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<mods:namePart>Wei, Meicai</mods:namePart>
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<treatment ID-GBIF-Taxon="152032829" LSID="urn:lsid:zoobank.org:act:1B386FA8-BF03-4AD1-A979-C0FC4C900077" httpUri="http://treatment.plazi.org/id/C40A5512983BB765D5E7F40FD464CDDD" lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86">
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<taxonomicName LSID="http://species-id.net/wiki/Anhoplocampa_yunanensis" authority="Haris &amp; Roller, 1999" authorityName="Haris &amp; Roller" authorityYear="1999" genus="Anhoplocampa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anhoplocampa yunanensis" pageId="5" pageNumber="86" rank="species" species="yunanensis">
<pageBreakToken pageId="5" pageNumber="86" start="start">Anhoplocampa</pageBreakToken>
<treatment id="C40A5512983BB765D5E7F40FD464CDDD" ID-GBIF-Taxon="152032829" LSID="urn:lsid:zoobank.org:act:1B386FA8-BF03-4AD1-A979-C0FC4C900077" httpUri="http://treatment.plazi.org/id/C40A5512983BB765D5E7F40FD464CDDD" lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86" scope_family="Tenthredinidae" scope_order="Hymenoptera">
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<paragraph id="70783B75E0CA6199B0F64571E623C89C" pageId="5" pageNumber="86">
<taxonomicName id="3673EAF0512BE66151EBA7B82244D657" LSID="http://species-id.net/wiki/Anhoplocampa_yunanensis" authority="Haris &amp; Roller, 1999" authorityName="Haris &amp; Roller" authorityYear="1999" genus="Anhoplocampa" lsidName="Anhoplocampa yunanensis" pageId="5" pageNumber="86" rank="species" species="yunanensis">
<pageBreakToken id="150938F9471F7D8DAD00F3A65775BD95" pageId="5" pageNumber="86" start="start">Anhoplocampa</pageBreakToken>
yunanensis (Haris &amp; Roller, 1999)
</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="86">comb. n.</taxonomicNameLabel>
<taxonomicNameLabel id="307B4A087B6F16C44A2D3E85B59F80C1" pageId="5" pageNumber="86">comb. n.</taxonomicNameLabel>
Figs 7-13
</paragraph>
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<subSubSection pageId="5" pageNumber="86" type="reference_group">
<paragraph pageId="5" pageNumber="86">
<taxonomicName class="Insecta" family="Tenthredinidae" genus="Trichiocampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trichiocampus yunanensis" order="Hymenoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="yunanensis">Trichiocampus yunanensis</taxonomicName>
<subSubSection id="3ACDCDE3412C8DFCD7F8EFFB1FDF0E61" pageId="5" pageNumber="86" type="reference_group">
<paragraph id="557E3C46F735B5B40927C58F74075B5B" pageId="5" pageNumber="86">
<taxonomicName id="09ACF8D2F4EEDC348BCCB54C62FBFA07" genus="Trichiocampus" lsidName="Trichiocampus yunanensis" pageId="5" pageNumber="86" rank="species" species="yunanensis">Trichiocampus yunanensis</taxonomicName>
Haris &amp; Roller, 1999: 231-232.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="86" type="description">
<paragraph pageId="5" pageNumber="86">Description.</paragraph>
<paragraph pageId="5" pageNumber="86">Female (Fig. 7). Body length 7 mm (excluding antenna, cerci and sheath). Body orange, extreme narrow margin of antennal socket, propleuron, parapsis of mesothorax largely, narrow posterior margin of mesoscutellum, parapsis of metanotum, posterior of metascutellum, ventral half of mesopleuron, ventral margin of metapleuron, black; legs black, apical 1/3 of fore femur, all tibiae and tarsi yellow brown. Wings hyaline, apical third weakly infuscate, vein C pale brown, other veins and pterostigma black brown, anterior breadth of a feebly defined smoky macula about half length of pterostigma. Body hairs silver brown.</paragraph>
<paragraph pageId="5" pageNumber="86">
<subSubSection id="93B7D8EBC77EAD7F6C1CE9AA7CF960A7" pageId="5" pageNumber="86" type="description">
<paragraph id="57A15979D06F71681CA11FC287B52C82" pageId="5" pageNumber="86">Description.</paragraph>
<paragraph id="5E5608BD087B2AF231E42806CB209CD0" pageId="5" pageNumber="86">Female (Fig. 7). Body length 7 mm (excluding antenna, cerci and sheath). Body orange, extreme narrow margin of antennal socket, propleuron, parapsis of mesothorax largely, narrow posterior margin of mesoscutellum, parapsis of metanotum, posterior of metascutellum, ventral half of mesopleuron, ventral margin of metapleuron, black; legs black, apical 1/3 of fore femur, all tibiae and tarsi yellow brown. Wings hyaline, apical third weakly infuscate, vein C pale brown, other veins and pterostigma black brown, anterior breadth of a feebly defined smoky macula about half length of pterostigma. Body hairs silver brown.</paragraph>
<paragraph id="00ED513C5B9796069689569869ECE27A" pageId="5" pageNumber="86">
Body shiny, dorsal side of pronotum distinctly punctured, dorsal half of mesopleuron distinctly microsculptured (Fig. 9). Clypeus distinctly and roundly emarginated to about half length of clypeus (Fig. 10); malar space about 1.15
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="F9B09099FB5537BE2B3CB5D21C92A685" originalValue="×">x</normalizedToken>
diameter of lateral ocellus; postocellar area flat, 2.2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="6DF407E2C55D9FB671B411A67E959CC3" originalValue="×">x</normalizedToken>
as broad as long, with a deep middle fovea and a shallow middle furrow, posterior of postocellar area not declined; postocellar furrow broad and shallow; lateral furrows deep and very broad, weakly divergent backwards; temple flat; frontal basin shallow, 1.1
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9CCD78F67BFB94C8DAD1E0C5EB0ED807" originalValue="×">x</normalizedToken>
as broad as long, with depth about 0.13
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9F8BD27FFCDD3EB2AB92D33464DB5D31" originalValue="×">x</normalizedToken>
breadth of basin; frontal walls distinctly elevated, not very sharp (Fig. 8); distance between eyes at level of antennal sockets 1.8
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="A3101E2E7DFED93A32B73C0866B04C4C" originalValue="×">x</normalizedToken>
wider than greater diameter of eye. Antenna stout, 1.2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="5204D26E2B6059722D44960586BFBD62" originalValue="×">x</normalizedToken>
length of abdomen, much shorter than thorax and abdomen together, tapering towards apex (Fig. 7). Prepectus, epicnemium and dorsal part of mesepisternum as Fig. 9. Claw with inner tooth clearly shorter and slightly broader than outer tooth (Fig. 11). Forewing: cell 2Rs slightly shorter than 1Rs, vein cu-a joining cell 1M at middle. Cerci and apical sheath in dorsal view as Fig. 12, cerci about 3.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="64F43D36D5EA29519B7EAFD59E19AA1C" originalValue="×">x</normalizedToken>
as long as broad, not beyond apex of sheath. Lancet slender, with 14 annuli and 16 serrulae, annuli simple, without annular teeth (Fig. 13).
</paragraph>
<paragraph pageId="5" pageNumber="86">Male. Unknown.</paragraph>
<caption pageId="5" pageNumber="86">
<paragraph pageId="5" pageNumber="86">
<paragraph id="26E412BC532B40803CD9F0331AC021B3" pageId="5" pageNumber="86">Male. Unknown.</paragraph>
<caption id="BC3FB65803843B8D92CBE0A6D71F425A" pageId="5" pageNumber="86">
<paragraph id="44B0AAA2716F225B3D7847E15506EC3C" pageId="5" pageNumber="86">
Figures 7-13.
<taxonomicName genus="Anhoplocampa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anhoplocampa yunanensis" pageId="5" pageNumber="86" rank="species" species="yunanensis">Anhoplocampa yunanensis</taxonomicName>
<taxonomicName id="B0A074DEE5167327E108AA9E603A4C24" genus="Anhoplocampa" lsidName="Anhoplocampa yunanensis" pageId="5" pageNumber="86" rank="species" species="yunanensis">Anhoplocampa yunanensis</taxonomicName>
(Haris &amp; Roller, 1999) comb. n., specimen from Xiaozhongdian, Yunnan 7 Adult female, dorsal view (scale bar = 1mm) 8 Head, dorsal view 9 Prepectus, epicnemium and dorsal part of mesepisternum 10 Clypeus 11 Claw 12 Cerci and apical sheath, dorsal view 13 Lancet
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="5" pageNumber="86" type="distribution">
<paragraph pageId="5" pageNumber="86">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="86">China (Yunnan).</paragraph>
<subSubSection id="D939C454DF08778EFA10D394684F7EBE" pageId="5" pageNumber="86" type="distribution">
<paragraph id="485D34ACA47739450D68B245D85EBE52" pageId="5" pageNumber="86">Distribution.</paragraph>
<paragraph id="3194AC6697AA4016B36FC4600D73B04E" pageId="5" pageNumber="86">China (Yunnan).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="86" type="specimens examined">
<paragraph pageId="5" pageNumber="86">Specimens examined.</paragraph>
<paragraph pageId="5" pageNumber="86">1♀, holotype: &quot;China, Yunan [Yunnan], 24-29. VI. 93, 50 km N. of Lijieng [Lijiang], Yulongshan, Nat. Res., E. Jednek, O. Sausa leg.&quot; (HNHM); 1 female, &quot;Xiaozhongdian, Shangri-La, Yunnan, China, alt. 3000m, 19 July 2004, Xiao Wei coll.&quot; (CSCS).</paragraph>
<subSubSection id="C0F08464D1A6163943C5D6AF00AB6FAB" pageId="5" pageNumber="86" type="specimens examined">
<paragraph id="24DBE58BB204CFFF26822AA5FA0C311C" pageId="5" pageNumber="86">Specimens examined.</paragraph>
<paragraph id="F7EB096D5A0313CE0FD14D75E10A9E1E" pageId="5" pageNumber="86">1♀, holotype: &quot;China, Yunan [Yunnan], 24-29. VI. 93, 50 km N. of Lijieng [Lijiang], Yulongshan, Nat. Res., E. Jednek, O. Sausa leg.&quot; (HNHM); 1 female, &quot;Xiaozhongdian, Shangri-La, Yunnan, China, alt. 3000m, 19 July 2004, Xiao Wei coll.&quot; (CSCS).</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86" type="remarks">
<paragraph pageId="5" pageNumber="86">Remarks.</paragraph>
<paragraph pageId="5" pageNumber="86">
<subSubSection id="EF944B1647C75F7AAB0B2DC6FFC543CE" lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86" type="remarks">
<paragraph id="F4E234E586D387E47A433CF34DF8B29F" pageId="5" pageNumber="86">Remarks.</paragraph>
<paragraph id="0C1AF8742AB2AFFAEF42A8E698A02BDA" pageId="5" pageNumber="86">
This species is easily separated from the other
<taxonomicName genus="Anhoplocampa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anhoplocampa" pageId="5" pageNumber="86" rank="genus">Anhoplocampa</taxonomicName>
<taxonomicName id="B054FDA008D2340702FFFB801B50F9FB" genus="Anhoplocampa" lsidName="Anhoplocampa" pageId="5" pageNumber="86" rank="genus">Anhoplocampa</taxonomicName>
species by the orange body; antenna stout and much shorter than body; forewing with a weak cross band below pterostigma; postocellar area not declined and with deep and broad lateral furrows; the upper half of mesopleuron distinctly microsculptured; cerci shorter; clypeus deeply incised; claw with inner tooth clearly shorter than outer tooth; cu-a of fore wing meeting cell 1M at about middle; and female lancet slender with 14 simple annuli.
</paragraph>
<paragraph lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86">
<bibRefCitation author="Haris, A" journalOrPublisher="Folia Entomologica Hungarica" pageId="8" pageNumber="89" pagination="231 - 237" title="Four new sawfly species from Yunan (Hymenoptera: Tenthredinidae)." volume="60" year="1999">Haris and Roller (1999)</bibRefCitation>
<paragraph id="880B759FE69C464FD082AC9D6042D0DE" lastPageId="6" lastPageNumber="87" pageId="5" pageNumber="86">
<bibRefCitation id="97981F2490B63A0DF84373A77C14D2BA" author="Haris, A" journalOrPublisher="Folia Entomologica Hungarica" pageId="8" pageNumber="89" pagination="231 - 237" title="Four new sawfly species from Yunan (Hymenoptera: Tenthredinidae)." volume="60" year="1999">Haris and Roller (1999)</bibRefCitation>
placed this species in
<taxonomicName class="Insecta" family="Tenthredinidae" genus="Trichiocampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trichiocampus" order="Hymenoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="genus">Trichiocampus</taxonomicName>
<taxonomicName id="56AFA8A19080B9BAD0F565BAFA3D16FD" genus="Trichiocampus" lsidName="Trichiocampus" pageId="5" pageNumber="86" rank="genus">Trichiocampus</taxonomicName>
. However, left mandible in lateral view bearing a strongly swollen base and narrowing to a thin blade-like apex
<pageBreakToken pageId="6" pageNumber="87" start="start">,</pageBreakToken>
<pageBreakToken id="4DC25D9A74BE65AEAACE2D640F37CFF0" pageId="6" pageNumber="87" start="start">,</pageBreakToken>
the long vein R+M, the narrow and strongly elevated epicnemium with a deep and strongly curved furrow, and the distinctly carinate frontal walls show that it is member of
<taxonomicName genus="Anhoplocampa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anhoplocampa" pageId="6" pageNumber="87" rank="genus">Anhoplocampa</taxonomicName>
<taxonomicName id="20620C08E282980FAA3BE8F4E5596C37" genus="Anhoplocampa" lsidName="Anhoplocampa" pageId="6" pageNumber="87" rank="genus">Anhoplocampa</taxonomicName>
and does not belong to
<taxonomicName class="Insecta" family="Tenthredinidae" genus="Trichiocampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trichiocampus" order="Hymenoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="genus">Trichiocampus</taxonomicName>
<taxonomicName id="29904ED3E3394B1CFF0D7C5C1969511F" genus="Trichiocampus" lsidName="Trichiocampus" pageId="6" pageNumber="87" rank="genus">Trichiocampus</taxonomicName>
.
</paragraph>
<paragraph pageId="6" pageNumber="87">
<paragraph id="0F00AA6224FD5EC87D1663AABBDE0AC4" pageId="6" pageNumber="87">
The weak cross-band of the forewing, the comparatively short antenna, the short cerci, the slender lancet with simple annuli, the flat temple and postocellar area of
<taxonomicName genus="Anhoplocampa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anhoplocampa yunanensis" pageId="6" pageNumber="87" rank="species" species="yunanensis">Anhoplocampa yunanensis</taxonomicName>
<taxonomicName id="F055283A2210BAA3DB4053166BDD0F0B" genus="Anhoplocampa" lsidName="Anhoplocampa yunanensis" pageId="6" pageNumber="87" rank="species" species="yunanensis">Anhoplocampa yunanensis</taxonomicName>
show that it is morphologically remote from
<taxonomicName genus="Anhoplocampa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anhoplocampa bicoloricornis" pageId="6" pageNumber="87" rank="species" species="bicoloricornis">Anhoplocampa bicoloricornis</taxonomicName>
<taxonomicName id="174C5EB6648E09251E12319D60DF706D" genus="Anhoplocampa" lsidName="Anhoplocampa bicoloricornis" pageId="6" pageNumber="87" rank="species" species="bicoloricornis">Anhoplocampa bicoloricornis</taxonomicName>
and
<taxonomicName genus="Anhoplocampa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anhoplocampa fumosa" pageId="6" pageNumber="87" rank="species" species="fumosa">Anhoplocampa fumosa</taxonomicName>
<taxonomicName id="4DE111D15B54093CF2AA3B3878171D99" genus="Anhoplocampa" lsidName="Anhoplocampa fumosa" pageId="6" pageNumber="87" rank="species" species="fumosa">Anhoplocampa fumosa</taxonomicName>
.
</paragraph>
</subSubSection>

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Genus
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<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
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differs from all other liphistiid genera by the presence of a tibial apophysis on male palp (Figs
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,
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<figureCitation id="4FA10D1AB7B9813FC77016BF8F0F1BCF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genital anatomy of Liphistius pyinoolwin sp. nov. A, H palp prolateral view B, I palp ventral view C, J palp retrolateral view D-G palp distal view A-C XUX- 2018 - 089 D-G XUX- 2018 - 110 B H-J XUX- 2018 - 098; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure3" httpUri="https://binary.pensoft.net/fig/531601" pageId="0" pageNumber="41">3H-J</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genital anatomy of Liphistius pyinoolwin sp. nov. (specimens from AMNH, collected by W. Sedgwick) A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view A-G AMNH _ IZC 00356855; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure4" httpUri="https://binary.pensoft.net/fig/531602" pageId="0" pageNumber="41">4A-C</figureCitation>
<figureCitation id="5EFD01742145C0C8F1D5C2EAC958E8B6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genital anatomy of Liphistius pyinoolwin sp. nov. (specimens from AMNH, collected by W. Sedgwick) A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view A-G AMNH _ IZC 00356855; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure4" httpUri="https://binary.pensoft.net/fig/531602" pageId="0" pageNumber="41">4A-C</figureCitation>
8A-C), and by the presence of a poreplate and a median receptacular cluster in female genitalia (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genital anatomy of Liphistius pyinoolwin sp. nov. A-C plate ventral view D-F vulva dorsal view G-I vulva ventral view A, D, G XUX- 2018 - 094 B, E, H XUX- 2018 - 096 C, F, I XUX- 2018 - 104; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure5" httpUri="https://binary.pensoft.net/fig/531603" pageId="0" pageNumber="41">5D-I</figureCitation>
<figureCitation id="0B3B17DF2655FAA30D22C7242C46A62C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female genital anatomy of Liphistius pyinoolwin sp. nov. A-C plate ventral view D-F vulva dorsal view G-I vulva ventral view A, D, G XUX- 2018 - 094 B, E, H XUX- 2018 - 096 C, F, I XUX- 2018 - 104; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure5" httpUri="https://binary.pensoft.net/fig/531603" pageId="0" pageNumber="41">5D-I</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genital anatomy of Liphistius pyinoolwin sp. nov. A-C plate ventral view D-F vulva dorsal view G-I vulva ventral view A, D, G XUX- 2018 - 105 B, E, H XUX- 2018 - 109 C, F, I XUX- 2018 - 110; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure6" httpUri="https://binary.pensoft.net/fig/531604" pageId="0" pageNumber="41">6D-I</figureCitation>
<figureCitation id="26EFA11E5462E44C4BC4E75B67164858" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female genital anatomy of Liphistius pyinoolwin sp. nov. A-C plate ventral view D-F vulva dorsal view G-I vulva ventral view A, D, G XUX- 2018 - 105 B, E, H XUX- 2018 - 109 C, F, I XUX- 2018 - 110; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure6" httpUri="https://binary.pensoft.net/fig/531604" pageId="0" pageNumber="41">6D-I</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female genital anatomy of Liphistius pyinoolwin sp. nov. (specimens from AMNH, collected by W. Sedgwick) A plate ventral view B, D-F vulva dorsal view C, G-I vulva ventral view A, D, G AMNH _ IZC 00356859 B, C AMNH _ IZC 00356856 E, H AMNH _ IZC 00356857 F, I AMNH _ IZC 00356858; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure7" httpUri="https://binary.pensoft.net/fig/531605" pageId="0" pageNumber="41">7B-I</figureCitation>
<figureCitation id="CC0DCB455F553DF5CEC743D3F506692F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female genital anatomy of Liphistius pyinoolwin sp. nov. (specimens from AMNH, collected by W. Sedgwick) A plate ventral view B, D-F vulva dorsal view C, G-I vulva ventral view A, D, G AMNH _ IZC 00356859 B, C AMNH _ IZC 00356856 E, H AMNH _ IZC 00356857 F, I AMNH _ IZC 00356858; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure7" httpUri="https://binary.pensoft.net/fig/531605" pageId="0" pageNumber="41">7B-I</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genital anatomy of Liphistius birmanicus (Thorell, 1897) A-C plate ventral view D-F vulva dorsal view G-I vulva ventral view A, D, G ARAMYN- 497 B, E, H ARAMYN- 501 C, F, I ARAMYN- 506; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure9" httpUri="https://binary.pensoft.net/fig/531607" pageId="0" pageNumber="41">9D-I</figureCitation>
<figureCitation id="8F0ED524088B3AEC47C6DB8A0B70407F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genital anatomy of Liphistius birmanicus (Thorell, 1897) A-C plate ventral view D-F vulva dorsal view G-I vulva ventral view A, D, G ARAMYN- 497 B, E, H ARAMYN- 501 C, F, I ARAMYN- 506; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure9" httpUri="https://binary.pensoft.net/fig/531607" pageId="0" pageNumber="41">9D-I</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female genital anatomy of Liphistius birmanicus (Thorell, 1897) A, B plate ventral view C, D, G vulva dorsal view E, F, H vulva ventral view A, C, E ARAMYN- 091 B, D, F ARAMYN- 095 G, H lectotype (specimen from MCSNG, collected by L. Fea); Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure10" httpUri="https://binary.pensoft.net/fig/531608" pageId="0" pageNumber="41">10C-G</figureCitation>
<figureCitation id="0C45230076E308C247C9C596C76126E3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female genital anatomy of Liphistius birmanicus (Thorell, 1897) A, B plate ventral view C, D, G vulva dorsal view E, F, H vulva ventral view A, C, E ARAMYN- 091 B, D, F ARAMYN- 095 G, H lectotype (specimen from MCSNG, collected by L. Fea); Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1031.59102.figure10" httpUri="https://binary.pensoft.net/fig/531608" pageId="0" pageNumber="41">10C-G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="distribution">
<paragraph pageId="0" pageNumber="41">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="41">Indonesia (Sumatra), Laos, Malaysia, Myanmar, and Thailand.</paragraph>
<subSubSection id="387A843472C7D257D453AE8D0ABF8752" pageId="0" pageNumber="41" type="distribution">
<paragraph id="2028AE5BA55A73080223F054A9472E2A" pageId="0" pageNumber="41">Distribution.</paragraph>
<paragraph id="0B101006EE4F8D8003654BEE7DFD17E5" pageId="0" pageNumber="41">Indonesia (Sumatra), Laos, Malaysia, Myanmar, and Thailand.</paragraph>
</subSubSection>
</treatment>
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@ -1,640 +1,640 @@
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchoeidestenhelia prote gen. et sp. nov., female A rostrum and antennule B antenna. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchoeidestenhelia prote gen. et sp. nov., female A maxilla B maxilliped (arrow indicates minute intercoxal sclerite). Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure6" httpUri="https://binary.pensoft.net/fig/472436" pageId="0" pageNumber="41">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lonchoeidestenhelia prote gen. et sp. nov., female A P 1, anterior B P 2, anterior. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure7" httpUri="https://binary.pensoft.net/fig/472437" pageId="0" pageNumber="41">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lonchoeidestenhelia prote gen. et sp. nov., female A P 3, anterior B P 4, anterior C P 5, anterior. Scale bars: 50 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">, 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lonchoeidestenhelia prote gen. et sp. nov., intersex individual A P 5 - bearing somite and double genital-somite, lateral B P 5 - bearing somite and double genital-somite, ventral. Scale bar: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure11" httpUri="https://binary.pensoft.net/fig/472441" pageId="0" pageNumber="41">, 11</figureCitation>
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<taxonomicNameLabel id="4828E4430EC836971AF60A11917D60B0" pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
<figureCitation id="2FA151A9DBE2D71148D4AE00A2EE0C81" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">Figs 2</figureCitation>
<figureCitation id="E4342A036682F5799BE83CEDEADAFB9F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">, 3</figureCitation>
<figureCitation id="344B44BD9A68EBBAB4A96DF19FC010B0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchoeidestenhelia prote gen. et sp. nov., female A rostrum and antennule B antenna. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41">, 4</figureCitation>
<figureCitation id="2F75D4D662C9DE2154EE742F3F139315" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">, 5</figureCitation>
<figureCitation id="4BABDECC3DDD133AA256F28F4CC239A6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchoeidestenhelia prote gen. et sp. nov., female A maxilla B maxilliped (arrow indicates minute intercoxal sclerite). Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure6" httpUri="https://binary.pensoft.net/fig/472436" pageId="0" pageNumber="41">, 6</figureCitation>
<figureCitation id="1E6BD2D474CA1D1C2A00F36B94A209D2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lonchoeidestenhelia prote gen. et sp. nov., female A P 1, anterior B P 2, anterior. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure7" httpUri="https://binary.pensoft.net/fig/472437" pageId="0" pageNumber="41">, 7</figureCitation>
<figureCitation id="37F1C3AEDED161BB7F954F8F194390FD" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lonchoeidestenhelia prote gen. et sp. nov., female A P 3, anterior B P 4, anterior C P 5, anterior. Scale bars: 50 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41">, 8</figureCitation>
<figureCitation id="6038AAB149E0E22BD6D52C5802FB3413" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">, 9</figureCitation>
<figureCitation id="0E94E9C9FDF87AF2E423BCC9B1350F8D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">, 10</figureCitation>
<figureCitation id="7601C494541439C9A5CC3DDA8747287C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lonchoeidestenhelia prote gen. et sp. nov., intersex individual A P 5 - bearing somite and double genital-somite, lateral B P 5 - bearing somite and double genital-somite, ventral. Scale bar: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure11" httpUri="https://binary.pensoft.net/fig/472441" pageId="0" pageNumber="41">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="materials_examined">
<paragraph pageId="0" pageNumber="41">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="41">
<subSubSection id="E981D9E10476CF7675073A672D5F7D45" pageId="0" pageNumber="41" type="materials_examined">
<paragraph id="AD27BF21A3859696E2DB7F84C553DB24" pageId="0" pageNumber="41">Specimens examined.</paragraph>
<paragraph id="819DBFAABFAB22827276CE0B024F3BAB" pageId="0" pageNumber="41">
One female holotype (ICML-EMUCOP-180119-01), one male allotype (ICML-EMUCOP-180119-02), and 14 paratypes (10 females and four males) (ICML-EMUCOP-180119-03) preserved in alcohol, and two female (ICML-EMUCOP-180119-05, ICML-EMUCOP-180119-06) and one male (ICML-EMUCOP-180119-07) paratypes dissected and mounted onto 11, six and seven slides, respectively, all from the type locality; six paratypes (two females and four males) (ICML-EMUCOP-180119-04) from stn 1, preserved in alcohol; one female paratype partially dissected (ICML-EMUCOP-180119-08) (P1-P4 dissected and mounted onto one slide, the rest left intact and preserved in alcohol), and nine paratypes (eight females and one male) (ICML-EMUCOP-180119-09) from stn 10, preserved in alcohol; 18 Jan. 2019. S.
<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
<normalizedToken id="F99D1532190C7AD551E785005B137C44" originalValue="Gómez">Gomez</normalizedToken>
leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="materials_examined">
<paragraph pageId="0" pageNumber="41">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="41">
<subSubSection id="595569CF35CFCBAA379F42B029BFD3C4" pageId="0" pageNumber="41" type="materials_examined">
<paragraph id="AF9E04229D6C3911BD47A200C2F5EBFC" pageId="0" pageNumber="41">Additional material examined.</paragraph>
<paragraph id="EB613D80205878C15D7332042250100B" pageId="0" pageNumber="41">
One intersexual individual partially dissected (ICML-EMUCOP-180119-10) (P1-P4 dissected and mounted onto one slide, the rest left intact and preserved in alcohol) from stn 4; 18 Jan. 2019. S.
<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
<normalizedToken id="B2D8352D9F8694E44D0E65C812B2D998" originalValue="Gómez">Gomez</normalizedToken>
leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="differential diagnosis">
<paragraph pageId="0" pageNumber="41">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="41">
<taxonomicName authorityName="Brady" authorityYear="1880" lsidName="" pageId="0" pageNumber="41" rank="subFamily" subFamily="Stenheliinae">Stenheliinae</taxonomicName>
<subSubSection id="10336AA5ADB3D36560F68F6F0BFC21F3" pageId="0" pageNumber="41" type="differential diagnosis">
<paragraph id="C2006C623AA1295AD2ABF89561836317" pageId="0" pageNumber="41">Differential diagnosis.</paragraph>
<paragraph id="4886E9D2AEC251F1EF45DC962CCE9433" pageId="0" pageNumber="41">
<taxonomicName id="D150E2092D81A7AC7FCF1974624CE30F" authorityName="Brady" authorityYear="1880" lsidName="" pageId="0" pageNumber="41" rank="subFamily" subFamily="Stenheliinae">Stenheliinae</taxonomicName>
. Anal operculum present, with minute spinules along posterior margin. P1 EXP2 without inner armature. P2-P4 EXP1 with inner seta. Armature formula of P2 EXP3 and P3 EXP3, 123 and 223, respectively. Female P2ENP3 with outer apical spinous process, with displacement of medial and inner apical setae to subapical inner margin, the latter setae normal, not swollen basally. Female P5 baseoendopod without modified setae. Male P2ENP2 without hyaline flange; outer apical sigmoid, bipinnate, flagellate process with incomplete suture indicating original articulation with the supporting segment, with longitudinal row of modified lanceolate spinules proximally. Outer spine of the male P4ENP3 normal, not sexually dimorphic. Male P5EXP and baseoendopod not fused; exopod with two outermost elements modified into spines. Innermost seta of the male P6 normal.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Description.</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">
<emphasis italics="true" pageId="0" pageNumber="41">Female</emphasis>
<subSubSection id="A80AB929A8BBA6A63A6A380163557B55" pageId="0" pageNumber="41" type="description">
<paragraph id="7E34CCD70C54448779FE20D5A11135E8" pageId="0" pageNumber="41">Description.</paragraph>
<paragraph id="24A94C75DDAB73A03AD339CDBF68109C" pageId="0" pageNumber="41">
<emphasis id="4A8389A5DD2215184F5797AD3ECFCE41" bold="true" pageId="0" pageNumber="41">
<emphasis id="EE3835F4130978432D7DF3EC02EF5D46" italics="true" pageId="0" pageNumber="41">Female</emphasis>
.
</emphasis>
Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 415
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="B5A0B76B69FD7F5DD44CC5F4427566F4" originalValue="µm">µm</normalizedToken>
to 563
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="3B9160DF8F6907C0D18FF69786DB01A2" originalValue="µm">µm</normalizedToken>
(mean, 491
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="F9CA66295F3A7441665816DD421592E7" originalValue="µm">µm</normalizedToken>
; n, 12; total body length of holotype, 563
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="D3120FF04DEF80DFFD30B8100D8A7178" originalValue="µm">µm</normalizedToken>
); habitus pyriform, widest at posterior end of cephalothorax in dorsal view, tapering posteriad (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
<figureCitation id="2F100AAFB4B967908A7B62D4037C6974" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Prosome</emphasis>
<paragraph id="1648284779A9ADAA37DF5A6713BDCB8C" pageId="0" pageNumber="41">
<emphasis id="C08112C34BBBEFAD5A2676B7F4975FBC" bold="true" pageId="0" pageNumber="41">Prosome</emphasis>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, B</figureCitation>
<figureCitation id="094C9123B924B38F1E6D8CB1D41D2A3C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, B</figureCitation>
): Consisting of cephalothorax with fused first pedigerous somite, and second to fourth free pedigerous somites, the latter without expansions nor spinular ornamentation; posterior hyaline frill of cephalothorax and pedigerous somites plain, of fourth pedigerous somite visibly narrower; integument smooth, weakly sclerotized.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 2.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="D60B85CBE3B56E82DAAA48F0771013FD" doi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41" start="Figure 2" startId="F2">
<paragraph id="7BFDDDB8319C6C2B89D4087DF4F29787" pageId="0" pageNumber="41">
<emphasis id="D8AA0F250299C6183883A5BD22A82CE9" bold="true" pageId="0" pageNumber="41">Figure 2.</emphasis>
<taxonomicName id="FB08005E45A4EB06FA83DBB3E84C1E37" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="806DFA455783F8951DCE1255A542EB33" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="AF6FEB6D3D301A78C43F05970D16922F" bold="true" pageId="0" pageNumber="41">A</emphasis>
habitus, dorsal
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="4E3B8AFD0646E469B4C5A4CA999070AC" bold="true" pageId="0" pageNumber="41">B</emphasis>
anal somite and left caudal ramus, dorsal
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
<emphasis id="113DE5D0F3313D2F6C8DA62DFED99D85" bold="true" pageId="0" pageNumber="41">C</emphasis>
habitus, lateral
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
<emphasis id="564C0B55C7C663126FBAFEB4242DA707" bold="true" pageId="0" pageNumber="41">D</emphasis>
left caudal ramus, lateral. Scale bars: 200
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="8DBACCC9CF7C12879B6C31C11DB21722" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A, C</emphasis>
<emphasis id="4FEFE5428B7BF6D8F324831FBAF1A5BB" bold="true" pageId="0" pageNumber="41">A, C</emphasis>
); 100
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="29981EDEC45713FF3B24678061C7F647" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">B, D</emphasis>
<emphasis id="F5F96032A6A761D54494C9F05D7FF0E0" bold="true" pageId="0" pageNumber="41">B, D</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Urosome</emphasis>
<paragraph id="CD6F42E56867505AADE0A73DA752EED6" pageId="0" pageNumber="41">
<emphasis id="445F5DB3545D81517F45B3212E492E9B" bold="true" pageId="0" pageNumber="41">Urosome</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, B</figureCitation>
<figureCitation id="976F9C13929CA62F7A65EFAEB417D571" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
<figureCitation id="58542A31FF8C90ACCCA18FA5FE22D25F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
): Consisting of fifth pedigerous somite (first urosomite), genital double-somite (genital (second urosomite) and third urosomites fused), two free urosomites, and anal somite; urosomites without expansions laterally nor dorsally; integument weakly sclerotized.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 3.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="9486E31BD59BA00535B86D66770F8CAB" doi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41" start="Figure 3" startId="F3">
<paragraph id="B57C50F8A48C75C324D8135781653720" pageId="0" pageNumber="41">
<emphasis id="6A6C15EACB61853A2321A556FDFFF776" bold="true" pageId="0" pageNumber="41">Figure 3.</emphasis>
<taxonomicName id="E61AD0872266C699F1C9DB76138E12B0" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="7D6D69115441D5CDEE4D7A2DA44F860D" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="00C81221D4835A3B36A64FFD4E352EE6" bold="true" pageId="0" pageNumber="41">A</emphasis>
urosome, dorsal (P5-bearing somite omitted)
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="F704FCC17A353D3E53B7C38CE84FDD82" bold="true" pageId="0" pageNumber="41">B</emphasis>
urosome, lateral (P5-bearing somite omitted)
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
<emphasis id="605FCD2D443761D8147AF33BBD9D5B37" bold="true" pageId="0" pageNumber="41">C</emphasis>
urosome, ventral (P5-bearing somite omitted). Scale bars: 100
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="E3AC8BFA116902673AA4BDC910223151" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A-C</emphasis>
<emphasis id="75634ED5A4F8C1DDDF0834E893B5FBD3" bold="true" pageId="0" pageNumber="41">A-C</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">First urosomite</emphasis>
<paragraph id="B7A9403D5B6D90BA905C3D3252E59FBE" pageId="0" pageNumber="41">
<emphasis id="5291341E53D18D269F92C23938F6E746" bold="true" pageId="0" pageNumber="41">First urosomite</emphasis>
(fifth pedigerous somite): Visibly narrower than preceding somites in dorsal view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
<figureCitation id="8371C5E0FCC2E82875EAB622C3ABD10C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
), without spinular ornamentation, with dorsolateral sensilla as shown (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, C</figureCitation>
<figureCitation id="D333EF3E5F4960994921B23ADE12F2CA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Genital double-somite</emphasis>
<paragraph id="8C6E7519EB5D34763AC0980172E06206" pageId="0" pageNumber="41">
<emphasis id="625251EEB5DE93BE5C6F2177C568D062" bold="true" pageId="0" pageNumber="41">Genital double-somite</emphasis>
: Slightly wider than long, widest at proximal half; with dorsolateral internal rib marking original division between second (genital) and third urosomite (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, C</figureCitation>
<figureCitation id="AE09BC52D823529A7E781670DA055597" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A, B</figureCitation>
<figureCitation id="0163E875BC93B3135B0520DBC5E4DF54" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A, B</figureCitation>
), completely fused dorsally (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
<figureCitation id="0EE9EB9B66E240A2C5BBC62AF59A458C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A</figureCitation>
<figureCitation id="10A938EB31F2E24A04C1E04F58ED8FEF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A</figureCitation>
) and ventrally (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3C</figureCitation>
<figureCitation id="E864072B106A7A115D660D05E9BF0F8E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3C</figureCitation>
); proximal half (genital somite) without spinular ornamentation, with posterior sensilla as depicted; distal half (third urosomite) with short transverse row of small dorsolateral spinules (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
<figureCitation id="2CE78F331884A3012BDDEE4F32FF8DF8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A</figureCitation>
<figureCitation id="39B87B2606A4611A92C8C369540BF941" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A</figureCitation>
), with sensilla as shown.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Fourth urosomite</emphasis>
<paragraph id="1F660ECC6DBDE0C8EC97D58D7BF9A2C0" pageId="0" pageNumber="41">
<emphasis id="041B89A07F94B5C9ADE87055FF955C89" bold="true" pageId="0" pageNumber="41">Fourth urosomite</emphasis>
: With spinular ornamentation (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
<figureCitation id="1D4E0353B25C63A25569A193FC267FD6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
) as in distal half of genital double-somite, with sensilla as shown.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Fifth urosomite</emphasis>
<paragraph id="ECA28BF00C515C065AF0C8E6F0352CF3" pageId="0" pageNumber="41">
<emphasis id="D35BA8D7893909D2CCD2136DD2F2E363" bold="true" pageId="0" pageNumber="41">Fifth urosomite</emphasis>
: With spinular ornamentation (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
<figureCitation id="60FA603DE0F6F17F6F9B5C5CF9F2E741" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
) as in distal half of genital double-somite, without sensilla.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Anal somite</emphasis>
<paragraph id="FADAE4E9F1946A23C1B98F528C58C728" pageId="0" pageNumber="41">
<emphasis id="9BF09C533281F47B39C395AD59AEBC18" bold="true" pageId="0" pageNumber="41">Anal somite</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A-C</figureCitation>
<figureCitation id="74FFA972E1CCBF80735F0C52AA018B21" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A-C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
<figureCitation id="26DA5EBD0FA97C8AC3FE7B9003FB06D6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
): Twice as long as broad, maximum breadth measured proximally; maximum length measured at the middle from anterior margin of somite to distal margin of anal operculum, with row of dorsolateral spinules close to joint with caudal rami, with one lateral pore on each side (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2C</figureCitation>
<figureCitation id="2997FA50478E3F01DDE78F11F5194CC8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A, B</figureCitation>
<figureCitation id="B4C5D72059A7C61EBD50D8199F2D618A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A, B</figureCitation>
); anal operculum rounded, with minute spinules along its posterior margin, with one sensillum on each side; ventrally cleft medially, with one pore on each side, with spinular row close to joint with caudal rami (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3C</figureCitation>
<figureCitation id="2600CEB4B279394B1874F8742DD7DC03" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Caudal rami</emphasis>
<paragraph id="7480DEDBC1F21205F0F0B0CF5D9AE70E" pageId="0" pageNumber="41">
<emphasis id="B97D47422E38A868B1D43167EDB9FEAE" bold="true" pageId="0" pageNumber="41">Caudal rami</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A-D</figureCitation>
<figureCitation id="367B3AEFF1F599C87B24384B2D10744B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A-D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
<figureCitation id="8BE0F1FB11070AEED8C9FC95D178F3D0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A-C</figureCitation>
): Typically divergent (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, B</figureCitation>
<figureCitation id="0372602241AE99ED2829A259F084F998" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A, B</figureCitation>
), but sometimes parallel (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A, C</figureCitation>
<figureCitation id="EB8214E5C3B9AD9A5CDCF72F3FA46772" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3A, C</figureCitation>
), cylindrical, twice as long as broad, and slightly longer than anal somite; each ramus with one lateral (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2C, D</figureCitation>
<figureCitation id="00FAD271A24BB321AA3D839BB8E92D47" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2C, D</figureCitation>
) and one ventral pore (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3B, C</figureCitation>
<figureCitation id="CF91387BC829C447D4CF91B161064815" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3B, C</figureCitation>
); with spinules at base of setae I and II (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2C, D</figureCitation>
<figureCitation id="C0D278BEA1B8234293AFA5B93249304E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2C, D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3B</figureCitation>
<figureCitation id="4BB76ADF831EF34B779509B31FA52ED3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3B</figureCitation>
), and ventrally close to insertion site of seta III (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3C</figureCitation>
<figureCitation id="58157DC5AC1E974B420B7907BA07F617" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3C</figureCitation>
); with seven elements as follows: seta I spine-like, ventral to seta II, the former visibly shorter, both arising subdistally on lateral margin; seta III ventral, subdistal, slightly longer than seta II; seta IV and V situated distally, with fracture plane, rat-tail-like; seta VI issuing at inner distal corner; dorsal seta VII triarticulate at base, situated subdistally close to inner margin.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Rostrum</emphasis>
<paragraph id="720CF4CF8EAC2A3B74EDADB2418B4B20" pageId="0" pageNumber="41">
<emphasis id="DB389754F059972C27578CBDD35A8076" bold="true" pageId="0" pageNumber="41">Rostrum</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
<figureCitation id="E063796599FFF109DA87F3E7F4360332" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lonchoeidestenhelia prote gen. et sp. nov., female A habitus, dorsal B anal somite and left caudal ramus, dorsal C habitus, lateral D left caudal ramus, lateral. Scale bars: 200 µm (A, C); 100 µm (B, D)." figureDoi="10.3897/zookeys.987.52906.figure2" httpUri="https://binary.pensoft.net/fig/472432" pageId="0" pageNumber="41">2A</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchoeidestenhelia prote gen. et sp. nov., female A rostrum and antennule B antenna. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41">4A</figureCitation>
<figureCitation id="1BE2CCC88C62CC7D1E66B6A9EEFF40C0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchoeidestenhelia prote gen. et sp. nov., female A rostrum and antennule B antenna. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41">4A</figureCitation>
): Trapezoidal, not fused to cephalothorax, reaching middle of second antennular segment, bifid, with medial pore and two subdistal sensilla.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 4.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="C58F15CDE6ABA51BB73BD368BB142F06" doi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41" start="Figure 4" startId="F4">
<paragraph id="75EBB98E1BA0889752BC2D1E5DC2BA08" pageId="0" pageNumber="41">
<emphasis id="EE1C74535B2C7F9A49E6DC9308C66C19" bold="true" pageId="0" pageNumber="41">Figure 4.</emphasis>
<taxonomicName id="5E061CDB977F9CF2473BFDCF3FE113A9" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="36146A65597512A8A887482670376878" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="BC5898EA5E08951B85075623CF721B67" bold="true" pageId="0" pageNumber="41">A</emphasis>
rostrum and antennule
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="8109ACF8EA5B9A901E0D8AA0D864B8AD" bold="true" pageId="0" pageNumber="41">B</emphasis>
antenna. Scale bars: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="4E5B36C41639C97D9BCF4033DF4B743E" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A, B</emphasis>
<emphasis id="EF64DA608C7C4404E7567D6111CE922C" bold="true" pageId="0" pageNumber="41">A, B</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Antennule</emphasis>
<paragraph id="6A354A6E5CB85DB2427533E3472BA576" pageId="0" pageNumber="41">
<emphasis id="47A93F5B710D951EC6D276B4C456CD68" bold="true" pageId="0" pageNumber="41">Antennule</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchoeidestenhelia prote gen. et sp. nov., female A rostrum and antennule B antenna. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41">4A</figureCitation>
<figureCitation id="321395D56706C7426DB4E82171D63488" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchoeidestenhelia prote gen. et sp. nov., female A rostrum and antennule B antenna. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41">4A</figureCitation>
): Seven-segmented; all segments smooth, except for first segment with spinular row. All setae smooth except for one and two pinnate setae on first and second segments, respectively; second and third segments each with one seta with fracture plane; sixth and seventh segments each with two articulated setae. Armature formula: 1(1); 2(11); 3(8); 4(5+(1+ae)), 5(3); 6(8); 7(4+acro). Acrothek consisting of two setae and one slender aesthetasc fused at their bases.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Antenna</emphasis>
<paragraph id="E9F3841205137E7B21671735747C0B69" pageId="0" pageNumber="41">
<emphasis id="A43581249AC524567DF71512C44AE756" bold="true" pageId="0" pageNumber="41">Antenna</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchoeidestenhelia prote gen. et sp. nov., female A rostrum and antennule B antenna. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41">4B</figureCitation>
<figureCitation id="00B107BE8A9A0D9E27291ECFE5F8B3CD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lonchoeidestenhelia prote gen. et sp. nov., female A rostrum and antennule B antenna. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure4" httpUri="https://binary.pensoft.net/fig/472434" pageId="0" pageNumber="41">4B</figureCitation>
