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<document id="55260E8849D2E07570B1CF5B6655FB8C" ID-DOI="10.5281/zenodo.186485" ID-GBIF-Dataset="2eec0d6a-510b-44e7-a8e3-602dc5a88838" ID-ISSN="1175-5326" ID-Zenodo-Dep="186485" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461052423066" checkinUser="plazi" docAuthor="Joly, Luis J." docDate="2009" docId="E2083F003B00FFA3FF74FE99FB3F6702" docLanguage="en" docName="zt02048p064.pdf" docOrigin="Zootaxa 2048" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cyclocephala dichroa Dechambre 1992" docType="treatment" docVersion="7" lastPageNumber="62" masterDocId="1E3147783B0DFFACFFE3FB23FFC16441" masterDocTitle="Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae)" masterLastPageNumber="64" masterPageNumber="47" pageNumber="60" updateTime="1698579509211" updateUser="plazi">
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<mods:title id="EBAA8CB45AA6233872BDBA49B39880EA">Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
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<mods:namePart id="A85B8F123F1B3607B6F78443665A53BE">Joly, Luis J.</mods:namePart>
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<treatment id="E2083F003B00FFA3FF74FE99FB3F6702" ID-DOI="http://doi.org/10.5281/zenodo.6216466" ID-GBIF-Taxon="119612266" ID-Zenodo-Dep="6216466" LSID="urn:lsid:plazi:treatment:E2083F003B00FFA3FF74FE99FB3F6702" httpUri="http://treatment.plazi.org/id/E2083F003B00FFA3FF74FE99FB3F6702" lastPageId="15" lastPageNumber="62" pageId="13" pageNumber="60">
<subSubSection id="22BBDD9D3B00FFA1FF74FE99FC4D61B6" pageId="13" pageNumber="60" type="nomenclature">
<paragraph id="6A1E8E163B00FFA1FF74FE99FDBE6195" blockId="13.[151,908,1466,1527]" box="[151,639,1466,1492]" pageId="13" pageNumber="60">
<heading id="3156397A3B00FFA1FF74FE99FDBE6195" bold="true" box="[151,639,1466,1492]" fontSize="11" level="1" pageId="13" pageNumber="60" reason="1">
<taxonomicName id="ADA1F5953B00FFA1FF74FE99FDBE6195" authority="Dechambre, 1992" authorityName="Dechambre" authorityYear="1992" box="[151,639,1466,1492]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="60" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B00FFA1FF74FE99FDBE6195" bold="true" box="[151,639,1466,1492]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF74FE99FE5C6195" bold="true" box="[151,413,1466,1492]" italics="true" pageId="13" pageNumber="60">Cyclocephala dichroa</emphasis>
<bibRefCitation id="0E30F3E73B00FFA1FE46FE99FDBE6195" author="Dechambre" box="[421,639,1466,1492]" pageId="13" pageNumber="64" refString="Dechambre, R. - P. (1992) Nouveaux Cyclocephalini des genres Cyclocephala et Aspidolea (Col. Dynastidae). Les Coleopteres du Monde, 14, 57 - 75, color plate 12." type="journal article" year="1992">Dechambre, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF74FEFEFC4D61B6" blockId="13.[151,908,1466,1527]" box="[151,908,1501,1527]" pageId="13" pageNumber="60">
(
<figureCitation id="F29A92933B00FFA1FF7CFEFEFF3261B6" box="[159,243,1501,1527]" captionStart="FIGURE 2" captionStartId="4.[151,255,1865,1889]" captionTargetBox="[373,1218,1132,1833]" captionTargetId="figure@4.[366,1220,1130,1841]" captionTargetPageId="4" captionText="FIGURE 2. Distribution of Cyclocephala dichroa (circles), and C. bicolorata (squares)." httpUri="https://zenodo.org/record/186487/files/figure.png" pageId="13" pageNumber="60">Figs. 2</figureCitation>
,
<figureCitation id="F29A92933B00FFA1FF1CFEFEFEB761B6" box="[255,374,1501,1527]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="13" pageNumber="60">11–14, 22</figureCitation>
,
<figureCitation id="F29A92933B00FFA1FE61FEFEFE0F61B6" box="[386,462,1501,1527]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="13" pageNumber="60">29–30</figureCitation>
,
<figureCitation id="F29A92933B00FFA1FE39FEFEFE3B61B6" box="[474,506,1501,1527]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="13" pageNumber="60">39</figureCitation>
, 46–47,
<figureCitation id="F29A92933B00FFA1FDBEFEFEFDBB61B6" box="[605,634,1501,1527]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="13" pageNumber="60">53</figureCitation>
–54,
<figureCitation id="F29A92933B00FFA1FD56FEFEFD1461B6" box="[693,725,1501,1527]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="13" pageNumber="60">60</figureCitation>
, 65–66, 71–72)
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B00FFA3FF74FD09FBB3650D" lastPageId="15" lastPageNumber="62" pageId="13" pageNumber="60" type="description">
<paragraph id="6A1E8E163B00FFA1FF74FD09FE8B62D3" blockId="13.[151,1437,1578,2030]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF74FD09FE806205" bold="true" box="[151,321,1578,1604]" pageId="13" pageNumber="60">Bibliography.</emphasis>
<taxonomicName id="ADA1F5953B00FFA1FEABFD08FC876205" authority="Dechambre 1992: 67" authorityName="Dechambre" authorityPageNumber="67" authorityYear="1992" box="[328,838,1578,1604]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="60" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B00FFA1FEABFD08FD8B6205" box="[328,586,1579,1604]" italics="true" pageId="13" pageNumber="60">Cyclocephala dichroa</emphasis>
<bibRefCitation id="0E30F3E73B00FFA1FDB2FD09FCDF6205" author="Dechambre" box="[593,798,1578,1604]" pageId="13" pageNumber="64" refString="Dechambre, R. - P. (1992) Nouveaux Cyclocephalini des genres Cyclocephala et Aspidolea (Col. Dynastidae). Les Coleopteres du Monde, 14, 57 - 75, color plate 12." type="journal article" year="1992">Dechambre 1992</bibRefCitation>
: 67
</taxonomicName>
(description of male; compared with
<taxonomicName id="ADA1F5953B00FFA1FB1EFD08FAB26205" box="[1277,1395,1578,1604]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="60" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B00FFA1FB1EFD08FAD16205" box="[1277,1296,1579,1604]" italics="true" pageId="13" pageNumber="60">C</emphasis>
.
<emphasis id="58D552043B00FFA1FAFCFD08FAB26205" box="[1311,1395,1579,1604]" italics="true" pageId="13" pageNumber="60">bicolor</emphasis>
</taxonomicName>
,
<taxonomicName id="ADA1F5953B00FFA1FA62FD08FED0622B" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="60" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B00FFA1FA62FD08FA556205" box="[1409,1428,1579,1604]" italics="true" pageId="13" pageNumber="60">C</emphasis>
.
<emphasis id="58D552043B00FFA1FF74FD72FED0622B" box="[151,273,1617,1642]" italics="true" pageId="13" pageNumber="60">bicolorata</emphasis>
</taxonomicName>
),
<figureCitation id="F29A92933B00FFA1FEC9FD72FEBC622A" box="[298,381,1617,1643]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="13" pageNumber="60">Fig. 31</figureCitation>
(genitalia),
<collectingCountry id="12B6CE863B00FFA1FDEFFD72FD7C622A" box="[524,701,1617,1643]" name="French Guiana" pageId="13" pageNumber="60">French Guiana</collectingCountry>
.
<typeStatus id="B51A30B43B00FFA1FD2AFD72FCF7622A" box="[713,822,1617,1643]" pageId="13" pageNumber="60" type="holotype">Holotype</typeStatus>
male, MNHN.
<typeStatus id="B51A30B43B00FFA1FC0CFD72FBE8622A" box="[1007,1065,1617,1643]" pageId="13" pageNumber="60">Type</typeStatus>
locality:
<collectingCountry id="12B6CE863B00FFA1FB7AFD72FA8B622A" box="[1177,1354,1617,1643]" name="French Guiana" pageId="13" pageNumber="60">French Guiana</collectingCountry>
: Route du Kaw, PK12.
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF26FDBDFAC06347" blockId="13.[151,1437,1578,2030]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26FDBDFE9762F9" bold="true" box="[197,342,1694,1720]" pageId="13" pageNumber="60">Description</emphasis>
. Short, oval, moderately shiny. Brownish red, posterior part of clypeus, frons brownish red or black, vertex black, elytra testaceous, sometimes with base and a reddish brown posteriorly narrowed sutural band; dorsal surface and pygidium glabrous, ventral pubescence sparse.
<emphasis id="58D552043B00FFA1FC05FDC8FB816344" bold="true" box="[998,1088,1771,1797]" pageId="13" pageNumber="60">Length</emphasis>
: 10.0–13.0 mm.
</paragraph>
<paragraph id="6A1E8E163B00FFA2FF26FC31FAAC64F3" blockId="13.[151,1437,1578,2030]" lastBlockId="14.[151,1437,152,2034]" lastPageId="14" lastPageNumber="61" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26FC31FEDC636D" bold="true" box="[197,285,1810,1836]" pageId="13" pageNumber="60">MALE</emphasis>
:
<emphasis id="58D552043B00FFA1FECFFC31FE50636D" bold="true" box="[300,401,1810,1836]" pageId="13" pageNumber="60">Clypeus</emphasis>
(
<figureCitation id="F29A92933B00FFA1FE43FC31FE32636D" box="[416,499,1810,1836]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="13" pageNumber="60">Fig. 11</figureCitation>
) about 2.4 times as wide across base as long; sides straight, strongly convergent to truncate and not reflexed apex, shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view slightly convex on apical half; surface finely alutaceous, with fine, shallow and widely annulate punctures, annulation wide, not well defined posteriorly, sometimes confluent, in which case appearing as irregular and fine, more or less transverse striae. Frontoclypeal suture (
<figureCitation id="F29A92933B00FFA1FAE1FC8EFA976386" box="[1282,1366,1965,1991]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="13" pageNumber="60">Fig. 11</figureCitation>
) thin, not well defined, feebly bisinuate, but not pointed posteriorly at middle.
<emphasis id="58D552043B00FFA1FBCEFCF0FBB863AC" bold="true" box="[1069,1145,2003,2029]" pageId="13" pageNumber="60">Frons</emphasis>
: width about 2.4 times transverse diameter of eye, finely alutaceous; punctures stronger and more discrete than those on clypeus.
</paragraph>
<paragraph id="6A1E8E163B03FFA2FF26FB9DFD74664F" blockId="14.[151,1437,152,2034]" pageId="14" pageNumber="61">
<emphasis id="58D552043B03FFA2FF26FB9DFE906499" bold="true" box="[197,337,190,216]" pageId="14" pageNumber="61">Mandibles</emphasis>
with external border sinuate, apex projecting externally.
<emphasis id="58D552043B03FFA2FBC6FB9DFB586499" bold="true" box="[1061,1177,190,216]" pageId="14" pageNumber="61">Maxillae</emphasis>
(
<figureCitation id="F29A92933B03FFA2FB4FFB9DFAC96499" box="[1196,1288,190,216]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="14" pageNumber="61">Fig. 14</figureCitation>
) with galea moderately robust, with two apical, one median and three basal teeth, the basal teeth very small and arising from a common base; stipes projected anteriorly on inner side and with a brush of fine yellowish brown setae on apex of projection; maxillary palpi (
<figureCitation id="F29A92933B03FFA2FD85FA11FD7D650D" box="[614,700,306,332]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="14" pageNumber="61">Fig. 22</figureCitation>
) moderately robust, second palpomere widened towards apex, longer than third, fourth fusiform, more than 3.0 times as long as preceding.
<emphasis id="58D552043B03FFA2FBA3FA7BFB676533" bold="true" box="[1088,1190,344,370]" pageId="14" pageNumber="61">Labium</emphasis>
(
<figureCitation id="F29A92933B03FFA2FB54FA7AFAD16532" box="[1207,1296,345,371]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="14" pageNumber="61">Fig. 13</figureCitation>
) with sides slightly convex on basal two-thirds, not constricted at middle of apical third; apex uniformly convex on ventral side, more arcuately produced, with small emargination at middle on oral side, fimbriate with long, fine, yellowish brown setae; surface almost flat, impunctate on disc, with one longitudinal series of long bristles on each side converging towards base.
</paragraph>
<paragraph id="6A1E8E163B03FFA2FF26F939FAF66675" blockId="14.[151,1437,152,2034]" box="[197,1335,538,564]" pageId="14" pageNumber="61">
<emphasis id="58D552043B03FFA2FF26F939FEFF6675" bold="true" box="[197,318,538,564]" pageId="14" pageNumber="61">Antennae</emphasis>
with 10 antennomeres, club almost 2.0 times the length of antennomeres 2–7 together.
</paragraph>
<paragraph id="6A1E8E163B03FFA2FF26F963FCC167D1" blockId="14.[151,1437,152,2034]" pageId="14" pageNumber="61">
<emphasis id="58D552043B03FFA2FF26F963FE82661B" bold="true" box="[197,323,576,602]" pageId="14" pageNumber="61">Pronotum</emphasis>
1.5 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected at middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface finely alutaceous (
<date id="1E1FA8D63B03FFA2FEC4F996FEA0668E" box="[295,353,693,719]" pageId="14" pageNumber="61" value="1940-10">40X</date>
); punctures annulate, moderately fine and dense, the distance between most punctures on disc as long as or longer than diameter of punctures, punctures gradually larger toward sides, the diameter of a puncture about 2.0 times the diameter of those on center of disc, more dense on basal angles where the distance between most punctures is as long as or shorter than diameter of a puncture; all surface also covered with numerous minute punctures (
<date id="1E1FA8D63B03FFA2FDD8F873FDB0672B" box="[571,625,848,874]" pageId="14" pageNumber="61" value="1950-10">50X</date>
or more).
<emphasis id="58D552043B03FFA2FD17F86CFCB76728" bold="true" box="[756,886,847,873]" pageId="14" pageNumber="61">Scutellum</emphasis>
triangular, sides feebly arcuate, apex briefly rounded; punctures similar to those on pronotal disc.
</paragraph>
<paragraph id="6A1E8E163B03FFA2FF26F8BEFE7F6039" blockId="14.[151,1437,152,2034]" pageId="14" pageNumber="61">
<emphasis id="58D552043B03FFA2FF26F8BEFED767F6" bold="true" box="[197,278,925,951]" pageId="14" pageNumber="61">Elytra</emphasis>
1.1 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones weak; a feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident, also with numerous micropunctures (
<date id="1E1FA8D63B03FFA2FF7CFF1BFF126013" box="[159,211,1080,1106]" pageId="14" pageNumber="61" value="1950-10">50X</date>
or more) scattered all over the surface; sutural interstria flat, a little elevated on apical half. Epipleura narrow, more so apically.
</paragraph>
<paragraph id="6A1E8E163B03FFA2FF26FFA6FD1B617B" blockId="14.[151,1437,152,2034]" pageId="14" pageNumber="61">
<emphasis id="58D552043B03FFA2FF26FFA6FE8560DE" bold="true" box="[197,324,1157,1183]" pageId="14" pageNumber="61">Pygidium</emphasis>
almost 2.0 times as wide as long; in posterior view the inferior borders almost straight, converging to feebly convex-truncate apex in an angle distinctly greater than 90°; in lateral view (
<figureCitation id="F29A92933B03FFA2FAD8FF8FFA526087" box="[1339,1427,1196,1222]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="14" pageNumber="61">Fig. 65</figureCitation>
) convex, more strongly on apical half; surface smooth, shiny, with large annulate punctures, more dense on lateral angles; on apex more shining, the punctures somewhat smaller and less dense, the distances between most punctures 1.0–2.0 times their own diameter.
</paragraph>
<paragraph id="6A1E8E163B03FFA2FF26FE65FDDB62A2" blockId="14.[151,1437,152,2034]" pageId="14" pageNumber="61">
<emphasis id="58D552043B03FFA2FF26FE65FEFB6121" bold="true" box="[197,314,1350,1376]" pageId="14" pageNumber="61">Protibiae</emphasis>
(
<figureCitation id="F29A92933B03FFA2FEA8FE65FE636121" box="[331,418,1350,1376]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="14" pageNumber="61">Fig. 29</figureCitation>
) tridentate, small basal tooth separated from the intermediate tooth by a distance 2.0 times as long as that between intermediate and anterior teeth; intermediate and anterior teeth well developed, basal tooth acute and anteriorly projected; apical spur almost straight, surpassing apex of first tarsomere.
<emphasis id="58D552043B03FFA2FF74FE99FF3C6195" bold="true" box="[151,253,1466,1492]" pageId="14" pageNumber="61">Protarsi</emphasis>
(
<figureCitation id="F29A92933B03FFA2FEEEFE98FEA36194" box="[269,354,1467,1493]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="14" pageNumber="61">Fig. 39</figureCitation>
) enlarged; tarsomere 5 shorter than 1–4 together, without ventro-internal carina, ventral side with inconspicuous longitudinal striae on inner side of apex; tarsomere 4 longitudinally striated on about apical half, tarsomere 3 usually without apical striae, sometimes (2 out of
<specimenCount id="7CA7459F3B03FFA2FBF3FD2BFBAF6263" box="[1040,1134,1544,1570]" pageId="14" pageNumber="61" type="male">6 males</specimenCount>
) with 1–2 striae on inner side of apex; largest claw (
<figureCitation id="F29A92933B03FFA2FE2EFD0DFD986209" box="[461,601,1582,1608]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="14" pageNumber="61">Figs. 46–47</figureCitation>
) wide, flattened, moderately widely cleft at apex, the external border of small branch continuous with internal curvature of the claw.
<emphasis id="58D552043B03FFA2FC99FD76FBCA622E" bold="true" box="[890,1035,1621,1647]" pageId="14" pageNumber="61">Metafemur</emphasis>
(
<figureCitation id="F29A92933B03FFA2FBFEFD76FBB4622E" box="[1053,1141,1621,1647]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="14" pageNumber="61">Fig. 53</figureCitation>
) 2.0 times as long as its maximum width.
<emphasis id="58D552043B03FFA2FE8DFD58FE3862D4" bold="true" box="[366,505,1659,1685]" pageId="14" pageNumber="61">Metatibiae</emphasis>
(
<figureCitation id="F29A92933B03FFA2FDE8FD5FFDA262D7" box="[523,611,1660,1686]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="14" pageNumber="61">Fig. 53</figureCitation>
) robust, widened at apex, bristles on diagonal carina and apex very strong.
<emphasis id="58D552043B03FFA2FF0CFD81FEA862FD" bold="true" box="[239,361,1698,1724]" pageId="14" pageNumber="61">Metatarsi</emphasis>
subequal in length to tibiae, with first tarsomere widened towards apex, with apical bristles similar to those of apex of tibiae.
</paragraph>
<paragraph id="6A1E8E163B03FFA2FF26FDCCFCD16371" blockId="14.[151,1437,152,2034]" pageId="14" pageNumber="61">
<emphasis id="58D552043B03FFA2FF26FDCCFE0E6348" bold="true" box="[197,463,1775,1801]" pageId="14" pageNumber="61">Prosternal projection</emphasis>
high, apex alutaceous, anterior part transversely oval, externally pointed; posterior part with posterior border fimbriate with long bristles.
</paragraph>
<paragraph id="6A1E8E163B03FFA2FF26FC1EFB4E63B3" blockId="14.[151,1437,152,2034]" pageId="14" pageNumber="61">
<emphasis id="58D552043B03FFA2FF26FC1EFE0E6316" bold="true" box="[197,463,1853,1879]" pageId="14" pageNumber="61">Pronotal hypomera</emphasis>
alutaceous, with sparse, erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large, dense, confluent, annulate punctures; disc of metasternum smooth, with small, scattered punctures; metacoxae with small, moderately dense punctures, more dense and confluent on posterior and external borders. Ventrites with moderately dense annulate punctures, some open posteriorly; second ventrite with strong, triangular carina along middle, from basal quarter to apex.
</paragraph>
<paragraph id="6A1E8E163B02FFA3FF26FBB4FE1E64F3" blockId="15.[151,1437,151,835]" box="[197,479,151,178]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FBB4FEFC64F0" bold="true" box="[197,317,151,177]" pageId="15" pageNumber="62">Aedeagus</emphasis>
:
<figureCitation id="F29A92933B02FFA3FEAEFBBBFE1A64F3" box="[333,475,152,178]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="15" pageNumber="62">Figs. 71–72</figureCitation>
.
</paragraph>
<paragraph id="6A1E8E163B02FFA3FF26FB9DFBB3650D" blockId="15.[151,1437,151,835]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FB9DFE866499" bold="true" box="[197,327,190,216]" pageId="15" pageNumber="62">FEMALE</emphasis>
: differs from the male in the following characters: punctation generally stronger, that of pygidium of the same size; protarsi slender; protibiae
<figureCitation id="F29A92933B02FFA3FCF9FBC6FCB364BE" box="[794,882,229,255]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="15" pageNumber="62">Fig. 30</figureCitation>
; metafemora (
<figureCitation id="F29A92933B02FFA3FBFEFBC6FBB564BE" box="[1053,1140,229,255]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="15" pageNumber="62">Fig. 54</figureCitation>
) somewhat more robust; metatibiae
<figureCitation id="F29A92933B02FFA3FEFAFA2FFEB06567" box="[281,369,268,294]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="15" pageNumber="62">Fig. 54</figureCitation>
; pygidium (
<figureCitation id="F29A92933B02FFA3FE1DFA2FFD946567" box="[510,597,268,294]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="15" pageNumber="62">Fig. 66</figureCitation>
) flattened in lateral view; epipleura (
<figureCitation id="F29A92933B02FFA3FBE8FA2FFBA36567" box="[1035,1122,268,294]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="15" pageNumber="62">Fig. 60</figureCitation>
) strongly narrowed before anterior border of metacoxae, then widened up to level of middle of second ventrite.
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B02FFA3FF26FA7BFECD675D" pageId="15" pageNumber="62" type="distribution">
<paragraph id="6A1E8E163B02FFA3FF26FA7BFF3465DB" blockId="15.[151,1437,151,835]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FA7BFEA16533" bold="true" box="[197,352,344,370]" pageId="15" pageNumber="62">Distribution</emphasis>
:
<collectingCountry id="12B6CE863B02FFA3FE93FA7AFE2C6532" box="[368,493,345,371]" name="Venezuela" pageId="15" pageNumber="62">Venezuela</collectingCountry>
,
<collectingCountry id="12B6CE863B02FFA3FE1AFA7AFD6D6532" box="[505,684,345,371]" name="French Guiana" pageId="15" pageNumber="62">French Guiana</collectingCountry>
,
<collectingCountry id="12B6CE863B02FFA3FD5BFA7AFCC26532" box="[696,771,345,371]" name="Brazil" pageId="15" pageNumber="62">Brazil</collectingCountry>
, and
<collectingCountry id="12B6CE863B02FFA3FCA0FA7AFCBC6532" box="[835,893,345,371]" name="Peru" pageId="15" pageNumber="62">Peru</collectingCountry>
.
<collectingCountry id="12B6CE863B02FFA3FC6AFA7AFBC76532" box="[905,1030,345,371]" name="Venezuela" pageId="15" pageNumber="62">Venezuela</collectingCountry>
,
<collectingCountry id="12B6CE863B02FFA3FBF1FA7AFB9C6532" box="[1042,1117,345,371]" name="Brazil" pageId="15" pageNumber="62">Brazil</collectingCountry>
, and
<collectingCountry id="12B6CE863B02FFA3FB7EFA7AFB126532" box="[1181,1235,345,371]" name="Peru" pageId="15" pageNumber="62">Peru</collectingCountry>
are new country records.
</paragraph>
<paragraph id="6A1E8E163B02FFA3FF26FA85FECD675D" blockId="15.[151,1437,151,835]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FA85FE6D6581" bold="true" box="[197,428,422,448]" pageId="15" pageNumber="62">Specimens studied</emphasis>
:
<collectingCountry id="12B6CE863B02FFA3FE5EFA85FDB56581" box="[445,628,422,448]" name="Venezuela" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FE5EFA85FDB56581" bold="true" box="[445,628,422,448]" pageId="15" pageNumber="62">VENEZUELA</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B02FFA3FD67FA85FCC66581" bold="true" box="[644,775,422,448]" pageId="15" pageNumber="62">Amazonas</emphasis>
: Culebra, P. N. Duida-Marahuaca,
<geoCoordinate id="0F95E8D13B02FFA3FB4AFA85FAD06581" box="[1193,1297,422,448]" direction="north" orientation="latitude" pageId="15" pageNumber="62" precision="925" value="3.7333333">3º 44’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B02FFA3FAFDFA85FA596581" box="[1310,1432,422,448]" direction="west" orientation="longitude" pageId="15" pageNumber="62" precision="925" value="-65.76667">65º 46’ W</geoCoordinate>
,
<date id="1E1FA8D63B02FFA3FF74FAEEFEF465A6" box="[151,309,461,487]" pageId="15" pageNumber="62" value="1992-01-24" valueMax="1992-01-26" valueMin="1992-01-24">24-26-I-1992</date>
, Exped. Terramar, J. Clavijo-A. Chacón, 1 F (MIZA). Pto. Ayacucho,
<date id="1E1FA8D63B02FFA3FB91FAEEFB3C65A6" box="[1138,1277,461,487]" pageId="15" pageNumber="62" value="1967-04-22">22-IV-1967</date>
, P. J. Salinas, 1 M (MIZA);
<date id="1E1FA8D63B02FFA3FEA3FAD7FE30664F" box="[320,497,500,526]" pageId="15" pageNumber="62" value="1982-12-04" valueMax="1982-12-15" valueMin="1982-12-04">4-15-XII-1982</date>
, G. Romero, 4 F (MIZA). Rio Baría,
<geoCoordinate id="0F95E8D13B02FFA3FC53FAD7FBD6664F" box="[944,1047,500,526]" direction="north" orientation="latitude" pageId="15" pageNumber="62" precision="925" value="0.9166667">0º 55’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B02FFA3FBC7FAD7FB5E664F" box="[1060,1183,500,526]" direction="west" orientation="longitude" pageId="15" pageNumber="62" precision="925" value="-66.166664">66º 10’ W</geoCoordinate>
,
<quantity id="AD5923F33B02FFA3FB4FFAD7FB3B664F" box="[1196,1274,500,526]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="62" unit="m" value="140.0">140 m</quantity>
, L. J. Joly-A. Chacón, 1 F (MIZA).
<emphasis id="58D552043B02FFA3FE7FF939FE3F6675" bold="true" box="[412,510,538,564]" pageId="15" pageNumber="62">Barinas</emphasis>
: Reserva Forestal de Ticoporo,
<quantity id="AD5923F33B02FFA3FC94F939FC056675" box="[887,964,538,564]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="62" unit="m" value="230.0">230 m</quantity>
,
<date id="1E1FA8D63B02FFA3FC33F939FB9A6675" box="[976,1115,538,564]" pageId="15" pageNumber="62" value="1968-04-10">10-IV-1968</date>
, en la luz, F. Fernández
<collectingCountry id="12B6CE863B02FFA3FA9AF939FA516675" box="[1401,1424,538,564]" name="Italy" pageId="15" pageNumber="62">Y</collectingCountry>
.- L. J. Joly, 1 F (MIZA).
<emphasis id="58D552043B02FFA3FE4EF963FDCB661B" bold="true" box="[429,522,576,602]" pageId="15" pageNumber="62">Bolívar</emphasis>
: El Barroso, Río Matú,
<date id="1E1FA8D63B02FFA3FCC4F962FC51661A" box="[807,912,577,603]" pageId="15" pageNumber="62" value="1972-01-07">7-I-1972</date>
, J. Salcedo, 1 F (MIZA). Los Pijiguaos,
<quantity id="AD5923F33B02FFA3FA93F962FF7366C3" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="62" unit="m" value="600.0">600 m</quantity>
,
<date id="1E1FA8D63B02FFA3FF5CF94BFEAF66C3" box="[191,366,616,642]" pageId="15" pageNumber="62" value="1992-06-03" valueMax="1992-06-13" valueMin="1992-06-03">3-13-VI-1992</date>
, Exped. MIZA, 1 M (MIZA). Río Guaniamo,
<geoCoordinate id="0F95E8D13B02FFA3FC52F94BFBDF66C3" box="[945,1054,616,642]" direction="north" orientation="latitude" pageId="15" pageNumber="62" precision="925" value="6.75">6º 45’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B02FFA3FBCFF94BFB6D66C3" box="[1068,1196,616,642]" direction="west" orientation="longitude" pageId="15" pageNumber="62" precision="925" value="-66.01667">66º 01’ W</geoCoordinate>
,
<date id="1E1FA8D63B02FFA3FB59F94BFA9A66C3" box="[1210,1371,616,642]" pageId="15" pageNumber="62" value="1979-05-08" valueMax="1979-05-12" valueMin="1979-05-08">8-12-V-1979</date>
, 1 M (MIZA).
<emphasis id="58D552043B02FFA3FEE7F9ADFE5066E9" bold="true" box="[260,401,654,680]" pageId="15" pageNumber="62">Portuguesa</emphasis>
: Biscucuy,
<date id="1E1FA8D63B02FFA3FDF8F9ADFD6566E9" box="[539,676,654,680]" pageId="15" pageNumber="62" value="1981-04-13">13-IV-1981</date>
, 1 M (MIZA).
<emphasis id="58D552043B02FFA3FCB6F9ADFC7966E9" bold="true" box="[853,952,654,680]" pageId="15" pageNumber="62">Táchira</emphasis>
: Nula,
<date id="1E1FA8D63B02FFA3FBF3F9ADFB4D66E9" box="[1040,1164,654,680]" pageId="15" pageNumber="62" value="1967-03-09">9-III-1967</date>
, L. J. Joly-V. Obregón, 2 M, 1 F (MHNLS).
<collectingCountry id="12B6CE863B02FFA3FE69F996FE3B668E" box="[394,506,693,719]" name="Brazil" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FE69F996FE3B668E" bold="true" box="[394,506,693,719]" pageId="15" pageNumber="62">BRAZIL</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B02FFA3FDE9F996FD87668E" bold="true" box="[522,582,693,719]" pageId="15" pageNumber="62">Pará</emphasis>
: Cachimbo,
<quantity id="AD5923F33B02FFA3FD3FF996FCE8668E" box="[732,809,693,719]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="62" unit="m" value="400.0">400 m</quantity>
, 14/
<date id="1E1FA8D63B02FFA3FCB9F996FC27668E" box="[858,998,693,719]" pageId="15" pageNumber="62" value="1955-09-21">21-IX-1955</date>
, L. Travassos &amp; S. Oliveira, 5 M, 1 F (HAHC).
<collectingCountry id="12B6CE863B02FFA3FEEFF9F8FE9D66B4" box="[268,348,731,757]" name="Peru" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FEEFF9F8FE9D66B4" bold="true" box="[268,348,731,757]" pageId="15" pageNumber="62">PERU</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B02FFA3FE88F9F8FE1C66B4" bold="true" box="[363,477,731,757]" pageId="15" pageNumber="62">Huánuco</emphasis>
: Tingo María,
<date id="1E1FA8D63B02FFA3FD6DF9FFFCCE66B7" box="[654,783,732,758]" pageId="15" pageNumber="62" value="1947-05-17">17-V-1947</date>
,
<quantity id="AD5923F33B02FFA3FCF9F9FFFCB966B7" box="[794,888,732,758]" metricMagnitude="2" metricUnit="m" metricValue="6.7056" pageId="15" pageNumber="62" unit="ft" value="2200.0">2.200 ft</quantity>
, J. C. Pallister Coll. Donor Frank Johnson, 1 F (AMNH).
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B02FFA3FF26F80BFB3F6702" box="[197,1278,808,835]" pageId="15" pageNumber="62" type="description">
<paragraph id="6A1E8E163B02FFA3FF26F80BFB3F6702" blockId="15.[151,1437,151,835]" box="[197,1278,808,835]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26F80BFE456703" bold="true" box="[197,388,808,834]" pageId="15" pageNumber="62">Other localities</emphasis>
:
<collectingCountry id="12B6CE863B02FFA3FE77F80BFD4C6703" box="[404,653,808,834]" name="French Guiana" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FE77F80BFD4C6703" bold="true" box="[404,653,808,834]" pageId="15" pageNumber="62">FRENCH GUIANA</emphasis>
</collectingCountry>
: Route de Kau; Saül; St. Laurent (
<bibRefCitation id="0E30F3E73B02FFA3FBC2F80AFB306702" author="Dechambre" box="[1057,1265,809,835]" pageId="15" pageNumber="64" refString="Dechambre, R. - P. (1992) Nouveaux Cyclocephalini des genres Cyclocephala et Aspidolea (Col. Dynastidae). Les Coleopteres du Monde, 14, 57 - 75, color plate 12." type="journal article" year="1992">Dechambre 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

232
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<mods:title id="C95888CD46CA973CFD5CBFAA96EB03F4">Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
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<subSubSection id="22BBDD9D3B01FFA0FF74F939FDEC6616" pageId="12" pageNumber="59" type="nomenclature">
<paragraph id="6A1E8E163B01FFA0FF74F939FD9D6675" blockId="12.[151,604,538,599]" box="[151,604,538,564]" pageId="12" pageNumber="59">
<heading id="3156397A3B01FFA0FF74F939FD9D6675" bold="true" box="[151,604,538,564]" fontSize="11" level="1" pageId="12" pageNumber="59" reason="1">
<taxonomicName id="ADA1F5953B01FFA0FF74F939FD9D6675" authority="Ratcliffe, 1992" authorityName="Ratcliffe" authorityYear="1992" box="[151,604,538,564]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="59" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B01FFA0FF74F939FD9D6675" bold="true" box="[151,604,538,564]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF74F939FE606675" bold="true" box="[151,417,538,564]" italics="true" pageId="12" pageNumber="59">Cyclocephala sarahae</emphasis>
<bibRefCitation id="0E30F3E73B01FFA0FE4BF939FD9D6675" author="Ratcliffe" box="[424,604,538,564]" pageId="12" pageNumber="64" refString="Ratcliffe, B. C. (1992) New species and country records of Brazilian Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae). Tidschrift voor Entomologie, 135 (2), 179 - 190." type="journal article" year="1992">Ratcliffe, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF74F91EFDEC6616" blockId="12.[151,604,538,599]" box="[151,557,573,599]" pageId="12" pageNumber="59">
(
<figureCitation id="F29A92933B01FFA0FF7CF91EFF306616" box="[159,241,573,599]" captionStart="FIGURE 1" captionStartId="4.[151,255,1052,1076]" captionTargetBox="[378,1217,320,1022]" captionTargetId="figure@4.[367,1219,313,1027]" captionTargetPageId="4" captionText="FIGURE 1. Distribution of Cyclocephala bicolor (circles), C. anibali, sp. n. (triangle), and C. sarahae (square)." httpUri="https://zenodo.org/record/186486/files/figure.png" pageId="12" pageNumber="59">Figs. 1</figureCitation>
,
<figureCitation id="F29A92933B01FFA0FF1CF91EFEDF6616" box="[255,286,573,599]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="12" pageNumber="59">21</figureCitation>
,
<figureCitation id="F29A92933B01FFA0FEC8F91EFE8A6616" box="[299,331,573,599]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="12" pageNumber="59">28</figureCitation>
,
<figureCitation id="F29A92933B01FFA0FEB4F91EFE0E6616" box="[343,463,573,599]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="12" pageNumber="59">37–38, 43</figureCitation>
,
<figureCitation id="F29A92933B01FFA0FE39F91EFE3B6616" box="[474,506,573,599]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="12" pageNumber="59">57</figureCitation>
,
<figureCitation id="F29A92933B01FFA0FDE4F91EFDE46616" box="[519,549,573,599]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="12" pageNumber="59">75</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B01FFA1FF74F9A9FF3C60AC" lastPageId="13" lastPageNumber="60" pageId="12" pageNumber="59" type="description">
<paragraph id="6A1E8E163B01FFA0FF74F9A9FCCC66B3" blockId="12.[151,1437,650,2030]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF74F9A9FE8B66E5" bold="true" box="[151,330,650,676]" pageId="12" pageNumber="59">Bibliography.</emphasis>
<taxonomicName id="ADA1F5953B01FFA0FEB0F9A9FCA466E5" authority="Ratcliffe 1992: 187" authorityName="Ratcliffe" authorityPageNumber="187" authorityYear="1992" box="[339,869,650,676]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="59" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B01FFA0FEB0F9A9FDA866E2" box="[339,617,650,675]" italics="true" pageId="12" pageNumber="59">Cyclocephala sarahae</emphasis>
<bibRefCitation id="0E30F3E73B01FFA0FD90F9A9FCEB66E5" author="Ratcliffe" box="[627,810,650,676]" pageId="12" pageNumber="64" refString="Ratcliffe, B. C. (1992) New species and country records of Brazilian Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae). Tidschrift voor Entomologie, 135 (2), 179 - 190." type="journal article" year="1992">Ratcliffe 1992</bibRefCitation>
: 187
</taxonomicName>
(description male, female; compared with
<taxonomicName id="ADA1F5953B01FFA0FA63F9A9FF2D668B" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="59" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B01FFA0FA63F9A9FA5266E2" box="[1408,1427,650,675]" italics="true" pageId="12" pageNumber="59">C</emphasis>
.
<emphasis id="58D552043B01FFA0FF74F992FF2D668B" box="[151,236,689,714]" italics="true" pageId="12" pageNumber="59">bicolor</emphasis>
</taxonomicName>
,
<taxonomicName id="ADA1F5953B01FFA0FF1FF992FE40668B" box="[252,385,689,715]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="59" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="58D552043B01FFA0FF1FF992FECE668B" box="[252,271,689,714]" italics="true" pageId="12" pageNumber="59">C</emphasis>
.
<emphasis id="58D552043B01FFA0FEFCF992FE40668B" box="[287,385,689,714]" italics="true" pageId="12" pageNumber="59">testacea</emphasis>
</taxonomicName>
ab.
<emphasis id="58D552043B01FFA0FE56F992FDCC668B" box="[437,525,689,714]" italics="true" pageId="12" pageNumber="59">ovulum</emphasis>
),
<figureCitation id="F29A92933B01FFA0FDC4F992FDAF668A" box="[551,622,689,715]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="12" pageNumber="59">Fig. 6</figureCitation>
(photograph),
<figureCitation id="F29A92933B01FFA0FCC2F992FC6E668A" box="[801,943,689,715]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="12" pageNumber="59">Figs. 27–28</figureCitation>
(genitalia).
<typeStatus id="B51A30B43B01FFA0FBA1F992FB70668A" box="[1090,1201,689,715]" pageId="12" pageNumber="59" type="holotype">Holotype</typeStatus>
male UNSM.
<typeStatus id="B51A30B43B01FFA0FA82F992FA5D668A" box="[1377,1436,689,715]" pageId="12" pageNumber="59">Type</typeStatus>
locality:
<collectingCountry id="12B6CE863B01FFA0FF1DF9FBFE8766B3" box="[254,326,728,754]" name="Brazil" pageId="12" pageNumber="59">Brazil</collectingCountry>
(Amazonas, Rio Demeni, nr. equator).
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF26F9DDFBBD677E" blockId="12.[151,1437,650,2030]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26F9DDFEA56759" bold="true" box="[197,356,766,792]" pageId="12" pageNumber="59">Description.</emphasis>
Short, oval, moderately shiny. Brownish red, frons and vertex black, elytra testaceous. Dorsal surface and pygidium glabrous, ventral pubescence sparse.
<emphasis id="58D552043B01FFA0FC47F806FC3F677E" bold="true" box="[932,1022,805,831]" pageId="12" pageNumber="59">Length</emphasis>
: 11.4 mm.
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF26F868FCDF6035" blockId="12.[151,1437,650,2030]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26F868FEE16724" bold="true" box="[197,288,843,869]" pageId="12" pageNumber="59">MALE</emphasis>
:
<emphasis id="58D552043B01FFA0FED2F868FE5A6724" bold="true" box="[305,411,843,869]" pageId="12" pageNumber="59">Clypeus</emphasis>
about 2.8 times as wide across base as long; sides straight, strongly convergent to truncate and feebly reflexed apex; shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view flat to slightly concave on apical half; surface finely alutaceous, with shallow and widely annulate punctures, the annulation wide, not well defined posteriorly, most punctures confluent, appearing as irregular and fine, more or less transverse striae. Frontoclypeal suture thin, bisinuate, not pointed posteriorly at middle.
<emphasis id="58D552043B01FFA0FDCBFF2EFDB06066" bold="true" box="[552,625,1037,1063]" pageId="12" pageNumber="59">Frons</emphasis>
: width about 2.8 times transverse diameter of eye, finely alutaceous; punctures smaller than those of clypeus, somewhat transverse by confluence of punctures, most punctures separated by distances more or less their own diameter.
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF26FFA3FC2F61C2" blockId="12.[151,1437,650,2030]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26FFA3FE8860DB" bold="true" box="[197,329,1152,1178]" pageId="12" pageNumber="59">Mandibles</emphasis>
with external border sinuate, apex projecting externally.
<emphasis id="58D552043B01FFA0FC0AFFA3FB9760DB" bold="true" box="[1001,1110,1152,1178]" pageId="12" pageNumber="59">Maxillae</emphasis>
not visible; maxillary palpi (
<figureCitation id="F29A92933B01FFA0FF43FF8BFF3B6083" box="[160,250,1192,1218]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="12" pageNumber="59">Fig. 21</figureCitation>
) moderately robust, second palpomere longer than third, scarcely widened towards apex, fourth fusiform, 3.5 times longer than preceding.
<emphasis id="58D552043B01FFA0FD6AFFEDFD2D60A9" bold="true" box="[649,748,1230,1256]" pageId="12" pageNumber="59">Labium</emphasis>
narrow, lateral borders slightly arched on basal two-thirds, slightly constricted at middle of apical third; apex truncate on middle of ventral side, more arcuately produced and with small emargination at middle on oral side, and fimbriate with long, fine blackish brown setae; surface convex on anterior part, depressed on center of base, basal two-thirds with strong, dense piliferous punctures, the setae long; anterior third with minute, scattered punctures.
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF26FEACFAEB61EB" blockId="12.[151,1437,650,2030]" box="[197,1322,1423,1450]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26FEACFEFF61E8" bold="true" box="[197,318,1423,1449]" pageId="12" pageNumber="59">Antennae</emphasis>
with 10 antennomeres, club about 2.0 times the length of antennomeres 2–7 together.
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF26FE95FC916347" blockId="12.[151,1437,650,2030]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26FE95FE826191" bold="true" box="[197,323,1462,1488]" pageId="12" pageNumber="59">Pronotum</emphasis>
1.5 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected at middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface finely alutaceous (
<date id="1E1FA8D63B01FFA0FEC4FD09FEA06205" box="[295,353,1578,1604]" pageId="12" pageNumber="59" value="1940-10">40X</date>
); punctures annulate, moderately fine and dense, the distance between most punctures on disc as long as or longer than diameter of punctures, punctures gradually larger towards sides, the diameter of a puncture at basal angle at least 2.0 times the diameter of those on center of disc, more dense on basal angles, where the distance between most punctures is as large or smaller than the diameter of a puncture; surface also covered with numerous minute punctures (
<date id="1E1FA8D63B01FFA0FD7DFDE6FD12629E" box="[670,723,1733,1759]" pageId="12" pageNumber="59" value="1950-10">50X</date>
or more).
<emphasis id="58D552043B01FFA0FCB6FDE6FC17629E" bold="true" box="[853,982,1733,1759]" pageId="12" pageNumber="59">Scutellum</emphasis>
triangular, sides feebly arcuate, apex briefly rounded; punctures similar to those of pronotal disc.
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF26FC31FE7F63AF" blockId="12.[151,1437,650,2030]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26FC31FED7636D" bold="true" box="[197,278,1810,1836]" pageId="12" pageNumber="59">Elytra</emphasis>
1.2 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones weak; a feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident, also with numerous minute punctures (
<date id="1E1FA8D63B01FFA0FF7CFC8EFF126386" box="[159,211,1965,1991]" pageId="12" pageNumber="59" value="1950-10">50X</date>
or more) scattered over all the surface; sutural interstria flat, weakly elevated on apical half. Epipleura narrow, more so apically.
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF26FBB4FE03650D" blockId="13.[151,1437,151,1376]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26FBB4FEFC64F0" bold="true" box="[197,317,151,177]" pageId="13" pageNumber="60">Pygidium</emphasis>
2.2 times as wide as long; in posterior view the inferior borders slightly convex, converging to widely rounded apex in an angle distinctly greater than 90°, but less than in
<taxonomicName id="ADA1F5953B00FFA1FBC3FB9CFB596499" box="[1056,1176,190,216]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="60" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B00FFA1FBC3FB9CFBF26499" box="[1056,1075,191,216]" italics="true" pageId="13" pageNumber="60">C</emphasis>
.
<emphasis id="58D552043B00FFA1FBA0FB9CFB596499" box="[1091,1176,191,216]" italics="true" pageId="13" pageNumber="60">bicolor</emphasis>
</taxonomicName>
; apex not truncate; in lateral view uniformly convex; surface smooth, shining, with large annulate punctures, more dense on lateral angles; shiner apically, the punctures smaller and less dense, distances between most punctures more than 2.0 times their own diameter.
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF26FA7BFDB7672B" blockId="13.[151,1437,151,1376]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26FA7BFEF96533" bold="true" box="[197,312,344,370]" pageId="13" pageNumber="60">Protibiae</emphasis>
(
<figureCitation id="F29A92933B00FFA1FEABFA7AFE5F6532" box="[328,414,345,371]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="13" pageNumber="60">Fig. 28</figureCitation>
) tridentate, small basal tooth separated from intermediate tooth by a distance 1.5 times as long as that between the intermediate and the anterior teeth; intermediate and anterior teeth well developed, basal tooth with its anterior border more or less perpendicular to longitudinal axis of tibiae; apical spur almost straight, slightly surpassing apex of first tarsomere.
<emphasis id="58D552043B00FFA1FD1DFAEEFCA565A6" bold="true" box="[766,868,461,487]" pageId="13" pageNumber="60">Protarsi</emphasis>
(
<figureCitation id="F29A92933B00FFA1FC96FAEEFC3665A6" box="[885,1015,461,487]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="13" pageNumber="60">Fig. 37–38</figureCitation>
) enlarged; tarsomere 5 shorter than 1–4 together, without ventro–internal carina, ventral side with very short and very fine longitudinal striae on inner side of apex; tarsomere 4 longitudinally striate on apical half, tarsomere 3 striated on apical third; largest claw (
<figureCitation id="F29A92933B00FFA1FF02F962FEF8661A" box="[225,313,577,603]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="13" pageNumber="60">Fig. 43</figureCitation>
) wide, flattened, narrowly cleft at apex, the external border of small branch continuous with internal curvature of the claw. All
<emphasis id="58D552043B00FFA1FDD0F944FD4C66C0" bold="true" box="[563,653,615,641]" pageId="13" pageNumber="60">femora</emphasis>
almost uniformly, not densely pilose, especially the mesofemora, on which, in addition to the normal longitudinal series of piliferous punctures close to anterior and posterior borders (seen in natural position of the leg), with numerous piliferous punctures over all the surface.
<emphasis id="58D552043B00FFA1FF74F9F8FEE466B4" bold="true" box="[151,293,731,757]" pageId="13" pageNumber="60">Metafemur</emphasis>
(
<figureCitation id="F29A92933B00FFA1FED6F9FFFE4B66B7" box="[309,394,732,758]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="13" pageNumber="60">Fig. 57</figureCitation>
) 2.3 times as long as its maximum width;
<emphasis id="58D552043B00FFA1FC62F9F8FBC466B4" bold="true" box="[897,1029,731,757]" pageId="13" pageNumber="60">metatibiae</emphasis>
(
<figureCitation id="F29A92933B00FFA1FBF5F9FFFBAA66B7" box="[1046,1131,732,758]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="13" pageNumber="60">Fig. 57</figureCitation>
) robust, widened at apex, bristles on diagonal carina broken in the
<typeStatus id="B51A30B43B00FFA1FD66F821FD30675D" box="[645,753,770,796]" pageId="13" pageNumber="60" type="holotype">holotype</typeStatus>
, and those of apex very strong (mostly missing from the
<typeStatus id="B51A30B43B00FFA1FF74F80AFEC66702" box="[151,263,809,835]" pageId="13" pageNumber="60" type="holotype">holotype</typeStatus>
).
<emphasis id="58D552043B00FFA1FEF8F80BFE586703" bold="true" box="[283,409,808,834]" pageId="13" pageNumber="60">Metatarsi</emphasis>
subequal in length to tibiae, with first tarsomere widened towards apex, with apical bristles similar to those of apex of tibiae.
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF26F855FB8667F6" blockId="13.[151,1437,151,1376]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26F855FE0E67D1" bold="true" box="[197,463,886,912]" pageId="13" pageNumber="60">Prosternal projection</emphasis>
high, apex alutaceous, anterior part transversely oval, externally pointed; posterior part shorter but wider than anterior, posterior border fimbriate with long bristles.
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF26F8E0FA8A6013" blockId="13.[151,1437,151,1376]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26F8E0FE11679C" bold="true" box="[197,464,963,989]" pageId="13" pageNumber="60">Pronotal hypomera</emphasis>
alutaceous, with sparse erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large, dense, confluent, annulate punctures; disc of metasternum smooth, with small, scattered punctures; metacoxae with punctures similar to pleurae but smaller and more disperse. Ventrites with annulate, moderately dense punctures, punctures smaller and sparser on apical ventrites.
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF26FF7DFE666039" blockId="13.[151,1437,151,1376]" box="[197,423,1118,1144]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26FF7DFEFC6039" bold="true" box="[197,317,1118,1144]" pageId="13" pageNumber="60">Aedeagus</emphasis>
:
<figureCitation id="F29A92933B00FFA1FEAEFF7DFE656039" box="[333,420,1118,1144]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="13" pageNumber="60">Fig. 75</figureCitation>
.
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF26FFA6FF3C60AC" blockId="13.[151,1437,151,1376]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26FFA6FE8760DE" bold="true" box="[197,326,1157,1183]" pageId="13" pageNumber="60">FEMALE</emphasis>
: differs from the male in the following characters: punctation generally stronger; protarsi slender; metafemora more robust; epipleura differences with
<taxonomicName id="ADA1F5953B00FFA1FC6FFF8FFBCB6084" box="[908,1034,1196,1222]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="60" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B00FFA1FC6FFF8FFC5E6084" box="[908,927,1196,1221]" italics="true" pageId="13" pageNumber="60">C</emphasis>
.
<emphasis id="58D552043B00FFA1FC52FF8FFBCB6084" box="[945,1034,1196,1221]" italics="true" pageId="13" pageNumber="60">bicolor</emphasis>
</taxonomicName>
not observed when studying the allotype.
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B00FFA1FF26FFDBFDB76121" pageId="13" pageNumber="60" type="distribution">
<paragraph id="6A1E8E163B00FFA1FF26FFDBFC066152" blockId="13.[151,1437,151,1376]" box="[197,967,1272,1299]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26FFDBFE9F6153" bold="true" box="[197,350,1272,1298]" pageId="13" pageNumber="60">Distribution</emphasis>
:
<collectingCountry id="12B6CE863B00FFA1FE8EFFDAFE786152" box="[365,441,1273,1299]" name="Brazil" pageId="13" pageNumber="60">Brazil</collectingCountry>
: Amazonas (only known from
<typeStatus id="B51A30B43B00FFA1FCC7FFDAFC966152" box="[804,855,1273,1299]" pageId="13" pageNumber="60">type</typeStatus>
locality).
</paragraph>
<paragraph id="6A1E8E163B00FFA1FF26FE3CFDB76121" blockId="13.[151,1437,151,1376]" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FF26FE3CFE716178" bold="true" box="[197,432,1311,1337]" pageId="13" pageNumber="60">Specimens studied</emphasis>
:
<collectingCountry id="12B6CE863B00FFA1FE22FE3CFDF26178" box="[449,563,1311,1337]" name="Brazil" pageId="13" pageNumber="60">
<emphasis id="58D552043B00FFA1FE22FE3CFDF26178" bold="true" box="[449,563,1311,1337]" pageId="13" pageNumber="60">BRAZIL</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B00FFA1FDA7FE3CFD096178" bold="true" box="[580,712,1311,1337]" pageId="13" pageNumber="60">Amazonas</emphasis>
: Rio Demeni, N. R. Equator,
<date id="1E1FA8D63B00FFA1FBD3FE03FB61617B" box="[1072,1184,1312,1338]" pageId="13" pageNumber="60" value="1978-04-10">IV-10-78</date>
, L. A. Lacey,
<specimenCount id="7CA7459F3B00FFA1FAA8FE03FA5D617B" box="[1355,1436,1312,1338]" pageId="13" pageNumber="60" type="male">1 male</specimenCount>
(
<typeStatus id="B51A30B43B00FFA1FF7CFE65FECD6121" box="[159,268,1350,1376]" pageId="13" pageNumber="60" type="holotype">holotype</typeStatus>
),
<specimenCount id="7CA7459F3B00FFA1FEFCFE65FE446121" box="[287,389,1350,1376]" pageId="13" pageNumber="60" type="female">1 female</specimenCount>
(allotype) (UNSM).
</paragraph>
</subSubSection>
</treatment>
</document>

255
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@ -0,0 +1,255 @@
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<mods:title id="3B9EA644DB1ADAB52EBDE53C4521681F">Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
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<mods:date id="D3C49D1E9C62B3B5FCDA27F4342A7CD9">2009</mods:date>
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<subSubSection id="22BBDD9D3B02FFA3FF74F8BEFC8F679B" pageId="15" pageNumber="62" type="nomenclature">
<paragraph id="6A1E8E163B02FFA3FF74F8BEFDF567F6" blockId="15.[151,846,925,986]" box="[151,564,925,951]" pageId="15" pageNumber="62">
<heading id="3156397A3B02FFA3FF74F8BEFDF567F6" bold="true" box="[151,564,925,951]" fontSize="11" level="1" pageId="15" pageNumber="62" reason="1">
<emphasis id="58D552043B02FFA3FF74F8BEFDF567F6" bold="true" box="[151,564,925,951]" pageId="15" pageNumber="62">
<taxonomicName id="ADA1F5953B02FFA3FF74F8BEFE5467F6" box="[151,405,925,951]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="62" phylum="Arthropoda" rank="species" species="anibali" status="sp. nov.">
<emphasis id="58D552043B02FFA3FF74F8BEFE5467F6" bold="true" box="[151,405,925,951]" italics="true" pageId="15" pageNumber="62">Cyclocephala anibali</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="43E6EF7F3B02FFA3FE47F8BEFDF567F6" box="[420,564,925,951]" pageId="15" pageNumber="62" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E8E163B02FFA3FF74F8E3FC8F679B" blockId="15.[151,846,925,986]" box="[151,846,960,986]" pageId="15" pageNumber="62">
<figureCitation id="F29A92933B02FFA3FF74F8E3FF26679B" box="[151,231,960,986]" captionStart="FIGURE 1" captionStartId="4.[151,255,1052,1076]" captionTargetBox="[378,1217,320,1022]" captionTargetId="figure@4.[367,1219,313,1027]" captionTargetPageId="4" captionText="FIGURE 1. Distribution of Cyclocephala bicolor (circles), C. anibali, sp. n. (triangle), and C. sarahae (square)." httpUri="https://zenodo.org/record/186486/files/figure.png" pageId="15" pageNumber="62">Figs. 1</figureCitation>
,
<figureCitation id="F29A92933B02FFA3FF16F8E3FEAC679B" box="[245,365,960,986]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="15" pageNumber="62">15–18, 23</figureCitation>
,
<figureCitation id="F29A92933B02FFA3FE9AF8E3FE04679B" box="[377,453,960,986]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="15" pageNumber="62">31–32</figureCitation>
,
<figureCitation id="F29A92933B02FFA3FE32F8E3FE30679B" box="[465,497,960,986]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="15" pageNumber="62">40</figureCitation>
, 48,
<figureCitation id="F29A92933B02FFA3FDCAF8E3FDB5679B" box="[553,628,960,986]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="15" pageNumber="62">55–56</figureCitation>
,
<figureCitation id="F29A92933B02FFA3FD63F8E3FD61679B" box="[640,672,960,986]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="15" pageNumber="62">61</figureCitation>
, 67–68, 76–77
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B02FFA3FF74FF2EFB056035" pageId="15" pageNumber="62" type="materials_examined">
<paragraph id="6A1E8E163B02FFA3FF74FF2EFB056035" blockId="15.[151,1437,1037,2030]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF74FF2EFE456066" bold="true" box="[151,388,1037,1063]" pageId="15" pageNumber="62">
<typeStatus id="B51A30B43B02FFA3FF74FF2EFEFE6066" box="[151,319,1037,1063]" pageId="15" pageNumber="62" type="holotype">HOLOTYPE</typeStatus>
male
</emphasis>
,
<emphasis id="58D552043B02FFA3FE76FF2EFC9F6066" bold="true" box="[405,862,1037,1063]" pageId="15" pageNumber="62">
ALLOTYPE female.
<collectingCountry id="12B6CE863B02FFA3FD46FF2EFC9F6066" box="[677,862,1037,1063]" name="Venezuela" pageId="15" pageNumber="62">VENEZUELA</collectingCountry>
</emphasis>
:
<emphasis id="58D552043B02FFA3FC8CFF2EFC016066" bold="true" box="[879,960,1037,1063]" pageId="15" pageNumber="62">Apure</emphasis>
: Capanaparo, Hato Los Turpiales,
<date id="1E1FA8D63B02FFA3FA85FF2EFF17600F" pageId="15" pageNumber="62" value="1980-05-08">8-V- 1980</date>
, E. Osuna (
<collectionCode id="0CB016D33B02FFA3FE81FF17FE74600F" LSID="urn:lsid:biocol.org:col:33834" box="[354,437,1076,1102]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="15" pageNumber="62">MIZA</collectionCode>
).
<typeStatus id="B51A30B43B02FFA3FE2AFF10FDB6600C" box="[457,631,1075,1101]" pageId="15" pageNumber="62" type="paratype">
<emphasis id="58D552043B02FFA3FE2AFF10FDB6600C" bold="true" box="[457,631,1075,1101]" pageId="15" pageNumber="62">PARATYPES</emphasis>
</typeStatus>
.
<collectingCountry id="12B6CE863B02FFA3FD65FF10FCFE600C" box="[646,831,1075,1101]" name="Venezuela" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FD65FF10FCFE600C" bold="true" box="[646,831,1075,1101]" pageId="15" pageNumber="62">VENEZUELA</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B02FFA3FCB3FF10FC60600C" bold="true" box="[848,929,1075,1101]" pageId="15" pageNumber="62">Apure</emphasis>
: Fundo La Florida [
<geoCoordinate id="0F95E8D13B02FFA3FB77FF17FAF4600F" box="[1172,1333,1076,1102]" direction="north" orientation="latitude" pageId="15" pageNumber="62" precision="15" value="6.875">6° 52’ 30” N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B02FFA3FAA1FF17FF286035" direction="west" orientation="longitude" pageId="15" pageNumber="62" precision="15" value="-68.96222">68° 57’ 44” W</geoCoordinate>
], cr. río Quitaparo,
<geoCoordinate id="0F95E8D13B02FFA3FE2FFF79FDED6035" box="[460,556,1114,1140]" direction="north" orientation="latitude" pageId="15" pageNumber="62" precision="925" value="7.0833335">7° 05’N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B02FFA3FDDAFF79FD6A6035" box="[569,683,1114,1140]" direction="west" orientation="longitude" pageId="15" pageNumber="62" precision="925" value="-68.6">68° 36’W</geoCoordinate>
,
<date id="1E1FA8D63B02FFA3FD54FF79FCAD6035" box="[695,876,1114,1140]" pageId="15" pageNumber="62" value="2001-11-12" valueMax="2001-11-13" valueMin="2001-11-12">12-13-XI-2001</date>
, E. Osuna, 1 M, 1 F (
<collectionCode id="0CB016D33B02FFA3FB85FF79FB796035" LSID="urn:lsid:biocol.org:col:33834" box="[1126,1208,1114,1140]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="15" pageNumber="62">MIZA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B02FFBCFF26FFA3FCD66209" lastPageId="16" lastPageNumber="63" pageId="15" pageNumber="62" type="description">
<paragraph id="6A1E8E163B02FFA3FF26FFA3FEC760A9" blockId="15.[151,1437,1037,2030]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FFA3FDD960DB" bold="true" box="[197,536,1152,1178]" pageId="15" pageNumber="62">
Description.
<typeStatus id="B51A30B43B02FFA3FE87FFA3FE1760DB" box="[356,470,1152,1178]" pageId="15" pageNumber="62" type="holotype">Holotype</typeStatus>
male
</emphasis>
. Short, oval, moderately shiny. Brownish red, frons, except anterior part, and vertex black, elytra testaceous. Dorsal surface and pygidium glabrous; ventral pubescence sparse.
<emphasis id="58D552043B02FFA3FADBFF84FA526080" bold="true" box="[1336,1427,1191,1217]" pageId="15" pageNumber="62">Length</emphasis>
: 11.0 mm.
</paragraph>
<paragraph id="6A1E8E163B02FFA3FF26FFD6FC2C61B6" blockId="15.[151,1437,1037,2030]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FFD6FEEF614E" bold="true" box="[197,302,1269,1295]" pageId="15" pageNumber="62">Clypeus</emphasis>
(
<figureCitation id="F29A92933B02FFA3FEDCFFD6FE58614E" box="[319,409,1269,1295]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="15" pageNumber="62">Fig. 15</figureCitation>
) about 2.3 times as wide across base as long; sides straight, strongly convergent to truncate and arcuately reflexed apex; shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view flat to slightly concave on apical half; surface finely alutaceous, with fine, discrete, shallow, punctures thinner and sparser toward apex and anterior half of sides. Frontoclypeal suture thin, bisinuate and angularly projecting posteriorly at middle.
<emphasis id="58D552043B02FFA3FC20FEACFBCD61E8" bold="true" box="[963,1036,1423,1449]" pageId="15" pageNumber="62">Frons</emphasis>
: width about 2.8 times transverse diameter of eye, finely alutaceous; punctures, except on smooth longitudinal middle line, fine, confluent by pairs so they appear as transverse, stronger and not confluent posteriorly.
</paragraph>
<paragraph id="6A1E8E163B02FFA3FF26FD20FD39636D" blockId="15.[151,1437,1037,2030]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FD20FE90625C" bold="true" box="[197,337,1539,1565]" pageId="15" pageNumber="62">Mandibles</emphasis>
with external border sinuate, apex projecting externally.
<emphasis id="58D552043B02FFA3FBC6FD20FB58625C" bold="true" box="[1061,1177,1539,1565]" pageId="15" pageNumber="62">Maxillae</emphasis>
(
<figureCitation id="F29A92933B02FFA3FB4FFD27FAC9625F" box="[1196,1288,1540,1566]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="15" pageNumber="62">Fig. 18</figureCitation>
) with galea moderately robust, with two apical, one median, and three basal teeth, the basal teeth very small and arising from a common base; stipes anteriorly projected on inner side, and with a brush of fine yellowish brown setae on apex of projection; maxillary palpi (
<figureCitation id="F29A92933B02FFA3FD88FD5BFD0262D3" box="[619,707,1656,1682]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="15" pageNumber="62">Fig. 23</figureCitation>
) moderately robust, second palpomere subcylindrical, longer than third, fourth fusiform, more than 3.0 times as long as preceding.
<emphasis id="58D552043B02FFA3FC02FDBDFB8762F9" bold="true" box="[993,1094,1694,1720]" pageId="15" pageNumber="62">Labium</emphasis>
(
<figureCitation id="F29A92933B02FFA3FBB4FDBDFB6E62F9" box="[1111,1199,1694,1720]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="15" pageNumber="62">Fig. 17</figureCitation>
) with sides slightly convex on basal two-thirds, not constricted at middle of apical third; apex convex and widely emarginated at middle, and fimbriate with long, yellowish brown setae; surface almost flat, with one longitudinal series of long bristles on each side, converging towards base.
