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<taxonomicName LSID="http://zoobank.org/A20109F5-2355-42D5-A6B7-7F11F2B00598" authority="Gadallah &amp; Soliman" class="Insecta" family="Bradynobaenidae" genus="Micatagla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Micatagla allaqiensis" order="Hymenoptera" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="allaqiensis">Micatagla allaqiensis Gadallah &amp; Soliman</taxonomicName>
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Figs 19-24
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<paragraph pageId="6" pageNumber="77">Material examined.</paragraph>
<paragraph pageId="6" pageNumber="77">
<subSubSection id="7F25A9237BF62CC4604C9EE9EEC041AC" pageId="6" pageNumber="77" type="material examined">
<paragraph id="8228F3B66C84A58525D0D366C9D0589F" pageId="6" pageNumber="77">Material examined.</paragraph>
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Holotype ♀: Southern Egypt, Wadi Allaqi, Aswan [
<geoCoordinate direction="north" orientation="latitude" precision="15" value="22.83889">22°50'20&quot;N</geoCoordinate>
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,
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<geoCoordinate id="8C57C70B2AD168DE5D6BBCCA41F07515" direction="east" orientation="longitude" precision="15" value="33.198334">33°11'54&quot;E</geoCoordinate>
], 20.xii.2012 (leg. Ahmed M. Soliman) [CUE].
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<paragraph pageId="6" pageNumber="77">Description.</paragraph>
<paragraph pageId="6" pageNumber="77">Female (Holotype). Body length 8 mm.</paragraph>
<paragraph pageId="6" pageNumber="77">Colour. Medium yellowish brown, except antennal flagellum brown, mandible reddish brown distally; fore tibial spur light red, mid and hind tibial spurs waxy white; mid and hind coxae with yellow apices; T2 with apical yellow band widened medially, T3 black with apical yellowish band, S3 black on basal two-thirds, pygidial area dark brown with black carinae.</paragraph>
<paragraph pageId="6" pageNumber="77">Pubescence. Body as well as legs rather densely clothed with erect to suberect white hairs (recumbent on face, denser on clypeus and metasomal segments 4-5 than elsewhere). T1-3 with apical fringe of compact inward directed white hairs, that is weak on T1; S2 and S5 with apical fringe of obviously scattered white hairs.</paragraph>
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<paragraph id="AF7129EE4AC5B31ABFEE8B59F070B992" pageId="6" pageNumber="77">Description.</paragraph>
<paragraph id="95C543D3CAE4A541857F5BAD8354E63B" pageId="6" pageNumber="77">Female (Holotype). Body length 8 mm.</paragraph>
<paragraph id="3DDE44B61D8C6743BC1A513217AB9508" pageId="6" pageNumber="77">Colour. Medium yellowish brown, except antennal flagellum brown, mandible reddish brown distally; fore tibial spur light red, mid and hind tibial spurs waxy white; mid and hind coxae with yellow apices; T2 with apical yellow band widened medially, T3 black with apical yellowish band, S3 black on basal two-thirds, pygidial area dark brown with black carinae.</paragraph>
<paragraph id="247E750EE5295F54C022C81708B2FEBC" pageId="6" pageNumber="77">Pubescence. Body as well as legs rather densely clothed with erect to suberect white hairs (recumbent on face, denser on clypeus and metasomal segments 4-5 than elsewhere). T1-3 with apical fringe of compact inward directed white hairs, that is weak on T1; S2 and S5 with apical fringe of obviously scattered white hairs.</paragraph>
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Head. (Fig. 20): In dorsal view a little wider than pronotum and abruptly converging behind eyes; vertex gently sloping posteriorly; occiput feebly convex; vertex, face and gena shallowly sparsely punctate, punctures on gena more dispersed; ocular orbit deeply punctate; eye small, subspherical and prominent, located above midline between upper margin of clypeus and vertex; malar space and antennal scrobe polished; distance between antennal tubercles noticeably shorter than tubercle length; clypeus strongly bent downward, polished apically, punctate basally; gena with obsolescent obtuse tubercle behind eye immediately below lower ocular margin; malar space markedly long (slightly shorter than longitudinal eye diameter); mandible slender, sickle-
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, edentate; palpi with long and slender segments. Scape of antenna 2.35
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as long as F1, gently and evenly convex from above; flagellomeres polished, F1 scarcely longer than F2, F2 as long as F3, F2-7 flattened beneath.
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<paragraph id="19CD6E770BAE1D685EADBC3B4C242E63" pageId="7" pageNumber="78">
Mesosoma (Fig. 21). Pronotum 0.4
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="61EF7EA24016DACE9F7AE717E98B5653" originalValue="×">x</normalizedToken>
as long as its maximal width, with anterior face abruptly declivous, humeral angle gently rounded and posterior margin broadly concave; dorsally deeply foveate-reticulate; horizontally carinate laterally; remainder of mesosomal dorsum foveate-striate. Propodeal posterior face gently declivous and impunctate. Mesopleuron deeply foveate-reticulate; metapleuron faintly horizontally carinate. Mesosternum strongly bidentate in front of hind coxae. Inner hind tibial spur long (0.85
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<normalizedToken id="23EA8E6553522856782EEB7673014F5B" originalValue="×">x</normalizedToken>
as long as basal tarsomere).
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Metasoma. T1 (Fig. 22) widened posteriorly (pear-shaped), as long as its maximal width, densely foveate, abruptly declivous along posterior rim; T2 semicircular, 0.6
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="1E7447B3F049614280BD81A4AC70B8F6" originalValue="×">x</normalizedToken>
as long as wide, strongly petiolate anteriorly; T3 narrowed anteriorly forming a constriction with T2; T2 and T3 with ellipsoid foveae separated by longitudinal ridges (Fig. 23), foveae on T3 slimmer than those on T2, ridges on apical fourth of T3 fine and closer to each other than elsewhere; T6 (pygidium) subtriangular, with longitudinal interrupted widely spaced carinae, bordered laterally with small sharp teeth that progressively reduce in size and become blunt distally, apical ones rounded (Fig. 24).
<pageBreakToken pageId="8" pageNumber="79" start="start">S</pageBreakToken>
<pageBreakToken id="C55C43D02C4D93BE1237B180B0341005" pageId="8" pageNumber="79" start="start">S</pageBreakToken>
1 smooth, transversely carinate basally; S2 deeply punctate-reticulate (impunctate on anterior declivity and posteromedially); S3 densely punctate laterally, impunctate medially; S4 and S5 with few scattered punctures; S2-5 with a row of punctures along their apical margins.
</paragraph>
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<paragraph pageId="8" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="79">The specific name originates from Wadi Allaqi (Aswan, southern Egypt), the type locality.</paragraph>
<subSubSection id="80AAAEC2ADF6DB08204177E923758F0F" pageId="8" pageNumber="79" type="etymology">
<paragraph id="4EEC499049785E470DB24CDE7EB57E9D" pageId="8" pageNumber="79">Etymology.</paragraph>
<paragraph id="EC695197CAC8AD208135A0DC3093A782" pageId="8" pageNumber="79">The specific name originates from Wadi Allaqi (Aswan, southern Egypt), the type locality.</paragraph>
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<mods:namePart>Spirin, Viacheslav</mods:namePart>
<mods:namePart id="D89A3D3BBECAE7301E22F0D03EE72CD3">Spirin, Viacheslav</mods:namePart>
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</mods:role>
<mods:namePart>Vlasak, Josef</mods:namePart>
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<mods:namePart>Rivoire, Bernard</mods:namePart>
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<mods:namePart>Stenroos, Soili</mods:namePart>
<mods:namePart id="E38FACF79698829E4132F80B29D9EF0D">Stenroos, Soili</mods:namePart>
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<mods:namePart>Hibbett, David S.</mods:namePart>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>19</mods:number>
<mods:part id="9705330B8B76EA451E96B008EFB3711A">
<mods:date id="5479A673AC57215290AF88F3579F1618">2017</mods:date>
<mods:detail id="2B2335DC2E681DE7A3C00767C5734FDB" type="volume">
<mods:number id="EDEC790399219B65D59D080B3D91FC81">19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>43</mods:start>
<mods:end>44</mods:end>
<mods:extent id="0F5E62BBAA35CA744D395DEDA813A171" unit="page">
<mods:start id="7CE8685957ABC161F804DC04448C4ABD">43</mods:start>
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<mods:identifier type="Pensoft-Pub">1314-4049-19-43</mods:identifier>
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<mods:identifier id="01997A48139A5675A7E425B9EE3869E9" type="DOI">http://dx.doi.org/10.3897/mycokeys.19.11562</mods:identifier>
<mods:identifier id="A9ACE71EEED522BD25A61E367EF2E34D" type="Pensoft-Pub">1314-4049-19-43</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127884467" LSID="urn:lsid:plazi:treatment:6D0FC1B0D5E9AD5DED23B15C45A87EC3" httpUri="http://treatment.plazi.org/id/6D0FC1B0D5E9AD5DED23B15C45A87EC3" lastPageNumber="43" pageId="0" pageNumber="43">
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<taxonomicName LSID="819500" authority="Miettinen &amp; Spirin" class="Agaricomycetes" family="Phanerochaetaceae" genus="Ceriporia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ceriporia mpurii" order="Polyporales" pageId="0" pageNumber="43" phylum="Basidiomycota" rank="species" species="mpurii">Ceriporia mpurii Miettinen &amp; Spirin</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
<treatment id="6D0FC1B0D5E9AD5DED23B15C45A87EC3" ID-GBIF-Taxon="127884467" LSID="urn:lsid:plazi:treatment:6D0FC1B0D5E9AD5DED23B15C45A87EC3" httpUri="http://treatment.plazi.org/id/6D0FC1B0D5E9AD5DED23B15C45A87EC3" lastPageNumber="43" pageId="0" pageNumber="43" scope_family="Phanerochaetaceae" scope_order="Polyporales" scope_phylum="Basidiomycota">
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<paragraph id="C0FDBCB118CBA116003D851CCDE668FD" pageId="0" pageNumber="43">
<taxonomicName id="681448E04C17B5B9DD2615FF2BDC5BD8" LSID="819500" authority="Miettinen &amp; Spirin" class="Agaricomycetes" family="Phanerochaetaceae" genus="Ceriporia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ceriporia mpurii" order="Polyporales" pageId="0" pageNumber="43" phylum="Basidiomycota" rank="species" species="mpurii">Ceriporia mpurii Miettinen &amp; Spirin</taxonomicName>
<taxonomicNameLabel id="1FA0B2B18EBCF5CB8A5341D318AC4C06" pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="description">
<paragraph pageId="0" pageNumber="43">Validating description.</paragraph>
<paragraph pageId="0" pageNumber="43">
<subSubSection id="07C3919CEC07B3D3017FE0B26232A5E7" pageId="0" pageNumber="43" type="description">
<paragraph id="3E3F95BF4115356676DCF4B32CE6BBBF" pageId="0" pageNumber="43">Validating description.</paragraph>
<paragraph id="FAB4870D5E93C903D3AEDC48A795D77A" pageId="0" pageNumber="43">
Under '
<taxonomicName class="Agaricomycetes" family="Phanerochaetaceae" genus="Ceriporia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ceriporia mpurii" order="Polyporales" pageId="0" pageNumber="43" phylum="Basidiomycota" rank="species" species="mpurii">Ceriporia mpurii</taxonomicName>
<taxonomicName id="F53DF5075B3482A64515C0A8EC822B07" class="Agaricomycetes" family="Phanerochaetaceae" genus="Ceriporia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ceriporia mpurii" order="Polyporales" pageId="0" pageNumber="43" phylum="Basidiomycota" rank="species" species="mpurii">Ceriporia mpurii</taxonomicName>
Miettinen &amp;
<normalizedToken originalValue="Spirin">Spirin'</normalizedToken>
<normalizedToken id="4FCD804093EBC209FDB72FF5DB6A1665" originalValue="Spirin">Spirin'</normalizedToken>
in MycoKeys 17: 35 (2016), nom. inval. (Art. 40.7 - no holotype specified).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="materials_examined">
<paragraph pageId="0" pageNumber="43">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="43">
<subSubSection id="301B635E7372AF1A174AF11AE3F90346" pageId="0" pageNumber="43" type="materials_examined">
<paragraph id="30B62D028CDC96F6CBD65A4112227F3B" pageId="0" pageNumber="43">Holotype.</paragraph>
<paragraph id="10881B2B91CDF7437218AAE58D0DD0ED" pageId="0" pageNumber="43">
Indonesia. Papua Barat: Saukorem, Minjanbiat, -0.5755°: 133.1447°, lowland primary forest, fallen trunk of
<taxonomicName class="Magnoliopsida" family="Anacardiaceae" genus="Spondias" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Spondias" order="Sapindales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="genus">Spondias</taxonomicName>
<taxonomicName id="418ABAC711B0C9B328FA61FA39213823" class="Magnoliopsida" family="Anacardiaceae" genus="Spondias" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Spondias" order="Sapindales" pageId="0" pageNumber="43" phylum="Tracheophyta" rank="genus">Spondias</taxonomicName>
(40 cm in diameter, decay stage 4/5), 3 Nov 2010, Miettinen 14381 (MAN, isotypes H, ANDA).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="notes">
<paragraph pageId="0" pageNumber="43">Note.</paragraph>
<paragraph pageId="0" pageNumber="43">The species epithet is corrected according to Art. 23.1 and Art. 60.12 Ex. 33.</paragraph>
<subSubSection id="490A3928D221D0DB12FC93D62E53FB6C" pageId="0" pageNumber="43" type="notes">
<paragraph id="699CFF46DAE5DDCDB02AC6BAC72600BA" pageId="0" pageNumber="43">Note.</paragraph>
<paragraph id="0735BD44EFFCD5708A41E54FC41281CD" pageId="0" pageNumber="43">The species epithet is corrected according to Art. 23.1 and Art. 60.12 Ex. 33.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,182 +1,181 @@
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<taxonomicName authority="Brunner von Wattenwyl, 1893" authorityName="Brunner von Wattenwyl" authorityYear="1893" class="Insecta" family="Phaneropteridae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva Brunner von Wattenwyl, 1893</taxonomicName>
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Fig. 15morphology, Figs 16, 17 song
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examined.
