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+
+
+
+Ancistrocerus shibuyai (Yasumatsu, 1938), stat. restit. (Hymenoptera: Vespidae: Eumeninae), an overlooked trans-Palaearctic species
+
+
+
+Author
+
+Fateryga, Alexander V.
+T. I. Vyazemsky Karadag Scientific Station-Nature Reserve of RAS-Branch of A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, Nauki Str. 24, Kurortnoye, 298188 Feodosiya, Russia.
+
+
+
+Author
+
+Proshchalykin, Maxim Yu.
+Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, 100 - let Vladivostoku Ave. 159, 690022 Vladivostok, Russia.
+
+text
+
+
+Zootaxa
+
+
+2024
+
+2024-11-12
+
+
+5537
+
+
+4
+
+
+527
+540
+
+
+
+
+http://dx.doi.org/10.11646/zootaxa.5537.4.5
+
+journal article
+10.11646/zootaxa.5537.4.5
+1175-5326
+14240360
+F2B1D954-3B04-49F6-8FF3-A124DEC913B2
+
+
+
+
+
+
+
+Ancistrocerus shibuyai
+(
+Yasumatsu, 1938
+)
+
+, stat. restit.
+
+
+
+
+
+
+(
+Figs 1A, B, E, G
+,
+2A, B, E, F
+)
+
+
+
+
+
+Odynerus shibuyai
+Yasumatsu, 1938
+
+[March]: 83,
+♀
+♂
+
+
+
+
+
+(type locality: “
+Osaka
+,
+Honshû
+” [
+Japan
+]);
+holotype
+,
+♂
+, Kyushu University, Fukuoka, Japan
+
+.
+
+
+
+
+
+
+Ancistrocerus shibuyai
+
+:
+
+van der Vecht & Fischer 1972: 117
+
+.
+
+
+
+
+
+Ancistrocerus trifasciatus shibuyai
+
+:
+
+Giordani Soika 1986: 147
+
+;
+
+Yamane 1990: 104
+
+, figs 184, 193, 199, 203, 204, 215;
+
+Kurzenko 2004: 203
+
+;
+
+Kim & Yamane 2009: 41
+
+, figs 5A–F.
+
+
+
+
+Ancistrocerus trifasciatus
+var.
+moeschleri
+Blüthgen, 1938
+
+[May]: 284,
+♀
+♂
+(type locality: “Klein Guja (Ostpreussen)” [
+Poland
+]);
+holotype
+,
+♀
+, Museum für Naturkunde,
+Berlin
+,
+Germany
+. Available under Art. 45.6.4 of
+ICZN (1999)
+.
+Syn. nov.
+
+
+
+
+Odynerus trifasciatus orientalis
+Kostylev, 1938: 305
+
+
+,
+♀
+♂
+(type locality: “
+Jakovlevka
+,
+Ussuri
+” [
+Russia
+,
+Primorskiy
+Territory
+]), nom. praeocc., nec
+
+O. orientalis
+Dalla Torre, 1889
+
+;
+lectotype
+(designated here),
+♀
+,
+ZISP
+.
+
+
+
+Ancistrocerus trifasciatus kostylevi
+van der Vecht
+
+in
+
+van der Vecht & Fischer 1972: 117
+
+, replacement name for
+
+Odynerus trifasciatus orientalis
+(
+Kostylev, 1938
+)
+
+, nec
+
+O. orientalis
+Dalla Torre, 1889
+
+; synonimized by
+
+Kurzenko 2004: 203
+
+.
+
+
+
+Ancistrocerus balticus
+Budrys & Orlovskytė
+
+in
+
+Budrys
+et al.
+2023
+
+, Supplement 3: 1,
+♀
+♂
+(type locality: “
+Lithuania
+,
+Alytus district
+, Punios šilas forest, 54°30ʹ48ʹʹN 24°02ʹ22ʹʹE”);
+holotype
+,
+♀
+,
+Nature Research Centre
+,
+Vilnius
+,
+Lithuania
+.
+Syn. nov
+.
+
+
+
+
+Differential diagnosis.
+Females of this species can be distinguished from the closely related
+
+A. trifasciatus
+
+by S
+2 in
+lateral view distinctly convex (
+Fig. 1E
+; vs. weakly convex to nearly flat in
+
+A. trifasciatus
+
+,
+Fig. 1F
+). Males can be distinguished by ventral lobe of the aedeagus apically rounded (
+Fig. 1G
+; vs. rather triangularly pointed in
+
+A. trifasciatus
+
+,
+Fig. 1H
+). An additional character is the coloration of the hind tibia: usually completely black in the female (
+Fig. 2A
+) (very rarely with small yellow spot,
+Fig. 2B
+) and completely black (
+Fig. 2E
+) to at least black dorsally (
+Fig. 2F
+) in the male. Other characters mentioned in the key by
+
+Budrys
+et al.
+(2023
+
+, Supplement 3) can be also applied but they are also variable and therefore less diagnostic.
+
+
+
+
+Remarks.
+The identity of
+
+Odynerus shibuyai
+
+is understood by us following
+Yamane (1990)
+and
+Kim & Yamane (2009)
+; the
+holotype
+was not available for study. It is absent from the drawer with the type specimens of taxa described by K. Yasumatsu in the Kyushu University and may be deposited among unsorted material (Toshiharu Mita, personal communication).