</paragraph>
<paragraph id="6A1E8E163B02FFA3FF26FC1BFAEB6312" blockId="15.[151,1437,1037,2030]" box="[197,1322,1848,1875]" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FC1BFEFF6313" bold="true" box="[197,318,1848,1874]" pageId="15" pageNumber="62">Antennae</emphasis>
with 10 antennomeres, club about 1.5 times the length of antennomeres 2–7 together.
</paragraph>
<paragraph id="6A1E8E163B02FFBCFF26FC7CFD17650D" blockId="15.[151,1437,1037,2030]" lastBlockId="16.[151,1437,152,2034]" lastPageId="16" lastPageNumber="63" pageId="15" pageNumber="62">
<emphasis id="58D552043B02FFA3FF26FC7CFE826338" bold="true" box="[197,323,1887,1913]" pageId="15" pageNumber="62">Pronotum</emphasis>
1.5 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected in middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface smooth (
<date id="1E1FA8D63B02FFA3FF7CFCF7FF1663AF" box="[159,215,2004,2030]" pageId="15" pageNumber="62" value="1940-10">40X</date>
); punctures annulate, moderately fine and dense, the distance between most punctures on disc as long as or longer than diameter of punctures, punctures gradually larger towards sides, the diameter of a puncture about 1.5 times the diameter of those on center of disc, more dense on basal angles, where the distance between most punctures is as large as or smaller than diameter of punctures; all surface also covered with numerous minute punctures (
<date id="1E1FA8D63B1DFFBCFE0DFA2FFDE36567" box="[494,546,268,294]" pageId="16" pageNumber="63" value="1950-10">50X</date>
or more).
<emphasis id="58D552043B1DFFBCFD43FA28FCDC6564" bold="true" box="[672,797,267,293]" pageId="16" pageNumber="63">Scutellum</emphasis>
triangular, sides feebly arcuate, apex briefly rounded; punctures much finer than those of pronotal disc.
</paragraph>
<paragraph id="6A1E8E163B1DFFBCFF26FA7BFE7F6675" blockId="16.[151,1437,152,2034]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26FA7BFED76533" bold="true" box="[197,278,344,370]" pageId="16" pageNumber="63">Elytra</emphasis>
1.2 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones slight; a feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident, also with numerous minute punctures (
<date id="1E1FA8D63B1DFFBCFF7CFAD7FF12664F" box="[159,211,500,526]" pageId="16" pageNumber="63" value="1950-10">50X</date>
or more) scattered over all the surface; sutural interstria flat, weakly elevated on apical half. Epipleura narrow, more so apically.
</paragraph>
<paragraph id="6A1E8E163B1DFFBCFF26F963FED0675D" blockId="16.[151,1437,152,2034]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26F963FEFC661B" bold="true" box="[197,317,576,602]" pageId="16" pageNumber="63">Pygidium</emphasis>
1.9 times as wide as long; in posterior view the inferior borders almost straight, converging to widely and feebly convex-truncate apex at an angle of about 90°, in lateral view (
<figureCitation id="F29A92933B1DFFBCFBBFF94BFB7366C3" box="[1116,1202,616,642]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="16" pageNumber="63">Fig. 67</figureCitation>
) uniformly convex, more strongly on apical half; surface smooth, shiny, with small and dense annulate punctures, distances between most punctures as large as or smaller than diameter of punctures on basal half; punctures on apical region much smaller, simple, the distances between most punctures much longer than the diameter of punctures.
</paragraph>
<paragraph id="6A1E8E163B1DFFBCFF26F80BFAEA60DE" blockId="16.[151,1437,152,2034]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26F80BFEF96703" bold="true" box="[197,312,808,834]" pageId="16" pageNumber="63">Protibiae</emphasis>
(
<figureCitation id="F29A92933B1DFFBCFEABF80AFE5F6702" box="[328,414,809,835]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="16" pageNumber="63">Fig. 31</figureCitation>
) tridentate, short basal tooth separated from intermediate tooth by a distance 1.2 times as long as that between intermediate and anterior teeth; intermediate and anterior teeth well developed, basal tooth with its anterior border more or less perpendicular to long axis of tibiae; apical spur almost straight, surpassing apex of first tarsomere.
<emphasis id="58D552043B1DFFBCFDD3F8BEFD5767F6" bold="true" box="[560,662,925,951]" pageId="16" pageNumber="63">Protarsi</emphasis>
(
<figureCitation id="F29A92933B1DFFBCFD45F8BEFD3D67F6" box="[678,764,925,951]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="16" pageNumber="63">Fig. 40</figureCitation>
) enlarged; tarsomere 5 shorter than 1–4 together, without ventro-internal carina, ventral side with very short and very fine longitudinal striae on inner side of apex; tarsomere 4 longitudinally striate on apical third, tarsomere 3 on apical fifth; largest claw (
<figureCitation id="F29A92933B1DFFBCFB0CF8C9FA886045" box="[1263,1353,1002,1028]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="16" pageNumber="63">Fig. 48</figureCitation>
) wide, flattened, moderately narrowly cleft at apex, external border of small branch continuous with internal curvature of claw.
<emphasis id="58D552043B1DFFBCFE93FF14FE3F6010" bold="true" box="[368,510,1079,1105]" pageId="16" pageNumber="63">Metafemur</emphasis>
(
<figureCitation id="F29A92933B1DFFBCFDEEFF1BFDA06013" box="[525,609,1080,1106]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="16" pageNumber="63">Fig. 55</figureCitation>
) 2.3 times as long as its maximum width.
<emphasis id="58D552043B1DFFBCFBB3FF14FB166010" bold="true" box="[1104,1239,1079,1105]" pageId="16" pageNumber="63">Metatibiae</emphasis>
(
<figureCitation id="F29A92933B1DFFBCFB05FF1BFAFA6013" box="[1254,1339,1080,1106]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="16" pageNumber="63">Fig. 55</figureCitation>
) robust, widened at apex, bristles on diagonal carina and apex very strong.
<emphasis id="58D552043B1DFFBCFC7EFF7DFBD66039" bold="true" box="[925,1047,1118,1144]" pageId="16" pageNumber="63">Metatarsi</emphasis>
subequal in length to tibiae, with first tarsomere scarcely widened towards apex, with apical bristles similar to those of apex of tibiae.
</paragraph>
<paragraph id="6A1E8E163B1DFFBCFF26FF88FB5160AC" blockId="16.[151,1437,152,2034]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26FF88FE1D6084" bold="true" box="[197,476,1195,1221]" pageId="16" pageNumber="63">Prosternal projection</emphasis>
high, apex alutaceous, anterior part transversely oval, not externally pointed; posterior part not wider than anterior one, posterior border fimbriate with long bristles.
</paragraph>
<paragraph id="6A1E8E163B1DFFBCFF26FFDBFC6C61C6" blockId="16.[151,1437,152,2034]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26FFDBFE116153" bold="true" box="[197,464,1272,1298]" pageId="16" pageNumber="63">Pronotal hypomera</emphasis>
alutaceous, with sparse erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large and very dense confluent, annulate punctures; disc of metasternum smooth, with small, disperse punctures; metacoxae with punctures similar to pleurae but smaller and more disperse. Ventrites with fine, annulate, moderately dense punctures.
</paragraph>
<paragraph id="6A1E8E163B1DFFBCFF26FEB0FE1E61EF" blockId="16.[151,1437,152,2034]" box="[197,479,1427,1454]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26FEB0FEFC61EC" bold="true" box="[197,317,1427,1453]" pageId="16" pageNumber="63">Aedeagus</emphasis>
:
<figureCitation id="F29A92933B1DFFBCFEAEFEB7FE1A61EF" box="[333,475,1428,1454]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="16" pageNumber="63">Figs. 76–77</figureCitation>
.
</paragraph>
<paragraph id="6A1E8E163B1DFFBCFF26FE99FCD66209" blockId="16.[151,1437,152,2034]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26FE99FE576195" bold="true" box="[197,406,1466,1492]" pageId="16" pageNumber="63">Allotype female</emphasis>
. differs from the male in the following characters: punctation generally stronger; protibiae as in
<figureCitation id="F29A92933B1DFFBCFEAEFEC2FE6961BA" box="[333,424,1505,1531]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="16" pageNumber="63">Fig. 32</figureCitation>
; protarsi slender; metafemora (
<figureCitation id="F29A92933B1DFFBCFCC5FEC2FC4161BA" box="[806,896,1505,1531]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="16" pageNumber="63">Fig. 56</figureCitation>
) somewhat more robust; epipleura (
<figureCitation id="F29A92933B1DFFBCFAD9FEC2FA5261BA" box="[1338,1427,1505,1531]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="16" pageNumber="63">Fig. 61</figureCitation>
) relatively wide in relation to other species in the group (see
<figureCitation id="F29A92933B1DFFBCFCBBFD2BFC266263" box="[856,999,1544,1570]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="16" pageNumber="63">Figs. 58–60</figureCitation>
), gradually narrowed to beginning of second ventrite; pygidium in lateral view as in
<figureCitation id="F29A92933B1DFFBCFD5EFD0DFCD26209" box="[701,787,1582,1608]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="16" pageNumber="63">Fig. 68</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B1DFFBCFF26FD76FBDD622E" box="[197,1052,1621,1647]" pageId="16" pageNumber="63" type="etymology">
<paragraph id="6A1E8E163B1DFFBCFF26FD76FBDD622E" blockId="16.[151,1437,152,2034]" box="[197,1052,1621,1647]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26FD76FE8B622E" bold="true" box="[197,330,1621,1647]" pageId="16" pageNumber="63">Etymology</emphasis>
. This species is named in honor of my friend Anibal Chacón.
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B1DFFBCFF26FD58FE2C62D7" box="[197,493,1659,1686]" pageId="16" pageNumber="63" type="distribution">
<paragraph id="6A1E8E163B1DFFBCFF26FD58FE2C62D7" blockId="16.[151,1437,152,2034]" box="[197,493,1659,1686]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26FD58FE9F62D4" bold="true" box="[197,350,1659,1685]" pageId="16" pageNumber="63">Distribution</emphasis>
.
<collectingCountry id="12B6CE863B1DFFBCFE8EFD5FFE2862D7" box="[365,489,1660,1686]" name="Venezuela" pageId="16" pageNumber="63">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B1DFFBDFF26FD81FE5D650D" lastPageId="17" lastPageNumber="64" pageId="16" pageNumber="63" type="diagnosis">
<paragraph id="6A1E8E163B1DFFBCFF26FD81FCBA638A" blockId="16.[151,1437,152,2034]" pageId="16" pageNumber="63">
<emphasis id="58D552043B1DFFBCFF26FD81FEFF62FD" bold="true" box="[197,318,1698,1724]" pageId="16" pageNumber="63">Diagnosis</emphasis>
.
<taxonomicName id="ADA1F5953B1DFFBCFEAEFD80FD8862FD" box="[333,585,1699,1724]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="63" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B1DFFBCFEAEFD80FD8862FD" box="[333,585,1699,1724]" italics="true" pageId="16" pageNumber="63">Cyclocephala anibali</emphasis>
</taxonomicName>
can be differentiated from all other species in the
<taxonomicName id="ADA1F5953B1DFFBCFB7DFD80FA5D62FD" box="[1182,1436,1699,1724]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="63" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B1DFFBCFB7DFD80FA5D62FD" box="[1182,1436,1699,1724]" italics="true" pageId="16" pageNumber="63">Cyclocephala bicolor</emphasis>
</taxonomicName>
species group by the following combination of character states: Surface of clypeus finely alutaceous with discrete punctures; frontoclypeal suture posteriorly projecting angularly; antennal club about 1.5 times as long as antennomeres 2–7 together; punctures on scutellum very fine, distinctly finer than those on middle of base of pronotum; punctures on basal half of pygidium small and dense, distinctly finer on apical half; prosternal projection with anterior part not externally pointed; basal tooth of protibia not forwardly directed, its anterior border more or less perpendicular to longitudinal axis of tibia; pygidium about 1.9 times as wide as long. The aedeagus is relatively wide with apical external angles rounded
</paragraph>
<paragraph id="6A1E8E163B1DFFBDFF25FCFBFE5D650D" blockId="16.[151,1437,152,2034]" lastBlockId="17.[151,1437,152,332]" lastPageId="17" lastPageNumber="64" pageId="16" pageNumber="63">
<taxonomicName id="ADA1F5953B1DFFBCFF25FCFBFE0C63B0" box="[198,461,2008,2033]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="63" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B1DFFBCFF25FCFBFE0C63B0" box="[198,461,2008,2033]" italics="true" pageId="16" pageNumber="63">Cyclocephala sarahae</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="ADA1F5953B1DFFBCFD42FCFBFCD663B0" box="[673,791,2008,2034]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="63" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B1DFFBCFD42FCFBFD7563B0" box="[673,692,2008,2033]" italics="true" pageId="16" pageNumber="63">C</emphasis>
.
<emphasis id="58D552043B1DFFBCFD27FCFBFCD663B0" box="[708,791,2008,2033]" italics="true" pageId="16" pageNumber="63">anibali</emphasis>
</taxonomicName>
, but it can be differentiated because most punctures on clypeus are confluent, the frontoclypeal suture is sinuate, not posteriorly projecting angularly, the antennal club about 2.0 times as long as antennomeres 2–7 together, punctures on scutellum similar to those on pronotal disc, punctures on basal half of pygidium large, smaller on apical half, the prosternal projection with anterior part externally pointed, the pygidium about 2.2 times as wide as long and the aedeagus is pointed on external apical angles.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="6A1E8E163B05FFA4FF74FC3EFDB56376" blockId="8.[151,894,1821,1882]" box="[151,628,1821,1847]" pageId="8" pageNumber="55">
<heading id="3156397A3B05FFA4FF74FC3EFDB56376" bold="true" box="[151,628,1821,1847]" fontSize="11" level="1" pageId="8" pageNumber="55" reason="1">
<taxonomicName id="ADA1F5953B05FFA4FF74FC3EFDB56376" authority="Endrodi, 1964" authorityName="Endrodi" authorityYear="1964" box="[151,628,1821,1847]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B05FFA4FF74FC3EFDB56376" bold="true" box="[151,628,1821,1847]" pageId="8" pageNumber="55">
<emphasis id="58D552043B05FFA4FF74FC3EFE7B6376" bold="true" box="[151,442,1821,1847]" italics="true" pageId="8" pageNumber="55">Cyclocephala bicolorata</emphasis>
<bibRefCitation id="0E30F3E73B05FFA4FE21FC3EFDB56376" author="Endrodi" box="[450,628,1821,1847]" pageId="8" pageNumber="64" refString="Endrodi, S. (1964) Eine Reihe von neuen Cyclocephala - Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica (n. s.), 17 (31), 433 - 470." type="journal article" year="1964">Endrödi, 1964</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="6A1E8E163B05FFA4FF74FC63FCBF631B" blockId="8.[151,894,1821,1882]" box="[151,894,1856,1882]" pageId="8" pageNumber="55">
(
<figureCitation id="F29A92933B05FFA4FF7CFC63FF32631B" box="[159,243,1856,1882]" captionStart="FIGURE 2" captionStartId="4.[151,255,1865,1889]" captionTargetBox="[373,1218,1132,1833]" captionTargetId="figure@4.[366,1220,1130,1841]" captionTargetPageId="4" captionText="FIGURE 2. Distribution of Cyclocephala dichroa (circles), and C. bicolorata (squares)." httpUri="https://zenodo.org/record/186487/files/figure.png" pageId="8" pageNumber="55">Figs. 2</figureCitation>
,
<figureCitation id="F29A92933B05FFA4FF1CFC63FEA9631B" box="[255,360,1856,1882]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="8" pageNumber="55">7–10, 20</figureCitation>
,
<figureCitation id="F29A92933B05FFA4FE97FC63FE00631B" box="[372,449,1856,1882]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="8" pageNumber="55">26–27</figureCitation>
,
<figureCitation id="F29A92933B05FFA4FE2FFC63FE2D631B" box="[460,492,1856,1882]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="8" pageNumber="55">36</figureCitation>
, 44–45,
<figureCitation id="F29A92933B05FFA4FDB3FC63FDAD631B" box="[592,620,1856,1882]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="8" pageNumber="55">51</figureCitation>
–52,
<figureCitation id="F29A92933B05FFA4FD44FC63FD06631B" box="[679,711,1856,1882]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="8" pageNumber="55">59</figureCitation>
, 63–64, 73–74)
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B05FFA0FF74FCAEFDD964BE" lastPageId="12" lastPageNumber="59" pageId="8" pageNumber="55" type="description">
<paragraph id="6A1E8E163B05FFA5FF74FCAEFC026499" blockId="8.[151,1437,1933,2037]" lastBlockId="9.[151,1437,152,294]" lastPageId="9" lastPageNumber="56" pageId="8" pageNumber="55">
<emphasis id="58D552043B05FFA4FF74FCAEFE8563E6" bold="true" box="[151,324,1933,1959]" pageId="8" pageNumber="55">Bibliography.</emphasis>
<taxonomicName id="ADA1F5953B05FFA4FEAEFCADFCA363E6" authority="Endrodi 1964: 441" authorityName="Endrodi" authorityPageNumber="441" authorityYear="1964" box="[333,866,1933,1959]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B05FFA4FEAEFCADFDB663E6" box="[333,631,1934,1959]" italics="true" pageId="8" pageNumber="55">Cyclocephala bicolorata</emphasis>
<bibRefCitation id="0E30F3E73B05FFA4FD9CFCAEFCE963E6" author="Endrodi" box="[639,808,1933,1959]" pageId="8" pageNumber="64" refString="Endrodi, S. (1964) Eine Reihe von neuen Cyclocephala - Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica (n. s.), 17 (31), 433 - 470." type="journal article" year="1964">Endrödi 1964</bibRefCitation>
: 441
</taxonomicName>
(description, male; compared with
<taxonomicName id="ADA1F5953B05FFA4FAF3FCADFA4A63E6" box="[1296,1419,1933,1959]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B05FFA4FAF3FCADFAE263E6" box="[1296,1315,1934,1959]" italics="true" pageId="8" pageNumber="55">C</emphasis>
.
<emphasis id="58D552043B05FFA4FAD7FCADFA4A63E6" box="[1332,1419,1934,1959]" italics="true" pageId="8" pageNumber="55">bicolor</emphasis>
</taxonomicName>
),
<figureCitation id="F29A92933B05FFA4FF74FC97FF2C638F" box="[151,237,1972,1998]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="8" pageNumber="55">Fig. 10</figureCitation>
(genitalia).
<typeStatus id="B51A30B43B05FFA4FE62FC97FE30638F" box="[385,497,1972,1998]" pageId="8" pageNumber="55" type="holotype">Holotype</typeStatus>
male Col. Endrödi.
<typeStatus id="B51A30B43B05FFA4FD0BFC97FCE5638F" box="[744,804,1972,1998]" pageId="8" pageNumber="55">Type</typeStatus>
locality:
<collectingCountry id="12B6CE863B05FFA4FC74FC97FBD6638F" box="[919,1047,1972,1998]" name="Venezuela" pageId="8" pageNumber="55">Venezuela</collectingCountry>
: Llanos.
<bibRefCitation id="0E30F3E73B05FFA4FB66FC97FAEF638F" author="Endrodi" box="[1157,1326,1972,1998]" pageId="8" pageNumber="64" refString="Endrodi, S. (1966) Monographie der Dynastinae (Coleoptera, Lamellicornia). I Teil. Entomologishe Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 33, 1 - 460." type="journal article" year="1966">Endrödi 1966</bibRefCitation>
: 91 (key, male), 143 (key, female), 157 (description, male), Fig. 388 (genitalia),
<collectingCountry id="12B6CE863B05FFA4FC19FCF8FBBB63B4" box="[1018,1146,2011,2037]" name="Venezuela" pageId="8" pageNumber="55">Venezuela</collectingCountry>
.
<bibRefCitation id="0E30F3E73B05FFA4FB65FCF8FA9B63B4" author="Dechambre" box="[1158,1370,2011,2037]" pageId="8" pageNumber="64" refString="Dechambre, R. - P. (1979) Missions entomologiques en Guyane et au Bresil (Coleoptera Dynastidae) Revue Francaise d'Entomologie. (N. S.), 1 (4), 160 - 168." type="journal article" year="1979">Dechambre 1979</bibRefCitation>
: 160,
<collectingCountry id="12B6CE863B04FFA5FF74FBBBFEDD64F3" box="[151,284,152,178]" name="Venezuela" pageId="9" pageNumber="56">Venezuela</collectingCountry>
,
<collectingCountry id="12B6CE863B04FFA5FECAFBBBFEBA64F3" box="[297,379,152,178]" name="Brazil" pageId="9" pageNumber="56">Brazil</collectingCountry>
.
<bibRefCitation id="0E30F3E73B04FFA5FE64FBBBFDF864F3" author="Endrodi" box="[391,569,152,178]" pageId="9" pageNumber="64" refString="Endrodi, S. (1985) The Dynastinae of the World. Dr. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi 1985</bibRefCitation>
: 110 (key, male), 146 (key, female), Fig. 650 (genitalia),
<collectingCountry id="12B6CE863B04FFA5FAF7FBBBFA5864F3" box="[1300,1433,152,178]" name="Venezuela" pageId="9" pageNumber="56">Venezuela</collectingCountry>
.
<bibRefCitation id="0E30F3E73B04FFA5FF74FB9DFEA56499" author="Dechambre" box="[151,356,190,216]" pageId="9" pageNumber="64" refString="Dechambre, R. - P. (1992) Nouveaux Cyclocephalini des genres Cyclocephala et Aspidolea (Col. Dynastidae). Les Coleopteres du Monde, 14, 57 - 75, color plate 12." type="journal article" year="1992">Dechambre 1992</bibRefCitation>
: 67 (compared with
<taxonomicName id="ADA1F5953B04FFA5FDB7FB9CFD116499" box="[596,720,190,216]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B04FFA5FDB7FB9CFDA66499" box="[596,615,191,216]" italics="true" pageId="9" pageNumber="56">C</emphasis>
.
<emphasis id="58D552043B04FFA5FD95FB9CFD116499" box="[630,720,191,216]" italics="true" pageId="9" pageNumber="56">dichroa</emphasis>
</taxonomicName>
),
<figureCitation id="F29A92933B04FFA5FD0AFB9DFCFA6499" box="[745,827,190,216]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="9" pageNumber="56">Fig. 33</figureCitation>
(genitalia).
</paragraph>
<paragraph id="6A1E8E163B04FFA5FF26FBC6FA856567" blockId="9.[151,1437,152,294]" pageId="9" pageNumber="56">
<emphasis id="58D552043B04FFA5FF26FBC6FEA264BE" bold="true" box="[197,355,229,255]" pageId="9" pageNumber="56">Description</emphasis>
. Short, oval, moderately shiny. Red to brownish red; frons and vertex black, elytra testaceous. Dorsal surface and pygidium glabrous, ventral pubescence sparse.
<emphasis id="58D552043B04FFA5FBC8FA28FB446564" bold="true" box="[1067,1157,267,293]" pageId="9" pageNumber="56">Length</emphasis>
: 11.0–11.8 mm.
</paragraph>
<caption id="3EDEDE9E3B04FFA5FF74FD7CFD39629E" httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="9" pageNumber="56" targetBox="[368,1218,387,1607]" targetPageId="9">
<paragraph id="6A1E8E163B04FFA5FF74FD7CFD39629E" blockId="9.[151,1437,1631,1759]" pageId="9" pageNumber="56">
<emphasis id="58D552043B04FFA5FF74FD7CFEB16236" bold="true" box="[151,368,1631,1655]" pageId="9" pageNumber="56">FIGURES 49–57.</emphasis>
Figs. 49–50.
<taxonomicName id="ADA1F5953B04FFA5FDF7FD43FCD56236" box="[532,788,1632,1655]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B04FFA5FDF7FD43FCD56236" box="[532,788,1632,1655]" italics="true" pageId="9" pageNumber="56">Cyclocephala bicolor</emphasis>
</taxonomicName>
: 49) metaleg, male; 50) metaleg, female. Figs. 51–52.
<taxonomicName id="ADA1F5953B04FFA5FF74FDA0FE5E62DB" box="[151,415,1667,1690]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B04FFA5FF74FDA0FE5E62DB" box="[151,415,1667,1690]" italics="true" pageId="9" pageNumber="56">Cyclocephala bicolorata</emphasis>
</taxonomicName>
: 51) metaleg, male; 52) metaleg female. Figs. 53–54.
<taxonomicName id="ADA1F5953B04FFA5FC3EFDA0FB0962DB" box="[989,1224,1667,1690]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B04FFA5FC3EFDA0FB0962DB" box="[989,1224,1667,1690]" italics="true" pageId="9" pageNumber="56">Cyclocephala dichroa</emphasis>
</taxonomicName>
: 53) metaleg, male; 54) metaleg, female. Figs. 55–56.
<taxonomicName id="ADA1F5953B04FFA5FDEFFD86FD3262FD" box="[524,755,1701,1724]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B04FFA5FDEFFD86FD3262FD" box="[524,755,1701,1724]" italics="true" pageId="9" pageNumber="56">Cyclocephala anibali</emphasis>
</taxonomicName>
: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57.
<taxonomicName id="ADA1F5953B04FFA5FF13FDEBFE1E629E" box="[240,479,1736,1759]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B04FFA5FF13FDEBFE1E629E" box="[240,479,1736,1759]" italics="true" pageId="9" pageNumber="56">Cyclocephala sarahae</emphasis>
</taxonomicName>
metaleg, male (holotype).
</paragraph>
</caption>
<paragraph id="6A1E8E163B04FFA6FF26FC2EFAD26499" blockId="9.[151,1436,1805,2025]" lastBlockId="10.[151,1436,152,216]" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56">
<emphasis id="58D552043B04FFA5FF26FC2EFEDC6366" bold="true" box="[197,285,1805,1831]" pageId="9" pageNumber="56">MALE</emphasis>
:
<emphasis id="58D552043B04FFA5FECFFC2EFE506366" bold="true" box="[300,401,1805,1831]" pageId="9" pageNumber="56">Clypeus</emphasis>
(
<figureCitation id="F29A92933B04FFA5FE41FC2EFE286366" box="[418,489,1805,1831]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="9" pageNumber="56">Fig. 7</figureCitation>
) about 2.4 times as wide across base as long; sides straight, strongly convergent to truncated and feebly reflexed apex; shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view flat to slightly concave on apical half; surface finely alutaceous, with punctures moderately strong, discrete, somewhat transverse (especially on base), dense, the distance between most punctures smaller than the diameter of the puncture, more dense towards apex. Frontoclypeal suture (
<figureCitation id="F29A92933B04FFA5FF7CFCECFF2463A8" box="[159,229,1999,2025]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="9" pageNumber="56">Fig. 7</figureCitation>
) thin, slightly arcuately projecting posteriorly.
<emphasis id="58D552043B04FFA5FCEFFCEDFC9463A9" bold="true" box="[780,853,1998,2024]" pageId="9" pageNumber="56">Frons</emphasis>
width about 2.5 times transverse diameter of eye, finely alutaceous; punctures similar to those of clypeus, but somewhat more scattered, except on a narrow, smooth, transverse band behind frontoclypeal suture and another longitudinal band along midline.
</paragraph>
<caption id="3EDEDE9E3B07FFA6FF74FDC8FF1063B4" httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="10" pageNumber="57" targetBox="[300,1286,266,1745]" targetPageId="10">
<paragraph id="6A1E8E163B07FFA6FF74FDC8FF1063B4" blockId="10.[151,1437,1771,2037]" pageId="10" pageNumber="57">
<emphasis id="58D552043B07FFA6FF74FDC8FE926342" bold="true" box="[151,339,1771,1795]" pageId="10" pageNumber="57">FIGURE 58–77.</emphasis>
Figs. 58–61. Female epipleurae (ventral view): 58)
<emphasis id="58D552043B07FFA6FC71FDC8FB436343" box="[914,1154,1771,1794]" italics="true" pageId="10" pageNumber="57">
<taxonomicName id="ADA1F5953B07FFA6FC71FDC8FBBF6343" box="[914,1150,1771,1794]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="bicolor">Cyclocephala bicolor</taxonomicName>
,
</emphasis>
59)
<taxonomicName id="ADA1F5953B07FFA6FB55FDC8FA886343" box="[1206,1353,1771,1794]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B07FFA6FB55FDC8FB096343" box="[1206,1224,1771,1794]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FB34FDC8FA886343" box="[1239,1353,1771,1794]" italics="true" pageId="10" pageNumber="57">bicolorata</emphasis>
</taxonomicName>
, 60)
<taxonomicName id="ADA1F5953B07FFA6FA60FDC8FF2D6364" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B07FFA6FA60FDC8FA546343" box="[1411,1429,1771,1794]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FF74FC2DFF2D6364" box="[151,236,1806,1829]" italics="true" pageId="10" pageNumber="57">dichroa</emphasis>
</taxonomicName>
, 61)
<taxonomicName id="ADA1F5953B07FFA6FEC4FC2DFE566364" box="[295,407,1806,1829]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B07FFA6FEC4FC2DFEF86364" box="[295,313,1806,1829]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FEABFC2DFE566364" box="[328,407,1806,1829]" italics="true" pageId="10" pageNumber="57">anibali</emphasis>
</taxonomicName>
. Figs. 62–68. Pygidium (lateral view): 62)
<taxonomicName id="ADA1F5953B07FFA6FC9EFC2DFC2C6364" box="[893,1005,1806,1829]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B07FFA6FC9EFC2DFC4E6364" box="[893,911,1806,1829]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FC7DFC2DFC2C6364" box="[926,1005,1806,1829]" italics="true" pageId="10" pageNumber="57">bicolor</emphasis>
</taxonomicName>
, male; 63)
<taxonomicName id="ADA1F5953B07FFA6FB8DFC2DFAC26364" box="[1134,1283,1806,1829]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B07FFA6FB8DFC2DFB416364" box="[1134,1152,1806,1829]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FB6CFC2DFAC26364" box="[1167,1283,1806,1829]" italics="true" pageId="10" pageNumber="57">bicolorata</emphasis>
</taxonomicName>
, male; 64)
<taxonomicName id="ADA1F5953B07FFA6FA60FC2DFEC76309" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B07FFA6FA60FC2DFA546364" box="[1411,1429,1806,1829]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FF74FC12FEC76309" box="[151,262,1841,1864]" italics="true" pageId="10" pageNumber="57">bicolorata</emphasis>
</taxonomicName>
, female; 65)
<taxonomicName id="ADA1F5953B07FFA6FE77FC12FDC76309" box="[404,518,1840,1864]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B07FFA6FE77FC12FE676309" box="[404,422,1841,1864]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FE57FC12FDC76309" box="[436,518,1841,1864]" italics="true" pageId="10" pageNumber="57">dichroa</emphasis>
</taxonomicName>
, male; 66)
<taxonomicName id="ADA1F5953B07FFA6FD63FC12FD306309" box="[640,753,1840,1864]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B07FFA6FD63FC12FD536309" box="[640,658,1841,1864]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FD7CFC12FD306309" box="[671,753,1841,1864]" italics="true" pageId="10" pageNumber="57">dichroa</emphasis>
</taxonomicName>
, female; 67)
<taxonomicName id="ADA1F5953B07FFA6FC63FC12FC2A6309" box="[896,1003,1840,1864]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B07FFA6FC63FC12FC536309" box="[896,914,1841,1864]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FC7CFC12FC2A6309" box="[927,1003,1841,1864]" italics="true" pageId="10" pageNumber="57">anibali</emphasis>
</taxonomicName>
, male (holotype); 68)
<taxonomicName id="ADA1F5953B07FFA6FB38FC12FA876309" box="[1243,1350,1840,1864]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B07FFA6FB38FC12FB2C6309" box="[1243,1261,1841,1864]" italics="true" pageId="10" pageNumber="57">C</emphasis>
.
<emphasis id="58D552043B07FFA6FB19FC12FA876309" box="[1274,1350,1841,1864]" italics="true" pageId="10" pageNumber="57">anibali</emphasis>
</taxonomicName>
, female (allotype). Figs. 69–70.
<taxonomicName id="ADA1F5953B07FFA6FE40FC70FD50632B" box="[419,657,1875,1898]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B07FFA6FE40FC70FD50632B" box="[419,657,1875,1898]" italics="true" pageId="10" pageNumber="57">Cyclocephala bicolor</emphasis>
</taxonomicName>
: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71–72.
<taxonomicName id="ADA1F5953B07FFA6FF74FC55FE4963CC" box="[151,392,1910,1933]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B07FFA6FF74FC55FE4963CC" box="[151,392,1910,1933]" italics="true" pageId="10" pageNumber="57">Cyclocephala dichroa</emphasis>
</taxonomicName>
: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73–74.
<taxonomicName id="ADA1F5953B07FFA6FB6DFC55FA5C63CC" box="[1166,1437,1910,1933]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B07FFA6FB6DFC55FA5C63CC" box="[1166,1437,1910,1933]" italics="true" pageId="10" pageNumber="57">Cyclocephala bicolorata</emphasis>
</taxonomicName>
(holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75.
<taxonomicName id="ADA1F5953B07FFA6FBF1FCBAFAD563F1" box="[1042,1300,1945,1968]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B07FFA6FBF1FCBAFAD563F1" box="[1042,1300,1945,1968]" italics="true" pageId="10" pageNumber="57">Cyclocephala sarahae</emphasis>
</taxonomicName>
(holotype), parameres, caudal view. Figs. 76–77.
<taxonomicName id="ADA1F5953B07FFA6FDC4FC98FCCD6393" box="[551,780,1979,2002]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B07FFA6FDC4FC98FCCD6393" box="[551,780,1979,2002]" italics="true" pageId="10" pageNumber="57">Cyclocephala anibali</emphasis>
</taxonomicName>
(holotype): 76) parameres, caudal view; 77) aedeagus, lateral view.
</paragraph>
</caption>
<paragraph id="6A1E8E163B06FFA7FF26FBB4FEF665A6" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26FBB4FE9064F0" bold="true" box="[197,337,151,177]" pageId="11" pageNumber="58">Mandibles</emphasis>
with external border sinuate, apex projecting externally.
<emphasis id="58D552043B06FFA7FBC6FBB4FB5864F0" bold="true" box="[1061,1177,151,177]" pageId="11" pageNumber="58">Maxillae</emphasis>
(
<figureCitation id="F29A92933B06FFA7FB4FFBBBFAC964F3" box="[1196,1288,152,178]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="11" pageNumber="58">Fig. 10</figureCitation>
) with galea moderately strong, with two apical, one median and three basal teeth, the basal teeth very small and arising from a common base; stipes projected anteriorly on inner side and with a brush of fine, yellowish brown setae on apex of projection; maxillary palpi (
<figureCitation id="F29A92933B06FFA7FD85FA2FFD7D6567" box="[614,700,268,294]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="11" pageNumber="58">Fig. 20</figureCitation>
) moderately robust, second palpomere with a small projection at base, longer than third, fourth palpomere fusiform, 3.0 times as long as preceding.
<emphasis id="58D552043B06FFA7FB7BFA11FB3C650D" bold="true" box="[1176,1277,306,332]" pageId="11" pageNumber="58">Labium</emphasis>
(
<figureCitation id="F29A92933B06FFA7FAEDFA11FA97650D" box="[1294,1366,306,332]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="11" pageNumber="58">Fig. 9</figureCitation>
) with sides slightly convex on basal two-thirds, not constricted at the middle of apical half; apex truncate on ventral side, with two small projections at middle on oral side, and fimbriated with long, fine, yellowish brown setae; surface without punctures on disc, with one longitudinal series of long bristles on each side, converging towards base.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26FAD0FAEB664F" blockId="11.[151,1437,151,2034]" box="[197,1322,499,526]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26FAD0FEFF664C" bold="true" box="[197,318,499,525]" pageId="11" pageNumber="58">Antennae</emphasis>
with 10 antennomeres, club about 2.0 times the length of antennomeres 2–7 together.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26F939FCC1672B" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26F939FE826675" bold="true" box="[197,323,538,564]" pageId="11" pageNumber="58">Pronotum</emphasis>
1.4 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected at middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface finely alutaceous (
<date id="1E1FA8D63B06FFA7FEC4F9ADFEA066E9" box="[295,353,654,680]" pageId="11" pageNumber="58" value="1940-10">40X</date>
); punctures annulate, moderately fine and dense, the distance between most punctures on disc as long as or longer than diameter of punctures, punctures gradually larger toward sides, diameter of a puncture about 2.0 times the diameter of those on center of disc, more dense on basal angles, where the distance between most punctures is as long as or shorter than the diameter of a puncture; surface also covered with numerous minute punctures (
<date id="1E1FA8D63B06FFA7FDD8F80AFDB06702" box="[571,625,809,835]" pageId="11" pageNumber="58" value="1950-10">50X</date>
or more).
<emphasis id="58D552043B06FFA7FD17F80BFCB76703" bold="true" box="[756,886,808,834]" pageId="11" pageNumber="58">Scutellum</emphasis>
triangular, sides feebly arcuate, apex briefly rounded; punctures similar to those on pronotal disc.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26F855FE7F6013" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26F855FED767D1" bold="true" box="[197,278,886,912]" pageId="11" pageNumber="58">Elytra</emphasis>
1.2 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones weak; with feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident; also with numerous minute punctures (
<date id="1E1FA8D63B06FFA7FF7CFF32FF14606A" box="[159,213,1041,1067]" pageId="11" pageNumber="58" value="1950-10">50X</date>
or more) scattered over all surface; sutural interstria flat, weakly elevated on apical half. Epipleura narrow, more so apically.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26FF7DFBA36152" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26FF7DFEFC6039" bold="true" box="[197,317,1118,1144]" pageId="11" pageNumber="58">Pygidium</emphasis>
2.0 times as wide as long; in posterior view the inferior borders straight, converging to narrow and feebly convex-truncate apex in an angle distinctly greater than 90°; in lateral view (
<figureCitation id="F29A92933B06FFA7FB36FFA6FAEE60DE" box="[1237,1327,1157,1183]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="11" pageNumber="58">Fig. 63</figureCitation>
) slightly convex, more strongly convex on apical half; surface finely alutaceous, with abundant minute punctures and with large annulate punctures, more dense on lateral angles; punctures on apex distinctly smaller and less dense, the distances between most punctures more than 2.0 times its own diameter.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26FE3CFB0562FC" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26FE3CFEFB6178" bold="true" box="[197,314,1311,1337]" pageId="11" pageNumber="58">Protibiae</emphasis>
(
<figureCitation id="F29A92933B06FFA7FEA8FE03FE63617B" box="[331,418,1312,1338]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="11" pageNumber="58">Fig. 26</figureCitation>
) tridentate, small basal tooth separated from the intermediate tooth by a distance 1.4 times as long as that between the intermediate and anterior teeth; intermediate and anterior teeth well developed, basal tooth with its anterior border more or less perpendicular to longitudinal axis of tibiae; apical spur almost straight, surpassing apex of first tarsomere.
<emphasis id="58D552043B06FFA7FCCFFEB0FC5361EC" bold="true" box="[812,914,1427,1453]" pageId="11" pageNumber="58">Protarsi</emphasis>
(
<figureCitation id="F29A92933B06FFA7FC40FEB7FC3861EF" box="[931,1017,1428,1454]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="11" pageNumber="58">Fig. 36</figureCitation>
) enlarged; tarsomere 5 shorter than 1–4 together, without ventro-internal carina, ventral side with very short and very fine longitudinal striae on inner side of apex; tarsomere 4 longitudinally striated on less than apical half; tarsomere 3 striated only on inner side of apical half; largest claw (
<figureCitation id="F29A92933B06FFA7FD8AFD2BFD3A6263" box="[617,763,1544,1570]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="11" pageNumber="58">Figs. 44–45</figureCitation>
) wide, flattened, moderately narrowly cleft at apex, the external border of small branch continuous with internal curvature of the claw.
<emphasis id="58D552043B06FFA7FBA6FD0DFB126209" bold="true" box="[1093,1235,1582,1608]" pageId="11" pageNumber="58">Metafemur</emphasis>
(
<figureCitation id="F29A92933B06FFA7FB07FD0DFAFB6209" box="[1252,1338,1582,1608]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="11" pageNumber="58">Fig. 51</figureCitation>
) robust, 2.0 times as long as its maximum width, its anterior border strongly convex.
<emphasis id="58D552043B06FFA7FBA1FD76FB0C622E" bold="true" box="[1090,1229,1621,1647]" pageId="11" pageNumber="58">Metatibiae</emphasis>
(
<figureCitation id="F29A92933B06FFA7FB3DFD76FAF6622E" box="[1246,1335,1621,1647]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="11" pageNumber="58">Fig. 51</figureCitation>
) robust, widened at apex, bristles on diagonal carina and apex very strong.
<emphasis id="58D552043B06FFA7FC7EFD58FBD662D4" bold="true" box="[925,1047,1659,1685]" pageId="11" pageNumber="58">Metatarsi</emphasis>
subequal in length to tibiae, with first tarsomere widened towards apex, with apical bristles similar to those of apex of tibiae.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26FDEAFB86634B" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26FDEAFE0E62A2" bold="true" box="[197,463,1737,1763]" pageId="11" pageNumber="58">Prosternal projection</emphasis>
high, apex alutaceous, anterior part transversely oval, externally pointed; posterior part shorter but wider than anterior, posterior border fimbriate with long bristles.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26FC35FA8163E4" blockId="11.[151,1437,151,2034]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26FC35FE116371" bold="true" box="[197,464,1814,1840]" pageId="11" pageNumber="58">Pronotal hypomera</emphasis>
alutaceous, with sparse erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large, dense confluent, annulate punctures; disc of metasternum smooth, with small, scattered punctures; metacoxae with punctures similar to those of pleurae but smaller and more disperse. Ventrites with annulate, moderately dense punctures, smaller and sparser on apical ventrites.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26FC92FE1E638A" blockId="11.[151,1437,151,2034]" box="[197,479,1969,1995]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26FC92FEFC638A" bold="true" box="[197,317,1969,1995]" pageId="11" pageNumber="58">Aedeagus</emphasis>
:
<figureCitation id="F29A92933B06FFA7FEAEFC92FE1A638A" box="[333,475,1969,1995]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="11" pageNumber="58">Figs. 73–74</figureCitation>
.
</paragraph>
<paragraph id="6A1E8E163B06FFA7FF26FCF4FA5D63B3" blockId="11.[151,1437,151,2034]" box="[197,1436,2007,2034]" pageId="11" pageNumber="58">
<emphasis id="58D552043B06FFA7FF26FCF4FE8363B0" bold="true" box="[197,322,2007,2033]" pageId="11" pageNumber="58">FEMALE</emphasis>
: differs from the male in the following characters: punctation generally stronger; protibiae
<figureCitation id="F29A92933B06FFA7FA8EFCFBFA5D63B3" box="[1389,1436,2008,2034]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="11" pageNumber="58">Fig.</figureCitation>
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF74FBBBFDD964BE" blockId="12.[151,1437,152,448]" pageId="12" pageNumber="59">
27; protarsi slender; metafemora (
<figureCitation id="F29A92933B01FFA0FDC7FBBBFDB864F3" box="[548,633,152,178]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="12" pageNumber="59">Fig. 52</figureCitation>
) somewhat stronger; metatibiae
<figureCitation id="F29A92933B01FFA0FC14FBBBFB8F64F3" box="[1015,1102,152,178]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="12" pageNumber="59">Fig. 52</figureCitation>
; epipleura (
<figureCitation id="F29A92933B01FFA0FB34FBBBFAED64F3" box="[1239,1324,152,178]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="12" pageNumber="59">Fig. 59</figureCitation>
) strongly narrowed at level of anterior border of metacoxa, then widened up to level of middle of second ventrite; pygidium in lateral view
<figureCitation id="F29A92933B01FFA0FE5EFBC6FDD564BE" box="[445,532,229,255]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="12" pageNumber="59">Fig. 64</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B01FFA0FF26FA28FB556532" pageId="12" pageNumber="59" type="distribution">
<paragraph id="6A1E8E163B01FFA0FF26FA28FD856567" blockId="12.[151,1437,152,448]" box="[197,580,267,294]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26FA28FE9F6564" bold="true" box="[197,350,267,293]" pageId="12" pageNumber="59">Distribution</emphasis>
:
<collectingCountry id="12B6CE863B01FFA0FE8EFA2FFE286567" box="[365,489,268,294]" name="Venezuela" pageId="12" pageNumber="59">Venezuela</collectingCountry>
,
<collectingCountry id="12B6CE863B01FFA0FE17FA2FFD816567" box="[500,576,268,294]" name="Brazil" pageId="12" pageNumber="59">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="6A1E8E163B01FFA0FF26FA11FB556532" blockId="12.[151,1437,152,448]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26FA11FE70650D" bold="true" box="[197,433,306,332]" pageId="12" pageNumber="59">Specimens studied</emphasis>
:
<collectingCountry id="12B6CE863B01FFA0FE22FA11FDBB650D" box="[449,634,306,332]" name="Venezuela" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FE22FA11FDBB650D" bold="true" box="[449,634,306,332]" pageId="12" pageNumber="59">VENEZUELA</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B01FFA0FD68FA11FD1D650D" bold="true" box="[651,732,306,332]" pageId="12" pageNumber="59">Apure</emphasis>
: Hato El Frío, Fundo El Ceibote,
<quantity id="AD5923F33B01FFA0FB9AFA11FB09650D" box="[1145,1224,306,332]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="59" unit="m" value="100.0">100 m</quantity>
,
<date id="1E1FA8D63B01FFA0FB36FA11FA99650D" box="[1237,1368,306,332]" pageId="12" pageNumber="59" value="1975-05-29">29-V-1975</date>
, C. J. Rosales, 1 F (MIZA).
<emphasis id="58D552043B01FFA0FE78FA7BFE396533" bold="true" box="[411,504,344,370]" pageId="12" pageNumber="59">Bolívar</emphasis>
: Guri, Río Caroní,
<date id="1E1FA8D63B01FFA0FD3FFA7AFCA46532" box="[732,869,345,371]" pageId="12" pageNumber="59" value="1968-04-11">11-IV-1968</date>
, J. Salcedo, 1 M (MIZA).
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B01FFA0FF26FA5CFF226581" pageId="12" pageNumber="59" type="description">
<paragraph id="6A1E8E163B01FFA0FF26FA5CFF226581" blockId="12.[151,1437,152,448]" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FF26FA5CFE4A65D8" bold="true" box="[197,395,383,409]" pageId="12" pageNumber="59">Other localities</emphasis>
:
<collectingCountry id="12B6CE863B01FFA0FE7FFA5CFD9765D8" box="[412,598,383,409]" name="Venezuela" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FE7FFA5CFD9765D8" bold="true" box="[412,598,383,409]" pageId="12" pageNumber="59">VENEZUELA</emphasis>
</collectingCountry>
: Llanos.
<collectingCountry id="12B6CE863B01FFA0FD2AFA5CFCFD65D8" box="[713,828,383,409]" name="Brazil" pageId="12" pageNumber="59">
<emphasis id="58D552043B01FFA0FD2AFA5CFCFD65D8" bold="true" box="[713,828,383,409]" pageId="12" pageNumber="59">BRAZIL</emphasis>
</collectingCountry>
: São Felix do Xingu (Moyen Xingu) (
<bibRefCitation id="0E30F3E73B01FFA0FAF3FAA3FF176581" author="Dechambre" pageId="12" pageNumber="64" refString="Dechambre, R. - P. (1979) Missions entomologiques en Guyane et au Bresil (Coleoptera Dynastidae) Revue Francaise d'Entomologie. (N. S.), 1 (4), 160 - 168." type="journal article" year="1979">Dechambre 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

240
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<mods:title id="122583E5ABE822744E1423D04B1FE2EC">Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
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<paragraph id="6A1E8E163B0CFFADFF74FD81FAFD62FD" blockId="1.[151,1340,1698,1724]" box="[151,1340,1698,1724]" pageId="1" pageNumber="48">
<heading id="3156397A3B0CFFADFF74FD81FAFD62FD" bold="true" box="[151,1340,1698,1724]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<emphasis id="58D552043B0CFFADFF74FD81FAFD62FD" bold="true" box="[151,1340,1698,1724]" pageId="1" pageNumber="48">
Key for identification of the species of the
<taxonomicName id="ADA1F5953B0CFFADFD47FD81FC6562FD" box="[676,932,1698,1724]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B0CFFADFD47FD81FC6562FD" bold="true" box="[676,932,1698,1724]" italics="true" pageId="1" pageNumber="48">Cyclocephala bicolor</emphasis>
</taxonomicName>
group of the genus
<taxonomicName id="ADA1F5953B0CFFADFB7AFD81FAFD62FD" box="[1177,1340,1698,1724]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="58D552043B0CFFADFB7AFD81FAFD62FD" bold="true" box="[1177,1340,1698,1724]" italics="true" pageId="1" pageNumber="48">Cyclocephala</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<keyStep id="D15596B33B0CFFADFF74FDCFFA5A6362" pageId="1" pageNumber="48">
<paragraph id="6A1E8E163B0CFFADFF74FDCFFA5A6342" blockId="1.[151,1437,1772,2019]" box="[151,1435,1772,1795]" pageId="1" pageNumber="48">
<keyLead id="D1502D233B0CFFADFF74FDCFFA5A6342" box="[151,1435,1772,1795]" pageId="1" pageNumber="48">1. Anterior tarsi enlarged (males) ..................................................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0CFFADFF74FC2FFA5A6362" blockId="1.[151,1437,1772,2019]" box="[151,1435,1804,1827]" pageId="1" pageNumber="48">
<keyLead id="D1502D233B0CFFADFF74FC2FFA5A6362" box="[151,1435,1804,1827]" pageId="1" pageNumber="48">- Anterior tarsi slender (females) .................................................................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<paragraph id="6A1E8E163B0CFFADFF74FC0FFA5D63C2" blockId="1.[151,1437,1772,2019]" pageId="1" pageNumber="48">
<keyStep id="D15596B33B0CFFADFF74FC0FFA5D63C2" pageId="1" pageNumber="48">
<keyLead id="D1502D233B0CFFADFF74FC0FFA5D63C2" pageId="1" pageNumber="48">
2. Second ventrite with a triangular, flat projection on posterior half; basal tooth of protibiae (
<figureCitation id="F29A92933B0CFFADFB30FC0FFAE76302" box="[1235,1318,1836,1859]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="1" pageNumber="48">Fig. 29</figureCitation>
) distinctly directed forward, separated from middle tooth by a distance about 2.0 times longer than that separating middle from anterior tooth; clypeus on apical half slightly convex in lateral view (
<figureCitation id="F29A92933B0CFFADFC44FC4FFC3563C2" box="[935,1012,1900,1923]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="1" pageNumber="48">Fig. 12</figureCitation>
) .......................
<taxonomicName id="ADA1F5953B0CFFADFB40FC4FFA5D63C2" authority="Dechambre." authorityName="Dechambre." box="[1187,1436,1900,1923]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B0CFFADFB40FC4FFB7463C2" box="[1187,1205,1900,1923]" italics="true" pageId="1" pageNumber="48">C</emphasis>
.
<emphasis id="58D552043B0CFFADFB21FC4FFAD563C2" box="[1218,1300,1900,1923]" italics="true" pageId="1" pageNumber="48">dichroa</emphasis>
Dechambre.
</taxonomicName>
</keyLead>
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</paragraph>
<paragraph id="6A1E8E163B0CFFADFF74FCAFFA5C63A2" blockId="1.[151,1437,1772,2019]" pageId="1" pageNumber="48">
<keyLead id="D1502D233B0CFFADFF74FCAFFA5C63A2" pageId="1" pageNumber="48">
- Second ventrite without triangular projection; basal tooth of protibiae (
<figureCitation id="F29A92933B0CFFADFC54FCAFFB4463E2" box="[951,1157,1932,1955]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="1" pageNumber="48">Figs. 24, 26, 28, 31</figureCitation>
) with anterior border usually perpendicular to longitudinal axis of tibiae and separated from the middle tooth by a distance 1.5 times or less than that separating middle from anterior tooth; clypeus on apical half more or less flat to slightly concave in lateral
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF24FBBBFA5A654E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
view (
<figureCitation id="F29A92933B0FFFAEFEEAFBBBFE5764EE" box="[265,406,152,175]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Figs. 4, 8, 16</figureCitation>
) .................................................................................................................................................... 3
<keyStep id="D15596B33B0FFFAEFF74FB9BFA5A654E" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FB9BFA5A654E" pageId="2" pageNumber="49">
3. Frontoclypeal suture (
<figureCitation id="F29A92933B0FFFAEFE5BFB9BFDC6648E" box="[440,519,184,207]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 15</figureCitation>
) posteriorly projecting angularly; punctation on scutellum very fine, much more so than on middle of base of pronotum; punctation on apical region of pygidium very fine, distinctly finer than that of basal region, which is moderately fine ...............................................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB0BFBDBFA92654E" box="[1256,1363,248,271]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="anibali" status="sp. nov.">
<emphasis id="58D552043B0FFFAEFB0BFBDBFB3B654E" box="[1256,1274,248,271]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFAE4FBDBFA92654E" box="[1287,1363,248,271]" italics="true" pageId="2" pageNumber="49">anibali</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="43E6EF7F3B0FFFAEFA83FBD4FA5A654E" box="[1376,1435,247,271]" pageId="2" pageNumber="49" rank="species">
<emphasis id="58D552043B0FFFAEFA83FBD4FA5A654E" bold="true" box="[1376,1435,247,271]" pageId="2" pageNumber="49">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FA3BFA5A652E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FA3BFA5A652E" pageId="2" pageNumber="49">
- Frontoclypeal suture (
<figureCitation id="F29A92933B0FFFAEFE50FA3BFDFF656E" box="[435,574,280,303]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Figs. 3, 7, 11</figureCitation>
) posteriorly projecting arcuately or almost straight; punctation of scutellum fine, scarcely finer than that on center of pronotal base; punctation on apical region of pygidium fine, scarcely smaller than that of basal region, which is strong to moderately strong .................................................................................. 4
</keyLead>
</paragraph>
<keyStep id="D15596B33B0FFFAEFF74FA5BFA5D666E" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74FA5BFA5A658E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FA5BFA5A658E" pageId="2" pageNumber="49">
4. Punctures on base of clypeus not discrete, closely set, mostly confluent, appearing as irregular and fine, more or less transversely striate; frontoclypeal suture posteriorly projecting arcuately (
<figureCitation id="F29A92933B0FFFAEFC31FABBFBD265EE" box="[978,1043,408,431]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 3</figureCitation>
); surface of pygidium smooth (
<date id="1E1FA8D63B0FFFAEFABDFABBFA4E65EE" box="[1374,1423,408,431]" pageId="2" pageNumber="49" value="1950-10">50X</date>
) ...................................................................................................................................................................................... 5
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FAFBFA5D666E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FAFBFA5D666E" pageId="2" pageNumber="49">
- Punctures on base of clypeus moderately strong, discrete, somewhat transverse, the distance between most punctures smaller than the diameter of puncture; frontoclypeal suture very slightly arcuately projecting posteriorly (
<figureCitation id="F29A92933B0FFFAEFA92FADBFF17666E" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 7</figureCitation>
); surface of pygidium finely alutaceous (
<date id="1E1FA8D63B0FFFAEFD6FF93BFD7C666E" box="[652,701,536,559]" pageId="2" pageNumber="49" value="1950-10">50X</date>
) ........................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB52F93BFA5D666E" authority="Endrodi" authorityName="Endrodi" box="[1201,1436,536,559]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B0FFFAEFB52F93BFB02666E" box="[1201,1219,536,559]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFB33F93BFAFE666E" box="[1232,1343,536,559]" italics="true" pageId="2" pageNumber="49">bicolorata</emphasis>
Endrödi
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="D15596B33B0FFFAEFF74F91BFA5D676E" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74F91BFA5D66EE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F91BFA5D66EE" pageId="2" pageNumber="49">
5. Labium (
<figureCitation id="F29A92933B0FFFAEFECAF91BFEA8660E" box="[297,361,568,591]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 5</figureCitation>
) with strong, dense piliferous punctures on basal two-thirds; metatibiae (
<figureCitation id="F29A92933B0FFFAEFB91F91BFB33660E" box="[1138,1266,568,591]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="2" pageNumber="49">Figs. 49–50</figureCitation>
) with piliferous punctures almost restricted to those of normal longitudinal series close to posterior border; distances between most punctures on apex of pygidium usually smaller than 2.0 times the diameter of punctures; apex of pygidium (in posterior view) feebly convex-truncate.
<emphasis id="58D552043B0FFFAEFDD5F9BBFB2966EE" box="[566,1256,664,687]" italics="true" pageId="2" pageNumber="49">
....................................................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB35F9BBFB2966EE" authority="Laporte" authorityName="Laporte" box="[1238,1256,664,687]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolor">C</taxonomicName>
</emphasis>
.
<emphasis id="58D552043B0FFFAEFB16F9BBFA8366EE" box="[1269,1346,664,687]" italics="true" pageId="2" pageNumber="49">bicolor</emphasis>
Laporte
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74F99BFA5D676E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F99BFA5D676E" pageId="2" pageNumber="49">
- Labium with one longitudinal series of long bristles on each side, converging towards base; mesotibia and especially metatibiae with numerous piliferous punctures over all surface, in addition to those of normal longitudinal series; distances between most punctures on apex of pygidium usually 2.0 times or more larger than the diameter of punctures; apex of pygidium (in posterior view) rounded.....................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB27F83BFA5D676E" authority="Ratcliffe" authorityName="Ratcliffe" box="[1220,1436,792,815]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B0FFFAEFB27F83BFB17676E" box="[1220,1238,792,815]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFB00F83BFAF8676E" box="[1251,1337,792,815]" italics="true" pageId="2" pageNumber="49">sarahae</emphasis>
Ratcliffe
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="D15596B33B0FFFAEFF74F81BFA5A67AE" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74F81BFA5D67CE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F81BFA5D67CE" pageId="2" pageNumber="49">
6. Basal tooth of protibiae distinctly anteriorly projected, separated from the middle tooth by a distance about 2.0 times larger than that separating the middle from the anterior (
<figureCitation id="F29A92933B0FFFAEFCFDF87BFCAC672E" box="[798,877,856,879]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="2" pageNumber="49">Fig. 30</figureCitation>
); clypeus convex in lateral view (
<figureCitation id="F29A92933B0FFFAEFB37F87BFAE0672E" box="[1236,1313,856,879]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 12</figureCitation>
) ................
<emphasis id="58D552043B0FFFAEFF2DF85BFB7A67CE" box="[206,1211,888,911]" italics="true" pageId="2" pageNumber="49">
....................................................................................................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB4AF85BFB7A67CE" authority="Dechambre" authorityName="Dechambre" box="[1193,1211,888,911]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="dichroa">C</taxonomicName>
</emphasis>
.
<emphasis id="58D552043B0FFFAEFB2BF85BFADB67CE" box="[1224,1306,888,911]" italics="true" pageId="2" pageNumber="49">dichroa</emphasis>
Dechambre
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74F8BBFA5A67AE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F8BBFA5A67AE" pageId="2" pageNumber="49">
- Basal tooth of protibiae more diagonal and/or its anterior border more or less perpendicular to longitudinal axis of tibiae, separated from the middle tooth by a distance 1.5 or less than that separating the middle from the anterior (
<figureCitation id="F29A92933B0FFFAEFF2CF8FBFE6067AE" box="[207,417,984,1007]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="2" pageNumber="49">Figs. 25, 27, 29, 32</figureCitation>
); clypeus more or less flat in lateral view (
<figureCitation id="F29A92933B0FFFAEFC84F8FBFC3567AE" box="[871,1012,984,1007]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Figs. 4, 8, 16</figureCitation>
) .......................................................... 7
</keyLead>
</paragraph>
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<keyStep id="D15596B33B0FFFAEFF74F8DBFA5A600E" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74F8DBFA5A604E" blockId="2.[151,1437,152,1263]" box="[151,1435,1015,1039]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74F8DBFA5A604E" box="[151,1435,1015,1039]" pageId="2" pageNumber="49">
7. Punctures on apical region of pygidium distinctly smaller than those of basal region. ......................
<taxonomicName id="ADA1F5953B0FFFAEFB0DF8DBFA98604E" box="[1262,1369,1016,1039]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="anibali" status="sp. nov.">
<emphasis id="58D552043B0FFFAEFB0DF8DBFAC1604E" box="[1262,1280,1016,1039]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFAEDF8DBFA98604E" box="[1294,1369,1016,1039]" italics="true" pageId="2" pageNumber="49">anibali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E6EF7F3B0FFFAEFA83F8D4FA5A604E" box="[1376,1435,1015,1039]" pageId="2" pageNumber="49" rank="species">
<emphasis id="58D552043B0FFFAEFA83F8D4FA5A604E" bold="true" box="[1376,1435,1015,1039]" pageId="2" pageNumber="49">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FF3BFA5A600E" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FF3BFA5A600E" pageId="2" pageNumber="49">- Punctures on apical region of pygidium scarcely smaller than those of basal region, which are strong to moderately strong ........................................................................................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="D15596B33B0FFFAEFF74FF7BFA5D60AE" pageId="2" pageNumber="49">
<paragraph id="6A1E8E163B0FFFAEFF74FF7BFA5D60CE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FF7BFA5D60CE" pageId="2" pageNumber="49">
8. Pygidium feebly convex in lateral view, surface smooth, punctures more or less strong almost to apex, on apical region somewhat smaller, usually dense, sometimes scattered .......................................................
<taxonomicName id="ADA1F5953B0FFFAEFB35FF5BFA5D60CE" authority="Laporte" authorityName="Laporte" box="[1238,1436,1144,1167]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B0FFFAEFB35FF5BFB2960CE" box="[1238,1256,1144,1167]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFB16FF5BFA8360CE" box="[1269,1346,1144,1167]" italics="true" pageId="2" pageNumber="49">bicolor</emphasis>
Laporte
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FFBBFA5D60AE" blockId="2.[151,1437,152,1263]" pageId="2" pageNumber="49">
<keyLead id="D1502D233B0FFFAEFF74FFBBFA5D60AE" pageId="2" pageNumber="49">
- Pygidium strongly convex in lateral view, surface finely alutaceous (
<date id="1E1FA8D63B0FFFAEFC54FFBBFC2860EE" box="[951,1001,1176,1199]" pageId="2" pageNumber="49" value="1950-10">50X</date>
), punctures more or less strong at base, smaller on apex, most distances between punctures 2.0 or more times larger than diameter of puncture. .................... ....................................................................................................................................................