</paragraph>
<paragraph lastPageId="5" lastPageNumber="44" pageId="4" pageNumber="43">
<paragraph id="DC90128F80927D6FBB54621F4C7A3EF1" lastPageId="5" lastPageNumber="44" pageId="4" pageNumber="43">
- For the behavioral studies four males and three females (out of series of nine each) were used, all offspring from animals from Thailand, Chiang Mai, Doi Suthep-Pui,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="18.8">18°48'N</geoCoordinate>
<geoCoordinate id="3CA213EB8073E4CDCC1F4918A80D4ACD" direction="north" orientation="latitude" precision="925" value="18.8">18°48'N</geoCoordinate>
,
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<geoCoordinate id="0D64B3BA0EB04B835E27BC490B4609D6" direction="east" orientation="longitude" precision="925" value="98.916664">98°55'E</geoCoordinate>
, 1100-1150 m a.s.l., 13.04.1995, ex ovo, bred in lab., leg. S. Ingrisch. Stridulatory files in six males,
<pageBreakToken pageId="5" pageNumber="44" start="start">China</pageBreakToken>
<pageBreakToken id="583BFFE64BD992807962ADDD3354577A" pageId="5" pageNumber="44" start="start">China</pageBreakToken>
, Yunnan Province, Jinping County, Fenshuiling National Nature Reserve,
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,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="103.23377">103.23377388°N</geoCoordinate>
<geoCoordinate id="5AE768162D4DF5A2F80BA9CCDAAE6A79" direction="north" orientation="latitude" precision="1" value="103.23377">103.23377388°N</geoCoordinate>
, August 2012.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44" type="description">
<paragraph pageId="5" pageNumber="44">Description.</paragraph>
<paragraph lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44">
<subSubSection id="7B46D00CFF07215C0F3EC8959BA75D44" lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44" type="description">
<paragraph id="1E1C844057D9461ABB7C5EE706C0D41A" pageId="5" pageNumber="44">Description.</paragraph>
<paragraph id="50E7AD8B83CE635E153DDCDED7530152" lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44">
- The species has been sufficiently redescribed by
<bibRefCitation author="Liu, CX" journalOrPublisher="Raffles Bulletin of Zoology" pageId="10" pageNumber="49" pagination="37 - 43" title="The genus Ectadia Brunner von Wattenwyl (Orthoptera: Tettigoniidae: Phaneropterinae), with descriptions of three new species from China." volume="52" year="2004">Liu et al. 2004</bibRefCitation>
<bibRefCitation id="C116351FBE9957E57A263572E83DF03A" author="Liu, CX" journalOrPublisher="Raffles Bulletin of Zoology" pageId="10" pageNumber="49" pagination="37 - 43" title="The genus Ectadia Brunner von Wattenwyl (Orthoptera: Tettigoniidae: Phaneropterinae), with descriptions of three new species from China." volume="52" year="2004">Liu et al. 2004</bibRefCitation>
. It may be added that the fore wings of the male are vaulted, bulging laterally in proximal half of tegmen
<pageBreakToken pageId="6" pageNumber="45" start="start">length</pageBreakToken>
<pageBreakToken id="66EEE203E0B543C57B38B1937E243822" pageId="6" pageNumber="45" start="start">length</pageBreakToken>
, flat with rounded tip thereafter; those of the female are shortened with acute tip. In both sexes the hind wings surpass the fore wings. The species shows a green-brown color polymorphism in both sexes. Of the specimens collected in the field, two males and two females were green and two males and one female brown. The brown color is of lighter shade in females (more ochreous) than in males (medium brown with dark pattern). In the offspring of a green female bred in lab, 19 males and 12 females became green, four males and nine females brown. Body, legs, fore wings and the projecting part of the hind wings were either all green or all brown, in two females pale yellowish brown. The broad dorsal field of the male tegmen including the stridulatory area was always dark brown in both color morphs while the narrow dorsal field of the female tegmen was yellow in green females or pale brown bordered by a dark brown line in brown females. In resting position,
<taxonomicName genus="E." lsidName="E. fulva" pageId="6" pageNumber="45" rank="species" species="fulva">E. fulva</taxonomicName>
<taxonomicName id="D540004D95E92F09727A0EF4E99AC92C" lsidName="E. fulva" pageId="6" pageNumber="45" rank="species" species="fulva">E. fulva</taxonomicName>
sits with the antennae and the fore legs stretched anteriorly in the body axis, the mid and the hind legs are stretched oblique posteriorly, while the abdomen and the wings are pointing dorso-posteriorly. That behavior resembles the situation in
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Ducetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ducetia" order="Orthoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">Ducetia</taxonomicName>
<taxonomicName id="F0AF1A666D2F7C84FFE6C521ED5FE73C" class="Insecta" family="Tettigoniidae" genus="Ducetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ducetia" order="Orthoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">Ducetia</taxonomicName>
(e.g.
<taxonomicName genus="D." lsidName="D. japonica" pageId="6" pageNumber="45" rank="species" species="japonica">D. japonica</taxonomicName>
<taxonomicName id="7EEB8369AF7B17D3C70378B12DA3242E" lsidName="D. japonica" pageId="6" pageNumber="45" rank="species" species="japonica">D. japonica</taxonomicName>
) or
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Elimaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elimaea" order="Orthoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">Elimaea</taxonomicName>
<taxonomicName id="65DCE2119635ED1885B3839A37DC56EC" class="Insecta" family="Tettigoniidae" genus="Elimaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elimaea" order="Orthoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">Elimaea</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="46" pageId="6" pageNumber="45" type="song.">
<paragraph pageId="6" pageNumber="45">Song.</paragraph>
<paragraph pageId="6" pageNumber="45">
<subSubSection id="9888C7C6F572676117936300E5CB4E88" lastPageId="7" lastPageNumber="46" pageId="6" pageNumber="45" type="song.">
<paragraph id="830BD89FAF493AF4AF26ACFEA8033A85" pageId="6" pageNumber="45">Song.</paragraph>
<paragraph id="30DCA7F5F509CE5A9C3DB1385E1DF6DA" pageId="6" pageNumber="45">
- Time-amplitude-pattern. The male calling song consisted of song units each lasting about 1.5 s repeated after an interval of about 4 s (Fig. 16A). Each was made of a series of microsyllables (about 15), repeated with 20 Hz, followed by a long series of impulses with decreasing intervals. This song unit is produced by opening the tegmina very widely, then closing and opening them several times only in part, resulting in the microsyllables. The microsyllables are not tick-like sounds as in
<taxonomicName genus="E." lsidName="E. diuturna" pageId="6" pageNumber="45" rank="species" species="diuturna">E. diuturna</taxonomicName>
<taxonomicName id="4EA6E8EE56AC950D306B7BF89633C076" lsidName="E. diuturna" pageId="6" pageNumber="45" rank="species" species="diuturna">E. diuturna</taxonomicName>
sp. n., but consist of compact series each with a few impulses. Then the male closed the tegmina completely and very slowly. The contact of the scraper with the fine basal part of the file produced a long series of heavily damped impulses (Fig. 16; see also fig. 7 in Heller et al. 2014).
</paragraph>
<paragraph pageId="7" pageNumber="46">
<pageBreakToken pageId="7" pageNumber="46" start="start">The</pageBreakToken>
<paragraph id="4D6E065D5F4A354BF76F844CC431F0C7" pageId="7" pageNumber="46">
<pageBreakToken id="5D81D5C95E72042263CE65D495C58B9B" pageId="7" pageNumber="46" start="start">The</pageBreakToken>
female responded immediately after the end of the male impulse series (Fig. 16), typically with one or a few impulses, occasionally with one or a small series up to a few hundred ms later in addition. Sometimes responses were observed even before the male series had ended.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46" type="carrier frequency.">
<paragraph pageId="7" pageNumber="46">Carrier frequency.</paragraph>
<paragraph pageId="7" pageNumber="46">- Both parts of the song unit had a quite similar spectral composition with two peaks (Fig. 17). Besides a narrow low-frequent peak around 10 kHz there was a broad maximum at about 50-60 kHz. The female response showed a peak at 20 kHz, in width and placement intermediate between the two male peaks.</paragraph>
<paragraph pageId="8" pageNumber="47">
<pageBreakToken pageId="8" pageNumber="47" start="start">Besides</pageBreakToken>
<subSubSection id="BB1774EFF4E1F96FFA45F80B7E022972" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46" type="carrier frequency.">
<paragraph id="CCCA915F5531CB22C87021ABB168B07C" pageId="7" pageNumber="46">Carrier frequency.</paragraph>
<paragraph id="8454E9DD41035D0B3A139618912865E5" pageId="7" pageNumber="46">- Both parts of the song unit had a quite similar spectral composition with two peaks (Fig. 17). Besides a narrow low-frequent peak around 10 kHz there was a broad maximum at about 50-60 kHz. The female response showed a peak at 20 kHz, in width and placement intermediate between the two male peaks.</paragraph>
<paragraph id="9187D488930CD09836854B950D46A43E" pageId="8" pageNumber="47">
<pageBreakToken id="7F78DC53A2934D2EBAD91F471AB54908" pageId="8" pageNumber="47" start="start">Besides</pageBreakToken>
the acoustical signals soundless vibratory body movements were observed in both sexes.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="47" type="mating.">
<paragraph pageId="8" pageNumber="47">Mating.</paragraph>
<paragraph pageId="8" pageNumber="47">
<subSubSection id="B0D9F2D8C562CE1685B4C40F225FC640" pageId="8" pageNumber="47" type="mating.">
<paragraph id="7090F376753482FDA9DA59368E2E0511" pageId="8" pageNumber="47">Mating.</paragraph>
<paragraph id="8B7C74CCDD8990041C6D45EFD8FB868D" pageId="8" pageNumber="47">
- In eight tests, a male (mean body mass 199
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="04A05ADABD4C601FC41B4EBA1590971C" originalValue="±">+/-</normalizedToken>
16 mg; n=7) and a female (mean body mass 626
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="1D26FFE91A752C0A9EB67F58859AF27D" originalValue="±">+/-</normalizedToken>
71 mg; n = 6) were placed together for mating. Four couples mated with the males transferring only very small spermatophores (1.3
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="FC58BB861A09E2D78C10ED22EB868776" originalValue="±">+/-</normalizedToken>
0.6 mg; no data on mating duration available; Fig. 18). Obviously the spermatophores consisted only from a pair of ampullas without spermatophylax.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47" type="nymphs.">
<paragraph pageId="8" pageNumber="47">Nymphs.</paragraph>
<paragraph lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47">
<subSubSection id="286C2581D6FA3A6F93E436A8C67A6F80" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47" type="nymphs.">
<paragraph id="D352316394FACED84CB05939A8186C57" pageId="8" pageNumber="47">Nymphs.</paragraph>
<paragraph id="F0A56BDD8B0FCE08889B34DCD3CA9A18" lastPageId="9" lastPageNumber="48" pageId="8" pageNumber="47">
- Postembryonic development occured over six nymphal instars as in many other
<taxonomicName lsidName="" pageId="8" pageNumber="47" rank="subfamily" subfamily="Phaneropterinae">Phaneropterinae</taxonomicName>
<taxonomicName id="0DF23CB84E7A2D53DDD9BAFEB1D3E170" lsidName="" pageId="8" pageNumber="47" rank="subfamily" subfamily="Phaneropterinae">Phaneropterinae</taxonomicName>
(Fig. 19). Development
<pageBreakToken pageId="9" pageNumber="48" start="start">from</pageBreakToken>
<pageBreakToken id="24282B9BD443DBB74369BA335CD042C2" pageId="9" pageNumber="48" start="start">from</pageBreakToken>
hatching to adult moult took 52-68 days at 20-23°C (
<bibRefCitation author="Ingrisch, S" journalOrPublisher="Tijdschrift voor Entomologie" pageId="10" pageNumber="49" pagination="65 - 108" title="A review of the Elimaeini In Western Indonesia, Malay Peninsula and Thailand (Ensifera, Phaneropteridae)." url="https://doi.org/10.1163/22119434-99900006" volume="141" year="1998">Ingrisch 1998</bibRefCitation>
<bibRefCitation id="D7D176B54B3875DBAD96E016B71DDCF2" author="Ingrisch, S" journalOrPublisher="Tijdschrift voor Entomologie" pageId="10" pageNumber="49" pagination="65 - 108" title="A review of the Elimaeini In Western Indonesia, Malay Peninsula and Thailand (Ensifera, Phaneropteridae)." url="https://doi.org/10.1163/22119434-99900006" volume="141" year="1998">Ingrisch 1998</bibRefCitation>
). Nymphs can be green or light brown. First instars appear green colored.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="48" type="distribution">
<paragraph pageId="9" pageNumber="48">Distribution</paragraph>
<paragraph pageId="9" pageNumber="48">- See Fig. 1.</paragraph>
<caption pageId="9" pageNumber="48">
<paragraph pageId="9" pageNumber="48">
<subSubSection id="7E8B2C2E7332600500D30989EAA9B5CC" pageId="9" pageNumber="48" type="distribution">
<paragraph id="E850B8DAA50DD765BEED6A394B2BECD1" pageId="9" pageNumber="48">Distribution</paragraph>
<paragraph id="2EFD737CF2EB0335BA40864E8B50431C" pageId="9" pageNumber="48">- See Fig. 1.</paragraph>
<caption id="DF5685D67D082A295EB0FD0BF7258105" pageId="9" pageNumber="48">
<paragraph id="A73B8014EB086371725E743F6D83B5C6" pageId="9" pageNumber="48">
Figure 15. Habitus of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva</taxonomicName>
<taxonomicName id="3E11D3830EBAD050A30A8186BD62FE44" class="Insecta" family="Tettigoniidae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva</taxonomicName>
. A. male, B. female.
</paragraph>
</caption>
<caption pageId="9" pageNumber="48">
<paragraph pageId="9" pageNumber="48">
<caption id="B781B58E3C98EE026FB44CF8F8D2E3FE" pageId="9" pageNumber="48">
<paragraph id="5884D76DA99E43A9F65535B54BC02A53" pageId="9" pageNumber="48">
Figure 16. Male calling song A. and female response B. of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva</taxonomicName>
<taxonomicName id="3318488DE97D28AA34F240236302A3A7" class="Insecta" family="Tettigoniidae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva</taxonomicName>
. Oscillograms of stridulatory movement and song [synchronous registration of left tegmen movement and sound (upper line: upward deflection represents opening, downward closing; lower line: sound)]. Male and female were recorded separately.
