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<mods:title>Electronoyesella antiqua Simutnik, gen. et sp. nov. (Chalcidoidea, Encyrtidae) from Rovno amber</mods:title>
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<mods:title id="2BE3C2D97038FF2C717137CEE95B2EB9">Electronoyesella antiqua Simutnik, gen. et sp. nov. (Chalcidoidea, Encyrtidae) from Rovno amber</mods:title>
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<mods:namePart>Simutnik, Serguei A.</mods:namePart>
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<mods:affiliation>I. I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev 01601, Ukraine</mods:affiliation>
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<mods:nameIdentifier type="email">simutnik@gmail.com</mods:nameIdentifier>
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<mods:namePart id="48F6C3B4988067F1CB1E2B68727AB82E">Simutnik, Serguei A.</mods:namePart>
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<mods:affiliation id="89F34D4E15D22BCC094D8FF7DFEC3CA3">I. I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev 01601, Ukraine</mods:affiliation>
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<mods:nameIdentifier id="7647AF5A0B76FA79F4917190A99092D0" type="email">simutnik@gmail.com</mods:nameIdentifier>
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<mods:namePart>Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation>I. I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev 01601, Ukraine</mods:affiliation>
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<mods:namePart id="6D3245AD265AF0F1C5D6DDD083E6088A">Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation id="A26B7FFBF0903450149DBF547009F9D2">I. I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev 01601, Ukraine</mods:affiliation>
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<mods:namePart>Vasilenko, Dmitry V.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4827-7290</mods:nameIdentifier>
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<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Cherepovets State University, Cherepovets, Vologda Region 162600, Russia</mods:affiliation>
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<mods:namePart id="96793BEB433BE518D2694DE90F50D618">Vasilenko, Dmitry V.</mods:namePart>
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<mods:affiliation id="596BEBEDF27387B9E74D6076823FF445">A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Cherepovets State University, Cherepovets, Vologda Region 162600, Russia</mods:affiliation>
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<mods:date>2022</mods:date>
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<mods:number>94</mods:number>
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<treatment LSID="urn:lsid:zoobank.org:act:5BFACFC4-6923-4E7B-B034-95DEEE352D78" httpUri="http://treatment.plazi.org/id/D1357CD7C5065E60928CB9DD88D82FCE" lastPageNumber="105" pageId="0" pageNumber="105">
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<treatment id="D1357CD7C5065E60928CB9DD88D82FCE" ID-GBIF-Taxon="206461055" LSID="urn:lsid:zoobank.org:act:5BFACFC4-6923-4E7B-B034-95DEEE352D78" httpUri="http://treatment.plazi.org/id/D1357CD7C5065E60928CB9DD88D82FCE" lastPageNumber="105" pageId="0" pageNumber="105" scope_family="Encyrtidae" scope_subFamily="Tetracneminae" scope_superFamily="Chalcidoidea">
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<paragraph id="BB1C952AB1F729E212EF08D83809E4CC" pageId="0" pageNumber="105">
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Genus
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<taxonomicName LSID="https://zoobank.org/5BFACFC4-6923-4E7B-B034-95DEEE352D78" authority="Simutnik" authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" rank="genus" status="gen. nov.">Electronoyesella Simutnik</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="105">gen. nov.</taxonomicNameLabel>
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||||
<taxonomicName id="2FE64860C6DA98851EF2C271626B64BC" LSID="https://zoobank.org/5BFACFC4-6923-4E7B-B034-95DEEE352D78" authority="Simutnik" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus" status="gen. nov.">Electronoyesella Simutnik</taxonomicName>
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||||
<taxonomicNameLabel id="BB36C40AD51B4D58EB399AC08860769C" pageId="0" pageNumber="105">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="105" type="type species">
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<paragraph pageId="0" pageNumber="105">Type species.</paragraph>
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||||
<paragraph pageId="0" pageNumber="105">
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||||
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
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||||
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
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<subSubSection id="8BA27B439A60B45075672CAF1D0DED95" pageId="0" pageNumber="105" type="type species">
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<paragraph id="7FF4028F7A8114348F950F604016D299" pageId="0" pageNumber="105">Type species.</paragraph>
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<paragraph id="47781282BD1A84C106DD740159C52493" pageId="0" pageNumber="105">
|
||||
<taxonomicName id="4D4C1C312B50A79A3393DE2E6FB18557" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
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||||
<emphasis id="D296E2B86017238369F80FC56772DDA8" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
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</taxonomicName>
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Simutnik, sp. nov.