<taxonomicName id="ADA1F5953B0FFFAEFB52FFFBFA5D60AE" authority="Endrodi" authorityName="Endrodi" box="[1201,1436,1240,1263]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B0FFFAEFB52FFFBFB0260AE" box="[1201,1219,1240,1263]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFB33FFFBFAFE60AE" box="[1232,1343,1240,1263]" italics="true" pageId="2" pageNumber="49">bicolorata</emphasis>
Endrödi
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

766
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<mods:title id="ACE1E80B423EB90DEEA12A09E8D87271">Review of the species in the Cyclocephala bicolor Laporte species group (Coleoptera: Scarabaeidae: Dynastinae)</mods:title>
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<paragraph id="6A1E8E163B0FFFAEFF74FE66FD8A611E" blockId="2.[151,836,1349,1410]" box="[151,587,1349,1375]" pageId="2" pageNumber="49">
<heading id="3156397A3B0FFFAEFF74FE66FD8A611E" bold="true" box="[151,587,1349,1375]" fontSize="11" level="1" pageId="2" pageNumber="49" reason="1">
<taxonomicName id="ADA1F5953B0FFFAEFF74FE66FD8A611E" authority="Laporte, 1840" authorityName="Laporte" authorityYear="1840" box="[151,587,1349,1375]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B0FFFAEFF74FE66FD8A611E" bold="true" box="[151,587,1349,1375]" pageId="2" pageNumber="49">
<emphasis id="58D552043B0FFFAEFF74FE66FE54611E" bold="true" box="[151,405,1349,1375]" italics="true" pageId="2" pageNumber="49">Cyclocephala bicolor</emphasis>
<bibRefCitation id="0E30F3E73B0FFFAEFE7EFE66FD8A611E" author="Laporte" box="[413,587,1349,1375]" pageId="2" pageNumber="64" refString="Laporte, F. L. de (1840) Histoire Naturelle des Insectes Coleopteres, volume 2. Dumeril, Paris, 563 pp., 36 plates." type="book" year="1840">Laporte, 1840</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF74FE4BFC8561C3" blockId="2.[151,836,1349,1410]" box="[151,836,1384,1410]" pageId="2" pageNumber="49">
(
<figureCitation id="F29A92933B0FFFAEFF7CFE4BFF3061C3" box="[159,241,1384,1410]" captionStart="FIGURE 1" captionStartId="4.[151,255,1052,1076]" captionTargetBox="[378,1217,320,1022]" captionTargetId="figure@4.[367,1219,313,1027]" captionTargetPageId="4" captionText="FIGURE 1. Distribution of Cyclocephala bicolor (circles), C. anibali, sp. n. (triangle), and C. sarahae (square)." httpUri="https://zenodo.org/record/186486/files/figure.png" pageId="2" pageNumber="49">Figs. 1</figureCitation>
,
<figureCitation id="F29A92933B0FFFAEFF1CFE4BFE9861C3" box="[255,345,1384,1410]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">3–6, 19</figureCitation>
,
<figureCitation id="F29A92933B0FFFAEFE86FE4BFDA061C3" box="[357,609,1384,1410]" captionStart-0="FIGURE 24 – 33" captionStart-1="FIGURE 34 – 48" captionStartId-0="7.[151,261,1850,1874]" captionStartId-1="8.[151,260,1509,1533]" captionTargetBox-0="[285,1302,542,1826]" captionTargetBox-1="[303,1283,197,1485]" captionTargetId-0="figure@7.[285,1302,542,1826]" captionTargetId-1="figure@8.[303,1283,197,1485]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." captionText-1="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri-0="https://zenodo.org/record/186489/files/figure.png" httpUri-1="https://zenodo.org/record/186490/files/figure.png" pageId="2" pageNumber="49">24–25, 33–35, 41–42</figureCitation>
,
<figureCitation id="F29A92933B0FFFAEFD8EFE4BFD7861C3" box="[621,697,1384,1410]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="2" pageNumber="49">49–50</figureCitation>
,
<figureCitation id="F29A92933B0FFFAEFD26FE4BFCFD61C3" box="[709,828,1384,1410]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="2" pageNumber="49">58, 69–70</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B0FFFA9FF74FE96FF396499" lastPageId="5" lastPageNumber="52" pageId="2" pageNumber="49" type="description">
<paragraph id="6A1E8E163B0FFFAEFF74FE96FC32636A" blockId="2.[151,1437,1461,2028]" pageId="2" pageNumber="49">
<emphasis id="58D552043B0FFFAEFF74FE96FE87618E" bold="true" box="[151,326,1461,1487]" pageId="2" pageNumber="49">Bibliography.</emphasis>
<taxonomicName id="ADA1F5953B0FFFAEFEACFE96FCFF618E" authority="Laporte 1840: 124" authorityName="Laporte" authorityPageNumber="124" authorityYear="1840" box="[335,830,1461,1487]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B0FFFAEFEACFE96FD95618F" box="[335,596,1461,1486]" italics="true" pageId="2" pageNumber="49">Cyclocephala bicolor</emphasis>
<bibRefCitation id="0E30F3E73B0FFFAEFDBEFE96FCC4618E" author="Laporte" box="[605,773,1461,1487]" pageId="2" pageNumber="64" refString="Laporte, F. L. de (1840) Histoire Naturelle des Insectes Coleopteres, volume 2. Dumeril, Paris, 563 pp., 36 plates." type="book" year="1840">Laporte 1840</bibRefCitation>
: 124
</taxonomicName>
(description).
<typeStatus id="B51A30B43B0FFFAEFC11FE96FBEF618E" box="[1010,1070,1461,1487]" pageId="2" pageNumber="49">Type</typeStatus>
(sex?) MNHN.
<typeStatus id="B51A30B43B0FFFAEFB17FE96FAF1618E" box="[1268,1328,1461,1487]" pageId="2" pageNumber="49">Type</typeStatus>
locality: Cayenne.
<bibRefCitation id="0E30F3E73B0FFFAEFEEDFEFFFE1961B7" author="Burmeister" box="[270,472,1500,1526]" pageId="2" pageNumber="64" refString="Burmeister, H. (1847) Handbuch der Entomologie, volume 5, 1 - 584. Berlin." type="journal article" year="1847">Burmeister 1847</bibRefCitation>
: 60 (description), Cayenne.
<bibRefCitation id="0E30F3E73B0FFFAEFCCBFEFFFBC061B7" author="Prudhomme" box="[808,1025,1500,1526]" pageId="2" pageNumber="64" refString="Prudhomme, M. (1906) Catalogue des coleopteres de la Guyane Francaise recuillis par M. Prudhomme de 1870 a 1906. Impr. Governement, Cayenne, 46 pp." type="book" year="1906">Prudhomme 1906</bibRefCitation>
: 13,
<collectingCountry id="12B6CE863B0FFFAEFBD9FEFFFB2C61B7" box="[1082,1261,1500,1526]" name="French Guiana" pageId="2" pageNumber="49">French Guiana</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFB1AFEFFFA5661B7" author="Bodkin" box="[1273,1431,1500,1526]" pageId="2" pageNumber="64" refString="Bodkin, G. E. (1919) Notes on the Coleoptera of British Guiana. Entomologist Monthly Magazine, 55, 210 - 219." type="journal article" year="1919">Bodkin 1919</bibRefCitation>
: 215,
<collectingCountry id="12B6CE863B0FFFAEFF32FD21FE40625D" box="[209,385,1538,1564]" name="Guyana" pageId="2" pageNumber="49">British Guiana</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFE68FD21FDDE625D" author="Hohne" box="[395,543,1538,1564]" pageId="2" pageNumber="64" refString="Hohne, W. (1923) Neue Cyclocephalen (Col. Dyn.). Deutsche Entomologische Zeitschrift, 1923 (4), 345 - 375." type="journal article" year="1923">Höhne 1923</bibRefCitation>
: 359 (compared with
<taxonomicName id="ADA1F5953B0FFFAEFCF8FD20FC48625D" box="[795,905,1538,1564]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="vestita">
<emphasis id="58D552043B0FFFAEFCF8FD20FCEF625D" box="[795,814,1539,1564]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFCDEFD20FC48625D" box="[829,905,1539,1564]" italics="true" pageId="2" pageNumber="49">vestita</emphasis>
</taxonomicName>
).
<bibRefCitation id="0E30F3E73B0FFFAEFC42FD21FBF5625D" author="Arrow" box="[929,1076,1538,1564]" pageId="2" pageNumber="64" refString="Arrow, G. J. (1937) Dynastinae. Coleopterorum Catalogus, 21, part 156, 3 - 124." type="journal article" year="1937">Arrow 1937</bibRefCitation>
: 8,
<collectingCountry id="12B6CE863B0FFFAEFBB8FD21FB75625D" box="[1115,1204,1538,1564]" name="Guyana" pageId="2" pageNumber="49">Guiana</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFB5DFD21FA59625D" author="Blackwelder" box="[1214,1432,1538,1564]" pageId="2" pageNumber="64" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America, Part. 2. Bulletin of the United States National Museum, 185, 189 - 341." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
: 251,
<collectingCountry id="12B6CE863B0FFFAEFF30FD0AFEEC6202" box="[211,301,1577,1603]" name="Guyana" pageId="2" pageNumber="49">Guiana</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFEDAFD0AFE206202" author="Endrodi" box="[313,481,1577,1603]" pageId="2" pageNumber="64" refString="Endrodi, S. (1964) Eine Reihe von neuen Cyclocephala - Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica (n. s.), 17 (31), 433 - 470." type="journal article" year="1964">Endrödi 1964</bibRefCitation>
: 442 (compared with
<taxonomicName id="ADA1F5953B0FFFAEFD04FD0AFC476203" box="[743,902,1577,1603]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B0FFFAEFD04FD0AFD3B6203" box="[743,762,1577,1602]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFCE9FD0AFC476203" box="[778,902,1577,1602]" italics="true" pageId="2" pageNumber="49">bicolorata</emphasis>
</taxonomicName>
),
<figureCitation id="F29A92933B0FFFAEFC43FD0AFC346202" box="[928,1013,1577,1603]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 11</figureCitation>
(genitalia).
<bibRefCitation id="0E30F3E73B0FFFAEFB64FD0AFAEE6202" author="Endrodi" box="[1159,1327,1577,1603]" pageId="2" pageNumber="64" refString="Endrodi, S. (1966) Monographie der Dynastinae (Coleoptera, Lamellicornia). I Teil. Entomologishe Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 33, 1 - 460." type="journal article" year="1966">Endrödi 1966</bibRefCitation>
: 91 (key, male), 140, 143 (key, female), 156 (description male, female), Figs. 386–397 (genitalia),
<collectingCountry id="12B6CE863B0FFFAEFB22FD73FAFF622B" box="[1217,1342,1616,1642]" name="Venezuela" pageId="2" pageNumber="49">Venezuela</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFAA8FD73FF3762D1" name="Guyana" pageId="2" pageNumber="49">British Guiana</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFEE0FD55FE0262D1" box="[259,451,1654,1680]" name="French Guiana" pageId="2" pageNumber="49">French Guiana</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFE33FD55FD8D62D1" box="[464,588,1654,1680]" name="Suriname" pageId="2" pageNumber="49">Suriname</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFDB9FD55FD6D62D1" box="[602,684,1654,1680]" name="Brazil" pageId="2" pageNumber="49">Brazil</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFD59FD55FCA162D1" author="Gruner" box="[698,864,1654,1680]" pageId="2" pageNumber="64" refString="Gruner, L. (1971) Scarabaeidae Melolonthinae, Dynastinae, Rutelinae, Cetoniinae [Coleoptera] recoltes en Guyane Francaise par la Mission du Museum National d'Histoire Naturelle. Annales de la Societe Entomologique de France (N. S.), 7 (4), 843 - 848." type="journal article" year="1971">Gruner 1971</bibRefCitation>
: 845,
<collectingCountry id="12B6CE863B0FFFAEFC4DFD55FBAC62D1" box="[942,1133,1654,1680]" name="French Guiana" pageId="2" pageNumber="49">French Guiana</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFB98FD55FA9962D1" author="Dechambre" box="[1147,1368,1654,1680]" pageId="2" pageNumber="64" refString="Dechambre, R. - P. (1979) Missions entomologiques en Guyane et au Bresil (Coleoptera Dynastidae) Revue Francaise d'Entomologie. (N. S.), 1 (4), 160 - 168." type="journal article" year="1979">Dechambre 1979</bibRefCitation>
: 160,
<collectingCountry id="12B6CE863B0FFFAEFF74FDBEFEDB62F6" box="[151,282,1693,1719]" name="Venezuela" pageId="2" pageNumber="49">Venezuela</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFEC5FDBEFE2162F6" box="[294,480,1693,1719]" name="Guyana" pageId="2" pageNumber="49">British Guiana</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFE0DFDBEFD6962F6" box="[494,680,1693,1719]" name="French Guiana" pageId="2" pageNumber="49">French Guiana</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFD55FDBEFCF162F6" box="[694,816,1693,1719]" name="Suriname" pageId="2" pageNumber="49">Suriname</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFCDEFDBEFC4C62F6" box="[829,909,1693,1719]" name="Brazil" pageId="2" pageNumber="49">Brazil</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFC7AFDBEFB8662F6" author="Endrodi" box="[921,1095,1693,1719]" pageId="2" pageNumber="64" refString="Endrodi, S. (1985) The Dynastinae of the World. Dr. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi 1985</bibRefCitation>
: 109 (key, male), 146 (key, female), Figs. 648–649 (genitalia),
<collectingCountry id="12B6CE863B0FFFAEFDD5FDE7FD26629F" box="[566,743,1732,1758]" name="French Guiana" pageId="2" pageNumber="49">French Guiana</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFD10FDE7FCA7629F" box="[755,870,1732,1758]" name="Suriname" pageId="2" pageNumber="49">Suriname</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFC91FDE7FC11629F" box="[882,976,1732,1758]" name="Guyana" pageId="2" pageNumber="49">Guyana</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFC3EFDE7FBE9629F" box="[989,1064,1732,1758]" name="Brazil" pageId="2" pageNumber="49">Brazil</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFBD0FDE7FB21629F" author="Ratcliffe" box="[1075,1248,1732,1758]" pageId="2" pageNumber="64" refString="Ratcliffe, B. C. (1992) New species and country records of Brazilian Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae). Tidschrift voor Entomologie, 135 (2), 179 - 190." type="journal article" year="1992">Ratcliffe 1992</bibRefCitation>
: 187 (compared with
<taxonomicName id="ADA1F5953B0FFFAEFF31FDC8FE906345" box="[210,337,1770,1796]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B0FFFAEFF31FDC8FF246345" box="[210,229,1771,1796]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFF10FDC8FE906345" box="[243,337,1771,1796]" italics="true" pageId="2" pageNumber="49">sarahae</emphasis>
</taxonomicName>
).
<bibRefCitation id="0E30F3E73B0FFFAEFE89FDC9FDEB6345" author="Lachaume" box="[362,554,1770,1796]" pageId="2" pageNumber="64" refString="Lachaume, G. (1992) Dynastidae americains. Coleopteres du Monde, 14, 1 - 56, plates A - C + 1 - 11." type="journal article" year="1992">Lachaume 1992</bibRefCitation>
: 16, plate 1,
<figureCitation id="F29A92933B0FFFAEFD5FFDC9FCD06345" box="[700,785,1770,1796]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="2" pageNumber="49">Fig. 28</figureCitation>
,
<collectingCountry id="12B6CE863B0FFFAEFCFFFDC9FCBB6345" box="[796,890,1770,1796]" name="Guyana" pageId="2" pageNumber="49">Guyana</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFC65FDC9FBF46345" box="[902,1077,1770,1796]" name="French Guiana" pageId="2" pageNumber="49">French Guiana</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFBA2FDC9FB756345" box="[1089,1204,1770,1796]" name="Suriname" pageId="2" pageNumber="49">Suriname</collectingCountry>
,
<collectingCountry id="12B6CE863B0FFFAEFB5CFDC9FACB6345" box="[1215,1290,1770,1796]" name="Brazil" pageId="2" pageNumber="49">Brazil</collectingCountry>
.
<bibRefCitation id="0E30F3E73B0FFFAEFAF6FDC9FF17636A" author="Dechambre" pageId="2" pageNumber="64" refString="Dechambre, R. - P. (1992) Nouveaux Cyclocephalini des genres Cyclocephala et Aspidolea (Col. Dynastidae). Les Coleopteres du Monde, 14, 57 - 75, color plate 12." type="journal article" year="1992">Dechambre 1992</bibRefCitation>
: 67 (compared with
<taxonomicName id="ADA1F5953B0FFFAEFE26FC32FD83636B" box="[453,578,1809,1835]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B0FFFAEFE26FC32FE19636B" box="[453,472,1809,1834]" italics="true" pageId="2" pageNumber="49">C</emphasis>
.
<emphasis id="58D552043B0FFFAEFE0BFC32FD83636B" box="[488,578,1809,1834]" italics="true" pageId="2" pageNumber="49">dichroa</emphasis>
</taxonomicName>
),
<figureCitation id="F29A92933B0FFFAEFDB9FC32FD6C636A" box="[602,685,1809,1835]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="2" pageNumber="49">Fig. 32</figureCitation>
(genitalia), plate 1,
<figureCitation id="F29A92933B0FFFAEFC7AFC32FC31636A" box="[921,1008,1809,1835]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="2" pageNumber="49">Fig. 28</figureCitation>
.
</paragraph>
<paragraph id="6A1E8E163B0FFFAEFF26FC14FB716339" blockId="2.[151,1437,1461,2028]" pageId="2" pageNumber="49">
<emphasis id="58D552043B0FFFAEFF26FC14FEA56310" bold="true" box="[197,356,1847,1873]" pageId="2" pageNumber="49">Description.</emphasis>
Short, oval, moderately shiny. Brownish red, frons and vertex black, elytra testaceous. Dorsal surface and pygidium glabrous, ventral pubescence sparse.
<emphasis id="58D552043B0FFFAEFC47FC7DFC3F6339" bold="true" box="[932,1022,1886,1912]" pageId="2" pageNumber="49">Length</emphasis>
: 9.0–13.0 mm.
</paragraph>
<paragraph id="6A1E8E163B0FFFAFFF26FCA6FE96650D" blockId="2.[151,1437,1461,2028]" lastBlockId="3.[151,1437,152,2034]" lastPageId="3" lastPageNumber="50" pageId="2" pageNumber="49">
<emphasis id="58D552043B0FFFAEFF26FCA6FEDC63DE" bold="true" box="[197,285,1925,1951]" pageId="2" pageNumber="49">MALE</emphasis>
:
<emphasis id="58D552043B0FFFAEFECFFCA6FE5063DE" bold="true" box="[300,401,1925,1951]" pageId="2" pageNumber="49">Clypeus</emphasis>
(
<figureCitation id="F29A92933B0FFFAEFE41FCA6FE2863DE" box="[418,489,1925,1951]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="2" pageNumber="49">Fig. 3</figureCitation>
) about 2.3 times as wide across base as long; sides straight, strongly convergent to truncated and feebly reflexed apex; shallowly sulcate longitudinally along middle and shallowly depressed laterally before apex, in lateral view flat to slightly concave on apical half; surface finely alutaceous, with fine, shallow and widely annulate punctures, annulation wide, not well defined posteriorly, usually confluent, appearing as irregular and fine, more or less transverse striae; punctures simple toward apex and anterior half of sides. Frontoclypeal suture thin, arcuately projecting posteriorly.
<emphasis id="58D552043B0EFFAFFC26FBC6FBCE64BE" bold="true" box="[965,1039,229,255]" pageId="3" pageNumber="50">Frons</emphasis>
width about 2.5 times transverse diameter of eye, finely alutaceous; punctures similar to those of clypeus, but less dense, somewhat thinner and better delimited.
</paragraph>
<paragraph id="6A1E8E163B0EFFAFFF26FA7BFC3666C3" blockId="3.[151,1437,152,2034]" pageId="3" pageNumber="50">
<emphasis id="58D552043B0EFFAFFF26FA7BFE936533" bold="true" box="[197,338,344,370]" pageId="3" pageNumber="50">Mandibles</emphasis>
with external border sinuate, apex projecting externally.
<emphasis id="58D552043B0EFFAFFBD2FA7BFB676533" bold="true" box="[1073,1190,344,370]" pageId="3" pageNumber="50">Maxillae</emphasis>
(
<figureCitation id="F29A92933B0EFFAFFB5BFA7AFAC76532" box="[1208,1286,345,371]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="3" pageNumber="50">Fig. 6</figureCitation>
) with galea moderately robust, with two apical, one median, and three basal teeth; basal teeth very small and arising from common base; stipes projected anteriorly on inner side and with a brush of fine, black setae on apex of projection; maxillary palpi (
<figureCitation id="F29A92933B0EFFAFFE02FAEEFDF765A6" box="[481,566,461,487]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="3" pageNumber="50">Fig. 19</figureCitation>
) moderately robust, second palpomere longer than third, weakly widened towards apex, fourth palpomere fusiform, 2.5 times longer than preceding.
<emphasis id="58D552043B0EFFAFFBE4FAD0FBAB664C" bold="true" box="[1031,1130,499,525]" pageId="3" pageNumber="50">Labium</emphasis>
(
<figureCitation id="F29A92933B0EFFAFFB99FAD7FB01664F" box="[1146,1216,500,526]" captionStart="FIGURES 3 – 23" captionStartId="5.[151,271,1668,1692]" captionTargetBox="[276,1310,351,1643]" captionTargetId="figure@5.[276,1312,351,1643]" captionTargetPageId="5" captionText="FIGURES 3 – 23. Figs. 3 – 6. Cyclocephala bicolor: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7 – 10. Cyclocephala bicolorata: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11 – 14. Cyclocephala dichroa: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15 – 18. Cyclocephala anibali (holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19 – 23. Maxillary palpi: 19) C. bicolor, 20) C. bicolorata, 21) C. sarahae (holotype), 22) C. dichroa, 23) C. anibali (holotype)." httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="3" pageNumber="50">Fig. 5</figureCitation>
) with sides almost parallel on basal two-thirds, constricted at middle of apical third; apex truncate on middle and fimbriate with long, fine yellowish-golden setae; surface almost flat, with some very small punctures on disc and one longitudinal series of long bristles on each side, converging towards base.
</paragraph>
<paragraph id="6A1E8E163B0EFFAFFF26F9ADFA9E66E9" blockId="3.[151,1437,152,2034]" box="[197,1375,654,680]" pageId="3" pageNumber="50">
<emphasis id="58D552043B0EFFAFFF26F9ADFEFF66E9" bold="true" box="[197,318,654,680]" pageId="3" pageNumber="50">Antennae</emphasis>
with 10 antennomeres, club more than 2.0 times the length of antennomeres 2–7 together.
</paragraph>
<paragraph id="6A1E8E163B0EFFAFFF26F996FC906045" blockId="3.[151,1437,152,2034]" pageId="3" pageNumber="50">
<emphasis id="58D552043B0EFFAFFF26F996FE82668E" bold="true" box="[197,323,693,719]" pageId="3" pageNumber="50">Pronotum</emphasis>
1.5 times as wide as long, widest near base, sides more strongly convergent to apex on anterior half; anterior border widely and arcuately projected at middle; basal angles widely rounded; basal border without marginal bead, almost uniformly arcuate, slightly more projected in central third; surface smooth (
<date id="1E1FA8D63B0EFFAFFF7CF80AFF166702" box="[159,215,809,835]" pageId="3" pageNumber="50" value="1940-10">40X</date>
); punctures annulate, moderately fine and dense, the distance between most punctures on disc, as long as or longer than diameter of punctures, punctures gradually larger and denser towards sides, the diameter of a puncture at basal angle at least 2.0 times the diameter of those on center of disc, more dense on basal angles, where the distance between most punctures is as large or smaller than the diameter of a puncture; all surface also covered with numerous minute punctures (
<date id="1E1FA8D63B0EFFAFFD5CF8E7FD32679F" box="[703,755,964,990]" pageId="3" pageNumber="50" value="1950-10">50X</date>
or more).
<emphasis id="58D552043B0EFFAFFC8EF8E0FC2A679C" bold="true" box="[877,1003,963,989]" pageId="3" pageNumber="50">Scutellum</emphasis>
triangular, sides feebly arcuate, apex briefly rounded; punctures similar to those of pronotal disc.
</paragraph>
<paragraph id="6A1E8E163B0EFFAFFF26FF33FE7F60AC" blockId="3.[151,1437,152,2034]" pageId="3" pageNumber="50">
<emphasis id="58D552043B0EFFAFFF26FF33FED7606B" bold="true" box="[197,278,1040,1066]" pageId="3" pageNumber="50">Elytra</emphasis>
1.1 times as long as wide, arcuately widened from first quarter to middle, then rounded to apex, sutural angle well defined, but rounded; humeral and apical umbones weak, with a feeble, short, elongate tumescence behind humeral umbones on external border; surface smooth, with large, shallow, annulate punctures, much smaller on sides and apex, double rows not evident, additional numerous minute punctures (
<date id="1E1FA8D63B0EFFAFFF7CFF8FFF146087" box="[159,213,1196,1222]" pageId="3" pageNumber="50" value="1950-10">50X</date>
or more) scattered over the surface; sutural interstria flat, slightly elevated on apical half. Epipleura narrow, more so apically.
</paragraph>
<paragraph id="6A1E8E163B0EFFAFFF26FFDBFE0061EF" blockId="3.[151,1437,152,2034]" pageId="3" pageNumber="50">
<emphasis id="58D552043B0EFFAFFF26FFDBFEFC6153" bold="true" box="[197,317,1272,1298]" pageId="3" pageNumber="50">Pygidium</emphasis>
2.0 times as wide as long; in posterior view the inferior borders slightly convex, converging to feebly convexly truncated apex at an angle distinctly greater than 90°; in lateral view (
<figureCitation id="F29A92933B0EFFAFFB5AFE03FAD3617B" box="[1209,1298,1312,1338]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="3" pageNumber="50">Fig. 62</figureCitation>
) uniformly convex; surface smooth, shining, with large annulate punctures, punctures more dense on lateral angles; on apex more shining, the punctures smaller and less dense, the distances between most punctures less than 2.0 times their own diameter.
</paragraph>
<paragraph id="6A1E8E163B0EFFAFFF26FE99FAC16316" blockId="3.[151,1437,152,2034]" pageId="3" pageNumber="50">
<emphasis id="58D552043B0EFFAFFF26FE99FEFB6195" bold="true" box="[197,314,1466,1492]" pageId="3" pageNumber="50">Protibiae</emphasis>
(fig. 24) tridentate, small basal tooth separated from the intermediate tooth by a distance 1.5 times as long as that between the intermediate and the anterior teeth; intermediate and anterior teeth well developed, basal tooth with its anterior border more or less perpendicular to long axis of tibiae; apical spur almost straight, surpassing the apex of the first tarsomere.
<emphasis id="58D552043B0EFFAFFCA3FD0DFC676209" bold="true" box="[832,934,1582,1608]" pageId="3" pageNumber="50">Protarsi</emphasis>
(
<figureCitation id="F29A92933B0EFFAFFC55FD0DFBF96209" box="[950,1080,1582,1608]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="3" pageNumber="50">Fig. 34–35</figureCitation>
) enlarged; tarsomere 5 shorter than 1–4 together, without ventrointernal carina, ventral side with short and fine longitudinal striae on inner side of apex and sometimes also on inner side of base; tarsomere 4 longitudinally striated on about apical half, tarsomere 3 on apical third; largest claw (
<figureCitation id="F29A92933B0EFFAFFD6BFD80FCCD62FC" box="[648,780,1699,1725]" captionStart="FIGURE 34 – 48" captionStartId="8.[151,260,1509,1533]" captionTargetBox="[303,1283,197,1485]" captionTargetId="figure@8.[303,1283,197,1485]" captionTargetPageId="8" captionText="FIGURE 34 – 48. Figs. 34 – 35. Cyclocephala bicolor male: 34) protarsus, lateral; 35) protarsus, ventral. Fig. 36. Cyclocephala bicolorata male protarsus, lateral. Figs. 37 – 38. Cyclocephala sarahae male (holotype): 37) protarsus, lateral; 38) protarsus, ventral. Fig. 39. Cyclocephala dichroa male protarsus, lateral. Fig. 40. Cyclocephala anibali male (holotype) protarsus, lateral. Figs. 41 – 42. Cyclocephala bicolor, male: 41) largest claw, apical; 42) largest claw, lateral. Fig. 43. Cyclocephala sarahae male (holotype) largest claw, apical. Figs. 44 – 45. Cyclocephala bicolorata, male: 44) largest claw, apical; 45) largest claw, lateral. Figs. 46 – 47. Cyclocephala dichroa, male: 46) largest claw, apical; 47) largest claw, lateral. Fig. 48. Cyclocephala anibali male (holotype) largest claw, apical." httpUri="https://zenodo.org/record/186490/files/figure.png" pageId="3" pageNumber="50">Fig. 41–42</figureCitation>
) wide, flattened, moderately widely cleft at apex, both branches feebly divergent (external border of small branch not continuous with internal curvature of the claw).
<emphasis id="58D552043B0EFFAFFF08FDCCFEBC6348" bold="true" box="[235,381,1775,1801]" pageId="3" pageNumber="50">Metafemur</emphasis>
(
<figureCitation id="F29A92933B0EFFAFFE6DFDD3FE29634B" box="[398,488,1776,1802]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="3" pageNumber="50">Fig. 49</figureCitation>
) about 2.3 times as long as its maximum width.
<emphasis id="58D552043B0EFFAFFBDCFDCCFB0A6348" bold="true" box="[1087,1227,1775,1801]" pageId="3" pageNumber="50">Metatibiae</emphasis>
(
<figureCitation id="F29A92933B0EFFAFFB3EFDD3FAF6634B" box="[1245,1335,1776,1802]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="3" pageNumber="50">Fig. 49</figureCitation>
) robust, widened at apex, bristles on diagonal carina and apex strong.
<emphasis id="58D552043B0EFFAFFC8BFC35FC236371" bold="true" box="[872,994,1814,1840]" pageId="3" pageNumber="50">Metatarsi</emphasis>
(
<figureCitation id="F29A92933B0EFFAFFC11FC35FB866371" box="[1010,1095,1814,1840]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="3" pageNumber="50">Fig. 33</figureCitation>
) subequal in length to tibiae, with first tarsomere widened towards apex, with apical bristles similar to those of apex of tibiae.
</paragraph>
<paragraph id="6A1E8E163B0EFFAFFF26FC40FB8663E4" blockId="3.[151,1437,152,2034]" pageId="3" pageNumber="50">
<emphasis id="58D552043B0EFFAFFF26FC40FE0E633C" bold="true" box="[197,463,1891,1917]" pageId="3" pageNumber="50">Prosternal projection</emphasis>
high, apex alutaceous, anterior part transversely oval, externally pointed; posterior part shorter but wider than anterior, posterior border fimbriate with long bristles.
</paragraph>
<paragraph id="6A1E8E163B0EFFA8FF26FC92FEC564BE" blockId="3.[151,1437,152,2034]" lastBlockId="4.[151,1436,152,255]" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
<emphasis id="58D552043B0EFFAFFF26FC92FE0E638A" bold="true" box="[197,463,1969,1995]" pageId="3" pageNumber="50">Pronotal hypomera</emphasis>
alutaceous, with sparse, erect setae. Mesopleura, metapleura, and sides of metasternum alutaceous, with large and very dense confluent, annulate punctures; disc of metasternum smooth, with small, scattered punctures; metacoxae with punctures similar to those of pleurae but smaller and more disperse. Ventrites with annulate, moderately dense punctures, punctures smaller and sparser on apical ventrites.
</paragraph>
<caption id="3EDEDE9E3B09FFA8FF74FF3FFA896072" box="[151,1352,1052,1076]" httpUri="https://zenodo.org/record/186486/files/figure.png" pageId="4" pageNumber="51" targetBox="[378,1217,320,1022]" targetPageId="4">
<paragraph id="6A1E8E163B09FFA8FF74FF3FFA896072" blockId="4.[151,1352,1052,1076]" box="[151,1352,1052,1076]" pageId="4" pageNumber="51">
<emphasis id="58D552043B09FFA8FF74FF3FFED86075" bold="true" box="[151,281,1052,1076]" pageId="4" pageNumber="51">FIGURE 1.</emphasis>
Distribution of
<taxonomicName id="ADA1F5953B09FFA8FE27FF3EFD6B6075" box="[452,682,1053,1076]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B09FFA8FE27FF3EFD6B6075" box="[452,682,1053,1076]" italics="true" pageId="4" pageNumber="51">Cyclocephala bicolor</emphasis>
</taxonomicName>
(circles),
<taxonomicName id="ADA1F5953B09FFA8FCF5FF3EFC406075" box="[790,897,1052,1076]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="anibali" status="sp. nov.">
<emphasis id="58D552043B09FFA8FCF5FF3EFCE96075" box="[790,808,1053,1076]" italics="true" pageId="4" pageNumber="51">C</emphasis>
.
<emphasis id="58D552043B09FFA8FCD6FF3EFC406075" box="[821,897,1053,1076]" italics="true" pageId="4" pageNumber="51">anibali</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="43E6EF7F3B09FFA8FC6EFF3FFC086075" box="[909,969,1052,1076]" pageId="4" pageNumber="51" rank="species">
<emphasis id="58D552043B09FFA8FC6EFF3FFC086075" bold="true" box="[909,969,1052,1076]" pageId="4" pageNumber="51">sp. n.</emphasis>
</taxonomicNameLabel>
(triangle), and
<taxonomicName id="ADA1F5953B09FFA8FB8DFF3EFB226075" box="[1134,1251,1052,1076]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B09FFA8FB8DFF3EFB416075" box="[1134,1152,1053,1076]" italics="true" pageId="4" pageNumber="51">C</emphasis>
.
<emphasis id="58D552043B09FFA8FB6EFF3EFB226075" box="[1165,1251,1053,1076]" italics="true" pageId="4" pageNumber="51">sarahae</emphasis>
</taxonomicName>
(square).
</paragraph>
</caption>
<caption id="3EDEDE9E3B09FFA8FF74FC6AFB876321" box="[151,1094,1865,1889]" httpUri="https://zenodo.org/record/186487/files/figure.png" pageId="4" pageNumber="51" targetBox="[373,1218,1132,1833]" targetPageId="4">
<paragraph id="6A1E8E163B09FFA8FF74FC6AFB876321" blockId="4.[151,1094,1865,1889]" box="[151,1094,1865,1889]" pageId="4" pageNumber="51">
<emphasis id="58D552043B09FFA8FF74FC6AFED86320" bold="true" box="[151,281,1865,1889]" pageId="4" pageNumber="51">FIGURE 2.</emphasis>
Distribution of
<taxonomicName id="ADA1F5953B09FFA8FE27FC6AFD6E6321" box="[452,687,1865,1888]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B09FFA8FE27FC6AFD6E6321" box="[452,687,1865,1888]" italics="true" pageId="4" pageNumber="51">Cyclocephala dichroa</emphasis>
</taxonomicName>
(circles), and
<taxonomicName id="ADA1F5953B09FFA8FCABFC6AFC176321" box="[840,982,1865,1888]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B09FFA8FCABFC6AFC9B6321" box="[840,858,1865,1888]" italics="true" pageId="4" pageNumber="51">C</emphasis>
.
<emphasis id="58D552043B09FFA8FC84FC6AFC176321" box="[871,982,1865,1888]" italics="true" pageId="4" pageNumber="51">bicolorata</emphasis>
</taxonomicName>
(squares).
</paragraph>
</caption>
<paragraph id="6A1E8E163B09FFA8FF26FCADFE1E63E9" blockId="4.[151,1437,1934,2038]" box="[197,479,1934,1960]" pageId="4" pageNumber="51">
<emphasis id="58D552043B09FFA8FF26FCADFEFC63E9" bold="true" box="[197,317,1934,1960]" pageId="4" pageNumber="51">Aedeagus</emphasis>
:
<figureCitation id="F29A92933B09FFA8FEAEFCADFE1A63E9" box="[333,475,1934,1960]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="4" pageNumber="51">Figs. 69–70</figureCitation>
.
</paragraph>
<paragraph id="6A1E8E163B09FFA9FF26FC96FF396499" blockId="4.[151,1437,1934,2038]" lastBlockId="5.[151,1437,152,294]" lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51">
<emphasis id="58D552043B09FFA8FF26FC96FE86638E" bold="true" box="[197,327,1973,1999]" pageId="4" pageNumber="51">FEMALE</emphasis>
: differs from the male in the following characters: punctation generally stronger, that of pygidium of the same size; protibiae
<figureCitation id="F29A92933B09FFA8FDADFCFFFD6463B7" box="[590,677,2012,2038]" captionStart="FIGURE 24 – 33" captionStartId="7.[151,261,1850,1874]" captionTargetBox="[285,1302,542,1826]" captionTargetId="figure@7.[285,1302,542,1826]" captionTargetPageId="7" captionText="FIGURE 24 – 33. Figs. 24 – 25. Cyclocephala bicolor: 24) protibiae, male; 25) protibiae, female. Figs. 26 – 27. Cyclocephala bicolorata: 26) protibiae, male; 27) protibiae, female. Fig. 28 Cyclocephala sarahae protibiae, male (holotype). Figs. 29 – 30. Cyclocephala dichroa: 29) protibiae, male; 30) protibiae, female. Figs. 31 – 32. Cyclocephala anibali: 31) protibiae, male (holotype); 32) protibiae, female (allotype). Fig. 33. Cyclocephala bicolor metatarsus." httpUri="https://zenodo.org/record/186489/files/figure.png" pageId="4" pageNumber="51">Fig. 25</figureCitation>
; protarsi slender; posterior femora (
<figureCitation id="F29A92933B09FFA8FBADFCFFFB6463B7" box="[1102,1189,2012,2038]" captionStart="FIGURES 49 – 57" captionStartId="9.[151,278,1631,1655]" captionTargetBox="[368,1218,387,1607]" captionTargetId="figure@9.[368,1219,387,1607]" captionTargetPageId="9" captionText="FIGURES 49 – 57. Figs. 49 – 50. Cyclocephala bicolor: 49) metaleg, male; 50) metaleg, female. Figs. 51 – 52. Cyclocephala bicolorata: 51) metaleg, male; 52) metaleg female. Figs. 53 – 54. Cyclocephala dichroa: 53) metaleg, male; 54) metaleg, female. Figs. 55 – 56. Cyclocephala anibali: 55) metaleg, male (holotype); 56) metaleg, female (allotype). Fig. 57. Cyclocephala sarahae metaleg, male (holotype)." httpUri="https://zenodo.org/record/186491/files/figure.png" pageId="4" pageNumber="51">Fig. 50</figureCitation>
) somewhat stronger; epipleura (
<figureCitation id="F29A92933B08FFA9FEF9FBBBFEB464F3" box="[282,373,152,178]" captionStart="FIGURE 58 – 77" captionStartId="10.[151,256,1771,1795]" captionTargetBox="[300,1286,266,1745]" captionTargetId="figure@10.[300,1286,266,1745]" captionTargetPageId="10" captionText="FIGURE 58 – 77. Figs. 58 – 61. Female epipleurae (ventral view): 58) Cyclocephala bicolor, 59) C. bicolorata, 60) C. dichroa, 61) C. anibali. Figs. 62 – 68. Pygidium (lateral view): 62) C. bicolor, male; 63) C. bicolorata, male; 64) C. bicolorata, female; 65) C. dichroa, male; 66) C. dichroa, female; 67) C. anibali, male (holotype); 68) C. anibali, female (allotype). Figs. 69 – 70. Cyclocephala bicolor: 69) parameres, caudal view; 70) aedeagus, lateral view. Figs. 71 – 72. Cyclocephala dichroa: 71) parameres, caudal view; 72) aedeagus, lateral view. Figs. 73 – 74. Cyclocephala bicolorata (holotype): 73) parameres, caudal view; 74) aedeagus, lateral view. Fig. 75. Cyclocephala sarahae (holotype), parameres, caudal view. Figs. 76 – 77. Cyclocephala anibali (holotype): 76) parameres, caudal view; 77) aedeagus, lateral view." httpUri="https://zenodo.org/record/186492/files/figure.png" pageId="5" pageNumber="52">Fig. 58</figureCitation>
) strongly narrowed at level of metacoxa, then widened up to level of middle of fourth ventrite.
</paragraph>
</subSubSection>
<subSubSection id="22BBDD9D3B08FFABFF26FBC6FC4D65A6" lastPageId="7" lastPageNumber="54" pageId="5" pageNumber="52" type="distribution">
<paragraph id="6A1E8E163B08FFA9FF26FBC6FCCE6567" blockId="5.[151,1437,152,294]" pageId="5" pageNumber="52">
<emphasis id="58D552043B08FFA9FF26FBC6FEA964BE" bold="true" box="[197,360,229,255]" pageId="5" pageNumber="52">Distribution</emphasis>
:
<collectingCountry id="12B6CE863B08FFA9FE99FBC6FE3F64BE" box="[378,510,229,255]" name="Venezuela" pageId="5" pageNumber="52">Venezuela</collectingCountry>
,
<collectingCountry id="12B6CE863B08FFA9FDE8FBC6FD4864BE" box="[523,649,229,255]" name="Colombia" pageId="5" pageNumber="52">Colombia</collectingCountry>
,
<collectingCountry id="12B6CE863B08FFA9FD74FBC6FD3A64BE" box="[663,763,229,255]" name="Guyana" pageId="5" pageNumber="52">Guyana</collectingCountry>
,
<collectingCountry id="12B6CE863B08FFA9FCEBFBC6FC4264BE" box="[776,899,229,255]" name="Suriname" pageId="5" pageNumber="52">Suriname</collectingCountry>
,
<collectingCountry id="12B6CE863B08FFA9FC73FBC6FB8C64BE" box="[912,1101,229,255]" name="French Guiana" pageId="5" pageNumber="52">French Guiana</collectingCountry>
,
<collectingCountry id="12B6CE863B08FFA9FBB8FBC6FB6A64BE" box="[1115,1195,229,255]" name="Brazil" pageId="5" pageNumber="52">Brazil</collectingCountry>
,
<collectingCountry id="12B6CE863B08FFA9FB5BFBC6FB3464BE" box="[1208,1269,229,255]" name="Peru" pageId="5" pageNumber="52">Peru</collectingCountry>
, and
<collectingCountry id="12B6CE863B08FFA9FADBFBC6FA5864BE" box="[1336,1433,229,255]" name="Bolivia" pageId="5" pageNumber="52">Bolivia</collectingCountry>
.
<collectingCountry id="12B6CE863B08FFA9FF74FA2FFECF6567" box="[151,270,268,294]" name="Colombia" pageId="5" pageNumber="52">Colombia</collectingCountry>
,
<collectingCountry id="12B6CE863B08FFA9FEFAFA2FFE936567" box="[281,338,268,294]" name="Peru" pageId="5" pageNumber="52">Peru</collectingCountry>
, and
<collectingCountry id="12B6CE863B08FFA9FE6CFA2FFE246567" box="[399,485,268,294]" name="Bolivia" pageId="5" pageNumber="52">Bolivia</collectingCountry>
are new country records.
</paragraph>
<caption id="3EDEDE9E3B08FFA9FF74FDA7FD0F6367" httpUri="https://zenodo.org/record/186488/files/figure.png" pageId="5" pageNumber="52" targetBox="[276,1310,351,1643]" targetPageId="5">
<paragraph id="6A1E8E163B08FFA9FF74FDA7FD0F6367" blockId="5.[151,1437,1668,1830]" pageId="5" pageNumber="52">
<emphasis id="58D552043B08FFA9FF74FDA7FE9562DD" bold="true" box="[151,340,1668,1692]" pageId="5" pageNumber="52">FIGURES 3–23.</emphasis>
Figs. 3–6.
<taxonomicName id="ADA1F5953B08FFA9FE32FDA7FD7C62DA" box="[465,701,1668,1691]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B08FFA9FE32FDA7FD7C62DA" box="[465,701,1668,1691]" italics="true" pageId="5" pageNumber="52">Cyclocephala bicolor</emphasis>
</taxonomicName>
: 3) head frontal, 4) head lateral, 5) labium, 6) maxilla. Figs. 7–10.
<taxonomicName id="ADA1F5953B08FFA9FF74FD84FE5E62FF" box="[151,415,1703,1726]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B08FFA9FF74FD84FE5E62FF" box="[151,415,1703,1726]" italics="true" pageId="5" pageNumber="52">Cyclocephala bicolorata</emphasis>
</taxonomicName>
: 7) head frontal, 8) head lateral, 9) labium, 10) maxilla. Figs. 11–14.
<taxonomicName id="ADA1F5953B08FFA9FB63FD84FAAD62FF" box="[1152,1388,1703,1726]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B08FFA9FB63FD84FAAD62FF" box="[1152,1388,1703,1726]" italics="true" pageId="5" pageNumber="52">Cyclocephala dichroa</emphasis>
</taxonomicName>
: 11) head frontal, 12) head lateral, 13) labium, 14) maxilla. Figs. 15–18.
<taxonomicName id="ADA1F5953B08FFA9FC99FDE9FBA062A0" box="[890,1121,1738,1761]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B08FFA9FC99FDE9FBA062A0" box="[890,1121,1738,1761]" italics="true" pageId="5" pageNumber="52">Cyclocephala anibali</emphasis>
</taxonomicName>
(holotype): 15) head frontal, 16) head lateral, 17) labium, 18) maxilla. Figs. 19–23. Maxillary palpi: 19)
<taxonomicName id="ADA1F5953B08FFA9FC5EFDCFFBE86342" box="[957,1065,1772,1795]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="58D552043B08FFA9FC5EFDCFFC0E6342" box="[957,975,1772,1795]" italics="true" pageId="5" pageNumber="52">C</emphasis>
.
<emphasis id="58D552043B08FFA9FC3EFDCFFBE86342" box="[989,1065,1772,1795]" italics="true" pageId="5" pageNumber="52">bicolor</emphasis>
</taxonomicName>
, 20)
<taxonomicName id="ADA1F5953B08FFA9FB82FDCFFB2E6342" box="[1121,1263,1772,1795]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="58D552043B08FFA9FB82FDCFFBB26342" box="[1121,1139,1772,1795]" italics="true" pageId="5" pageNumber="52">C</emphasis>
.
<emphasis id="58D552043B08FFA9FB63FDCFFB2E6342" box="[1152,1263,1772,1795]" italics="true" pageId="5" pageNumber="52">bicolorata</emphasis>
</taxonomicName>
, 21)
<taxonomicName id="ADA1F5953B08FFA9FAC4FDCFFA5D6342" box="[1319,1436,1772,1795]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="58D552043B08FFA9FAC4FDCFFAF86342" box="[1319,1337,1772,1795]" italics="true" pageId="5" pageNumber="52">C</emphasis>
.
<emphasis id="58D552043B08FFA9FAA5FDCFFA5D6342" box="[1350,1436,1772,1795]" italics="true" pageId="5" pageNumber="52">sarahae</emphasis>
</taxonomicName>
(holotype), 22)
<taxonomicName id="ADA1F5953B08FFA9FEDEFC2CFE6F6367" box="[317,430,1807,1830]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="dichroa">
<emphasis id="58D552043B08FFA9FEDEFC2CFE8E6367" box="[317,335,1807,1830]" italics="true" pageId="5" pageNumber="52">C</emphasis>
.
<emphasis id="58D552043B08FFA9FEBFFC2CFE6F6367" box="[348,430,1807,1830]" italics="true" pageId="5" pageNumber="52">dichroa</emphasis>
</taxonomicName>
, 23)
<taxonomicName id="ADA1F5953B08FFA9FE05FC2CFD916367" box="[486,592,1807,1830]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="anibali">
<emphasis id="58D552043B08FFA9FE05FC2CFE396367" box="[486,504,1807,1830]" italics="true" pageId="5" pageNumber="52">C</emphasis>
.
<emphasis id="58D552043B08FFA9FDE6FC2CFD916367" box="[517,592,1807,1830]" italics="true" pageId="5" pageNumber="52">anibali</emphasis>
</taxonomicName>
(holotype).
</paragraph>
</caption>
<paragraph id="6A1E8E163B08FFA9FF26FC77FA5D63A3" blockId="5.[151,1437,1876,2018]" pageId="5" pageNumber="52">
<emphasis id="58D552043B08FFA9FF26FC77FE7E632F" bold="true" box="[197,447,1876,1902]" pageId="5" pageNumber="52">Specimens studied</emphasis>
:
<collectingCountry id="12B6CE863B08FFA9FE33FC77FD51632F" box="[464,656,1876,1902]" name="Venezuela" pageId="5" pageNumber="52">
<emphasis id="58D552043B08FFA9FE33FC77FD51632F" bold="true" box="[464,656,1876,1902]" pageId="5" pageNumber="52">VENEZUELA</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B08FFA9FD40FC77FCEF632F" bold="true" box="[675,814,1876,1902]" pageId="5" pageNumber="52">Amazonas</emphasis>
: Coromoto, Dto. Atures,
<date id="1E1FA8D63B08FFA9FB9BFC77FAC5632F" box="[1144,1284,1876,1902]" pageId="5" pageNumber="52" value="1979-06-01">VI-1979, 1</date>
F (HAHC). Culebra, Caño Negro, P. N. Duida-Marahuaca,
<geoCoordinate id="0F95E8D13B08FFA9FD03FC58FC8D63D4" box="[736,844,1915,1941]" direction="north" orientation="latitude" pageId="5" pageNumber="52" precision="925" value="3.55">3º 33’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B08FFA9FCB9FC58FC1863D4" box="[858,985,1915,1941]" direction="west" orientation="longitude" pageId="5" pageNumber="52" precision="925" value="-65.916664">65º 55’ W</geoCoordinate>
,
<date id="1E1FA8D63B08FFA9FC05FC58FB4863D4" box="[998,1161,1915,1941]" pageId="5" pageNumber="52" value="1983-10-06" valueMax="1983-10-16" valueMin="1983-10-06">6-16-X-1983</date>
, Exped. Terramar, 4 F (MIZA). La Esmeralda, LF Station,
<date id="1E1FA8D63B08FFA9FDAAFC81FD0F63FD" box="[585,718,1954,1980]" pageId="5" pageNumber="52" value="1997-10-01">01-X-1997</date>
, leg. Krimse, 1 F (MIZA). Pto. Ayacucho,
<date id="1E1FA8D63B08FFA9FB31FC81FA9E63FD" box="[1234,1375,1954,1980]" pageId="5" pageNumber="52" value="1967-04-22">22-IV-1967</date>
, P. J. Anduze, 1 M (HAHC);
<date id="1E1FA8D63B08FFA9FE5AFCEBFDAF63A3" box="[441,622,1992,2018]" pageId="5" pageNumber="52" value="1992-12-04" valueMax="1992-12-15" valueMin="1992-12-04">4-15-XII-1992</date>
, G. Romero, 1 F (MIZA). San Carlos de Río Negro,
<date id="1E1FA8D63B08FFA9FB12FCEBFAB063A3" box="[1265,1393,1992,2018]" pageId="5" pageNumber="52" value="1982-07-13">7-13-1982</date>
, A.
</paragraph>
<paragraph id="6A1E8E163B0BFFABFF74FBBBFECD650D" blockId="6.[151,1437,152,2034]" lastBlockId="7.[151,1437,151,487]" lastPageId="7" lastPageNumber="54" pageId="6" pageNumber="53">
Chacón-G. Yépez, 1 M, 15 F (MIZA); idem,
<geoCoordinate id="0F95E8D13B0BFFAAFD41FBBBFCC664F3" box="[674,775,152,178]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="1.9166667">1º 55’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFCF1FBBBFC4864F3" box="[786,905,152,178]" direction="west" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="-67.01667">67º 01’ W</geoCoordinate>
,
<date id="1E1FA8D63B0BFFAAFC77FBBBFBF964F3" box="[916,1080,152,178]" pageId="6" pageNumber="53" value="1984-03-04" valueMax="1984-03-14" valueMin="1984-03-04">4-14-III-1984</date>
, J. Clavijo-J.
<collectingCountry id="12B6CE863B0BFFAAFB34FBBBFB3864F3" box="[1239,1273,152,178]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="53">De</collectingCountry>
Marmels, 1 F (MIZA). San Fernando de Atabapo,
<date id="1E1FA8D63B0BFFAAFDA1FB9DFD226499" box="[578,739,190,216]" pageId="6" pageNumber="53" value="1982-08-17">17-VIII-1982</date>
, CEUM Fac. Agronomía, 1 F (MIZA).
<collectingCountry id="12B6CE863B0BFFAAFB57FB9DFA836499" box="[1204,1346,190,216]" name="Saint Lucia" pageId="6" pageNumber="53">Santa Lucía</collectingCountry>
,
<date id="1E1FA8D63B0BFFAAFAADFB9DFF3C64BE" pageId="6" pageNumber="53" value="1982-11-15" valueMax="1982-11-21" valueMin="1982-11-15">15-21- XI-1982</date>
, A. Chacón-G. Yépez, 1 F (MIZA). Talud Cerro Aracamuni,
<geoCoordinate id="0F95E8D13B0BFFAAFC31FBC6FB8064BE" box="[978,1089,229,255]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="1.4833333">1 º 29’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFBADFBC6FB0964BE" box="[1102,1224,229,255]" direction="west" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="-65.63333">65º 38’ W</geoCoordinate>
,
<quantity id="AD5923F33B0BFFAAFB37FBC6FAE364BE" box="[1236,1314,229,255]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="53" unit="m" value="600.0">600 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFACCFBC6FF166567" pageId="6" pageNumber="53" value="1987-10-18" valueMax="1987-10-24" valueMin="1987-10-18">18-24-X- 1987</date>
, Exped. Terramar, 3 M, 4 F (MIZA).
<emphasis id="58D552043B0BFFAAFD44FA28FCC66564" bold="true" box="[679,775,267,293]" pageId="6" pageNumber="53">Bolívar</emphasis>
: Anacoco,
<quantity id="AD5923F33B0BFFAAFC75FA2FFC166567" box="[918,983,268,294]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="53" unit="m" value="60.0">60 m</quantity>
,
<geoCoordinate id="0F95E8D13B0BFFAAFC07FA2FFB806567" box="[996,1089,268,294]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="6.0833335">6º 5’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFBADFA2FFB786567" box="[1102,1209,268,294]" direction="east" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="61.133335">
61º 8’
<collectingCountry id="12B6CE863B0BFFAAFB42FA2FFB786567" box="[1185,1209,268,294]" name="Iceland" pageId="6" pageNumber="53">O</collectingCountry>
</geoCoordinate>
,
<date id="1E1FA8D63B0BFFAAFB24FA2FFA596567" box="[1223,1432,268,294]" pageId="6" pageNumber="53" value="1979-08-10" valueMax="1979-08-30" valueMin="1979-08-10">10-30-VIII-1979</date>
, Expedición La Salle, 3 M, 2 F (MIZA). Canaima, Río Carrao, (Campamento Yuri),
<date id="1E1FA8D63B0BFFAAFB8AFA11FAC2650D" box="[1129,1283,306,332]" pageId="6" pageNumber="53" value="1972-02-09" valueMax="1972-02-15" valueMin="1972-02-09">9-15-II-1972</date>
, J. M. Ayala, 1 F (JMAL). El Paují, 100 Km
<collectingCountry id="12B6CE863B0BFFAAFDE9FA7AFDDE6532" box="[522,543,345,371]" name="Iceland" pageId="6" pageNumber="53">O</collectingCountry>
Sta. Elena,
<quantity id="AD5923F33B0BFFAAFD4EFA7AFCC86532" box="[685,777,345,371]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="53" unit="m" value="1100.0">1100 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFCF6FA7AFCB26532" box="[789,883,345,371]" pageId="6" pageNumber="53" value="1980-04-05">5-IV-80</date>
, 2 M, 2 F (CCBM). El Dorado-Sta. Elena, Km
<quantity id="AD5923F33B0BFFAAFF74FAA3FED265DB" box="[151,275,384,410]" metricMagnitude="1" metricUnit="m" metricValue="8.815999999999999" pageId="6" pageNumber="53" unit="m" value="88.16">88, 160 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFEC3FAA3FE7165DB" box="[288,432,384,410]" pageId="6" pageNumber="53" value="1967-09-28">28-IX-1967</date>
, C. J. Rosales-M. Gelbez-L. Rodríguez V., 2 F (MIZA);
<date id="1E1FA8D63B0BFFAAFBB8FAA3FB2B65DB" box="[1115,1258,384,410]" pageId="6" pageNumber="53" value="1967-09-27">27-IX-1967</date>
, L. J. Joly, 1 F (MHNLS). El Dorado-Sta. Elena, Km
<quantity id="AD5923F33B0BFFAAFD82FA85FD2A6581" box="[609,747,422,448]" metricMagnitude="2" metricUnit="m" metricValue="1.0946" pageId="6" pageNumber="53" unit="m" value="109.46">109, 460 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFD1BFA85FCB66581" box="[760,887,422,448]" pageId="6" pageNumber="53" value="1964-09-06">6-IX-1964</date>
, P. J. Salinas, 1 F (MIZA). Carret. El Dorado- Sta. Elena, Km
<quantity id="AD5923F33B0BFFAAFEB9FAEEFE1865A6" box="[346,473,461,487]" metricMagnitude="1" metricUnit="m" metricValue="8.809999999999999" pageId="6" pageNumber="53" unit="m" value="88.1">88, 100 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFE04FAEEFDBB65A6" box="[487,634,461,487]" pageId="6" pageNumber="53" value="1967-09-27">27-IX-1967</date>
, L. J. Joly, 1 F (MHNLS); idem,
<quantity id="AD5923F33B0BFFAAFBF5FAEEFBA665A6" box="[1046,1127,461,487]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="53" unit="m" value="200.0">200 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFB97FAEEFB1765A6" box="[1140,1238,461,487]" pageId="6" pageNumber="53" value="1980-04-07">7-IV-80</date>
, C. Bordón, 1 F (CCBM). 5 Km E Caicara,
<date id="1E1FA8D63B0BFFAAFE35FAD7FDA3664F" box="[470,610,500,526]" pageId="6" pageNumber="53" value="1996-06-11">11-VI-1996</date>
, B. D. Gill, 1 M (BDGC). Carret. Caicara-San Juan de Manapiare, Km
<quantity id="AD5923F33B0BFFAAFF74F939FEDE6675" box="[151,287,538,564]" metricMagnitude="2" metricUnit="m" metricValue="1.703" pageId="6" pageNumber="53" unit="m" value="170.3">170, 300 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFEC9F939FE296675" box="[298,488,538,564]" pageId="6" pageNumber="53" value="1973-12-21" valueMax="1973-12-30" valueMin="1973-12-21">21-30-XII-1973</date>
, J. L. García, 1 M (HAHC). Cerro Guaiquinima,
<quantity id="AD5923F33B0BFFAAFBC9F939FB476675" box="[1066,1158,538,564]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="53" unit="m" value="1000.0">1000 m</quantity>
,
<geoCoordinate id="0F95E8D13B0BFFAAFB71F939FB366675" box="[1170,1271,538,564]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="5.883333">5º 53‘ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFAE0F939FAB96675" box="[1283,1400,538,564]" direction="east" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="63.5">
63º 30’
<collectingCountry id="12B6CE863B0BFFAAFA83F939FAB96675" box="[1376,1400,538,564]" name="Iceland" pageId="6" pageNumber="53">O</collectingCountry>
</geoCoordinate>
,
<date id="1E1FA8D63B0BFFAAFA67F939FEC8661A" pageId="6" pageNumber="53" value="1990-02-03" valueMax="1990-02-04" valueMin="1990-02-03">3- 4-II-1990</date>
, Exped. FUDECI, J.
<collectingCountry id="12B6CE863B0BFFAAFDE2F962FDE2661A" box="[513,547,577,603]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="53">De</collectingCountry>
Marmels-A. Chacón, 2 F (MIZA). Guri, Río Caroní,
<quantity id="AD5923F33B0BFFAAFB74F962FB24661A" box="[1175,1253,577,603]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="53" unit="m" value="100.0">100 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFB13F962FABC661A" box="[1264,1405,577,603]" pageId="6" pageNumber="53" value="1966-11-16">16-XI-1966</date>
, J. &amp; B. Bechyne-E. Osuna, 3 F (MIZA);
<date id="1E1FA8D63B0BFFAAFD8EF94BFD3B66C3" box="[621,762,616,642]" pageId="6" pageNumber="53" value="1968-04-11">11-IV-1968</date>
, J. Salcedo, 1 M, 1 F (MIZA). Kamarata,
<quantity id="AD5923F33B0BFFAAFB1AF94BFA8866C3" box="[1273,1353,616,642]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="53" unit="m" value="450.0">450 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFAB5F94BFF1966E9" pageId="6" pageNumber="53" value="1978-02-24">24-II- 1978</date>
, L. J. Joly, 2 M, 5 F (MIZA). Kanarakuni, Alto Caura,
<quantity id="AD5923F33B0BFFAAFC6CF9ADFC2066E9" box="[911,993,654,680]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="53" unit="m" value="450.0">450 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFC0DF9ADFB6D66E9" box="[1006,1196,654,680]" pageId="6" pageNumber="53" value="1964-09-10" valueMax="1964-09-13" valueMin="1964-09-10">10-13-IX-1964</date>
, F. Fernández
<collectingCountry id="12B6CE863B0BFFAAFA80F9ADFABC66E9" box="[1379,1405,654,680]" name="Italy" pageId="6" pageNumber="53">Y</collectingCountry>
.-J. Bechyne, 5 F (MIZA). 35 Km N Las Trincheras,
<date id="1E1FA8D63B0BFFAAFD02F996FCAE668E" box="[737,879,693,719]" pageId="6" pageNumber="53" value="1987-06-17">17-VI-1987</date>
, S. &amp; J. Peck, woodland, forest edge, UV trap, 15 Km N Corocito, 2 M (HAHC). Los Pijiguaos,
<quantity id="AD5923F33B0BFFAAFCE5F9FFFC9966B7" box="[774,856,732,758]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="53" unit="m" value="190.0">190 m</quantity>
, Exped. MIZA, UCV,
<date id="1E1FA8D63B0BFFAAFB91F9FFFAE066B7" box="[1138,1313,732,758]" pageId="6" pageNumber="53" value="1992-06-03" valueMax="1992-06-13" valueMin="1992-06-03">3-13-VI-1992</date>
, 4 M, 4 F (MIZA). Pendare, Río Parguaza,
<geoCoordinate id="0F95E8D13B0BFFAAFDC4F821FD40675D" box="[551,641,770,796]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="6.1">6º 6’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFD6CF821FD3D675D" box="[655,764,770,796]" direction="west" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="-67.083336">67º 5’ W</geoCoordinate>
,
<date id="1E1FA8D63B0BFFAAFCEBF821FC63675D" box="[776,930,770,796]" pageId="6" pageNumber="53" value="1997-06-06" valueMax="1997-06-07" valueMin="1997-06-06">6-7-VI-1997</date>
, E. Osuna-A. Chacón-F. Rojas, 1 M, 38 F (MIZA). Río Caroní, Salto Las Babas,
<date id="1E1FA8D63B0BFFAAFD83F80AFCC26702" box="[608,771,809,835]" pageId="6" pageNumber="53" value="1983-04-08" valueMax="1983-04-10" valueMin="1983-04-08">8-10-IV-1983</date>
, Exped. Inst. Zool. Agric., 5 M, 5 F (MIZA). Río Caura, El Playón,
<date id="1E1FA8D63B0BFFAAFEFBF873FE65672B" box="[280,420,848,874]" pageId="6" pageNumber="53" value="1978-11-23">23-XI-1978</date>
, 3 F (MIZA). Río Caura, Salto Pará,
<quantity id="AD5923F33B0BFFAAFCB5F873FC62672B" box="[854,931,848,874]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="53" unit="m" value="250.0">250 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFC4CF873FBC1672B" box="[943,1024,848,874]" bridgedPair="-" pageId="6" pageNumber="53" value="1921-11">21-XI-</date>
178, B. Bechyné, 3 F; idem,
<date id="1E1FA8D63B0BFFAAFAADF873FEC067D1" pageId="6" pageNumber="53" value="1978-11-20" valueMax="1978-11-22" valueMin="1978-11-20">20-22- XI-1978</date>
, A. Chacón, 1 F (MIZA). Río Caura, Jabillal,
<quantity id="AD5923F33B0BFFAAFCDFF855FC4D67D1" box="[828,908,886,912]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="53" unit="m" value="100.0">100 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFC79F855FBED67D1" box="[922,1068,886,912]" pageId="6" pageNumber="53" value="1978-11-25">25-XI-1978</date>
, A. Chacón, 2 F (MIZA). Río Guaniamo,
<geoCoordinate id="0F95E8D13B0BFFAAFEFCF8BEFE4467F6" box="[287,389,925,951]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="6.75">6º 45’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFE72F8BEFDC567F6" box="[401,516,925,951]" direction="east" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="66.01667">
66º 01’
<collectingCountry id="12B6CE863B0BFFAAFE0EF8BEFDC567F6" box="[493,516,925,951]" name="Iceland" pageId="6" pageNumber="53">O</collectingCountry>
</geoCoordinate>
,
<quantity id="AD5923F33B0BFFAAFDF3F8BEFD9F67F6" box="[528,606,925,951]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="53" unit="m" value="160.0">160 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFD8AF8BEFCD067F6" box="[617,785,925,951]" pageId="6" pageNumber="53" value="1979-05-25" valueMax="1979-05-28" valueMin="1979-05-25">25-28-V-1979</date>
, J. Clavijo-A. Chacón-G. Yépez, 3 M, 2 F (MIZA). Río
<collectingCountry id="12B6CE863B0BFFAAFF74F8E7FF3F679F" box="[151,254,964,990]" name="Paraguay" pageId="6" pageNumber="53">Paragua</collectingCountry>
, E Río Cachicamo,
<quantity id="AD5923F33B0BFFAAFE14F8E7FD88679F" box="[503,585,964,990]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="6" pageNumber="53" unit="m" value="425.0">425 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFDB5F8E7FD26679F" box="[598,743,964,990]" pageId="6" pageNumber="53" value="1983-04-11">11-IV-1983</date>
, Exped. Inst. Zool. Agric., 1 F (MIZA). San Ignacio de Yuruaní, Gran Sabana,
<quantity id="AD5923F33B0BFFAAFE44F8C9FE346045" box="[423,501,1002,1028]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="6" pageNumber="53" unit="m" value="830.0">830 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFDE3F8C9FD4B6045" box="[512,650,1002,1028]" pageId="6" pageNumber="53" value="1988-04-19">19-IV-1988</date>
, A. Chacón, 1 M (MIZA). Suapure, Caura River,
<date id="1E1FA8D63B0BFFAAFB31F8C9FA926045" box="[1234,1363,1002,1028]" pageId="6" pageNumber="53" value="1899-05-27">V-27-1899</date>
, E. A. Klages, 1 M (HAHC). Uonquen,
<quantity id="AD5923F33B0BFFAAFDFEFF32FDAB606A" box="[541,618,1041,1067]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="53" unit="m" value="850.0">850 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFD95FF32FD32606A" box="[630,755,1041,1067]" pageId="6" pageNumber="53" value="1966-03-04">4-III-1966</date>
, A. Pérez, 1 F (MIZA).