+
+
+
+Ancistrocerus trifasciatus moeschleri
+
+was placed by
+van der Vecht & Fischer (1972)
+among synonyms of
+
+A. trifasciatus trifasciatus
+
+following
+Blüthgen (1961)
+. The
+holotype
+of
+
+A. trifasciatus moeschleri
+
+was not available for our study; however, its diagnosis, particularly completely black mid and hind legs in the female (
+Blüthgen 1938
+), suggests that it is conspecific rather with
+
+A. shibuyai
+
+; therefore, we hereby recognize
+
+A. trifasciatus moeschleri
+
+as a synonym of
+
+A. shibuyai
+
+, and not as a synonym of
+
+A. trifasciatus
+
+.
+
+Budrys
+et al.
+(2023
+
+, Supplement 3: 6) supposed that
+
+A. trifasciatus moeschleri
+
+was a melanistic variety of
+
+A. trifasciatus
+
+; however, this is not so probable because there are no specific conditions (e.g., mountains with high elevation) in the type locality of
+
+A. trifasciatus moeschleri
+
+.
+
+Odynerus shibuyai
+
+,
+
+A. trifasciatus moeschleri
+
+, and
+
+O. trifasciatus orientalis
+
+were described in the same year. We do not know the exact date of publication for
+Kostylev’s (1938)
+paper but his name is anyway invalid due to the homonymy with
+
+O. orientalis
+Dalla Torre, 1889
+
+(Art. 57.2 of
+ICZN 1999
+). Among two remaining names,
+Yasumatsu’s (1938)
+paper was published in March (dating from
+Pulawski 2024
+), while
+Blüthgen’s (1938)
+paper was published in May (dating from http://hbs.bishopmuseum.org/dating/sherbornia/konowia.html). Thus, the valid name for the species previously known as both
+
+A. trifasciatus shibuyai
+
+and
+
+A. balticus
+
+is
+
+A. shibuyai
+
+.
+
+
+Kostylev (1938: 305)
+indicated the type material of
+
+O. trifasciatus orientalis
+
+as follows: “Type:
+♀
+Jakovlevka, Ussuri,
+10/IX 1926
+, Djakonov et Filipjev;
+♂
+—Jakovlevka, Ussuri,
+21/VI 1926
+, Djakonov et Filipjev”. Therefore, it is unclear whether he designated the female only as the type (the
+holotype
+according to Art. 73.1.1 of
+ICZN 1999
+) or both specimens (
+syntypes
+according to Art. 73.2 of
+ICZN 1999
+). He also reported (
+Kostylev 1938: 302
+) that the type specimens of the taxa described in his paper were deposited in ZMMU; however, we failed to find corresponding specimens of
+
+O. trifasciatus orientalis
+
+in that collection but discovered them in ZISP. These
+two specimens
+correspond in full to Kostylev’s description; however, they were not properly labeled. The female specimen is designated here as the
+lectotype
+in accordance with Art. 74.7.3 of
+ICZN (1999)
+, for the purpose of nomenclature stability and correct application of the name
+
+O. trifasciatus orientalis
+
+.
+
+
+
+FIGURE 1
+.
+
+Ancistrocerus shibuyai
+(Yasumatsu)
+
+(
+A
+,
+B
+,
+E
+,
+G
+) and
+
+A. trifasciatus
+(Müller)
+
+(
+C
+,
+D
+,
+F
+,
+H
+).
+A
+,
+E
+, ♀, Primorskiy Territory;
+B
+, ♂, Nizhniy Novgorod Province;
+C
+,
+F
+, ♀, Nizhniy Novgorod Province;
+D
+,
+H
+, ♂, Kabardino-Balkaria;
+G
+, ♂, Novosibirsk Province.
+A–D
+, habitus in dorsal view;
+E
+,
+F
+, antero-ventral part of metasoma in lateral view;
+G
+,
+H
+, aedeagus in lateral view (vl, ventral lobe). Scale bars: 1 mm.
+
+
+
+
+Budrys
+et al.
+(2023: 15
+
+; Supplement 3: 6) also supposed that
+
+A. balticus
+
+could be considered a subspecies of
+
+A. shibuyai
+
+if further study will reveal their continuous trans-Palaearctic distribution and such a distribution is shown by the present data. We, however, do not support the idea to recognize subspecies in vespid wasps. As introduced by
+Nixon & Wheeler (1990)
+and later applied to vespids by various authors (e.g.
+Carpenter 1988
+;
+
+Tan
+et al.
+2014
+
+;
+Hermes & de Oliveira 2016
+;
+Selis 2018
+;
+
+Fateryga
+et al.
+2021
+
+;
+Selis & Carpenter 2023
+;
+
+Selis
+et al.
+2024
+
+), the subspecific rank has no place in a taxonomic system based on phylogeny and therefore the numerous taxa considered as subspecies must be revised to ascertain their actual status, whether as simple synonyms or as distinct species.
+
+
+
+
+
+Material examined.
+LECTOTYPE
+Of
+
+ODYNERUS TRIFASCIATUS ORIENTALIS
+(DESIGNATED HERE)
+
+:
+
+♀
+, labeled “Яковлевка Спас. у. / Уссур. кр. [Yakovlevka, Spassk Uezd, Ussuriysk Territory] 10.IX 926 / ДЬЯконв ФилипЬев [leg. Dyakonov and Filipyev] // Пасека / Квашука [Kvashuk’s apiary] //
+
+Odynerus
+
+♀
+/
+
+trifasciatus
+
+F / G. Kostylev det. 1928 //
+Lectotype
+♀
+/
+
+Odynerus trifasciatus
+
+/
+
+orientalis
+Kostylev, 1938
+
+/ des. Fateryga, 2024 [red label]” [
+ZISP
+].
+
+PARALECTOTYPE
+Of
+
+ODYNERUS TRIFASCIATUS ORIENTALIS
+
+:
+
+Same data as the lectotype but collected on 21.
+VI
+,
+1 ♂
+[
+ZISP
+].