): Allobasis as long as free endopodal segment, inner margin with long spinules on proximal third, with one abexopodal seta arising midway inner margin. Exopod three-segmented, issuing proximally; first and third segments longest, each 3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="2648103BF80AB202E32886F362FD36E2" originalValue="×">x</normalizedToken>
as long as wide, second segment shortest, ca. 1.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B52EA0D023CD4EE1E382BE45E3B13C93" originalValue="×">x</normalizedToken>
as long as broad; first and second segments with one subdistal pinnate seta each, first segment with spinular row as shown, second segment unornamented; third segment with one lateral proximal pinnate element, and three distal setae, of which two fused basally, with spinular row as depicted. Free endopodal segment elongate, inner margin with spinular row proximally, subdistally with curved row of strong spinules, with medial and subdistal outer fringes; armature consisting of two lateral spines and two accompanying setae, one non-geniculate inner distal element, three distal geniculate spines (of which innermost shortest) and one slender seta, and one outer distal geniculate seta fused basally to slender element.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Mandible</emphasis>
<paragraph id="3E4A77A9533B314A14C7424A519D2F7E" pageId="0" pageNumber="41">
<emphasis id="E346B17D81630D389B5E809D184783E1" bold="true" pageId="0" pageNumber="41">Mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">5A</figureCitation>
<figureCitation id="89B828762ECDF5C8C4FAD538098FA497" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">5A</figureCitation>
): With relatively short coxa. Gnathobasis wide; with two strong bicuspidate teeth, several smaller bicuspidate teeth, some spinules, and one lanceolate element accompanied by seta. Basis elongate, tapering distally; with transverse spinular rows as shown; with three subdistal setae. Exopod arising from short pedestal, one-segmented, elongate, ca. 4
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="A5534099FC1D87FEA88F4E4E8E7F3C40" originalValue="×">x</normalizedToken>
as long as wide, tapering distally; with three lateral and three apical slender setae, none of which fused basally. Endopod recurved, twisted over exopod; with three lateral setae, and five distal elements (one short seta and long pinnate element fused basally and to endopod, indicated with an asterisk in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">5A</figureCitation>
<figureCitation id="E11D0E7515D7CFC5C295B4C0F167CA1F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">5A</figureCitation>
, one slender pinnate and one strong bare element, and one long element fused to endopod and with hyaline flange in middle part, indicated with an asterisk in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">5A</figureCitation>
<figureCitation id="3F7BEE7DCCCAFE3BDC310B2985BF89A3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">5A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 5.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="7365CF7FD4AA13A8D4F6B244C38DD325" doi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41" start="Figure 5" startId="F5">
<paragraph id="4DA0823D6EE720134AD67CC842BCC1D3" pageId="0" pageNumber="41">
<emphasis id="A9FBA2897FEA98A4DAE4C3961D335FE2" bold="true" pageId="0" pageNumber="41">Figure 5.</emphasis>
<taxonomicName id="09B1171E4368E2FC358F81C6A99221B7" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="43AE4ABE8BC1D2DEBE656A995FB31EDC" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="49C4FAD70197D410539F6EF11C26B3E3" bold="true" pageId="0" pageNumber="41">A</emphasis>
mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus)
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="1C6FDBF5976BF9186AE48D7789B16184" bold="true" pageId="0" pageNumber="41">B</emphasis>
maxillule. Scale bars: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="2171368B639C1BCBFFC806830A4C3A3A" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A, B</emphasis>
<emphasis id="DC8E627F91F44F36DB9F420C26D8075C" bold="true" pageId="0" pageNumber="41">A, B</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Maxillule</emphasis>
<paragraph id="973BA532B3891F06F29B4BC55FF6FD33" pageId="0" pageNumber="41">
<emphasis id="131325B95555C0A8E3187EB837C78C4B" bold="true" pageId="0" pageNumber="41">Maxillule</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">5B</figureCitation>
<figureCitation id="59EE226BBD9B30E342445B7F0B3E34ED" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lonchoeidestenhelia prote gen. et sp. nov., female A mandible (asterisks indicate two elements fused basally and to ramus, and single element fused to ramus) B maxillule. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure5" httpUri="https://binary.pensoft.net/fig/472435" pageId="0" pageNumber="41">5B</figureCitation>
): Arthrite of praecoxa with two surface setae and seven bare distal elements (one of which a slender seta arising next to ventralmost spine), one spinulose dorsal spine, and one lateral spinulose recurved seta. Coxal endite with three setae. Basis with two endites; proximal endite with four, distal endite with three slender setae. Exopod and endopod fused basally, separated from basis, one-segmented; endopod larger than exopod, with four setae; exopod small, with two setae.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Maxilla</emphasis>
<paragraph id="4E56BD4043BC5549944DBC807FF421E8" pageId="0" pageNumber="41">
<emphasis id="B1C16010BD8452CC47C23021E978C17C" bold="true" pageId="0" pageNumber="41">Maxilla</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchoeidestenhelia prote gen. et sp. nov., female A maxilla B maxilliped (arrow indicates minute intercoxal sclerite). Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure6" httpUri="https://binary.pensoft.net/fig/472436" pageId="0" pageNumber="41">6A</figureCitation>
<figureCitation id="C9F9F674BCBEA45699428B54F10F297B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchoeidestenhelia prote gen. et sp. nov., female A maxilla B maxilliped (arrow indicates minute intercoxal sclerite). Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure6" httpUri="https://binary.pensoft.net/fig/472436" pageId="0" pageNumber="41">6A</figureCitation>
): With large syncoxa with outer spinules as shown; with three endites; proximal endite smallest, with one proximal and two distal setae; middle and distal endites elongate, the latter slightly longer, with three elements each as figured. Basis drawn out into strong claw, additionally with strong spine and two slender setae, one of which arising from elongate setophore. Endopod one-segmented, 2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="FE269EB5B76D3306D4615BB756A34333" originalValue="×">x</normalizedToken>
as long as wide, with six slender setae.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure6" httpUri="https://binary.pensoft.net/fig/472436" pageId="0" pageNumber="41" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 6.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="9619AC6E8E64FE59A66C68AF50FD758A" doi="10.3897/zookeys.987.52906.figure6" httpUri="https://binary.pensoft.net/fig/472436" pageId="0" pageNumber="41" start="Figure 6" startId="F6">
<paragraph id="85418DD4572FF1565EB07C3572A322A9" pageId="0" pageNumber="41">
<emphasis id="80B33880F00474146C3FB14E7A83592D" bold="true" pageId="0" pageNumber="41">Figure 6.</emphasis>
<taxonomicName id="654DCFFD6558E005F732CC9A9E9B2AB1" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="FD7D18C6B27FA386630384BE770EB2B8" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="71E3E6AB89516042EDF08E3545338621" bold="true" pageId="0" pageNumber="41">A</emphasis>
maxilla
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="4EA5370257AB4045E457CF48533AB581" bold="true" pageId="0" pageNumber="41">B</emphasis>
maxilliped (arrow indicates minute intercoxal sclerite). Scale bars: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="735427DED8AFEC44B0503C4757B9E62D" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A, B</emphasis>
<emphasis id="4204EBF0D0D28B1AD53B1C10C5D5A2D0" bold="true" pageId="0" pageNumber="41">A, B</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Maxilliped</emphasis>
<paragraph id="F31BEC58E220F9DC6090BF073BCBD958" pageId="0" pageNumber="41">
<emphasis id="464DC6F6DC2B78174D8373EE2C1672B6" bold="true" pageId="0" pageNumber="41">Maxilliped</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchoeidestenhelia prote gen. et sp. nov., female A maxilla B maxilliped (arrow indicates minute intercoxal sclerite). Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure6" httpUri="https://binary.pensoft.net/fig/472436" pageId="0" pageNumber="41">6B</figureCitation>
<figureCitation id="A2678C1C97888B49F2AD6663D1F84C5D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lonchoeidestenhelia prote gen. et sp. nov., female A maxilla B maxilliped (arrow indicates minute intercoxal sclerite). Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure6" httpUri="https://binary.pensoft.net/fig/472436" pageId="0" pageNumber="41">6B</figureCitation>
): Subchelate. Syncoxa rectangular, ca. 1.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9CFEB10F1718172A399A3A1A9A670163" originalValue="×">x</normalizedToken>
as long as wide, outer margin irregular and with medial protrusion; with spinules as shown; with one bare and two spinulose strong elements, of which the bare seta and one spinulose element subdistal and both at the same level, the other arising distally from long pedestal. Basis shorter than syncoxa, oval; with inner and outer spinules as depicted, and two slender distal setae, one of which visibly shorter. Endopod one-segmented, slender, with one claw-like element and one seta.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P1</emphasis>
<paragraph id="69131BC8E7CDED99B171B7582E481132" pageId="0" pageNumber="41">
<emphasis id="5BC114294C98230C755924991E3BB2AC" bold="true" pageId="0" pageNumber="41">P1</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lonchoeidestenhelia prote gen. et sp. nov., female A P 1, anterior B P 2, anterior. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure7" httpUri="https://binary.pensoft.net/fig/472437" pageId="0" pageNumber="41">7A</figureCitation>
<figureCitation id="EF4F3CDC5B679AF0C558E1C787421B54" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lonchoeidestenhelia prote gen. et sp. nov., female A P 1, anterior B P 2, anterior. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure7" httpUri="https://binary.pensoft.net/fig/472437" pageId="0" pageNumber="41">7A</figureCitation>
): Intercoxal sclerite narrow and elongate, without surface ornamentation. Praecoxa large, triangular, unornamented. Coxa quadrate, with spinular rows as shown. Basis with inner robust and strongly spinulose spine, and outer slender pinnate element; with strong spinules at the bases of inner and outer elements and between rami, and with long slender inner spinules. Exopod three-segmented, reaching tip of first endopodal segment, situated at a lower level than ENP; first segment longest, third segment shortest; all segments without outer nor inner distal processes; with spinular ornamentation as shown; first and second segments without inner seta, third segment with four elements. Endopod three-segmented, situated distally on medial circular outgrowth of basis and at a higher level than EXP; ENP1:EXP length ratio 0.9, ENP1 ca. 1.4
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="947A8FDD2A2FB172705E4E5DAA253468" originalValue="×">x</normalizedToken>
as long as ENP2 and ENP3 combined; ENP1 and ENP3 without, ENP2 with outer acute distal process; segments with spinular ornamentation as figured; ENP1 with pinnate inner seta arising subdistally; ENP2 with one slender inner seta; ENP3 with three elements (one slender inner seta, one apical pinnate element, and one apical outer spine.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure7" httpUri="https://binary.pensoft.net/fig/472437" pageId="0" pageNumber="41" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 7.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="811DDE3F10663E749A9339ECA382C828" doi="10.3897/zookeys.987.52906.figure7" httpUri="https://binary.pensoft.net/fig/472437" pageId="0" pageNumber="41" start="Figure 7" startId="F7">
<paragraph id="C210C3771085C148FA356E8C3EBC9A94" pageId="0" pageNumber="41">
<emphasis id="8AE77203E6208564B2582EEAD6AE639F" bold="true" pageId="0" pageNumber="41">Figure 7.</emphasis>
<taxonomicName id="F85EFD0235DCF39D3AE6BAD551E337BD" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="8689288776EBBAFE3867DAF8200302BC" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="EF48B8AD6558D59ABD3CF2C63D27C096" bold="true" pageId="0" pageNumber="41">A</emphasis>
P1, anterior
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="D723563E04CE87EB7E20F1B0D74FBAFD" bold="true" pageId="0" pageNumber="41">B</emphasis>
P2, anterior. Scale bars: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="7426D08EE9C11A29C7574739CA4F9003" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A, B</emphasis>
<emphasis id="4642B17059DD1C9702DB5E1F5BAC32A2" bold="true" pageId="0" pageNumber="41">A, B</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P2</emphasis>
<paragraph id="6F809667150082EDFDB10856833C8687" pageId="0" pageNumber="41">
<emphasis id="50168D1C0831D521D734C6365FD1DF6B" bold="true" pageId="0" pageNumber="41">P2</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lonchoeidestenhelia prote gen. et sp. nov., female A P 1, anterior B P 2, anterior. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure7" httpUri="https://binary.pensoft.net/fig/472437" pageId="0" pageNumber="41">7B</figureCitation>
<figureCitation id="8A200D178413B43B7D2CCAE37A6A9748" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lonchoeidestenhelia prote gen. et sp. nov., female A P 1, anterior B P 2, anterior. Scale bars: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure7" httpUri="https://binary.pensoft.net/fig/472437" pageId="0" pageNumber="41">7B</figureCitation>
): Intercoxal sclerite not transversely elongate, trapezoidal, with strong pointed process on distal outer corners, without surface ornamentation. Praecoxa small, triangular, unornamented. Coxa massive, quadrate, with outer spinules proximally and subdistally, with subdistal spinules and one pore close to inner margin. Basis with outer setiform element and strong acute process between rami and on inner distal corner, the latter with slender spinules proximally. Exopod three-segmented, reaching slightly beyond ENP3; EXP1 and EXP2 with inner distal frill, with outer acute distal process, with spinular ornamentation as shown, and with one inner seta; EXP3 with processes as shown, with small outer spinules at base of proximal outer spine, with subdistal pore, with one inner and two apical setae, and three outer spines. Endopod three-segmented; segments with spinules as shown; ENP1 shortest, with subdistal inner pore, with inner and outer acute processes, the former slightly larger, with one slender short inner seta; ENP2 and ENP3 subequal in length, the former with outer acute and small inner process, with two inner setae subequal in length; ENP3 with distal processes as shown, with subdistal inner pore, with one inner seta, two inner apical elements, and one outer distal spine.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P3</emphasis>
<paragraph id="DA1D04ACC4099B558649367231525AFE" pageId="0" pageNumber="41">
<emphasis id="3471A3BBCCE1557C2F777CB80A7145B4" bold="true" pageId="0" pageNumber="41">P3</emphasis>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lonchoeidestenhelia prote gen. et sp. nov., female A P 3, anterior B P 4, anterior C P 5, anterior. Scale bars: 50 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41">8A</figureCitation>
<figureCitation id="615C8A4F2D030BDDD4B436BC2D3FE3B2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lonchoeidestenhelia prote gen. et sp. nov., female A P 3, anterior B P 4, anterior C P 5, anterior. Scale bars: 50 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41">8A</figureCitation>
): Intercoxal sclerite not transversely elongate; trapezoidal; wider than in P2; with strong pointed process on distal outer corners; without surface ornamentation. Praecoxa triangular, small. Coxa as in P2 but without inner spinules. Basis as in P2, but with somewhat more slender outer seta. Exopod three-segmented, slightly longer than ENP; segments with spinules as shown; EXP1 and EXP2 with outer acute distal process, without pores, with inner distal frill, and with one inner seta each; EXP3 with outer subdistal pore, with two inner setae, two apical elements, and three outer spines. Endopod three-segmented; spinular ornamentation of segments as depicted; ENP1 shortest, ENP3 longest; ENP1 with small outer and inner distal processes, with inner seta; ENP2 with well-developed outer and small inner distal process, with inner pinnate seta; ENP3 with distal processes as shown, with subdistal outer pore, with two inner and two apical setae, and one outer apical spine.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 8.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="69A436AFF4C501227BA5F8DEACC2E3A2" doi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41" start="Figure 8" startId="F8">
<paragraph id="9EE473B04041B4240CAE2E785F3740A3" pageId="0" pageNumber="41">
<emphasis id="0E8E7C4D91B95A33C7413E4D5FF3E175" bold="true" pageId="0" pageNumber="41">Figure 8.</emphasis>
<taxonomicName id="C1A13537453636E68D29B4B208591A65" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="10FADE01AB93C7E5D265E5F00594FBC7" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., female
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="FADCB4BBE1067FFBE36D723FC7A3D8F8" bold="true" pageId="0" pageNumber="41">A</emphasis>
P3, anterior
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="EF086BB614848694207106FC0A1AAE7B" bold="true" pageId="0" pageNumber="41">B</emphasis>
P4, anterior
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
<emphasis id="7C05548088C3FB689AE2A4FEDB3B7125" bold="true" pageId="0" pageNumber="41">C</emphasis>
P5, anterior. Scale bars: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="00DF1154217659B4EDDB03EA0342EA84" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A-C</emphasis>
<emphasis id="172E169C168012A9EB7B8DB79301CCED" bold="true" pageId="0" pageNumber="41">A-C</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P4</emphasis>
<paragraph id="369D206A080B1974DEA9F49B41B7317C" pageId="0" pageNumber="41">
<emphasis id="9B2106E918DAE82D2428A1DC0A7CADA8" bold="true" pageId="0" pageNumber="41">P4</emphasis>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lonchoeidestenhelia prote gen. et sp. nov., female A P 3, anterior B P 4, anterior C P 5, anterior. Scale bars: 50 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41">8B</figureCitation>
<figureCitation id="DCB6580D84AC5042D50EBE44284EAB37" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lonchoeidestenhelia prote gen. et sp. nov., female A P 3, anterior B P 4, anterior C P 5, anterior. Scale bars: 50 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41">8B</figureCitation>
): Intercoxal sclerite not transversely elongate; trapezoidal; smaller than in P3; with strong pointed process on distal outer corners; without surface ornamentation. Praecoxa, coxa and basis as in P3 except for comparatively smaller inner distal process of basis. Exopod three-segmented, longer than ENP; spinular ornamentation of segments as shown; EXP1 and EXP2 with outer distal process less developed than in P3, without pores, with inner distal frill, and with inner seta; EXP3 with subdistal outer pore, with three inner setae of which medial visibly stronger, two apical elements, and three outer spines. Endopod three-segmented, reaching tip of EXP2; spinular ornamentation of segments as shown; ENP1 shortest, ENP3 longest; ENP1 with small outer distal process, without pore, with inner pinnate seta; ENP2 with well-developed outer distal process, armature as in previous segment; ENP3 with distal processes as shown, with subdistal inner pore, with two inner setae, two apical elements, and one outer apical spine.
</paragraph>
<paragraph pageId="0" pageNumber="41">Setal formula of swimming legs as follows:</paragraph>
<paragraph pageId="0" pageNumber="41">
<table pageId="0" pageNumber="41">
<tr pageId="0" pageNumber="41">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="41" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">P1</th>
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">P2</th>
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">P3</th>
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">P4</th>
<paragraph id="9AB993C2FDE514B4E90F6B5505D16E7B" pageId="0" pageNumber="41">Setal formula of swimming legs as follows:</paragraph>
<paragraph id="FA18AFF0EE05C71650E83F31CF5F6B7E" pageId="0" pageNumber="41">
<table id="4D28040B166BB4C6713877E78E3B38C9" pageId="0" pageNumber="41">
<tr id="B537E29CC20237C14DE09B9EAB8542CF" pageId="0" pageNumber="41">
<th id="910837B5853600FA14255D17A43AF11B" colspan="1" isEmpty="true" pageId="0" pageNumber="41" rowspan="1">-</th>
<th id="3066F6AA185EABA7E04DF5B012DEF315" colspan="1" pageId="0" pageNumber="41" rowspan="1">P1</th>
<th id="E8DDE40BA2AF5D36D21A9B76D38D44DD" colspan="1" pageId="0" pageNumber="41" rowspan="1">P2</th>
<th id="3DD67981C08DD1CE30B942F06EDCF6F7" colspan="1" pageId="0" pageNumber="41" rowspan="1">P3</th>
<th id="29A14B38D3E715710BBED30043BAAD63" colspan="1" pageId="0" pageNumber="41" rowspan="1">P4</th>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">EXP</emphasis>
<tr id="1A0962795B32FD709350E6BBF5BC5D5F" pageId="0" pageNumber="41">
<td id="FF20CC23BA675A6B5A08373597D46212" colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis id="0C7AE23524ABFD24C83E537242CA069C" bold="true" pageId="0" pageNumber="41">EXP</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">0,0,022</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,123</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,223</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,323</td>
<td id="F627E7A5C094D2E385B320F0A5F7325C" colspan="1" pageId="0" pageNumber="41" rowspan="1">0,0,022</td>
<td id="84026623B42E903883DDBB70A809FA5C" colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,123</td>
<td id="4F375D8D81279B44A04DAF065CEB7930" colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,223</td>
<td id="5F16BD3EFCDB28F28BE533896D0574CC" colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,323</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">ENP</emphasis>
<tr id="238BCF643B15CAB3C0DC04E889700807" pageId="0" pageNumber="41">
<td id="FCDA371AF5C3FCB1D6C2B00CAFDFBBDE" colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis id="1973E1F756F49E877113B040D0453AA3" bold="true" pageId="0" pageNumber="41">ENP</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,111</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1,2,121</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,221</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,221</td>
<td id="F402D6318DDFA2F659F18547DD8C0DB2" colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,111</td>
<td id="250416A942C9AD211823FE9E71DDA45E" colspan="1" pageId="0" pageNumber="41" rowspan="1">1,2,121</td>
<td id="8C8FEE6ED0FFC44B74775493F647391D" colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,221</td>
<td id="1219D5832951C126F0154C24ACD00832" colspan="1" pageId="0" pageNumber="41" rowspan="1">1,1,221</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P5</emphasis>
<paragraph id="5959C90962A99A19226C8EDF1336B15F" pageId="0" pageNumber="41">
<emphasis id="D4A9610C49BD3E2752C2942ED3F58C56" bold="true" pageId="0" pageNumber="41">P5</emphasis>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lonchoeidestenhelia prote gen. et sp. nov., female A P 3, anterior B P 4, anterior C P 5, anterior. Scale bars: 50 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41">8C</figureCitation>
<figureCitation id="8E702699B3F140536697F5C3E2462DF4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lonchoeidestenhelia prote gen. et sp. nov., female A P 3, anterior B P 4, anterior C P 5, anterior. Scale bars: 50 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure8" httpUri="https://binary.pensoft.net/fig/472438" pageId="0" pageNumber="41">8C</figureCitation>
): Baseoendopod transversely elongate; with five setae, of which outermost shortest and set close to adjoining element, all setae whip-like without any transformation. Exopod oval, with some outer proximal spinules, with six setae, of which fourth from outer to inner margin shortest.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P6</emphasis>
<paragraph id="DC78ACE0F4E22FA06B58CD116D3C5A84" pageId="0" pageNumber="41">
<emphasis id="F151B87799401A159721C4150D37278B" bold="true" pageId="0" pageNumber="41">P6</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3B, C</figureCitation>
<figureCitation id="E90D501574DA63649979D7C8EE361DE4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lonchoeidestenhelia prote gen. et sp. nov., female A urosome, dorsal (P 5 - bearing somite omitted) B urosome, lateral (P 5 - bearing somite omitted) C urosome, ventral (P 5 - bearing somite omitted). Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure3" httpUri="https://binary.pensoft.net/fig/472433" pageId="0" pageNumber="41">3B, C</figureCitation>
): Represented by a minute flap covering ventrolateral genital aperture, fused to somite, without surface ornamentation, with one slender seta.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">
<emphasis italics="true" pageId="0" pageNumber="41">Male</emphasis>
<paragraph id="7FF1EA572514CA187A5A6704C0E2C326" pageId="0" pageNumber="41">
<emphasis id="1B0A0C57641E306812E60CB7EB4B0F9C" bold="true" pageId="0" pageNumber="41">
<emphasis id="2B979533ECAE80166095F0B3C9ABFC55" italics="true" pageId="0" pageNumber="41">Male</emphasis>
.
</emphasis>
Total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 289
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="05ECB253AEC54F9A4781FFF43B15959B" originalValue="µm">µm</normalizedToken>
to 460
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="A921F41B139811199AF3525880F810EF" originalValue="µm">µm</normalizedToken>
(mean, 377
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="1E6E46CECB5B45FD76A03216C98FBAF7" originalValue="µm">µm</normalizedToken>
; n, 7; total body length of allotype, 415
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="2B147C2BFB5CDB4A274B454619C06FD5" originalValue="µm">µm</normalizedToken>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Prosome</emphasis>
<paragraph id="3FD3A2D2390ECAF790C9AE67CEB382FE" pageId="0" pageNumber="41">
<emphasis id="448C74721BBB9C0AA812AD5CFA301F9E" bold="true" pageId="0" pageNumber="41">Prosome</emphasis>
: As in female.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Urosome</emphasis>
<paragraph id="9486EFDD0AF447431587CB486C142261" pageId="0" pageNumber="41">
<emphasis id="73A57651FBE9E85A580383F8D124E604" bold="true" pageId="0" pageNumber="41">Urosome</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">9A-C</figureCitation>
<figureCitation id="7D4F8667D573AB62182F577E05DE4D9A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">9A-C</figureCitation>
): Largely as in female except for genital somite and third urosomite separated, and for lateral and ventral spinular ornamentation.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 9.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="EE03442ABF74B1AAA27A87A899534B1C" doi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41" start="Figure 9" startId="F9">
<paragraph id="5E964099D579EBE0BFA10B629360C2B3" pageId="0" pageNumber="41">
<emphasis id="5421828F4DBC14D746547D0B4F2E33CE" bold="true" pageId="0" pageNumber="41">Figure 9.</emphasis>
<taxonomicName id="9D18ED2FA88BBDB5D9F5EE2A7DBF2767" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="811D97A12A7C1C702E5DB5B1864E3CB5" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., male
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="2FBA15856E2089117A75D87E318CAA51" bold="true" pageId="0" pageNumber="41">A</emphasis>
urosome, dorsal
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="BEB16E1F5346F02E0F833B4032CA30AD" bold="true" pageId="0" pageNumber="41">B</emphasis>
urosome, lateral
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
<emphasis id="9CAD5316D36A70DC680F4719EA069495" bold="true" pageId="0" pageNumber="41">C</emphasis>
urosome, ventral. Scale bars: 100
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="DA3CC7178AA4F5ACE1B27CA984CD7C3E" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A-C</emphasis>
<emphasis id="EBE174AC9DFFB5A5F68B5E73FA570CAA" bold="true" pageId="0" pageNumber="41">A-C</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Caudal rami</emphasis>
<paragraph id="1120C4E93CFEBA42FA4046F891DE2278" pageId="0" pageNumber="41">
<emphasis id="896710AC4878019BF3BF7F931D31DEA8" bold="true" pageId="0" pageNumber="41">Caudal rami</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">9A-C</figureCitation>
<figureCitation id="44B62A81DDAE1E979714A93811B73B6A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">9A-C</figureCitation>
): As in female.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Sexual dimorphism</emphasis>
<paragraph id="78691847AA6CA41DF57832A2405B9593" pageId="0" pageNumber="41">
<emphasis id="445F04FD8891FAEA556E8B72410EA33F" bold="true" pageId="0" pageNumber="41">Sexual dimorphism</emphasis>
: Expressed in A1, P2ENP, P5, and P6.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Antennule</emphasis>
<paragraph id="D784D04578DE0E1855977F684EE5CCEE" pageId="0" pageNumber="41">
<emphasis id="F9A5F56DC78C398283A8FEA85BE9A464" bold="true" pageId="0" pageNumber="41">Antennule</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">10A, B</figureCitation>
<figureCitation id="019B2C1D483F6761864984393ACF9AC7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">10A, B</figureCitation>
): 10-segmented, haplocer. All segments smooth except for first and seventh segment with spinules as shown. All setae smooth except for one and two pinnate setae on first and second segments, respectively, and one and two modified spine-like elements on seventh and eighth segments. Armature formula: 1(1); 2(11); 3(7+ae); 4(2), 5(5+(1+ae)); 6(1); 7(3); 8(3); 9(4);10(4+acro). Acrothek consisting of two setae and one minute aesthetasc fused at their bases.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 10.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="19747C55198386B0C4ECA74F0423F9C9" doi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41" start="Figure 10" startId="F10">
<paragraph id="483794E5E24ED4BF55C980347C3505C8" pageId="0" pageNumber="41">
<emphasis id="FEDCBF5CDABD1460C84ACDBE6BDB7D5C" bold="true" pageId="0" pageNumber="41">Figure 10.</emphasis>
<taxonomicName id="18B08A46A3DAE6280474DD2F64871042" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="FF31A41797260C6F7FF9FF98FB462075" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., male
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="352CE0B437219AC80C0AA0B15A8E1942" bold="true" pageId="0" pageNumber="41">A</emphasis>
rostrum antennule with armature
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="21BAF8D97429511DEC2486A96D1BB0EE" bold="true" pageId="0" pageNumber="41">B</emphasis>
rostrum and antennule without armature
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
<emphasis id="6724956566D782D0543637AD6AD4A081" bold="true" pageId="0" pageNumber="41">C</emphasis>
P2ENP, anterior
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
<emphasis id="18982B335B3DF5602B9849C541144954" bold="true" pageId="0" pageNumber="41">D</emphasis>
P5, anterior
<emphasis bold="true" pageId="0" pageNumber="41">E</emphasis>
<emphasis id="4A2B8F6177694A110887067EEC36140E" bold="true" pageId="0" pageNumber="41">E</emphasis>
P6, anterior. Scale bars: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="9DAD737C48AFA7C5EDC5017A71BD48A2" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A-E</emphasis>
<emphasis id="86C0CAFEA9B2430900571CF775B4E21D" bold="true" pageId="0" pageNumber="41">A-E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Antenna, mandible, maxillule, maxilla, and maxilliped</emphasis>
<paragraph id="38FC8790A2F61F4ACFFB0934398903CC" pageId="0" pageNumber="41">
<emphasis id="C11C12409B3F5C949A51F9896DD6923A" bold="true" pageId="0" pageNumber="41">Antenna, mandible, maxillule, maxilla, and maxilliped</emphasis>
: As in female.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P1</emphasis>
<paragraph id="92CA1BC3426130B1EEF867707F1AF2B5" pageId="0" pageNumber="41">
<emphasis id="3F464B4D099015B0A9924D28128FB738" bold="true" pageId="0" pageNumber="41">P1</emphasis>
: As in female.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P2</emphasis>
<paragraph id="C186732AE60BD7047ED2DDD067600611" pageId="0" pageNumber="41">
<emphasis id="BB18B5BDC850A7A1B9A7E9FC5EE130ED" bold="true" pageId="0" pageNumber="41">P2</emphasis>
: EXP (not shown) as in female. ENP (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">10C</figureCitation>
<figureCitation id="91F432104679EB7DC05CA6E5AAF029D0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">10C</figureCitation>
) sexually dimorphic; basis, coxa and ENP1 largely as in female; ENP2 transformed, proximal half globular, distal half triangular, with inner notch indicating former division between ENP2 and ENP3, proximal half with inner long setules, distal half with subdistal pore, proximal half with two inner (one proximal and one medial) small setae, distal half with one inner subdistal strong pinnate element, and one apical sigmoid pinnate process with proximal row of outer spinules modified into lanceolate ornaments.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P3 and P4</emphasis>
<paragraph id="8883DF2B63B3890C769A62BA812712BC" pageId="0" pageNumber="41">
<emphasis id="FA8D15DEF1F5C9B17688CCAEB89B3B02" bold="true" pageId="0" pageNumber="41">P3 and P4</emphasis>
: As in female.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P5</emphasis>
<paragraph id="E2C7E8B9C6670A226883DBC6672E7B86" pageId="0" pageNumber="41">
<emphasis id="AE44D25CDC1BBE4D76280B95EACB0979" bold="true" pageId="0" pageNumber="41">P5</emphasis>
(Figs
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">9B, C</figureCitation>
<figureCitation id="2BF972578C069633F2AB9A4DCA07853D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">9B, C</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">10D</figureCitation>
<figureCitation id="1812093F3463E3BCD7B6AFEFFF04418F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">10D</figureCitation>
): Baseoendopods of both legs fused medially forming a continuous plate; each endopodal lobe with two setae, of which inner well-developed, outer small. Exopod small, discrete; with four elements, of which two outermost spine-like, third element from outer to inner margin shortest, innermost seta longest.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">P6</emphasis>
<paragraph id="5BDB24C904EEBB9EA345DA6B0E887D18" pageId="0" pageNumber="41">
<emphasis id="9C5D74EF2896ADAA783C1705C4AB8F0A" bold="true" pageId="0" pageNumber="41">P6</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">9B, C</figureCitation>
<figureCitation id="1B10137E309B5B3420B7B5538F8BD56D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lonchoeidestenhelia prote gen. et sp. nov., male A urosome, dorsal B urosome, lateral C urosome, ventral. Scale bars: 100 µm (A-C)." figureDoi="10.3897/zookeys.987.52906.figure9" httpUri="https://binary.pensoft.net/fig/472439" pageId="0" pageNumber="41">9B, C</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">10E</figureCitation>
<figureCitation id="3D2F9F9C783DA470B642BC9B37F28A2E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lonchoeidestenhelia prote gen. et sp. nov., male A rostrum antennule with armature B rostrum and antennule without armature C P 2 ENP, anterior D P 5, anterior E P 6, anterior. Scale bars: 50 µm (A-E)." figureDoi="10.3897/zookeys.987.52906.figure10" httpUri="https://binary.pensoft.net/fig/472440" pageId="0" pageNumber="41">10E</figureCitation>
): Asymmetrical, only one leg functional; each leg with three setae, of which medial longest, inner and outer elements subequal in length.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Variability.</emphasis>
<paragraph id="47986F1933689FE9417D207826D38231" pageId="0" pageNumber="41">
<emphasis id="FE43BF93FE04116797DFFDFD7E043FE4" bold="true" pageId="0" pageNumber="41">Variability.</emphasis>
One intersexual specimen (ICML-EMUCOP-180119-10) possesses female antennules, displays genital double-somite (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lonchoeidestenhelia prote gen. et sp. nov., intersex individual A P 5 - bearing somite and double genital-somite, lateral B P 5 - bearing somite and double genital-somite, ventral. Scale bar: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure11" httpUri="https://binary.pensoft.net/fig/472441" pageId="0" pageNumber="41">11B</figureCitation>
<figureCitation id="DFB041CF0CD29FFF1B9A4EC109B2B117" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lonchoeidestenhelia prote gen. et sp. nov., intersex individual A P 5 - bearing somite and double genital-somite, lateral B P 5 - bearing somite and double genital-somite, ventral. Scale bar: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure11" httpUri="https://binary.pensoft.net/fig/472441" pageId="0" pageNumber="41">11B</figureCitation>
), and lacks dimorphism in swimming legs, but the P5 (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lonchoeidestenhelia prote gen. et sp. nov., intersex individual A P 5 - bearing somite and double genital-somite, lateral B P 5 - bearing somite and double genital-somite, ventral. Scale bar: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure11" httpUri="https://binary.pensoft.net/fig/472441" pageId="0" pageNumber="41">11A, B</figureCitation>
<figureCitation id="A5E991D1940D9D15C6D3FB93775A6618" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lonchoeidestenhelia prote gen. et sp. nov., intersex individual A P 5 - bearing somite and double genital-somite, lateral B P 5 - bearing somite and double genital-somite, ventral. Scale bar: 50 µm (A, B)." figureDoi="10.3897/zookeys.987.52906.figure11" httpUri="https://binary.pensoft.net/fig/472441" pageId="0" pageNumber="41">11A, B</figureCitation>
) seems more of the male type with exopod bearing four elements (but two outer elements long and seta-like), and both baseoendopods fused medially and with two setae (outer small, inner long and pinnate), and the P6 possesses two setae.
</paragraph>
<caption doi="10.3897/zookeys.987.52906.figure11" httpUri="https://binary.pensoft.net/fig/472441" pageId="0" pageNumber="41" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 11.</emphasis>
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
<caption id="B7B64BE9F83920490FB941F6DD4B8D2B" doi="10.3897/zookeys.987.52906.figure11" httpUri="https://binary.pensoft.net/fig/472441" pageId="0" pageNumber="41" start="Figure 11" startId="F11">
<paragraph id="8FF22E7F9320C88518A5FCB94E580E5B" pageId="0" pageNumber="41">
<emphasis id="61DC6B8DF98974435B5C70C7AEE10727" bold="true" pageId="0" pageNumber="41">Figure 11.</emphasis>
<taxonomicName id="47A0728B65CE4F0703DC386A5F0EBAA3" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia prote" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="prote">
<emphasis id="C610FE1995692913514F23861FD1DF90" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia prote</emphasis>
</taxonomicName>
gen. et sp. nov., intersex individual
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
<emphasis id="DD1537DEE961DF14E85653DDE79DDB1E" bold="true" pageId="0" pageNumber="41">A</emphasis>
P5-bearing somite and double genital-somite, lateral
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
<emphasis id="1F3E2B9DF9430875509F7E684AE47D14" bold="true" pageId="0" pageNumber="41">B</emphasis>
P5-bearing somite and double genital-somite, ventral. Scale bar: 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="D6CE09878DBB92B53723C7CB5361A728" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="41">A, B</emphasis>
<emphasis id="F6047048135A8A042474E5F710A671FF" bold="true" pageId="0" pageNumber="41">A, B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="etymology">
<paragraph pageId="0" pageNumber="41">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="41">
<subSubSection id="0A0064186743D3027DADEC8A3BC2DEC7" pageId="0" pageNumber="41" type="etymology">
<paragraph id="7EE6E89101E13B2A65B630B22ABA01D7" pageId="0" pageNumber="41">Etymology.</paragraph>
<paragraph id="6F60D3F2D01691AD10844364E50ED93E" pageId="0" pageNumber="41">
The specific epithet from the Greek
<emphasis italics="true" pageId="0" pageNumber="41">
<normalizedToken originalValue="πρώτη">πρώτη</normalizedToken>
<emphasis id="6832E5A3B6BC33C58B53FB6E7A758686" italics="true" pageId="0" pageNumber="41">
<normalizedToken id="6D421FA0AA84418FA3E0ABD95D7FBD2D" originalValue="πρώτη">πρώτη</normalizedToken>
</emphasis>
, prṓtē, first, makes reference to the first species of
<taxonomicName authorityName="Gómez" authorityYear="2020" class="Hexanauplia" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia</emphasis>
<taxonomicName id="91C80173F82401EAABEBDFC009E6F49C" authorityName="Gómez" authorityYear="2020" class="Copepoda" family="Miraciidae" genus="Lonchoeidestenhelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lonchoeidestenhelia" order="Harpacticoida" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="6EEADBE6D4DACE8D971657F7AD6AEBD7" italics="true" pageId="0" pageNumber="41">Lonchoeidestenhelia</emphasis>
</taxonomicName>
gen. nov. described so far. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="type locality">
<paragraph pageId="0" pageNumber="41">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="41">
<subSubSection id="A8475AF868A90CBF0159F2AA93BEB7F1" pageId="0" pageNumber="41" type="type locality">
<paragraph id="ED76F547576F2677708461BD6A9DDD50" pageId="0" pageNumber="41">Type locality.</paragraph>
<paragraph id="4132FE70AC16DC94AC986654D6AE485E" pageId="0" pageNumber="41">
Mexico, Sinaloa State:
<normalizedToken originalValue="Urías">Urias</normalizedToken>
<normalizedToken id="A9C8769E90AEC291F7CC9D05C8E8175A" originalValue="Urías">Urias</normalizedToken>
estuary, stn 2,
<geoCoordinate degrees="23.1587" direction="north" orientation="latitude" precision="5" value="23.1587">23.1587°N</geoCoordinate>
<geoCoordinate id="290726D2E99F44C716E3E7628F7CC1C8" degrees="23.1587" direction="north" orientation="latitude" precision="5" value="23.1587">23.1587°N</geoCoordinate>
,
<geoCoordinate degrees="106.3326" direction="west" orientation="longitude" precision="5" value="-106.3326">106.3326°W</geoCoordinate>
<geoCoordinate id="6295A8C0E81463023B965A3639AF5A7E" degrees="106.3326" direction="west" orientation="longitude" precision="5" value="-106.3326">106.3326°W</geoCoordinate>
, depth 1.8 m, organic carbon content 3.99%, organic matter content 6.86%, sand 80.42%, clay 8.29%, silt 11.28%.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="other localities">
<paragraph pageId="0" pageNumber="41">Other localities.</paragraph>
<paragraph pageId="0" pageNumber="41">
<subSubSection id="92F11EE4BE3FDFE37EBA4D4CC603A5A1" pageId="0" pageNumber="41" type="other localities">
<paragraph id="7730E76484B4FC70F125E888ACA848D3" pageId="0" pageNumber="41">Other localities.</paragraph>
<paragraph id="65A1F9EDE0C5AE3233FCE8434C70CB5E" pageId="0" pageNumber="41">
Mexico, Sinaloa State:
<normalizedToken originalValue="Urías">Urias</normalizedToken>
<normalizedToken id="051C1C26582EBB3CD7A63D66E7879585" originalValue="Urías">Urias</normalizedToken>
estuary, stn 1:
<geoCoordinate degrees="23.15194" direction="north" orientation="latitude" precision="1" value="23.15194">23.15194°N</geoCoordinate>
<geoCoordinate id="A5FAAE8F9687B46D1B496D80400E160B" degrees="23.15194" direction="north" orientation="latitude" precision="1" value="23.15194">23.15194°N</geoCoordinate>
,
<geoCoordinate degrees="106.3128" direction="west" orientation="longitude" precision="5" value="-106.3128">106.3128°W</geoCoordinate>
<geoCoordinate id="D9ABF795FC5741F6BE1C9C842A369A6C" degrees="106.3128" direction="west" orientation="longitude" precision="5" value="-106.3128">106.3128°W</geoCoordinate>
, depth 1.5 m, organic carbon content 3.74%, organic matter content 6.43%, sand 25.31%, clay 35.75%, silt 38.94%, stn 4:
<geoCoordinate degrees="23.1840" direction="north" orientation="latitude" precision="5" value="23.184">23.1840°N</geoCoordinate>
<geoCoordinate id="5C16D22AA5C98EBE1A586E144A3306B7" degrees="23.1840" direction="north" orientation="latitude" precision="5" value="23.184">23.1840°N</geoCoordinate>
,
<geoCoordinate degrees="106.3579" direction="west" orientation="longitude" precision="5" value="-106.3579">106.3579°W</geoCoordinate>
<geoCoordinate id="1B114C8CF9731F929A6AA314EFFFF825" degrees="106.3579" direction="west" orientation="longitude" precision="5" value="-106.3579">106.3579°W</geoCoordinate>
, depth 0.7 m, organic carbon content 1.13%, organic matter content 1.94%, sand 82.44%, clay 8.27%, silt 9.29%, stn 7:
<geoCoordinate degrees="23.2174" direction="north" orientation="latitude" precision="5" value="23.2174">23.2174°N</geoCoordinate>
<geoCoordinate id="CAD3F114DA2C8FAB6FB0B4B7ACC97240" degrees="23.2174" direction="north" orientation="latitude" precision="5" value="23.2174">23.2174°N</geoCoordinate>
,
<geoCoordinate degrees="106.3917" direction="west" orientation="longitude" precision="5" value="-106.3917">106.3917°W</geoCoordinate>
<geoCoordinate id="064ADB9AE3C9D1F2FF833B6A30FCD96A" degrees="106.3917" direction="west" orientation="longitude" precision="5" value="-106.3917">106.3917°W</geoCoordinate>
, depth 3.7 m, organic carbon content 5.59%, organic matter content 9.62%, sand 10.78%, clay 37.54%, silt 51.68%, stn 9:
<geoCoordinate degrees="23.1904" direction="north" orientation="latitude" precision="5" value="23.1904">23.1904°N</geoCoordinate>
<geoCoordinate id="C3F81E0E346240142D544397E763154E" degrees="23.1904" direction="north" orientation="latitude" precision="5" value="23.1904">23.1904°N</geoCoordinate>
,
<geoCoordinate degrees="106.4121" direction="west" orientation="longitude" precision="5" value="-106.4121">106.4121°W</geoCoordinate>
<geoCoordinate id="139C5AA57F11231A21B549B8AEDD5EF3" degrees="106.4121" direction="west" orientation="longitude" precision="5" value="-106.4121">106.4121°W</geoCoordinate>
, depth 5.4 m, organic carbon content 1.41%, organic matter content 2.43%, sand 64.81%, clay 8.09%, silt 27.11%, and stn 10:
<geoCoordinate degrees="23.1815" direction="north" orientation="latitude" precision="5" value="23.1815">23.1815°N</geoCoordinate>
<geoCoordinate id="8A5932201839735973A3936AFB853471" degrees="23.1815" direction="north" orientation="latitude" precision="5" value="23.1815">23.1815°N</geoCoordinate>
,
<geoCoordinate degrees="106.4214" direction="west" orientation="longitude" precision="5" value="-106.4214">106.4214°W</geoCoordinate>
<geoCoordinate id="1C1BA505C6E6A7E24680DF7237406A98" degrees="106.4214" direction="west" orientation="longitude" precision="5" value="-106.4214">106.4214°W</geoCoordinate>
, depth 6.0 m, organic carbon content 1.2%, organic matter content 2.07%, sand 69.12%, clay 7.91%, silt 22.97%.