<emphasis id="58D552043B0BFFAAFBEFFF33FBAE606B" bold="true" box="[1036,1135,1040,1066]" pageId="6" pageNumber="53">Táchira</emphasis>
: Navay,
<quantity id="AD5923F33B0BFFAAFB3AFF32FAE7606A" box="[1241,1318,1041,1067]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="53" unit="m" value="200.0">200 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFAD2FF32FA56606A" box="[1329,1431,1041,1067]" pageId="6" pageNumber="53" value="1978-10-30">30-X-78</date>
, C. Bordón, 1 F (CCBM).
<collectingCountry id="12B6CE863B0BFFAAFE20FF14FDA86010" box="[451,617,1079,1105]" name="Colombia" pageId="6" pageNumber="53">
<emphasis id="58D552043B0BFFAAFE20FF14FDA86010" bold="true" box="[451,617,1079,1105]" pageId="6" pageNumber="53">COLOMBIA</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B0BFFAAFD9BFF14FD236010" bold="true" box="[632,738,1079,1105]" pageId="6" pageNumber="53">Caquetá</emphasis>
, Río Caquetá at Tres Esquinas,
<date id="1E1FA8D63B0BFFAAFBB4FF1BFB0E6013" box="[1111,1231,1080,1106]" pageId="6" pageNumber="53" value="1969-01-18">18-I-1969</date>
, R. E. Dietz IV, 1 M (NMNH).
<collectingCountry id="12B6CE863B0BFFAAFED6FF7DFE726039" box="[309,435,1118,1144]" name="Guyana" pageId="6" pageNumber="53">
<emphasis id="58D552043B0BFFAAFED6FF7DFE726039" bold="true" box="[309,435,1118,1144]" pageId="6" pageNumber="53">GUYANA</emphasis>
</collectingCountry>
:
<collectingCountry id="12B6CE863B0BFFAAFE20FF7DFDB26039" box="[451,627,1118,1144]" name="Guyana" pageId="6" pageNumber="53">British Guiana</collectingCountry>
, 4 M, 1 F (AMNH), 1 M (MIZA).
<collectingCountry id="12B6CE863B0BFFAAFBF3FF7DFB456039" box="[1040,1156,1118,1144]" name="Guyana" pageId="6" pageNumber="53">Demerara</collectingCountry>
(Bartica), R. J. Crew, 1 M, 3 F (NMNH). Kamakusa, Jan, 1923, 1 M, 1 F, (AMNH). Kartabo, Bartica District,
<date id="1E1FA8D63B0BFFAAFB52FFA6FAF260DE" box="[1201,1331,1157,1183]" pageId="6" pageNumber="53" value="1920-09-07">7-IX-1920</date>
, Wm. L. Beebe collection, 1 F (AMNH);
<date id="1E1FA8D63B0BFFAAFDF6FF8FFD576087" box="[533,662,1196,1222]" pageId="6" pageNumber="53" value="1920-10-11">11-X-1920</date>
, Gift of N.
<collectingCountry id="12B6CE863B0BFFAAFCFDFF8FFCF26087" box="[798,819,1196,1222]" name="Italy" pageId="6" pageNumber="53">Y</collectingCountry>
. Zool. Soc. 1 F (AMNH). Mazaruni, Potaro District, Takuku Mountains,
<geoCoordinate id="0F95E8D13B0BFFAAFE67FFF0FE2A60AC" box="[388,491,1235,1261]" direction="north" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="6.25">6º 15’ N</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFE1BFFF0FDA260AC" box="[504,611,1235,1261]" direction="west" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="-59.083332">59º 5’ W</geoCoordinate>
,
<date id="1E1FA8D63B0BFFAAFD93FFF0FD3860AC" box="[624,761,1235,1261]" pageId="6" pageNumber="53" value="1983-12-07">7-XII-1983</date>
, at blacklight in forest clearing near streams, Earthwatch Research Expedition, P. J. Spangler &amp; W. E. Steiner, 1 M (NMNH). Penal settlement, Bartica District, 1917, Gift of N.
<collectingCountry id="12B6CE863B0BFFAAFEF7FE03FEE8617B" box="[276,297,1312,1338]" name="Italy" pageId="6" pageNumber="53">Y</collectingCountry>
. Zool. Soc., 1 F (AMNH). Rupununi, XI, 1913, 1 F (AMNH). Rockstone,
<collectingCountry id="12B6CE863B0BFFAAFB40FE03FADD617B" box="[1187,1308,1312,1338]" name="Guyana" pageId="6" pageNumber="53">Essequibo</collectingCountry>
River, 4 F (NMNH). Tumatumari, 1918, 19 F (AMNH), 1 F (MIZA); idem, VIII, 1913, 1 M, 17 F (AMNH). Tumatumari, Potaro R.,
<date id="1E1FA8D63B0BFFAAFE50FE4EFD8161C6" box="[435,576,1389,1415]" pageId="6" pageNumber="53" value="1927-06-29">29-VI-1927</date>
, Cornell Unv. Lot 760 Sub 117, 1 F (HAHC).
<collectingCountry id="12B6CE863B0BFFAAFBBCFE4EFB3E61C6" box="[1119,1279,1389,1415]" name="Suriname" pageId="6" pageNumber="53">
<emphasis id="58D552043B0BFFAAFBBCFE4EFB3E61C6" bold="true" box="[1119,1279,1389,1415]" pageId="6" pageNumber="53">SURINAME</emphasis>
</collectingCountry>
: Aibinia,
<date id="1E1FA8D63B0BFFAAFA96FE4EFECD61EF" pageId="6" pageNumber="53" value="1975-07-29">29- VII-1975</date>
, D. Engleman, at light, 1 F (UNSM). Brokopando District, Brownsberg Natuurpark, Mazaroni Plateau,
<quantity id="AD5923F33B0BFFAAFF1FFE98FE446194" box="[252,389,1467,1493]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="53" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400–500 m</quantity>
,
<date id="1E1FA8D63B0BFFAAFE73FE98FD8E6194" box="[400,591,1467,1493]" pageId="6" pageNumber="53" value="1982-08-20">20 August 1982</date>
, W. E. Steiner, Earthwatch
<collectingCountry id="12B6CE863B0BFFAAFC71FE98FBC36194" box="[914,1026,1467,1493]" name="Suriname" pageId="6" pageNumber="53">Suriname</collectingCountry>
Expedition,
<date id="1E1FA8D63B0BFFAAFB79FE98FAF26194" box="[1178,1331,1467,1493]" pageId="6" pageNumber="53" value="1982-08">August 1982</date>
, Collins, Early, Oberman, Pollock, Putnem, Steiner, 4 M, 2 F (NMNH). Moengo Boven, Cottica,
<date id="1E1FA8D63B0BFFAAFB57FEC2FAF761BA" box="[1204,1334,1505,1531]" pageId="6" pageNumber="53" value="1927-05-16">16-V-1927</date>
, Cornell Univ., Lot 760, Sub. 60, 1 F (NMNH).
<collectingCountry id="12B6CE863B0BFFAAFD88FD24FCA76260" box="[619,870,1543,1569]" name="French Guiana" pageId="6" pageNumber="53">
<emphasis id="58D552043B0BFFAAFD88FD24FCA76260" bold="true" box="[619,870,1543,1569]" pageId="6" pageNumber="53">FRENCH GUIANA</emphasis>
</collectingCountry>
: Kaw Rd. PK 37,
<date id="1E1FA8D63B0BFFAAFBA7FD2BFBA16263" box="[1092,1120,1544,1570]" pageId="6" pageNumber="53" value="1995-08-19">19</date>
-VIII-1995, 1 M (UNSM). Maroni River, 1 F (NMNH). Route de Petit Saut PK9
<date id="1E1FA8D63B0BFFAAFCE9FD0DFC6B6209" box="[778,938,1582,1608]" pageId="6" pageNumber="53" value="2000-08-25">25-VIII-2000</date>
,
<collectingCountry id="12B6CE863B0BFFAAFC56FD0DFC0B6209" box="[949,970,1582,1608]" name="Italy" pageId="6" pageNumber="53">Y</collectingCountry>
. Ponchel, 5 M (HAHC). St. Jean, W. M. Schaus col. 7 F (NMNH). 13 Km W D-5, Risq. Rd.
<date id="1E1FA8D63B0BFFAAFD1CFD76FC5E622E" box="[767,927,1621,1647]" pageId="6" pageNumber="53" value="1995-08-14">14-VIII-1995</date>
, J. E. Wappes, 3 M, 6 F (UNSM);
<date id="1E1FA8D63B0BFFAAFADAFD76FF1462D7" pageId="6" pageNumber="53" value="1995-08-22">22-VIII- 1995</date>
, 2 M, 1 F (UNSM), 1 M, 1 F (HAHC).
<collectingCountry id="12B6CE863B0BFFAAFD55FD58FCE962D4" box="[694,808,1659,1685]" name="Brazil" pageId="6" pageNumber="53">
<emphasis id="58D552043B0BFFAAFD55FD58FCE962D4" bold="true" box="[694,808,1659,1685]" pageId="6" pageNumber="53">BRAZIL</emphasis>
</collectingCountry>
: Robinson bequest, 1929, 1F (NMNH).
<emphasis id="58D552043B0BFFAAFAF3FD58FA5562D4" bold="true" box="[1296,1428,1659,1685]" pageId="6" pageNumber="53">Amazonas</emphasis>
: AM 0 10, Km 244,
<date id="1E1FA8D63B0BFFAAFE90FD80FE5362FC" box="[371,402,1699,1725]" pageId="6" pageNumber="53" value="1977-01-20">20</date>
-I-1977, B. C. Ratcliffe, 7 M, 6 F (UNSM). AM 0 10, Km 268,
<date id="1E1FA8D63B0BFFAAFB8AFD80FB4462FC" box="[1129,1157,1699,1725]" pageId="6" pageNumber="53" value="1977-01-18">18</date>
-I-1977, B. C. Ratcliffe, 3 F (UNSM). Manaus,
<date id="1E1FA8D63B0BFFAAFE45FDEAFDCD62A2" box="[422,524,1737,1763]" pageId="6" pageNumber="53" value="1983-10-30">30-X-83</date>
, C. Bordón, 2 M, 1 F (CCBM);
<date id="1E1FA8D63B0BFFAAFC60FDEAFC3A62A2" box="[899,1019,1737,1763]" pageId="6" pageNumber="53" value="1978-01-13">13-I-1978</date>
, B. C. Ratcliffe, 4 M, 2 F (UNSM);
<date id="1E1FA8D63B0BFFAAFF74FDD3FEE2634B" box="[151,291,1776,1802]" pageId="6" pageNumber="53" value="1978-11-13">13-XI-1978</date>
, B. C. Ratcliffe, 3 M, 3 F (UNSM). CEPLAC, 30 Km NE Manaus,
<date id="1E1FA8D63B0BFFAAFBA5FDD3FB1C634B" box="[1094,1245,1776,1802]" pageId="6" pageNumber="53" value="1976-12-12">12-XII-1976</date>
, B. C. Ratcliffe, 1 M, 1 F (UNSM).
<emphasis id="58D552043B0BFFAAFE67FC35FE206371" bold="true" box="[388,481,1814,1840]" pageId="6" pageNumber="53">Amapá</emphasis>
: Serra Lombard Lomao,
<date id="1E1FA8D63B0BFFAAFCF4FC35FC7C6371" box="[791,957,1814,1840]" pageId="6" pageNumber="53" value="1961-08-24">24-VIII-1961</date>
, J. &amp; B. Bechyne, 3 F (MIZA).
<emphasis id="58D552043B0BFFAAFAA6FC35FA526371" bold="true" box="[1349,1427,1814,1840]" pageId="6" pageNumber="53">Ceará</emphasis>
: Fortaleza,
<date id="1E1FA8D63B0BFFAAFEFBFC1EFE406316" box="[280,385,1853,1879]" pageId="6" pageNumber="53" value="1943-09">IX-1943</date>
, at light, Mac Creary, 1 F (NMNH).
<emphasis id="58D552043B0BFFAAFCA5FC1EFC2C6316" bold="true" box="[838,1005,1853,1879]" pageId="6" pageNumber="53">Mato Grosso</emphasis>
: Reserva Humboldt,
<geoCoordinate id="0F95E8D13B0BFFAAFB10FC1EFAA66316" box="[1267,1383,1853,1879]" direction="south" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="-10.183333">10º 11’ S</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFA96FC1EFF1B633F" direction="west" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="-59.8">59º 48’W</geoCoordinate>
,
<date id="1E1FA8D63B0BFFAAFF04FC47FE67633F" box="[231,422,1892,1918]" pageId="6" pageNumber="53" value="1977-07-13" valueMax="1977-07-14" valueMin="1977-07-13">13-14-VII-1977</date>
, B. C. Ratcliffe, 1 F (UNSM).
<emphasis id="58D552043B0BFFAAFCFBFC40FC94633C" bold="true" box="[792,853,1891,1917]" pageId="6" pageNumber="53">Pará</emphasis>
: Braçança,
<date id="1E1FA8D63B0BFFAAFC01FC47FB9A633F" box="[994,1115,1892,1918]" pageId="6" pageNumber="53" value="1949-01-20">20-I-1949</date>
, J. M. Díaz, 1 M (HAHC). Rio Negro, Santa Isabel,
<date id="1E1FA8D63B0BFFAAFE58FCA8FDD663E4" box="[443,535,1931,1957]" pageId="6" pageNumber="53" value="1930-10">X-1930</date>
, Holt, Blake &amp; Agostini, 5 F (NMNH). Cachimbo,
<quantity id="AD5923F33B0BFFAAFB8EFCA8FB7863E4" box="[1133,1209,1931,1957]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="53" unit="m" value="400.0">400 m</quantity>
, 14/
<date id="1E1FA8D63B0BFFAAFB08FCA8FAB663E4" box="[1259,1399,1931,1957]" pageId="6" pageNumber="53" value="1955-09-21">21-IX-1955</date>
, L. Travassos &amp; Oliveira, 5 M, 1 F (HAHC). Rio Xingu Camp.,
<geoCoordinate id="0F95E8D13B0BFFAAFCB9FC92FC79638A" box="[858,952,1969,1995]" direction="south" orientation="latitude" pageId="6" pageNumber="53" precision="925" value="-3.65">3º 39’ S</geoCoordinate>
,
<geoCoordinate id="0F95E8D13B0BFFAAFC26FC92FBFD638A" box="[965,1084,1969,1995]" direction="west" orientation="longitude" pageId="6" pageNumber="53" precision="925" value="-52.366665">52º 22’ W</geoCoordinate>
, ca 60 Km S Altamira,
<date id="1E1FA8D63B0BFFAAFAAFFC92FF1763B3" pageId="6" pageNumber="53" value="1986-10-01" valueMax="1986-10-07" valueMin="1986-10-01">1-7-X- 1986</date>
, 1F (NMNH). Santarem,
<date id="1E1FA8D63B0BFFAAFDEEFCFBFDB763B3" box="[525,630,2008,2034]" pageId="6" pageNumber="53" value="1933-09">IX-1933</date>
, L. C. Scaramuzza, 2 F (NMNH).
<emphasis id="58D552043B0BFFAAFBF8FCF4FB7F63B0" bold="true" box="[1051,1214,2007,2033]" pageId="6" pageNumber="53">Pernambuco</emphasis>
: Recife,
<date id="1E1FA8D63B0BFFAAFACDFCFBFA5963B3" box="[1326,1432,2008,2034]" pageId="6" pageNumber="53" value="1950-10-10">10-X-50</date>
, Asua leg, 1 F (HAHC).
<emphasis id="58D552043B0AFFABFE55FBB4FD0264F0" bold="true" box="[438,707,151,177]" pageId="7" pageNumber="54">Río Grande do Norte</emphasis>
:
<collectingCountry id="12B6CE863B0AFFABFD37FBBBFCD664F3" box="[724,791,152,178]" name="South Africa" pageId="7" pageNumber="54">Natal</collectingCountry>
, VIII-X-1954, 1 M (HAHC).
<emphasis id="58D552043B0AFFABFB98FBB4FB3764F0" bold="true" box="[1147,1270,151,177]" pageId="7" pageNumber="54">Rondonia</emphasis>
: Porto Velho, Rio Madeira,
<collectingCountry id="12B6CE863B0AFFABFEDBFB9DFEBD6499" box="[312,380,190,216]" name="Isle of Man" pageId="7" pageNumber="54">Mann</collectingCountry>
&amp;
<collectingCountry id="12B6CE863B0AFFABFE42FB9DFE286499" box="[417,489,190,216]" name="United States Pacific Island Wildlife Refuges" pageId="7" pageNumber="54">Baker</collectingCountry>
, 1918, 1 F (NMNH).
<emphasis id="58D552043B0AFFABFD07FB9DFCD86499" bold="true" box="[740,793,190,216]" pageId="7" pageNumber="54">Terr</emphasis>
.
<emphasis id="58D552043B0AFFABFCCAFB9DFCA56499" bold="true" box="[809,868,190,216]" pageId="7" pageNumber="54">Acre</emphasis>
: Río Branco,
<date id="1E1FA8D63B0AFFABFBEBFB9DFB496499" box="[1032,1160,190,216]" pageId="7" pageNumber="54" value="1952-05-15">15-V-1952</date>
, M. Alvarenga col., 1 F (HAHC). Ro. (?): Santa Cruz da Serra,
<date id="1E1FA8D63B0AFFABFD8CFBC6FD3564BE" box="[623,756,229,255]" pageId="7" pageNumber="54" value="1985-08-20">20-VIII-85</date>
, J. P. Caldwell, 3 F (HAHC). I have also seen specimens from
<collectingCountry id="12B6CE863B0AFFABFF3AFA28FE9B6564" box="[217,346,267,293]" name="Bolivia" pageId="7" pageNumber="54">
<emphasis id="58D552043B0AFFABFF3AFA28FE9B6564" bold="true" box="[217,346,267,293]" pageId="7" pageNumber="54">BOLIVIA</emphasis>
</collectingCountry>
: Beni, Guayamerin (AMNH).
<collectingCountry id="12B6CE863B0AFFABFD2CFA28FC806564" box="[719,833,267,293]" name="Brazil" pageId="7" pageNumber="54">
<emphasis id="58D552043B0AFFABFD2CFA28FC806564" bold="true" box="[719,833,267,293]" pageId="7" pageNumber="54">BRAZIL</emphasis>
</collectingCountry>
:
<emphasis id="58D552043B0AFFABFCB1FA28FC166564" bold="true" box="[850,983,267,293]" pageId="7" pageNumber="54">Amazonas</emphasis>
: Pará (AMNH).
<collectingCountry id="12B6CE863B0AFFABFB47FA28FB346564" box="[1188,1269,267,293]" name="Peru" pageId="7" pageNumber="54">
<emphasis id="58D552043B0AFFABFB47FA28FB346564" bold="true" box="[1188,1269,267,293]" pageId="7" pageNumber="54">PERU</emphasis>
</collectingCountry>
: Tingo María (AMNH).
</paragraph>
<paragraph id="6A1E8E163B0AFFABFF26FA7BFC4D65A6" blockId="7.[151,1437,151,487]" pageId="7" pageNumber="54">
<emphasis id="58D552043B0AFFABFF26FA7BFE456533" bold="true" box="[197,388,344,370]" pageId="7" pageNumber="54">Other localities</emphasis>
:
<collectingCountry id="12B6CE863B0AFFABFE77FA7BFDF26533" box="[404,563,344,370]" name="Suriname" pageId="7" pageNumber="54">
<emphasis id="58D552043B0AFFABFE77FA7BFDF26533" bold="true" box="[404,563,344,370]" pageId="7" pageNumber="54">SURINAME</emphasis>
</collectingCountry>
: Pakira-Iméné; Paramaribo;
<collectingCountry id="12B6CE863B0AFFABFC64FA7AFC3F6532" box="[903,1022,345,371]" name="Guyana" pageId="7" pageNumber="54">Demerara</collectingCountry>
.
<collectingCountry id="12B6CE863B0AFFABFBE9FA7BFAC56533" box="[1034,1284,344,370]" name="French Guiana" pageId="7" pageNumber="54">
<emphasis id="58D552043B0AFFABFBE9FA7BFAC56533" bold="true" box="[1034,1284,344,370]" pageId="7" pageNumber="54">FRENCH GUIANA</emphasis>
</collectingCountry>
: Acarouany; Ante cume Pata (Haut Kialo); Carbet République; Cayenne; Fleuve Oyapock, du Saut-Maripa à Massikiri; Kourou fôret; Mana; Organabo; route Conte; Saint Élie; Sinnamary.
<collectingCountry id="12B6CE863B0AFFABFC22FA85FBF06581" box="[961,1073,422,448]" name="Brazil" pageId="7" pageNumber="54">
<emphasis id="58D552043B0AFFABFC22FA85FBF06581" bold="true" box="[961,1073,422,448]" pageId="7" pageNumber="54">BRAZIL</emphasis>
</collectingCountry>
: Gorotire; São Felix do Xingu (Moyen Xingu) (
<bibRefCitation id="0E30F3E73B0AFFABFEBEFAEEFDC365A6" author="Endrodi" box="[349,514,461,487]" pageId="7" pageNumber="64" refString="Endrodi, S. (1966) Monographie der Dynastinae (Coleoptera, Lamellicornia). I Teil. Entomologishe Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 33, 1 - 460." type="journal article" year="1966">Endrödi 1966</bibRefCitation>
,
<bibRefCitation id="0E30F3E73B0AFFABFDEDFAEEFD6465A6" author="Gruner" box="[526,677,461,487]" pageId="7" pageNumber="64" refString="Gruner, L. (1971) Scarabaeidae Melolonthinae, Dynastinae, Rutelinae, Cetoniinae [Coleoptera] recoltes en Guyane Francaise par la Mission du Museum National d'Histoire Naturelle. Annales de la Societe Entomologique de France (N. S.), 7 (4), 843 - 848." type="journal article" year="1971">Gruner 1971</bibRefCitation>
,
<bibRefCitation id="0E30F3E73B0AFFABFD51FAEEFCBE65A6" author="Dechambre" box="[690,895,461,487]" pageId="7" pageNumber="64" refString="Dechambre, R. - P. (1979) Missions entomologiques en Guyane et au Bresil (Coleoptera Dynastidae) Revue Francaise d'Entomologie. (N. S.), 1 (4), 160 - 168." type="journal article" year="1979">Dechambre 1979</bibRefCitation>
).
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<taxonomicName id="DE24506060630BBFE5C0DB4D11A6E3D9" ID-CoL="4FY7S" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33604" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole auropilosa Mayr" lsidName-HNS="Pheidole auropilosa Mayr" order="Hymenoptera" pageNumber="658" phylum="Arthropoda" rank="species" species="auropilosa">Pheidole auropilosa</taxonomicName>
Mayr
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<taxonomicName id="E513F8E342C0E351C65A6BB8C6F13A44" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33604" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole auropilosa Mayr" lsidName-HNS="Pheidole auropilosa Mayr" order="Hymenoptera" pageNumber="658" phylum="Arthropoda" rank="species" species="auropilosa">Pheidole auropilosa</taxonomicName>
Mayr 1887: 596.
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<paragraph id="605ADF29E86AD1A949F680D0641609A2" pageNumber="658">Types Naturhist. Mus. Wien.</paragraph>
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etymology L
<taxonomicName id="9561565EF0D8CCF807FFA1A19D06F13F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33604" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole auropilosa Mayr" lsidName-HNS="Pheidole auropilosa Mayr" order="Hymenoptera" pageNumber="658" phylum="Arthropoda" rank="species" species="auropilosa">auropilosa</taxonomicName>
, golden (yellow) hair.
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Diagnosis A very large, yellow member of the
<taxonomicName id="1A3644329609F1650CA717E5952F79EB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34070" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tristis (Smith)" lsidName-HNS="Pheidole tristis (Smith)" order="Hymenoptera" pageNumber="658" phylum="Arthropoda" rank="species" species="tristis">tristis</taxonomicName>
group strongly distinguished by the following traits.
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<subSubSection id="21F3C1CFFB1DDD595FEE35C44E22B93D" type="description">
<paragraph id="A150C996EA132E17FEE5EE301AC436BA" pageNumber="658">Major: humerus seen in dorsal-oblique view very prominent, bluntly right-angular; a convex process extrudes from center of postpetiolar venter in side view; postpetiolar node diamond-shaped from above; anterior two-thirds of dorsal surface of head densely covered by longitudinal carinulae, those originating from the anterior margin of the lateral clypeal border curving outward toward the eye and those originating from the posterior frontal carinae traveling to the sides of the head.</paragraph>
<paragraph id="A0B1F5295279EA96FE137D47A270496B" pageNumber="658">Minor: occipital border broad, lacking nuchal collar; postpetiole diamond-shaped from above.</paragraph>
<paragraph id="A43E4A77700DBD2C729B80307D49950A" pageNumber="658">Measurements (mm) Lectotype major: HW 2.06, HL 2.06, SL 1.10, EL 0.22, PW 0.86.</paragraph>
<paragraph id="DCE16F67BF565AFA2AFE6B582ABC0B20" pageNumber="658">Paralectotype minor: HW 0.84, HL 0.84, SL 1.04, EL 0.14, PW 0.52.</paragraph>
<paragraph id="DF1262EF9045E2FE62264E93E283B44B" pageNumber="658">Color Major: body light reddish yellow, shaded slightly darker on the gaster.</paragraph>
<paragraph id="D85BDC85BFD565CB85B1282BE92492A3" pageNumber="658">Minor: concolorous dark yellow.</paragraph>
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<paragraph id="246DEBB0630268AC7A8F822650E39D1F" pageNumber="658">Range Southeastern Brazil, from Espirito Santo south to Rio Grande do Sul, and Misiones, Argentina.</paragraph>
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<paragraph id="27ECC0A19E06CD1C79B4697C4FC86464" pageNumber="658">Biology Unknown.</paragraph>
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<paragraph id="D64EBE06225AD6F8BE42E4D2F8A7FBAC" pageNumber="658">Figure Upper: lectotype, major. Lower: paralectotype, minor. BRAZIL: Santa Catarina. Scale bars = 1 mm.</paragraph>
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<taxonomicName id="ADA14D22284C7F03F7A0FBE7B006EB29" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[151,530,1096,1122]" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="haitensis">
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<emphasis id="58D5EAB3284C7F03F7A0FBE7B375EB29" bold="true" box="[151,353,1096,1122]" italics="true" pageId="19" pageNumber="20">Rolleia haitensis</emphasis>
Bartsch, 1946
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<figureCitation id="F29A2A24284C7F03F7A0FBC4B32EEBCF" box="[151,314,1131,1156]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="19" pageNumber="20">Figures 5 A–F</figureCitation>
,
<date id="1E1F1061284C7F03F673FBC3B376EBC8" box="[324,354,1132,1155]" pageId="19" pageNumber="20">
<figureCitation id="F29A2A24284C7F03F673FBC3B346EBC8" box="[324,338,1132,1155]" captionStart="FIGURE 7" captionStartId="20.[151,250,1604,1627]" captionTargetBox="[287,1301,204,1571]" captionTargetId="figure@20.[287,1302,193,1574]" captionTargetPageId="20" captionText="FIGURE 7. Distributions are approximate. Base map courtesy of NASA. A. Abbottella (Abbottella) calliotropis new species. B. Abbottella (Abbottella) diadema new species. C. Abbottella (Abbottella) dichroa new species. D. Leiabbottella soluta (Pfeiffer, 1852). E. Leiabbottella thompsoni new species. F. Abbottella (Abbottella) abbotti Bartsch, 1946. G. Abbottella (Abbottella) crossei (Pilsbry, 1933). H. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. I. Rolleia haitensis Bartsch, 1946." httpUri="https://zenodo.org/record/283735/files/figure.png" pageId="19" pageNumber="20">7</figureCitation>
I
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<typeStatus id="B51A8803284C7F03F7A0FB1BB2C6EB87" box="[151,210,1204,1228]" pageId="19" pageNumber="20">Type</typeStatus>
material.
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USNM 504088,
<typeStatus id="B51A8803284C7F03F532FB1CB07FEB80" box="[517,619,1203,1227]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
.
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<typeStatus id="B51A8803284C7F03F7FEFB77B316EBBB" box="[201,258,1240,1264]" pageId="19" pageNumber="20">Type</typeStatus>
locality.
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Ennery,
<collectingCountry id="12B67631284C7F03F6FCFB78B012EBA4" box="[459,518,1239,1263]" name="Haiti" pageId="19" pageNumber="20">Haiti</collectingCountry>
.
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<paragraph id="6A1E36A1284C7F03F7FEFB54B1B7EA10" blockId="19.[151,1436,1203,1696]" pageId="19" pageNumber="20">
<emphasis id="58D5EAB3284C7F03F7FEFB54B3EBEA5F" bold="true" box="[201,511,1275,1300]" pageId="19" pageNumber="20">Other material examined.</emphasis>
UF 32863, 9 km NE of “Dufailly” [= Duffalty], Centre Department,
<collectingCountry id="12B67631284C7F03F238FB54B75EEA58" box="[1295,1354,1275,1299]" name="Haiti" pageId="19" pageNumber="20">Haiti</collectingCountry>
, at
<quantity id="AD599B44284C7F03F245FB54B2A4EA7C" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="20" unit="m" value="280.0">280 m</quantity>
; UF 32777, hill W of waterfall,
<quantity id="AD599B44284C7F03F522FAB0B059EA7C" box="[533,589,1311,1335]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="km" value="2.0">2 km</quantity>
W of “Saut-d’Eau” [= Haut Saut De Eau], Centre Department,
<collectingCountry id="12B67631284C7F03F225FAB0B759EA7C" box="[1298,1357,1311,1335]" name="Haiti" pageId="19" pageNumber="20">Haiti</collectingCountry>
, at
<quantity id="AD599B44284C7F03F245FAB0B2A4EA10" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="19" pageNumber="20" unit="m" value="440.0">440 m</quantity>
; UF 236258, 2 km SW of Trou Forban, L’Ouest Department,
<collectingCountry id="12B67631284C7F03F454FAECB18BEA10" box="[867,927,1347,1371]" name="Haiti" pageId="19" pageNumber="20">Haiti</collectingCountry>
.
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<subSubSection id="22BB652A284C7F03F7FEFAC8B17EE9D4" pageId="19" pageNumber="20" type="distribution">
<paragraph id="6A1E36A1284C7F03F7FEFAC8B6BFE95B" blockId="19.[151,1436,1203,1696]" pageId="19" pageNumber="20">
<emphasis id="58D5EAB3284C7F03F7FEFAC8B3E0EACB" bold="true" box="[201,500,1383,1408]" pageId="19" pageNumber="20">Distribution and habitat.</emphasis>
Apparently limited to the Montagnes Noires range. Specimens were found in talus and in leaf litter in cut over forest, some in deep, wet ravines. Confusingly, there is a Dufailly and a Saut-d’Eau in the vicinity of Ennery (the
<typeStatus id="B51A8803284C7F03F6FFFA1FB3ECEA83" box="[456,504,1456,1480]" pageId="19" pageNumber="20">type</typeStatus>
locality) in Artibonite Department, and indeed Saut-d’Eau has a famous waterfall, but the correct UF localities are Duffalty and Haut Saut De Eau based on the details of the collection records. These records extend the range of this species ca.
<quantity id="AD599B44284C7F03F54AFA58B0D0E944" box="[637,708,1527,1551]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="20" unit="km" value="80.0">80 km</quantity>
to the southeast along this mountain range.
</paragraph>
<paragraph id="6A1E36A1284C7F03F7FEF9B4B17EE9D4" blockId="19.[151,1436,1203,1696]" pageId="19" pageNumber="20">
<emphasis id="58D5EAB3284C7F03F7FEF9B4B05BE97F" bold="true" box="[201,591,1563,1588]" pageId="19" pageNumber="20">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D22284C7F03F56CF9B4B108E978" box="[603,796,1563,1587]" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="haitensis">
<emphasis id="58D5EAB3284C7F03F56CF9B4B108E978" box="[603,796,1563,1587]" italics="true" pageId="19" pageNumber="20">Rolleia haitensis</emphasis>
</taxonomicName>
has a more adnate final whorl than does
<taxonomicName id="ADA14D22284C7F03F222F9B3B358E91C" authority="Maltzan, 1888" authorityName="Maltzan" authorityYear="1888" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="martensi">
<emphasis id="58D5EAB3284C7F03F222F9B3B788E978" box="[1301,1436,1564,1587]" italics="true" pageId="19" pageNumber="20">R. martensi</emphasis>
(Maltzan, 1888)
</taxonomicName>
from the Ennery region of
<collectingCountry id="12B67631284C7F03F5B7F990B0ACE91C" box="[640,696,1599,1623]" name="Haiti" pageId="19" pageNumber="20">Haiti</collectingCountry>
and has more numerous axial threads.
<taxonomicName id="ADA14D22284C7F03F347F990B717E91C" box="[1136,1283,1599,1623]" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="oberi">
<emphasis id="58D5EAB3284C7F03F347F990B717E91C" box="[1136,1283,1599,1623]" italics="true" pageId="19" pageNumber="20">Rolleia oberi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24284C7F03F225F990B766E91C" box="[1298,1394,1599,1623]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="19" pageNumber="20">Figure 5</figureCitation>
G– N) is smaller than
<taxonomicName id="ADA14D22284C7F03F65CF9CBB3FBE930" box="[363,495,1635,1659]" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="haitensis">
<emphasis id="58D5EAB3284C7F03F65CF9CBB3FBE930" box="[363,495,1635,1659]" italics="true" pageId="19" pageNumber="20">R. haitensis</emphasis>
</taxonomicName>
or
<taxonomicName id="ADA14D22284C7F03F520F9CBB08DE930" box="[535,665,1636,1659]" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="martensi">
<emphasis id="58D5EAB3284C7F03F520F9CBB08DE930" box="[535,665,1636,1659]" italics="true" pageId="19" pageNumber="20">R. martensi</emphasis>
</taxonomicName>
and has the axial sculpture produced into erect lamellae rather than threads; the outer lip is much more expanded in
<taxonomicName id="ADA14D22284C7F03F584F926B119E9D4" box="[691,781,1671,1696]" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="oberi">
<emphasis id="58D5EAB3284C7F03F584F926B119E9D4" box="[691,781,1671,1696]" italics="true" pageId="19" pageNumber="20">R. oberi</emphasis>
</taxonomicName>
as well.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0C27D0E3FA3E7F2CFB60CF6891660CF3">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B7284C7F05F7A0F95CB6C5EEE7" ID-DOI="http://doi.org/10.5281/zenodo.6164960" ID-GBIF-Taxon="127664021" ID-Zenodo-Dep="6164960" LSID="urn:lsid:plazi:treatment:E20887B7284C7F05F7A0F95CB6C5EEE7" httpUri="http://treatment.plazi.org/id/E20887B7284C7F05F7A0F95CB6C5EEE7" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection id="22BB652A284C7F03F7A0F95CB360E865" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="6A1E36A1284C7F03F7A0F95CB040E846" blockId="19.[151,596,1779,1838]" box="[151,596,1779,1805]" pageId="19" pageNumber="20">
<heading id="315681CD284C7F03F7A0F95CB040E846" bold="true" box="[151,596,1779,1805]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="ADA14D22284C7F03F7A0F95CB040E846" authority="Watters &amp; Duffy, 2010" authorityName="Watters &amp; Duffy" authorityYear="2010" box="[151,596,1779,1805]" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="oberi">
<emphasis id="58D5EAB3284C7F03F7A0F95CB040E846" bold="true" box="[151,596,1779,1805]" pageId="19" pageNumber="20">
<emphasis id="58D5EAB3284C7F03F7A0F95CB327E846" bold="true" box="[151,307,1779,1805]" italics="true" pageId="19" pageNumber="20">Rolleia oberi</emphasis>
Watters &amp; Duffy, 2010
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A1284C7F03F7A0F8B9B360E865" blockId="19.[151,596,1779,1838]" box="[151,372,1814,1838]" pageId="19" pageNumber="20">
<figureCitation id="F29A2A24284C7F03F7A0F8B9B314E865" box="[151,256,1814,1838]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="19" pageNumber="20">Figures 5</figureCitation>
G–N, 8 G
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284C7F05F7A0F8F1B194EE57" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="6A1E36A1284C7F03F7A0F8F1B038E8D1" blockId="19.[151,1436,1885,2018]" pageId="19" pageNumber="20">
<emphasis id="58D5EAB3284C7F03F7A0F8F1B35DE83D" bold="true" box="[151,329,1885,1910]" pageId="19" pageNumber="20">
<typeStatus id="B51A8803284C7F03F7A0F8F1B2C6E83D" box="[151,210,1886,1910]" pageId="19" pageNumber="20">Type</typeStatus>
material.
</emphasis>
UF 434775,
<typeStatus id="B51A8803284C7F03F6DEF8F1B05AE83D" box="[489,590,1886,1910]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
. UF 434776, 3
<typeStatus id="B51A8803284C7F03F43BF8F0B16FE83C" box="[780,891,1887,1911]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
; BMSM 17970, 2
<typeStatus id="B51A8803284C7F03F36CF8F0B6DEE83C" box="[1115,1226,1887,1911]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
; OSUM 35489, 2
<typeStatus id="B51A8803284C7F03F7A0F82CB312E8D0" box="[151,262,1923,1947]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
; all from the
<typeStatus id="B51A8803284C7F03F6ACF82CB3DFE8D0" box="[411,459,1923,1947]" pageId="19" pageNumber="20">type</typeStatus>
locality.
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<typeStatus id="B51A8803284C7F03F7FEF809B316E8F5" box="[201,258,1958,1982]" pageId="19" pageNumber="20">Type</typeStatus>
locality.
</emphasis>
On road half way between Santiago de los Caballeros and Puerto Plata, El Puerto, La Has, Puerto Plata Province,
<collectingCountry id="12B67631284C7F03F6A0F865B097E8A9" box="[407,643,1994,2018]" name="Dominican Republic" pageId="19" pageNumber="20">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B44284C7F03F59EF865B122E8A9" box="[681,822,1994,2018]" metricMagnitude="2" metricUnit="m" metricValue="9.15" metricValueMax="10.0" metricValueMin="8.3" pageId="19" pageNumber="20" unit="m" value="915.0" valueMax="1000.0" valueMin="830.0">830–1000 m</quantity>
in the Cordillera Septentrional.
</paragraph>
<caption id="3EDE6629284B7F04F7A0F9EBB33FE99E" httpUri="https://zenodo.org/record/283735/files/figure.png" pageId="20" pageNumber="21" targetBox="[287,1301,204,1571]" targetPageId="20">
<paragraph id="6A1E36A1284B7F04F7A0F9EBB33FE99E" blockId="20.[151,1437,1604,2025]" pageId="20" pageNumber="21">
<emphasis id="58D5EAB3284B7F04F7A0F9EBB307E911" bold="true" box="[151,275,1604,1627]" pageId="20" pageNumber="21">FIGURE 7.</emphasis>
Distributions are approximate. Base map courtesy of NASA. A.
<taxonomicName id="ADA14D22284B7F04F49BF9EBB706E911" box="[940,1298,1604,1626]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="calliotropis" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F49BF9EBB601E911" box="[940,1045,1604,1626]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F312F9EBB69AE911" box="[1061,1166,1604,1626]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F3A9F9EBB706E911" box="[1182,1298,1604,1626]" italics="true" pageId="20" pageNumber="21">calliotropis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8284B7F04F22FF9EAB782E910" box="[1304,1430,1604,1627]" pageId="20" pageNumber="21" rank="species">
<emphasis id="58D5EAB3284B7F04F22FF9EAB782E910" bold="true" box="[1304,1430,1604,1627]" pageId="20" pageNumber="21">new species</emphasis>
</taxonomicNameLabel>
. B.
<taxonomicName id="ADA14D22284B7F04F78EF9CCB01CE932" box="[185,520,1635,1657]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="diadema" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F78EF9CCB336E932" box="[185,290,1635,1657]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F602F9CCB38AE932" box="[309,414,1635,1657]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F686F9CCB01CE932" box="[433,520,1635,1657]" italics="true" pageId="20" pageNumber="21">diadema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8284B7F04F524F9CCB087E932" box="[531,659,1634,1657]" pageId="20" pageNumber="21" rank="species">
<emphasis id="58D5EAB3284B7F04F524F9CCB087E932" bold="true" box="[531,659,1634,1657]" pageId="20" pageNumber="21">new species</emphasis>
</taxonomicNameLabel>
. C.
<taxonomicName id="ADA14D22284B7F04F5F1F9CCB619E932" box="[710,1037,1635,1657]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="dichroa" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F5F1F9CCB124E932" box="[710,816,1635,1657]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F474F9CCB1B8E932" box="[835,940,1635,1657]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F488F9CCB619E932" box="[959,1037,1635,1657]" italics="true" pageId="20" pageNumber="21">dichroa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8284B7F04F32FF9CCB68CE932" box="[1048,1176,1634,1657]" pageId="20" pageNumber="21" rank="species">
<emphasis id="58D5EAB3284B7F04F32FF9CCB68CE932" bold="true" box="[1048,1176,1634,1657]" pageId="20" pageNumber="21">new species</emphasis>
</taxonomicNameLabel>
. D.
<taxonomicName id="ADA14D22284B7F04F3FBF9CCB32FE9DC" authority="Pfeiffer, 1852" authorityName="Pfeiffer" authorityYear="1852" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="soluta">
<emphasis id="58D5EAB3284B7F04F3FBF9CCB788E932" box="[1228,1436,1635,1657]" italics="true" pageId="20" pageNumber="21">Leiabbottella soluta</emphasis>
(Pfeiffer, 1852)
</taxonomicName>
. E.
<taxonomicName id="ADA14D22284B7F04F65DF92EB072E9DC" box="[362,614,1665,1687]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="thompsoni" status="sp. nov.">
<emphasis id="58D5EAB3284B7F04F65DF92EB072E9DC" box="[362,614,1665,1687]" italics="true" pageId="20" pageNumber="21">Leiabbottella thompsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8284B7F04F546F92DB0E6E9D3" box="[625,754,1665,1688]" pageId="20" pageNumber="21" rank="species">
<emphasis id="58D5EAB3284B7F04F546F92DB0E6E9D3" bold="true" box="[625,754,1665,1688]" pageId="20" pageNumber="21">new species</emphasis>
</taxonomicNameLabel>
. F.
<taxonomicName id="ADA14D22284B7F04F416F92EB717E9DC" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[801,1283,1665,1687]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="abbotti" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F416F92EB19EE9DC" box="[801,906,1665,1687]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F4A9F92EB613E9DC" box="[926,1031,1665,1687]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F32BF92EB670E9DC" box="[1052,1124,1665,1687]" italics="true" pageId="20" pageNumber="21">abbotti</emphasis>
Bartsch, 1946
</taxonomicName>
. G.
<taxonomicName id="ADA14D22284B7F04F205F92EB019E9FC" authority="Pilsbry, 1933" authorityName="Pilsbry" authorityYear="1933" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="crossei" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F205F92EB788E9DC" box="[1330,1436,1665,1687]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F7A8F90FB31DE9FD" box="[159,265,1696,1718]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F62BF90EB377E9FD" box="[284,355,1697,1718]" italics="true" pageId="20" pageNumber="21">crossei</emphasis>
(Pilsbry, 1933)
</taxonomicName>
. H.
<taxonomicName id="ADA14D22284B7F04F577F90FB6D0E9FD" authority="Watters &amp; Duffy, 2010" authorityName="Watters &amp; Duffy" authorityYear="2010" box="[576,1220,1696,1718]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="milleacantha" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F577F90FB0BEE9FD" box="[576,682,1696,1718]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F589F90FB133E9FD" box="[702,807,1696,1718]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F40CF90FB1ABE9FD" box="[827,959,1696,1718]" italics="true" pageId="20" pageNumber="21">milleacantha</emphasis>
Watters &amp; Duffy, 2010
</taxonomicName>
. I.
<taxonomicName id="ADA14D22284B7F04F3DAF90FB333E99E" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="haitensis">
<emphasis id="58D5EAB3284B7F04F3DAF90FB788E9FD" box="[1261,1436,1696,1718]" italics="true" pageId="20" pageNumber="21">Rolleia haitensis</emphasis>
Bartsch, 1946
</taxonomicName>
.
</paragraph>
</caption>
<caption id="3EDE6629284B7F04F7A0F972B3FFE825" pageId="20" pageNumber="21">
<paragraph id="6A1E36A1284B7F04F7A0F972B3FFE825" blockId="20.[151,1437,1604,2025]" pageId="20" pageNumber="21">
<emphasis id="58D5EAB3284B7F04F7A0F972B300E9B8" bold="true" box="[151,276,1757,1780]" pageId="20" pageNumber="21">FIGURE 8.</emphasis>
Distributions are approximate. Base map courtesy of NASA. A.
<taxonomicName id="ADA14D22284B7F04F48AF971B6FAE9BF" box="[957,1262,1758,1780]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="nitens" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F48AF971B632E9BF" box="[957,1062,1758,1780]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F30FF971B6B5E9BF" box="[1080,1185,1758,1780]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F385F970B6FAE9BF" box="[1202,1262,1759,1780]" italics="true" pageId="20" pageNumber="21">nitens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8284B7F04F3C0F971B761E9BF" box="[1271,1397,1757,1780]" pageId="20" pageNumber="21" rank="species">
<emphasis id="58D5EAB3284B7F04F3C0F971B761E9BF" bold="true" box="[1271,1397,1757,1780]" pageId="20" pageNumber="21">new species</emphasis>
</taxonomicNameLabel>
. B.
<taxonomicName id="ADA14D22284B7F04F7A0F953B3E5E859" box="[151,497,1788,1810]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="tenebrosa" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F7A0F953B315E859" box="[151,257,1788,1810]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F624F953B368E859" box="[275,380,1788,1810]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F6B9F953B3E5E859" box="[398,497,1788,1810]" italics="true" pageId="20" pageNumber="21">tenebrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8284B7F04F6CBF952B068E858" box="[508,636,1788,1811]" pageId="20" pageNumber="21" rank="species">
<emphasis id="58D5EAB3284B7F04F6CBF952B068E858" bold="true" box="[508,636,1788,1811]" pageId="20" pageNumber="21">new species</emphasis>
</taxonomicNameLabel>
. C.
<taxonomicName id="ADA14D22284B7F04F59AF953B66BE859" authority="Watters, 2010" authorityName="Watters" authorityYear="2010" box="[685,1151,1788,1810]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="aenea" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F59AF953B103E859" box="[685,791,1788,1810]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F41EF953B186E859" box="[809,914,1788,1810]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F492F952B1F5E859" box="[933,993,1789,1810]" italics="true" pageId="20" pageNumber="21">aenea</emphasis>
Watters, 2010
</taxonomicName>
. D.
<taxonomicName id="ADA14D22284B7F04F399F953B38EE87A" authority="Pfeiffer, 1858" authorityName="Pfeiffer" authorityYear="1858" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="wilhelmi" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F399F953B70CE859" box="[1198,1304,1788,1810]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F21DF953B787E859" box="[1322,1427,1788,1810]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F7A0F8B4B2FAE87A" box="[151,238,1819,1841]" italics="true" pageId="20" pageNumber="21">wilhelmi</emphasis>
(Pfeiffer, 1858)
</taxonomicName>
. E.
<taxonomicName id="ADA14D22284B7F04F6F1F8B3B129E87A" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[454,829,1819,1841]" class="Gastropoda" family="Pomatiidae" genus="Lagopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="lagopoma">
<emphasis id="58D5EAB3284B7F04F6F1F8B3B0B5E87A" box="[454,673,1819,1841]" italics="true" pageId="20" pageNumber="21">Lagopoma lagopoma</emphasis>
Bartsch, 1946
</taxonomicName>
. F.
<taxonomicName id="ADA14D22284B7F04F451F8B4B6FCE87A" authority="Watters, 2010" authorityName="Watters" authorityYear="2010" box="[870,1256,1819,1841]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="galaxius">
<emphasis id="58D5EAB3284B7F04F451F8B4B1F9E87A" box="[870,1005,1819,1841]" italics="true" pageId="20" pageNumber="21">Leiabbottella</emphasis>
<emphasis id="58D5EAB3284B7F04F4C1F8B4B658E87A" box="[1014,1100,1819,1841]" italics="true" pageId="20" pageNumber="21">galaxius</emphasis>
Watters, 2010
</taxonomicName>
. G.
<taxonomicName id="ADA14D22284B7F04F223F8B4B38DE804" authority="Watters &amp; Duffy, 2010" authorityName="Watters &amp; Duffy" authorityYear="2010" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oberi">
<emphasis id="58D5EAB3284B7F04F223F8B4B78FE87A" box="[1300,1435,1819,1841]" italics="true" pageId="20" pageNumber="21">Rolleia oberi</emphasis>
Watters &amp; Duffy, 2010
</taxonomicName>
. H.
<taxonomicName id="ADA14D22284B7F04F6E6F896B62AE804" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[465,1086,1849,1871]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="subSpecies" species="adolfi" subGenus="Abbottella" subSpecies="peninsularis">
<emphasis id="58D5EAB3284B7F04F6E6F896B02FE804" box="[465,571,1849,1871]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F566F896B0AEE804" box="[593,698,1849,1871]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F5E6F896B183E804" box="[721,919,1849,1871]" italics="true" pageId="20" pageNumber="21">adolfi peninsularis</emphasis>
Bartsch, 1946
</taxonomicName>
. I.
<taxonomicName id="ADA14D22284B7F04F35BF896B3F2E825" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="bombardopolensis" subGenus="Gundlachtudora">
<emphasis id="58D5EAB3284B7F04F35BF896B6C2E804" box="[1132,1238,1849,1871]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F3DBF896B787E804" box="[1260,1427,1849,1871]" italics="true" pageId="20" pageNumber="21">Gundlachtudora</emphasis>
)
<emphasis id="58D5EAB3284B7F04F7A0F8F7B345E825" box="[151,337,1880,1902]" italics="true" pageId="20" pageNumber="21">bombardopolensis</emphasis>
Bartsch, 1946
</taxonomicName>
.
</paragraph>
</caption>
<caption id="3EDE6629284B7F04F7ACF8D9B02BE8A2" pageId="20" pageNumber="21">
<paragraph id="6A1E36A1284B7F04F7ACF8D9B02BE8A2" blockId="20.[151,1437,1604,2025]" pageId="20" pageNumber="21">
<emphasis id="58D5EAB3284B7F04F7ACF8D9B303E8C7" bold="true" box="[155,279,1910,1933]" pageId="20" pageNumber="21">FIGURE 9.</emphasis>
Distributions are approximate. Base map courtesy of NASA. A.
<taxonomicName id="ADA14D22284B7F04F498F8D8B78DE8C6" authority="Pfeiffer, 1858" authorityName="Pfeiffer" authorityYear="1858" box="[943,1433,1911,1933]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="rosaliae" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F498F8D8B60DE8C6" box="[943,1049,1911,1933]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F310F8D8B684E8C6" box="[1063,1168,1911,1933]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F397F8D8B6E6E8C6" box="[1184,1266,1911,1933]" italics="true" pageId="20" pageNumber="21">rosaliae</emphasis>
(Pfeiffer, 1858)
</taxonomicName>
. B.
<taxonomicName id="ADA14D22284B7F04F78CF839B0C5E8E0" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[187,721,1941,1964]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="samanensis" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F78CF839B331E8E7" box="[187,293,1942,1964]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F60DF839B3B7E8E7" box="[314,419,1942,1964]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F68EF838B039E8E7" box="[441,557,1943,1964]" italics="true" pageId="20" pageNumber="21">samanensis</emphasis>
Bartsch, 1946
</taxonomicName>
. C.
<taxonomicName id="ADA14D22284B7F04F432F839B6E9E8E0" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[773,1277,1941,1964]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="sanchezi" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F432F839B17AE8E7" box="[773,878,1942,1964]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F4B3F839B1F9E8E7" box="[900,1005,1942,1964]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F335F839B64DE8E7" box="[1026,1113,1942,1964]" italics="true" pageId="20" pageNumber="21">sanchezi</emphasis>
Bartsch, 1946
</taxonomicName>
. D.
<taxonomicName id="ADA14D22284B7F04F205F839B39EE881" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="adolfi" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F205F839B788E8E7" box="[1330,1436,1942,1964]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F7A8F81BB31DE881" box="[159,265,1972,1994]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F613F81BB39EE881" box="[292,394,1972,1994]" italics="true" pageId="20" pageNumber="21">cf. adolfi</emphasis>
</taxonomicName>
<taxonomicName id="ADA14D22284B7F04F6ABF81BB082E881" authority="(Pfeiffer, 1852)" authorityName="Pfeiffer" authorityYear="1852" baseAuthorityName="Pfeiffer" baseAuthorityYear="1852" box="[412,662,1972,1994]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="adolfi">adolfi (Pfeiffer, 1852)</taxonomicName>
. E.
<taxonomicName id="ADA14D22284B7F04F5E3F81BB6ECE881" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[724,1272,1972,1994]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="sosuaensis" subGenus="Abbottella">
<emphasis id="58D5EAB3284B7F04F5E3F81BB129E881" box="[724,829,1972,1994]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F46FF81BB1D5E881" box="[856,961,1972,1994]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284B7F04F4EAF81AB65DE881" box="[989,1097,1973,1994]" italics="true" pageId="20" pageNumber="21">sosuaensis</emphasis>
Bartsch, 1946
</taxonomicName>
. F.
<taxonomicName id="ADA14D22284B7F04F205F81BB3A2E8A2" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="paradoxa" status="sp. nov." subGenus="Gundlachtudora">
<emphasis id="58D5EAB3284B7F04F205F81BB788E881" box="[1330,1436,1972,1994]" italics="true" pageId="20" pageNumber="21">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284B7F04F7A8F87CB352E8A2" box="[159,326,2003,2025]" italics="true" pageId="20" pageNumber="21">Gundlachtudora</emphasis>
)
<emphasis id="58D5EAB3284B7F04F662F87CB3A2E8A2" box="[341,438,2003,2025]" italics="true" pageId="20" pageNumber="21">paradoxa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8284B7F04F68BF87CB02DE8A2" box="[444,569,2002,2025]" pageId="20" pageNumber="21" rank="species">
<emphasis id="58D5EAB3284B7F04F68BF87CB02DE8A2" bold="true" box="[444,569,2002,2025]" pageId="20" pageNumber="21">new species</emphasis>
</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="6A1E36A1284A7F05F7FEFF38B194EE57" blockId="21.[151,1436,151,428]" pageId="21" pageNumber="22">
<emphasis id="58D5EAB3284A7F05F7FEFF38B014EFFB" bold="true" box="[201,512,151,176]" pageId="21" pageNumber="22">Other material examined.</emphasis>
UF 216195, 7 km SSW of Yásica Abajo, Loma el Indio Viejo, near Puerto Plata/ Santiago Province boundary,
<collectingCountry id="12B67631284A7F05F6D5FF13B0DAEF9F" box="[482,718,188,212]" name="Dominican Republic" pageId="21" pageNumber="22">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B44284A7F05F5C2FF13B154EF9F" box="[757,832,188,212]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="21" pageNumber="22" unit="m" value="800.0">800 m</quantity>
; UF 216186, 1 km NW of Laja de Yaroa, near Puerto Plata/Santiago Province boundary,
<collectingCountry id="12B67631284A7F05F514FF70B104EFBC" box="[547,784,223,247]" name="Dominican Republic" pageId="21" pageNumber="22">Dominican Republic</collectingCountry>
; UF 236228, Laja de Yaroa, Loma de Puerto, near Puerto Plata/Santiago Province boundary,
<collectingCountry id="12B67631284A7F05F517FEABB118EE57" box="[544,780,260,284]" name="Dominican Republic" pageId="21" pageNumber="22">Dominican Republic</collectingCountry>
, at
<specimenCount id="7CA7FD28284A7F05F405FEABB194EE57" box="[818,896,260,284]" pageId="21" pageNumber="22" type="generic">
<quantity id="AD599B44284A7F05F405FEABB168EE57" box="[818,892,260,284]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="21" pageNumber="22" unit="m" value="700.0">700 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284A7F05F7FEFE88B6C5EEE7" pageId="21" pageNumber="22" type="distribution">
<paragraph id="6A1E36A1284A7F05F7FEFE88B3F9EECC" blockId="21.[151,1436,151,428]" pageId="21" pageNumber="22">
<emphasis id="58D5EAB3284A7F05F7FEFE88B3E1EE0B" bold="true" box="[201,501,295,320]" pageId="21" pageNumber="22">Distribution and habitat.</emphasis>
This species has only been collected off the road from Puerto Plata to Santiago de los Caballeros, which crosses the Cordillera Septentrional. The extent of its distribution along that mountain range is unknown. Locally common.
</paragraph>
<paragraph id="6A1E36A1284A7F05F7FEFE3CB6C5EEE7" blockId="21.[151,1436,151,428]" box="[201,1233,403,428]" pageId="21" pageNumber="22">
<emphasis id="58D5EAB3284A7F05F7FEFE3CB054EEE7" bold="true" box="[201,576,403,428]" pageId="21" pageNumber="22">Comparison with other species.</emphasis>
See Comparison under
<taxonomicName id="ADA14D22284A7F05F47BFE3AB1C4EEE7" box="[844,976,404,428]" class="Gastropoda" family="Pomatiidae" genus="Rolleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="haitensis">
<emphasis id="58D5EAB3284A7F05F47BFE3AB1C4EEE7" box="[844,976,404,428]" italics="true" pageId="21" pageNumber="22">R. haitensis</emphasis>
</taxonomicName>
above (
<figureCitation id="F29A2A24284A7F05F31FFE3BB6D1EEE7" box="[1064,1221,404,428]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="21" pageNumber="22">Figure 5 A–F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="2B9F30EE2FE1AE5D7E3F1797E184D827">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<mods:namePart id="C655D581E9FB4B4B8071E3BAA1877782">Watters, G. Thomas</mods:namePart>
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<mods:date id="4A14921B6E77E4D803335D793B6A4801">2013</mods:date>
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<treatment id="E20887B7284E7F01F7A0FC01B0DFE962" ID-DOI="http://doi.org/10.5281/zenodo.6164954" ID-GBIF-Taxon="127663995" ID-Zenodo-Dep="6164954" LSID="urn:lsid:plazi:treatment:E20887B7284E7F01F7A0FC01B0DFE962" httpUri="http://treatment.plazi.org/id/E20887B7284E7F01F7A0FC01B0DFE962" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="22BB652A284E7F01F7A0FC01B371ECA2" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="6A1E36A1284E7F01F7A0FC01B050EC83" blockId="17.[151,580,942,1001]" box="[151,580,942,968]" pageId="17" pageNumber="18">
<heading id="315681CD284E7F01F7A0FC01B050EC83" bold="true" box="[151,580,942,968]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="ADA14D22284E7F01F7A0FC01B050EC83" authority="Pfeiffer, 1852" authorityName="Pfeiffer" authorityYear="1852" box="[151,580,942,968]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="soluta">
<emphasis id="58D5EAB3284E7F01F7A0FC01B050EC83" bold="true" box="[151,580,942,968]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7A0FC01B391EC83" bold="true" box="[151,389,942,968]" italics="true" pageId="17" pageNumber="18">Leiabbottella soluta</emphasis>
(Pfeiffer, 1852)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A1284E7F01F7A0FC7EB371ECA2" blockId="17.[151,580,942,1001]" box="[151,357,977,1001]" pageId="17" pageNumber="18">
<figureCitation id="F29A2A24284E7F01F7A0FC7EB314ECA2" box="[151,256,977,1001]" captionStart="FIGURE 2. A – E" captionStartId="9.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@9.[151,1436,193,1582]" captionTargetPageId="9" captionText="FIGURE 2. A – E. Leiabbottella thompsoni new species. A – C. Holotype (UF 456799), 13.7 mm width. D. Paratype (UF 236225), 14.1 mm width. E. Paratype (UF 236225), 13.6 mm width. F – I. Leiabbottella soluta (Pfeiffer, 1852). F. Illustration from Pfeiffer (1854: pl. 39, fig. 8). G. UF 216196, 11.9 mm width. H. UF 216196, 13.9 mm width. I. UF 216196, 13.8 mm width. J – L. Leiabbottella galaxius Watters, 2010. Paratype (OSUM 35491), 8.4 mm width." httpUri="https://zenodo.org/record/283730/files/figure.png" pageId="17" pageNumber="18">Figures 2</figureCitation>
F–I, 7 D
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284E7F01F7A0FBB5B367EA42" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="6A1E36A1284E7F01F7A0FBB5B3C1EB7A" blockId="17.[151,1437,1049,1577]" box="[151,469,1049,1074]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7A0FBB5B351EB79" bold="true" box="[151,325,1049,1074]" pageId="17" pageNumber="18">
<typeStatus id="B51A8803284E7F01F7A0FBB5B2C6EB79" box="[151,210,1050,1074]" pageId="17" pageNumber="18">Type</typeStatus>
material.
</emphasis>
Not located.
</paragraph>
<paragraph id="6A1E36A1284E7F01F7FEFB92B69CEB8A" blockId="17.[151,1437,1049,1577]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7FEFB92B37CEB1D" bold="true" box="[201,360,1085,1110]" pageId="17" pageNumber="18">
<typeStatus id="B51A8803284E7F01F7FEFB92B316EB1E" box="[201,258,1085,1109]" pageId="17" pageNumber="18">Type</typeStatus>
locality.
</emphasis>
Island of Santo
<collectingCountry id="12B67631284E7F01F51CFB92B08CEB1E" box="[555,664,1085,1109]" name="Dominican Republic" pageId="17" pageNumber="18">Domingo</collectingCountry>
. Restricted by Watters (2012) to the southern edge of Los Haitises Mountains, N of Majagual, ca.