</paragraph>
</caption>
<caption pageId="9" pageNumber="48">
<paragraph pageId="9" pageNumber="48">
<caption id="C35F26238CCAC4E61DCEEFE3F7E08FAD" pageId="9" pageNumber="48">
<paragraph id="F94F37DC9CF295F3807F71315BBDF063" pageId="9" pageNumber="48">
Figure 17. Power spectra of different parts of the male calling song and of the female response of
<taxonomicName genus="E." lsidName="E. fulva" pageId="9" pageNumber="48" rank="species" species="fulva">E. fulva</taxonomicName>
<taxonomicName id="38AB486CA097EF7FC3C8D691437CB23A" lsidName="E. fulva" pageId="9" pageNumber="48" rank="species" species="fulva">E. fulva</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="9" pageNumber="48">
<paragraph pageId="9" pageNumber="48">
<caption id="15E53F90915EEAFEAE0AEA87E7D27EA6" pageId="9" pageNumber="48">
<paragraph id="F1C0B8A6403830DEB931A7B3E11F9778" pageId="9" pageNumber="48">
Figure 18. Female
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva</taxonomicName>
<taxonomicName id="3E888B0898D85746E82B0D56B67A65F8" class="Insecta" family="Tettigoniidae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva</taxonomicName>
with spermatophore. A. overview, B. detail.
</paragraph>
</caption>
<caption pageId="9" pageNumber="48">
<paragraph pageId="9" pageNumber="48">
<caption id="5F233CFA9A7C934547C5958E7A03B015" pageId="9" pageNumber="48">
<paragraph id="E323A129CC568A08A0F8E989CC2F76F8" pageId="9" pageNumber="48">
Figure 19. Nymphs of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva</taxonomicName>
<taxonomicName id="5E97B56AB647A71B8847463031C229D8" class="Insecta" family="Tettigoniidae" genus="Ectadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectadia fulva" order="Orthoptera" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fulva">Ectadia fulva</taxonomicName>
. A. First stage, B. second stage, C. third stage, D. 4th stage, E. 5th stage, male, F. 5th stage, female, G. 6th stage, female, H, I. 6th stage, male.
</paragraph>
</caption>

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@ -1,225 +1,225 @@
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<mods:namePart>Grego 2, Jozef</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
<mods:namePart id="1010A0164B3BCB25CFE4D5DA421224C7">Grego 2, Jozef</mods:namePart>
<mods:affiliation id="7066C13D6C380B653231A608CBEC6B1B">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="3CCC0337DE0D8D7854FAF68F149D443F" type="personal">
<mods:role id="1045BC2EE25EA5DA210EBE5F296FFB31">
<mods:roleTerm id="C89F3BCEC0A1BEB075EA309D2DEE5201">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rysiewska 1, Aleksandra</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
<mods:namePart id="0420015D0DD064CB5C50B06571F2480B">Rysiewska 1, Aleksandra</mods:namePart>
<mods:affiliation id="A4FDE625BB9E884CE2653EE3A9EEEA9D">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="6E3B8AE6F75425CCFE3295168FBA987B" type="personal">
<mods:role id="D1F3A67A1E788E0CDB971DF71426F5FE">
<mods:roleTerm id="5529CADAD5D66B9CCB0730E034050CC8">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Osikowski 3, Artur</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
<mods:namePart id="81AED5189DA7770D9E943F72758AB744">Osikowski 3, Artur</mods:namePart>
<mods:affiliation id="3AB1841A13D0837FA32C25450D25856E">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
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<mods:roleTerm id="A20BBF0AB4D97A75E644B39106C22379">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hofman 4, Sebastian</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
<mods:namePart id="01C37C48B16404D89419E970780F4941">Hofman 4, Sebastian</mods:namePart>
<mods:affiliation id="55A1F7DE09EAC15D3E55218A7A32C839">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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<mods:title>Subterranean Biology</mods:title>
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<mods:title id="549EAD3E5196374218C0DC11A9E0084B">Subterranean Biology</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-02-16</mods:number>
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<mods:number>37</mods:number>
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<mods:number id="1332E34ACF54E0DCBD9B3B27DA877522">37</mods:number>
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<subSection pageId="0" pageNumber="89" type="systematic part">
<treatment ID-GBIF-Taxon="177423715" LSID="urn:lsid:zoobank.org:act:36A97AFE-52B6-4F52-A17E-348C5CF604F3" httpUri="http://treatment.plazi.org/id/6D1C9FCF30635F25A8EFF712C4204DF3" lastPageNumber="89" pageId="0" pageNumber="89">
<subSubSection pageId="0" pageNumber="89" type="nomenclature">
<paragraph pageId="0" pageNumber="89">
<taxonomicName LSID="http://zoobank.org/36A97AFE-52B6-4F52-A17E-348C5CF604F3" authority="Grego &amp; Rysiewska, sp. nov." authorityName="Grego &amp; Rysiewska, sp. nov." class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia stygorumina" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="stygorumina">Horatia stygorumina Grego &amp; Rysiewska, sp. nov.</taxonomicName>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">Figure 4F-H</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of H. ozimeci (A) juvenile specimen from the type locality (SBMNH 635080) and H. stygorumina (B) juvenile shell from cave Mali Rumin (SBMNH 635079). SEM SBMNH by Vanessa Delnavaz." figureDoi="10.3897/subtbiol.37.61573.figure5" httpUri="https://binary.pensoft.net/fig/509944" pageId="0" pageNumber="89">, 5B</figureCitation>
<subSection id="C48CDCAC11F9490F75C70E29A8358131" pageId="0" pageNumber="89" type="systematic part">
<treatment id="6D1C9FCF30635F25A8EFF712C4204DF3" ID-GBIF-Taxon="177423715" LSID="urn:lsid:zoobank.org:act:36A97AFE-52B6-4F52-A17E-348C5CF604F3" httpUri="http://treatment.plazi.org/id/6D1C9FCF30635F25A8EFF712C4204DF3" lastPageNumber="89" pageId="0" pageNumber="89" scope_family="Hydrobiidae">
<subSubSection id="8D529A63140A987B4AF262B43051D2F1" pageId="0" pageNumber="89" type="nomenclature">
<paragraph id="1936AB6449F748E14FE05B97DB6ACEC1" pageId="0" pageNumber="89">
<taxonomicName id="E5E37FD94194F9E96D816A3ACD54C964" ID-CoL="7RPS2" LSID="http://zoobank.org/36A97AFE-52B6-4F52-A17E-348C5CF604F3" authority="Grego &amp; Rysiewska, sp. nov." authorityName="Grego &amp; Rysiewska, sp. nov." class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia stygorumina" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="stygorumina">Horatia stygorumina Grego &amp; Rysiewska, sp. nov.</taxonomicName>
<figureCitation id="2AC0710C260B49C1BD8F59146D163034" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">Figure 4F-H</figureCitation>
<figureCitation id="6A7A1D986A48CA8425434E73D229201B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of H. ozimeci (A) juvenile specimen from the type locality (SBMNH 635080) and H. stygorumina (B) juvenile shell from cave Mali Rumin (SBMNH 635079). SEM SBMNH by Vanessa Delnavaz." figureDoi="10.3897/subtbiol.37.61573.figure5" httpUri="https://binary.pensoft.net/fig/509944" pageId="0" pageNumber="89">, 5B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="type locality">
<paragraph pageId="0" pageNumber="89">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="89">
<subSubSection id="18A274DABE73CFB617D9F1B7D2A6A199" pageId="0" pageNumber="89" type="type locality">
<paragraph id="54FA8DB43EF8EE67053F877B148F4311" pageId="0" pageNumber="89">Type locality.</paragraph>
<paragraph id="C8922D63DCAA0F0847EAACFC44230E3A" pageId="0" pageNumber="89">
Mali Rumin cave, Rumin, Sinj District, Croatia,
<geoCoordinate degrees="43" direction="north" minutes="46" orientation="latitude" precision="1" seconds="50.88" value="43.7808">43°46'50.88&quot;N</geoCoordinate>
<geoCoordinate id="99111D0C19A20F85983AF860EC5342E6" degrees="43" direction="north" minutes="46" orientation="latitude" precision="1" seconds="50.88" value="43.7808">43°46'50.88&quot;N</geoCoordinate>
,
<geoCoordinate degrees="16" direction="east" minutes="38" orientation="longitude" precision="1" seconds="56.75" value="16.649097">16°38'56.75&quot;E</geoCoordinate>
<geoCoordinate id="0BDE7384A542FE34A26677BBC1A6A778" degrees="16" direction="east" minutes="38" orientation="longitude" precision="1" seconds="56.75" value="16.649097">16°38'56.75&quot;E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="holotype">
<paragraph pageId="0" pageNumber="89">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="89">
<subSubSection id="50FAEC36A9CEA4D16064E6FD621B84EF" pageId="0" pageNumber="89" type="holotype">
<paragraph id="775FCE3E23D60C3B43978880D20B4C2C" pageId="0" pageNumber="89">Holotype.</paragraph>
<paragraph id="DA50AC718AF77FE7FB9BB6403F2A7083" pageId="0" pageNumber="89">
Dry specimen, J. Grego, A. Falniowski, M.
<normalizedToken originalValue="Olšavský">Olsavsky</normalizedToken>
<normalizedToken id="D8A9A4FA819D58D07694213DD480F924" originalValue="Olšavský">Olsavsky</normalizedToken>
and J.
<normalizedToken originalValue="Olšavská">Olsavska</normalizedToken>
<normalizedToken id="363667991B7CC2917AA5DE91DCA67D8F" originalValue="Olšavská">Olsavska</normalizedToken>
leg., August 5-th 2020, OZRM 11602 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4F</figureCitation>
<figureCitation id="73435E839AC20F11076DE5125AD94853" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="paratypes">
<paragraph pageId="0" pageNumber="89">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="89">
<subSubSection id="E53EF6B9A9BAEDCB45B990662D44EA80" pageId="0" pageNumber="89" type="paratypes">
<paragraph id="624D95AD9EA1A1BD5DCC2F9108D1794D" pageId="0" pageNumber="89">Paratypes.</paragraph>
<paragraph id="B5A4EB71DF97017D24FFB55EA92EA225" pageId="0" pageNumber="89">
From Type locality: J. Grego, A. Falniowski, M.
<normalizedToken originalValue="Olšavský">Olsavsky</normalizedToken>
<normalizedToken id="110BC8045EF39C35C65E44585023D7AA" originalValue="Olšavský">Olsavsky</normalizedToken>
and J.
<normalizedToken originalValue="Olšavská">Olsavska</normalizedToken>
<normalizedToken id="E4363FFBE46B3FF77F2AEB72185B7B3D" originalValue="Olšavská">Olsavska</normalizedToken>
leg., August 5-th 2020, OZRM 11603/1 specimen; NHMW 113608/1 specimen, JG F1526/160 specimen; Type locality, J. Grego, G. Jakab and B.
<normalizedToken originalValue="Šmída">Smida</normalizedToken>
<normalizedToken id="C034EB839B4E632EE203A7F00101BCE6" originalValue="Šmída">Smida</normalizedToken>
leg. March 17-th 2017, JG F0736/26 specimens.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="materials_examined">
<paragraph pageId="0" pageNumber="89">Other material.</paragraph>
<paragraph pageId="0" pageNumber="89">
<subSubSection id="B94B6CE0CE8306AC9B0CFBC79E1DF280" pageId="0" pageNumber="89" type="materials_examined">
<paragraph id="BF728C38A96895322D160A1B592CA87E" pageId="0" pageNumber="89">Other material.</paragraph>
<paragraph id="A0C154421091843F6C4CF7F814FF33E8" pageId="0" pageNumber="89">
Morphologically similar stygobiotic
<taxonomicName authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia</emphasis>
<taxonomicName id="DED9B63D02C4D4BC3BC717C7E2B151C8" authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis id="32479DA807BF67F04A754BCA53E2B2A1" italics="true" pageId="0" pageNumber="89">Horatia</emphasis>
</taxonomicName>
shells were found in following spring localities (springs at left tributaries of Upper Cetina River: Vrelo Kosinac 1, Sinjski Obrovac, Sinj district, Croatia,
<geoCoordinate degrees="43" direction="north" minutes="43" orientation="latitude" precision="1" seconds="40.89" value="43.728027">43°43'40.89&quot;N</geoCoordinate>
<geoCoordinate id="E9C17C7C041B8BD8D1AE78CCD9ACE5C1" degrees="43" direction="north" minutes="43" orientation="latitude" precision="1" seconds="40.89" value="43.728027">43°43'40.89&quot;N</geoCoordinate>
,
<geoCoordinate degrees="16" direction="east" minutes="42" orientation="longitude" precision="1" seconds="2.29" value="16.700638">16°42'2.29&quot;E</geoCoordinate>
<geoCoordinate id="E9FABB9BD602E3FD9F6BC57147F51F65" degrees="16" direction="east" minutes="42" orientation="longitude" precision="1" seconds="2.29" value="16.700638">16°42'2.29&quot;E</geoCoordinate>
JG/8; Vrelo Kosinac 2, Sinjski Obrovac, Sinj District, Croatia,
<geoCoordinate degrees="43" direction="north" minutes="43" orientation="latitude" precision="1" seconds="43.11" value="43.72864">43°43'43.11&quot;N</geoCoordinate>
<geoCoordinate id="2E215C0682A3D78214B3EA4F1C82E4DE" degrees="43" direction="north" minutes="43" orientation="latitude" precision="1" seconds="43.11" value="43.72864">43°43'43.11&quot;N</geoCoordinate>
,
<geoCoordinate degrees="16" direction="east" minutes="42" orientation="longitude" precision="1" seconds="0.83" value="16.700232">16°42'0.83&quot;E</geoCoordinate>
<geoCoordinate id="B9454DA5C8D26389AE9F2EE837E8602D" degrees="16" direction="east" minutes="42" orientation="longitude" precision="1" seconds="0.83" value="16.700232">16°42'0.83&quot;E</geoCoordinate>
JG/4; Grabske Mlinice, Grab, Sinj District, Croatia,
<geoCoordinate degrees="43" direction="north" minutes="38" orientation="latitude" precision="1" seconds="27.26" value="43.640907">43°38'27.26&quot;N</geoCoordinate>
<geoCoordinate id="B45A355E72DD27E89742C49CBEDC7D84" degrees="43" direction="north" minutes="38" orientation="latitude" precision="1" seconds="27.26" value="43.640907">43°38'27.26&quot;N</geoCoordinate>
,
<geoCoordinate degrees="16" direction="east" minutes="46" orientation="longitude" precision="1" seconds="13.57" value="16.770435">16°46'13.57&quot;E</geoCoordinate>
<geoCoordinate id="800B97513E90C77FDB08B2D33C87104C" degrees="16" direction="east" minutes="46" orientation="longitude" precision="1" seconds="13.57" value="16.770435">16°46'13.57&quot;E</geoCoordinate>
JG/3). For the time being we treat those as
<taxonomicName genus="H." lsidName="H. stygorumina" pageId="0" pageNumber="89" rank="species" species="stygorumina">H. cf. stygorumina</taxonomicName>
<taxonomicName id="04EE80E52186177C588C9407E6BA9602" lsidName="H. stygorumina" pageId="0" pageNumber="89" rank="species" species="stygorumina">H. cf. stygorumina</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="diagnosis">
<paragraph pageId="0" pageNumber="89">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="89">
<subSubSection id="977F3F156ACE636257B960D99D6FE7A7" pageId="0" pageNumber="89" type="diagnosis">
<paragraph id="83623849D5254560EFAA6231C567261E" pageId="0" pageNumber="89">Diagnosis.</paragraph>
<paragraph id="E41D59D514033E390D71D87865DC5843" pageId="0" pageNumber="89">
Shell minute, trochiform, distinguished from the other species of
<taxonomicName authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia</emphasis>
<taxonomicName id="E24131ED025CFDFFFA0CF91285140479" authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis id="81F83D85A9835FFC977445BBB70AC64D" italics="true" pageId="0" pageNumber="89">Horatia</emphasis>
</taxonomicName>
by its very wide umbilicus showing earlier whorls inside. From the
<taxonomicName genus="H." lsidName="H. ozimeci" pageId="0" pageNumber="89" rank="species" species="ozimeci">
<emphasis italics="true" pageId="0" pageNumber="89">H. ozimeci</emphasis>
<taxonomicName id="A4AA53020CE1516EFABBD0CC19390EA9" lsidName="H. ozimeci" pageId="0" pageNumber="89" rank="species" species="ozimeci">
<emphasis id="184F7CC925980EECB6234DAFB6FA5CFB" italics="true" pageId="0" pageNumber="89">H. ozimeci</emphasis>
</taxonomicName>
s. nov. distinguished by higher conical spire and suture reaching the aperture, as well by more swollen protoconch and by slight sinuation at labral columellar margin. From stygobiotic cf.