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</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="105" type="species composition">
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||||
<paragraph pageId="0" pageNumber="105">Species composition.</paragraph>
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||||
<paragraph pageId="0" pageNumber="105">Type species only.</paragraph>
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||||
<subSubSection id="96BE1B450DB54EA1577F9C3948C46F7A" pageId="0" pageNumber="105" type="species composition">
|
||||
<paragraph id="095622053C6502F28CF75A3761A4C36C" pageId="0" pageNumber="105">Species composition.</paragraph>
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||||
<paragraph id="460E66A0117C413410A26E7FAF9BB549" pageId="0" pageNumber="105">Type species only.</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="105" type="etymology">
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||||
<paragraph pageId="0" pageNumber="105">Etymology.</paragraph>
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||||
<paragraph pageId="0" pageNumber="105">
|
||||
<subSubSection id="A4AEF73D4BA5BEA182157ED30ACB506A" pageId="0" pageNumber="105" type="etymology">
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<paragraph id="7EF73D8B90F3D448DABB5862EBFD153B" pageId="0" pageNumber="105">Etymology.</paragraph>
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||||
<paragraph id="CD30F87DF38A702B734731E7062B5B55" pageId="0" pageNumber="105">
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||||
The new genus is named in honor of John Stuart Noyes, who first saw the presence of the structures similar to paratergites in the new fossil. The name is a combination of
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||||
<normalizedToken originalValue="“electrum”">"electrum"</normalizedToken>
|
||||
<normalizedToken id="F42FAE025CD7EC23AB5FA3B5F7CFEA65" originalValue="“electrum”">"electrum"</normalizedToken>
|
||||
(Latin:
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">electrum</emphasis>
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<emphasis id="863CDBABD00CFEEBC6E82042C1E614D9" italics="true" pageId="0" pageNumber="105">electrum</emphasis>
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= amber) and
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||||
<normalizedToken originalValue="“Noyes”">"Noyes"</normalizedToken>
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||||
<normalizedToken id="8E40F497773CC9BF8B47FE7742F3465B" originalValue="“Noyes”">"Noyes"</normalizedToken>
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. The genus name is a feminine noun.
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||||
</paragraph>
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||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="105" type="diagnosis">
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<paragraph pageId="0" pageNumber="105">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="105">
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<emphasis bold="true" pageId="0" pageNumber="105">Female.</emphasis>
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<subSubSection id="89A4BECEDE95E4C46FB686B4FD86194B" pageId="0" pageNumber="105" type="diagnosis">
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||||
<paragraph id="76484F38E545958AF84387272F40EB64" pageId="0" pageNumber="105">Diagnosis.</paragraph>
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||||
<paragraph id="B4ABE433999ABCDA0650DA2BC887229C" pageId="0" pageNumber="105">
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<emphasis id="280D6EB5E42230D71303019284533E57" bold="true" pageId="0" pageNumber="105">Female.</emphasis>
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Body not flattened; frontovertex as long as broad, with four vertical rows of piliferous punctures (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3A</figureCitation>
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||||
<figureCitation id="6DBD7BD4E8369B005D2E7BD71C42AE00" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3A</figureCitation>
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); ocelli forming right angled triangle; horizontal row of three large, deep cells under each torulus (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
|
||||
<figureCitation id="4E64E8D8E7D8DC06026D79B7BA984680" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
|
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); clypeus and interantennal prominence intricately sculptured; mandibles tridentate with middle tooth longest; scape much more than 3
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||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="B5BD34B1CF8F0DB1B39698939CDBB2D6" originalValue="×">x</normalizedToken>
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as long as broad; F1 a little longer than broad to quadrate; notauli present only anteriorly as small but distinct depressions (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3B</figureCitation>