<quantity id="AD599B44284E7F01F6C4FBCEB02DEB32" box="[499,569,1121,1145]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="km" value="12.0">12 km</quantity>
NW of Sabana Grande de Boyá, Monte Plata Province,
<collectingCountry id="12B67631284E7F01F39AFBCEB78DEB32" box="[1197,1433,1121,1145]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
. However, this is now believed to be a separate species,
<taxonomicName id="ADA14D22284E7F01F43AFB2AB60BEBD6" box="[781,1055,1157,1181]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="thompsoni" status="sp. nov.">
<emphasis id="58D5EAB3284E7F01F43AFB2AB60BEBD6" box="[781,1055,1157,1181]" italics="true" pageId="17" pageNumber="18">Leiabbottella thompsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8284E7F01F310FB29B6A5EBD6" box="[1063,1201,1157,1181]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
. The
<typeStatus id="B51A8803284E7F01F3C6FB29B735EBD5" box="[1265,1313,1158,1182]" pageId="17" pageNumber="18">type</typeStatus>
locality of
<taxonomicName id="ADA14D22284E7F01F7A0FB05B2EEEB8A" box="[151,250,1193,1217]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="soluta">
<emphasis id="58D5EAB3284E7F01F7A0FB05B2EEEB8A" box="[151,250,1193,1217]" italics="true" pageId="17" pageNumber="18">L. soluta</emphasis>
</taxonomicName>
is herein corrected to Yásica Abajo, Puerto Plata Province,
<collectingCountry id="12B67631284E7F01F4AFFB06B691EB8A" box="[920,1157,1193,1217]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
.
</paragraph>
<paragraph id="6A1E36A1284E7F01F7FEFB62B367EA42" blockId="17.[151,1437,1049,1577]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7FEFB62B01CEBAD" bold="true" box="[201,520,1229,1254]" pageId="17" pageNumber="18">Other material examined.</emphasis>
UF 216196, 7 km SSW of Yásica Abajo, Puerto Plata Province,
<collectingCountry id="12B67631284E7F01F228FB62B2EBEA42" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
, at
<specimenCount id="7CA7FD28284E7F01F612FB5EB367EA42" box="[293,371,1265,1289]" pageId="17" pageNumber="18" type="generic">
<quantity id="AD599B44284E7F01F612FB5EB37BEA42" box="[293,367,1265,1289]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="18" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284E7F01F7FEFABAB705EA3E" pageId="17" pageNumber="18" type="distribution">
<paragraph id="6A1E36A1284E7F01F7FEFABAB615EA1A" blockId="17.[151,1437,1049,1577]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7FEFABAB3ECEA65" bold="true" box="[201,504,1301,1326]" pageId="17" pageNumber="18">Distribution and habitat.</emphasis>
Known only from Yásica Abajo in the Cordillera Septentrional, which is on the road over the mountains between Puerto Plata and Santiago de los Caballeros.
</paragraph>
<paragraph id="6A1E36A1284E7F01F7FEFAF2B705EA3E" blockId="17.[151,1437,1049,1577]" box="[201,1297,1373,1398]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7FEFAF2B054EA3D" bold="true" box="[201,576,1373,1398]" pageId="17" pageNumber="18">Comparison with other species.</emphasis>
See comparison under
<taxonomicName id="ADA14D22284E7F01F471FAF2B64CEA3E" box="[838,1112,1373,1397]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="thompsoni">
<emphasis id="58D5EAB3284E7F01F471FAF2B64CEA3E" box="[838,1112,1373,1397]" italics="true" pageId="17" pageNumber="18">Leiabbottella thompsoni</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24284E7F01F351FAF2B710EA3E" box="[1126,1284,1373,1397]" captionStart="FIGURE 2. A – E" captionStartId="9.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@9.[151,1436,193,1582]" captionTargetPageId="9" captionText="FIGURE 2. A – E. Leiabbottella thompsoni new species. A – C. Holotype (UF 456799), 13.7 mm width. D. Paratype (UF 236225), 14.1 mm width. E. Paratype (UF 236225), 13.6 mm width. F – I. Leiabbottella soluta (Pfeiffer, 1852). F. Illustration from Pfeiffer (1854: pl. 39, fig. 8). G. UF 216196, 11.9 mm width. H. UF 216196, 13.9 mm width. I. UF 216196, 13.8 mm width. J – L. Leiabbottella galaxius Watters, 2010. Paratype (OSUM 35491), 8.4 mm width." httpUri="https://zenodo.org/record/283730/files/figure.png" pageId="17" pageNumber="18">Figure 2 A–E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284E7F01F7FEFA2DB0DFE962" pageId="17" pageNumber="18" type="discussion">
<paragraph id="6A1E36A1284E7F01F7FEFA2DB0DFE962" blockId="17.[151,1437,1049,1577]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7FEFA2DB329EAD1" bold="true" box="[201,317,1410,1434]" pageId="17" pageNumber="18">Remarks.</emphasis>
Watters (2012) identified Pfeiffer’s
<taxonomicName id="ADA14D22284E7F01F5E1FA2EB1D1EAD2" box="[726,965,1409,1433]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="solutus">
<emphasis id="58D5EAB3284E7F01F5E1FA2EB1D1EAD2" box="[726,965,1409,1433]" italics="true" pageId="17" pageNumber="18">Choanopoma solutus</emphasis>
</taxonomicName>
with a specimen from the Majugual area. At the time the presence of another, closely related species was unknown. Examination of over
<specimenCount id="7CA7FD28284E7F01F3E1FA0AB769EAF6" box="[1238,1405,1445,1469]" pageId="17" pageNumber="18" type="generic">200 specimens</specimenCount>
of this species has revealed that the description and figure of
<taxonomicName id="ADA14D22284E7F01F41AFA65B18AEAAA" box="[813,926,1481,1505]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="solutus">
<emphasis id="58D5EAB3284E7F01F41AFA65B18AEAAA" box="[813,926,1481,1505]" italics="true" pageId="17" pageNumber="18">C. solutus</emphasis>
</taxonomicName>
better match the second species from Yásica Abajo in the Cordillera Septentrional (compare
<figureCitation id="F29A2A24284E7F01F59AFA42B102E94E" box="[685,790,1517,1541]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="17" pageNumber="18">Figures 3</figureCitation>
F and G). The Majugual species in the Cordillera Central is here described as new,
<taxonomicName id="ADA14D22284E7F01F684F9BEB0D0E962" box="[435,708,1553,1577]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="thompsoni">
<emphasis id="58D5EAB3284E7F01F684F9BEB0D0E962" box="[435,708,1553,1577]" italics="true" pageId="17" pageNumber="18">Leiabbottella thompsoni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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Bartsch, 1946
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<figureCitation id="F29A2A24284E7F01F7A0FE3DB314EEE1" box="[151,256,402,426]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="17" pageNumber="18">Figures 4</figureCitation>
Y–CC, 8E
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locality.
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USNM 356198,
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.
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locality.
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Laguna,
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N of Samaná, Samaná Province,
<collectingCountry id="12B67631284E7F01F4ADFE51B691ED5D" box="[922,1157,510,534]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
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<emphasis id="58D5EAB3284E7F01F7FEFD8DB012ED70" bold="true" box="[201,518,546,571]" pageId="17" pageNumber="18">Other material examined.</emphasis>
UF 216120, El Valle, Samaná Province,
<collectingCountry id="12B67631284E7F01F4D8FD8DB6CBED71" box="[1007,1247,546,570]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
; UF 216201, El Valle, Samaná Province,
<collectingCountry id="12B67631284E7F01F699FDE9B08EED15" box="[430,666,582,606]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
; UF 236121, El Valle, Samaná Province,
<collectingCountry id="12B67631284E7F01F350FDE9B746ED15" box="[1127,1362,582,606]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
; GTW 10081a, El Valle, Samaná Province,
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.
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<emphasis id="58D5EAB3284E7F01F7FEFD21B3E3EDEC" bold="true" box="[201,503,654,679]" pageId="17" pageNumber="18">Distribution and habitat.</emphasis>
Found along the base of the high limestone cliffs in the region of El Valle in the Sierra de Samaná of the Cordillera Septentrional, possibly along the south-western edge of this outcrop as well. This species is found in association with
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and
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<emphasis id="58D5EAB3284E7F01F430FD79B189EDA5" box="[775,925,726,750]" italics="true" pageId="17" pageNumber="18">A. sosuaensis</emphasis>
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.
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<emphasis id="58D5EAB3284E7F01F7FEFD55B053EC58" bold="true" box="[201,583,762,787]" pageId="17" pageNumber="18">Comparison with other species.</emphasis>
This species is characterized by the peculiarly folded auricle and notched outer lip.
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<emphasis id="58D5EAB3284E7F01F632FCB1B3F6EC7D" box="[261,482,798,822]" italics="true" pageId="17" pageNumber="18">Abbottella sanchezi</emphasis>
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(
<figureCitation id="F29A2A24284E7F01F6C6FCB1B046EC7D" box="[497,594,798,822]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="17" pageNumber="18">Figure 4</figureCitation>
J–N) occasionally has a similar auricle but lacks the notch. However, it is questionable whether these minor features of the outer lip deserve a genus-level name.
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Watters, 2010
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J–L, 8 F
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284E7F01F7A0F972B393E8BA" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="6A1E36A1284E7F01F7A0F972B035E852" blockId="17.[151,1436,1757,2033]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7A0F972B35CE9BE" bold="true" box="[151,328,1757,1782]" pageId="17" pageNumber="18">
<typeStatus id="B51A8803284E7F01F7A0F972B2C4E9BE" box="[151,208,1757,1781]" pageId="17" pageNumber="18">Type</typeStatus>
material.
</emphasis>
UF 434779,
<typeStatus id="B51A8803284E7F01F6DBF972B046E9BE" box="[492,594,1757,1781]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
. UF 434780, 1
<typeStatus id="B51A8803284E7F01F425F971B162E9BD" box="[786,886,1758,1782]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
; BMSM 17972, 2
<typeStatus id="B51A8803284E7F01F36EF971B6DCE9BD" box="[1113,1224,1758,1782]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
; OSUM 35491, 1
<typeStatus id="B51A8803284E7F01F7A0F8ADB2EFE851" box="[151,251,1794,1818]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
; all from the
<typeStatus id="B51A8803284E7F01F6A7F8ADB3D4E851" box="[400,448,1794,1818]" pageId="17" pageNumber="18">type</typeStatus>
locality.
</paragraph>
<paragraph id="6A1E36A1284E7F01F7FEF88AB1C3E82A" blockId="17.[151,1436,1757,2033]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7FEF88AB37CE875" bold="true" box="[201,360,1829,1854]" pageId="17" pageNumber="18">
<typeStatus id="B51A8803284E7F01F7FEF88AB310E876" box="[201,260,1829,1853]" pageId="17" pageNumber="18">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B67631284E7F01F658F88AB04FE876" box="[367,603,1829,1853]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
, Samaná Province, Samaná Peninsula, along Rt. 5 between Santa Bárbara de Samaná and Sánchez, in the southern foothills of the Sierra de Samaná.
</paragraph>
<paragraph id="6A1E36A1284E7F01F7FEF8C2B393E8BA" blockId="17.[151,1436,1757,2033]" pageId="17" pageNumber="18">
<emphasis id="58D5EAB3284E7F01F7FEF8C2B01DE8CD" bold="true" box="[201,521,1901,1926]" pageId="17" pageNumber="18">Other material examined.</emphasis>
UF 236226,
<quantity id="AD599B44284E7F01F599F8C2B115E8CE" box="[686,769,1901,1925]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="km" value="0.5">0.5 km</quantity>
NNE of Caro Clara, María
<collectingCountry id="12B67631284E7F01F36CF8C2B6AEE8CE" box="[1115,1210,1901,1925]" name="Trinidad and Tobago" pageId="17" pageNumber="18">Trinidad</collectingCountry>
Sánchez Province,
<collectingCountry id="12B67631284E7F01F7A0F83EB393E8E2" box="[151,391,1937,1961]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B44284E7F01F68FF83EB3EEE8E2" box="[440,506,1937,1961]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="m" value="30.0">30 m</quantity>
; UF 216122, 3 km W of Cabrar [Cabrera], María
<collectingCountry id="12B67631284E7F01F36BF83EB6AFE8E2" box="[1116,1211,1937,1961]" name="Trinidad and Tobago" pageId="17" pageNumber="18">Trinidad</collectingCountry>
Sánchez Province,
<collectingCountry id="12B67631284E7F01F7A0F81AB399E886" box="[151,397,1973,1997]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
; UF 216192, 10 km WNW of Cabrar [Cabrera], María
<collectingCountry id="12B67631284E7F01F363F81AB6A0E886" box="[1108,1204,1973,1997]" name="Trinidad and Tobago" pageId="17" pageNumber="18">Trinidad</collectingCountry>
Sánchez Province,
<collectingCountry id="12B67631284E7F01F7A0F876B397E8BA" box="[151,387,2009,2033]" name="Dominican Republic" pageId="17" pageNumber="18">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284D7F02F7FEFF38B705EE74" pageId="18" pageNumber="19" type="distribution">
<paragraph id="6A1E36A1284D7F02F7FEFF38B0A5EE57" blockId="18.[151,1436,151,320]" pageId="18" pageNumber="19">
<emphasis id="58D5EAB3284D7F02F7FEFF38B3E3EFFB" bold="true" box="[201,503,151,176]" pageId="18" pageNumber="19">Distribution and habitat.</emphasis>
Originally described from the south coast of the Samaná Peninsula, new records extend the range to the Cabrera region on the north coast west of the peninsula. This is a region of limestone cliffs and karst ridges with cool, sheltered crevices in tropical forests. Specimens were only found on cliffs, under large boulders, and in solution pits, never under talus.
</paragraph>
<paragraph id="6A1E36A1284D7F02F7FEFE88B705EE74" blockId="18.[151,1436,151,320]" box="[201,1297,295,320]" pageId="18" pageNumber="19">
<emphasis id="58D5EAB3284D7F02F7FEFE88B054EE0B" bold="true" box="[201,576,295,320]" pageId="18" pageNumber="19">Comparison with other species.</emphasis>
See comparison under
<taxonomicName id="ADA14D22284D7F02F471FE87B64CEE0B" box="[838,1112,296,320]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="thompsoni">
<emphasis id="58D5EAB3284D7F02F471FE87B64CEE0B" box="[838,1112,296,320]" italics="true" pageId="18" pageNumber="19">Leiabbottella thompsoni</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24284D7F02F351FE88B710EE74" box="[1126,1284,295,319]" captionStart="FIGURE 2. A – E" captionStartId="9.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@9.[151,1436,193,1582]" captionTargetPageId="9" captionText="FIGURE 2. A – E. Leiabbottella thompsoni new species. A – C. Holotype (UF 456799), 13.7 mm width. D. Paratype (UF 236225), 14.1 mm width. E. Paratype (UF 236225), 13.6 mm width. F – I. Leiabbottella soluta (Pfeiffer, 1852). F. Illustration from Pfeiffer (1854: pl. 39, fig. 8). G. UF 216196, 11.9 mm width. H. UF 216196, 13.9 mm width. I. UF 216196, 13.8 mm width. J – L. Leiabbottella galaxius Watters, 2010. Paratype (OSUM 35491), 8.4 mm width." httpUri="https://zenodo.org/record/283730/files/figure.png" pageId="18" pageNumber="19">Figure 2 A–E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="5251CB428A8BEAE523F3D81BBD551981" ID-DOI="10.11646/zootaxa.3646.1.1" ID-GBIF-Dataset="9b6ec282-ff71-4189-81cb-f63930d210de" ID-ISSN="1175-5326" ID-Zenodo-Dep="283728" ID-ZooBank="55F70B5D-79AD-4C77-9164-EFE7126316E8" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460598948263" checkinUser="plazi" docAuthor="Watters, G. Thomas" docDate="2013" docId="E20887B7284F7F00F7A0FC01B128E84A" docLanguage="en" docName="zt03646p022.pdf" docOrigin="Zootaxa 3646 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Abbottella (Abbottella) wilhelmi Pfeiffer 1858" docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="1E31FFCF285F7F10F737FFAFB214EF4B" masterDocTitle="New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="17" updateTime="1698327222889" updateUser="plazi">
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<mods:title id="355A8BA6620E232FE8ACC0AD359C04EF">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="28B4B3A7E5EF1542698005DDA1D510D3">Watters, G. Thomas</mods:namePart>
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<mods:date id="C90B9D058A2E90D845992C997C86E22A">2013</mods:date>
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<treatment id="E20887B7284F7F00F7A0FC01B128E84A" ID-DOI="http://doi.org/10.5281/zenodo.6164948" ID-GBIF-Taxon="127664017" ID-Zenodo-Dep="6164948" LSID="urn:lsid:plazi:treatment:E20887B7284F7F00F7A0FC01B128E84A" httpUri="http://treatment.plazi.org/id/E20887B7284F7F00F7A0FC01B128E84A" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="22BB652A284F7F00F7A0FC01B374ECA2" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="6A1E36A1284F7F00F7A0FC01B0CCEC83" blockId="16.[151,728,942,1001]" box="[151,728,942,968]" pageId="16" pageNumber="17">
<heading id="315681CD284F7F00F7A0FC01B0CCEC83" bold="true" box="[151,728,942,968]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="ADA14D22284F7F00F7A0FC01B0CCEC83" authority="Pfeiffer, 1858" authorityName="Pfeiffer" authorityYear="1858" box="[151,728,942,968]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="wilhelmi" subGenus="Abbottella">
<emphasis id="58D5EAB3284F7F00F7A0FC01B0CCEC83" bold="true" box="[151,728,942,968]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7A0FC01B307EC83" bold="true" box="[151,275,942,968]" italics="true" pageId="16" pageNumber="17">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284F7F00F613FC01B3B4EC83" bold="true" box="[292,416,942,968]" italics="true" pageId="16" pageNumber="17">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284F7F00F687FC01B00CEC83" bold="true" box="[432,536,942,968]" italics="true" pageId="16" pageNumber="17">wilhelmi</emphasis>
(Pfeiffer, 1858)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A1284F7F00F7A0FC7EB374ECA2" blockId="16.[151,728,942,1001]" box="[151,352,977,1001]" pageId="16" pageNumber="17">Plates 4 T–X, 8 D</paragraph>
</subSubSection>
<subSubSection id="22BB652A284F7F00F7A0FBB5B310EAD2" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="6A1E36A1284F7F00F7A0FBB5B3C1EB7A" blockId="16.[151,1437,1049,1793]" box="[151,469,1049,1074]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7A0FBB5B351EB79" bold="true" box="[151,325,1049,1074]" pageId="16" pageNumber="17">
<typeStatus id="B51A8803284F7F00F7A0FBB5B2C6EB79" box="[151,210,1050,1074]" pageId="16" pageNumber="17">Type</typeStatus>
material.
</emphasis>
Not located.
</paragraph>
<paragraph id="6A1E36A1284F7F00F7FEFB92B3D7EB32" blockId="16.[151,1437,1049,1793]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7FEFB92B372EB1D" bold="true" box="[201,358,1085,1110]" pageId="16" pageNumber="17">
<typeStatus id="B51A8803284F7F00F7FEFB92B316EB1E" box="[201,258,1085,1109]" pageId="16" pageNumber="17">Type</typeStatus>
locality.
</emphasis>
<emphasis id="58D5EAB3284F7F00F658FB91B034EB1E" box="[367,544,1085,1109]" italics="true" pageId="16" pageNumber="17">
In insulae
<collectingCountry id="12B67631284F7F00F6D0FB91B034EB1E" box="[487,544,1086,1109]" name="Haiti" pageId="16" pageNumber="17">Haiti</collectingCountry>
</emphasis>
. But Pfeiffer continues that it “was collected under stones in moist places near Puerto Plata” [translation].
</paragraph>
<paragraph id="6A1E36A1284F7F00F7FEFB2AB310EAD2" blockId="16.[151,1437,1049,1793]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7FEFB2AB01EEBD5" bold="true" box="[201,522,1157,1182]" pageId="16" pageNumber="17">Other material examined.</emphasis>
UF 216149, 3 km N of Madre Vieja, Loma Blanca, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F7A0FB06B39CEB8A" box="[151,392,1193,1217]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 216150, 1 km N of Madre Vieja, Loma Blanca, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F217FB06B2EBEBAE" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B44284F7F00F613FB62B374EBAE" box="[292,352,1229,1253]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="17" unit="m" value="80.0">80 m</quantity>
; UF 216151, 18 km SE of Rio San Juan, María
<collectingCountry id="12B67631284F7F00F445FB62B1C6EBAE" box="[882,978,1229,1253]" name="Trinidad and Tobago" pageId="16" pageNumber="17">Trinidad</collectingCountry>
Sanchéz Province,
<collectingCountry id="12B67631284F7F00F39AFB62B783EBAE" box="[1197,1431,1229,1253]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B44284F7F00F783FB5EB2E5EA42" box="[180,241,1265,1289]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="17" unit="m" value="70.0">70 m</quantity>
; UF 184441, Puerto Plata, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F419FB5EB609EA42" box="[814,1053,1265,1289]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 158837, Puerto Plata, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F67FFABAB021EA66" box="[328,565,1301,1325]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 119145, Puerto Plata, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F353FABAB745EA66" box="[1124,1361,1301,1325]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; GTW 8273a, 15273b, Pico Isabel De Torres, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F46CFA96B65DEA1A" box="[859,1097,1337,1361]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; GTW 15273a, Nagua, María
<collectingCountry id="12B67631284F7F00F7A0FAF2B2E2EA3E" box="[151,246,1373,1397]" name="Trinidad and Tobago" pageId="16" pageNumber="17">Trinidad</collectingCountry>
Sanchéz Province,
<collectingCountry id="12B67631284F7F00F6D7FAF2B0C4EA3E" box="[480,720,1373,1397]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; Grego coll., Puerto Plata, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F217FAF2B314EAD2" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284F7F00F7FEFA0AB128E84A" pageId="16" pageNumber="17" type="distribution">
<paragraph id="6A1E36A1284F7F00F7FEFA0AB3EFE93A" blockId="16.[151,1437,1049,1793]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7FEFA0AB01FEAF5" bold="true" box="[201,523,1445,1470]" pageId="16" pageNumber="17">Distribution and habitat.</emphasis>
Specimens have been reported from isolated portions of the Cordillera Septentrional at Pico Isabel de Torres near Puerto Plata, Loma Blanca at El Choco, and the north side of the outcrop between Rio San Juan and Cabrera. Thus a large portion of its range is contained within the Parque Nacional Isabel de Torres and the Parque Nacional El Choco. Specimens were collected in rainforest on the Pico Isabel de Torres and elsewhere under limestone rubble in a grassy area with royal palms and evergreens in rolling, karsted hills, in some areas cut over for pasture.
</paragraph>
<paragraph id="6A1E36A1284F7F00F7FEF9D2B128E84A" blockId="16.[151,1437,1049,1793]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7FEF9D2B056E9DD" bold="true" box="[201,578,1661,1686]" pageId="16" pageNumber="17">Comparison with other species.</emphasis>
See Comparison under
<taxonomicName id="ADA14D22284F7F00F464F9D2B634E9DE" box="[851,1056,1661,1685]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="adolfi">
<emphasis id="58D5EAB3284F7F00F464F9D2B634E9DE" box="[851,1056,1661,1685]" italics="true" pageId="16" pageNumber="17">A. cf. adolfi adolfi</emphasis>
</taxonomicName>
above (
<figureCitation id="F29A2A24284F7F00F34CF9D2B6CFE9DE" box="[1147,1243,1661,1685]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="16" pageNumber="17">Figure 3</figureCitation>
G–I).
<taxonomicName id="ADA14D22284F7F00F210F9D2B2E3E9F2" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="sanchezi">
<emphasis id="58D5EAB3284F7F00F210F9D2B2E3E9F2" italics="true" pageId="16" pageNumber="17">Abbottella sanchezi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24284F7F00F63EF90EB379E9F2" box="[265,365,1697,1721]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="16" pageNumber="17">Figure 4</figureCitation>
J–N) is very similar but the spiral sculpture tends to form a carina at the periphery in that species.
<taxonomicName id="ADA14D22284F7F00F7CDF96AB3CDE996" box="[250,473,1733,1757]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="wilhelmi">
<emphasis id="58D5EAB3284F7F00F7CDF96AB3CDE996" box="[250,473,1733,1757]" italics="true" pageId="16" pageNumber="17">Abbottella wilhelmi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24284F7F00F6DBF96AB044E996" box="[492,592,1733,1757]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="16" pageNumber="17">Figure 4</figureCitation>
T–X) tends to have stronger, wider spiral cords than other species of a similar size and lacks keel-like spiral cords in the umbilicus.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="B85CBF62E4122C373FF224BF468015FF">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B7284F7F00F7A0FF38B1EBEC11" ID-DOI="http://doi.org/10.5281/zenodo.6164946" ID-GBIF-Taxon="127664016" ID-Zenodo-Dep="6164946" LSID="urn:lsid:plazi:treatment:E20887B7284F7F00F7A0FF38B1EBEC11" httpUri="http://treatment.plazi.org/id/E20887B7284F7F00F7A0FF38B1EBEC11" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="22BB652A284F7F00F7A0FF38B379EF99" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="6A1E36A1284F7F00F7A0FF38B0F0EFFA" blockId="16.[151,740,151,210]" box="[151,740,151,177]" pageId="16" pageNumber="17">
<heading id="315681CD284F7F00F7A0FF38B0F0EFFA" bold="true" box="[151,740,151,177]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="ADA14D22284F7F00F7A0FF38B0F0EFFA" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[151,740,151,177]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="sosuaensis" subGenus="Abbottella">
<emphasis id="58D5EAB3284F7F00F7A0FF38B0F0EFFA" bold="true" box="[151,740,151,177]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7A0FF38B307EFFA" bold="true" box="[151,275,151,177]" italics="true" pageId="16" pageNumber="17">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284F7F00F613FF38B3B4EFFA" bold="true" box="[292,416,151,177]" italics="true" pageId="16" pageNumber="17">Abbottella</emphasis>
)
<emphasis id="58D5EAB3284F7F00F687FF38B025EFFA" bold="true" box="[432,561,151,177]" italics="true" pageId="16" pageNumber="17">sosuaensis</emphasis>
Bartsch, 1946
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A1284F7F00F7A0FF15B379EF99" blockId="16.[151,740,151,210]" box="[151,365,186,210]" pageId="16" pageNumber="17">
<figureCitation id="F29A2A24284F7F00F7A0FF15B314EF99" box="[151,256,186,210]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="16" pageNumber="17">Figures 4</figureCitation>
O–S, 9 E
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284F7F00F7A0FEACB3ECEDC9" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="6A1E36A1284F7F00F7A0FEACB07BEE51" blockId="16.[151,1437,258,859]" box="[151,623,258,283]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7A0FEACB351EE50" bold="true" box="[151,325,258,283]" pageId="16" pageNumber="17">
<typeStatus id="B51A8803284F7F00F7A0FEACB2C6EE50" box="[151,210,259,283]" pageId="16" pageNumber="17">Type</typeStatus>
material.
</emphasis>
USNM 336768,
<typeStatus id="B51A8803284F7F00F532FEADB07FEE51" box="[517,619,258,282]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="6A1E36A1284F7F00F7FEFE88B1E8EE75" blockId="16.[151,1437,258,859]" box="[201,1020,294,319]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7FEFE88B371EE74" bold="true" box="[201,357,294,319]" pageId="16" pageNumber="17">
<typeStatus id="B51A8803284F7F00F7FEFE88B316EE74" box="[201,258,295,319]" pageId="16" pageNumber="17">Type</typeStatus>
locality.
</emphasis>
Sosúa,
<quantity id="AD599B44284F7F00F689FE89B00BEE75" box="[446,543,294,318]" metricMagnitude="4" metricUnit="m" metricValue="2.5749504" pageId="16" pageNumber="17" unit="mi" value="16.0">16 miles</quantity>
east of Puerto Plata,
<collectingCountry id="12B67631284F7F00F43BFE89B1ECEE75" box="[780,1016,294,318]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
.
</paragraph>
<paragraph id="6A1E36A1284F7F00F7FEFEE5B3ECEDC9" blockId="16.[151,1437,258,859]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7FEFEE5B3EAEE28" bold="true" box="[201,510,330,355]" pageId="16" pageNumber="17">Other material examined.</emphasis>
UF 218136, 3 km W of Puerto Plata, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F39CFEE5B78CEE29" box="[1195,1432,330,354]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 216124, 7 km E and
<quantity id="AD599B44284F7F00F683FEC1B3FAEECD" box="[436,494,366,390]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="km" value="2.0">2 km</quantity>
S of Sosúa, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F4B1FEC1B667EECD" box="[902,1139,366,390]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 216137, 4 km SW of Sosúa, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F531FE3DB0EEEEE1" box="[518,762,402,426]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 216182, 4 km E of Sosúa, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F7A0FE19B393EE85" box="[151,391,438,462]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 216179, 7 km E of Sosúa, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F324FE19B710EE85" box="[1043,1284,438,462]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 215993, Sosúa, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F6D1FE75B0C6EEB9" box="[486,722,474,498]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 216176, 1 km NE of Rio San Juan, María
<collectingCountry id="12B67631284F7F00F3EEFE75B72DEEB9" box="[1241,1337,474,498]" name="Trinidad and Tobago" pageId="16" pageNumber="17">Trinidad</collectingCountry>
Sanchéz Province,
<collectingCountry id="12B67631284F7F00F639FE51B014ED5D" box="[270,512,510,534]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; UF 216110,
<quantity id="AD599B44284F7F00F59FFE51B0EFED5D" box="[680,763,510,534]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="km" value="0.5">0.5 km</quantity>
NNE of Cano Claro, María
<collectingCountry id="12B67631284F7F00F36CFE51B6AEED5D" box="[1115,1210,510,534]" name="Trinidad and Tobago" pageId="16" pageNumber="17">Trinidad</collectingCountry>
Sanchéz Province,
<collectingCountry id="12B67631284F7F00F7A0FD8DB390ED71" box="[151,388,546,570]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B44284F7F00F699FD8DB3F9ED71" box="[430,493,546,570]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="17" unit="m" value="30.0">30 m</quantity>
; UF 216120, El Valle, Samaná Province,
<collectingCountry id="12B67631284F7F00F4FBFD8DB6ADED71" box="[972,1209,546,570]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; GTW 13728b, Rio San Juan, María
<collectingCountry id="12B67631284F7F00F66EFDE9B3ACED15" box="[345,440,582,606]" name="Trinidad and Tobago" pageId="16" pageNumber="17">Trinidad</collectingCountry>
Sanchéz Province,
<collectingCountry id="12B67631284F7F00F5ABFDE9B19EED15" box="[668,906,582,606]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
; GTW 9431a, outskirts of Sosúa, Puerto Plata Province,
<collectingCountry id="12B67631284F7F00F63FFDC5B3E0EDC9" box="[264,500,618,642]" name="Dominican Republic" pageId="16" pageNumber="17">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A284F7F00F7FEFD21B1EBEC11" pageId="16" pageNumber="17" type="distribution">
<paragraph id="6A1E36A1284F7F00F7FEFD21B3EFEC59" blockId="16.[151,1437,258,859]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7FEFD21B3E1EDEC" bold="true" box="[201,501,654,679]" pageId="16" pageNumber="17">Distribution and habitat.</emphasis>
Known from Puerto Plata through Cano Claro on the north slope of the Cordillera Septentrional and at El Valle on the Samaná Peninsula. Part of its range lies within the Parque Nacional El Choco. Specimens have been found under rocks in pasture along limestone rolling hills and under limestone rubble at the base of cliffs. Locally common.
</paragraph>
<paragraph id="6A1E36A1284F7F00F7FEFCB1B1EBEC11" blockId="16.[151,1437,258,859]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7FEFCB1B057EC7C" bold="true" box="[201,579,798,823]" pageId="16" pageNumber="17">Comparison with other species.</emphasis>
Although closely resembling
<taxonomicName id="ADA14D22284F7F00F4AEFCB1B62EEC7C" box="[921,1082,798,823]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="samanensis">
<emphasis id="58D5EAB3284F7F00F4AEFCB1B62EEC7C" box="[921,1082,798,823]" italics="true" pageId="16" pageNumber="17">A. samanensis</emphasis>
</taxonomicName>
(
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F–I) and
<taxonomicName id="ADA14D22284F7F00F22CFCB1B788EC7D" box="[1307,1436,798,822]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="sanchezi">
<emphasis id="58D5EAB3284F7F00F22CFCB1B788EC7D" box="[1307,1436,798,822]" italics="true" pageId="16" pageNumber="17">A. sanchezi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24284F7F00F7A9FCEDB2EAEC11" box="[158,254,834,858]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="16" pageNumber="17">Figure 4</figureCitation>
J–N),
<taxonomicName id="ADA14D22284F7F00F67DFCECB3F5EC10" box="[330,481,835,859]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="sosuaensis">
<emphasis id="58D5EAB3284F7F00F67DFCECB3F5EC10" box="[330,481,835,859]" italics="true" pageId="16" pageNumber="17">A. sosuaensis</emphasis>
</taxonomicName>
lacks the keel-like spiral cords in the umbilicus.
</paragraph>
</subSubSection>
</treatment>
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108
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<emphasis id="58D5EAB3284F7F00F7A0F8E5B19FE82F" bold="true" box="[151,907,1866,1892]" pageId="16" pageNumber="17">
<emphasis id="58D5EAB3284F7F00F7A0F8E5B307E82F" bold="true" box="[151,275,1866,1892]" italics="true" pageId="16" pageNumber="17">Abbottella</emphasis>
(
<emphasis id="58D5EAB3284F7F00F613F8E5B3F8E82F" bold="true" box="[292,492,1866,1892]" italics="true" pageId="16" pageNumber="17">Gundlachtudora</emphasis>
)
<emphasis id="58D5EAB3284F7F00F6CAF8E5B0CDE82F" bold="true" box="[509,729,1866,1892]" italics="true" pageId="16" pageNumber="17">bompardopolensis</emphasis>
Bartsch, 1946
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<figureCitation id="F29A2A24284F7F00F7A0F8C2B2E1E8CE" box="[151,245,1901,1925]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="16" pageNumber="17">Figure 1</figureCitation>
DD,
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material.
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USNM 504085,
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<typeStatus id="B51A8803284F7F00F7FEF875B316E8B9" box="[201,258,2010,2034]" pageId="16" pageNumber="17">Type</typeStatus>
locality.
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Crevices in rocks in a ravine a little west of Bombardopolis,
<collectingCountry id="12B67631284F7F00F322F876B645E8BA" box="[1045,1105,2009,2033]" name="Haiti" pageId="16" pageNumber="17">Haiti</collectingCountry>
.
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<emphasis id="58D5EAB3284E7F01F7FEFF38B3EAEFFB" bold="true" box="[201,510,151,176]" pageId="17" pageNumber="18">Other material examined.</emphasis>
UF 33037, 8 km W of Terre Neuve, Artibonite Department,
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<emphasis id="58D5EAB3284E7F01F7FEFF14B3E6EF9F" bold="true" box="[201,498,187,212]" pageId="17" pageNumber="18">Distribution and habitat.</emphasis>
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.
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<emphasis id="58D5EAB3284E7F01F7FEFEACB054EE57" bold="true" box="[201,576,259,284]" pageId="17" pageNumber="18">Comparison with other species.</emphasis>
See under
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<emphasis id="58D5EAB3284E7F01F58AFEABB153EE57" box="[701,839,260,284]" italics="true" pageId="17" pageNumber="18">A. paradoxa</emphasis>
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above (
<figureCitation id="F29A2A24284E7F01F497FEABB614EE57" box="[928,1024,260,284]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="17" pageNumber="18">Figure 1</figureCitation>
Z–CC).
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<treatment id="E20887B728507F1FF7A0FB29B00CE826" ID-DOI="http://doi.org/10.5281/zenodo.6164944" ID-GBIF-Taxon="127664015" ID-Zenodo-Dep="6164944" LSID="urn:lsid:plazi:treatment:E20887B728507F1FF7A0FB29B00CE826" httpUri="http://treatment.plazi.org/id/E20887B728507F1FF7A0FB29B00CE826" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="22BB652A28507F1FF7A0FB29B37EEB8A" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="6A1E36A128507F1FF7A0FB29B0DEEBEB" blockId="15.[151,714,1158,1217]" box="[151,714,1158,1184]" pageId="15" pageNumber="16">
<heading id="315681CD28507F1FF7A0FB29B0DEEBEB" bold="true" box="[151,714,1158,1184]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="ADA14D2228507F1FF7A0FB29B0DEEBEB" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[151,714,1158,1184]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="sanchezi" subGenus="Abbottella">
<emphasis id="58D5EAB328507F1FF7A0FB29B0DEEBEB" bold="true" box="[151,714,1158,1184]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7A0FB29B307EBEB" bold="true" box="[151,275,1158,1184]" italics="true" pageId="15" pageNumber="16">Abbottella</emphasis>
(
<emphasis id="58D5EAB328507F1FF613FB29B3B4EBEB" bold="true" box="[292,416,1158,1184]" italics="true" pageId="15" pageNumber="16">Abbottella</emphasis>
)
<emphasis id="58D5EAB328507F1FF687FB29B00CEBEB" bold="true" box="[432,536,1158,1184]" italics="true" pageId="15" pageNumber="16">sanchezi</emphasis>
Bartsch, 1946
</emphasis>
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</heading>
</paragraph>
<paragraph id="6A1E36A128507F1FF7A0FB06B37EEB8A" blockId="15.[151,714,1158,1217]" box="[151,362,1193,1217]" pageId="15" pageNumber="16">
<figureCitation id="F29A2A2428507F1FF7A0FB06B314EB8A" box="[151,256,1193,1217]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="15" pageNumber="16">Figures 4</figureCitation>
J–N, 9 C
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28507F1FF7A0FB5DB653E906" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="6A1E36A128507F1FF7A0FB5DB07BEA42" blockId="15.[151,1437,1265,1901]" box="[151,623,1265,1290]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7A0FB5DB351EA41" bold="true" box="[151,325,1265,1290]" pageId="15" pageNumber="16">
<typeStatus id="B51A880328507F1FF7A0FB5DB2C6EA41" box="[151,210,1266,1290]" pageId="15" pageNumber="16">Type</typeStatus>
material.
</emphasis>
USNM 504092,
<typeStatus id="B51A880328507F1FF532FB5EB07FEA42" box="[517,619,1265,1289]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="6A1E36A128507F1FF7FEFABAB1A1EA66" blockId="15.[151,1437,1265,1901]" box="[201,949,1301,1326]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFABAB371EA65" bold="true" box="[201,357,1301,1326]" pageId="15" pageNumber="16">
<typeStatus id="B51A880328507F1FF7FEFABAB316EA66" box="[201,258,1301,1325]" pageId="15" pageNumber="16">Type</typeStatus>
locality.
</emphasis>
<quantity id="AD599B4428507F1FF659FABAB3D4EA66" box="[366,448,1301,1325]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="15" pageNumber="16" unit="mi" value="2.0">2 miles</quantity>
northwest of Sánchez,
<collectingCountry id="12B6763128507F1FF5F1FABAB1A6EA66" box="[710,946,1301,1325]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
.
</paragraph>
<paragraph id="6A1E36A128507F1FF7FEFA96B653E906" blockId="15.[151,1437,1265,1901]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFA96B015EA19" bold="true" box="[201,513,1337,1362]" pageId="15" pageNumber="16">Other material examined.</emphasis>
UF 215988, 5 km ENE of Sánchez, Samaná Province,
<collectingCountry id="12B6763128507F1FF349FA96B778EA1A" box="[1150,1388,1337,1361]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; UF 236224, 2 km W of Sánchez, Samaná Province,
<collectingCountry id="12B6763128507F1FF5FCFAF2B1AEEA3E" box="[715,954,1373,1397]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; UF 215991, 3 km E of Samaná, Samaná Province,
<collectingCountry id="12B6763128507F1FF63CFA2EB3E8EAD2" box="[267,508,1409,1433]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; UF 215992, 15 km W of Samaná, Samaná Province,
<collectingCountry id="12B6763128507F1FF34DFA2EB77EEAD2" box="[1146,1386,1409,1433]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; UF 215989, 9 km E of Las Terreras, Samaná Province,
<collectingCountry id="12B6763128507F1FF5DFFA0AB1C2EAF6" box="[744,982,1445,1469]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; UF 215990, 3 km SW of Las Terreras, Samaná Province,
<collectingCountry id="12B6763128507F1FF643FA66B072EAAA" box="[372,614,1481,1505]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; UF 216143, 2 km W of Los Cacaos, Samaná Province,
<collectingCountry id="12B6763128507F1FF228FA66B314E94E" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; UF 216154, 40 km [
<quantity id="AD599B4428507F1FF535FA42B052E94E" box="[514,582,1517,1541]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="km" value="4.0">4 km</quantity>
?] NE of Sánchez, Samaná Province,
<collectingCountry id="12B6763128507F1FF4C8FA42B6FBE94E" box="[1023,1263,1517,1541]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
,
<quantity id="AD599B4428507F1FF3C9FA42B758E94E" box="[1278,1356,1517,1541]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="15" pageNumber="16" unit="m" value="270.0">270 m</quantity>
; GTW 7020a, near cave, Samaná Province,
<collectingCountry id="12B6763128507F1FF574F9BEB127E962" box="[579,819,1553,1577]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; GTW 14791a, Samaná Peninsula, Samaná Province,
<collectingCountry id="12B6763128507F1FF7A0F99AB397E906" box="[151,387,1589,1613]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; Grego coll., Sánchez, Samaná Province,
<collectingCountry id="12B6763128507F1FF461F99AB657E906" box="[854,1091,1589,1613]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28507F1FF7FEF9F6B00CE826" pageId="15" pageNumber="16" type="distribution">
<paragraph id="6A1E36A128507F1FF7FEF9F6B2D6E84A" blockId="15.[151,1437,1265,1901]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEF9F6B3EAE939" bold="true" box="[201,510,1625,1650]" pageId="15" pageNumber="16">Distribution and habitat.</emphasis>
Sierra de Samaná of the Cordillera Septentrional. Both coasts of the Samaná Peninsula, from at least Sánchez to Los Cacaos in the south, and Las Terrenas in the north. Most specimens were collected in association with limestone boulders, leaf litter, and talus in tropical forest, also near caves; some sites have been cleared for cocoa groves. I believe the record for UF 216154 of
<quantity id="AD599B4428507F1FF4DFF96AB624E996" box="[1000,1072,1733,1757]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="km" value="40.0">40 km</quantity>
NE of Sánchez is an error for
<quantity id="AD599B4428507F1FF2B9F96AB2AAE84A" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="km" value="4.0">4 km</quantity>
.
</paragraph>
<paragraph id="6A1E36A128507F1FF7FEF8A2B00CE826" blockId="15.[151,1437,1265,1901]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEF8A2B051E86D" bold="true" box="[201,581,1805,1830]" pageId="15" pageNumber="16">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D2228507F1FF57AF8A2B13EE86E" box="[589,810,1805,1829]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="wilhelmi">
<emphasis id="58D5EAB328507F1FF57AF8A2B13EE86E" box="[589,810,1805,1829]" italics="true" pageId="15" pageNumber="16">Abbottella wilhelmi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428507F1FF40CF8A2B188E86E" box="[827,924,1805,1829]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="15" pageNumber="16">Figure 4</figureCitation>
T–X) is very similar but the spiral sculpture does not form a keel at the periphery.
<taxonomicName id="ADA14D2228507F1FF508F89EB10FE802" box="[575,795,1841,1865]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="sanchezi">
<emphasis id="58D5EAB328507F1FF508F89EB10FE802" box="[575,795,1841,1865]" italics="true" pageId="15" pageNumber="16">Abbottella sanchezi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428507F1FF41DF89EB19EE802" box="[810,906,1841,1865]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="15" pageNumber="16">Figure 4</figureCitation>
J–N) differs from
<taxonomicName id="ADA14D2228507F1FF36BF89EB6E8E802" box="[1116,1276,1841,1865]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="samanensis">
<emphasis id="58D5EAB328507F1FF36BF89EB6E8E802" box="[1116,1276,1841,1865]" italics="true" pageId="15" pageNumber="16">A. samanensis</emphasis>
</taxonomicName>
in lacking the enrolled axial side of the outer lip.
</paragraph>
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109
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<taxonomicName id="ADA14D2228507F1FF7A0FD20B0FAEDE2" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[151,750,655,681]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="samanensis" subGenus="Abbottella">
<emphasis id="58D5EAB328507F1FF7A0FD20B0FAEDE2" bold="true" box="[151,750,655,681]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7A0FD20B307EDE2" bold="true" box="[151,275,655,681]" italics="true" pageId="15" pageNumber="16">Abbottella</emphasis>
(
<emphasis id="58D5EAB328507F1FF613FD20B3B4EDE2" bold="true" box="[292,416,655,681]" italics="true" pageId="15" pageNumber="16">Abbottella</emphasis>
)
<emphasis id="58D5EAB328507F1FF687FD20B028EDE2" bold="true" box="[432,572,655,681]" italics="true" pageId="15" pageNumber="16">samanensis</emphasis>
Bartsch, 1946
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A128507F1FF7A0FD1DB370ED81" blockId="15.[151,750,655,714]" box="[151,356,690,714]" pageId="15" pageNumber="16">
<figureCitation id="F29A2A2428507F1FF7A0FD1DB314ED81" box="[151,256,690,714]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="15" pageNumber="16">Figures 4</figureCitation>
F–I, 9 B
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28507F1FF7A0FD54B126ECE9" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="6A1E36A128507F1FF7A0FD54B07BEC59" blockId="15.[151,1437,762,1075]" box="[151,623,762,787]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7A0FD54B351EC58" bold="true" box="[151,325,762,787]" pageId="15" pageNumber="16">
<typeStatus id="B51A880328507F1FF7A0FD54B2C6EC58" box="[151,210,763,787]" pageId="15" pageNumber="16">Type</typeStatus>
material.
</emphasis>
USNM 504090,
<typeStatus id="B51A880328507F1FF532FD55B07FEC59" box="[517,619,762,786]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="6A1E36A128507F1FF7FEFCB0B111EC7D" blockId="15.[151,1437,762,1075]" box="[201,773,798,823]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFCB0B371EC7C" bold="true" box="[201,357,798,823]" pageId="15" pageNumber="16">
<typeStatus id="B51A880328507F1FF7FEFCB0B316EC7C" box="[201,258,799,823]" pageId="15" pageNumber="16">Type</typeStatus>
locality.
</emphasis>
Cape Samaná,
<collectingCountry id="12B6763128507F1FF522FCB1B115EC7D" box="[533,769,798,822]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
.
</paragraph>
<paragraph id="6A1E36A128507F1FF7FEFCEDB126ECE9" blockId="15.[151,1437,762,1075]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFCEDB3E9EC10" bold="true" box="[201,509,834,859]" pageId="15" pageNumber="16">Other material examined.</emphasis>
GTW 7085d, along Rt. 5 between Santa Bárbara de Samaná and Sánchez, Samaná Province,
<collectingCountry id="12B6763128507F1FF63BFCC9B3E9EC35" box="[268,509,870,894]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; GTW 7085c, Sánchez, Samaná Province,
<collectingCountry id="12B6763128507F1FF4CFFCC9B6FCEC35" box="[1016,1256,870,894]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
; GTW 14791a, Samaná Peninsula, Samaná Province,
<collectingCountry id="12B6763128507F1FF574FC25B13BECE9" box="[579,815,906,930]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28507F1FF7FEFC01B73AEB79" pageId="15" pageNumber="16" type="distribution">
<paragraph id="6A1E36A128507F1FF7FEFC01B110ECA1" blockId="15.[151,1437,762,1075]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFC01B3E1EC8C" bold="true" box="[201,501,942,967]" pageId="15" pageNumber="16">Distribution and habitat.</emphasis>
Southern coast of the Samaná Peninsula in the Sierra de Samaná of the Cordillera Septentrional. Live snails have been found in leaf litter.
</paragraph>
<paragraph id="6A1E36A128507F1FF7FEFC59B73AEB79" blockId="15.[151,1437,762,1075]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFC59B058EB44" bold="true" box="[201,588,1014,1039]" pageId="15" pageNumber="16">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D2228507F1FF560FC59B14CEB45" box="[599,856,1014,1038]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="samanensis">
<emphasis id="58D5EAB328507F1FF560FC59B14CEB45" box="[599,856,1014,1038]" italics="true" pageId="15" pageNumber="16">Abbottella samanensis</emphasis>
</taxonomicName>
is immediately identified by its peculiar enrolled outer lip on the axial side.
<taxonomicName id="ADA14D2228507F1FF6F6FBB4B089EB78" box="[449,669,1051,1075]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="sanchezi">
<emphasis id="58D5EAB328507F1FF6F6FBB4B089EB78" box="[449,669,1051,1075]" italics="true" pageId="15" pageNumber="16">Abbottella sanchezi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428507F1FF59CFBB5B11FEB79" box="[683,779,1050,1074]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="15" pageNumber="16">Figure 4</figureCitation>
J–N) is very similar but lacks this characteristic.
</paragraph>
</subSubSection>
</treatment>
</document>

193
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<treatment id="E20887B728527F1DF7A0FC25B13FE86E" ID-DOI="http://doi.org/10.5281/zenodo.6164938" ID-GBIF-Taxon="127664013" ID-Zenodo-Dep="6164938" LSID="urn:lsid:plazi:treatment:E20887B728527F1DF7A0FC25B13FE86E" httpUri="http://treatment.plazi.org/id/E20887B728527F1DF7A0FC25B13FE86E" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="22BB652A28527F1DF7A0FC25B391EC8E" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="6A1E36A128527F1DF7A0FC25B17BECEF" blockId="13.[151,879,906,965]" box="[151,879,906,932]" pageId="13" pageNumber="14">
<heading id="315681CD28527F1DF7A0FC25B17BECEF" bold="true" box="[151,879,906,932]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="ADA14D2228527F1DF7A0FC25B17BECEF" authority="Watters &amp; Duffy, 2010" authorityName="Watters &amp; Duffy" authorityYear="2010" box="[151,879,906,932]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="milleacantha" subGenus="Abbottella">
<emphasis id="58D5EAB328527F1DF7A0FC25B17BECEF" bold="true" box="[151,879,906,932]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7A0FC25B307ECEF" bold="true" box="[151,275,906,932]" italics="true" pageId="13" pageNumber="14">Abbottella</emphasis>
(
<emphasis id="58D5EAB328527F1DF613FC25B3B4ECEF" bold="true" box="[292,416,906,932]" italics="true" pageId="13" pageNumber="14">Abbottella</emphasis>
)
<emphasis id="58D5EAB328527F1DF687FC25B05AECEF" bold="true" box="[432,590,906,932]" italics="true" pageId="13" pageNumber="14">milleacantha</emphasis>
Watters &amp; Duffy, 2010
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A128527F1DF7A0FC02B391EC8E" blockId="13.[151,879,906,965]" box="[151,389,941,965]" pageId="13" pageNumber="14">
<figureCitation id="F29A2A2428527F1DF7A0FC02B314EC8E" box="[151,256,941,965]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="13" pageNumber="14">
Figures
<date id="1E1F106128527F1DF7C5FC02B314EC8E" box="[242,256,941,965]" pageId="13" pageNumber="14">3</date>
</figureCitation>
X–BB, 7 H
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28527F1DF7A0FC5AB060EA3E" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="6A1E36A128527F1DF7A0FC5AB165EB45" blockId="13.[151,1437,1013,1829]" box="[151,881,1013,1038]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7A0FC5AB351EB45" bold="true" box="[151,325,1013,1038]" pageId="13" pageNumber="14">
<typeStatus id="B51A880328527F1DF7A0FC5AB2C6EB46" box="[151,210,1013,1037]" pageId="13" pageNumber="14">Type</typeStatus>
material.
</emphasis>
UF 420728,
<typeStatus id="B51A880328527F1DF6EFFC5AB02AEB46" box="[472,574,1013,1037]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
. OSUM 32478, 1
<typeStatus id="B51A880328527F1DF43EFC59B179EB45" box="[777,877,1014,1038]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="6A1E36A128527F1DF7FEFBB5B15AEB7A" blockId="13.[151,1437,1013,1829]" box="[201,846,1049,1074]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7FEFBB5B371EB79" bold="true" box="[201,357,1049,1074]" pageId="13" pageNumber="14">
<typeStatus id="B51A880328527F1DF7FEFBB5B316EB79" box="[201,258,1050,1074]" pageId="13" pageNumber="14">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B6763128527F1DF659FBB6B04DEB7A" box="[366,601,1049,1073]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
, northeast
<collectingCountry id="12B6763128527F1DF5E5FBB6B0EFEB7A" box="[722,763,1049,1073]" name="Iceland" pageId="13" pageNumber="14">Isla</collectingCountry>
Saona.
</paragraph>
<paragraph id="6A1E36A128527F1DF7FEFB92B060EA3E" blockId="13.[151,1437,1013,1829]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7FEFB92B014EB1D" bold="true" box="[201,512,1085,1110]" pageId="13" pageNumber="14">Other material examined.</emphasis>
UF 216132, 2 km SE of El Peñon, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF39DFB92B78CEB1E" box="[1194,1432,1085,1109]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 216155, 2 km SE of El Peñon, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF418FBCEB60FEB32" box="[815,1051,1121,1145]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 33415,
<quantity id="AD599B4428527F1DF392FBCEB715EB32" box="[1189,1281,1121,1145]" metricMagnitude="4" metricUnit="m" metricValue="1.7899999999999998" pageId="13" pageNumber="14" unit="km" value="17.9">17.9 km</quantity>
NW of Punta Cana, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF6C7FB2AB0C8EBD6" box="[496,732,1157,1181]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 215971, 4 km SW of Punta Cana, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF7A0FB06B397EB8A" box="[151,387,1193,1217]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 215967, 3 km SW of Juanillo, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF317FB06B718EB8A" box="[1056,1292,1193,1217]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 216181, Boca de Yuma, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF56CFB62B153EBAE" box="[603,839,1229,1253]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 244456, Boca de Yuma, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF7A0FB5EB393EA42" box="[151,391,1265,1289]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 216161,
<collectingCountry id="12B6763128527F1DF518FB5EB04CEA42" box="[559,600,1265,1289]" name="Iceland" pageId="13" pageNumber="14">Isla</collectingCountry>
Saona, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF4ECFB5EB6D9EA42" box="[987,1229,1265,1289]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 216160,
<collectingCountry id="12B6763128527F1DF243FB5EB788EA42" box="[1396,1436,1265,1289]" name="Iceland" pageId="13" pageNumber="14">Isla</collectingCountry>
Saona, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF6C8FABAB0F8EA66" box="[511,748,1301,1325]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 216165,
<collectingCountry id="12B6763128527F1DF4B0FABAB1BBEA66" box="[903,943,1301,1325]" name="Iceland" pageId="13" pageNumber="14">Isla</collectingCountry>
Saona, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF217FABAB314EA1A" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; Grego coll., Guaraguao, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF470FA96B623EA1A" box="[839,1079,1337,1361]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; GTW 15254a, Guaraguao, La Altagracia Province,
<collectingCountry id="12B6763128527F1DF6B3FAF2B064EA3E" box="[388,624,1373,1397]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28527F1DF7FEFA2EB13FE86E" pageId="13" pageNumber="14" type="distribution">
<paragraph id="6A1E36A128527F1DF7FEFA2EB1D1E962" blockId="13.[151,1437,1013,1829]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7FEFA2EB3E1EAD1" bold="true" box="[201,501,1409,1434]" pageId="13" pageNumber="14">Distribution and habitat.</emphasis>
Described from
<collectingCountry id="12B6763128527F1DF582FA2EB0C9EAD2" box="[693,733,1409,1433]" name="Iceland" pageId="13" pageNumber="14">Isla</collectingCountry>
Saona, this species is now known to occur along the adjacent coast of mainland La Altagracia Province as well. It seems to be particularly associated with limestone ridges such as the one that runs from El Peñon to El Cabo and around the coast to Boca de Yuma and further to the southeastern shore of the peninsula. A portion of its range is within the Parque Nacional Del Este. Snails live in sub-mesic to mesic forests along these limestone ridges under blocks and in leaf litter.
</paragraph>
<paragraph id="6A1E36A128527F1DF7FEF99AB13FE86E" blockId="13.[151,1437,1013,1829]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7FEF99AB054E905" bold="true" box="[201,576,1589,1614]" pageId="13" pageNumber="14">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D2228527F1DF570F99AB141E906" box="[583,853,1589,1613]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="milleacantha">
<emphasis id="58D5EAB328527F1DF570F99AB141E906" box="[583,853,1589,1613]" italics="true" pageId="13" pageNumber="14">Abbottella milleacantha</emphasis>
</taxonomicName>
belongs to a group of small species having a dense, minute sculpture of pustules or points. It does not have the sculpture arranged in obvious keels as in
<taxonomicName id="ADA14D2228527F1DF223F9F6B761E93A" box="[1300,1397,1625,1649]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="gabbi">
<emphasis id="58D5EAB328527F1DF223F9F6B761E93A" box="[1300,1397,1625,1649]" italics="true" pageId="13" pageNumber="14">A. gabbi</emphasis>
</taxonomicName>
,
<taxonomicName id="ADA14D2228527F1DF2B3F9F6B2E6E9DE" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="rosaliae">
<emphasis id="58D5EAB328527F1DF2B3F9F6B2E6E9DE" italics="true" pageId="13" pageNumber="14">A. rosaliae</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428527F1DF637F9D2B38AE9DE" box="[256,414,1661,1685]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="13" pageNumber="14">Figure 4 A–E</figureCitation>
),
<taxonomicName id="ADA14D2228527F1DF686F9D2B00BE9DE" box="[433,543,1661,1685]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="abbotti">
<emphasis id="58D5EAB328527F1DF686F9D2B00BE9DE" box="[433,543,1661,1685]" italics="true" pageId="13" pageNumber="14">A. abbotti</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428527F1DF519F9D2B0DFE9DE" box="[558,715,1661,1685]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="13" pageNumber="14">Figure 3 A–E</figureCitation>
), or
<taxonomicName id="ADA14D2228527F1DF5CAF9D2B17EE9DE" box="[765,874,1661,1685]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="crossei">
<emphasis id="58D5EAB328527F1DF5CAF9D2B17EE9DE" box="[765,874,1661,1685]" italics="true" pageId="13" pageNumber="14">A. crossei</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428527F1DF44FF9D2B1CCE9DE" box="[888,984,1661,1685]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="13" pageNumber="14">Figure 3</figureCitation>
S–W). The sculpture is finer and denser (ca. 120 axial threads on final whorl) than in
<taxonomicName id="ADA14D2228527F1DF592F90EB127E9F2" box="[677,819,1697,1721]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="tentorium">
<emphasis id="58D5EAB328527F1DF592F90EB127E9F2" box="[677,819,1697,1721]" italics="true" pageId="13" pageNumber="14">A. tentorium</emphasis>
</taxonomicName>
(ca. 60 axial threads),
<taxonomicName id="ADA14D2228527F1DF309F90EB6B6E9F2" box="[1086,1186,1697,1721]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="aenea">
<emphasis id="58D5EAB328527F1DF309F90EB6B6E9F2" box="[1086,1186,1697,1721]" italics="true" pageId="13" pageNumber="14">A. aenea</emphasis>
</taxonomicName>
(ca. 90 axial threads;
<figureCitation id="F29A2A2428527F1DF7A0F96AB2E2E996" box="[151,246,1733,1757]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="13" pageNumber="14">Figure 3</figureCitation>
N–R), or
<taxonomicName id="ADA14D2228527F1DF65BF96AB3CAE996" box="[364,478,1733,1757]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="urbana">
<emphasis id="58D5EAB328527F1DF65BF96AB3CAE996" box="[364,478,1733,1757]" italics="true" pageId="13" pageNumber="14">A. urbana</emphasis>
</taxonomicName>
(ca. 70 axial threads). From
<taxonomicName id="ADA14D2228527F1DF41DF96AB199E996" box="[810,909,1733,1757]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="aenea">
<emphasis id="58D5EAB328527F1DF41DF96AB199E996" box="[810,909,1733,1757]" italics="true" pageId="13" pageNumber="14">A. aenea</emphasis>
</taxonomicName>
it also differs in color (copper in
<taxonomicName id="ADA14D2228527F1DF227F96AB767E996" box="[1296,1395,1733,1757]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="aenea">
<emphasis id="58D5EAB328527F1DF227F96AB767E996" box="[1296,1395,1733,1757]" italics="true" pageId="13" pageNumber="14">A. aenea</emphasis>
</taxonomicName>
vs. white in
<taxonomicName id="ADA14D2228527F1DF7CBF946B3BAE84A" box="[252,430,1769,1793]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="milleacantha">
<emphasis id="58D5EAB328527F1DF7CBF946B3BAE84A" box="[252,430,1769,1793]" italics="true" pageId="13" pageNumber="14">A. milleacantha</emphasis>
</taxonomicName>
) and in the form of the sculpture, which arises as separate points from the surface in
<taxonomicName id="ADA14D2228527F1DF2B3F946B2CEE86E" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="aenea">
<emphasis id="58D5EAB328527F1DF2B3F946B2CEE86E" italics="true" pageId="13" pageNumber="14">A. aenea</emphasis>
</taxonomicName>
but arises from the spiral threads in
<taxonomicName id="ADA14D2228527F1DF544F8A2B130E86E" box="[627,804,1805,1829]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="milleacantha">
<emphasis id="58D5EAB328527F1DF544F8A2B130E86E" box="[627,804,1805,1829]" italics="true" pageId="13" pageNumber="14">A. milleacantha</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="E20887B728527F1DF7A0FF38B1CAEC7D" ID-DOI="http://doi.org/10.5281/zenodo.6164936" ID-GBIF-Taxon="127664010" ID-Zenodo-Dep="6164936" LSID="urn:lsid:plazi:treatment:E20887B728527F1DF7A0FF38B1CAEC7D" httpUri="http://treatment.plazi.org/id/E20887B728527F1DF7A0FF38B1CAEC7D" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="22BB652A28527F1DF7A0FF38B367EF99" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="6A1E36A128527F1DF7A0FF38B0ABEFFA" blockId="13.[151,703,151,210]" box="[151,703,151,177]" pageId="13" pageNumber="14">
<heading id="315681CD28527F1DF7A0FF38B0ABEFFA" bold="true" box="[151,703,151,177]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="ADA14D2228527F1DF7A0FF38B0ABEFFA" authority="Pilsbry, 1933" authorityName="Pilsbry" authorityYear="1933" box="[151,703,151,177]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="crossei" subGenus="Abbottella">
<emphasis id="58D5EAB328527F1DF7A0FF38B0ABEFFA" bold="true" box="[151,703,151,177]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7A0FF38B307EFFA" bold="true" box="[151,275,151,177]" italics="true" pageId="13" pageNumber="14">Abbottella</emphasis>
(
<emphasis id="58D5EAB328527F1DF613FF38B3B4EFFA" bold="true" box="[292,416,151,177]" italics="true" pageId="13" pageNumber="14">Abbottella</emphasis>
)
<emphasis id="58D5EAB328527F1DF687FF38B017EFFA" bold="true" box="[432,515,151,177]" italics="true" pageId="13" pageNumber="14">crossei</emphasis>
(Pilsbry, 1933)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A128527F1DF7A0FF15B367EF99" blockId="13.[151,703,151,210]" box="[151,371,186,210]" pageId="13" pageNumber="14">
<figureCitation id="F29A2A2428527F1DF7A0FF15B314EF99" box="[151,256,186,210]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="13" pageNumber="14">Figures 3</figureCitation>
S–W, 7 G
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28527F1DF7A0FEACB393ED5D" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="6A1E36A128527F1DF7A0FEACB17DEE50" blockId="13.[151,1436,258,822]" box="[151,873,258,283]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7A0FEACB351EE50" bold="true" box="[151,325,258,283]" pageId="13" pageNumber="14">
<typeStatus id="B51A880328527F1DF7A0FEACB2C6EE50" box="[151,210,259,283]" pageId="13" pageNumber="14">Type</typeStatus>
material.
