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@ -55,9 +58,9 @@
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(
<figureCitation id="136F2A54EC107A11FF64FCFD7011FCDE" box="[159,260,799,825]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="1" pageNumber="585">Figs 12</figureCitation>
<figureCitation id="136F2A54EC107A11FF64FCFD7011FCDE" box="[159,260,799,825]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="1" pageNumber="585">Figs 12</figureCitation>
,
<figureCitation id="136F2A54EC107A11FEEBFCFD702EFCDE" box="[272,315,799,825]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." pageId="1" pageNumber="585">68</figureCitation>
<figureCitation id="136F2A54EC107A11FEEBFCFD702EFCDE" box="[272,315,799,825]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="1" pageNumber="585">68</figureCitation>
)
</paragraph>
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@ -112,7 +115,7 @@ of
</taxonomicName>
<specimenCount id="9D52FD58EC107A11FBCCFC057559FBE6" box="[1079,1100,999,1025]" count="1" pageId="1" pageNumber="585" type="male"></specimenCount>
(
<figureCitation id="136F2A54EC107A11FBA7FC05758AFBE6" box="[1116,1183,999,1025]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="1" pageNumber="585">Fig. 1</figureCitation>
<figureCitation id="136F2A54EC107A11FBA7FC05758AFBE6" box="[1116,1183,999,1025]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="1" pageNumber="585">Fig. 1</figureCitation>
; dissected): Kashmir, Aru | lg.H.Franz,
<date id="FFEA1011EC107A11FEA4FBE970C6FBC2" box="[351,467,1035,1061]" pageId="1" pageNumber="585" value="1977-10">Okt. 1977</date>
&lt;printed&gt;, TYPE &lt;red, printed&gt;,
@ -148,7 +151,7 @@ of
</taxonomicName>
<specimenCount id="9D52FD58EC107A11FD0DFB79721EFB52" box="[758,779,1179,1205]" count="1" pageId="1" pageNumber="585" type="male"></specimenCount>
(
<figureCitation id="136F2A54EC107A11FCE0FB797277FB52" box="[795,866,1179,1205]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="1" pageNumber="585">Fig. 2</figureCitation>
<figureCitation id="136F2A54EC107A11FCE0FB797277FB52" box="[795,866,1179,1205]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="1" pageNumber="585">Fig. 2</figureCitation>
; dissected): Kashmir, Chandanwari | b.Pahalgam,
<quantity id="4CAC9B34EC107A11FF6CFB5D7025FB3E" box="[151,304,1215,1241]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="1" pageNumber="585" unit="m" value="2950.0" valueMax="3100.0" valueMin="2800.0">2800-3100 m</quantity>
| lg.H.Franz,
@ -196,7 +199,7 @@ of
</paragraph>
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Habitus as in
<figureCitation id="136F2A54EC107A11FE9BFA5970D1FA32" box="[352,452,1467,1493]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="1" pageNumber="585">Figs 12</figureCitation>
<figureCitation id="136F2A54EC107A11FE9BFA5970D1FA32" box="[352,452,1467,1493]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="1" pageNumber="585">Figs 12</figureCitation>
. Body reddish-brown, with distinctly darker head and abdomen (pronotum and elytra of
<typeStatus id="54EF8873EC107A11FF6CFA3D71ECFA1E" box="[151,249,1503,1529]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
of
@ -232,7 +235,7 @@ of
with fused five rows of serial punctures; abdominal tergites with very fine and sparse punctation.