</paragraph>
</subSubSection>

View file

@ -1,105 +1,105 @@
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<paragraph pageId="3" pageNumber="4">
<taxonomicName LSID="http://species-id.net/wiki/Seriana_equata" authority="Singh, 1969" authorityName="Singh" authorityYear="1969" class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Seriana equata (Singh, 1969)</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="4">rec. n.</taxonomicNameLabel>
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<paragraph id="F76E898A008C694A2D61DC6CFF4725BF" pageId="3" pageNumber="4">
<taxonomicName id="3696877B6F39067DEEF40C5C93A0D1C4" ID-CoL="4WTB9" LSID="http://species-id.net/wiki/Seriana_equata" authority="Singh, 1969" authorityName="Singh" authorityYear="1969" class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Seriana equata (Singh, 1969)</taxonomicName>
<taxonomicNameLabel id="D65FB1D28EAABA457D167CA3B19B48CC" pageId="3" pageNumber="4">rec. n.</taxonomicNameLabel>
Figures 11-17
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<subSubSection pageId="3" pageNumber="4" type="reference_group">
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Cicadellidae" genus="Zygina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygina equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Zygina equata</taxonomicName>
<subSubSection id="053C2D849003403ADB1E4A35E0CAF585" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="552AEF093D0970F10C37E7B064971321" pageId="3" pageNumber="4">
<taxonomicName id="CBEE2CAEF08151FB36B128099B705D69" class="Insecta" family="Cicadellidae" genus="Zygina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygina equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Zygina equata</taxonomicName>
Singh, 1969: 344, figs 20-23
</paragraph>
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Cicadellidae" genus="Empoascanara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Empoascanara equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Empoascanara equata</taxonomicName>
<paragraph id="7BB138DC7F49A560E91F6AD186954BBD" pageId="3" pageNumber="4">
<taxonomicName id="B7993BB9D77685D3274A96BF2D35D2CD" class="Insecta" family="Cicadellidae" genus="Empoascanara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Empoascanara equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Empoascanara equata</taxonomicName>
(Singh, 1969) (
<bibRefCitation author="Sohi, AS" journalOrPublisher="Entomon" pageId="4" pageNumber="5" pagination="203 - 205" title="New combinations of some Typhlocybines (Homoptera, Cicadellidae, Typhlocybinae) from India." volume="1" year="1976">Sohi 1976</bibRefCitation>
<bibRefCitation id="EA5BE1E6ECB2B534E3A417DA3F404F31" author="Sohi, AS" journalOrPublisher="Entomon" pageId="4" pageNumber="5" pagination="203 - 205" title="New combinations of some Typhlocybines (Homoptera, Cicadellidae, Typhlocybinae) from India." volume="1" year="1976">Sohi 1976</bibRefCitation>
: 204)
</paragraph>
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Seriana equata</taxonomicName>
<paragraph id="73C66D48A6BEDA72EA1EEE2D4CE775C4" pageId="3" pageNumber="4">
<taxonomicName id="3232D4EEC7372BB481831A1427D966E5" class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Seriana equata</taxonomicName>
(Singh, 1969) (
<bibRefCitation author="Dworakowska, I" journalOrPublisher="Bulletin de l'Academie Polonaise des Sciences, Serie des Sciences Biologiques" pageId="4" pageNumber="5" pagination="521 - 530" title="On some Typhlocybinae from India (Auchenorrhyncha, Cicadellidae)." volume="23" year="1975">Dworakowska and Viraktamath 1975</bibRefCitation>
<bibRefCitation id="9CF8E106FB8441DE483CF31C432E417F" author="Dworakowska, I" journalOrPublisher="Bulletin de l'Academie Polonaise des Sciences, Serie des Sciences Biologiques" pageId="4" pageNumber="5" pagination="521 - 530" title="On some Typhlocybinae from India (Auchenorrhyncha, Cicadellidae)." volume="23" year="1975">Dworakowska and Viraktamath 1975</bibRefCitation>
: 529, no figures)
</paragraph>
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana punjabensis" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="punjabensis">Seriana punjabensis</taxonomicName>
<paragraph id="94CC6FE490001268118B36752703D2EF" pageId="3" pageNumber="4">
<taxonomicName id="4F16512379504C88C83AD9FD4CD540B0" class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana punjabensis" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="punjabensis">Seriana punjabensis</taxonomicName>
Dworakowska, Nagaich &amp; Singh, 1978: 246, figs 38-42 (Syn. by
<bibRefCitation author="Sohi, AS" journalOrPublisher="Oriental Insects" pageId="4" pageNumber="5" pagination="159 - 213" title="A review of the indian Typhlocybinae (Homoptera: Cicadellidae) from India." volume="17" year="1983">Sohi and Dworakowska 1983</bibRefCitation>
<bibRefCitation id="013963D7A8F496B29BC1F0D9F986C8ED" author="Sohi, AS" journalOrPublisher="Oriental Insects" pageId="4" pageNumber="5" pagination="159 - 213" title="A review of the indian Typhlocybinae (Homoptera: Cicadellidae) from India." volume="17" year="1983">Sohi and Dworakowska 1983</bibRefCitation>
: 180)
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="material examined">
<paragraph pageId="3" pageNumber="4">Material examined.</paragraph>
<paragraph pageId="3" pageNumber="4">One male, China: Yunnan Prov., Xishuangbanna, Original Forest Park, 21 July 2008, coll. YUEHUA SONG; one male, China: Henan Prov., Luanchuan, Heyu, 19 August 2008, coll. JIANDA LI.</paragraph>
<caption pageId="3" pageNumber="4">
<paragraph pageId="3" pageNumber="4">
<subSubSection id="30D1E4CEE20B6969FCDC2EA6EA18F6DA" pageId="3" pageNumber="4" type="material examined">
<paragraph id="CDE819F1834484682E93D63EE821E7E2" pageId="3" pageNumber="4">Material examined.</paragraph>
<paragraph id="7C14C7ABFB0E3B1B70BE210D9AEB7EB1" pageId="3" pageNumber="4">One male, China: Yunnan Prov., Xishuangbanna, Original Forest Park, 21 July 2008, coll. YUEHUA SONG; one male, China: Henan Prov., Luanchuan, Heyu, 19 August 2008, coll. JIANDA LI.</paragraph>
<caption id="AF971D72DAF2156F08F86BFE2A4FA109" pageId="3" pageNumber="4">
<paragraph id="83680653BF2981F3AF03B6FD8842DD73" pageId="3" pageNumber="4">
Figures 11-21.
<taxonomicName class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Seriana equata</taxonomicName>
<taxonomicName id="6D370AA77F5F965221A44408761315B9" class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana equata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="equata">Seriana equata</taxonomicName>
(Singh, 1969) rec. n. (after
<bibRefCitation author="Dworakowska, I" journalOrPublisher="Bulletin de l'Academie Polonaise des Sciences, Serie des Sciences Biologiques" pageId="4" pageNumber="5" pagination="243 - 249" title="Kapsa simlensis sp. n. from India and some other Typhlocybinae (Auchenorrhyncha, Cicadellidae)." volume="26" year="1978">Dworakowska et al. 1978</bibRefCitation>
<bibRefCitation id="34612FC85DAE1B1BE8F762D79A600B42" author="Dworakowska, I" journalOrPublisher="Bulletin de l'Academie Polonaise des Sciences, Serie des Sciences Biologiques" pageId="4" pageNumber="5" pagination="243 - 249" title="Kapsa simlensis sp. n. from India and some other Typhlocybinae (Auchenorrhyncha, Cicadellidae)." volume="26" year="1978">Dworakowska et al. 1978</bibRefCitation>
) 11 Head and thorax, dorsal view 12 Face 13 Fore wing 14 Pygofer lobe, lateral view 15 Pygofer dorsal appendage 16 Aedeagus, lateral view 17 Aedeagus, ventral view;
<taxonomicName class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana indefinita" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="indefinita">Seriana indefinita</taxonomicName>
<taxonomicName id="D0DA82E828D05FE08B2BB6193CC0EA35" class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana indefinita" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="indefinita">Seriana indefinita</taxonomicName>
Dworakowska, 1971 18 Aedeagus, lateral view 19 Aedeagus, ventral view;
<taxonomicName class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana ochrata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ochrata">Seriana ochrata</taxonomicName>
<taxonomicName id="17093BF3867C94ABB8C07F457C1E307C" class="Insecta" family="Cicadellidae" genus="Seriana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Seriana ochrata" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ochrata">Seriana ochrata</taxonomicName>
Dworakowska, 1971 20 Aedeagus, lateral view 21 Aedeagus, ventral view. (after
<bibRefCitation author="Dworakowska, I" journalOrPublisher="Bulletin de l'Academie Polonaise des Sciences, Serie des Sciences Biologiques" pageId="4" pageNumber="5" pagination="341 - 350" title="On some genera of Erythroneurini (Cicadellidae, Typhlocybinae) from the Oriental Region." volume="19" year="1971">Dworakowska 1971</bibRefCitation>
<bibRefCitation id="559BA58EDAFEA5C3352DBDFA14C88D8E" author="Dworakowska, I" journalOrPublisher="Bulletin de l'Academie Polonaise des Sciences, Serie des Sciences Biologiques" pageId="4" pageNumber="5" pagination="341 - 350" title="On some genera of Erythroneurini (Cicadellidae, Typhlocybinae) from the Oriental Region." volume="19" year="1971">Dworakowska 1971</bibRefCitation>
)
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="host plant">
<paragraph pageId="3" pageNumber="4">Host plant.</paragraph>
<paragraph pageId="3" pageNumber="4">
<subSubSection id="4A47371E8305D6C781042FF90B359818" pageId="3" pageNumber="4" type="host plant">
<paragraph id="128A8DEF549C9A1C8A817EB4D07D7676" pageId="3" pageNumber="4">Host plant.</paragraph>
<paragraph id="FD4E326A9C5F61B2E9942767B588D6FE" pageId="3" pageNumber="4">
Grasses, potato, black gram, cowpea, Egyptian clover, groundnut, linseed, lucerne, musk melon, spinach, sweet potato, sunnhemp (
<bibRefCitation author="Sohi, AS" journalOrPublisher="Oriental Insects" pageId="4" pageNumber="5" pagination="159 - 213" title="A review of the indian Typhlocybinae (Homoptera: Cicadellidae) from India." volume="17" year="1983">Sohi and Dworakowska 1983</bibRefCitation>
<bibRefCitation id="6293565A0BE2661B6C5DEAD1C1E6D521" author="Sohi, AS" journalOrPublisher="Oriental Insects" pageId="4" pageNumber="5" pagination="159 - 213" title="A review of the indian Typhlocybinae (Homoptera: Cicadellidae) from India." volume="17" year="1983">Sohi and Dworakowska 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="distribution">
<paragraph pageId="3" pageNumber="4">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="4">India; China (Henan, Yunnan).</paragraph>
<subSubSection id="E5FFF94F4E48A0F1F917D17A436FA862" pageId="3" pageNumber="4" type="distribution">
<paragraph id="C96B200C3815779B0A82118FAB813A1B" pageId="3" pageNumber="4">Distribution.</paragraph>
<paragraph id="345C0C10C730D35B4BB463528F01FF1D" pageId="3" pageNumber="4">India; China (Henan, Yunnan).</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.70.52043</mods:identifier>
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<mods:identifier id="442BD2DCC08CE814961E4CFB3F3AC033" type="DOI">http://dx.doi.org/10.3897/mycokeys.70.52043</mods:identifier>
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<treatment ID-GBIF-Taxon="165857307" LSID="urn:lsid:plazi:treatment:C429487BB6395CF2A5B5E45B14E51180" httpUri="http://treatment.plazi.org/id/C429487BB6395CF2A5B5E45B14E51180" lastPageNumber="19" pageId="0" pageNumber="19">
<subSubSection pageId="0" pageNumber="19" type="nomenclature">
<paragraph pageId="0" pageNumber="19">
<taxonomicName LSID="C429487B-B639-5CF2-A5B5-E45B14E51180" authority="(S. Hughes) Subram., J. Indian bot. Soc. 35: 78 (1956)" authorityName="Subram., J. Indian bot. Soc. 35: 78" authorityYear="1956" baseAuthorityName="S. Hughes" baseAuthorityYear="1956" class="Sordariomycetes" family="Didymosphaeriaceae" genus="Spegazzinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spegazzinia deightonii" order="Pleosporales" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="deightonii">Spegazzinia deightonii (S. Hughes) Subram., J. Indian bot. Soc. 35: 78 (1956)</taxonomicName>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Spegazzinia deightonii (MFLU 19 - 2908) a-c fungal colonies on host surface d conidiophore mother cell of α conidia e-g α conidia i a developmental stage of β conidia h, k conidia l colonies on PDA after 28 days showing sporulation j, m-p β conidia. Scale bars: 500 μm (a), 200 μm (b), 50 μm (c), 20 μm (e-h), 10 μm (d, k, m-p), 5 μm (i, j)." figureDoi="10.3897/mycokeys.70.52043.figure2" httpUri="https://binary.pensoft.net/fig/434238" pageId="0" pageNumber="19">Figure 2</figureCitation>
<treatment id="C429487BB6395CF2A5B5E45B14E51180" ID-GBIF-Taxon="165857307" LSID="urn:lsid:plazi:treatment:C429487BB6395CF2A5B5E45B14E51180" httpUri="http://treatment.plazi.org/id/C429487BB6395CF2A5B5E45B14E51180" lastPageNumber="19" pageId="0" pageNumber="19" scope_order="Pleosporales">
<subSubSection id="1D93453DE0663B2ABBF1785287B9AC0C" pageId="0" pageNumber="19" type="nomenclature">
<paragraph id="71C009C59EDA70F56A5DF40BA4809AB9" pageId="0" pageNumber="19">
<taxonomicName id="103EBA7B195A43C4AA0A3CA5CCD720D3" LSID="C429487B-B639-5CF2-A5B5-E45B14E51180" authority="(S. Hughes) Subram., J. Indian bot. Soc. 35: 78 (1956)" authorityName="Subram., J. Indian bot. Soc. 35: 78" authorityYear="1956" baseAuthorityName="S. Hughes" baseAuthorityYear="1956" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Spegazzinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spegazzinia deightonii" order="Pleosporales" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="deightonii">Spegazzinia deightonii (S. Hughes) Subram., J. Indian bot. Soc. 35: 78 (1956)</taxonomicName>
<figureCitation id="749C957079CB37CAF27798BBE90D7221" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Spegazzinia deightonii (MFLU 19 - 2908) a-c fungal colonies on host surface d conidiophore mother cell of α conidia e-g α conidia i a developmental stage of β conidia h, k conidia l colonies on PDA after 28 days showing sporulation j, m-p β conidia. Scale bars: 500 μm (a), 200 μm (b), 50 μm (c), 20 μm (e-h), 10 μm (d, k, m-p), 5 μm (i, j)." figureDoi="10.3897/mycokeys.70.52043.figure2" httpUri="https://binary.pensoft.net/fig/434238" pageId="0" pageNumber="19">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="description">
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
<paragraph pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">Saprobic</emphasis>
<subSubSection id="51BB40795E4C2201EB59621C78834BD9" pageId="0" pageNumber="19" type="description">
<paragraph id="819309CCE49BF1F635A40BCCDBE6C7DD" pageId="0" pageNumber="19">Description.</paragraph>
<paragraph id="37E4BE50B7348FDB231A8FC5D1BC11E0" pageId="0" pageNumber="19">
<emphasis id="A02F253A925A88CB4D046A5B5DAA98A5" italics="true" pageId="0" pageNumber="19">Saprobic</emphasis>
on dead leaves of
<taxonomicName class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Musa</emphasis>
<taxonomicName id="E6A9CA8EAD50A8FDB3EFCEDDB2252D05" class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="13B90325653D48B33E2344E2C9088844" italics="true" pageId="0" pageNumber="19">Musa</emphasis>
</taxonomicName>
sp.
<emphasis bold="true" pageId="0" pageNumber="19">Sexual morph</emphasis>
<emphasis id="5EFE5F745C761ED69102A9F0899403BC" bold="true" pageId="0" pageNumber="19">Sexual morph</emphasis>
Undetermined.
<emphasis bold="true" pageId="0" pageNumber="19">Asexual morph</emphasis>
<emphasis id="6F1215206F8020B57C9803FF451562BC" bold="true" pageId="0" pageNumber="19">Asexual morph</emphasis>
Hyphomycetous.
<emphasis italics="true" pageId="0" pageNumber="19">Sporodochia</emphasis>
<emphasis id="C0298133BAA930BBCD2C2F916CBBE32C" italics="true" pageId="0" pageNumber="19">Sporodochia</emphasis>
powder like, dark, dense, dry, 1-3 mm diameter.
<emphasis italics="true" pageId="0" pageNumber="19">Conidiophore mother cells</emphasis>
<emphasis id="609FE69108ADAE684104023185E857DE" italics="true" pageId="0" pageNumber="19">Conidiophore mother cells</emphasis>
3.5-6.8
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="52C7FA4068E9F3D69F7FF4BBB91BD0BF" originalValue="×">x</normalizedToken>
2.5-5.0
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="DEF7EDFDB3F74F46E7C964BE75286B47" originalValue="μm">μm</normalizedToken>
(
<emphasis italics="true" pageId="0" pageNumber="19"></emphasis>
<emphasis id="1AC499C03063EAB86183C700A0D36811" italics="true" pageId="0" pageNumber="19"></emphasis>
= 5.59
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="BA3630822D5E67998CE3173F350E81A4" originalValue="×">x</normalizedToken>
4.15
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="98993A036A4E9408586D50946F23E98C" originalValue="μm">μm</normalizedToken>
, n = 6), hyaline to light brown, subspherical or doliiform.
<emphasis italics="true" pageId="0" pageNumber="19">Conidiophores</emphasis>
<emphasis id="3677AA11579F4D814B968C33CC057324" italics="true" pageId="0" pageNumber="19">Conidiophores</emphasis>
long or short and give rise to two types of conidia referred here as
<normalizedToken originalValue="α">α</normalizedToken>
<normalizedToken id="2D65B0BE4F5A11EB7A409B0221CE637E" originalValue="α">α</normalizedToken>
and
<normalizedToken originalValue="β">β</normalizedToken>
<normalizedToken id="1FF0DE10F53803437D8C1B1F6E3160FF" originalValue="β">β</normalizedToken>
.
<emphasis italics="true" pageId="0" pageNumber="19">
<emphasis id="3E283645FD14A410F0C06E3A204BEA78" italics="true" pageId="0" pageNumber="19">
Conidiophores of
<normalizedToken originalValue="α">α</normalizedToken>
<normalizedToken id="A17F78BCE7DF630195AA6F183C1D5D02" originalValue="α">α</normalizedToken>
conidia
</emphasis>
up to 48-120
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="AF594147F82BFBD3966A41D0292FAC27" originalValue="×">x</normalizedToken>
1-2
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="8779E78EE58E9249DAF01921A621D301" originalValue="μm">μm</normalizedToken>
(
<emphasis italics="true" pageId="0" pageNumber="19"></emphasis>
<emphasis id="B77E6A8BEE6B8ECD598E9A8063F1802E" italics="true" pageId="0" pageNumber="19"></emphasis>
= 95.3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="7984B2F89A84DFA7F55DCCC1AEB860A7" originalValue="×">x</normalizedToken>
1.6
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="08EC70D893BDAEBA40DB818AF9E30420" originalValue="μm">μm</normalizedToken>
, n = 20) long, erect or flexuous, narrow, verruculose, unbranched, hyaline to golden-brown.
<emphasis italics="true" pageId="0" pageNumber="19">
<emphasis id="DAF94005FD0747D1017542268560D1FC" italics="true" pageId="0" pageNumber="19">
Conidiophores of
<normalizedToken originalValue="β">β</normalizedToken>
<normalizedToken id="48713EFB4F45687256CBFB27D8E71381" originalValue="β">β</normalizedToken>
conidia
</emphasis>
initially hyaline, light brown to brown at maturity, very short and slightly bent, 1.6-2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="7265E26E4E1AA22BF72594703DA903D4" originalValue="×">x</normalizedToken>
2.5-3
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="04577CE347E7CBDF2B5634E8897F610C" originalValue="μm">μm</normalizedToken>
(
<emphasis italics="true" pageId="0" pageNumber="19"></emphasis>
<emphasis id="99463828593C889D9B75BB195683474D" italics="true" pageId="0" pageNumber="19"></emphasis>
= 1.8
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="10269D62ADEBF0A64A1F282ED719ADA5" originalValue="×">x</normalizedToken>
2.6
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="0780DB197F3C984A0A41D25B6B5731BC" originalValue="μm">μm</normalizedToken>
, n =10).
<emphasis italics="true" pageId="0" pageNumber="19">Conidiogenous cell development</emphasis>
<emphasis id="A82B78A4F8DF296B6DF59C5D037DB248" italics="true" pageId="0" pageNumber="19">Conidiogenous cell development</emphasis>
basauxic, forming a single, terminal holoblastic conidium at the apex of conidiophore.
<emphasis italics="true" pageId="0" pageNumber="19">Conidial development</emphasis>
<emphasis id="E430DEAF537C30C2FBBAB885B89AE391" italics="true" pageId="0" pageNumber="19">Conidial development</emphasis>
holoblastic.
<emphasis italics="true" pageId="0" pageNumber="19">Conidia</emphasis>
<emphasis id="0F7BDAD60BA56E4ADEEE7797CC83C8D8" italics="true" pageId="0" pageNumber="19">Conidia</emphasis>
two types:
<normalizedToken originalValue="α">α</normalizedToken>
<emphasis italics="true" pageId="0" pageNumber="19">conidia</emphasis>
<normalizedToken id="02616372BDDD3CC7010DDA8BDA3929B1" originalValue="α">α</normalizedToken>
<emphasis id="D97A28BFD20B6809928C37ED8D25DE83" italics="true" pageId="0" pageNumber="19">conidia</emphasis>
stellate, 18-28
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="641A26DC896E7301C1048B253625F4A6" originalValue="×">x</normalizedToken>
17-29
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="CDE099D30165DA184C38D5895141DB73" originalValue="μm">μm</normalizedToken>
(
<emphasis italics="true" pageId="0" pageNumber="19"></emphasis>
<emphasis id="2656C06CEC6F557A65BD0BBD7ED1B504" italics="true" pageId="0" pageNumber="19"></emphasis>
= 25.1
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="81005223960A2103B5169D3FB196C8CE" originalValue="×">x</normalizedToken>
23.3
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="AE28226D5CBCF67997B3ACDC5A3CD3D4" originalValue="μm">μm</normalizedToken>
, n = 25), solitary, globose to variously shaped, with spines 4-6
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="B26FB867512483458CEDFD1DF4BFCE2F" originalValue="μm">μm</normalizedToken>
long, 4-8-celled, frequently 4- to 6-celled, deeply constricted at the septa.
<normalizedToken originalValue="β">β</normalizedToken>
<emphasis italics="true" pageId="0" pageNumber="19">conidia</emphasis>
<normalizedToken id="EE062C6E2A682385D7D1C50537831F76" originalValue="β">β</normalizedToken>
<emphasis id="C045B0F64F8E203FFBE45F63767B0AB8" italics="true" pageId="0" pageNumber="19">conidia</emphasis>
disc-shaped, initially hyaline, light brown to dark brown at maturity, 8-celled, 16-21
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B3B7634A90886F7E55301FF99EDD1794" originalValue="×">x</normalizedToken>
11-14
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="CB9F9D4D0B5E48FD2451F513B00D8A4E" originalValue="μm">μm</normalizedToken>
(
<emphasis italics="true" pageId="0" pageNumber="19"></emphasis>
<emphasis id="F83BEECF2895E6468B71D29F792D834F" italics="true" pageId="0" pageNumber="19"></emphasis>
= 19.2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="26D71029FA0A8E71971C31460670F62D" originalValue="×">x</normalizedToken>
14.6
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="ACE3F3253EB24938B74153FD5C88B177" originalValue="μm">μm</normalizedToken>
, n = 25), flat from both sides with short and blunt spines, frequently with attached conidiogenous cells when splitting from the conidiophores.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="description">
<paragraph pageId="0" pageNumber="19">Culture characteristics.</paragraph>
<paragraph pageId="0" pageNumber="19">Conidia germinating on PDA within 13-14 h. Colonies growing on PDA, reaching a diameter of 55 mm after 14 d at 25 °C, raised, moderately dense, undulate margin, middle grey, periphery brownish grey and olive green at immature stage; reverse white to greyish white.</paragraph>
<subSubSection id="3E8F81F907CF45B384C0D54BB40B561E" pageId="0" pageNumber="19" type="description">
<paragraph id="6790EAD49B7FCEE169836652317C64F0" pageId="0" pageNumber="19">Culture characteristics.</paragraph>
<paragraph id="2322065A0F3655446498A9E3295E0D0D" pageId="0" pageNumber="19">Conidia germinating on PDA within 13-14 h. Colonies growing on PDA, reaching a diameter of 55 mm after 14 d at 25 °C, raised, moderately dense, undulate margin, middle grey, periphery brownish grey and olive green at immature stage; reverse white to greyish white.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="materials_examined">
<paragraph pageId="0" pageNumber="19">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="19">
<subSubSection id="29E534FBB1027C13A140EA27EB4EACAF" pageId="0" pageNumber="19" type="materials_examined">
<paragraph id="C571096B734FC7EB7807FBF66B70471C" pageId="0" pageNumber="19">Material examined.</paragraph>
<paragraph id="A4B34F103F57B2E646EEEF546B41E9FA" pageId="0" pageNumber="19">
Thailand, Chiang Rai Province, Doi Thun, on a dead leaf of
<taxonomicName class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Musa</emphasis>
<taxonomicName id="AF89AC32059F355D159E063584343B40" class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="6072C9EFD16DBCCD319D69B1115A76C0" italics="true" pageId="0" pageNumber="19">Musa</emphasis>
</taxonomicName>
sp. (
<taxonomicName family="Musaceae" lsidName="" pageId="0" pageNumber="19" rank="family">Musaceae</taxonomicName>
<taxonomicName id="80B2EC371C033B2780B26D389B4E3093" class="Liliopsida" family="Musaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Zingiberales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="family">Musaceae</taxonomicName>
), 7 December 2018, M.C. Samarakoon, BNS 072 (MFLU 19-2908), living culture MFLUCC 20-0002.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="notes">
<paragraph pageId="0" pageNumber="19">Notes.</paragraph>
<paragraph pageId="0" pageNumber="19">
<taxonomicName authorityName="Subram., J. Indian bot. Soc. 35: 78" authorityYear="1956" baseAuthorityName="S. Hughes" class="Sordariomycetes" family="Apiosporaceae" genus="Spegazzinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spegazzinia deightonii" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="deightonii">
<emphasis italics="true" pageId="0" pageNumber="19">Spegazzinia deightonii</emphasis>
<subSubSection id="4948037FE263C760781D8E6C7FE0D78D" pageId="0" pageNumber="19" type="notes">
<paragraph id="01E7952D32A66ACEEA5336782FF06BB6" pageId="0" pageNumber="19">Notes.</paragraph>
<paragraph id="45C430213A96244BF4533CB33410C978" pageId="0" pageNumber="19">
<taxonomicName id="47FADD801B460E199EE9E195836454BD" authorityName="Subram., J. Indian bot. Soc. 35: 78" authorityYear="1956" baseAuthorityName="S. Hughes" class="Sordariomycetes" family="Apiosporaceae" genus="Spegazzinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spegazzinia deightonii" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="deightonii">
<emphasis id="07D8906E0940FCD99B0FDCE93C1AF467" italics="true" pageId="0" pageNumber="19">Spegazzinia deightonii</emphasis>
</taxonomicName>
MFLUCC 20-0002 clustered with
<taxonomicName lsidName="S. deightonii" pageId="0" pageNumber="19" rank="species" species="deightonii">
<emphasis italics="true" pageId="0" pageNumber="19">S. deightonii</emphasis>
<taxonomicName id="85DC6CC63DF5047B40101619603DB9E4" lsidName="S. deightonii" pageId="0" pageNumber="19" rank="species" species="deightonii">
<emphasis id="CE49EE4F080D96B2AE835D769E77AD50" italics="true" pageId="0" pageNumber="19">S. deightonii</emphasis>
</taxonomicName>
(yone 66, yone 212) with significant statistical support (Figure
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree revealed by RAxML from an analysis of SSU, LSU and ITS and TEF 1 - α sequence data of selected genera of family Didymosphaeriaceae, showing the phylogenetic position of Spegazzinia musae (MFLUCC 20 - 0001) and S. deightonii (MFLUCC 20 - 0002). ML bootstrap supports (≥ 60 %) and Bayesian posterior probabilities (≥ 0.95 BYPP) are given above in the branches, respectively. The tree was rooted with Pleospora herbarum and Stemphylium botryosum (Pleosporaceae). Strains generated in this study are indicated in red-bold. Ex-type species are indicated in bold. The scale bar represents the expected number of nucleotide substitutions per site. A best scoring RAxML tree is shown with a final ML optimization likelihood value of - 13516.66. The matrix had 795 distinct alignment patterns, with 33.60 % of undetermined characters or gaps. Estimated base frequencies were: A = 0.239862, C = 0.245185, G = 0.277025, T = 0.237927; substitution rates AC = 1.626982, AG = 2.468452, AT = 1.211822, CG = 1.092437, CT = 6.295657, GT = 1.000000; proportion of invariable sites I = 0.484119; gamma distribution shape parameter α = 0.445929." figureDoi="10.3897/mycokeys.70.52043.figure1" httpUri="https://binary.pensoft.net/fig/434237" pageId="0" pageNumber="19">1</figureCitation>
<figureCitation id="0FB59D9479BD1E9A256C2B1B3E577AA7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree revealed by RAxML from an analysis of SSU, LSU and ITS and TEF 1 - α sequence data of selected genera of family Didymosphaeriaceae, showing the phylogenetic position of Spegazzinia musae (MFLUCC 20 - 0001) and S. deightonii (MFLUCC 20 - 0002). ML bootstrap supports (≥ 60 %) and Bayesian posterior probabilities (≥ 0.95 BYPP) are given above in the branches, respectively. The tree was rooted with Pleospora herbarum and Stemphylium botryosum (Pleosporaceae). Strains generated in this study are indicated in red-bold. Ex-type species are indicated in bold. The scale bar represents the expected number of nucleotide substitutions per site. A best scoring RAxML tree is shown with a final ML optimization likelihood value of - 13516.66. The matrix had 795 distinct alignment patterns, with 33.60 % of undetermined characters or gaps. Estimated base frequencies were: A = 0.239862, C = 0.245185, G = 0.277025, T = 0.237927; substitution rates AC = 1.626982, AG = 2.468452, AT = 1.211822, CG = 1.092437, CT = 6.295657, GT = 1.000000; proportion of invariable sites I = 0.484119; gamma distribution shape parameter α = 0.445929." figureDoi="10.3897/mycokeys.70.52043.figure1" httpUri="https://binary.pensoft.net/fig/434237" pageId="0" pageNumber="19">1</figureCitation>
). All the strains of
<taxonomicName lsidName="S. deightonii" pageId="0" pageNumber="19" rank="species" species="deightonii">
<emphasis italics="true" pageId="0" pageNumber="19">S. deightonii</emphasis>
<taxonomicName id="32220C9B2B32D090B119168C16F210D6" lsidName="S. deightonii" pageId="0" pageNumber="19" rank="species" species="deightonii">
<emphasis id="FC38E93C9892BD9DF7CA23868CA85C54" italics="true" pageId="0" pageNumber="19">S. deightonii</emphasis>
</taxonomicName>
described in
<bibRefCitation author="Ellis, MB" journalOrPublisher="Cryptogamie Mycologie" pageId="0" pageNumber="19" refId="B16" refString="Ellis, MB, 1971. Dematiaceous hyphomycetes. Commonwealth Mycological Institute, Kew." title="Dematiaceous hyphomycetes. Commonwealth Mycological Institute, Kew." year="1971">Ellis (1971)</bibRefCitation>
<bibRefCitation id="760E5E85256ADEEC3BCAF514D1038AEC" author="Ellis, MB" journalOrPublisher="Cryptogamie Mycologie" pageId="0" pageNumber="19" refId="B16" refString="Ellis, MB, 1971. Dematiaceous hyphomycetes. Commonwealth Mycological Institute, Kew." title="Dematiaceous hyphomycetes. Commonwealth Mycological Institute, Kew." year="1971">Ellis (1971)</bibRefCitation>
and
<bibRefCitation DOI="doi.org/10.1016/j.simyco.2015.10.002" author="Tanaka, K" journalOrPublisher="Studies in mycology" pageId="0" pageNumber="19" refId="B61" refString="Tanaka, K, Hirayama, K, Yonezawa, H, Sato, G, Toriyabe, A, Kudo, H, Hashimoto, A, Matsumura, M, Harada, Y, Kurihara, Y, Shirouzu, T, 2015. Revision of the Massarineae (Pleosporales, Dothideomycetes). Studies in mycology 82: 75-136. ttps:// doi.org/10.1016/j.simyco.2015.10.002" title="Revision of the Massarineae (Pleosporales, Dothideomycetes). Studies in mycology 82: 75 - 136. ttps: //" url="doi.org/10.1016/j.simyco.2015.10.002" year="2015">Tanaka et al. (2015)</bibRefCitation>
<bibRefCitation id="9441E615BE1B2BC224AC4C37F539BBAC" DOI="doi.org/10.1016/j.simyco.2015.10.002" author="Tanaka, K" journalOrPublisher="Studies in mycology" pageId="0" pageNumber="19" refId="B61" refString="Tanaka, K, Hirayama, K, Yonezawa, H, Sato, G, Toriyabe, A, Kudo, H, Hashimoto, A, Matsumura, M, Harada, Y, Kurihara, Y, Shirouzu, T, 2015. Revision of the Massarineae (Pleosporales, Dothideomycetes). Studies in mycology 82: 75-136. ttps:// doi.org/10.1016/j.simyco.2015.10.002" title="Revision of the Massarineae (Pleosporales, Dothideomycetes). Studies in mycology 82: 75 - 136. ttps: //" url="doi.org/10.1016/j.simyco.2015.10.002" year="2015">Tanaka et al. (2015)</bibRefCitation>
have similar morphological features with our strain such as dark brown, 8-celled, disked-shaped, spiny conidia. With morphological and multigene phylogenetic support, we report a new host record of
<taxonomicName lsidName="S. deightonii" pageId="0" pageNumber="19" rank="species" species="deightonii">
<emphasis italics="true" pageId="0" pageNumber="19">S. deightonii</emphasis>
<taxonomicName id="37F41CEF725794571C08ED243300F992" lsidName="S. deightonii" pageId="0" pageNumber="19" rank="species" species="deightonii">
<emphasis id="04A2322ECCF403D8C7060FF64CAAB41B" italics="true" pageId="0" pageNumber="19">S. deightonii</emphasis>
</taxonomicName>
from
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<emphasis italics="true" pageId="0" pageNumber="19">Musa</emphasis>
<taxonomicName id="A3C6E441A6A00F1CBC233623DBDD8737" class="Liliopsida" family="Musaceae" genus="Musa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Musa" order="Zingiberales" pageId="0" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="0E22FBDBCDFBDC0A3CF3D163FB2FEC43" italics="true" pageId="0" pageNumber="19">Musa</emphasis>
</taxonomicName>
sp.
</paragraph>
<caption doi="10.3897/mycokeys.70.52043.figure2" httpUri="https://binary.pensoft.net/fig/434238" pageId="0" pageNumber="19" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">Figure 2.</emphasis>
<taxonomicName authorityName="Subram., J. Indian bot. Soc. 35: 78" authorityYear="1956" baseAuthorityName="S. Hughes" class="Sordariomycetes" family="Apiosporaceae" genus="Spegazzinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spegazzinia deightonii" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="deightonii">
<emphasis italics="true" pageId="0" pageNumber="19">Spegazzinia deightonii</emphasis>
<caption id="9DC8E4B46313B5ADFB68BA60732B29CE" doi="10.3897/mycokeys.70.52043.figure2" httpUri="https://binary.pensoft.net/fig/434238" pageId="0" pageNumber="19" start="Figure 2" startId="F2">
<paragraph id="4AC25DED3C12BB76C320C13AF75CE2FC" pageId="0" pageNumber="19">
<emphasis id="6468E0B62E7ED2B07995FAF0DB96CC91" bold="true" pageId="0" pageNumber="19">Figure 2.</emphasis>
<taxonomicName id="374C67FBC63E380C360016D90B8FEF47" authorityName="Subram., J. Indian bot. Soc. 35: 78" authorityYear="1956" baseAuthorityName="S. Hughes" class="Sordariomycetes" family="Apiosporaceae" genus="Spegazzinia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spegazzinia deightonii" pageId="0" pageNumber="19" phylum="Ascomycota" rank="species" species="deightonii">
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(MFLU 19-2908)
<emphasis bold="true" pageId="0" pageNumber="19">a-c</emphasis>
<emphasis id="A17B205A1BE17396AF9E152B02C61523" bold="true" pageId="0" pageNumber="19">a-c</emphasis>
fungal colonies on host surface
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<emphasis id="2D7B2EC01A4C2DEDDDBB30C128E7C2B5" bold="true" pageId="0" pageNumber="19">d</emphasis>
conidiophore mother cell of
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conidia
<emphasis bold="true" pageId="0" pageNumber="19">e-g</emphasis>
<normalizedToken originalValue="α">α</normalizedToken>
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a developmental stage of
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conidia
<emphasis bold="true" pageId="0" pageNumber="19">h, k</emphasis>
<emphasis id="DE494F66035338F97DB890FF32E5F4D9" bold="true" pageId="0" pageNumber="19">h, k</emphasis>
conidia
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colonies on PDA after 28 days showing sporulation
<emphasis bold="true" pageId="0" pageNumber="19">j, m-p</emphasis>
<normalizedToken originalValue="β">β</normalizedToken>
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conidia. Scale bars: 500μm (
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<emphasis id="BA0176A8E4308E3116C9FC05274C8CD3" bold="true" pageId="0" pageNumber="19">a</emphasis>
), 200μm (
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<emphasis id="6FCE0EB6612381494F200B67EB3D8ABF" bold="true" pageId="0" pageNumber="19">b</emphasis>
), 50
<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis id="E4FF9BA8AEE72CA32416C7E995A8A586" bold="true" pageId="0" pageNumber="19">c</emphasis>
), 20μm (
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<emphasis id="8A1A00DC539391FA41476B856CD3399A" bold="true" pageId="0" pageNumber="19">e-h</emphasis>
), 10μm (
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<emphasis id="3C5FF933F6A61CD168A7E64742D862B2" bold="true" pageId="0" pageNumber="19">d, k, m-p</emphasis>
), 5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="C608EE539BA26A5461DFF1866960BF98" originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="19">i, j</emphasis>
<emphasis id="EBA4FAB4D12CA3D0EBDD93658CD2499F" bold="true" pageId="0" pageNumber="19">i, j</emphasis>
).
</paragraph>
</caption>

View file

@ -1,151 +1,151 @@
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Figs 2, 15-20
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<paragraph pageId="6" pageNumber="7">
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<paragraph id="9BD9A67EF8FF294FFC9A79C1AD01FF5A" pageId="6" pageNumber="7">
<taxonomicName id="64FCA66BDE3BFF7D763FAE16E6B81EBA" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus caussaneli" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caussaneli">Cryptopygus caussaneli</taxonomicName>
Thibaud, 1996
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<paragraph pageId="6" pageNumber="7">
<taxonomicName class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus riebi" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="riebi">Cryptopygus riebi</taxonomicName>
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Barra, 1997 n. syn.
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<paragraph pageId="6" pageNumber="7">Material.</paragraph>
<paragraph pageId="6" pageNumber="7">
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<paragraph id="AB609CB7EFC4993F44178E9A48A9214F" pageId="6" pageNumber="7">Material.</paragraph>
<paragraph id="B271999C4F7201B58E15DD0891DC2173" pageId="6" pageNumber="7">
Holotype and paratypes of
<taxonomicName class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus caussaneli" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caussaneli">Cryptopygus caussaneli</taxonomicName>
<taxonomicName id="5CAC586C99A9253B7C512779775BEB68" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus caussaneli" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caussaneli">Cryptopygus caussaneli</taxonomicName>
: Mauritania (Aftout-es-Sahel); holotype and paratypes of
<taxonomicName class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus riebi" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="riebi">Cryptopygus riebi</taxonomicName>
<taxonomicName id="4A1285CBBC1B9BB0FD5E0984DE118D0E" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus riebi" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="riebi">Cryptopygus riebi</taxonomicName>
: South Africa (Sodwana Bay, Natal Province); two paratypes of
<taxonomicName class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus axayacatl" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="axayacatl">Cryptopygus axayacatl</taxonomicName>
<taxonomicName id="4069C4C645F85986C4C02772AFC6F4DE" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus axayacatl" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="axayacatl">Cryptopygus axayacatl</taxonomicName>
Palacios-Vargas &amp; Thibaud, 2001: Mexico (Guerrero, Acapulco); material of J.-M.Thibaud identified by him as
<taxonomicName class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus caussaneli" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caussaneli">Cryptopygus caussaneli</taxonomicName>
<taxonomicName id="F2AA8BBC1B8D806A591251734718E376" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus caussaneli" order="Entomobryomorpha" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caussaneli">Cryptopygus caussaneli</taxonomicName>
from Senegal, Morocco, Congo, Madagascar, Maurice and Mayotte Islands. More precise labels in the associated publications (
<bibRefCitation author="Palacios-Vargas, JG" journalOrPublisher="Revue francaise d'Entomologie" pageId="13" pageNumber="14" pagination="159 - 166" title="Three new species of Mexican littoral Collembola of genera Willemia, Cryptopygus and Isotogastrura (Hypogastruridae, Isotomidae, Isotogastruridae)." volume="23" year="2001">Palacios-Vargas &amp; Thibaud, 2001</bibRefCitation>
<bibRefCitation id="B3144D1A8BD0E4FF2F5C192207E7A0BB" author="Palacios-Vargas, JG" journalOrPublisher="Revue francaise d'Entomologie" pageId="13" pageNumber="14" pagination="159 - 166" title="Three new species of Mexican littoral Collembola of genera Willemia, Cryptopygus and Isotogastrura (Hypogastruridae, Isotomidae, Isotogastruridae)." volume="23" year="2001">Palacios-Vargas &amp; Thibaud, 2001</bibRefCitation>
;
<bibRefCitation author="Thibaud, JM" journalOrPublisher="Revue francaise d'Entomologie" pageId="14" pageNumber="15" pagination="41 - 48" title="Collemboles interstitiels des sables littoraux du Senegal." volume="28" year="2006">Thibaud &amp; Ndiaye, 2006</bibRefCitation>
<bibRefCitation id="250488ED2CEBB6A8889C166229308196" author="Thibaud, JM" journalOrPublisher="Revue francaise d'Entomologie" pageId="14" pageNumber="15" pagination="41 - 48" title="Collemboles interstitiels des sables littoraux du Senegal." volume="28" year="2006">Thibaud &amp; Ndiaye, 2006</bibRefCitation>
;
<bibRefCitation author="Thibaud, JM" journalOrPublisher="Revue francaise d'Entomologie" pageId="14" pageNumber="15" pagination="63 - 67" title="Collemboles interstitiels des sables littoraux du Maroc - II." volume="28" year="2006">Thibaud &amp; Boumezzough, 2006</bibRefCitation>
<bibRefCitation id="4378A44524B2F6700687588DF5648B3A" author="Thibaud, JM" journalOrPublisher="Revue francaise d'Entomologie" pageId="14" pageNumber="15" pagination="63 - 67" title="Collemboles interstitiels des sables littoraux du Maroc - II." volume="28" year="2006">Thibaud &amp; Boumezzough, 2006</bibRefCitation>
;
<bibRefCitation author="Thibaud, JM" journalOrPublisher="Annales de la Societe entomologique de France" pageId="14" pageNumber="15" pagination="503 - 519" title="Les Collemboles des sables littoraux de Madagascar." volume="44" year="2008">Thibaud, 2008</bibRefCitation>
<bibRefCitation id="5143A371C210A9FA2C78C0F2062EC109" author="Thibaud, JM" journalOrPublisher="Annales de la Societe entomologique de France" pageId="14" pageNumber="15" pagination="503 - 519" title="Les Collemboles des sables littoraux de Madagascar." volume="44" year="2008">Thibaud, 2008</bibRefCitation>
). All specimens are kept in the Museum national
<normalizedToken originalValue="dHistoire">d'Histoire</normalizedToken>
<normalizedToken id="442E724C9231D02BF7FCD2361D79E6AF" originalValue="dHistoire">d'Histoire</normalizedToken>
naturelle of Paris (France).