</emphasis>
ANSP 7951,
<typeStatus id="B51A880328527F1DF6D7FEADB052EE51" box="[480,582,258,282]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
. ANSP 373769,
<typeStatus id="B51A880328527F1DF436FEACB171EE50" box="[769,869,259,283]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="6A1E36A128527F1DF7FEFE88B133EE29" blockId="13.[151,1436,258,822]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7FEFE88B373EE74" bold="true" box="[201,359,294,319]" pageId="13" pageNumber="14">
<typeStatus id="B51A880328527F1DF7FEFE88B316EE74" box="[201,258,295,319]" pageId="13" pageNumber="14">Type</typeStatus>
locality.
</emphasis>
Santo
<collectingCountry id="12B6763128527F1DF68CFE89B03CEE75" box="[443,552,294,318]" name="Dominican Republic" pageId="13" pageNumber="14">Domingo</collectingCountry>
. Bartsch (1946: 133) believed Pilsbry’s specimen came from “somewhere on the Samaná Peninsula,” which is shown here to be correct.
</paragraph>
<paragraph id="6A1E36A128527F1DF7FEFEC1B393ED5D" blockId="13.[151,1436,258,822]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7FEFEC1B013EECC" bold="true" box="[201,519,366,391]" pageId="13" pageNumber="14">Other material examined.</emphasis>
UF 216146, 4 km E of Samaná, Samaná Province,
<collectingCountry id="12B6763128527F1DF34FFEC1B77CEECD" box="[1144,1384,366,390]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 216134, 8 km S of Las Galeras, Samaná Province,
<collectingCountry id="12B6763128527F1DF5DDFE3DB1CDEEE1" box="[746,985,402,426]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 216139,
<quantity id="AD599B4428527F1DF34EFE3DB6DDEEE1" box="[1145,1225,402,426]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="13" pageNumber="14" unit="km" value="2.9">2.9 km</quantity>
S of Las Galeras, Samaná Province,
<collectingCountry id="12B6763128527F1DF645FE19B077EE85" box="[370,611,438,462]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; UF 216128,
<quantity id="AD599B4428527F1DF43DFE19B149EE85" box="[778,861,438,462]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="13" pageNumber="14" unit="km" value="6.5">6.5 km</quantity>
S of Las Galeras, Samaná Province,
<collectingCountry id="12B6763128527F1DF217FE19B314EEB9" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
; GTW 9432a, limestone area, at side of road to Cape Samaná,
<quantity id="AD599B4428527F1DF4ECFE75B63FEEB9" box="[987,1067,474,498]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="14" unit="km" value="0.6">0.6 km</quantity>
before Cape, Samaná Province,
<collectingCountry id="12B6763128527F1DF7A0FE51B397ED5D" box="[151,387,510,534]" name="Dominican Republic" pageId="13" pageNumber="14">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28527F1DF7FEFD8DB1CAEC7D" pageId="13" pageNumber="14" type="distribution">
<paragraph id="6A1E36A128527F1DF7FEFD8DB609ED81" blockId="13.[151,1436,258,822]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7FEFD8DB3EDED70" bold="true" box="[201,505,546,571]" pageId="13" pageNumber="14">Distribution and habitat.</emphasis>
This species is found along the coastal limestone bluff on the eastern tip of the Samaná Peninsula in the Sierra de Samaná of the Cordillera Septentrional, possibly to Sánchez. Bartsch (1946) added localities at
<quantity id="AD599B4428527F1DF65FFDC5B3A2EDC9" box="[360,438,618,642]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="14" unit="km" value="0.8">0.8 km</quantity>
from the sea,
<quantity id="AD599B4428527F1DF561FDC5B0B0EDC9" box="[598,676,618,642]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="13" pageNumber="14" unit="km" value="2.4">2.4 km</quantity>
E of Puerto Francés, Samaná Province and at a cave
<quantity id="AD599B4428527F1DF3CDFDC5B75CEDC9" box="[1274,1352,618,642]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="14" unit="km" value="3.2">3.2 km</quantity>
NW of Sánchez, Samaná Province. I have not seen the Sánchez specimens, which seem to be out of the range of this species. Specimens have been found at the base of limestone outcrops and cliffs.
</paragraph>
<paragraph id="6A1E36A128527F1DF7FEFD79B1CAEC7D" blockId="13.[151,1436,258,822]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7FEFD79B055EDA4" bold="true" box="[201,577,726,751]" pageId="13" pageNumber="14">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D2228527F1DF57EFD79B107EDA5" box="[585,787,726,750]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="crossei">
<emphasis id="58D5EAB328527F1DF57EFD79B107EDA5" box="[585,787,726,750]" italics="true" pageId="13" pageNumber="14">Abbottella crossei</emphasis>
</taxonomicName>
belongs to the group of small, heavily sculptured taxa on the Samaná Peninsula. It differs in having finer sculpture and less developed keels. It never has the fimbriated lip usually found in
<taxonomicName id="ADA14D2228527F1DF663FCB1B3DAEC7D" box="[340,462,798,822]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="rosaliae">
<emphasis id="58D5EAB328527F1DF663FCB1B3DAEC7D" box="[340,462,798,822]" italics="true" pageId="13" pageNumber="14">A. rosaliae</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428527F1DF6EBFCB1B063EC7D" box="[476,631,798,822]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="13" pageNumber="14">Figure 4 A–E</figureCitation>
) and
<taxonomicName id="ADA14D2228527F1DF581FCB1B130EC7D" box="[694,804,798,822]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="abbotti">
<emphasis id="58D5EAB328527F1DF581FCB1B130EC7D" box="[694,804,798,822]" italics="true" pageId="13" pageNumber="14">A. abbotti</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428527F1DF404FCB1B1C7EC7D" box="[819,979,798,822]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="13" pageNumber="14">Figure 3 A–E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="084562FCF98DA36B75284520366E3D27">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B728527F1FF7A0F8C1B2FDED77" ID-DOI="http://doi.org/10.5281/zenodo.6164940" ID-GBIF-Taxon="127664011" ID-Zenodo-Dep="6164940" LSID="urn:lsid:plazi:treatment:E20887B728527F1FF7A0F8C1B2FDED77" httpUri="http://treatment.plazi.org/id/E20887B728527F1FF7A0F8C1B2FDED77" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="22BB652A28527F1DF7A0F8C1B365E8E1" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="6A1E36A128527F1DF7A0F8C1B0C4E8C3" blockId="13.[151,720,1902,1962]" box="[151,720,1902,1928]" pageId="13" pageNumber="14">
<heading id="315681CD28527F1DF7A0F8C1B0C4E8C3" bold="true" box="[151,720,1902,1928]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="ADA14D2228527F1DF7A0F8C1B0C4E8C3" authority="Pfeiffer, 1858" authorityName="Pfeiffer" authorityYear="1858" box="[151,720,1902,1928]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="rosaliae" subGenus="Abbottella">
<emphasis id="58D5EAB328527F1DF7A0F8C1B0C4E8C3" bold="true" box="[151,720,1902,1928]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7A0F8C1B307E8C3" bold="true" box="[151,275,1902,1928]" italics="true" pageId="13" pageNumber="14">Abbottella</emphasis>
(
<emphasis id="58D5EAB328527F1DF613F8C1B3B4E8C3" bold="true" box="[292,416,1902,1928]" italics="true" pageId="13" pageNumber="14">Abbottella</emphasis>
)
<emphasis id="58D5EAB328527F1DF687F8C1B004E8C3" bold="true" box="[432,528,1902,1928]" italics="true" pageId="13" pageNumber="14">rosaliae</emphasis>
(Pfeiffer, 1858)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A128527F1DF7A0F83EB365E8E1" blockId="13.[151,720,1902,1962]" box="[151,369,1937,1962]" pageId="13" pageNumber="14">
<figureCitation id="F29A2A2428527F1DF7A0F83EB329E8E1" box="[151,317,1937,1962]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="13" pageNumber="14">Figures 4 A–E</figureCitation>
,
<figureCitation id="F29A2A2428527F1DF67FF83EB342E8E2" box="[328,342,1937,1961]" captionStart="FIGURE 9" captionStartId="20.[155,253,1910,1933]" captionText="FIGURE 9. Distributions are approximate. Base map courtesy of NASA. A. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). B. Abbottella (Abbottella) samanensis Bartsch, 1946. C. Abbottella (Abbottella) sanchezi Bartsch, 1946. D. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). E. Abbottella (Abbottella) sosuaensis Bartsch, 1946. F. Abbottella (Gundlachtudora) paradoxa new species." pageId="13" pageNumber="14">9</figureCitation>
A
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28527F1FF7A0F875B640EF9F" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="6A1E36A128527F1DF7A0F875B3C1E8BA" blockId="13.[151,469,2009,2034]" box="[151,469,2009,2034]" pageId="13" pageNumber="14">
<emphasis id="58D5EAB328527F1DF7A0F875B351E8B9" bold="true" box="[151,325,2009,2034]" pageId="13" pageNumber="14">
<typeStatus id="B51A880328527F1DF7A0F875B2C6E8B9" box="[151,210,2010,2034]" pageId="13" pageNumber="14">Type</typeStatus>
material.
</emphasis>
Not located.
</paragraph>
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<paragraph id="6A1E36A128517F1EF7A0F9ECB338E87B" blockId="14.[151,1437,1603,1841]" pageId="14" pageNumber="15">
<emphasis id="58D5EAB328517F1EF7A0F9ECB300E912" bold="true" box="[151,276,1603,1626]" pageId="14" pageNumber="15">FIGURE 4.</emphasis>
A–E.
<taxonomicName id="ADA14D2228517F1EF660F9EBB150E911" authority="Pfeiffer, 1858" authorityName="Pfeiffer" authorityYear="1858" box="[343,836,1604,1626]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="rosaliae" subGenus="Abbottella">
<emphasis id="58D5EAB328517F1EF660F9EBB3D4E911" box="[343,448,1604,1626]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
(
<emphasis id="58D5EAB328517F1EF6E6F9EBB02EE911" box="[465,570,1604,1626]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
)
<emphasis id="58D5EAB328517F1EF57DF9EBB088E911" box="[586,668,1604,1626]" italics="true" pageId="14" pageNumber="15">rosaliae</emphasis>
(Pfeiffer, 1858)
</taxonomicName>
. A–C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F–I.
<taxonomicName id="ADA14D2228517F1EF576F9CCB628E933" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[577,1084,1634,1657]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="samanensis" subGenus="Abbottella">
<emphasis id="58D5EAB328517F1EF576F9CCB0BEE932" box="[577,682,1635,1657]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
(
<emphasis id="58D5EAB328517F1EF58EF9CCB136E932" box="[697,802,1635,1657]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
)
<emphasis id="58D5EAB328517F1EF406F9CBB1B2E932" box="[817,934,1636,1657]" italics="true" pageId="14" pageNumber="15">samanensis</emphasis>
Bartsch, 1946
</taxonomicName>
. F–H. GTW 7085c, 8.7 mm width. I. GTW 7085d, 8.0 mm width. J–N.
<taxonomicName id="ADA14D2228517F1EF525F92EB1E3E9DC" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[530,1015,1665,1687]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="sanchezi" subGenus="Abbottella">
<emphasis id="58D5EAB328517F1EF525F92EB06FE9DC" box="[530,635,1665,1687]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
(
<emphasis id="58D5EAB328517F1EF5BBF92EB0E1E9DC" box="[652,757,1665,1687]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
)
<emphasis id="58D5EAB328517F1EF431F92EB149E9DC" box="[774,861,1665,1687]" italics="true" pageId="14" pageNumber="15">sanchezi</emphasis>
Bartsch, 1946
</taxonomicName>
. J–L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O–S.
<taxonomicName id="ADA14D2228517F1EF41EF90FB738E9FD" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[809,1324,1696,1718]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="sosuaensis" subGenus="Abbottella">
<emphasis id="58D5EAB328517F1EF41EF90FB186E9FD" box="[809,914,1696,1718]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
(
<emphasis id="58D5EAB328517F1EF492F90FB61BE9FD" box="[933,1039,1696,1718]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
)
<emphasis id="58D5EAB328517F1EF316F90EB699E9FD" box="[1057,1165,1697,1718]" italics="true" pageId="14" pageNumber="15">sosuaensis</emphasis>
Bartsch, 1946
</taxonomicName>
. O–Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431a, 7.6 mm width. T–X.
<taxonomicName id="ADA14D2228517F1EF365F910B32FE9B8" authority="Pfeiffer, 1858" authorityName="Pfeiffer" authorityYear="1858" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="wilhelmi" subGenus="Abbottella">
<emphasis id="58D5EAB328517F1EF365F910B6AFE99E" box="[1106,1211,1727,1749]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
(
<emphasis id="58D5EAB328517F1EF3FCF910B720E99E" box="[1227,1332,1727,1749]" italics="true" pageId="14" pageNumber="15">Abbottella</emphasis>
)
<emphasis id="58D5EAB328517F1EF273F910B78FE99E" box="[1348,1435,1727,1749]" italics="true" pageId="14" pageNumber="15">wilhelmi</emphasis>
(Pfeiffer, 1858)
</taxonomicName>
. T–V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y–CC.
<taxonomicName id="ADA14D2228517F1EF7A0F952B012E859" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[151,518,1788,1810]" class="Gastropoda" family="Pomatiidae" genus="Lagopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="lagopoma">
<emphasis id="58D5EAB328517F1EF7A0F952B37BE859" box="[151,367,1788,1810]" italics="true" pageId="14" pageNumber="15">Lagopoma lagopoma</emphasis>
Bartsch, 1946
</taxonomicName>
. Y–AA. GTW 10081a, 9.7 mm width. BB. GTW 10081a, 7.6 mm width. CC. UF 216120, 7.7 mm width.
</paragraph>
</caption>
<paragraph id="6A1E36A128517F1EF7FEF8F7B2E8E8DF" blockId="14.[151,1436,1879,2012]" pageId="14" pageNumber="15">
<emphasis id="58D5EAB328517F1EF7FEF8F7B378E83B" bold="true" box="[201,364,1879,1904]" pageId="14" pageNumber="15">
<typeStatus id="B51A880328517F1EF7FEF8F7B310E83B" box="[201,260,1880,1904]" pageId="14" pageNumber="15">Type</typeStatus>
locality.
</emphasis>
<emphasis id="58D5EAB328517F1EF64FF8F6B025E83B" box="[376,561,1880,1904]" italics="true" pageId="14" pageNumber="15">
In insulae
<collectingCountry id="12B6763128517F1EF6CFF8F6B025E83B" box="[504,561,1881,1904]" name="Haiti" pageId="14" pageNumber="15">Haiti</collectingCountry>
</emphasis>
. Here restricted to
<collectingCountry id="12B6763128517F1EF42BF8F7B1D3E83B" box="[796,967,1880,1904]" name="Dominican Republic" pageId="14" pageNumber="15">the Dominican</collectingCountry>
Republic, Puerto Plata Province, Loma Catalina.
</paragraph>
<paragraph id="6A1E36A128517F1FF7FEF830B640EF9F" blockId="14.[151,1436,1879,2012]" lastBlockId="15.[151,1436,151,572]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="58D5EAB328517F1EF7FEF830B3EAE8F3" bold="true" box="[201,510,1951,1976]" pageId="14" pageNumber="15">Other material examined.</emphasis>
UF 216191, Loma Catalina,
<quantity id="AD599B4428517F1EF471F80FB16BE8F3" box="[838,895,1952,1976]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="km" value="6.0">6 km</quantity>
SE of Sosuá, Puerto Plata Province,
<collectingCountry id="12B6763128517F1EF228F80FB2EBE897" name="Dominican Republic" pageId="14" pageNumber="15">Dominican Republic</collectingCountry>
,
<quantity id="AD599B4428517F1EF63DF86BB352E897" box="[266,326,1988,2012]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="15" unit="m" value="80.0">80 m</quantity>
; UF 216171, Loma Catalina,
<quantity id="AD599B4428517F1EF5A4F86BB0D8E897" box="[659,716,1988,2012]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="15" unit="km" value="9.0">9 km</quantity>
SSE of Sosuá, Puerto Plata Province,
<collectingCountry id="12B6763128517F1EF34CF86BB772E897" box="[1147,1382,1988,2012]" name="Dominican Republic" pageId="14" pageNumber="15">Dominican Republic</collectingCountry>
,
<quantity id="AD599B4428517F1FF245F86BB2A4EFE4" lastPageId="15" lastPageNumber="16" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="15" unit="m" value="210.0">210 m</quantity>
; UF 216125, Loma Catalina,
<quantity id="AD599B4428507F1FF525FF38B048EFE4" box="[530,604,151,175]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="km" value="11.0">11 km</quantity>
SE of Sosuá, Puerto Plata Province,
<collectingCountry id="12B6763128507F1FF32DFF38B71EEFE4" box="[1050,1290,151,175]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
,
<quantity id="AD599B4428507F1FF22DFF38B77CEFE4" box="[1306,1384,151,175]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="16" unit="m" value="260.0">260 m</quantity>
; UF 216135, 7 km E,
<quantity id="AD599B4428507F1FF66FFF13B385EF9F" box="[344,401,188,212]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="km" value="2.0">2 km</quantity>
S of Sosuá, Puerto Plata Province,
<collectingCountry id="12B6763128507F1FF42AFF13B61DEF9F" box="[797,1033,188,212]" name="Dominican Republic" pageId="15" pageNumber="16">Dominican Republic</collectingCountry>
,
<specimenCount id="7CA7FD2828507F1FF323FF13B640EF9F" box="[1044,1108,188,212]" pageId="15" pageNumber="16" type="generic">
<quantity id="AD599B4428507F1FF323FF13B644EF9F" box="[1044,1104,188,212]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="16" unit="m" value="30.0">30 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28507F1FF7FEFF70B1F0EEE7" pageId="15" pageNumber="16" type="distribution">
<paragraph id="6A1E36A128507F1FF7FEFF70B0ACEE2F" blockId="15.[151,1436,151,572]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFF70B3ECEFB3" bold="true" box="[201,504,223,248]" pageId="15" pageNumber="16">Distribution and habitat.</emphasis>
Collections at UF enable us to limit Pfeiffer’s imprecise “
<emphasis id="58D5EAB328507F1FF3A9FF4EB747EFB3" box="[1182,1363,224,248]" italics="true" pageId="15" pageNumber="16">
In insulae
<collectingCountry id="12B6763128507F1FF22DFF4EB747EFB3" box="[1306,1363,225,248]" name="Haiti" pageId="15" pageNumber="16">Haiti</collectingCountry>
</emphasis>
.” The species is known only from Loma Catalina, an outcrop of the Cordillera Septentrional southeast of Sosuá. Much of this area is contained in the Parque Nacional El Choco. The
<typeStatus id="B51A880328507F1FF46BFE87B198EE0B" box="[860,908,296,320]" pageId="15" pageNumber="16">type</typeStatus>
locality is herein restricted to
<collectingCountry id="12B6763128507F1FF3C4FE88B78FEE74" box="[1267,1435,295,319]" name="Dominican Republic" pageId="15" pageNumber="16">the Dominican</collectingCountry>
Republic, Puerto Plata Province, Loma Catalina.
</paragraph>
<paragraph id="6A1E36A128507F1FF7FEFEC0B1F0EEE7" blockId="15.[151,1436,151,572]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFEC0B05CEEC3" bold="true" box="[201,584,367,392]" pageId="15" pageNumber="16">Comparison with other species.</emphasis>
This species is similar to
<taxonomicName id="ADA14D2228507F1FF44CFEDFB1F8EEC3" box="[891,1004,368,392]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="abbotti">
<emphasis id="58D5EAB328507F1FF44CFEDFB1F8EEC3" box="[891,1004,368,392]" italics="true" pageId="15" pageNumber="16">A. abbotti</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428507F1FF4C9FEC0B6B4EECC" box="[1022,1184,367,391]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="15" pageNumber="16">Figure 3 A–E</figureCitation>
) but differs in having finer, less serrate sculpture, less developed keels, and a less fluted outer lip.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28507F1FF7FEFE17B2FDED77" pageId="15" pageNumber="16" type="discussion">
<paragraph id="6A1E36A128507F1FF7FEFE17B2FDED77" blockId="15.[151,1436,151,572]" pageId="15" pageNumber="16">
<emphasis id="58D5EAB328507F1FF7FEFE17B329EE9B" bold="true" box="[201,317,440,464]" pageId="15" pageNumber="16">Remarks.</emphasis>
This species is perhaps unique among
<taxonomicName id="ADA14D2228507F1FF5C2FE17B17EEE9B" box="[757,874,440,464]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328507F1FF5C2FE17B17EEE9B" box="[757,874,440,464]" italics="true" pageId="15" pageNumber="16">Abbottella</emphasis>
</taxonomicName>
in possessing a periostracum although it is lost in most specimens. On fresh examples this periostracum forms short bristles at the junctures of the axial and spiral sculpture. The degree of fluting and serration of the outer lip varies greatly—some specimens show virtually no fluting.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="E20887B728537F1CF7A0F99AB6D1E827" ID-DOI="http://doi.org/10.5281/zenodo.6164934" ID-GBIF-Taxon="127664008" ID-Zenodo-Dep="6164934" LSID="urn:lsid:plazi:treatment:E20887B728537F1CF7A0F99AB6D1E827" httpUri="http://treatment.plazi.org/id/E20887B728537F1CF7A0F99AB6D1E827" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection id="22BB652A28537F1CF7A0F99AB366E924" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="6A1E36A128537F1CF7A0F99AB0B8E904" blockId="12.[151,684,1589,1647]" box="[151,684,1589,1615]" pageId="12" pageNumber="13">
<heading id="315681CD28537F1CF7A0F99AB0B8E904" bold="true" box="[151,684,1589,1615]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="ADA14D2228537F1CF7A0F99AB0B8E904" authority="Watters, 2010" authorityName="Watters" authorityYear="2010" box="[151,684,1589,1615]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="aenea" subGenus="Abbottella">
<emphasis id="58D5EAB328537F1CF7A0F99AB0B8E904" bold="true" box="[151,684,1589,1615]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7A0F99AB307E904" bold="true" box="[151,275,1589,1615]" italics="true" pageId="12" pageNumber="13">Abbottella</emphasis>
(
<emphasis id="58D5EAB328537F1CF613F99AB3B4E904" bold="true" box="[292,416,1589,1615]" italics="true" pageId="12" pageNumber="13">Abbottella</emphasis>
)
<emphasis id="58D5EAB328537F1CF687F99AB3ECE904" bold="true" box="[432,504,1589,1615]" italics="true" pageId="12" pageNumber="13">aenea</emphasis>
Watters, 2010
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A128537F1CF7A0F9F8B366E924" blockId="12.[151,684,1589,1647]" box="[151,370,1623,1647]" pageId="12" pageNumber="13">
<figureCitation id="F29A2A2428537F1CF7A0F9F8B314E924" box="[151,256,1623,1647]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="12" pageNumber="13">Figures 3</figureCitation>
N–R, 8 C
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28537F1CF7A0F90FB1B7E86F" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="6A1E36A128537F1CF7A0F90FB31EE996" blockId="12.[151,1436,1695,1900]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7A0F90FB35CE9F3" bold="true" box="[151,328,1695,1720]" pageId="12" pageNumber="13">
<typeStatus id="B51A880328537F1CF7A0F90FB2C4E9F3" box="[151,208,1696,1720]" pageId="12" pageNumber="13">Type</typeStatus>
material.
</emphasis>
UF 434777,
<typeStatus id="B51A880328537F1CF6DBF930B046E9FC" box="[492,594,1695,1719]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
. UF 434778, 1
<typeStatus id="B51A880328537F1CF425F90FB162E9F3" box="[786,886,1696,1720]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
; BMSM 17971, 2
<typeStatus id="B51A880328537F1CF36EF90FB6DCE9F3" box="[1113,1224,1696,1720]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
; OSUM 35490, 2
<typeStatus id="B51A880328537F1CF7A0F96AB312E996" box="[151,262,1733,1757]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
.
</paragraph>
<paragraph id="6A1E36A128537F1CF7FEF947B1EAE9B4" blockId="12.[151,1436,1695,1900]" box="[201,1022,1767,1792]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7FEF947B371E84B" bold="true" box="[201,357,1767,1792]" pageId="12" pageNumber="13">
<typeStatus id="B51A880328537F1CF7FEF947B316E84B" box="[201,258,1768,1792]" pageId="12" pageNumber="13">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B6763128537F1CF659F948B04DE9B4" box="[366,601,1767,1791]" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
, La Altagracia Province, Punta Cana.
</paragraph>
<paragraph id="6A1E36A128537F1CF7FEF8A4B1B7E86F" blockId="12.[151,1436,1695,1900]" box="[201,931,1803,1829]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7FEF8A4B3EAE86F" bold="true" box="[201,510,1803,1828]" pageId="12" pageNumber="13">Other material examined.</emphasis>
GTW 14181a, from the
<typeStatus id="B51A880328537F1CF425F8A2B156E86E" box="[786,834,1805,1829]" pageId="12" pageNumber="13">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28537F1CF7FEF880B786E80C" box="[201,1426,1839,1865]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="6A1E36A128537F1CF7FEF880B786E80C" blockId="12.[151,1436,1695,1900]" box="[201,1426,1839,1865]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7FEF880B3E7E803" bold="true" box="[201,499,1839,1864]" pageId="12" pageNumber="13">Distribution and habitat.</emphasis>
Known only from the
<typeStatus id="B51A880328537F1CF5C4F89EB137E802" box="[755,803,1841,1865]" pageId="12" pageNumber="13">type</typeStatus>
locality. Specimens were found under moldy leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28537F1CF7FEF8FBB6D1E827" box="[201,1221,1876,1900]" pageId="12" pageNumber="13" type="discussion">
<paragraph id="6A1E36A128537F1CF7FEF8FBB6D1E827" blockId="12.[151,1436,1695,1900]" box="[201,1221,1876,1900]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7FEF8FBB329E827" bold="true" box="[201,317,1876,1900]" pageId="12" pageNumber="13">Remarks.</emphasis>
This species is included here for comparison. See under
<taxonomicName id="ADA14D2228537F1CF48CF8FBB678E827" box="[955,1132,1876,1900]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="milleacantha">
<emphasis id="58D5EAB328537F1CF48CF8FBB678E827" box="[955,1132,1876,1900]" italics="true" pageId="12" pageNumber="13">A. milleacantha</emphasis>
</taxonomicName>
, below.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E214A4CBBAE23CA9927AA325FD980C32">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B728537F1CF7A0FCB1B115EAAA" ID-DOI="http://doi.org/10.5281/zenodo.6164932" ID-GBIF-Taxon="127664009" ID-Zenodo-Dep="6164932" LSID="urn:lsid:plazi:treatment:E20887B728537F1CF7A0FCB1B115EAAA" httpUri="http://treatment.plazi.org/id/E20887B728537F1CF7A0FCB1B115EAAA" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection id="22BB652A28537F1CF7A0FCB1B37CEC12" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="6A1E36A128537F1CF7A0FCB1B151EC73" blockId="12.[151,837,798,857]" box="[151,837,798,824]" pageId="12" pageNumber="13">
<heading id="315681CD28537F1CF7A0FCB1B151EC73" bold="true" box="[151,837,798,824]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="ADA14D2228537F1CF7A0FCB1B151EC73" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[151,837,798,824]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="adolfi" subGenus="Abbottella" subSpecies="peninsularis">
<emphasis id="58D5EAB328537F1CF7A0FCB1B151EC73" bold="true" box="[151,837,798,824]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7A0FCB1B307EC73" bold="true" box="[151,275,798,824]" italics="true" pageId="12" pageNumber="13">Abbottella</emphasis>
(
<emphasis id="58D5EAB328537F1CF613FCB1B3B4EC73" bold="true" box="[292,416,798,824]" italics="true" pageId="12" pageNumber="13">Abbottella</emphasis>
)
<emphasis id="58D5EAB328537F1CF687FCB1B080EC73" bold="true" box="[432,660,798,824]" italics="true" pageId="12" pageNumber="13">adolfi peninsularis</emphasis>
Bartsch, 1946
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A128537F1CF7A0FCEEB37CEC12" blockId="12.[151,837,798,857]" box="[151,360,833,857]" pageId="12" pageNumber="13">
<figureCitation id="F29A2A2428537F1CF7A0FCEEB314EC12" box="[151,256,833,857]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="12" pageNumber="13">Figures 3</figureCitation>
J–L, 8 H
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28537F1CF7A0FC25B701EBD6" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="6A1E36A128537F1CF7A0FC25B07BECEA" blockId="12.[151,1436,905,1505]" box="[151,623,905,930]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7A0FC25B351ECE9" bold="true" box="[151,325,905,930]" pageId="12" pageNumber="13">
<typeStatus id="B51A880328537F1CF7A0FC25B2C6ECE9" box="[151,210,906,930]" pageId="12" pageNumber="13">Type</typeStatus>
material.
</emphasis>
USNM 504095,
<typeStatus id="B51A880328537F1CF532FC26B07FECEA" box="[517,619,905,929]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="6A1E36A128537F1CF7FEFC02B630EC8E" blockId="12.[151,1436,905,1505]" box="[201,1060,941,966]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7FEFC02B371EC8D" bold="true" box="[201,357,941,966]" pageId="12" pageNumber="13">
<typeStatus id="B51A880328537F1CF7FEFC02B316EC8E" box="[201,258,941,965]" pageId="12" pageNumber="13">Type</typeStatus>
locality.
</emphasis>
On the trail from Samaná to Rio San Juan, Samaná Peninsula.
</paragraph>
<paragraph id="6A1E36A128537F1CF7FEFC7EB701EBD6" blockId="12.[151,1436,905,1505]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7FEFC7EB012ECA1" bold="true" box="[201,518,977,1002]" pageId="12" pageNumber="13">Other material examined.</emphasis>
UF 216164, 8 km W of Samaná, Samaná Province,
<collectingCountry id="12B6763128537F1CF34EFC7EB77EECA2" box="[1145,1386,977,1001]" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
; UF 215974, 10 km E of Las Terrenas, Samaná Province,
<collectingCountry id="12B6763128537F1CF5DAFC5AB1C1EB46" box="[749,981,1013,1037]" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
at sea level; UF 215975, 8 km WNW of Sánchez, Samaná Province,
<collectingCountry id="12B6763128537F1CF6E2FBB6B0D6EB7A" box="[469,706,1049,1073]" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
; UF 215976, 11 km E of Sánchez, Samaná Province,
<collectingCountry id="12B6763128537F1CF217FBB6B314EB1E" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
; UF 215979a, b,
<quantity id="AD599B4428537F1CF6F4FB91B3E8EB1E" box="[451,508,1085,1109]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="km" value="7.0">7 km</quantity>
E of Sánchez, Samaná Province,
<collectingCountry id="12B6763128537F1CF44CFB92B673EB1E" box="[891,1127,1085,1109]" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
; UF 215978a, b,
<quantity id="AD599B4428537F1CF21CFB92B771EB1E" box="[1323,1381,1085,1109]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="km" value="5.0">5 km</quantity>
E of Sánchez, Samaná Province,
<collectingCountry id="12B6763128537F1CF6D6FBCEB0C6EB32" box="[481,722,1121,1145]" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
; UF 215977, 10 km SW of Las Terrenas, Samaná Province,
<collectingCountry id="12B6763128537F1CF7A0FB2AB397EBD6" box="[151,387,1157,1181]" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
; GTW 9430a, several km W of Sánchez, Samaná Province,
<collectingCountry id="12B6763128537F1CF313FB2AB705EBD6" box="[1060,1297,1157,1181]" name="Dominican Republic" pageId="12" pageNumber="13">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28537F1CF7FEFB06B115EAAA" pageId="12" pageNumber="13" type="distribution">
<paragraph id="6A1E36A128537F1CF7FEFB06B007EA65" blockId="12.[151,1436,905,1505]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7FEFB06B3E1EB89" bold="true" box="[201,501,1193,1218]" pageId="12" pageNumber="13">Distribution and habitat.</emphasis>
Records place this species closer to Sánchez than to Rio San Juan in the Sierra de Samaná of the Cordillera Septentrional; also on the north coast of the peninsula near Las Terrenas. Specimens have been found at the base of limestone mogotes and chimneys in lowland rain forests on damp, algae-covered rocks; some areas are now cacao groves.
</paragraph>
<paragraph id="6A1E36A128537F1CF7FEFA96B115EAAA" blockId="12.[151,1436,905,1505]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7FEFA96B058EA19" bold="true" box="[201,588,1337,1362]" pageId="12" pageNumber="13">Comparison with other species.</emphasis>
This subspecies is very similar to
<taxonomicName id="ADA14D2228537F1CF4DDFA96B66AEA1A" box="[1002,1150,1337,1361]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="newcombi">
<emphasis id="58D5EAB328537F1CF4DDFA96B66AEA1A" box="[1002,1150,1337,1361]" italics="true" pageId="12" pageNumber="13">A. newcombi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428537F1CF3A7FA96B6E0EA1A" box="[1168,1268,1337,1361]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="12" pageNumber="13">Figure 3</figureCitation>
M), a species described without an exact locality. Both species have two prominent spiral cords in the umbilicus, nearly identical sculpture, and similar dark coloration.
<taxonomicName id="ADA14D2228537F1CF561FA2EB152EAD2" box="[598,838,1409,1433]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="newcombi">
<emphasis id="58D5EAB328537F1CF561FA2EB152EAD2" box="[598,838,1409,1433]" italics="true" pageId="12" pageNumber="13">Abbottella newcombi</emphasis>
</taxonomicName>
has a lower spire and less developed outer lip than does
<taxonomicName id="ADA14D2228537F1CF7E4FA0AB38EEAF6" box="[211,410,1445,1469]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="adolfi" subSpecies="peninsularis">
<emphasis id="58D5EAB328537F1CF7E4FA0AB38EEAF6" box="[211,410,1445,1469]" italics="true" pageId="12" pageNumber="13">A. a. peninsularis</emphasis>
</taxonomicName>
, but additional material may indicate that the two taxa are not distinct from each other, in which case the name
<taxonomicName id="ADA14D2228537F1CF6B1FA66B003EAAA" box="[390,535,1481,1505]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="newcombi">
<emphasis id="58D5EAB328537F1CF6B1FA66B003EAAA" box="[390,535,1481,1505]" italics="true" pageId="12" pageNumber="13">A. newcombi</emphasis>
</taxonomicName>
would have priority.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="37A1093AF3B562433B7361822B6A6667">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B728537F1CF7A0FF38B746ED81" ID-DOI="http://doi.org/10.5281/zenodo.6164930" ID-GBIF-Taxon="127664006" ID-Zenodo-Dep="6164930" LSID="urn:lsid:plazi:treatment:E20887B728537F1CF7A0FF38B746ED81" httpUri="http://treatment.plazi.org/id/E20887B728537F1CF7A0FF38B746ED81" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection id="22BB652A28537F1CF7A0FF38B371EF99" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="6A1E36A128537F1CF7A0FF38B13CEFFA" blockId="12.[151,808,151,210]" box="[151,808,151,177]" pageId="12" pageNumber="13">
<heading id="315681CD28537F1CF7A0FF38B13CEFFA" bold="true" box="[151,808,151,177]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="58D5EAB328537F1CF7A0FF38B13CEFFA" bold="true" box="[151,808,151,177]" pageId="12" pageNumber="13">
<taxonomicName id="ADA14D2228537F1CF7A0FF38B008EFFA" box="[151,540,151,177]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="adolfi" subGenus="Abbottella">
<emphasis id="58D5EAB328537F1CF7A0FF38B307EFFA" bold="true" box="[151,275,151,177]" italics="true" pageId="12" pageNumber="13">Abbottella</emphasis>
(
<emphasis id="58D5EAB328537F1CF613FF38B3B4EFFA" bold="true" box="[292,416,151,177]" italics="true" pageId="12" pageNumber="13">Abbottella</emphasis>
)
<emphasis id="58D5EAB328537F1CF687FF38B008EFFA" bold="true" box="[432,540,151,177]" italics="true" pageId="12" pageNumber="13">cf. adolfi</emphasis>
</taxonomicName>
<taxonomicName id="ADA14D2228537F1CF514FF38B13CEFFA" authority="Pfeiffer, 1852" authorityName="Pfeiffer" authorityYear="1852" box="[547,808,151,177]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="adolfi">adolfi (Pfeiffer, 1852)</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A128537F1CF7A0FF15B371EF99" blockId="12.[151,808,151,210]" box="[151,357,186,210]" pageId="12" pageNumber="13">
<figureCitation id="F29A2A2428537F1CF7A0FF15B314EF99" box="[151,256,186,210]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="12" pageNumber="13">Figures 3</figureCitation>
F–I, 9 D
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28537F1CF7A0FEACB746ED81" pageId="12" pageNumber="13" type="discussion">
<paragraph id="6A1E36A128537F1CF7A0FEACB746ED81" blockId="12.[151,1437,258,715]" pageId="12" pageNumber="13">
<emphasis id="58D5EAB328537F1CF7A0FEACB31FEE50" bold="true" box="[151,267,259,283]" pageId="12" pageNumber="13">Remarks.</emphasis>
The
<typeStatus id="B51A880328537F1CF673FEACB360EE50" box="[324,372,259,283]" pageId="12" pageNumber="13">type</typeStatus>
of
<taxonomicName id="ADA14D2228537F1CF6ADFEACB06CEE50" box="[410,632,259,283]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="adolfi">
<emphasis id="58D5EAB328537F1CF6ADFEACB06CEE50" box="[410,632,259,283]" italics="true" pageId="12" pageNumber="13">Choanopoma adolfi</emphasis>
</taxonomicName>
collected by Sallé has apparently been lost. Pfeiffer illustrated it (1854: pl. 48, figs. 5-8); figures 5–7 are only outline drawings but figure 8 is a color figure (refigured here as
<figureCitation id="F29A2A2428537F1CF220FE89B761EE75" box="[1303,1397,294,318]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="12" pageNumber="13">Figure 3</figureCitation>
F). Bartsch (1946) had no material he could refer to this species and reproduced Pfeiffer’s translated description and his figure 8. As Bartsch noted, Pfeiffer’s
<typeStatus id="B51A880328537F1CF559FEC0B08AEECC" box="[622,670,367,391]" pageId="12" pageNumber="13">type</typeStatus>
locality of
<collectingCountry id="12B6763128537F1CF413FEC1B148EECD" box="[804,860,366,390]" name="Haiti" pageId="12" pageNumber="13">Haiti</collectingCountry>
must refer to
<collectingCountry id="12B6763128537F1CF4C8FEC1B6B2EECD" box="[1023,1190,366,390]" name="Dominican Republic" pageId="12" pageNumber="13">the Dominican</collectingCountry>
Republic, which was known as
<collectingCountry id="12B6763128537F1CF63FFE3DB354EEE1" box="[264,320,402,426]" name="Haiti" pageId="12" pageNumber="13">Haiti</collectingCountry>
at the time, because Sallé did not collect in
<collectingCountry id="12B6763128537F1CF41BFE3DB170EEE1" box="[812,868,402,426]" name="Haiti" pageId="12" pageNumber="13">Haiti</collectingCountry>
proper. The taxon shown here seems to best fit the description and figure of
<taxonomicName id="ADA14D2228537F1CF683FE18B003EE85" box="[436,535,438,462]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="adolfi">
<emphasis id="58D5EAB328537F1CF683FE18B003EE85" box="[436,535,438,462]" italics="true" pageId="12" pageNumber="13">C. adolfi</emphasis>
</taxonomicName>
of all the
<taxonomicName id="ADA14D2228537F1CF5BDFE19B0EBEE85" box="[650,767,438,462]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328537F1CF5BDFE19B0EBEE85" box="[650,767,438,462]" italics="true" pageId="12" pageNumber="13">Abbottella</emphasis>
</taxonomicName>
I have examined. It is from the Rio San Juan area in María
<collectingCountry id="12B6763128537F1CF7A0FE75B2E2EEB9" box="[151,246,474,498]" name="Trinidad and Tobago" pageId="12" pageNumber="13">Trinidad</collectingCountry>
Sánchez Province (UF 216151, GTW 7020c). However, if these specimens are indeed
<taxonomicName id="ADA14D2228537F1CF3FBFE74B739EEB8" box="[1228,1325,475,499]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="adolfi">
<emphasis id="58D5EAB328537F1CF3FBFE74B739EEB8" box="[1228,1325,475,499]" italics="true" pageId="12" pageNumber="13">A. adolfi</emphasis>
</taxonomicName>
it is most closely related to
<taxonomicName id="ADA14D2228537F1CF66AFE51B3C8ED5D" box="[349,476,510,534]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="wilhelmi">
<emphasis id="58D5EAB328537F1CF66AFE51B3C8ED5D" box="[349,476,510,534]" italics="true" pageId="12" pageNumber="13">A. wilhelmi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428537F1CF6DCFE51B05FED5D" box="[491,587,510,534]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="12" pageNumber="13">Figure 4</figureCitation>
T–X), a species found just to the west of Rio San Juan.
<taxonomicName id="ADA14D2228537F1CF3F7FE51B788ED5D" box="[1216,1436,510,534]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="wilhelmi">
<emphasis id="58D5EAB328537F1CF3F7FE51B788ED5D" box="[1216,1436,510,534]" italics="true" pageId="12" pageNumber="13">Abbottella wilhelmi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428537F1CF7A9FD8DB2EAED71" box="[158,254,546,570]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="12" pageNumber="13">Figure 4</figureCitation>
T–X) is very similar to
<taxonomicName id="ADA14D2228537F1CF53AFD8CB084ED70" box="[525,656,547,571]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="adolfi">
<emphasis id="58D5EAB328537F1CF53AFD8CB084ED70" box="[525,656,547,571]" italics="true" pageId="12" pageNumber="13">A. cf. adolfi</emphasis>
</taxonomicName>
but is smaller and has a different operculum. The operculum of
<taxonomicName id="ADA14D2228537F1CF256FD8CB2CDED15" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="adolfi">
<emphasis id="58D5EAB328537F1CF256FD8CB2CDED15" italics="true" pageId="12" pageNumber="13">A. cf. adolfi</emphasis>
</taxonomicName>
is composed of a broad, cup-like, oblique lamella; the operculum of
<taxonomicName id="ADA14D2228537F1CF4E0FDE9B642ED15" box="[983,1110,582,606]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="wilhelmi">
<emphasis id="58D5EAB328537F1CF4E0FDE9B642ED15" box="[983,1110,582,606]" italics="true" pageId="12" pageNumber="13">A. wilhelmi</emphasis>
</taxonomicName>
has a nearly vertical lamella.
<taxonomicName id="ADA14D2228537F1CF7A0FDC4B36CEDC8" box="[151,376,619,643]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="sanchezi">
<emphasis id="58D5EAB328537F1CF7A0FDC4B36CEDC8" box="[151,376,619,643]" italics="true" pageId="12" pageNumber="13">Abbottella sanchezi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428537F1CF6BCFDC5B3FBEDC9" box="[395,495,618,642]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="12" pageNumber="13">Figure 4</figureCitation>
J–N) is also similar but has a vertical lamella on the operculum and the spiral sculpture tends to form a carina at the periphery. Bartsch (1946) described a subspecies from the trail from Samaná to Rio San Juan,
<taxonomicName id="ADA14D2228537F1CF662FD1CB00FED80" box="[341,539,691,715]" class="Gastropoda" family="Pomatiidae" genus="Choanopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="subSpecies" species="adolfi" subSpecies="peninsularis">
<emphasis id="58D5EAB328537F1CF662FD1CB00FED80" box="[341,539,691,715]" italics="true" pageId="12" pageNumber="13">A. a. peninsularis</emphasis>
</taxonomicName>
(see below,
<figureCitation id="F29A2A2428537F1CF591FD1DB111ED81" box="[678,773,690,714]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="12" pageNumber="13">Figure 3</figureCitation>
J–L), which differs in having an unbanded outer lip.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="3F9ED89EC79DF7EAA96B5A691560E4BF">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B728557F1AF7A0FB91B080E802" ID-DOI="http://doi.org/10.5281/zenodo.6164928" ID-GBIF-Taxon="127664007" ID-Zenodo-Dep="6164928" LSID="urn:lsid:plazi:treatment:E20887B728557F1AF7A0FB91B080E802" httpUri="http://treatment.plazi.org/id/E20887B728557F1AF7A0FB91B080E802" lastPageNumber="11" pageId="10" pageNumber="11">
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<paragraph id="6A1E36A128557F1AF7A0FB91B0A1EB13" blockId="10.[151,693,1086,1146]" box="[151,693,1086,1112]" pageId="10" pageNumber="11">
<heading id="315681CD28557F1AF7A0FB91B0A1EB13" bold="true" box="[151,693,1086,1112]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="ADA14D2228557F1AF7A0FB91B0A1EB13" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[151,693,1086,1112]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="abbotti" subGenus="Abbottella">
<emphasis id="58D5EAB328557F1AF7A0FB91B0A1EB13" bold="true" box="[151,693,1086,1112]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7A0FB91B307EB13" bold="true" box="[151,275,1086,1112]" italics="true" pageId="10" pageNumber="11">Abbottella</emphasis>
(
<emphasis id="58D5EAB328557F1AF613FB91B3B4EB13" bold="true" box="[292,416,1086,1112]" italics="true" pageId="10" pageNumber="11">Abbottella</emphasis>
)
<emphasis id="58D5EAB328557F1AF687FB91B017EB13" bold="true" box="[432,515,1086,1112]" italics="true" pageId="10" pageNumber="11">abbotti</emphasis>
Bartsch, 1946
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6A1E36A128557F1AF7A0FBCEB379EB32" blockId="10.[151,693,1086,1146]" box="[151,365,1121,1146]" pageId="10" pageNumber="11">
<figureCitation id="F29A2A2428557F1AF7A0FBCEB329EB31" box="[151,317,1121,1146]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="10" pageNumber="11">Figures 3 A–E</figureCitation>
,
<figureCitation id="F29A2A2428557F1AF67FFBCDB342EB32" box="[328,342,1122,1145]" captionStart="FIGURE 7" captionStartId="20.[151,250,1604,1627]" captionTargetBox="[287,1301,204,1571]" captionTargetId="figure@20.[287,1302,193,1574]" captionTargetPageId="20" captionText="FIGURE 7. Distributions are approximate. Base map courtesy of NASA. A. Abbottella (Abbottella) calliotropis new species. B. Abbottella (Abbottella) diadema new species. C. Abbottella (Abbottella) dichroa new species. D. Leiabbottella soluta (Pfeiffer, 1852). E. Leiabbottella thompsoni new species. F. Abbottella (Abbottella) abbotti Bartsch, 1946. G. Abbottella (Abbottella) crossei (Pilsbry, 1933). H. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. I. Rolleia haitensis Bartsch, 1946." httpUri="https://zenodo.org/record/283735/files/figure.png" pageId="10" pageNumber="11">7</figureCitation>
F
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28557F1AF7A0FB05B16BEAD2" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="6A1E36A128557F1AF7A0FB05B07BEB8A" blockId="10.[151,1436,1193,1865]" box="[151,623,1193,1218]" pageId="10" pageNumber="11">
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<typeStatus id="B51A880328557F1AF7A0FB05B2C6EB89" box="[151,210,1194,1218]" pageId="10" pageNumber="11">Type</typeStatus>
material.
</emphasis>
USNM 504108,
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.
</paragraph>
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<emphasis id="58D5EAB328557F1AF7FEFB62B37DEBAD" bold="true" box="[201,361,1229,1254]" pageId="10" pageNumber="11">
<typeStatus id="B51A880328557F1AF7FEFB62B310EBAE" box="[201,260,1229,1253]" pageId="10" pageNumber="11">Type</typeStatus>
locality.
</emphasis>
Near Laguna, Samaná Bay [
<emphasis id="58D5EAB328557F1AF582FB61B0C0EBAE" box="[693,724,1230,1253]" italics="true" pageId="10" pageNumber="11">sic</emphasis>
, probable error for Samaná Peninsula, see below],
<collectingCountry id="12B6763128557F1AF228FB62B314EA42" name="Dominican Republic" pageId="10" pageNumber="11">Dominican Republic</collectingCountry>
.
</paragraph>
<paragraph id="6A1E36A128557F1AF7FEFABAB16BEAD2" blockId="10.[151,1436,1193,1865]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7FEFABAB3EAEA65" bold="true" box="[201,510,1301,1326]" pageId="10" pageNumber="11">Other material examined.</emphasis>
UF 215981, El Valle, Samaná Province,
<collectingCountry id="12B6763128557F1AF4FEFABAB6A0EA66" box="[969,1204,1301,1325]" name="Dominican Republic" pageId="10" pageNumber="11">Dominican Republic</collectingCountry>
; UF 215983, 2 km S of El Valle, Samaná Province,
<collectingCountry id="12B6763128557F1AF6C0FA96B0F0EA1A" box="[503,740,1337,1361]" name="Dominican Republic" pageId="10" pageNumber="11">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B4428557F1AF439FA96B158EA1A" box="[782,844,1337,1361]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="20.0">20 m</quantity>
; UF 215980, 2 km S of El Valle, Samaná Province,
<collectingCountry id="12B6763128557F1AF7A0FAF2B39DEA3E" box="[151,393,1373,1397]" name="Dominican Republic" pageId="10" pageNumber="11">Dominican Republic</collectingCountry>
; UF 215982, hills E and W of El Valle, Samaná Province,
<collectingCountry id="12B6763128557F1AF36DFAF2B75FEA3E" box="[1114,1355,1373,1397]" name="Dominican Republic" pageId="10" pageNumber="11">Dominican Republic</collectingCountry>
; GTW 10829a, Cayo Levantado, Samaná Province,
<collectingCountry id="12B6763128557F1AF5B8FA2EB16FEAD2" box="[655,891,1409,1433]" name="Dominican Republic" pageId="10" pageNumber="11">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28557F1AF7FEFA0AB080E802" pageId="10" pageNumber="11" type="distribution">
<paragraph id="6A1E36A128557F1AF7FEFA0AB38FE996" blockId="10.[151,1436,1193,1865]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7FEFA0AB3E6EAF5" bold="true" box="[201,498,1445,1470]" pageId="10" pageNumber="11">Distribution and habitat.</emphasis>
This species occurs in the Sierra de Samaná of the Cordillera Septentrional. Bartsch (1946) did not give any additional information on the
<typeStatus id="B51A880328557F1AF438FA65B12BEAA9" box="[783,831,1482,1506]" pageId="10" pageNumber="11">type</typeStatus>
locality in his description of this species. However, under his description of
<taxonomicName id="ADA14D2228557F1AF691FA41B080E94E" box="[422,660,1517,1541]" class="Gastropoda" family="Pomatiidae" genus="Lagopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="lagopoma">
<emphasis id="58D5EAB328557F1AF691FA41B080E94E" box="[422,660,1517,1541]" italics="true" pageId="10" pageNumber="11">Lagopoma lagopoma</emphasis>
</taxonomicName>
, he stated that Laguna was
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north of Samaná. This is probably Punta Laguna Salada, which is adjacent to El Valle, the source of most of our material. Contrary to Bartsch’s description, Laguna is not on Samaná Bay but on the north coast of the peninsula; this was probably an error for Samaná Peninsula. All of our records are for the El Valle and the Las Galeras areas on the northeast coast of the peninsula. This species is probably distributed around the base of the outcrop between these two areas, a distance of ca.
<quantity id="AD599B4428557F1AF7D5F90EB33BE9F2" box="[226,303,1697,1721]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="km" value="20.0">20 km</quantity>
. Snails occur on and at the base of the steep limestone walls in cool, shaded areas with abundant vegetation and mosses.
</paragraph>
<paragraph id="6A1E36A128557F1AF7FEF946B080E802" blockId="10.[151,1436,1193,1865]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7FEF946B046E849" bold="true" box="[201,594,1769,1794]" pageId="10" pageNumber="11">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D2228557F1AF568F946B125E84A" box="[607,817,1769,1793]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="abbotti">
<emphasis id="58D5EAB328557F1AF568F946B125E84A" box="[607,817,1769,1793]" italics="true" pageId="10" pageNumber="11">Abbottella abbotti</emphasis>
</taxonomicName>
is one of a group of small species on the Samaná Peninsula characterized by thorny sculpture. It differs from similar species in its deeply fimbriated outer lip and coarse sculpture arranged in prominent keels.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="EA15B44A28F59B1FA6EFCB197568A9E6" ID-DOI="10.11646/zootaxa.3646.1.1" ID-GBIF-Dataset="9b6ec282-ff71-4189-81cb-f63930d210de" ID-ISSN="1175-5326" ID-Zenodo-Dep="283728" ID-ZooBank="55F70B5D-79AD-4C77-9164-EFE7126316E8" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460598948263" checkinUser="plazi" docAuthor="Watters, G. Thomas" docDate="2013" docId="E20887B728577F1AF7A0F96EB693ECEF" docLanguage="en" docName="zt03646p022.pdf" docOrigin="Zootaxa 3646 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leiabbottella thompsoni Watters, 2013, new species" docType="treatment" docVersion="6" lastPageNumber="11" masterDocId="1E31FFCF285F7F10F737FFAFB214EF4B" masterDocTitle="New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="9" updateTime="1698327222889" updateUser="plazi">
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<mods:title id="C70E1CC7B584F4726530FA9B42E3515D">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<mods:namePart id="E1CD68AD15F9E421778A901CC06EDA8E">Watters, G. Thomas</mods:namePart>
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<mods:date id="1E7082E2F705D59DA335D4F56CD9101A">2013</mods:date>
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<treatment id="E20887B728577F1AF7A0F96EB693ECEF" ID-DOI="http://doi.org/10.5281/zenodo.6164926" ID-GBIF-Taxon="127663993" ID-Zenodo-Dep="6164926" LSID="urn:lsid:plazi:treatment:E20887B728577F1AF7A0F96EB693ECEF" httpUri="http://treatment.plazi.org/id/E20887B728577F1AF7A0F96EB693ECEF" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="22BB652A28577F18F7A0F96EB37AE9B7" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="6A1E36A128577F18F7A0F96EB046E991" blockId="8.[151,594,1728,1788]" box="[151,594,1728,1755]" pageId="8" pageNumber="9">
<heading id="315681CD28577F18F7A0F96EB046E991" bold="true" box="[151,594,1728,1755]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="58D5EAB328577F18F7A0F96EB046E991" bold="true" box="[151,594,1728,1755]" pageId="8" pageNumber="9">
<taxonomicName id="ADA14D2228577F18F7A0F96EB3AFE990" box="[151,443,1729,1755]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="thompsoni" status="sp. nov.">
<emphasis id="58D5EAB328577F18F7A0F96EB3AFE990" bold="true" box="[151,443,1729,1755]" italics="true" pageId="8" pageNumber="9">Leiabbottella thompsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C828577F18F6F5F96FB046E991" box="[450,594,1728,1754]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A128577F18F7A0F94BB37AE9B7" blockId="8.[151,594,1728,1788]" box="[151,366,1764,1788]" pageId="8" pageNumber="9">
<figureCitation id="F29A2A2428577F18F7A0F94BB329E9B7" box="[151,317,1764,1788]" captionStart="FIGURE 2. A – E" captionStartId="9.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@9.[151,1436,193,1582]" captionTargetPageId="9" captionText="FIGURE 2. A – E. Leiabbottella thompsoni new species. A – C. Holotype (UF 456799), 13.7 mm width. D. Paratype (UF 236225), 14.1 mm width. E. Paratype (UF 236225), 13.6 mm width. F – I. Leiabbottella soluta (Pfeiffer, 1852). F. Illustration from Pfeiffer (1854: pl. 39, fig. 8). G. UF 216196, 11.9 mm width. H. UF 216196, 13.9 mm width. I. UF 216196, 13.8 mm width. J – L. Leiabbottella galaxius Watters, 2010. Paratype (OSUM 35491), 8.4 mm width." httpUri="https://zenodo.org/record/283730/files/figure.png" pageId="8" pageNumber="9">Figures 2 A–E</figureCitation>
,
<figureCitation id="F29A2A2428577F18F67FF94AB342E9B7" box="[328,342,1765,1788]" captionStart="FIGURE 7" captionStartId="20.[151,250,1604,1627]" captionTargetBox="[287,1301,204,1571]" captionTargetId="figure@20.[287,1302,193,1574]" captionTargetPageId="20" captionText="FIGURE 7. Distributions are approximate. Base map courtesy of NASA. A. Abbottella (Abbottella) calliotropis new species. B. Abbottella (Abbottella) diadema new species. C. Abbottella (Abbottella) dichroa new species. D. Leiabbottella soluta (Pfeiffer, 1852). E. Leiabbottella thompsoni new species. F. Abbottella (Abbottella) abbotti Bartsch, 1946. G. Abbottella (Abbottella) crossei (Pilsbry, 1933). H. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. I. Rolleia haitensis Bartsch, 1946." httpUri="https://zenodo.org/record/283735/files/figure.png" pageId="8" pageNumber="9">7</figureCitation>
E
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28577F19F7A0F883B314E8A7" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="6A1E36A128577F18F7A0F883B2E5E82C" blockId="8.[151,1436,1835,1968]" pageId="8" pageNumber="9">
<emphasis id="58D5EAB328577F18F7A0F883B352E80F" bold="true" box="[151,326,1835,1860]" pageId="8" pageNumber="9">
<typeStatus id="B51A880328577F18F7A0F883B2C6E80F" box="[151,210,1836,1860]" pageId="8" pageNumber="9">Type</typeStatus>
material.
</emphasis>
UF 456799,
<typeStatus id="B51A880328577F18F6E8F883B050E80F" box="[479,580,1836,1860]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
; length,
<quantity id="AD599B4428577F18F590F883B115E80F" box="[679,769,1836,1860]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="8" pageNumber="9" unit="mm" value="7.6">7.6 mm</quantity>
; width,
<quantity id="AD599B4428577F18F46AF883B1D2E80F" box="[861,966,1836,1860]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="8" pageNumber="9" unit="mm" value="13.7">13.7 mm</quantity>
. UF 236225, 91
<typeStatus id="B51A880328577F18F3B0F882B6E1E80E" box="[1159,1269,1837,1861]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
, from the
<typeStatus id="B51A880328577F18F25BF882B788E80E" box="[1388,1436,1837,1861]" pageId="8" pageNumber="9">type</typeStatus>
locality.
</paragraph>
<paragraph id="6A1E36A128577F18F7FEF8DBB0B6E8FB" blockId="8.[151,1436,1835,1968]" pageId="8" pageNumber="9">
<emphasis id="58D5EAB328577F18F7FEF8DBB37EE8C7" bold="true" box="[201,362,1907,1932]" pageId="8" pageNumber="9">
<typeStatus id="B51A880328577F18F7FEF8DBB310E8C7" box="[201,260,1908,1932]" pageId="8" pageNumber="9">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B6763128577F18F643F8DBB076E8C7" box="[372,610,1908,1932]" name="Dominican Republic" pageId="8" pageNumber="9">Dominican Republic</collectingCountry>
, Monte Plata Province,
<quantity id="AD599B4428577F18F44EF8DBB1A0E8C7" box="[889,948,1908,1932]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="km" value="5.0">5 km</quantity>
N of Majagual, at
<specimenCount id="7CA7FD2828577F18F3A2F8DBB6F2E8C7" box="[1173,1254,1908,1932]" pageId="8" pageNumber="9" type="generic">
<quantity id="AD599B4428577F18F3A2F8DBB6F6E8C7" box="[1173,1250,1908,1932]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="m" value="150.0">150 m</quantity>
.
</specimenCount>
ca.
<geoCoordinate id="0F95506628577F18F22DF8DBB790E8C7" box="[1306,1412,1908,1932]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="19.09">19.09° N</geoCoordinate>
, -
<geoCoordinate id="0F95506628577F18F7A0F837B317E8FB" box="[151,259,1944,1968]" direction="west" orientation="longitude" pageId="8" pageNumber="9" precision="555" value="-69.83">69.83° W</geoCoordinate>
. See Remarks under
<taxonomicName id="ADA14D2228577F18F6D9F836B044E8FB" box="[494,592,1944,1968]" class="Gastropoda" family="Pomatiidae" genus="Lagopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="soluta">
<emphasis id="58D5EAB328577F18F6D9F836B044E8FB" box="[494,592,1944,1968]" italics="true" pageId="8" pageNumber="9">L. soluta</emphasis>
</taxonomicName>
below.
</paragraph>
<caption id="3EDE662928567F19F7A0F9ECB626E9FD" httpUri="https://zenodo.org/record/283730/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1435,193,1582]" targetPageId="9">
<paragraph id="6A1E36A128567F19F7A0F9ECB626E9FD" blockId="9.[151,1436,1603,1718]" pageId="9" pageNumber="10">
<emphasis id="58D5EAB328567F19F7A0F9ECB303E912" bold="true" box="[151,279,1603,1626]" pageId="9" pageNumber="10">FIGURE 2.</emphasis>
A–E.
<taxonomicName id="ADA14D2228567F19F656F9EBB04FE911" box="[353,603,1604,1626]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="thompsoni" status="sp. nov.">
<emphasis id="58D5EAB328567F19F656F9EBB04FE911" box="[353,603,1604,1626]" italics="true" pageId="9" pageNumber="10">Leiabbottella thompsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C828567F19F551F9EBB0F2E911" box="[614,742,1603,1626]" pageId="9" pageNumber="10" rank="species">
<emphasis id="58D5EAB328567F19F551F9EBB0F2E911" bold="true" box="[614,742,1603,1626]" pageId="9" pageNumber="10">new species</emphasis>
</taxonomicNameLabel>
. A–C. Holotype (UF 456799), 13.7 mm width. D. Paratype (UF 236225), 14.1 mm width. E. Paratype (UF 236225), 13.6 mm width. F–I.
<taxonomicName id="ADA14D2228567F19F4A5F9CCB713E933" authority="Pfeiffer, 1852" authorityName="Pfeiffer" authorityYear="1852" box="[914,1287,1634,1657]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="soluta">
<emphasis id="58D5EAB328567F19F4A5F9CCB64BE932" box="[914,1119,1635,1657]" italics="true" pageId="9" pageNumber="10">Leiabbottella soluta</emphasis>
(Pfeiffer, 1852)
</taxonomicName>
. F. Illustration from Pfeiffer (1854: pl. 39, fig. 8). G. UF 216196, 11.9 mm width. H. UF 216196, 13.9 mm width. I. UF 216196, 13.8 mm width. J–L.
<taxonomicName id="ADA14D2228567F19F627F90FB09DE9FE" authority="Watters, 2010" authorityName="Watters" authorityYear="2010" box="[272,649,1696,1718]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="galaxius">
<emphasis id="58D5EAB328567F19F627F90FB3E7E9FD" box="[272,499,1696,1718]" italics="true" pageId="9" pageNumber="10">Leiabbottella galaxius</emphasis>
Watters, 2010
</taxonomicName>
. Paratype (OSUM 35491), 8.4 mm width.