</paragraph>
<paragraph id="8BEB36D1EC107A13FF3CF88972BAFF32" blockId="1.[151,1437,998,2033]" lastBlockId="3.[151,1437,151,1293]" lastPageId="3" lastPageNumber="587" pageId="1" pageNumber="585">Head 1.31.4 times as broad as long, indistinctly elevated in middle, with very short and shallow anteocellar foveae about as long as diameter of ocellus; anterior portion between antennal insertion and anterior margins of eyes slightly concave. Eyes very large, convex. Ocelli large, well visible, situated at about level of posterior margins of eyes; distance between ocelli slightly greater than distance between ocellus and posterior margin of eyes. Apical segment of maxillary palpi distinctly more than twice as long as preceding segment, from widest basal third gradually narrowed toward subacute apex. Antennae reaching basal portion of elytra when reclined; basal antennomere ovoid, slightly less than three times as long as broad, antennomere 2 distinctly shorter than basal antennomere, 3 narrower and slightly longer than 2, 4 small, about as long as broad, 56 distinctly longer and broader than 4, 78 slightly broader than 6, 910 indistinctly longer and broader than 8, apical antennomere about as long as two preceding antennomeres, from middle gradually narrowed toward subacute apex.</paragraph>
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<paragraph id="8BEB36D1EC137A12FF6CF93C70D3F8FD" blockId="2.[151,1437,1758,1819]" pageId="2" pageNumber="586">
<emphasis id="B920EAC3EC137A12FF6CF93C702EF911" bold="true" box="[151,315,1758,1782]" pageId="2" pageNumber="586">FIGURES 15.</emphasis>
Habitus of
@ -277,9 +280,9 @@ of
<paragraph id="8BEB36D1EC127A13FF3CFE1D744AFDDA" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="587">Abdomen about as broad as elytra or slightly broader, with two small transverse tomentose wing-folding spots in middle of abdominal tergite V; apical margin of abdominal tergite VII with distinct narrow palisade fringe.</paragraph>
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Male. Apical margins of abdominal tergite VIII and sternite VIII widely and rather deeply concave. Aedeagus with wide basal portion, gradually narrowed apically, with narrow median lobe and small rounded apex; parameres moderately wide, distinctly broadened in apical portions, slightly not reaching apex of median lobe; internal sac narrow and moderately long (
<figureCitation id="136F2A54EC127A13FE18FD517356FD2A" box="[483,579,691,717]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." pageId="3" pageNumber="587">Fig. 6, 8</figureCitation>
<figureCitation id="136F2A54EC127A13FE18FD517356FD2A" box="[483,579,691,717]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="3" pageNumber="587">Fig. 6, 8</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="136F2A54EC127A13FC38FD517512FD2A" box="[963,1031,691,717]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." pageId="3" pageNumber="587">Fig. 7</figureCitation>
<figureCitation id="136F2A54EC127A13FC38FD517512FD2A" box="[963,1031,691,717]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="3" pageNumber="587">Fig. 7</figureCitation>
.
</paragraph>
<paragraph id="8BEB36D1EC127A13FF3CFD357080FD16" blockId="3.[151,1437,151,1293]" box="[199,405,727,753]" pageId="3" pageNumber="587">Female unstudied.</paragraph>

View file

@ -1,45 +1,48 @@
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@ -55,9 +58,9 @@
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<paragraph id="8BEB36D1EC127A13FF6CFA9D7020FA7E" blockId="3.[151,637,1370,1433]" box="[151,309,1407,1433]" pageId="3" pageNumber="587">
(
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<figureCitation id="136F2A54EC127A13FF64FA9D71F2FA7E" box="[159,231,1407,1433]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="3" pageNumber="587">Figs 3</figureCitation>
,
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<figureCitation id="136F2A54EC127A13FF0FFA9D7038FA7E" box="[244,301,1407,1433]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="3" pageNumber="587">910</figureCitation>
)
</paragraph>
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@ -87,7 +90,7 @@
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(
<figureCitation id="136F2A54EC127A13FD9DF9EB73B8F9C4" box="[614,685,1545,1571]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="3" pageNumber="587">Fig. 3</figureCitation>
<figureCitation id="136F2A54EC127A13FD9DF9EB73B8F9C4" box="[614,685,1545,1571]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="3" pageNumber="587">Fig. 3</figureCitation>
; dissected prior to the present study: the aedeagus and the apical genital segment are glued on the same card with the specimen): Umg.Alm Darghari | b. Maharigaon, 4000m &lt;printed&gt;, Gebiet von Jumla | Westnepal,lg.H.Franz &lt;printed&gt;, TYPE &lt;red, printed&gt;,
<taxonomicName id="4C544D52EC127A13FB88F9B37409F98C" authorityName="Maklin" authorityYear="1880" box="[1139,1308,1617,1643]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="genus">Mannerheimia</taxonomicName>
| emodensa | H. COIFFAIT det. 19[printed]76 &lt;handwritten in black&gt; (
@ -133,7 +136,7 @@ dissected),
)
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;
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<materialsCitation id="3B3C3C8CEC127A13FB06F903754AF8F8" collectionCode="NMW" collectorName="H. Franz" country="Nepal" pageId="3" pageNumber="587" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D52FD58EC127A13FB06F9037438F91B" box="[1277,1325,1761,1788]" count="1" pageId="3" pageNumber="587" type="male">1 ♂</specimenCount>
:
<collectorName id="26A15307EC127A13FABBF903701DF8F8" pageId="3" pageNumber="587">Dzunda Khola-Tal</collectorName>
@ -148,7 +151,7 @@ b.