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph pageId="7" pageNumber="8">
<pageBreakToken pageId="7" pageNumber="8" start="start">Redescription</pageBreakToken>
<subSubSection id="87FE24078C6BF78D419943B540B0EA04" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="9AA633C92DF79832493C4BF09730DAD5" pageId="7" pageNumber="8">
<pageBreakToken id="60F1E159DE024CE80669C038D53C9169" pageId="7" pageNumber="8" start="start">Redescription</pageBreakToken>
.
</paragraph>
<paragraph pageId="7" pageNumber="8">
<paragraph id="4057A3C0302C43E33FA2646FFB41AF23" pageId="7" pageNumber="8">
White, without eyes. Size up to 0.4 mm. Cuticle smooth. PAO about 1,9 as long as inner edge of U.III and shorter (0.7) than width of Ant.I. Outer (Fig. 20) and inner mouthparts principally as in
<taxonomicName class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus projectus" order="Entomobryomorpha" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="projectus">Pauropygus projectus</taxonomicName>
<taxonomicName id="27CCF5CC6D9F9AEEB7D695E9EE991E9B" class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus projectus" order="Entomobryomorpha" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="projectus">Pauropygus projectus</taxonomicName>
. Ventral side of head with 5+5 postlabial chaetae. Ant.I with 14-15 chaetae, 1 ventro-basal microchaeta (bms; dorsal bms not differentiated), and 2 thick ventral sensilla (s), short and long. Ant.II with 3 bms and 1 thick laterodistal s. Ant.III without bms and with 5 distal s of which two inner thicker and longer than outer ones. Male antennal
<normalizedToken originalValue="“spurs”">&quot;spurs&quot;</normalizedToken>
<normalizedToken id="93FF0C2F84EA1C751CC90B89E3A54DC3" originalValue="“spurs”">&quot;spurs&quot;</normalizedToken>
present on Ant.II, III and basal part of Ant.IV.
</paragraph>
<paragraph pageId="7" pageNumber="8">
<paragraph id="3333C242DD2F6BD591909EE68306F1FB" pageId="7" pageNumber="8">
Dorsal axial chaetom of Th.
<normalizedToken originalValue="IIAbd">II-Abd</normalizedToken>
<normalizedToken id="E2C9BF74E14F5B0179310F40DAA6BD7B" originalValue="IIAbd">II-Abd</normalizedToken>
.III as 10,6/4,4(3),4(3). Thorax without ventral chaetae. Macrochaetae smooth, with the whole number as 1(2),1(2)/3,3,3,4 (Th.II-Abd.IV). Medial macrochaetae on Th.II-Abd.III hardly differentiated. Medial macrochaetae on Abd.V about 0.4 as long as dens. All sensilla of thorax and medial sensilla of abdomen nearly as long as ordinary chaetae and hardly visible, lateral sensilla on Abd.III-V shorter than ordinary chaetae. Microsensilla short but well visible. Sensillar formula 31/11221 (s), 00/100 (ms) (Fig. 15-18). Sensilla in posterior position. (Fig. 16).
</paragraph>
<paragraph pageId="8" pageNumber="9">
<pageBreakToken pageId="8" pageNumber="9" start="start">Unguis</pageBreakToken>
<paragraph id="4B817DE19F770FE411521E6522AAC6E7" pageId="8" pageNumber="9">
<pageBreakToken id="58950C3903E28782F428414499683CFE" pageId="8" pageNumber="9" start="start">Unguis</pageBreakToken>
of normal shape, without inner tooth, two broad unequal lateral teeth. Upper subcoxa of Leg I-III with 1,1,4 chaetae, lower subcoxa with 1,6-7,7-8. Ti.I-III with one chaeta lost in basic set (20, 21,&gt;25). Chaetae x and B5 on Ti. III in males unmodified. Distal tibiotarsal tenent chaetae on Ti.I-III (1-2-2) well developed, not clavate, about 1.1-1.2 as long as U.III. Each tibiotarsus with one additional tenent chaeta at middle part. Ti.III with one stick-like chaetae (A7) in distal ring, its shape slightly varies (Fig. 19). Tenent hairs (1,2,2) about 1.1-1.2 as long as U.III (Fig. 19). Pretarsus with two chaetae. Ventral tube with 4+4 laterodistal and 4 posterior chaetae in one transversal row. Tenaculum with 4+4 teeth and 1 chaeta. Anterior furcal subcoxa with 12-15 chaetae, posterior with 6 ones. Manubrium principally as in previous species. Dens slender, anteriorly normally with 11 chaetae. Posterior side of dens slightly crenulated in the medial part, with 5 chaetae of which 3 basal and 2 at the medial part (short inner and long outer) set together on low papillum. Mucro slender with two teeth of unequal size. Ratio of manubrium: dens: mucro = 3.2-3.6: 4.1-4.3: 1. Anal lobes without microchaetae.
</paragraph>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
<caption id="D9C6CC5F7BB232DACF6EC9B6C8C74BEF" pageId="8" pageNumber="9">
<paragraph id="FA206A648FF148932ECB2210493B861F" pageId="8" pageNumber="9">
Figures 15-18.
<taxonomicName class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus caussaneli" order="Entomobryomorpha" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="caussaneli">Pauropygus caussaneli</taxonomicName>
<taxonomicName id="755D52B37F5AB7AA655FF7E53DEC94E5" class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus caussaneli" order="Entomobryomorpha" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="caussaneli">Pauropygus caussaneli</taxonomicName>
. 15 sensillar chaetotaxy 16 posterior part of abdomen 17 lateral part of Abd.I 18 lateral part of Th.III. s sensillum, ms microsensillum.
</paragraph>
</caption>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
<caption id="E287E434693B615347251C7DDFEC7B5C" pageId="8" pageNumber="9">
<paragraph id="395DBE492A42D7F23F8CA79785E26CC4" pageId="8" pageNumber="9">
Figures 19-20.
<taxonomicName class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus caussaneli" order="Entomobryomorpha" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="caussaneli">Pauropygus caussaneli</taxonomicName>
<taxonomicName id="A75AC7A35042102E7A3C99892D2A437B" class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus caussaneli" order="Entomobryomorpha" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="caussaneli">Pauropygus caussaneli</taxonomicName>
. 19 distal half of Leg III 20 anterior part of head, lateral view (group of basolateral chaetae marked).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="affinity">
<paragraph pageId="9" pageNumber="10">
<pageBreakToken pageId="9" pageNumber="10" start="start">Affinity</pageBreakToken>
<subSubSection id="097FCAE2C9F0B10D131879AAC966E589" pageId="9" pageNumber="10" type="affinity">
<paragraph id="9FCCC612378DF02B8272AE16DF8C23C1" pageId="9" pageNumber="10">
<pageBreakToken id="CF6749F201892709BA6B89C83157E071" pageId="9" pageNumber="10" start="start">Affinity</pageBreakToken>
.
</paragraph>
<paragraph pageId="9" pageNumber="10">
<paragraph id="0881861D9A178E64AA944E017E418AAF" pageId="9" pageNumber="10">
See above for differences with
<taxonomicName class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus projectus" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="projectus">Pauropygus projectus</taxonomicName>
<taxonomicName id="24EBDEF88EB9CCAB1EBA9C089EDD3B2F" class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus projectus" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="projectus">Pauropygus projectus</taxonomicName>
sp. n. After our study of type material of
<taxonomicName class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus riebi" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="riebi">Cryptopygus riebi</taxonomicName>
<taxonomicName id="D4582A7D4EB844812DE5143F7FB6FFB9" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus riebi" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="riebi">Cryptopygus riebi</taxonomicName>
(South Africa) and two paratypes of
<taxonomicName class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus axayacatl" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="axayacatl">Cryptopygus axayacatl</taxonomicName>
<taxonomicName id="EB62F0D8CB9C505DAE941F84C0952227" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus axayacatl" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="axayacatl">Cryptopygus axayacatl</taxonomicName>
(Mexico) we concluded that these species only complement the wide distribution area of
<taxonomicName class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus caussaneli" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caussaneli">Pauropygus caussaneli</taxonomicName>
<taxonomicName id="AA324DDA588ED83165241FF5FD437802" class="Collembola" family="Isotomidae" genus="Pauropygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pauropygus caussaneli" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="caussaneli">Pauropygus caussaneli</taxonomicName>
. Holotype and other paratypes of
<taxonomicName class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus axayacatl" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="axayacatl">Cryptopygus axayacatl</taxonomicName>
<taxonomicName id="4FEA50386DAEEE31E16C85F504BA07A8" class="Collembola" family="Isotomidae" genus="Cryptopygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptopygus axayacatl" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="axayacatl">Cryptopygus axayacatl</taxonomicName>
support its species status, but in the genus
<taxonomicName class="Collembola" family="Isotomidae" genus="Proisotomodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Proisotomodes" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Proisotomodes</taxonomicName>
<taxonomicName id="8FBCF3AA34685FA64097869583D09014" class="Collembola" family="Isotomidae" genus="Proisotomodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Proisotomodes" order="Entomobryomorpha" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Proisotomodes</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="distribution">
<paragraph pageId="9" pageNumber="10">Distribution.</paragraph>
<paragraph pageId="9" pageNumber="10">Probably widely distributed on sandy seashores of tropics and subtro-pics. Recorded from the coast of Indian and Atlantic Oceans (Africa, Central America).</paragraph>
<subSubSection id="BCB412D401AD66D84821704AE84FC028" pageId="9" pageNumber="10" type="distribution">
<paragraph id="C13CEE09232FEB0D8F18DAF681852A9D" pageId="9" pageNumber="10">Distribution.</paragraph>
<paragraph id="374E23DEA488FB9AAA8930472349437A" pageId="9" pageNumber="10">Probably widely distributed on sandy seashores of tropics and subtro-pics. Recorded from the coast of Indian and Atlantic Oceans (Africa, Central America).</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,77 +1,77 @@
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sichuana Ahrens, 2000
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<taxonomicName class="Insecta" family="Melolonthidae" genus="Gastroserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastroserica sichuana" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="sichuana">Gastroserica sichuana</taxonomicName>
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<taxonomicName id="5CA96E9F4F37542E5B262F1DD1B71CDA" class="Insecta" family="Scarabaeidae" genus="Gastroserica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastroserica sichuana" order="Coleoptera" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="sichuana">Gastroserica sichuana</taxonomicName>
Ahrens, 2000: 95.
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<subSubSection pageId="9" pageNumber="32" type="material examined">
<paragraph pageId="9" pageNumber="32">Material examined.</paragraph>
<paragraph pageId="9" pageNumber="32">1♂&quot;Baoguosi, Mt. Emei Shan, Sichuan, 550-750m, 2.5.1957, Wang Zongyuan leg.&quot; (IZAS); 1♂&quot;Shiping, Fengdu, Sichuan, 610m, 3.6.1994, Li Wenzhu leg.&quot; (IZAS); 2♀♀&quot;Shiping, Fengdu, Sichuan, 610m, 2-3.6.1994, Zhang Youwei leg.&quot; (IZAS); 1♀&quot;Nanmuyuan, Changshou, Sichuan, 450m, 9.6.1994, Zhang Youwei leg.&quot; (IZAS); 3♀♀&quot;Mt. Emei Shan, Sichuan, 800-1000m, 9-10.5.1957, Wang Zongyuan leg.&quot; (IZAS); 1♀&quot;Qiaoting, Wanxian, Sichuan, 1300m, 27.6.1974, Han Yinheng leg.&quot; (IZAS).</paragraph>
<subSubSection id="AD2C0853BED2F209B6F5C233192004B9" pageId="9" pageNumber="32" type="material examined">
<paragraph id="431A9E2944E1DF202EFE3634F23C90E0" pageId="9" pageNumber="32">Material examined.</paragraph>
<paragraph id="E2E286C851EA0A14931037EABE65474B" pageId="9" pageNumber="32">1♂&quot;Baoguosi, Mt. Emei Shan, Sichuan, 550-750m, 2.5.1957, Wang Zongyuan leg.&quot; (IZAS); 1♂&quot;Shiping, Fengdu, Sichuan, 610m, 3.6.1994, Li Wenzhu leg.&quot; (IZAS); 2♀♀&quot;Shiping, Fengdu, Sichuan, 610m, 2-3.6.1994, Zhang Youwei leg.&quot; (IZAS); 1♀&quot;Nanmuyuan, Changshou, Sichuan, 450m, 9.6.1994, Zhang Youwei leg.&quot; (IZAS); 3♀♀&quot;Mt. Emei Shan, Sichuan, 800-1000m, 9-10.5.1957, Wang Zongyuan leg.&quot; (IZAS); 1♀&quot;Qiaoting, Wanxian, Sichuan, 1300m, 27.6.1974, Han Yinheng leg.&quot; (IZAS).</paragraph>
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<subSubSection pageId="9" pageNumber="32" type="distribution">
<paragraph pageId="9" pageNumber="32">Distribution.</paragraph>
<paragraph pageId="9" pageNumber="32">Fig. 5.</paragraph>
<subSubSection id="CAC38F4D7AE459DF6CB6325D48A7A92F" pageId="9" pageNumber="32" type="distribution">
<paragraph id="2882047E5A6502975F3FFAF19CE72F50" pageId="9" pageNumber="32">Distribution.</paragraph>
<paragraph id="9503B519630BD07FD838051F42CD331B" pageId="9" pageNumber="32">Fig. 5.</paragraph>
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Genus
<taxonomicName LSID="http://species-id.net/wiki/Coccus" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus" order="Hemiptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">Coccus Linnaeus, 1758</taxonomicName>
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<taxonomicName class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus" order="Hemiptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">Coccus</taxonomicName>
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<taxonomicName id="CA88383F7B336444318720B205D46E51" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus" order="Hemiptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="genus">Coccus</taxonomicName>
Linnaeus, 1758: 455-457. Type species:
<taxonomicName class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus hesperidum" order="Hemiptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="hesperidum">Coccus hesperidum</taxonomicName>
<taxonomicName id="9AE6ABFCDDCA9E6C41B8DEC6FCD5B4EB" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus hesperidum" order="Hemiptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="hesperidum">Coccus hesperidum</taxonomicName>
Linnaeus, 1758.
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<paragraph pageId="1" pageNumber="60">Generic diagnosis.</paragraph>
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<paragraph id="196317DB17964458BD06651BC5F53E93" pageId="1" pageNumber="60">Generic diagnosis.</paragraph>
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Adult female. Body oval, elongate or pyriform; usually flat, moderately convex, or nearly hemispherical. Dorsum. Derm membranous; gradually i
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<pageBreakToken id="BA2D97735F83B9E184828DE6FEB66033" pageId="2" pageNumber="61" start="start">ncreasing</pageBreakToken>
in thickness during ageing. Dorsal setae setose, spinose, clavate or cylindrical. Dorsal tubular ducts and dorsal submarginal tubercles present or absent. Preopercular pores present or absent, varying in number and shape. Anal plates together quadrate. Anal ring with 6 or 8 setae. Margin. Marginal setae simple to branched; stigmatic clefts deep or shallow, each with 2-8 stigmatic spines. Venter. Antennae 2-8 segmented. Legs well developed or reduced; with or without a tibio-tarsal articulatory sclerosis. Spiracular disc-pores with 5 loculi. Pregenital disc-pores mainly with 10 loculi, present around anal opening, or extending anteriorly, even as far as head. Ventral tubular ducts present or absent; when present located medially, or forming a sparse or dense submarginal band.
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<subSubSection pageId="2" pageNumber="61" type="key to adult female coccus known from china">
<paragraph pageId="2" pageNumber="61">
<subSubSection id="C09E19075888CADF123203E539C9DB11" pageId="2" pageNumber="61" type="key to adult female coccus known from china">
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Key to adult female
<taxonomicName class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus" order="Hemiptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Coccus</taxonomicName>
<taxonomicName id="37AADB32F5F7E0981FA8257DFA3DEB19" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus" order="Hemiptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">Coccus</taxonomicName>
known from China
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<taxonomicName class="Insecta" family="Coccidae" genus="Ceroplastes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceroplastes takanoi" order="Hemiptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="takanoi">Ceroplastes takanoi</taxonomicName>
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<taxonomicName id="DBA807102412DE33AB028803D4DA7E80" class="Insecta" family="Coccidae" genus="Ceroplastes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceroplastes takanoi" order="Hemiptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="takanoi">Ceroplastes takanoi</taxonomicName>
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<paragraph pageId="0" pageNumber="21">
<taxonomicName LSID="http://zoobank.org/0624CCB0-DC63-40F9-A7B7-7F8627B491BB" authority="Poyarkov, Gorin &amp; Scherz" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus" status="gen. nov.">Nanohyla Poyarkov, Gorin &amp; Scherz</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="21">gen. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Palmar views of hands (above) and thenar views of feet (below) of the representative Microhyla s. str. and Nanohyla gen. nov. species: N. annamensis (A, B), N. arboricola (C, D), M. minuta (E, F), and M. tetrix (G, H). Arrow indicates outer metatarsal tubercle. Not to scale. Line drawings by Valentina D. Kretova." figureDoi="10.3897/zse.97.57968.figure10" httpUri="https://binary.pensoft.net/fig/496424" pageId="0" pageNumber="21">Figs 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Members of the new genus Nanohyla gen. nov. in life (males): N. annectens from Genting Highlands, Pahang, Malaysia (A), N. annamensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (B), N. arboricola from Chu Yang Sin N. P., Dak Lak, Vietnam (C), N. hongiaoensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (D), N. marmorata from Kon Chu Rang N. R., Gia Lai, Vietnam (E), N. nanapollexa from Kon Plong, Kon Tum, Vietnam (F), N. perparva from Gunung Mulu, Sarawak, Malaysia (G), N. petrigena from Gunung Mulu, Sarawak, Malaysia (H), and N. pulchella from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (I). Insets show tympanic area of the each species; white arrow points at the tympanic rim of the external tympanum. Photos by Nikolay A. Poyarkov (A-S, D-F, I), Vu Dang Hoang Nguyen (D), and Indraneil Das (G, H)." figureDoi="10.3897/zse.97.57968.figure11" httpUri="https://binary.pensoft.net/fig/496425" pageId="0" pageNumber="21">, 11</figureCitation>
<treatment id="C444A1B69DF558E6B6CCBE2DA94E5A81" ID-GBIF-Taxon="182236432" LSID="urn:lsid:zoobank.org:act:0624CCB0-DC63-40F9-A7B7-7F8627B491BB" httpUri="http://treatment.plazi.org/id/C444A1B69DF558E6B6CCBE2DA94E5A81" lastPageNumber="21" pageId="0" pageNumber="21" scope_family="Microhylidae" scope_order="Anura" scope_subFamily="Microhylinae">
<subSubSection id="CFBFFB5B38180C6F2528C2FFF269F6DA" pageId="0" pageNumber="21" type="nomenclature">
<paragraph id="2B0D5F7A8B147087ECC82A481506CA45" pageId="0" pageNumber="21">
<taxonomicName id="7DA409442F8102BCA029B9BF225C75D1" LSID="http://zoobank.org/0624CCB0-DC63-40F9-A7B7-7F8627B491BB" authority="Poyarkov, Gorin &amp; Scherz" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus" status="gen. nov.">Nanohyla Poyarkov, Gorin &amp; Scherz</taxonomicName>
<taxonomicNameLabel id="12F95ADFADC24FD9389989FD9D989DCB" pageId="0" pageNumber="21">gen. nov.</taxonomicNameLabel>
<figureCitation id="27D69197E8426938860E8BFE3A3DEC74" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Palmar views of hands (above) and thenar views of feet (below) of the representative Microhyla s. str. and Nanohyla gen. nov. species: N. annamensis (A, B), N. arboricola (C, D), M. minuta (E, F), and M. tetrix (G, H). Arrow indicates outer metatarsal tubercle. Not to scale. Line drawings by Valentina D. Kretova." figureDoi="10.3897/zse.97.57968.figure10" httpUri="https://binary.pensoft.net/fig/496424" pageId="0" pageNumber="21">Figs 10</figureCitation>
<figureCitation id="C61573D19E93EC59129D9674F2893F49" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Members of the new genus Nanohyla gen. nov. in life (males): N. annectens from Genting Highlands, Pahang, Malaysia (A), N. annamensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (B), N. arboricola from Chu Yang Sin N. P., Dak Lak, Vietnam (C), N. hongiaoensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (D), N. marmorata from Kon Chu Rang N. R., Gia Lai, Vietnam (E), N. nanapollexa from Kon Plong, Kon Tum, Vietnam (F), N. perparva from Gunung Mulu, Sarawak, Malaysia (G), N. petrigena from Gunung Mulu, Sarawak, Malaysia (H), and N. pulchella from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (I). Insets show tympanic area of the each species; white arrow points at the tympanic rim of the external tympanum. Photos by Nikolay A. Poyarkov (A-S, D-F, I), Vu Dang Hoang Nguyen (D), and Indraneil Das (G, H)." figureDoi="10.3897/zse.97.57968.figure11" httpUri="https://binary.pensoft.net/fig/496425" pageId="0" pageNumber="21">, 11</figureCitation>
; Suppl. material 5: Table S5
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="chresonymy">
<paragraph pageId="0" pageNumber="21">Chresonymy.</paragraph>
<paragraph pageId="0" pageNumber="21">
<taxonomicName authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
<subSubSection id="DC03D5CA4839622A64AEA0BEE913C937" pageId="0" pageNumber="21" type="chresonymy">
<paragraph id="36B3A97F64033D8426DEFB0E313DFFAD" pageId="0" pageNumber="21">Chresonymy.</paragraph>
<paragraph id="88C8ADAD0DAA8E001E00D5BF7C7E1D85" pageId="0" pageNumber="21">
<taxonomicName id="D035684420E99717E054232DE1A6CDAE" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="7B1792377BF6F2FF817B39C4ADA2888E" italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
</taxonomicName>
(partim)-
<bibRefCitation DOI="https://doi.org/10.1080/00222930008678356" author="Boulenger, GA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="21" refId="B19" refString="Boulenger, GA, 1900. Descriptions of new batrachians and reptiles from the Larut Hills, Perak. Annals and Magazine of Natural History, Series 7 6: 186-193. https://doi.org/10.1080/00222930008678356" title="Descriptions of new batrachians and reptiles from the Larut Hills, Perak. Annals and Magazine of Natural History, Series 7 6: 186 - 193." url="https://doi.org/10.1080/00222930008678356" year="1900">Boulenger 1900</bibRefCitation>
<bibRefCitation id="F03ED9E3E226C4FFCE0C801DFEF7DB83" DOI="https://doi.org/10.1080/00222930008678356" author="Boulenger, GA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="21" refId="B19" refString="Boulenger, GA, 1900. Descriptions of new batrachians and reptiles from the Larut Hills, Perak. Annals and Magazine of Natural History, Series 7 6: 186-193. https://doi.org/10.1080/00222930008678356" title="Descriptions of new batrachians and reptiles from the Larut Hills, Perak. Annals and Magazine of Natural History, Series 7 6: 186 - 193." url="https://doi.org/10.1080/00222930008678356" year="1900">Boulenger 1900</bibRefCitation>
;
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<bibRefCitation id="85DABC55E6EF026E6D04B0E9E8E7B8A8" author="Smith, M" journalOrPublisher="Journal of the Natural History Society of Siam" pageId="0" pageNumber="21" pagination="47 - 53" refId="B141" refString="Smith, M, 1923. Notes on reptiles and batrachians from Siam and Indo-China (No. 2). Journal of the Natural History Society of Siam 6 (1): 47 - 53" title="Notes on reptiles and batrachians from Siam and Indo-China (No. 2)." volume="6" year="1923">Smith 1923</bibRefCitation>
;
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;
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<bibRefCitation id="C27B5BE4BD36350A4A3DB81F97F650B9" author="Inger, RF" journalOrPublisher="Malayan Nature Journal, Kuala Lumpur" pageId="0" pageNumber="21" pagination="229 - 243" refId="B75" refString="Inger, RF, 1989. Four new species of frogs from Borneo. Malayan Nature Journal, Kuala Lumpur 42: 229 - 243" title="Four new species of frogs from Borneo." volume="42" year="1989">Inger 1989</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen 2004</bibRefCitation>
<bibRefCitation id="AB7C7AB81C7FE6DCC68E5EDA3F194E40" DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen 2004</bibRefCitation>
;
<bibRefCitation author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
<bibRefCitation id="49DA4C9234F9C53170F4AC6C48F10F96" author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
;
<bibRefCitation author="Hoang, VC" journalOrPublisher="Asian Herpetological Research" pageId="0" pageNumber="21" pagination="1 - 22" refId="B69" refString="Hoang, VC, Nguyen, TT, Luong, MA, Nguyen, QT, Orlov, NL, Chen, Y, Wang, B, Jiang, J, 2020. A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam. Asian Herpetological Research 11 (3): 1 - 22" title="A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam." volume="11" year="2020">Hoang et al. 2020</bibRefCitation>
<bibRefCitation id="75E975B7E0A0AA942FA7E38280DA0736" author="Hoang, VC" journalOrPublisher="Asian Herpetological Research" pageId="0" pageNumber="21" pagination="1 - 22" refId="B69" refString="Hoang, VC, Nguyen, TT, Luong, MA, Nguyen, QT, Orlov, NL, Chen, Y, Wang, B, Jiang, J, 2020. A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam. Asian Herpetological Research 11 (3): 1 - 22" title="A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam." volume="11" year="2020">Hoang et al. 2020</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="21">
<taxonomicName authority="(Microhyla)" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla (Microhyla)" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="subGenus" subGenus="Microhyla">Microhyla (Microhyla)</taxonomicName>
<paragraph id="6D4DE587CF00AD97D2E4C4E905A84C5F" pageId="0" pageNumber="21">
<taxonomicName id="CD2378758B6967439F5163F2ED69BF0F" authority="(Microhyla)" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla (Microhyla)" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="subGenus" subGenus="Microhyla">Microhyla (Microhyla)</taxonomicName>
(partim)-
<bibRefCitation author="Dubois, A" journalOrPublisher="Alytes, Paris" pageId="0" pageNumber="21" pagination="1 - 9" refId="B37" refString="Dubois, A, 1987. Miscelanea taxinomica batrachologica, II. Alytes, Paris 6: 1 - 9" title="Miscelanea taxinomica batrachologica, II." volume="6" year="1987">Dubois 1987</bibRefCitation>
<bibRefCitation id="67D486B3A1134E9335BD52746EEEC350" author="Dubois, A" journalOrPublisher="Alytes, Paris" pageId="0" pageNumber="21" pagination="1 - 9" refId="B37" refString="Dubois, A, 1987. Miscelanea taxinomica batrachologica, II. Alytes, Paris 6: 1 - 9" title="Miscelanea taxinomica batrachologica, II." volume="6" year="1987">Dubois 1987</bibRefCitation>
(as a part of the subgenus
<taxonomicName authority="Microhyla" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla (Microhyla)" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="subGenus" subGenus="Microhyla">Microhyla Microhyla</taxonomicName>
<taxonomicName id="1924213F4FC26EFAD09F584549DFFA77" authority="Microhyla" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla (Microhyla)" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="subGenus" subGenus="Microhyla">Microhyla Microhyla</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="type species">
<paragraph pageId="0" pageNumber="21">Type species.</paragraph>
<paragraph pageId="0" pageNumber="21">
<taxonomicName authorityName="Boulenger" authorityYear="1900" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla annectens" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="annectens">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla annectens</emphasis>
<subSubSection id="7FDE68861A427ACDD38EE05B75B36CD3" pageId="0" pageNumber="21" type="type species">
<paragraph id="D1B2D8195FD40AFC1D52313F594E16F7" pageId="0" pageNumber="21">Type species.</paragraph>
<paragraph id="1416041C6FDBD28160F64CF874F3080A" pageId="0" pageNumber="21">
<taxonomicName id="2BD6D3CFFE260362DA65D30F342CAA77" authorityName="Boulenger" authorityYear="1900" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla annectens" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="annectens">
<emphasis id="C464409615AADA3165C2FDACA11D5C5E" italics="true" pageId="0" pageNumber="21">Microhyla annectens</emphasis>
</taxonomicName>
Boulenger, 1900.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="etymology">
<paragraph pageId="0" pageNumber="21">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="21">
<subSubSection id="61E187741BFE0BCF0ED2AEF34FC9E7C0" pageId="0" pageNumber="21" type="etymology">
<paragraph id="B57A4AD864239F992969203EE1F86389" pageId="0" pageNumber="21">Etymology.</paragraph>
<paragraph id="FD2D19B979FB566D9C97929BABD42A40" pageId="0" pageNumber="21">
The genus name is derived from the Greek
<emphasis italics="true" pageId="0" pageNumber="21">
<normalizedToken originalValue="νᾶνος">νᾶνος</normalizedToken>
<emphasis id="B71BF84311BBD13C3E6BD013C3397D39" italics="true" pageId="0" pageNumber="21">
<normalizedToken id="A111E600A55A4BB2F2A66E643AF95BA1" originalValue="νᾶνος">νᾶνος</normalizedToken>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="21">nanos</emphasis>
<emphasis id="D680C34C291988AB3AE53DB5368DE4F6" italics="true" pageId="0" pageNumber="21">nanos</emphasis>
), meaning
<normalizedToken originalValue="“dwarf”">&quot;dwarf&quot;</normalizedToken>
<normalizedToken id="749D855D2B5ABB75E29AF1B923EA71F7" originalValue="“dwarf”">&quot;dwarf&quot;</normalizedToken>
,
<normalizedToken originalValue="“pygmy”">&quot;pygmy&quot;</normalizedToken>
<normalizedToken id="34FB1DF6B0A6E65C286256F5D8CB5D5A" originalValue="“pygmy”">&quot;pygmy&quot;</normalizedToken>
, and the mythological figure, Hylas (Ancient Greek:
<emphasis italics="true" pageId="0" pageNumber="21">
<normalizedToken originalValue="Ὕλας">Ὕλας</normalizedToken>
<emphasis id="A5885031D76459C085AC7841B2432590" italics="true" pageId="0" pageNumber="21">
<normalizedToken id="76E46227E3D0753D49EE0CF62AEEA15F" originalValue="Ὕλας">Ὕλας</normalizedToken>
</emphasis>
), which is probably derived from the Ancient Greek verb &quot;
<emphasis italics="true" pageId="0" pageNumber="21">
<normalizedToken originalValue="ὕλαω">ὕλαω</normalizedToken>
<emphasis id="EE46304D371862C7B2587D918F344248" italics="true" pageId="0" pageNumber="21">
<normalizedToken id="EAD01F186933AFF51162D24BD531F5C9" originalValue="ὕλαω">ὕλαω</normalizedToken>
</emphasis>
&quot; meaning &quot;to bark&quot; (
<bibRefCitation author="Bourret, R" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="21" refId="B22" refString="Bourret, R, 1942. Les Batraciens de l'Indochine. Institut Oceanographique de l'Indochine, Hanoi (in French)." title="Les Batraciens de l'Indochine. Institut Oceanographique de l'Indochine, Hanoi (in French)." year="1942">Bourret 1942</bibRefCitation>
<bibRefCitation id="8FA93F1D04359798754EB06A17340EFC" author="Bourret, R" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="21" refId="B22" refString="Bourret, R, 1942. Les Batraciens de l'Indochine. Institut Oceanographique de l'Indochine, Hanoi (in French)." title="Les Batraciens de l'Indochine. Institut Oceanographique de l'Indochine, Hanoi (in French)." year="1942">Bourret 1942</bibRefCitation>
). In classical mythology, Hylas, son of King Theiodamas, was a youth who served as
<normalizedToken originalValue="Heracles">Heracles'</normalizedToken>
<normalizedToken id="A8C3D1AD01B1D5CC470AEE3D81C86868" originalValue="Heracles">Heracles'</normalizedToken>
companion, lover, and servant. Heracles took Hylas with him on the
<normalizedToken originalValue="Argonauts">Argonauts'</normalizedToken>
<normalizedToken id="7EE9CC7D45350F0F1A17BF63CF8BC091" originalValue="Argonauts">Argonauts'</normalizedToken>
expedition, during which Hylas was kidnapped by nymphs of the spring in Pegae, Mysia, and turned into an echo. Heracles left the ship and was searching for Hylas for a great length of time, calling his name: &quot;
<emphasis italics="true" pageId="0" pageNumber="21">His adjunxit Hylan nautae quo fonte relictum / Clamassent ut littus Hyla! Hyla! omne sonaret</emphasis>
<emphasis id="8FB73957F0DB7D977F8E4F96577E2B74" italics="true" pageId="0" pageNumber="21">His adjunxit Hylan nautae quo fonte relictum / Clamassent ut littus Hyla! Hyla! omne sonaret</emphasis>
&quot; (&quot;The mariners cried on Hylas till the shore / Then Re-echoed Hylas! Hylas! soothed...&quot;;
<bibRefCitation DOI="https://doi.org/10.4159/DLCL.virgil-eclogues.1916" author="Virgil" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" refId="B162" refString="Virgil, 1916. Eclogues. Georgics. Aeneid: Books 1-6. Translated by H. Rushton Fairclough. Revised by G. P. Goold. Loeb Classical Library 63. Cambridge, MA: Harvard University Press. https://doi.org/10.4159/DLCL.virgil-eclogues.1916" title="Eclogues. Georgics. Aeneid: Books 1 - 6. Translated by H. Rushton Fairclough. Revised by G. P. Goold. Loeb Classical Library 63. Cambridge, MA: Harvard University Press." url="https://doi.org/10.4159/DLCL.virgil-eclogues.1916" year="1916">Virgil 1916</bibRefCitation>
<bibRefCitation id="F0755A789FE271039FFD6C806AE609C7" DOI="https://doi.org/10.4159/DLCL.virgil-eclogues.1916" author="Virgil" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" refId="B162" refString="Virgil, 1916. Eclogues. Georgics. Aeneid: Books 1-6. Translated by H. Rushton Fairclough. Revised by G. P. Goold. Loeb Classical Library 63. Cambridge, MA: Harvard University Press. https://doi.org/10.4159/DLCL.virgil-eclogues.1916" title="Eclogues. Georgics. Aeneid: Books 1 - 6. Translated by H. Rushton Fairclough. Revised by G. P. Goold. Loeb Classical Library 63. Cambridge, MA: Harvard University Press." url="https://doi.org/10.4159/DLCL.virgil-eclogues.1916" year="1916">Virgil 1916</bibRefCitation>
, Ecl. 6, 43). The genus name refers to the small body size (&lt;25 mm) of all known
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="E9F18DB91C3D1C8AC186C4CA80C66D04" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="092965A0AB81AE71150208ADBFDDCE55" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
species, while maintaining resemblance to its sister genus
<taxonomicName authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
<taxonomicName id="CC7BCDB8A4815CF526558414E693CA86" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="641C7528B46430446BBB12E3348DDF98" italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
</taxonomicName>
, from which it is separated herein. The new genus name is feminine in gender.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="suggested common name">
<paragraph pageId="0" pageNumber="21">Suggested common name.</paragraph>
<paragraph pageId="0" pageNumber="21">Pygmy Narrow-mouthed Frogs.</paragraph>
<subSubSection id="7CB14EA8E5E3C9AD9FE0190CC352AEC9" pageId="0" pageNumber="21" type="suggested common name">
<paragraph id="D6700E5CD58339FB46A3A63823D61C14" pageId="0" pageNumber="21">Suggested common name.</paragraph>
<paragraph id="77D07CB7D55A49AED40F4F737E3B6E46" pageId="0" pageNumber="21">Pygmy Narrow-mouthed Frogs.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="taxonomic content">
<paragraph pageId="0" pageNumber="21">Taxonomic content.</paragraph>
<paragraph pageId="0" pageNumber="21">
<subSubSection id="7F8581A42EA1A71954AD3392A6575923" pageId="0" pageNumber="21" type="taxonomic content">
<paragraph id="DA97BEB5783D0E1565482E278B18E2E8" pageId="0" pageNumber="21">Taxonomic content.</paragraph>
<paragraph id="952F719D51A8C90B71A9A39DFB720E6D" pageId="0" pageNumber="21">
Nine species, including:
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" baseAuthorityName="Smith" baseAuthorityYear="1923" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla annamensis" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="annamensis">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla annamensis</emphasis>
<taxonomicName id="507B3DF81F1215F33DF404D1C483A9FC" baseAuthorityName="Smith" baseAuthorityYear="1923" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla annamensis" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="annamensis">
<emphasis id="FF9CD0E85987C9E398FE7AABF840B60A" italics="true" pageId="0" pageNumber="21">Nanohyla annamensis</emphasis>
</taxonomicName>
comb. nov. (Smith, 1923);
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" baseAuthorityName="Boulenger" baseAuthorityYear="1900" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla annectens" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="annectens">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla annectens</emphasis>
<taxonomicName id="4FE11FAE1B3D219034E5CC9AD2E4DF98" baseAuthorityName="Boulenger" baseAuthorityYear="1900" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla annectens" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="annectens">
<emphasis id="E22289C8800483E06F10037BB9F35207" italics="true" pageId="0" pageNumber="21">Nanohyla annectens</emphasis>
</taxonomicName>
comb. nov. (Boulenger, 1900);
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla arboricola" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="arboricola">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla arboricola</emphasis>
<taxonomicName id="152DC36485B4DC6D019731D641C70BE2" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla arboricola" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="arboricola">
<emphasis id="5D6A2C0E5F88DB0E3269302772F179F2" italics="true" pageId="0" pageNumber="21">Nanohyla arboricola</emphasis>
</taxonomicName>
comb. nov. (Poyarkov, Vassilieva, Orlov, Galoyan, Tran, Le, Kretova &amp; Geissler, 2014);
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" baseAuthorityName="Hoang, Nguyen, Luong, Nguyen, Chen, Wang &amp; Jiang" baseAuthorityYear="2020" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla hongiaoensis" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="hongiaoensis">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla hongiaoensis</emphasis>
<taxonomicName id="4E6C3B4A87BFB5CFAC16350FAFE0E9F5" baseAuthorityName="Hoang, Nguyen, Luong, Nguyen, Chen, Wang &amp; Jiang" baseAuthorityYear="2020" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla hongiaoensis" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="hongiaoensis">
<emphasis id="863132B9B3CE7111F57087274C7D25B3" italics="true" pageId="0" pageNumber="21">Nanohyla hongiaoensis</emphasis>
</taxonomicName>
comb. nov. (Hoang, Nguyen, Luong, Nguyen, Orlov, Chen, Wang &amp; Jiang, 2020);
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla marmorata" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="marmorata">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla marmorata</emphasis>
<taxonomicName id="847784697DE5791911AF70605CF43911" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla marmorata" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="marmorata">
<emphasis id="BFDA8D1883D13BE0C5E6E3CD71ED2B23" italics="true" pageId="0" pageNumber="21">Nanohyla marmorata</emphasis>
</taxonomicName>
comb. nov. (Bain &amp; Nguyen, 2004);
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" baseAuthorityName="Bain &amp; Truong" baseAuthorityYear="2004" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla nanapollexa" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="nanapollexa">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla nanapollexa</emphasis>
<taxonomicName id="E02568F1CD68022E8B46945895761C99" baseAuthorityName="Bain &amp; Truong" baseAuthorityYear="2004" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla nanapollexa" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="nanapollexa">
<emphasis id="1563E9D31114B0DF62F327EF7202C0C7" italics="true" pageId="0" pageNumber="21">Nanohyla nanapollexa</emphasis>
</taxonomicName>
comb. nov. (Bain &amp; Nguyen, 2004);
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" baseAuthorityName="Inger &amp; Frogner" baseAuthorityYear="1979" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla petrigena" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="petrigena">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla petrigena</emphasis>
<taxonomicName id="707F4F7DA2CC26C424A9A1EA972C0107" baseAuthorityName="Inger &amp; Frogner" baseAuthorityYear="1979" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla petrigena" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="petrigena">
<emphasis id="F3544EF68FDF84F8643C0A93F951E588" italics="true" pageId="0" pageNumber="21">Nanohyla petrigena</emphasis>
</taxonomicName>
comb. nov. (Inger &amp; Frogner, 1979);
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" baseAuthorityName="Inger &amp; Frogner" baseAuthorityYear="1979" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla perparva" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="perparva">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla perparva</emphasis>
<taxonomicName id="71ED7764195354C1E4DD61556ECD85EF" baseAuthorityName="Inger &amp; Frogner" baseAuthorityYear="1979" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla perparva" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="perparva">
<emphasis id="C2B47EFB288281B751F4839A659E2D6B" italics="true" pageId="0" pageNumber="21">Nanohyla perparva</emphasis>
</taxonomicName>
comb. nov. (Inger &amp; Frogner, 1979); and
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla pulchella" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="pulchella">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla pulchella</emphasis>
<taxonomicName id="166B21FD4B856596A775960D4EAAECEF" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla pulchella" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="pulchella">
<emphasis id="E4F7AAD5C19BEF632DEC5465B248734F" italics="true" pageId="0" pageNumber="21">Nanohyla pulchella</emphasis>
</taxonomicName>
comb. nov. (Poyarkov, Vassilieva, Orlov, Galoyan, Tran, Le, Kretova &amp; Geissler, 2014). Photos of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="4EBD28EBCCAE269406E16FA5DE478DB0" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="90F3F0F813511418EDA72D853DA0EFA6" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. members are presented in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Members of the new genus Nanohyla gen. nov. in life (males): N. annectens from Genting Highlands, Pahang, Malaysia (A), N. annamensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (B), N. arboricola from Chu Yang Sin N. P., Dak Lak, Vietnam (C), N. hongiaoensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (D), N. marmorata from Kon Chu Rang N. R., Gia Lai, Vietnam (E), N. nanapollexa from Kon Plong, Kon Tum, Vietnam (F), N. perparva from Gunung Mulu, Sarawak, Malaysia (G), N. petrigena from Gunung Mulu, Sarawak, Malaysia (H), and N. pulchella from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (I). Insets show tympanic area of the each species; white arrow points at the tympanic rim of the external tympanum. Photos by Nikolay A. Poyarkov (A-S, D-F, I), Vu Dang Hoang Nguyen (D), and Indraneil Das (G, H)." figureDoi="10.3897/zse.97.57968.figure11" httpUri="https://binary.pensoft.net/fig/496425" pageId="0" pageNumber="21">11</figureCitation>
<figureCitation id="21C99ECE23D371F02CEBF051693947A0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Members of the new genus Nanohyla gen. nov. in life (males): N. annectens from Genting Highlands, Pahang, Malaysia (A), N. annamensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (B), N. arboricola from Chu Yang Sin N. P., Dak Lak, Vietnam (C), N. hongiaoensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (D), N. marmorata from Kon Chu Rang N. R., Gia Lai, Vietnam (E), N. nanapollexa from Kon Plong, Kon Tum, Vietnam (F), N. perparva from Gunung Mulu, Sarawak, Malaysia (G), N. petrigena from Gunung Mulu, Sarawak, Malaysia (H), and N. pulchella from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (I). Insets show tympanic area of the each species; white arrow points at the tympanic rim of the external tympanum. Photos by Nikolay A. Poyarkov (A-S, D-F, I), Vu Dang Hoang Nguyen (D), and Indraneil Das (G, H)." figureDoi="10.3897/zse.97.57968.figure11" httpUri="https://binary.pensoft.net/fig/496425" pageId="0" pageNumber="21">11</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="diagnosis">
<paragraph pageId="0" pageNumber="21">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="21">
<subSubSection id="10D32964839D47E1656CF160E752342D" pageId="0" pageNumber="21" type="diagnosis">
<paragraph id="149C816B0B62164ED7C2B7BB0304CEDF" pageId="0" pageNumber="21">Diagnosis.</paragraph>
<paragraph id="35DE7857EA7B12B756B948B831AF9F64" pageId="0" pageNumber="21">
The new genus is assigned to the subfamily
<taxonomicName authorityName="Günther" authorityYear="1858" lsidName="" pageId="0" pageNumber="21" rank="subFamily" subFamily="Microhylinae">Microhylinae</taxonomicName>
<taxonomicName id="42CA09451071BF13F2B926D4A362BF66" authorityName="Günther" authorityYear="1858" lsidName="" pageId="0" pageNumber="21" rank="subFamily" subFamily="Microhylinae">Microhylinae</taxonomicName>
on the basis of phylogenetic affinities and the following combination of morphological character states: vomers small, confined to the anterior and medial margins of choanae; clavicles and, in most cases, procoracoids absent, maxillary arcade edentate (
<bibRefCitation author="Parker, HW" journalOrPublisher="Trustees of the British Museum, London" pageId="0" pageNumber="21" refId="B111" refString="Parker, HW, 1934. Monograph of the Frogs of the Family Microhylidae. Trustees of the British Museum, London" title="Monograph of the Frogs of the Family Microhylidae." year="1934">Parker 1934</bibRefCitation>
<bibRefCitation id="B5EC2C971CFED6EBB53B3CE0A1BD2423" author="Parker, HW" journalOrPublisher="Trustees of the British Museum, London" pageId="0" pageNumber="21" refId="B111" refString="Parker, HW, 1934. Monograph of the Frogs of the Family Microhylidae. Trustees of the British Museum, London" title="Monograph of the Frogs of the Family Microhylidae." year="1934">Parker 1934</bibRefCitation>
).