<taxonomicName authorityName="Schutt" authorityYear="1961" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia knorri" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="knorri">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia knorri</emphasis>
<taxonomicName id="D47CA73D64C1CDAB8481D77C1C0401D9" authorityName="Schutt" authorityYear="1961" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia knorri" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="knorri">
<emphasis id="C534166321006960F1D41DC968AA93A4" italics="true" pageId="0" pageNumber="89">Horatia knorri</emphasis>
</taxonomicName>
different by much wider umbilicus and by aperture shape less declined from the columella in its labral profile.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="description">
<paragraph pageId="0" pageNumber="89">Description.</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Shell</emphasis>
<subSubSection id="02CB414B46A172B9F30A0CC55976BFC3" pageId="0" pageNumber="89" type="description">
<paragraph id="7BBFFFEAD5BCF542DB7DB05F1577A63C" pageId="0" pageNumber="89">Description.</paragraph>
<paragraph id="5C57DFC740B4387B48E0D15577123B0F" pageId="0" pageNumber="89">
<emphasis id="C10223EA67204AA3C9458F252351511E" bold="true" italics="true" pageId="0" pageNumber="89">Shell</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4F</figureCitation>
<figureCitation id="418A829BA4078B6441E96FFFBB5C05A5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4F</figureCitation>
) 2.08 mm high and 1.57 mm broad, conical, fresh shells milky white and translucent, with 3 rounded inflated whorls and deep suture, the whorls regularly tapering towards the aperture. Spire conically elevated, body whorl prominent. Smooth shell surface covered with blunt regular axial growth lines more prominent at the body whorl. Aperture almost rounded, tear-shaped, shortly attached to the body whorl by weak grove. Labral lateral profile backward protruded almost straight, columellar margin slightly sinuated at its middle. Umbilicus open.
</paragraph>
<caption doi="10.3897/subtbiol.37.61573.figure6" httpUri="https://binary.pensoft.net/fig/509945" pageId="0" pageNumber="89" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" pageId="0" pageNumber="89">Figure 6.</emphasis>
<caption id="F555744DD5A850E3A69A5467D09FF94E" doi="10.3897/subtbiol.37.61573.figure6" httpUri="https://binary.pensoft.net/fig/509945" pageId="0" pageNumber="89" start="Figure 6" startId="F6">
<paragraph id="5C8857FCDCB20B0DB8C46EBF455AC14C" pageId="0" pageNumber="89">
<emphasis id="A5023D50F9F6FA9DBF644633894C49F0" bold="true" pageId="0" pageNumber="89">Figure 6.</emphasis>
Maximum Likelihood tree inferred from mitochondrial COI. Bootstrap support above 60% and corresponding Bayesian probabilities are given. Bold indicates newly obtained sequences.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Measurements</emphasis>
<paragraph id="55D577DB6E20AC50493C8B0D1E7FA7DD" pageId="0" pageNumber="89">
<emphasis id="2C7F1FB24527C0553238FAD0B2DF8FE1" bold="true" italics="true" pageId="0" pageNumber="89">Measurements</emphasis>
of holotype of
<taxonomicName genus="H." lsidName="H. stygorumina" pageId="0" pageNumber="89" rank="species" species="stygorumina">
<emphasis italics="true" pageId="0" pageNumber="89">H. stygorumina</emphasis>
<taxonomicName id="D28CFF3604735CABEAE3799513007628" lsidName="H. stygorumina" pageId="0" pageNumber="89" rank="species" species="stygorumina">
<emphasis id="64288CDC53838CC9BF47AA44354C9845" italics="true" pageId="0" pageNumber="89">H. stygorumina</emphasis>
</taxonomicName>
: Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Shell measurements of the Horatia: I-IV H. ozimeci (I sequenced specimen, II holotype, in bold; III-IV paratypes); V-VII sequenced specimens of H. klecakiana; VIII H. stygorumina, holotype, in bold. Measurements are shown in Fig. 4 J." httpUri="http://table.plazi.org/id/B70765E2784600EB05E676C7B53632D9" pageId="0" pageNumber="89" tableUuid="B70765E2784600EB05E676C7B53632D9">2</tableCitation>
<tableCitation id="5DC0533BD4CFC860FA23AF5A6968001E" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Shell measurements of the Horatia: I-IV H. ozimeci (I sequenced specimen, II holotype, in bold; III-IV paratypes); V-VII sequenced specimens of H. klecakiana; VIII H. stygorumina, holotype, in bold. Measurements are shown in Fig. 4 J." httpUri="http://table.plazi.org/id/B70765E2784600EB05E676C7B53632D9" pageId="0" pageNumber="89" tableUuid="B70765E2784600EB05E676C7B53632D9">2</tableCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" pageId="0" pageNumber="89">
<emphasis italics="true" pageId="0" pageNumber="89">Soft parts morphology and anatomy</emphasis>
<paragraph id="B1B334525E0C71EC85FADB2E859987C6" pageId="0" pageNumber="89">
<emphasis id="319CDD21668E2DF46BEB7367E960CD46" bold="true" pageId="0" pageNumber="89">
<emphasis id="43D3FDC722645A833FC0AAE31827869E" italics="true" pageId="0" pageNumber="89">Soft parts morphology and anatomy</emphasis>
.
</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="etymology">
<paragraph pageId="0" pageNumber="89">Derivatio nominis.</paragraph>
<paragraph pageId="0" pageNumber="89">
<subSubSection id="B26DAE203ACA15AA960C97C0014778C8" pageId="0" pageNumber="89" type="etymology">
<paragraph id="D87944FD2A3853B642089C2D3DE85772" pageId="0" pageNumber="89">Derivatio nominis.</paragraph>
<paragraph id="85C16BF8AA3932E5EAF318FF9DB08EBC" pageId="0" pageNumber="89">
The specific epithet
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia stygorumina" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="stygorumina">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia stygorumina</emphasis>
<taxonomicName id="4D1C81218FF03B9186D73C0D25B60F14" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia stygorumina" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="stygorumina">
<emphasis id="C66D9B4C5104854561F9B87BD4E697C8" italics="true" pageId="0" pageNumber="89">Horatia stygorumina</emphasis>
</taxonomicName>
is derived from the stygobiont habitat and from the type locality: cave Mali Rumin also referring to the name of the nearby settlement Rumin.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="distribution">
<paragraph pageId="0" pageNumber="89">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="89">
<subSubSection id="E7D7A9017E305FAB23412A2FF2F39F99" pageId="0" pageNumber="89" type="distribution">
<paragraph id="B6DA858194B310C2858D061491AD469C" pageId="0" pageNumber="89">Distribution and habitat.</paragraph>
<paragraph id="FF1B267725D96DABB89B53916D4E2D68" pageId="0" pageNumber="89">
Known only from the type locality, where empty shells can be found in the cave sediments, as well as in the sandy sediments of the intermittent spring. The shells are washed out from their stygobiont habitat. The cave is 50 m long, with two branches, acting as an intermittent overflow of the larger permanent spring Vrilo Rumin, situated 730 m east-southeast. Both springs are draining karstwater from middle part of Livansko Polje Basin (Bosnia and Herzegovina) Under Dinara Mts. towards upper Cetina River Valley. The following
<taxonomicName class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
<taxonomicName id="13890E232C6A1F1ECA4A2419752F696A" class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
were detected in the locality:
<taxonomicName authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia klecakiana" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia klecakiana</emphasis>
<taxonomicName id="EB21A2550240FA7DAACBFF1784B2BA30" authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia klecakiana" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="klecakiana">
<emphasis id="DFB4AB08A4E765CD4CED4CD7245F4407" italics="true" pageId="0" pageNumber="89">Horatia klecakiana</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Frauenfeld" baseAuthorityYear="1863" genus="Orientalina" lsidName="Orientalina curta subsp. germari" pageId="0" pageNumber="89" rank="subSpecies" species="curta" subSpecies="germari">
<emphasis italics="true" pageId="0" pageNumber="89">Orientalina curta germari</emphasis>
<taxonomicName id="A04D130419706EE6B5C0C5A8A1F85C06" baseAuthorityName="Frauenfeld" baseAuthorityYear="1863" class="Gastropoda" family="Hydrobiidae" genus="Orientalina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientalina curta subsp. germari" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="subSpecies" species="curta" subSpecies="germari">
<emphasis id="5C8A49D29A2AA84A9ADE917B403F4047" italics="true" pageId="0" pageNumber="89">Orientalina curta germari</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Kuščer" baseAuthorityYear="1933" class="Gastropoda" family="Hydrobiidae" genus="Kerkia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Kerkia jadertina subsp. sinjana" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="subSpecies" species="jadertina" subSpecies="sinjana">
<emphasis italics="true" pageId="0" pageNumber="89">Kerkia jadertina sinjana</emphasis>
<taxonomicName id="A159825A297125F717BD73227ECC4AC3" baseAuthorityName="Kuščer" baseAuthorityYear="1933" class="Gastropoda" family="Hydrobiidae" genus="Kerkia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Kerkia jadertina subsp. sinjana" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="subSpecies" species="jadertina" subSpecies="sinjana">
<emphasis id="21363F30B4080D3582FB643B1C89FCB8" italics="true" pageId="0" pageNumber="89">Kerkia jadertina sinjana</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Kuščer">Kuscer</normalizedToken>
<normalizedToken id="AC387C5DDEF3421FCE3DCC331530518F" originalValue="Kuščer">Kuscer</normalizedToken>
, 1933) and a Montesieriid species of
<taxonomicName authorityName="Pavlovic" authorityYear="1913" class="Gastropoda" family="Hydrobiidae" genus="Paladilhiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paladilhiopsis" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Paladilhiopsis</emphasis>
<taxonomicName id="4CBD51EF11520EB233C649A6B843B1CD" authorityName="Pavlovic" authorityYear="1913" class="Gastropoda" family="Hydrobiidae" genus="Paladilhiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paladilhiopsis" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis id="9638D791AA26AA0BA05DA718A56EE1AC" italics="true" pageId="0" pageNumber="89">Paladilhiopsis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Pavlović">Pavlovic</normalizedToken>
<normalizedToken id="B37B582B8D44390BAFC3B6C084B8658A" originalValue="Pavlović">Pavlovic</normalizedToken>
, 1913 and/or
<taxonomicName authorityName="Brusina" authorityYear="1906" class="Gastropoda" family="Hydrobiidae" genus="Lanzaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lanzaia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Lanzaia</emphasis>
<taxonomicName id="D0ADA72C172935B7B8D727CD6198D7C7" authorityName="Brusina" authorityYear="1906" class="Gastropoda" family="Hydrobiidae" genus="Lanzaia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lanzaia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis id="B0A15BB082DA068418E18E0D04A43E15" italics="true" pageId="0" pageNumber="89">Lanzaia</emphasis>
</taxonomicName>
Brusina, 1906.