|
||||
<figureCitation id="092B36E959E230B834459AAF61A032D2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3B</figureCitation>
|
||||
: arrow), without visible lines; filum spinosum absent; covering setae present; marginal vein 5
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="CDBE6FD1B9D37B52CD11F6AE1D69AC8C" originalValue="×">x</normalizedToken>
|
||||
as long as broad; postmarginal vein 1.5
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="B43625FCFEF4185E4F87E047997EF010" originalValue="×">x</normalizedToken>
|
||||
as long as marginal; costal cell of hind wing with line of long setae (Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
<figureCitation id="6CF563AC4F11075A4E7E60346AF4AAAA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
: ls1), longest of which located alongside parastigma (Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
<figureCitation id="4827CB5090AE460B2C555AEB33A55F6B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
: pst); row of thickened setae present alongside hyaline spur vein of hind wing (Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
<figureCitation id="939EDFE0B9D67C087A5F903504539CE0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
: ls2, spv); apex of metatibia with one peg (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
|
||||
<figureCitation id="E1D370D7EAA018F5EB1B39C246A39086" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
|
||||
: arrows); cerci located in apical third of metasoma; paratergite-like, sclerotized, separated part of Mt8 present alongside margin of syntergum (Figs
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6A, B</figureCitation>
|
||||
<figureCitation id="0761D54D9AA64E70249CCC1323AD073E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6A, B</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7C</figureCitation>
|
||||
<figureCitation id="A64D8A3E7EB6B1A5F5CB241CB94B36C6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7C</figureCitation>
|
||||
: ptrg?); apex of hypopygium reaching a little way past apex of last gastral tergum.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="105">
|
||||
<emphasis bold="true" pageId="0" pageNumber="105">Male.</emphasis>
|
||||
<paragraph id="41D8BDB56BA30BA2E21ED7602E66D559" pageId="0" pageNumber="105">
|
||||
<emphasis id="C4C1CEED9E8A08945ACFE54310A5D73F" bold="true" pageId="0" pageNumber="105">Male.</emphasis>
|
||||
Unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="105" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="105">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="105">
|
||||
<subSubSection id="F02802ABD8EF9F450CEB736D901CA73D" pageId="0" pageNumber="105" type="remarks">
|
||||
<paragraph id="FD878A5A4D9E1B6E83C5D2785FE99423" pageId="0" pageNumber="105">Remarks.</paragraph>
|
||||
<paragraph id="DA7A83BA7125AEC3FFA5C3403CD07AFA" pageId="0" pageNumber="105">
|
||||
Placement of
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella</emphasis>
|
||||
<taxonomicName id="DCFD29B8ACF966D376D6DBFD587919CB" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="C938C6255F4B4DC9C8C3D88C5D436A78" italics="true" pageId="0" pageNumber="105">Electronoyesella</emphasis>
|
||||
</taxonomicName>
|
||||
gen. nov. in
|
||||
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
|
||||
<taxonomicName id="5B613C555EFFF85410F486CB24F955CB" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
|
||||
is supported by the absence of the filum spinosum of linea calva, the tridentate mandibles with the middle tooth being the longest, the hypopygium reaching a little way past the apex of the syntergum, and the presence of the sclerotized, ribbon-like, separated part of Mt8 along the margin of the syntergum (Figs
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6A, B</figureCitation>
|
||||
<figureCitation id="D7E26ED4DC90417A2636792397786541" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6A, B</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7C</figureCitation>
|
||||
<figureCitation id="A578548D254E0D8AA5FAE8E31429AD38" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7C</figureCitation>
|
||||
: ptrg?). This structure closely resembles the paratergites of extant
|
||||
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
|
||||
<taxonomicName id="96D0D90170DA605271228897361AE882" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7A, B</figureCitation>
|
||||
<figureCitation id="77995153EB092B9840E69F0F1F2A6763" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7A, B</figureCitation>
|
||||
: ptrg) and may be morphological evidence for the existence of this subfamily in the late Eocene.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="105">
|
||||
<paragraph id="D5CC5AFE8FB8122905F5B6420BE23389" pageId="0" pageNumber="105">
|
||||
The hind wing of the new genus has a single line of long setae alongside the entire costal cell (Fig.