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="3B0447CE0B015440FD202D22E5DBA030" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="C54D60846B94592018A17BA49851CD13" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. differs from other
<taxonomicName authorityName="Günther" authorityYear="1858" lsidName="" pageId="0" pageNumber="21" rank="subFamily" subFamily="Microhylinae">Microhylinae</taxonomicName>
<taxonomicName id="A0ED14B4722CD5861E41FA82D96B911C" authorityName="Günther" authorityYear="1858" lsidName="" pageId="0" pageNumber="21" rank="subFamily" subFamily="Microhylinae">Microhylinae</taxonomicName>
genera by the following combination of osteological character states: (1) frontoparietals fused with exoccipitals; (2) exoccipitals fused with each other (incomplete fusion in
<taxonomicName genus="N." lsidName="N. pulchella" pageId="0" pageNumber="21" rank="species" species="pulchella">
<emphasis italics="true" pageId="0" pageNumber="21">N. pulchella</emphasis>
<taxonomicName id="4020F863C43BAD4919AB24ED25855206" lsidName="N. pulchella" pageId="0" pageNumber="21" rank="species" species="pulchella">
<emphasis id="A8C70550A150AA5D004F7E1F1B799749" italics="true" pageId="0" pageNumber="21">N. pulchella</emphasis>
</taxonomicName>
); (3) neopalatines present; (4) sphenethmoids completely fused with parasphenoid (incomplete fusion in
<taxonomicName genus="N." lsidName="N. pulchella" pageId="0" pageNumber="21" rank="species" species="pulchella">
<emphasis italics="true" pageId="0" pageNumber="21">N. pulchella</emphasis>
<taxonomicName id="27C3A64053BD4C0DFF18C5CBBFB8E51A" lsidName="N. pulchella" pageId="0" pageNumber="21" rank="species" species="pulchella">
<emphasis id="66EA762CF81EDBA66825B7CEDC0257C5" italics="true" pageId="0" pageNumber="21">N. pulchella</emphasis>
</taxonomicName>
); (5) crista parotica entirely cartilaginous; (6) otic ramus of squamosal well-developed; (7) tympanic annulus well-developed; (8) transverse processes of presacral vertebrae with the following orientation: IV and V posterolaterally, II, VII and VIII anterolaterally, III and VI at right angle to body axis; (9) clavicles absent; (10) omosternum present, cartilaginous; (11) prehallux cartilaginous; (12) terminal phalanges of longest fingers and toes T-shaped. The combination of diagnostic external morphological characters includes: (13) small to extremely small frogs (adult SVL 11.8-25.8 mm); (14) snout rounded or pointed in profile; (15) supratympanic fold present; (16) ridge on posterior sides of choanae absent; (17) first finger (FI) length less than
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="DF5D63B91BE81CEB0E29F083B030B398" originalValue="½">1/2</normalizedToken>
FII or reduced to a nub; (18) finger discs present, at least on FII-FIV; (19) dorsal median longitudinal grooves on finger discs generally present (with the exception of
<taxonomicName genus="N." lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
<taxonomicName id="761150F841D03CADD8DCB442E8415CF8" lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis id="CB7DF60576173F8C90010BB7854CDA02" italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
</taxonomicName>
); (20) toes dorsolaterally flattened, prominent discs present; (21) dorsal median longitudinal grooves on toe discs present; (22) metatarsal tubercle single (inner metatarsal tubercle present, outer absent); (23) dorsomedial line absent; (24) superciliary tubercles absent; (25) tibiotarsal articulation of adpressed hindlimb reaching well beyond snout; (26) toe webbing well-developed (at least one-half webbed); (27) skin on dorsum feebly granular to tubercular; (28) tympanum externally distinct at least in males (
<taxonomicName genus="N." lsidName="N. annamensis" pageId="0" pageNumber="21" rank="species" species="annamensis">
<emphasis italics="true" pageId="0" pageNumber="21">N. annamensis</emphasis>
<taxonomicName id="3600BEA510B280F0B44DCDB77CED216B" lsidName="N. annamensis" pageId="0" pageNumber="21" rank="species" species="annamensis">
<emphasis id="277B528B17DCFB8B4B942C548D65E022" italics="true" pageId="0" pageNumber="21">N. annamensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. annectens" pageId="0" pageNumber="21" rank="species" species="annectens">
<emphasis italics="true" pageId="0" pageNumber="21">N. annectens</emphasis>
<taxonomicName id="D3786F32F23905444702200A6D90E19F" lsidName="N. annectens" pageId="0" pageNumber="21" rank="species" species="annectens">
<emphasis id="077B4347ED02090640B737798DB03F80" italics="true" pageId="0" pageNumber="21">N. annectens</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
<taxonomicName id="6147D067EC1D5CD361E51444A268B7DB" lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis id="B5638E633E14302092371DBF87086F2C" italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
<taxonomicName id="C00B3EF1453592A255483640BB7DB23D" lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis id="7B1B5A74DB9577975BAC06EEDEE4DB6A" italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. nanapollexa" pageId="0" pageNumber="21" rank="species" species="nanapollexa">
<emphasis italics="true" pageId="0" pageNumber="21">N. nanapollexa</emphasis>
<taxonomicName id="262E546FBBBD8FAAC5F1195129044076" lsidName="N. nanapollexa" pageId="0" pageNumber="21" rank="species" species="nanapollexa">
<emphasis id="263B2D441603899E7E49336695767E24" italics="true" pageId="0" pageNumber="21">N. nanapollexa</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. pulchella" pageId="0" pageNumber="21" rank="species" species="pulchella">
<emphasis italics="true" pageId="0" pageNumber="21">N. pulchella</emphasis>
<taxonomicName id="88D20278049395EE7334D9F41CC3067D" lsidName="N. pulchella" pageId="0" pageNumber="21" rank="species" species="pulchella">
<emphasis id="4CC3179FDED98DB73EACDA8A34D13C69" italics="true" pageId="0" pageNumber="21">N. pulchella</emphasis>
</taxonomicName>
) or barely distinct (
<taxonomicName genus="N." lsidName="N. hongiaoensis" pageId="0" pageNumber="21" rank="species" species="hongiaoensis">
<emphasis italics="true" pageId="0" pageNumber="21">N. hongiaoensis</emphasis>
<taxonomicName id="65C2FAA71380509D6DFBBC70D46E9450" lsidName="N. hongiaoensis" pageId="0" pageNumber="21" rank="species" species="hongiaoensis">
<emphasis id="D2C98E17928FF78A33B58629F90788E7" italics="true" pageId="0" pageNumber="21">N. hongiaoensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
<taxonomicName id="6E69E3A6AC71F92140E36889BD1082AC" lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis id="31296ED4EB9DCAC506526AEBB14DCB62" italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. petrigena" pageId="0" pageNumber="21" rank="species" species="petrigena">
<emphasis italics="true" pageId="0" pageNumber="21">N. petrigena</emphasis>
<taxonomicName id="93C1DB36BDC2114489454225EF6574C4" lsidName="N. petrigena" pageId="0" pageNumber="21" rank="species" species="petrigena">
<emphasis id="FD3CB4599E81D3F082563DB7DA1D0D30" italics="true" pageId="0" pageNumber="21">N. petrigena</emphasis>
</taxonomicName>
); (29) terrestrial or scansorial semi-arboreal microhabitat preference.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="phylogenetic definition">
<paragraph pageId="0" pageNumber="21">Phylogenetic definition.</paragraph>
<paragraph pageId="0" pageNumber="21">
<subSubSection id="318955541EB8564C7848061D70566AB5" pageId="0" pageNumber="21" type="phylogenetic definition">
<paragraph id="F344C53C03793243814C5EF685437EFF" pageId="0" pageNumber="21">Phylogenetic definition.</paragraph>
<paragraph id="0E37C02F6C320AE2EC76CACEEB5DFD1D" pageId="0" pageNumber="21">
The genus
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="B774D1610CCAAEA7F0376DC6FD61C9D8" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="C098001193EF18E634A58BA9BE6C8F5E" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. includes all species sharing a more recent common ancestor with
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" baseAuthorityName="Boulenger" baseAuthorityYear="1900" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla annectens" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="annectens">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla annectens</emphasis>
<taxonomicName id="188C75CDF379DB355DCD9A5DE8562EC2" baseAuthorityName="Boulenger" baseAuthorityYear="1900" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla annectens" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="annectens">
<emphasis id="DAA184DFBE01CE3CF8789036277C2280" italics="true" pageId="0" pageNumber="21">Nanohyla annectens</emphasis>
</taxonomicName>
than with
<taxonomicName authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla achatina" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="achatina">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla achatina</emphasis>
<taxonomicName id="85CD988BF0E796318259E6C1D90EB788" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla achatina" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="achatina">
<emphasis id="EA4D3D0945EA6067E1AA7B171D5CBDB3" italics="true" pageId="0" pageNumber="21">Microhyla achatina</emphasis>
</taxonomicName>
and
<taxonomicName class="Amphibia" family="Microhylidae" genus="Glyphoglossus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphoglossus molossus" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="molossus">
<emphasis italics="true" pageId="0" pageNumber="21">Glyphoglossus molossus</emphasis>
<taxonomicName id="AEFC66C446B6E24F0B0E9523168B746F" class="Amphibia" family="Microhylidae" genus="Glyphoglossus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphoglossus molossus" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="molossus">
<emphasis id="5BC664EBBE5DB56827D4F90963E05690" italics="true" pageId="0" pageNumber="21">Glyphoglossus molossus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="distribution">
<paragraph pageId="0" pageNumber="21">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="21">
<subSubSection id="964538D7FAB4D62E4A02C01181CAB486" pageId="0" pageNumber="21" type="distribution">
<paragraph id="D9DC199FCB16BF4C59358BF8AD7A798E" pageId="0" pageNumber="21">Distribution.</paragraph>
<paragraph id="568291CF46672ECDCE581201241A68B5" pageId="0" pageNumber="21">
The distribution area of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="A7E5BC7C7DBDC1ED95DAC07F2A5F3B8D" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="9D8BF279A0CDE24E203A2877245B4381" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. covers montane forests of the Annamite (Truong Son) Mountains in Vietnam, eastern Laos, and north-eastern Cambodia, the Titiwangsa Mountain Range in the southernmost Thailand and peninsular Malaysia, mountains of Borneo (including Sabah and Sarawak of Malaysia, Brunei, and Kalimantan of Indonesia) and the Sulu Archipelago of the Philippines (see Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution ranges of the three clades of the Microhyla - Glyphoglossus assemblage. Distribution area of Microhyla I is shown in yellow, of Microhyla II in red, and of Glyphoglossus in blue. Distributional data from Gorin et al. (2020). Question mark denotes the unconfirmed record of &quot; Microhyla annamensis &quot; from Khao Sebab in eastern Thailand by Taylor (1962)." figureDoi="10.3897/zse.97.57968.figure1" httpUri="https://binary.pensoft.net/fig/496415" pageId="0" pageNumber="21">1</figureCitation>
<figureCitation id="6E832DB02C06C1BBA3F1EC852662B135" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution ranges of the three clades of the Microhyla - Glyphoglossus assemblage. Distribution area of Microhyla I is shown in yellow, of Microhyla II in red, and of Glyphoglossus in blue. Distributional data from Gorin et al. (2020). Question mark denotes the unconfirmed record of &quot; Microhyla annamensis &quot; from Khao Sebab in eastern Thailand by Taylor (1962)." figureDoi="10.3897/zse.97.57968.figure1" httpUri="https://binary.pensoft.net/fig/496415" pageId="0" pageNumber="21">1</figureCitation>
). The occurrence of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="A4D12F327934047807EA53E573084E22" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="9C80473CC1D186E27A64FACB0D3A4983" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. in Cardamom Mountains in eastern Thailand (the record of &quot;
<taxonomicName genus="M." lsidName="M. annamensis" pageId="0" pageNumber="21" rank="species" species="annamensis">
<emphasis italics="true" pageId="0" pageNumber="21">M. annamensis</emphasis>
<taxonomicName id="D3858EA92DFF2456423780743326E6FE" lsidName="M. annamensis" pageId="0" pageNumber="21" rank="species" species="annamensis">
<emphasis id="05B1D054329BA616B5E191169498C76F" italics="true" pageId="0" pageNumber="21">M. annamensis</emphasis>
</taxonomicName>
&quot; from Khao Sebab by
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.13347" author="Taylor, EH" journalOrPublisher="University of Kansas Science Bulletin" pageId="0" pageNumber="21" pagination="265 - 599" refId="B149" refString="Taylor, EH, 1962. The amphibian fauna of Thailand. University of Kansas Science Bulletin 43: 265 - 599, DOI: https://doi.org/10.5962/bhl.part.13347" title="The amphibian fauna of Thailand." url="https://doi.org/10.5962/bhl.part.13347" volume="43" year="1962">Taylor [1962]</bibRefCitation>
<bibRefCitation id="F101E209B2111F7AB3DFCB82D799AD4C" DOI="https://doi.org/10.5962/bhl.part.13347" author="Taylor, EH" journalOrPublisher="University of Kansas Science Bulletin" pageId="0" pageNumber="21" pagination="265 - 599" refId="B149" refString="Taylor, EH, 1962. The amphibian fauna of Thailand. University of Kansas Science Bulletin 43: 265 - 599, DOI: https://doi.org/10.5962/bhl.part.13347" title="The amphibian fauna of Thailand." url="https://doi.org/10.5962/bhl.part.13347" volume="43" year="1962">Taylor [1962]</bibRefCitation>
, see Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution ranges of the three clades of the Microhyla - Glyphoglossus assemblage. Distribution area of Microhyla I is shown in yellow, of Microhyla II in red, and of Glyphoglossus in blue. Distributional data from Gorin et al. (2020). Question mark denotes the unconfirmed record of &quot; Microhyla annamensis &quot; from Khao Sebab in eastern Thailand by Taylor (1962)." figureDoi="10.3897/zse.97.57968.figure1" httpUri="https://binary.pensoft.net/fig/496415" pageId="0" pageNumber="21">1</figureCitation>
<figureCitation id="8E28131D3C0D0736FAF2817672778414" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution ranges of the three clades of the Microhyla - Glyphoglossus assemblage. Distribution area of Microhyla I is shown in yellow, of Microhyla II in red, and of Glyphoglossus in blue. Distributional data from Gorin et al. (2020). Question mark denotes the unconfirmed record of &quot; Microhyla annamensis &quot; from Khao Sebab in eastern Thailand by Taylor (1962)." figureDoi="10.3897/zse.97.57968.figure1" httpUri="https://binary.pensoft.net/fig/496415" pageId="0" pageNumber="21">1</figureCitation>
) is questionable (see
<bibRefCitation author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
<bibRefCitation id="11BFBE38E724F581746A6C7835B2075F" author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4834.4.2" author="Poyarkov, NA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="502 - 522" refId="B119" refString="Poyarkov, NA, Solovyeva, EN, Nguyen, TV, Geissler, P, 2020a. On the taxonomic status of three enigmatic Indochinese frog species (Amphibia: Anura) described by L. G. Andersson. Zootaxa 4834 (4): 502 - 522, DOI: https://doi.org/10.11646/zootaxa.4834.4.2" title="On the taxonomic status of three enigmatic Indochinese frog species (Amphibia: Anura) described by L. G. Andersson." url="https://doi.org/10.11646/zootaxa.4834.4.2" volume="4834" year="2020 a">2020a</bibRefCitation>
<bibRefCitation id="701E2A9C3A8453A2DFCB15FCEC69019E" DOI="https://doi.org/10.11646/zootaxa.4834.4.2" author="Poyarkov, NA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="502 - 522" refId="B119" refString="Poyarkov, NA, Solovyeva, EN, Nguyen, TV, Geissler, P, 2020a. On the taxonomic status of three enigmatic Indochinese frog species (Amphibia: Anura) described by L. G. Andersson. Zootaxa 4834 (4): 502 - 522, DOI: https://doi.org/10.11646/zootaxa.4834.4.2" title="On the taxonomic status of three enigmatic Indochinese frog species (Amphibia: Anura) described by L. G. Andersson." url="https://doi.org/10.11646/zootaxa.4834.4.2" volume="4834" year="2020 a">2020a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="morphological comparison">
<paragraph pageId="0" pageNumber="21">Morphological comparison.</paragraph>
<paragraph pageId="0" pageNumber="21">
<subSubSection id="65D40E5F636915E47FBA18FD49DA960C" pageId="0" pageNumber="21" type="morphological comparison">
<paragraph id="A5F7356F22B10C6F7C47339F4E0BD7A1" pageId="0" pageNumber="21">Morphological comparison.</paragraph>
<paragraph id="2F31901DE160A9EDA36DAF2845E82363" pageId="0" pageNumber="21">
The new genus
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="4CAF65F53FBDD58CB92ADC8BC48D89BD" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="7EBA95A103970E59403B4540346357F5" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. differs from its sister genus
<taxonomicName authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
<taxonomicName id="4E5908C1C6C421261E7AA06245AE8DE6" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="58379FD14195E0BB996F08A95CDE9CA1" italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
</taxonomicName>
Tschudi, 1838
<emphasis italics="true" pageId="0" pageNumber="21">s. str.</emphasis>
<emphasis id="C41A7AAD6D151F1B073E52705B7500A3" italics="true" pageId="0" pageNumber="21">s. str.</emphasis>
by the well-developed (vs poorly-developed) otic ramus of the squamosal, frontoparietals and exoccipitals fused (vs separated or slightly fused), exoccipitals fused with each other (vs always separated), omosternum present (vs usually absent), sphenethmoid and parasphenoid fused completely or partially (vs separated), cartilaginous crista parotica (vs mineralized posteriorly), cartilaginous prehallux (vs mineralized), tympanum externally visible or barely visible (vs concealed beneath skin), inner metatarsal tubercle well-developed, outer generally absent (vs two metatarsal tubercles well-developed), and in having digits dorso ventrally flattened, FI often reduced to a nub or shortened (vs variably longer). The new genus differs from the closely related genus
<taxonomicName authorityName="Guenther" authorityYear="1869" class="Amphibia" family="Microhylidae" genus="Glyphoglossus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphoglossus" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Glyphoglossus</emphasis>
<taxonomicName id="3B256707196E7FA6B3584599747C8F43" authorityName="Guenther" authorityYear="1869" class="Amphibia" family="Microhylidae" genus="Glyphoglossus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphoglossus" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="76410E4BF7D47322355FF76AAA2D3AD6" italics="true" pageId="0" pageNumber="21">Glyphoglossus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Günther">Guenther</normalizedToken>
<normalizedToken id="40821BDA393052C34DACE40C31697791" originalValue="Günther">Guenther</normalizedToken>
, 1869 by its smaller adult size with SVL &lt;25mm (vs SVL&gt; 25mm), skull longer than wide or almost equal (vs wider than long), alary process of premaxilla oriented slightly anteriorly (vs posteriorly), neopalatines present (vs obscured by vomers), vomers small, indistinct (vs large, well-developed), omosternum present (vs absent), terminal phalanges T-shaped (vs simple), tibio-tarsal articulation reaching well beyond snout (vs to the anterior border of the eye, or less), by body habitus short, triangular-shaped (vs stout, balloon-shaped), and by inner metatarsal tubercle not enlarged (vs enlarged, shovel-shaped).
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="C900E2E93D2E2615AC027D936E5D791C" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="971F2A0DF28C4B2C302AB58FDB2A650A" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. differs from
<taxonomicName authorityName="Gray" authorityYear="1831" class="Amphibia" family="Microhylidae" genus="Kaloula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaloula" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Kaloula</emphasis>
<taxonomicName id="B399B3467B8A02097D687713EEA3C841" authorityName="Gray" authorityYear="1831" class="Amphibia" family="Microhylidae" genus="Kaloula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaloula" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="2610B3881A392E6CA901398F1FED643B" italics="true" pageId="0" pageNumber="21">Kaloula</emphasis>
</taxonomicName>
Gray, 1831 by its much smaller adult body size SVL &lt;25 mm (vs SVL&gt; 38 mm), procoracoids absent (vs present), postchoanal portion of vomer absent (vs present), neopalatines present (vs obscured), prehallux formed by two elements (vs one), tibio-tarsal articulation reaching well beyond snout (vs to shoulder), absence (vs presence) of ridge on posterior margin of choanae, inner metatarsal tubercle not enlarged (vs enlarged and spatulate), and by body habitus short, triangular-shaped (vs robust). The new genus can be distinguished from
<taxonomicName class="Reptilia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uperodon" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Uperodon</emphasis>
<taxonomicName id="6F0CB9B9D4ABB49381D92A28E287E62E" class="Reptilia" family="Microhylidae" genus="Uperodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uperodon" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="20DE06A76466F14CE931C9A953109187" italics="true" pageId="0" pageNumber="21">Uperodon</emphasis>
</taxonomicName>
<normalizedToken originalValue="Duméril">Dumeril</normalizedToken>
<normalizedToken id="FE01B8FF7DE2633BA129D44ABD84E167" originalValue="Duméril">Dumeril</normalizedToken>
&amp; Bibron, 1841 by its smaller adult size, SVL &lt;25 mm (vs SVL&gt; 34 mm), postchoanal portion of vomer absent (vs present), neopalatines present (vs obscured), tibio-tarsal articulation reaching well beyond snout (vs posterior border of eye, or less), absence (vs presence) of ridge on posterior margins of choanae, inner metatarsal tubercle not enlarged (vs enlarged or spatulate), and by body habitus short, triangular-shaped (vs robust and globular).
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="E6649C36F85F1395C075EBF840863BBF" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="20BB64B5666B8935C68F43CCC6F9660F" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. differs from
<taxonomicName class="Amphibia" family="Microhylidae" genus="Phrynella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phrynella" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Phrynella</emphasis>
<taxonomicName id="710F2B2E88F33B1D681374FBB7D5D1A2" class="Amphibia" family="Microhylidae" genus="Phrynella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phrynella" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="FED7265DB4DF2F5DACC7CAA703E7B6FE" italics="true" pageId="0" pageNumber="21">Phrynella</emphasis>
</taxonomicName>
Boulenger, 1887 by its smaller adult size, SVL &lt;25 mm (vs SVL&gt; 30 mm), medial process of the prechoanal part of vomer absent (vs present), neopalatines present (vs absent), procoracoids absent (vs present), vertebral column diplasiocoelus (vs procoelus), metatarsal tubercules separate (vs united), by tibio-tarsal articulation reaching well beyond snout (vs to tympanic region), by body habitus short, triangular-shaped (vs robust and flattened), and by generally dull brownish coloration of inguinal and dorsal surfaces (vs greenish coloration of dorsum and bright-red coloration of inguinal area, and ventral surfaces of limbs). The new genus further differs from
<taxonomicName authorityName="Parker" authorityYear="1934" class="Amphibia" family="Microhylidae" genus="Metaphrynella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Metaphrynella" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Metaphrynella</emphasis>
<taxonomicName id="9638D81E9F34817B6A9E31F6D529126A" authorityName="Parker" authorityYear="1934" class="Amphibia" family="Microhylidae" genus="Metaphrynella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Metaphrynella" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="CC72D6527DF7D7074887141904041949" italics="true" pageId="0" pageNumber="21">Metaphrynella</emphasis>
</taxonomicName>
Parker, 1934 by its smaller adult size, SVL &lt;25 mm (vs SVL&gt; 25 mm), skull longer than wide or almost equal (vs wider than long), neopalatines present (vs absent), omosternum present (vs absent), vertebral column diplasiocoelus (vs procoelus), tibio-tarsal articulation reaching well beyond snout (vs to tympanic region), absence (vs presence) of a ridge on posterior margins of choanae, metatarsal tubercules separate (vs united and enlarged), and by finger webbing absent (vs present). The new genus differs from
<taxonomicName authorityName="Garg &amp; Biju" authorityYear="2019" class="Amphibia" family="Microhylidae" genus="Mysticellus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mysticellus" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Mysticellus</emphasis>
<taxonomicName id="2A9D4332BBA3D77529D4E341B445AF75" authorityName="Garg &amp; Biju" authorityYear="2019" class="Amphibia" family="Microhylidae" genus="Mysticellus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mysticellus" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="14558B9C3630EAD182E870D224283C8D" italics="true" pageId="0" pageNumber="21">Mysticellus</emphasis>
</taxonomicName>
Garg &amp; Biju, 2019 by its short triangular-shaped body habitus (vs slender), supratympanic fold present (vs absent), finger and toe tips enlarged with prominent discs (vs slightly enlarged), toe webbing well-developed (vs rudimentary), supernumerary carpal tubercles absent (vs prominent subarticular tubercles alternating with additional smaller tubercles), and the two prominent blackish-brown
<normalizedToken originalValue="false-eye">'false-eye'</normalizedToken>
<normalizedToken id="FA0E57BE29C81C331CB02743CCD1539C" originalValue="false-eye">'false-eye'</normalizedToken>
inguinal spots absent (vs present).
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="A9E045F35F41AF267ECBA124080F0F21" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="67DB91ACB4D4EDF30B7C903CAA5209E5" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. differs from
<taxonomicName authorityName="Dubois" authorityYear="1987" class="Amphibia" family="Microhylidae" genus="Micryletta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micryletta" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Micryletta</emphasis>
<taxonomicName id="D86238242C77CD8015A5E7690D16481E" authorityName="Dubois" authorityYear="1987" class="Amphibia" family="Microhylidae" genus="Micryletta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micryletta" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="1DE61BAF3C38FF8097E166EC8502889C" italics="true" pageId="0" pageNumber="21">Micryletta</emphasis>
</taxonomicName>
Dubois, 1987 by its snout longer than eye diameter, and having eye less (vs more) prominent in lateral and dorsal aspects, finger and toe tips enlarged with prominent discs (vs slightly enlarged), toe webbing well-developed (vs rudimentary or absent), supernumerary carpal tubercles absent (vs present), omosternum present (vs absent), neopalatines present (vs absent), tibio-tarsal articulation reaching well beyond snout (vs to anterior border eye, or less), supratympanic fold present (vs absent), and body habitus short, triangular-shaped (vs slender). Finally, the new genus is distinguished from
<taxonomicName class="Amphibia" family="Microhylidae" genus="Chaperina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Chaperina" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Chaperina</emphasis>
<taxonomicName id="BF1A7B48AF066B761F4F4D5CDCCE82C0" class="Amphibia" family="Microhylidae" genus="Chaperina" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Chaperina" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="E66D67C69FC157183FD37939C0F3F80E" italics="true" pageId="0" pageNumber="21">Chaperina</emphasis>
</taxonomicName>
Mocquard, 1892 by clavicles and procoracoids absent (vs present), postchoanal portion of vomer absent (vs present), omosternum present (vs absent), terminal phalanges T-shaped (vs simple), tibiotarsal articulation reaching well beyond snout (vs anterior border of eye), belly dull-colored (vs bright saffron-yellow belly with dark pattern), and by absence of spine-like projections on limbs (vs a long, narrow dermal spine projecting from calcaneus).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="larval morphology">
<paragraph pageId="0" pageNumber="21">Larval morphology.</paragraph>
<paragraph pageId="0" pageNumber="21">
<subSubSection id="698756140807A39FF4FD1EB424DF3324" pageId="0" pageNumber="21" type="larval morphology">
<paragraph id="EF0EA074BFFEC53DBB460ACF91E3A6AE" pageId="0" pageNumber="21">Larval morphology.</paragraph>
<paragraph id="80DF861CE6A6528340EE70FC224F9239" pageId="0" pageNumber="21">
Description of the larval stages of the
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="AE4FBB5DE926A73833AC5DFCC0718053" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="951F0ED4BBB0BDD506241EA26D504F3C" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. members are sparse and often not detailed.
<bibRefCitation author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. (2014)</bibRefCitation>
<bibRefCitation id="80A7E1DDD1DC61F6E55848BAC4AE7D64" author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. (2014)</bibRefCitation>
provided descriptions, photos and illustrations of tadpole morphology for
<taxonomicName genus="N." lsidName="N. annamensis" pageId="0" pageNumber="21" rank="species" species="annamensis">
<emphasis italics="true" pageId="0" pageNumber="21">N. annamensis</emphasis>
<taxonomicName id="47B757DA28302F7C5F4B5EC44577A1F7" lsidName="N. annamensis" pageId="0" pageNumber="21" rank="species" species="annamensis">
<emphasis id="3CA0ACDB315AB174362D513AEF0206E0" italics="true" pageId="0" pageNumber="21">N. annamensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
<taxonomicName id="4712B82DACB8493CB9D52689EAF39E08" lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis id="DEF2E0D55DE020C8AB0CB5538299A955" italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
</taxonomicName>
and
<taxonomicName genus="N." lsidName="N. pulchella" pageId="0" pageNumber="21" rank="species" species="pulchella">
<emphasis italics="true" pageId="0" pageNumber="21">N. pulchella</emphasis>
<taxonomicName id="A64AEEE6376AB10128A17E0FDC5FE525" lsidName="N. pulchella" pageId="0" pageNumber="21" rank="species" species="pulchella">
<emphasis id="254F7581B6BB6706E8DE104DD0AC7B53" italics="true" pageId="0" pageNumber="21">N. pulchella</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4247.4.4" author="Vassilieva, AB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="413 - 428" refId="B159" refString="Vassilieva, AB, Trounov, VL, Poyarkov, NA, Galoyan, EA, 2017. The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development. Zootaxa 4247 (4): 413 - 428, DOI: https://doi.org/10.11646/zootaxa.4247.4.4" title="The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development." url="https://doi.org/10.11646/zootaxa.4247.4.4" volume="4247" year="2017">Vassilieva et al. (2017)</bibRefCitation>
<bibRefCitation id="1B8A6D7F7A75E55B436A9C38FC5DFAAC" DOI="https://doi.org/10.11646/zootaxa.4247.4.4" author="Vassilieva, AB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="413 - 428" refId="B159" refString="Vassilieva, AB, Trounov, VL, Poyarkov, NA, Galoyan, EA, 2017. The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development. Zootaxa 4247 (4): 413 - 428, DOI: https://doi.org/10.11646/zootaxa.4247.4.4" title="The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development." url="https://doi.org/10.11646/zootaxa.4247.4.4" volume="4247" year="2017">Vassilieva et al. (2017)</bibRefCitation>
provided a detailed description of development, larval morphology and anatomy for
<taxonomicName genus="N." lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
<taxonomicName id="9C26C4EF356CAE3787D61BAA4EB88EFC" lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis id="6D53F20C31D394A8FEED5D8B038B53CB" italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.15625/0866-7160/v38n2.7738" author="Le, TD" journalOrPublisher="Tap Chi Sinh Hoc [Journal of Biology], Hanoi" pageId="0" pageNumber="21" pagination="154 - 161" refId="B87" refString="Le, TD, Hoa, TH, Luong, MA, Nguyen, QT, 2016. Advertisment call and description of the tadpole of Microhyla marmorata Bain &amp; Nguyen, 2004 from Xuan Son National Park, Phu Tho Province. Tap Chi Sinh Hoc [Journal of Biology], Hanoi 38 (2): 154 - 161, DOI: https://doi.org/10.15625/0866-7160/v38n2.7738" title="Advertisment call and description of the tadpole of Microhyla marmorata Bain &amp; Nguyen, 2004 from Xuan Son National Park, Phu Tho Province." url="https://doi.org/10.15625/0866-7160/v38n2.7738" volume="38" year="2016">Le et al. (2016)</bibRefCitation>
<bibRefCitation id="789D85A7871F29F77975B5D16B83588F" DOI="https://doi.org/10.15625/0866-7160/v38n2.7738" author="Le, TD" journalOrPublisher="Tap Chi Sinh Hoc [Journal of Biology], Hanoi" pageId="0" pageNumber="21" pagination="154 - 161" refId="B87" refString="Le, TD, Hoa, TH, Luong, MA, Nguyen, QT, 2016. Advertisment call and description of the tadpole of Microhyla marmorata Bain &amp; Nguyen, 2004 from Xuan Son National Park, Phu Tho Province. Tap Chi Sinh Hoc [Journal of Biology], Hanoi 38 (2): 154 - 161, DOI: https://doi.org/10.15625/0866-7160/v38n2.7738" title="Advertisment call and description of the tadpole of Microhyla marmorata Bain &amp; Nguyen, 2004 from Xuan Son National Park, Phu Tho Province." url="https://doi.org/10.15625/0866-7160/v38n2.7738" volume="38" year="2016">Le et al. (2016)</bibRefCitation>
provided a brief description of tadpole morphology of
<taxonomicName genus="N." lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
<taxonomicName id="9A8FAB8E033908E36DC9C7C8B17ED0D1" lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis id="C0A29E7EEF2E8A77525B41AE8C0DC330" italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
</taxonomicName>
.
<bibRefCitation author="Leong, TM" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="21" pagination="609 - 620" refId="B90" refString="Leong, TM, 2004. Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura). Raffles Bulletin of Zoology 52 (2): 609 - 620" title="Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura)." volume="52" year="2004">Leong (2004)</bibRefCitation>
<bibRefCitation id="0733302240C827CEE0CAE7BAF64E654F" author="Leong, TM" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="21" pagination="609 - 620" refId="B90" refString="Leong, TM, 2004. Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura). Raffles Bulletin of Zoology 52 (2): 609 - 620" title="Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura)." volume="52" year="2004">Leong (2004)</bibRefCitation>
provided a short description and photographs of larval and metamorph morphology for
<taxonomicName genus="N." lsidName="N. annectens" pageId="0" pageNumber="21" rank="species" species="annectens">
<emphasis italics="true" pageId="0" pageNumber="21">N. annectens</emphasis>
<taxonomicName id="3F74F18F7C46B92E4D3597BE0A954304" lsidName="N. annectens" pageId="0" pageNumber="21" rank="species" species="annectens">
<emphasis id="E5F16ACB338293F2F6EA9F4BE6EF62D2" italics="true" pageId="0" pageNumber="21">N. annectens</emphasis>
</taxonomicName>
. Brief descriptions and figures depicting larvae of
<taxonomicName genus="N." lsidName="N. petrigena" pageId="0" pageNumber="21" rank="species" species="petrigena">
<emphasis italics="true" pageId="0" pageNumber="21">N. petrigena</emphasis>
<taxonomicName id="50D76A5ADF6B6ED85356A9C5B1B5231C" lsidName="N. petrigena" pageId="0" pageNumber="21" rank="species" species="petrigena">
<emphasis id="D0DE5ACDF4E6BB954F320E23AD27E8D4" italics="true" pageId="0" pageNumber="21">N. petrigena</emphasis>
</taxonomicName>
and
<taxonomicName genus="N." lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
<taxonomicName id="53C90B9739329601EB869B56C5290F4B" lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis id="D1AFA24D8027657BB01E711F8A204300" italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
</taxonomicName>
are found in the original description of these species by
<bibRefCitation author="Inger, RF" journalOrPublisher="The Sarawak Museum Journal" pageId="0" pageNumber="21" pagination="311 - 322" refId="B74" refString="Inger, RF, Frogner, KJ, 1979. New species of narrow-mouth frogs (genus Microhyla) from Borneo. The Sarawak Museum Journal 27 (48): 311 - 322" title="New species of narrow-mouth frogs (genus Microhyla) from Borneo." volume="27" year="1979">Inger and Frogner (1979)</bibRefCitation>
<bibRefCitation id="0093CE559671F8D5DF19CDCA2F67417B" author="Inger, RF" journalOrPublisher="The Sarawak Museum Journal" pageId="0" pageNumber="21" pagination="311 - 322" refId="B74" refString="Inger, RF, Frogner, KJ, 1979. New species of narrow-mouth frogs (genus Microhyla) from Borneo. The Sarawak Museum Journal 27 (48): 311 - 322" title="New species of narrow-mouth frogs (genus Microhyla) from Borneo." volume="27" year="1979">Inger and Frogner (1979)</bibRefCitation>
, as well as in Inger and Steubing (2005) and
<bibRefCitation author="Haas, A" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="21" publicationUrl="http://http://www.frogsofborneo.org" refId="B58" refString="Haas, A, Das, I, Hertwig, ST, 2020.: Frogs of Borneo - The frogs of East Malaysia and their larval forms. [Electronic Database accessible at] http://http://www.frogsofborneo.org" title=": Frogs of Borneo - The frogs of East Malaysia and their larval forms. [Electronic Database accessible at]" url="http://http://www.frogsofborneo.org" year="2020">Haas et al. (2020)</bibRefCitation>
<bibRefCitation id="A4B8FB21F65462EA005E25D14F88DF7B" author="Haas, A" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="21" publicationUrl="http://http://www.frogsofborneo.org" refId="B58" refString="Haas, A, Das, I, Hertwig, ST, 2020.: Frogs of Borneo - The frogs of East Malaysia and their larval forms. [Electronic Database accessible at] http://http://www.frogsofborneo.org" title=": Frogs of Borneo - The frogs of East Malaysia and their larval forms. [Electronic Database accessible at]" url="http://http://www.frogsofborneo.org" year="2020">Haas et al. (2020)</bibRefCitation>
. Larval stages of
<taxonomicName genus="N." lsidName="N. hongiaoensis" pageId="0" pageNumber="21" rank="species" species="hongiaoensis">
<emphasis italics="true" pageId="0" pageNumber="21">N. hongiaoensis</emphasis>
<taxonomicName id="32D955C2704BAE51609029C8EE8A731A" lsidName="N. hongiaoensis" pageId="0" pageNumber="21" rank="species" species="hongiaoensis">
<emphasis id="AC6319D32F0BDC03A73BB72704E8784C" italics="true" pageId="0" pageNumber="21">N. hongiaoensis</emphasis>
</taxonomicName>
and
<taxonomicName genus="N." lsidName="N. nanapollexa" pageId="0" pageNumber="21" rank="species" species="nanapollexa">
<emphasis italics="true" pageId="0" pageNumber="21">N. nanapollexa</emphasis>
<taxonomicName id="2EC23D98BA592952A26FF8EB01C7A7BE" lsidName="N. nanapollexa" pageId="0" pageNumber="21" rank="species" species="nanapollexa">
<emphasis id="09BE70F6D6B18C1317AF9643657F463D" italics="true" pageId="0" pageNumber="21">N. nanapollexa</emphasis>
</taxonomicName>
remain unknown.