)
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;
<materialsCitation id="3B3C3C8CEC127A13FB90F8E773E9F8A4" collectionCode="NMW" collectorName="Dampelek &amp; Jumla' &amp; H. Franz'" country="Nepal" location="Gebiet des Rarasees Westnepal" pageId="3" pageNumber="587" specimenCount="1" specimenCount-female="1">
<materialsCitation id="3B3C3C8CEC127A13FB90F8E773E9F8A4" collectionCode="NMW" collectorName="H. Franz" country="Nepal" location="Gebiet des Rarasees Westnepal" pageId="3" pageNumber="587" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D52FD58EC127A13FB90F8E7758CF8C7" box="[1131,1177,1797,1824]" count="1" pageId="3" pageNumber="587" type="female">1 ♀</specimenCount>
:
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@ -180,14 +183,14 @@ Measurements (n=20): HW: 0.470.53; HL: 0.310.33; OL: 0.140.15; AL (
</paragraph>
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Habitus as in
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<figureCitation id="136F2A54EC127A13FE90F85B70A7F834" box="[363,434,1977,2003]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="3" pageNumber="587">Fig. 3</figureCitation>
. Body yellow-brown to reddish-brown, with distinctly darker head (abdomen of some specimens slightly darker than forebody; lateral and basal portions of pronotum and apical portion of elytra slightly paler in
<typeStatus id="54EF8873EC157A14FF0AFF757046FF56" box="[241,339,151,177]" pageId="4" pageNumber="588" type="holotype">holotype</typeStatus>
and in some
<typeStatus id="54EF8873EC157A14FE1DFF757342FF56" box="[486,599,151,177]" pageId="4" pageNumber="588" type="paratype">paratypes</typeStatus>
); antennomeres 511 or 611 brown; mouthparts, antennomeres 14 or 15 and paratergites yellow. Forebody without microsculpture; abdomen with fine and dense transverse microreticulation. Head with moderately fine and sparse punctation, distinctly denser in middle; punctation of pronotum denser, larger and deeper than that in middle portion of head, finer and sparser in middle, and sometimes with narrow impunctate portion in middle or in mediobasal third; scutellum with one to four fine punctures in middle or without them; punctation of elytra denser, slightly larger and deeper than that on pronotum, punctures in middle of each elytron forming five to six vague and tangled rows of serial punctures, but some specimens with markedly coarse surface without longitudinal rows between punctures.
</paragraph>
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<paragraph id="8BEB36D1EC157A14FF6CF8407593F839" blockId="4.[151,1437,1954,2015]" pageId="4" pageNumber="588">
<emphasis id="B920EAC3EC157A14FF6CF840705DF85D" bold="true" box="[151,328,1954,1978]" pageId="4" pageNumber="588">FIGURES 610.</emphasis>
Aedeagus of
@ -216,9 +219,9 @@ Aedeagus of
<paragraph id="8BEB36D1EC147A15FF3CFE1D708DFDDA" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">Abdomen about as broad as elytra, with two small oval or transverse tomentose wing-folding patches in middle of abdominal tergite V.</paragraph>
<paragraph id="8BEB36D1EC147A15FF3CFDA5753DFD2A" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">
Male. Apical margin of abdominal tergite VIII straight or slightly sinuate. Apical margin of abdominal sternite VIII widely sinuate. Aedeagus with wide basal portion, gradually narrowed apically, with relatively narrow median lobe and small rounded apex; parameres with wide apical portions, slightly exceeding apex of median lobe; internal sac narrow and moderately long (
<figureCitation id="136F2A54EC147A15FDF5FD517347FD2A" box="[526,594,691,717]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." pageId="5" pageNumber="589">Fig. 9</figureCitation>
<figureCitation id="136F2A54EC147A15FDF5FD517347FD2A" box="[526,594,691,717]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="5" pageNumber="589">Fig. 9</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="136F2A54EC147A15FC29FD517531FD2A" box="[978,1060,691,717]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." pageId="5" pageNumber="589">Fig. 10</figureCitation>
<figureCitation id="136F2A54EC147A15FC29FD517531FD2A" box="[978,1060,691,717]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="5" pageNumber="589">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="8BEB36D1EC147A15FF3CFD35709EFCF2" blockId="5.[151,1437,150,1221]" pageId="5" pageNumber="589">Female. Apical margin of abdominal tergite VIII straight or somewhat rounded. Apical margin of abdominal sternite VIII rounded.</paragraph>