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
<figureCitation id="6C63DD4D58A2CEE52E70747A5A985342" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
: ls1) as in most extant
|
||||
<taxonomicName authorityName="Howard" authorityYear="1890" class="Insecta" family="Tanaostigmatidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Tanaostigmatidae</taxonomicName>
|
||||
<taxonomicName id="F3E625BBF08190E63030C4E7684F19A9" authorityName="Howard" authorityYear="1890" class="Insecta" family="Tanaostigmatidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Tanaostigmatidae</taxonomicName>
|
||||
. These setae vary in length, the longest located along the parastigma of the hind wing. In fossil
|
||||
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
|
||||
<taxonomicName id="28D73FE4294CD644EE998D3FB005F507" baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
|
||||
, the same line of long setae has been recorded in
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2002" class="Insecta" family="Encyrtidae" genus="Eocencnemus" kingdom="Animalia" lsidName="Eocencnemus sugonjaevi" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="sugonjaevi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Eocencnemus sugonjaevi</emphasis>
|
||||
<taxonomicName id="79F248C94504F6566206780CB2168040" authorityName="Simutnik" authorityYear="2002" class="Insecta" family="Encyrtidae" genus="Eocencnemus" kingdom="Animalia" lsidName="Eocencnemus sugonjaevi" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="sugonjaevi">
|
||||
<emphasis id="A31626CC08767D3E94953D4C70722AD2" italics="true" pageId="0" pageNumber="105">Eocencnemus sugonjaevi</emphasis>
|
||||
</taxonomicName>
|
||||
Simutnik, 2002,
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2015" class="Insecta" family="Encyrtidae" genus="Sulia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sulia glaesaria" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="glaesaria">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Sulia glaesaria</emphasis>
|
||||
<taxonomicName id="93DBF20C39900B1FC64FDA08C03CEFB0" authorityName="Simutnik" authorityYear="2015" class="Insecta" genus="Sulia" kingdom="Animalia" lsidName="Sulia glaesaria" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="glaesaria">
|
||||
<emphasis id="5C4CFB588BE5C71E1CEE4D571F7F4148" italics="true" pageId="0" pageNumber="105">Sulia glaesaria</emphasis>
|
||||
</taxonomicName>
|
||||
Simutnik, 2015 (Simutnik et al. 2021), and is also known in several undescribed specimens without filum spinosum. These differ from the new genus by the absence of a vertical rows of large piliferous punctures on the frontovertex.
|
||||
<taxonomicName lsidName="E. sugonjaevi" pageId="0" pageNumber="105" rank="species" species="sugonjaevi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">E. sugonjaevi</emphasis>
|
||||
<taxonomicName id="99072558D395CBE643BB93F8D5B48AAE" lsidName="E. sugonjaevi" pageId="0" pageNumber="105" rank="species" species="sugonjaevi">
|
||||
<emphasis id="A3D45D9072F1582B9C95756E13988CB9" italics="true" pageId="0" pageNumber="105">E. sugonjaevi</emphasis>
|
||||
</taxonomicName>
|
||||
also differs by its short, ring-like F1.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="105">
|
||||
<paragraph id="9A7F8AB1A241D6AE9F8378D54FD5E79C" pageId="0" pageNumber="105">
|
||||
The same line of long setae alongside the costal cell of the hind wing is present in few extant
|
||||
<taxonomicName lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
|
||||
<taxonomicName id="31A943E196560D11E8B454934DA41688" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
|
||||
:
|
||||
<taxonomicName authorityName="Ashmead" authorityYear="1904" class="Insecta" family="Encyrtidae" genus="Exoristobia" kingdom="Animalia" lsidName="Exoristobia" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Exoristobia</emphasis>
|
||||
<taxonomicName id="27D726502D85C2EDC59FEB9CE4EB3028" authorityName="Ashmead" authorityYear="1904" class="Insecta" family="Encyrtidae" genus="Exoristobia" kingdom="Animalia" lsidName="Exoristobia" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="72594E2DF734754AD4DAB289DE8AF2A6" italics="true" pageId="0" pageNumber="105">Exoristobia</emphasis>