</paragraph>
<paragraph pageId="0" pageNumber="21">
<paragraph id="1F6EDC28F690689A1D0800C7EAB49039" pageId="0" pageNumber="21">
As with almost all larvae in
<taxonomicName class="Amphibia" family="Microhylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="family">Microhylidae</taxonomicName>
<taxonomicName id="7D083FEC96F79E48E3519C94F5854ACD" class="Amphibia" family="Microhylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="family">Microhylidae</taxonomicName>
, labial teeth and mandibles are absent from the oral discs of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="CDCFA2E23A1BEC69986F8E5049290DC8" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B82D331E2D938001967C9D3F1C451A02" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
tadpoles. Most species of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="C851A072D49E1202A718928BEBEC959C" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="C7184B2FD62E2AC40D63127A5D2D25F5" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
have larval morphology resembling that of many pond-breeding
<taxonomicName authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
<taxonomicName id="A4C754B5019D48E748540A62832FACDE" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="6CE390768316D44D5B2AA1133CB39AF7" italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
</taxonomicName>
species (
<bibRefCitation author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
<bibRefCitation id="0FF3FA9997FB9365B4B2D07A422631B2" author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
) with rather short-tailed transparent or semi-transparent
<normalizedToken originalValue="Ortons">Orton's</normalizedToken>
<normalizedToken id="3BF6840FE297B46541F51818ECB2D1D7" originalValue="Ortons">Orton's</normalizedToken>
type II tadpoles (
<bibRefCitation DOI="https://doi.org/10.2307/2411661" author="Orton, GL" journalOrPublisher="Systematic Zoology" pageId="0" pageNumber="21" pagination="63 - 75" refId="B108" refString="Orton, GL, 1953. The systematics of vertebrate larvae. Systematic Zoology 2 (2): 63 - 75, DOI: https://doi.org/10.2307/2411661" title="The systematics of vertebrate larvae." url="https://doi.org/10.2307/2411661" volume="2" year="1953">Orton 1953</bibRefCitation>
<bibRefCitation id="E60D5F944BEFCE1F66844D05607A48BE" DOI="https://doi.org/10.2307/2411661" author="Orton, GL" journalOrPublisher="Systematic Zoology" pageId="0" pageNumber="21" pagination="63 - 75" refId="B108" refString="Orton, GL, 1953. The systematics of vertebrate larvae. Systematic Zoology 2 (2): 63 - 75, DOI: https://doi.org/10.2307/2411661" title="The systematics of vertebrate larvae." url="https://doi.org/10.2307/2411661" volume="2" year="1953">Orton 1953</bibRefCitation>
), that are mid-water column (neustonic) feeders with comparatively unexpanded lower labium and anteriorly directed terminal mouths, lateral orientation of eyes, spiraculum located in a medial position on the venter, spiracular flap with crenulate margins, and tail lacking terminal filament (
<bibRefCitation author="Altig, R" journalOrPublisher="Herpetological Monographs" pageId="0" pageNumber="21" pagination="81 - 109" refId="B4" refString="Altig, R, Johnston, GF, 1989. Guilds of anuran larvae: relationships among developmental modes, morphologies, and habitats. Herpetological Monographs 3: 81 - 109" title="Guilds of anuran larvae: relationships among developmental modes, morphologies, and habitats." volume="3" year="1989">Altig and Johnston 1989</bibRefCitation>
<bibRefCitation id="ABD075872428CEE2CE414791B9526789" author="Altig, R" journalOrPublisher="Herpetological Monographs" pageId="0" pageNumber="21" pagination="81 - 109" refId="B4" refString="Altig, R, Johnston, GF, 1989. Guilds of anuran larvae: relationships among developmental modes, morphologies, and habitats. Herpetological Monographs 3: 81 - 109" title="Guilds of anuran larvae: relationships among developmental modes, morphologies, and habitats." volume="3" year="1989">Altig and Johnston 1989</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1093/molbev/mss075" author="Donnelly, MA" journalOrPublisher="Novitates" pageId="0" pageNumber="21" pagination="1 - 19" refId="B35" refString="Donnelly, MA, de Sa, RO, Guyer, C, 1990. Description of the tadpoles of Gastrophryne pictiventris and Nelsonophryne aterrima (Anura: Microhylidae), with a review of morphological variation in free-swimming microhylid larvae. Novitates 2976: 1 - 19, DOI: https://doi.org/10.1093/molbev/mss075" title="Description of the tadpoles of Gastrophryne pictiventris and Nelsonophryne aterrima (Anura: Microhylidae), with a review of morphological variation in free-swimming microhylid larvae." url="https://doi.org/10.1093/molbev/mss075" volume="2976" year="1990">Donnelly et al. 1990</bibRefCitation>
<bibRefCitation id="6E9E2A7D27CC03C82EC4E0C1A8DB2E33" DOI="https://doi.org/10.1093/molbev/mss075" author="Donnelly, MA" journalOrPublisher="Novitates" pageId="0" pageNumber="21" pagination="1 - 19" refId="B35" refString="Donnelly, MA, de Sa, RO, Guyer, C, 1990. Description of the tadpoles of Gastrophryne pictiventris and Nelsonophryne aterrima (Anura: Microhylidae), with a review of morphological variation in free-swimming microhylid larvae. Novitates 2976: 1 - 19, DOI: https://doi.org/10.1093/molbev/mss075" title="Description of the tadpoles of Gastrophryne pictiventris and Nelsonophryne aterrima (Anura: Microhylidae), with a review of morphological variation in free-swimming microhylid larvae." url="https://doi.org/10.1093/molbev/mss075" volume="2976" year="1990">Donnelly et al. 1990</bibRefCitation>
;
<bibRefCitation author="Leong, TM" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="21" pagination="609 - 620" refId="B90" refString="Leong, TM, 2004. Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura). Raffles Bulletin of Zoology 52 (2): 609 - 620" title="Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura)." volume="52" year="2004">Leong 2004</bibRefCitation>
<bibRefCitation id="D0AC86F56877F949383FB26BD35BE4B7" author="Leong, TM" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="21" pagination="609 - 620" refId="B90" refString="Leong, TM, 2004. Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura). Raffles Bulletin of Zoology 52 (2): 609 - 620" title="Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura)." volume="52" year="2004">Leong 2004</bibRefCitation>
). In contrast, many species of
<taxonomicName authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
<taxonomicName id="67EB97E6111121432E0EF73774E17E68" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="7B235F98D50A6C1FF64BAD89ECB6F63F" italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="0" pageNumber="21">s. str.</emphasis>
<emphasis id="87E2E5A74972DC16D55EFF54AFEE841B" italics="true" pageId="0" pageNumber="21">s. str.</emphasis>
are surface suspension feeders, and demonstrate greatly expanded lower labium and dorso-terminal mouth orientation; they may have terminal filament on tail and smooth margins of spiracular flap (e.g.,
<bibRefCitation author="Leong, TM" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="21" pagination="609 - 620" refId="B90" refString="Leong, TM, 2004. Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura). Raffles Bulletin of Zoology 52 (2): 609 - 620" title="Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura)." volume="52" year="2004">Leong 2004</bibRefCitation>
<bibRefCitation id="2E8A70BB835840A84ABF5459E6EE01BF" author="Leong, TM" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="21" pagination="609 - 620" refId="B90" refString="Leong, TM, 2004. Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura). Raffles Bulletin of Zoology 52 (2): 609 - 620" title="Larval descriptions of some poorly known tadpoles from Peninsular Malaysia (Amphibia: Anura)." volume="52" year="2004">Leong 2004</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1675.1.5" author="Hendrix, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="67 - 68" refId="B67" refString="Hendrix, R, Gawor, A, Vences, M, Ziegler, T, 2008. The tadpole of the narrow-mouthed frog Microhyla fissipes from Vietnam (Anura: Microhylidae). Zootaxa 1675 (1): 67 - 68, DOI: https://doi.org/10.11646/zootaxa.1675.1.5" title="The tadpole of the narrow-mouthed frog Microhyla fissipes from Vietnam (Anura: Microhylidae)." url="https://doi.org/10.11646/zootaxa.1675.1.5" volume="1675" year="2008">Hendrix et al. 2008</bibRefCitation>
<bibRefCitation id="023D88B3F21242EAA717E091F5CB23BE" DOI="https://doi.org/10.11646/zootaxa.1675.1.5" author="Hendrix, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="67 - 68" refId="B67" refString="Hendrix, R, Gawor, A, Vences, M, Ziegler, T, 2008. The tadpole of the narrow-mouthed frog Microhyla fissipes from Vietnam (Anura: Microhylidae). Zootaxa 1675 (1): 67 - 68, DOI: https://doi.org/10.11646/zootaxa.1675.1.5" title="The tadpole of the narrow-mouthed frog Microhyla fissipes from Vietnam (Anura: Microhylidae)." url="https://doi.org/10.11646/zootaxa.1675.1.5" volume="1675" year="2008">Hendrix et al. 2008</bibRefCitation>
;
<bibRefCitation author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
<bibRefCitation id="9394D26D1CC42095F1C30DDC51A65D48" author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="21">
<paragraph id="8DDC4AD3C72BF44E280514E3A836435A" pageId="0" pageNumber="21">
A peculiar exception is the case of
<taxonomicName genus="N." lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
<taxonomicName id="7549817409A3D7236A4D458BAB035D52" lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis id="86F0A3236A1B4D02DBD41002AB3E6228" italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
</taxonomicName>
, which is an obligate phytotelm-breeding species that reproduces in water-filled tree hollows (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4247.4.4" author="Vassilieva, AB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="413 - 428" refId="B159" refString="Vassilieva, AB, Trounov, VL, Poyarkov, NA, Galoyan, EA, 2017. The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development. Zootaxa 4247 (4): 413 - 428, DOI: https://doi.org/10.11646/zootaxa.4247.4.4" title="The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development." url="https://doi.org/10.11646/zootaxa.4247.4.4" volume="4247" year="2017">Vassilieva et al. 2017</bibRefCitation>
<bibRefCitation id="00021EAF77EBF73CE3024649A776E07B" DOI="https://doi.org/10.11646/zootaxa.4247.4.4" author="Vassilieva, AB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="413 - 428" refId="B159" refString="Vassilieva, AB, Trounov, VL, Poyarkov, NA, Galoyan, EA, 2017. The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development. Zootaxa 4247 (4): 413 - 428, DOI: https://doi.org/10.11646/zootaxa.4247.4.4" title="The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development." url="https://doi.org/10.11646/zootaxa.4247.4.4" volume="4247" year="2017">Vassilieva et al. 2017</bibRefCitation>
). The oophagous tadpoles of this species differ from larvae of pond-dwelling
<taxonomicName authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
<taxonomicName id="96C4C8D56CA40C36A5221572108DBE88" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="A1E57F63BE3538BECC3F60716FE53332" italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
</taxonomicName>
and
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="C529E7766E7A262AE831FE962294C364" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="BC1BB7BD4CB7F5063617CB984F429653" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
species in many aspects, including external morphology (extremely long tails, dorsolateral position of the eyes, dark pigmentation), morphology of digestive tract (large, extensible stomach with comparatively short intestine), and characteristic oral morphology (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4247.4.4" author="Vassilieva, AB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="413 - 428" refId="B159" refString="Vassilieva, AB, Trounov, VL, Poyarkov, NA, Galoyan, EA, 2017. The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development. Zootaxa 4247 (4): 413 - 428, DOI: https://doi.org/10.11646/zootaxa.4247.4.4" title="The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development." url="https://doi.org/10.11646/zootaxa.4247.4.4" volume="4247" year="2017">Vassilieva et al. 2017</bibRefCitation>
<bibRefCitation id="90943B05755D65574C3338E86DACDD94" DOI="https://doi.org/10.11646/zootaxa.4247.4.4" author="Vassilieva, AB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="413 - 428" refId="B159" refString="Vassilieva, AB, Trounov, VL, Poyarkov, NA, Galoyan, EA, 2017. The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development. Zootaxa 4247 (4): 413 - 428, DOI: https://doi.org/10.11646/zootaxa.4247.4.4" title="The phytotelm tadpoles of Microhyla arboricola (Anura: Microhylidae) from Vietnam, with comments on reproductive biology and development." url="https://doi.org/10.11646/zootaxa.4247.4.4" volume="4247" year="2017">Vassilieva et al. 2017</bibRefCitation>
).
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" baseAuthorityName="Bain &amp; Truong" baseAuthorityYear="2004" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla nanapollexa" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="nanapollexa">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla nanapollexa</emphasis>
<taxonomicName id="CF63F7698EA68291342BB7FD135FBAA6" baseAuthorityName="Bain &amp; Truong" baseAuthorityYear="2004" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla nanapollexa" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="nanapollexa">
<emphasis id="F6F7B0FA38C3AB89F30BB76AC9473159" italics="true" pageId="0" pageNumber="21">Nanohyla nanapollexa</emphasis>
</taxonomicName>
was suggested as phytotelm-breeder as a single specimen of this species was recorded in a water-filled tree hollow (
<bibRefCitation DOI="https://doi.org/10.7717/peerj.9411" author="Gorin, VA" journalOrPublisher="Hoffmann, Stuttgart" pageId="0" pageNumber="21" refId="B54" refString="Gorin, VA, Solovyeva, EN, Hasan, MK, Okamiya, H, Karunarathna, DMSS, Pawangkhanant, P, de Silva, A, Juthong, W, Milto, KD, Nguyen, LT, Suwannapoom, C, Haas, A, Bickford, DP, Das, I, Poyarkov, NA, 2020. A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ 8: 2-47 [e9411]. https://doi.org/10.7717/peerj.9411" title="A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ 8: 2 - 47 [e 9411]." url="https://doi.org/10.7717/peerj.9411" year="2020">Gorin et al. 2020</bibRefCitation>
<bibRefCitation id="B9642A25E4B184437CD2B5B0D5467BC1" DOI="https://doi.org/10.7717/peerj.9411" author="Gorin, VA" journalOrPublisher="Hoffmann, Stuttgart" pageId="0" pageNumber="21" refId="B54" refString="Gorin, VA, Solovyeva, EN, Hasan, MK, Okamiya, H, Karunarathna, DMSS, Pawangkhanant, P, de Silva, A, Juthong, W, Milto, KD, Nguyen, LT, Suwannapoom, C, Haas, A, Bickford, DP, Das, I, Poyarkov, NA, 2020. A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ 8: 2-47 [e9411]. https://doi.org/10.7717/peerj.9411" title="A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ 8: 2 - 47 [e 9411]." url="https://doi.org/10.7717/peerj.9411" year="2020">Gorin et al. 2020</bibRefCitation>
), although the details of reproductive biology and tadpole morphology of this species are still unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="taxonomic comment">
<paragraph pageId="0" pageNumber="21">Taxonomic comment.</paragraph>
<paragraph pageId="0" pageNumber="21">
<taxonomicName class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla pulverata" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="pulverata">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla pulverata</emphasis>
<subSubSection id="1C0883C58EADABEE86104405CD24B228" pageId="0" pageNumber="21" type="taxonomic comment">
<paragraph id="1AB7876F89FF5D52729B81DF79DC80AD" pageId="0" pageNumber="21">Taxonomic comment.</paragraph>
<paragraph id="E41A933724890C4A9F929416673E4F50" pageId="0" pageNumber="21">
<taxonomicName id="0AA682F2098AE62909A86B12B6ACF243" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla pulverata" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="species" species="pulverata">
<emphasis id="4DF5BBDF227DB6E2492C553F6B40A9BF" italics="true" pageId="0" pageNumber="21">Microhyla pulverata</emphasis>
</taxonomicName>
Bain &amp; Nguyen, 2004 was considered a junior synonym of
<taxonomicName genus="N." lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
<taxonomicName id="F8D6E537BD7E5B33A7BA15360D151A33" lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis id="F6B5C0706784081BFF78D3389396EC0B" italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
</taxonomicName>
based on the phylogenetic results of
<bibRefCitation DOI="https://doi.org/10.7717/peerj.9411" author="Gorin, VA" journalOrPublisher="Hoffmann, Stuttgart" pageId="0" pageNumber="21" refId="B54" refString="Gorin, VA, Solovyeva, EN, Hasan, MK, Okamiya, H, Karunarathna, DMSS, Pawangkhanant, P, de Silva, A, Juthong, W, Milto, KD, Nguyen, LT, Suwannapoom, C, Haas, A, Bickford, DP, Das, I, Poyarkov, NA, 2020. A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ 8: 2-47 [e9411]. https://doi.org/10.7717/peerj.9411" title="A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ 8: 2 - 47 [e 9411]." url="https://doi.org/10.7717/peerj.9411" year="2020">Gorin et al. (2020)</bibRefCitation>
<bibRefCitation id="63D6C9966F80FD866887E375F82CC74F" DOI="https://doi.org/10.7717/peerj.9411" author="Gorin, VA" journalOrPublisher="Hoffmann, Stuttgart" pageId="0" pageNumber="21" refId="B54" refString="Gorin, VA, Solovyeva, EN, Hasan, MK, Okamiya, H, Karunarathna, DMSS, Pawangkhanant, P, de Silva, A, Juthong, W, Milto, KD, Nguyen, LT, Suwannapoom, C, Haas, A, Bickford, DP, Das, I, Poyarkov, NA, 2020. A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ 8: 2-47 [e9411]. https://doi.org/10.7717/peerj.9411" title="A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ 8: 2 - 47 [e 9411]." url="https://doi.org/10.7717/peerj.9411" year="2020">Gorin et al. (2020)</bibRefCitation>
; the same study also reported on three putative candidate species within
<taxonomicName genus="N." lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
<taxonomicName id="29494CC9147164038AB7E14A271304C3" lsidName="N. arboricola" pageId="0" pageNumber="21" rank="species" species="arboricola">
<emphasis id="1D3139CDCD54301B5E19F5E0B72F2CD4" italics="true" pageId="0" pageNumber="21">N. arboricola</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
<taxonomicName id="2B430BECB555305A88E969C55D37045E" lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis id="C33BA0E3CAA02F2F08B7547E71DE825A" italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
</taxonomicName>
, and
<taxonomicName genus="N." lsidName="N. petrigena" pageId="0" pageNumber="21" rank="species" species="petrigena">
<emphasis italics="true" pageId="0" pageNumber="21">N. petrigena</emphasis>
<taxonomicName id="B206F921D19DC2435D6D312576364F2C" lsidName="N. petrigena" pageId="0" pageNumber="21" rank="species" species="petrigena">
<emphasis id="685B8AA90905870E1BBA722DD57E1F0C" italics="true" pageId="0" pageNumber="21">N. petrigena</emphasis>
</taxonomicName>
, indicating that our knowledge on diversity of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="8B7D687EC9F622F5232350B3B217BBCF" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="39CEC35DBA731ECACE870C61B445EEFB" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
is still incomplete.
</paragraph>
<paragraph pageId="0" pageNumber="21">
<paragraph id="E2A09BB49355C7B654FBFE0FB317CD2A" pageId="0" pageNumber="21">
Certain variation in diagnostically important characters of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="C63D6C7DA657B5BDEFF0412DD8DE4C63" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="73914C52546DFEFB37DE7C8F51A266C1" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. requires further comments.
<bibRefCitation DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen (2004)</bibRefCitation>
<bibRefCitation id="FAE035D6C9D9E89D03E6013B0B6CC7A6" DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen (2004)</bibRefCitation>
reported on significant variation in size and shape of the outer metatarsal tubercle in
<taxonomicName genus="N." lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
<taxonomicName id="BDB6F3400A801F5A320942B0B52FF17D" lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis id="205D81FBBE464FDDDAE2F4D8F2A30937" italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
</taxonomicName>
which was reported to vary from almost indistinct to
<normalizedToken originalValue="“conical.”">&quot;conical.&quot;</normalizedToken>
<normalizedToken id="843C05CAA26677EC51A85FEB2EDB09C7" originalValue="“conical.”">&quot;conical.&quot;</normalizedToken>
We have examined a large series of
<taxonomicName genus="N." lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
<taxonomicName id="276750DD9382E2561E4E019F87EA5222" lsidName="N. marmorata" pageId="0" pageNumber="21" rank="species" species="marmorata">
<emphasis id="BA920EF5C8686C6B253259B247545230" italics="true" pageId="0" pageNumber="21">N. marmorata</emphasis>
</taxonomicName>
(see
<bibRefCitation author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
<bibRefCitation id="2E2C2C106C948A735DF838912D248634" author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. 2014</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4543.4.4" author="Nguyen, LT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="549 - 580" refId="B102" refString="Nguyen, LT, Poyarkov, NA, Nguyen, TT, Nguyen, TA, Nguyen, VH, Gorin, VA, Murphy, RW, Nguyen, SN, 2019. A new species of the genus Microhyla Tschudi, 1838 (Amphibia: Anura: Microhylidae) from Tay Nguyen Plateau, Central Vietnam. Zootaxa 4543 (4): 549 - 580, DOI: https://doi.org/10.11646/zootaxa.4543.4.4" title="A new species of the genus Microhyla Tschudi, 1838 (Amphibia: Anura: Microhylidae) from Tay Nguyen Plateau, Central Vietnam." url="https://doi.org/10.11646/zootaxa.4543.4.4" volume="4543" year="2019">Nguyen et al. 2019</bibRefCitation>
<bibRefCitation id="A10FB6AA1E3A3062AF93F890F4D218C2" DOI="https://doi.org/10.11646/zootaxa.4543.4.4" author="Nguyen, LT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="549 - 580" refId="B102" refString="Nguyen, LT, Poyarkov, NA, Nguyen, TT, Nguyen, TA, Nguyen, VH, Gorin, VA, Murphy, RW, Nguyen, SN, 2019. A new species of the genus Microhyla Tschudi, 1838 (Amphibia: Anura: Microhylidae) from Tay Nguyen Plateau, Central Vietnam. Zootaxa 4543 (4): 549 - 580, DOI: https://doi.org/10.11646/zootaxa.4543.4.4" title="A new species of the genus Microhyla Tschudi, 1838 (Amphibia: Anura: Microhylidae) from Tay Nguyen Plateau, Central Vietnam." url="https://doi.org/10.11646/zootaxa.4543.4.4" volume="4543" year="2019">Nguyen et al. 2019</bibRefCitation>
) and found that in this species the outer metatarsal tubercle usually is not discernable or is indistinct; we assume that this discrepancy might be explained with the differences in preservation of specimens examined by us and by
<bibRefCitation DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen (2004)</bibRefCitation>
<bibRefCitation id="173A9FFFE1C0037954234CFB44CC033E" DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen (2004)</bibRefCitation>
.
<bibRefCitation author="Hoang, VC" journalOrPublisher="Asian Herpetological Research" pageId="0" pageNumber="21" pagination="1 - 22" refId="B69" refString="Hoang, VC, Nguyen, TT, Luong, MA, Nguyen, QT, Orlov, NL, Chen, Y, Wang, B, Jiang, J, 2020. A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam. Asian Herpetological Research 11 (3): 1 - 22" title="A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam." volume="11" year="2020">Hoang et al. (2020)</bibRefCitation>
<bibRefCitation id="ED948FEFF46CB1C4F73637848026B844" author="Hoang, VC" journalOrPublisher="Asian Herpetological Research" pageId="0" pageNumber="21" pagination="1 - 22" refId="B69" refString="Hoang, VC, Nguyen, TT, Luong, MA, Nguyen, QT, Orlov, NL, Chen, Y, Wang, B, Jiang, J, 2020. A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam. Asian Herpetological Research 11 (3): 1 - 22" title="A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam." volume="11" year="2020">Hoang et al. (2020)</bibRefCitation>
reported two metatarsal tubercles in their diagnosis of
<taxonomicName genus="N." lsidName="N. hongiaoensis" pageId="0" pageNumber="21" rank="species" species="hongiaoensis">
<emphasis italics="true" pageId="0" pageNumber="21">N. hongiaoensis</emphasis>
<taxonomicName id="3FDD178676D870AEB6F90F32E82F4532" lsidName="N. hongiaoensis" pageId="0" pageNumber="21" rank="species" species="hongiaoensis">
<emphasis id="BBB3AD5C572477B985FB5B944A43E73A" italics="true" pageId="0" pageNumber="21">N. hongiaoensis</emphasis>
</taxonomicName>
, however in the holotype description they refer to the outer metatarsal tubercle as
<normalizedToken originalValue="“indistinct;”">&quot;indistinct;&quot;</normalizedToken>
<normalizedToken id="44D89F5264D8D369850AEB081BBB5B33" originalValue="“indistinct;”">&quot;indistinct;&quot;</normalizedToken>
it is also not discernable in their photo of
<normalizedToken originalValue="holotypes">holotype's</normalizedToken>
<normalizedToken id="83E172F29B6528F0694588344632CA06" originalValue="holotypes">holotype's</normalizedToken>
foot (
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<bibRefCitation id="3C2F5C2F394AEC07A8B27A2B0207CE60" author="Hoang, VC" journalOrPublisher="Asian Herpetological Research" pageId="0" pageNumber="21" pagination="1 - 22" refId="B69" refString="Hoang, VC, Nguyen, TT, Luong, MA, Nguyen, QT, Orlov, NL, Chen, Y, Wang, B, Jiang, J, 2020. A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam. Asian Herpetological Research 11 (3): 1 - 22" title="A New species of Microhyla (Amphibia: Anura: Microhylidae) from Langbian Plateau, Central Vietnam." volume="11" year="2020">Hoang et al. 2020</bibRefCitation>
:fig. 3F). In all the remaining species of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="61F91E802110F9A26313865B8F7176FB" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="A0A5F91FF160FB437D39F63EF56D03A5" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov. it is absent, and we therefore consider this state to be diagnostic for the genus (in comparison to
<taxonomicName authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
<taxonomicName id="F912B2446B2B895E74FFC058AEC088A9" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microhyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="461B1E336459F0511E5586C5424D2AA7" italics="true" pageId="0" pageNumber="21">Microhyla</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="0" pageNumber="21">s. str.</emphasis>
<emphasis id="ED7A6074A5E07A401A08829BA91ABC68" italics="true" pageId="0" pageNumber="21">s. str.</emphasis>
, which has two metatarsal tubercles in all species but
<taxonomicName genus="M." lsidName="M. maculifera" pageId="0" pageNumber="21" rank="species" species="maculifera">
<emphasis italics="true" pageId="0" pageNumber="21">M. maculifera</emphasis>
<taxonomicName id="A9D396283567DACA7E34380275C7721C" lsidName="M. maculifera" pageId="0" pageNumber="21" rank="species" species="maculifera">
<emphasis id="B352696B2E00DAC63EF01978A9DBE4AC" italics="true" pageId="0" pageNumber="21">M. maculifera</emphasis>
</taxonomicName>
, see comment below). It is not clear why
<bibRefCitation DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen (2004)</bibRefCitation>
<bibRefCitation id="080785439D905FF1067577AB87C3A762" DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen (2004)</bibRefCitation>
, or
<bibRefCitation author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. (2014</bibRefCitation>
<bibRefCitation id="D1D58CAD689F10383A53727F452A2D2B" author="Poyarkov, NA" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="21" pagination="89 - 148" refId="B120" refString="Poyarkov, NA, Vassilieva, AB, Orlov, NL, Galoyan, EA, Tran, D, Le, DTT, Kretova, VD, Geissler, P, 2014. Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species. Russian Journal of Herpetology 21 (2): 89 - 148" title="Taxonomy and distribution of narrow-mouth frogs of the genus Microhyla Tschudi, 1838 (Anura: Microhylidae) from Vietnam with descriptions of five new species." volume="21" year="2014">Poyarkov et al. (2014</bibRefCitation>
; and other preceding studies) did not recognize the presence of externally visible tympanum in most of species of the genus (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Members of the new genus Nanohyla gen. nov. in life (males): N. annectens from Genting Highlands, Pahang, Malaysia (A), N. annamensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (B), N. arboricola from Chu Yang Sin N. P., Dak Lak, Vietnam (C), N. hongiaoensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (D), N. marmorata from Kon Chu Rang N. R., Gia Lai, Vietnam (E), N. nanapollexa from Kon Plong, Kon Tum, Vietnam (F), N. perparva from Gunung Mulu, Sarawak, Malaysia (G), N. petrigena from Gunung Mulu, Sarawak, Malaysia (H), and N. pulchella from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (I). Insets show tympanic area of the each species; white arrow points at the tympanic rim of the external tympanum. Photos by Nikolay A. Poyarkov (A-S, D-F, I), Vu Dang Hoang Nguyen (D), and Indraneil Das (G, H)." figureDoi="10.3897/zse.97.57968.figure11" httpUri="https://binary.pensoft.net/fig/496425" pageId="0" pageNumber="21">11</figureCitation>
<figureCitation id="1E3BB4CE98969B4329EA1D07631FB74E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Members of the new genus Nanohyla gen. nov. in life (males): N. annectens from Genting Highlands, Pahang, Malaysia (A), N. annamensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (B), N. arboricola from Chu Yang Sin N. P., Dak Lak, Vietnam (C), N. hongiaoensis from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (D), N. marmorata from Kon Chu Rang N. R., Gia Lai, Vietnam (E), N. nanapollexa from Kon Plong, Kon Tum, Vietnam (F), N. perparva from Gunung Mulu, Sarawak, Malaysia (G), N. petrigena from Gunung Mulu, Sarawak, Malaysia (H), and N. pulchella from Bidoup - Nui Ba N. P., Lam Dong, Vietnam (I). Insets show tympanic area of the each species; white arrow points at the tympanic rim of the external tympanum. Photos by Nikolay A. Poyarkov (A-S, D-F, I), Vu Dang Hoang Nguyen (D), and Indraneil Das (G, H)." figureDoi="10.3897/zse.97.57968.figure11" httpUri="https://binary.pensoft.net/fig/496425" pageId="0" pageNumber="21">11</figureCitation>
). In species of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="3AB31895E23E38CD29CDB8C033C38DCE" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="ABBE836E7D143B889EC52A9D34C71BE2" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov., smaller tubercles and other dermal structures of the skin become flattened and less distinct after fixation and preservation; this has also been reported in other anurans (
<bibRefCitation DOI="https://doi.org/10.1080/00222933.2020.1804005" author="Poyarkov, NA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="221 - 252" refId="B117" refString="Poyarkov, NA, Rowley, JJL, Gogoleva, SS, Vassilieva, AB, Galoyan, EA, Orlov, NL, 2015. A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam. Zootaxa 3931 (2): 221 - 252, DOI: https://doi.org/10.1080/00222933.2020.1804005" title="A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam." url="https://doi.org/10.1080/00222933.2020.1804005" volume="3931" year="2015">Poyarkov et al. 2015</bibRefCitation>
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,
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.672.10624" author="Poyarkov, NA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="21" pagination="49 - 120" refId="B116" refString="Poyarkov, NA, Duong, TV, Orlov, NL, Gogoleva, SS, Vassilieva, AB, Nguyen, LT, Nguyen, VHD, Nguyen, SN, Che, J, Mahony, S, 2017. Molecular, morphological and acoustic assessment of the genus Ophryophryne (Anura, Megophryidae) from Langbian Plateau, southern Vietnam, with description of a new species. ZooKeys 672: 49 - 120, DOI: https://doi.org/10.3897/zookeys.672.10624" title="Molecular, morphological and acoustic assessment of the genus Ophryophryne (Anura, Megophryidae) from Langbian Plateau, southern Vietnam, with description of a new species." url="https://doi.org/10.3897/zookeys.672.10624" volume="672" year="2017">2017</bibRefCitation>
<bibRefCitation id="0B0A09ABCFFACD76DF627F4EE11E07E3" DOI="https://doi.org/10.3897/zookeys.672.10624" author="Poyarkov, NA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="21" pagination="49 - 120" refId="B116" refString="Poyarkov, NA, Duong, TV, Orlov, NL, Gogoleva, SS, Vassilieva, AB, Nguyen, LT, Nguyen, VHD, Nguyen, SN, Che, J, Mahony, S, 2017. Molecular, morphological and acoustic assessment of the genus Ophryophryne (Anura, Megophryidae) from Langbian Plateau, southern Vietnam, with description of a new species. ZooKeys 672: 49 - 120, DOI: https://doi.org/10.3897/zookeys.672.10624" title="Molecular, morphological and acoustic assessment of the genus Ophryophryne (Anura, Megophryidae) from Langbian Plateau, southern Vietnam, with description of a new species." url="https://doi.org/10.3897/zookeys.672.10624" volume="672" year="2017">2017</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.24272/j.issn.2095-8137.2019.044" author="Poyarkov, NA" journalOrPublisher="Zoological Research" pageId="0" pageNumber="21" pagination="244 - 276" refId="B121" refString="Poyarkov, NA, Gorin, VA, Zaw, T, Kretova, VD, Gogoleva, SI, Pawangkhanant, P, Che, J, 2019. On the road to Mandalay: contribution to the Microhyla Tschudi, 1838 (Amphibia: Anura: Microhylidae) fauna of Myanmar with description of two new species. Zoological Research 40 (4): 244 - 276, DOI: https://doi.org/10.24272/j.issn.2095-8137.2019.044" title="On the road to Mandalay: contribution to the Microhyla Tschudi, 1838 (Amphibia: Anura: Microhylidae) fauna of Myanmar with description of two new species." url="https://doi.org/10.24272/j.issn.2095-8137.2019.044" volume="40" year="2019">2019</bibRefCitation>
<bibRefCitation id="5D0984B61260DDE61CEA87BE3FA935BD" DOI="https://doi.org/10.24272/j.issn.2095-8137.2019.044" author="Poyarkov, NA" journalOrPublisher="Zoological Research" pageId="0" pageNumber="21" pagination="244 - 276" refId="B121" refString="Poyarkov, NA, Gorin, VA, Zaw, T, Kretova, VD, Gogoleva, SI, Pawangkhanant, P, Che, J, 2019. On the road to Mandalay: contribution to the Microhyla Tschudi, 1838 (Amphibia: Anura: Microhylidae) fauna of Myanmar with description of two new species. Zoological Research 40 (4): 244 - 276, DOI: https://doi.org/10.24272/j.issn.2095-8137.2019.044" title="On the road to Mandalay: contribution to the Microhyla Tschudi, 1838 (Amphibia: Anura: Microhylidae) fauna of Myanmar with description of two new species." url="https://doi.org/10.24272/j.issn.2095-8137.2019.044" volume="40" year="2019">2019</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2009.01.014" author="Nguyen, LT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="1 - 21" refId="B101" refString="Nguyen, LT, Poyarkov, NA, Le DT, Vo BD, Phan, HT, Duong, TV, Murphy, RW, Nguyen, SN, 2018. A new species of Leptolalax (Anura: Megophryidae) from Son Tra Peninsula, central Vietnam. Zootaxa 4388 (1): 1 - 21, DOI: https://doi.org/10.1016/j.ympev.2009.01.014" title="A new species of Leptolalax (Anura: Megophryidae) from Son Tra Peninsula, central Vietnam." url="https://doi.org/10.1016/j.ympev.2009.01.014" volume="4388" year="2018">Nguyen et al. 2018</bibRefCitation>
<bibRefCitation id="3493C55D24F39F878A05174E6DB36027" DOI="https://doi.org/10.1016/j.ympev.2009.01.014" author="Nguyen, LT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="21" pagination="1 - 21" refId="B101" refString="Nguyen, LT, Poyarkov, NA, Le DT, Vo BD, Phan, HT, Duong, TV, Murphy, RW, Nguyen, SN, 2018. A new species of Leptolalax (Anura: Megophryidae) from Son Tra Peninsula, central Vietnam. Zootaxa 4388 (1): 1 - 21, DOI: https://doi.org/10.1016/j.ympev.2009.01.014" title="A new species of Leptolalax (Anura: Megophryidae) from Son Tra Peninsula, central Vietnam." url="https://doi.org/10.1016/j.ympev.2009.01.014" volume="4388" year="2018">Nguyen et al. 2018</bibRefCitation>
,
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,
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). It is likely that the presence of the tympanum was artifactually concealed from
<bibRefCitation DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen (2004)</bibRefCitation>
<bibRefCitation id="A5E58D7FB3B9D76D89D901FB01B3B782" DOI="https://doi.org/10.1643/CH-04-020R2" author="Bain, RH" journalOrPublisher="Copeia" pageId="0" pageNumber="21" pagination="507 - 524" refId="B7" refString="Bain, RH, Nguyen, QT, 2004. Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes. Copeia 2004 (3): 507 - 524, DOI: https://doi.org/10.1643/CH-04-020R2" title="Three new species of narrow-mouth frogs (Genus: Microhyla) from Indochina, with comments on Microhyla annamensis and Microhyla palmipes." url="https://doi.org/10.1643/CH-04-020R2" volume="2004" year="2004">Bain and Nguyen (2004)</bibRefCitation>
, since their description was based exclusively on museum specimens. In some species of
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="10FAB20F43CBFFD3D40B26A55CA3579E" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="9F080F5A7173327C0D4BA2356E179AB7" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov., we were not able to detect an externally visible tympanum (
<taxonomicName genus="N." lsidName="N. hongiaoensis" pageId="0" pageNumber="21" rank="species" species="hongiaoensis">
<emphasis italics="true" pageId="0" pageNumber="21">N. hongiaoensis</emphasis>
<taxonomicName id="0D93A73533BAC350428601EB279A1B1C" lsidName="N. hongiaoensis" pageId="0" pageNumber="21" rank="species" species="hongiaoensis">
<emphasis id="5C45CE558CAE2252B8D832C8F7F402AD" italics="true" pageId="0" pageNumber="21">N. hongiaoensis</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
<taxonomicName id="192F2F9AA8AE6181D6D9B43C6D9E03A7" lsidName="N. perparva" pageId="0" pageNumber="21" rank="species" species="perparva">
<emphasis id="23317C383E952A65695FAC1784B31FC9" italics="true" pageId="0" pageNumber="21">N. perparva</emphasis>
</taxonomicName>
,
<taxonomicName genus="N." lsidName="N. petrigena" pageId="0" pageNumber="21" rank="species" species="petrigena">
<emphasis italics="true" pageId="0" pageNumber="21">N. petrigena</emphasis>
<taxonomicName id="A13B62A1669C749197E6F434D35C12D5" lsidName="N. petrigena" pageId="0" pageNumber="21" rank="species" species="petrigena">
<emphasis id="26B5F4C728EDD9669C3CBC2883CED710" italics="true" pageId="0" pageNumber="21">N. petrigena</emphasis>
</taxonomicName>
). It is not clear whether this reflects an actual character state in these species, or if this apparent state relates to the small sample size of specimens and photographs available to us. Further studies are needed to clarify variation of the external tympanum in
<taxonomicName authority="Poyarkov, Gorin &amp; Scherz, 2021" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
<taxonomicName id="65AEA5198FB7095E3F20F80844872C42" authorityName="Poyarkov, Gorin &amp; Scherz" authorityYear="2021" class="Amphibia" family="Microhylidae" genus="Nanohyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanohyla" order="Anura" pageId="0" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="A9D09A4F15A53B27CEA0C032391C3C2B" italics="true" pageId="0" pageNumber="21">Nanohyla</emphasis>
</taxonomicName>
gen. nov.
</paragraph>

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@ -1,298 +1,298 @@
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<paragraph pageId="0" pageNumber="73">Untersuchte Exemplare.</paragraph>
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<paragraph id="6A285FAE546243EC330D269B1895B35B" pageId="0" pageNumber="73">Untersuchte Exemplare.</paragraph>
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FR
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, leg.
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; DNA Barcode
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;
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Gaedike Nr. 9727; coll.
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vom gleichen Fundort,
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Bryner Nr.
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(Abb.
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,
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): Spannweite 10-11 mm; Stirnschopf gelbbraun, Labialpalpus nach oben aufgebogen, die beiden basalen Glieder
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dunkelbraun, Innenseite fast
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, 2. Glied sehr lang, mit einzelnen Borsten, 3. Glied kurz, spitz, komplett
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; Antenne dunkelbraun, Scapus ohne Pecten; Thorax und Tegulae dunkelbraun, Spitzen der Tegulae gelbbraun;
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dunkelbraun, mit einer cremefarbenen (Abb.
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) oder hell rostbraunen Zeichnung (Abb.
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) in Form einer langgezogenen
<normalizedToken originalValue="Fläche">Flaeche</normalizedToken>
<normalizedToken id="C70BF298A242CA8F4791FC3303EFB00A" originalValue="Fläche">Flaeche</normalizedToken>
von der Basis zwischen Zelle und Dorsum bis zum Beginn der Fransen und einem dreieckigen Fleck bei 2/3 von der Costa bis zur Zelle, bei 1/3 in der Mitte ein fast schwarzer Strich; vor dem Apex an der Costa drei kurze cremefarbene Streifen; Fransen in der Mitte mit einer dunklen Schuppenlinie;
<normalizedToken originalValue="Hinterflügel">Hinterfluegel</normalizedToken>
<normalizedToken id="232C3CF7B87F9E258B237F16B63AB25A" originalValue="Hinterflügel">Hinterfluegel</normalizedToken>
dunkelbraun.
</paragraph>
<caption doi="10.3897/alpento.4.51727.abbildung5-6" httpUri="https://binary.pensoft.net/fig/412013" pageId="0" pageNumber="73" start="Abbildung 5, 6" startId="F2">
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" pageId="0" pageNumber="73">Abbildung 5, 6.</emphasis>
<taxonomicName authorityName="Gaedike &amp; Bryner" authorityYear="2020" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nemapogon helveticola" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="helveticola">
<emphasis italics="true" pageId="0" pageNumber="73">Nemapogon helveticola</emphasis>
<caption id="A583123E930636FA66E06C9A2A3E166E" doi="10.3897/alpento.4.51727.abbildung5-6" httpUri="https://binary.pensoft.net/fig/412013" pageId="0" pageNumber="73" start="Abbildung 5, 6" startId="F2">
<paragraph id="E3F1DAB33496A9AE438CFCA2625DEDC1" pageId="0" pageNumber="73">
<emphasis id="6BF2B30E409FA00669D2DB0FAA49AA66" bold="true" pageId="0" pageNumber="73">Abbildung 5, 6.</emphasis>
<taxonomicName id="2D819F37A144BDFB0051477BAA993670" authorityName="Gaedike &amp; Bryner" authorityYear="2020" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nemapogon helveticola" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="helveticola">
<emphasis id="2FE2C20B3B80158995F7472E6D915019" italics="true" pageId="0" pageNumber="73">Nemapogon helveticola</emphasis>
</taxonomicName>
sp. nov.: Imago (
<emphasis bold="true" pageId="0" pageNumber="73">5</emphasis>
<emphasis id="7C2F9E538DF7982257F5FF527E128A81" bold="true" pageId="0" pageNumber="73">5</emphasis>
- Holotypus;
<emphasis bold="true" pageId="0" pageNumber="73">6</emphasis>
<emphasis id="84FC614649A01F7274D54436F40A4427" bold="true" pageId="0" pageNumber="73">6</emphasis>
-
<taxonomicName genus="Paratypus" lsidName="Paratypus" pageId="0" pageNumber="73" rank="genus">Paratypus</taxonomicName>
<taxonomicName id="A27D7AD5E05A9648A66AC7FDCDA48FE2" genus="Paratypus" lsidName="Paratypus" pageId="0" pageNumber="73" rank="genus">Paratypus</taxonomicName>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="männliche genitalien">
<paragraph pageId="0" pageNumber="73">
<normalizedToken originalValue="Männliche">Maennliche</normalizedToken>
<subSubSection id="BD354880CCF340889940E63EE8168A1C" pageId="0" pageNumber="73" type="männliche genitalien">
<paragraph id="524CCCC714A09F55B80E3F1959AC768E" pageId="0" pageNumber="73">
<normalizedToken id="676D8D5F4A45325735D334C053D46204" originalValue="Männliche">Maennliche</normalizedToken>
Genitalien
</paragraph>
<paragraph pageId="0" pageNumber="73">
<paragraph id="2CA93DFFA5D77D75298F5F23F7D9EEAD" pageId="0" pageNumber="73">
(Abb.
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<figureCitation id="FF2D83443ABB35D2E1327321CB4CCC47" captionStart="Abbildung 7, 8" captionStartId="F3" captionText="Abbildung 7, 8. Nemapogon helveticola sp. nov.: maennliches Genital (7 - Holotypus; 8 - Paratypus)." figureDoi="10.3897/alpento.4.51727.abbildung7-8" httpUri="https://binary.pensoft.net/fig/412014" pageId="0" pageNumber="73">7</figureCitation>
,
<figureCitation captionStart="Abbildung 7, 8" captionStartId="F3" captionText="Abbildung 7, 8. Nemapogon helveticola sp. nov.: maennliches Genital (7 - Holotypus; 8 - Paratypus)." figureDoi="10.3897/alpento.4.51727.abbildung7-8" httpUri="https://binary.pensoft.net/fig/412014" pageId="0" pageNumber="73">8</figureCitation>
<figureCitation id="D11C3CCC743EA95156B55176B179FB0A" captionStart="Abbildung 7, 8" captionStartId="F3" captionText="Abbildung 7, 8. Nemapogon helveticola sp. nov.: maennliches Genital (7 - Holotypus; 8 - Paratypus)." figureDoi="10.3897/alpento.4.51727.abbildung7-8" httpUri="https://binary.pensoft.net/fig/412014" pageId="0" pageNumber="73">8</figureCitation>
): Uncus mit zwei spitzen, beborsteten Socii; lateral je ein mit
<normalizedToken originalValue="kräftigen">kraeftigen</normalizedToken>
<normalizedToken id="9A8B2ED51226DD06B2E79AF4FFFF25B2" originalValue="kräftigen">kraeftigen</normalizedToken>
Borsten besetzter Fortsatz, aufgeklappt (siehe Abb.
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<figureCitation id="61B67C601ADCCBA80C0118F8A82523CE" captionStart="Abbildung 7, 8" captionStartId="F3" captionText="Abbildung 7, 8. Nemapogon helveticola sp. nov.: maennliches Genital (7 - Holotypus; 8 - Paratypus)." figureDoi="10.3897/alpento.4.51727.abbildung7-8" httpUri="https://binary.pensoft.net/fig/412014" pageId="0" pageNumber="73">8</figureCitation>
) abgerundet; unterhalb des beborsteten Fortsatzes eine kleine deutlich
<normalizedToken originalValue="stärker">staerker</normalizedToken>
<normalizedToken id="A124707987161708B1E178F8499A1C25" originalValue="stärker">staerker</normalizedToken>
sklerotisierte
<normalizedToken originalValue="Fläche">Flaeche</normalizedToken>
<normalizedToken id="DEF5A9E540A04E077BE60D26396E7493" originalValue="Fläche">Flaeche</normalizedToken>
, die vielleicht als Gnathosrest gedeutet werden
<normalizedToken originalValue="könnte">koennte</normalizedToken>
<normalizedToken id="251C1D7E2227F9FCEFAE809DBDCB0419" originalValue="könnte">koennte</normalizedToken>
; Tegumen und Vinculum breit, Saccus schmal, relativ kurz; Valve mehr oder weniger parallelseitig, apikale
<normalizedToken originalValue="Hälfte">Haelfte</normalizedToken>
<normalizedToken id="9B2233B279267E27A4538A3A43030B20" originalValue="Hälfte">Haelfte</normalizedToken>
schmaler, Apex stumpf, Ecken abgrundet, subapikal auf der Innenseite mit einem kurzen, stark sklerotisierten dreieckigen Zahn; Transtilla fast halb so lang wie die Valve, Digitus schmal, beborstet; Phallus etwas
<normalizedToken originalValue="länger">laenger</normalizedToken>
<normalizedToken id="7FE828621B78C59496C36C39FD77C9EE" originalValue="länger">laenger</normalizedToken>
als Valve + Transtilla, apikal mit sehr feiner Bedornung, hinter der Basis etwas abgesetzt.
