699FF97FFC9FED683B80537FB15" blockId="26.[274,631,1183,1208]" box="[274,631,1183,1208]" pageId="26" pageNumber="1271">
TRIBE
<taxonomicName id="ADF74D1AFF97FFC9FE9A83B80537FB15" ID-CoL="L26" authority="POPOV, 1970" authorityName="POPOV" authorityYear="1970" box="[350,631,1183,1208]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Naucorini">
NAUCORINI
<bibRefCitation id="0E664B68FF97FFC9FE2283B80537FB15" author="Popov YA" box="[486,631,1183,1208]" pageId="26" pageNumber="1271" pagination="93 - 98" refId="ref23866" refString="Popov YA. 1970. Notes on the classification of the recent Naucoridae (Heteroptera: Nepomorpha). Bulletin of the Polish Academy of Sciences (Biological Sciences) 18: 93 - 98." type="journal article" year="1970">POPOV, 1970</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF97FFC9FE5783EC06B6FB4E" box="[403,502,1227,1252]" pageId="26" pageNumber="1271" type="description">
<paragraph id="6A483699FF97FFC9FE5783EC06B6FB4E" blockId="26.[403,502,1227,1252]" box="[403,502,1227,1252]" pageId="26" pageNumber="1271">
<figureCitation id="F2CC2A1CFF97FFC9FE5783EC06B6FB4E" box="[403,502,1227,1252]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="26" pageNumber="1271">FIG. 10H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF97FFC9FF5583DD06F5F98E" pageId="26" pageNumber="1271" type="diagnosis">
<paragraph id="6A483699FF97FFC9FF5583DD06F5F98E" blockId="26.[145,761,1273,1571]" pageId="26" pageNumber="1271">
<emphasis id="5883EA8BFF97FFC9FF5583DD0648FAA2" box="[145,264,1274,1295]" italics="true" pageId="26" pageNumber="1271">Diagnosis:</emphasis>
As has been the dilemma with the previous concept of the subfamily
<taxonomicName id="ADF74D1AFF97FFC9FE08823F051BFA83" authorityName="LEACH" authorityYear="1815" box="[460,603,1304,1326]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
, apomorphic characters to define the tribe as considered here are not evident. The dramatically asymmetrical parameres are consistent with all other tribes in
<taxonomicName id="ADF74D1AFF97FFC9FDAC825305B5FA27" authorityName="LEACH" authorityYear="1815" box="[616,757,1396,1418]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
. It lacks the relatively broad left paramere without a pronounced lobe on the left side which is characteristic of
<taxonomicName id="ADF74D1AFF97FFC9FF6B82F70666FA4B" authority="All" authorityName="Sites" authorityYear="2022" box="[175,294,1488,1510]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1269" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Idiocarini">Idiocarini</taxonomicName>
. It lacks the anteromedial tooth of the prosternum, which is characteristic of all but one member of
<taxonomicName id="ADF74D1AFF97FFC9FECB812A06F1F98E" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[271,433,1549,1571]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1258" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Asthenocorini">Asthenocorini</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF97FFC8FF55816B058EFE9A" lastPageId="27" lastPageNumber="1272" pageId="26" pageNumber="1271" type="discussion">
<paragraph id="6A483699FF97FFC9FF55816B04B8FB3C" blockId="26.[145,762,1612,1910]" lastBlockId="26.[809,1425,197,1170]" pageId="26" pageNumber="1271">
<emphasis id="5883EA8BFF97FFC9FF55816B0652F9CC" box="[145,274,1612,1633]" italics="true" pageId="26" pageNumber="1271">Comments:</emphasis>
The major clade at the root of
<taxonomicName id="ADF74D1AFF97FFC9FDB5816B05B8F9CF" authorityName="LEACH" authorityYear="1815" box="[625,760,1612,1634]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
includes two species of
<taxonomicName id="ADF74D1AFF97FFC9FE59814C0546F92D" authorityName="Geoffroy" authorityYear="1762" box="[413,518,1643,1664]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FE59814C0546F92D" box="[413,518,1643,1664]" italics="true" pageId="26" pageNumber="1271">Naucoris</emphasis>
</taxonomicName>
and two populations of the closely related
<taxonomicName id="ADF74D1AFF97FFC9FE5A81AE05FBF933" baseAuthorityName="Stal" baseAuthorityYear="1860" box="[414,699,1673,1694]" class="Insecta" family="Naucoridae" genus="Thurselinus" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FE5A81AE05FBF933" box="[414,699,1673,1694]" italics="true" pageId="26" pageNumber="1271">Thurselinus scutellaris</emphasis>
</taxonomicName>
, and forms the tribe
<taxonomicName id="ADF74D1AFF97FFC9FE80818F06FCF913" authorityName="POPOV" authorityYear="1970" box="[324,444,1704,1726]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini</taxonomicName>
. Although most treatments have considered the species as
<taxonomicName id="ADF74D1AFF97FFC9FDC081E00673F957" authority=", Zettel (2001)" authorityName=", Zettel" authorityYear="2001" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FDC081E005B3F976" box="[516,755,1734,1756]" italics="true" pageId="26" pageNumber="1271">Naucoris scutellaris</emphasis>
,
<bibRefCitation id="0E664B68FF97FFC9FF5581C20673F957" author="Zettel H" box="[145,307,1765,1787]" pageId="26" pageNumber="1271" pagination="1085 - 1095" refId="ref25492" refString="Zettel H. 2001. Die indische Zwergschwimmwanze Nanonaucoris gen. n., eine neue Gattung der Naucorini (Heteroptera: Naucoridae) aus Sudindien. Linzer biologische Beitrage 33: 1085 - 1095." type="journal article" year="2001">Zettel (2001)</bibRefCitation>
</taxonomicName>
resurrected
<bibRefCitation id="0E664B68FF97FFC9FE1F81C205E5F957" author="Distant WL" box="[475,677,1765,1787]" pageId="26" pageNumber="1271" pagination="258 - 259" refId="ref21779" refString="Distant WL. 1904. New Rhynchota-Cryptocerata. The Entomologist 37: 258 - 259." type="journal article" year="1904">Distant’s (1904)</bibRefCitation>
genus
<taxonomicName id="ADF74D1AFF97FFC9FF5580230663F8B4" authorityName="Distant" authorityYear="1904" box="[145,291,1796,1817]" class="Insecta" family="Naucoridae" genus="Thurselinus" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FF5580230663F8B4" box="[145,291,1796,1817]" italics="true" pageId="26" pageNumber="1271">Thurselinus</emphasis>
</taxonomicName>
in which to place
<taxonomicName id="ADF74D1AFF97FFC9FDCC802305EAF8B4" authorityName=", Zettel" authorityYear="2001" box="[520,682,1796,1817]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FDCC802305EAF8B4" box="[520,682,1796,1817]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
based on connexival corner shape and paramere condition.
<taxonomicName id="ADF74D1AFF97FFC9FF55806506D5F8FB" authorityName="Fabricius" authorityYear="1798" box="[145,405,1857,1879]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="5883EA8BFF97FFC9FF55806506D5F8FB" box="[145,405,1857,1879]" italics="true" pageId="26" pageNumber="1271">Naucoris maculatus</emphasis>
</taxonomicName>
is the nominate species of the family and its relationship with
<taxonomicName id="ADF74D1AFF97FFC9FDA0804705B9F8D8" authorityName="Distant" authorityYear="1904" box="[612,761,1888,1909]" class="Insecta" family="Naucoridae" genus="Thurselinus" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FDA0804705B9F8D8" box="[612,761,1888,1909]" italics="true" pageId="26" pageNumber="1271">Thurselinus</emphasis>
</taxonomicName>
renders the clade paraphyletic in this topology; thus,
<taxonomicName id="ADF74D1AFF97FFC9FCED87C304F9FF54" authorityName="Distant" authorityYear="1904" box="[809,953,228,249]" class="Insecta" family="Naucoridae" genus="Thurselinus" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FCED87C304F9FF54" box="[809,953,228,249]" italics="true" pageId="26" pageNumber="1271">Thurselinus</emphasis>
</taxonomicName>
is here re-synonymized with
<taxonomicName id="ADF74D1AFF97FFC9FAE087C302D0FF54" authorityName="Geoffroy" authorityYear="1762" box="[1316,1424,228,249]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FAE087C302D0FF54" box="[1316,1424,228,249]" italics="true" pageId="26" pageNumber="1271">Naucoris</emphasis>
</taxonomicName>
as originally described by
<bibRefCitation id="0E664B68FF97FFC9FBAD862503AEFEB5" author="Stal C" box="[1129,1262,258,280]" pageId="26" pageNumber="1271" pagination="1851 - 1853" refId="ref24975" refString="Stal C. 1860. Zoologi. I. Insecta. Hemiptera. Species nova descripsit. In: Kongliga Svenska fregatten Eugenies resa omkring jorden 1851 - 1853 under befal af C. A. Virgin aren 1851 - 1853. Stockholm: Norstedt &amp; Soner." type="book chapter" year="1860">Stål (1860)</bibRefCitation>
. Further, the
<specimenCount id="7CF1FD10FF97FFC9FCED86050498FE9A" box="[809,984,289,311]" pageId="26" pageNumber="1271" type="generic">two specimens</specimenCount>
identified as
<taxonomicName id="ADF74D1AFF97FFC9FB4586050260FE9B" authorityName=", Zettel" authorityYear="2001" box="[1153,1312,289,311]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FB4586050260FE9B" box="[1153,1312,289,311]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
are from
<collectingCountry id="12E07609FF97FFC9FCED866704D8FEFB" box="[809,920,320,342]" name="Australia" pageId="26" pageNumber="1271">Australia</collectingCountry>
and
<collectingCountry id="12E07609FF97FFC9FC0E86670374FEFB" box="[970,1076,320,342]" name="Thailand" pageId="26" pageNumber="1271">Thailand</collectingCountry>
and are of sufficient molecular evolutionary distance that they apparently are not conspecific. The population from
<collectingCountry id="12E07609FF97FFC9FB2C865A021AFE3E" box="[1256,1370,381,403]" name="Australia" pageId="26" pageNumber="1271">Australia</collectingCountry>
was first described as
<taxonomicName id="ADF74D1AFF97FFC9FBCD86BB0421FE62" authority="Polhemus, 1984" authorityName="J. Polhemus" authorityYear="1984" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="rhizomatus">
<emphasis id="5883EA8BFF97FFC9FBCD86BB024AFE1C" box="[1033,1290,412,433]" italics="true" pageId="26" pageNumber="1271">Naucoris rhizomatus</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FAD086BB0421FE62" author="Polhemus JT" pageId="26" pageNumber="1271" pagination="157 - 160" refId="ref23810" refString="Polhemus JT. 1984. A review of the Naucorinae of Australia (Heteroptera: Naucoridae). Journal of the Australian Entomological Society 23: 157 - 160." type="journal article" year="1984">Polhemus, 1984</bibRefCitation>
</taxonomicName>
, but was later synonymized with
<taxonomicName id="ADF74D1AFF97FFC9FB31869C03E8FE43" authority="(Polhemus &amp; Polhemus, 2013)" baseAuthorityName="Polhemus &amp; Polhemus" baseAuthorityYear="2013" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FB31869C02D0FE62" box="[1269,1424,442,464]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
(
<bibRefCitation id="0E664B68FF97FFC9FCF686FE03DCFE43" author="Polhemus DA &amp; Polhemus JT" box="[818,1180,473,495]" pageId="26" pageNumber="1271" pagination="665 - 686" refId="ref23727" refString="Polhemus DA, Polhemus JT. 2013. Guide to the aquatic Heteroptera of Singapore and Peninsular Malaysia. XI. Infraorder Nepomorpha - families Naucoridae and Aphelocheiridae. Raffles Bulletin of Zoology 61: 665 - 686." type="journal article" year="2013">Polhemus &amp; Polhemus, 2013</bibRefCitation>
)
</taxonomicName>
. Because the
<typeStatus id="B54C883BFF97FFC9FA9F86FE02CFFE43" box="[1371,1423,473,494]" pageId="26" pageNumber="1271">type</typeStatus>
locality for the species is Java, it is uncertain if either the Australian or Thai specimens represent actual
<taxonomicName id="ADF74D1AFF97FFC9FCED85110485FDE7" baseAuthorityName="Polhemus &amp; Polhemus" baseAuthorityYear="2013" box="[809,965,565,587]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FCED85110485FDE7" box="[809,965,565,587]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
. Comparison with material from Java is needed to determine appropriate name changes. Assuming the Australian and Thai specimens included in the analysis here are not conspecific, one of three nomenclatural scenarios is possible: the Thai population is unique and should be described,
<taxonomicName id="ADF74D1AFF97FFC9FCED85C9048DFCAF" authorityName="J. Polhemus" authorityYear="1984" box="[809,973,749,771]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="rhizomatus">
<emphasis id="5883EA8BFF97FFC9FCED85C9048DFCAF" box="[809,973,749,771]" italics="true" pageId="26" pageNumber="1271">N. rhizomatus</emphasis>
</taxonomicName>
should be reinstated as a valid species as originally described, or both if neither population is conspecific with
<taxonomicName id="ADF74D1AFF97FFC9FBCE840C03E6FC92" baseAuthorityName="Polhemus &amp; Polhemus" baseAuthorityYear="2013" box="[1034,1190,810,832]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FBCE840C03E6FC92" box="[1034,1190,810,832]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
from Java. Further complicating the identity of
<taxonomicName id="ADF74D1AFF97FFC9FB4B846D0274FCF3" baseAuthorityName="Polhemus &amp; Polhemus" baseAuthorityYear="2013" box="[1167,1332,841,863]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FB4B846D0274FCF3" box="[1167,1332,841,863]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
is that Shruti Paripatyadar (Pune,
<collectingCountry id="12E07609FF97FFC9FBBB844F0380FCD3" box="[1151,1216,872,894]" name="India" pageId="26" pageNumber="1271">India</collectingCountry>
) has populations of putative
<taxonomicName id="ADF74D1AFF97FFC9FC7F84A00320FC36" baseAuthorityName="Polhemus &amp; Polhemus" baseAuthorityYear="2013" box="[955,1120,902,924]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FC7F84A00320FC36" box="[955,1120,902,924]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
from
<collectingRegion id="A833F87BFF97FFC9FB6984A10210FC31" box="[1197,1360,902,924]" country="India" name="Maharashtra" pageId="26" pageNumber="1271">Maharashtra</collectingRegion>
with the typical spined left paramere but with a right paramere that differs. Also,
<taxonomicName id="ADF74D1AFF97FFC9FBBA84E3025EFC74" baseAuthorityName="Polhemus &amp; Polhemus" baseAuthorityYear="2013" box="[1150,1310,964,985]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FBBA84E3025EFC74" box="[1150,1310,964,985]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
has been recorded from the
<collectingCountry id="12E07609FF97FFC9FBD884C503EDFC55" box="[1052,1197,994,1016]" name="Philippines" pageId="26" pageNumber="1271">Philippines</collectingCountry>
and also has the spined left paramere (
<taxonomicName id="ADF74D1AFF97FFC9FBFD83260388FBBB" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1081,1224,1025,1047]" genus="Zettel" pageId="26" pageNumber="1285" rank="genus" status="gen. nov.">Zettel, 1999</taxonomicName>
). Thus, it seems that the widespread
<taxonomicName id="ADF74D1AFF97FFC9FBD2830703EEFB98" baseAuthorityName="Polhemus &amp; Polhemus" baseAuthorityYear="2013" box="[1046,1198,1056,1077]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FBD2830703EEFB98" box="[1046,1198,1056,1077]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
might be a complex of cryptic species. If so, several synonyms exist that might be available for Indian and Sri Lankan species (
<bibRefCitation id="0E664B68FF97FFC9FCF5835B04A8FB3C" author="Lundblad O" box="[817,1000,1148,1170]" pageId="26" pageNumber="1271" pagination="263 - 489" refId="ref22883" refString="Lundblad O. 1933. Zur Kenntnis der aquatilen und semiaquatilen Hemipteren von Sumatra, Java und Bali. Archiv fur Hydrobiologie, Supplement 12: 1 - 195, 263 - 489, 21 plates." type="book chapter" year="1933">Lundblad, 1933</bibRefCitation>
).
</paragraph>
<paragraph id="6A483699FF97FFC8FCED839D058EFE9A" blockId="26.[809,1425,1210,1907]" lastBlockId="27.[163,779,197,311]" lastPageId="27" lastPageNumber="1272" pageId="26" pageNumber="1271">
<emphasis id="5883EA8BFF97FFC9FCED839D0498FB7D" box="[809,984,1210,1232]" italics="true" pageId="26" pageNumber="1271">Included taxa:</emphasis>
The distribution of
<taxonomicName id="ADF74D1AFF97FFC9FB17839C0233FB62" authorityName="Fabricius" authorityYear="1798" box="[1235,1395,1210,1232]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="5883EA8BFF97FFC9FB17839C0233FB62" box="[1235,1395,1210,1232]" italics="true" pageId="26" pageNumber="1271">N. maculatus</emphasis>
</taxonomicName>
is Palaearctic,
<taxonomicName id="ADF74D1AFF97FFC9FC7483FD0301FB43" baseAuthorityName="Polhemus &amp; Polhemus" baseAuthorityYear="2013" box="[944,1089,1241,1263]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="5883EA8BFF97FFC9FC7483FD0301FB43" box="[944,1089,1241,1263]" italics="true" pageId="26" pageNumber="1271">N. scutellaris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF97FFC9FBB183FD03B1FB43" authorityName="La Rivers" authorityYear="1974" box="[1141,1265,1241,1263]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="sigaloeis">
<emphasis id="5883EA8BFF97FFC9FBB183FD03B1FB43" box="[1141,1265,1241,1263]" italics="true" pageId="26" pageNumber="1271">N. sigaloeis</emphasis>
</taxonomicName>
are Southeast Asian,
<taxonomicName id="ADF74D1AFF97FFC9FCBC83DF0358FAA0" authorityName="J. Polhemus" authorityYear="1984" box="[888,1048,1272,1293]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="rhizomatus">
<emphasis id="5883EA8BFF97FFC9FCBC83DF0358FAA0" box="[888,1048,1272,1293]" italics="true" pageId="26" pageNumber="1271">N. rhizomatus</emphasis>
</taxonomicName>
is Australian and five additional species of
<taxonomicName id="ADF74D1AFF97FFC9FC668230034CFA81" authorityName="Geoffroy" authorityYear="1762" box="[930,1036,1303,1324]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF97FFC9FC668230034CFA81" box="[930,1036,1303,1324]" italics="true" pageId="26" pageNumber="1271">Naucoris</emphasis>
</taxonomicName>
have not been mentioned above. Because of the apparent vagility of the ancestral stock and precarious action of assigning these additional five species to groups without compelling non-molecular evidence, the following are not treated here and are
<emphasis id="5883EA8BFF97FFC9FCED82970482FA69" box="[809,962,1455,1477]" italics="true" pageId="26" pageNumber="1271">incertae sedis</emphasis>
:
<taxonomicName id="ADF74D1AFF97FFC9FC0B8297025EFA68" authority="Spinola, 1837" authorityName="Spinola" authorityYear="1837" box="[975,1310,1455,1477]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="5883EA8BFF97FFC9FC0B8297033CFA68" box="[975,1148,1456,1477]" italics="true" pageId="26" pageNumber="1271">Naucoris acuta</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FB478288025EFA68" author="Spinola M" box="[1155,1310,1455,1477]" pageId="26" pageNumber="1271" refId="ref24867" refString="Spinola M. 1837. Essai sur les genres d'insectes appartenants a l'ordre des Hemipteres, Lin. ou Rhyngotes, Fab. et a la section des Heteropteres, Dufour. Chez Yves Graviers, Genes, 383 pp." type="book" year="1837">Spinola, 1837</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FAEE8297025AFA4E" authority="D. Polhemus &amp; J. Polhemus, 2013" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="2013" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="5883EA8BFF97FFC9FAEE829704CAFA49" italics="true" pageId="26" pageNumber="1271">Naucoris minutus</emphasis>
D. Polhemus &amp; J.
<bibRefCitation id="0E664B68FF97FFC9FBA782E9025AFA4E" author="Polhemus DA &amp; Polhemus JT" box="[1123,1306,1486,1508]" pageId="26" pageNumber="1271" pagination="665 - 686" refId="ref23727" refString="Polhemus DA, Polhemus JT. 2013. Guide to the aquatic Heteroptera of Singapore and Peninsular Malaysia. XI. Infraorder Nepomorpha - families Naucoridae and Aphelocheiridae. Raffles Bulletin of Zoology 61: 665 - 686." type="journal article" year="2013">Polhemus, 2013</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FAEC82E80314F9AE" authority="Stal, 1854" authorityName="Stal" authorityYear="1854" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="5883EA8BFF97FFC9FAEC82E8048EF9AF" italics="true" pageId="26" pageNumber="1271">Naucoris obscuripennis</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FC1C82CB0314F9AE" author="Stal C" box="[984,1108,1516,1539]" pageId="26" pageNumber="1271" pagination="231 - 235" refId="ref24908" refString="Stal C. 1854. Nya Hemiptera. Ofversigt af Kongliga Svenska Vetenskaps-Akademiens Forhandlingar, Stockholm 11: 231 - 235." type="journal article" year="1854">Stål, 1854</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF97FFC9FBA782CA0322F98C" authority="Zettel, Nieser &amp; D. Polhemus, 1999" authorityName="Zettel, Nieser &amp; D. Polhemus" authorityYear="1999" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="5883EA8BFF97FFC9FBA782CA027AF9AF" box="[1123,1338,1517,1538]" italics="true" pageId="26" pageNumber="1271">Naucoris pumilus</emphasis>
Zettel, Nieser &amp; D.Polhemus, 1999
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF97FFC9FB5E812B04FFF9ED" authority="Fieber, 1851" authorityName="Fieber" authorityYear="1851" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="5883EA8BFF97FFC9FB5E812B02D1F98C" box="[1178,1425,1548,1569]" italics="true" pageId="26" pageNumber="1271">Naucoris sumatranus</emphasis>
<bibRefCitation id="0E664B68FF97FFC9FCED810D04FFF9ED" author="Fieber FX" box="[809,959,1578,1600]" pageId="26" pageNumber="1271" refId="ref21899" refString="Fieber FX. 1851. Genera Hydrocoridum secundum ordinem naturalem in familias disposita. Actis Regiae Bohemicae Societatis Scientiarum. Prague: J. G. Calve." type="book" year="1851">Fieber, 1851</bibRefCitation>
</taxonomicName>
. The identity of
<taxonomicName id="ADF74D1AFF97FFC9FB56810C03B8F9ED" authorityName="Spinola" authorityYear="1837" box="[1170,1272,1579,1600]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="5883EA8BFF97FFC9FB56810C03B8F9ED" box="[1170,1272,1579,1600]" italics="true" pageId="26" pageNumber="1271">N. acuta</emphasis>
</taxonomicName>
is unknown because the eleven-word original description and lack of authoritatively determined specimens render the species unrecognizable based on available information, thus this is a
<emphasis id="5883EA8BFF97FFC9FC248182038AF917" box="[992,1226,1701,1722]" italics="true" pageId="26" pageNumber="1271">species inquirenda</emphasis>
. Figures in the original description of
<taxonomicName id="ADF74D1AFF97FFC9FBFD81E303FDF974" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="2013" box="[1081,1213,1732,1753]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="5883EA8BFF97FFC9FBFD81E303FDF974" box="[1081,1213,1732,1753]" italics="true" pageId="26" pageNumber="1271">N. minutus</emphasis>
</taxonomicName>
, and descriptions of
<taxonomicName id="ADF74D1AFF97FFC9FC8281C40346F95A" authorityName="Stal" authorityYear="1854" box="[838,1030,1762,1784]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="5883EA8BFF97FFC9FC8281C40346F95A" box="[838,1030,1762,1784]" italics="true" pageId="26" pageNumber="1271">N. obscuripennis</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF97FFC9FB8581C40381F95A" authorityName="Zettel, Nieser &amp; D. Polhemus" authorityYear="1999" box="[1089,1217,1762,1784]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="5883EA8BFF97FFC9FB8581C40381F95A" box="[1089,1217,1762,1784]" italics="true" pageId="26" pageNumber="1271">N. pumilus</emphasis>
</taxonomicName>
(see
<taxonomicName id="ADF74D1AFF97FFC9FB3A81C502C8F95A" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1278,1416,1762,1784]" genus="Zettel" pageId="26" pageNumber="1285" rank="genus" status="gen. nov.">Zettel, 1999</taxonomicName>
) show a lobe on the left side of the left paramere. These species are therefore not members of
<taxonomicName id="ADF74D1AFF97FFC9FB3D80380232F898" authority="All" authorityName="Sites" authorityYear="2022" box="[1273,1394,1823,1845]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1269" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Idiocarini">Idiocarini</taxonomicName>
. A female
<typeStatus id="B54C883BFF97FFC9FC47801904B0F8F9" box="[899,1008,1854,1876]" pageId="26" pageNumber="1271" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="ADF74D1AFF97FFC9FBDD8018038CF8F9" authorityName="Fieber" authorityYear="1851" box="[1049,1228,1855,1876]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="1271" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="5883EA8BFF97FFC9FBDD8018038CF8F9" box="[1049,1228,1855,1876]" italics="true" pageId="26" pageNumber="1271">N. sumatranus</emphasis>
</taxonomicName>
was designated based on the only remaining specimen from the type series (
<taxonomicName id="ADF74D1AFF96FFC8FF3387E206C2FF77" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[247,386,197,219]" genus="Zettel" pageId="27" pageNumber="1285" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF96FFC8FF3387E206C2FF77" author="Zettel H" box="[247,386,197,219]" pageId="27" pageNumber="1272" pagination="65 - 68" refId="ref25575" refString="Zettel H. 2011. The type of the enigmatic creeping water bug Naucoris sumatranus Fieber, 1851 (Hemiptera: Heteroptera: Naucoridae). Zootaxa 2981: 65 - 68." type="journal article" year="2011">Zettel, 2011</bibRefCitation>
</taxonomicName>
), if more than
<specimenCount id="7CF1FD10FF96FFC8FDF087E10590FF76" box="[564,720,197,219]" pageId="27" pageNumber="1272" type="generic">one specimen</specimenCount>
ever existed, thus male genitalia are not available for this species. Additional research is needed to determine proper taxonomic placement of these five species.
</paragraph>
</subSubSection>
</treatment>
</document>

107
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TRIBE
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</paragraph>
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<figureCitation id="F2CC2A1CFF98FFC6FB8182C903D8F9AB" box="[1093,1176,1518,1543]" captionStart="Figure 4" captionStartId="9.[163,249,1379,1401]" captionTargetBox="[163,781,195,1339]" captionTargetId="figure-575@9.[163,781,195,1339]" captionTargetPageId="9" captionText="Figure 4. Foretibiae and foretarsi of Laccocorinae: Ctenipocoris asiaticus (a), Heleolaccocoris ovatus (b), Namtokocoris dalanta (c)." figureDoi="http://doi.org/10.5281/zenodo.6994615" httpUri="https://zenodo.org/record/6994615/files/figure.png" pageId="21" pageNumber="1266">FIG. 4C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF98FFC5FCFF8102052FFDAB" lastPageId="22" lastPageNumber="1267" pageId="21" pageNumber="1266" type="description">
<paragraph id="6A483699FF98FFC6FCFF810202D7F9F4" blockId="21.[827,1443,1572,1625]" pageId="21" pageNumber="1266">
<emphasis id="5883EA8BFF98FFC6FCFF810203C8F997" box="[827,1160,1572,1594]" italics="true" pageId="21" pageNumber="1266">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF98FFC6FC4E816402D7F9F4" box="[906,1431,1603,1625]" pageId="21" pageNumber="1266">046C1346-835E-4C5D-B4B1-D5BC902FFF12</uuid>
</paragraph>
<paragraph id="6A483699FF98FFC6FCFF81A503A6F8C3" blockId="21.[827,1443,1666,1902]" pageId="21" pageNumber="1266">
<emphasis id="5883EA8BFF98FFC6FCFF81A504FDF93A" box="[827,957,1666,1687]" italics="true" pageId="21" pageNumber="1266">Diagnosis:</emphasis>
Diagnostic features were provided by
<bibRefCitation id="0E664B68FF98FFC6FCFF818703C6F918" author="Sites RW &amp; Vitheepradit A" box="[827,1158,1696,1718]" pageId="21" pageNumber="1266" pagination="1 - 29" refId="ref24696" refString="Sites RW, Vitheepradit A. 2007. Namtokocoris, a new genus of Naucoridae (Hemiptera: Heteroptera) in waterfalls of Indochina, with descriptions of six new species. Zootaxa 1588: 1 - 29." type="journal article" year="2007">
<taxonomicName id="ADF74D1AFF98FFC6FCFF81870436F91B" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[827,886,1696,1718]" genus="Sites" pageId="21" pageNumber="1270" rank="genus" status="gen. nov.">Sites</taxonomicName>
&amp; Vitheepradit (2007)
</bibRefCitation>
as follows: (1) a pair of prominent scutellar protuberances; (2) the prosternal midline bears an expansive, thin, plate-like carina; (3) the forelegs of both sexes have a one-segmented tarsus apparently fused with the tibia; (4) the forelegs have a single pretarsal claw; and (5) prominent linear series of stout hairs occur on the hemelytra.
</paragraph>
<paragraph id="6A483699FF9BFFC5FF5587E10555FE3E" blockId="22.[145,762,197,403]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FF5587E106FDFF77" box="[145,445,197,219]" italics="true" pageId="22" pageNumber="1267">
Comments:
<taxonomicName id="ADF74D1AFF9BFFC5FEDA87E206FDFF77" authorityName="Sites" authorityYear="2007" box="[286,445,197,218]" class="Insecta" family="Naucoridae" genus="Namtokocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">Namtokocoris</taxonomicName>
</emphasis>
occurs on the vertical rock surfaces of waterfalls in Indochina and is recognized here as the new tribe
<taxonomicName id="ADF74D1AFF9BFFC5FEB986250566FEB5" authorityName="Sites" authorityYear="2022" box="[381,550,258,280]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="tribe" tribe="Namtokocorini">Namtokocorini</taxonomicName>
. The designation of single-genus tribes for
<taxonomicName id="ADF74D1AFF9BFFC5FE5286060572FE9B" authorityName="Sites" authorityYear="2007" box="[406,562,289,310]" class="Insecta" family="Naucoridae" genus="Namtokocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9BFFC5FE5286060572FE9B" box="[406,562,289,310]" italics="true" pageId="22" pageNumber="1267">Namtokocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9BFFC5FDA8860605B8FE9B" authorityName="Montandon" authorityYear="1897" box="[620,760,289,310]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9BFFC5FDA8860605B8FE9B" box="[620,760,289,310]" italics="true" pageId="22" pageNumber="1267">Ctenipocoris</emphasis>
</taxonomicName>
is necessary to create a consistent taxonomic framework for the subfamily with the inclusion of the more diverse tribes
<taxonomicName id="ADF74D1AFF9BFFC5FF1C865A0618FE3E" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[216,344,381,403]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1258" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Heleocorini">Heleocorini</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9BFFC5FE4B865A0551FE3E" authorityName="POPOV" authorityYear="1970" box="[399,529,381,403]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1266" phylum="Arthropoda" rank="tribe" tribe="Laccocorini">Laccocorini</taxonomicName>
.