+ADDITIONAL MATERIAL:
+294 ♀
+,
+158 ♂
+.
+AUSTRIA
+: Hausbach bei Innerbreitenau.
+UKRAINE
+:
+Kiev
+.
+RUSSIA
+.
+
+EUROPEAN PART
+,
+NORTH-WEST
+.
+Saint Petersburg
+and
+Leningrad Province
+
+: Kolomäggi [Kolomyagi];
+Saint Petersburg
+; Udelnaya Station; Zamanilovka.
+
+EUROPEAN PART
+,
+CENTRAL
+
+.
+
+Belgorod Province
+
+: Staryy Oskol.
+
+Moscow
+and
+Moscow Province
+
+: Kosino; Krylatskoye; Kuntsevo; Mytishchi; Ruza; Vicinity of
+Moscow
+.
+
+Nizhniy Novgorod Province
+
+:
+Ardatov District
+, Mukhtolovo;
+Arzamas District
+, vicinity of Staraya Pustyn;
+Balakhna District
+, Konevo;
+Bogorodsk District
+, Laksha;
+Borskoye District
+, vicinity of Elesino; Dzerzhinsk, Svyatoye Lake,
+56.213°N
+,
+43.3912°E
+;
+Kstovo District
+, vicinity of Vetchak; Kstovo,
+50.178°N
+,
+44.174°E
+;
+Lyskovo District
+, Krasnobobrovskoye Forestry;
+Lyskovo District
+, vicinity of Belozerikha;
+Navashino District
+, vicinity of Rodyakovo;
+Nizhniy Novgorod
+, car factory;
+Nizhniy Novgorod
+, Priokskiy District;
+Nizhniy Novgorod
+, Striginskiy Bor Park;
+Semenov District
+, vicinity of Khakhaly,
+56.664°N
+,
+44.658°E
+;
+Shakhunya District
+, Shakhunya;
+Sharanga District
+, Krasnaya Gorka;
+Sharanga District
+, vicinity of Tanayka;
+Sharanga District
+, Vtoryye Nikolayevskiye,
+56°59.763′N
+,
+46°30.420′E
+;
+Sosnovskoye District
+, vicinity of Bechikha, Kirkideyevo Lake,
+55°42.121′N
+,
+43°26.046′E
+;
+Tonkino District
+, Aleshino;
+Tonkino District
+, Tonkino;
+Volodarskiy District
+, vicinity of Reshetikha;
+Vyksa District
+, Vyksa; ibid.,
+55.319945°N
+,
+42.213395°E
+.
+
+Ryazan Province
+
+: Dankov Uezd, [Miloslavskoye District] Gremyachka.
+
+Tula Province
+
+: Strakhovo.
+
+Tver Province
+
+: Bologoye.
+
+Yaroslavl Province
+
+: Berditsino;
+Yaroslavl
+.
+
+EUROPEAN PART
+,
+EAST
+
+.
+
+Chuvashia
+
+:
+Alatyr District
+, Bezdna River;
+Alatyr District
+, Prisurskiy Reserve;
+Yadrin District
+,
+1 km
+SSE Nikitino.
+
+Mari El
+
+:
+Mari-Turek District
+, vicinity of Mariyetz,
+56°30.866′N
+,
+49°49.414′E
+;
+Novyy Toryal District
+, Novyy Toryal,
+56°59.765′N
+,
+48°43.700′E
+;
+Orshanka District
+, Ilinka,
+56°47.695′N
+,
+47°57.349′E
+;
+Orshanka District
+, vicinity of Norka,
+56°58.633′N
+,
+48°03.495′E
+.
+
+URALS
+.
+Bashkortostan
+
+: Shulgan-Tash Reserve, Kapov Cordon, Belaya River valley; Vicinity of Birsk.
+
+Orenburg Province
+
+: Vicinity of
+Orenburg
+.
+
+Perm
+Territory
+
+:
+Perm
+.
+WESTERN SIBERIA
+.
+
+Altai Republic
+
+:
+15 km
+SW Artybash;
+24 km
+NWW Aktash, Chuya River valley,
+50°21′31″N
+,
+87°16′15″E
+; Artybash, Teletskoye Lake; ibid.,
+51°49′N
+,
+87°15′E
+,
+443 m
+; Chulyshman River, right bank; Teletskoye Lake, Kyga River; Ulagan Valley, Kaltar-Ilen; Vicinity of Artybash; Yaylyu.
+
+Altai
+Territory
+
+: Barnaul, vicinity of Yuzhnyy; Vicinity of Barnaul.
+
+Kemerovo Province
+
+:
+5–7 km
+upper Bolshoy Tesh River mouth, ~
+8 km
+NEE Kuzedeyevo;
+Krapivinskiy District
+,
+8 km
+SSW Saltymakovo,
+54°45′46″N
+,
+87°01′27″E
+,
+150 m
+;
+Novokuznetsk District
+,
+7 km
+E Kuzedeyevo,
+53°18′N
+,
+87°19′E
+.
+
+Novosibirsk Province
+
+:
+Barabinsk District
+, Tandovo Lake, Suchun Peninsula;
+Moshkovo District
+,
+5 km
+E Zherebtsovo;
+Novosibirsk
+;
+Novosibirsk District
+, vicinity of Komarovka;
+Novosibirsk
+, Akademgorodok; ibid.,
+54°50′35″N
+,
+83°05′28″E
+;
+Novosibirsk
+, Botanical Garden,
+55°03′N
+,
+82°52′E
+;
+Toguchin District
+,
+6 km
+SE Shmakovo, Kruteikha River; Vicinity of Akademgorodok, water reservoir coast.