</paragraph>
<caption doi="10.3897/alpento.4.51727.abbildung7-8" httpUri="https://binary.pensoft.net/fig/412014" pageId="0" pageNumber="73" start="Abbildung 7, 8" startId="F3">
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" pageId="0" pageNumber="73">Abbildung 7, 8.</emphasis>
<taxonomicName authorityName="Gaedike &amp; Bryner" authorityYear="2020" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nemapogon helveticola" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="helveticola">
<emphasis italics="true" pageId="0" pageNumber="73">Nemapogon helveticola</emphasis>
<caption id="A3DF02E4500FDB81B19540B773ECCA7A" doi="10.3897/alpento.4.51727.abbildung7-8" httpUri="https://binary.pensoft.net/fig/412014" pageId="0" pageNumber="73" start="Abbildung 7, 8" startId="F3">
<paragraph id="17EC965BDC5C575EFD91A3DFEC0E6ACD" pageId="0" pageNumber="73">
<emphasis id="DEE8BD77E48C55FD3535BD6A7E4358EA" bold="true" pageId="0" pageNumber="73">Abbildung 7, 8.</emphasis>
<taxonomicName id="22E734B31CD02E8D7605A96EACF414E3" authorityName="Gaedike &amp; Bryner" authorityYear="2020" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nemapogon helveticola" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="helveticola">
<emphasis id="86248EF31AE955891AAA72D37DB8D204" italics="true" pageId="0" pageNumber="73">Nemapogon helveticola</emphasis>
</taxonomicName>
sp. nov.:
<normalizedToken originalValue="männliches">maennliches</normalizedToken>
<normalizedToken id="76AE6A075C02BB78D0A9C228873277DC" originalValue="männliches">maennliches</normalizedToken>
Genital (
<emphasis bold="true" pageId="0" pageNumber="73">7</emphasis>
<emphasis id="286E262122AF0879EFE6527FEF4411AF" bold="true" pageId="0" pageNumber="73">7</emphasis>
- Holotypus;
<emphasis bold="true" pageId="0" pageNumber="73">8</emphasis>
<emphasis id="3E08779BC50EA373B7636AF154E06EB8" bold="true" pageId="0" pageNumber="73">8</emphasis>
-
<taxonomicName genus="Paratypus" lsidName="Paratypus" pageId="0" pageNumber="73" rank="genus">Paratypus</taxonomicName>
<taxonomicName id="20D13936BB7574CBAEA7360786E66F33" genus="Paratypus" lsidName="Paratypus" pageId="0" pageNumber="73" rank="genus">Paratypus</taxonomicName>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="weibliche genitalien">
<paragraph pageId="0" pageNumber="73">Weibliche Genitalien.</paragraph>
<paragraph pageId="0" pageNumber="73">Unbekannt.</paragraph>
<subSubSection id="43A34DCCEDC9D2B6305FC045323F7925" pageId="0" pageNumber="73" type="weibliche genitalien">
<paragraph id="0710012F077FECF72132BB4C44FEA2EF" pageId="0" pageNumber="73">Weibliche Genitalien.</paragraph>
<paragraph id="3F8459CEB390393617A4F1FACDE7D782" pageId="0" pageNumber="73">Unbekannt.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="differentialdiagnose">
<paragraph pageId="0" pageNumber="73">Differentialdiagnose.</paragraph>
<paragraph pageId="0" pageNumber="73">
<subSubSection id="026116EEB1BEC7825F0FA32D2B0705DC" pageId="0" pageNumber="73" type="differentialdiagnose">
<paragraph id="4E57CC9B78D85E8B0D73DD81CBAA4105" pageId="0" pageNumber="73">Differentialdiagnose.</paragraph>
<paragraph id="D19B55DBCFB58A071531CE0879947DB8" pageId="0" pageNumber="73">
Die
<normalizedToken originalValue="für">fuer</normalizedToken>
<normalizedToken id="93CE98455D6BB801AB02E6FDF8436CD2" originalValue="für">fuer</normalizedToken>
fast alle Arten der Gattung typische
<normalizedToken originalValue="Flügelzeichnung">Fluegelzeichnung</normalizedToken>
<normalizedToken id="83E545F4E3A3BA68A9CB15888E62D10E" originalValue="Flügelzeichnung">Fluegelzeichnung</normalizedToken>
aus zumeist hellen (oft
<normalizedToken originalValue="weißen">weissen</normalizedToken>
<normalizedToken id="F78B60E94667D14E7162C78A4D87F35A" originalValue="weißen">weissen</normalizedToken>
bis cremefarbenen) sowie dunklen (gelbbraunen, braunen bis schwarzen) Flecken und
<normalizedToken originalValue="bindenähnlichen">bindenaehnlichen</normalizedToken>
<normalizedToken id="F4C367F1901162CE019B422BC0797A29" originalValue="bindenähnlichen">bindenaehnlichen</normalizedToken>
Streifen ist hier fast vollkommen
<normalizedToken originalValue="überdeckt">ueberdeckt</normalizedToken>
<normalizedToken id="CED065CD25511DB2359BB09D2589004C" originalValue="überdeckt">ueberdeckt</normalizedToken>
mit dunkelbraunen Schuppen. Gewisse
<normalizedToken originalValue="Ähnlichkeiten">Aehnlichkeiten</normalizedToken>
<normalizedToken id="4C22FE2F8BAC3DC282FAA819E0F95235" originalValue="Ähnlichkeiten">Aehnlichkeiten</normalizedToken>
sind mit den beiden Arten
<taxonomicName genus="N." lsidName="N. cloacella" pageId="0" pageNumber="73" rank="species" species="cloacella">
<emphasis italics="true" pageId="0" pageNumber="73">N. cloacella</emphasis>
<taxonomicName id="8946C604AC56B57235370197006C8F5C" lsidName="N. cloacella" pageId="0" pageNumber="73" rank="species" species="cloacella">
<emphasis id="D484E545F0608DF0F218A522DC3C09CF" italics="true" pageId="0" pageNumber="73">N. cloacella</emphasis>
</taxonomicName>
(Haworth, 1828) und
<taxonomicName genus="N." lsidName="N. koenigi" pageId="0" pageNumber="73" rank="species" species="koenigi">
<emphasis italics="true" pageId="0" pageNumber="73">N. koenigi</emphasis>
<taxonomicName id="A244D30B9FF728EBB562FD5D0F42141A" lsidName="N. koenigi" pageId="0" pageNumber="73" rank="species" species="koenigi">
<emphasis id="F283DA0AE74F16FF0038749F48438AAA" italics="true" pageId="0" pageNumber="73">N. koenigi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Capuşe">Capuse</normalizedToken>
<normalizedToken id="F16AB7ED05D7752515C5729B03C16CED" originalValue="Capuşe">Capuse</normalizedToken>
, 1967 vorhanden, bei denen aber die Zeichnungselemente deutlicher sichtbar sind.
</paragraph>
<paragraph pageId="0" pageNumber="73">
<paragraph id="C3CEDDF44F11B621359990971B3B506A" pageId="0" pageNumber="73">
Im Genitalbau ist das charakteristische Merkmal der Gattung, die Valve mit Digitus, vorhanden. Das Fehlen von Gnathosarmen ist untypisch, bisher ist nur
<taxonomicName authorityName="Gaedike" authorityYear="2000" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nemapogon agnathosella" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="agnathosella">
<emphasis italics="true" pageId="0" pageNumber="73">Nemapogon agnathosella</emphasis>
<taxonomicName id="F8DB9E792A18A6DC2AE22F32E09104E5" authorityName="Gaedike" authorityYear="2000" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nemapogon agnathosella" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="agnathosella">
<emphasis id="E94B7CCA2C0782611F81319641D111C2" italics="true" pageId="0" pageNumber="73">Nemapogon agnathosella</emphasis>
</taxonomicName>
Gaedike, 2000 aus dem Fernen Osten Russlands bekannt, wo auch keine Gnathosarme vorhanden sind. Es
<normalizedToken originalValue="wäre">waere</normalizedToken>
<normalizedToken originalValue="möglich">moeglich</normalizedToken>
<normalizedToken id="80261607C6463D2FB24C07F012B69442" originalValue="wäre">waere</normalizedToken>
<normalizedToken id="7623FF206994893638F42F09FF3D51D0" originalValue="möglich">moeglich</normalizedToken>
, dass die beiden lateralen beborsteten
<normalizedToken originalValue="Fortsätze">Fortsaetze</normalizedToken>
<normalizedToken id="C2D1698C438DD62D4361DBAB701DF2DA" originalValue="Fortsätze">Fortsaetze</normalizedToken>
die Funktion der Gnathosarme
<normalizedToken originalValue="übernommen">uebernommen</normalizedToken>
<normalizedToken id="F658401ECB045A2C15BA7EDE0E0E272D" originalValue="übernommen">uebernommen</normalizedToken>
haben. Bei einigen Arten (
<taxonomicName genus="N." lsidName="N. falstriella" pageId="0" pageNumber="73" rank="species" species="falstriella">
<emphasis italics="true" pageId="0" pageNumber="73">N. falstriella</emphasis>
<taxonomicName id="C6944D5F65683F0F174BDDC5C135CDD7" lsidName="N. falstriella" pageId="0" pageNumber="73" rank="species" species="falstriella">
<emphasis id="FC6D8EDD8E91A1AEA33405F6FFA095D9" italics="true" pageId="0" pageNumber="73">N. falstriella</emphasis>
</taxonomicName>
(Haas, 1881),
<taxonomicName genus="N." lsidName="N. caucasicus" pageId="0" pageNumber="73" rank="species" species="caucasicus">
<emphasis italics="true" pageId="0" pageNumber="73">N. caucasicus</emphasis>
<taxonomicName id="C46B5DDFC2B98D32DCCD890D04E5FE1E" lsidName="N. caucasicus" pageId="0" pageNumber="73" rank="species" species="caucasicus">
<emphasis id="EEE558F4375CE427B96E982E9830CAF5" italics="true" pageId="0" pageNumber="73">N. caucasicus</emphasis>
</taxonomicName>
Zagulajev, 1964 und
<taxonomicName genus="N." lsidName="N. echinata" pageId="0" pageNumber="73" rank="species" species="echinata">
<emphasis italics="true" pageId="0" pageNumber="73">N. echinata</emphasis>
<taxonomicName id="491509DF734BEF91533E3C330081A962" lsidName="N. echinata" pageId="0" pageNumber="73" rank="species" species="echinata">
<emphasis id="09216F13F00693AA9B22FE5492B806E9" italics="true" pageId="0" pageNumber="73">N. echinata</emphasis>
</taxonomicName>
Gaedike, 2000) sind laterale bestachelte
<normalizedToken originalValue="Uncusfortsätze">Uncusfortsaetze</normalizedToken>
<normalizedToken id="E5D7FA4519C7A72409D9AAC88512B777" originalValue="Uncusfortsätze">Uncusfortsaetze</normalizedToken>
(Socii) vorhanden, die denen bei
<taxonomicName genus="N." lsidName="N. helveticola" pageId="0" pageNumber="73" rank="species" species="helveticola">
<emphasis italics="true" pageId="0" pageNumber="73">N. helveticola</emphasis>
<taxonomicName id="73CD7595BCD98DC1368AF5E32D4968B5" lsidName="N. helveticola" pageId="0" pageNumber="73" rank="species" species="helveticola">
<emphasis id="55CFB2DB191F46B754EC623C62DBD9FC" italics="true" pageId="0" pageNumber="73">N. helveticola</emphasis>
</taxonomicName>
<normalizedToken originalValue="ähneln">aehneln</normalizedToken>
<normalizedToken id="1F3E7030074A6C62ADA81CC050CFE5BE" originalValue="ähneln">aehneln</normalizedToken>
, diese Arten haben aber alle Gnathosarme (
<bibRefCitation DOI="https://doi.org/10.1163/9789004289161_002" author="Gaedike, R" pageId="0" pageNumber="73" refId="B1" refString="Gaedike, R, 2015. Tineidae I (Dryadaulinae, Hapsiferinae, Euplocaminae, Scardiinae, Nemapogoninae and Meessiinae). - Brill NV, Leiden: 1-308. [6 col. pls, numerous b/w figs (Microlepidoptera of Europe 7)] https://doi.org/10.1163/9789004289161_002" title="Tineidae I (Dryadaulinae, Hapsiferinae, Euplocaminae, Scardiinae, Nemapogoninae and Meessiinae). - Brill NV, Leiden: 1 - 308. [6 col. pls, numerous b / w figs (Microlepidoptera of Europe 7)]" url="https://doi.org/10.1163/9789004289161_002" year="2015">Gaedike 2015</bibRefCitation>
<bibRefCitation id="7BE325C67C42B179F7BB75E402669F18" DOI="https://doi.org/10.1163/9789004289161_002" author="Gaedike, R" pageId="0" pageNumber="73" refId="B1" refString="Gaedike, R, 2015. Tineidae I (Dryadaulinae, Hapsiferinae, Euplocaminae, Scardiinae, Nemapogoninae and Meessiinae). - Brill NV, Leiden: 1-308. [6 col. pls, numerous b/w figs (Microlepidoptera of Europe 7)] https://doi.org/10.1163/9789004289161_002" title="Tineidae I (Dryadaulinae, Hapsiferinae, Euplocaminae, Scardiinae, Nemapogoninae and Meessiinae). - Brill NV, Leiden: 1 - 308. [6 col. pls, numerous b / w figs (Microlepidoptera of Europe 7)]" url="https://doi.org/10.1163/9789004289161_002" year="2015">Gaedike 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="dna barcode">
<paragraph pageId="0" pageNumber="73">DNA Barcode</paragraph>
<paragraph pageId="0" pageNumber="73">
<subSubSection id="7DA2C80A34FD79F402A786EF7FE1FEFF" pageId="0" pageNumber="73" type="dna barcode">
<paragraph id="8BB3CDBBFB315D2B5FD387C17FBFD161" pageId="0" pageNumber="73">DNA Barcode</paragraph>
<paragraph id="98E3C3ECF9BB33DE51D3A2176081FFB4" pageId="0" pageNumber="73">
(
<normalizedToken originalValue="persönliche">persoenliche</normalizedToken>
<normalizedToken id="8E9C0A09DDE922A908FFFAD9375BEF02" originalValue="persönliche">persoenliche</normalizedToken>
Mitteilung von Peter Huemer): Vom Holotypus konnte eine DNA-Barcode Sequenz von 641 Basenpaaren ermittelt werden. Die genetisch
<normalizedToken originalValue="nächst">naechst</normalizedToken>
<normalizedToken id="65E487780BD256F0E2C0D4258B405117" originalValue="nächst">naechst</normalizedToken>
stehenden, erfolgreich sequenzierten
<normalizedToken originalValue="europäischen">europaeischen</normalizedToken>
<normalizedToken id="AF7042FF79FBF8296810BD73EAA8EC4F" originalValue="europäischen">europaeischen</normalizedToken>
Arten in BOLD sind
<taxonomicName genus="N." lsidName="N. lagodechiellus" pageId="0" pageNumber="73" rank="species" species="lagodechiellus">
<emphasis italics="true" pageId="0" pageNumber="73">N. lagodechiellus</emphasis>
<taxonomicName id="77BEAC9CB180E3490C7A69F4C4421E72" lsidName="N. lagodechiellus" pageId="0" pageNumber="73" rank="species" species="lagodechiellus">
<emphasis id="65A5046B332E67DF667CC0CFF0070CFC" italics="true" pageId="0" pageNumber="73">N. lagodechiellus</emphasis>
</taxonomicName>
Zagulajev, 1962,
<taxonomicName genus="N." lsidName="N. orientalis" pageId="0" pageNumber="73" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="73">N. orientalis</emphasis>
<taxonomicName id="F6B4BD816DBCD92057BF13F37EBBCEFE" lsidName="N. orientalis" pageId="0" pageNumber="73" rank="species" species="orientalis">
<emphasis id="34B6AAC023E210AA40C34668130B486F" italics="true" pageId="0" pageNumber="73">N. orientalis</emphasis>
</taxonomicName>
Petersen, 1961 und
<taxonomicName genus="N." lsidName="N. cloacella" pageId="0" pageNumber="73" rank="species" species="cloacella">
<emphasis italics="true" pageId="0" pageNumber="73">N. cloacella</emphasis>
<taxonomicName id="2AC45749E4F01F6AA9FA3DFDD1E4F5C0" lsidName="N. cloacella" pageId="0" pageNumber="73" rank="species" species="cloacella">
<emphasis id="031FC15C91B589836A7305680412F773" italics="true" pageId="0" pageNumber="73">N. cloacella</emphasis>
</taxonomicName>
(Haworth, 1828) mit jeweils etwa 9 % minimaler Distanz. Die Distanzen zum
<normalizedToken originalValue="nächsten">naechsten</normalizedToken>
<normalizedToken id="F2DC02E664BB848F4BD8B65136F3D489" originalValue="nächsten">naechsten</normalizedToken>
Nachbarn betragen hingegen bei 32 Arten in BOLD im Vergleich lediglich durchschnittlich 5,97% (min. 3,3%, max. 12.29%). Die genetische Sequenz der neuen Art sowie geographische Informationen und Bildmaterial finden sich im
<normalizedToken originalValue="öffentlichen">oeffentlichen</normalizedToken>
<normalizedToken id="FAF01FC0AAD533534FFE5C83ED2BA9A4" originalValue="öffentlichen">oeffentlichen</normalizedToken>
Dataset DS-NEMAHELV &quot;
<taxonomicName authorityName="Gaedike &amp; Bryner" authorityYear="2020" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nemapogon helveticola" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="helveticola">Nemapogon helveticola</taxonomicName>
<taxonomicName id="F15A33B943737C08B06FD0323ACFD575" authorityName="Gaedike &amp; Bryner" authorityYear="2020" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nemapogon helveticola" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="helveticola">Nemapogon helveticola</taxonomicName>
sp. nov.&quot; in BOLD (www.boldsystems.org).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="etymology">
<paragraph pageId="0" pageNumber="73">Derivatio nominis.</paragraph>
<paragraph pageId="0" pageNumber="73">
<subSubSection id="55BA3E4CD0436BFEC00939223FD7B827" pageId="0" pageNumber="73" type="etymology">
<paragraph id="01586F75BE74E3E10A3AFE3AA0BD487E" pageId="0" pageNumber="73">Derivatio nominis.</paragraph>
<paragraph id="0A69378B41E3A4E55EA4D3746B0E9FF0" pageId="0" pageNumber="73">
Der Name
<taxonomicName authorityName="Gaedike &amp; Bryner" authorityYear="2020" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="helveticola" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="helveticola">
<emphasis italics="true" pageId="0" pageNumber="73">helveticola</emphasis>
<taxonomicName id="E88DCAC4C75BF0CFA2EE03A5DE2A2B7A" authorityName="Gaedike &amp; Bryner" authorityYear="2020" class="Insecta" family="Tineidae" genus="Nemapogon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="helveticola" order="Lepidoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="helveticola">
<emphasis id="4ABA1B871DCB67CD39732CD661AF3132" italics="true" pageId="0" pageNumber="73">helveticola</emphasis>
</taxonomicName>
ist von
<normalizedToken originalValue="„Helvetia“">&quot;Helvetia&quot;</normalizedToken>
<normalizedToken id="F7C79318023B636678960C80A423CE9E" originalValue="„Helvetia“">&quot;Helvetia&quot;</normalizedToken>
, dem Land des Erstfundes, abgeleitet.
</paragraph>
</subSubSection>

View file

@ -1,163 +1,163 @@
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<mods:title id="5BBA356844482ACB1698DC4F45B6AB0F">Halocoryza Alluaud 1919, sea-side beetles of the Indian, Atlantic (sensu lato), and Pacific Oceans: a generic synopsis and description of a remarkable new species from Baja California Sur, Mexico (Coleoptera, Carabidae, Scaritini, Clivinina)</mods:title>
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<treatment ID-GBIF-Taxon="152030631" LSID="urn:lsid:plazi:treatment:C4881D51D1D9D7E852050143CA0A9778" httpUri="http://treatment.plazi.org/id/C4881D51D1D9D7E852050143CA0A9778" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection pageId="3" pageNumber="4" type="nomenclature">
<paragraph pageId="3" pageNumber="4">
<taxonomicName LSID="C4881D51-D1D9-D7E8-5205-0143CA0A9778" authority="(Darlington, 1939)" authorityName="Darlington" authorityYear="1939" baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halocoryza arenaria" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="arenaria">Halocoryza arenaria (Darlington, 1939)</taxonomicName>
<figureCitation captionStart="" captionStartId="F1" captionText=". Figure 1 - 2. 1 Color image, showing habitus of Halocoryza acapulcana Whitehead, 1966, dorsal aspect, ABL = 2.4 mm, ADP 127171; Acapulco, Mexico 2 Color image, showing habitus of Halocoryza arenaria (Darlington, 1939), dorsal aspect, ABL = 2.9 mm, ADP 116798; St. Johns, Virgin Islands." httpUri="https://binary.pensoft.net/fig/17265" pageId="3" pageNumber="4">Figs 2</figureCitation>
<figureCitation captionStart="Figure 48" captionStartId="F3" captionText="Figure 4 - 8. 4 Line drawing, showing habitus of Halocoryza acapulcana Whitehead, 1966, dorsal aspect, ABL = 2.4 mm; Acapulco, Mexico; modified from Whitehead 1966 5 Line drawing, showing habitus of Halocoryza arenaria (Darlington, 1939), dorsal aspect, ABL = 2.9 mm; Rio Piedras, Puerto Rico; modified from Whitehead, 1966 6 Line drawing, showing habitus of Halocoryza whiteheadiana sp. n., dorsal aspect, ABL = 2.9 mm; ABL = 2.8 mm, ADP 127139; Isla Del Carmen, Mexico 7 Line drawing, showing male genitalia, median lobe, of Halocoryza acapulcana Whitehead, 1966, left lateral aspect, 0.41 mm (dorsal margin of basal lobe to apex); Acapulco, Mexico; modified from Whitehead 1966 8 Line drawing, showing male genitalia, median lobe, Halocoryza arenaria (Darlington, 1939), left lateral aspect with internal sac everted, 0.41 mm (dorsal margin of basal lobe to apex); Rio Piedras, Puerto Rico; modified from Whitehead, 1966." httpUri="https://binary.pensoft.net/fig/17267" pageId="3" pageNumber="4">5, 8</figureCitation>
<treatment id="C4881D51D1D9D7E852050143CA0A9778" ID-GBIF-Taxon="152030631" LSID="urn:lsid:plazi:treatment:C4881D51D1D9D7E852050143CA0A9778" httpUri="http://treatment.plazi.org/id/C4881D51D1D9D7E852050143CA0A9778" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Clivinina" scope_tribe="Scaritini">
<subSubSection id="8AC7CA360418F50821D4DC07D2665AEE" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="615D40F0FC28AA771F02F9CFDB915A80" pageId="3" pageNumber="4">
<taxonomicName id="875106AEF47D30F5E9169084827C44A9" ID-CoL="3JF3P" LSID="C4881D51-D1D9-D7E8-5205-0143CA0A9778" authority="(Darlington, 1939)" authorityName="Darlington" authorityYear="1939" baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halocoryza arenaria" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="arenaria">Halocoryza arenaria (Darlington, 1939)</taxonomicName>
<figureCitation id="271FE9BAC8C5C559A544E91C0A4698E1" captionStart="" captionStartId="F1" captionText=". Figure 1 - 2. 1 Color image, showing habitus of Halocoryza acapulcana Whitehead, 1966, dorsal aspect, ABL = 2.4 mm, ADP 127171; Acapulco, Mexico 2 Color image, showing habitus of Halocoryza arenaria (Darlington, 1939), dorsal aspect, ABL = 2.9 mm, ADP 116798; St. Johns, Virgin Islands." httpUri="https://binary.pensoft.net/fig/17265" pageId="3" pageNumber="4">Figs 2</figureCitation>
<figureCitation id="0E4A74CC2E58E00BB42B5EEF8F36A583" captionStart="Figure 48" captionStartId="F3" captionText="Figure 4 - 8. 4 Line drawing, showing habitus of Halocoryza acapulcana Whitehead, 1966, dorsal aspect, ABL = 2.4 mm; Acapulco, Mexico; modified from Whitehead 1966 5 Line drawing, showing habitus of Halocoryza arenaria (Darlington, 1939), dorsal aspect, ABL = 2.9 mm; Rio Piedras, Puerto Rico; modified from Whitehead, 1966 6 Line drawing, showing habitus of Halocoryza whiteheadiana sp. n., dorsal aspect, ABL = 2.9 mm; ABL = 2.8 mm, ADP 127139; Isla Del Carmen, Mexico 7 Line drawing, showing male genitalia, median lobe, of Halocoryza acapulcana Whitehead, 1966, left lateral aspect, 0.41 mm (dorsal margin of basal lobe to apex); Acapulco, Mexico; modified from Whitehead 1966 8 Line drawing, showing male genitalia, median lobe, Halocoryza arenaria (Darlington, 1939), left lateral aspect with internal sac everted, 0.41 mm (dorsal margin of basal lobe to apex); Rio Piedras, Puerto Rico; modified from Whitehead, 1966." httpUri="https://binary.pensoft.net/fig/17267" pageId="3" pageNumber="4">5, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="reference_group">
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Carabidae" genus="Schizogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="arenaria">Schizogenius arenaria</taxonomicName>
<subSubSection id="C5E3917AD682A420C8AA460E6169B050" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="F89A2546BC24DCE9225FDD91F3DE19E3" pageId="3" pageNumber="4">
<taxonomicName id="E6FF89E6BD46D992F7E28FDCCC25AF88" class="Insecta" family="Carabidae" genus="Schizogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="arenaria">Schizogenius arenaria</taxonomicName>
Darlington, 1939: 84
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="common name">
<paragraph pageId="3" pageNumber="4">Common name.</paragraph>
<paragraph pageId="3" pageNumber="4">
<emphasis bold="true" pageId="3" pageNumber="4">Sand Saline Catarrh Beetle</emphasis>
<subSubSection id="F6030C0344EB39E761B6D102E236E717" pageId="3" pageNumber="4" type="common name">
<paragraph id="8BA9532ADFA9935A23159782DA974AB8" pageId="3" pageNumber="4">Common name.</paragraph>
<paragraph id="D033627FA04EB544457D6FE65109D4A9" pageId="3" pageNumber="4">
<emphasis id="C7F109C12376294A8D51EE08DC10C232" bold="true" pageId="3" pageNumber="4">Sand Saline Catarrh Beetle</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="geographic distribution">
<paragraph pageId="3" pageNumber="4">Geographic Distribution.</paragraph>
<paragraph pageId="3" pageNumber="4">
<subSubSection id="5544728C55D3C37EE4A6F7F966EA17E3" pageId="3" pageNumber="4" type="geographic distribution">
<paragraph id="07F2111A476B644A2FF6CE8BEA47128B" pageId="3" pageNumber="4">Geographic Distribution.</paragraph>
<paragraph id="12D634256BDA7E90E66144D2AAA5C740" pageId="3" pageNumber="4">
Native, New World. Barbados; Brazil; Dominican Republic; Grenada; Guadeloupe; Jamaica;
<normalizedToken originalValue="México">Mexico</normalizedToken>
<normalizedToken id="AA8FD9DBC1563D70368B1B4492035130" originalValue="México">Mexico</normalizedToken>
- QR, YC;
<normalizedToken originalValue="Panamá">Panama</normalizedToken>
<normalizedToken id="0CB61A5FD3D6E90227280C9C4BBACD62" originalValue="Panamá">Panama</normalizedToken>
; Puerto Rico; USA - FL; Virgin Islands - St. John, St. Thomas; natural invasive, Africa - Cameroon.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="way of life">
<paragraph pageId="3" pageNumber="4">Way of Life.</paragraph>
<paragraph lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis bold="true" pageId="3" pageNumber="4">Macrohabitat:</emphasis>
<subSubSection id="95032CE150F34576D1C35E54D07EFA33" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="way of life">
<paragraph id="52FAA7D7284F8E4306A8EC6F4C70AF1B" pageId="3" pageNumber="4">Way of Life.</paragraph>
<paragraph id="C690270CAE36AD9BED8ED271B32444F8" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="94EEA1B6BBE5625C44B4350FCF69454F" bold="true" pageId="3" pageNumber="4">Macrohabitat:</emphasis>
Lowlands, sea level - 1 meter altitude, on sea beaches and in the intertidal area, at or near the high tide line, and in mangrove swamps.
<emphasis bold="true" pageId="4" pageNumber="5">
<pageBreakToken pageId="4" pageNumber="5" start="start">Microhabitat</pageBreakToken>
<emphasis id="39B61A185CB870A1D84C28AAB01EE124" bold="true" pageId="4" pageNumber="5">
<pageBreakToken id="32C6A349DFC2590B5A99BF401AE34CAC" pageId="4" pageNumber="5" start="start">Microhabitat</pageBreakToken>
:
</emphasis>
Adults are ground-dwelling on exposed wet substrate consisting of coquina-coral cemented by very fine silt or sand and covered with seaweed mats.
<emphasis bold="true" pageId="4" pageNumber="5">Dispersal abilities:</emphasis>
<emphasis id="C93D9E8330A968B435624656A86FD91A" bold="true" pageId="4" pageNumber="5">Dispersal abilities:</emphasis>
Wing-polymorphic: macropterous form probably capable of flight; brachypterous form, consequently flightless thus vagility limited to walking or running; both forms slow runners.
<emphasis bold="true" pageId="4" pageNumber="5">Seasonal occurrence:</emphasis>
<emphasis id="2221B7F15A2396ACD26D5499BBACE006" bold="true" pageId="4" pageNumber="5">Seasonal occurrence:</emphasis>
Adults have been found in March - April, July, and October.
<emphasis bold="true" pageId="4" pageNumber="5">Behavior:</emphasis>
<emphasis id="0C9395C23B1A9D9A69A720E9DF7841A1" bold="true" pageId="4" pageNumber="5">Behavior:</emphasis>
Adults are nocturnal predaceous halobionts and take cover in the sand or under drift and piles of seaweed on the beach. Populations of this species are associated with the centipede
<taxonomicName authorityName="Crabill" authorityYear="1959" class="Chilopoda" family="Schendylidae" genus="Pectiniunguis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Geophilomorpha" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="halirrhytus">
<emphasis italics="true" pageId="4" pageNumber="5">Pectiniunguis halirrhytus</emphasis>
<taxonomicName id="A0954170B52C49F138B7E99DCFE3776A" authorityName="Crabill" authorityYear="1959" class="Chilopoda" family="Schendylidae" genus="Pectiniunguis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Geophilomorpha" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="halirrhytus">
<emphasis id="AD3846B3AD7EC80D8439BEF3FB1C6E23" italics="true" pageId="4" pageNumber="5">Pectiniunguis halirrhytus</emphasis>
</taxonomicName>
Crabill. In the northern part of their range, adults overwinter in the substrate; in the southern part, they likely aestivate during the dry season in the substrate.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/17266" pageId="4" pageNumber="5" start="Figure 3" startId="F2">
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" pageId="4" pageNumber="5">Figure 3.</emphasis>
<caption id="E9B548E355549EB18D3D9BF6F44B39F2" httpUri="https://binary.pensoft.net/fig/17266" pageId="4" pageNumber="5" start="Figure 3" startId="F2">
<paragraph id="3649C803777E373BD8F79CD260E101AC" pageId="4" pageNumber="5">
<emphasis id="AACBFC4717CF5F52B2BA7F40DA4AEB58" bold="true" pageId="4" pageNumber="5">Figure 3.</emphasis>
Color image, showing habitus of
<taxonomicName authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="whiteheadiana">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza whiteheadiana</emphasis>
<taxonomicName id="929193C4E83B89F21BACC0C55B78188D" authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="whiteheadiana">
<emphasis id="0EA2937EA26800E1D7D6DA416AA02231" italics="true" pageId="4" pageNumber="5">Halocoryza whiteheadiana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="5">sp. n.</emphasis>
<emphasis id="7BCD6AA8ABD44D605EE18CC775D845F2" bold="true" pageId="4" pageNumber="5">sp. n.</emphasis>
, 3a, left lateral aspect, 3b, dorsal aspect, ABL = 2.9mm; Holotype: Isla Del Carmen, BJ,
<normalizedToken originalValue="México">Mexico</normalizedToken>
<normalizedToken id="CA6572A50B93234F04E0AC2C4FEEF985" originalValue="México">Mexico</normalizedToken>
.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/17267" pageId="4" pageNumber="5" start="Figure 48" startId="F3">
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" pageId="4" pageNumber="5">Figure 4-8.</emphasis>
<emphasis bold="true" pageId="4" pageNumber="5">4</emphasis>
<caption id="81858F4E6691ADD7FF123308F80994D2" httpUri="https://binary.pensoft.net/fig/17267" pageId="4" pageNumber="5" start="Figure 48" startId="F3">
<paragraph id="B16168BD4E4D9717EE3C41983985F195" pageId="4" pageNumber="5">
<emphasis id="B1C035F8A21D5F402F14129658FEAE16" bold="true" pageId="4" pageNumber="5">Figure 4-8.</emphasis>
<emphasis id="25686A709DE63FEBEA3357A3F7A216F2" bold="true" pageId="4" pageNumber="5">4</emphasis>
Line drawing, showing habitus of
<taxonomicName authorityName="Whitehead" authorityYear="1966" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="acapulcana">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza acapulcana</emphasis>
<taxonomicName id="373337E9C491C2DF1E7AB9EE47572D32" authorityName="Whitehead" authorityYear="1966" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="acapulcana">
<emphasis id="FFF4E55ECE129E1C962118A20F3B8596" italics="true" pageId="4" pageNumber="5">Halocoryza acapulcana</emphasis>
</taxonomicName>
Whitehead, 1966, dorsal aspect, ABL = 2.4mm; Acapulco,
<normalizedToken originalValue="México">Mexico</normalizedToken>
<normalizedToken id="270671477F984EBEB8FCCD6A3DC3FCE6" originalValue="México">Mexico</normalizedToken>
; modified from
<bibRefCitation author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead 1966</bibRefCitation>
<emphasis bold="true" pageId="4" pageNumber="5">5</emphasis>
<bibRefCitation id="F64CAC4D5B8ECF8FCEAA234E4CA28A7A" author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead 1966</bibRefCitation>
<emphasis id="16728D15B69FC36C5989CC05744CC0B9" bold="true" pageId="4" pageNumber="5">5</emphasis>
Line drawing, showing habitus of
<taxonomicName baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza arenaria</emphasis>
<taxonomicName id="18B5B6F8627ABE3D420063A2D62451C7" baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="998FF5889B08B2E7C6604FA291A91E21" italics="true" pageId="4" pageNumber="5">Halocoryza arenaria</emphasis>
</taxonomicName>
(Darlington, 1939), dorsal aspect, ABL = 2.9mm; Rio Piedras, Puerto Rico; modified from Whitehead, 1966
<emphasis bold="true" pageId="4" pageNumber="5">6</emphasis>
<emphasis id="F01C3EF0A22245DBEA264F1C57F5BA05" bold="true" pageId="4" pageNumber="5">6</emphasis>
Line drawing, showing habitus of
<taxonomicName authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="whiteheadiana">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza whiteheadiana</emphasis>
<taxonomicName id="BD89DD9B56B4DA893B419BF4C3DF2ED8" authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="whiteheadiana">
<emphasis id="F5C35DEB50CF56A628BC5E7889FDDD8D" italics="true" pageId="4" pageNumber="5">Halocoryza whiteheadiana</emphasis>
</taxonomicName>
sp. n., dorsal aspect, ABL = 2.9mm; ABL = 2.8mm,ADP127139; Isla Del Carmen,
<normalizedToken originalValue="México">Mexico</normalizedToken>
<emphasis bold="true" pageId="4" pageNumber="5">7</emphasis>
<normalizedToken id="67F2A75C51AB39B36F3D98F9A7762592" originalValue="México">Mexico</normalizedToken>
<emphasis id="167C6376F5CE9CBE9293E0F1B610F4FC" bold="true" pageId="4" pageNumber="5">7</emphasis>
Line drawing, showing male genitalia, median lobe, of
<taxonomicName authorityName="Whitehead" authorityYear="1966" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="acapulcana">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza acapulcana</emphasis>
<taxonomicName id="4250A3A22B2085CAFD76496E23690888" authorityName="Whitehead" authorityYear="1966" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="acapulcana">
<emphasis id="AE8FD217C2C41935B86CAEEEC9397711" italics="true" pageId="4" pageNumber="5">Halocoryza acapulcana</emphasis>
</taxonomicName>
Whitehead, 1966, left lateral aspect, 0.41mm (dorsal margin of basal lobe to apex); Acapulco,
<normalizedToken originalValue="México">Mexico</normalizedToken>
<normalizedToken id="4CE01248592AD705371A12084D8F5FDB" originalValue="México">Mexico</normalizedToken>
; modified from
<bibRefCitation author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead 1966</bibRefCitation>
<emphasis bold="true" pageId="4" pageNumber="5">8</emphasis>
<bibRefCitation id="CC45B66C5AB5628D1708F7FE87F66F55" author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead 1966</bibRefCitation>
<emphasis id="528B9849A12FC9F2986F987836C2E6DC" bold="true" pageId="4" pageNumber="5">8</emphasis>
Line drawing, showing male genitalia, median lobe,
<taxonomicName baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza arenaria</emphasis>
<taxonomicName id="F695C98CFB3D9D667C60137EB512FFC0" baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="D4E41D2461E7EBA7BA524DD4CB69B7FC" italics="true" pageId="4" pageNumber="5">Halocoryza arenaria</emphasis>
</taxonomicName>
(Darlington, 1939), left lateral aspect with internal sac everted, 0.41mm (dorsal margin of basal lobe to apex); Rio Piedras, Puerto Rico; modified from Whitehead, 1966.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="references">
<paragraph pageId="4" pageNumber="5">References.</paragraph>
<paragraph pageId="4" pageNumber="5">
<bibRefCitation author="Bruneau de Mire, P" editor="Erwin, TL" journalOrPublisher="Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b. v. Publishers, The Hague" pageId="8" pageNumber="9" pagination="327 - 330" refId="B3" refString="Bruneau de Mire, P, 1979. Trans-Atlantic dispersal: Several examples of colonization of the Gulf of Biafra by Middle American stocks of Carabidae. In: Erwin, TL, Ball, GE, Whitehead, DR, Halpern, AL, Eds., Carabid Beetles: Their evolution, natural history, and classification. Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b.v. Publishers, The Hague: 327 - 330" title="Trans-Atlantic dispersal: Several examples of colonization of the Gulf of Biafra by Middle American stocks of Carabidae." volumeTitle="Carabid Beetles: Their evolution, natural history, and classification." year="1979">
<subSubSection id="CA90502345505019B6EAA480B114557A" pageId="4" pageNumber="5" type="references">
<paragraph id="66CD805146793C132D9B059E94406C6B" pageId="4" pageNumber="5">References.</paragraph>
<paragraph id="FA454DB4B2985938CBDFDE6CA9A28C8A" pageId="4" pageNumber="5">
<bibRefCitation id="F86048000972F60B8C05F0F2B6981D8C" author="Bruneau de Mire, P" editor="Erwin, TL" journalOrPublisher="Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b. v. Publishers, The Hague" pageId="8" pageNumber="9" pagination="327 - 330" refId="B3" refString="Bruneau de Mire, P, 1979. Trans-Atlantic dispersal: Several examples of colonization of the Gulf of Biafra by Middle American stocks of Carabidae. In: Erwin, TL, Ball, GE, Whitehead, DR, Halpern, AL, Eds., Carabid Beetles: Their evolution, natural history, and classification. Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b.v. Publishers, The Hague: 327 - 330" title="Trans-Atlantic dispersal: Several examples of colonization of the Gulf of Biafra by Middle American stocks of Carabidae." volumeTitle="Carabid Beetles: Their evolution, natural history, and classification." year="1979">
Bruneau de
<normalizedToken originalValue="Miré">Mire</normalizedToken>
<normalizedToken id="235427D80E80B964DE515AF22B30A824" originalValue="Miré">Mire</normalizedToken>
(1979)
</bibRefCitation>
,
<bibRefCitation author="Nichols, SW" journalOrPublisher="Cornell University" pageId="8" pageNumber="9" refId="B14" refString="Nichols, SW, 1988. Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Cornell University" title="Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae)" year="1988">Nichols (1988</bibRefCitation>
<bibRefCitation id="AAF68AA7857A661EAA1886C77E12FF61" author="Nichols, SW" journalOrPublisher="Cornell University" pageId="8" pageNumber="9" refId="B14" refString="Nichols, SW, 1988. Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Cornell University" title="Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae)" year="1988">Nichols (1988</bibRefCitation>
, Ph.D. dissertation),
<bibRefCitation author="Peck, SB" journalOrPublisher="Arthropods of Florida and Neighboring Land Areas" pageId="8" pageNumber="9" refId="B16" refString="Peck, SB, Thomas, MC, 1998. A distributional checklist of the beetles (Coleoptera) of Florida. Arthropods of Florida and Neighboring Land Areas 16, 180 pp." title="A distributional checklist of the beetles (Coleoptera) of Florida" volume="16" year="1998">Peck and Thomas (1998)</bibRefCitation>
<bibRefCitation id="87432A9588CA7BE0842D8AB0D48B448D" author="Peck, SB" journalOrPublisher="Arthropods of Florida and Neighboring Land Areas" pageId="8" pageNumber="9" refId="B16" refString="Peck, SB, Thomas, MC, 1998. A distributional checklist of the beetles (Coleoptera) of Florida. Arthropods of Florida and Neighboring Land Areas 16, 180 pp." title="A distributional checklist of the beetles (Coleoptera) of Florida" volume="16" year="1998">Peck and Thomas (1998)</bibRefCitation>
, (
<bibRefCitation author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead (1966</bibRefCitation>
<bibRefCitation id="EAD3CF077D4E4ABF6D299BE6D9A96B4C" author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead (1966</bibRefCitation>
, 1969).
</paragraph>
</subSubSection>

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@ -1,103 +1,103 @@
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Wooldridge, 1993: 359-360 (orig. descr.).
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<subSubSection pageId="0" pageNumber="51" type="materials_examined">
<paragraph pageId="0" pageNumber="51">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="51">
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<collectingCountry name="Philippines">Philippines</collectingCountry>
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<paragraph id="C73AEE9AE7F0DE825B63BBB346A93C39" pageId="0" pageNumber="51">Additional material examined.</paragraph>
<paragraph id="DBA3190BBFE7699F520479D8C500077A" pageId="0" pageNumber="51">
<materialsCitation id="28A06F644CF9BB9ED9D64DFBC0739BC9" accessionNumber="OK316810" collectingDate="1995-01-01" collectorName="Mendoza, GenBank" country="Philippines" elevation="100" latitude="12.796945" location="Sablayan" longLatPrecision="21" longitude="120.909164" specimenCount="♂">
<collectingCountry id="CD0DEC80B0435D1201115459A4693241" name="Philippines">Philippines</collectingCountry>
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(AdMU), Occ. Mindoro,
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, small limestone river; rootpacks, dist. primary forest; ca.
<geoCoordinate degrees="12" direction="north" minutes="47" orientation="latitude" precision="15" seconds="49" value="12.796945">12°47'49&quot;N</geoCoordinate>
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,
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<geoCoordinate id="0C1BA68A4DBB833ED6165D0A087869D2" degrees="120" direction="east" minutes="54" orientation="longitude" precision="15" seconds="33" value="120.909164">120°54'33&quot;E</geoCoordinate>
; ca.
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
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<quantity id="3C074973848656B9357516A68BCB68FE" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
a.s.l.
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;
<collectingDate value="1995-01-01">01 Jan. 1995</collectingDate>
<collectingDate id="551B2571F2F88F47728361D63FBAE44F" value="1995-01-01">01 Jan. 1995</collectingDate>
, leg.
<collectorName>Mendoza</collectorName>
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<collectorName id="C2C0988FC4A4968E738D5C0F6C763110">Mendoza</collectorName>
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;
<collectorName>GenBank</collectorName>
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:
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<accessionNumber id="04AB5CFA7CA05468B4324A968C4D4C69" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OK316810">OK316810</accessionNumber>
; BOLD: COLPH064-21
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.
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<paragraph pageId="0" pageNumber="51">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="51">
<subSubSection id="7451448A5438F879BCEB6F99CAF52266" pageId="0" pageNumber="51" type="remarks">
<paragraph id="4820D8AA93BA55E31A0750016E113C00" pageId="0" pageNumber="51">Remarks.</paragraph>
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Assignment of the specimen to this taxon was based primarily on genital characters. No remarkable difference was noted compared to the original description. The material used for the molecular analysis in this study was collected 90 km south of one of the localities of the paratypes, but on the same island. Known distribution of
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<emphasis italics="true" pageId="0" pageNumber="51">B. ferax</emphasis>
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<emphasis id="0BC46036A3C17A0D72DF1BB1ED58263D" italics="true" pageId="0" pageNumber="51">B. ferax</emphasis>
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includes the Philippine Islands of Mindoro and Mindanao.