|
||||
</taxonomicName>
|
||||
Ashmead, 1904 (Simutnik et al. 2021),
|
||||
<taxonomicName authorityName="Ashmead" authorityYear="1900" class="Insecta" family="Encyrtidae" genus="Rhytidothorax" kingdom="Animalia" lsidName="Rhytidothorax" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Rhytidothorax</emphasis>
|
||||
<taxonomicName id="AB467BCF388DC368362498906E1D8322" authorityName="Ashmead" authorityYear="1900" class="Insecta" family="Encyrtidae" genus="Rhytidothorax" kingdom="Animalia" lsidName="Rhytidothorax" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A182331F5BE36FE2E509D23E3252DC3E" italics="true" pageId="0" pageNumber="105">Rhytidothorax</emphasis>
|
||||
</taxonomicName>
|
||||
Ashmead, 1900, and some other
|
||||
<taxonomicName authorityName="Howard" authorityYear="1895" lsidName="" pageId="0" pageNumber="105" rank="tribe" tribe="Bothriothoracini">Bothriothoracini</taxonomicName>
|
||||
<taxonomicName id="C3A15AF1042C54D0E1A62AD0ADBE222C" authorityName="Howard" authorityYear="1895" lsidName="" pageId="0" pageNumber="105" rank="tribe" tribe="Bothriothoracini">Bothriothoracini</taxonomicName>
|
||||
Howard, 1895 (J. S. Noyes, pers. comm. 2022). In
|
||||
<taxonomicName authorityName="Mercet" authorityYear="1921" class="Insecta" family="Encyrtidae" genus="Aphycoides" kingdom="Animalia" lsidName="Aphycoides" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Aphycoides</emphasis>
|
||||
<taxonomicName id="0A7A05C57FD84AC8799E01EFE338CECF" authorityName="Mercet" authorityYear="1921" class="Insecta" family="Encyrtidae" genus="Aphycoides" kingdom="Animalia" lsidName="Aphycoides" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="52CCEE2E007F94B2E0C444F5E9264338" italics="true" pageId="0" pageNumber="105">Aphycoides</emphasis>
|
||||
</taxonomicName>
|
||||
Mercet, 1921 these setae are sparser and shorter. Within
|
||||
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
|
||||
<taxonomicName id="8E13175CE0C281F81C4A594647136B15" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
|
||||
, a somewhat similar line of setae is present in
|
||||
<taxonomicName class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
|
||||
<taxonomicName id="40C44355C912D590F636DC652FEE5DD8" class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="9A30840D6DFD1710CEE234C53D885AF4" italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
|
||||
</taxonomicName>
|
||||
Walker, 1837 and
|
||||
<taxonomicName authorityName="Hoffer" authorityYear="1954" class="Insecta" family="Encyrtidae" genus="Moraviella" kingdom="Animalia" lsidName="Moraviella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Moraviella</emphasis>
|
||||
<taxonomicName id="10D1274CEA45601D83BE3C5E732595CC" authorityName="Hoffer" authorityYear="1954" class="Insecta" family="Encyrtidae" genus="Moraviella" kingdom="Animalia" lsidName="Moraviella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="1A3BA6BB3E358AC9D12995BFE4D73F9A" italics="true" pageId="0" pageNumber="105">Moraviella</emphasis>
|
||||
</taxonomicName>
|
||||
Hoffer, 1954, but their setae do not vary in length. A line or several lines of setae are present in
|
||||
<taxonomicName authorityName="Walker" authorityYear="1846" class="Insecta" family="Encyrtidae" genus="Aenasius" kingdom="Animalia" lsidName="Aenasius" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Aenasius</emphasis>
|
||||
<taxonomicName id="9686E8E29DB8BE698F8AB961A213E0B5" authorityName="Walker" authorityYear="1846" class="Insecta" family="Encyrtidae" genus="Aenasius" kingdom="Animalia" lsidName="Aenasius" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="AC16C23C448C96451754F7B792E04DD5" italics="true" pageId="0" pageNumber="105">Aenasius</emphasis>
|
||||
</taxonomicName>
|
||||
Walker, 1846;
|
||||
<taxonomicName authorityName="Howard" authorityYear="1898" class="Insecta" family="Encyrtidae" genus="Blepyrus" kingdom="Animalia" lsidName="Blepyrus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Blepyrus</emphasis>
|
||||
<taxonomicName id="194F9E1D6F42031F9F31C16FDD4D08A8" authorityName="Howard" authorityYear="1898" class="Insecta" family="Encyrtidae" genus="Blepyrus" kingdom="Animalia" lsidName="Blepyrus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="043B3E18A8659126859D359A709DE43D" italics="true" pageId="0" pageNumber="105">Blepyrus</emphasis>