</paragraph>
<paragraph id="6A483699FF9BFFC5FF558693052FFDAB" blockId="22.[145,761,436,519]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FF558693067BFE64" box="[145,315,436,457]" italics="true" pageId="22" pageNumber="1267">Included taxa:</emphasis>
This tribe is represented by only the genus
<taxonomicName id="ADF74D1AFF9BFFC5FF1D86F50636FE4A" authorityName="Sites" authorityYear="2007" box="[217,374,466,487]" class="Insecta" family="Naucoridae" genus="Namtokocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9BFFC5FF1D86F50636FE4A" box="[217,374,466,487]" italics="true" pageId="22" pageNumber="1267">Namtokocoris</emphasis>
</taxonomicName>
and seven described species (
<bibRefCitation id="0E664B68FF9BFFC5FD0486F5063FFDAB" author="Sites RW &amp; Vitheepradit A" pageId="22" pageNumber="1267" pagination="1 - 29" refId="ref24696" refString="Sites RW, Vitheepradit A. 2007. Namtokocoris, a new genus of Naucoridae (Hemiptera: Heteroptera) in waterfalls of Indochina, with descriptions of six new species. Zootaxa 1588: 1 - 29." type="journal article" year="2007">
<taxonomicName id="ADF74D1AFF9BFFC5FD0486F505B9FE45" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[704,761,466,488]" genus="Sites" pageId="22" pageNumber="1270" rank="genus" status="gen. nov.">Sites</taxonomicName>
&amp; Vitheepradit, 2007
</bibRefCitation>
;
<bibRefCitation id="0E664B68FF9BFFC5FE4F86D60520FDAB" author="Vitheepradit A" box="[395,608,497,519]" pageId="22" pageNumber="1267" pagination="592 - 596" refId="ref25254" refString="Vitheepradit A. 2017. A new species of Namtokocoris Sites (Hemiptera: Naucoridae) from Thailand. Zootaxa 4320: 592 - 596." type="journal article" year="2017">Vitheepradit, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="22ED6512FF98FFC6FC6B87E3026EFF70" box="[943,1326,196,221]" pageId="21" pageNumber="1266" type="nomenclature">
<paragraph id="6A483699FF98FFC6FC6B87E3026EFF70" blockId="21.[943,1326,196,221]" box="[943,1326,196,221]" pageId="21" pageNumber="1266">
TRIBE
<taxonomicName id="ADF74D1AFF98FFC6FC3F87E3026EFF70" ID-CoL="KX3" authority="POPOV, 1970" authorityName="POPOV" authorityYear="1970" box="[1019,1326,196,221]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="tribe" tribe="Laccocorini">
LACCOCORINI
<bibRefCitation id="0E664B68FF98FFC6FB5987E3026EFF70" author="Popov YA" box="[1181,1326,196,221]" pageId="21" pageNumber="1266" pagination="93 - 98" refId="ref23866" refString="Popov YA. 1970. Notes on the classification of the recent Naucoridae (Heteroptera: Nepomorpha). Bulletin of the Polish Academy of Sciences (Biological Sciences) 18: 93 - 98." type="journal article" year="1970">POPOV, 1970</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF98FFC6FCBA87D7021FFEA4" box="[894,1375,240,265]" pageId="21" pageNumber="1266" type="description">
<paragraph id="6A483699FF98FFC6FCBA87D7021FFEA4" blockId="21.[894,1375,240,265]" box="[894,1375,240,265]" pageId="21" pageNumber="1266">
<figureCitation id="F2CC2A1CFF98FFC6FCBA87D70498FEAA" box="[894,984,240,265]" captionStart="Figure 3" captionStartId="8.[145,230,1468,1490]" captionTargetBox="[172,733,195,1428]" captionTargetId="figure-532@8.[172,733,195,1428]" captionTargetPageId="8" captionText="Figure 3. Mouthpart positions: recessed rostrum of Cheirochela thailandana (a); lateral views of Laccocoris spurcus (b) and Australambrysus crenulatus (c). White arrows indicate positions of labrum with respect to the functional anterior margin of the head." figureDoi="http://doi.org/10.5281/zenodo.6994613" httpUri="https://zenodo.org/record/6994613/files/figure.png" pageId="21" pageNumber="1266">FIGS 3B</figureCitation>
,
<figureCitation id="F2CC2A1CFF98FFC6FC2187D7035EFEAA" box="[997,1054,240,265]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="21" pageNumber="1266">5C, D</figureCitation>
,
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<figureCitation id="F2CC2A1CFF98FFC6FBEF87D70379FEA4" box="[1067,1081,240,265]" captionStart="Figure 6" captionStartId="10.[145,228,1329,1351]" captionTargetBox="[216,689,828,1289]" captionTargetId="figure-415@10.[216,689,828,1289]" captionTargetPageId="10" captionText="Figure 6. Laccocoris spurcus showing the transverse sulcus at the base of the labrum, which is characteristic of most Laccocorini." figureDoi="http://doi.org/10.5281/zenodo.6994621" httpUri="https://zenodo.org/record/6994621/files/figure.png" pageId="21" pageNumber="1266">6</figureCitation>
,
<figureCitation id="F2CC2A1CFF98FFC6FB8387D70332FEAA" box="[1095,1138,240,265]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="21" pageNumber="1266">10E</figureCitation>
</geoCoordinate>
,
<figureCitation id="F2CC2A1CFF98FFC6FBBA87D703EAFEAA" box="[1150,1194,240,265]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="21" pageNumber="1266">11E</figureCitation>
,
<figureCitation id="F2CC2A1CFF98FFC6FB7287D703BEFEAA" box="[1206,1278,240,265]" captionStart="Figure 14" captionStartId="19.[163,244,765,787]" captionTargetBox="[163,1442,195,726]" captionTargetId="figure-559@19.[163,1442,195,726]" captionTargetPageId="19" captionText="Figure 14. Revised generic representation of the former genus Heleocoris. Dorsal habitus of Heleocoris vicinus (a), Laccocoris minisculus (b), Heleolaccocoris ovatus (c), Tsingala humeralis (d)." figureDoi="http://doi.org/10.5281/zenodo.6994645" httpUri="https://zenodo.org/record/6994645/files/figure.png" pageId="21" pageNumber="1266">14B, D</figureCitation>
,
<figureCitation id="F2CC2A1CFF98FFC6FACF87D70276FEAA" box="[1291,1334,240,265]" captionStart="Figure 15" captionStartId="20.[145,226,1055,1077]" captionTargetBox="[145,1425,195,1015]" captionTargetId="figure-409@20.[145,1425,195,1015]" captionTargetPageId="20" captionText="Figure 15. Revised generic representation of the former genus Laccocoris. Dorsal habitus of Laccocoris spurcus (a), Heleolaccocoris horvathi (b)." figureDoi="http://doi.org/10.5281/zenodo.6994647" httpUri="https://zenodo.org/record/6994647/files/figure.png" pageId="21" pageNumber="1266">15A</figureCitation>
,
<figureCitation id="F2CC2A1CFF98FFC6FA8787D7021FFEA4" box="[1347,1375,240,265]" captionStart="Figure 20" captionStartId="28.[146,226,1107,1129]" captionTargetBox="[145,1425,195,1067]" captionTargetId="figure-347@28.[145,1425,195,1067]" captionTargetPageId="28" captionText="Figure 20. Tsingala humeralis male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994662" httpUri="https://zenodo.org/record/6994662/files/figure.png" pageId="21" pageNumber="1266">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF98FFC6FCFF8601049FFD21" pageId="21" pageNumber="1266" type="diagnosis">
<paragraph id="6A483699FF98FFC6FCFF8601049FFD21" blockId="21.[827,1443,294,653]" pageId="21" pageNumber="1266">
<emphasis id="5883EA8BFF98FFC6FCFF860104FCFE96" box="[827,956,294,315]" italics="true" pageId="21" pageNumber="1266">Diagnosis:</emphasis>
A unifying feature for all members of
<taxonomicName id="ADF74D1AFF98FFC6FCFF866304FBFEF7" authorityName="POPOV" authorityYear="1970" box="[827,955,324,346]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="tribe" tribe="Laccocorini">Laccocorini</taxonomicName>
except
<taxonomicName id="ADF74D1AFF98FFC6FBCE866203D0FEF7" authorityName="Montandon" authorityYear="1897" box="[1034,1168,325,346]" class="Insecta" family="Naucoridae" genus="Aneurocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FBCE866203D0FEF7" box="[1034,1168,325,346]" italics="true" pageId="21" pageNumber="1266">Aneurocoris</emphasis>
</taxonomicName>
is that the anteroventral seta row on the metafemur gradually extends toward the middle of the ventral surface basally (
<figureCitation id="F2CC2A1CFF98FFC6FAE186A50235FE3A" box="[1317,1397,385,408]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="21" pageNumber="1266">Fig. 5d</figureCitation>
), or if there is an inflection in the direction, it occurs in the distal half of the femur. The spinose setae of the posteroventral row on the mesofemur are not tightly packed and are separated from each other at least by the width of a seta (
<figureCitation id="F2CC2A1CFF98FFC6FBDD853C0321FD9C" box="[1049,1121,539,561]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="21" pageNumber="1266">Fig. 5c</figureCitation>
) and number approximately 40–60. The labrum has a transverse sulcus at the base (
<figureCitation id="F2CC2A1CFF98FFC6FC87857F04C3FDC3" box="[835,899,600,622]" captionStart="Figure 6" captionStartId="10.[145,228,1329,1351]" captionTargetBox="[216,689,828,1289]" captionTargetId="figure-415@10.[216,689,828,1289]" captionTargetPageId="10" captionText="Figure 6. Laccocoris spurcus showing the transverse sulcus at the base of the labrum, which is characteristic of most Laccocorini." figureDoi="http://doi.org/10.5281/zenodo.6994621" httpUri="https://zenodo.org/record/6994621/files/figure.png" pageId="21" pageNumber="1266">Fig. 6</figureCitation>
), although the sulcus appears to have been lost in
<taxonomicName id="ADF74D1AFF98FFC6FC9C855F0498FD20" authorityName="Montandon" authorityYear="1897" box="[856,984,632,653]" class="Insecta" family="Naucoridae" genus="Temnocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FC9C855F0498FD20" box="[856,984,632,653]" italics="true" pageId="21" pageNumber="1266">Temnocoris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF98FFC6FCFF85910489FA2B" pageId="21" pageNumber="1266" type="discussion">
<paragraph id="6A483699FF98FFC6FCFF85910330FB5E" blockId="21.[827,1444,694,1267]" pageId="21" pageNumber="1266">
<emphasis id="5883EA8BFF98FFC6FCFF85910488FD66" box="[827,968,694,715]" italics="true" pageId="21" pageNumber="1266">Comments:</emphasis>
Because
<taxonomicName id="ADF74D1AFF98FFC6FB9485910209FD66" baseAuthorityName="Stal" baseAuthorityYear="1856" box="[1104,1353,694,715]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="spurcus">
<emphasis id="5883EA8BFF98FFC6FB9485910209FD66" box="[1104,1353,694,715]" italics="true" pageId="21" pageNumber="1266">Laccocoris spurcus</emphasis>
</taxonomicName>
is the
<typeStatus id="B54C883BFF98FFC6FCFF85F20430FD47" box="[827,880,725,746]" pageId="21" pageNumber="1266">type</typeStatus>
species for the subfamily, this clade is here recognized as tribe
<taxonomicName id="ADF74D1AFF98FFC6FBD985D403E3FCA4" authorityName="POPOV" authorityYear="1970" box="[1053,1187,755,777]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="tribe" tribe="Laccocorini">Laccocorini</taxonomicName>
. This tribe is endemic to Africa,
<collectingCountry id="12E07609FF98FFC6FC6B8435037EFC85" box="[943,1086,786,808]" name="Madagascar" pageId="21" pageNumber="1266">Madagascar</collectingCountry>
and the Middle East. Generic level relationships among
<taxonomicName id="ADF74D1AFF98FFC6FBBB8416024EFCEB" authorityName="Montandon" authorityYear="1897" box="[1151,1294,817,838]" class="Insecta" family="Naucoridae" genus="Aneurocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FBBB8416024EFCEB" box="[1151,1294,817,838]" italics="true" pageId="21" pageNumber="1266">Aneurocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF98FFC6FADB841602DCFCEB" authorityName="Stal" authorityYear="1856" box="[1311,1436,817,838]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FADB841602DCFCEB" box="[1311,1436,817,838]" italics="true" pageId="21" pageNumber="1266">Laccocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF98FFC6FCFF847704FEFCC8" authorityName="Montandon" authorityYear="1897" box="[827,958,848,869]" class="Insecta" family="Naucoridae" genus="Temnocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FCFF847704FEFCC8" box="[827,958,848,869]" italics="true" pageId="21" pageNumber="1266">Temnocoris</emphasis>
</taxonomicName>
and the new genus
<taxonomicName id="ADF74D1AFF98FFC6FB6C8468024EFCC9" authorityName="Sites" authorityYear="2022" box="[1192,1294,847,868]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FB6C8468024EFCC9" box="[1192,1294,847,868]" italics="true" pageId="21" pageNumber="1266">Tsingala</emphasis>
</taxonomicName>
are strongly supported (bs = 100). The clade containing
<taxonomicName id="ADF74D1AFF98FFC6FAD9844902E3FC2E" authorityName="Montandon" authorityYear="1897" box="[1309,1443,878,899]" class="Insecta" family="Naucoridae" genus="Aneurocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FAD9844902E3FC2E" box="[1309,1443,878,899]" italics="true" pageId="21" pageNumber="1266">Aneurocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF98FFC6FCAB84AA04A9FC0F" authorityName="Stal" authorityYear="1856" box="[879,1001,909,930]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FCAB84AA04A9FC0F" box="[879,1001,909,930]" italics="true" pageId="21" pageNumber="1266">Laccocoris</emphasis>
</taxonomicName>
from mainland Africa have a unique fracture (
<figureCitation id="F2CC2A1CFF98FFC6FC76848C0354FC6D" box="[946,1044,939,961]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="21" pageNumber="1266">Fig. 10e</figureCitation>
) across the hemelytra from the distal ends of the embolia through the tip of the claval commissure (
<bibRefCitation id="0E664B68FF98FFC6FC1984CF03A9FC50" author="Mbogho AY &amp; Sites RW" box="[989,1257,1000,1022]" pageId="21" pageNumber="1266" pagination="513 - 542" refId="ref22951" refString="Mbogho AY, Sites RW. 2013. Naucoridae Leach, 1815 (Hemiptera: Heteroptera) of Tanzania. African Invertebrates 54: 513 - 542." type="journal article" year="2013">
Mbogho &amp;
<taxonomicName id="ADF74D1AFF98FFC6FBA784CF03A9FC50" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1123,1257,1000,1022]" genus="Sites" pageId="21" pageNumber="1270" rank="genus" status="gen. nov.">Sites, 2013</taxonomicName>
</bibRefCitation>
). This fracture is present in both sexes of
<taxonomicName id="ADF74D1AFF98FFC6FB5C8320036CFB91" authority="Montandon, 1897 b" authorityName="Montandon" authorityYear="1897" class="Insecta" family="Naucoridae" genus="Aneurocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="5883EA8BFF98FFC6FB5C832002E3FBB1" box="[1176,1443,1031,1052]" italics="true" pageId="21" pageNumber="1266">Aneurocoris insolitus</emphasis>
<bibRefCitation id="0E664B68FF98FFC6FCFF8301036CFB91" author="Montandon AL" box="[827,1068,1062,1084]" pageId="21" pageNumber="1266" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
</taxonomicName>
and mostly only females of
<taxonomicName id="ADF74D1AFF98FFC6FCFF836204F3FBF7" authorityName="Stal" authorityYear="1856" box="[827,947,1093,1114]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FCFF836204F3FBF7" box="[827,947,1093,1114]" italics="true" pageId="21" pageNumber="1266">Laccocoris</emphasis>
</taxonomicName>
, although it is also present in some males. The Middle Eastern
<taxonomicName id="ADF74D1AFF98FFC6FBFC83440431FB3A" authority="Walker, 1870" authorityName="Walker" authorityYear="1870" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="minisculus">
<emphasis id="5883EA8BFF98FFC6FBFC83440200FBD5" box="[1080,1344,1123,1144]" italics="true" pageId="21" pageNumber="1266">Heleocoris minisculus</emphasis>
<bibRefCitation id="0E664B68FF98FFC6FA8D83440431FB3A" author="Walker F" pageId="21" pageNumber="1266" pagination="2378 - 2381" refId="ref25280" refString="Walker F. 1870. A list of the Hemiptera collected by J. K. Lord, Esq., in Egypt, along the African shore of the Red Sea and in Arabia; with descriptions of the species new to science. The Zoologist: a Popular Miscellany of Natural History, Series 2 5: 2378 - 2381." type="journal article" year="1870">Walker, 1870</bibRefCitation>
</taxonomicName>
also bears this fracture and is here transferred to
<taxonomicName id="ADF74D1AFF98FFC6FCFF838604F2FB1B" authorityName="Stal" authorityYear="1856" box="[827,946,1185,1206]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FCFF838604F2FB1B" box="[827,946,1185,1206]" italics="true" pageId="21" pageNumber="1266">Laccocoris</emphasis>
</taxonomicName>
as
<taxonomicName id="ADF74D1AFF98FFC6FC1D83860261FB18" authority="(Walker, 1870)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Walker" baseAuthorityYear="1870" box="[985,1313,1184,1206]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="minisculus" status="comb. nov.">
<emphasis id="5883EA8BFF98FFC6FC1D83860334FB18" box="[985,1140,1184,1206]" italics="true" pageId="21" pageNumber="1266">L. minisculus</emphasis>
(
<bibRefCitation id="0E664B68FF98FFC6FB4783870259FB18" author="Walker F" box="[1155,1305,1184,1206]" pageId="21" pageNumber="1266" pagination="2378 - 2381" refId="ref25280" refString="Walker F. 1870. A list of the Hemiptera collected by J. K. Lord, Esq., in Egypt, along the African shore of the Red Sea and in Arabia; with descriptions of the species new to science. The Zoologist: a Popular Miscellany of Natural History, Series 2 5: 2378 - 2381." type="journal article" year="1870">Walker, 1870</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF98FFC6FAEC838702DFFB1B" box="[1320,1439,1184,1206]" pageId="21" pageNumber="1266" rank="species">comb. nov.</taxonomicNameLabel>
, but the fracture is absent in the clade of
<emphasis id="5883EA8BFF98FFC6FAC8839802CDFB79" box="[1292,1421,1215,1236]" italics="true" pageId="21" pageNumber="1266">Temnocoris</emphasis>
+
<taxonomicName id="ADF74D1AFF98FFC6FCFF83FA04DFFB5F" authorityName="Sites" authorityYear="2022" box="[827,927,1245,1266]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FCFF83FA04DFFB5F" box="[827,927,1245,1266]" italics="true" pageId="21" pageNumber="1266">Tsingala</emphasis>
</taxonomicName>
from
<collectingCountry id="12E07609FF98FFC6FC2583FA032CFB5E" box="[993,1132,1245,1267]" name="Madagascar" pageId="21" pageNumber="1266">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="6A483699FF98FFC6FCFF82330489FA2B" blockId="21.[827,1443,1300,1414]" pageId="21" pageNumber="1266">
<emphasis id="5883EA8BFF98FFC6FCFF82330328FA84" box="[827,1128,1300,1322]" italics="true" pageId="21" pageNumber="1266">
Included taxa:
<taxonomicName id="ADF74D1AFF98FFC6FC2682330328FA84" authorityName="Montandon" authorityYear="1897" box="[994,1128,1300,1321]" class="Insecta" family="Naucoridae" genus="Aneurocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">Aneurocoris</taxonomicName>
</emphasis>
(known by only two species in Lake Tanganyika) and
<taxonomicName id="ADF74D1AFF98FFC6FB938214038CFAE5" authorityName="Stal" authorityYear="1856" box="[1111,1228,1331,1352]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FB938214038CFAE5" box="[1111,1228,1331,1352]" italics="true" pageId="21" pageNumber="1266">Laccocoris</emphasis>
</taxonomicName>
are from mainland Africa, and
<taxonomicName id="ADF74D1AFF98FFC6FC0182750309FACA" authorityName="Montandon" authorityYear="1897" box="[965,1097,1362,1383]" class="Insecta" family="Naucoridae" genus="Temnocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FC0182750309FACA" box="[965,1097,1362,1383]" italics="true" pageId="21" pageNumber="1266">Temnocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF98FFC6FB41827603ADFACB" authorityName="Sites" authorityYear="2022" box="[1157,1261,1361,1382]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FB41827603ADFACB" box="[1157,1261,1361,1382]" italics="true" pageId="21" pageNumber="1266">Tsingala</emphasis>
</taxonomicName>
are endemic to
<collectingCountry id="12E07609FF98FFC6FCFF82570486FA2B" box="[827,966,1392,1414]" name="Madagascar" pageId="21" pageNumber="1266">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

261
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<treatment id="E25E878FFF99FFC6FC6281220554F8D9" ID-DOI="http://doi.org/10.5281/zenodo.6994721" ID-GBIF-Taxon="199315262" ID-Zenodo-Dep="6994721" LSID="urn:lsid:plazi:treatment:E25E878FFF99FFC6FC6281220554F8D9" httpUri="http://treatment.plazi.org/id/E25E878FFF99FFC6FC6281220554F8D9" lastPageId="21" lastPageNumber="1266" pageId="20" pageNumber="1265">
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<paragraph id="6A483699FF99FFC7FC6281220254F9B0" blockId="20.[934,1300,1541,1566]" box="[934,1300,1541,1566]" pageId="20" pageNumber="1265">
<heading id="310081F5FF99FFC7FC6281220254F9B0" box="[934,1300,1541,1566]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="1265" reason="3">
TRIBE
<taxonomicName id="ADF74D1AFF99FFC7FC36812203CFF9B6" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1010,1167,1541,1565]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Heleocorini">HELEOCORINI</taxonomicName>
<emphasis id="5883EA8BFF99FFC7FB52812E0254F9B0" bold="true" box="[1174,1300,1541,1565]" pageId="20" pageNumber="1265">
<taxonomicNameLabel id="43B057F0FF99FFC7FB52812E0254F9B0" box="[1174,1300,1541,1565]" pageId="20" pageNumber="1265" rank="tribe">TRIB. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF99FFC6FC5A81160554F8D9" lastPageId="21" lastPageNumber="1266" pageId="20" pageNumber="1265" type="description">
<paragraph id="6A483699FF99FFC7FC5A8116025BF9E7" blockId="20.[926,1307,1585,1610]" box="[926,1307,1585,1610]" pageId="20" pageNumber="1265">
<figureCitation id="F2CC2A1CFF99FFC7FC5A811604B8F9E4" box="[926,1016,1585,1610]" captionStart="Figure 4" captionStartId="9.[163,249,1379,1401]" captionTargetBox="[163,781,195,1339]" captionTargetId="figure-575@9.[163,781,195,1339]" captionTargetPageId="9" captionText="Figure 4. Foretibiae and foretarsi of Laccocorinae: Ctenipocoris asiaticus (a), Heleolaccocoris ovatus (b), Namtokocoris dalanta (c)." figureDoi="http://doi.org/10.5281/zenodo.6994615" httpUri="https://zenodo.org/record/6994615/files/figure.png" pageId="20" pageNumber="1265">FIGS 4B</figureCitation>
,
<figureCitation id="F2CC2A1CFF99FFC7FBC18116037DF9E4" box="[1029,1085,1585,1610]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="20" pageNumber="1265">5A, B</figureCitation>
,
<figureCitation id="F2CC2A1CFF99FFC7FB8E81160326F9E7" box="[1098,1126,1585,1610]" captionStart="Figure 12" captionStartId="17.[163,243,870,892]" captionTargetBox="[163,1443,195,830]" captionTargetId="figure-534@17.[163,1443,195,830]" captionTargetPageId="17" captionText="Figure 12. Mesotibia and mesotarsus of male and female Heleolaccocoris ovatus showing a sexually dimorphic ventral tomentose pad of hairs." figureDoi="http://doi.org/10.5281/zenodo.6994637" httpUri="https://zenodo.org/record/6994637/files/figure.png" pageId="20" pageNumber="1265">12</figureCitation>
,
<figureCitation id="F2CC2A1CFF99FFC7FBB0811603FBF9E4" box="[1140,1211,1585,1610]" captionStart="Figure 14" captionStartId="19.[163,244,765,787]" captionTargetBox="[163,1442,195,726]" captionTargetId="figure-559@19.[163,1442,195,726]" captionTargetPageId="19" captionText="Figure 14. Revised generic representation of the former genus Heleocoris. Dorsal habitus of Heleocoris vicinus (a), Laccocoris minisculus (b), Heleolaccocoris ovatus (c), Tsingala humeralis (d)." figureDoi="http://doi.org/10.5281/zenodo.6994645" httpUri="https://zenodo.org/record/6994645/files/figure.png" pageId="20" pageNumber="1265">14A, C</figureCitation>
,
<figureCitation id="F2CC2A1CFF99FFC7FB03811603B2F9E4" box="[1223,1266,1585,1610]" captionStart="Figure 15" captionStartId="20.[145,226,1055,1077]" captionTargetBox="[145,1425,195,1015]" captionTargetId="figure-409@20.[145,1425,195,1015]" captionTargetPageId="20" captionText="Figure 15. Revised generic representation of the former genus Laccocoris. Dorsal habitus of Laccocoris spurcus (a), Heleolaccocoris horvathi (b)." figureDoi="http://doi.org/10.5281/zenodo.6994647" httpUri="https://zenodo.org/record/6994647/files/figure.png" pageId="20" pageNumber="1265">15B</figureCitation>
,
<figureCitation id="F2CC2A1CFF99FFC7FB3B8116025BF9E7" box="[1279,1307,1585,1610]" captionStart="Figure 21" captionStartId="29.[164,242,1876,1898]" captionTargetBox="[163,1443,921,1837]" captionTargetId="figure-364@29.[163,1443,921,1837]" captionTargetPageId="29" captionText="Figure 21. Heleolaccocoris horvathi male: dorsal habitus (a), genital capsule with terga IX and X removed (b)." figureDoi="http://doi.org/10.5281/zenodo.6994664" httpUri="https://zenodo.org/record/6994664/files/figure.png" pageId="20" pageNumber="1265">21</figureCitation>
</paragraph>
<paragraph id="6A483699FF99FFC7FCED814F02C1F936" blockId="20.[809,1426,1639,1691]" pageId="20" pageNumber="1265">
<emphasis id="5883EA8BFF99FFC7FCED814F0337F9D0" box="[809,1143,1639,1661]" italics="true" pageId="20" pageNumber="1265">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF99FFC7FCB381A202C1F936" box="[887,1409,1669,1691]" pageId="20" pageNumber="1265">8CBB9ABB-857E-4540-8213-1B1648FDF868</uuid>
</paragraph>
<paragraph id="6A483699FF99FFC6FCED81E206A1FE7D" blockId="20.[809,1425,1732,1907]" lastBlockId="21.[163,779,197,464]" lastPageId="21" lastPageNumber="1266" pageId="20" pageNumber="1265">
<emphasis id="5883EA8BFF99FFC7FCED81E204EFF977" box="[809,943,1733,1754]" italics="true" pageId="20" pageNumber="1265">Diagnosis:</emphasis>
This group includes some taxa with extreme habitat specializations, thus a simple characterization of the tribe is not possible. A feature characteristic for all species except
<taxonomicName id="ADF74D1AFF99FFC7FB1F80070322F8F9" authority="Sites &amp; Zettel, 2011" authorityName="Sites &amp; Zettel" authorityYear="2011" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="5883EA8BFF99FFC7FB1F80070433F8F9" italics="true" pageId="20" pageNumber="1265">Pogonocaudina indica</emphasis>
<bibRefCitation id="0E664B68FF99FFC7FCBE80180322F8F9" author="Sites RW &amp; Zettel H &amp; Arunachalam M" box="[890,1122,1855,1877]" pageId="20" pageNumber="1265" pagination="1 - 17" refId="ref24813" refString="Sites RW, Zettel H, Arunachalam M. 2011. Waterfallinhabiting Naucoridae (Hemiptera: Heteroptera) of southern India and Sri Lanka: Pogonocaudina Sites and Zettel, n. gen., and a review of Diaphorocoris with descriptions of two new species. Zootaxa 2760: 1 - 17." type="journal article" year="2011">Sites &amp; Zettel, 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF99FFC7FB59801804B2F8DE" authority="La Rivers, 1969" authorityName="La Rivers" authorityYear="1969" class="Insecta" family="Naucoridae" genus="Decarloa" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="5883EA8BFF99FFC7FB59801802D1F8F9" box="[1181,1425,1855,1876]" italics="true" pageId="20" pageNumber="1265">Decarloa darlingtoni</emphasis>
<bibRefCitation id="0E664B68FF99FFC7FCED807904B2F8DE" author="La Rivers I" box="[809,1010,1885,1907]" pageId="20" pageNumber="1265" pagination="1 - 12" refId="ref22401" refString="La Rivers I. 1969. New naucorid taxa (Hemiptera). Biological Society of Nevada Occasional Papers 20: 1 - 12." type="journal article" year="1969">La Rivers, 1969</bibRefCitation>
</taxonomicName>
is that the spinose setae of the posteroventral row on the mesofemur are tightly spaced such that adjacent setae or their sockets are in contact or nearly so (
<figureCitation id="F2CC2A1CFF98FFC6FDCC8625051EFEB5" box="[520,606,258,280]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="21" pageNumber="1266">Fig. 5a</figureCitation>
) and number approximately 60–90. In addition, in all except the clade containing
<taxonomicName id="ADF74D1AFF98FFC6FE028667057AFEF8" authorityName="La Rivers" authorityYear="1969" box="[454,570,320,341]" class="Insecta" family="Naucoridae" genus="Decarloa" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FE028667057AFEF8" box="[454,570,320,341]" italics="true" pageId="21" pageNumber="1266">Decarloa</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF98FFC6FD958667044BFEF8" authorityName="Montandon" authorityYear="1897" box="[593,779,320,341]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FD958667044BFEF8" box="[593,779,320,341]" italics="true" pageId="21" pageNumber="1266">Diaphorocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF98FFC6FF1A86780629FED9" authorityName="La Rivers" authorityYear="1974" box="[222,361,351,372]" class="Insecta" family="Naucoridae" genus="Interocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FF1A86780629FED9" box="[222,361,351,372]" italics="true" pageId="21" pageNumber="1266">Interocoris</emphasis>
</taxonomicName>
, the anteroventral seta row on the metafemur is noticeably arcuate or with a posteromesal direction change in the basal third as it extends basally (
<figureCitation id="F2CC2A1CFF98FFC6FE46869D0691FE7D" box="[386,465,442,464]" captionStart="Figure 5" captionStartId="10.[145,223,735,757]" captionTargetBox="[145,1426,195,695]" captionTargetId="figure-384@10.[145,1426,195,695]" captionTargetPageId="10" captionText="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." figureDoi="http://doi.org/10.5281/zenodo.6994617" httpUri="https://zenodo.org/record/6994617/files/figure.png" pageId="21" pageNumber="1266">Fig. 5b</figureCitation>
).
</paragraph>
<paragraph id="6A483699FF98FFC6FF6786DE0630FBDE" blockId="21.[163,780,505,1723]" pageId="21" pageNumber="1266">
<emphasis id="5883EA8BFF98FFC6FF6786DE0668FDA3" box="[163,296,505,526]" italics="true" pageId="21" pageNumber="1266">Comments:</emphasis>
Because the clades have a high degree of fidelity with respect to geography and the
<typeStatus id="B54C883BFF98FFC6FD12853F0449FD80" box="[726,777,536,557]" pageId="21" pageNumber="1266">type</typeStatus>
species is from
<collectingCountry id="12E07609FF98FFC6FEA585110687FDE1" box="[353,455,566,588]" name="India" pageId="21" pageNumber="1266">Mumbai</collectingCountry>
, the
<taxonomicName id="ADF74D1AFF98FFC6FDC1851105C1FDE6" authorityName="Stal" authorityYear="1876" box="[517,641,566,587]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FDC1851105C1FDE6" box="[517,641,566,587]" italics="true" pageId="21" pageNumber="1266">Heleocoris</emphasis>
</taxonomicName>
clade from
<collectingCountry id="12E07609FF98FFC6FF67857207A1FDC6" box="[163,225,597,619]" name="India" pageId="21" pageNumber="1266">India</collectingCountry>
remains unchanged as the genus
<taxonomicName id="ADF74D1AFF98FFC6FDA585720599FDC7" authorityName="Stal" authorityYear="1876" box="[609,729,597,618]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FDA585720599FDC7" box="[609,729,597,618]" italics="true" pageId="21" pageNumber="1266">Heleocoris</emphasis>
</taxonomicName>
(see Comments section above) and the containing group is the new tribe
<taxonomicName id="ADF74D1AFF98FFC6FEA685B506ABFD05" authorityName="Sites" authorityYear="2022" box="[354,491,658,680]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="tribe" tribe="Heleocorini">Heleocorini</taxonomicName>
. Representing this clade in the analyses are
<taxonomicName id="ADF74D1AFF98FFC6FE4C859607A9FD4B" authority="Montandon, 1897 b" authorityName="Montandon" authorityYear="1897" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="5883EA8BFF98FFC6FE4C85960538FD6B" box="[392,632,689,710]" italics="true" pageId="21" pageNumber="1266">Heleocoris elongatus</emphasis>
<bibRefCitation id="0E664B68FF98FFC6FDBB859607A9FD4B" author="Montandon AL" pageId="21" pageNumber="1266" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF98FFC6FF3C85F70596FD48" authority="Montandon, 1908" authorityName="Montandon" authorityYear="1908" box="[248,726,720,742]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="5883EA8BFF98FFC6FF3C85F706BFFD48" box="[248,511,720,741]" italics="true" pageId="21" pageNumber="1266">Heleocoris rotundatus</emphasis>
<bibRefCitation id="0E664B68FF98FFC6FDC385F70596FD48" author="Montandon AL" box="[519,726,720,742]" pageId="21" pageNumber="1266" pagination="299 - 304" refId="ref23198" refString="Montandon AL. 1908. Nouvelle especes d'Hemipteres aquatiques. Annales Historico-Naturales Musei Nationalis Hungarici 6: 299 - 304." type="journal article" year="1908">Montandon, 1908</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF98FFC6FF6785C90518FCA9" authority="Montandon, 1910 b" authorityName="Montandon" authorityYear="1910" box="[163,600,750,772]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="5883EA8BFF98FFC6FF6785C90637FCA9" box="[163,375,750,772]" italics="true" pageId="21" pageNumber="1266">Heleocoris vicinus</emphasis>
<bibRefCitation id="0E664B68FF98FFC6FEBA85C90518FCA9" author="Montandon AL" box="[382,600,750,772]" pageId="21" pageNumber="1266" pagination="652 - 658" refId="ref23277" refString="Montandon AL. 1910 b. Especes nouvelles ou peu connues d'Hydrococrises de l'Inde. Bulletin de la Societe des Sciences de Bucarest, Roumanie 19: 652 - 658." type="journal article" year="1910">Montandon, 1910b</bibRefCitation>
</taxonomicName>
. Species in the clades containing
<taxonomicName id="ADF74D1AFF98FFC6FEBA842A06B9FC8F" authorityName="Stal" authorityYear="1876" box="[382,505,781,802]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FEBA842A06B9FC8F" box="[382,505,781,802]" italics="true" pageId="21" pageNumber="1266">Heleocoris</emphasis>
</taxonomicName>
from
<collectingCountry id="12E07609FF98FFC6FD87842A0595FC8E" box="[579,725,781,803]" name="Madagascar" pageId="21" pageNumber="1266">Madagascar</collectingCountry>
and Southeast Asia are here transferred to the newly erected genera
<taxonomicName id="ADF74D1AFF98FFC6FE96846D06FAFCF2" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[338,442,842,863]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF98FFC6FE96846D06FAFCF2" box="[338,442,842,863]" italics="true" pageId="21" pageNumber="1266">Tsingala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF98FFC6FE7B846C0562FCCD" box="[447,546,843,864]" pageId="21" pageNumber="1266" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="ADF74D1AFF98FFC6FD9C846D044AFCF2" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[600,778,842,863]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF98FFC6FD9C846D044AFCF2" box="[600,778,842,863]" italics="true" pageId="21" pageNumber="1266">Heleolaccocoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF98FFC6FF67844D0650FCD2" box="[163,272,874,895]" pageId="21" pageNumber="1266" rank="genus">gen. nov.</taxonomicNameLabel>
, respectively (described below).
<taxonomicName id="ADF74D1AFF98FFC6FD5B844E044AFCD3" authorityName="Sites" authorityYear="2022" box="[671,778,873,894]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FD5B844E044AFCD3" box="[671,778,873,894]" italics="true" pageId="21" pageNumber="1266">Tsingala</emphasis>
</taxonomicName>
was recovered with its sister genus
<taxonomicName id="ADF74D1AFF98FFC6FD8184AF0589FC30" authorityName="Montandon" authorityYear="1897" box="[581,713,904,925]" class="Insecta" family="Naucoridae" genus="Temnocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FD8184AF0589FC30" box="[581,713,904,925]" italics="true" pageId="21" pageNumber="1266">Temnocoris</emphasis>
</taxonomicName>
, both endemic to
<collectingCountry id="12E07609FF98FFC6FEE9848106FEFC11" box="[301,446,934,956]" name="Madagascar" pageId="21" pageNumber="1266">Madagascar</collectingCountry>
(red-coloured clades in
<figureCitation id="F2CC2A1CFF98FFC6FD1A848107FFFC77" captionStart="Figure 13" captionStartId="18.[145,223,1057,1079]" captionTargetBox="[149,1425,195,1017]" captionTargetId="figure-432@18.[145,1425,195,1017]" captionTargetPageId="18" captionText="Figure 13. Detail of Most Probable Phylogeny dendogram of Laccocorinae.The red-coloured clade includes the Madagascar genera Temnocoris and Tsingala gen. nov.(formerly Heleocoris). The blue-coloured clade represents Heleolaccocoris gen.nov., which comprises species of the former genera Heleocoris and Laccocoris. All species of Tsingala gen. nov. and Heleolaccocoris gen. nov. are new combinations. Geographic associations for major clades within and among tribes are indicated. The red box in the inset shows the part of Figure 9 depicted in this figure." figureDoi="http://doi.org/10.5281/zenodo.6994641" httpUri="https://zenodo.org/record/6994641/files/figure.png" pageId="21" pageNumber="1266">Fig. 13</figureCitation>
), in the tribe
<taxonomicName id="ADF74D1AFF98FFC6FEAA84E206BCFC76" authorityName="POPOV" authorityYear="1970" box="[366,508,965,987]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="tribe" tribe="Laccocorini">Laccocorini</taxonomicName>
. Species of
<taxonomicName id="ADF74D1AFF98FFC6FD4884E1044BFC76" authorityName="Stal" authorityYear="1856" box="[652,779,966,987]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FD4884E1044BFC76" box="[652,779,966,987]" italics="true" pageId="21" pageNumber="1266">Laccocoris</emphasis>
</taxonomicName>
from Borneo, Sumatra, the
<collectingCountry id="12E07609FF98FFC6FE1384C3051CFC57" box="[471,604,996,1018]" name="Philippines" pageId="21" pageNumber="1266">Philippines</collectingCountry>
and
<taxonomicName id="ADF74D1AFF98FFC6FD5584C306ACFBBA" authority="Montandon, 1910" authorityName="Montandon" authorityYear="1910" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="dissidens">
<emphasis id="5883EA8BFF98FFC6FD5584C30652FBBA" italics="true" pageId="21" pageNumber="1266">Laccocoris dissidens</emphasis>
Montandon, 1910
</taxonomicName>
, described from
<collectingCountry id="12E07609FF98FFC6FD7683240446FBB5" box="[690,774,1027,1048]" name="Myanmar" pageId="21" pageNumber="1266">Burma</collectingCountry>
, are transferred to
<taxonomicName id="ADF74D1AFF98FFC6FE498306050AFB9B" authorityName="Sites" authorityYear="2022" box="[397,586,1057,1078]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FE498306050AFB9B" box="[397,586,1057,1078]" italics="true" pageId="21" pageNumber="1266">Heleolaccocoris</emphasis>
</taxonomicName>
along with the Southeast Asian species of
<taxonomicName id="ADF74D1AFF98FFC6FE278367051BFBF8" authorityName="Stal" authorityYear="1876" box="[483,603,1088,1109]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FE278367051BFBF8" box="[483,603,1088,1109]" italics="true" pageId="21" pageNumber="1266">Heleocoris</emphasis>
</taxonomicName>
(blue-coloured clades in
<figureCitation id="F2CC2A1CFF98FFC6FED583790620FBDE" box="[273,352,1118,1140]" captionStart="Figure 13" captionStartId="18.[145,223,1057,1079]" captionTargetBox="[149,1425,195,1017]" captionTargetId="figure-432@18.[145,1425,195,1017]" captionTargetPageId="18" captionText="Figure 13. Detail of Most Probable Phylogeny dendogram of Laccocorinae.The red-coloured clade includes the Madagascar genera Temnocoris and Tsingala gen. nov.(formerly Heleocoris). The blue-coloured clade represents Heleolaccocoris gen.nov., which comprises species of the former genera Heleocoris and Laccocoris. All species of Tsingala gen. nov. and Heleolaccocoris gen. nov. are new combinations. Geographic associations for major clades within and among tribes are indicated. The red box in the inset shows the part of Figure 9 depicted in this figure." figureDoi="http://doi.org/10.5281/zenodo.6994641" httpUri="https://zenodo.org/record/6994641/files/figure.png" pageId="21" pageNumber="1266">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="6A483699FF98FFC6FF7F835A07ADF916" blockId="21.[163,780,505,1723]" pageId="21" pageNumber="1266">
Among the Indian waterfall-inhabiting species,
<taxonomicName id="ADF74D1AFF98FFC6FF6783BB06F1FB1C" authorityName="Sites &amp; Zettel" authorityYear="2011" box="[163,433,1180,1201]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="5883EA8BFF98FFC6FF6783BB06F1FB1C" box="[163,433,1180,1201]" italics="true" pageId="21" pageNumber="1266">Pogonocaudina indica</emphasis>
</taxonomicName>
was recovered as a derived member of a clade of
<taxonomicName id="ADF74D1AFF98FFC6FE7D839D0528FB62" authorityName="Montandon" authorityYear="1897" box="[441,616,1210,1231]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FE7D839D0528FB62" box="[441,616,1210,1231]" italics="true" pageId="21" pageNumber="1266">Diaphorocoris</emphasis>
</taxonomicName>
. Although it exhibits morphological features supporting its distinction as a genus-level taxon, including singlesegmented fore tarsi in both sexes, its evolutionary relationship with species of
<taxonomicName id="ADF74D1AFF98FFC6FE188212053CFAE7" authorityName="Montandon" authorityYear="1897" box="[476,636,1333,1354]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FE188212053CFAE7" box="[476,636,1333,1354]" italics="true" pageId="21" pageNumber="1266">Diaphorocoris</emphasis>
</taxonomicName>
require that the genus be synonymized. As such, the monotypic
<taxonomicName id="ADF74D1AFF98FFC6FF6782550614FA2A" authorityName="Sites &amp; Zettel" authorityYear="2011" box="[163,340,1394,1415]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FF6782550614FA2A" box="[163,340,1394,1415]" italics="true" pageId="21" pageNumber="1266">Pogonocaudina</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName id="ADF74D1AFF98FFC6FDA38255044AFA2A" authorityName="Montandon" authorityYear="1897" box="[615,778,1394,1415]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FDA38255044AFA2A" box="[615,778,1394,1415]" italics="true" pageId="21" pageNumber="1266">Diaphorocoris</emphasis>
</taxonomicName>
with the species becoming
<taxonomicName id="ADF74D1AFF98FFC6FE3C82B606CFFA68" authority="(Sites &amp; Zettel, 2011)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Sites &amp; Zettel" baseAuthorityYear="2011" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="indicus" status="comb. nov.">
<emphasis id="5883EA8BFF98FFC6FE3C82B6044AFA0B" box="[504,778,1425,1446]" italics="true" pageId="21" pageNumber="1266">Diaphorocoris indicus</emphasis>
(
<bibRefCitation id="0E664B68FF98FFC6FF6F829706C8FA68" author="Sites RW &amp; Zettel H &amp; Arunachalam M" box="[171,392,1456,1478]" pageId="21" pageNumber="1266" pagination="1 - 17" refId="ref24813" refString="Sites RW, Zettel H, Arunachalam M. 2011. Waterfallinhabiting Naucoridae (Hemiptera: Heteroptera) of southern India and Sri Lanka: Pogonocaudina Sites and Zettel, n. gen., and a review of Diaphorocoris with descriptions of two new species. Zootaxa 2760: 1 - 17." type="journal article" year="2011">Sites &amp; Zettel, 2011</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF98FFC6FE5182970547FA68" box="[405,519,1456,1478]" pageId="21" pageNumber="1266" rank="species">comb. nov.</taxonomicNameLabel>
Nested in this group of Indian taxa are
<taxonomicName id="ADF74D1AFF98FFC6FE9882E8044AFA4E" authority="(Usinger, 1935)" baseAuthorityName="Usinger" baseAuthorityYear="1935" box="[348,778,1486,1508]" class="Insecta" family="Naucoridae" genus="Interocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="5883EA8BFF98FFC6FE9882E80515FA49" box="[348,597,1487,1508]" italics="true" pageId="21" pageNumber="1266">Interocoris mexicanus</emphasis>
(
<bibRefCitation id="0E664B68FF98FFC6FDA182E90442FA49" author="Usinger RL" box="[613,770,1486,1508]" pageId="21" pageNumber="1266" pagination="133 - 136" refId="ref25136" refString="Usinger RL. 1935. A second American species of the naucorid subfamily Laccocorinae (Hemiptera). Revista de Entomologia 5: 133 - 136." type="journal article" year="1935">Usinger, 1935</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF98FFC6FF1182CA0682F9AF" authorityName="La Rivers" authorityYear="1969" box="[213,450,1517,1538]" class="Insecta" family="Naucoridae" genus="Decarloa" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="5883EA8BFF98FFC6FF1182CA0682F9AF" box="[213,450,1517,1538]" italics="true" pageId="21" pageNumber="1266">Decarloa darlingtoni</emphasis>
</taxonomicName>
, both from northern tropical America (Mesoamerica and Hispaniola, respectively). These two monotypic genera have long been enigmatic because there is little representation of this subfamily in the New World. That they are most closely related to taxa from
<collectingCountry id="12E07609FF98FFC6FEFA81A1063DF931" box="[318,381,1670,1692]" name="India" pageId="21" pageNumber="1266">India</collectingCountry>
raises new questions about their origin.
</paragraph>
<paragraph id="6A483699FF98FFC6FF6781C30554F8D9" blockId="21.[163,779,1764,1908]" pageId="21" pageNumber="1266">
<emphasis id="5883EA8BFF98FFC6FF6781C3068BF954" box="[163,459,1764,1785]" italics="true" pageId="21" pageNumber="1266">
Included taxa:
<taxonomicName id="ADF74D1AFF98FFC6FE9181C3068BF954" authorityName="Stal" authorityYear="1876" box="[341,459,1764,1785]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">Heleocoris</taxonomicName>
</emphasis>
and
<taxonomicName id="ADF74D1AFF98FFC6FDC281C305E8F954" authorityName="Montandon" authorityYear="1897" box="[518,680,1764,1785]" class="Insecta" family="Naucoridae" genus="Diaphorocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FDC281C305E8F954" box="[518,680,1764,1785]" italics="true" pageId="21" pageNumber="1266">Diaphorocoris</emphasis>
</taxonomicName>
occur in
<collectingCountry id="12E07609FF98FFC6FF67802507A5F8B5" box="[163,229,1794,1816]" name="India" pageId="21" pageNumber="1266">India</collectingCountry>
;
<taxonomicName id="ADF74D1AFF98FFC6FF36802506E9F8BA" authorityName="Sites" authorityYear="2022" box="[242,425,1794,1815]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FF36802506E9F8BA" box="[242,425,1794,1815]" italics="true" pageId="21" pageNumber="1266">Heleolaccocoris</emphasis>
</taxonomicName>
occurs from the easternmost Indian states and Indochina through the Sunda Islands to the
<collectingCountry id="12E07609FF98FFC6FE9480670699F8FB" box="[336,473,1856,1878]" name="Philippines" pageId="21" pageNumber="1266">Philippines</collectingCountry>
.
<taxonomicName id="ADF74D1AFF98FFC6FE228067050FF8F8" authorityName="La Rivers" authorityYear="1969" box="[486,591,1856,1877]" class="Insecta" family="Naucoridae" genus="Decarloa" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FE228067050FF8F8" box="[486,591,1856,1877]" italics="true" pageId="21" pageNumber="1266">Decarloa</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF98FFC6FD488067044BF8F8" authorityName="La Rivers" authorityYear="1974" box="[652,779,1856,1877]" class="Insecta" family="Naucoridae" genus="Interocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="1266" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF98FFC6FD488067044BF8F8" box="[652,779,1856,1877]" italics="true" pageId="21" pageNumber="1266">Interocoris</emphasis>
</taxonomicName>
are American monotypic genera.