+
+Omsk Province
+
+:
+Cherlak District
+, Tatarka,
+55.587°N
+,
+75.023°E
+.
+
+Tomsk Province
+
+:
+Kargasok District
+, vicinity of Ust-Chizhapka;
+Tomsk
+, Stepanovka.
+EASTERN SIBERIA
+.
+
+Buryatia
+
+: Selenginskiy Uezd, Gusinoye Lake.
+
+Irkutsk Province
+
+:
+Irkutsk
+; Lena River headspring, Ada River; Listvinichnoye [Listvyanka], Baikal Lake; Ushalovka River,
+9 km
+E
+Irkutsk
+.
+
+Khakassia
+
+: Matur.
+
+Krasnoyarsk
+Territory
+
+:
+Krasnoyarsk
+; Yenisseysk.
+Zabaykalskiy Territory
+: 100 versts from Nerchinsk;
+25 km
+NE Argunsk; 30 versts from Chita, Shirokaya Pad’;
+35 km
+NWW Borzi, Dauria Reserve, Adon-Chelon Mts.;
+60 km
+SE Nizhniy Tsasuchey, Dauria Reserve, vicinity of Teli Cordon; Argun River valley, ~
+10 km
+S Uryupino; Khilok; Onon River valley,
+10–12 km
+upper Nizhniy Tsasuchey, vicinity of Kudon Lake; S vicinity of Nerchinskiy Zavod; SE Nizhniy Tsasuchey; Vicinity of Nizhniy Tsasuchey, Onon River valley, from Verkhniy Tsasuchey to the bridge.
+
+FAR
+EAST
+.
+Amur Province
+
+:
+3 km
+E Uril; Gribovka, Arkhara River; Khingan Reserve,
+27 km
+W Arkhara, vicinity of Yuzhnyy Cordon; Khingan Reserve, Antonovskoye Forestry, vicinity of Kleshenskoye Cordon; Khingan Reserve, Arkhara; Khingan Reserve, Khingan Forestry, Erakta; Khingan Reserve, Khingan Forestry, Mt. Pukhovaya.
+
+Jewish
+Autonomous Province
+
+: Bastak Reserve,
+48°59′N
+,
+132°54′E
+.
+
+Kamchatka
+Territory
+
+: Atlasovo;
+Kamchatka
+[without details]; Kozyrevsk.
+
+Khabarovsk
+Territory
+
+:
+20 km
+W Nikolayevsk-on-Amur, vicinity of Arkhangelskoye,
+53°10.5′N
+,
+140°24′E
+;
+44 km
+NEE Komsomolsk, Komsomolskiy Reverve, Bichi Cordon;
+Amur
+River mouth, Bolonskiy Reserve, Kirpu Cordon,
+49°30.4′N
+,
+136°02′E
+; Anyuy River valley, national park, Bogbasu Cordon,
+49°25.5′N
+,
+137°42.7′E
+; Bolshekhekhtsirskiy Reserve, vicinity of Bychikha,
+48°17′N
+,
+134°48′E
+; ibid., 48°17–18′N, 134°49– 50′E; ibid.,
+48°18′N
+,
+134°49′E
+; Botchinskiy Reserve, Teplyy Kluch Cordon,
+48°18′N
+,
+139°36′E
+, ~
+283 m
+; Bureinskiy Reserve, Bureya River headspring, Strelka Cordon,
+51°33′N
+,
+134°03′E
+; Gorin River, inflow of
+Amur
+;
+Khabarovsk
+; Komsomolsk-on-Amur; Komsomolsk-on-Amur, Siliinskiy Park,
+50°34′N
+,
+137°03′E
+; Malmyzh, right inflow of
+Amur
+; Nizhnetambovskoye, right bank of
+Amur
+River; Sikhote-Alin, Botchi River; Sikhote-Alin, Botchinskiy Reserve, upper reaches of Mulpa River,
+7 km
+E Teplyy Kluch Cordon,
+48°19′N
+,
+139°38′E
+; Sikhote-Alin, Botchinskiy Reserve, upper reaches of Mulpa River, Teplyy Kluch Cordon,
+48°18′N
+,
+139°35′E
+,
+250 m
+; Sikhote-Alin, Botchinskiy Reserve, upper reaches of Mulpa River, vicinity of Teplyy Kluch Cordon,
+48°18′N
+,
+139°35′E
+,
+250–300 m
+;
+Solnechnyy District
+, vicinity of Amut Lake,
+50°47′N
+,
+136°23′E
+,
+925 m
+; Upper reaches of Bureya River,
+51°08′N
+,
+133°01′E
+.
+
+Primorskiy
+Territory
+
+:
+5 km
+W Lazo;
+50 km
+SW Terney, Cheremukhovaya River; Behind Manchzhur; Bolshaya Ussurka River,
+20 km
+NW Melnichnoye; Bolshaya Ussurka River, vicinity of Gogolevka; E Ussuriysk; Gamov Peninsula, Vityaz,
+42°36′29″N
+,
+131°11′36″E
+; Kamen-Rybolov; Khanka Lake; Kiyevka; Lazo; Lazovskiy Reserve; Lazovskiy Reserve,
+10 km
+W Preobrazheniye; Lazovskiy Reserve, Amerika Place; Lazovskiy Reserve, Tachengouza Bay; Lazovskiy Reserve, Zarya Bay; Okeanskaya Station; Petrov Bay; Sikhote-Alin, Koi River, 52 versts from mouth; Spassk; Spassk Uezd, Yakovlevka; Suchan [Partizansk];
+Suchan District
+, Monakino;
+Suchan District
+, Sitsa River; Suchan Railroad, Tigrovoy Station; Tigrovoy; Vicinity of Anisimovka; Vicinity of Preobrazheniye, Khanikheza Place; Vicinity of Preobrazheniye, Sya Ukhe Place; Vicinity of Vladivostok; Vladivistok, S Zolotoy Rog Bay; Vladivostok; Vladivostok, Sedanka; Vladivostok, Zolotoy Rog Bay.