</paragraph>

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@ -1,232 +1,231 @@
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<mods:namePart>Zawisza, Marlena</mods:namePart>
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<mods:date>2017</mods:date>
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<treatment ID-GBIF-Taxon="137663109" LSID="urn:lsid:zoobank.org:act:C8F33D89-AEDF-45FB-B748-2078A73A5F2B" httpUri="http://treatment.plazi.org/id/6D471A9A923AF607F520311B881C9D04" lastPageNumber="45" pageId="0" pageNumber="45">
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<taxonomicName LSID="http://zoobank.org/C8F33D89-AEDF-45FB-B748-2078A73A5F2B" authority="Skarzynski &amp; Kahrarian" class="Collembola" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypogastrura ellisi" order="Collembola" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ellisi">
<treatment id="6D471A9A923AF607F520311B881C9D04" ID-GBIF-Taxon="137663109" LSID="urn:lsid:zoobank.org:act:C8F33D89-AEDF-45FB-B748-2078A73A5F2B" httpUri="http://treatment.plazi.org/id/6D471A9A923AF607F520311B881C9D04" lastPageNumber="45" pageId="0" pageNumber="45" scope_class="Collembola" scope_family="Hypogastruridae">
<subSubSection id="128C5136716D06E9E261B57617BBF211" pageId="0" pageNumber="45" type="nomenclature">
<paragraph id="6570A93205FB5FF8CCB412FC75AB3D6A" pageId="0" pageNumber="45">
<taxonomicName id="3724A85E37E997D53543FC66CBE1429E" LSID="http://zoobank.org/C8F33D89-AEDF-45FB-B748-2078A73A5F2B" authority="Skarzynski &amp; Kahrarian" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hypogastrura ellisi" order="Collembola" pageId="0" pageNumber="45" rank="species" species="ellisi">
Hypogastrura ellisi
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
<normalizedToken id="762CC1C804F2B0537B7ECCD135D6C485" originalValue="Skarżyński">Skarzynski</normalizedToken>
&amp; Kahrarian
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. n.</taxonomicNameLabel>
<taxonomicNameLabel id="AE96463276C2FB114FF781238C358610" pageId="0" pageNumber="45">sp. n.</taxonomicNameLabel>
Figs 1-3, 4-8, 9-10
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">Type material.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="4DE6857CE47D0FB8EE94C8CFAE605A3C" pageId="0" pageNumber="45" type="type material">
<paragraph id="2A7460B70AA135AED8FF4C3CF651233B" pageId="0" pageNumber="45">Type material.</paragraph>
<paragraph id="4919AC6B6C4D9338EF8F4F1073F42577" pageId="0" pageNumber="45">
Holotype: female on slide, litter in oak forest, Zagros Mountains, Dalab mountain (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="33.566666">33°34'N</geoCoordinate>
<geoCoordinate id="4F43819F4661431D3DE2B95266E941B9" direction="north" orientation="latitude" precision="925" value="33.566666">33°34'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="47.516666">47°31'E</geoCoordinate>
<geoCoordinate id="912672D751235A3BB1A0F1DFEF809BE4" direction="east" orientation="longitude" precision="925" value="47.516666">47°31'E</geoCoordinate>
/ 1700 m a.s.l.), Kohdasht County, Lorestan Province, Iran, 4.XII.2013, leg. M. Kahrarian. Paratypes: 4 females, 1 male, same data as holotype; 1 female, 1 male, litter in oak forest, Zagros Mountains, Sorkhdom mountain (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="33.566666">33°34'N</geoCoordinate>
<geoCoordinate id="C700533799E09F767A61C51F8B432091" direction="north" orientation="latitude" precision="925" value="33.566666">33°34'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="47.533333">47°32'E</geoCoordinate>
<geoCoordinate id="5B0ECD3AEDB45272248AE7838ED94827" direction="east" orientation="longitude" precision="925" value="47.533333">47°32'E</geoCoordinate>
/ 1650 m a.s.l.), Kohdasht County, Lorestan Province, Iran, 14.XI.2013, leg. M. Kahrarian; 2 females, 1 male, litter in oak forest, Zagros Mountains, near Patogh ghaut (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="34.416668">34°25'N</geoCoordinate>
<geoCoordinate id="66020CF164FAFF7E347C1BB189677D35" direction="north" orientation="latitude" precision="925" value="34.416668">34°25'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="46.0">46°00'E</geoCoordinate>
<geoCoordinate id="A266AE233275F03DB049A90084A20110" direction="east" orientation="longitude" precision="925" value="46.0">46°00'E</geoCoordinate>
/ 1030 m a.s.l.), Sarpol-e-zahab County, Kermanshah Province, Iran, 9.II.2014, leg. M. Kahrarian; 1 male, oak forest, Zagros Mountains, near Shabankareh village (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="34.866665">34°52'N</geoCoordinate>
<geoCoordinate id="4BDEAC5CC7824973BBB89E60A875701F" direction="north" orientation="latitude" precision="925" value="34.866665">34°52'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="46.5">46°30'E</geoCoordinate>
<geoCoordinate id="8739B9F4BA6F8B1C83A2E6BB4F333AE7" direction="east" orientation="longitude" precision="925" value="46.5">46°30'E</geoCoordinate>
/ 1600 m a.s.l.), Paveh County, Kermanshah Province, Iran, 20.I.2014, leg. M. Kahrarian. Holotype and 7 paratypes deposited at the Department of Agronomy, Kermanshah Branch, Islamic Azad University, Kermanshah, Iran and 4 paratypes deposited in the collection of the Institute of Environmental Biology, University of
<normalizedToken originalValue="Wrocław">Wroclaw</normalizedToken>
<normalizedToken id="DD3C2A95FFA74FED39853E8BBCAD6CD1" originalValue="Wrocław">Wroclaw</normalizedToken>
, Poland.
</paragraph>
<caption pageId="0" pageNumber="45">
<paragraph pageId="0" pageNumber="45">
<caption id="E214839EE564E151A1F71D1C86C3C7AD" pageId="0" pageNumber="45">
<paragraph id="D2426BBA711FA2A30E377D7217AF3197" pageId="0" pageNumber="45">
Figures 1-3.
<taxonomicName class="Collembola" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypogastrura ellisi" order="Poduromorpha" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ellisi">Hypogastrura ellisi</taxonomicName>
<taxonomicName id="307AF2964A130A4730D0143BBAF5ACEE" class="Collembola" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypogastrura ellisi" order="Poduromorpha" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ellisi">Hypogastrura ellisi</taxonomicName>
sp. n. 1 chaetotaxy of head and Th. I 2 chaetotaxy of Th.
<normalizedToken originalValue="IIIII">II-III</normalizedToken>
<normalizedToken id="39B8B397C534489D9CEDC8B1B8A45B10" originalValue="IIIII">II-III</normalizedToken>
3 chaetotaxy of Abd.
<normalizedToken originalValue="IIIVI">III-VI</normalizedToken>
<normalizedToken id="4BDE23597D19E8452194007E20020F2B" originalValue="IIIVI">III-VI</normalizedToken>
. Scale bar: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="materials_examined">
<paragraph pageId="0" pageNumber="45">Other material.</paragraph>
<paragraph pageId="0" pageNumber="45">Greece, Lesbos, leg. Ellis (deposited at the Naturalis Biodiversity Center, Netherland): 22 females, 6 males (21038-21054, 21056-21059, 21062-21064, 21066-21068), Antissa, 30.X.1973, 973240; 18 females, 23 males (21060, 21078-21117), Antissa, 30.X.1973, 973243; 1 male (21061), Antissa, 30.X.1973, 973244; 1 male (21037), Gavathas, 31.X.1973, 973247.</paragraph>
<subSubSection id="E3EBBD490E56303C2661801C99DD49C7" pageId="0" pageNumber="45" type="materials_examined">
<paragraph id="C6407FEBAEED90FDB126CEFB34579F45" pageId="0" pageNumber="45">Other material.</paragraph>
<paragraph id="2708E713CC67FF41BCCB436322B08C0F" pageId="0" pageNumber="45">Greece, Lesbos, leg. Ellis (deposited at the Naturalis Biodiversity Center, Netherland): 22 females, 6 males (21038-21054, 21056-21059, 21062-21064, 21066-21068), Antissa, 30.X.1973, 973240; 18 females, 23 males (21060, 21078-21117), Antissa, 30.X.1973, 973243; 1 male (21061), Antissa, 30.X.1973, 973244; 1 male (21037), Gavathas, 31.X.1973, 973247.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="DDF5C5ACD6B383910041460B49CFF7C2" pageId="0" pageNumber="45" type="etymology">
<paragraph id="7D4470122C43026CAF62FB13A22E7DBA" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="80CA5ED73ED10B85143F6117E188E44D" pageId="0" pageNumber="45">
Dedicated to Dr. Willem N. Ellis, an excellent specialist in
<taxonomicName lsidName="" order="Collembola" pageId="0" pageNumber="45" rank="order">Collembola</taxonomicName>
<taxonomicName id="90CF3B72B5363E3E62CEA4BD301D1BBE" lsidName="" order="Collembola" pageId="0" pageNumber="45" rank="order">Collembola</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="diagnosis">
<paragraph pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="45">Habitus typical of genus. Ant. IV with three lateral and one dorsal long thin and curved blunt sensilla (sometimes 1-2 more in dorsal group, longer and less curved). Postantennal organ equal to, or slightly larger than, nearest ocellus. Labrum with distinct apical papillae. Tibiotarsi with one clavate tenent hair. Empodial lamellae broad. Ventral tube with 4 + 4 setae. Retinaculum with 4 + 4 teeth. Dens with fine, uniform granulation and seven setae. Mucro with comparatively high outer lamella. Anal spines small, situated on low basal papillae.</paragraph>
<subSubSection id="99A8F24ED13C43ACD4682A21113C276F" pageId="0" pageNumber="45" type="diagnosis">
<paragraph id="34A5F58719F9388616349D93F660741B" pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph id="80F807958C0448A60ECD38559B84064A" pageId="0" pageNumber="45">Habitus typical of genus. Ant. IV with three lateral and one dorsal long thin and curved blunt sensilla (sometimes 1-2 more in dorsal group, longer and less curved). Postantennal organ equal to, or slightly larger than, nearest ocellus. Labrum with distinct apical papillae. Tibiotarsi with one clavate tenent hair. Empodial lamellae broad. Ventral tube with 4 + 4 setae. Retinaculum with 4 + 4 teeth. Dens with fine, uniform granulation and seven setae. Mucro with comparatively high outer lamella. Anal spines small, situated on low basal papillae.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">Body length 1.1-1.6 mm. Habitus typical of the genus. Color in alcohol pale brown dorsally and yellowish ventrally, eye-patches dark. Granulation fine and uniform, 12-20 granules between setae p1 on Abd. V.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="94149F263B5D28DA91CF7C3D27BA5B8B" pageId="0" pageNumber="45" type="description">
<paragraph id="B96DF8F656E44DA6F381F3F3BF574EBD" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="6C27163421BF302A20CD81A8F63E18EF" pageId="0" pageNumber="45">Body length 1.1-1.6 mm. Habitus typical of the genus. Color in alcohol pale brown dorsally and yellowish ventrally, eye-patches dark. Granulation fine and uniform, 12-20 granules between setae p1 on Abd. V.</paragraph>
<paragraph id="4C8F4CCA51D4D034D9810A41B5C681EF" pageId="0" pageNumber="45">
Chaetotaxy of head typical of the genus, with complete set of v-setae (Fig. 1). Setae slightly differentiated in length, especially on last abdominal segments, smooth and rather thick and stiff. Body sensilla (s) about 2-3 times longer than ordinary setae, fine and smooth. Dorsal chaetotaxy of Th.
<normalizedToken originalValue="IIII">I-III</normalizedToken>
<normalizedToken id="A7772D57F10273C44DC65FF5A0AAA898" originalValue="IIII">I-III</normalizedToken>
and Abd.
<normalizedToken originalValue="IIIVI">III-VI</normalizedToken>
<normalizedToken id="4F88ACD2608558580EA471711EC401D9" originalValue="IIIVI">III-VI</normalizedToken>
as in Figs 1-3. Th. I with 3 + 3 setae. Th. II with setae m2 absent, m3 present or absent and m4, m6 present. Th. III with setae m2 and m3 absent and setae m4 and m6 present. Abd. IV with setae p3 present, p7 absent and increased number of m-setae. On Abd. V setae p2 present and m-setae absent. Subcoxae I, II, III with 1, 3, 3 setae respectively. Microsensillum on Th. II present.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<paragraph id="AE6CCEA84057288C49F181F2A334958B" pageId="0" pageNumber="45">
Ant. IV with simple apical vesicle, subapical organite (or), microsensillum (ms), three lateral and one dorsal long thin and curved blunt sensilla (sometimes 1-2 more in dorsal group, longer and less curved, marked with an asterisk in Fig. 4) and 5-10 short pointed setae in ventral file (Fig. 5). Ant. III-organ with two long (outer) and two short (inner) sensilla (Fig. 4). Microsensillum on Ant. III present. Ant. I with seven setae (seta
<normalizedToken originalValue="p">p'</normalizedToken>
<normalizedToken id="FAEF07C4672666E24305E5521C99A95B" originalValue="p">p'</normalizedToken>
absent).
</paragraph>
<caption pageId="0" pageNumber="45">
<paragraph pageId="0" pageNumber="45">
<caption id="9D22CCFD996A8A7380E09F5A53C3728F" pageId="0" pageNumber="45">
<paragraph id="72B01C6077D715557EC6BA01DB46086D" pageId="0" pageNumber="45">
Figures 4-8.