</paragraph>
</subSubSection>
</treatment>
</document>

341
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SUBFAMILY
<taxonomicName id="ADF74D1AFF9AFFC4FE47828705F3FA14" ID-CoL="JYC" authority="LEACH, 1815" authorityName="LEACH" authorityYear="1815" box="[387,691,1440,1465]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">
NAUCORINAE
<bibRefCitation id="0E664B68FF9AFFC4FDE5828705F3FA14" author="Leach WE" box="[545,691,1440,1465]" pageId="23" pageNumber="1268" pagination="57 - 192" refId="ref22574" refString="Leach WE. 1815. Entomology. In: Brewster D, ed. The Edinburgh encyclopedia, Vol. 9. Edinburgh: William Blackwood, 57 - 192." type="book chapter" year="1815">LEACH, 1815</bibRefCitation>
</taxonomicName>
</paragraph>
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<figureCitation id="F2CC2A1CFF9AFFC4FEF782EB06DDFA49" box="[307,413,1484,1509]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="23" pageNumber="1268">FIGS 10H</figureCitation>
,
<figureCitation id="F2CC2A1CFF9AFFC4FE6E82EB0697FA49" box="[426,471,1484,1509]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="23" pageNumber="1268">11H</figureCitation>
,
<figureCitation id="F2CC2A1CFF9AFFC4FE2082EB0540FA48" box="[484,512,1484,1509]" captionStart="Figure 17" captionStartId="23.[827,914,1816,1838]" captionTargetBox="[929,1337,1400,1776]" captionTargetId="figure-735@23.[929,1337,1400,1776]" captionTargetPageId="23" captionText="Figure 17. Lateral view of head and prothorax of Philippinocoris usingeri showing anteromedian tooth of prosternum (white arrow)." figureDoi="http://doi.org/10.5281/zenodo.6994654" httpUri="https://zenodo.org/record/6994654/files/figure.png" pageId="23" pageNumber="1268">17</figureCitation>
,
<figureCitation id="F2CC2A1CFF9AFFC4FDC982EB0569FA48" box="[525,553,1484,1509]" captionStart="Figure 19" captionStartId="25.[164,245,774,796]" captionTargetBox="[163,1444,195,734]" captionTargetId="figure-573@25.[163,1444,195,734]" captionTargetPageId="25" captionText="Figure 19. Revised generic representation of the former genus Naucoris. Dorsal habitus of Naucoris maculatus (a), Australonaucoris congrex (b), Afronaucoris parvulus (c), Indonaucoris minutus sp. nov. (d)." figureDoi="http://doi.org/10.5281/zenodo.6994658" httpUri="https://zenodo.org/record/6994658/files/figure.png" pageId="23" pageNumber="1268">19</figureCitation>
,
<figureCitation id="F2CC2A1CFF9AFFC4FDF382EB053DFA48" box="[567,637,1484,1509]" captionStart-0="Figure 22" captionStart-1="Figure 23" captionStart-2="Figure 24" captionStart-3="Figure 25" captionStartId-0="31.[164,242,1851,1873]" captionStartId-1="33.[163,241,986,1008]" captionStartId-2="35.[163,243,1352,1374]" captionStartId-3="36.[144,224,1075,1097]" captionTargetBox-0="[163,1444,895,1813]" captionTargetBox-1="[163,1444,195,946]" captionTargetBox-2="[243,1363,195,1313]" captionTargetBox-3="[224,1344,195,1035]" captionTargetId-0="figure-311@31.[163,1444,895,1813]" captionTargetId-1="figure-408@33.[163,1444,195,946]" captionTargetId-2="figure-243@35.[243,1363,195,1313]" captionTargetId-3="figure-24@36.[224,1344,195,1035]" captionTargetPageId-0="31" captionTargetPageId-1="33" captionTargetPageId-2="35" captionTargetPageId-3="36" captionText-0="Figure 22. Afronaucoris obscuratus male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c), distal part of aedeagus (d). Sizes are not proportional among figures." captionText-1="Figure 23. Australonaucoris congrex male: dorsal habitus (a), genital capsule with parameres and terga IX and X removed (b), left paramere (c), right paramere (d). Sizes are not proportional among figures." captionText-2="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." captionText-3="Figure 25. Tamhini stream, the presumed type locality of Indonaucoris minutus gen. nov., sp. nov., in December 2019. Photo courtesy of Shruti Paripatyadar." figureDoi-0="http://doi.org/10.5281/zenodo.6994668" figureDoi-1="http://doi.org/10.5281/zenodo.6994670" figureDoi-2="http://doi.org/10.5281/zenodo.6994674" figureDoi-3="http://doi.org/10.5281/zenodo.6994676" httpUri-0="https://zenodo.org/record/6994668/files/figure.png" httpUri-1="https://zenodo.org/record/6994670/files/figure.png" httpUri-2="https://zenodo.org/record/6994674/files/figure.png" httpUri-3="https://zenodo.org/record/6994676/files/figure.png" pageId="23" pageNumber="1268">22–25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9AFFC4FF60812405A0F9B5" box="[164,736,1538,1560]" pageId="23" pageNumber="1268" type="type_taxon">
<paragraph id="6A483699FF9AFFC4FF60812405A0F9B5" blockId="23.[164,736,1538,1560]" box="[164,736,1538,1560]" pageId="23" pageNumber="1268">
<emphasis id="5883EA8BFF9AFFC4FF6081240674F9B5" box="[164,308,1539,1560]" italics="true" pageId="23" pageNumber="1268">
<typeStatus id="B54C883BFF9AFFC4FF6081240799F9B5" box="[164,217,1539,1560]" pageId="23" pageNumber="1268">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF9AFFC4FE8481240599F9BA" ID-CoL="745Y9" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[320,729,1538,1560]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="5883EA8BFF9AFFC4FE8481240565F9BA" box="[320,549,1538,1560]" italics="true" pageId="23" pageNumber="1268">Naucoris maculatus</emphasis>
<bibRefCitation id="0E664B68FF9AFFC4FDE881250599F9BA" author="Fabricius JC" box="[556,729,1538,1560]" pageId="23" pageNumber="1268" pagination="511 - 572" refId="ref21860" refString="Fabricius JC. 1798. Ryngota. Os rostro: vagina articulata. Supplementum Entomologiae Systematicae 12: 511 - 572." type="journal article" year="1798">Fabricius, 1798</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9AFFCBFF60816606ECFB1F" lastPageId="24" lastPageNumber="1269" pageId="23" pageNumber="1268" type="discussion">
<paragraph id="6A483699FF9AFFC4FF6081660300FD0A" blockId="23.[164,781,1601,1899]" lastBlockId="23.[828,1445,198,680]" pageId="23" pageNumber="1268">
<emphasis id="5883EA8BFF9AFFC4FF60816606D1F9FB" box="[164,401,1601,1622]" italics="true" pageId="23" pageNumber="1268">Taxonomic history:</emphasis>
The nominate subfamily was recognized as the division Naucoraria by
<bibRefCitation id="0E664B68FF9AFFC4FD488178044DF9D8" author="Stal C" box="[652,781,1631,1653]" pageId="23" pageNumber="1268" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">Stål (1876)</bibRefCitation>
to include
<taxonomicName id="ADF74D1AFF9AFFC4FEF1815906E4F93E" authorityName="Stal" authorityYear="1861" box="[309,420,1662,1683]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FEF1815906E4F93E" box="[309,420,1662,1683]" italics="true" pageId="23" pageNumber="1268">Ilyocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FE7D81580507F939" authorityName="Signoret" authorityYear="1861" box="[441,583,1663,1684]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FE7D81580507F939" box="[441,583,1663,1684]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FD9881580590F939" authorityName="Geoffroy" authorityYear="1762" box="[604,720,1663,1684]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FD9881580590F939" box="[604,720,1663,1684]" italics="true" pageId="23" pageNumber="1268">Naucoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9AFFC4FF6081BA0650F91F" baseAuthorityName="La Rivers" baseAuthorityYear="1956" box="[164,272,1693,1714]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FF6081BA0650F91F" box="[164,272,1693,1714]" italics="true" pageId="23" pageNumber="1268">Pelocoris</emphasis>
</taxonomicName>
. The group was referenced as subfamily
<taxonomicName id="ADF74D1AFF9AFFC4FF60819C066FF97C" authorityName="LEACH" authorityYear="1815" box="[164,303,1723,1745]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
by
<bibRefCitation id="0E664B68FF9AFFC4FE9A819C057CF97C" author="Montandon AL" box="[350,572,1723,1745]" pageId="23" pageNumber="1268" pagination="414 - 422" refId="ref23172" refString="Montandon AL. 1900. Hemipteres exotiques nouveaux ou peu connus des collections du Musee National Hongrois. Termeszetrajzi Fuzetek 23: 414 - 422." type="journal article" year="1900">Montandon (1900)</bibRefCitation>
when describing
<taxonomicName id="ADF74D1AFF9AFFC4FF6081FC06DEF95D" box="[164,414,1755,1776]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="species" species="parviceps">
<emphasis id="5883EA8BFF9AFFC4FF6081FC06DEF95D" box="[164,414,1755,1776]" italics="true" pageId="23" pageNumber="1268">Macrocoris parviceps</emphasis>
</taxonomicName>
. Since then, additional genera have been described and placed in
<taxonomicName id="ADF74D1AFF9AFFC4FD8F81DE0597F8A2" authorityName="LEACH" authorityYear="1815" box="[587,727,1785,1807]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
, but the subfamily was not based on any synapomorphies and the monotypy has been considered doubtful (e.g.
<bibRefCitation id="0E664B68FF9AFFC4FF60807206E3F8C7" author="Sites RW &amp; Mbogho AY" box="[164,419,1877,1899]" pageId="23" pageNumber="1268" pagination="1 - 39" refId="ref24588" refString="Sites RW, Mbogho AY. 2012. Revision of the African genus Neomacrocoris (Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Zootaxa 3555: 1 - 39." type="journal article" year="2012">
<taxonomicName id="ADF74D1AFF9AFFC4FF608072079EF8C6" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[164,222,1877,1899]" genus="Sites" pageId="23" pageNumber="1270" rank="genus" status="gen. nov.">Sites</taxonomicName>
&amp; Mbogho, 2012
</bibRefCitation>
). The genus
<taxonomicName id="ADF74D1AFF9AFFC4FDFD80720598F8C7" authorityName="De Carlo" authorityYear="1963" box="[569,728,1877,1898]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FDFD80720598F8C7" box="[569,728,1877,1898]" italics="true" pageId="23" pageNumber="1268">Carvalhoiella</emphasis>
</taxonomicName>
was described in
<taxonomicName id="ADF74D1AFF9AFFC4FC1787E1026BFF76" authority="(De Carlo, 1963)" authorityName="Stal" authorityYear="1862" baseAuthorityName="De Carlo" baseAuthorityYear="1963" box="[979,1323,198,220]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">
Ambrysinae (
<bibRefCitation id="0E664B68FF9AFFC4FBB687E10261FF71" author="De Carlo JA" box="[1138,1313,198,220]" pageId="23" pageNumber="1268" pagination="9 - 11" refId="ref21695" refString="De Carlo JA. 1963. Un nuevo genero y una nueva especie de la subfamilia Ambrysinae (Hemiptera: Naucoridae). Revista de la Sociedad Entomologica Argentina 24: 9 - 11." type="journal article" year="1963">De Carlo, 1963</bibRefCitation>
)
</taxonomicName>
, then was transferred to
<taxonomicName id="ADF74D1AFF9AFFC4FC2087C203BEFF57" authority="(Nieser, 1975)" authorityName="POPOV" authorityYear="1970" baseAuthorityName="Nieser" baseAuthorityYear="1975" box="[996,1278,229,251]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">
Naucorini (
<bibRefCitation id="0E664B68FF9AFFC4FBA387C203B5FF56" author="Nieser N" box="[1127,1269,229,251]" pageId="23" pageNumber="1268" pagination="1 - 303" refId="ref23396" refString="Nieser N. 1975. The water bugs (Heteroptera: Nepomorpha) of the Guyana Region. Studies on the Fauna of Suriname and other Guyanas 16: 1 - 303." type="journal article" year="1975">Nieser, 1975</bibRefCitation>
)
</taxonomicName>
, but was later transferred back to
<taxonomicName id="ADF74D1AFF9AFFC4FBE0862402E0FEB5" authority="(Nieser et al., 1999)" authorityName="Stal" authorityYear="1862" baseAuthorityName="Nieser" baseAuthorityYear="1999" box="[1060,1440,259,281]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">
Ambrysinae (
<bibRefCitation id="0E664B68FF9AFFC4FB05862402D5FEB5" author="Nieser N &amp; Pelli A &amp; Melo AL" box="[1217,1429,259,281]" pageId="23" pageNumber="1268" pagination="157 - 163" refId="ref23491" refString="Nieser N, Pelli A, Melo AL. 1999. Two new Ambrysinae (Heteroptera: Naucoridae) from Minas Gerais, Brazil. Acta Societatis Zoologicae Bohemoslovacae 63: 157 - 163." type="journal article" year="1999">
Nieser
<emphasis id="5883EA8BFF9AFFC4FAD18623020FFEB5" box="[1301,1359,259,281]" italics="true" pageId="23" pageNumber="1268">et al.</emphasis>
, 1999
</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="ADF74D1AFF9AFFC4FCF88605048FFE9A" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[828,975,290,312]" genus="Zettel" pageId="23" pageNumber="1285" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF9AFFC4FCF88605048FFE9A" author="Zettel H" box="[828,975,290,312]" pageId="23" pageNumber="1268" pagination="1085 - 1095" refId="ref25492" refString="Zettel H. 2001. Die indische Zwergschwimmwanze Nanonaucoris gen. n., eine neue Gattung der Naucorini (Heteroptera: Naucoridae) aus Sudindien. Linzer biologische Beitrage 33: 1085 - 1095." type="journal article" year="2001">Zettel (2001)</bibRefCitation>
</taxonomicName>
suggested the tribe
<taxonomicName id="ADF74D1AFF9AFFC4FB788605026AFE95" box="[1212,1322,290,312]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="tribe" tribe="Ilyocorini">Ilyocorini</taxonomicName>
to contain
<taxonomicName id="ADF74D1AFF9AFFC4FCF8866604E3FEFB" authorityName="Stal" authorityYear="1861" box="[828,931,321,342]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FCF8866604E3FEFB" box="[828,931,321,342]" italics="true" pageId="23" pageNumber="1268">Ilyocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FC708666035FFEFB" baseAuthorityName="La Rivers" baseAuthorityYear="1956" box="[948,1055,321,342]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FC708666035FFEFB" box="[948,1055,321,342]" italics="true" pageId="23" pageNumber="1268">Pelocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9AFFC4FB9B866603ABFEFB" box="[1119,1259,321,342]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FB9B866603ABFEFB" box="[1119,1259,321,342]" italics="true" pageId="23" pageNumber="1268">Placomerus</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9AFFC4FAEF866602E3FEFA" authorityName="POPOV" authorityYear="1970" box="[1323,1443,321,343]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini</taxonomicName>
to contain
<taxonomicName id="ADF74D1AFF9AFFC4FC7986780314FED9" authorityName="Usinger" authorityYear="1937" box="[957,1108,351,372]" class="Insecta" family="Naucoridae" genus="Asthenocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FC7986780314FED9" box="[957,1108,351,372]" italics="true" pageId="23" pageNumber="1268">Asthenocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FBA0864703A7FED8" authorityName="Signoret" authorityYear="1861" box="[1124,1255,352,373]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FBA0864703A7FED8" box="[1124,1255,352,373]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FB33864702DEFED8" authorityName="Zettel" authorityYear="2001" box="[1271,1438,352,373]" class="Insecta" family="Naucoridae" genus="Nanonaucoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FB33864702DEFED8" box="[1271,1438,352,373]" italics="true" pageId="23" pageNumber="1268">Nanonaucoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FCF8865804E5FE39" authorityName="Geoffroy" authorityYear="1762" box="[828,933,383,404]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FCF8865804E5FE39" box="[828,933,383,404]" italics="true" pageId="23" pageNumber="1268">Naucoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FC7086580320FE39" authorityName="Montandon" authorityYear="1913" box="[948,1120,383,404]" class="Insecta" family="Naucoridae" genus="Neomacrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FC7086580320FE39" box="[948,1120,383,404]" italics="true" pageId="23" pageNumber="1268">Neomacrocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FBAA86590261FE3E" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="1987" box="[1134,1313,382,403]" class="Insecta" family="Naucoridae" genus="Philippinocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FBAA86590261FE3E" box="[1134,1313,382,403]" italics="true" pageId="23" pageNumber="1268">Philippinocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FAEB865902E4FE3E" authorityName="La Rivers" authorityYear="1969" box="[1327,1444,382,403]" class="Insecta" family="Naucoridae" genus="Stalocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FAEB865902E4FE3E" box="[1327,1444,382,403]" italics="true" pageId="23" pageNumber="1268">Stalocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9AFFC4FCB486BA03E5FE1F" authority="Distant, 1904" authorityName="Distant" authorityYear="1904" box="[880,1189,413,435]" class="Insecta" family="Naucoridae" genus="Thurselinus" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FCB486BA04BDFE1F" box="[880,1021,413,434]" italics="true" pageId="23" pageNumber="1268">Thurselinus</emphasis>
<bibRefCitation id="0E664B68FF9AFFC4FBC186BA03E5FE1F" author="Distant WL" box="[1029,1189,413,435]" pageId="23" pageNumber="1268" pagination="258 - 259" refId="ref21779" refString="Distant WL. 1904. New Rhynchota-Cryptocerata. The Entomologist 37: 258 - 259." type="journal article" year="1904">Distant, 1904</bibRefCitation>
</taxonomicName>
. In the description of
<taxonomicName id="ADF74D1AFF9AFFC4FCF8869C0328FE7D" authority=", Zettel (2007)" authorityName=", Zettel" authorityYear="2007" box="[828,1128,443,465]" class="Insecta" family="Naucoridae" genus="Halmaheria" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FCF8869C0487FE7D" box="[828,967,443,464]" italics="true" pageId="23" pageNumber="1268">Halmaheria</emphasis>
,
<bibRefCitation id="0E664B68FF9AFFC4FC11869C0328FE7D" author="Zettel H" box="[981,1128,443,465]" pageId="23" pageNumber="1268" pagination="89 - 94" refId="ref25528" refString="Zettel H. 2007. Halmaheria skalei gen. n. et sp. n., eine neue Schwimmwanze (Insecta: Heteroptera: Naucoridae) von der Insel Halmahera, Indonesien. Annalen des Naturhistorischen Museums in Wien 108 B: 89 - 94." type="journal article" year="2007">Zettel (2007)</bibRefCitation>
</taxonomicName>
noted a series of attributes suggesting a close relationship of this new genus with both Sagocorini (
<taxonomicName id="ADF74D1AFF9AFFC4FBF986DE03AEFDA2" authorityName="MONTANDON" authorityYear="1897" box="[1085,1262,505,527]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1260" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
) and
<taxonomicName id="ADF74D1AFF9AFFC4FAF486DE0495FD80" authority="(Naucorinae)" authorityName="POPOV" authorityYear="1970" baseAuthorityName="Naucorinae" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini (Naucorinae)</taxonomicName>
. Therefore, he transferred the Philippine genera of Sagocorini (
<taxonomicName id="ADF74D1AFF9AFFC4FB818511039DFDE6" authorityName="Usinger" authorityYear="1937" box="[1093,1245,566,587]" class="Insecta" family="Naucoridae" genus="Asthenocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FB818511039DFDE6" box="[1093,1245,566,587]" italics="true" pageId="23" pageNumber="1268">Asthenocoris</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9AFFC4FB29851102E5FDE6" authorityName="D. Polhemus &amp; J. Polhemus" authorityYear="1987" box="[1261,1445,566,587]" class="Insecta" family="Naucoridae" genus="Philippinocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FB29851102E5FDE6" box="[1261,1445,566,587]" italics="true" pageId="23" pageNumber="1268">Philippinocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9AFFC4FCB4857204A5FDC7" authorityName="La Rivers" authorityYear="1969" box="[880,997,597,618]" class="Insecta" family="Naucoridae" genus="Stalocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FCB4857204A5FDC7" box="[880,997,597,618]" italics="true" pageId="23" pageNumber="1268">Stalocoris</emphasis>
</taxonomicName>
) to
<taxonomicName id="ADF74D1AFF9AFFC4FBD0857203CCFDC6" authorityName="POPOV" authorityYear="1970" box="[1044,1164,597,619]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini</taxonomicName>
. Recently,
<taxonomicName id="ADF74D1AFF9AFFC4FACC8572022FFDC7" baseAuthorityName="La Rivers" baseAuthorityYear="1956" box="[1288,1391,597,618]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FACC8572022FFDC7" box="[1288,1391,597,618]" italics="true" pageId="23" pageNumber="1268">Pelocoris</emphasis>
</taxonomicName>
was transferred from
<taxonomicName id="ADF74D1AFF9AFFC4FC3B855403C4FD24" authorityName="LEACH" authorityYear="1815" box="[1023,1156,627,649]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
to
<taxonomicName id="ADF74D1AFF9AFFC4FB608554037CFD0A" authority="(ReynosoVelasco &amp; Sites, 2021)" authorityName="Stal" authorityYear="1862" baseAuthorityName="ReynosoVelasco &amp; Sites" baseAuthorityYear="2021" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">
Ambrysinae (ReynosoVelasco &amp;
<bibRefCitation id="0E664B68FF9AFFC4FC7785B50370FD0A" author="Sites RW" box="[947,1072,658,680]" pageId="23" pageNumber="1268" pagination="72 - 94" refId="ref24528" refString="Sites RW. 2021. A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species. Zootaxa 4958: 72 - 94." type="journal article" year="2021">Sites, 2021</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="6A483699FF9AFFCBFCF885F606ECFB1F" blockId="23.[828,1445,721,1233]" lastBlockId="24.[145,760,1150,1203]" lastPageId="24" lastPageNumber="1269" pageId="23" pageNumber="1268">
<emphasis id="5883EA8BFF9AFFC4FCF885F60358FD4B" box="[828,1048,721,742]" italics="true" pageId="23" pageNumber="1268">Revised taxonomy:</emphasis>
In my Most Probable Phylogeny dendrogram (
<figureCitation id="F2CC2A1CFF9AFFC4FC1B85C80362FCA9" box="[991,1058,751,773]" captionStart="Figure 9" captionStartId="12.[145,223,1678,1700]" captionTargetBox="[308,1266,197,1636]" captionTargetId="figure-82@12.[302,1268,195,1638]" captionTargetPageId="12" captionText="Figure 9. Most Probable Phylogeny dendogram based on Bayesian inference with substitutions from maximum likelihood for Laccocorinae (except Ctenipocorini trib. nov. and Idiocarini trib. nov.). Closer views of Laccocorinae and Naucorinae individually can be seen in Figures 13 and 18, respectively." figureDoi="http://doi.org/10.5281/zenodo.6994629" httpUri="https://zenodo.org/record/6994629/files/figure.png" pageId="23" pageNumber="1268">Fig. 9</figureCitation>
), I substituted the ML topology of Clade O, the predominantly New Guinean clade, into the BI dendrogram to represent what I consider to be the most plausible, supported evolutionary relationships within the subfamily. The revised subfamily
<taxonomicName id="ADF74D1AFF9AFFC4FC0984AE0325FC33" authorityName="LEACH" authorityYear="1815" box="[973,1125,905,927]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
presented in the Most Probable Phylogeny has four major clades (
<figureCitation id="F2CC2A1CFF9AFFC4FA87848002D4FC11" box="[1347,1428,935,957]" captionStart="Figure 18" captionStartId="24.[145,223,1007,1029]" captionTargetBox="[147,1427,197,966]" captionTargetId="figure-376@24.[145,1429,195,967]" captionTargetPageId="24" captionText="Figure 18. Detail of Most Probable Phylogeny dendogram of Naucorinae, based primarily on Bayesian inference, but with maximum likelihood topology substituted for Idiocarini trib. nov. All species of Afronaucoris gen. nov. and Australonaucoris gen. nov. are new combinations. The red box in the inset shows the part of Figure 9 depicted in this figure." figureDoi="http://doi.org/10.5281/zenodo.6994656" httpUri="https://zenodo.org/record/6994656/files/figure.png" pageId="23" pageNumber="1268">Fig. 18</figureCitation>
), each of which represents a tribe and includes a species of
<taxonomicName id="ADF74D1AFF9AFFC4FC7C84C20364FC57" authorityName="Geoffroy" authorityYear="1762" box="[952,1060,997,1018]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FC7C84C20364FC57" box="[952,1060,997,1018]" italics="true" pageId="23" pageNumber="1268">Naucoris</emphasis>
</taxonomicName>
(
<figureCitation id="F2CC2A1CFF9AFFC4FBF184C203C7FC57" box="[1077,1159,997,1019]" captionStart="Figure 19" captionStartId="25.[164,245,774,796]" captionTargetBox="[163,1444,195,734]" captionTargetId="figure-573@25.[163,1444,195,734]" captionTargetPageId="25" captionText="Figure 19. Revised generic representation of the former genus Naucoris. Dorsal habitus of Naucoris maculatus (a), Australonaucoris congrex (b), Afronaucoris parvulus (c), Indonaucoris minutus sp. nov. (d)." figureDoi="http://doi.org/10.5281/zenodo.6994658" httpUri="https://zenodo.org/record/6994658/files/figure.png" pageId="23" pageNumber="1268">Fig. 19</figureCitation>
) at its root with strong support. Because the genus
<taxonomicName id="ADF74D1AFF9AFFC4FB5083230240FBB4" authorityName="Geoffroy" authorityYear="1762" box="[1172,1280,1028,1049]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FB5083230240FBB4" box="[1172,1280,1028,1049]" italics="true" pageId="23" pageNumber="1268">Naucoris</emphasis>
</taxonomicName>
, as currently conceived, is clearly polyphyletic, new genera will be necessary for species of
<taxonomicName id="ADF74D1AFF9AFFC4FB95836603F9FBFB" authorityName="Geoffroy" authorityYear="1762" box="[1105,1209,1089,1110]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FB95836603F9FBFB" box="[1105,1209,1089,1110]" italics="true" pageId="23" pageNumber="1268">Naucoris</emphasis>
</taxonomicName>
in all but one clade. Because these species were previously considered congeners, morphological distinctions supporting their recognition as multiple genera are not evident in some cases. Nonetheless, the molecular evidence presented here that
<taxonomicName id="ADF74D1AFF95FFCBFE45835806AAFB39" authorityName="Geoffroy" authorityYear="1762" box="[385,490,1151,1172]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFE45835806AAFB39" box="[385,490,1151,1172]" italics="true" pageId="24" pageNumber="1269">Naucoris</emphasis>
</taxonomicName>
as currently conceived is polyphyletic is strong.
</paragraph>
</subSubSection>
<caption id="3E886611FF9AFFC4FCFF803F0310F8C4" ID-DOI="http://doi.org/10.5281/zenodo.6994654" ID-Zenodo-Dep="6994654" httpUri="https://zenodo.org/record/6994654/files/figure.png" pageId="23" pageNumber="1268" startId="23.[827,914,1816,1838]" targetBox="[929,1337,1400,1776]" targetPageId="23">
<paragraph id="6A483699FF9AFFC4FCFF803F0310F8C4" blockId="23.[825,1441,1816,1897]" pageId="23" pageNumber="1268">
<emphasis id="5883EA8BFF9AFFC4FCFF803F0482F880" bold="true" box="[827,962,1816,1838]" pageId="23" pageNumber="1268">Figure 17.</emphasis>
Lateral view of head and prothorax of
<taxonomicName id="ADF74D1AFF9AFFC4FCFD80120302F8E6" baseAuthorityName="La Rivers" baseAuthorityYear="1970" box="[825,1090,1845,1867]" class="Insecta" family="Naucoridae" genus="Philippinocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="species" species="usingeri">
<emphasis id="5883EA8BFF9AFFC4FCFD80120302F8E6" box="[825,1090,1845,1867]" italics="true" pageId="23" pageNumber="1268">Philippinocoris usingeri</emphasis>
</taxonomicName>
showing anteromedian tooth of prosternum (white arrow).
</paragraph>
</caption>
<caption id="3E886611FF95FFCBFF5584C803A4FB92" ID-DOI="http://doi.org/10.5281/zenodo.6994656" ID-Zenodo-Dep="6994656" httpUri="https://zenodo.org/record/6994656/files/figure.png" pageId="24" pageNumber="1269" startId="24.[145,223,1007,1029]" targetBox="[147,1427,197,966]" targetPageId="24">
<paragraph id="6A483699FF95FFCBFF5584C803A4FB92" blockId="24.[145,1425,1007,1087]" pageId="24" pageNumber="1269">
<emphasis id="5883EA8BFF95FFCBFF5584C80646FBA9" bold="true" box="[145,262,1007,1029]" pageId="24" pageNumber="1269">Figure 18.</emphasis>
Detail of Most Probable Phylogeny dendogram of
<taxonomicName id="ADF74D1AFF95FFCBFCD284C804D1FBA8" authorityName="LEACH" authorityYear="1815" box="[790,913,1007,1029]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
, based primarily on Bayesian inference, but with maximum likelihood topology substituted for
<taxonomicName id="ADF74D1AFF95FFCBFDA8832B0592FB8F" authority="All" authorityName="Sites" authorityYear="2022" box="[620,722,1036,1058]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Idiocarini">Idiocarini</taxonomicName>
<taxonomicNameLabel id="43B057F0FF95FFCBFD13832B0473FB8F" box="[727,819,1036,1058]" pageId="24" pageNumber="1269" rank="tribe">trib. nov.</taxonomicNameLabel>
All species of
<taxonomicName id="ADF74D1AFF95FFCBFC00832B030BFB8F" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[964,1099,1036,1058]" class="Insecta" family="Naucoridae" genus="Afronaucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF95FFCBFC00832B030BFB8F" box="[964,1099,1036,1058]" italics="true" pageId="24" pageNumber="1269">Afronaucoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF95FFCBFB95832B03EBFB8F" box="[1105,1195,1036,1058]" pageId="24" pageNumber="1269" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="ADF74D1AFF95FFCBFB1A832B02D1FB8F" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1246,1425,1036,1058]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF95FFCBFB1A832B02D1FB8F" box="[1246,1425,1036,1058]" italics="true" pageId="24" pageNumber="1269">Australonaucoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF95FFCBFF55830E07ABFB92" box="[145,235,1065,1087]" pageId="24" pageNumber="1269" rank="genus">gen. nov.</taxonomicNameLabel>
are new combinations. The red box in the inset shows the part of Figure 9 depicted in this figure.
</paragraph>
</caption>
<subSubSection id="22ED6512FF95FFCBFF5583FB0548F9EF" pageId="24" pageNumber="1269" type="diagnosis">
<paragraph id="6A483699FF95FFCBFF5583FB0548F9EF" blockId="24.[145,761,1244,1603]" pageId="24" pageNumber="1269">
<emphasis id="5883EA8BFF95FFCBFF5583FB064EFB5C" box="[145,270,1244,1265]" italics="true" pageId="24" pageNumber="1269">Diagnosis:</emphasis>
With the recent transfer of
<taxonomicName id="ADF74D1AFF95FFCBFDAA83FB0598FB5C" baseAuthorityName="La Rivers" baseAuthorityYear="1956" box="[622,728,1244,1265]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFDAA83FB0598FB5C" box="[622,728,1244,1265]" italics="true" pageId="24" pageNumber="1269">Pelocoris</emphasis>
</taxonomicName>
to
<taxonomicName id="ADF74D1AFF95FFCBFF5583DD065FFABD" authorityName="Stal" authorityYear="1862" box="[145,287,1274,1296]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
,
<taxonomicName id="ADF74D1AFF95FFCBFEEE83DD06CCFAA2" authorityName="Stal" authorityYear="1861" box="[298,396,1274,1295]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFEEE83DD06CCFAA2" box="[298,396,1274,1295]" italics="true" pageId="24" pageNumber="1269">Ilyocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF95FFCBFE0383DD050DFAA2" box="[455,589,1274,1295]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFE0383DD050DFAA2" box="[455,589,1274,1295]" italics="true" pageId="24" pageNumber="1269">Placomerus</emphasis>
</taxonomicName>
to
<taxonomicName id="ADF74D1AFF95FFCBFDB683DD05B4FABD" authorityName="Sites" authorityYear="2022" box="[626,756,1274,1296]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1261" phylum="Arthropoda" rank="subFamily" status="stat. nov." subFamily="Ilyocorinae">Ilyocorinae</taxonomicName>
, and
<taxonomicName id="ADF74D1AFF95FFCBFF01823D0606FA82" authorityName="Signoret" authorityYear="1861" box="[197,326,1306,1327]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFF01823D0606FA82" box="[197,326,1306,1327]" italics="true" pageId="24" pageNumber="1269">Macrocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF95FFCBFE46823D056EFA82" authorityName="Montandon" authorityYear="1913" box="[386,558,1306,1327]" class="Insecta" family="Naucoridae" genus="Neomacrocoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFE46823D056EFA82" box="[386,558,1306,1327]" italics="true" pageId="24" pageNumber="1269">Neomacrocoris</emphasis>
</taxonomicName>
to
<taxonomicName id="ADF74D1AFF95FFCBFD90823E05B4FA82" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[596,756,1305,1327]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1260" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Macrocorinae">Macrocorinae</taxonomicName>
, a clear synapomorphy among genera in
<taxonomicName id="ADF74D1AFF95FFCBFDB4821F05B8FAE3" authorityName="LEACH" authorityYear="1815" box="[624,760,1336,1358]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
is now evident. The male parameres are large, crisscrossing and dramatically asymmetrical in all species. More specifically, the left paramere overlaps the right and is broader and slightly shorter than the right. The right paramere tends to have a sulcus or elongated fossa on its right side to accommodate the left side of the aedeagus shaft. Both parameres wrap around and embrace the aedeagus (
<figureCitation id="F2CC2A1CFF95FFCBFE58810A06B8F9EE" box="[412,504,1581,1603]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="24" pageNumber="1269">Fig. 11h</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF95FFCBFF558143048BFB1E" pageId="24" pageNumber="1269" type="discussion">
<paragraph id="6A483699FF95FFCBFF558143048BFB1E" blockId="24.[145,760,1636,1903]" lastBlockId="24.[809,1424,1150,1203]" pageId="24" pageNumber="1269">
<emphasis id="5883EA8BFF95FFCBFF558143065DF9D4" box="[145,285,1636,1657]" italics="true" pageId="24" pageNumber="1269">Comments:</emphasis>
Because each of these tribe groups has one or more species of
<taxonomicName id="ADF74D1AFF95FFCBFE2A81A4051EF935" authorityName="Geoffroy" authorityYear="1762" box="[494,606,1667,1688]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFE2A81A4051EF935" box="[494,606,1667,1688]" italics="true" pageId="24" pageNumber="1269">Naucoris</emphasis>
</taxonomicName>
at the root, this suggests that an especially vagile ancestral
<taxonomicName id="ADF74D1AFF95FFCBFF5581E707BCF978" authorityName="Geoffroy" authorityYear="1762" box="[145,252,1728,1749]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="1269" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF95FFCBFF5581E707BCF978" box="[145,252,1728,1749]" italics="true" pageId="24" pageNumber="1269">Naucoris</emphasis>
</taxonomicName>
-like taxon dispersed widely, giving rise to multiple independent lineages of naucorids. This concept was astutely hypothesized by
<bibRefCitation id="0E664B68FF95FFCBFDA281DA0620F89C" author="Polhemus DA &amp; Polhemus JT" pageId="24" pageNumber="1269" pagination="265 - 269" refId="ref23665" refString="Polhemus DA, Polhemus JT. 1987. A new genus of Naucoridae (Hemiptera) from the Philippines, with comments on zoogeography. Pan-Pacific Entomologist 63: 265 - 269." type="journal article" year="1987">Polhemus &amp; Polhemus (1987)</bibRefCitation>
when considering the origins of the Philippine and New Guinean stock based on head structure and male genitalia. We now have strong corroborating molecular evidence for these relationships.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="3B81F9D7095B0357F69DB5A7EF488B6C">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:date id="693BC45FAED1DC204256D5438D03777D">2022</mods:date>
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<paragraph id="6A483699FF9BFFC5FCA583860218FB14" blockId="22.[865,1368,1185,1210]" box="[865,1368,1185,1210]" pageId="22" pageNumber="1267">
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SUBFAMILY
<taxonomicName id="ADF74D1AFF9BFFC5FC2C838603EBFB1A" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1000,1195,1185,1209]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Macrocorinae">MACROCORINAE</taxonomicName>
<emphasis id="5883EA8BFF9BFFC5FB7683830218FB14" bold="true" box="[1202,1368,1185,1209]" pageId="22" pageNumber="1267">
<taxonomicNameLabel id="43B057F0FF9BFFC5FB7683830218FB14" box="[1202,1368,1185,1209]" pageId="22" pageNumber="1267" rank="subFamily">SUBFAM. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="22ED6512FF9BFFC4FBC883EA0662FAE2" lastPageId="23" lastPageNumber="1268" pageId="22" pageNumber="1267" type="description">
<paragraph id="6A483699FF9BFFC5FBC883EA03EEFB49" blockId="22.[1036,1198,1229,1254]" box="[1036,1198,1229,1254]" pageId="22" pageNumber="1267">
<figureCitation id="F2CC2A1CFF9BFFC5FBC883EA0335FB49" box="[1036,1141,1229,1254]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="22" pageNumber="1267">FIGS 10G</figureCitation>
,
<figureCitation id="F2CC2A1CFF9BFFC5FB4683EA03EEFB49" box="[1154,1198,1229,1254]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="22" pageNumber="1267">11G</figureCitation>
</paragraph>
<paragraph id="6A483699FF9BFFC5FCED82240238FA9A" blockId="22.[809,1425,1282,1335]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FCED82240336FAB5" box="[809,1142,1282,1304]" italics="true" pageId="22" pageNumber="1267">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF9BFFC5FCB382060238FA9A" box="[887,1400,1313,1335]" pageId="22" pageNumber="1267">75C4C846-27F8-42DC-91B4-7812C7789D36</uuid>
</paragraph>
<paragraph id="6A483699FF9BFFC4FCED824607B4FE7C" blockId="22.[809,1386,1376,1398]" lastBlockId="23.[164,781,198,465]" lastPageId="23" lastPageNumber="1268" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FCED824604F9FAD8" box="[809,953,1376,1398]" italics="true" pageId="22" pageNumber="1267">
<typeStatus id="B54C883BFF9BFFC5FCED8246041EFADB" box="[809,862,1377,1398]" pageId="22" pageNumber="1267">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF9BFFC5FC0182470223FADB" authority="Signoret, 1861" authorityName="Signoret" authorityYear="1861" box="[965,1379,1376,1398]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="species" species="flavicollis">
<emphasis id="5883EA8BFF9BFFC5FC01824703F7FAD8" box="[965,1207,1376,1397]" italics="true" pageId="22" pageNumber="1267">Macrocoris flavicollis</emphasis>
<bibRefCitation id="0E664B68FF9BFFC5FB7A82470223FADB" author="Signoret V" box="[1214,1379,1376,1398]" pageId="22" pageNumber="1267" pagination="917 - 972" refId="ref24335" refString="Signoret V. 1861. Faune des Hemipteres de Madagascar (Suite et fin.). 2 Partie. Heteropteres. Annales de la Societe Entomologique de France, Serie 3 8: 917 - 972." type="journal article" year="1861">Signoret, 1861</bibRefCitation>
</taxonomicName>
.