+
+Sakhalin
+
+:
+10 km
+N Tymovskoye;
+10 km
+S Dolinsk;
+10 km
+S Dolinsk, Sokol;
+20 km
+N Nevelsk;
+20 km
+NW Aniva; Kholmsk; Kirillovo;
+Makarov District
+, Tikhaya River; Novoaleksandrovsk; Ozerskiy;
+Sakhalin
+[without details]; Schmidt Peninsula,
+70 km
+N Okha; Susunayskiy, Anna River mouth; Tunaycha Lake; Tym River, Voskresenovka; Vicinity of Shebunino; Yuzhno-Sakhalinsk.
+Kuril Islands
+: Iturup, Dobroye Nachalo Bay; Kunashir,
+2 km
+NE Tretyakovo; Kunashir,
+7 km
+S Lagunnoye Lake; Kunashir, Kislaya River; Kunashir, Mendeleyevo; Kunashir, Stolbchatyy Cape; Kunashir, Tretyakovo; Kunashir, vicinity of Alekhino; Kunashir, vicinity of Dubovoye; Kunashir, vicinity of Yuzhno-Kurilsk; Kunashir, W coast, Lagunnoye; Kunashir, W coast, Nazarovo.
+KAZAKHSTAN
+.
+
+Abai Province
+
+: Semipalatinsk [Semey].
+
+Akmola Province
+
+: Kokshetau.
+MONGOLIA
+:
+Arkhangai Aimag
+,
+48 km
+S Tevshrulekh,
+2000 m
+; Central [
+Töv
+] Aimag, Zaysan Place, S slope of Mt. Bogdo-Ula.
+SOUTH KOREA
+: Simgok-ri, Sinbukmyeon, Pocheon-si; Yongdae-ri, Buk-myeon, Inje-gun.
+JAPAN
+:
+Hokkaido
+, Daisetsuzan Mts.
+
+
+
+
+Distribution.
+Based on the studied material, the species is confirmed to *
+Austria
+, *
+Ukraine
+,
+Russia
+(*European part, *Urals, *Western Siberia, Eastern Siberia, Far East), *
+Kazakhstan
+,
+Mongolia, Korean
+Peninsula, and
+Japan
+(
+Fig. 3
+). Also known from
+Poland
+(
+Blüthgen 1938
+, as
+
+Ancistrocerus trifasciatus
+var.
+moeschleri
+
+),
+Lithuania
+(
+
+Budrys
+et al.
+2023
+
+, as
+
+A. balticus
+
+), and
+China
+(North-East) (
+Giordani Soika 1986
+, as
+
+A. trifasciatus orientalis
+
+;
+Kim & Yamane 2009
+;
+
+Tan
+et al.
+2018
+
+, both as
+
+A. trifasciatus shibuyai
+
+). The distribution of
+
+A. shibuyai
+
+in Europe is apparently underestimated.
+
+
+
+
\ No newline at end of file
diff --git a/data/B6/2A/87/B62A8782FFD5D17445EAFB3AFB80FE8E.xml b/data/B6/2A/87/B62A8782FFD5D17445EAFB3AFB80FE8E.xml
new file mode 100644
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--- /dev/null
+++ b/data/B6/2A/87/B62A8782FFD5D17445EAFB3AFB80FE8E.xml
@@ -0,0 +1,919 @@
+
+
+
+Ancistrocerus shibuyai (Yasumatsu, 1938), stat. restit. (Hymenoptera: Vespidae: Eumeninae), an overlooked trans-Palaearctic species
+
+
+
+Author
+
+Fateryga, Alexander V.
+T. I. Vyazemsky Karadag Scientific Station-Nature Reserve of RAS-Branch of A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS, Nauki Str. 24, Kurortnoye, 298188 Feodosiya, Russia.
+
+
+
+Author
+
+Proshchalykin, Maxim Yu.
+Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, 100 - let Vladivostoku Ave. 159, 690022 Vladivostok, Russia.
+
+text
+
+
+Zootaxa
+
+
+2024
+
+2024-11-12
+
+
+5537
+
+
+4
+
+
+527
+540
+
+
+
+
+http://dx.doi.org/10.11646/zootaxa.5537.4.5
+
+journal article
+10.11646/zootaxa.5537.4.5
+1175-5326
+14240360
+F2B1D954-3B04-49F6-8FF3-A124DEC913B2
+
+
+
+
+
+
+
+Ancistrocerus trifasciatus
+(
+Müller, 1776
+)
+
+
+
+
+
+
+
+(
+Figs 1C, D, F, H
+;
+2C, D, G, H
+)
+
+
+
+
+
+
+
+Vespa trifasciata
+Müller, 1776: 163
+
+
+(
+type
+locality: “Daniae et Norvegiae” [
+Denmark
+and
+Norway
+]);
+type
+depository unknown.
+
+
+
+?
+
+
+Vespa trifasciata
+Fabricius, 1787: 290
+
+
+(type locality: “Habitat Hafniae” [
+Denmark
+]), nom. praeocc., nec
+
+V. trifasciata
+Müller, 1776
+
+;
+syntype
+(s),
+♀
+, Natural History Museum
+Denmark
+, Uniersity of Copenhagen, Copenhagen,
+Denmark
+.