</paragraph>
</caption>
<paragraph id="6A1E36A128567F19F7FEF954B314E8A7" blockId="9.[151,1436,1787,2028]" pageId="9" pageNumber="10">
<emphasis id="58D5EAB328567F19F7FEF954B3E8E85F" bold="true" box="[201,508,1787,1812]" pageId="9" pageNumber="10">Other material examined.</emphasis>
UF 216194,
<specimenCount id="7CA7FD2828567F19F5B9F953B13DE85F" box="[654,809,1788,1812]" pageId="9" pageNumber="10" type="generic">18 specimens</specimenCount>
,
<quantity id="AD599B4428567F19F404F953B17FE85F" box="[819,875,1788,1812]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="km" value="5.0">5 km</quantity>
N of Majagual, Monte Plata Province,
<collectingCountry id="12B6763128567F19F228F953B314E873" name="Dominican Republic" pageId="9" pageNumber="10">Dominican Republic</collectingCountry>
; UF 236230,
<specimenCount id="7CA7FD2828567F19F6ADF88FB021E873" box="[410,565,1824,1848]" pageId="9" pageNumber="10" type="generic">29 specimens</specimenCount>
,
<quantity id="AD599B4428567F19F576F88FB06FE873" box="[577,635,1824,1848]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="km" value="2.0">2 km</quantity>
N of Majagual, Monte Plata Province,
<collectingCountry id="12B6763128567F19F30EF88FB731E873" box="[1081,1317,1824,1848]" name="Dominican Republic" pageId="9" pageNumber="10">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B4428567F19F27AF88FB783E873" box="[1357,1431,1824,1848]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="m" value="150.0">150 m</quantity>
; UF 216189,
<specimenCount id="7CA7FD2828567F19F615F8EBB3A8E817" box="[290,444,1860,1884]" pageId="9" pageNumber="10" type="generic">30 specimens</specimenCount>
,
<quantity id="AD599B4428567F19F6F0F8EAB3EBE817" box="[455,511,1860,1884]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="10" unit="km" value="7.0">7 km</quantity>
N of Majagual, Monte Plata Province,
<collectingCountry id="12B6763128567F19F484F8EBB689E817" box="[947,1181,1860,1884]" name="Dominican Republic" pageId="9" pageNumber="10">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B4428567F19F3F5F8EBB718E817" box="[1218,1292,1860,1884]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="10" unit="m" value="105.0">105 m</quantity>
; UF 216127,
<specimenCount id="7CA7FD2828567F19F7A0F8C7B327E8CB" box="[151,307,1896,1920]" pageId="9" pageNumber="10" type="generic">26 specimens</specimenCount>
,
<quantity id="AD599B4428567F19F608F8C7B393E8CB" box="[319,391,1896,1920]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="10" unit="km" value="13.0">13 km</quantity>
NW of Sabana Grande de Boyá, Monte Plata Province,
<collectingCountry id="12B6763128567F19F33CF8C7B6E3E8CB" box="[1035,1271,1896,1920]" name="Dominican Republic" pageId="9" pageNumber="10">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B4428567F19F217F8C7B77FE8CB" box="[1312,1387,1896,1920]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="10" unit="m" value="250.0">250 m</quantity>
; UF 249105,
<specimenCount id="7CA7FD2828567F19F7CFF823B391E8EF" box="[248,389,1932,1956]" pageId="9" pageNumber="10" type="generic">6 specimens</specimenCount>
, N of Majagual, ca.
<quantity id="AD599B4428567F19F551F823B0B8E8EF" box="[614,684,1932,1956]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="10" unit="km" value="12.0">12 km</quantity>
NW of Sabana Grande de Boyá, Monte Plata Province,
<collectingCountry id="12B6763128567F19F228F823B2EBE883" name="Dominican Republic" pageId="9" pageNumber="10">Dominican Republic</collectingCountry>
, at
<quantity id="AD599B4428567F19F610F81FB365E883" box="[295,369,1968,1992]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="m" value="150.0">150 m</quantity>
; UF 216193,
<specimenCount id="7CA7FD2828567F19F53BF81FB0BCE883" box="[524,680,1968,1992]" pageId="9" pageNumber="10" type="generic">22 specimens</specimenCount>
,
<quantity id="AD599B4428567F19F583F81FB0F9E883" box="[692,749,1968,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="km" value="1.0">1 km</quantity>
N of Majagual, Monte Plata Province,
<collectingCountry id="12B6763128567F19F39BF81FB78CE883" box="[1196,1432,1968,1992]" name="Dominican Republic" pageId="9" pageNumber="10">Dominican Republic</collectingCountry>
, at
<specimenCount id="7CA7FD2828567F19F785F87BB314E8A7" box="[178,256,2004,2028]" pageId="9" pageNumber="10" type="generic">
<quantity id="AD599B4428567F19F785F87BB2E8E8A7" box="[178,252,2004,2028]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="m" value="200.0">200 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28557F1AF7FEFF38B746EFB3" pageId="10" pageNumber="11" type="distribution">
<paragraph id="6A1E36A128557F1AF7FEFF38B746EFB3" blockId="10.[151,1437,151,933]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7FEFF38B3E7EFFB" bold="true" box="[201,499,151,176]" pageId="10" pageNumber="11">Distribution and habitat.</emphasis>
Known only from the karstic region of Los Haitises, which separates the Cordillera Central from the Cordillera Oriental. The species lives at the base of deeply karsted limestone knolls and in ravines in tropical forest under debris and leaf litter. A portion of this distribution has been planted as coffee groves.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28557F1AF7FEFEACB776EDCF" pageId="10" pageNumber="11" type="description">
<paragraph id="6A1E36A128557F1AF7FEFEACB776EDCF" blockId="10.[151,1437,151,933]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7FEFEACB34EEE57" bold="true" box="[201,346,259,284]" pageId="10" pageNumber="11">Description.</emphasis>
Shell large for genus (largest specimen,
<quantity id="AD599B4428557F1AF40EFEAAB187EE57" box="[825,915,261,284]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="10" pageNumber="11" unit="mm" value="7.7">7.7 mm</quantity>
total length x
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total width; smallest specimen,
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total length x
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total width;
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,
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total length including peristome x
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total width including peristome), depressed, nearly planispiral, whorls adnate except just before lip. Umbilicus very wide, ca. 35% of total width, open to earliest whorls. Protoconch of ca. 1.5 erect whorls but demarcation between protoconch and teleoconch not well-defined. Teleoconch of 3.25 whorls. Shell smooth except for numerous (ca. 50–60 on final whorl), incised axial lines between which are microscopic threads. Spiral sculpture absent except for the faintest trace of umbilical cords in a few specimens. Suture deeply indented. Aperture double, circular (
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maximum width in
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), adnate to the previous whorl. Inner lip smooth, exserted, tube-like. Outer lip broadly expanded, sinuate, adherent to the previous whorl, forming a broad, prominent, cup-shaped auricle. Shell dirty white, tan, or bronze, somewhat shiny, patterned with narrow, often interrupted, brown spiral bands, apparent on both sides of outer lip and on inner lip as well. Operculum multispiral with an oblique, calcareous lamella.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28557F1AF7FEFD20B709EC17" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="6A1E36A128557F1AF7FEFD20B0F8ED87" blockId="10.[151,1437,151,933]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7FEFD20B3F5EDE3" bold="true" box="[201,481,655,680]" pageId="10" pageNumber="11">Variation in specimens.</emphasis>
<specimenCount id="7CA7FD2828557F1AF6DEFD20B09BEDEC" box="[489,655,655,679]" pageId="10" pageNumber="11" type="generic">223 specimens</specimenCount>
seen. Specimens are uniform in shell characteristics, varying only in the strength of the color bands and background color.
</paragraph>
<paragraph id="6A1E36A128557F1AF7FEFD78B709EC17" blockId="10.[151,1437,151,933]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7FEFD78B052EDBB" bold="true" box="[201,582,727,752]" pageId="10" pageNumber="11">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D2228557F1AF579FD77B1FDEDA4" authority="Pfeiffer, 1852" authorityName="Pfeiffer" authorityYear="1852" box="[590,1001,727,752]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="soluta">
<emphasis id="58D5EAB328557F1AF579FD77B125EDBB" box="[590,817,728,752]" italics="true" pageId="10" pageNumber="11">Leiabbottella soluta</emphasis>
(Pfeiffer, 1852)
</taxonomicName>
(
<figureCitation id="F29A2A2428557F1AF4CEFD78B64FEDA4" box="[1017,1115,727,751]" captionStart="FIGURE 2. A – E" captionStartId="9.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@9.[151,1436,193,1582]" captionTargetPageId="9" captionText="FIGURE 2. A – E. Leiabbottella thompsoni new species. A – C. Holotype (UF 456799), 13.7 mm width. D. Paratype (UF 236225), 14.1 mm width. E. Paratype (UF 236225), 13.6 mm width. F – I. Leiabbottella soluta (Pfeiffer, 1852). F. Illustration from Pfeiffer (1854: pl. 39, fig. 8). G. UF 216196, 11.9 mm width. H. UF 216196, 13.9 mm width. I. UF 216196, 13.8 mm width. J – L. Leiabbottella galaxius Watters, 2010. Paratype (OSUM 35491), 8.4 mm width." httpUri="https://zenodo.org/record/283730/files/figure.png" pageId="10" pageNumber="11">Figure 2</figureCitation>
F–I) is closely related to
<taxonomicName id="ADA14D2228557F1AF2B2FD76B318EC5F" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="thompsoni">
<emphasis id="58D5EAB328557F1AF2B2FD76B318EC5F" italics="true" pageId="10" pageNumber="11">L. thompsoni</emphasis>
</taxonomicName>
.
<taxonomicName id="ADA14D2228557F1AF62BFD53B015EC5F" box="[284,513,764,788]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="soluta">
<emphasis id="58D5EAB328557F1AF62BFD53B015EC5F" box="[284,513,764,788]" italics="true" pageId="10" pageNumber="11">Leiabbottella soluta</emphasis>
</taxonomicName>
tends to be less planispiral, with relatively wider whorls, and has a less adnate outer lip auricle than does
<taxonomicName id="ADA14D2228557F1AF688FC8EB046EC73" box="[447,594,800,824]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="thompsoni">
<emphasis id="58D5EAB328557F1AF688FC8EB046EC73" box="[447,594,800,824]" italics="true" pageId="10" pageNumber="11">L. thompsoni</emphasis>
</taxonomicName>
.
<taxonomicName id="ADA14D2228557F1AF568FC8FB12BEC73" box="[607,831,800,824]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="soluta">
<emphasis id="58D5EAB328557F1AF568FC8FB12BEC73" box="[607,831,800,824]" italics="true" pageId="10" pageNumber="11">Leiabbottella soluta</emphasis>
</taxonomicName>
occurs over
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to the northwest in the Cordillera Septentrional.
<taxonomicName id="ADA14D2228557F1AF60DFCEBB021EC17" box="[314,565,836,860]" class="Gastropoda" family="Annulariidae" genus="Leiabbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="galaxius">
<emphasis id="58D5EAB328557F1AF60DFCEBB021EC17" box="[314,565,836,860]" italics="true" pageId="10" pageNumber="11">Leiabbottella galaxius</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428557F1AF574FCEBB0B7EC17" box="[579,675,836,860]" captionStart="FIGURE 2. A – E" captionStartId="9.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@9.[151,1436,193,1582]" captionTargetPageId="9" captionText="FIGURE 2. A – E. Leiabbottella thompsoni new species. A – C. Holotype (UF 456799), 13.7 mm width. D. Paratype (UF 236225), 14.1 mm width. E. Paratype (UF 236225), 13.6 mm width. F – I. Leiabbottella soluta (Pfeiffer, 1852). F. Illustration from Pfeiffer (1854: pl. 39, fig. 8). G. UF 216196, 11.9 mm width. H. UF 216196, 13.9 mm width. I. UF 216196, 13.8 mm width. J – L. Leiabbottella galaxius Watters, 2010. Paratype (OSUM 35491), 8.4 mm width." httpUri="https://zenodo.org/record/283730/files/figure.png" pageId="10" pageNumber="11">Figure 2</figureCitation>
J–L) is smaller and has a much less developed outer lip.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28557F1AF7FEFCC7B693ECEF" pageId="10" pageNumber="11" type="etymology">
<paragraph id="6A1E36A128557F1AF7FEFCC7B693ECEF" blockId="10.[151,1437,151,933]" pageId="10" pageNumber="11">
<emphasis id="58D5EAB328557F1AF7FEFCC7B35BECCB" bold="true" box="[201,335,872,896]" pageId="10" pageNumber="11">Etymology.</emphasis>
Named for Dr. Fred Thompson, Curator, UF, whose unparalleled collections in Hispaniola form the basis of our understanding of this group and the terrestrial snail fauna there in general.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D97F4DD4F0FF654FCC2D33BBA4DEF18D">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B728587F17F7A0FABFB02FE9ED" ID-DOI="http://doi.org/10.5281/zenodo.6164922" ID-GBIF-Taxon="127663999" ID-Zenodo-Dep="6164922" LSID="urn:lsid:plazi:treatment:E20887B728587F17F7A0FABFB02FE9ED" httpUri="http://treatment.plazi.org/id/E20887B728587F17F7A0FABFB02FE9ED" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="22BB652A28587F17F7A0FABFB168EA27" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="6A1E36A128587F17F7A0FABFB0EAEA61" blockId="7.[151,766,1296,1323]" box="[151,766,1296,1323]" pageId="7" pageNumber="8">
<heading id="315681CD28587F17F7A0FABFB0EAEA61" bold="true" box="[151,766,1296,1323]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="58D5EAB328587F17F7A0FABFB0EAEA61" bold="true" box="[151,766,1296,1323]" pageId="7" pageNumber="8">
Subgenus
<taxonomicName id="ADA14D2228587F17F620FABEB0EAEA61" authority="Torre &amp; Bartsch, 1941" authorityName="Torre &amp; Bartsch" authorityYear="1941" box="[279,766,1296,1323]" class="Gastropoda" family="Pomatiidae" genus="Gundlachtudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328587F17F620FABEB3CBEA60" bold="true" box="[279,479,1297,1323]" italics="true" pageId="7" pageNumber="8">Gundlachtudora</emphasis>
Torre &amp; Bartsch, 1941
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</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A128587F17F7A0FAF8B168EA27" blockId="7.[151,892,1367,1389]" box="[151,892,1367,1389]" pageId="7" pageNumber="8">
<typeStatus id="B51A880328587F17F7A0FAF8B2D3EA27" box="[151,199,1367,1388]" pageId="7" pageNumber="8">Type</typeStatus>
species by original designation
<taxonomicName id="ADA14D2228587F17F6CCFAF7B16CEA27" authority="Pfeiffer, 1859" authorityName="Pfeiffer" authorityYear="1859" box="[507,888,1367,1389]" class="Gastropoda" family="Assimineidae" genus="Cyclostoma" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="decoloratum">
<emphasis id="58D5EAB328587F17F6CCFAF7B0FFEA26" box="[507,747,1368,1389]" italics="true" pageId="7" pageNumber="8">Cyclostoma decoloratum</emphasis>
Pfeiffer, 1859
</taxonomicName>
.
</paragraph>
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<subSubSection id="22BB652A28587F17F7A0FA3DB75AEAE1" box="[151,1358,1426,1451]" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="6A1E36A128587F17F7A0FA3DB75AEAE1" blockId="7.[151,1437,1426,1702]" box="[151,1358,1426,1451]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7A0FA3DB305EAE0" bold="true" box="[151,273,1426,1451]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
Like
<emphasis id="58D5EAB328587F17F665FA3CB3E5EAE0" box="[338,497,1427,1451]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="ADA14D2228587F17F665FA3CB3D3EAE0" box="[338,455,1427,1451]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus" sensu="stricto">Abbottella</taxonomicName>
<taxonomicNameLabel id="43E657C828587F17F6F9FA3BB3E5EAE0" box="[462,497,1428,1451]" pageId="7" pageNumber="8" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
but operculum with reflected lamellae. Operculum may not fit into aperture.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28587F17F7F0FA18B02FE9ED" pageId="7" pageNumber="8" type="discussion">
<paragraph id="6A1E36A128587F17F7F0FA18B02FE9ED" blockId="7.[151,1437,1426,1702]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7F0FA18B32FEA84" bold="true" box="[199,315,1463,1487]" pageId="7" pageNumber="8">Remarks.</emphasis>
Torre &amp; Bartsch (1941) had created
<taxonomicName id="ADA14D2228587F17F5EDFA19B187EA85" box="[730,915,1462,1486]" class="Gastropoda" family="Pomatiidae" genus="Gundlachtudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328587F17F5EDFA19B187EA85" box="[730,915,1462,1486]" italics="true" pageId="7" pageNumber="8">Gundlachtudora</emphasis>
</taxonomicName>
for a Cuban
<taxonomicName id="ADA14D2228587F17F311FA19B68EEA85" box="[1062,1178,1462,1486]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328587F17F311FA19B68EEA85" box="[1062,1178,1462,1486]" italics="true" pageId="7" pageNumber="8">Abbottella</emphasis>
</taxonomicName>
-like species having an operculum with a reflected lamella forming a continuous surface. Bartsch (1946) then created
<taxonomicName id="ADA14D2228587F17F3E0FA73B74FEAB8" box="[1239,1371,1500,1523]" class="Gastropoda" family="Pomatiidae" genus="Petasipoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328587F17F3E0FA73B74FEAB8" box="[1239,1371,1500,1523]" italics="true" pageId="7" pageNumber="8">Petasipoma</emphasis>
</taxonomicName>
for a Haitian species with the same characteristics. Watters (2006) synonymized the two genera and relegated
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<emphasis id="58D5EAB328587F17F7A0F98CB344E970" box="[151,336,1571,1595]" italics="true" pageId="7" pageNumber="8">Gundlachtudora</emphasis>
</taxonomicName>
to a subgenus of
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<emphasis id="58D5EAB328587F17F515F98CB082E970" box="[546,662,1571,1595]" italics="true" pageId="7" pageNumber="8">Abbottella</emphasis>
</taxonomicName>
. The species described here also has this characteristic operculum. However, it may be that these opercular features have evolved independently and the taxa are not actually closely related. Additional phylogenetic work is required to answer this question. For this reason this new species is placed in
<taxonomicName id="ADA14D2228587F17F783F921B379E9ED" box="[180,365,1678,1702]" class="Gastropoda" family="Pomatiidae" genus="Gundlachtudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328587F17F783F921B379E9ED" box="[180,365,1678,1702]" italics="true" pageId="7" pageNumber="8">Gundlachtudora</emphasis>
</taxonomicName>
with reservations.
</paragraph>
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<document id="7B8C98DE626B50A87BEE62ABDD693EB5" ID-DOI="10.11646/zootaxa.3646.1.1" ID-GBIF-Dataset="9b6ec282-ff71-4189-81cb-f63930d210de" ID-ISSN="1175-5326" ID-Zenodo-Dep="283728" ID-ZooBank="55F70B5D-79AD-4C77-9164-EFE7126316E8" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460598948263" checkinUser="plazi" docAuthor="Watters, G. Thomas" docDate="2013" docId="E20887B728587F17F7A0FF38B1FFEB89" docLanguage="en" docName="zt03646p022.pdf" docOrigin="Zootaxa 3646 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Abbottella (Abbottella) tenebrosa Watters, 2013, new species" docType="treatment" docVersion="6" lastPageNumber="8" masterDocId="1E31FFCF285F7F10F737FFAFB214EF4B" masterDocTitle="New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="8" updateTime="1698327222889" updateUser="plazi">
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<mods:title id="905000D9BF6DAEE10FC71A34B99C73BD">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<mods:namePart id="7FF49543EDE89C2A03DFD9C85D597C17">Watters, G. Thomas</mods:namePart>
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<mods:date id="BC008A959FAF78AFF5B0518BD59F54C8">2013</mods:date>
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<treatment id="E20887B728587F17F7A0FF38B1FFEB89" ID-DOI="http://doi.org/10.5281/zenodo.6164920" ID-GBIF-Taxon="127664004" ID-Zenodo-Dep="6164920" LSID="urn:lsid:plazi:treatment:E20887B728587F17F7A0FF38B1FFEB89" httpUri="http://treatment.plazi.org/id/E20887B728587F17F7A0FF38B1FFEB89" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="22BB652A28587F17F7A0FF38B37AEF99" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="6A1E36A128587F17F7A0FF38B0A9EFFA" blockId="7.[151,701,151,210]" box="[151,701,151,177]" pageId="7" pageNumber="8">
<heading id="315681CD28587F17F7A0FF38B0A9EFFA" bold="true" box="[151,701,151,177]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="58D5EAB328587F17F7A0FF38B0A9EFFA" bold="true" box="[151,701,151,177]" pageId="7" pageNumber="8">
<taxonomicName id="ADA14D2228587F17F7A0FF38B031EFFA" box="[151,549,151,177]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tenebrosa" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB328587F17F7A0FF38B307EFFA" bold="true" box="[151,275,151,177]" italics="true" pageId="7" pageNumber="8">Abbottella</emphasis>
(
<emphasis id="58D5EAB328587F17F613FF38B3B4EFFA" bold="true" box="[292,416,151,177]" italics="true" pageId="7" pageNumber="8">Abbottella</emphasis>
)
<emphasis id="58D5EAB328587F17F687FF38B031EFFA" bold="true" box="[432,549,151,177]" italics="true" pageId="7" pageNumber="8">tenebrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C828587F17F51AFF38B0A9EFFA" box="[557,701,151,177]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A128587F17F7A0FF15B37AEF99" blockId="7.[151,701,151,210]" box="[151,366,186,210]" pageId="7" pageNumber="8">
<figureCitation id="F29A2A2428587F17F7A0FF15B314EF99" box="[151,256,186,210]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="7" pageNumber="8">Figures 1</figureCitation>
U–Y, 8 B
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28587F17F7A0FEACB06EEEE1" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="6A1E36A128587F17F7A0FEACB163EE75" blockId="7.[151,1437,258,1219]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7A0FEACB350EE50" bold="true" box="[151,324,258,283]" pageId="7" pageNumber="8">
<typeStatus id="B51A880328587F17F7A0FEACB2C6EE50" box="[151,210,259,283]" pageId="7" pageNumber="8">Type</typeStatus>
material.
</emphasis>
UF 456796,
<typeStatus id="B51A880328587F17F6E1FEADB028EE51" box="[470,572,258,282]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
; length,
<quantity id="AD599B4428587F17F5ADFEADB0E6EE51" box="[666,754,258,282]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="8" unit="mm" value="7.6">7.6 mm</quantity>
; width,
<quantity id="AD599B4428587F17F47CFEADB1B7EE51" box="[843,931,258,282]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="7" pageNumber="8" unit="mm" value="8.9">8.9 mm</quantity>
. UF 456797, 15
<typeStatus id="B51A880328587F17F36BFEACB6DFEE50" box="[1116,1227,259,283]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
; OSUM 37271, 12
<typeStatus id="B51A880328587F17F7A0FE88B312EE74" box="[151,262,295,319]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
; BMSM 17937, 16
<typeStatus id="B51A880328587F17F6D4FE88B046EE74" box="[483,594,295,319]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
; all from the
<typeStatus id="B51A880328587F17F5D1FE88B102EE74" box="[742,790,295,319]" pageId="7" pageNumber="8">type</typeStatus>
locality.
</paragraph>
<paragraph id="6A1E36A128587F17F7FEFEE4B78DEE29" blockId="7.[151,1437,258,1219]" box="[201,1433,330,355]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7FEFEE4B371EE28" bold="true" box="[201,357,330,355]" pageId="7" pageNumber="8">
<typeStatus id="B51A880328587F17F7FEFEE4B316EE28" box="[201,258,331,355]" pageId="7" pageNumber="8">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B6763128587F17F659FEE5B04DEE29" box="[366,601,330,354]" name="Dominican Republic" pageId="7" pageNumber="8">Dominican Republic</collectingCountry>
, Puerto Plata Province, El Choco, near Cabarete. ca.
<geoCoordinate id="0F95506628587F17F39CFEE5B707EE29" box="[1195,1299,330,354]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="555" value="19.74">19.74° N</geoCoordinate>
, -
<geoCoordinate id="0F95506628587F17F211FEE5B780EE29" box="[1318,1428,330,354]" direction="west" orientation="longitude" pageId="7" pageNumber="8" precision="555" value="-70.42">70.42° W</geoCoordinate>
.
</paragraph>
<paragraph id="6A1E36A128587F17F7FEFEC1B06EEEE1" blockId="7.[151,1437,258,1219]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7FEFEC1B016EECC" bold="true" box="[201,514,366,391]" pageId="7" pageNumber="8">Other material examined.</emphasis>
UF 456798, 12 immature or weathered specimens, from the
<typeStatus id="B51A880328587F17F388FEC0B6FBEECC" box="[1215,1263,367,391]" pageId="7" pageNumber="8">type</typeStatus>
locality; GTW 7020b,
<specimenCount id="7CA7FD2828587F17F7DCFE3DB36CEEE1" box="[235,376,402,426]" pageId="7" pageNumber="8" type="generic">2 specimens</specimenCount>
, from the
<typeStatus id="B51A880328587F17F6DEFE3CB00DEEE0" box="[489,537,403,427]" pageId="7" pageNumber="8">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28587F17F7FEFE19B706EE85" box="[201,1298,438,463]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="6A1E36A128587F17F7FEFE19B706EE85" blockId="7.[151,1437,258,1219]" box="[201,1298,438,463]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7FEFE19B3E7EE84" bold="true" box="[201,499,438,463]" pageId="7" pageNumber="8">Distribution and habitat.</emphasis>
Known only from the
<typeStatus id="B51A880328587F17F5C4FE18B137EE84" box="[755,803,439,463]" pageId="7" pageNumber="8">type</typeStatus>
locality on limestone outcrops of El Choco.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28587F17F7FEFE75B691EC8D" pageId="7" pageNumber="8" type="description">
<paragraph id="6A1E36A128587F17F7FEFE75B691EC8D" blockId="7.[151,1437,258,1219]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7FEFE75B34EEEB8" bold="true" box="[201,346,474,499]" pageId="7" pageNumber="8">Description.</emphasis>
Shell large for genus (largest specimen,
<quantity id="AD599B4428587F17F47CFE75B1B3EEB9" box="[843,935,474,498]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="7" pageNumber="8" unit="mm" value="8.6">8.6 mm</quantity>
total length x
<quantity id="AD599B4428587F17F357FE75B6DDEEB9" box="[1120,1225,474,498]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="7" pageNumber="8" unit="mm" value="10.1">10.1 mm</quantity>
maximum width; smallest specimen,
<quantity id="AD599B4428587F17F643FE51B3D8ED5D" box="[372,460,510,534]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="7" pageNumber="8" unit="mm" value="6.6">6.6 mm</quantity>
total length x
<quantity id="AD599B4428587F17F545FE50B0DEED5D" box="[626,714,511,534]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="7" pageNumber="8" unit="mm" value="7.7">7.7 mm</quantity>
width;
<typeStatus id="B51A880328587F17F415FE51B193ED5D" box="[802,903,510,534]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
,
<quantity id="AD599B4428587F17F4A4FE51B1FEED5D" box="[915,1002,510,534]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="8" unit="mm" value="7.6">7.6 mm</quantity>
total length including peristome x
<quantity id="AD599B4428587F17F24EFE51B2D7ED71" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="7" pageNumber="8" unit="mm" value="8.9">8.9 mm</quantity>
total width including peristome), moderately high turbinate, whorls adnate except just before lip. Umbilicus wide, ca. 33% of total width, open to earliest whorls. Protoconch of 1.5–1.75 smooth, white or dark brown whorls with a white tip, but demarcation between protoconch and teleoconch not well-defined. Teleoconch of 2.75–3.0 whorls. Axial sculpture nearly obsolete, of numerous, crowded, low, microscopic lamellae, ca. 150 on last whorl. Spiral sculpture of ca. 50 low cords, occasionally of two sizes. Intersections of sculpture form nearly obsolete beads or scales. Whorls rounded with no indication of keels. Cords only slightly stronger in umbilicus. Suture narrowly, deeply channeled, finely serrate. Aperture double, circular (
<quantity id="AD599B4428587F17F496FD55B1ECEC59" box="[929,1016,762,786]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="7" pageNumber="8" unit="mm" value="4.3">4.3 mm</quantity>
maximum width in
<typeStatus id="B51A880328587F17F3EDFD55B754EC59" box="[1242,1344,762,786]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
), adnate with final whorl. Inner lip smooth, barely exserted. Outer lip expanded, slightly narrower facing the umbilicus, somewhat undulating, consisting of numerous, fine, fused lamellae, with a prominent auricle fused to previous whorl; outer part of lip somewhat cup-shaped, inner part perpendicular to whorl. Shell uniformly dirty white to dark reddish brown, outer lip white, shell without bands except on lip. Rare specimens white on top of whorls, brown on bottom. Operculum multispiral with an oblique, cup-shaped calcareous lamella.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28587F17F7FEFC7DB763EBD5" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="6A1E36A128587F17F7FEFC7DB057EB45" blockId="7.[151,1437,258,1219]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7FEFC7DB3FFECA0" bold="true" box="[201,491,978,1003]" pageId="7" pageNumber="8">Variation in specimens.</emphasis>
<specimenCount id="7CA7FD2828587F17F6CFFC7DB082ECA1" box="[504,662,978,1002]" pageId="7" pageNumber="8" type="generic">58 specimens</specimenCount>
seen. Specimens vary in color from white to brown, with some individuals half white and half brown.
</paragraph>
<paragraph id="6A1E36A128587F17F7FEFBB5B763EBD5" blockId="7.[151,1437,258,1219]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7FEFBB5B043EB78" bold="true" box="[201,599,1050,1075]" pageId="7" pageNumber="8">Comparison with other species.</emphasis>
This species differs from
<taxonomicName id="ADA14D2228587F17F494FBB4B63EEB78" box="[931,1066,1051,1075]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="sanchezi">
<emphasis id="58D5EAB328587F17F494FBB4B63EEB78" box="[931,1066,1051,1075]" italics="true" pageId="7" pageNumber="8">A. sanchezi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428587F17F377FBB5B6BCEB79" box="[1088,1192,1050,1074]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="7" pageNumber="8">Figure 4</figureCitation>
J–N) and
<taxonomicName id="ADA14D2228587F17F203FBB4B336EB1D" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="subSpecies" species="adolfi" subSpecies="peninsularis">
<emphasis id="58D5EAB328587F17F203FBB4B336EB1D" italics="true" pageId="7" pageNumber="8">A. adolfi peninsularis</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428587F17F618FB91B39BEB1D" box="[303,399,1086,1110]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="7" pageNumber="8">Figure 3</figureCitation>
J–L) in having nearly obsolete sculpture and in lacking the prominent spiral cords delimiting the umbilicus. It differs from the adjacent
<taxonomicName id="ADA14D2228587F17F549FBCCB0E2EB30" box="[638,758,1123,1147]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="mellosa">
<emphasis id="58D5EAB328587F17F549FBCCB0E2EB30" box="[638,758,1123,1147]" italics="true" pageId="7" pageNumber="8">A. mellosa</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428587F17F430FBCDB17EEB31" box="[775,874,1122,1146]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="7" pageNumber="8">Figure 1</figureCitation>
T) in its larger size and lack of prominent spiral cords in the umbilicus. It differs from
<taxonomicName id="ADA14D2228587F17F575FB29B0A3EBD5" box="[578,695,1158,1182]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="dichroa">
<emphasis id="58D5EAB328587F17F575FB29B0A3EBD5" box="[578,695,1158,1182]" italics="true" pageId="7" pageNumber="8">A. dichroa</emphasis>
</taxonomicName>
in having obsolete sculpture and an oblique opercular lamella.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28587F17F7FEFB04B1FFEB89" box="[201,1003,1194,1219]" pageId="7" pageNumber="8" type="etymology">
<paragraph id="6A1E36A128587F17F7FEFB04B1FFEB89" blockId="7.[151,1437,258,1219]" box="[201,1003,1194,1219]" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7FEFB04B344EB88" bold="true" box="[201,336,1195,1219]" pageId="7" pageNumber="8">Etymology.</emphasis>
L.
<emphasis id="58D5EAB328587F17F642FB04B3F9EB88" box="[373,493,1195,1219]" italics="true" pageId="7" pageNumber="8">tenebrosus</emphasis>
, dark, gloomy, in reference to the shell color.
</paragraph>
</subSubSection>
</treatment>
</document>

191
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<mods:title id="583055F013C338D85A5328F3F28A53D0">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B728587F18F7A0F955B71FEBA3" ID-DOI="http://doi.org/10.5281/zenodo.6164924" ID-GBIF-Taxon="127664005" ID-Zenodo-Dep="6164924" LSID="urn:lsid:plazi:treatment:E20887B728587F18F7A0F955B71FEBA3" httpUri="http://treatment.plazi.org/id/E20887B728587F18F7A0F955B71FEBA3" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="22BB652A28587F17F7A0F955B36AE87E" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="6A1E36A128587F17F7A0F955B112E85F" blockId="7.[151,774,1786,1845]" box="[151,774,1786,1812]" pageId="7" pageNumber="8">
<heading id="315681CD28587F17F7A0F955B112E85F" bold="true" box="[151,774,1786,1812]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="58D5EAB328587F17F7A0F955B112E85F" bold="true" box="[151,774,1786,1812]" pageId="7" pageNumber="8">
<taxonomicName id="ADA14D2228587F17F7A0F955B07BE85F" box="[151,623,1786,1812]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="paradoxa" status="sp. nov." subGenus="Gundlachtudora">
<emphasis id="58D5EAB328587F17F7A0F955B307E85F" bold="true" box="[151,275,1786,1812]" italics="true" pageId="7" pageNumber="8">Abbottella</emphasis>
(
<emphasis id="58D5EAB328587F17F613F955B3F8E85F" bold="true" box="[292,492,1786,1812]" italics="true" pageId="7" pageNumber="8">Gundlachtudora</emphasis>
)
<emphasis id="58D5EAB328587F17F6CAF955B07BE85F" bold="true" box="[509,623,1786,1812]" italics="true" pageId="7" pageNumber="8">paradoxa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C828587F17F541F955B112E85F" box="[630,774,1786,1812]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A128587F17F7A0F8B2B36AE87E" blockId="7.[151,774,1786,1845]" box="[151,382,1821,1845]" pageId="7" pageNumber="8">
<figureCitation id="F29A2A2428587F17F7A0F8B2B314E87E" box="[151,256,1821,1845]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="7" pageNumber="8">Figures 1</figureCitation>
Z–CC, 9 F
</paragraph>
</subSubSection>
<paragraph id="6A1E36A128587F17F7A0F8CAB313E8A2" pageId="7" pageNumber="8">
<table id="18A1C401285880EFF7A0F8CAB788E8A2" box="[151,1436,1893,2025]" gridcols="3" gridrows="4" pageId="7" pageNumber="8">
<tr id="D49134E3285880EFF7A0F8CAB788E835" box="[151,1436,1893,1918]" gridrow="0" pageId="7" pageNumber="8">
<th id="97405D9F285880EFF7A0F8CAB0DAE835" box="[151,718,1893,1918]" gridcol="0" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7A0F8CAB351E836" bold="true" box="[151,325,1893,1918]" pageId="7" pageNumber="8">Type material.</emphasis>
UF 456812, holotype; length, 7.8
</th>
<th id="97405D9F285880EFF5E2F8CAB19DE835" box="[725,905,1893,1918]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">mm; width, 9.0</th>
<th id="97405D9F285880EFF4A6F8CAB788E835" box="[913,1436,1893,1918]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">mm. UF 456813, 22 paratypes, from the type</th>
</tr>
<tr id="D49134E3285880EFF7A0F826B788E8EA" box="[151,1436,1929,1953]" gridrow="1" pageId="7" pageNumber="8" rowspan-1="1" rowspan-2="1">
<th id="97405D9F285880EFF7A0F826B0DAE8EA" box="[151,718,1929,1953]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">locality.</th>
</tr>
<tr id="D49134E3285880EFF7A0F802B788E88D" box="[151,1436,1965,1990]" gridrow="2" pageId="7" pageNumber="8">
<th id="97405D9F285880EFF7A0F802B0DAE88D" box="[151,718,1965,1990]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">
<emphasis id="58D5EAB328587F17F7FEF802B373E88D" bold="true" box="[201,359,1965,1990]" pageId="7" pageNumber="8">Type locality.</emphasis>
Dominican Republic, Santiago
</th>
<td id="97405D9F285880EFF5E2F802B19DE88D" box="[725,905,1965,1990]" gridcol="1" gridrow="2" pageId="7" pageNumber="8">Province, Loma</td>
<td id="97405D9F285880EFF4A6F802B788E88D" box="[913,1436,1965,1990]" gridcol="2" gridrow="2" pageId="7" pageNumber="8">Diego de Ocampo, at 1200 m. ca. 19.62° N, -</td>
</tr>
<tr id="D49134E3285880EFF7A0F87EB788E8A2" box="[151,1436,2001,2025]" gridrow="3" pageId="7" pageNumber="8" rowspan-1="1" rowspan-2="1">
<th id="97405D9F285880EFF7A0F87EB0DAE8A2" box="[151,718,2001,2025]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">70.76° W.</th>
</tr>
</table>
</paragraph>
<subSubSection id="22BB652A28577F18F7FEFF38B72DEFE4" box="[201,1337,151,176]" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="6A1E36A128577F18F7FEFF38B72DEFE4" blockId="8.[151,1437,151,1256]" box="[201,1337,151,176]" pageId="8" pageNumber="9">
<emphasis id="58D5EAB328577F18F7FEFF38B3EAEFFB" bold="true" box="[201,510,151,176]" pageId="8" pageNumber="9">Other material examined.</emphasis>
UF 216119, 20 weathered or immature specimens, from the
<typeStatus id="B51A880328577F18F390FF37B6C3EFFB" box="[1191,1239,152,176]" pageId="8" pageNumber="9">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28577F18F7FEFF14B1AEEE74" pageId="8" pageNumber="9" type="distribution">
<paragraph id="6A1E36A128577F18F7FEFF14B1AEEE74" blockId="8.[151,1437,151,1256]" pageId="8" pageNumber="9">
<emphasis id="58D5EAB328577F18F7FEFF14B3E0EF9F" bold="true" box="[201,500,187,212]" pageId="8" pageNumber="9">Distribution and habitat.</emphasis>
This species occurs at Loma Diego de Ocampo of the Cordillera Septentrional, on the mountain crest in dense forest with thick vegetative debris, mosses, and epiphytes. Loma Diego de Ocampo is located north of Santiago de los Caballeros and is the highest point of the Cordillera Septentrional at
<quantity id="AD599B4428577F18F201FEABB78CEE57" box="[1334,1432,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="m" value="1200.0">1,200 m</quantity>
. Snails were collected on limestone rocks at this peak. Locally common.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28577F18F7FEFEE4B3AAEC34" pageId="8" pageNumber="9" type="description">
<paragraph id="6A1E36A128577F18F7FEFEE4B3AAEC34" blockId="8.[151,1437,151,1256]" pageId="8" pageNumber="9">
<emphasis id="58D5EAB328577F18F7FEFEE4B34EEE2F" bold="true" box="[201,346,331,356]" pageId="8" pageNumber="9">Description.</emphasis>
Shell small (largest specimen,
<quantity id="AD599B4428577F18F58CFEE3B106EE2F" box="[699,786,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="8" pageNumber="9" unit="mm" value="7.9">7.9 mm</quantity>
total length x
<quantity id="AD599B4428577F18F481FEE3B619EE2F" box="[950,1037,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="8" pageNumber="9" unit="mm" value="9.1">9.1 mm</quantity>
total width; smallest specimen,
<quantity id="AD599B4428577F18F24EFEE3B2D7EECC" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="8" pageNumber="9" unit="mm" value="5.9">5.9 mm</quantity>
total length x
<quantity id="AD599B4428577F18F659FEC0B3D3EECC" box="[366,455,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="9" unit="mm" value="6.8">6.8 mm</quantity>
total width;
<typeStatus id="B51A880328577F18F56AFEC0B0D6EECC" box="[605,706,367,391]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
,
<quantity id="AD599B4428577F18F5F8FEC0B13CEECC" box="[719,808,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="8" pageNumber="9" unit="mm" value="7.8">7.8 mm</quantity>
total length including peristome x 9.0 mm total width including peristome), turbinate, whorls adnate except just before lip. Umbilicus wide, ca. 30% of total width, open to earliest whorls. Protoconch of ca. 1.5 erect whorls but demarcation between protoconch and teleoconch not welldefined. Teleoconch of 2.75–3.25 whorls. Axial sculpture of two sizes. The primary sculpture consists of 50–65 low, erect lamellae, widely spaced on the last ½ whorl, more closely spaced elsewhere. Secondary sculpture of 3– 10 microscopic lamellae between primary sculpture. Spiral sculpture of ca. 30 low, weak cords between suture and into umbilicus. Intersections of primary axial and spiral sculpture form minute, hollow points or scallops, often obsolete on base. Suture narrowly channeled, serrate. Aperture double, circular (
<quantity id="AD599B4428577F18F364FDC3B6A4EDCF" box="[1107,1200,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="9" unit="mm" value="4.2">4.2 mm</quantity>
maximum width in
<typeStatus id="B51A880328577F18F7A0FD20B2E9EDEC" box="[151,253,655,679]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
), solute from final whorl. Inner lip smooth, very narrowly exserted. Outer lip expanded, composed of numerous coarse lamellae, very narrow at the 11 o’clock position, then forming a broad auricle. Shell dirty white/ tan patterned with narrow, pale brown, spiral bands, apparent on both sides of outer lip. Operculum multispiral with a calcareous lamella; lamella arises vertically, arches over distally, and overlaps with previous whorl to form a continuous plate. The edge of this plate at the 11 o’clock position overlaps the lip and in most specimens the operculum cannot be withdrawn into the aperture. However, in a few specimens this edge has been broken off and the operculum withdrawn.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28577F18F7FEFC24B62DEBEB" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="6A1E36A128577F18F7FEFC24B1EBEC8C" blockId="8.[151,1437,151,1256]" pageId="8" pageNumber="9">
<emphasis id="58D5EAB328577F18F7FEFC24B3F7ECEF" bold="true" box="[201,483,907,932]" pageId="8" pageNumber="9">Variation in specimens.</emphasis>
<specimenCount id="7CA7FD2828577F18F6DBFC23B092ECEF" box="[492,646,908,932]" pageId="8" pageNumber="9" type="generic">43 specimens</specimenCount>
seen. Specimens are uniform in shell characteristics, varying only in the number of primary axial lamellae (50–65) and strength of the color bands.
</paragraph>
<paragraph id="6A1E36A128577F18F7FEFC7CB62DEBEB" blockId="8.[151,1437,151,1256]" pageId="8" pageNumber="9">
<emphasis id="58D5EAB328577F18F7FEFC7CB059ECA7" bold="true" box="[201,589,979,1004]" pageId="8" pageNumber="9">Comparison with other species.</emphasis>
The structure of the operculum separates this from all other
<taxonomicName id="ADA14D2228577F18F210FC7BB78FECA7" box="[1319,1435,980,1004]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328577F18F210FC7BB78FECA7" box="[1319,1435,980,1004]" italics="true" pageId="8" pageNumber="9">Abbottella</emphasis>
</taxonomicName>
except
<taxonomicName id="ADA14D2228577F18F7DEFC57B3C3EB5B" box="[233,471,1016,1040]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="bombardopolensis">
<emphasis id="58D5EAB328577F18F7DEFC57B3C3EB5B" box="[233,471,1016,1040]" italics="true" pageId="8" pageNumber="9">A. bombardopolensis</emphasis>
</taxonomicName>
from
<collectingCountry id="12B6763128577F18F528FC58B043EB44" box="[543,599,1015,1039]" name="Haiti" pageId="8" pageNumber="9">Haiti</collectingCountry>
and
<taxonomicName id="ADA14D2228577F18F5A6FC57B138EB5B" box="[657,812,1016,1040]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="decolorata">
<emphasis id="58D5EAB328577F18F5A6FC57B138EB5B" box="[657,812,1016,1040]" italics="true" pageId="8" pageNumber="9">A. decolorata</emphasis>
</taxonomicName>
from
<collectingCountry id="12B6763128577F18F443FC58B1A6EB44" box="[884,946,1015,1039]" name="Cuba" pageId="8" pageNumber="9">Cuba</collectingCountry>
, the only other recognized members of the subgenus
<taxonomicName id="ADA14D2228577F18F63DFBB3B3D7EB7F" box="[266,451,1052,1076]" class="Gastropoda" family="Pomatiidae" genus="Gundlachtudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328577F18F63DFBB3B3D7EB7F" box="[266,451,1052,1076]" italics="true" pageId="8" pageNumber="9">Gundlachtudora</emphasis>
</taxonomicName>
.
<taxonomicName id="ADA14D2228577F18F6E1FBB3B130EB7F" box="[470,804,1052,1076]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="bombardopolensis">
<emphasis id="58D5EAB328577F18F6E1FBB3B130EB7F" box="[470,804,1052,1076]" italics="true" pageId="8" pageNumber="9">Abbottella bombardopolensis</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428577F18F401FBB3B18EEB7F" box="[822,922,1052,1076]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="8" pageNumber="9">Figure 1</figureCitation>
DD) is known from Bombardopolis on the western-most tip of the northern Haitian peninsula to Terre Neuve in the Cordillera Centrale over
<quantity id="AD599B4428577F18F3DBFB90B756EB1C" box="[1260,1346,1087,1111]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="km" value="200.0">200 km</quantity>
west of
<taxonomicName id="ADA14D2228577F18F7A0FBCBB335EB37" box="[151,289,1124,1148]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="paradoxa">
<emphasis id="58D5EAB328577F18F7A0FBCBB335EB37" box="[151,289,1124,1148]" italics="true" pageId="8" pageNumber="9">A. paradoxa</emphasis>
</taxonomicName>
. Both species have nearly identical sculpture but the outer lip of
<taxonomicName id="ADA14D2228577F18F337FBCBB69FEB37" box="[1024,1163,1124,1148]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="paradoxa">
<emphasis id="58D5EAB328577F18F337FBCBB69FEB37" box="[1024,1163,1124,1148]" italics="true" pageId="8" pageNumber="9">A. paradoxa</emphasis>
</taxonomicName>
is widely expanded and notched; the outer lip of
<taxonomicName id="ADA14D2228577F18F69CFB27B08CEBEB" box="[427,664,1160,1184]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="bombardopolensis">
<emphasis id="58D5EAB328577F18F69CFB27B3A8EBEB" box="[427,444,1160,1184]" italics="true" pageId="8" pageNumber="9">A</emphasis>
.
<emphasis id="58D5EAB328577F18F6FDFB27B08CEBEB" box="[458,664,1160,1184]" italics="true" pageId="8" pageNumber="9">bombardopolensis</emphasis>
</taxonomicName>
is narrower but uniformly expanded.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28577F18F7FEFB03B71FEBA3" pageId="8" pageNumber="9" type="etymology">
<paragraph id="6A1E36A128577F18F7FEFB03B71FEBA3" blockId="8.[151,1437,151,1256]" pageId="8" pageNumber="9">
<emphasis id="58D5EAB328577F18F7FEFB03B344EB8F" bold="true" box="[201,336,1196,1220]" pageId="8" pageNumber="9">Etymology.</emphasis>
L.
<emphasis id="58D5EAB328577F18F6BDFB03B014EB8F" box="[394,512,1196,1220]" italics="true" pageId="8" pageNumber="9">paradoxus</emphasis>
, strange, contrary to expectation; the position of this species, whether in
<taxonomicName id="ADA14D2228577F18F7A0FB7FB344EBA3" box="[151,336,1232,1256]" class="Gastropoda" family="Pomatiidae" genus="Gundlachtudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328577F18F7A0FB7FB344EBA3" box="[151,336,1232,1256]" italics="true" pageId="8" pageNumber="9">Gundlachtudora</emphasis>
</taxonomicName>
or not, is problematic. It’s presence in
<collectingCountry id="12B6763128577F18F433FB7FB1BEEBA3" box="[772,938,1232,1256]" name="Dominican Republic" pageId="8" pageNumber="9">the Dominican</collectingCountry>
Republic was also unexpected.
</paragraph>
</subSubSection>
</treatment>
</document>

212
data/E2/08/87/E20887B728597F16F7A0FEC0B6B2E801.xml Normal file
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@ -0,0 +1,212 @@
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<mods:title id="0C6F4F332DBD4CD85AF0D428D2083D5B">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B728597F16F7A0FEC0B6B2E801" ID-DOI="http://doi.org/10.5281/zenodo.6164918" ID-GBIF-Taxon="127664002" ID-Zenodo-Dep="6164918" LSID="urn:lsid:plazi:treatment:E20887B728597F16F7A0FEC0B6B2E801" httpUri="http://treatment.plazi.org/id/E20887B728597F16F7A0FEC0B6B2E801" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection id="22BB652A28597F16F7A0FEC0B3A3EEE0" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="6A1E36A128597F16F7A0FEC0B085EEC2" blockId="6.[151,657,367,427]" box="[151,657,367,393]" pageId="6" pageNumber="7">
<heading id="315681CD28597F16F7A0FEC0B085EEC2" bold="true" box="[151,657,367,393]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="58D5EAB328597F16F7A0FEC0B085EEC2" bold="true" box="[151,657,367,393]" pageId="6" pageNumber="7">
<taxonomicName id="ADA14D2228597F16F7A0FEC0B3EEEEC2" box="[151,506,367,393]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="nitens" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB328597F16F7A0FEC0B307EEC2" bold="true" box="[151,275,367,393]" italics="true" pageId="6" pageNumber="7">Abbottella</emphasis>
(
<emphasis id="58D5EAB328597F16F613FEC0B3B4EEC2" bold="true" box="[292,416,367,393]" italics="true" pageId="6" pageNumber="7">Abbottella</emphasis>
)
<emphasis id="58D5EAB328597F16F687FEC0B3EEEEC2" bold="true" box="[432,506,367,393]" italics="true" pageId="6" pageNumber="7">nitens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C828597F16F536FEC0B085EEC2" box="[513,657,367,393]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A128597F16F7A0FE3DB3A3EEE0" blockId="6.[151,657,367,427]" box="[151,439,402,427]" pageId="6" pageNumber="7">
<figureCitation id="F29A2A2428597F16F7A0FE3DB314EEE1" box="[151,256,402,426]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="6" pageNumber="7">Figures 1</figureCitation>
P–S, 5 R,S, 8 A
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28597F16F7A0FE74B788EDC9" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="6A1E36A128597F16F7A0FE74B2E5ED5D" blockId="6.[151,1436,474,1867]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7A0FE74B351EEB8" bold="true" box="[151,325,474,499]" pageId="6" pageNumber="7">
<typeStatus id="B51A880328597F16F7A0FE74B2C4EEB8" box="[151,208,475,499]" pageId="6" pageNumber="7">Type</typeStatus>
material.
</emphasis>
UF 456806,
<typeStatus id="B51A880328597F16F6D5FE75B05CEEB9" box="[482,584,474,498]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
; length,
<quantity id="AD599B4428597F16F59CFE74B113EEB9" box="[683,775,475,498]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="6" pageNumber="7" unit="mm" value="7.7">7.7 mm</quantity>
; width,
<quantity id="AD599B4428597F16F452FE75B1D4EEB9" box="[869,960,474,498]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="7" unit="mm" value="7.8">7.8 mm</quantity>
. UF 456808, 15
<typeStatus id="B51A880328597F16F3B3FE74B6E6EEB8" box="[1156,1266,475,499]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
, from the
<typeStatus id="B51A880328597F16F25BFE74B788EEB8" box="[1388,1436,475,499]" pageId="6" pageNumber="7">type</typeStatus>
locality.
</paragraph>
<paragraph id="6A1E36A128597F16F7FEFD8CB313ED15" blockId="6.[151,1436,474,1867]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFD8CB37EED70" bold="true" box="[201,362,546,571]" pageId="6" pageNumber="7">
<typeStatus id="B51A880328597F16F7FEFD8CB310ED70" box="[201,260,547,571]" pageId="6" pageNumber="7">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B6763128597F16F644FD8DB075ED71" box="[371,609,546,570]" name="Dominican Republic" pageId="6" pageNumber="7">Dominican Republic</collectingCountry>
, La Altagracia Province,
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S of Higüey, at
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<quantity id="AD599B4428597F16F3A1FD8DB6F6ED71" box="[1174,1250,546,570]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="m" value="100.0">100 m</quantity>
.
</specimenCount>
ca.
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, -
<geoCoordinate id="0F95506628597F16F7A0FDE9B317ED15" box="[151,259,582,606]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="555" value="-68.71">68.71° W</geoCoordinate>
.
</paragraph>
<paragraph id="6A1E36A128597F16F7FEFDC5B788EDC9" blockId="6.[151,1436,474,1867]" box="[201,1436,618,643]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFDC5B3EAEDC8" bold="true" box="[201,510,618,643]" pageId="6" pageNumber="7">Other material examined.</emphasis>
UF 216167, 23 immature, broken, or weathered specimens, from the
<typeStatus id="B51A880328597F16F23CFDC4B72FEDC8" box="[1291,1339,619,643]" pageId="6" pageNumber="7">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28597F16F7FEFD21B63BEDA5" pageId="6" pageNumber="7" type="distribution">
<paragraph id="6A1E36A128597F16F7FEFD21B63BEDA5" blockId="6.[151,1436,474,1867]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFD21B3ECEDEC" bold="true" box="[201,504,654,679]" pageId="6" pageNumber="7">Distribution and habitat.</emphasis>
Known only from the
<typeStatus id="B51A880328597F16F432FD20B121EDEC" box="[773,821,655,679]" pageId="6" pageNumber="7">type</typeStatus>
locality under rocks among boulders, on a limestone bluff west of Highway 4. Further collecting may show that it generally occurs along this bluff, which runs parallel to the Rio Yuma on the west, from the
<typeStatus id="B51A880328597F16F57EFD78B06DEDA4" box="[585,633,727,751]" pageId="6" pageNumber="7">type</typeStatus>
locality to Boca de Yuma on the coast.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28597F16F7FEFD55B37CEBAD" pageId="6" pageNumber="7" type="description">
<paragraph id="6A1E36A128597F16F7FEFD55B37CEBAD" blockId="6.[151,1436,474,1867]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFD55B34EEC58" bold="true" box="[201,346,762,787]" pageId="6" pageNumber="7">Description.</emphasis>
Shell small (largest specimen,
<quantity id="AD599B4428597F16F58CFD55B106EC59" box="[699,786,762,786]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="7" unit="mm" value="8.5">8.5 mm</quantity>
total length x
<quantity id="AD599B4428597F16F481FD55B619EC59" box="[950,1037,762,786]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="7" unit="mm" value="8.6">8.6 mm</quantity>
total width; smallest specimen,
<quantity id="AD599B4428597F16F24EFD55B2D7EC7D" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="6" pageNumber="7" unit="mm" value="5.9">5.9 mm</quantity>
total length x
<quantity id="AD599B4428597F16F659FCB1B3D3EC7D" box="[366,455,798,822]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="7" unit="mm" value="6.1">6.1 mm</quantity>
total width;
<typeStatus id="B51A880328597F16F56AFCB1B0D6EC7D" box="[605,706,798,822]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
,
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total length including peristome x
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total width including peristome), turbinate, compact, whorls adnate. Umbilicus wide, ca. 30% of total width, open to earliest whorls. Protoconch of ca. 1.5 smooth whorls clearly demarcated from teleoconch. Teleoconch of 3–3.25 whorls. Axial sculpture of ca. 110 regularly spaced, low, fine lamellae, more widely spaced in umbilicus. Spiral sculpture of ca. 50 low, weak cords, slightly stronger in umbilicus. Intersections of axial and spiral sculpture form minute pustules or points, often nearly obsolete, elongated into blades at the suture but lamellae not fused. Suture very narrowly channeled, serrate. Aperture double, circular (
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maximum width in
<typeStatus id="B51A880328597F16F35CFC59B6C5EB45" box="[1131,1233,1014,1038]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
), adnate with the auricle joining the final whorl. Inner lip smooth, slightly exserted. Outer lip narrowly, uniformly expanded, composed of numerous lamellae, auriculate. Base color of shell uniformly white or metallic tan or bronze, usually with very faint, tan spots arranged in ca. 6 spiral rows. Bands present on both sides of the outer lip. Sculpture the same color as the base except near the suture where the blades and a few rows of sculpture may be white; sculpture within the umbilicus may be white as well. Operculum multispiral with a calcareous lamella, arising obliquely then becoming vertical.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28597F16F7FEFB5DB1CBEAA9" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="6A1E36A128597F16F7FEFB5DB08BEA65" blockId="6.[151,1436,474,1867]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFB5DB3F5EA40" bold="true" box="[201,481,1266,1291]" pageId="6" pageNumber="7">Variation in specimens.</emphasis>
<specimenCount id="7CA7FD2828597F16F6DEFB5DB096EA41" box="[489,642,1266,1290]" pageId="6" pageNumber="7" type="generic">39 specimens</specimenCount>
seen. Shells vary in color from brown to white; sculpture varies from minute pustules to being nearly imperceptible.
</paragraph>
<paragraph id="6A1E36A128597F16F7FEFA95B1CBEAA9" blockId="6.[151,1436,474,1867]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFA95B02AEA18" bold="true" box="[201,574,1338,1363]" pageId="6" pageNumber="7">Comparison with other species.</emphasis>
This species bears a striking resemblance to
<taxonomicName id="ADA14D2228597F16F304FA94B6BCEA18" box="[1075,1192,1339,1363]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="mellosa">
<emphasis id="58D5EAB328597F16F304FA94B6BCEA18" box="[1075,1192,1339,1363]" italics="true" pageId="6" pageNumber="7">A. mellosa</emphasis>
</taxonomicName>
from Sósua (
<figureCitation id="F29A2A2428597F16F20AFA95B788EA19" box="[1341,1436,1338,1362]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="6" pageNumber="7">Figure 1</figureCitation>
T), from which it differs in its more rounded, capacious whorls and greater number of spiral cords (ca.
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<taxonomicName id="ADA14D2228597F16F2B3FAF1B2FAEAD0" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="mellosa">
<emphasis id="58D5EAB328597F16F2B3FAF1B2FAEAD0" italics="true" pageId="6" pageNumber="7">A. mellosa</emphasis>
</taxonomicName>
vs. ca.
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<taxonomicName id="ADA14D2228597F16F6B5FA2CB3F0EAD0" box="[386,484,1411,1435]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="nitens">
<emphasis id="58D5EAB328597F16F6B5FA2CB3F0EAD0" box="[386,484,1411,1435]" italics="true" pageId="6" pageNumber="7">A. nitens</emphasis>
</taxonomicName>
). Both have the same obsolete sculpture and metallic sheen. However,
<taxonomicName id="ADA14D2228597F16F237FA2CB776EAD0" box="[1280,1378,1411,1435]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="nitens">
<emphasis id="58D5EAB328597F16F237FA2CB776EAD0" box="[1280,1378,1411,1435]" italics="true" pageId="6" pageNumber="7">A. nitens</emphasis>
</taxonomicName>
does not seem to be related to the other La Altagracia Province taxa, such as
<taxonomicName id="ADA14D2228597F16F4FBFA09B66AEAF5" box="[972,1150,1446,1470]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="milleacantha">
<emphasis id="58D5EAB328597F16F4FBFA09B66AEAF5" box="[972,1150,1446,1470]" italics="true" pageId="6" pageNumber="7">A. milleacantha</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A2428597F16F3B9FA09B6FBEAF5" box="[1166,1263,1446,1470]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="6" pageNumber="7">
Figure
<date id="1E1F106128597F16F3D6FA09B6FBEAF5" box="[1249,1263,1446,1470]" pageId="6" pageNumber="7">3</date>
</figureCitation>
X–BB), which are characterized by their small, compact shells and fine, prickly sculpture.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28597F16F7FEFA40B12CE86D" pageId="6" pageNumber="7" type="discussion">
<paragraph id="6A1E36A128597F16F7FEFA40B12CE86D" blockId="6.[151,1436,474,1867]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFA40B329E94C" bold="true" box="[201,317,1519,1543]" pageId="6" pageNumber="7">Remarks.</emphasis>
Like other
<taxonomicName id="ADA14D2228597F16F6E4FA41B05CE94D" box="[467,584,1518,1542]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB328597F16F6E4FA41B05CE94D" box="[467,584,1518,1542]" italics="true" pageId="6" pageNumber="7">Abbottella</emphasis>
</taxonomicName>
from the eastern part of the island, numerous examples bear signs of predation. Most have been broken into on the dorsal side of the final whorl (
<figureCitation id="F29A2A2428597F16F322F9BDB66CE961" box="[1045,1144,1554,1578]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="6" pageNumber="7">Figure 5</figureCitation>
R). A few, including the
<typeStatus id="B51A880328597F16F7A0F999B2E8E905" box="[151,252,1590,1614]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
, have a small, circular hole bored through the shell (
<figureCitation id="F29A2A2428597F16F463F999B1A2E905" box="[852,950,1590,1614]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="6" pageNumber="7">Figure 5</figureCitation>
S). Similar holes have been reported from other terrestrial snail shells and may be the result of predation by the larvae of drilid or lampyrid beetles (Harry, 1950). The
<typeStatus id="B51A880328597F16F629F9D1B394E9DD" box="[286,384,1662,1686]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
also has a web that nearly occludes the umbilicus (
<figureCitation id="F29A2A2428597F16F4ECF9D1B62AE9DD" box="[987,1086,1662,1686]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="6" pageNumber="7">Figure 1</figureCitation>
R). Within the umbilicus was found a pseudoscorpion, family
<taxonomicName id="ADA14D2228597F16F525F90DB061E9F1" box="[530,629,1698,1722]" class="Arachnida" family="Olpiidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Olpiidae</taxonomicName>
, subfamily Hesperolpiinae. The
<typeStatus id="B51A880328597F16F4C4F90DB641E9F1" box="[1011,1109,1698,1722]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
was the only recently dead specimen (operculum still in place) and no such pseudoscorpion association was found in any of the other
<specimenCount id="7CA7FD2828597F16F2B7F969B318E849" pageId="6" pageNumber="7" type="generic">42 specimens</specimenCount>
examined, which were all dead, mostly weathered specimens. It appears that the pseudoscorpion may have been commensal within the umbilicus of the live snail.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28597F16F7FEF89CB6B2E801" box="[201,1190,1842,1867]" pageId="6" pageNumber="7" type="etymology">
<paragraph id="6A1E36A128597F16F7FEF89CB6B2E801" blockId="6.[151,1436,474,1867]" box="[201,1190,1842,1867]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEF89CB35BE800" bold="true" box="[201,335,1843,1867]" pageId="6" pageNumber="7">Etymology.</emphasis>
L.
<emphasis id="58D5EAB328597F16F642F89BB3B9E800" box="[373,429,1844,1867]" italics="true" pageId="6" pageNumber="7">niteo</emphasis>
, shine, glitter, in reference to the metallic sheen of most specimens.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="E20887B7285A7F16F7A0FD8BB6C6EE57" ID-DOI="http://doi.org/10.5281/zenodo.6164916" ID-GBIF-Taxon="127664003" ID-Zenodo-Dep="6164916" LSID="urn:lsid:plazi:treatment:E20887B7285A7F16F7A0FD8BB6C6EE57" httpUri="http://treatment.plazi.org/id/E20887B7285A7F16F7A0FD8BB6C6EE57" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<paragraph id="6A1E36A1285A7F15F7A0FD8BB0B0ED76" blockId="5.[151,676,547,606]" box="[151,676,547,574]" pageId="5" pageNumber="6">
<heading id="315681CD285A7F15F7A0FD8BB0B0ED76" bold="true" box="[151,676,547,574]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="58D5EAB3285A7F15F7A0FD8BB0B0ED76" bold="true" box="[151,676,547,574]" pageId="5" pageNumber="6">
<taxonomicName id="ADA14D22285A7F15F7A0FD8BB019ED75" box="[151,525,547,574]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="dichroa" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3285A7F15F7A0FD8BB307ED75" bold="true" box="[151,275,548,574]" italics="true" pageId="5" pageNumber="6">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285A7F15F613FD8BB3B4ED75" bold="true" box="[292,416,548,574]" italics="true" pageId="5" pageNumber="6">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285A7F15F687FD8BB019ED75" bold="true" box="[432,525,548,574]" italics="true" pageId="5" pageNumber="6">dichroa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8285A7F15F523FD8CB0B0ED76" box="[532,676,547,573]" pageId="5" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A1285A7F15F7A0FDE9B3BEED15" blockId="5.[151,676,547,606]" box="[151,426,582,606]" pageId="5" pageNumber="6">
<figureCitation id="F29A2A24285A7F15F7A0FDE9B314ED15" box="[151,256,582,606]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="5" pageNumber="6">Figures 1</figureCitation>
K–O, 5 Q, 7 C
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285A7F15F7A0FD20B3EDEC8D" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="6A1E36A1285A7F15F7A0FD20B2E5ED81" blockId="5.[151,1437,654,2011]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7A0FD20B351EDEC" bold="true" box="[151,325,654,679]" pageId="5" pageNumber="6">
<typeStatus id="B51A8803285A7F15F7A0FD20B2C4EDEC" box="[151,208,655,679]" pageId="5" pageNumber="6">Type</typeStatus>
material.