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<taxonomicName LSID="http://species-id.net/wiki/Chinavia_brasicola" authority="Rolston, 1983" authorityName="Rolston" authorityYear="1983" class="Insecta" family="Pentatomidae" genus="Chinavia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chinavia brasicola" order="Hemiptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="brasicola">Chinavia brasicola (Rolston, 1983)</taxonomicName>
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Figs 13-18
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<paragraph pageId="6" pageNumber="65">Remarks.</paragraph>
<paragraph pageId="6" pageNumber="65">
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Adults have general body color green, connexivum red-orange with black maculae on posterolateral angles (
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). The distribution includes the Brazilian states of
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Paulo, Santa Catarina and Rio Grande do Sul (
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). The single sitting record is a fern
<taxonomicName class="Polypodiopsida" family="Dennstaedtiaceae" genus="Dennstaedtia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dennstaedtia globulifera" order="Polypodiales" pageId="6" pageNumber="65" phylum="Tracheophyta" rank="species" species="globulifera">Dennstaedtia globulifera</taxonomicName>
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(Poir.) Hieron (
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) (
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). The morphometric parameters of nymphal instars are shown in Table 4.
</paragraph>
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<paragraph pageId="6" pageNumber="65">
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<paragraph id="4848488CA98EC13F2DF460866160BD1B" pageId="6" pageNumber="65">
Table 4. Morphometric traits of nymphs of
<taxonomicName class="Insecta" family="Pentatomidae" genus="Chinavia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chinavia brasicola" order="Hemiptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="brasicola">Chinavia brasicola</taxonomicName>
<taxonomicName id="6E3DC46489C33FB28E49F177B26D99C9" genus="Chinavia" lsidName="Chinavia brasicola" pageId="6" pageNumber="65" rank="species" species="brasicola">Chinavia brasicola</taxonomicName>
(Rolston, 1983) (n = 15) (mean
<normalizedToken originalValue="±">+/-</normalizedToken>
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standard deviation, mm).
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="65">
<table pageId="6" pageNumber="65">
<tr pageId="6" pageNumber="65">
<th colspan="1" pageId="6" pageNumber="65" rowspan="1">Measures</th>
<th colspan="1" pageId="6" pageNumber="65" rowspan="1">1st instar</th>
<th colspan="1" pageId="6" pageNumber="65" rowspan="1">2nd instar</th>
<th colspan="1" pageId="6" pageNumber="65" rowspan="1">3rd instar</th>
<th colspan="1" pageId="6" pageNumber="65" rowspan="1">4th instar</th>
<th colspan="1" pageId="6" pageNumber="65" rowspan="1">5th instar</th>
<paragraph id="AAE8EDCB01A268B235E543D5C4762417" pageId="6" pageNumber="65">
<table id="F042B2E7728906D0341E7C8722A1CA6E" pageId="6" pageNumber="65">
<tr id="DCA6269E0D127E553DA55D64A27D0619" pageId="6" pageNumber="65">
<th id="6E5F56B0063D01B5D1B44E11A3A29384" colspan="1" pageId="6" pageNumber="65" rowspan="1">Measures</th>
<th id="1C17461033C0F86635C7D6D9811E739D" colspan="1" pageId="6" pageNumber="65" rowspan="1">1st instar</th>
<th id="708DC5A437D9F43AB6CAF4097F19CB21" colspan="1" pageId="6" pageNumber="65" rowspan="1">2nd instar</th>
<th id="95A24329647D7A50907AB9652B5DBECD" colspan="1" pageId="6" pageNumber="65" rowspan="1">3rd instar</th>
<th id="D37126CFD33C6472DA834FA2AA2B7BEE" colspan="1" pageId="6" pageNumber="65" rowspan="1">4th instar</th>
<th id="3C102D17C3A5BE8C2388EF83B7EC8B1B" colspan="1" pageId="6" pageNumber="65" rowspan="1">5th instar</th>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="65" type="first instar">
<paragraph pageId="6" pageNumber="65">First instar</paragraph>
<paragraph pageId="6" pageNumber="65">(Fig. 14). Body round and convex surface without punctuation. General coloration dark brown to black. Head conical and strongly declivent; black, with median orange macula, which extends from the posterior portion of the head to the anterior margin of mesonotum; clypeus with apex obtuse, slightly surpassing the mandibular plates, these subtriangular shaped. Ocelli absent. Antennae black; intersegmental areas with orange color; antennal segments with short and uniformly distributed hairs. Antennal segment I shortest and antennal segment IV longest. Antennal segments III and IV subequal in size. Rostrum black, reaching anterior margin of urosternite II. Thorax mostly dark, except for the orange median macula. Legs black, with hairs uniformly distributed on all segments, tibiae dorsally flattened, tarsi with two segments, a pair of tarsal claws and pulvili. Abdomen predominantly black, with three pairs (3+3) of white maculae, located between lateral and median plates. Median dorsal plates black; ostioles on median dorsal plates I-III. Lateral plates semicircular, black, without punctuation, adjacent to lateral margin of each segment. Spiracles near anterior margin of lateral plates, on urosternites II to VIII. From urosternites III to VII, 1+1 trichobothria placed medially of an imaginary line across spiracles and near posterior margin of each segment.</paragraph>
<subSubSection id="0463B5A776B8E92A593571F3C7D707EE" pageId="6" pageNumber="65" type="first instar">
<paragraph id="1429454804755B96921420985741627A" pageId="6" pageNumber="65">First instar</paragraph>
<paragraph id="09DD872A074780EF7D41F51DBF9E3DC3" pageId="6" pageNumber="65">(Fig. 14). Body round and convex surface without punctuation. General coloration dark brown to black. Head conical and strongly declivent; black, with median orange macula, which extends from the posterior portion of the head to the anterior margin of mesonotum; clypeus with apex obtuse, slightly surpassing the mandibular plates, these subtriangular shaped. Ocelli absent. Antennae black; intersegmental areas with orange color; antennal segments with short and uniformly distributed hairs. Antennal segment I shortest and antennal segment IV longest. Antennal segments III and IV subequal in size. Rostrum black, reaching anterior margin of urosternite II. Thorax mostly dark, except for the orange median macula. Legs black, with hairs uniformly distributed on all segments, tibiae dorsally flattened, tarsi with two segments, a pair of tarsal claws and pulvili. Abdomen predominantly black, with three pairs (3+3) of white maculae, located between lateral and median plates. Median dorsal plates black; ostioles on median dorsal plates I-III. Lateral plates semicircular, black, without punctuation, adjacent to lateral margin of each segment. Spiracles near anterior margin of lateral plates, on urosternites II to VIII. From urosternites III to VII, 1+1 trichobothria placed medially of an imaginary line across spiracles and near posterior margin of each segment.</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65" type="second instar">
<paragraph pageId="6" pageNumber="65">Second instar</paragraph>
<paragraph lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">
<subSubSection id="E9F975AFCAB69499F88C70C2C9314112" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65" type="second instar">
<paragraph id="05518828BF698B0EA77B1686D014296B" pageId="6" pageNumber="65">Second instar</paragraph>
<paragraph id="C91D83079DAB460BFA942618178B8B1C" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">
(Fig. 15). Body oval and less convex than first instar. General color black, with punctuation on the dorsum of the head, thorax, median and lateral plates. Head less declivent than in previous instar. Clypeus obtuse at apex, subequal in size to the mandibular plates, which are broader than that observed in previous instar. Eyes almost as wide as base of clypeus. Rostrum black, surpassing the anterior margin
<pageBreakToken pageId="7" pageNumber="66" start="start">of</pageBreakToken>
<pageBreakToken id="EABAECDD6ADD66CAD4D4422AEAD95FEA" pageId="7" pageNumber="66" start="start">of</pageBreakToken>
urosternite III. Thorax with 1+1 orange maculae along the margins of pronotum; lateral margins of pro- and mesonotum slightly deflected. Legs black, hairs more dense on tibiae. Abdomen predominantly black, maculae distributed as follows: one, white, round maculae, located anteriad of first median plate; a pair (1+1) of white maculae, located on each side of the first lateral plates; a pair (1+1) of large, rounded, orange maculae, located between lateral plates and first median plate and four pairs (4+4) white, located between lateral and median plates, the third pair is the largest. Median and lateral dorsal plates black punctured. Urosternites III to VII with 2+2 trichobothria, one trichobothrium medially of the spiracular line and the other along that line. Other characteristics as described for the first instar.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="66" type="third instar">
<paragraph pageId="7" pageNumber="66">Third instar</paragraph>
<paragraph pageId="7" pageNumber="66">(Fig. 16). Antennal segment I shorter, segments II and IV subequal in length, larger than segment III. Thorax punctured, predominantly black, except an orange maculae along each margin. Pronotum with margins slightly deflected and serrate. Legs black, except for the area between the femur and tibia, which has light brown color; hairs denser on the ventral surface of tibia and tarsus. Abdomen with a round white macula anteriad of first median plate; one pair (1+1) of orange maculae between the first median plate and lateral plates, and four pairs (4+4) of white maculae located between median and lateral plates of tergites IV-VII. Other characters as described for the previous instar.</paragraph>
<subSubSection id="441F052C06E26B53C4617F09D085AECB" pageId="7" pageNumber="66" type="third instar">
<paragraph id="F6035A17267803E6036A96935508DA32" pageId="7" pageNumber="66">Third instar</paragraph>
<paragraph id="A4BD59D7DF90CD75B63D3E7F320361FB" pageId="7" pageNumber="66">(Fig. 16). Antennal segment I shorter, segments II and IV subequal in length, larger than segment III. Thorax punctured, predominantly black, except an orange maculae along each margin. Pronotum with margins slightly deflected and serrate. Legs black, except for the area between the femur and tibia, which has light brown color; hairs denser on the ventral surface of tibia and tarsus. Abdomen with a round white macula anteriad of first median plate; one pair (1+1) of orange maculae between the first median plate and lateral plates, and four pairs (4+4) of white maculae located between median and lateral plates of tergites IV-VII. Other characters as described for the previous instar.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="66" type="fourth instar">
<paragraph pageId="7" pageNumber="66">Fourth instar</paragraph>
<paragraph pageId="7" pageNumber="66">(Fig. 17). Body oval, less convex than in earlier instars, predominantly black. Head less declivent than in previous instar, black, with punctures. Some specimens with an orange band in the middle of each mandibular plates. Antennae with abundant hairs on segments III and IV. Thorax predominantly black, with orange maculae at margins of pro- and mesonotum. Pronotum trapezoidal; mesonotum rectangular, posterior margin wide, V-shaped, denoting the formation of scutellum. Wing pads slightly developed, reaching posterior margin of metanotum. Dorsal abdominal maculae with the same color and distribution as observed in third instar, but wider. Lateral plates semicircular, predominantly orange, slightly punctured. Other characteristics as described in the previous instars.</paragraph>
<subSubSection id="E4D27D102C8EDE2B1B90DDF5D6D825DA" pageId="7" pageNumber="66" type="fourth instar">
<paragraph id="2C39E6F60B81D116A4071F7E97C0756D" pageId="7" pageNumber="66">Fourth instar</paragraph>
<paragraph id="DD5498451E593DD825DA1C46084F7AD9" pageId="7" pageNumber="66">(Fig. 17). Body oval, less convex than in earlier instars, predominantly black. Head less declivent than in previous instar, black, with punctures. Some specimens with an orange band in the middle of each mandibular plates. Antennae with abundant hairs on segments III and IV. Thorax predominantly black, with orange maculae at margins of pro- and mesonotum. Pronotum trapezoidal; mesonotum rectangular, posterior margin wide, V-shaped, denoting the formation of scutellum. Wing pads slightly developed, reaching posterior margin of metanotum. Dorsal abdominal maculae with the same color and distribution as observed in third instar, but wider. Lateral plates semicircular, predominantly orange, slightly punctured. Other characteristics as described in the previous instars.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="67" type="fifth instar">
<paragraph pageId="8" pageNumber="67">
<pageBreakToken pageId="8" pageNumber="67" start="start">Fifth</pageBreakToken>
<subSubSection id="409D920CF0FDC971A1C71DB64675CCFE" pageId="8" pageNumber="67" type="fifth instar">
<paragraph id="820585514A683D204E24325A3AE73640" pageId="8" pageNumber="67">
<pageBreakToken id="0708EC842DB25051DD3313231407FCE2" pageId="8" pageNumber="67" start="start">Fifth</pageBreakToken>
instar
</paragraph>
<paragraph pageId="8" pageNumber="67">(Fig. 18). Body oval to pyriform, predominantly black. Head flat; mandibular plates wide, each with orange band present in some individuals. Thorax predominantly black, 1+1 orange maculae on pronotum and mesonotum along the anterolateral margins, median macula orange. Pronotum wider; mesonotum more developed; scutellum well delimited. Wing pads well developed, surpassing the middle of abdominal segment III. Legs black, hairs denser ventrally. Abdomen black, coarsely punctured with maculae dorsally distributed as follows: one pair (1+1) of orange maculae near first median plate, with another white, semicircular macula between the preceding two maculae; and four pairs (4+4) of white maculae, located near lateral plates, on segments IV to VII; lateral plates with semicircular orange maculae, outlined in black. Urosternite VIII split longitudinally in females and entire in males. Other characteristics as described in the previous instars.</paragraph>
<caption pageId="8" pageNumber="67">
<paragraph pageId="8" pageNumber="67">
<paragraph id="B279ABC9700812A633A115FB803839F6" pageId="8" pageNumber="67">(Fig. 18). Body oval to pyriform, predominantly black. Head flat; mandibular plates wide, each with orange band present in some individuals. Thorax predominantly black, 1+1 orange maculae on pronotum and mesonotum along the anterolateral margins, median macula orange. Pronotum wider; mesonotum more developed; scutellum well delimited. Wing pads well developed, surpassing the middle of abdominal segment III. Legs black, hairs denser ventrally. Abdomen black, coarsely punctured with maculae dorsally distributed as follows: one pair (1+1) of orange maculae near first median plate, with another white, semicircular macula between the preceding two maculae; and four pairs (4+4) of white maculae, located near lateral plates, on segments IV to VII; lateral plates with semicircular orange maculae, outlined in black. Urosternite VIII split longitudinally in females and entire in males. Other characteristics as described in the previous instars.</paragraph>
<caption id="EAF9951308A3230D043B2AA76CAE0BE1" pageId="8" pageNumber="67">
<paragraph id="792B6D147FAA20F205DE1F054DA6B170" pageId="8" pageNumber="67">
Figures 13-18.
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<taxonomicName id="92964E08210D047EB88FA79D3520DA8A" genus="Chinavia" lsidName="Chinavia brasicola" pageId="8" pageNumber="67" rank="species" species="brasicola">Chinavia brasicola</taxonomicName>
(Rolston, 1983). 13 Adult 14 First instar 15 Second instar 16 Third instar 17 Fourth instar 18 Fifth instar.
</paragraph>
</caption>

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<taxonomicName LSID="http://species-id.net/wiki/Hemilamprops_normani" authority="Bonnier, 1896" authorityName="Bonnier" authorityYear="1896" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani Bonnier, 1896</taxonomicName>
<treatment id="C4C82B0C2CCF0C5DBD3EBD5D70A7E168" ID-GBIF-Taxon="152037606" LSID="urn:lsid:plazi:treatment:C4C82B0C2CCF0C5DBD3EBD5D70A7E168" httpUri="http://treatment.plazi.org/id/C4C82B0C2CCF0C5DBD3EBD5D70A7E168" lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76" scope_family="Lampropidae">
<subSubSection id="2D356613CB4B58AC3FA7BD9825042229" pageId="3" pageNumber="76" type="nomenclature">
<paragraph id="059C1556F56C9689958E38F137372681" pageId="3" pageNumber="76">
<taxonomicName id="54759086976C66522C5E3E88AC1D3862" ID-CoL="3KPDX" LSID="http://species-id.net/wiki/Hemilamprops_normani" authority="Bonnier, 1896" authorityName="Bonnier" authorityYear="1896" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani Bonnier, 1896</taxonomicName>
Fig. 1A, B
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="76" type="reference_group">
<paragraph pageId="3" pageNumber="76">
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
<subSubSection id="CF4FF149DF2C1AA3515CAC96552545BD" pageId="3" pageNumber="76" type="reference_group">
<paragraph id="0BBD3E48B47EF12CFBF7EB4EFFA3D17C" pageId="3" pageNumber="76">
<taxonomicName id="57D7C853C7ED4DC88D8CCFF49A35811A" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
-
<bibRefCitation author="Bonnier, JJ" journalOrPublisher="Annales de l'Universitee de Lyon" pageId="9" pageNumber="82" pagination="528 - 562" title="Edriophthalmes. Resultats Scientifiques de la Campagne du &quot; Caudan &quot; dans le Golfe de Goscogne, Aout-Septembre 1895." volume="26" year="1896">Bonnier 1896</bibRefCitation>
<bibRefCitation id="F11B6B3F86451EEF1E1066840CB19A81" author="Bonnier, JJ" journalOrPublisher="Annales de l'Universitee de Lyon" pageId="9" pageNumber="82" pagination="528 - 562" title="Edriophthalmes. Resultats Scientifiques de la Campagne du &quot; Caudan &quot; dans le Golfe de Goscogne, Aout-Septembre 1895." volume="26" year="1896">Bonnier 1896</bibRefCitation>
, pp. 546-549, pl. 29 fig. 3.
</paragraph>
<paragraph pageId="3" pageNumber="76">
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops cristata" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="cristata">Hemilamprops cristata</taxonomicName>
<paragraph id="479BD1A9E5F089A40C7D387FE4AEBD64" pageId="3" pageNumber="76">
<taxonomicName id="E778F4D5ACAFF3A50022193716C15677" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops cristata" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="cristata">Hemilamprops cristata</taxonomicName>
-
<bibRefCitation author="Calman, WT" journalOrPublisher="Fishery Ireland Scientific Investigations" pageId="9" pageNumber="82" pagination="1 - 52" title="The marine fauna of the west coast of Ireland. Part IV. Cumacea." volume="1" year="1905">Calman 1905</bibRefCitation>
<bibRefCitation id="F34E98A6E19136EB256DDCAFBE67927D" author="Calman, WT" journalOrPublisher="Fishery Ireland Scientific Investigations" pageId="9" pageNumber="82" pagination="1 - 52" title="The marine fauna of the west coast of Ireland. Part IV. Cumacea." volume="1" year="1905">Calman 1905</bibRefCitation>
, p. 41, 49 [nec
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops cristata" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="cristata">Hemilamprops cristata</taxonomicName>
<taxonomicName id="5499F0A8E015FB846B28C60DF1A56E26" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops cristata" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="cristata">Hemilamprops cristata</taxonomicName>
(Sars, 1870)].
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="76" type="material examined">
<paragraph pageId="3" pageNumber="76">Material examined.</paragraph>
<paragraph pageId="3" pageNumber="76">
<subSubSection id="0706DB2BC18774BEC523BAA5FD5CBFC6" pageId="3" pageNumber="76" type="material examined">
<paragraph id="23BE5F79118EFEA1B95B86929F5557DC" pageId="3" pageNumber="76">Material examined.</paragraph>
<paragraph id="91A45C1602EC58C7BC91BFE812F99572" pageId="3" pageNumber="76">
Cap Ferret Canyon, Bay of Biscay, ESSAIS II: stn TS10,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="44.551666">44°33.10'N</geoCoordinate>
<geoCoordinate id="960AF434407FC56AE6F3DE10D1FA0BA4" direction="north" orientation="latitude" precision="9" value="44.551666">44°33.10'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-2.2188334">2°13.13'W</geoCoordinate>
<geoCoordinate id="F888A03F74FFDC97BEDD88595AC13D43" direction="west" orientation="longitude" precision="9" value="-2.2188334">2°13.13'W</geoCoordinate>
, 791-790 m, 18/05/89, 3 mancas, 2 pread. female, 1 ad. male; stn TS11,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="44.54817">44°32.89'N</geoCoordinate>
<geoCoordinate id="72F4590B57042928849C0E8C55D9005A" direction="north" orientation="latitude" precision="9" value="44.54817">44°32.89'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-2.2373333">2°14.24'W</geoCoordinate>
<geoCoordinate id="3517BFF1AAEB189B5BF3D5E9286F7785" direction="west" orientation="longitude" precision="9" value="-2.2373333">2°14.24'W</geoCoordinate>
, 923-924 m, 18/05/89, 6 mancas, 2 pread. males; stn TS13,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="44.56983">44°34.19'N</geoCoordinate>
<geoCoordinate id="DAC6A48FE303A5206FADB34F26F75788" direction="north" orientation="latitude" precision="9" value="44.56983">44°34.19'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-2.2696667">2°16.18'W</geoCoordinate>
<geoCoordinate id="E9B9A31F6AE371743DC5B07AFA766EB8" direction="west" orientation="longitude" precision="9" value="-2.2696667">2°16.18'W</geoCoordinate>
, 1097-1099 m, 17/05/89, 4 mancas, 2 imm. males. J.-C. Sorbe leg.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="76" type="remarks">
<paragraph pageId="3" pageNumber="76">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="76">
<subSubSection id="E49D0F65999CCE5B0E5354BFE7B73A10" pageId="3" pageNumber="76" type="remarks">
<paragraph id="4AEE17C504D11F7EDF036F1085FF1F1D" pageId="3" pageNumber="76">Remarks.</paragraph>
<paragraph id="F5760CC34769C2C5D97417E0527017C1" pageId="3" pageNumber="76">
Although
<bibRefCitation author="Calman, WT" journalOrPublisher="Fishery Ireland Scientific Investigations" pageId="9" pageNumber="82" pagination="1 - 52" title="The marine fauna of the west coast of Ireland. Part IV. Cumacea." volume="1" year="1905">Calman (1905)</bibRefCitation>
<bibRefCitation id="2BEF6FC5F644C958C65C76D4B9876E7A" author="Calman, WT" journalOrPublisher="Fishery Ireland Scientific Investigations" pageId="9" pageNumber="82" pagination="1 - 52" title="The marine fauna of the west coast of Ireland. Part IV. Cumacea." volume="1" year="1905">Calman (1905)</bibRefCitation>
suggested the synonymy between
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops cristatus" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="cristatus">Hemilamprops cristatus</taxonomicName>
<taxonomicName id="9006667052BEB65746407BD904950107" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops cristatus" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="cristatus">Hemilamprops cristatus</taxonomicName>
and
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
<taxonomicName id="1B2F029B924BBA9D1A1D5E847FCD8036" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
, other authors (
<bibRefCitation author="Sars, GO" journalOrPublisher="Cumacea. Bergen Museum, Christiania, Oslo" pageId="9" pageNumber="82" title="An acount of the Crustacea of Norway. Vol. 3." year="1900">Sars 1900</bibRefCitation>
<bibRefCitation id="7367BC64783EAFE22D8BFE0D850D858A" author="Sars, GO" journalOrPublisher="Cumacea. Bergen Museum, Christiania, Oslo" pageId="9" pageNumber="82" title="An acount of the Crustacea of Norway. Vol. 3." year="1900">Sars 1900</bibRefCitation>
;
<bibRefCitation author="Hansen, H" journalOrPublisher="The Danish Ingolf-Expedition" pageId="9" pageNumber="82" pagination="1 - 86" title="Cumacea. Crustacea Malacostraca 4." volume="3" year="1920">Hansen 1920</bibRefCitation>
<bibRefCitation id="3BEFCB4D36C9501FA74D6EDA00549BB4" author="Hansen, H" journalOrPublisher="The Danish Ingolf-Expedition" pageId="9" pageNumber="82" pagination="1 - 86" title="Cumacea. Crustacea Malacostraca 4." volume="3" year="1920">Hansen 1920</bibRefCitation>
;
<bibRefCitation author="Fage, L" journalOrPublisher="Resultats des Campagnes Scientifiques" pageId="9" pageNumber="82" pagination="1 - 50" title="Cumaces et Leptostraces provenants des campagnes scientifiques du Prince Albert 1 er de Monaco." volume="77" year="1929">Fage 1929</bibRefCitation>
<bibRefCitation id="46F94B73CC2E665B8559429F42EE96E1" author="Fage, L" journalOrPublisher="Resultats des Campagnes Scientifiques" pageId="9" pageNumber="82" pagination="1 - 50" title="Cumaces et Leptostraces provenants des campagnes scientifiques du Prince Albert 1 er de Monaco." volume="77" year="1929">Fage 1929</bibRefCitation>
,
<bibRefCitation author="Fage, L" journalOrPublisher="Bulletin de l'Institute oceanographique, Monaco" pageId="9" pageNumber="82" pagination="1 - 14" title="Les Cumaces de la Mediterranee. Remarques systematiques et biologiques." volume="783" year="1940">1940</bibRefCitation>
<bibRefCitation id="CD4A5BD85126F5BD8953580F3A52D9D7" author="Fage, L" journalOrPublisher="Bulletin de l'Institute oceanographique, Monaco" pageId="9" pageNumber="82" pagination="1 - 14" title="Les Cumaces de la Mediterranee. Remarques systematiques et biologiques." volume="783" year="1940">1940</bibRefCitation>
) consider them as valid species, which is the criterion followed here. It is possible that the presence of
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Mesolamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesolamprops denticulatus" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="denticulatus">Mesolamprops denticulatus</taxonomicName>
<taxonomicName id="10CFA2C2ED7C93F67A7F3F2A5FBE64F0" class="Malacostraca" family="Lampropidae" genus="Mesolamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesolamprops denticulatus" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="denticulatus">Mesolamprops denticulatus</taxonomicName>
in the Bay of Biscay, together with the two species of
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="genus">Hemilamprops</taxonomicName>
<taxonomicName id="0F26E52B67D9740202207F12F575CBDB" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="genus">Hemilamprops</taxonomicName>
, led to the confusion between these three species.
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
<taxonomicName id="291A9E4785C9DE3BA7534A8066C46F40" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
can be distinguished from the other two species by its higher number of lateral setae on the telson (6-8). Moreover, the three terminal setae of the telsonof
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops cristatus" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="cristatus">Hemilamprops cristatus</taxonomicName>
<taxonomicName id="E418C41FB0F112BDAF596FC3DFAE972D" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops cristatus" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="cristatus">Hemilamprops cristatus</taxonomicName>
are of the same length, while in
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
<taxonomicName id="2A90263D9676D325E755915F714F4B5B" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
the central one is the longest.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76" type="distribution">
<paragraph pageId="3" pageNumber="76">Distribution.</paragraph>
<paragraph lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76">
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
<subSubSection id="B0521371B5A0109B9C81F43057FB0647" lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76" type="distribution">
<paragraph id="5FD933A212AA7B3947D6702F3801A021" pageId="3" pageNumber="76">Distribution.</paragraph>
<paragraph id="F58DC2BD82C7EFC0B22550C866340E8B" lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76">
<taxonomicName id="2DF799B0465F00A295A731BB9118D62B" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
is known to inhabit the waters of the Bay of Biscay (
<bibRefCitation author="Bonnier, JJ" journalOrPublisher="Annales de l'Universitee de Lyon" pageId="9" pageNumber="82" pagination="528 - 562" title="Edriophthalmes. Resultats Scientifiques de la Campagne du &quot; Caudan &quot; dans le Golfe de Goscogne, Aout-Septembre 1895." volume="26" year="1896">Bonnier 1896</bibRefCitation>
<bibRefCitation id="2BEEB27BE67B17630201259E5FCF4B19" author="Bonnier, JJ" journalOrPublisher="Annales de l'Universitee de Lyon" pageId="9" pageNumber="82" pagination="528 - 562" title="Edriophthalmes. Resultats Scientifiques de la Campagne du &quot; Caudan &quot; dans le Golfe de Goscogne, Aout-Septembre 1895." volume="26" year="1896">Bonnier 1896</bibRefCitation>
;
<bibRefCitation pageId="3" pageNumber="76">Jones 1985</bibRefCitation>
<bibRefCitation id="FBB114578A6AEB4A07BDB2E8DBB6FD1A" pageId="3" pageNumber="76">Jones 1985</bibRefCitation>
), the west of Ireland (
<bibRefCitation author="Calman, WT" journalOrPublisher="Fishery Ireland Scientific Investigations" pageId="9" pageNumber="82" pagination="1 - 52" title="The marine fauna of the west coast of Ireland. Part IV. Cumacea." volume="1" year="1905">Calman 1905</bibRefCitation>
<bibRefCitation id="57F854F13A954A687031749AB9B1ECAE" author="Calman, WT" journalOrPublisher="Fishery Ireland Scientific Investigations" pageId="9" pageNumber="82" pagination="1 - 52" title="The marine fauna of the west coast of Ireland. Part IV. Cumacea." volume="1" year="1905">Calman 1905</bibRefCitation>
), the Azores Islands (
<bibRefCitation author="Fage, L" journalOrPublisher="Resultats des Campagnes Scientifiques" pageId="9" pageNumber="82" pagination="1 - 50" title="Cumaces et Leptostraces provenants des campagnes scientifiques du Prince Albert 1 er de Monaco." volume="77" year="1929">Fage 1929</bibRefCitation>
<bibRefCitation id="15FD1BB7C58971F486E16B73D83365FA" author="Fage, L" journalOrPublisher="Resultats des Campagnes Scientifiques" pageId="9" pageNumber="82" pagination="1 - 50" title="Cumaces et Leptostraces provenants des campagnes scientifiques du Prince Albert 1 er de Monaco." volume="77" year="1929">Fage 1929</bibRefCitation>
) and the Mediterranean Sea (
<bibRefCitation author="Fage, L" journalOrPublisher="Bulletin de l'Institute oceanographique, Monaco" pageId="9" pageNumber="82" pagination="1 - 14" title="Les Cumaces de la Mediterranee. Remarques systematiques et biologiques." volume="783" year="1940">Fage 1940</bibRefCitation>
<bibRefCitation id="DD7054363C3672F839534CBB86F29E7B" author="Fage, L" journalOrPublisher="Bulletin de l'Institute oceanographique, Monaco" pageId="9" pageNumber="82" pagination="1 - 14" title="Les Cumaces de la Mediterranee. Remarques systematiques et biologiques." volume="783" year="1940">Fage 1940</bibRefCitation>
;
<bibRefCitation author="Reyss, D" journalOrPublisher="Crustaceana" pageId="9" pageNumber="82" pagination="216 - 222" title="Cumaces Resultats scientifiques de la campagne &quot; Polymede II &quot; du NO &quot; Jean Charcot &quot; en Mer Ionienne et en Mer Egee (Avril-Mai 1972)." url="10.1163/156854074X00442" volume="27" year="1974 b">Reyss 1974b</bibRefCitation>
<bibRefCitation id="1168FC973A65605512565BA86D5B7298" author="Reyss, D" journalOrPublisher="Crustaceana" pageId="9" pageNumber="82" pagination="216 - 222" title="Cumaces Resultats scientifiques de la campagne &quot; Polymede II &quot; du NO &quot; Jean Charcot &quot; en Mer Ionienne et en Mer Egee (Avril-Mai 1972)." url="10.1163/156854074X00442" volume="27" year="1974 b">Reyss 1974b</bibRefCitation>
). According to
<bibRefCitation pageId="3" pageNumber="76">Jones (1985)</bibRefCitation>
<bibRefCitation id="0F46BD247056431BD5B99A4B24AADFFF" pageId="3" pageNumber="76">Jones (1985)</bibRefCitation>
, in the Bay of Biscay this species inhabits bottoms between 280 and 3000 m. This wide bathymetric range, however, should not be assumed with complete
<pageBreakToken pageId="4" pageNumber="77" start="start">certainty</pageBreakToken>
<pageBreakToken id="2885B5E722A41230C1835F1FDAC419DC" pageId="4" pageNumber="77" start="start">certainty</pageBreakToken>
, since it is possible that
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
<taxonomicName id="FC872F5A954C2A54B98E9C1D92BB53E2" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
has been confused with
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops denticulatus" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="denticulatus">Hemilamprops denticulatus</taxonomicName>
<taxonomicName id="17511B3096F10370E2A16473D80FA124" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops denticulatus" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="denticulatus">Hemilamprops denticulatus</taxonomicName>
, at least in its most shallow distribution. During this study
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
<taxonomicName id="FA3D922DA156203C71FF7DC3F99A83CD" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
was always collected in waters deeper than 700 m.
</paragraph>
<caption pageId="4" pageNumber="77">
<paragraph pageId="4" pageNumber="77">
<caption id="F39FBC154403753FB66109FE21370A9E" pageId="4" pageNumber="77">
<paragraph id="6A60CEACCE27C4189F6B96CDDD199949" pageId="4" pageNumber="77">
Figure 1.
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
<taxonomicName id="828710CC6CA9A82448951F93EFD8B8A0" class="Malacostraca" family="Lampropidae" genus="Hemilamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemilamprops normani" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="normani">Hemilamprops normani</taxonomicName>
Bonnier, 1896: A preadult female, telson and left uropod peduncle B manca, telson and right uropod.
<taxonomicName class="Malacostraca" family="Lampropidae" genus="Mesolamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesolamprops denticulatus" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="denticulatus">Mesolamprops denticulatus</taxonomicName>
<taxonomicName id="3FBD0E91316E51754C86CEFF7C328B92" class="Malacostraca" family="Lampropidae" genus="Mesolamprops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesolamprops denticulatus" order="Cumacea" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="denticulatus">Mesolamprops denticulatus</taxonomicName>
Ledoyer, 1983 C preadult female, telson and right uropod. Scale bar: 0.2 mm.
</paragraph>
</caption>

View file

@ -1,80 +1,79 @@
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<paragraph pageId="12" pageNumber="73">
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<taxonomicNameLabel pageId="12" pageNumber="73">sp. n.</taxonomicNameLabel>
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<paragraph id="CF208349B4F5843F76C6D7314432DBCA" pageId="12" pageNumber="73">
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<taxonomicNameLabel id="1642096CEF9A486E3BE78EE3A2B9A818" pageId="12" pageNumber="73">sp. n.</taxonomicNameLabel>
Figs 33-43
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="type material">
<paragraph pageId="12" pageNumber="73">Type material.</paragraph>
<paragraph pageId="12" pageNumber="73">
<subSubSection id="A950263338D697AF6CF9DF53A3089393" pageId="12" pageNumber="73" type="type material">
<paragraph id="7EA67A48FCABCDC2EE5120F9C18FF00F" pageId="12" pageNumber="73">Type material.</paragraph>
<paragraph id="2EB5413AD22D15C86B36C4D02862FE41" pageId="12" pageNumber="73">
Holotype, ♀ (RMNH), &quot;Indonesia: N. Sumatra, Ketambe, c 400 m, near N. P. Gn. Leuser, Mal. trap, vi.1994, Y. v. Nierop &amp; C. v. Achterberg,
<normalizedToken originalValue="RMNH95”">RMNH'95&quot;</normalizedToken>
<normalizedToken id="C77DAE091911E020267F08AD81857A9A" originalValue="RMNH95”">RMNH'95&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="diagnosis">
<paragraph pageId="12" pageNumber="73">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="73">Clypeus with rather narrow anchor-shaped black patch medially (Fig. 41); (Fig. 36) and comparatively wide ventrally (Fig. 37); first discal cell of fore wing subhyaline (Fig. 35); hind tibia black; metasoma comparatively slender in dorsal view (Fig. 33); ovipositor with comparatively widely separated serrations (Fig. 40). Differs from the other known species by the rather narrow anchor-shaped black patch of the clypeus, the narrowly interrupted ivory transverse stripe on the pronotum and the dark brown hind tarsus.</paragraph>
<subSubSection id="4830069442486F0E83FCED11942519C9" pageId="12" pageNumber="73" type="diagnosis">
<paragraph id="AE2C8434C5921EB791DA48DF194272BD" pageId="12" pageNumber="73">Diagnosis.</paragraph>
<paragraph id="FEF61A8A221C046255FF7041D38878C8" pageId="12" pageNumber="73">Clypeus with rather narrow anchor-shaped black patch medially (Fig. 41); (Fig. 36) and comparatively wide ventrally (Fig. 37); first discal cell of fore wing subhyaline (Fig. 35); hind tibia black; metasoma comparatively slender in dorsal view (Fig. 33); ovipositor with comparatively widely separated serrations (Fig. 40). Differs from the other known species by the rather narrow anchor-shaped black patch of the clypeus, the narrowly interrupted ivory transverse stripe on the pronotum and the dark brown hind tarsus.</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="74" pageId="12" pageNumber="73" type="description">
<paragraph pageId="12" pageNumber="73">Description.</paragraph>
<paragraph pageId="12" pageNumber="73">Holotype, ♀, length of body 13.9 mm (of fore wing 9.8 mm).</paragraph>
<paragraph pageId="13" pageNumber="74">
<pageBreakToken pageId="13" pageNumber="74" start="start">Head</pageBreakToken>
<subSubSection id="9899FF990E25BEF8C8D07D7B054284D4" lastPageId="13" lastPageNumber="74" pageId="12" pageNumber="73" type="description">
<paragraph id="101B81DA3CD5CE857E5DE0B16AC88E53" pageId="12" pageNumber="73">Description.</paragraph>
<paragraph id="3071FBE639DB20225511AAE2C775F912" pageId="12" pageNumber="73">Holotype, ♀, length of body 13.9 mm (of fore wing 9.8 mm).</paragraph>
<paragraph id="D61D5603B90CB0F8638ACD39F123C8AA" pageId="13" pageNumber="74">
<pageBreakToken id="3C8A32743EED355A5AB009ADDF0FC67B" pageId="13" pageNumber="74" start="start">Head</pageBreakToken>
. Antenna with 12 segments and penultimate segment 1.2 times as wide as apical segment in dorsal view (Fig. 38); frons moderately reticulate; vertex coarsely reticulate-punctate (Fig. 42); temple coarsely punctate; malar space densely punctulate; head narrowed behind eyes (Fig. 42); clypeus rather coarsely reticulate and with nearly complete median crest (Fig. 41).
</paragraph>
<paragraph pageId="13" pageNumber="74">Mesosoma. Length of mesosoma 1.6 times its height (Fig. 34); mesopleuron largely coarsely reticulate-punctate; metapleuron densely punctulate anteriorly and coarsely obliquely rugose posteriorly, with a wide smooth shiny band above it (Fig. 37); pronotum, mesoscutum, scutellum and metanotum coarsely reticulate-punctate, interspaces between punctures of pronotum and metanotum medio-dorsally present, sparsely punctulate and usually 0.5-1.0 times as wide as punctures (Fig. 36); metanotum medially moderately convex and not protruding above level of scutellum (Fig. 34); entire propodeum densely and rather coarsely reticulate-rugose, medially coarser than laterally (Fig. 36).</paragraph>
<paragraph pageId="13" pageNumber="74">Wings. Fore wing: vein 2m-cu just postfurcal (Fig. 35).</paragraph>
<paragraph pageId="13" pageNumber="74">Legs. Hind basitarsus comparatively slender (Fig. 39).</paragraph>
<paragraph pageId="13" pageNumber="74">Metasoma. Metasoma comparatively slender in dorsal view (Fig. 33); basal tergites finely punctate and shiny, with smooth interspaces wider than diameter of punctures (Fig. 33); hypopygium as long as fifth sternite ventrally (Fig. 34); ovipositor with rather widely separated serrations (Fig. 40).</paragraph>
<paragraph pageId="13" pageNumber="74">Colour. Black; ivory: pair of wide c-shaped lateral patches on clypeus (resulting in a comparatively narrow black anchor medially; Fig. 41), patch at inner orbita up to top of incision of eye (Fig. 41), shelf of frons anteriorly between antennal sockets, temple largely except narrowly dorsally (Fig. 43), transverse stripe on pronotum (but comparatively narrowly interrupted medially (Fig. 36) and ventrally comparatively wide (Fig. 37)), small patch on pronotum postero-dorsally, elongate patch on mesopleuron antero-dorsally, patch on border of propodeum and metapleuron, small patch near tegula, axilla, lateral patch on metanotum, elongate apical patch on femora, fore femur largely ventrally and elongate basal patch on fore and middle tibiae and outer side of fore tibia subapically largely; metasoma orange red; inner side of fore femur and tibia brown; tarsi largely, veins and pterostigma, dark brown; fore coxa densely silvery setose; first submarginal cell apically, marginal cell and second and third submarginal cells of fore wing dark brown, area below it brown and remainder largely subhyaline (Fig. 35).</paragraph>
<paragraph pageId="13" pageNumber="74">Male. Unknown.</paragraph>
<paragraph id="921A21AE4732F0E8D0F791F981336743" pageId="13" pageNumber="74">Mesosoma. Length of mesosoma 1.6 times its height (Fig. 34); mesopleuron largely coarsely reticulate-punctate; metapleuron densely punctulate anteriorly and coarsely obliquely rugose posteriorly, with a wide smooth shiny band above it (Fig. 37); pronotum, mesoscutum, scutellum and metanotum coarsely reticulate-punctate, interspaces between punctures of pronotum and metanotum medio-dorsally present, sparsely punctulate and usually 0.5-1.0 times as wide as punctures (Fig. 36); metanotum medially moderately convex and not protruding above level of scutellum (Fig. 34); entire propodeum densely and rather coarsely reticulate-rugose, medially coarser than laterally (Fig. 36).</paragraph>
<paragraph id="E7E07B534E3B2CD32BDE91E865157D46" pageId="13" pageNumber="74">Wings. Fore wing: vein 2m-cu just postfurcal (Fig. 35).</paragraph>
<paragraph id="D21DB9096275F92A1C9B21F26D0F0404" pageId="13" pageNumber="74">Legs. Hind basitarsus comparatively slender (Fig. 39).</paragraph>
<paragraph id="4E60702322BD2644E4FD5F301F019294" pageId="13" pageNumber="74">Metasoma. Metasoma comparatively slender in dorsal view (Fig. 33); basal tergites finely punctate and shiny, with smooth interspaces wider than diameter of punctures (Fig. 33); hypopygium as long as fifth sternite ventrally (Fig. 34); ovipositor with rather widely separated serrations (Fig. 40).</paragraph>
<paragraph id="C469471D4EADA338D981A5A24F7E417C" pageId="13" pageNumber="74">Colour. Black; ivory: pair of wide c-shaped lateral patches on clypeus (resulting in a comparatively narrow black anchor medially; Fig. 41), patch at inner orbita up to top of incision of eye (Fig. 41), shelf of frons anteriorly between antennal sockets, temple largely except narrowly dorsally (Fig. 43), transverse stripe on pronotum (but comparatively narrowly interrupted medially (Fig. 36) and ventrally comparatively wide (Fig. 37)), small patch on pronotum postero-dorsally, elongate patch on mesopleuron antero-dorsally, patch on border of propodeum and metapleuron, small patch near tegula, axilla, lateral patch on metanotum, elongate apical patch on femora, fore femur largely ventrally and elongate basal patch on fore and middle tibiae and outer side of fore tibia subapically largely; metasoma orange red; inner side of fore femur and tibia brown; tarsi largely, veins and pterostigma, dark brown; fore coxa densely silvery setose; first submarginal cell apically, marginal cell and second and third submarginal cells of fore wing dark brown, area below it brown and remainder largely subhyaline (Fig. 35).</paragraph>
<paragraph id="36B383EE4DE7AE67D5A029CA2E6E1130" pageId="13" pageNumber="74">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="74" type="distribution">
<paragraph pageId="13" pageNumber="74">Distribution.</paragraph>
<paragraph pageId="13" pageNumber="74">Indonesia (Sumatra).</paragraph>
<subSubSection id="F7BF102B53488527974C6E9CFA1C1E1C" pageId="13" pageNumber="74" type="distribution">
<paragraph id="7908100301187F6E369E54BEDDC22F43" pageId="13" pageNumber="74">Distribution.</paragraph>
<paragraph id="08AE6908B32EC4A32EC4C024C0846C73" pageId="13" pageNumber="74">Indonesia (Sumatra).</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="74" type="etymology">
<paragraph pageId="13" pageNumber="74">Etymology.</paragraph>
<paragraph pageId="13" pageNumber="74">Named in honour of one of the collectors, Yvonne van Nierop (Leiden) for all her collecting efforts in N. Sumatra.</paragraph>
<subSubSection id="92DBD5D2D400067CAD1610F309DA4910" pageId="13" pageNumber="74" type="etymology">
<paragraph id="BBD56372E1B2EEEC1B63ECC8A07CF069" pageId="13" pageNumber="74">Etymology.</paragraph>
<paragraph id="E7C575BF080E2E281A7A1545A8B1C0B0" pageId="13" pageNumber="74">Named in honour of one of the collectors, Yvonne van Nierop (Leiden) for all her collecting efforts in N. Sumatra.</paragraph>
</subSubSection>
</treatment>
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