View file

@ -1,45 +1,48 @@
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<paragraph id="8BEB36D1EC147A15FF6CFAF47370FAD7" blockId="5.[151,613,1302,1365]" box="[151,613,1302,1329]" pageId="5" pageNumber="589">
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@ -55,7 +58,7 @@
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<paragraph id="8BEB36D1EC147A15FF6CFAD9700FFAB2" blockId="5.[151,613,1302,1365]" box="[151,282,1339,1365]" pageId="5" pageNumber="589">
(
<figureCitation id="136F2A54EC147A15FF64FAD971FDFAB2" box="[159,232,1339,1365]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="5" pageNumber="589">Figs 4</figureCitation>
<figureCitation id="136F2A54EC147A15FF64FAD971FDFAB2" box="[159,232,1339,1365]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="5" pageNumber="589">Figs 4</figureCitation>
, 12)
</paragraph>
</subSubSection>
@ -85,7 +88,7 @@
<emphasis id="B920EAC3EC147A15FF6CFA2770ABFA38" bold="true" box="[151,446,1477,1503]" pageId="5" pageNumber="589">Type material examined.</emphasis>
<typeStatus id="54EF8873EC147A15FE3FFA277339FA38" box="[452,556,1477,1503]" pageId="5" pageNumber="589" type="holotype">Holotype</typeStatus>
(
<figureCitation id="136F2A54EC147A15FDC0FA27736AFA38" box="[571,639,1477,1503]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="5" pageNumber="589">Fig. 4</figureCitation>
<figureCitation id="136F2A54EC147A15FDC0FA27736AFA38" box="[571,639,1477,1503]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="5" pageNumber="589">Fig. 4</figureCitation>
; dissected prior to the present study: the aedeagus and the apical genital segment are glued on the same card with the specimen): Kashmir,Chandanwari | b. Pahalgam,
<quantity id="4CAC9B34EC147A15FB07FA0B749EF9E4" box="[1276,1419,1513,1539]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="5" pageNumber="589" unit="m" value="2950.0" valueMax="3100.0" valueMin="2800.0">2800-3100m</quantity>
| lg.H.Franz,
@ -243,7 +246,7 @@ Measurements (n=90): HW: 0.530.60; HL: 0.300.39; OL: 0.150.17; AL (
<typeStatus id="54EF8873EC147A15FBBDF85F75BBF830" box="[1094,1198,1981,2007]" pageId="5" pageNumber="589" type="holotype">holotype</typeStatus>
: 3.37).
</paragraph>
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<emphasis id="B920EAC3EC177A16FF6CFB71700BFB4C" bold="true" box="[151,286,1171,1195]" pageId="6" pageNumber="590">FIGURE 11.</emphasis>
Distribution of
@ -255,9 +258,9 @@ in the Himalayan Region.
</caption>
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Body convex, elongate (
<figureCitation id="136F2A54EC177A16FE25FB1C7337FAFF" box="[478,546,1278,1304]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="6" pageNumber="590">Fig. 4</figureCitation>
<figureCitation id="136F2A54EC177A16FE25FB1C7337FAFF" box="[478,546,1278,1304]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="6" pageNumber="590">Fig. 4</figureCitation>
and
<figureCitation id="136F2A54EC177A16FDA6FB1C73B5FAF0" box="[605,672,1277,1304]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." pageId="6" pageNumber="590">
<figureCitation id="136F2A54EC177A16FDA6FB1C73B5FAF0" box="[605,672,1277,1304]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="6" pageNumber="590">
Fig.
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@ -306,7 +309,7 @@ and Kashmir to
) and
<collectingCountry id="F3437641EC167A17FF3CFE71701FFE4A" box="[199,266,403,429]" name="Nepal" pageId="7" pageNumber="591">Nepal</collectingCountry>
(
<figureCitation id="136F2A54EC167A17FEE2FE71707FFE4A" box="[281,362,403,429]" captionStart="FIGURE 11" captionStartId="6.[151,250,1171,1195]" captionTargetBox="[151,1435,181,1147]" captionTargetId="figure-16@6.[151,1435,181,1147]" captionTargetPageId="6" captionText="FIGURE 11. Distribution of Mannerheimia obscura in the Himalayan Region." pageId="7" pageNumber="591">Fig. 11</figureCitation>
<figureCitation id="136F2A54EC167A17FEE2FE71707FFE4A" box="[281,362,403,429]" captionStart="FIGURE 11" captionStartId="6.[151,250,1171,1195]" captionTargetBox="[151,1435,181,1147]" captionTargetId="figure-16@6.[151,1435,181,1147]" captionTargetPageId="6" captionText="FIGURE 11. Distribution of Mannerheimia obscura in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.12731016" httpUri="https://zenodo.org/record/12731016/files/figure.png" pageId="7" pageNumber="591">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BEB36D1EC167A17FF3CFE54729DFE12" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="591">
@ -341,7 +344,7 @@ recorded it from