</emphasis>
UF 456801,
<typeStatus id="B51A8803285A7F15F6D5FD21B053EDED" box="[482,583,654,678]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
; length,
<quantity id="AD599B44285A7F15F59BFD21B113EDED" box="[684,775,654,678]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="mm" value="6.5">6.5 mm</quantity>
; width,
<quantity id="AD599B44285A7F15F452FD21B1D5EDED" box="[869,961,654,678]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="5" pageNumber="6" unit="mm" value="9.9">9.9 mm</quantity>
. UF 216112, 31
<typeStatus id="B51A8803285A7F15F3B3FD20B6E6EDEC" box="[1156,1266,655,679]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
, from the
<typeStatus id="B51A8803285A7F15F25BFD20B788EDEC" box="[1388,1436,655,679]" pageId="5" pageNumber="6">type</typeStatus>
locality.
</paragraph>
<paragraph id="6A1E36A1285A7F15F7FEFD78B3FBEC59" blockId="5.[151,1437,654,2011]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEFD78B37CEDA4" bold="true" box="[201,360,726,751]" pageId="5" pageNumber="6">
<typeStatus id="B51A8803285A7F15F7FEFD78B316EDA4" box="[201,258,727,751]" pageId="5" pageNumber="6">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B67631285A7F15F645FD79B074EDA5" box="[370,608,726,750]" name="Dominican Republic" pageId="5" pageNumber="6">Dominican Republic</collectingCountry>
, Samaná Province, just E of El Limón,
<quantity id="AD599B44285A7F15F31BFD79B661EDA5" box="[1068,1141,726,750]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="km" value="10.0">10 km</quantity>
E of Las Terrenas, at sea level. ca.
<geoCoordinate id="0F955066285A7F15F635FD55B37EEC59" box="[258,362,762,786]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="555" value="19.29">19.29° N</geoCoordinate>
, -
<geoCoordinate id="0F955066285A7F15F64AFD55B3FFEC59" box="[381,491,762,786]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="555" value="-69.44">69.44° W</geoCoordinate>
.
</paragraph>
<paragraph id="6A1E36A1285A7F15F7FEFCB1B3EDEC8D" blockId="5.[151,1437,654,2011]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEFCB1B014EC7C" bold="true" box="[201,512,798,823]" pageId="5" pageNumber="6">Other material examined.</emphasis>
UF 216112, 4 broken, weathered, or immature specimens, from the
<typeStatus id="B51A8803285A7F15F23FFCB0B72CEC7C" box="[1288,1336,799,823]" pageId="5" pageNumber="6">type</typeStatus>
locality; UF 216116,
<specimenCount id="7CA7FD28285A7F15F605FCEDB34FEC11" box="[306,347,834,858]" pageId="5" pageNumber="6" type="generic">118</specimenCount>
specimens,
<quantity id="AD599B44285A7F15F6C2FCECB021EC11" box="[501,565,834,858]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="km" value="7.0">7 km</quantity>
E of Sánchez, Samaná Province,
<collectingCountry id="12B67631285A7F15F4EAFCEDB6C6EC11" box="[989,1234,834,858]" name="Dominican Republic" pageId="5" pageNumber="6">Dominican Republic</collectingCountry>
; UF 236229,
<specimenCount id="7CA7FD28285A7F15F2B6FCEDB31BEC35" pageId="5" pageNumber="6" type="generic">18 specimens</specimenCount>
,
<quantity id="AD599B44285A7F15F62BFCC9B371EC35" box="[284,357,870,894]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="6" unit="km" value="11.0">11 km</quantity>
E of Sánchez, Samaná Province,
<collectingCountry id="12B67631285A7F15F5DAFCC9B1C8EC35" box="[749,988,870,894]" name="Dominican Republic" pageId="5" pageNumber="6">Dominican Republic</collectingCountry>
; UF 216114,
<specimenCount id="7CA7FD28285A7F15F34DFCC9B70CEC35" box="[1146,1304,870,894]" pageId="5" pageNumber="6" type="generic">10 specimens</specimenCount>
,
<quantity id="AD599B44285A7F15F212FCC9B775EC35" box="[1317,1377,870,894]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="km" value="5.0">5 km</quantity>
E of Sánchez, Samaná Province,
<collectingCountry id="12B67631285A7F15F6DFFC25B0CFECE9" box="[488,731,906,930]" name="Dominican Republic" pageId="5" pageNumber="6">Dominican Republic</collectingCountry>
; Grego coll.,
<specimenCount id="7CA7FD28285A7F15F4B0FC25B604ECE9" box="[903,1040,906,930]" pageId="5" pageNumber="6" type="generic">1 specimen</specimenCount>
, Nagua, María
<collectingCountry id="12B67631285A7F15F3E6FC25B724ECE9" box="[1233,1328,906,930]" name="Trinidad and Tobago" pageId="5" pageNumber="6">Trinidad</collectingCountry>
Sánchez Province,
<collectingCountry id="12B67631285A7F15F63FFC01B3E1EC8D" box="[264,501,942,966]" name="Dominican Republic" pageId="5" pageNumber="6">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285A7F15F7FEFC7DB113EB1D" pageId="5" pageNumber="6" type="distribution">
<paragraph id="6A1E36A1285A7F15F7FEFC7DB113EB1D" blockId="5.[151,1437,654,2011]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEFC7DB3ECECA0" bold="true" box="[201,504,978,1003]" pageId="5" pageNumber="6">Distribution and habitat.</emphasis>
Known from the
<typeStatus id="B51A8803285A7F15F5FEFC7CB0EDECA0" box="[713,761,979,1003]" pageId="5" pageNumber="6">type</typeStatus>
locality on the north coast of the Sierra de Samaná of the Cordillera Septentrional to as far west as Nagua, also the region of Sánchez at three sites from
<quantity id="AD599B44285A7F15F3E2FC59B72CEB45" box="[1237,1336,1014,1038]" metricMagnitude="3" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7.0" pageId="5" pageNumber="6" unit="km" value="9.0" valueMax="11.0" valueMin="7.0">7–11 km</quantity>
E of the town along the south shore of the Samaná Peninsula. Individuals were found crawling on damp, algae-covered rocks on a shaded limestone hillside in a tropical forest.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285A7F15F7FEFBCDB6F6E939" pageId="5" pageNumber="6" type="description">
<paragraph id="6A1E36A1285A7F15F7FEFBCDB6F6E939" blockId="5.[151,1437,654,2011]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEFBCDB34EEB30" bold="true" box="[201,346,1122,1147]" pageId="5" pageNumber="6">Description.</emphasis>
Shell small (largest specimen,
<quantity id="AD599B44285A7F15F580FBCDB119EB31" box="[695,781,1122,1146]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="5" pageNumber="6" unit="mm" value="6.7">6.7 mm</quantity>
total length x
<quantity id="AD599B44285A7F15F499FBCDB605EB31" box="[942,1041,1122,1146]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="5" pageNumber="6" unit="mm" value="10.2">10.2 mm</quantity>
total width; smallest specimen,
<quantity id="AD599B44285A7F15F24EFBCDB2D7EBD5" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="6" unit="mm" value="5.3">5.3 mm</quantity>
total length x 8.0 mm total width;
<typeStatus id="B51A8803285A7F15F56AFB29B0D6EBD5" box="[605,706,1158,1182]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
,
<quantity id="AD599B44285A7F15F5F8FB29B13CEBD5" box="[719,808,1158,1182]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="mm" value="6.5">6.5 mm</quantity>
total length including peristome x
<quantity id="AD599B44285A7F15F388FB29B70CEBD5" box="[1215,1304,1158,1182]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="5" pageNumber="6" unit="mm" value="9.9">9.9 mm</quantity>
total width including peristome), turbinate, whorls adnate except just before lip. Umbilicus wide, ca. 37% of total width, open to earliest whorls. Protoconch of ca. 1.25 smooth whorls but demarcation between protoconch and teleoconch not well-defined. Teleoconch of 3–3.5 whorls. Axial sculpture of ca. 100 extremely fine lamellae of two strengths on the final whorl, more widely spaced in earlier whorls, often arranged in groups of 1–7 primary lamellae with irregular “empty” spaces interspersed that may be growth stops. Spiral sculpture of 30–36 low, weak cords, slightly stronger in umbilicus, occasionally stronger at periphery and elsewhere. Intersections of primary axial and spiral sculpture form minute elongated pustules or fine prickly sculpture; the sculpture appears more scalloped than serrate on the final whorl but often is more pronounced and prickly on earlier whorls. Suture narrowly channeled, serrate. Aperture double, circular (
<quantity id="AD599B44285A7F15F517FA65B06DEAA9" box="[544,633,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="6" unit="mm" value="4.4">4.4 mm</quantity>
maximum width in
<typeStatus id="B51A8803285A7F15F454FA65B1DEEAA9" box="[867,970,1482,1506]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
), solute or barely adnate with the auricle joining the final whorl, deflected to various degrees. Inner lip smooth, well-exserted. Outer lip expanded, composed of numerous lamellae, narrowest at the 10–11 o’clock position, auriculate. Base color of shell uniformly white or brown, rarely light tan, without any indication of bands on the whorls, although bands are apparent on both sides of outer lip. Sculpture white. Operculum multispiral with a calcareous, vertical lamella.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285A7F15F7FEF9D1B09EE891" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="6A1E36A1285A7F15F7FEF9D1B6C8E995" blockId="5.[151,1437,654,2011]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEF9D1B3F0E9DC" bold="true" box="[201,484,1662,1687]" pageId="5" pageNumber="6">Variation in specimens.</emphasis>
<specimenCount id="7CA7FD28285A7F15F6DBF9D1B081E9DD" box="[492,661,1662,1686]" pageId="5" pageNumber="6" type="generic">183 specimens</specimenCount>
seen. The most obvious variation is in color; this species occurs in two colors, white or brown, with little evidence of intergrades, although all have a white outer lip banded with brown. Specimens also vary in the degree of deflection of the aperture and the height of the spire.
</paragraph>
<paragraph id="6A1E36A1285A7F15F7FEF945B09EE891" blockId="5.[151,1437,654,2011]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEF945B02BE848" bold="true" box="[201,575,1770,1795]" pageId="5" pageNumber="6">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D22285A7F15F572F944B180E848" box="[581,916,1771,1795]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="subSpecies" species="adolfi" subSpecies="peninsularis">
<emphasis id="58D5EAB3285A7F15F572F944B180E848" box="[581,916,1771,1795]" italics="true" pageId="5" pageNumber="6">Abbottella adolfi peninsularis</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24285A7F15F495F945B616E849" box="[930,1026,1770,1794]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="5" pageNumber="6">Figure 3</figureCitation>
J–L) and
<taxonomicName id="ADA14D22285A7F15F345F944B6E4E848" box="[1138,1264,1771,1795]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="sanchezi">
<emphasis id="58D5EAB3285A7F15F345F944B6E4E848" box="[1138,1264,1771,1795]" italics="true" pageId="5" pageNumber="6">A. sanchezi</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24285A7F15F3C8F945B74BE849" box="[1279,1375,1770,1794]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="5" pageNumber="6">Figure 4</figureCitation>
J–N) have much finer, denser sculpture resembling sandpaper and the umbilicus of those species has two or more strong spiral cords;
<taxonomicName id="ADA14D22285A7F15F61DF89CB3B4E800" box="[298,416,1843,1867]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="dichroa">
<emphasis id="58D5EAB3285A7F15F61DF89CB3B4E800" box="[298,416,1843,1867]" italics="true" pageId="5" pageNumber="6">A. dichroa</emphasis>
</taxonomicName>
lacks these cords.
<taxonomicName id="ADA14D22285A7F15F54EF89CB17AE800" box="[633,878,1843,1867]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="sosuaensis">
<emphasis id="58D5EAB3285A7F15F54EF89CB17AE800" box="[633,878,1843,1867]" italics="true" pageId="5" pageNumber="6">Abbottella sosuaensis</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24285A7F15F44AF89DB1CAE801" box="[893,990,1842,1866]" captionStart="FIGURE 4. A – E" captionStartId="14.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@14.[151,1436,193,1582]" captionTargetPageId="14" captionText="FIGURE 4. A – E. Abbottella (Abbottella) rosaliae (Pfeiffer, 1858). A – C. UF 216191, 9.2 mm width. D. UF 216135, 9.9 mm width. E. UF 216125, 9.3 mm width. F – I. Abbottella (Abbottella) samanensis Bartsch, 1946. F – H. GTW 7085 c, 8.7 mm width. I. GTW 7085 d, 8.0 mm width. J – N. Abbottella (Abbottella) sanchezi Bartsch, 1946. J – L. UF 216154, 9.1 mm width. M. UF 216154, 8.1 mm width. N. UF 216154, 7.6 mm width. O – S. Abbottella (Abbottella) sosuaensis Bartsch, 1946. O – Q. UF 218136, 6.9 mm width. R. UF 218136, 7.7 mm width. S. GTW 9431 a, 7.6 mm width. T – X. Abbottella (Abbottella) wilhelmi (Pfeiffer, 1858). T – V. UF 216150, 8.5 mm width. W. UF 216150, 8 mm width. X. UF 216150, 7.6 mm width. Y – CC. Lagopoma lagopoma Bartsch, 1946. Y – AA. GTW 10081 a, 9.7 mm width. BB. GTW 10081 a, 7.6 mm width. CC. UF 216120, 7.7 mm width." httpUri="https://zenodo.org/record/283732/files/figure.png" pageId="5" pageNumber="6">Figure 4</figureCitation>
O–S) has fewer, almost obsolete, axial lamellae, the sculpture is weakly beaded rather than scalloped or prickly, the outer lip is narrower and deflected abaperturally. In
<taxonomicName id="ADA14D22285A7F15F660F8D4B3F2E8D8" box="[343,486,1915,1939]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="tenebrosa">
<emphasis id="58D5EAB3285A7F15F660F8D4B3F2E8D8" box="[343,486,1915,1939]" italics="true" pageId="5" pageNumber="6">A. tenebrosa</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24285A7F15F6C2F8D5B042E8D9" box="[501,598,1914,1938]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="5" pageNumber="6">Figure 1</figureCitation>
U–Y) the axial sculpture is obsolete and the opercular lamella is oblique. The peculiar sculpture of
<taxonomicName id="ADA14D22285A7F15F6FDF830B057E8FC" box="[458,579,1951,1975]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="dichroa">
<emphasis id="58D5EAB3285A7F15F6FDF830B057E8FC" box="[458,579,1951,1975]" italics="true" pageId="5" pageNumber="6">A. dichroa</emphasis>
</taxonomicName>
, arranged in “blocks” separated by smooth areas, is shared by
<emphasis id="58D5EAB3285A7F15F200F830B3A2E890" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="ADA14D22285A7F15F200F830B330E890" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="subSpecies" species="adolfi" subSpecies="peninsularis">A. adolfi peninsularis</taxonomicName>
,
<taxonomicName id="ADA14D22285A7F15F618F86CB3A5E890" box="[303,433,1987,2011]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="sanchezi">A. sanchezi</taxonomicName>
,
</emphasis>
and
<taxonomicName id="ADA14D22285A7F15F6DBF86CB097E890" box="[492,643,1987,2011]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="sosuaensis">
<emphasis id="58D5EAB3285A7F15F6DBF86CB097E890" box="[492,643,1987,2011]" italics="true" pageId="5" pageNumber="6">A. sosuaensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28597F16F7FEFF37B3B6EFBC" pageId="6" pageNumber="7" type="discussion">
<paragraph id="6A1E36A128597F16F7FEFF37B3B6EFBC" blockId="6.[151,1437,151,284]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFF37B329EFFB" bold="true" box="[201,317,152,176]" pageId="6" pageNumber="7">Remarks.</emphasis>
This species is peculiar in having two color morphs in the same population: white or brown. Like other species on the Samaná Peninsula, specimens have been broken through on the first half of the final whorl by a predator (
<figureCitation id="F29A2A2428597F16F621FF70B362EFBC" box="[278,374,223,247]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="6" pageNumber="7">Figure 5</figureCitation>
Q).
</paragraph>
</subSubSection>
<subSubSection id="22BB652A28597F16F7FEFEABB6C6EE57" box="[201,1234,260,284]" pageId="6" pageNumber="7" type="etymology">
<paragraph id="6A1E36A128597F16F7FEFEABB6C6EE57" blockId="6.[151,1437,151,284]" box="[201,1234,260,284]" pageId="6" pageNumber="7">
<emphasis id="58D5EAB328597F16F7FEFEABB344EE57" bold="true" box="[201,336,260,284]" pageId="6" pageNumber="7">Etymology.</emphasis>
Gr.
<emphasis id="58D5EAB328597F16F6B7FEABB382EE57" box="[384,406,260,284]" italics="true" pageId="6" pageNumber="7">di</emphasis>
-, two + Gr.
<emphasis id="58D5EAB328597F16F528FEABB04BEE57" box="[543,607,260,284]" italics="true" pageId="6" pageNumber="7">chroa</emphasis>
, color of the skin; in reference to the two color morphs.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="6EC878113A164BC7C1FED958F5E506AD">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B7285C7F15F7A0FBCBB0B3EE84" ID-DOI="http://doi.org/10.5281/zenodo.6164914" ID-GBIF-Taxon="127664001" ID-Zenodo-Dep="6164914" LSID="urn:lsid:plazi:treatment:E20887B7285C7F15F7A0FBCBB0B3EE84" httpUri="http://treatment.plazi.org/id/E20887B7285C7F15F7A0FBCBB0B3EE84" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<subSubSection id="22BB652A285C7F13F7A0FBCBB381EBD5" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="6A1E36A1285C7F13F7A0FBCBB0BAEB36" blockId="3.[151,686,1123,1182]" box="[151,686,1123,1150]" pageId="3" pageNumber="4">
<heading id="315681CD285C7F13F7A0FBCBB0BAEB36" bold="true" box="[151,686,1123,1150]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="58D5EAB3285C7F13F7A0FBCBB0BAEB36" bold="true" box="[151,686,1123,1150]" pageId="3" pageNumber="4">
<taxonomicName id="ADA14D22285C7F13F7A0FBCBB003EB35" box="[151,535,1123,1150]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="diadema" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3285C7F13F7A0FBCBB307EB35" bold="true" box="[151,275,1124,1150]" italics="true" pageId="3" pageNumber="4">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285C7F13F613FBCBB3B4EB35" bold="true" box="[292,416,1124,1150]" italics="true" pageId="3" pageNumber="4">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285C7F13F687FBCBB003EB35" bold="true" box="[432,535,1124,1150]" italics="true" pageId="3" pageNumber="4">diadema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8285C7F13F529FBCCB0BAEB36" box="[542,686,1123,1149]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A1285C7F13F7A0FB29B381EBD5" blockId="3.[151,686,1123,1182]" box="[151,405,1158,1182]" pageId="3" pageNumber="4">
<figureCitation id="F29A2A24285C7F13F7A0FB29B314EBD5" box="[151,256,1158,1182]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="3" pageNumber="4">Figures 1</figureCitation>
F–J, 5 P, 7 B
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285C7F13F7A0FB60B6CCEA19" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="6A1E36A1285C7F13F7A0FB60B2E5EA41" blockId="3.[151,1437,1230,1974]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7A0FB60B35CEBAC" bold="true" box="[151,328,1230,1255]" pageId="3" pageNumber="4">
<typeStatus id="B51A8803285C7F13F7A0FB60B2C6EBAC" box="[151,210,1231,1255]" pageId="3" pageNumber="4">Type</typeStatus>
material.
</emphasis>
UF 456814,
<typeStatus id="B51A8803285C7F13F6D2FB61B05FEBAD" box="[485,587,1230,1254]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
; length,
<quantity id="AD599B44285C7F13F586FB61B119EBAD" box="[689,781,1230,1254]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="4" unit="mm" value="5.1">5.1 mm</quantity>
; width, 6.0 mm. UF 456815, 5
<typeStatus id="B51A8803285C7F13F3B5FB60B6E4EBAC" box="[1154,1264,1231,1255]" pageId="3" pageNumber="4" type="paratype">paratypes</typeStatus>
, from the
<typeStatus id="B51A8803285C7F13F25BFB60B788EBAC" box="[1388,1436,1231,1255]" pageId="3" pageNumber="4">type</typeStatus>
locality.
</paragraph>
<paragraph id="6A1E36A1285C7F13F7FEFAB8B748EA65" blockId="3.[151,1437,1230,1974]" box="[201,1372,1302,1327]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7FEFAB8B371EA64" bold="true" box="[201,357,1302,1327]" pageId="3" pageNumber="4">
<typeStatus id="B51A8803285C7F13F7FEFAB8B316EA64" box="[201,258,1303,1327]" pageId="3" pageNumber="4">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B67631285C7F13F659FAB9B04DEA65" box="[366,601,1302,1326]" name="Dominican Republic" pageId="3" pageNumber="4">Dominican Republic</collectingCountry>
, Samaná Province, Cabo Cabrón, at
<specimenCount id="7CA7FD28285C7F13F4C4FAB9B655EA65" box="[1011,1089,1302,1326]" pageId="3" pageNumber="4" type="generic">
<quantity id="AD599B44285C7F13F4C4FAB9B629EA65" box="[1011,1085,1302,1326]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="4" unit="m" value="280.0">280 m</quantity>
.
</specimenCount>
ca.
<geoCoordinate id="0F955066285C7F13F359FAB9B6C2EA65" box="[1134,1238,1302,1326]" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="555" value="19.34">19.34° N</geoCoordinate>
, -
<geoCoordinate id="0F955066285C7F13F3DEFAB9B74CEA65" box="[1257,1368,1302,1326]" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="555" value="-69.25">69.25° W</geoCoordinate>
.
</paragraph>
<paragraph id="6A1E36A1285C7F13F7FEFA95B6CCEA19" blockId="3.[151,1437,1230,1974]" box="[201,1240,1338,1363]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7FEFA95B3EAEA18" bold="true" box="[201,510,1338,1363]" pageId="3" pageNumber="4">Other material examined.</emphasis>
UF 215985,
<specimenCount id="7CA7FD28285C7F13F5A6FA95B138EA19" box="[657,812,1338,1362]" pageId="3" pageNumber="4" type="generic">44 specimens</specimenCount>
, from the
<typeStatus id="B51A8803285C7F13F4AAFA94B1D9EA18" box="[925,973,1339,1363]" pageId="3" pageNumber="4">type</typeStatus>
locality (see Remarks).
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285C7F13F7FEFAF1B15DEAA8" pageId="3" pageNumber="4" type="distribution">
<paragraph id="6A1E36A1285C7F13F7FEFAF1B15DEAA8" blockId="3.[151,1437,1230,1974]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7FEFAF1B3ECEA3C" bold="true" box="[201,504,1374,1399]" pageId="3" pageNumber="4">Distribution and habitat.</emphasis>
Known only from the
<typeStatus id="B51A8803285C7F13F432FAF0B121EA3C" box="[773,821,1375,1399]" pageId="3" pageNumber="4">type</typeStatus>
locality. This region is on the northeastern tip of the Samaná Peninsula; it is part of the Sierra de Samaná of the Cordillera Septentrional and is locally known as Loma Travesada, which name appears on the original label. Specimens were found under limestone boulders on a slope in a virgin rain forest. This is a very isolated area of the country.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285C7F13F7FEFA41B0FBE8FD" pageId="3" pageNumber="4" type="description">
<paragraph id="6A1E36A1285C7F13F7FEFA41B0FBE8FD" blockId="3.[151,1437,1230,1974]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7FEFA41B34EE94C" bold="true" box="[201,346,1518,1543]" pageId="3" pageNumber="4">Description.</emphasis>
Shell very small (largest specimen,
<typeStatus id="B51A8803285C7F13F5C3FA41B14DE94D" box="[756,857,1518,1542]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
,
<quantity id="AD599B44285C7F13F453FA41B1AEE94D" box="[868,954,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="4" unit="mm" value="5.1">5.1 mm</quantity>
total length including peristome x 6.0 mm total width including peristome; smallest specimen,
<quantity id="AD599B44285C7F13F5DEF9BDB155E961" box="[745,833,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="4" unit="mm" value="3.1">3.1 mm</quantity>
total length x
<quantity id="AD599B44285C7F13F4DEF9BDB655E961" box="[1001,1089,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="4" unit="mm" value="3.7">3.7 mm</quantity>
total width), turbinate, whorls adnate except just before lip. Umbilicus wide, ca. 33% of total width, open to earliest whorls. Protoconch of ca. 1.5 minutely pustulose whorls but demarcation between protoconch and teleoconch not well-defined. Teleoconch of 2.5–2.75 whorls. Axial sculpture of numerous, crowded, low lamellae, slightly wider on spiral keels. Sculpture is more widely spaced on earliest whorls. Spiral sculpture present only as keels; 1–3 keels between the suture and the periphery; one on the periphery; one bounding the umbilicus;
<quantity id="AD599B44285C7F13F47EF969B19BE995" box="[841,911,1734,1758]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="7.62" metricValueMin="0.0" pageId="3" pageNumber="4" unit="in" value="1.5" valueMax="3.0" valueMin="0.0">0–3 in</quantity>
the umbilicus. The axial sculpture on the keels forms a very dense, finely serrate edge. On the whole the sculpture has a frosted aspect. Suture narrowly channeled, serrate. Aperture double, circular (
<quantity id="AD599B44285C7F13F52FF8A1B07BE86D" box="[536,623,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="4" unit="mm" value="2.7">2.7 mm</quantity>
maximum width in
<typeStatus id="B51A8803285C7F13F465F8A1B1ACE86D" box="[850,952,1806,1830]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
), solute from final whorl. Inner lip smooth, narrowly exserted. Outer lip slightly expanded, slightly narrower facing the umbilicus, consisting of numerous fused lamellae, with two auricles 180° apart. Axial sculpture does not extend onto outer lip. Shell uniformly dirty white/tan or patterned with vague brown, spiral bands, most apparent on both sides of outer lip. Operculum multispiral with an oblique, erect, calcareous lamella.
</paragraph>
</subSubSection>
<caption id="3EDE6629285B7F14F7A0F9ECB2CCE8C7" httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="4" pageNumber="5" targetBox="[151,1435,193,1582]" targetPageId="4">
<paragraph id="6A1E36A1285B7F14F7A0F9ECB2CCE8C7" blockId="4.[151,1437,1603,1932]" pageId="4" pageNumber="5">
<emphasis id="58D5EAB3285B7F14F7A0F9ECB300E912" bold="true" box="[151,276,1603,1626]" pageId="4" pageNumber="5">FIGURE 1.</emphasis>
A–E.
<taxonomicName id="ADA14D22285B7F14F660F9EBB0AAE911" box="[343,702,1604,1626]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="calliotropis" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3285B7F14F660F9EBB3D4E911" box="[343,448,1604,1626]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285B7F14F6E6F9EBB02EE911" box="[465,570,1604,1626]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285B7F14F57DF9EBB0AAE911" box="[586,702,1604,1626]" italics="true" pageId="4" pageNumber="5">calliotropis</emphasis>
</taxonomicName>
<emphasis id="58D5EAB3285B7F14F5F2F9EBB15DE911" bold="true" box="[709,841,1603,1626]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="43E657C8285B7F14F5F2F9EBB150E911" box="[709,836,1603,1626]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
A–C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F–J.
<taxonomicName id="ADA14D22285B7F14F4A4F9CCB6CEE932" box="[915,1242,1635,1657]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="diadema" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3285B7F14F4A4F9CCB1E8E932" box="[915,1020,1635,1657]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285B7F14F33CF9CCB660E932" box="[1035,1140,1635,1657]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285B7F14F3B4F9CCB6CEE932" box="[1155,1242,1635,1657]" italics="true" pageId="4" pageNumber="5">diadema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8285B7F14F3D7F9CCB748E932" box="[1248,1372,1634,1657]" pageId="4" pageNumber="5" rank="species">
<emphasis id="58D5EAB3285B7F14F3D7F9CCB748E932" bold="true" box="[1248,1372,1634,1657]" pageId="4" pageNumber="5">new species</emphasis>
</taxonomicNameLabel>
. F–H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K–O.
<taxonomicName id="ADA14D22285B7F14F7A0F90FB3C3E9FD" box="[151,471,1696,1718]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="dichroa" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3285B7F14F7A0F90FB314E9FD" box="[151,256,1696,1718]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285B7F14F627F90FB36DE9FD" box="[272,377,1696,1718]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285B7F14F6BEF90FB3C3E9FD" box="[393,471,1696,1718]" italics="true" pageId="4" pageNumber="5">dichroa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8285B7F14F6EAF90FB04EE9FD" box="[477,602,1695,1718]" pageId="4" pageNumber="5" rank="species">
<emphasis id="58D5EAB3285B7F14F6EAF90FB04EE9FD" bold="true" box="[477,602,1695,1718]" pageId="4" pageNumber="5">new species</emphasis>
</taxonomicNameLabel>
. K–M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P–T.
<taxonomicName id="ADA14D22285B7F14F58EF910B1F1E99E" box="[697,997,1727,1749]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="nitens" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3285B7F14F58EF910B136E99E" box="[697,802,1727,1749]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285B7F14F406F910B18EE99E" box="[817,922,1727,1749]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285B7F14F49EF96FB1F1E99E" box="[937,997,1728,1749]" italics="true" pageId="4" pageNumber="5">nitens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8285B7F14F4DBF910B67CE99E" box="[1004,1128,1726,1749]" pageId="4" pageNumber="5" rank="species">
<emphasis id="58D5EAB3285B7F14F4DBF910B67CE99E" bold="true" box="[1004,1128,1726,1749]" pageId="4" pageNumber="5">new species</emphasis>
</taxonomicNameLabel>
. P–R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808),8.3 mm width. T.
<taxonomicName id="ADA14D22285B7F14F398F972B3C8E85A" authority="Watters &amp; Duffy, 2010" authorityName="Watters &amp; Duffy" authorityYear="2010" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="mellosa" subGenus="Abbottella">
<emphasis id="58D5EAB3285B7F14F398F972B70CE9B8" box="[1199,1304,1757,1779]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285B7F14F21DF972B787E9B8" box="[1322,1427,1757,1779]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285B7F14F7A0F953B2F2E859" box="[151,230,1788,1810]" italics="true" pageId="4" pageNumber="5">mellosa</emphasis>
Watters &amp; Duffy, 2010
</taxonomicName>
. Holotype (UF 420729), 7.9 mm width. U–Y.
<taxonomicName id="ADA14D22285B7F14F48DF953B704E859" box="[954,1296,1788,1810]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="tenebrosa" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3285B7F14F48DF953B637E859" box="[954,1059,1788,1810]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285B7F14F304F953B689E859" box="[1075,1181,1788,1810]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285B7F14F39AF953B704E859" box="[1197,1296,1788,1810]" italics="true" pageId="4" pageNumber="5">tenebrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8285B7F14F22FF953B782E859" box="[1304,1430,1787,1810]" pageId="4" pageNumber="5" rank="species">
<emphasis id="58D5EAB3285B7F14F22FF953B782E859" bold="true" box="[1304,1430,1787,1810]" pageId="4" pageNumber="5">new species</emphasis>
</taxonomicNameLabel>
. U–W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z–CC.
<taxonomicName id="ADA14D22285B7F14F603F896B0DBE804" box="[308,719,1849,1871]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="paradoxa" status="sp. nov." subGenus="Gundlachtudora">
<emphasis id="58D5EAB3285B7F14F603F896B38AE804" box="[308,414,1849,1871]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285B7F14F685F896B04DE804" box="[434,601,1849,1871]" italics="true" pageId="4" pageNumber="5">Gundlachtudora</emphasis>
)
<emphasis id="58D5EAB3285B7F14F559F896B0DBE804" box="[622,719,1849,1871]" italics="true" pageId="4" pageNumber="5">paradoxa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8285B7F14F5ECF895B149E81B" box="[731,861,1849,1872]" pageId="4" pageNumber="5" rank="species">
<emphasis id="58D5EAB3285B7F14F5ECF895B149E81B" bold="true" box="[731,861,1849,1872]" pageId="4" pageNumber="5">new species</emphasis>
</taxonomicNameLabel>
. Z–BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD.
<taxonomicName id="ADA14D22285B7F14F565F8F7B6C6E825" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[594,1234,1880,1902]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="bombardopolensis" subGenus="Gundlachtudora">
<emphasis id="58D5EAB3285B7F14F565F8F7B0AFE825" box="[594,699,1880,1902]" italics="true" pageId="4" pageNumber="5">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285B7F14F5FCF8F7B166E825" box="[715,882,1880,1902]" italics="true" pageId="4" pageNumber="5">Gundlachtudora</emphasis>
)
<emphasis id="58D5EAB3285B7F14F4B6F8F7B62FE825" box="[897,1083,1880,1902]" italics="true" pageId="4" pageNumber="5">bombardopolensis</emphasis>
Bartsch, 1946
</taxonomicName>
. UF 33037, 7.0 mm width.
</paragraph>
</caption>
<subSubSection id="22BB652A285A7F15F7FEFF38B6EFEE2F" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="6A1E36A1285A7F15F7FEFF38B778EFBC" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEFF38B3F5EFFB" bold="true" box="[201,481,151,176]" pageId="5" pageNumber="6">Variation in specimens.</emphasis>
<specimenCount id="7CA7FD28285A7F15F6DFFF38B095EFE4" box="[488,641,151,175]" pageId="5" pageNumber="6" type="generic">50 specimens</specimenCount>
seen. The degree of coloration varies from uniformly white to banded with brown. The number of spiral keels (3–7) varies but there is always a keel between the suture and periphery, at the periphery, and bounding the umbilicus. There is a tendency for larger specimens to have more spiral cords.
</paragraph>
<paragraph id="6A1E36A1285A7F15F7FEFEACB6EFEE2F" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEFEACB055EE57" bold="true" box="[201,577,259,284]" pageId="5" pageNumber="6">Comparison with other species.</emphasis>
This species displays so many unique characteristics that it should likely be placed in its own genus. It is one of the smallest
<taxonomicName id="ADA14D22285A7F15F5FEFE87B12AEE0B" box="[713,830,296,320]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB3285A7F15F5FEFE87B12AEE0B" box="[713,830,296,320]" italics="true" pageId="5" pageNumber="6">Abbottella</emphasis>
</taxonomicName>
yet found and has the fewest number of whorls. The curious combination of tightly packed axial sculpture and spiral cords is unlike any other annulariid.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285A7F15F7FEFEDFB6B7EEE7" pageId="5" pageNumber="6" type="discussion">
<paragraph id="6A1E36A1285A7F15F7FEFEDFB6B7EEE7" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEFEDFB329EEC3" bold="true" box="[201,317,368,392]" pageId="5" pageNumber="6">Remarks.</emphasis>
The 44 additional specimens range from fresh dead to weathered specimens but all are broken in the same manner as
<taxonomicName id="ADA14D22285A7F15F666FE3BB3E4EEE7" box="[337,496,404,428]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="calliotropis">
<emphasis id="58D5EAB3285A7F15F666FE3BB3E4EEE7" box="[337,496,404,428]" italics="true" pageId="5" pageNumber="6">A. calliotropis</emphasis>
</taxonomicName>
(see Remarks there) by an unidentified predator (
<figureCitation id="F29A2A24285A7F15F32BFE3BB668EEE7" box="[1052,1148,404,428]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="5" pageNumber="6">Figure 5</figureCitation>
P).
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285A7F15F7FEFE17B0B3EE84" box="[201,679,439,464]" pageId="5" pageNumber="6" type="etymology">
<paragraph id="6A1E36A1285A7F15F7FEFE17B0B3EE84" blockId="5.[151,1436,151,464]" box="[201,679,439,464]" pageId="5" pageNumber="6">
<emphasis id="58D5EAB3285A7F15F7FEFE17B344EE9B" bold="true" box="[201,336,440,464]" pageId="5" pageNumber="6">Etymology.</emphasis>
Gr.
<emphasis id="58D5EAB3285A7F15F6B7FE17B3C6EE9B" box="[384,466,440,464]" italics="true" pageId="5" pageNumber="6">diadem</emphasis>
, headband, crown.
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="315681CD285D7F12F7A0FBA0B0D8EB62" bold="true" box="[151,716,1039,1065]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="58D5EAB3285D7F12F7A0FBA0B0D8EB62" bold="true" box="[151,716,1039,1065]" pageId="2" pageNumber="3">
Genus
<taxonomicName id="ADA14D22285D7F12F7D9FBA0B0D8EB62" authority="Henderson &amp; Bartsch, 1920" authorityName="Henderson &amp; Bartsch" authorityYear="1920" box="[238,716,1039,1065]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB3285D7F12F7D9FBA0B37EEB62" bold="true" box="[238,362,1039,1065]" italics="true" pageId="2" pageNumber="3">Abbottella</emphasis>
Henderson &amp; Bartsch, 1920
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</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A1285D7F12F7A0FBF9B0CEEB24" blockId="2.[151,730,1110,1135]" box="[151,730,1110,1135]" pageId="2" pageNumber="3">
<emphasis id="58D5EAB3285D7F12F7A0FBF9B0CEEB24" bold="true" box="[151,730,1110,1135]" pageId="2" pageNumber="3">
Subgenus
<taxonomicName id="ADA14D22285D7F12F626FBF9B0CEEB24" authority="Henderson &amp; Bartsch, 1920" authorityName="Henderson &amp; Bartsch" authorityYear="1920" box="[273,730,1110,1135]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="58D5EAB3285D7F12F626FBF9B393EB24" bold="true" box="[273,391,1110,1135]" italics="true" pageId="2" pageNumber="3">Abbottella</emphasis>
Henderson &amp; Bartsch, 1920
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="6A1E36A1285D7F12F7A0FB30B60DEBFD" blockId="2.[151,1436,1182,1350]" box="[151,1049,1182,1207]" pageId="2" pageNumber="3">
<typeStatus id="B51A8803285D7F12F7A0FB30B2DBEBFC" box="[151,207,1183,1207]" pageId="2" pageNumber="3">Type</typeStatus>
species by original designation
<taxonomicName id="ADA14D22285D7F12F501FB31B601EBFD" authority="Crosse, 1873" authorityName="Crosse" authorityYear="1873" box="[566,1045,1182,1206]" class="Gastropoda" family="Pomatiidae" genus="Chondropoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="moreletianum">
<emphasis id="58D5EAB3285D7F12F501FB31B16DEBFD" box="[566,889,1182,1206]" italics="true" pageId="2" pageNumber="3">Chondropoma moreletianum</emphasis>
Crosse, 1873
</taxonomicName>
.
</paragraph>
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<paragraph id="6A1E36A1285D7F12F7F0FB6DB012EA0D" blockId="2.[151,1436,1182,1350]" pageId="2" pageNumber="3">
<emphasis id="58D5EAB3285D7F12F7F0FB6DB355EB90" bold="true" box="[199,321,1218,1243]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Small (generally &lt;
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in width), depressed, widely umbilicate helicoid. Widely expanded, often fimbriated peristome. Axial and spiral sculpture usually present, usually forming cusps or serrations at their intersections. Multispiral operculum with erect, calcified spiral lamella. Radula with bicuspid inner marginal; inner cusp much larger than outer one.
</paragraph>
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</treatment>
</document>

238
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@ -0,0 +1,238 @@
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<mods:title id="25EAE7DC95546E1D5FEB74574CBD4291">New taxa and distributional notes on Abbottella and related taxa (Gastropoda: Littorinoidea: Annulariidae)</mods:title>
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<treatment id="E20887B7285D7F13F7A0FA35B10CEB44" ID-DOI="http://doi.org/10.5281/zenodo.6164912" ID-GBIF-Taxon="127663998" ID-Zenodo-Dep="6164912" LSID="urn:lsid:plazi:treatment:E20887B7285D7F13F7A0FA35B10CEB44" httpUri="http://treatment.plazi.org/id/E20887B7285D7F13F7A0FA35B10CEB44" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="22BB652A285D7F12F7A0FA35B3BCEA9D" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="6A1E36A1285D7F12F7A0FA35B0DAEAFF" blockId="2.[151,718,1434,1494]" box="[151,718,1434,1460]" pageId="2" pageNumber="3">
<heading id="315681CD285D7F12F7A0FA35B0DAEAFF" bold="true" box="[151,718,1434,1460]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="58D5EAB3285D7F12F7A0FA35B0DAEAFF" bold="true" box="[151,718,1434,1460]" pageId="2" pageNumber="3">
<taxonomicName id="ADA14D22285D7F12F7A0FA35B023EAFF" box="[151,567,1434,1460]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="calliotropis" status="sp. nov." subGenus="Abbottella">
<emphasis id="58D5EAB3285D7F12F7A0FA35B307EAFF" bold="true" box="[151,275,1434,1460]" italics="true" pageId="2" pageNumber="3">Abbottella</emphasis>
(
<emphasis id="58D5EAB3285D7F12F613FA35B3B4EAFF" bold="true" box="[292,416,1434,1460]" italics="true" pageId="2" pageNumber="3">Abbottella</emphasis>
)
<emphasis id="58D5EAB3285D7F12F687FA35B023EAFF" bold="true" box="[432,567,1434,1460]" italics="true" pageId="2" pageNumber="3">calliotropis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43E657C8285D7F12F509FA35B0DAEAFF" box="[574,718,1434,1460]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6A1E36A1285D7F12F7A0FA12B3BCEA9D" blockId="2.[151,718,1434,1494]" box="[151,424,1469,1494]" pageId="2" pageNumber="3">
<figureCitation id="F29A2A24285D7F12F7A0FA12B329EA9D" box="[151,317,1469,1494]" captionStart="FIGURE 1. A – E" captionStartId="4.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@4.[151,1436,193,1582]" captionTargetPageId="4" captionText="FIGURE 1. A – E. Abbottella (Abbottella) calliotropis new species. A – C. Holotype (UF 456810), 7.5 mm width. D. Paratype (UF 216131), 6.8 mm width. E. Paratype (UF 216131), 7.0 mm width. F – J. Abbottella (Abbottella) diadema new species. F – H. Holotype (UF 456814), 6.0 mm width. I. Paratype (UF 456815), 4.6 mm width. J. Paratype (UF 456815), 3.7 mm width. K – O. Abbottella (Abbottella) dichroa new species. K – M. Holotype (UF 456801), 9.9 mm width. N. Paratype (UF 216112), 8.8 mm width. O. Paratype (UF 216112), 9.4 mm width. P – T. Abbottella (Abbottella) nitens new species. P – R. Holotype (UF 456806), 7.8 mm width. Note pseudoscorpion web in umbilicus. S. Paratype (UF 456808), 8.3 mm width. T. Abbottella (Abbottella) mellosa Watters &amp; Duffy, 2010. Holotype (UF 420729), 7.9 mm width. U – Y. Abbottella (Abbottella) tenebrosa new species. U – W. Holotype (UF 456796), 8.9 mm width. X. Paratype (OSUM 37271), 8.6 mm width. Y. Paratype (OSUM 37271), 8.3 mm width. Z – CC. Abbottella (Gundlachtudora) paradoxa new species. Z – BB. Holotype (UF 456812), 9.0 mm width. CC. Paratype (UF 456813), 8.9 mm width. DD. Abbottella (Gundlachtudora) bombardopolensis Bartsch, 1946. UF 33037, 7.0 mm width." httpUri="https://zenodo.org/record/283729/files/figure.png" pageId="2" pageNumber="3">Figures 1 A–E</figureCitation>
,
<figureCitation id="F29A2A24285D7F12F67FFA12B342EA9E" box="[328,342,1469,1493]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="2" pageNumber="3">5</figureCitation>
O, 7 A
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285D7F12F7A0F9AAB6E2E99A" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="6A1E36A1285D7F12F7A0F9AAB2E5E90A" blockId="2.[151,1436,1541,2034]" pageId="2" pageNumber="3">
<emphasis id="58D5EAB3285D7F12F7A0F9AAB353E956" bold="true" box="[151,327,1541,1566]" pageId="2" pageNumber="3">
<typeStatus id="B51A8803285D7F12F7A0F9AAB2C4E956" box="[151,208,1541,1565]" pageId="2" pageNumber="3">Type</typeStatus>
material.
</emphasis>
UF 456810,
<typeStatus id="B51A8803285D7F12F6D5F9AAB05CE956" box="[482,584,1541,1565]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
; length,
<quantity id="AD599B44285D7F12F59CF9AAB113E956" box="[683,775,1541,1565]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="3" unit="mm" value="6.3">6.3 mm</quantity>
; width,
<quantity id="AD599B44285D7F12F452F9AAB1D4E956" box="[869,960,1541,1565]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="3" unit="mm" value="7.5">7.5 mm</quantity>
. UF 216131, 59
<typeStatus id="B51A8803285D7F12F3B3F9A9B6E6E955" box="[1156,1266,1542,1566]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
, from the
<typeStatus id="B51A8803285D7F12F25BF9A9B788E955" box="[1388,1436,1542,1566]" pageId="2" pageNumber="3">type</typeStatus>
locality.
</paragraph>
<paragraph id="6A1E36A1285D7F12F7FEF9E2B327E9C2" blockId="2.[151,1436,1541,2034]" pageId="2" pageNumber="3">
<emphasis id="58D5EAB3285D7F12F7FEF9E2B37CE92D" bold="true" box="[201,360,1613,1638]" pageId="2" pageNumber="3">
<typeStatus id="B51A8803285D7F12F7FEF9E2B310E92E" box="[201,260,1613,1637]" pageId="2" pageNumber="3">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="12B67631285D7F12F658F9E2B04FE92E" box="[367,603,1613,1637]" name="Dominican Republic" pageId="2" pageNumber="3">Dominican Republic</collectingCountry>
, La Altagracia Province, along Rio Yuna,
<quantity id="AD599B44285D7F12F300F9E2B664E92E" box="[1079,1136,1613,1637]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="km" value="2.0">2 km</quantity>
S of La Guana. ca.
<geoCoordinate id="0F955066285D7F12F266F9E2B2BAE9C2" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="555" value="18.79">18.79° N</geoCoordinate>
, -
<geoCoordinate id="0F955066285D7F12F7F6F9DEB33BE9C2" box="[193,303,1649,1673]" direction="west" orientation="longitude" pageId="2" pageNumber="3" precision="555" value="-68.67">68.67° W</geoCoordinate>
.
</paragraph>
<paragraph id="6A1E36A1285D7F12F7FEF93AB6E2E99A" blockId="2.[151,1436,1541,2034]" pageId="2" pageNumber="3">
<emphasis id="58D5EAB3285D7F12F7FEF93AB01DE9E5" bold="true" box="[201,521,1685,1710]" pageId="2" pageNumber="3">Other material examined.</emphasis>
UF 216131,
<specimenCount id="7CA7FD28285D7F12F59AF93AB127E9E6" box="[685,819,1685,1709]" pageId="2" pageNumber="3" type="juvenile">17 juvenile</specimenCount>
, broken, or weathered specimens, all from the
<typeStatus id="B51A8803285D7F12F25BF939B788E9E5" box="[1388,1436,1686,1710]" pageId="2" pageNumber="3">type</typeStatus>
locality; GTW 7085a,
<specimenCount id="7CA7FD28285D7F12F6A4F916B000E99A" box="[403,532,1721,1745]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
,
<quantity id="AD599B44285D7F12F528F916B04CE99A" box="[543,600,1721,1745]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="3" unit="km" value="8.0">8 km</quantity>
E of Higüey, La Altagracia Province,
<collectingCountry id="12B67631285D7F12F331F916B6E6E99A" box="[1030,1266,1721,1745]" name="Dominican Republic" pageId="2" pageNumber="3">Dominican Republic</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285D7F12F7FEF972B1CFE852" pageId="2" pageNumber="3" type="distribution">
<paragraph id="6A1E36A1285D7F12F7FEF972B1CFE852" blockId="2.[151,1436,1541,2034]" pageId="2" pageNumber="3">
<emphasis id="58D5EAB3285D7F12F7FEF972B3E0E9BD" bold="true" box="[201,500,1757,1782]" pageId="2" pageNumber="3">Distribution and habitat.</emphasis>
Probably along the eastern edge of the easternmost outcrops of the cordillera in La Altagracia Province. Specimens were found under rocks in adjacent fields.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285D7F13F7FEF88AB0B9EE84" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="6A1E36A1285D7F13F7FEF88AB0B9EE84" blockId="2.[151,1436,1541,2034]" lastBlockId="3.[151,1437,151,1040]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="58D5EAB3285D7F12F7FEF88AB34EE875" bold="true" box="[201,346,1829,1854]" pageId="2" pageNumber="3">Description.</emphasis>
Shell small (largest specimen, 7.0 mm total length x
<quantity id="AD599B44285D7F12F481F88AB619E876" box="[950,1037,1829,1853]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="2" pageNumber="3" unit="mm" value="8.3">8.3 mm</quantity>
total width; smallest specimen,
<quantity id="AD599B44285D7F12F24EF88AB2D7E82A" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="3" unit="mm" value="4.6">4.6 mm</quantity>
total length x
<quantity id="AD599B44285D7F12F659F8E6B3D3E82A" box="[366,455,1865,1889]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="3" unit="mm" value="5.5">5.5 mm</quantity>
total width;
<typeStatus id="B51A8803285D7F12F56AF8E6B0D6E82A" box="[605,706,1865,1889]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
,
<quantity id="AD599B44285D7F12F5F8F8E6B13CE82A" box="[719,808,1865,1889]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="3" unit="mm" value="6.3">6.3 mm</quantity>
total length including peristome x
<quantity id="AD599B44285D7F12F388F8E6B70CE82A" box="[1215,1304,1865,1889]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="3" unit="mm" value="7.5">7.5 mm</quantity>
total width including peristome), depressed turbinate, whorls adnate except just before lip. Umbilicus wide, ca. 33% of total width, open to earliest whorls. Protoconch of ca. 1.25–1.5 smooth whorls but demarcation between protoconch and teleoconch not well-defined. Teleoconch of 3–3.25 whorls. Axial sculpture of 64–90 threads only indicated by their sculpture over the spiral cords. Irregular “empty” spaces interspersed may be growth stops. Spiral sculpture of 10– 17 low, weak cords; generally 5–9 above the periphery, one peripheral, 4–7 below periphery into umbilicus. Peripheral and umbilical cords strongest. Intersections of primary axial and spiral sculpture form minute pustules or denticles, most developed as serrate denticles on sutural, peripheral, and umbilical cords, as minute pustules elsewhere. Suture deeply channeled, bounded by serrate cords above and below. Aperture double, circular (
<quantity id="AD599B44285C7F13F272FEABB788EE57" box="[1349,1436,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="4" unit="mm" value="3.5">3.5 mm</quantity>
maximum width in
<typeStatus id="B51A8803285C7F13F641FE88B3C8EE74" box="[374,476,295,319]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
), solute or barely adnate to final whorl. Inner lip smooth, moderately exserted. Outer lip expanded, composed of numerous lamellae, narrowest at the 11 o’clock position, slightly auriculate. Base color of shell white, tan, or dark brown patterned with diffuse, broken, brown spiral bands, often reduced to spots, bands apparent on both sides of outer lip; denticles and pustules white. Operculum multispiral with an calcareous lamella; lamella arises obliquely, then curves to vertical.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285C7F13F7FEFE74B316EC17" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="6A1E36A1285C7F13F7FEFE74B01BED5C" blockId="3.[151,1437,151,1040]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7FEFE74B3F6EEBF" bold="true" box="[201,482,475,500]" pageId="3" pageNumber="4">Variation in specimens.</emphasis>
<specimenCount id="7CA7FD28285C7F13F6DDFE73B097EEBF" box="[490,643,476,500]" pageId="3" pageNumber="4" type="generic">78 specimens</specimenCount>
seen. Shells vary in degree of coloration and number of axial threads (64–90) and spiral cords (10–17).
</paragraph>
<paragraph id="6A1E36A1285C7F13F7FEFD8CB316EC17" blockId="3.[151,1437,151,1040]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7FEFD8CB051ED77" bold="true" box="[201,581,547,572]" pageId="3" pageNumber="4">Comparison with other species.</emphasis>
<taxonomicName id="ADA14D22285C7F13F57AFD8BB15FED77" box="[589,843,548,572]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="calliotropis">
<emphasis id="58D5EAB3285C7F13F57AFD8BB15FED77" box="[589,843,548,572]" italics="true" pageId="3" pageNumber="4">Abbottella calliotropis</emphasis>
</taxonomicName>
is part of the group of small, turbinate species with fine, prickly or pustulose sculpture. They occur in the easternmost parts of the island. This species is most similar to
<taxonomicName id="ADA14D22285C7F13F784FDC3B335EDCF" box="[179,289,620,644]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="crossei">
<emphasis id="58D5EAB3285C7F13F784FDC3B335EDCF" box="[179,289,620,644]" italics="true" pageId="3" pageNumber="4">A. crossei</emphasis>
</taxonomicName>
(
<figureCitation id="F29A2A24285C7F13F618FDC3B39AEDCF" box="[303,398,620,644]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="3" pageNumber="4">Figure 3</figureCitation>
S–W) but the sculpture is not as prickly;
<taxonomicName id="ADA14D22285C7F13F46AFDC3B1DFEDCF" box="[861,971,620,644]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="crossei">
<emphasis id="58D5EAB3285C7F13F46AFDC3B1DFEDCF" box="[861,971,620,644]" italics="true" pageId="3" pageNumber="4">A. crossei</emphasis>
</taxonomicName>
has two keels on the body whorl whereas
<taxonomicName id="ADA14D22285C7F13F7A0FD3FB32DEDE3" box="[151,313,656,680]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="calliotropis">
<emphasis id="58D5EAB3285C7F13F7A0FD3FB32DEDE3" box="[151,313,656,680]" italics="true" pageId="3" pageNumber="4">A. calliotropis</emphasis>
</taxonomicName>
has only one.
<taxonomicName id="ADA14D22285C7F13F6DFFD3FB61DEDEC" authority="Bartsch, 1946" authorityName="Bartsch" authorityYear="1946" box="[488,1033,655,680]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="subSpecies" species="moreletiana" subSpecies="wetmorei">
<emphasis id="58D5EAB3285C7F13F6DFFD3FB14AEDE3" box="[488,862,656,680]" italics="true" pageId="3" pageNumber="4">Abbottella moreletiana wetmorei</emphasis>
Bartsch, 1946
</taxonomicName>
, from Pelican Key off the Samaná Peninsula, has similar coloration but has 39 strong axial lamellar ribs on the final whorl in comparison with the 64– 90 nearly obsolete axial threads in
<taxonomicName id="ADA14D22285C7F13F52CFD77B0AEEDBB" box="[539,698,728,752]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="calliotropis">
<emphasis id="58D5EAB3285C7F13F52CFD77B0AEEDBB" box="[539,698,728,752]" italics="true" pageId="3" pageNumber="4">A. calliotropis</emphasis>
</taxonomicName>
.
<taxonomicName id="ADA14D22285C7F13F5FFFD77B1D7EDBB" box="[712,963,728,752]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="calliotropis">
<emphasis id="58D5EAB3285C7F13F5FFFD77B1D7EDBB" box="[712,963,728,752]" italics="true" pageId="3" pageNumber="4">Abbottella calliotropis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="ADA14D22285C7F13F361FD77B783EDA4" authority="Pfeiffer, 1850" authorityName="Pfeiffer" authorityYear="1850" box="[1110,1431,727,752]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="tentorium">
<emphasis id="58D5EAB3285C7F13F361FD77B6F6EDBB" box="[1110,1250,728,752]" italics="true" pageId="3" pageNumber="4">A. tentorium</emphasis>
(Pfeiffer, 1850)
</taxonomicName>
,
<taxonomicName id="ADA14D22285C7F13F7A0FD53B3A1EC5F" authority="Watters, 2012" authorityName="Watters" authorityYear="2012" box="[151,437,764,788]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="urbana">
<emphasis id="58D5EAB3285C7F13F7A0FD53B31EEC5F" box="[151,266,764,788]" italics="true" pageId="3" pageNumber="4">A. urbana</emphasis>
Watters, 2012
</taxonomicName>
,
<taxonomicName id="ADA14D22285C7F13F6F3FD53B0C6EC5F" authority="Watters, 2010" authorityName="Watters" authorityYear="2010" box="[452,722,764,788]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="aenea">
<emphasis id="58D5EAB3285C7F13F6F3FD53B03DEC5F" box="[452,553,764,788]" italics="true" pageId="3" pageNumber="4">A. aenea</emphasis>
Watters, 2010
</taxonomicName>
(
<figureCitation id="F29A2A24285C7F13F5D3FD53B153EC5F" box="[740,839,764,788]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="3" pageNumber="4">Figure 3</figureCitation>
N–R), and
<taxonomicName id="ADA14D22285C7F13F4E3FD53B788EC5F" authority="Watters &amp; Duffy, 2010" authorityName="Watters &amp; Duffy" authorityYear="2010" box="[980,1436,764,788]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="milleacantha">
<emphasis id="58D5EAB3285C7F13F4E3FD53B69CEC5F" box="[980,1160,764,788]" italics="true" pageId="3" pageNumber="4">A. milleacantha</emphasis>
Watters &amp; Duffy, 2010
</taxonomicName>
(
<figureCitation id="F29A2A24285C7F13F7A9FCB0B310EC7C" box="[158,260,799,823]" captionStart="FIGURE 3. A – E" captionStartId="11.[151,250,1603,1626]" captionTargetBox="[151,1435,193,1582]" captionTargetId="figure@11.[151,1436,193,1582]" captionTargetPageId="11" captionText="FIGURE 3. A – E. Abbottella (Abbottella) abbotti Bartsch, 1946. A – C. UF 215980, 5.6 mm width. D. UF 215983, 6.0 mm width. E. UF 215981, 5.2 mm width. F. Choanopoma adolfi Pfeiffer, 1852. Pfeiffer, 1854: pl. 48, fig. 8. G – I. Abbottella (Abbottella) cf. adolfi adolfi (Pfeiffer, 1852). GTW 7020 c, 5 mm width. J – L. Abbottella (Abbottella) adolfi peninsularis Bartsch, 1946. GTW 9430 a, 7.8 mm width. M. Abbottella (Abbottella) newcombi (Crosse, 1873). Syntype (MNHN 5436), 5.7 mm length. Photo courtesy of MNHN. N – R. Abbottella (Abbottella) aenea Watters, 2010. N – P. GTW 14181 a, 6.5 mm width. Q. GTW 14181 a, 6.1 mm width. R. Paratype (OSUM 35490), 6.0 mm width. S – W. Abbottella (Abbottella) crossei (Pilsbry, 1933). S – U. GTW 9432 a, 7.3 mm width. V. UF 216146, 7.6 mm width. W. UF 216146, 7.2 mm width. X – BB. Abbottella (Abbottella) milleacantha Watters &amp; Duffy, 2010. X – Z. Holotype (UF 420728), 7.5 mm width. AA. Grego, 7.4 mm width. BB. Grego, 8.1 mm width." httpUri="https://zenodo.org/record/283731/files/figure.png" pageId="3" pageNumber="4">
Figure
<date id="1E1F1061285C7F13F7C1FCB0B310EC7C" box="[246,260,799,823]" pageId="3" pageNumber="4">3</date>
</figureCitation>
X–BB), in having fewer spiral cords, more pronounced denticles, and a peripheral keel of larger denticles.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285C7F13F7FEFCC7B088ECA7" pageId="3" pageNumber="4" type="discussion">
<paragraph id="6A1E36A1285C7F13F7FEFCC7B088ECA7" blockId="3.[151,1437,151,1040]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7FEFCC7B329ECCB" bold="true" box="[201,317,872,896]" pageId="3" pageNumber="4">Remarks.</emphasis>
Some specimens have been damaged in the same manner—the shell is broken open on the first half of the final whorl (
<figureCitation id="F29A2A24285C7F13F65FFC23B3DCECEF" box="[360,456,908,932]" captionStart="FIGURE 5. A – F" captionStartId="18.[151,250,1786,1809]" captionTargetBox="[151,1435,375,1764]" captionTargetId="figure@18.[151,1436,375,1764]" captionTargetPageId="18" captionText="FIGURE 5. A – F. Rolleia haitensis Bartsch, 1946. A – C. Holotype (USNM 504088), 12.0 mm width. Photo courtesy of USNM. D. UF 32777, 15.4 mm width. E. UF 32777, 13.7 mm width. F. UF 32863, 13.2 mm width. G – N. Rolleia oberi Watters &amp; Duffy, 2010. G – I. Holotype (UF 434775), 10.3 mm width. J. UF 236228, 10.2 mm width. K. UF 216195, 11.0 mm width. L. UF 216195, 11.5 mm width. M. UF 216195, 10.7 mm width. N. UF 216195, 11.3 mm width. O. Abbottella (Abbottella) calliotropis new species. UF 216131, 7.1 mm width. Note predation damage. P. Abbottella (Abbottella) diadema new species. UF 215985, 5.0 mm width. Note predation damage. Q. Abbottella (Abbottella) dichroa new species. UF 216112, 8.3 mm width. Note predation damage. R, S. Abbottella (Abbottella) nitens new species. R. UF 216167, 7.9 mm width. Note predation damage. S. Holotype (UF 456806), 7.8 mm width. Note predation damage." httpUri="https://zenodo.org/record/283733/files/figure.png" pageId="3" pageNumber="4">Figure 5</figureCitation>
O). This suggests that all were killed by the same
<typeStatus id="B51A8803285C7F13F4C9FC22B63AECEE" box="[1022,1070,909,933]" pageId="3" pageNumber="4">type</typeStatus>
of predator. This same breakage is seen in
<taxonomicName id="ADA14D22285C7F13F63DFC1FB39FEC83" box="[266,395,944,968]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="diadema">
<emphasis id="58D5EAB3285C7F13F63DFC1FB39FEC83" box="[266,395,944,968]" italics="true" pageId="3" pageNumber="4">A. diadema</emphasis>
</taxonomicName>
,
<taxonomicName id="ADA14D22285C7F13F6ADFC1FB005EC83" box="[410,529,944,968]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="dichroa">
<emphasis id="58D5EAB3285C7F13F6ADFC1FB005EC83" box="[410,529,944,968]" italics="true" pageId="3" pageNumber="4">A. dichroa</emphasis>
</taxonomicName>
, and
<emphasis id="58D5EAB3285C7F13F566FC1FB0AFEC83" box="[593,699,944,968]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="ADA14D22285C7F13F566FC1FB0A3EC83" box="[593,695,944,968]" class="Gastropoda" family="Pomatiidae" genus="Abbottella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="nitens">A. nitens</taxonomicName>
.
</emphasis>
It is curious that this damage seems to be limited to the Samaná Peninsula and eastern La Altagracia Province.
</paragraph>
</subSubSection>
<subSubSection id="22BB652A285C7F13F7FEFC57B10CEB44" box="[201,792,1015,1040]" pageId="3" pageNumber="4" type="etymology">
<paragraph id="6A1E36A1285C7F13F7FEFC57B10CEB44" blockId="3.[151,1437,151,1040]" box="[201,792,1015,1040]" pageId="3" pageNumber="4">
<emphasis id="58D5EAB3285C7F13F7FEFC57B344EB5B" bold="true" box="[201,336,1016,1040]" pageId="3" pageNumber="4">Etymology.</emphasis>
Gr.
<emphasis id="58D5EAB3285C7F13F6B7FC57B3D7EB5B" box="[384,451,1016,1040]" italics="true" pageId="3" pageNumber="4">kallos</emphasis>
, beauty + Gr.
<emphasis id="58D5EAB3285C7F13F553FC56B08AEB5B" box="[612,670,1017,1040]" italics="true" pageId="3" pageNumber="4">trope</emphasis>
, a turning.
</paragraph>
</subSubSection>
</treatment>
</document>