<taxonomicName class="Collembola" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypogastrura ellisi" order="Poduromorpha" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ellisi">Hypogastrura ellisi</taxonomicName>
<taxonomicName id="5E787EF919D3BFAEDB63995A825CD89C" class="Collembola" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypogastrura ellisi" order="Poduromorpha" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ellisi">Hypogastrura ellisi</taxonomicName>
sp. n. 4 chaetotaxy of dorsal side of Ant. IV 5 chaetotaxy of ventral side of Ant. IV 6 postantennal organ and neighbor ocelli 7 claw III 8 dens and mucro. Abbreviations in text. Scale bars: 0.01 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
<paragraph id="9FE74A1B3D9406EFCF0A67FB7DA1BCE0" pageId="0" pageNumber="45">
Ocelli 8 + 8. Postantennal organ equal to, or slightly larger than, neighboring ocellus, with four subequal lobes. Accessory boss present (Fig. 6). Labrum with six distinct apical papillae (four large and two smaller, Figs 9-10). Labral setae 5, 5, 4, prelabrals 4. Maxillary head of the
<taxonomicName lsidName="H. tullbergi" pageId="0" pageNumber="45" rank="species" species="tullbergi">H. tullbergi</taxonomicName>
<taxonomicName id="D2AEB1E4189FFC2DEFEC6C7F6E91113A" lsidName="H. tullbergi" pageId="0" pageNumber="45" rank="species" species="tullbergi">H. tullbergi</taxonomicName>
type (
<bibRefCitation author="Fjellberg, A" journalOrPublisher="Annales de la Societe Royale Zoologique de Belgique" pageId="3" pageNumber="48" pagination="89 - 99" title="Maxillary structures in Hypogastruridae (Collembola)." volume="114" year="1984">Fjellberg 1984</bibRefCitation>
<bibRefCitation id="E2FF653D9327511BB7F926EA3ABC0E7D" author="Fjellberg, A" journalOrPublisher="Annales de la Societe Royale Zoologique de Belgique" pageId="3" pageNumber="48" pagination="89 - 99" title="Maxillary structures in Hypogastruridae (Collembola)." volume="114" year="1984">Fjellberg 1984</bibRefCitation>
) and labium as in
<bibRefCitation author="Fjellberg, A" journalOrPublisher="Zoologischer Anzeiger" pageId="3" pageNumber="48" pagination="309 - 330" title="The Labial Palp in Collembola." volume="237" year="1999">Fjellberg (1999</bibRefCitation>
<bibRefCitation id="5188066BF3806658B8DAE7D048C7D8E9" author="Fjellberg, A" journalOrPublisher="Zoologischer Anzeiger" pageId="3" pageNumber="48" pagination="309 - 330" title="The Labial Palp in Collembola." volume="237" year="1999">Fjellberg (1999</bibRefCitation>
: fig. 6). Outer lobe of maxilla with two sublobal hairs.
</paragraph>
<paragraph pageId="0" pageNumber="45">Tibiotarsi I, II, III with 19, 19, 18 setae respectively. Apical seta A1 long and clavate. Claws with small inner tooth. Empodial appendage with broad basal lamella and apical filament reaching slightly beyond inner tooth of unguis (Fig. 7).</paragraph>
<paragraph pageId="0" pageNumber="45">Ventral tube with four setae on each side. Retinaculum with 4 + 4 teeth.</paragraph>
<paragraph pageId="0" pageNumber="45">Furca well developed (ratio dens + mucro/inner edge of claws III 2.4-2.7). Dorsal side of dens with fine, uniform granulation and seven setae. Mucro with relatively high outer lamella. Ratio dens/mucro 2.0-2.3 (Fig. 8).</paragraph>
<paragraph pageId="0" pageNumber="45">Anal spines small, situated on low basal papillae (Fig. 3), ratio anal spine/basal papilla 0.7-1.1, ratio anal spine + basal papilla/inner edge of claws III 0.5-0.6.</paragraph>
<paragraph id="EB63BD1500A08662180397999D4E3560" pageId="0" pageNumber="45">Tibiotarsi I, II, III with 19, 19, 18 setae respectively. Apical seta A1 long and clavate. Claws with small inner tooth. Empodial appendage with broad basal lamella and apical filament reaching slightly beyond inner tooth of unguis (Fig. 7).</paragraph>
<paragraph id="8513F4B7EEA0078A38B42F7EC8353B09" pageId="0" pageNumber="45">Ventral tube with four setae on each side. Retinaculum with 4 + 4 teeth.</paragraph>
<paragraph id="8DF394ACD6DCDE7FDA9F5151DC8C8C9F" pageId="0" pageNumber="45">Furca well developed (ratio dens + mucro/inner edge of claws III 2.4-2.7). Dorsal side of dens with fine, uniform granulation and seven setae. Mucro with relatively high outer lamella. Ratio dens/mucro 2.0-2.3 (Fig. 8).</paragraph>
<paragraph id="2AF657C74FAE1BA5E16961F077B4AC3E" pageId="0" pageNumber="45">Anal spines small, situated on low basal papillae (Fig. 3), ratio anal spine/basal papilla 0.7-1.1, ratio anal spine + basal papilla/inner edge of claws III 0.5-0.6.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="remarks">
<paragraph pageId="0" pageNumber="45">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="BA356AD4A639AA9F4B86551E191923E9" pageId="0" pageNumber="45" type="remarks">
<paragraph id="9C57F70F674D490DD5676FDB177917C7" pageId="0" pageNumber="45">Remarks.</paragraph>
<paragraph id="60C77D98245FE97E05FE8629C0C91707" pageId="0" pageNumber="45">
It is difficult to find a right place for
<taxonomicName lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
<taxonomicName id="B1C028CEA18BF79AA198794671A54087" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n. within the genus. This species does not key to any of the groups in
<bibRefCitation author="Skarzynski, D" journalOrPublisher="Soil Organisms" pageId="3" pageNumber="48" pagination="77 - 83" title="Reassessment of the taxonomic position of Hypogastruramonticola Stach, 1946 (Collembola, Hypogastruridae)." volume="81" year="2009">
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
<bibRefCitation id="D4C8A0BD13F6ED7CAA476E1E4030B71F" author="Skarzynski, D" journalOrPublisher="Soil Organisms" pageId="3" pageNumber="48" pagination="77 - 83" title="Reassessment of the taxonomic position of Hypogastruramonticola Stach, 1946 (Collembola, Hypogastruridae)." volume="81" year="2009">
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(2009)
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. However, having weakly differentiated blunt Ant. IV sensilla, one tenent hair on the tibiotarsi, broad empodial lamellae, dens without tooth-like granules and ventro-apical swelling and a mucro without a distinct subapical tooth, it can be compared with some representatives of the
<taxonomicName lsidName="trybomi" pageId="0" pageNumber="45" rank="species" species="trybomi">trybomi</taxonomicName>
<taxonomicName id="D6C8D1303BE9550A8166A93E73BA2D0A" lsidName="trybomi" pageId="0" pageNumber="45" rank="species" species="trybomi">trybomi</taxonomicName>
or
<taxonomicName lsidName="monticola" pageId="0" pageNumber="45" rank="species" species="monticola">monticola</taxonomicName>
<taxonomicName id="9489F39115ECA9C25EC5DEE08994FCC7" lsidName="monticola" pageId="0" pageNumber="45" rank="species" species="monticola">monticola</taxonomicName>
groups as well as
<taxonomicName lsidName="H. aterrima" pageId="0" pageNumber="45" rank="species" species="aterrima">H. aterrima</taxonomicName>
<taxonomicName id="0449EAB89F460F5A5C131F83792BD444" lsidName="H. aterrima" pageId="0" pageNumber="45" rank="species" species="aterrima">H. aterrima</taxonomicName>
Yosii, 1972, which has an isolated position within the genus.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<paragraph id="A77AD1D8E5167A77B7D08A0A861305C7" pageId="0" pageNumber="45">
Undoubtedly,
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<taxonomicName id="3918FE8E090FC3BEA362D14872838D26" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n. is the most similar to
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="BC73F23D9F7F3A78B4A9AF180664BAAE" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
, considered as a member of the
<taxonomicName lsidName="trybomi" pageId="0" pageNumber="45" rank="species" species="trybomi">trybomi</taxonomicName>
<taxonomicName id="79569E9DC848EE9DBFC0CA4BE5CAAE05" lsidName="trybomi" pageId="0" pageNumber="45" rank="species" species="trybomi">trybomi</taxonomicName>
group. Most noticeably they differ in the shape of labral apical papillae (
<taxonomicName lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
<taxonomicName id="FE6FB604B1B1A01782F5E121D9410D8B" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n. - convex, strong, well visible, Figs 9-10;
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="53A2C100C7777A1E9914C082EBA33781" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
- flat, delicate, hardly visible, Figs 11-13). Apart from this
<taxonomicName lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
<taxonomicName id="B135299BE29A562617E051732C554FAF" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n. lacks setae m2 on Th. II (present in
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="A650C3DA42AF5B0D00EE8CC2F2191EB1" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
) and possesses smaller anal spines (the ratio of anal spine + basal papilla/inner edge of claws III 0.5-0.6 in
<taxonomicName lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
<taxonomicName id="112105048A540AFAB3CBCF9E45AFBBEC" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n. vs 0.75-1.1 in
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="A210C1480E020E7BBBF5AD2DA947FD83" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
), 5-10 short pointed setae in the ventral file on Ant. IV (Fig. 5) (
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="C1A16D357CE16CC623F56A73047163DB" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
- approx. ten short and stiff sensilla, truncate at apex, Fig. 14), and a mucro with a relatively high outer lamella (both inner and outer lamellae are low in
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="D80F7A10D28638042CBE9B90837467BC" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
).
</paragraph>
<caption pageId="0" pageNumber="45">
<paragraph pageId="0" pageNumber="45">
<caption id="3598A3EFC0A3F4B288369954792751FB" pageId="0" pageNumber="45">
<paragraph id="CBECE0F9A3AAE557BBE119720AAFBA3D" pageId="0" pageNumber="45">
Figures 9-14.
<taxonomicName class="Collembola" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypogastrura ellisi" order="Poduromorpha" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ellisi">Hypogastrura ellisi</taxonomicName>
<taxonomicName id="6EB4A86028950EDA2A096E9DACA2BABC" class="Collembola" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypogastrura ellisi" order="Poduromorpha" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ellisi">Hypogastrura ellisi</taxonomicName>
sp. n. 9 labrum 10 apical part of labrum,
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="D790E64B7FAF830FE78404E30A23DACA" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
: 11 labrum, specimen from Azerbaijan 12 apical part of labrum, specimen from Crete 13 apical papillae, specimen from Crete 14 chaetotaxy of ventral side of Ant. IV. Black arrows indicate apical papillae. Scale bars: 0.01 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
<paragraph id="3BEB0B3F817A76120B88EB39D64AEA54" pageId="0" pageNumber="45">
Both species have similar general ranges of distribution (
<taxonomicName lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
<taxonomicName id="7194E3086142E07EDEF7A6C7C807F9EE" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n.: Iran, Greece - Lesbos;
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="C448D39CF9E7D9432B7A1DE62D15F8C6" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
: Greece - Crete, Lesbos and Azerbaijan), but on a local scale they co-occur only in Lesbos. In
<normalizedToken originalValue="Elliss">Ellis's</normalizedToken>
<normalizedToken id="C7EAAAFD0810CF547C60005AF80DE104" originalValue="Elliss">Ellis's</normalizedToken>
material from this island, numerous
<taxonomicName lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
<taxonomicName id="BD5E10D931D7C8C785A16F9F56CF48AA" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n. from two localities and rare
<taxonomicName lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
<taxonomicName id="54E7390E0AB83AF007D19A12F0FF0EA4" lsidName="H. tethyca" pageId="0" pageNumber="45" rank="species" species="tethyca">H. tethyca</taxonomicName>
individuals from five sites were found. Nevertheless, they were isolated spatially. Unfortunately, due to incomplete collecting data, we do not know whether these populations differ in habitat preferences.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<paragraph id="84C8D7852AF043D5DCDFCC34AB707DCB" pageId="0" pageNumber="45">
The new species is easy to distinguish from the members of the
<taxonomicName lsidName="monticola" pageId="0" pageNumber="45" rank="species" species="monticola">monticola</taxonomicName>
<taxonomicName id="2EEB6E3D60B7338EB74B8B0C21E241CB" lsidName="monticola" pageId="0" pageNumber="45" rank="species" species="monticola">monticola</taxonomicName>
group by the absence of m-setae on Abd. V (vs present) and the size of the postantennal organ, which is equal to, or slightly larger than, the neighboring ocellus (vs 1.5-2 times larger than ocellus).
<taxonomicName lsidName="H. aterrima" pageId="0" pageNumber="45" rank="species" species="aterrima">H. aterrima</taxonomicName>
<taxonomicName id="12A37D41A2F8BBC5F04259FE4B5E7796" lsidName="H. aterrima" pageId="0" pageNumber="45" rank="species" species="aterrima">H. aterrima</taxonomicName>
can also be easily separated from
<taxonomicName lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
<taxonomicName id="717F9A9B11A6F557786B37F2865B54C2" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n. due to tridentate retinaculum (vs quadridentate), minute anal spines, slightly larger than surrounding granules (vs large, the ratio of anal spine + basal papilla/inner edge of claws III 0.5-0.6 in
<taxonomicName lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
<taxonomicName id="7A1570A8D66AB497FA5E34F535AB7DC5" lsidName="H. ellisi" pageId="0" pageNumber="45" rank="species" species="ellisi">H. ellisi</taxonomicName>
sp. n.), the presence of setae m2 on Th. II, and the absence of setae m6 on Th.
<normalizedToken originalValue="IIIII">II-III</normalizedToken>
<normalizedToken id="6961E332EAD3A86555A704C31BACF407" originalValue="IIIII">II-III</normalizedToken>
.
</paragraph>
</subSubSection>

View file

@ -1,294 +1,294 @@
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<paragraph pageId="0" pageNumber="105">
<taxonomicName LSID="http://zoobank.org/65D4AC75-488B-4A29-9773-5E0F5FB25449" authority="Sischo &amp; Hadfield, 2021" authorityName="Sischo &amp; Hadfield" authorityYear="2021" class="Gastropoda" family="Partulidae" genus="Partula" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Partula lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis" status="sp. nov.">Partula lutaensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="105">sp. nov.</taxonomicNameLabel>
<treatment id="6D583946B90057479D87A83059996E69" ID-GBIF-Taxon="182216098" LSID="urn:lsid:zoobank.org:act:65D4AC75-488B-4A29-9773-5E0F5FB25449" httpUri="http://treatment.plazi.org/id/6D583946B90057479D87A83059996E69" lastPageNumber="105" pageId="0" pageNumber="105" scope_class="Gastropoda" scope_family="Partulidae" scope_subClass="Heterobranchia">
<subSubSection id="B4F9739D03090F1BA30F9E1A61F102E4" pageId="0" pageNumber="105" type="nomenclature">
<paragraph id="040FAA007C5CD932D0C90E0514333A7A" pageId="0" pageNumber="105">
<taxonomicName id="DB23709C48962A9D978D7775021A62C7" ID-CoL="8H9DG" LSID="http://zoobank.org/65D4AC75-488B-4A29-9773-5E0F5FB25449" authority="Sischo &amp; Hadfield, 2021" authorityName="Sischo &amp; Hadfield" authorityYear="2021" class="Gastropoda" family="Partulidae" genus="Partula" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Partula lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis" status="sp. nov.">Partula lutaensis</taxonomicName>
<taxonomicNameLabel id="D53E3FF41289B56623AC3D7608B2B860" pageId="0" pageNumber="105">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="type material">
<paragraph pageId="0" pageNumber="105">Type material.</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Holotype</emphasis>
<subSubSection id="D0C6AB60E7C0EC3287BAC7A64FACBA3B" pageId="0" pageNumber="105" type="type material">
<paragraph id="2B97F74BC8BFA13A567BD5E753544DEA" pageId="0" pageNumber="105">Type material.</paragraph>
<paragraph id="8E8DCB93E1D3BCC76C0410A2BEC56233" pageId="0" pageNumber="105">
<emphasis id="5FBD7290652E5E3D526C52F6CD282D86" bold="true" pageId="0" pageNumber="105">
<emphasis id="7182529FBD20C62374BA77DDCE6ADF89" italics="true" pageId="0" pageNumber="105">Holotype</emphasis>
.