and Kashmir, provided the photograph of the habitus (
<bibRefCitation id="EFC54B20EC167A17FD04FDA57289FD86" author="Shavrin, A. V." box="[767,924,583,609]" pageId="7" pageNumber="591" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021</bibRefCitation>
:
<figureCitation id="136F2A54EC167A17FC53FDA572F9FD86" box="[936,1004,583,609]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." pageId="7" pageNumber="591">Fig. 6</figureCitation>
<figureCitation id="136F2A54EC167A17FC53FDA572F9FD86" box="[936,1004,583,609]" captionStart="FIGURES 610" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 610. Aedeagus of Mannerheimia (Figs 67: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 910: M. emodensa (Nepal: Maharigaon): 6, 89parameral view, 7, 10lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="7" pageNumber="591">Fig. 6</figureCitation>
) and figures of the aedeagus (
<bibRefCitation id="EFC54B20EC167A17FAB8FDA571C6FD62" author="Shavrin, A. V." pageId="7" pageNumber="591" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021</bibRefCitation>
: Figs. 6162), and included this species to the key of the genus. Later, the same author recorded additional material from
@ -353,7 +356,7 @@ and Kashmir, provided the photograph of the habitus (
<emphasis id="B920EAC3EC167A17FDACFD6D7248FD4E" box="[599,861,655,681]" italics="true" pageId="7" pageNumber="591">Mannerheimia obscura</emphasis>
</taxonomicName>
is the most widely distributed species of the genus in the Himalayan Region (
<figureCitation id="136F2A54EC167A17FE44FD517304FD2A" box="[447,529,691,717]" captionStart="FIGURE 11" captionStartId="6.[151,250,1171,1195]" captionTargetBox="[151,1435,181,1147]" captionTargetId="figure-16@6.[151,1435,181,1147]" captionTargetPageId="6" captionText="FIGURE 11. Distribution of Mannerheimia obscura in the Himalayan Region." pageId="7" pageNumber="591">Fig. 11</figureCitation>
<figureCitation id="136F2A54EC167A17FE44FD517304FD2A" box="[447,529,691,717]" captionStart="FIGURE 11" captionStartId="6.[151,250,1171,1195]" captionTargetBox="[151,1435,181,1147]" captionTargetId="figure-16@6.[151,1435,181,1147]" captionTargetPageId="6" captionText="FIGURE 11. Distribution of Mannerheimia obscura in the Himalayan Region." figureDoi="http://doi.org/10.5281/zenodo.12731016" httpUri="https://zenodo.org/record/12731016/files/figure.png" pageId="7" pageNumber="591">Fig. 11</figureCitation>
). It is here recorded from
<collectingRegion id="4990F833EC167A17FCCEFD517517FD2A" box="[821,1026,691,717]" country="India" name="Himachal Pradesh" pageId="7" pageNumber="591">Himachal Pradesh</collectingRegion>
(

View file

@ -1,45 +1,48 @@
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(
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<figureCitation id="136F2A54EC167A17FF64FCA171F7FCBA" box="[159,226,834,861]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="7" pageNumber="591">Fig. 5</figureCitation>
)
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@ -81,7 +84,7 @@
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<specimenCount id="9D52FD58EC167A17FDCFFC2C735CFC0F" box="[564,585,974,1000]" count="1" pageId="7" pageNumber="591" type="male"></specimenCount>
(
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<figureCitation id="136F2A54EC167A17FDA3FC2F7389FC0F" box="[600,668,973,1000]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="7" pageNumber="591">Fig. 5</figureCitation>
; dissected prior to the present study: the aedeagus glued on the same card with the specimen):
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: Kashmir: Sonamarg | Himalaya-Hauptkette | Nichinai-Tal,
@ -125,7 +128,7 @@ from “
, Cachemire, Sonamarg…”. I study the
<typeStatus id="54EF8873EC167A17FEEEFB0F7062FAE0" box="[277,375,1261,1287]" pageId="7" pageNumber="591" type="holotype">holotype</typeStatus>
(
<figureCitation id="136F2A54EC167A17FE7DFB0F70DCFAEF" box="[390,457,1261,1288]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." pageId="7" pageNumber="591">Fig. 5</figureCitation>
<figureCitation id="136F2A54EC167A17FE7DFB0F70DCFAEF" box="[390,457,1261,1288]" captionStart="FIGURES 15" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 15. Habitus of Mannerheimia (holotypes): 1M. cachemirica, 2M. vicina, 3M. emodensa, 4M. obscura, 5M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="7" pageNumber="591">Fig. 5</figureCitation>
) from SF, but aedeagus is slightly damaged, thus I decide to not redescribe this species in the present study.
</paragraph>
</subSubSection>