|
||||
</taxonomicName>
|
||||
Howard, 1898;
|
||||
<taxonomicName class="Insecta" family="Encyrtidae" genus="Monodiscodes" kingdom="Animalia" lsidName="Monodiscodes" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Monodiscodes</emphasis>
|
||||
<taxonomicName id="7616D4C760640F31F48772A7F526BD2E" class="Insecta" family="Encyrtidae" genus="Monodiscodes" kingdom="Animalia" lsidName="Monodiscodes" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="9FED762A661C31A3EAABADF9637474EB" italics="true" pageId="0" pageNumber="105">Monodiscodes</emphasis>
|
||||
</taxonomicName>
|
||||
Hoffer, 1953, but they are also short.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="105">
|
||||
<paragraph id="7F648E4B6CD1DDEC6922E503E2266BBC" pageId="0" pageNumber="105">
|
||||
A line of long setae, but sparser and more or less equal in length originating from the membrane of the costal cell, similar to these of
|
||||
<taxonomicName class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
|
||||
<taxonomicName id="2B9162710B203B4A0B7BC49CC57336C0" class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3E63B880174D66C3A66F6021635ACA63" italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
|
||||
</taxonomicName>
|
||||
, are also found in the earliest known Sakhalinian amber encyrtids:
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2015" class="Insecta" family="Encyrtidae" genus="Sugonjaevia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sugonjaevia" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Sugonjaevia</emphasis>
|
||||
<taxonomicName id="B31CF6CEE5E6C7AB0AD1A62C313EADE4" authorityName="Simutnik" authorityYear="2015" class="Insecta" family="Encyrtidae" genus="Sugonjaevia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sugonjaevia" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="E3AB7DD4AA8E11B3B4CA78B146154116" italics="true" pageId="0" pageNumber="105">Sugonjaevia</emphasis>
|
||||
</taxonomicName>
|
||||
Simutnik, 2015,
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2020" class="Insecta" family="Encyrtidae" genus="Encyrtoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Encyrtoides" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Encyrtoides</emphasis>
|
||||
<taxonomicName id="F3A6F57D07CF85AE74810590A7AA33C9" authorityName="Simutnik" authorityYear="2020" class="Insecta" family="Encyrtidae" genus="Encyrtoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Encyrtoides" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="5E6AC01C5090902D1BAF2A233ECB002E" italics="true" pageId="0" pageNumber="105">Encyrtoides</emphasis>
|
||||
</taxonomicName>
|
||||
Simutnik, 2021, and
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2021" class="Insecta" family="Encyrtidae" genus="Sakhalinencyrtus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sakhalinencyrtus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Sakhalinencyrtus</emphasis>
|
||||
<taxonomicName id="3F96880BCE62C7A386CB164CC461FE20" authorityName="Simutnik" authorityYear="2021" class="Insecta" genus="Sakhalinencyrtus" kingdom="Animalia" lsidName="Sakhalinencyrtus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="34F57F08681E1CD1621A8EBBE99D14BE" italics="true" pageId="0" pageNumber="105">Sakhalinencyrtus</emphasis>
|
||||
</taxonomicName>
|
||||
Simutnik, 2021. This character state seems to be a feature of the stem group of
|
||||
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
|
||||
<taxonomicName id="1228600A15A05F61F09F9DFE8C3AA39D" baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="105">
|
||||
<paragraph id="3474E225DD0EA464A07D6B33DA4C8AF0" pageId="0" pageNumber="105">
|
||||
The first funicular segment seen in the majority of known Eocene encyrtid females, including the oldest described female from middle Eocene Sakhalinian amber (
|
||||
<bibRefCitation DOI="https://doi.org/10.1080/08912963.2020.1835887" author="Simutnik, SA" journalOrPublisher="Entomological Review" pageId="0" pageNumber="105" refId="B14" refString="Simutnik, SA, 2021. https://doi.org/10.1080/08912963.2020.1835887" url="https://doi.org/10.1080/08912963.2020.1835887" year="2021">Simutnik 2021</bibRefCitation>