+
+
+
+
+
+Odynerus trimarginatus
+Zetterstedt, 1838: 456
+
+
+,
+♀
+♂
+(type locality: “Lapponia … (Lappon.-Uplandia; Gotlandia pass.)” [
+Finland
+]);
+syntypes
+,
+Lund University
+,
+Sweden
+.
+
+
+
+
+
+
+Odynerus tricinctus
+Herrich-Schäffer, 1839: 20
+
+
+, [
+♀
+] (
+type
+locality not indicated, probably
+Germany
+), 1841: pl. 10;
+type
+material destroyed.
+
+
+
+
+
+Differential diagnosis.
+Females of this species can be distinguished from the closely related
+
+A. shibuyai
+
+by S
+2 in
+lateral view weakly convex to nearly flat (
+Fig. 1F
+; vs. distinctly convex in
+
+A. shibuyai
+
+,
+Fig. 1E
+). Males can be distinguished by ventral lobe of the aedeagus apically rather triangularly pointed (
+Fig. 1H
+; vs. rounded in
+
+A. shibuyai
+
+,
+Fig. 1G
+). An additional character is the coloration of the hind tibia: usually with extensive brown and yellow pattern in the female (
+Fig. 2C
+) (very rarely completely or near completely black,
+Fig. 2D
+) and usually almost entirely yellow in the male (
+Fig. 2G
+) (very rarely black dorsally, particularly in specimens from
+Kamchatka
+,
+Fig. 2H
+). Other characters mentioned in the key by
+
+Budrys
+et al.
+(2023
+
+, Supplement 3) can be also applied but they are also variable and therefore less diagnostic.
+
+
+
+
+Remarks.
+We follow the traditional treatment of
+
+A. trifasciatus
+
+(
+Blüthgen 1961
+;
+
+Budrys
+et al.
+2023
+
+) and its synonymy (
+Blüthgen 1938
+,
+1961
+;
+van der Vecht & Fischer 1972
+), except that
+
+A. trifasciatus moeschleri
+
+is treated here as a synonym of
+
+A. shibuyai
+
+(see above). It is, however, unclear from the original description of
+
+Vespa trifasciata
+Müller, 1776
+
+if it was the same species as commonly known at present as
+
+A. trifasciatus
+
+, or the species known at present as
+
+A. shibuyai
+
+. Since the type depository of
+
+V. trifasciata
+Müller, 1776
+
+is unknown (
+van der Vecht & Fischer 1972
+), the type material can be actually lost and we cannot clarify its diagnostic characters. In our opinion, a
+neotype
+should be designated to keep nomenclature stability (Art. 75.3 of
+ICZN 1999
+) but we do not have examined any specimen from the type locality (either
+Denmark
+or
+Norway
+). Such a specimen should be corresponding to the traditional treatment of
+
+A. trifasciatus
+
+(
+Blüthgen 1961
+;
+
+Budrys
+et al.
+2023
+
+; present paper) and, since
+
+A. trifasciatus
+
+and
+
+A. shibuyai
+
+can be easily distinguished by DNA barcoding (
+
+Budrys
+et al.
+2023
+
+), the COI-5P gene of a possible
+neotype
+should be barcoded prior to its designation (cf. Selis
+et al.
+2023).
+
+
+
+FIGURE 2
+. Left hind tibia of
+
+Ancistrocerus shibuyai
+(Yasumatsu)
+
+(
+A
+,
+B
+,
+E
+,
+F
+) and
+
+A. trifasciatus
+(Müller)
+
+(
+C
+,
+D
+,
+G
+,
+H
+) in dorso-lateral view.
+A–C
+, ♀, Nizhniy Novgorod Province;
+D
+, ♀, Kamchatka Territory;
+E
+,
+G
+, ♂♂, Nizhniy Novgorod Province;
+F
+, ♂, Novosibirsk Province;
+H
+, ♂, Kamchatka Territory. Scale bars: 1 mm.
+
+
+
+The name
+
+V. trifasciata
+Fabricius, 1787
+
+is also doubtful due to a lack of the description of important characters in the original publication. However, this name is anyway permanently invalid due to the homonymy with
+
+V. trifasciata
+Müller, 1776
+
+(Art. 57.2 of
+ICZN 1999
+). As for the names
+
+Odynerus trimarginatus
+
+and
+
+O. tricinctus
+
+, their descriptions are enough to clarify their conspecificity with
+
+A. trifasciatus
+
+(in its traditional treatment) due to the leg color specified (
+Zetterstedt 1838
+) or even illustrated (
+Herrich-Schäffer 1841
+).
+
+
+It is also to be noted that in
+Gusenleitner’s (1995)
+paper, fig. 4 labeled as
+
+A. trifasciatus
+
+actually depicts
+
+A. oviventris
+(
+Wesmael, 1836
+)
+
+, while fig. 5 labeled as
+
+A. oviventris
+
+depicts rather
+
+A. shibuyai
+
+than
+
+A. trifasciatus
+
+.
+
+
+
+
+Material examined.
+233 ♀
+,
+239 ♂
+.
+AUSTRIA
+: Hausbach bei Innerbreitenau; Vicinity of Linz; Zeissberg bei Freistadt.
+GERMANY
+: Vicinity of Dresden.
+SWEDEN
+:
+Stokholm
+.
+FINLAND
+: Rajala; Rajala, Saimaa Canal; Raucharanta, Saimaa Canal.
+CZECH REPUBLIC
+: Čelákovice.
+SLOVAKIA
+: Zádiel.
+POLAND
+: Danzig [Gdańsk];
+Silesia
+, Caralath [Siedlisko].
+ESTONIA
+: Hapsal [Haapsalu]; Narva; Verro.