<emphasis id="5883EA8BFF9AFFC4FF6087E006D9FF76" box="[164,409,198,220]" italics="true" pageId="23" pageNumber="1268">Taxonomic history:</emphasis>
The genera
<taxonomicName id="ADF74D1AFF9AFFC4FD8187E00591FF71" authorityName="Signoret" authorityYear="1861" box="[581,721,199,220]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FD8187E00591FF71" box="[581,721,199,220]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9AFFC4FF6087C20623FF57" authorityName="Montandon" authorityYear="1913" box="[164,355,229,250]" class="Insecta" family="Naucoridae" genus="Neomacrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FF6087C20623FF57" box="[164,355,229,250]" italics="true" pageId="23" pageNumber="1268">Neomacrocoris</emphasis>
</taxonomicName>
have been held in
<taxonomicName id="ADF74D1AFF9AFFC4FDAB87C20447FF56" authorityName="LEACH" authorityYear="1815" box="[623,775,229,251]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF9AFFC4FF6086240662FEB4" authorityName="POPOV" authorityYear="1970" box="[164,290,259,281]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini</taxonomicName>
(
<bibRefCitation id="0E664B68FF9AFFC4FEFC8625050EFEB4" author="Stys P &amp; Jansson A" box="[312,590,258,282]" pageId="23" pageNumber="1268" pagination="1 - 44" refId="ref25103" refString="Stys P, Jansson A. 1988. Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica 50: 1 - 44." type="journal article" year="1988">Štys &amp; Jansson, 1988</bibRefCitation>
;
<taxonomicName id="ADF74D1AFF9AFFC4FDA6862405BAFEB4" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[610,762,259,281]" genus="Zettel" pageId="23" pageNumber="1285" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF9AFFC4FDA6862405BAFEB4" author="Zettel H" box="[610,762,259,281]" pageId="23" pageNumber="1268" pagination="1085 - 1095" refId="ref25492" refString="Zettel H. 2001. Die indische Zwergschwimmwanze Nanonaucoris gen. n., eine neue Gattung der Naucorini (Heteroptera: Naucoridae) aus Sudindien. Linzer biologische Beitrage 33: 1085 - 1095." type="journal article" year="2001">Zettel, 2001</bibRefCitation>
</taxonomicName>
). Two species of
<taxonomicName id="ADF74D1AFF9AFFC4FEA5860406A7FE95" authorityName="Signoret" authorityYear="1861" box="[353,487,291,312]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FEA5860406A7FE95" box="[353,487,291,312]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
from
<collectingCountry id="12E07609FF9AFFC4FDF086050588FE95" box="[564,712,290,312]" name="Madagascar" pageId="23" pageNumber="1268">Madagascar</collectingCountry>
were originally described in
<taxonomicName id="ADF74D1AFF9AFFC4FE72866607BBFED9" authority="(Montandon, 1897 a)" baseAuthorityName="Montandon" baseAuthorityYear="1897" class="Insecta" genus="Pseudambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FE728666052EFEFB" box="[438,622,321,342]" italics="true" pageId="23" pageNumber="1268">Pseudambrysus</emphasis>
(
<bibRefCitation id="0E664B68FF9AFFC4FDBB866607B0FED9" author="Montandon AL" pageId="23" pageNumber="1268" pagination="6 - 23" refId="ref22979" refString="Montandon AL. 1897 a. Hemiptera Cryptocerata. Fam. Naucoridae. Sous-fam. Cryptocricinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 6 - 23." type="journal article" year="1897">Montandon, 1897a</bibRefCitation>
)
</taxonomicName>
; however, the genus was downgraded to subgenus by
<bibRefCitation id="0E664B68FF9AFFC4FE8486590559FE3E" author="Montandon AL" box="[320,537,382,404]" pageId="23" pageNumber="1268" pagination="414 - 422" refId="ref23172" refString="Montandon AL. 1900. Hemipteres exotiques nouveaux ou peu connus des collections du Musee National Hongrois. Termeszetrajzi Fuzetek 23: 414 - 422." type="journal article" year="1900">Montandon (1900)</bibRefCitation>
, then appeared as a synonym of
<taxonomicName id="ADF74D1AFF9AFFC4FEF586BA06F1FE1F" authorityName="Signoret" authorityYear="1861" box="[305,433,413,434]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FEF586BA06F1FE1F" box="[305,433,413,434]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
in the catalogue of
<bibRefCitation id="0E664B68FF9AFFC4FD5E86BA07ACFE7D" author="La Rivers I" pageId="23" pageNumber="1268" pagination="1 - 120" refId="ref22462" refString="La Rivers I. 1971. Studies of Naucoridae (Hemiptera). Biological Society of Nevada Memoirs 2: 1 - 120." type="journal article" year="1971">La Rivers (1971)</bibRefCitation>
.
</paragraph>
<caption id="3E886611FF9BFFC5FF5580720277F8C1" ID-DOI="http://doi.org/10.5281/zenodo.6994650" ID-Zenodo-Dep="6994650" box="[145,1335,1877,1900]" httpUri="https://zenodo.org/record/6994650/files/figure.png" pageId="22" pageNumber="1267" startId="22.[145,223,1877,1899]" targetBox="[355,1214,1461,1837]" targetPageId="22">
<paragraph id="6A483699FF9BFFC5FF5580720277F8C1" blockId="22.[145,1335,1877,1900]" box="[145,1335,1877,1900]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FF5580720646F8C7" bold="true" box="[145,262,1877,1899]" pageId="22" pageNumber="1267">Figure 16.</emphasis>
Ventrolateral aspect of
<taxonomicName id="ADF74D1AFF9BFFC5FDC1807105A6F8C1" authorityName="Montandon" authorityYear="1898" box="[517,742,1878,1900]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis id="5883EA8BFF9BFFC5FDC1807105A6F8C1" box="[517,742,1878,1900]" italics="true" pageId="22" pageNumber="1267">Limnocoris inornatus</emphasis>
</taxonomicName>
, showing meso- and metasternal carinae (white arrows).
</paragraph>
</caption>
<paragraph id="6A483699FF9AFFC4FF6086DD0622FD04" blockId="23.[164,780,506,681]" pageId="23" pageNumber="1268">
<emphasis id="5883EA8BFF9AFFC4FF6086DD06C1FDBD" box="[164,385,506,528]" italics="true" pageId="23" pageNumber="1268">Revised taxonomy:</emphasis>
Based on the evidence from the molecular analyses and morphology of the aedeagus, I here erect the new subfamily
<taxonomicName id="ADF74D1AFF9AFFC4FDD1851005F3FDE0" authorityName="Sites" authorityYear="2022" box="[533,691,567,589]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="subFamily" subFamily="Macrocorinae">Macrocorinae</taxonomicName>
to hold these two sister genera. As such,
<taxonomicName id="ADF74D1AFF9AFFC4FDD9857005DAFDC1" authorityName="Signoret" authorityYear="1861" box="[541,666,599,620]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FDD9857005DAFDC1" box="[541,666,599,620]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
(
<figureCitation id="F2CC2A1CFF9AFFC4FD6E85710444FDC6" box="[682,772,598,620]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="23" pageNumber="1268">
Fig.
<quantity id="AD0F9B7CFF9AFFC4FD1F85710444FDC6" box="[731,772,598,619]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="23" pageNumber="1268" unit="g" value="10.0">10g</quantity>
</figureCitation>
) and
<taxonomicName id="ADF74D1AFF9AFFC4FF1C855206C3FD27" authorityName="Montandon" authorityYear="1913" box="[216,387,629,650]" class="Insecta" family="Naucoridae" genus="Neomacrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FF1C855206C3FD27" box="[216,387,629,650]" italics="true" pageId="23" pageNumber="1268">Neomacrocoris</emphasis>
</taxonomicName>
are transferred from
<taxonomicName id="ADF74D1AFF9AFFC4FD418552044CFD26" authorityName="LEACH" authorityYear="1815" box="[645,780,629,651]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
to
<taxonomicName id="ADF74D1AFF9AFFC4FF0585B4061FFD04" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[193,351,659,681]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1260" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Macrocorinae">Macrocorinae</taxonomicName>
.
</paragraph>
<paragraph id="6A483699FF9AFFC4FF6085F40658FBDB" blockId="23.[164,781,722,1143]" pageId="23" pageNumber="1268">
<emphasis id="5883EA8BFF9AFFC4FF6085F40667FD45" box="[164,295,723,744]" italics="true" pageId="23" pageNumber="1268">Diagnosis:</emphasis>
Members of
<taxonomicName id="ADF74D1AFF9AFFC4FE1085F5053DFD45" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[468,637,722,744]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1260" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Macrocorinae">Macrocorinae</taxonomicName>
are among the most dorsoventrally robust in the family. Both genera have a dramatically asymmetrical aedeagus with a characteristic shape and substantial, heavily sclerotized, articulated vesica (
<figureCitation id="F2CC2A1CFF9AFFC4FD53846A05BCFCCF" box="[663,764,845,867]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="23" pageNumber="1268">
Fig.
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). Large, elliptical parameres flank the aedeagus in
<taxonomicName id="ADF74D1AFF9AFFC4FF0384AC060FFC0D" authorityName="Signoret" authorityYear="1861" box="[199,335,907,928]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FF0384AC060FFC0D" box="[199,335,907,928]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
(
<figureCitation id="F2CC2A1CFF9AFFC4FEA084AD0689FC32" box="[356,457,906,928]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="23" pageNumber="1268">
Fig.
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</figureCitation>
), whereas parameres are absent in
<emphasis id="5883EA8BFF9AFFC4FEDF848E0690FC13" box="[283,464,937,958]" italics="true" pageId="23" pageNumber="1268">
<taxonomicName id="ADF74D1AFF9AFFC4FEDF848E068CFC13" authorityName="Montandon" authorityYear="1913" box="[283,460,937,958]" class="Insecta" family="Naucoridae" genus="Neomacrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">Neomacrocoris</taxonomicName>
.
</emphasis>
The mesosternal carina is sufficiently setose to obscure the cuticular structure of the carina, which can be entire (
<taxonomicName id="ADF74D1AFF9AFFC4FDB384C00443FC51" authorityName="Signoret" authorityYear="1861" box="[631,771,999,1020]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FDB384C00443FC51" box="[631,771,999,1020]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
) or crenulate (
<taxonomicName id="ADF74D1AFF9AFFC4FE9E83220556FBB7" authorityName="Montandon" authorityYear="1913" box="[346,534,1029,1050]" class="Insecta" family="Naucoridae" genus="Neomacrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FE9E83220556FBB7" box="[346,534,1029,1050]" italics="true" pageId="23" pageNumber="1268">Neomacrocoris</emphasis>
</taxonomicName>
) (
<bibRefCitation id="0E664B68FF9AFFC4FDF2832207A2FB95" author="Sites RW &amp; Mbogho AY" pageId="23" pageNumber="1268" pagination="1 - 39" refId="ref24588" refString="Sites RW, Mbogho AY. 2012. Revision of the African genus Neomacrocoris (Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Zootaxa 3555: 1 - 39." type="journal article" year="2012">
<taxonomicName id="ADF74D1AFF9AFFC4FDF283220535FBB6" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[566,629,1029,1051]" genus="Sites" pageId="23" pageNumber="1270" rank="genus" status="gen. nov.">Sites</taxonomicName>
&amp; Mbogho, 2012
</bibRefCitation>
). The pygophore is greatly elongated at the midline to nearly half the aedeagus length (
<figureCitation id="F2CC2A1CFF9AFFC4FF6983460649FBDB" box="[173,265,1121,1143]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="23" pageNumber="1268">
Fig.
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).
</paragraph>
<paragraph id="6A483699FF9AFFC4FF6083870662FAE2" blockId="23.[164,780,1183,1359]" pageId="23" pageNumber="1268">
<emphasis id="5883EA8BFF9AFFC4FF608387066DFB18" box="[164,301,1184,1205]" italics="true" pageId="23" pageNumber="1268">Comments:</emphasis>
This subfamily is endemic to Africa, with
<taxonomicName id="ADF74D1AFF9AFFC4FF2483980620FB79" authorityName="Signoret" authorityYear="1861" box="[224,352,1215,1236]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FF2483980620FB79" box="[224,352,1215,1236]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
occurring on both the mainland and
<collectingCountry id="12E07609FF9AFFC4FF6083FA067AFB5E" box="[164,314,1245,1267]" name="Madagascar" pageId="23" pageNumber="1268">Madagascar</collectingCountry>
and
<taxonomicName id="ADF74D1AFF9AFFC4FEBA83FA0573FB5F" authorityName="Montandon" authorityYear="1913" box="[382,563,1245,1266]" class="Insecta" family="Naucoridae" genus="Neomacrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FEBA83FA0573FB5F" box="[382,563,1245,1266]" italics="true" pageId="23" pageNumber="1268">Neomacrocoris</emphasis>
</taxonomicName>
restricted to the African mainland.
<taxonomicName id="ADF74D1AFF9AFFC4FEBC83DB055FFABC" authorityName="Montandon" authorityYear="1913" box="[376,543,1276,1297]" class="Insecta" family="Naucoridae" genus="Neomacrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FEBC83DB055FFABC" box="[376,543,1276,1297]" italics="true" pageId="23" pageNumber="1268">Neomacrocoris</emphasis>
</taxonomicName>
was revised recently (
<bibRefCitation id="0E664B68FF9AFFC4FF68823D06E2FA82" author="Sites RW &amp; Mbogho AY" box="[172,418,1306,1328]" pageId="23" pageNumber="1268" pagination="1 - 39" refId="ref24588" refString="Sites RW, Mbogho AY. 2012. Revision of the African genus Neomacrocoris (Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Zootaxa 3555: 1 - 39." type="journal article" year="2012">
<taxonomicName id="ADF74D1AFF9AFFC4FF68823D07A5FA9D" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[172,229,1306,1328]" genus="Sites" pageId="23" pageNumber="1270" rank="genus" status="gen. nov.">Sites</taxonomicName>
&amp; Mbogho, 2012
</bibRefCitation>
), whereas
<taxonomicName id="ADF74D1AFF9AFFC4FDD8823C05D9FA9D" authorityName="Signoret" authorityYear="1861" box="[540,665,1307,1328]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="1268" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9AFFC4FDD8823C05D9FA9D" box="[540,665,1307,1328]" italics="true" pageId="23" pageNumber="1268">Macrocoris</emphasis>
</taxonomicName>
is in need of revision.
</paragraph>
</subSubSection>
</treatment>
</document>

159
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<paragraph id="6A483699FF9BFFC5FF22851C05E3FDF9" blockId="22.[230,675,571,596]" box="[230,675,571,596]" pageId="22" pageNumber="1267">
SUBFAMILY
<taxonomicName id="ADF74D1AFF9BFFC5FEA9851C05E3FDF9" ID-CoL="JTN" authority="STAL, 1876" authorityName="STAL" authorityYear="1876" box="[365,675,571,596]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Limnocorinae">
LIMNOCORINAE
<bibRefCitation id="0E664B68FF9BFFC5FDE0851C05E3FDF9" author="Stal C" box="[548,675,571,596]" pageId="22" pageNumber="1267" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">STÅL, 1876</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9BFFC5FEA585400568FD2D" box="[353,552,615,640]" pageId="22" pageNumber="1267" type="description">
<paragraph id="6A483699FF9BFFC5FEA585400568FD2D" blockId="22.[353,552,615,640]" box="[353,552,615,640]" pageId="22" pageNumber="1267">
<figureCitation id="F2CC2A1CFF9BFFC5FEA585400688FDD2" box="[353,456,615,640]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="22" pageNumber="1267">FIGS 10F</figureCitation>
,
<figureCitation id="F2CC2A1CFF9BFFC5FE11854006BFFDD2" box="[469,511,615,640]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="22" pageNumber="1267">11F</figureCitation>
,
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</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9BFFC5FF5585B905BAFB32" pageId="22" pageNumber="1267" type="materials_examined">
<paragraph id="6A483699FF9BFFC5FF5585B905CDFD1E" blockId="22.[145,653,669,691]" box="[145,653,669,691]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FF5585B90661FD1E" box="[145,289,670,691]" italics="true" pageId="22" pageNumber="1267">
<typeStatus id="B54C883BFF9BFFC5FF5585B90786FD1E" box="[145,198,670,691]" pageId="22" pageNumber="1267">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF9BFFC5FEE985B905C6FD1E" authority="Stal, 1858" authorityName="Stal" authorityYear="1858" box="[301,646,669,691]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="5883EA8BFF9BFFC5FEE985B9054DFD1E" box="[301,525,670,691]" italics="true" pageId="22" pageNumber="1267">Limnocoris insignis</emphasis>
<bibRefCitation id="0E664B68FF9BFFC5FDD085BA05C6FD1E" author="Stal C" box="[532,646,669,691]" pageId="22" pageNumber="1267" pagination="1 - 84" refId="ref24954" refString="Stal C. 1858. Bidrag till Rio Janeiro-traktens Hemipter-fauna. Kongliga Svenska Vetenskaps-Akademiens Handlingar 2: 1 - 84." type="journal article" year="1858">Stål, 1858</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="6A483699FF9BFFC5FF5585FB05BAFB32" blockId="22.[145,762,731,1183]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FF5585FB062CFD5C" box="[145,364,732,753]" italics="true" pageId="22" pageNumber="1267">Taxonomic history:</emphasis>
<bibRefCitation id="0E664B68FF9BFFC5FEBD85FC06B9FD5F" author="Stal C" box="[377,505,731,754]" pageId="22" pageNumber="1267" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">Stål (1876)</bibRefCitation>
proposed the division Limnocoraria to contain two genera,
<taxonomicName id="ADF74D1AFF9BFFC5FD9B85DD05B9FCA2" box="[607,761,762,783]" class="Insecta" family="Naucoridae" genus="Borborocoris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9BFFC5FD9B85DD05B9FCA2" box="[607,761,762,783]" italics="true" pageId="22" pageNumber="1267">Borborocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9BFFC5FF08843D061AFC82" authorityName="Stal" authorityYear="1858" box="[204,346,794,815]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9BFFC5FF08843D061AFC82" box="[204,346,794,815]" italics="true" pageId="22" pageNumber="1267">Limnocoris</emphasis>
</taxonomicName>
, that he described.
<bibRefCitation id="0E664B68FF9BFFC5FDAD843E07A4FCE0" author="Montandon AL" pageId="22" pageNumber="1267" pagination="1 - 8" refId="ref23040" refString="Montandon AL. 1897 c. Hemiptera Cryptocerata. Revision de le s. fam. ' Limnocorinae'. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino 12: 1 - 8." type="journal article" year="1897">Montandon (1897c)</bibRefCitation>
elevated Limnocoraria to subfamily status as
<taxonomicName id="ADF74D1AFF9BFFC5FF5584710671FCC1" authorityName="STAL" authorityYear="1876" box="[145,305,854,876]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="subFamily" subFamily="Limnocorinae">Limnocorinae</taxonomicName>
and synonymized the two genera. Two additional genera,
<taxonomicName id="ADF74D1AFF9BFFC5FEAB845206B3FC27" box="[367,499,885,906]" class="Insecta" family="Naucoridae" genus="Sattleriella" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9BFFC5FEAB845206B3FC27" box="[367,499,885,906]" italics="true" pageId="22" pageNumber="1267">Sattleriella</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9BFFC5FDF4845105F2FC26" box="[560,690,886,907]" class="Insecta" family="Naucoridae" genus="Usingerina" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9BFFC5FDF4845105F2FC26" box="[560,690,886,907]" italics="true" pageId="22" pageNumber="1267">Usingerina</emphasis>
</taxonomicName>
, have been described, but ultimately were also synonymized with
<taxonomicName id="ADF74D1AFF9BFFC5FF0B84940614FC65" authorityName="Stal" authorityYear="1858" box="[207,340,947,968]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9BFFC5FF0B84940614FC65" box="[207,340,947,968]" italics="true" pageId="22" pageNumber="1267">Limnocoris</emphasis>
</taxonomicName>
by
<bibRefCitation id="0E664B68FF9BFFC5FE4684950583FC65" author="Nieser N &amp; Lopez Ruf M" box="[386,707,946,968]" pageId="22" pageNumber="1267" pagination="261 - 328" refId="ref23454" refString="Nieser N, Lopez Ruf M. 2001. A review of Limnocoris Stal (Heteroptera: Naucoridae) in southern South America east of the Andes. Tijdschrift voor Entomologie 144: 261 - 328." type="journal article" year="2001">Nieser &amp; Lopez Ruf (2001)</bibRefCitation>
and
<bibRefCitation id="0E664B68FF9BFFC5FF5584F6063BFC4B" author="Sites RW &amp; Willig MR" box="[145,379,977,999]" pageId="22" pageNumber="1267" pagination="803 - 814" refId="ref24770" refString="Sites RW, Willig MR. 1994. Efficacy of mensural characters in discriminating among species of Naucoridae (Insecta: Hemiptera): multivariate approaches and ontogenetic perspectives. Annals of the Entomological Society of America 87: 803 - 814." type="journal article" year="1994">
<taxonomicName id="ADF74D1AFF9BFFC5FF5584F60789FC4A" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[145,201,977,999]" genus="Sites" pageId="22" pageNumber="1270" rank="genus" status="gen. nov.">Sites</taxonomicName>
&amp; Willig (1994)
</bibRefCitation>
, respectively. Thus, the subfamily is monogeneric.
<bibRefCitation id="0E664B68FF9BFFC5FE9784D706ADFBA8" author="Popov YA" box="[339,493,1008,1029]" pageId="22" pageNumber="1267" pagination="93 - 98" refId="ref23866" refString="Popov YA. 1970. Notes on the classification of the recent Naucoridae (Heteroptera: Nepomorpha). Bulletin of the Polish Academy of Sciences (Biological Sciences) 18: 93 - 98." type="journal article" year="1970">Popov (1970)</bibRefCitation>
considered this group as a tribe-level taxon, Limnocorini, in the subfamily
<taxonomicName id="ADF74D1AFF9BFFC5FF55830A0655FBEE" authorityName="LEACH" authorityYear="1815" box="[145,277,1069,1091]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
, but this classification scheme has not been followed by other specialists.
<taxonomicName id="ADF74D1AFF9BFFC5FE26836B05C0FBCF" authorityName="STAL" authorityYear="1876" box="[482,640,1100,1122]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="subFamily" subFamily="Limnocorinae">Limnocorinae</taxonomicName>
comprises 71 described species and many more awaiting description, including species included in this analysis.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9BFFC5FF5583EF05A1FAB6" pageId="22" pageNumber="1267" type="discussion">
<paragraph id="6A483699FF9BFFC5FF5583EF05A1FAB6" blockId="22.[145,760,1224,1307]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FF5583EF063AFB70" box="[145,378,1224,1245]" italics="true" pageId="22" pageNumber="1267">Revised taxonomy:</emphasis>
This is the only subfamily of
<taxonomicName id="ADF74D1AFF9BFFC5FF5583C1065AFB51" authorityName="LEACH" authorityYear="1815" box="[145,282,1254,1276]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="family">Naucoridae</taxonomicName>
that remains unchanged following this revision and that of
<bibRefCitation id="0E664B68FF9BFFC5FEB38222059BFAB7" author="Reynoso-Velasco D &amp; Sites RW" box="[375,731,1285,1307]" pageId="22" pageNumber="1267" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF9BFFC5FD908222059BFAB7" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[596,731,1285,1307]" genus="Sites" pageId="22" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9BFFC5FF55826303F9FE9B" pageId="22" pageNumber="1267" type="diagnosis">
<paragraph id="6A483699FF9BFFC5FF55826303F9FE9B" blockId="22.[145,761,1348,1400]" lastBlockId="22.[809,1425,197,311]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FF5582630650FAF4" box="[145,272,1348,1369]" italics="true" pageId="22" pageNumber="1267">Diagnosis:</emphasis>
This subfamily is distinguished by the strongly developed meso- and metasternal midventral carinae (
<figureCitation id="F2CC2A1CFF9BFFC5FC5C87E204AEFF76" box="[920,1006,197,219]" captionStart="Figure 16" captionStartId="22.[145,223,1877,1899]" captionTargetBox="[355,1214,1461,1837]" captionTargetId="figure-559@22.[355,1214,1461,1837]" captionTargetPageId="22" captionText="Figure 16. Ventrolateral aspect of Limnocoris inornatus, showing meso- and metasternal carinae (white arrows)." figureDoi="http://doi.org/10.5281/zenodo.6994650" httpUri="https://zenodo.org/record/6994650/files/figure.png" pageId="22" pageNumber="1267">Fig. 16</figureCitation>
), a condition that can be species specific. Many species are concolorous brownish-green, flattened and ovate to circular in outline, thus their bodies are saucer shaped (
<figureCitation id="F2CC2A1CFF9BFFC5FB90860603EAFE9A" box="[1108,1194,289,311]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="22" pageNumber="1267">Fig. 10f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9BFFC5FCED864702D2FB9A" pageId="22" pageNumber="1267" type="discussion">
<paragraph id="6A483699FF9BFFC5FCED864704B0FC33" blockId="22.[809,1426,352,1079]" pageId="22" pageNumber="1267">
<emphasis id="5883EA8BFF9BFFC5FCED864704E5FED8" box="[809,933,352,373]" italics="true" pageId="22" pageNumber="1267">Comments:</emphasis>
This subfamily is exclusively New World in distribution and is sister to
<taxonomicName id="ADF74D1AFF9BFFC5FB90865903BEFE39" authorityName="MONTANDON" authorityYear="1897" box="[1108,1278,382,404]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
, which also is restricted to the New World. Both subfamilies are more species rich in the Neotropics, with minimal incursions into the Nearctic region. The molecular distance of
<taxonomicName id="ADF74D1AFF9BFFC5FCED86DE0488FDA2" authorityName="STAL" authorityYear="1876" box="[809,968,505,527]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="subFamily" subFamily="Limnocorinae">Limnocorinae</taxonomicName>
in the dendrograms of both the BI and ML analyses was sufficiently great that the clade length was interrupted and shortened to be depicted on the graphics. These two subfamilies are sufficiently adapted to their particular environmental circumstances that morphological synapomorphies to link them are not evident; however, biogeographic and molecular evidence (pp = 0.90, bs = 99) provides compelling evidence for this sister group relationship. The North American and South American Andean faunas were each recently revised (
<bibRefCitation id="0E664B68FF9BFFC5FC49840B03E6FCEC" author="Rodrigues HDD &amp; Sites RW" box="[909,1190,812,834]" pageId="22" pageNumber="1267" pagination="451 - 497" refId="ref24078" refString="Rodrigues HDD, Sites RW. 2019. Revision of Limnocoris (Heteroptera: Nepomorpha: Naucoridae) of North America. Zootaxa 4629: 451 - 497." type="journal article" year="2019">
Rodrigues &amp;
<taxonomicName id="ADF74D1AFF9BFFC5FBE2840B03E6FCEC" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1062,1190,812,834]" genus="Sites" pageId="22" pageNumber="1270" rank="genus" status="gen. nov.">Sites, 2019</taxonomicName>
</bibRefCitation>
,
<bibRefCitation id="0E664B68FF9BFFC5FB76840B03A9FCEC" author="Rodrigues HDD &amp; Sites RW" box="[1202,1257,812,833]" pageId="22" pageNumber="1267" pagination="1 - 93" refId="ref24107" refString="Rodrigues HDD, Sites RW. 2021. Revision of Limnocoris Stal (Heteroptera: Nepomorpha: Naucoridae) of the tropical Andes. Zootaxa 4986: 1 - 93." type="journal article" year="2021">2021</bibRefCitation>
, respectively), with further revisions in progress for the Guiana Shield, Amazonian, and southern South American faunas by the same authors.
</paragraph>
<paragraph id="6A483699FF9BFFC5FC85848102D2FB9A" blockId="22.[809,1426,352,1079]" pageId="22" pageNumber="1267">
Although
<taxonomicName id="ADF74D1AFF9BFFC5FC7E8481031FFC11" authorityName="STAL" authorityYear="1876" box="[954,1119,934,956]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="1267" phylum="Arthropoda" rank="subFamily" subFamily="Limnocorinae">Limnocorinae</taxonomicName>
was recovered as clearly monophyletic, relationships among species and species groups in the subfamily were not well supported in either analysis. Further research will be necessary to determine evolutionary relationships in the subfamily.
</paragraph>
</subSubSection>
</treatment>
</document>

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SUBFAMILY
<taxonomicName id="ADF74D1AFF9CFFC2FBD38176020DF9C7" ID-CoL="JS3" authority="STAL, 1876" authorityName="STAL" authorityYear="1876" box="[1047,1357,1617,1642]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="subFamily" status="stat. nov." subFamily="Laccocorinae">
LACCOCORINAE
<bibRefCitation id="0E664B68FF9CFFC2FB0A8176020DF9C7" author="Stal C" box="[1230,1357,1617,1642]" pageId="17" pageNumber="1262" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">STÅL, 1876</bibRefCitation>
</taxonomicName>
</paragraph>
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<figureCitation id="F2CC2A1CFF9CFFC2FC43815A04A1F939" box="[903,993,1661,1686]" captionStart="Figure 3" captionStartId="8.[145,230,1468,1490]" captionTargetBox="[172,733,195,1428]" captionTargetId="figure-532@8.[172,733,195,1428]" captionTargetPageId="8" captionText="Figure 3. Mouthpart positions: recessed rostrum of Cheirochela thailandana (a); lateral views of Laccocoris spurcus (b) and Australambrysus crenulatus (c). White arrows indicate positions of labrum with respect to the functional anterior margin of the head." figureDoi="http://doi.org/10.5281/zenodo.6994613" httpUri="https://zenodo.org/record/6994613/files/figure.png" pageId="17" pageNumber="1262">FIGS 3B</figureCitation>
,
<figureCitation id="F2CC2A1CFF9CFFC2FC2A815A0358F93B" box="[1006,1048,1661,1686]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="9.[163,249,1379,1401]" captionStartId-1="10.[145,223,735,757]" captionStartId-2="10.[145,228,1329,1351]" captionTargetBox-0="[163,781,195,1339]" captionTargetBox-1="[145,1426,195,695]" captionTargetBox-2="[216,689,828,1289]" captionTargetId-0="figure-575@9.[163,781,195,1339]" captionTargetId-1="figure-384@10.[145,1426,195,695]" captionTargetId-2="figure-415@10.[216,689,828,1289]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="10" captionText-0="Figure 4. Foretibiae and foretarsi of Laccocorinae: Ctenipocoris asiaticus (a), Heleolaccocoris ovatus (b), Namtokocoris dalanta (c)." captionText-1="Figure 5. Heleocoris elongatus mesofemur (a), metafemur (b), Tsingala gen. nov. sp. C mesofemur (c), metafemur (d). White arrows indicate posteroventral spine rows, black arrows indicate anteroventral spine rows." captionText-2="Figure 6. Laccocoris spurcus showing the transverse sulcus at the base of the labrum, which is characteristic of most Laccocorini." figureDoi-0="http://doi.org/10.5281/zenodo.6994615" figureDoi-1="http://doi.org/10.5281/zenodo.6994617" figureDoi-2="http://doi.org/10.5281/zenodo.6994621" httpUri-0="https://zenodo.org/record/6994615/files/figure.png" httpUri-1="https://zenodo.org/record/6994617/files/figure.png" httpUri-2="https://zenodo.org/record/6994621/files/figure.png" pageId="17" pageNumber="1262">
4–
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</figureCitation>
,
<figureCitation id="F2CC2A1CFF9CFFC2FBE1815A0310F939" box="[1061,1104,1661,1686]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="17" pageNumber="1262">10E</figureCitation>
,
<figureCitation id="F2CC2A1CFF9CFFC2FB99815A03C8F939" box="[1117,1160,1661,1686]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="17" pageNumber="1262">11E</figureCitation>
,
<figureCitation id="F2CC2A1CFF9CFFC2FB51815A03F1F93B" box="[1173,1201,1661,1686]" captionStart="Figure 12" captionStartId="17.[163,243,870,892]" captionTargetBox="[163,1443,195,830]" captionTargetId="figure-534@17.[163,1443,195,830]" captionTargetPageId="17" captionText="Figure 12. Mesotibia and mesotarsus of male and female Heleolaccocoris ovatus showing a sexually dimorphic ventral tomentose pad of hairs." figureDoi="http://doi.org/10.5281/zenodo.6994637" httpUri="https://zenodo.org/record/6994637/files/figure.png" pageId="17" pageNumber="1262">12</figureCitation>
,
<figureCitation id="F2CC2A1CFF9CFFC2FB7A815A039AF93B" box="[1214,1242,1661,1686]" captionStart="Figure 14" captionStartId="19.[163,244,765,787]" captionTargetBox="[163,1442,195,726]" captionTargetId="figure-559@19.[163,1442,195,726]" captionTargetPageId="19" captionText="Figure 14. Revised generic representation of the former genus Heleocoris. Dorsal habitus of Heleocoris vicinus (a), Laccocoris minisculus (b), Heleolaccocoris ovatus (c), Tsingala humeralis (d)." figureDoi="http://doi.org/10.5281/zenodo.6994645" httpUri="https://zenodo.org/record/6994645/files/figure.png" pageId="17" pageNumber="1262">14</figureCitation>
,
<figureCitation id="F2CC2A1CFF9CFFC2FB2C815A0244F93B" box="[1256,1284,1661,1686]" captionStart="Figure 15" captionStartId="20.[145,226,1055,1077]" captionTargetBox="[145,1425,195,1015]" captionTargetId="figure-409@20.[145,1425,195,1015]" captionTargetPageId="20" captionText="Figure 15. Revised generic representation of the former genus Laccocoris. Dorsal habitus of Laccocoris spurcus (a), Heleolaccocoris horvathi (b)." figureDoi="http://doi.org/10.5281/zenodo.6994647" httpUri="https://zenodo.org/record/6994647/files/figure.png" pageId="17" pageNumber="1262">15</figureCitation>
,
<figureCitation id="F2CC2A1CFF9CFFC2FAD5815A026DF93B" box="[1297,1325,1661,1686]" captionStart="Figure 20" captionStartId="28.[146,226,1107,1129]" captionTargetBox="[145,1425,195,1067]" captionTargetId="figure-347@28.[145,1425,195,1067]" captionTargetPageId="28" captionText="Figure 20. Tsingala humeralis male: dorsal habitus (a), genital capsule with terga IX and X removed (b), left and right parameres (c). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994662" httpUri="https://zenodo.org/record/6994662/files/figure.png" pageId="17" pageNumber="1262">20</figureCitation>
,
<figureCitation id="F2CC2A1CFF9CFFC2FAFE815A0216F93B" box="[1338,1366,1661,1686]" captionStart="Figure 21" captionStartId="29.[164,242,1876,1898]" captionTargetBox="[163,1443,921,1837]" captionTargetId="figure-364@29.[163,1443,921,1837]" captionTargetPageId="29" captionText="Figure 21. Heleolaccocoris horvathi male: dorsal habitus (a), genital capsule with terga IX and X removed (b)." figureDoi="http://doi.org/10.5281/zenodo.6994664" httpUri="https://zenodo.org/record/6994664/files/figure.png" pageId="17" pageNumber="1262">21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9CFFC2FCFF8194026CF965" box="[827,1324,1714,1736]" pageId="17" pageNumber="1262" type="type_taxon">
<paragraph id="6A483699FF9CFFC2FCFF8194026CF965" blockId="17.[827,1324,1714,1736]" box="[827,1324,1714,1736]" pageId="17" pageNumber="1262">
<emphasis id="5883EA8BFF9CFFC2FCFF8194048BF965" box="[827,971,1715,1736]" italics="true" pageId="17" pageNumber="1262">
<typeStatus id="B54C883BFF9CFFC2FCFF81940430F965" box="[827,880,1715,1736]" pageId="17" pageNumber="1262">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF9CFFC2FC1381940265F965" ID-CoL="3RLYX" authority="Stal, 1856" authorityName="Stal" authorityYear="1856" box="[983,1317,1714,1736]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="species" species="spurcus">
<emphasis id="5883EA8BFF9CFFC2FC13819403ECF965" box="[983,1196,1715,1736]" italics="true" pageId="17" pageNumber="1262">Laccocoris spurcus</emphasis>
<bibRefCitation id="0E664B68FF9CFFC2FB7781950265F965" author="Stal C" box="[1203,1317,1714,1736]" pageId="17" pageNumber="1262" pagination="193 - 199" refId="ref24930" refString="Stal C. 1856. Hemiptera samlade af Victorin i Caplandet. Ofversigt af Kongliga Svenska Vetenskaps-Akademiens Forhandlingar 13: 193 - 199." type="journal article" year="1856">Stål, 1856</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9CFFC0FCFF81D5056CF929" lastPageId="19" lastPageNumber="1264" pageId="17" pageNumber="1262" type="discussion">
<paragraph id="6A483699FF9CFFC1FCFF81D506D0FA25" blockId="17.[827,1443,1777,1891]" lastBlockId="18.[145,761,1241,1416]" lastPageId="18" lastPageNumber="1263" pageId="17" pageNumber="1262">
<emphasis id="5883EA8BFF9CFFC2FCFF81D50356F8AB" box="[827,1046,1777,1799]" italics="true" pageId="17" pageNumber="1262">Taxonomic history:</emphasis>
<bibRefCitation id="0E664B68FF9CFFC2FBE781D603E3F8AA" author="Stal C" box="[1059,1187,1777,1799]" pageId="17" pageNumber="1262" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">Stål (1876)</bibRefCitation>
proposed the division Laccocoraria to contain two genera,
<taxonomicName id="ADF74D1AFF9CFFC2FB378037022DF888" authorityName="Stal" authorityYear="1876" box="[1267,1389,1808,1829]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FB378037022DF888" box="[1267,1389,1808,1829]" italics="true" pageId="17" pageNumber="1262">Heleocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9CFFC2FCFF800804FAF8E9" authorityName="Stal" authorityYear="1856" box="[827,954,1839,1860]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FCFF800804FAF8E9" box="[827,954,1839,1860]" italics="true" pageId="17" pageNumber="1262">Laccocoris</emphasis>
</taxonomicName>
, that he described.
<bibRefCitation id="0E664B68FF9CFFC2FB70800802E3F8E8" author="Montandon AL" box="[1204,1443,1839,1861]" pageId="17" pageNumber="1262" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon (1897b)</bibRefCitation>
e l e v a t e d L a c c o c o r a r i a t o s u b f a m i l y s t a t u s a s
<taxonomicName id="ADF74D1AFF9FFFC1FF5583FE0673FB42" authorityName="STAL" authorityYear="1876" box="[145,307,1241,1263]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
.
<bibRefCitation id="0E664B68FF9FFFC1FE8583FE06A0FB43" author="Popov YA" box="[321,480,1241,1262]" pageId="18" pageNumber="1263" pagination="93 - 98" refId="ref23866" refString="Popov YA. 1970. Notes on the classification of the recent Naucoridae (Heteroptera: Nepomorpha). Bulletin of the Polish Academy of Sciences (Biological Sciences) 18: 93 - 98." type="journal article" year="1970">Popov (1970)</bibRefCitation>
considered this group as a tribe-level taxon,
<taxonomicName id="ADF74D1AFF9FFFC1FE5F83DF0563FAA3" authorityName="POPOV" authorityYear="1970" box="[411,547,1272,1294]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1266" phylum="Arthropoda" rank="tribe" tribe="Laccocorini">Laccocorini</taxonomicName>
, within subfamily
<taxonomicName id="ADF74D1AFF9FFFC1FF5582310656FA81" authorityName="LEACH" authorityYear="1815" box="[145,278,1302,1324]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
, but this classification scheme has not been followed by other specialists. Presently, ten genera are recognized in the subfamily, with no suprageneric organization as tribes.