|
||||
<bibRefCitation id="4051C7040EDA53FFA4E9FD259D778473" DOI="https://doi.org/10.1080/08912963.2020.1835887" author="Simutnik, SA" journalOrPublisher="Entomological Review" pageId="0" pageNumber="105" refId="B14" refString="Simutnik, SA, 2021. https://doi.org/10.1080/08912963.2020.1835887" url="https://doi.org/10.1080/08912963.2020.1835887" year="2021">Simutnik 2021</bibRefCitation>
|
||||
), had an annular shape, or at least is broader than long. Of the 27 examined females of European and Sakhalinian ambers (some of which might belong to the same species), some undescribed, few have an F1 longer than broad. Among Eocene females without the filum spinosum (supposedly
|
||||
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
|
||||
<taxonomicName id="04441733732C1A12184C3E0836E5B16D" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
|
||||
), only in
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2015" class="Insecta" family="Encyrtidae" genus="Sulia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sulia" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Sulia</emphasis>
|
||||
<taxonomicName id="96EEF8A9D5E2E9D39DC6D36168311E5B" authorityName="Simutnik" authorityYear="2015" class="Insecta" genus="Sulia" kingdom="Animalia" lsidName="Sulia" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="590940F625468471970C26B8E1116379" italics="true" pageId="0" pageNumber="105">Sulia</emphasis>
|
||||
</taxonomicName>
|
||||
Simutnik, 2015 and
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Rovnopositor" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rovnopositor" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Rovnopositor</emphasis>
|
||||
<taxonomicName id="8BE37B60F180AE8BF48FD8548C1A5E6D" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Rovnopositor" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rovnopositor" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="4ED79A8DAD5AC1EDDDEB342205512472" italics="true" pageId="0" pageNumber="105">Rovnopositor</emphasis>
|
||||
</taxonomicName>
|
||||
Simutnik, 2022, is F1 about 1.5
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="2C6BCCAC905B28840155DC2FCBDA1633" originalValue="×">x</normalizedToken>
|
||||
as long as broad (the latter differs from the new genus by the absence of vertical rows of piliferous punctures on the frontovertex, its shorter marginal vein, the reduced gonostyli, and by its long, curved ovipositor stylet). F1 in
|
||||
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella</emphasis>
|
||||
<taxonomicName id="3F5E783D6320A69311873CBA51A871E0" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A1C6D3C7075A576650389F54FA865E8F" italics="true" pageId="0" pageNumber="105">Electronoyesella</emphasis>
|
||||
</taxonomicName>
|
||||
is slightly longer than broad (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3A, B</figureCitation>
|
||||
<figureCitation id="1458F11E4C7F2D66C7DFD5BB9E8A1E4D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="105">
|
||||
<paragraph id="D6EB9B2115AA743D8C6190237F1E9249" pageId="0" pageNumber="105">
|
||||
The new taxon further differs from most known
|
||||
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
|
||||
<taxonomicName id="965CBBEFF66B139E1329705A59DE30F6" baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
|
||||
by the presence of a peg or spicule originating from the round, deep pit at the apex of metatibia (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
|
||||
<figureCitation id="73197EA5A18D88F6B30F9EB57244506B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
|
||||
, arrows); and a row of thickened setae alongside the spur vein of the hind wing (Figs
|
||||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
<figureCitation id="62C2FFBD6FFB2CDD68A1954DF964986B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
|
||||
<figureCitation id="3036F2965E6F4763CC715E2A95ABB6C6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
|
||||
: spv). Such a row of setae is also absent in the middle Eocene encyrtids from the Sakhalinian amber.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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