+BELARUS
+:
+Brest Province
+, vicinity of Pinsk;
+Minsk Province
+, Loshnitsa; Pinsk;
+Vitebsk Province
+, Ambrosovichi;
+Vitebsk Province
+, Lepel District, Postrezhye;
+Vitebsk
+Uezd [
+Vitebsk Province
+without details].
+UKRAINE
+: Bila Tserkva, Alexandria Dendropark;
+Kharkov
+;
+Kiev
+; Mirgorod Uezd, Yereski;
+Poltava Province
+, Gadyach;
+Vinnytsia Province
+, Yanov [Ivaniv]; Zhitomir Province, Berdichev.
+MOLDOVA
+:
+Bendery
+, Kitskany.
+RUSSIA
+.
+
+EUROPEAN PART
+,
+NORTH-WEST
+.
+Novgorod Province
+
+: Borovichi; Valday Uezd.
+
+Pskov Province
+
+: Gdov Uezd, Glebova Gorka; Novorzhev Uezd, [Bezhanitsi District] Aparino; Novorzhev Uezd, [Bezhanitsi District] Kalinino; Sushchevo Station.
+
+Saint Petersburg
+and
+Leningrad Province
+
+: Baltic Port; Chernaya Lakhta; Duderhof Heights; Gatchina; Kolomäggi [Kolomyagi]; Levashovo; Luga; Martyshkino; Pargola [Pargolovo]; Peterhof; Popovka; Primorskaya Railroad, Borskaya [Borovaya] Station;
+Saint Petersburg
+;
+Saint Petersburg
+, Chernaya River;
+Saint Petersburg
+,
+Moscow
+Highway;
+Saint Petersburg
+, Smolenskoye Cemetery; Siverskiy, Mezhino; Terijoki [Zelenogorsk]; Tolmachevo; Udelnaya Station; Volkovo; Zamanilovka.
+
+EUROPEAN PART
+,
+NORTH
+.
+Arkhangelsk Province
+
+:
+Arkhangelsk
+; Mudyugskiy Beacon; Solovki [Solovetskiye Ostrova].
+
+Karelia
+
+: Petrozavodsk; Povenets.
+
+Murmansk Province
+
+: Kola, Kola River bank; Vicinity of
+Murmansk
+.
+
+Vologda Province
+
+: Gryazovets;
+Vologda
+.
+
+EUROPEAN PART
+,
+CENTRAL
+.
+Belgorod Province
+
+: Khotmyzhsk.
+
+Bryansk Province
+
+: Orlovskiye Dvoriki.
+
+Ivanovo Province
+
+:
+Ivanovo
+.
+
+Kostroma Province
+
+:
+Kostroma
+; Troitsa; Vicinity of
+Kostroma
+.
+
+Kursk Province
+
+: Vicinity of
+Kursk
+; Vicinity of
+Kursk
+, Selizovo.
+
+Mordovia
+
+: Ruzayevka District, vicinity of Boldovo.
+
+Moscow
+and
+Moscow Province
+
+: Izmailovo; Kashira Station; Kashira Uezd, Sredneye; Klyazma [Pushkino]; Krylatskoye; Leonovo;
+Moscow
+; Mytishchi; Nikolo-Peshnoshskiy Monastery; Peredelkino; Ruza; Valuyevo; Vicinity of
+Moscow
+; Zvenigorod.
+
+Nizhniy Novgorod Province
+
+: Arzamas District, vicinity of Pustyn; Arzamas District, vicinity of Staraya Pustyn; Balakhna District, Maloye Kozino; Kstovo District, vicinity of Igumnovo;
+Nizhniy Novgorod
+, Avtozavodskoy District;
+Nizhniy Novgorod
+, Priokskiy District;
+Nizhniy Novgorod
+, Striginskiy Bor Park; Tonkino District, Aleshino; Tonkino District, Tonkino; Vetluga District, Kalinino.
+
+Ryazan Province
+
+: Dankov Uezd, [Miloslavskoye District] Gremyachka.
+
+Tver Province
+
+: Bologoye; Doroshikha; Rzhev.
+
+Yaroslavl Province
+
+: Berditsino; Norskoye;
+Yaroslavl
+; Zhedenovo.
+
+EUROPEAN PART
+,
+EAST
+.
+Kirov Province
+
+: Vyatka Governorate.
+
+Samara Province
+
+: Kinel;
+Samara
+.
+
+EUROPEAN PART
+,
+SOUTH
+. Kherson Province
+
+: Vicinity of Kherson.
+Lugansk People Republic
+: Provalskaya Steppe.
+
+Volgograd Province
+
+: Kamyshin District, Shcherbatovka; Sarepta [
+Volgograd
+].
+
+EUROPEAN PART
+,
+NORTH CAUCASUS
+.
+Chechnya
+
+: Gudermes District, vicinity of Islakhan-Yurt.
+KabardinoBalkaria
+: Chegem Canyon, Bashil,
+2080 m
+.
+
+Karachay-Cherkessia
+
+:
+7 km
+N Rozhkao,
+43°53′05″N
+,
+40°56′55″E
+.
+
+Stavropol
+Territory
+
+: Bolshoy Zelenchuk River.
+
+EUROPEAN PART
+,
+CRIMEA
+
+: Belogorsk District, Suat River valley; Krasnolesye; Mt. Aypetri; Simferopol; Simferopol, Anatra; Yalta Reserve, Mt. Aypetrinskaya Yayla; Yalta, Mt. Lapata.
+
+URALS
+.
+Bashkortostan
+
+: Birsk; Irgizly.
+
+Chelyabinsk Province
+
+: Ilmenskiy Reserve; Zlatoust, Miass.