</emphasis>
Bishop Museum BPBM 284888 Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shell holotype of P. lutaensis sp. nov." figureDoi="10.3897/zookeys.1037.56303.figure3" httpUri="https://binary.pensoft.net/fig/543754" pageId="0" pageNumber="105">3</figureCitation>
<figureCitation id="BE23A29BA5495C647A790267666F9630" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shell holotype of P. lutaensis sp. nov." figureDoi="10.3897/zookeys.1037.56303.figure3" httpUri="https://binary.pensoft.net/fig/543754" pageId="0" pageNumber="105">3</figureCitation>
. Entire specimen collected by Jill Liske-Clark, 20/11/2014 from type locality.
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Paratypes</emphasis>
<emphasis id="69616D4BE15B21CB0878C97BC57B9B04" bold="true" pageId="0" pageNumber="105">
<emphasis id="5972CADABA5000E97F6497056EE32669" italics="true" pageId="0" pageNumber="105">Paratypes</emphasis>
.
</emphasis>
BPBM 284889, 2 specimens collected by Jill Liske-Clark, 20/11/2014, from type locality, and BPBM 284890, 3 specimens collected by Jill Liske-Clark, same date, from a second location on Rota Island, Commonwealth of the Northern Mariana Islands.
</paragraph>
<caption doi="10.3897/zookeys.1037.56303.figure3" httpUri="https://binary.pensoft.net/fig/543754" pageId="0" pageNumber="105" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 3.</emphasis>
<caption id="5E5BEC32F91D49DC92499E43D9669C91" doi="10.3897/zookeys.1037.56303.figure3" httpUri="https://binary.pensoft.net/fig/543754" pageId="0" pageNumber="105" start="Figure 3" startId="F3">
<paragraph id="995032D1E53EFFE1F528198101340802" pageId="0" pageNumber="105">
<emphasis id="FBAFECA4232C6414B0B4C9F658CE6ADC" bold="true" pageId="0" pageNumber="105">Figure 3.</emphasis>
Shell holotype of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
<taxonomicName id="E8B70395F834472A64BA99BAEDC73916" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="45B1EAD75BD7A7BB95AAEC1F0BD79A90" italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="type locality">
<paragraph pageId="0" pageNumber="105">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="105">Rota Island, Commonwealth of the Northern Mariana Islands (CNMI).</paragraph>
<subSubSection id="640DE8FDBFA8DCA412FE36C51ADEFCAC" pageId="0" pageNumber="105" type="type locality">
<paragraph id="FBD80658540B67C13A79E0C06B3E81A8" pageId="0" pageNumber="105">Type locality.</paragraph>
<paragraph id="A7F8533ACD58B07F61DA61AA0E8DA9DD" pageId="0" pageNumber="105">Rota Island, Commonwealth of the Northern Mariana Islands (CNMI).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="diagnosis">
<paragraph pageId="0" pageNumber="105">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Shell</emphasis>
<subSubSection id="FDC0E7C07F00D21929740942E17BAC5F" pageId="0" pageNumber="105" type="diagnosis">
<paragraph id="7028A89468CC10AEB2C09283FCFD5FAF" pageId="0" pageNumber="105">Diagnosis.</paragraph>
<paragraph id="92814200402168235FEEDE25D999E86B" pageId="0" pageNumber="105">
<emphasis id="DAD8229AD85A1CDB4153F6D107544B32" bold="true" pageId="0" pageNumber="105">
<emphasis id="61647D7B84D5B144F116BB86DCF23A47" italics="true" pageId="0" pageNumber="105">Shell</emphasis>
.
</emphasis>
Shell dextral, moderately thin, ovate-conic, slightly perforate; umbilicus open; whorls moderately convex, suture adpressed; aperture ovate-elongate, slightly oblique; outer lip reflexed, thick, glossy; parietal lip glossy with light or dark coloration; color of embryonic whorls and post-embryonic whorls variable from shades of brown, buff, white and yellow with prominent white subsutural band; Measurements (
<emphasis italics="true" pageId="0" pageNumber="105">N</emphasis>
<emphasis id="B289AF2A0191487379DBD3FE57A030E4" italics="true" pageId="0" pageNumber="105">N</emphasis>
= 48 specimens from five lots in BPBM collections from Rota): height (= length) 15.98 mm, s.d. 0.75 mm; width, 10.64 mm, s.d., 0.24 mm. See Figure
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left - Color morphs of Partula lutaensis sp. nov. found within a 10 x 10 - meter quadrat. Right - Closeup of a Partula lutaensis sp. nov. with a dark shell." figureDoi="10.3897/zookeys.1037.56303.figure4" httpUri="https://binary.pensoft.net/fig/543755" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="2940DB9869868B4A0955821AC2398F3D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left - Color morphs of Partula lutaensis sp. nov. found within a 10 x 10 - meter quadrat. Right - Closeup of a Partula lutaensis sp. nov. with a dark shell." figureDoi="10.3897/zookeys.1037.56303.figure4" httpUri="https://binary.pensoft.net/fig/543755" pageId="0" pageNumber="105">4</figureCitation>
for examples of shell color variation. Shell greatly resembles those of
<taxonomicName class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis italics="true" pageId="0" pageNumber="105">Partula gibba</emphasis>
<taxonomicName id="B669761CBD5D780383212EE8D2427E65" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis id="6FFE3EF4C41F9AD99BF47AA3A1AAF3A7" italics="true" pageId="0" pageNumber="105">Partula gibba</emphasis>
</taxonomicName>
on Guam and Saipan (
<bibRefCitation author="Crampton, HE" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="105" refId="B8" refString="Crampton, HE, 1925. Studies on the variation, distribution, and evolution of the genus Partula. The species of the Mariana Islands, Guam and Saipan. Carnegie Institution of Washington Publication 228a: 1-116." title="Studies on the variation, distribution, and evolution of the genus Partula. The species of the Mariana Islands, Guam and Saipan. Carnegie Institution of Washington Publication 228 a: 1 - 116." year="1925">Crampton 1925</bibRefCitation>
<bibRefCitation id="96FEE9CCCE8935C522979AAC858949F2" author="Crampton, HE" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="105" refId="B8" refString="Crampton, HE, 1925. Studies on the variation, distribution, and evolution of the genus Partula. The species of the Mariana Islands, Guam and Saipan. Carnegie Institution of Washington Publication 228a: 1-116." title="Studies on the variation, distribution, and evolution of the genus Partula. The species of the Mariana Islands, Guam and Saipan. Carnegie Institution of Washington Publication 228 a: 1 - 116." year="1925">Crampton 1925</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1037.56303.figure4" httpUri="https://binary.pensoft.net/fig/543755" pageId="0" pageNumber="105" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 4.</emphasis>
<caption id="58923E389B57A54EF04D1F79FED30678" doi="10.3897/zookeys.1037.56303.figure4" httpUri="https://binary.pensoft.net/fig/543755" pageId="0" pageNumber="105" start="Figure 4" startId="F4">
<paragraph id="3DAEE08EC51F7B33366FB567CE85088F" pageId="0" pageNumber="105">
<emphasis id="A661326B7E61C17DC69CD46D18C8F36C" bold="true" pageId="0" pageNumber="105">Figure 4.</emphasis>
Left - Color morphs of
<taxonomicName authorityName="Sischo &amp; Hadfield" authorityYear="2021" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">Partula lutaensis</emphasis>
<taxonomicName id="89DE0E6C42B1A2546569E5C17CDE6C02" authorityName="Sischo &amp; Hadfield" authorityYear="2021" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="60BEE74D3F78B18ADDE29B89D85C1445" italics="true" pageId="0" pageNumber="105">Partula lutaensis</emphasis>
</taxonomicName>
sp. nov. found within a 10
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="3699C5FE810D7646BD8999E60E52E5BD" originalValue="×">x</normalizedToken>
10-meter quadrat. Right - Closeup of a
<taxonomicName authorityName="Sischo &amp; Hadfield" authorityYear="2021" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">Partula lutaensis</emphasis>
<taxonomicName id="940282270A92BACDF9BA0238667B4C42" authorityName="Sischo &amp; Hadfield" authorityYear="2021" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="5BC4B939BA65D8B4FDBA1871C6C02BE8" italics="true" pageId="0" pageNumber="105">Partula lutaensis</emphasis>
</taxonomicName>
sp. nov. with a dark shell.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="distinguishing shells of p. lutaensis from those of p. langfordi">
<paragraph pageId="0" pageNumber="105">
<subSubSection id="C18C96CE09074F6249B34FC3174345D2" pageId="0" pageNumber="105" type="distinguishing shells of p. lutaensis from those of p. langfordi">
<paragraph id="81173A2DDC0FF022B9028C5E5B93DEF0" pageId="0" pageNumber="105">
Distinguishing shells of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
<taxonomicName id="C8EA18428375A8F3DFA49836B2D58112" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="56592CD0ABF31051E8636BA1CE6C34FC" italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
</taxonomicName>
from those of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
<taxonomicName id="7494B6A763159AEFA5A8F9B627C7F2C0" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis id="A66A66146A9BD7197784CB4CCE199B57" italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<paragraph id="882BDCFE61AD17D2D990F522C7AA9978" pageId="0" pageNumber="105">
Shell length of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
<taxonomicName id="4A38427AE53A9427BAE5064C226352DE" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis id="1910E3E22D7B8CAA4B5BDB392C2E9D9D" italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="105">M</emphasis>
<emphasis id="9772C600F32B3B07399F586D5ECE4D78" italics="true" pageId="0" pageNumber="105">M</emphasis>
= 13.83 mm, s.d. = 0.37 mm,
<emphasis italics="true" pageId="0" pageNumber="105">N</emphasis>
<emphasis id="DF14E1ED8CD0A224F48FA4A5CC9BB663" italics="true" pageId="0" pageNumber="105">N</emphasis>
= 47) was significantly shorter than the length of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
<taxonomicName id="F699E9A6B57035363637F6F3B6640283" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="442A8F3513DE9AFE01C7DD3E0A6F1882" italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="105">M</emphasis>
<emphasis id="F06ACCCB58F9E12D35735A73078F5DA0" italics="true" pageId="0" pageNumber="105">M</emphasis>
= 15.98 mm, s.d. = 0.75 mm,
<emphasis italics="true" pageId="0" pageNumber="105">N</emphasis>
<emphasis id="EC35EFB03F1D33923BD6CDE99522C3F0" italics="true" pageId="0" pageNumber="105">N</emphasis>
= 48), t(82) = -13.91, P &lt;0.001. Similarly, shell width of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
<taxonomicName id="E7B346E4069350CD3FCCA4A8F181A59F" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis id="9223654BBB9D75F54CE6F21C531B7471" italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="105">M</emphasis>
<emphasis id="36C81FCA24B0F14024C5424AF423E13A" italics="true" pageId="0" pageNumber="105">M</emphasis>
= 9.74 mm, s.d = 0.36 mm) was significantly less than that of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
<taxonomicName id="65DB416122358D4AF3D5E81E69737F71" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="758382115251643D264F8E58CDD2A4B1" italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
</taxonomicName>
sp. nov. (
<emphasis italics="true" pageId="0" pageNumber="105">M</emphasis>
<emphasis id="6CA9E033161AD89D09D2B93B4C00CCBC" italics="true" pageId="0" pageNumber="105">M</emphasis>
= 10.64 mm, s.d = 0.24 mm), t(90) = -8.02, P &lt;0.001 (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scattergram of Bishop Museum shell measurements comparing shell width (y-axis) by shell length (x-axis) of 48 P. langfordi from Aguigan and 47 P. lutaensis sp. nov. from Rota. The mean shell length and width plus standard deviation of each species are encircled. All shells were lipped indicating snails were mature and had reached terminal growth." figureDoi="10.3897/zookeys.1037.56303.figure5" httpUri="https://binary.pensoft.net/fig/543756" pageId="0" pageNumber="105">5</figureCitation>
<figureCitation id="7C9B9FC50B855877E5A37FDF84E309B2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scattergram of Bishop Museum shell measurements comparing shell width (y-axis) by shell length (x-axis) of 48 P. langfordi from Aguigan and 47 P. lutaensis sp. nov. from Rota. The mean shell length and width plus standard deviation of each species are encircled. All shells were lipped indicating snails were mature and had reached terminal growth." figureDoi="10.3897/zookeys.1037.56303.figure5" httpUri="https://binary.pensoft.net/fig/543756" pageId="0" pageNumber="105">5</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1037.56303.figure5" httpUri="https://binary.pensoft.net/fig/543756" pageId="0" pageNumber="105" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 5.</emphasis>
<caption id="5DCF449273086EA0D5A8756D5DB7EFEF" doi="10.3897/zookeys.1037.56303.figure5" httpUri="https://binary.pensoft.net/fig/543756" pageId="0" pageNumber="105" start="Figure 5" startId="F5">
<paragraph id="394F315DDF8569750D1FA7D98FA6DED6" pageId="0" pageNumber="105">
<emphasis id="F5A0DF188F49474C1028F53DB54997F9" bold="true" pageId="0" pageNumber="105">Figure 5.</emphasis>
Scattergram of Bishop Museum shell measurements comparing shell width (y-axis) by shell length (x-axis) of 48
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
<taxonomicName id="BB977AE3FF2A5E176FB9A2F15B3E33FE" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis id="927BC5801317DBA7BCD0F898F04B44F5" italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
</taxonomicName>
from Aguigan and 47
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
<taxonomicName id="A040F2722F78AFEEED8B8B42B63D1291" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="92381B0BB4D40FFAACC8395C8DCF0DE2" italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
</taxonomicName>
sp. nov. from Rota. The mean shell length and width plus standard deviation of each species are encircled. All shells were lipped indicating snails were mature and had reached terminal growth.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="male reproductive system">
<paragraph pageId="0" pageNumber="105">Male reproductive system.</paragraph>
<paragraph pageId="0" pageNumber="105">
<subSubSection id="B1C6B9ACE7280128D2965C3B9BB491A1" pageId="0" pageNumber="105" type="male reproductive system">
<paragraph id="1D8F11F121182D83AD094C432FD05B5B" pageId="0" pageNumber="105">Male reproductive system.</paragraph>
<paragraph id="334B888E4DE9A39B90AFA65144F688B0" pageId="0" pageNumber="105">
The male reproductive system of
<taxonomicName class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis italics="true" pageId="0" pageNumber="105">Partula gibba</emphasis>
<taxonomicName id="83E412A103E701C38D905E0C09644BAE" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis id="6E62CB37F5272B22D30EAE701FA3DC55" italics="true" pageId="0" pageNumber="105">Partula gibba</emphasis>
</taxonomicName>
figured by
<bibRefCitation author="Kondo, Y" journalOrPublisher="Pacific Science" pageId="0" pageNumber="105" refId="B15" refString="Kondo, Y, 1955. A revision of the family Partulidae (GastropodaPulmonata). PhD Thesis, Cambridge, Harvard University, Massachusetts." title="A revision of the family Partulidae (GastropodaPulmonata). PhD Thesis, Cambridge, Harvard University, Massachusetts." year="1955">Kondo (1955</bibRefCitation>