</paragraph>
<caption id="3E886611FF9FFFC1FF5583060454FB00" ID-DOI="http://doi.org/10.5281/zenodo.6994641" ID-Zenodo-Dep="6994641" httpUri="https://zenodo.org/record/6994641/files/figure.png" pageId="18" pageNumber="1263" startId="18.[145,223,1057,1079]" targetBox="[149,1425,195,1017]" targetPageId="18">
<paragraph id="6A483699FF9FFFC1FF5583060454FB00" blockId="18.[145,1425,1057,1197]" pageId="18" pageNumber="1263">
<emphasis id="5883EA8BFF9FFFC1FF5583060645FB9B" bold="true" box="[145,261,1057,1079]" pageId="18" pageNumber="1263">Figure 13.</emphasis>
Detail of Most Probable Phylogeny dendogram of
<taxonomicName id="ADF74D1AFF9FFFC1FCCA830604D8FB9A" authorityName="STAL" authorityYear="1876" box="[782,920,1057,1079]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
. The red-coloured clade includes the Madagascar genera
<taxonomicName id="ADF74D1AFF9FFFC1FF1983180613FBF8" authorityName="Montandon" authorityYear="1897" box="[221,339,1087,1109]" class="Insecta" family="Naucoridae" genus="Temnocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FF1983180613FBF8" box="[221,339,1087,1109]" italics="true" pageId="18" pageNumber="1263">Temnocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9FFFC1FE40831806A1FBF8" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[388,481,1087,1109]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF9FFFC1FE40831806A1FBF8" box="[388,481,1087,1109]" italics="true" pageId="18" pageNumber="1263">Tsingala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF9FFFC1FE218318057EFBF8" box="[485,574,1087,1109]" pageId="18" pageNumber="1263" rank="genus">gen. nov.</taxonomicNameLabel>
(formerly
<taxonomicName id="ADF74D1AFF9FFFC1FD6383180452FBF8" authorityName="Stal" authorityYear="1876" box="[679,786,1087,1109]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FD6383180452FBF8" box="[679,786,1087,1109]" italics="true" pageId="18" pageNumber="1263">Heleocoris</emphasis>
</taxonomicName>
). The blue-coloured clade represents
<taxonomicName id="ADF74D1AFF9FFFC1FB4B8318026EFBF8" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1167,1326,1087,1109]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF9FFFC1FB4B8318026EFBF8" box="[1167,1326,1087,1109]" italics="true" pageId="18" pageNumber="1263">Heleolaccocoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF9FFFC1FAF7831802CEFBF8" box="[1331,1422,1087,1109]" pageId="18" pageNumber="1263" rank="genus">gen. nov.</taxonomicNameLabel>
, which comprises species of the former genera
<taxonomicName id="ADF74D1AFF9FFFC1FDAC837B0593FBDF" authorityName="Stal" authorityYear="1876" box="[616,723,1116,1138]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FDAC837B0593FBDF" box="[616,723,1116,1138]" italics="true" pageId="18" pageNumber="1263">Heleocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9FFFC1FCC1837B0432FBDF" authorityName="Stal" authorityYear="1856" box="[773,882,1116,1138]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FCC1837B0432FBDF" box="[773,882,1116,1138]" italics="true" pageId="18" pageNumber="1263">Laccocoris</emphasis>
</taxonomicName>
. All species of
<taxonomicName id="ADF74D1AFF9FFFC1FBC2837B0323FBDF" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1030,1123,1116,1138]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF9FFFC1FBC2837B0323FBDF" box="[1030,1123,1116,1138]" italics="true" pageId="18" pageNumber="1263">Tsingala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF9FFFC1FBAC837B0381FBDF" box="[1128,1217,1116,1138]" pageId="18" pageNumber="1263" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="ADF74D1AFF9FFFC1FB36837B02D1FBDF" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1266,1425,1116,1138]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF9FFFC1FB36837B02D1FBDF" box="[1266,1425,1116,1138]" italics="true" pageId="18" pageNumber="1263">Heleolaccocoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF9FFFC1FF55835E07AAFB22" box="[145,234,1145,1167]" pageId="18" pageNumber="1263" rank="genus">gen. nov.</taxonomicNameLabel>
are new combinations. Geographic associations for major clades within and among tribes are indicated. The red box in the inset shows the part of Figure 9 depicted in this figure.
</paragraph>
</caption>
<paragraph id="6A483699FF9FFFC1FF558296057CF8B5" blockId="18.[145,761,1457,1908]" pageId="18" pageNumber="1263">
<emphasis id="5883EA8BFF9FFFC1FF5582960628FA6A" box="[145,360,1457,1479]" italics="true" pageId="18" pageNumber="1263">Revised taxonomy:</emphasis>
<taxonomicName id="ADF74D1AFF9FFFC1FEB18296054CFA6A" authorityName="STAL" authorityYear="1876" box="[373,524,1457,1479]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
remains unchanged with respect to constituent species, although synonymies, new genera and tribes are proposed here. In my Most Probable Phylogeny dendrogram (
<figureCitation id="F2CC2A1CFF9FFFC1FD6D812A05AAF98F" box="[681,746,1549,1571]" captionStart="Figure 9" captionStartId="12.[145,223,1678,1700]" captionTargetBox="[308,1266,197,1636]" captionTargetId="figure-82@12.[302,1268,195,1638]" captionTargetPageId="12" captionText="Figure 9. Most Probable Phylogeny dendogram based on Bayesian inference with substitutions from maximum likelihood for Laccocorinae (except Ctenipocorini trib. nov. and Idiocarini trib. nov.). Closer views of Laccocorinae and Naucorinae individually can be seen in Figures 13 and 18, respectively." figureDoi="http://doi.org/10.5281/zenodo.6994629" httpUri="https://zenodo.org/record/6994629/files/figure.png" pageId="18" pageNumber="1263">Fig. 9</figureCitation>
), I substituted the ML topology of the clade sister to
<taxonomicName id="ADF74D1AFF9FFFC1FF55816C065EF9CD" authorityName="Montandon" authorityYear="1897" box="[145,286,1611,1632]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FF55816C065EF9CD" box="[145,286,1611,1632]" italics="true" pageId="18" pageNumber="1263">Ctenipocoris</emphasis>
</taxonomicName>
to represent what I consider to be the most plausible evolutionary relationships in the subfamily.
<taxonomicName id="ADF74D1AFF9FFFC1FF5581A0066DF930" authorityName="STAL" authorityYear="1876" box="[145,301,1671,1693]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
is represented by four distinct major clades, which are herein recognized as tribe-level taxa (
<figureCitation id="F2CC2A1CFF9FFFC1FF5D81E207A9F977" box="[153,233,1733,1755]" captionStart="Figure 13" captionStartId="18.[145,223,1057,1079]" captionTargetBox="[149,1425,195,1017]" captionTargetId="figure-432@18.[145,1425,195,1017]" captionTargetPageId="18" captionText="Figure 13. Detail of Most Probable Phylogeny dendogram of Laccocorinae.The red-coloured clade includes the Madagascar genera Temnocoris and Tsingala gen. nov.(formerly Heleocoris). The blue-coloured clade represents Heleolaccocoris gen.nov., which comprises species of the former genera Heleocoris and Laccocoris. All species of Tsingala gen. nov. and Heleolaccocoris gen. nov. are new combinations. Geographic associations for major clades within and among tribes are indicated. The red box in the inset shows the part of Figure 9 depicted in this figure." figureDoi="http://doi.org/10.5281/zenodo.6994641" httpUri="https://zenodo.org/record/6994641/files/figure.png" pageId="18" pageNumber="1263">Fig. 13</figureCitation>
). Two of the predominant genera (
<taxonomicName id="ADF74D1AFF9FFFC1FD4581E205B9F977" authorityName="Stal" authorityYear="1876" box="[641,761,1733,1754]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FD4581E205B9F977" box="[641,761,1733,1754]" italics="true" pageId="18" pageNumber="1263">Heleocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9FFFC1FF0781C3067AF954" authorityName="Stal" authorityYear="1856" box="[195,314,1764,1785]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FF0781C3067AF954" box="[195,314,1764,1785]" italics="true" pageId="18" pageNumber="1263">Laccocoris</emphasis>
</taxonomicName>
) in the subfamily are polyphyletic and occur in multiple clades in two tribes.
</paragraph>
<paragraph id="6A483699FF9FFFC0FF6D800606C7FC09" blockId="18.[145,761,1457,1908]" lastBlockId="19.[163,780,880,1668]" lastPageId="19" lastPageNumber="1264" pageId="18" pageNumber="1263">
Species of
<taxonomicName id="ADF74D1AFF9FFFC1FEE4800606D6F89B" authorityName="Stal" authorityYear="1876" box="[288,406,1825,1846]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FEE4800606D6F89B" box="[288,406,1825,1846]" italics="true" pageId="18" pageNumber="1263">Heleocoris</emphasis>
</taxonomicName>
occur in three disparate clades (
<figureCitation id="F2CC2A1CFF9FFFC1FF5D801807ACF8F8" box="[153,236,1855,1877]" captionStart="Figure 13" captionStartId="18.[145,223,1057,1079]" captionTargetBox="[149,1425,195,1017]" captionTargetId="figure-432@18.[145,1425,195,1017]" captionTargetPageId="18" captionText="Figure 13. Detail of Most Probable Phylogeny dendogram of Laccocorinae.The red-coloured clade includes the Madagascar genera Temnocoris and Tsingala gen. nov.(formerly Heleocoris). The blue-coloured clade represents Heleolaccocoris gen.nov., which comprises species of the former genera Heleocoris and Laccocoris. All species of Tsingala gen. nov. and Heleolaccocoris gen. nov. are new combinations. Geographic associations for major clades within and among tribes are indicated. The red box in the inset shows the part of Figure 9 depicted in this figure." figureDoi="http://doi.org/10.5281/zenodo.6994641" httpUri="https://zenodo.org/record/6994641/files/figure.png" pageId="18" pageNumber="1263">Figs 13</figureCitation>
,
<figureCitation id="F2CC2A1CFF9FFFC1FF3C80670654F8F8" box="[248,276,1856,1877]" captionStart="Figure 14" captionStartId="19.[163,244,765,787]" captionTargetBox="[163,1442,195,726]" captionTargetId="figure-559@19.[163,1442,195,726]" captionTargetPageId="19" captionText="Figure 14. Revised generic representation of the former genus Heleocoris. Dorsal habitus of Heleocoris vicinus (a), Laccocoris minisculus (b), Heleolaccocoris ovatus (c), Tsingala humeralis (d)." figureDoi="http://doi.org/10.5281/zenodo.6994645" httpUri="https://zenodo.org/record/6994645/files/figure.png" pageId="18" pageNumber="1263">14</figureCitation>
). The
<typeStatus id="B54C883BFF9FFFC1FE9E806706CCF8F8" box="[346,396,1856,1877]" pageId="18" pageNumber="1263">type</typeStatus>
species is
<taxonomicName id="ADF74D1AFF9FFFC1FDC08018060DF8DE" authority="(Spinola, 1837)" baseAuthorityName="Spinola" baseAuthorityYear="1837" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="species" species="obliquatus">
<emphasis id="5883EA8BFF9FFFC1FDC0801805B8F8F9" box="[516,760,1855,1876]" italics="true" pageId="18" pageNumber="1263">Heleocoris obliquatus</emphasis>
(
<bibRefCitation id="0E664B68FF9FFFC1FF5E80790603F8DE" author="Spinola M" box="[154,323,1886,1908]" pageId="18" pageNumber="1263" refId="ref24867" refString="Spinola M. 1837. Essai sur les genres d'insectes appartenants a l'ordre des Hemipteres, Lin. ou Rhyngotes, Fab. et a la section des Heteropteres, Dufour. Chez Yves Graviers, Genes, 383 pp." type="book" year="1837">Spinola, 1837</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="12E07609FF9FFFC1FE5F807906BDF8D9" box="[411,509,1886,1908]" name="India" pageId="18" pageNumber="1263">Bombay</collectingCountry>
(
<collectingCountry id="12E07609FF9FFFC1FDD480790539F8D9" box="[528,633,1886,1908]" name="India" pageId="18" pageNumber="1263">Mumbai</collectingCountry>
,
<collectingCountry id="12E07609FF9FFFC1FD4D8079058BF8D9" box="[649,715,1886,1908]" name="India" pageId="18" pageNumber="1263">India</collectingCountry>
) as designated by
<bibRefCitation id="0E664B68FF9FFFC1FC1583FF0311FB42" author="Stal C" box="[977,1105,1240,1263]" pageId="18" pageNumber="1263" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">Stål (1876)</bibRefCitation>
, who based his designation on a specimen in the collection of Signoret.
<bibRefCitation id="0E664B68FF9FFFC1FAC983D0043FFA81" author="Montandon AL" pageId="18" pageNumber="1263" pagination="652 - 658" refId="ref23277" refString="Montandon AL. 1910 b. Especes nouvelles ou peu connues d'Hydrococrises de l'Inde. Bulletin de la Societe des Sciences de Bucarest, Roumanie 19: 652 - 658." type="journal article" year="1910">Montandon (1910b)</bibRefCitation>
questioned the existence of the
<typeStatus id="B54C883BFF9FFFC1FB2C8230025AFA81" box="[1256,1306,1303,1324]" pageId="18" pageNumber="1263">type</typeStatus>
specimen, which was based on a brief original description, and provided a redescription of the species based on a specimen in Distant’s collection from the Dawna Hills of eastern
<collectingCountry id="12E07609FF9FFFC1FC2982B60302FA0B" box="[1005,1090,1425,1446]" name="Myanmar" pageId="18" pageNumber="1263">Burma</collectingCountry>
. Given that most species in these groups are restricted to particular geographic regions, it is highly doubtful that the specimen upon which Montandon redescribed
<taxonomicName id="ADF74D1AFF9FFFC1FB5E82CA027BF9AF" baseAuthorityName="Spinola" baseAuthorityYear="1837" box="[1178,1339,1517,1538]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="species" species="obliquatus">
<emphasis id="5883EA8BFF9FFFC1FB5E82CA027BF9AF" box="[1178,1339,1517,1538]" italics="true" pageId="18" pageNumber="1263">H. obliquatus</emphasis>
</taxonomicName>
was in fact that species. Further vitiating this identification is the figure in
<bibRefCitation id="0E664B68FF9FFFC1FC2A810D039EF992" author="Distant WL" box="[1006,1246,1578,1600]" pageId="18" pageNumber="1263" refId="ref21795" refString="Distant WL. 1910. The fauna of British India, including Ceylon and Burma. Rhynchota. Vol. V. Heteroptera: Appendix. London: Taylor &amp; Francis." type="book" year="1910">Distant (1910: 322)</bibRefCitation>
of a specimen from ‘Lower Burma’ identified by Montandon as
<taxonomicName id="ADF74D1AFF9FFFC1FCED814F0490F9D1" baseAuthorityName="Spinola" baseAuthorityYear="1837" box="[809,976,1639,1661]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="species" species="obliquatus">
<emphasis id="5883EA8BFF9FFFC1FCED814F0490F9D1" box="[809,976,1639,1661]" italics="true" pageId="18" pageNumber="1263">H. obliquatus</emphasis>
</taxonomicName>
, which instead clearly is
<taxonomicName id="ADF74D1AFF9FFFC1FAD78140032EF931" authority="Montandon, 1897 b" authorityName="Montandon" authorityYear="1897" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="species" species="strabus">
<emphasis id="5883EA8BFF9FFFC1FAD7814004C3F936" italics="true" pageId="18" pageNumber="1263">Heleocoris strabus</emphasis>
<bibRefCitation id="0E664B68FF9FFFC1FC4881A1032EF931" author="Montandon AL" box="[908,1134,1670,1692]" pageId="18" pageNumber="1263" pagination="435 - 454" refId="ref23009" refString="Montandon AL. 1897 b. Hemiptera Cryptocerata. Fam. Naucoridae. Sous fam. Laccocorinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 435 - 454." type="journal article" year="1897">Montandon, 1897b</bibRefCitation>
</taxonomicName>
. As such,
<taxonomicName id="ADF74D1AFF9FFFC1FB2881A002D1F936" baseAuthorityName="Spinola" baseAuthorityYear="1837" box="[1260,1425,1670,1692]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="species" species="obliquatus">
<emphasis id="5883EA8BFF9FFFC1FB2881A002D1F936" box="[1260,1425,1670,1692]" italics="true" pageId="18" pageNumber="1263">H. obliquatus</emphasis>
</taxonomicName>
from
<collectingCountry id="12E07609FF9FFFC1FCB1818204A0F916" box="[885,992,1701,1723]" name="India" pageId="18" pageNumber="1263">Mumbai</collectingCountry>
remains enigmatic and nearly impossible to diagnose without an authoritatively identified specimen, and thus is a
<emphasis id="5883EA8BFF9FFFC1FB7181C402CBF95A" box="[1205,1419,1762,1784]" italics="true" pageId="18" pageNumber="1263">species inquirenda</emphasis>
. Nonetheless, because the
<typeStatus id="B54C883BFF9FFFC1FB99802603D0F8BB" box="[1117,1168,1793,1814]" pageId="18" pageNumber="1263">type</typeStatus>
locality was given as ‘Bombay’, I here consider the clade containing species of
<taxonomicName id="ADF74D1AFF9FFFC1FC87801904F9F8FE" authorityName="Stal" authorityYear="1876" box="[835,953,1854,1875]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FC87801904F9F8FE" box="[835,953,1854,1875]" italics="true" pageId="18" pageNumber="1263">Heleocoris</emphasis>
</taxonomicName>
from central
<collectingCountry id="12E07609FF9FFFC1FB8B801903CCF8F9" box="[1103,1164,1854,1876]" name="India" pageId="18" pageNumber="1263">India</collectingCountry>
(all but the far eastern states) to be valid
<taxonomicName id="ADF74D1AFF9FFFC1FC38807A0332F8DF" authorityName="Stal" authorityYear="1876" box="[1020,1138,1885,1906]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="1263" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9FFFC1FC38807A0332F8DF" box="[1020,1138,1885,1906]" italics="true" pageId="18" pageNumber="1263">Heleocoris</emphasis>
</taxonomicName>
. Other clades containing
<taxonomicName id="ADF74D1AFF9EFFC0FF6784570659FC28" authorityName="Stal" authorityYear="1876" box="[163,281,880,901]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FF6784570659FC28" box="[163,281,880,901]" italics="true" pageId="19" pageNumber="1264">Heleocoris</emphasis>
</taxonomicName>
from Africa and Indochina each must have the genus renamed.
</paragraph>
<caption id="3E886611FF9EFFC0FF6785DA0480FC9C" ID-DOI="http://doi.org/10.5281/zenodo.6994645" ID-Zenodo-Dep="6994645" httpUri="https://zenodo.org/record/6994645/files/figure.png" pageId="19" pageNumber="1264" startId="19.[163,244,765,787]" targetBox="[163,1442,195,726]" targetPageId="19">
<paragraph id="6A483699FF9EFFC0FF6785DA0480FC9C" blockId="19.[163,1443,765,817]" pageId="19" pageNumber="1264">
<emphasis id="5883EA8BFF9EFFC0FF6785DA065EFCBF" bold="true" box="[163,286,765,787]" pageId="19" pageNumber="1264">Figure 14.</emphasis>
Revised generic representation of the former genus
<taxonomicName id="ADF74D1AFF9EFFC0FCAF85D9049BFCB9" authorityName="Stal" authorityYear="1876" box="[875,987,766,788]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FCAF85D9049BFCB9" box="[875,987,766,788]" italics="true" pageId="19" pageNumber="1264">Heleocoris</emphasis>
</taxonomicName>
. Dorsal habitus of
<taxonomicName id="ADF74D1AFF9EFFC0FB7585D90237FCB9" authorityName="Montandon" authorityYear="1910" box="[1201,1399,766,788]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="5883EA8BFF9EFFC0FB7585D90237FCB9" box="[1201,1399,766,788]" italics="true" pageId="19" pageNumber="1264">Heleocoris vicinus</emphasis>
</taxonomicName>
(a),
<taxonomicName id="ADF74D1AFF9EFFC0FF67843C06C9FC9C" box="[163,393,795,817]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="species" species="minisculus">
<emphasis id="5883EA8BFF9EFFC0FF67843C06C9FC9C" box="[163,393,795,817]" italics="true" pageId="19" pageNumber="1264">Laccocoris minisculus</emphasis>
</taxonomicName>
(b),
<taxonomicName id="ADF74D1AFF9EFFC0FE72843C05DFFC9C" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[438,671,795,817]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="5883EA8BFF9EFFC0FE72843C05DFFC9C" box="[438,671,795,817]" italics="true" pageId="19" pageNumber="1264">Heleolaccocoris ovatus</emphasis>
</taxonomicName>
(c),
<taxonomicName id="ADF74D1AFF9EFFC0FD0E843C04D7FC9C" authorityName="Sites" authorityYear="2022" baseAuthorityName="Signoret" baseAuthorityYear="1860" box="[714,919,795,817]" class="Insecta" family="Naucoridae" genus="Tsingala" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="5883EA8BFF9EFFC0FD0E843C04D7FC9C" box="[714,919,795,817]" italics="true" pageId="19" pageNumber="1264">Tsingala humeralis</emphasis>
</taxonomicName>
(d).
</paragraph>
</caption>
<paragraph id="6A483699FF9EFFC0FF7F848A056CF929" blockId="19.[163,780,880,1668]" pageId="19" pageNumber="1264">
Species of
<taxonomicName id="ADF74D1AFF9EFFC0FEF0848906EAFC6E" authorityName="Stal" authorityYear="1856" box="[308,426,942,963]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FEF0848906EAFC6E" box="[308,426,942,963]" italics="true" pageId="19" pageNumber="1264">Laccocoris</emphasis>
</taxonomicName>
were recovered in two distinct clades within different tribes (
<figureCitation id="F2CC2A1CFF9EFFC0FDCF84EB0522FC4C" box="[523,610,972,994]" captionStart="Figure 13" captionStartId="18.[145,223,1057,1079]" captionTargetBox="[149,1425,195,1017]" captionTargetId="figure-432@18.[145,1425,195,1017]" captionTargetPageId="18" captionText="Figure 13. Detail of Most Probable Phylogeny dendogram of Laccocorinae.The red-coloured clade includes the Madagascar genera Temnocoris and Tsingala gen. nov.(formerly Heleocoris). The blue-coloured clade represents Heleolaccocoris gen.nov., which comprises species of the former genera Heleocoris and Laccocoris. All species of Tsingala gen. nov. and Heleolaccocoris gen. nov. are new combinations. Geographic associations for major clades within and among tribes are indicated. The red box in the inset shows the part of Figure 9 depicted in this figure." figureDoi="http://doi.org/10.5281/zenodo.6994641" httpUri="https://zenodo.org/record/6994641/files/figure.png" pageId="19" pageNumber="1264">Figs 13</figureCitation>
,
<figureCitation id="F2CC2A1CFF9EFFC0FDB584EB05CDFC4F" box="[625,653,972,994]" captionStart="Figure 15" captionStartId="20.[145,226,1055,1077]" captionTargetBox="[145,1425,195,1015]" captionTargetId="figure-409@20.[145,1425,195,1015]" captionTargetPageId="20" captionText="Figure 15. Revised generic representation of the former genus Laccocoris. Dorsal habitus of Laccocoris spurcus (a), Heleolaccocoris horvathi (b)." figureDoi="http://doi.org/10.5281/zenodo.6994647" httpUri="https://zenodo.org/record/6994647/files/figure.png" pageId="19" pageNumber="1264">15</figureCitation>
). The
<typeStatus id="B54C883BFF9EFFC0FD1C84EB044AFC4C" box="[728,778,972,993]" pageId="19" pageNumber="1264">type</typeStatus>
species is
<taxonomicName id="ADF74D1AFF9EFFC0FED384CC053AFBAD" authority="(Stal, 1856)" baseAuthorityName="Stal" baseAuthorityYear="1856" box="[279,634,1002,1024]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="species" species="spurcus">
<emphasis id="5883EA8BFF9EFFC0FED384CC06AFFBAD" box="[279,495,1003,1024]" italics="true" pageId="19" pageNumber="1264">Laccocoris spurcus</emphasis>
(
<bibRefCitation id="0E664B68FF9EFFC0FE3A84CD0532FBAD" author="Stal C" box="[510,626,1002,1024]" pageId="19" pageNumber="1264" pagination="193 - 199" refId="ref24930" refString="Stal C. 1856. Hemiptera samlade af Victorin i Caplandet. Ofversigt af Kongliga Svenska Vetenskaps-Akademiens Forhandlingar 13: 193 - 199." type="journal article" year="1856">Stål, 1856</bibRefCitation>
)
</taxonomicName>
from Africa; thus, the species of
<taxonomicName id="ADF74D1AFF9EFFC0FE5D832D0556FBB2" authorityName="Stal" authorityYear="1856" box="[409,534,1034,1055]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FE5D832D0556FBB2" box="[409,534,1034,1055]" italics="true" pageId="19" pageNumber="1264">Laccocoris</emphasis>
</taxonomicName>
from Africa remain unchanged. However, the species of
<taxonomicName id="ADF74D1AFF9EFFC0FD94830F058AFB90" authorityName="Stal" authorityYear="1856" box="[592,714,1064,1085]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FD94830F058AFB90" box="[592,714,1064,1085]" italics="true" pageId="19" pageNumber="1264">Laccocoris</emphasis>
</taxonomicName>
from Sundaland and the
<collectingCountry id="12E07609FF9EFFC0FEB1836106B3FBF1" box="[373,499,1094,1116]" name="Philippines" pageId="19" pageNumber="1264">Philippines</collectingCountry>
are here placed in a newly erected genus,
<taxonomicName id="ADF74D1AFF9EFFC0FE9783420545FBD7" authority="Because" authorityName="Sites" authorityYear="2022" box="[339,517,1125,1146]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5883EA8BFF9EFFC0FE9783420545FBD7" box="[339,517,1125,1146]" italics="true" pageId="19" pageNumber="1264">Heleolaccocoris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FF9EFFC0FDCA83410534FBD6" box="[526,628,1126,1147]" pageId="19" pageNumber="1264" rank="genus">gen. nov.</taxonomicNameLabel>
Because the well-supported ML topology recovered
<taxonomicName id="ADF74D1AFF9EFFC0FD9A83A3044BFB34" authorityName="Sites" authorityYear="2022" box="[606,779,1156,1177]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FD9A83A3044BFB34" box="[606,779,1156,1177]" italics="true" pageId="19" pageNumber="1264">Heleolaccocoris</emphasis>
</taxonomicName>
as sister to three species of Indochinese
<taxonomicName id="ADF74D1AFF9EFFC0FDB2838505ACFB1A" authorityName="Stal" authorityYear="1876" box="[630,748,1186,1207]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FDB2838505ACFB1A" box="[630,748,1186,1207]" italics="true" pageId="19" pageNumber="1264">Heleocoris</emphasis>
</taxonomicName>
in successively more inclusive clades, this necessitated either designation of additional genera to accommodate the clades of Indochinese
<taxonomicName id="ADF74D1AFF9EFFC0FE0B83D90505FABE" authorityName="Stal" authorityYear="1876" box="[463,581,1278,1299]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FE0B83D90505FABE" box="[463,581,1278,1299]" italics="true" pageId="19" pageNumber="1264">Heleocoris</emphasis>
</taxonomicName>
, or placing them all in
<taxonomicName id="ADF74D1AFF9EFFC0FF27823A06CCFA9F" authorityName="Sites" authorityYear="2022" box="[227,396,1309,1330]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FF27823A06CCFA9F" box="[227,396,1309,1330]" italics="true" pageId="19" pageNumber="1264">Heleolaccocoris</emphasis>
</taxonomicName>
as I have done here (
<figureCitation id="F2CC2A1CFF9EFFC0FDBF823A0586FA9F" box="[635,710,1309,1331]" captionStart="Figure 13" captionStartId="18.[145,223,1057,1079]" captionTargetBox="[149,1425,195,1017]" captionTargetId="figure-432@18.[145,1425,195,1017]" captionTargetPageId="18" captionText="Figure 13. Detail of Most Probable Phylogeny dendogram of Laccocorinae.The red-coloured clade includes the Madagascar genera Temnocoris and Tsingala gen. nov.(formerly Heleocoris). The blue-coloured clade represents Heleolaccocoris gen.nov., which comprises species of the former genera Heleocoris and Laccocoris. All species of Tsingala gen. nov. and Heleolaccocoris gen. nov. are new combinations. Geographic associations for major clades within and among tribes are indicated. The red box in the inset shows the part of Figure 9 depicted in this figure." figureDoi="http://doi.org/10.5281/zenodo.6994641" httpUri="https://zenodo.org/record/6994641/files/figure.png" pageId="19" pageNumber="1264">Fig. 13</figureCitation>
: bluecoloured clades). The previously recognized distinction between the two genera (
<taxonomicName id="ADF74D1AFF9EFFC0FE15827D0509FAC2" authorityName="Stal" authorityYear="1876" box="[465,585,1370,1391]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FE15827D0509FAC2" box="[465,585,1370,1391]" italics="true" pageId="19" pageNumber="1264">Heleocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9EFFC0FD43827C0441FADD" authorityName="Stal" authorityYear="1856" box="[647,769,1371,1392]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FD43827C0441FADD" box="[647,769,1371,1392]" italics="true" pageId="19" pageNumber="1264">Laccocoris</emphasis>
</taxonomicName>
) was based on the shape of the labrum as wider than long with a rounded apex or longer than wide with an acute apex. This has been generally considered a feature inconsistent with phylogeny by recent scientists who have studied the group. In a treatment of the
<taxonomicName id="ADF74D1AFF9EFFC0FD4F82D3044BF9A7" authorityName="LEACH" authorityYear="1815" box="[651,779,1524,1546]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="family">Naucoridae</taxonomicName>
in
<collectingCountry id="12E07609FF9EFFC0FF0481350673F985" box="[192,307,1554,1576]" name="Singapore" pageId="19" pageNumber="1264">Singapore</collectingCountry>
,
<bibRefCitation id="0E664B68FF9EFFC0FEFB813505CBF98A" author="Polhemus DA &amp; Polhemus JT" box="[319,651,1554,1576]" pageId="19" pageNumber="1264" pagination="665 - 686" refId="ref23727" refString="Polhemus DA, Polhemus JT. 2013. Guide to the aquatic Heteroptera of Singapore and Peninsular Malaysia. XI. Infraorder Nepomorpha - families Naucoridae and Aphelocheiridae. Raffles Bulletin of Zoology 61: 665 - 686." type="journal article" year="2013">Polhemus &amp; Polhemus (2013)</bibRefCitation>
considered the generic limits of
<taxonomicName id="ADF74D1AFF9EFFC0FE5E81160552F9EB" authorityName="Stal" authorityYear="1876" box="[410,530,1585,1606]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FE5E81160552F9EB" box="[410,530,1585,1606]" italics="true" pageId="19" pageNumber="1264">Heleocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9EFFC0FD8B81150589F9EA" authorityName="Stal" authorityYear="1856" box="[591,713,1586,1607]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FD8B81150589F9EA" box="[591,713,1586,1607]" italics="true" pageId="19" pageNumber="1264">Laccocoris</emphasis>
</taxonomicName>
to be indistinct and correctly predicted the possible synonymy of Asian
<taxonomicName id="ADF74D1AFF9EFFC0FEC681480635F929" authorityName="Stal" authorityYear="1856" box="[258,373,1647,1668]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FEC681480635F929" box="[258,373,1647,1668]" italics="true" pageId="19" pageNumber="1264">Laccocoris</emphasis>
</taxonomicName>
with
<taxonomicName id="ADF74D1AFF9EFFC0FE7081490565F92E" authorityName="Stal" authorityYear="1876" box="[436,549,1646,1667]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FE7081490565F92E" box="[436,549,1646,1667]" italics="true" pageId="19" pageNumber="1264">Heleocoris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9EFFC0FF6781940228FB34" pageId="19" pageNumber="1264" type="diagnosis">
<paragraph id="6A483699FF9EFFC0FF6781940228FB34" blockId="19.[163,780,1714,1890]" lastBlockId="19.[827,1443,880,1178]" pageId="19" pageNumber="1264">
<emphasis id="5883EA8BFF9EFFC0FF6781940663F965" box="[163,291,1715,1736]" italics="true" pageId="19" pageNumber="1264">Diagnosis:</emphasis>
This subfamily has a suite of unique diagnostic attributes, some of which exhibit variation among genera. The front of the head is folded posteroventrally such that the labrum is set back from the functional anterior margin of the head (
<figureCitation id="F2CC2A1CFF9EFFC0FD6F800A05BBF8EE" box="[683,763,1837,1859]" captionStart="Figure 3" captionStartId="8.[145,230,1468,1490]" captionTargetBox="[172,733,195,1428]" captionTargetId="figure-532@8.[172,733,195,1428]" captionTargetPageId="8" captionText="Figure 3. Mouthpart positions: recessed rostrum of Cheirochela thailandana (a); lateral views of Laccocoris spurcus (b) and Australambrysus crenulatus (c). White arrows indicate positions of labrum with respect to the functional anterior margin of the head." figureDoi="http://doi.org/10.5281/zenodo.6994613" httpUri="https://zenodo.org/record/6994613/files/figure.png" pageId="19" pageNumber="1264">Fig. 3b</figureCitation>
). Males of all genera have a dense tomentose pad ventrally over much of the mesotibia and tarsomeres 2 and 3 (
<figureCitation id="F2CC2A1CFF9EFFC0FC8784A904CEFC0E" box="[835,910,910,932]" captionStart="Figure 12" captionStartId="17.[163,243,870,892]" captionTargetBox="[163,1443,195,830]" captionTargetId="figure-534@17.[163,1443,195,830]" captionTargetPageId="17" captionText="Figure 12. Mesotibia and mesotarsus of male and female Heleolaccocoris ovatus showing a sexually dimorphic ventral tomentose pad of hairs." figureDoi="http://doi.org/10.5281/zenodo.6994637" httpUri="https://zenodo.org/record/6994637/files/figure.png" pageId="19" pageNumber="1264">Fig. 12</figureCitation>
), and to a lesser extent the protibia and tarsus, whereas in females the pads are greatly reduced. The prothoracic pretarsal claws are paired and articulated, except in
<taxonomicName id="ADF74D1AFF9EFFC0FC6884CD030BFC52" authorityName="Sites" authorityYear="2007" box="[940,1099,1002,1023]" class="Insecta" family="Naucoridae" genus="Namtokocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FC6884CD030BFC52" box="[940,1099,1002,1023]" italics="true" pageId="19" pageNumber="1264">Namtokocoris</emphasis>
</taxonomicName>
in which they are single and fused with the tarsus, which is the condition found in all other subfamilies (
<figureCitation id="F2CC2A1CFF9EFFC0FBD4830F031AFB90" box="[1040,1114,1064,1086]" captionStart="Figure 4" captionStartId="9.[163,249,1379,1401]" captionTargetBox="[163,781,195,1339]" captionTargetId="figure-575@9.[163,781,195,1339]" captionTargetPageId="9" captionText="Figure 4. Foretibiae and foretarsi of Laccocorinae: Ctenipocoris asiaticus (a), Heleolaccocoris ovatus (b), Namtokocoris dalanta (c)." figureDoi="http://doi.org/10.5281/zenodo.6994615" httpUri="https://zenodo.org/record/6994615/files/figure.png" pageId="19" pageNumber="1264">Fig. 4c</figureCitation>
). Males have two-segmented front tarsi and females only one, except in
<taxonomicName id="ADF74D1AFF9EFFC0FADC836002E2FBF1" authorityName="Montandon" authorityYear="1897" box="[1304,1442,1095,1116]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FADC836002E2FBF1" box="[1304,1442,1095,1116]" italics="true" pageId="19" pageNumber="1264">Ctenipocoris</emphasis>
</taxonomicName>
where both sexes have two, and in
<taxonomicName id="ADF74D1AFF9EFFC0FB158342022FFBD7" authorityName="Sites" authorityYear="2007" box="[1233,1391,1125,1146]" class="Insecta" family="Naucoridae" genus="Namtokocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FB158342022FFBD7" box="[1233,1391,1125,1146]" italics="true" pageId="19" pageNumber="1264">Namtokocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9EFFC0FCFF83A304A4FB34" authorityName="Sites &amp; Zettel" authorityYear="2011" box="[827,996,1156,1177]" class="Insecta" family="Naucoridae" genus="Pogonocaudina" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FCFF83A304A4FB34" box="[827,996,1156,1177]" italics="true" pageId="19" pageNumber="1264">Pogonocaudina</emphasis>
</taxonomicName>
where both sexes have one (
<figureCitation id="F2CC2A1CFF9EFFC0FAD983A3021AFB34" box="[1309,1370,1156,1178]" captionStart="Figure 4" captionStartId="9.[163,249,1379,1401]" captionTargetBox="[163,781,195,1339]" captionTargetId="figure-575@9.[163,781,195,1339]" captionTargetPageId="9" captionText="Figure 4. Foretibiae and foretarsi of Laccocorinae: Ctenipocoris asiaticus (a), Heleolaccocoris ovatus (b), Namtokocoris dalanta (c)." figureDoi="http://doi.org/10.5281/zenodo.6994615" httpUri="https://zenodo.org/record/6994615/files/figure.png" pageId="19" pageNumber="1264">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9EFFC0FCFF83920482FAC9" pageId="19" pageNumber="1264" type="discussion">
<paragraph id="6A483699FF9EFFC0FCFF83920482FAC9" blockId="19.[827,1443,1205,1380]" pageId="19" pageNumber="1264">
<emphasis id="5883EA8BFF9EFFC0FCFF83920486FB67" box="[827,966,1205,1226]" italics="true" pageId="19" pageNumber="1264">Comments:</emphasis>
This subfamily is circumtropical in distribution and is most common in Africa and Asia. It is not known from
<collectingCountry id="12E07609FF9EFFC0FBD483D5033FFAA5" box="[1040,1151,1266,1288]" name="Australia" pageId="19" pageNumber="1264">Australia</collectingCountry>
. Although
<taxonomicName id="ADF74D1AFF9EFFC0FB3F83D402C9FAA5" authorityName="Montandon" authorityYear="1897" box="[1275,1417,1267,1288]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9EFFC0FB3F83D402C9FAA5" box="[1275,1417,1267,1288]" italics="true" pageId="19" pageNumber="1264">Ctenipocoris</emphasis>
</taxonomicName>
is circumtropical in distribution, each of the other major clades in the subfamily has a distinct geographic association.