+
+Kurgan Province
+
+: Kurtamysh District, Pesyanoye.
+
+Orenburg Province
+
+: Buzuluk Uezd;
+Orenburg
+.
+
+Perm
+Territory
+
+:
+Perm
+.
+Sverdlov Province
+: Yekaterinburg.
+
+WESTERN SIBERIA.
+Altai Republic
+
+:
+14 km
+SE Aktash, Chuya River,
+50°13.984′N
+,
+87°45.471′E
+,
+1450 m
+; Teletskoye Lake; Teletskoye Lake, Kyga River; Vicinity of Artybash; Yazula, Chulyshman River.
+
+Altai
+Territory
+
+: Barnaul, vicinity of Yuzhnyy; Biysk Uezd, Lugovoye.
+
+Kemerovo Province
+
+:
+Kemerovo
+, Tsentralnyy District.
+Khanty-Mansiysk Autonomous Okrug
+: Yuganskiy Reserve, Yurt-Achimovy.
+
+Novosibirsk Province
+
+: Iskitim District, Yevsino;
+Novosibirsk District
+, vicinity of Komarovka;
+Novosibirsk
+, Zayeltsovskiy Forest; Ust-Tarka District, Yelanka.
+
+Omsk Province
+
+:
+Omsk
+.
+
+Tyumen Province
+
+:
+Tyumen
+.
+
+EASTERN SIBERIA.
+Irkutsk Province
+
+: Baikal Lake, 18 versts from Listvinichnoye [Listvyanka]; Padun [Bratsk].
+
+Khakassia
+
+: Abakan; Konsu River, N Abakan.
+
+Krasnoyarsk
+Territory
+
+:
+Krasnoyarsk
+; Vicinity of Norilsk, Talnakh.
+
+Yakutia
+
+: 7 versts below Tyuktyurskaya [Tekhtyur], left bank of Lena River; Ulakhan-An River mouth, left inflow of Lena River.
+Zabaykalskiy Territory
+: Amazar; Vicinity of Chita.
+
+FAR
+EAST
+.
+Amur Province
+
+: Khingan Reserve,
+27 km
+W Arkhara, vicinity of Yuzhnyy Cordon.
+
+Kamchatka
+Territory
+
+:
+8 km
+S Kozyrevsk; Atlasovo;
+Kamchatka
+[without details];
+Kamchatka
+River valley; Kichiga River; Kozyrevsk; Petropavlovsk; Yelizovo; Zhupanovo.
+
+Khabarovsk
+Territory
+
+:
+44 km
+NEE Komsomolsk, Komsomolskiy Reverve, Bichi Cordon;
+Amur
+Liman; Bureinskiy Reserve, Bureya River headspring, Strelka Cordon,
+51°33′N
+,
+134°03′E
+; Malmyzh, right inflow of
+Amur
+.
+
+Magadan Province
+
+: Ola River.
+
+Sakhalin
+
+: Langeri River mouth; Novoaleksandrovsk; Tunaycha Lake. ABKHAZIA: Sukhum.
+GEORGIA
+: Bakuriani; Batumi, vicinity of Botanical Garden; Kutaisi Governorate; Lagodechy [Lagodekhi].
+ARMENIA
+: Armenia [without details].
+AZERBAIJAN
+: Aresh Uezd, Geok-Topa;
+Nakhchivan
+AR,
+Shakhbuz
+, Bichenek,
+39°31′N
+,
+45°46′E
+,
+2000 m
+.
+KYRGYZSTAN
+:
+Fergana Province
+, Aslyk-Bob, 10400ʹ; Moldo-Too, Shavyr.
+KAZAKHSTAN
+.
+Abai Province
+: Semipalatinsk [Semey].
+
+Almaty Province
+
+: Trans-Ili Alatau, Turgen Place.
+
+East Kazakhstan Province
+
+:
+50 km
+SE Ust-Kamenogorsk, vicinity of Severnoye; Nikolayevka,
+30 km
+SE Georgiyevka [Kolbatau].
+Jambyl Province
+:
+25 km
+S Merke.
+Karaganda Province
+: Balkhash.
+
+North Kazakhstan Province
+
+: Petropavlovsk.
+
+
+
+
+Distribution.
+Based on the studied material, the species is confirmed to
+Austria
+,
+Germany
+,
+Sweden
+,
+Finland
+,
+Czech Republic
+,
+Slovakia
+,
+Poland
+,
+Estonia
+,
+Belarus
+,
+Ukraine
+,
+Moldova
+,
+Russia
+(European part, Urals, Western Siberia, Eastern Siberia, Far East), *
+Abkhazia
+, *
+Georgia
+, *
+Armenia
+, *
+Azerbaijan
+,
+Kyrgyzstan
+, and
+Kazakhstan
+(
+Fig. 3
+). This species is known to be widely distributed through Western, Northern, and Eastern Europe (e.g.,
+von Schulthess 1887
+–1897;
+Berland 1928
+;
+Benno 1957
+;
+Blüthgen 1961
+;
+Scobiola-Palade 1989
+;
+Pekkarinen & Huldén 1991
+;
+Schljachtenok & Gusenleitner 1996
+;
+Schneider 1997
+;
+Gusenleitner 2008
+;
+Amolin 2009
+;
+Archer 2011
+;
+
+Budrys
+et al.
+2023
+
+), although some of its records may actually refer to
+
+A. shibuyai
+
+. Also known from
+Italy
+(
+Bonelli 1970
+; Selis 2023) and
+Turkey
+(
+Giordani Soika 1970
+;
+Yildirim & Gusenleitner 2012
+).
+
+
+
+
\ No newline at end of file