</paragraph>
</subSubSection>
</treatment>
</document>

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SUBFAMILY
<taxonomicName id="ADF74D1AFF9DFFC3FBCA812203EFF9B6" authority="MONTANDON, 1897" authorityName="Sites" authorityYear="2022" box="[1038,1199,1541,1565]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="subFamily" status="stat. nov." subFamily="Ilyocorinae">ILYOCORINAE</taxonomicName>
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<taxonomicNameLabel id="43B057F0FF9DFFC3FB73812F0273F9B0" box="[1207,1331,1541,1565]" pageId="16" pageNumber="1261">STAT. REV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FC35811602C2F916" pageId="16" pageNumber="1261" type="description">
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<figureCitation id="F2CC2A1CFF9DFFC3FC358116037BF9E7" box="[1009,1083,1585,1610]" captionStart="Figure 7" captionStartId="11.[163,241,696,718]" captionTargetBox="[399,1207,195,656]" captionTargetId="figure-197@11.[399,1207,195,656]" captionTargetPageId="11" captionText="Figure 7. Valvulae of female Ilyocoris cimicoides with dentate margins. vlf = valvifer, vlv = valvula." figureDoi="http://doi.org/10.5281/zenodo.6994623" httpUri="https://zenodo.org/record/6994623/files/figure.png" pageId="16" pageNumber="1261">FIGS 7</figureCitation>
,
<figureCitation id="F2CC2A1CFF9DFFC3FB8D81160317F9E7" box="[1097,1111,1585,1610]" captionStart="Figure 8" captionStartId="11.[163,242,1486,1508]" captionTargetBox="[163,1443,800,1447]" captionTargetId="figure-171@11.[163,1443,800,1447]" captionTargetPageId="11" captionText="Figure 8. Mesofemora of Ilyocoris cimicoides (a) and Placomerus sp. (b) showing the posterodorsal margin flattened and fringed with a dense brush-line of hairs." figureDoi="http://doi.org/10.5281/zenodo.6994627" httpUri="https://zenodo.org/record/6994627/files/figure.png" pageId="16" pageNumber="1261">8</figureCitation>
,
<figureCitation id="F2CC2A1CFF9DFFC3FBA0811603D0F9E5" box="[1124,1168,1585,1610]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="16" pageNumber="1261">10D</figureCitation>
,
<figureCitation id="F2CC2A1CFF9DFFC3FB5981160389F9E5" box="[1181,1225,1585,1610]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="16" pageNumber="1261">11D</figureCitation>
</paragraph>
<paragraph id="6A483699FF9DFFC3FCED81A002C2F916" blockId="16.[809,1426,1670,1723]" pageId="16" pageNumber="1261">
<emphasis id="5883EA8BFF9DFFC3FCED81A00337F931" box="[809,1143,1670,1692]" italics="true" pageId="16" pageNumber="1261">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF9DFFC3FCB2818202C2F916" box="[886,1410,1701,1723]" pageId="16" pageNumber="1261">CBCF3B3B-1324-486F-BD2C-95230D825861</uuid>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FCED81C204A8F8B5" pageId="16" pageNumber="1261" type="type_taxon">
<paragraph id="6A483699FF9DFFC3FCED81C204A8F8B5" blockId="16.[809,1425,1764,1816]" pageId="16" pageNumber="1261">
<emphasis id="5883EA8BFF9DFFC3FCED81C204F9F954" box="[809,953,1764,1786]" italics="true" pageId="16" pageNumber="1261">
<typeStatus id="B54C883BFF9DFFC3FCED81C2041EF957" box="[809,862,1765,1786]" pageId="16" pageNumber="1261">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF9DFFC3FC0281C3022BF954" ID-CoL="3PH9Z" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[966,1387,1764,1786]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="species" species="cimicoides">
<emphasis id="5883EA8BFF9DFFC3FC0281C303E4F954" box="[966,1188,1764,1785]" italics="true" pageId="16" pageNumber="1261">Ilyocoris cimicoides</emphasis>
(
<bibRefCitation id="0E664B68FF9DFFC3FB7781C30222F957" author="Linnaeus C" box="[1203,1378,1764,1786]" pageId="16" pageNumber="1261" refId="ref22650" refString="Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Stockholm: Salvius." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
, as
<taxonomicName id="ADF74D1AFF9DFFC3FCED802404A1F8BA" authorityName="Linnaeus" authorityYear="1758" box="[809,993,1794,1816]" class="Insecta" family="Nepidae" genus="Nepa" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="species" species="cimicoides">
<emphasis id="5883EA8BFF9DFFC3FCED802404A1F8BA" box="[809,993,1794,1816]" italics="true" pageId="16" pageNumber="1261">Nepa cimicoides</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC2FCED80650668F8A9" lastPageId="17" lastPageNumber="1262" pageId="16" pageNumber="1261" type="discussion">
<paragraph id="6A483699FF9DFFC3FCED806502CFF8DB" blockId="16.[809,1424,1857,1910]" pageId="16" pageNumber="1261">
<heading id="310081F5FF9DFFC3FCED806502CFF8DB" centered="true" fontSize="9" level="2" pageId="16" pageNumber="1261" reason="2">
<emphasis id="5883EA8BFF9DFFC3FCED80650347F8FB" box="[809,1031,1857,1879]" italics="true" pageId="16" pageNumber="1261">Taxonomic history:</emphasis>
Previously, the genera
<taxonomicName id="ADF74D1AFF9DFFC3FAE8806604ECF8DB" authority="Stal, 1861" authorityName="Stal" authorityYear="1861" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FAE8806602D0F8FB" box="[1324,1424,1857,1878]" italics="true" pageId="16" pageNumber="1261">Ilyocoris</emphasis>
<bibRefCitation id="0E664B68FF9DFFC3FCED807804ECF8DB" author="Stal C" box="[809,940,1887,1910]" pageId="16" pageNumber="1261" pagination="195 - 212" refId="ref25023" refString="Stal C. 1861. Nova methodus familias quasdam Hemipterorum disponendi. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar 18: 195 - 212." type="journal article" year="1861">Stål, 1861</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9DFFC3FC7B80470311F8D8" authorityName="La Rivers" authorityYear="1956" box="[959,1105,1888,1909]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FC7B80470311F8D8" box="[959,1105,1888,1909]" italics="true" pageId="16" pageNumber="1261">Placomerus</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9DFFC3FB5380470248F8D8" authorityName="Stal" authorityYear="1876" box="[1175,1288,1888,1909]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FB5380470248F8D8" box="[1175,1288,1888,1909]" italics="true" pageId="16" pageNumber="1261">Pelocoris</emphasis>
</taxonomicName>
were held
</heading>
</paragraph>
<paragraph id="6A483699FF9CFFC2FF6787A706C8FF34" blockId="17.[163,392,128,155]" box="[163,392,128,155]" pageId="17" pageNumber="1262">
1262
<collectionCode id="0CE6AE5CFF9CFFC2FEC687A20655FF37" box="[258,277,133,154]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="1262">R</collectionCode>
.W. SITES
</paragraph>
<caption id="3E886611FF9CFFC2FF67844106D7FC34" ID-DOI="http://doi.org/10.5281/zenodo.6994637" ID-Zenodo-Dep="6994637" httpUri="https://zenodo.org/record/6994637/files/figure.png" pageId="17" pageNumber="1262" startId="17.[163,243,870,892]" targetBox="[163,1443,195,830]" targetPageId="17">
<paragraph id="6A483699FF9CFFC2FF67844106D7FC34" blockId="17.[163,1443,870,921]" pageId="17" pageNumber="1262">
<emphasis id="5883EA8BFF9CFFC2FF678441065BFCD6" bold="true" box="[163,283,870,892]" pageId="17" pageNumber="1262">Figure 12.</emphasis>
Mesotibia and mesotarsus of male and female
<taxonomicName id="ADF74D1AFF9CFFC2FCDE84410348FCD6" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[794,1032,870,892]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="5883EA8BFF9CFFC2FCDE84410348FCD6" box="[794,1032,870,892]" italics="true" pageId="17" pageNumber="1262">Heleolaccocoris ovatus</emphasis>
</taxonomicName>
showing a sexually dimorphic ventral tomentose pad of hairs.
</paragraph>
</caption>
<paragraph id="6A483699FF9CFFC2FF6784F706ECF9A0" blockId="17.[163,780,974,1550]" pageId="17" pageNumber="1262">
in
<taxonomicName id="ADF74D1AFF9CFFC2FF0184F70616FC4B" authorityName="LEACH" authorityYear="1815" box="[197,342,976,998]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF9CFFC2FEA784F7069DFC4B" authorityName="POPOV" authorityYear="1970" box="[355,477,976,998]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini</taxonomicName>
(
<bibRefCitation id="0E664B68FF9CFFC2FE3484E905BBFC48" author="Stys P &amp; Jansson A" box="[496,763,974,998]" pageId="17" pageNumber="1262" pagination="1 - 44" refId="ref25103" refString="Stys P, Jansson A. 1988. Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica 50: 1 - 44." type="journal article" year="1988">Štys &amp; Jansson, 1988</bibRefCitation>
). Later,
<bibRefCitation id="0E664B68FF9CFFC2FF3184C80535FBA9" author="Lopez Ruf ML &amp; Bachmann AO" box="[245,629,1007,1029]" pageId="17" pageNumber="1262" pagination="341 - 344" refId="ref22812" refString="Lopez Ruf ML, Bachmann AO. 1987. Sobre la classification de los Naucoroidea Americanos (Heteroptera). Revista de la Sociedad Entomologica Argentina 44: 341 - 344." type="journal article" year="1987">López Ruf &amp; Bachmann (1987)</bibRefCitation>
erected the tribe
<taxonomicName id="ADF74D1AFF9CFFC2FF23832A0621FB8E" box="[231,353,1037,1059]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="tribe" tribe="Pelocorini">Pelocorini</taxonomicName>
to hold
<taxonomicName id="ADF74D1AFF9CFFC2FE0E832A0517FB8F" authorityName="La Rivers" authorityYear="1956" box="[458,599,1037,1058]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FE0E832A0517FB8F" box="[458,599,1037,1058]" italics="true" pageId="17" pageNumber="1262">Placomerus</emphasis>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF9CFFC2FDAC832A0594FB8F" authorityName="Stal" authorityYear="1876" box="[616,724,1037,1058]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FDAC832A0594FB8F" box="[616,724,1037,1058]" italics="true" pageId="17" pageNumber="1262">Pelocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9CFFC2FF67830B0610FBEC" authorityName="De Carlo" authorityYear="1963" box="[163,336,1068,1089]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FF67830B0610FBEC" box="[163,336,1068,1089]" italics="true" pageId="17" pageNumber="1262">Carvalhoiella</emphasis>
</taxonomicName>
, separate from
<taxonomicName id="ADF74D1AFF9CFFC2FDE4830B05DDFBEF" authorityName="POPOV" authorityYear="1970" box="[544,669,1068,1090]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini</taxonomicName>
because they followed
<bibRefCitation id="0E664B68FF9CFFC2FEA4836C0506FBCD" author="De Carlo JA" box="[352,582,1099,1121]" pageId="17" pageNumber="1262" pagination="159 - 166" refId="ref21730" refString="De Carlo JA. 1971. Valor sistematico del estudio del aparato genital macho en los hemipteros acuaticos y semicuaticos. Division de la familia Naucoridae en dos familias. Caracteristica externa de la capsula genital. Revista de la Sociedad Entomologica Argentina 33: 159 - 166." type="journal article" year="1971">De Carlo’s (1971)</bibRefCitation>
hemispherical splitting of the family into two families,
<taxonomicName id="ADF74D1AFF9CFFC2FD47834E044AFBD2" authorityName="LEACH" authorityYear="1815" box="[643,778,1129,1151]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="family">Naucoridae</taxonomicName>
and Pelocoridae (= Limnocoridae). Although
<taxonomicName id="ADF74D1AFF9CFFC2FF678380060AFB11" authorityName="De Carlo" authorityYear="1963" box="[163,330,1191,1212]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FF678380060AFB11" box="[163,330,1191,1212]" italics="true" pageId="17" pageNumber="1262">Carvalhoiella</emphasis>
</taxonomicName>
clearly was misplaced in
<taxonomicName id="ADF74D1AFF9CFFC2FD4F83800446FB10" box="[651,774,1191,1213]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="tribe" tribe="Pelocorini">Pelocorini</taxonomicName>
, the genus had been transferred to
<taxonomicName id="ADF74D1AFF9CFFC2FD4C83E10449FB76" authorityName="POPOV" authorityYear="1970" box="[648,777,1221,1243]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1271" phylum="Arthropoda" rank="tribe" tribe="Naucorini">Naucorini</taxonomicName>
by
<bibRefCitation id="0E664B68FF9CFFC2FF0983C306C1FB57" author="Nieser N" box="[205,385,1252,1274]" pageId="17" pageNumber="1262" pagination="1 - 303" refId="ref23396" refString="Nieser N. 1975. The water bugs (Heteroptera: Nepomorpha) of the Guyana Region. Studies on the Fauna of Suriname and other Guyanas 16: 1 - 303." type="journal article" year="1975">Nieser (1975)</bibRefCitation>
, but later was appropriately transferred back to
<taxonomicName id="ADF74D1AFF9CFFC2FE4E82240447FAB5" authority="(Nieser et al., 1999)" authorityName="Stal" authorityYear="1862" baseAuthorityName="Nieser" baseAuthorityYear="1999" box="[394,775,1283,1305]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">
Ambrysinae (
<bibRefCitation id="0E664B68FF9CFFC2FDEC822405BCFAB5" author="Nieser N &amp; Pelli A &amp; Melo AL" box="[552,764,1283,1305]" pageId="17" pageNumber="1262" pagination="157 - 163" refId="ref23491" refString="Nieser N, Pelli A, Melo AL. 1999. Two new Ambrysinae (Heteroptera: Naucoridae) from Minas Gerais, Brazil. Acta Societatis Zoologicae Bohemoslovacae 63: 157 - 163." type="journal article" year="1999">
Nieser
<emphasis id="5883EA8BFF9CFFC2FDB8822405F6FAB5" box="[636,694,1283,1304]" italics="true" pageId="17" pageNumber="1262">et al.</emphasis>
, 1999
</bibRefCitation>
)
</taxonomicName>
. During this interim,
<bibRefCitation id="0E664B68FF9CFFC2FE598206044BFA9B" author="Lopez Ruf ML &amp; Bachmann AO" box="[413,779,1313,1335]" pageId="17" pageNumber="1262" pagination="341 - 344" refId="ref22812" refString="Lopez Ruf ML, Bachmann AO. 1987. Sobre la classification de los Naucoroidea Americanos (Heteroptera). Revista de la Sociedad Entomologica Argentina 44: 341 - 344." type="journal article" year="1987">López Ruf &amp; Bachmann (1987)</bibRefCitation>
followed Nieser in recognizing
<taxonomicName id="ADF74D1AFF9CFFC2FDE482670586FAF8" authorityName="De Carlo" authorityYear="1963" box="[544,710,1344,1365]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FDE482670586FAF8" box="[544,710,1344,1365]" italics="true" pageId="17" pageNumber="1262">Carvalhoiella</emphasis>
</taxonomicName>
to be a member of this group, which they recognized as
<taxonomicName id="ADF74D1AFF9CFFC2FF67825A065EFA3E" box="[163,286,1405,1427]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="tribe" tribe="Pelocorini">Pelocorini</taxonomicName>
. Following the transfer of
<taxonomicName id="ADF74D1AFF9CFFC2FDA4825A058AFA3F" authorityName="Stal" authorityYear="1876" box="[608,714,1405,1426]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FDA4825A058AFA3F" box="[608,714,1405,1426]" italics="true" pageId="17" pageNumber="1262">Pelocoris</emphasis>
</taxonomicName>
from
<taxonomicName id="ADF74D1AFF9CFFC2FF6782BB066EFA1F" authorityName="LEACH" authorityYear="1815" box="[163,302,1436,1458]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
:
<taxonomicName id="ADF74D1AFF9CFFC2FEFD82BB06EDFA1F" box="[313,429,1436,1458]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="tribe" tribe="Pelocorini">Pelocorini</taxonomicName>
to
<taxonomicName id="ADF74D1AFF9CFFC2FE1682BB0521FA1F" authorityName="Stal" authorityYear="1862" box="[466,609,1436,1458]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
, only
<taxonomicName id="ADF74D1AFF9CFFC2FD6382BB044AFA1C" authorityName="Stal" authorityYear="1861" box="[679,778,1436,1457]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FD6382BB044AFA1C" box="[679,778,1436,1457]" italics="true" pageId="17" pageNumber="1262">Ilyocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9CFFC2FF12829D061DFA62" authorityName="La Rivers" authorityYear="1956" box="[214,349,1466,1487]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FF12829D061DFA62" box="[214,349,1466,1487]" italics="true" pageId="17" pageNumber="1262">Placomerus</emphasis>
</taxonomicName>
remained in the tribe; thus the tribe
<taxonomicName id="ADF74D1AFF9CFFC2FF6782FE065BFA42" box="[163,283,1497,1519]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="tribe" tribe="Pelocorini">Pelocorini</taxonomicName>
necessarily became
<taxonomicName id="ADF74D1AFF9CFFC2FDDC82FE05CCFA42" box="[536,652,1497,1519]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1268" phylum="Arthropoda" rank="tribe" tribe="Ilyocorini">Ilyocorini</taxonomicName>
(ReynosoVelasco &amp;
<taxonomicName id="ADF74D1AFF9CFFC2FED982DF06DCF9A0" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[285,412,1528,1550]" genus="Sites" pageId="17" pageNumber="1270" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF9CFFC2FED982DF06DCF9A0" author="Sites RW" box="[285,412,1528,1550]" pageId="17" pageNumber="1262" pagination="72 - 94" refId="ref24528" refString="Sites RW. 2021. A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species. Zootaxa 4958: 72 - 94." type="journal article" year="2021">Sites, 2021</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="6A483699FF9CFFC2FF6781110668F8A9" blockId="17.[163,779,1590,1796]" pageId="17" pageNumber="1262">
<emphasis id="5883EA8BFF9CFFC2FF67811106D4F9E1" box="[163,404,1590,1612]" italics="true" pageId="17" pageNumber="1262">Revised taxonomy:</emphasis>
With the recent transfer of
<taxonomicName id="ADF74D1AFF9CFFC2FF678172064BF9C7" authorityName="Stal" authorityYear="1876" box="[163,267,1621,1642]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FF678172064BF9C7" box="[163,267,1621,1642]" italics="true" pageId="17" pageNumber="1262">Pelocoris</emphasis>
</taxonomicName>
from
<taxonomicName id="ADF74D1AFF9CFFC2FE968172069AF9C6" authorityName="LEACH" authorityYear="1815" box="[338,474,1621,1643]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
to
<taxonomicName id="ADF74D1AFF9CFFC2FDC5817206D9F924" authority="(ReynosoVelasco &amp; Sites, 2021)" authorityName="Stal" authorityYear="1862" baseAuthorityName="ReynosoVelasco &amp; Sites" baseAuthorityYear="2021" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">
Ambrysinae (ReynosoVelasco &amp;
<bibRefCitation id="0E664B68FF9CFFC2FED3815306D1F924" author="Sites RW" box="[279,401,1652,1674]" pageId="17" pageNumber="1262" pagination="72 - 94" refId="ref24528" refString="Sites RW. 2021. A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species. Zootaxa 4958: 72 - 94." type="journal article" year="2021">Sites, 2021</bibRefCitation>
)
</taxonomicName>
and clear association of
<taxonomicName id="ADF74D1AFF9CFFC2FD6F8153044BF924" authorityName="Stal" authorityYear="1861" box="[683,779,1652,1673]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FD6F8153044BF924" box="[683,779,1652,1673]" italics="true" pageId="17" pageNumber="1262">Ilyocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9CFFC2FF1E81B50627F90A" authorityName="La Rivers" authorityYear="1956" box="[218,359,1682,1703]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FF1E81B50627F90A" box="[218,359,1682,1703]" italics="true" pageId="17" pageNumber="1262">Placomerus</emphasis>
</taxonomicName>
as sister genera distinct from all others, I here remove these genera from
<taxonomicName id="ADF74D1AFF9CFFC2FD478196044AF96A" authorityName="LEACH" authorityYear="1815" box="[643,778,1713,1735]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
by elevating the tribe
<taxonomicName id="ADF74D1AFF9CFFC2FE5981F70549F94B" box="[413,521,1744,1766]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="tribe" tribe="Ilyocorini">Ilyocorini</taxonomicName>
to subfamily status as
<taxonomicName id="ADF74D1AFF9CFFC2FF6781C90665F8A9" authorityName="Sites" authorityYear="2022" box="[163,293,1774,1796]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1261" phylum="Arthropoda" rank="subFamily" status="stat. nov." subFamily="Ilyocorinae">Ilyocorinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9CFFC2FF678009021BFBD3" pageId="17" pageNumber="1262" type="diagnosis">
<paragraph id="6A483699FF9CFFC2FF678009021BFBD3" blockId="17.[163,779,1837,1890]" lastBlockId="17.[827,1444,976,1151]" pageId="17" pageNumber="1262">
<emphasis id="5883EA8BFF9CFFC2FF678009065CF8EE" box="[163,284,1838,1859]" italics="true" pageId="17" pageNumber="1262">Diagnosis:</emphasis>
The female genitalia are strongly dentate along the dorsal and lateral margins of valvulae 1 and lateral margins of valvulae 2 (
<figureCitation id="F2CC2A1CFF9CFFC2FAD684F7021AFC48" box="[1298,1370,976,998]" captionStart="Figure 7" captionStartId="11.[163,241,696,718]" captionTargetBox="[399,1207,195,656]" captionTargetId="figure-197@11.[399,1207,195,656]" captionTargetPageId="11" captionText="Figure 7. Valvulae of female Ilyocoris cimicoides with dentate margins. vlf = valvifer, vlv = valvula." figureDoi="http://doi.org/10.5281/zenodo.6994623" httpUri="https://zenodo.org/record/6994623/files/figure.png" pageId="17" pageNumber="1262">Fig. 7</figureCitation>
). The male parameres are twisted, contoured (López-Ruf &amp; Bachmann, 1991), elongated, overlapping and symmetrical or with only slight asymmetry (
<figureCitation id="F2CC2A1CFF9CFFC2FAFD830B02D5FBEF" box="[1337,1429,1068,1090]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="17" pageNumber="1262">Fig. 11d</figureCitation>
). The mesofemur posterodorsal margin is flattened and fringed with a dense brush-line of hairs (
<figureCitation id="F2CC2A1CFF9CFFC2FAC8834E020CFBD3" box="[1292,1356,1129,1151]" captionStart="Figure 8" captionStartId="11.[163,242,1486,1508]" captionTargetBox="[163,1443,800,1447]" captionTargetId="figure-171@11.[163,1443,800,1447]" captionTargetPageId="11" captionText="Figure 8. Mesofemora of Ilyocoris cimicoides (a) and Placomerus sp. (b) showing the posterodorsal margin flattened and fringed with a dense brush-line of hairs." figureDoi="http://doi.org/10.5281/zenodo.6994627" httpUri="https://zenodo.org/record/6994627/files/figure.png" pageId="17" pageNumber="1262">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9CFFC2FCFF838F0489F9A2" pageId="17" pageNumber="1262" type="discussion">
<paragraph id="6A483699FF9CFFC2FCFF838F0489F9A2" blockId="17.[827,1443,1192,1551]" pageId="17" pageNumber="1262">
<emphasis id="5883EA8BFF9CFFC2FCFF838F04FBFB10" box="[827,955,1192,1213]" italics="true" pageId="17" pageNumber="1262">Comments:</emphasis>
The condition of the mesofemur in which the posterodorsal margin is flattened and fringed with a dense brush-line of hairs (also present in many members of
<taxonomicName id="ADF74D1AFF9CFFC2FBE4822303FFFAB7" authorityName="STAL" authorityYear="1876" box="[1056,1215,1284,1306]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
) had been used to distinguish between
<taxonomicName id="ADF74D1AFF9CFFC2FBFE82050385FA9A" authorityName="La Rivers" authorityYear="1956" box="[1082,1221,1314,1335]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FBFE82050385FA9A" box="[1082,1221,1314,1335]" italics="true" pageId="17" pageNumber="1262">Placomerus</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9CFFC2FAC282050498FAFB" authority="(La Rivers, 1956)" baseAuthorityName="La Rivers" baseAuthorityYear="1956" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FAC282050231FA9A" box="[1286,1393,1314,1335]" italics="true" pageId="17" pageNumber="1262">Pelocoris</emphasis>
(
<bibRefCitation id="0E664B68FF9CFFC2FA408204048DFAFA" author="La Rivers I" pageId="17" pageNumber="1262" pagination="237 - 245" refId="ref22373" refString="La Rivers I. 1956. A new genus and species of naucorid from South America (Hemiptera). Entomological News 67: 237 - 245." type="journal article" year="1956">La Rivers, 1956</bibRefCitation>
)
</taxonomicName>
. Those two genera are superficially so similar that
<bibRefCitation id="0E664B68FF9CFFC2FC028247033FFADB" author="La Rivers I" box="[966,1151,1376,1398]" pageId="17" pageNumber="1262" pagination="237 - 245" refId="ref22373" refString="La Rivers I. 1956. A new genus and species of naucorid from South America (Hemiptera). Entomological News 67: 237 - 245." type="journal article" year="1956">La Rivers (1956)</bibRefCitation>
prepared a list of features to distinguish between them. However,
<taxonomicName id="ADF74D1AFF9CFFC2FACC8259022EFA3E" baseAuthorityName="La Rivers" baseAuthorityYear="1956" box="[1288,1390,1406,1427]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FACC8259022EFA3E" box="[1288,1390,1406,1427]" italics="true" pageId="17" pageNumber="1262">Pelocoris</emphasis>
</taxonomicName>
was shown recently to be distantly related to
<taxonomicName id="ADF74D1AFF9CFFC2FAD882BA02E2FA1F" authorityName="La Rivers" authorityYear="1956" box="[1308,1442,1437,1458]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FAD882BA02E2FA1F" box="[1308,1442,1437,1458]" italics="true" pageId="17" pageNumber="1262">Placomerus</emphasis>
</taxonomicName>
and is now in a different subfamily (
<taxonomicName id="ADF74D1AFF9CFFC2FAC4829B02D6FA7F" authorityName="Stal" authorityYear="1862" box="[1280,1430,1468,1490]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
).
<taxonomicName id="ADF74D1AFF9CFFC2FCFF82FD04DBFA42" authorityName="Stal" authorityYear="1861" box="[827,923,1498,1519]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FCFF82FD04DBFA42" box="[827,923,1498,1519]" italics="true" pageId="17" pageNumber="1262">Ilyocoris</emphasis>
</taxonomicName>
is Palaearctic and
<taxonomicName id="ADF74D1AFF9CFFC2FBB182FD02C4FA5D" authority="Neotropical" authorityName="Neotropical" box="[1141,1412,1498,1520]" class="Insecta" family="Naucoridae" genus="Placomerus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="1262" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9CFFC2FBB182FD03B9FA42" box="[1141,1273,1498,1519]" italics="true" pageId="17" pageNumber="1262">Placomerus</emphasis>
Neotropical
</taxonomicName>
in distribution.
</paragraph>
</subSubSection>
</treatment>
</document>

200
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<paragraph id="6A483699FF9DFFC3FF62820805A4FAEA" blockId="16.[166,740,1327,1352]" box="[166,740,1327,1352]" pageId="16" pageNumber="1261">
SUBFAMILY
<taxonomicName id="ADF74D1AFF9DFFC3FEE982080595FAE5" ID-CoL="626SY" authority="MONTANDON, 1897" authorityName="MONTANDON" authorityYear="1897" box="[301,725,1327,1352]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">CRYPHOCRICINAE MONTANDON, 1897</taxonomicName>
A
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FEB1827C0555FADE" box="[373,533,1371,1396]" pageId="16" pageNumber="1261" type="description">
<paragraph id="6A483699FF9DFFC3FEB1827C0555FADE" blockId="16.[373,533,1371,1396]" box="[373,533,1371,1396]" pageId="16" pageNumber="1261">
<figureCitation id="F2CC2A1CFF9DFFC3FEB1827C069DFADE" box="[373,477,1371,1396]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="16" pageNumber="1261">FIGS 10C</figureCitation>
,
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</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FF5582B5059BFA0A" box="[145,731,1425,1447]" pageId="16" pageNumber="1261" type="type_taxon">
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<emphasis id="5883EA8BFF9DFFC3FF5582B50661FA0A" box="[145,289,1426,1447]" italics="true" pageId="16" pageNumber="1261">
<typeStatus id="B54C883BFF9DFFC3FF5582B50786FA0A" box="[145,198,1426,1447]" pageId="16" pageNumber="1261">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF9DFFC3FEF082B60594FA0A" ID-CoL="6BK23" authority="Signoret, 1850" authorityName="Signoret" authorityYear="1850" box="[308,724,1425,1447]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="species" species="barozzii">
<emphasis id="5883EA8BFF9DFFC3FEF082B60568FA0B" box="[308,552,1425,1446]" italics="true" pageId="16" pageNumber="1261">Cryphocricos barozzii</emphasis>
<bibRefCitation id="0E664B68FF9DFFC3FDEB82B60594FA0A" author="Signoret V" box="[559,724,1425,1447]" pageId="16" pageNumber="1261" pagination="289 - 291" refId="ref24266" refString="Signoret V. 1850. Description d'un genre nouveau de l'ordre des Hemipteres heteropteres, et de la section des Hydrocoryses. Revue et Magasin de Zoologie Pure et Appliquee 2: 289 - 291." type="journal article" year="1850">Signoret, 1850</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FF5582F703F1FDC6" pageId="16" pageNumber="1261" type="discussion">
<paragraph id="6A483699FF9DFFC3FF5582F703C4FE42" blockId="16.[145,762,1488,1908]" lastBlockId="16.[809,1426,197,495]" pageId="16" pageNumber="1261">
<emphasis id="5883EA8BFF9DFFC3FF5582F70633FA48" box="[145,371,1488,1509]" italics="true" pageId="16" pageNumber="1261">Taxonomic history:</emphasis>
<bibRefCitation id="0E664B68FF9DFFC3FE4582F7052AFA48" author="Montandon AL" box="[385,618,1488,1510]" pageId="16" pageNumber="1261" pagination="6 - 23" refId="ref22979" refString="Montandon AL. 1897 a. Hemiptera Cryptocerata. Fam. Naucoridae. Sous-fam. Cryptocricinae. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 47: 6 - 23." type="journal article" year="1897">Montandon (1897a)</bibRefCitation>
erected the subfamily (originally misspelled as Cryptocricinae, but later emended) to include
<taxonomicName id="ADF74D1AFF9DFFC3FDD8812A05FFF98F" authorityName="Signoret" authorityYear="1850" box="[540,703,1549,1570]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FDD8812A05FFF98F" box="[540,703,1549,1570]" italics="true" pageId="16" pageNumber="1261">Cryphocricos</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9DFFC3FF55810B064EF9EC" authorityName="Stal" authorityYear="1862" box="[145,270,1580,1601]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FF55810B064EF9EC" box="[145,270,1580,1601]" italics="true" pageId="16" pageNumber="1261">Ambrysus</emphasis>
</taxonomicName>
, as well as the two therein-proposed genera
<taxonomicName id="ADF74D1AFF9DFFC3FF30816D062DF9F2" authorityName="Montandon" authorityYear="1897" box="[244,365,1610,1631]" class="Insecta" family="Naucoridae" genus="Idiocarus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FF30816D062DF9F2" box="[244,365,1610,1631]" italics="true" pageId="16" pageNumber="1261">Idiocarus</emphasis>
</taxonomicName>
(currently in
<taxonomicName id="ADF74D1AFF9DFFC3FDE9816D05AEF9CD" authorityName="MONTANDON" authorityYear="1897" box="[557,750,1610,1632]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1260" phylum="Arthropoda" rank="subFamily" subFamily="Cheirochelinae">Cheirochelinae</taxonomicName>
) and
<taxonomicName id="ADF74D1AFF9DFFC3FF0D814E06CAF9D3" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[201,394,1641,1662]" class="Insecta" genus="Pseudambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FF0D814E06CAF9D3" box="[201,394,1641,1662]" italics="true" pageId="16" pageNumber="1261">Pseudambrysus</emphasis>
</taxonomicName>
(currently a junior synonym of
<taxonomicName id="ADF74D1AFF9DFFC3FF7281AF0606F930" authorityName="Signoret" authorityYear="1861" box="[182,326,1672,1693]" class="Insecta" family="Naucoridae" genus="Macrocoris" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FF7281AF0606F930" box="[182,326,1672,1693]" italics="true" pageId="16" pageNumber="1261">Macrocoris</emphasis>
</taxonomicName>
in
<taxonomicName id="ADF74D1AFF9DFFC3FEBF81AF0552F930" authorityName="LEACH" authorityYear="1815" box="[379,530,1672,1694]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
).
<bibRefCitation id="0E664B68FF9DFFC3FDF781AF05B1F930" author="Usinger RL" box="[563,753,1672,1694]" pageId="16" pageNumber="1261" pagination="5 - 16" refId="ref25189" refString="Usinger RL. 1941. Key to the subfamilies of Naucoridae with a generic synopsis of the new subfamily Ambrysinae (Hemiptera). Annals of the Entomological Society of America 34: 5 - 16." type="journal article" year="1941">Usinger (1941</bibRefCitation>
,
<bibRefCitation id="0E664B68FF9DFFC3FF558180078CF911" author="Usinger RL" box="[145,204,1703,1724]" pageId="16" pageNumber="1261" pagination="329 - 343" refId="ref25226" refString="Usinger RL. 1947. Classification of the Cryphocricinae (Hemiptera: Naucoridae). Annals of the Entomological Society of America 40: 329 - 343." type="journal article" year="1947">1947</bibRefCitation>
) proposed a more restrictive classification scheme in which
<taxonomicName id="ADF74D1AFF9DFFC3FEA081E2055CF976" authorityName="MONTANDON" authorityYear="1897" box="[356,540,1733,1755]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
included only the genus
<taxonomicName id="ADF74D1AFF9DFFC3FF2581C3063EF954" authorityName="Signoret" authorityYear="1850" box="[225,382,1764,1785]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FF2581C3063EF954" box="[225,382,1764,1785]" italics="true" pageId="16" pageNumber="1261">Cryphocricos</emphasis>
</taxonomicName>
; he transferred
<taxonomicName id="ADF74D1AFF9DFFC3FD8E81C30582F954" authorityName="Stal" authorityYear="1862" box="[586,706,1764,1785]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FD8E81C30582F954" box="[586,706,1764,1785]" italics="true" pageId="16" pageNumber="1261">Ambrysus</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF9DFFC3FF5580250646F8BA" authorityName="De Carlo" authorityYear="1935" box="[145,262,1794,1815]" class="Insecta" family="Naucoridae" genus="Melloiella" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FF5580250646F8BA" box="[145,262,1794,1815]" italics="true" pageId="16" pageNumber="1261">Melloiella</emphasis>
</taxonomicName>
into his newly erected
<taxonomicName id="ADF74D1AFF9DFFC3FDDC802505E8F8B5" authorityName="Stal" authorityYear="1862" box="[536,680,1794,1816]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
.
<bibRefCitation id="0E664B68FF9DFFC3FD708024079BF89B" author="Popov YA" pageId="16" pageNumber="1261" pagination="93 - 98" refId="ref23866" refString="Popov YA. 1970. Notes on the classification of the recent Naucoridae (Heteroptera: Nepomorpha). Bulletin of the Polish Academy of Sciences (Biological Sciences) 18: 93 - 98." type="journal article" year="1970">Popov (1970)</bibRefCitation>
erected the tribe
<taxonomicName id="ADF74D1AFF9DFFC3FE708006052EF89A" authorityName="Popov" authorityYear="1970" box="[436,622,1825,1847]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="tribe" tribe="Cataractocorini">Cataractocorini</taxonomicName>
and placed it and Cryphocricini in
<taxonomicName id="ADF74D1AFF9DFFC3FE0F806705D1F8FB" authorityName="MONTANDON" authorityYear="1897" box="[459,657,1856,1878]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
.
<bibRefCitation id="0E664B68FF9DFFC3FD6580190605F8D9" author="Stys P &amp; Jansson A" pageId="16" pageNumber="1261" pagination="1 - 44" refId="ref25103" refString="Stys P, Jansson A. 1988. Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica 50: 1 - 44." type="journal article" year="1988">Štys &amp; Jansson (1988)</bibRefCitation>
placed all three groups (Ambrysini,
<taxonomicName id="ADF74D1AFF9DFFC3FCED87E204A3FF76" authorityName="Popov" authorityYear="1970" box="[809,995,197,219]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="tribe" tribe="Cataractocorini">Cataractocorini</taxonomicName>
, Cryphocricini) together as tribes in the subfamily
<taxonomicName id="ADF74D1AFF9DFFC3FC1F87C303D7FF57" authorityName="MONTANDON" authorityYear="1897" box="[987,1175,228,250]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
. This subfamily was revised recently (
<bibRefCitation id="0E664B68FF9DFFC3FBC3862502C7FEB5" author="Reynoso-Velasco D &amp; Sites RW" box="[1031,1415,258,280]" pageId="16" pageNumber="1261" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF9DFFC3FB39862502C7FEB5" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1277,1415,258,280]" genus="Sites" pageId="16" pageNumber="1270" rank="genus" status="gen. nov.">Sites, 2021</taxonomicName>
</bibRefCitation>
) and now includes only the genus
<taxonomicName id="ADF74D1AFF9DFFC3FB20860602C9FE9B" authorityName="Signoret" authorityYear="1850" box="[1252,1417,289,310]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FB20860602C9FE9B" box="[1252,1417,289,310]" italics="true" pageId="16" pageNumber="1261">Cryphocricos</emphasis>
</taxonomicName>
, agreeing with
<bibRefCitation id="0E664B68FF9DFFC3FC0A86670331FEF8" author="Usinger RL" box="[974,1137,320,342]" pageId="16" pageNumber="1261" pagination="5 - 16" refId="ref25189" refString="Usinger RL. 1941. Key to the subfamilies of Naucoridae with a generic synopsis of the new subfamily Ambrysinae (Hemiptera). Annals of the Entomological Society of America 34: 5 - 16." type="journal article" year="1941">Usinger (1941</bibRefCitation>
,
<bibRefCitation id="0E664B68FF9DFFC3FBBA866703F6FEF8" author="Usinger RL" box="[1150,1206,320,341]" pageId="16" pageNumber="1261" pagination="329 - 343" refId="ref25226" refString="Usinger RL. 1947. Classification of the Cryphocricinae (Hemiptera: Naucoridae). Annals of the Entomological Society of America 40: 329 - 343." type="journal article" year="1947">1947</bibRefCitation>
), with
<taxonomicName id="ADF74D1AFF9DFFC3FAC1866702D1FEFB" authorityName="Stal" authorityYear="1862" box="[1285,1425,320,342]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">Ambrysinae</taxonomicName>
comprising the other groups formerly held within
<taxonomicName id="ADF74D1AFF9DFFC3FCED865A04AAFE3E" authorityName="MONTANDON" authorityYear="1897" box="[809,1002,381,403]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
. Recently,
<taxonomicName id="ADF74D1AFF9DFFC3FBB0865A024AFE3F" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1140,1290,381,403]" genus="Sites" pageId="16" pageNumber="1270" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF9DFFC3FBB0865A024AFE3F" author="Sites RW" box="[1140,1290,381,403]" pageId="16" pageNumber="1261" pagination="72 - 94" refId="ref24528" refString="Sites RW. 2021. A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species. Zootaxa 4958: 72 - 94." type="journal article" year="2021">Sites (2021)</bibRefCitation>
</taxonomicName>
presented a review of the genus
<taxonomicName id="ADF74D1AFF9DFFC3FB9786BB03B9FE1C" authorityName="Signoret" authorityYear="1850" box="[1107,1273,412,433]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FB9786BB03B9FE1C" box="[1107,1273,412,433]" italics="true" pageId="16" pageNumber="1261">Cryphocricos</emphasis>
</taxonomicName>
along with descriptions of three new species, two of which are included in the analyses here.
</paragraph>
<paragraph id="6A483699FF9DFFC3FCED853F03F1FDC6" blockId="16.[809,1424,536,619]" pageId="16" pageNumber="1261">
<emphasis id="5883EA8BFF9DFFC3FCED853F0355FD80" box="[809,1045,536,557]" italics="true" pageId="16" pageNumber="1261">Revised taxonomy:</emphasis>
This subfamily was recently revised by
<bibRefCitation id="0E664B68FF9DFFC3FC608511024AFDE1" author="Reynoso-Velasco D &amp; Sites RW" box="[932,1290,566,588]" pageId="16" pageNumber="1261" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF9DFFC3FB468511024AFDE1" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1154,1290,566,588]" genus="Sites" pageId="16" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
. No further taxonomic action is proposed here.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FCED85B30393FC57" pageId="16" pageNumber="1261" type="diagnosis">
<paragraph id="6A483699FF9DFFC3FCED85B30393FC57" blockId="16.[809,1425,660,1019]" pageId="16" pageNumber="1261">
<emphasis id="5883EA8BFF9DFFC3FCED85B304EAFD04" box="[809,938,660,681]" italics="true" pageId="16" pageNumber="1261">Diagnosis:</emphasis>
Diagnostic features for the subfamily were provided by
<bibRefCitation id="0E664B68FF9DFFC3FC328595021EFD65" author="Reynoso-Velasco D &amp; Sites RW" box="[1014,1374,690,712]" pageId="16" pageNumber="1261" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF9DFFC3FB128595021EFD65" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1238,1374,690,712]" genus="Sites" pageId="16" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
and include: labium inserted near the anterior margin of the head, labrum well developed, maxillary plates produced anteriorly and reaching the labral apex, anterior margin of the pronotum with a deep concavity, pronotum with crenate lateral margins, prosternum exposed and at the same dorsoventral level as the propleura, alary polymorphism (
<taxonomicName id="ADF74D1AFF9DFFC3FC1A84AE031BFC33" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[990,1115,905,927]" genus="Sites" pageId="16" pageNumber="1270" rank="genus" status="gen. nov.">Sites, 1990</taxonomicName>
), hydrostatic sense organs adjacent to lateral margin on abdominal sterna II–VII, plastral respiration, glabrous abdomen and males with a long, tubular, coiled vesica (
<figureCitation id="F2CC2A1CFF9DFFC3FBAB84C20384FC57" box="[1135,1220,997,1019]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="16" pageNumber="1261">Fig. 11c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9DFFC3FCED830303A7FA65" pageId="16" pageNumber="1261" type="discussion">
<paragraph id="6A483699FF9DFFC3FCED830303A7FA65" blockId="16.[809,1425,1060,1480]" pageId="16" pageNumber="1261">
<emphasis id="5883EA8BFF9DFFC3FCED830304E7FB94" box="[809,935,1060,1081]" italics="true" pageId="16" pageNumber="1261">Comments:</emphasis>
This subfamily is exclusively New World in distribution and is represented by 16 described species, with additional undescribed species represented in these analyses. Although formal descriptions of these species are important, they are of limited value because of the difficulty in finding species-specific diagnostic features.
<bibRefCitation id="0E664B68FF9DFFC3FC5083FB0300FB5C" author="Usinger RL" box="[916,1088,1244,1266]" pageId="16" pageNumber="1261" pagination="329 - 343" refId="ref25226" refString="Usinger RL. 1947. Classification of the Cryphocricinae (Hemiptera: Naucoridae). Annals of the Entomological Society of America 40: 329 - 343." type="journal article" year="1947">Usinger (1947)</bibRefCitation>
used size- and shape-related features to diagnose species from Central and South America. However, in an attempt to determine the specific identity of
<taxonomicName id="ADF74D1AFF9DFFC3FBC1821F03DBFAE0" authorityName="Signoret" authorityYear="1850" box="[1029,1179,1336,1357]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="1261" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF9DFFC3FBC1821F03DBFAE0" box="[1029,1179,1336,1357]" italics="true" pageId="16" pageNumber="1261">Cryphocricos</emphasis>
</taxonomicName>
in
<collectingCountry id="12E07609FF9DFFC3FB06821F024BFAE3" box="[1218,1291,1336,1358]" name="Belize" pageId="16" pageNumber="1261">Belize</collectingCountry>
,
<bibRefCitation id="0E664B68FF9DFFC3FAD2821F0430FAC6" author="Sites RW &amp; Carrie R &amp; Shepard WD" pageId="16" pageNumber="1261" pagination="243 - 273" refId="ref24559" refString="Sites RW, Carrie R, Shepard WD. 2018. The Naucoridae (Heteroptera: Nepomorpha) of Belize. Aquatic Insects 39: 243 - 273." type="journal article" year="2018">
<taxonomicName id="ADF74D1AFF9DFFC3FAD2821F0210FAE3" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1302,1360,1336,1358]" genus="Sites" pageId="16" pageNumber="1270" rank="genus" status="gen. nov.">Sites</taxonomicName>
<emphasis id="5883EA8BFF9DFFC3FA93821F02D1FAE0" box="[1367,1425,1336,1357]" italics="true" pageId="16" pageNumber="1261">et al.</emphasis>
(2018)
</bibRefCitation>
were unable to reliably identify their specimens using Usinger’s criteria because intraspecific variation was sufficiently high that some populations contained individuals that keyed to three species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7BAB5D7E4074A5FECAAC6C94B02B465B">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:date id="A988F7BE47C00CD8AECFA640D1D61642">2022</mods:date>
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<subSubSection id="22ED6512FF9EFFC0FC6282BE0277FA1C" box="[934,1335,1432,1458]" pageId="19" pageNumber="1264" type="nomenclature">
<paragraph id="6A483699FF9EFFC0FC6282BE0277FA1C" blockId="19.[934,1335,1432,1458]" box="[934,1335,1432,1458]" pageId="19" pageNumber="1264">
<heading id="310081F5FF9EFFC0FC6282BE0277FA1C" box="[934,1335,1432,1458]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="1264" reason="3">
TRIBE
<taxonomicName id="ADF74D1AFF9EFFC0FC3682BF03F2FA02" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1010,1202,1432,1456]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1264" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Ctenipocorini">CTENIPOCORINI</taxonomicName>
<emphasis id="5883EA8BFF9EFFC0FB7D82BB0277FA1C" bold="true" box="[1209,1335,1433,1457]" pageId="19" pageNumber="1264">
<taxonomicNameLabel id="43B057F0FF9EFFC0FB7D82BB0277FA1C" box="[1209,1335,1433,1457]" pageId="19" pageNumber="1264" rank="tribe">TRIB. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="22ED6512FF9EFFC0FB8182E202C3F982" pageId="19" pageNumber="1264" type="description">
<paragraph id="6A483699FF9EFFC0FB8182E203D8FA71" blockId="19.[1093,1176,1477,1502]" box="[1093,1176,1477,1502]" pageId="19" pageNumber="1264">
<figureCitation id="F2CC2A1CFF9EFFC0FB8182E203D8FA71" box="[1093,1176,1477,1502]" captionStart="Figure 4" captionStartId="9.[163,249,1379,1401]" captionTargetBox="[163,781,195,1339]" captionTargetId="figure-575@9.[163,781,195,1339]" captionTargetPageId="9" captionText="Figure 4. Foretibiae and foretarsi of Laccocorinae: Ctenipocoris asiaticus (a), Heleolaccocoris ovatus (b), Namtokocoris dalanta (c)." figureDoi="http://doi.org/10.5281/zenodo.6994615" httpUri="https://zenodo.org/record/6994615/files/figure.png" pageId="19" pageNumber="1264">FIG. 4A</figureCitation>
</paragraph>
<paragraph id="6A483699FF9EFFC0FCFF82DC02C3F982" blockId="19.[827,1444,1530,1583]" pageId="19" pageNumber="1264">
<emphasis id="5883EA8BFF9EFFC0FCFF82DC03C9F9BD" box="[827,1161,1530,1552]" italics="true" pageId="19" pageNumber="1264">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFF9EFFC0FC4E813E02C3F982" box="[906,1411,1561,1583]" pageId="19" pageNumber="1264">12EF5622-B1D8-465E-8393-3311969AB761</uuid>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF9EFFC7FCFF817F06BAFAA2" lastPageId="20" lastPageNumber="1265" pageId="19" pageNumber="1264" type="diagnosis">
<paragraph id="6A483699FF9EFFC0FCFF817F02E3F8CE" blockId="19.[827,1443,1624,1891]" pageId="19" pageNumber="1264">
<emphasis id="5883EA8BFF9EFFC0FCFF817F0483F9C0" box="[827,963,1624,1645]" italics="true" pageId="19" pageNumber="1264">Diagnosis:</emphasis>
Males and females both have twosegmented fore tarsi with paired, movable pretarsal claws (
<figureCitation id="F2CC2A1CFF9EFFC0FC4F81B2049AF907" box="[907,986,1685,1707]" captionStart="Figure 4" captionStartId="9.[163,249,1379,1401]" captionTargetBox="[163,781,195,1339]" captionTargetId="figure-575@9.[163,781,195,1339]" captionTargetPageId="9" captionText="Figure 4. Foretibiae and foretarsi of Laccocorinae: Ctenipocoris asiaticus (a), Heleolaccocoris ovatus (b), Namtokocoris dalanta (c)." figureDoi="http://doi.org/10.5281/zenodo.6994615" httpUri="https://zenodo.org/record/6994615/files/figure.png" pageId="19" pageNumber="1264">Fig. 4a</figureCitation>
), which is the plesiomorphic condition. The nymphs have two meso- and metatarsal segments, another plesiomorphic condition, which distinguishes this tribe from the other tribes in
<taxonomicName id="ADF74D1AFF9EFFC0FB7B81D60214F8AA" authorityName="STAL" authorityYear="1876" box="[1215,1364,1777,1799]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
, which each have only a single segment. Adults have stout spines on the meso- and metatibiae. The aedeagus and parameres are symmetrical, which along with the
</paragraph>
<paragraph id="6A483699FF99FFC7FD3187A302D0FF36" blockId="20.[757,1424,128,155]" box="[757,1424,128,155]" pageId="20" pageNumber="1265">
PHYLOGENY OF
<taxonomicName id="ADF74D1AFF99FFC7FC0F87A3033CFF34" authorityName="LEACH" authorityYear="1815" box="[971,1148,132,153]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="family">NAUCORIDAE</taxonomicName>
(
<taxonomicName id="ADF74D1AFF99FFC7FB4E87A3026AFF34" box="[1162,1322,132,153]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Tracheophyta" rank="order">HEMIPTERA</taxonomicName>
) 1265
</paragraph>
<caption id="3E886611FF99FFC7FF55833806F7FBFF" ID-DOI="http://doi.org/10.5281/zenodo.6994647" ID-Zenodo-Dep="6994647" httpUri="https://zenodo.org/record/6994647/files/figure.png" pageId="20" pageNumber="1265" startId="20.[145,226,1055,1077]" targetBox="[145,1425,195,1015]" targetPageId="20">
<paragraph id="6A483699FF99FFC7FF55833806F7FBFF" blockId="20.[145,1424,1055,1107]" pageId="20" pageNumber="1265">
<emphasis id="5883EA8BFF99FFC7FF558338064BFB99" bold="true" box="[145,267,1055,1077]" pageId="20" pageNumber="1265">Figure 15.</emphasis>
Revised generic representation of the former genus
<taxonomicName id="ADF74D1AFF99FFC7FC9083380485FB98" authorityName="Stal" authorityYear="1856" box="[852,965,1055,1077]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FC9083380485FB98" box="[852,965,1055,1077]" italics="true" pageId="20" pageNumber="1265">Laccocoris</emphasis>
</taxonomicName>
. Dorsal habitus of
<taxonomicName id="ADF74D1AFF99FFC7FB5D83380226FB98" baseAuthorityName="Stal" baseAuthorityYear="1856" box="[1177,1382,1055,1077]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="species" species="spurcus">
<emphasis id="5883EA8BFF99FFC7FB5D83380226FB98" box="[1177,1382,1055,1077]" italics="true" pageId="20" pageNumber="1265">Laccocoris spurcus</emphasis>
</taxonomicName>
(a),
<taxonomicName id="ADF74D1AFF99FFC7FF55831A06CFFBFE" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1897" box="[145,399,1085,1107]" class="Insecta" family="Naucoridae" genus="Heleolaccocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="species" species="horvathi">
<emphasis id="5883EA8BFF99FFC7FF55831A06CFFBFE" box="[145,399,1085,1107]" italics="true" pageId="20" pageNumber="1265">Heleolaccocoris horvathi</emphasis>
</taxonomicName>
(b).
</paragraph>
</caption>
<paragraph id="6A483699FF99FFC7FF55835806BAFAA2" blockId="20.[145,760,1151,1295]" pageId="20" pageNumber="1265">pygophores, are dramatically different between species from the Old and New Worlds, and possibly also within each hemisphere. The body shape tends to be slightly wider at the middle and the fore femora less robust than in most other saucer bugs.</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF99FFC7FF55821E03C4FA64" pageId="20" pageNumber="1265" type="discussion">
<paragraph id="6A483699FF99FFC7FF55821E0502F9CF" blockId="20.[145,762,1336,1910]" pageId="20" pageNumber="1265">
<emphasis id="5883EA8BFF99FFC7FF55821E064CFAE3" box="[145,268,1337,1358]" italics="true" pageId="20" pageNumber="1265">Comments:</emphasis>
The two analyses yielded different topologies in the relationship of
<taxonomicName id="ADF74D1AFF99FFC7FEBF82700544FAC1" authorityName="Montandon" authorityYear="1897" box="[379,516,1367,1388]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FEBF82700544FAC1" box="[379,516,1367,1388]" italics="true" pageId="20" pageNumber="1265">Ctenipocoris</emphasis>
</taxonomicName>
with the remainder of the subfamily. The BI analysis recovered
<taxonomicName id="ADF74D1AFF99FFC7FDAF825105B8FA26" authorityName="Montandon" authorityYear="1897" box="[619,760,1398,1419]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FDAF825105B8FA26" box="[619,760,1398,1419]" italics="true" pageId="20" pageNumber="1265">Ctenipocoris</emphasis>
</taxonomicName>
as sister to the remainder of
<taxonomicName id="ADF74D1AFF99FFC7FE1A82B30532FA07" authorityName="STAL" authorityYear="1876" box="[478,626,1428,1450]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
and is here recognized also as the new tribe
<taxonomicName id="ADF74D1AFF99FFC7FDD982940580FA64" authorityName="Sites" authorityYear="2022" box="[541,704,1459,1481]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="tribe" tribe="Ctenipocorini">Ctenipocorini</taxonomicName>
. The hypothesis in ML is less likely from the standpoint of character evolution, because the alternative character state of a suite of characters descriptive of
<taxonomicName id="ADF74D1AFF99FFC7FDAD812805B9F988" authorityName="STAL" authorityYear="1876" box="[617,761,1551,1573]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1285" phylum="Arthropoda" rank="subFamily" subFamily="Laccocorinae">Laccocorinae</taxonomicName>
would have to have convergently evolved multiple times in other clades to support this topology.
</paragraph>
<paragraph id="6A483699FF99FFC7FF6D814C04BBFAA2" blockId="20.[145,762,1336,1910]" lastBlockId="20.[809,1425,1151,1295]" pageId="20" pageNumber="1265">
Although
<taxonomicName id="ADF74D1AFF99FFC7FED8814C06EBF92D" authorityName="Montandon" authorityYear="1897" box="[284,427,1643,1664]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FED8814C06EBF92D" box="[284,427,1643,1664]" italics="true" pageId="20" pageNumber="1265">Ctenipocoris</emphasis>
</taxonomicName>
is essentially circumtropical in lentic habitats, it is uncommonly collected. This widespread, disjunct distribution with few species and uncommon occurrence suggests that
<taxonomicName id="ADF74D1AFF99FFC7FDAF81E005B8F971" authorityName="Montandon" authorityYear="1897" box="[619,760,1735,1756]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FDAF81E005B8F971" box="[619,760,1735,1756]" italics="true" pageId="20" pageNumber="1265">Ctenipocoris</emphasis>
</taxonomicName>
might have previously been more diverse and was outcompeted by other taxa, such as
<taxonomicName id="ADF74D1AFF99FFC7FD85802305EAF8B4" baseAuthorityName="La Rivers" baseAuthorityYear="1956" box="[577,682,1796,1817]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FD85802305EAF8B4" box="[577,682,1796,1817]" italics="true" pageId="20" pageNumber="1265">Pelocoris</emphasis>
</taxonomicName>
in the New World and species of
<taxonomicName id="ADF74D1AFF99FFC7FE7680040576F894" authorityName="LEACH" authorityYear="1815" box="[434,566,1827,1849]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
in the Old World. In a treatment of
<taxonomicName id="ADF74D1AFF99FFC7FE9C806605B5F8FA" authority="Stal, 1854" authorityName="Stal" authorityYear="1854" box="[344,757,1857,1879]" class="Insecta" family="Nepidae" genus="Telmatotrephes" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FE9C80660545F8FB" box="[344,517,1857,1878]" italics="true" pageId="20" pageNumber="1265">Telmatotrephes</emphasis>
<bibRefCitation id="0E664B68FF99FFC7FDC88066053EF8FA" author="Stal C" box="[524,638,1857,1879]" pageId="20" pageNumber="1265" pagination="231 - 235" refId="ref24908" refString="Stal C. 1854. Nya Hemiptera. Ofversigt af Kongliga Svenska Vetenskaps-Akademiens Forhandlingar, Stockholm 11: 231 - 235." type="journal article" year="1854">Stål, 1854</bibRefCitation>
(Nepidae)
</taxonomicName>
, which similarly is distributed in both hemispheres and represented by relatively few species,
<bibRefCitation id="0E664B68FF99FFC7FB11835802D1FB39" author="Lansbury I" box="[1237,1425,1151,1173]" pageId="20" pageNumber="1265" pagination="271 - 286" refId="ref22220" refString="Lansbury I. 1972. A revision of the genus Telmatotrephes Stal (Hemiptera-Heteroptera, Nepidae). Zoologica Scripta 1: 271 - 286." type="journal article" year="1972">Lansbury (1972)</bibRefCitation>
also considered the discontinuous distribution to be evidence of a relict group in decline. A key to identify the Neotropical species of
<taxonomicName id="ADF74D1AFF99FFC7FB9383FC03A5FB5D" authorityName="Montandon" authorityYear="1897" box="[1111,1253,1243,1264]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FB9383FC03A5FB5D" box="[1111,1253,1243,1264]" italics="true" pageId="20" pageNumber="1265">Ctenipocoris</emphasis>
</taxonomicName>
was presented by
<bibRefCitation id="0E664B68FF99FFC7FC8F83DD04B5FAA2" author="Herrera F" box="[843,1013,1274,1295]" pageId="20" pageNumber="1265" pagination="338 - 344" refId="ref22045" refString="Herrera F. 2013. First record of the genus Ctenipocoris (Heteroptera: Naucoridae) in Central America, with a preliminary key to the American species and description of a new species. Zootaxa 3731: 338 - 344." type="journal article" year="2013">Herrera (2013)</bibRefCitation>
.
</paragraph>
<paragraph id="6A483699FF99FFC7FCED821F03C4FA64" blockId="20.[809,1425,1336,1481]" pageId="20" pageNumber="1265">
<emphasis id="5883EA8BFF99FFC7FCED821F0493FAE3" box="[809,979,1336,1358]" italics="true" pageId="20" pageNumber="1265">Included taxa:</emphasis>
This tribe is represented by only the genus
<taxonomicName id="ADF74D1AFF99FFC7FCB082700342FAC1" authorityName="Montandon" authorityYear="1897" box="[884,1026,1367,1388]" class="Insecta" family="Naucoridae" genus="Ctenipocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="1265" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF99FFC7FCB082700342FAC1" box="[884,1026,1367,1388]" italics="true" pageId="20" pageNumber="1265">Ctenipocoris</emphasis>
</taxonomicName>
and eight described species. A few additional undescribed species are represented in the Enns Entomology Museum, University of
<collectingRegion id="A833F87BFF99FFC7FAE682B302CCFA07" box="[1314,1420,1428,1450]" country="United States of America" name="Missouri" pageId="20" pageNumber="1265">Missouri</collectingRegion>
, and other museum collections.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="8EB54384972183A748F362C8EE8B4621" ID-CLB-Dataset="121760" ID-DOI="10.1093/zoolinnean/zlab105" ID-GBIF-Dataset="83d946c2-fa80-4948-9160-16625c240886" ID-ISSN="0024-4082" ID-Zenodo-Dep="6994599" ID-ZooBank="C06A1F94-AF08-4A21-B1F3-A0865FB1A8DF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660324512590" checkinUser="plazi" docAuthor="Sites, Robert W." docDate="2022" docId="E25E878FFFADFFF1FECF81A90645FE9A" docLanguage="en" docName="zlab105.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Australonaucoris Sites 2022, GEN. NOV." docType="treatment" docVersion="6" lastPageNumber="1279" masterDocId="1E67FFF7FF8DFFD3FFC487270740FFAD" masterDocTitle="Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)" masterLastPageNumber="1286" masterPageNumber="1245" pageNumber="1277" updateTime="1699366499196" updateUser="ExternalLinkService">
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<mods:title id="94E2FB609605122A4EF15AC6064C148E">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:date id="AB7CF4628942E1EBA3A71F2C26C964D9">2022</mods:date>
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<treatment id="E25E878FFFADFFF1FECF81A90645FE9A" ID-DOI="http://doi.org/10.5281/zenodo.6994717" ID-GBIF-Taxon="199315254" ID-Zenodo-Dep="6994717" LSID="urn:lsid:plazi:treatment:E25E878FFFADFFF1FECF81A90645FE9A" httpUri="http://treatment.plazi.org/id/E25E878FFFADFFF1FECF81A90645FE9A" lastPageId="34" lastPageNumber="1279" pageId="32" pageNumber="1277">
<subSubSection id="22ED6512FFADFFF3FECF81A906AEF97E" pageId="32" pageNumber="1277" type="nomenclature">
<paragraph id="6A483699FFADFFF3FECF81A9053FF90A" blockId="32.[267,639,1678,1703]" box="[267,639,1678,1703]" pageId="32" pageNumber="1277">
<heading id="310081F5FFADFFF3FECF81A9053FF90A" box="[267,639,1678,1703]" centered="true" fontSize="9" level="2" pageId="32" pageNumber="1277" reason="2">
<taxonomicName id="ADF74D1AFFADFFF3FECF81A90541F908" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[267,513,1678,1702]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="genus" status="gen. nov.">AUSTRALONAUCORIS</taxonomicName>
<taxonomicNameLabel id="43B057F0FFADFFF3FDCC81B6053FF90A" box="[520,639,1678,1703]" pageId="32" pageNumber="1277" rank="genus">
<emphasis id="5883EA8BFFADFFF3FDCC81B60538F909" bold="true" box="[520,632,1678,1702]" pageId="32" pageNumber="1277">GEN. NOV</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="6A483699FFADFFF3FE58819D06AEF97E" blockId="32.[412,494,1722,1747]" box="[412,494,1722,1747]" pageId="32" pageNumber="1277">
<figureCitation id="F2CC2A1CFFADFFF3FE58819D06AEF97E" box="[412,494,1722,1747]" captionStart="Figure 23" captionStartId="33.[163,241,986,1008]" captionTargetBox="[163,1444,195,946]" captionTargetId="figure-408@33.[163,1444,195,946]" captionTargetPageId="33" captionText="Figure 23. Australonaucoris congrex male: dorsal habitus (a), genital capsule with parameres and terga IX and X removed (b), left paramere (c), right paramere (d). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994670" httpUri="https://zenodo.org/record/6994670/files/figure.png" pageId="32" pageNumber="1277">FIG. 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FFADFFF2FF5581DC0209F8A4" lastPageId="33" lastPageNumber="1278" pageId="32" pageNumber="1277" type="description">
<paragraph id="6A483699FFADFFF3FF5581DC05A4F882" blockId="32.[145,761,1786,1839]" pageId="32" pageNumber="1277">
<emphasis id="5883EA8BFFADFFF3FF5581DC069DF8BD" box="[145,477,1786,1808]" italics="true" pageId="32" pageNumber="1277">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFFADFFF3FF1A803E05A4F882" box="[222,740,1817,1839]" pageId="32" pageNumber="1277">E165175A-EC63-4447-954B-EFE478B99A16</uuid>
</paragraph>
<paragraph id="6A483699FFADFFF3FF55807E0380FC8F" blockId="32.[145,734,1879,1902]" lastBlockId="32.[809,1425,197,1905]" pageId="32" pageNumber="1277">
<emphasis id="5883EA8BFFADFFF3FF55807E0661F8C0" box="[145,289,1880,1902]" italics="true" pageId="32" pageNumber="1277">
<typeStatus id="B54C883BFFADFFF3FF55807E0786F8C3" box="[145,198,1881,1902]" pageId="32" pageNumber="1277">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFFADFFF3FEE9807F0599F8C0" authority="(Stal, 1876)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Stal" baseAuthorityYear="1876" box="[301,729,1879,1902]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1277" phylum="Arthropoda" rank="species" species="congrex">
<emphasis id="5883EA8BFFADFFF3FEE9807F050DF8C3" box="[301,589,1880,1902]" italics="true" pageId="32" pageNumber="1277">Australonaucoris congrex</emphasis>
(
<bibRefCitation id="0E664B68FFADFFF3FD988070058EF8C3" author="Stal C" box="[604,718,1879,1902]" pageId="32" pageNumber="1277" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">Stål, 1876</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="5883EA8BFFADFFF3FCED87E10484FF76" box="[809,964,198,219]" italics="true" pageId="32" pageNumber="1277">Description:</emphasis>
Form ovate; widest at middle, the middle of embolia; moderately flattened for family. Head broad, about twice as wide as long, eyes and anterior margin continuously and evenly convex to anterolateral corners of pronotum. Eyes convergent anteriorly in anterior half, subparallel in posterior half; each eye narrowing anteriorly when viewed from above, anteroventral margin concave; narrow band of cuticle posterior and lateral to eye, widening slightly at anterolateral point. Labrum arising at front of head between inner margins of eyes; apex broadly rounded. Maxillary plates angled ventromedially, tips rounded, nearly attaining apex of labrum. Rostrum with three visible segments, with segments one and part of two concealed behind labrum. Antennae four segmented with segment 1 short and inconspicuous, 2 and 4 longer, 3 longest, segments 2–4 setose. Midventer of head with rounded, setose carina. Head and pronotum moderately flattened transversely.
</paragraph>
<paragraph id="6A483699FFADFFF3FC85840C042BFAE7" blockId="32.[809,1425,197,1905]" pageId="32" pageNumber="1277">Pronotum with anterior margin straight or slightly concave between eyes, then concavity becoming more pronounced as margin contours around eyes; lateral margins straight to slightly convex and convergent anteriorly; posterior margin mostly straight, with slight concavity medially and convexity laterally to angulate posterolateral corners. Scutellum triangular, approximately twice as wide as long, flattened, heavily punctate, lateral margins concave posteriorly and convex anteriorly. Hemelytra usually not covering tip of abdomen, leaving last segment exposed; narrowed such that lateral margins of abdominal terga are widely exposed. Embolium well defined with embolar suture, yellowish anteriorly, dark posteriorly; clavus well defined with claval and intraclaval sutures; corium, clavus and membrane punctate, darkly coloured, with sparse fine recumbent hairs.</paragraph>
<paragraph id="6A483699FFADFFF3FC858275048FF917" blockId="32.[809,1425,197,1905]" pageId="32" pageNumber="1277">Propleuron lateral to coxal cavity infuscated and setose; posterolaterally light coloured and glabrous; mesal margin subducting beneath (dorsad to) prosternellum. Probasisternum sharply carinate, carina continuous with rounded midventral carina of head. Mesosternum broadly carinate medially with a brush of long hairs and several isolated short, dark spines, which can be concealed by hairs. Metaxyphus with sharp median carina. Abdominal venter covered with long hairs except wide glabrous lateral band; laterosternites with paired, oval, glabrous depressions near spiracles.</paragraph>
<paragraph id="6A483699FFADFFF2FC8581E507B3FB52" blockId="32.[809,1425,197,1905]" lastBlockId="33.[163,779,1074,1647]" lastPageId="33" lastPageNumber="1278" pageId="32" pageNumber="1277">Profemur inflated, anterior margin with dorsal and ventral rows of golden hairs sandwiching two rows of short, dark spines. Protarsus single segmented with single, fixed, minute pretarsal claw. Meso- and metafemora with posteroventral row of short spines; those on mesofemur densely packed such that adjacent spines are touching or nearly touching; those on metafemur less densely packed, spaced approximately one spine width apart. Meso- and metatibiae with numerous rows of long dark red spines. Long golden swimming hairs sparse on mesotibia and tarsus, profuse on metatibia and tarsus.</paragraph>
<caption id="3E886611FFACFFF2FF6784FD0342FBA0" ID-DOI="http://doi.org/10.5281/zenodo.6994670" ID-Zenodo-Dep="6994670" httpUri="https://zenodo.org/record/6994670/files/figure.png" pageId="33" pageNumber="1278" startId="33.[163,241,986,1008]" targetBox="[163,1444,195,946]" targetPageId="33">
<paragraph id="6A483699FFACFFF2FF6784FD0342FBA0" blockId="33.[163,1442,986,1037]" pageId="33" pageNumber="1278">
<emphasis id="5883EA8BFFACFFF2FF6784FD0657FC42" bold="true" box="[163,279,986,1008]" pageId="33" pageNumber="1278">Figure 23.</emphasis>
<taxonomicName id="ADF74D1AFFACFFF2FEE684FD056AFC5D" authorityName="Sites" authorityYear="2022" baseAuthorityName="Stal" baseAuthorityYear="1876" box="[290,554,986,1008]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="congrex">
<emphasis id="5883EA8BFFACFFF2FEE684FD056AFC5D" box="[290,554,986,1008]" italics="true" pageId="33" pageNumber="1278">Australonaucoris congrex</emphasis>
</taxonomicName>
male: dorsal habitus (a), genital capsule with parameres and terga IX and X removed (b), left paramere (c), right paramere (d). Sizes are not proportional among figures.
</paragraph>
</caption>
<paragraph id="6A483699FFACFFF2FF7F822F0604F9C2" blockId="33.[163,779,1074,1647]" pageId="33" pageNumber="1278">Male genitalia with aedeagus long and slender, with right side angled apically and left side curving slightly to acuminate apex; parameres strongly asymmetrical, criss-crossing in basal two-thirds; left paramere overlapping right, broad, without pronounced lobe on left margin; right paramere slenderer, helically twisted, with elongate sulcus in distal fourth to embrace aedeagus; pygophore asymmetrical. Female mediosternite VII (subgenital plate) as wide as long, convex transversely, lateral margins with distinct concavity at middle, convergent toward broadly rounded apex.</paragraph>
<paragraph id="6A483699FFACFFF2FF6781A9044CF96F" blockId="33.[163,780,1677,1730]" pageId="33" pageNumber="1278">
<emphasis id="5883EA8BFFACFFF2FF6781A9065AF90E" box="[163,282,1678,1699]" italics="true" pageId="33" pageNumber="1278">Diagnosis:</emphasis>
The left paramere is without a pronounced lobe on the left side and the pygophore is asymmetrical.
</paragraph>
<paragraph id="6A483699FFACFFF2FF6781FC06C2F8C6" blockId="33.[163,779,1755,1899]" pageId="33" pageNumber="1278">
<emphasis id="5883EA8BFFACFFF2FF6781FC0661F95D" box="[163,289,1755,1776]" italics="true" pageId="33" pageNumber="1278">Etymology:</emphasis>
The genus name recognizes the Australian region in which these former members of
<taxonomicName id="ADF74D1AFFACFFF2FD5B81DD044BF8A2" authorityName="Geoffroy" authorityYear="1762" box="[671,779,1786,1807]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFACFFF2FD5B81DD044BF8A2" box="[671,779,1786,1807]" italics="true" pageId="33" pageNumber="1278">Naucoris</emphasis>
</taxonomicName>
occur, while still referencing the previous generic association. The prefix is derived from the Latin
<emphasis id="5883EA8BFFACFFF2FF6780720648F8C7" box="[163,264,1877,1898]" italics="true" pageId="33" pageNumber="1278">australis</emphasis>
, southern.
</paragraph>
<paragraph id="6A483699FFACFFF2FCFF83150209F8A4" blockId="33.[827,1445,1073,1892]" pageId="33" pageNumber="1278">
<emphasis id="5883EA8BFFACFFF2FCFF831504FDFBEA" box="[827,957,1074,1095]" italics="true" pageId="33" pageNumber="1278">Comments:</emphasis>
The left paramere is diagnostic for this former member of
<taxonomicName id="ADF74D1AFFACFFF2FBE0837603D2FBCB" authorityName="Geoffroy" authorityYear="1762" box="[1060,1170,1105,1126]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFACFFF2FBE0837603D2FBCB" box="[1060,1170,1105,1126]" italics="true" pageId="33" pageNumber="1278">Naucoris</emphasis>
</taxonomicName>
. Among other species of
<taxonomicName id="ADF74D1AFFACFFF2FC998348048FFB29" authorityName="Geoffroy" authorityYear="1762" box="[861,975,1135,1156]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFACFFF2FC998348048FFB29" box="[861,975,1135,1156]" italics="true" pageId="33" pageNumber="1278">Naucoris</emphasis>
</taxonomicName>
from
<collectingCountry id="12E07609FFACFFF2FBE5834803DCFB28" box="[1057,1180,1135,1157]" name="Australia" pageId="33" pageNumber="1278">Australia</collectingCountry>
, figures included in the original description of
<taxonomicName id="ADF74D1AFFACFFF2FB5683A90277FB0F" authorityName="Lansbury" authorityYear="1985" box="[1170,1335,1165,1187]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="subaureus">
<emphasis id="5883EA8BFFACFFF2FB5683A90277FB0F" box="[1170,1335,1165,1187]" italics="true" pageId="33" pageNumber="1278">N. subaureus</emphasis>
</taxonomicName>
and the redescription of
<taxonomicName id="ADF74D1AFFACFFF2FBC6838A03E0FB6C" authorityName="Montandon" authorityYear="1913" box="[1026,1184,1196,1218]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="5883EA8BFFACFFF2FBC6838A03E0FB6C" box="[1026,1184,1196,1218]" italics="true" pageId="33" pageNumber="1278">N. subopacus</emphasis>
</taxonomicName>
clearly show the left parameres each without a lobe on the left side and pygophore asymmetrical (Lansbury, 1985); thus, these species are transferred to
<taxonomicName id="ADF74D1AFFACFFF2FB9B822F0262FAB0" authorityName="Sites" authorityYear="2022" box="[1119,1314,1288,1309]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFACFFF2FB9B822F0262FAB0" box="[1119,1314,1288,1309]" italics="true" pageId="33" pageNumber="1278">Australonaucoris</emphasis>
</taxonomicName>
. Validation for the transfer of
<taxonomicName id="ADF74D1AFFACFFF2FBC9820003E1FA91" authorityName="Montandon" authorityYear="1913" box="[1037,1185,1319,1340]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="5883EA8BFFACFFF2FBC9820003E1FA91" box="[1037,1185,1319,1340]" italics="true" pageId="33" pageNumber="1278">N. subopacus</emphasis>
</taxonomicName>
was provided recently whereby it was recovered as sister to the clade with the New
<collectingCountry id="12E07609FFACFFF2FC6882430344FAD7" box="[940,1028,1380,1402]" name="Guinea-Bissau" pageId="33" pageNumber="1278">Guinea</collectingCountry>
endemic genera (Ye
<emphasis id="5883EA8BFFACFFF2FACD82420208FAD4" box="[1289,1352,1380,1402]" italics="true" pageId="33" pageNumber="1278">et al.</emphasis>
, 2020). The parameres of
<taxonomicName id="ADF74D1AFFACFFF2FBDC82A403D0FA35" authorityName="Lansbury" authorityYear="1991" box="[1048,1168,1411,1432]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="magela">
<emphasis id="5883EA8BFFACFFF2FBDC82A403D0FA35" box="[1048,1168,1411,1432]" italics="true" pageId="33" pageNumber="1278">N. magela</emphasis>
</taxonomicName>
were illustrated twice (figs 19, 20) by
<bibRefCitation id="0E664B68FFACFFF2FBCC82860399FA1A" author="Lansbury I" box="[1032,1241,1441,1463]" pageId="33" pageNumber="1278" pagination="103 - 114" refId="ref22277" refString="Lansbury I. 1991. Naucoridae and Notonectidae (HemipteraHeteroptera) of the Northern Territory, Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences 8: 103 - 114." type="journal article" year="1991">Lansbury (1991)</bibRefCitation>
. Although each illustration shows a dramatically different shape of the left paramere that cannot be resolved by considering viewing angles, neither illustration shows a pronounced lobe on the left side, suggesting that
<taxonomicName id="ADF74D1AFFACFFF2FA4E813A04CEF9FD" authorityName="Lansbury" authorityYear="1991" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="magela">
<emphasis id="5883EA8BFFACFFF2FA4E813A04CEF9FD" italics="true" pageId="33" pageNumber="1278">N. magela</emphasis>
</taxonomicName>
also should be transferred to
<taxonomicName id="ADF74D1AFFACFFF2FB1E811C02DDF9FD" authorityName="Sites" authorityYear="2022" box="[1242,1437,1595,1616]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFACFFF2FB1E811C02DDF9FD" box="[1242,1437,1595,1616]" italics="true" pageId="33" pageNumber="1278">Australonaucoris</emphasis>
</taxonomicName>
; thus, this species is provisionally transferred pending confirmation of paramere condition. The parameres of
<taxonomicName id="ADF74D1AFFACFFF2FC9D81B00340F901" authorityName="Stal" authorityYear="1876" box="[857,1024,1687,1708]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="australicus">
<emphasis id="5883EA8BFFACFFF2FC9D81B00340F901" box="[857,1024,1687,1708]" italics="true" pageId="33" pageNumber="1278">N. australicus</emphasis>
</taxonomicName>
have not been illustrated, so until specimens have been examined for paramere shape and other features, this species remains
<emphasis id="5883EA8BFFACFFF2FAC081F202DDF944" box="[1284,1437,1748,1770]" italics="true" pageId="33" pageNumber="1278">incertae sedis</emphasis>
. The following new combinations are proposed:
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FFACFFF1FC9780360645FE9A" lastPageId="34" lastPageNumber="1279" pageId="33" pageNumber="1278" type="reference_group">
<paragraph id="6A483699FFACFFF2FC9780360238F88A" blockId="33.[827,1445,1073,1892]" box="[851,1400,1809,1831]" pageId="33" pageNumber="1278">
<taxonomicName id="ADF74D1AFFACFFF2FC97803603BCF88B" authority="(Stal, 1876)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Stal" baseAuthorityYear="1876" box="[851,1276,1809,1831]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="congrex" status="comb. nov.">
<emphasis id="5883EA8BFFACFFF2FC9780360333F88A" box="[851,1139,1809,1831]" italics="true" pageId="33" pageNumber="1278">Australonaucoris congrex</emphasis>
(
<bibRefCitation id="0E664B68FFACFFF2FB46803603B5F88A" author="Stal C" box="[1154,1269,1809,1831]" pageId="33" pageNumber="1278" pagination="1 - 167" refId="ref25070" refString="Stal C. 1876. Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda. Hemiptera, jemte systematiska meddelanden. Kongliga Svenska VetenskapsAkademiens Handlingar 14: 1 - 167." type="journal article" year="1876">Stål, 1876</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFACFFF2FAC780360238F88A" box="[1283,1400,1809,1831]" pageId="33" pageNumber="1278" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFACFFF2FC9780170428F8C9" blockId="33.[827,1445,1073,1892]" pageId="33" pageNumber="1278">
<taxonomicName id="ADF74D1AFFACFFF2FC9780170216F8E8" authority="(Lansbury, 1991)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Lansbury" baseAuthorityYear="1991" box="[851,1366,1840,1862]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="1278" phylum="Arthropoda" rank="species" species="magela" status="comb. nov.">
<emphasis id="5883EA8BFFACFFF2FC97801703C1F8E8" box="[851,1153,1840,1861]" italics="true" pageId="33" pageNumber="1278">Australonaucoris magela</emphasis>
(
<bibRefCitation id="0E664B68FFACFFF2FB508017020DF8E8" author="Lansbury I" box="[1172,1357,1840,1862]" pageId="33" pageNumber="1278" pagination="103 - 114" refId="ref22277" refString="Lansbury I. 1991. Naucoridae and Notonectidae (HemipteraHeteroptera) of the Northern Territory, Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences 8: 103 - 114." type="journal article" year="1991">Lansbury, 1991</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFACFFF2FA9B80170428F8C9" pageId="33" pageNumber="1278" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFAFFFF1FF6D87E207FEFF54" blockId="34.[145,762,197,311]" pageId="34" pageNumber="1279">
<taxonomicName id="ADF74D1AFFAFFFF1FF6D87E205F1FF77" authority="(Lansbury, 1985)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Lansbury" baseAuthorityYear="1985" box="[169,689,197,219]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="1279" phylum="Arthropoda" rank="species" species="subaureus" status="comb. nov.">
<emphasis id="5883EA8BFFAFFFF1FF6D87E206A9FF77" box="[169,489,197,218]" italics="true" pageId="34" pageNumber="1279">Australonaucoris subaureus</emphasis>
(Lansbury, 1985)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFAFFFF1FD7C87E207FEFF54" pageId="34" pageNumber="1279" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6A483699FFAFFFF1FF6D86250645FE9A" blockId="34.[145,762,197,311]" pageId="34" pageNumber="1279">
<taxonomicName id="ADF74D1AFFAFFFF1FF6D862505B9FEB5" authority="(Montandon, 1913)" authorityName="Sites" authorityYear="2022" baseAuthorityName="Montandon" baseAuthorityYear="1913" box="[169,761,258,280]" class="Insecta" family="Naucoridae" genus="Australonaucoris" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="1279" phylum="Arthropoda" rank="species" species="subopacus" status="comb. nov.">
<emphasis id="5883EA8BFFAFFFF1FF6D86250545FEBA" box="[169,517,258,279]" italics="true" pageId="34" pageNumber="1279">Australonaucoris subopacus</emphasis>
(
<bibRefCitation id="0E664B68FFAFFFF1FDDD862505B0FEB5" author="Montandon AL" box="[537,752,258,280]" pageId="34" pageNumber="1279" pagination="219 - 224" refId="ref23328" refString="Montandon AL. 1913. Hemipteres aquatiques. Notes et descriptions de deux especes nouvelles. Bulletin de la Section Scientifique de l'Academie Roumaine 1: 219 - 224." type="journal article" year="1913">Montandon, 1913</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFAFFFF1FF5586060645FE9A" box="[145,261,289,311]" pageId="34" pageNumber="1279" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

108
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<mods:title id="72BA6DE5DDAC7948AF83E00FEC5A1842">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<subSubSection id="22ED6512FFAFFFF1FEEF86BD051FFE1F" box="[299,607,410,434]" pageId="34" pageNumber="1279" type="nomenclature">
<paragraph id="6A483699FFAFFFF1FEEF86BD051FFE1F" blockId="34.[299,607,410,434]" box="[299,607,410,434]" pageId="34" pageNumber="1279">
<heading id="310081F5FFAFFFF1FEEF86BD051FFE1F" box="[299,607,410,434]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="1279" reason="2">
<taxonomicName id="ADF74D1AFFAFFFF1FEEF86BD06A1FE1C" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[299,481,410,433]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="1279" phylum="Arthropoda" rank="genus" status="gen. nov.">INDONAUCORIS</taxonomicName>
<emphasis id="5883EA8BFFAFFFF1FE2C86BA051FFE1F" bold="true" box="[488,607,410,434]" pageId="34" pageNumber="1279">
<taxonomicNameLabel id="43B057F0FFAFFFF1FE2C86BA051FFE1F" box="[488,607,410,434]" pageId="34" pageNumber="1279" rank="genus">GEN. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FFAFFFF1FE4086E105B8FDE5" pageId="34" pageNumber="1279" type="description">
<paragraph id="6A483699FFAFFFF1FE4086E10546FE72" blockId="34.[388,518,454,479]" box="[388,518,454,479]" pageId="34" pageNumber="1279">
<figureCitation id="F2CC2A1CFFAFFFF1FE4086E1069CFE72" box="[388,476,454,479]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="34" pageNumber="1279">FIGS 24</figureCitation>
,
<figureCitation id="F2CC2A1CFFAFFFF1FE2E86E10546FE72" box="[490,518,454,479]" captionStart="Figure 25" captionStartId="36.[144,224,1075,1097]" captionTargetBox="[224,1344,195,1035]" captionTargetId="figure-24@36.[224,1344,195,1035]" captionTargetPageId="36" captionText="Figure 25. Tamhini stream, the presumed type locality of Indonaucoris minutus gen. nov., sp. nov., in December 2019. Photo courtesy of Shruti Paripatyadar." figureDoi="http://doi.org/10.5281/zenodo.6994676" httpUri="https://zenodo.org/record/6994676/files/figure.png" pageId="34" pageNumber="1279">25</figureCitation>
</paragraph>
<paragraph id="6A483699FFAFFFF1FF55853205B8FDE5" blockId="34.[145,761,532,584]" pageId="34" pageNumber="1279">
<emphasis id="5883EA8BFFAFFFF1FF558532069DFD87" box="[145,477,532,554]" italics="true" pageId="34" pageNumber="1279">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="1E510C4CFFAFFFF1FF25851505B8FDE5" box="[225,760,562,584]" pageId="34" pageNumber="1279">1E965D6D-14C1-4AE0-9A6E-C7D19EE8E41E</uuid>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FFAFFFF1FF55855505C3FD2A" box="[145,643,625,647]" pageId="34" pageNumber="1279" type="type_taxon">
<paragraph id="6A483699FFAFFFF1FF55855505C3FD2A" blockId="34.[145,643,625,647]" box="[145,643,625,647]" pageId="34" pageNumber="1279">
<emphasis id="5883EA8BFFAFFFF1FF5585550569FD2A" box="[145,553,625,647]" italics="true" pageId="34" pageNumber="1279">
<typeStatus id="B54C883BFFAFFFF1FF5585550786FD2A" box="[145,198,626,647]" pageId="34" pageNumber="1279">Type</typeStatus>
species:
<taxonomicName id="ADF74D1AFFAFFFF1FEE985560569FD2A" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[301,553,625,647]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="1279" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">Indonaucoris minutus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="43B057F0FFAFFFF1FDF4855505C3FD2A" box="[560,643,626,647]" pageId="34" pageNumber="1279" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FFAFFFF1FF5585970344FBB5" pageId="34" pageNumber="1279" type="description">
<paragraph id="6A483699FFAFFFF1FF5585970520FD49" blockId="34.[145,762,688,1906]" pageId="34" pageNumber="1279">
<emphasis id="5883EA8BFFAFFFF1FF558597065AFD68" box="[145,282,688,709]" italics="true" pageId="34" pageNumber="1279">Description:</emphasis>
Form nearly ovate; widest at middle, the embolia; moderately flattened for family.
</paragraph>
<paragraph id="6A483699FFAFFFF1FF6D85CA0546FB43" blockId="34.[145,762,688,1906]" pageId="34" pageNumber="1279">Head broad, eyes and anterior margin evenly convex to anterolateral corners of pronotum. Anterior margin of head with sparse row of elongate, lightcoloured setae. Eyes convergent anteriorly, angle of convergence subtly becoming greater in anterior half; anteroventral margin concave; narrow band of cuticle lateral to eye. Labrum arising at front of head between inner margins of eyes; apex broadly rounded. Maxillary plates angled ventromedially, tips almost truncate, but broadly rounded, not exceeding apex of labrum. Rostrum with three visible segments, with segments 1 and part of 2 concealed behind labrum. Antennae four segmented with segment 1 short and inconspicuous, 2 widest, 2–4 subequal in length. Midventer of head with anterior half roundedly carinate, posterior half flat. Head and pronotum moderately convex transversely.</paragraph>
<paragraph id="6A483699FFAFFFF1FF6D83D005BAF95B" blockId="34.[145,762,688,1906]" pageId="34" pageNumber="1279">Pronotum with anterior margin straight or slightly concave between eyes, then concavity becoming more pronounced as margin contours around eyes; lateral margins convex and convergent anteriorly; posterior margin mostly straight, with slight concavities medially and midlaterally before rounded posterolateral corners; sparse long hairs near lateral margin. Scutellum large, triangular, slightly wider than long, evenly domed, heavily punctate. Hemelytra extending to tip of abdomen; embolium well defined with embolar suture, yellowish anteriorly, darker posteriorly; clavus well defined with claval and intraclaval sutures; corium, clavus and membrane punctate and with subtle mottling; sparse long hairs evident on scutellum, clavus and corium, hairs not evident on membrane; a single light spot at middle of corium/membrane interface. Hindwings macropterous.</paragraph>
<paragraph id="6A483699FFAFFFF1FF6D80270365FE7D" blockId="34.[145,762,688,1906]" lastBlockId="34.[809,1425,197,985]" pageId="34" pageNumber="1279">Propleuron lateral to coxal cavity infuscated and setose; posterolaterally light coloured and glabrous; mesal margin roundedly triangular and deflexed at tip, suggesting that mesal margin would subduct beneath prosternellum in intact specimen. Probasisternum sharply carinate, carina widening posteriorly and apparently not continuous with midventral carina of head. Mesosternum on right side suggests at most a broad tumescence medially, left side missing. Metasternum missing entirely. Abdominal venter thickly covered with long hairs; laterosternites with paired, oval, glabrous depressions near spiracles. Wide glabrous lateral band.</paragraph>
<paragraph id="6A483699FFAFFFF1FC8586FE03BAFCF3" blockId="34.[809,1425,197,985]" pageId="34" pageNumber="1279">Profemur inflated, anterior margin with ventral row of golden hairs beneath two rows of short, dark spines; dorsal row of golden hairs imperceptible. Protarsus single-segmented with single, fixed, minute pretarsal claw. Meso- and metafemora with posteroventral row of short spines; those on mesofemur densely packed such that adjacent spines are touching or nearly touching, but with occasional interruptions; those on metafemur less densely packed, spaced approximately one spine width apart. Meso- and metatibiae with numerous rows of long dark red spines. Long golden swimming hairs imperceptible on mesotibia and tarsus, profuse on metatibia and tarsus.</paragraph>
<paragraph id="6A483699FFAFFFF1FC8584400337FC74" blockId="34.[809,1425,197,985]" pageId="34" pageNumber="1279">Aedeagus long and slender; parameres asymmetrical, criss-crossing; left paramere overlapping right, with pronounced lobe on left margin; right paramere with elongate sulcus in distal half.</paragraph>
<paragraph id="6A483699FFAFFFF1FCED83250344FBB5" blockId="34.[809,1028,1026,1048]" box="[809,1028,1026,1048]" pageId="34" pageNumber="1279">
<heading id="310081F5FFAFFFF1FCED83250344FBB5" box="[809,1028,1026,1048]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="1279" reason="2">
<emphasis id="5883EA8BFFAFFFF1FCED832504C1FBBA" box="[809,897,1026,1047]" italics="true" pageId="34" pageNumber="1279">Female:</emphasis>
Unknown.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FFAFFFF1FCED83660394FB1E" pageId="34" pageNumber="1279" type="etymology">
<paragraph id="6A483699FFAFFFF1FCED83660394FB1E" blockId="34.[809,1425,1089,1203]" pageId="34" pageNumber="1279">
<emphasis id="5883EA8BFFAFFFF1FCED836604EBFBFB" box="[809,939,1089,1110]" italics="true" pageId="34" pageNumber="1279">Etymology:</emphasis>
The genus name references the Indian region in which this insect was collected and the genus
<taxonomicName id="ADF74D1AFFAFFFF1FCED835804D6FB39" authorityName="Geoffroy" authorityYear="1762" box="[809,918,1151,1172]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="1279" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFAFFFF1FCED835804D6FB39" box="[809,918,1151,1172]" italics="true" pageId="34" pageNumber="1279">Naucoris</emphasis>
</taxonomicName>
which it closely resembles. The prefix is derived from the Latin
<emphasis id="5883EA8BFFAFFFF1FBF683BA0334FB1F" box="[1074,1140,1181,1202]" italics="true" pageId="34" pageNumber="1279">Indus</emphasis>
, Indian.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FFAFFFF1FCED83FB027AF9EE" pageId="34" pageNumber="1279" type="discussion">
<paragraph id="6A483699FFAFFFF1FCED83FB027AF9EE" blockId="34.[809,1425,1243,1603]" pageId="34" pageNumber="1279">
<emphasis id="5883EA8BFFAFFFF1FCED83FB04EBFB5C" box="[809,939,1244,1265]" italics="true" pageId="34" pageNumber="1279">Comments:</emphasis>
The single male specimen was dissected so that thoracic musculature could be extracted and used for the molecular analyses. Thus, it has incurred significant meso- and metasternal damage. The connection between the prothorax and mesothorax has been substantially weakened. Nonetheless, most other structures, including the genital capsule, are intact and five legs are available. Because of the condition of the specimen, it is preserved in 80% etOH. It would not pin or point well, and if card-mounted, future removal from the card to examine ventral features would likely cause further damage to this fragile specimen.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="9A1CB757959E27BB41B8283D4AD3E166" ID-CLB-Dataset="121760" ID-DOI="10.1093/zoolinnean/zlab105" ID-GBIF-Dataset="83d946c2-fa80-4948-9160-16625c240886" ID-ISSN="0024-4082" ID-Zenodo-Dep="6994599" ID-ZooBank="C06A1F94-AF08-4A21-B1F3-A0865FB1A8DF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660324512590" checkinUser="plazi" docAuthor="Sites, Robert W." docDate="2022" docId="E25E878FFFAFFFF7FC57815A0363FACC" docLanguage="en" docName="zlab105.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Indonaucoris minutus Sites 2022, SP. NOV." docType="treatment" docVersion="7" lastPageNumber="1281" masterDocId="1E67FFF7FF8DFFD3FFC487270740FFAD" masterDocTitle="Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)" masterLastPageNumber="1286" masterPageNumber="1245" pageNumber="1279" updateTime="1699366499196" updateUser="ExternalLinkService">
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<mods:title id="23D97EA6ED930C604EC42AF539ED6555">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:namePart id="3CCFE2595883759910ED0F93BB8F4BCE">Sites, Robert W.</mods:namePart>
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<mods:date id="81ACB0AE465267DC1FB77578DA466400">2022</mods:date>
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<treatment id="E25E878FFFAFFFF7FC57815A0363FACC" ID-DOI="http://doi.org/10.5281/zenodo.6994688" ID-GBIF-Taxon="199315253" ID-Zenodo-Dep="6994688" LSID="urn:lsid:plazi:treatment:E25E878FFFAFFFF7FC57815A0363FACC" httpUri="http://treatment.plazi.org/id/E25E878FFFAFFFF7FC57815A0363FACC" lastPageId="36" lastPageNumber="1281" pageId="34" pageNumber="1279">
<subSubSection id="22ED6512FFAFFFF1FC57815A02C3F957" pageId="34" pageNumber="1279" type="nomenclature">
<paragraph id="6A483699FFAFFFF1FC57815A0266F938" blockId="34.[915,1318,1661,1685]" box="[915,1318,1661,1685]" pageId="34" pageNumber="1279">
<heading id="310081F5FFAFFFF1FC57815A0266F938" box="[915,1318,1661,1685]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="1279" reason="2">
<taxonomicName id="ADF74D1AFFAFFFF1FC57815A03FDF93E" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[915,1213,1661,1684]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="1279" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">INDONAUCORIS MINUTUS</taxonomicName>
<emphasis id="5883EA8BFFAFFFF1FB0781A70266F938" bold="true" box="[1219,1318,1661,1685]" pageId="34" pageNumber="1279">
<taxonomicNameLabel id="43B057F0FFAFFFF1FB0781A70266F938" box="[1219,1318,1661,1685]" pageId="34" pageNumber="1279" rank="species">SP. NOV.</taxonomicNameLabel>
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<paragraph id="6A483699FFAFFFF1FCED81E102C3F957" blockId="34.[809,1425,1733,1786]" pageId="34" pageNumber="1279">
<emphasis id="5883EA8BFFAFFFF1FCED81E10335F976" box="[809,1141,1733,1755]" italics="true" pageId="34" pageNumber="1279">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
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</paragraph>
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<subSubSection id="22ED6512FFAFFFF7FCED80040363FACC" lastPageId="36" lastPageNumber="1281" pageId="34" pageNumber="1279" type="description">
<paragraph id="6A483699FFAFFFF0FCED8004037EF9DE" blockId="34.[809,1425,1826,1910]" lastBlockId="35.[827,1444,1476,1897]" lastPageId="35" lastPageNumber="1280" pageId="34" pageNumber="1279">
<emphasis id="5883EA8BFFAFFFF1FCED800404F5F895" box="[809,949,1827,1848]" italics="true" pageId="34" pageNumber="1279">Description:</emphasis>
See generic description; only additional details given here. Colour medium brown with yellowish markings. Head mostly yellow, with brown punctures (
<figureCitation id="F2CC2A1CFFAEFFF0FEEA82E206D0FA77" box="[302,400,1477,1499]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="35" pageNumber="1280">Fig. 24a</figureCitation>
). Pronotum yellow with brown punctures; transverse band in posterior third with irregular longitudinal stripes (
<figureCitation id="F2CC2A1CFFAEFFF0FDDC81250538F9BA" box="[536,632,1538,1560]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="35" pageNumber="1280">Fig. 24a</figureCitation>
). Scutellum yellowish, hemelytra brown with subtle mottling most evident on corium and membrane. Legs mostly yellowish, except brown protarsus (
<figureCitation id="F2CC2A1CFFAEFFF0FDA181790589F9D9" box="[613,713,1630,1652]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="35" pageNumber="1280">Fig. 24b</figureCitation>
) and tarsomere 3 of hindleg (
<figureCitation id="F2CC2A1CFFAEFFF0FE0C815A0567F93F" box="[456,551,1661,1683]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="35" pageNumber="1280">
Fig.
<quantity id="AD0F9B7CFFAEFFF0FE3A815A0567F93F" box="[510,551,1661,1682]" metricMagnitude="-2" metricUnit="kg" metricValue="2.4" pageId="35" pageNumber="1280" unit="g" value="24.0">24g</quantity>
</figureCitation>
). Abdominal terga mostly yellowish and sterna light brown. Hindwing extending to posterior half of tergum VIII. Mensural features (in mm) as follows: body length 5.08, width 2.76; head length 0.90, width 1.61; synthlipsis 0.66; labrum length 0.28, width 0.44; rostrum 0.38; antenna 0.44 (segments 1–4 0.04, 0.12, 0.14, 0.14, respectively); pronotum length 1.00, width 2.58; scutellum length 0.50, width 1.60; hemelytron (chord measurement) 3.44; embolium 0.48; foreleg: femur 1.02, tibia 0.68, tarsus 0.18; mesoleg: femur 1.10, tibia 0.84, tarsomere 1 0.10, tarsomere 2 0.14, tarsomere 3 0.18; hindleg: femur 1.22, tibia 1.36, tarsomere 1 0.12, tarsomere 2 0.26, tarsomere 3 0.27.
</paragraph>
<caption id="3E886611FFAEFFF0FF67826F0349FA35" ID-DOI="http://doi.org/10.5281/zenodo.6994674" ID-Zenodo-Dep="6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="35" pageNumber="1280" startId="35.[163,243,1352,1374]" targetBox="[243,1363,195,1313]" targetPageId="35">
<paragraph id="6A483699FFAEFFF0FF67826F0349FA35" blockId="35.[163,1443,1352,1432]" pageId="35" pageNumber="1280">
<emphasis id="5883EA8BFFAEFFF0FF67826F065CFAF0" bold="true" box="[163,284,1352,1374]" pageId="35" pageNumber="1280">Figure 24.</emphasis>
<taxonomicName id="ADF74D1AFFAEFFF0FEE3826F0555FAF3" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[295,533,1352,1374]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="1280" phylum="Arthropoda" rank="species" species="minutus" status="gen. nov., sp. nov.">
<emphasis id="5883EA8BFFAEFFF0FEE3826F0555FAF3" box="[295,533,1352,1374]" italics="true" pageId="35" pageNumber="1280">Indonaucoris minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFAEFFF0FDD9826F0596FAF3" box="[541,726,1352,1374]" pageId="35" pageNumber="1280" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures.
</paragraph>
</caption>
<paragraph id="6A483699FFAEFFF0FC97815A02E4F8C4" blockId="35.[827,1444,1476,1897]" pageId="35" pageNumber="1280">
Proctiger nearly twice as long as wide, brush of long setae along lateral margins, apex broadly rounded. Pygophore (sternum IX) conical posteriorly (basally), with thick brush of hairs around lateral and posterior margins, anterior margin broadly rounded (
<figureCitation id="F2CC2A1CFFAEFFF0FA8581D00409F881" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="35" pageNumber="1280">Fig. 24c, d</figureCitation>
). Aedeagus long and slender, slightly arcuate, with subapical membranous area on right side; parameres strongly asymmetrical, criss-crossing in basal half (Fig.
</paragraph>
<paragraph id="6A483699FFA9FFF7FD3187A302D0FF36" blockId="36.[757,1424,128,155]" box="[757,1424,128,155]" pageId="36" pageNumber="1281">
PHYLOGENY OF
<taxonomicName id="ADF74D1AFFA9FFF7FC0F87A3033CFF34" authorityName="LEACH" authorityYear="1815" box="[971,1148,132,153]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="1281" phylum="Arthropoda" rank="family">NAUCORIDAE</taxonomicName>
(
<taxonomicName id="ADF74D1AFFA9FFF7FB4E87A3026AFF34" box="[1162,1322,132,153]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Hemiptera" pageId="36" pageNumber="1281" phylum="Tracheophyta" rank="order">HEMIPTERA</taxonomicName>
) 1281
</paragraph>
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<paragraph id="6A483699FFA9FFF7FF5483140564FBCB" blockId="36.[144,1425,1075,1126]" pageId="36" pageNumber="1281">
<emphasis id="5883EA8BFFA9FFF7FF5483140648FBE5" bold="true" box="[144,264,1075,1097]" pageId="36" pageNumber="1281">Figure 25.</emphasis>
Tamhini stream, the presumed type locality of
<taxonomicName id="ADF74D1AFFA9FFF7FCCA831404BCFBE4" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[782,1020,1075,1097]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="1281" phylum="Arthropoda" rank="species" species="minutus" status="gen. nov., sp. nov.">
<emphasis id="5883EA8BFFA9FFF7FCCA831404BCFBE4" box="[782,1020,1075,1097]" italics="true" pageId="36" pageNumber="1281">Indonaucoris minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B057F0FFA9FFF7FBC7831403FCFBE4" box="[1027,1212,1075,1097]" pageId="36" pageNumber="1281" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
, in December 2019. Photo courtesy of Shruti Paripatyadar.
</paragraph>
</caption>
<paragraph id="6A483699FFA9FFF7FF5583B4056AFAA9" blockId="36.[145,761,1171,1285]" pageId="36" pageNumber="1281">
24d); left paramere overlapping right, with pronounced truncate lobe on left margin (
<figureCitation id="F2CC2A1CFFA9FFF7FE1A83960577FB6B" box="[478,567,1201,1223]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="36" pageNumber="1281">Fig. 24e</figureCitation>
); right paramere slenderer, more arcuate, with elongate sulcus in distal half to embrace aedeagus (
<figureCitation id="F2CC2A1CFFA9FFF7FE0583C8055AFAA9" box="[449,538,1263,1285]" captionStart="Figure 24" captionStartId="35.[163,243,1352,1374]" captionTargetBox="[243,1363,195,1313]" captionTargetId="figure-243@35.[243,1363,195,1313]" captionTargetPageId="35" captionText="Figure 24. Indonaucoris minutus gen. nov., sp. nov.: dorsal habitus (a), ventral habitus (b), genital capsule (c), genital capsule with terga IX and X removed (d), left and right parameres (e), dorsal and ventral views of mesothoracic leg (f), dorsal and ventral views of metathoracic leg (g). Sizes are not proportional among figures." figureDoi="http://doi.org/10.5281/zenodo.6994674" httpUri="https://zenodo.org/record/6994674/files/figure.png" pageId="36" pageNumber="1281">Fig. 24e</figureCitation>
).
</paragraph>
<paragraph id="6A483699FFA9FFF7FF55820A0525FA32" blockId="36.[145,760,1325,1439]" pageId="36" pageNumber="1281">
<emphasis id="5883EA8BFFA9FFF7FF55820A065EFAEF" box="[145,286,1325,1346]" italics="true" pageId="36" pageNumber="1281">Etymology:</emphasis>
This is one of the smallest species that would have been formerly placed in the genus
<taxonomicName id="ADF74D1AFFA9FFF7FF55824C07B8FA2D" authorityName="Geoffroy" authorityYear="1762" box="[145,248,1387,1408]" class="Insecta" family="Naucoridae" genus="Naucoris" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="1281" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFFA9FFF7FF55824C07B8FA2D" box="[145,248,1387,1408]" italics="true" pageId="36" pageNumber="1281">Naucoris</emphasis>
</taxonomicName>
. Therefore, the Latin epithet
<taxonomicName id="ADF74D1AFFA9FFF7FD8C824C05E7FA2D" authorityName="Sites" authorityYear="2022" box="[584,679,1387,1408]" class="Insecta" family="Naucoridae" genus="Indonaucoris" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="1281" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="5883EA8BFFA9FFF7FD8C824C05E7FA2D" box="[584,679,1387,1408]" italics="true" pageId="36" pageNumber="1281">minutus</emphasis>
</taxonomicName>
, small, is in reference to its relatively small size.
</paragraph>
<paragraph id="6A483699FFA9FFF7FF5582EF0646F9B7" blockId="36.[145,761,1480,1562]" pageId="36" pageNumber="1281">
<emphasis id="5883EA8BFFA9FFF7FF5582EF063DFA70" box="[145,381,1480,1501]" italics="true" pageId="36" pageNumber="1281">Material examined:</emphasis>
<materialsCitation id="DA9F3CC4FFA9FFF7FE4F82EF0646F9B7" ID-GBIF-Occurrence="3888705301" collectingDate="2004-09-28" country="India" latitude="18.44425" location="Mulshi" longLatPrecision="1" longitude="73.42757" municipality="Macropterous" pageId="36" pageNumber="1281" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
<typeStatus id="B54C883BFFA9FFF7FE4F82EF06BAFA73" box="[395,506,1480,1502]" pageId="36" pageNumber="1281" type="holotype">Holotype</typeStatus>
.
<collectingMunicipality id="8A2CACE3FFA9FFF7FDCC82EF05EEFA70" box="[520,686,1480,1501]" pageId="36" pageNumber="1281">Macropterous</collectingMunicipality>
male:
<collectingCountry id="12E07609FFA9FFF7FF5582C00790FA50" box="[145,208,1511,1533]" name="India" pageId="36" pageNumber="1281">India</collectingCountry>
:
<collectingRegion id="A833F87BFFA9FFF7FF1E82C00633FA50" box="[218,371,1511,1533]" country="India" name="Maharashtra" pageId="36" pageNumber="1281">Maharashtra</collectingRegion>
,
<location id="6F286042FFA9FFF7FEBA82C00690FA50" LSID="urn:lsid:plazi:treatment:E25E878FFFAFFFF7FC57815A0363FACC:6F286042FFA9FFF7FEBA82C00690FA50" box="[382,464,1511,1533]" country="India" latitude="18.44425" longLatPrecision="1" longitude="73.42757" municipality="Macropterous" name="Mulshi" pageId="36" pageNumber="1281" stateProvince="Maharashtra">Mulshi</location>
,
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,
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,
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.
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<emphasis id="5883EA8BFFA9FFF7FF5581620654F9F7" box="[145,276,1605,1626]" italics="true" pageId="36" pageNumber="1281">Repository:</emphasis>
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The
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is currently housed in the
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,
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of
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, and will be transferred to the
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(
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.
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<emphasis id="5883EA8BFFA9FFF7FF5581F8064FF959" box="[145,271,1759,1780]" italics="true" pageId="36" pageNumber="1281">Comments:</emphasis>
Only a single specimen was available to me at the time of this writing. The damage incurred during muscle extraction is regrettable, but the taxonomic importance of this specimen was not foreseen. My colleague Shruti Parapatyadar in Pune,
<collectingCountry id="12E07609FFA9FFF7FDBB807E0581F8C2" box="[639,705,1881,1903]" name="India" pageId="36" pageNumber="1281">India</collectingCountry>
, has sampled at the
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locality, but unfortunately she has not re-collected this species, although she provided a recent photo of the habitat (
<figureCitation id="F2CC2A1CFFA9FFF7FBBC83F70389FB4B" box="[1144,1225,1232,1254]" captionStart="Figure 25" captionStartId="36.[144,224,1075,1097]" captionTargetBox="[224,1344,195,1035]" captionTargetId="figure-24@36.[224,1344,195,1035]" captionTargetPageId="36" captionText="Figure 25. Tamhini stream, the presumed type locality of Indonaucoris minutus gen. nov., sp. nov., in December 2019. Photo courtesy of Shruti Paripatyadar." figureDoi="http://doi.org/10.5281/zenodo.6994676" httpUri="https://zenodo.org/record/6994676/files/figure.png" pageId="36" pageNumber="1281">Fig. 25</figureCitation>
). She noted that this is a seasonal stream that is subject to complete drying, especially after the 2004 construction of a retention wall that cuts off water flow shortly after the monsoon season ends.
</paragraph>
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<emphasis id="58839157475CFF90FF6EFF4BD6DE31A2" bold="true" box="[163,422,201,223]" italics="true" pageId="5" pageNumber="1152">Micreumenes argentatus</emphasis>
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Ƌ.
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: Ethiopia-S, Konso
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SE,
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,
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, 11.-
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, Ƌ, leg. J. Halada.
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<paragraph id="6A484D45475CFF90FF6EFE91D62C30D9" blockId="5.[163,1070,275,420]" pageId="5" pageNumber="1152">
Diese Art (
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) hat, ähnlich
<emphasis id="58839157475CFF90FDC1FE91D46D3054" box="[524,789,275,297]" italics="true" pageId="5" pageNumber="1152">Microdynerus mutarensis</emphasis>
GUSENLEITNER, 2002, ein ausgerandetes Hinterschildchen, aber dieses flacher ausgerandet. Weiters ist bei der hier beschriebenen Art das Ende der Horizontalfläche des Propodeums doppelspitzig (
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). Artspezifisch, ist der vollständig von einer anliegenden silbrigen Pubeszenz bedeckte Thorax.
</paragraph>
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Bei schwarzer Grundfarbe sind weiss gefärbt: der Clypeus mit verdunkelten Rändern, eine in der Mitte breit unterbrochene Binde auf dem Pronotum, kleine Flecken vorne auf den Tegulae, Endbinden auf den Tergiten 1 bis 3 sowie auf den Sterniten 2 und 3. Auf den Beinen sind gelb gefärbt: das Ende der Schenkel
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grösserem Umfang, bei den Schenkeln
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geringerem Bereich und die Schienen 1 und 2 aussen. Sehr kleine Flecken auf den Hüften der Beine 2 und 3. Die Flügel sind glasklar durchscheinend.
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Der Clypeus (
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) etwa so breit wie lang, sein Ausschnitt sehr schmal mit spitzen Ausschnittecken. Der Clypeus grob punktiert und dicht silbrig behaart. Die Stirn und der Scheitel sehr grob punktiert, die Schläfen sehr fein. Von der Seite gesehen der Scheitel eben. Die Fühlerschäfte fein punktuliert und der schmale Fühlerhaken recht zurückgeschlagen bis zur Basis des 1. Fühlergliedes. Das sehr grob punktierte Pronotum geht abgerundet auf die Vorderwand über, aber die Schultern spitz ausgebildet. Ebenso grob wie das Pronotum sind das Mesonotum, das Schildchen und die Mesopleuren punktiert. Das Hinterschildchen ist feiner punktiert und flach ausgerandet. Das Propodeum ist ähnlich dem Mesonotum punktiert, wobei der ebene, mittlere Horizontalbereich am distalen Ende in zwei Spitzen endet. Die Tegulae grob punktiert (aber kleiner als das Mesonotum), die seidig glänzenden Punktzwischenräume punktuliert. Die Beine matt.
</paragraph>
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Das 1. Tergit (
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) von oben gesehen seitlich flach ausgerandet, erweitert sich aber gegen das distale Ende. Das 1. Tergit unregelmässig grob punktiert mit seidig glänzenden Punktzwischenräumen. Das 2. Tergit gleichmässig punktiert mit matten Zwischenräumen, welche an den Seiten aber stärker glänzen. Der eingedrückte Endsaum des 2. Tergites durch grobe längliche Punktgruben gegittert und zwischen diesen in der Farbe der hellen Endbinde undurchsichtig pigmentiert. Ähnlich strukturiert auch das distale Ende des 3. Tergites. Die Tergite 4 bis 7 nicht punktiert. Die Sternite entsprechen in den Strukturen den jeweiligen Tergiten.
</paragraph>
<paragraph id="6A484D45475CFF90FF6EFB49D6BD3421" blockId="5.[163,1071,1227,1372]" pageId="5" pageNumber="1152">Die helle Behaarung auf der Stirn etwa in der Länge eines Durchmessers einer Ocelle. Der Scheitel wesentlich kürzer behaart. Der untere Bereich der Stirn und der gesamte Thorax von einer anliegenden silbrigen Pubeszenz bedeckt. Die Basis der Vorderschenkel und die Hüften 1 und 2 mit kurzer borstenartiger Behaarung. Das Abdomen mit mikroskopischer Pubeszenz.</paragraph>
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Länge:
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.
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Das
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ist unbekannt.
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<mods:namePart id="8722B39789F80D170DC22F698E4B5F9D">Gusenleitner, Josef</mods:namePart>
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<mods:typeOfResource id="18180E4880773009F4889E15DF036C81">text</mods:typeOfResource>
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<mods:title id="C47CBB3F9C032F5CB335D069A9797706">Linzer biologische Beiträge</mods:title>
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<mods:part id="140120B3AC1E35CE2CBD3A8E5FF2E9DE">
<mods:date id="984356077FA9EC92F507525017E727DE">2017</mods:date>
<mods:detail id="7D04AC0D9D817325758AF1A2C7311581" type="pubDate">
<mods:number id="F16A291CEED617E289D09D5A6DA0E83D">2017-12-11</mods:number>
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<mods:detail id="A9E23D03E207B90BB12A5EFE4C21B189" type="volume">
<mods:number id="61358476356B9C00CDF0D9AEC9FC3BC5">49</mods:number>
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<mods:number id="6BE436F804DB1BBBF743CB2F9A12D62C">2</mods:number>
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<mods:start id="F24A1BEC4EAAA8D8A745846DEA541776">1147</mods:start>
<mods:end id="D5101E6478C9835FAD17DBC1024BA14F">1155</mods:end>
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<mods:classification id="5B87369FFC402FB78CABE5E996C3452D">journal article</mods:classification>
<mods:identifier id="4674A73B5E88A0936939BA207C94DB46" type="DOI">10.5281/zenodo.5183200</mods:identifier>
<mods:identifier id="F4052D22811050BBD96CCACD26ABC9A5" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="88FB39B0C9D0B388C21151292F7964C1" type="Zenodo-Dep">5183200</mods:identifier>
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<treatment id="E25EFC53475EFF92FF6EFF4BD60334D1" ID-DOI="http://doi.org/10.5281/zenodo.5192090" ID-Zenodo-Dep="5192090" LSID="urn:lsid:plazi:treatment:E25EFC53475EFF92FF6EFF4BD60334D1" httpUri="http://treatment.plazi.org/id/E25EFC53475EFF92FF6EFF4BD60334D1" lastPageNumber="1154" pageId="7" pageNumber="1154">
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<paragraph id="6A484D45475EFF92FF6EFF4BD56031A2" blockId="7.[163,536,201,223]" box="[163,536,201,223]" pageId="7" pageNumber="1154">
<heading id="3100FA29475EFF92FF6EFF4BD56031A2" box="[163,536,201,223]" fontSize="8" level="2" pageId="7" pageNumber="1154" reason="2">
<emphasis id="58839157475EFF92FF6EFF4BD68F31A2" bold="true" box="[163,503,201,223]" pageId="7" pageNumber="1154">
<taxonomicName id="ADF736C6475EFF92FF6EFF4BD6DB31A2" authority="Gusenleitner, 2017" authorityName="Gusenleitner" authorityYear="2017" box="[163,419,201,223]" class="Insecta" family="Eumenidae" genus="Micreumenes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="1154" phylum="Arthropoda" rank="species" species="lamellatus" status="sp. nov.">
<emphasis id="58839157475EFF92FF6EFF4BD6DB31A2" bold="true" box="[163,419,201,223]" italics="true" pageId="7" pageNumber="1154">Micreumenes lamellatus</emphasis>
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</emphasis>
Ƌ.
</heading>
</paragraph>
<paragraph id="6A484D45475EFF92FF6EFF6CD7923066" blockId="7.[163,1070,238,283]" pageId="7" pageNumber="1154">
<typeStatus id="B54CF3E7475EFF92FF6EFF6CD67D3079" box="[163,261,238,260]" pageId="7" pageNumber="1154" type="holotype">Holotypus</typeStatus>
: Ethiopia-C, Shen Husen
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NW,
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<geoCoordinate id="0FC32B82475EFF92FD1FFF6CD4633079" box="[722,795,238,260]" degrees="40" direction="east" minutes="36" orientation="longitude" pageId="7" pageNumber="1154" precision="925" value="40.6">40°36'E</geoCoordinate>
,
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,
<date id="1E496B85475EFF92FCA2FF6CD4B93079" box="[879,961,238,260]" pageId="7" pageNumber="1154" value="2015-06-01">1.6.2015</date>
, Ƌ, leg. J. Halada.
</paragraph>
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<subSubSection id="22ED1ECE475EFF92FF6EFEABD60334D1" pageId="7" pageNumber="1154" type="description">
<paragraph id="6A484D45475EFF92FF6EFEABD5EF3007" blockId="7.[163,1070,297,378]" pageId="7" pageNumber="1154">
Diese Art (
<figureCitation id="F2CC51C0475EFF92FED9FEABD6103042" box="[276,360,297,319]" captionStart="Abb" captionStartId="8.[163,206,1065,1086]" captionTargetBox="[163,1070,197,1055]" captionTargetId="figure-0@8.[163,1070,197,1055]" captionTargetPageId="8" captionText="Abb. 16-18: (16) Micreumenes lamellatus nov.sp. ♀, Habitus; (17) Micreumenus lamellatus nov.sp. ♀, Endlamelle des 2. Tergites; (18) Micreumenus lamellatus nov.sp. ♀, Clypeus." figureDoi="http://doi.org/10.5281/zenodo.5183210" httpUri="https://zenodo.org/record/5183210/files/figure.png" pageId="7" pageNumber="1154">Abb. 16</figureCitation>
) ist besonders durch die lamellenartige, steil nach oben aufgerichtete Endlamelle (
<figureCitation id="F2CC51C0475EFF92FEE7FEC4D6FB3021" box="[298,387,326,348]" captionStart="Abb" captionStartId="8.[163,206,1065,1086]" captionTargetBox="[163,1070,197,1055]" captionTargetId="figure-0@8.[163,1070,197,1055]" captionTargetPageId="8" captionText="Abb. 16-18: (16) Micreumenes lamellatus nov.sp. ♀, Habitus; (17) Micreumenus lamellatus nov.sp. ♀, Endlamelle des 2. Tergites; (18) Micreumenus lamellatus nov.sp. ♀, Clypeus." figureDoi="http://doi.org/10.5281/zenodo.5183210" httpUri="https://zenodo.org/record/5183210/files/figure.png" pageId="7" pageNumber="1154">Abb. 17</figureCitation>
) des 2. Tergites charakterisiert. Ich kenne keine Art innerhalb dieser Gattung, welche dieses Merkmal aufweist.
</paragraph>
<paragraph id="6A484D45475EFF92FF6EFE0BD4B0334F" blockId="7.[163,1070,393,562]" pageId="7" pageNumber="1154">Bei schwarzer Grundfarbe sind weiss gefärbt: der Clypeus, die Unterseite der Fühlerschäfte, kleine Flecken auf den Schultern, Flecken auf den Tegulae, Endbinden auf den Tergiten 1 und 2 sowie auf dem 2. Sternit. Flecken an den Enden der Schenkel 1 und 2 und die Schienen 1 und 2 aussen. Rötlich gefärbt sind die Mandibeln, die Unterseiten der Fühler-Endglieder, die Tegulae, das Ende des 1. Sternites und die Beine soweit sie nicht weiss gezeichnet sind. Die Flügel sind glasklar durchscheinend.</paragraph>
<paragraph id="6A484D45475EFF92FF6EFDC3D46632CA" blockId="7.[163,1071,577,951]" pageId="7" pageNumber="1154">
Der Clypeus (
<figureCitation id="F2CC51C0475EFF92FEF8FDC3D6F3332A" box="[309,395,577,599]" captionStart="Abb" captionStartId="8.[163,206,1065,1086]" captionTargetBox="[163,1070,197,1055]" captionTargetId="figure-0@8.[163,1070,197,1055]" captionTargetPageId="8" captionText="Abb. 16-18: (16) Micreumenes lamellatus nov.sp. ♀, Habitus; (17) Micreumenus lamellatus nov.sp. ♀, Endlamelle des 2. Tergites; (18) Micreumenus lamellatus nov.sp. ♀, Clypeus." figureDoi="http://doi.org/10.5281/zenodo.5183210" httpUri="https://zenodo.org/record/5183210/files/figure.png" pageId="7" pageNumber="1154">Abb. 18</figureCitation>
) breiter als lang (2,0: 1,7), sein Ausschnitt flach (Breite: Tiefe =1,0: 0,3) und so breit wie der Abstand der Fühlergruben. Die gröbere Punktierung des Clypeus wegen der dichten silbrigen Pubeszenz kaum zu erkennen. Grob punktiert sind die Stirn und der Scheitel. Die Schläfen feiner skulpturiert, die Fühlerschäfte weitläufig punktuliert. Die Fühlerendglieder sehr zart, kurz, schwach gebogen und spitz. Das Pronotum gröber als die Stirn und dicht punktiert, mit einer Kante gegen die Vorderwand, die Schultern spitz. Ebenso grob und dicht das Mesonotum und das Schildchen punktiert. Die Mesopleuren etwas feiner und weitläufiger punktiert. Das grob skulpturierte Hinterschildchen ausgerandet. Das Propodeum kraterartig skulpturiert wobei der ebene, mittlere Horizontalbereich am distalen Ende in zwei Spitzen endet. Die Skulptur der Konkavität und der Aussenwände des Propodeums gegenüber seinen anderen Abschnitten feiner. Die Tegulae breit mit durchscheinendem Aussenrand und in der Mitte erhabenen Längsstreifen. Die Beine seidig glänzend.
</paragraph>
<paragraph id="6A484D45475EFF92FF6EFC45D66C3599" blockId="7.[163,1070,966,1252]" pageId="7" pageNumber="1154">
In der Aufsicht verläuft das 1. Tergit parallel, erweitert sich am distalen Ende. Vor diesem Ende eine Querrinne ausgebildet. Das 1. Tergit unregelmässig grob punktiert mit glänzenden Punktzwischenräumen. Das 2. Tergit regelmässig und grob punktiert, die schmalen Punktzwischenräume punktuliert. Der eingedrückte Endsaum des 2. Tergites durch grobe längliche Punktgruben gegittert und zwischen diesen in der Farbe der hellen Endbinde undurchsichtig pigmentiert. Das Ende dieses durchsichtigen Endsaumes steil nach oben lamellenartig aufgerichtet (
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). Die Tergite 3 bis 7 ohne Punkte. Das 2. Sternit ebenso grob wie das 2. Tergit gestaltet, aber weitläufiger punktiert. Der eingedrückte Endsaum dieses Sternites nicht lamellenartig ausgeprägt. Die Sternite 3 bis 7 punktlos.
</paragraph>
<paragraph id="6A484D45475EFF92FF6EFB76D36B341F" blockId="7.[163,1070,1268,1378]" pageId="7" pageNumber="1154">Die unteren Bereiche der Stirn und die Schläfen von einer silbrigen, anliegenden Pubeszenz bedeckt. Der obere Abschnitt der Stirn und der Scheitel mit extrem kurzer Behaarung (⅓ eines Ocellen-Durchmessers). Der Thorax und das Abdomen mit mikroskopischer Pubeszenz, nur die Mesopleuren mit anliegender silbriger Pubeszenz.</paragraph>
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Länge:
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.
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Das
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ist unbekannt.
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</treatment>
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