<bibRefCitation id="A540756C1366838DFF5BDFE988B2A0BE" author="Kondo, Y" journalOrPublisher="Pacific Science" pageId="0" pageNumber="105" refId="B15" refString="Kondo, Y, 1955. A revision of the family Partulidae (GastropodaPulmonata). PhD Thesis, Cambridge, Harvard University, Massachusetts." title="A revision of the family Partulidae (GastropodaPulmonata). PhD Thesis, Cambridge, Harvard University, Massachusetts." year="1955">Kondo (1955</bibRefCitation>
,
<bibRefCitation author="Kondo, Y" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="105" pagination="73 - 90" publicationUrl="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" refId="B17" refString="Kondo, Y, 1970. Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata). Bishop Museum Occasional Papers 24 (5): 73 - 90, http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" title="Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata)." url="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" volume="24" year="1970">1970</bibRefCitation>
<bibRefCitation id="3AA7734CCA2E846719097B377570F8B7" author="Kondo, Y" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="105" pagination="73 - 90" publicationUrl="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" refId="B17" refString="Kondo, Y, 1970. Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata). Bishop Museum Occasional Papers 24 (5): 73 - 90, http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" title="Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata)." url="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" volume="24" year="1970">1970</bibRefCitation>
) and
<bibRefCitation author="Gerlach, J" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="105" refId="B9" refString="Gerlach, J, 2016. Icons of evolution: Pacific Island tree-snails, family Partulidae. Cambridge Phelsuma Press, 336 pp." title="Icons of evolution: Pacific Island tree-snails, family Partulidae. Cambridge Phelsuma Press, 336 pp." year="2016">Gerlach (2016)</bibRefCitation>
<bibRefCitation id="E1386036EDB1C3DBACE4D5F62AA5B250" author="Gerlach, J" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="105" refId="B9" refString="Gerlach, J, 2016. Icons of evolution: Pacific Island tree-snails, family Partulidae. Cambridge Phelsuma Press, 336 pp." title="Icons of evolution: Pacific Island tree-snails, family Partulidae. Cambridge Phelsuma Press, 336 pp." year="2016">Gerlach (2016)</bibRefCitation>
is highly variable. In specimens we examined (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Outline drawings comparing the male reproductive tracks of four extant Partula species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of Partula langfordi is adapted from figure 5 of Kondo (1970). Note, Kondo found no difference between the male reproductive tracts of P. gibba and P. langfordi." figureDoi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105">6A</figureCitation>
<figureCitation id="BFC0F2D68FB909D163F4467DB33C99FC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Outline drawings comparing the male reproductive tracks of four extant Partula species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of Partula langfordi is adapted from figure 5 of Kondo (1970). Note, Kondo found no difference between the male reproductive tracts of P. gibba and P. langfordi." figureDoi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105">6A</figureCitation>
), the vas deferens entered the penis very near its top, leaving the upper portion of the penis, attached to the retractor muscle and called the caecum by
<bibRefCitation author="Gerlach, J" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="105" refId="B9" refString="Gerlach, J, 2016. Icons of evolution: Pacific Island tree-snails, family Partulidae. Cambridge Phelsuma Press, 336 pp." title="Icons of evolution: Pacific Island tree-snails, family Partulidae. Cambridge Phelsuma Press, 336 pp." year="2016">Gerlach (2016)</bibRefCitation>
<bibRefCitation id="B90C0D1EEB0763013E9868E9092A7514" author="Gerlach, J" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="105" refId="B9" refString="Gerlach, J, 2016. Icons of evolution: Pacific Island tree-snails, family Partulidae. Cambridge Phelsuma Press, 336 pp." title="Icons of evolution: Pacific Island tree-snails, family Partulidae. Cambridge Phelsuma Press, 336 pp." year="2016">Gerlach (2016)</bibRefCitation>
, to be very short. In specimens of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. radiolata" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="radiolata">
<emphasis italics="true" pageId="0" pageNumber="105">P. radiolata</emphasis>
<taxonomicName id="53910DE69A222FF1571356847749B6E7" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. radiolata" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="radiolata">
<emphasis id="9B14E658F94219FDE707B9805490FE73" italics="true" pageId="0" pageNumber="105">P. radiolata</emphasis>
</taxonomicName>
, the entry of the vas deferens was about 1/5 to 1/4 of the length of the penis below the retractor-muscle attachment (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Outline drawings comparing the male reproductive tracks of four extant Partula species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of Partula langfordi is adapted from figure 5 of Kondo (1970). Note, Kondo found no difference between the male reproductive tracts of P. gibba and P. langfordi." figureDoi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105">6B</figureCitation>
<figureCitation id="21ACF1EA047614F13BF21ED2C1A8C880" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Outline drawings comparing the male reproductive tracks of four extant Partula species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of Partula langfordi is adapted from figure 5 of Kondo (1970). Note, Kondo found no difference between the male reproductive tracts of P. gibba and P. langfordi." figureDoi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105">6B</figureCitation>
). However, the male duct was distinctive in
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
<taxonomicName id="8D7C4A8979D8784DAA737C9341A81710" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="2F30172F8E1A4A5EF574DE8BCCC839C2" italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
</taxonomicName>
sp. nov. by the bulge or shoulder at the top of an expanded caecum, proximal to the vas deferens. In the male system of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
<taxonomicName id="D2336493480A545F68BB1FDF695D65D7" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="D3BCA29094372BC9681377AD61E3C61D" italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
</taxonomicName>
sp. nov., the attachment of the vas deferens was consistently more distal than in either
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis italics="true" pageId="0" pageNumber="105">P. gibba</emphasis>
<taxonomicName id="54CAD640F7EF5004B12AC1BA3EBE2752" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis id="ACC558E5324783E7BC572A81187180C9" italics="true" pageId="0" pageNumber="105">P. gibba</emphasis>
</taxonomicName>
or
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. radiolata" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="radiolata">
<emphasis italics="true" pageId="0" pageNumber="105">P. radiolata</emphasis>
<taxonomicName id="CA6B9D0D0F87658B800D70B96B433723" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. radiolata" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="radiolata">
<emphasis id="89A075D3E26978F6EFF46B9BE6A44548" italics="true" pageId="0" pageNumber="105">P. radiolata</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Outline drawings comparing the male reproductive tracks of four extant Partula species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of Partula langfordi is adapted from figure 5 of Kondo (1970). Note, Kondo found no difference between the male reproductive tracts of P. gibba and P. langfordi." figureDoi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105">6C</figureCitation>
<figureCitation id="4634552C8533CC3314DED598094857EA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Outline drawings comparing the male reproductive tracks of four extant Partula species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of Partula langfordi is adapted from figure 5 of Kondo (1970). Note, Kondo found no difference between the male reproductive tracts of P. gibba and P. langfordi." figureDoi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105">6C</figureCitation>
); the insertion was close to 1/3 of the length of the penis below the retractor-muscle attachment. In no other regard were there any distinctive differences among the penial structures of these three species. The lower attachment of the vas deferens to the retractor muscle in both
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
<taxonomicName id="6EAE9E51D5CB33C3A500131A1BD8A188" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="6AB020886FB7512C8D125700D9F7F884" italics="true" pageId="0" pageNumber="105">P. lutaensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. radiolata" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="radiolata">
<emphasis italics="true" pageId="0" pageNumber="105">P. radiolata</emphasis>
<taxonomicName id="69C3172925546F1C2D3C75A621C05C31" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. radiolata" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="radiolata">
<emphasis id="11B6DFFD01AC6FC23C80589398F926A4" italics="true" pageId="0" pageNumber="105">P. radiolata</emphasis>
</taxonomicName>
is concordant with their placement as sister taxa in phylogenetic reconstructions of the group.
<bibRefCitation author="Kondo, Y" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="105" pagination="73 - 90" publicationUrl="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" refId="B17" refString="Kondo, Y, 1970. Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata). Bishop Museum Occasional Papers 24 (5): 73 - 90, http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" title="Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata)." url="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" volume="24" year="1970">Kondo (1970)</bibRefCitation>
<bibRefCitation id="D32B3D1EBD3C828411619E99A402D9FE" author="Kondo, Y" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="105" pagination="73 - 90" publicationUrl="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" refId="B17" refString="Kondo, Y, 1970. Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata). Bishop Museum Occasional Papers 24 (5): 73 - 90, http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" title="Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata)." url="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" volume="24" year="1970">Kondo (1970)</bibRefCitation>
found no differences between the male reproductive tract of
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
<taxonomicName id="38A817BBCD0AEF6BEEC184DBC55FC9E1" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis id="0D9D0C49940C3966A0E5AF2EC803F6FF" italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
</taxonomicName>
and
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis italics="true" pageId="0" pageNumber="105">P. gibba</emphasis>
<taxonomicName id="32747DF18FC742B3837AE139489D449B" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis id="880C5783EDDD24D61F4EF729BF41D5C6" italics="true" pageId="0" pageNumber="105">P. gibba</emphasis>
</taxonomicName>
. We include a modified version of his drawing with ours for comparison (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Outline drawings comparing the male reproductive tracks of four extant Partula species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of Partula langfordi is adapted from figure 5 of Kondo (1970). Note, Kondo found no difference between the male reproductive tracts of P. gibba and P. langfordi." figureDoi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105">6D</figureCitation>
<figureCitation id="DD824217F01D40AB962A9123A041A276" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Outline drawings comparing the male reproductive tracks of four extant Partula species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of Partula langfordi is adapted from figure 5 of Kondo (1970). Note, Kondo found no difference between the male reproductive tracts of P. gibba and P. langfordi." figureDoi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105">6D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 6.</emphasis>
<caption id="1B0995BB91F74D3131B09145E5BF2A11" doi="10.3897/zookeys.1037.56303.figure6" httpUri="https://binary.pensoft.net/fig/543757" pageId="0" pageNumber="105" start="Figure 6" startId="F6">
<paragraph id="C5574BC2AA53A4968AE1F39DD98D68D5" pageId="0" pageNumber="105">
<emphasis id="2922E790B5DAFB9CD7110D4F66F5BC34" bold="true" pageId="0" pageNumber="105">Figure 6.</emphasis>
Outline drawings comparing the male reproductive tracks of four extant
<taxonomicName authorityName="Ferussac" authorityYear="1821" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Partula</emphasis>
<taxonomicName id="102320F43A0654165076B4CD7D2ADBF0" authorityName="Ferussac" authorityYear="1821" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="genus">
<emphasis id="BF68719A6532691F6AFF5E45DFAE62BB" italics="true" pageId="0" pageNumber="105">Partula</emphasis>
</taxonomicName>
species from the Mariana Islands. Male anatomy abbreviations are as follows: retractor muscle (RM), caecum (C), penis (P), vas deferens (VD), uterus (U). The figure of
<taxonomicName authorityName="Y.Kondo" authorityYear="1970" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis italics="true" pageId="0" pageNumber="105">Partula langfordi</emphasis>
<taxonomicName id="F37091E5B9341C6DC596FBE5CADB6296" authorityName="Y.Kondo" authorityYear="1970" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis id="442C49B442CC1CD39CB74ADCAEDB16B0" italics="true" pageId="0" pageNumber="105">Partula langfordi</emphasis>
</taxonomicName>
is adapted from figure 5 of
<bibRefCitation author="Kondo, Y" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="105" pagination="73 - 90" publicationUrl="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" refId="B17" refString="Kondo, Y, 1970. Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata). Bishop Museum Occasional Papers 24 (5): 73 - 90, http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" title="Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata)." url="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" volume="24" year="1970">Kondo (1970)</bibRefCitation>
<bibRefCitation id="1F2D194FE1D53A1EC79C81077F0221AC" author="Kondo, Y" journalOrPublisher="Bishop Museum Occasional Papers" pageId="0" pageNumber="105" pagination="73 - 90" publicationUrl="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" refId="B17" refString="Kondo, Y, 1970. Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata). Bishop Museum Occasional Papers 24 (5): 73 - 90, http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" title="Some aspects of Mariana Islands Partulidae (Mollusca, Pulmonata)." url="http://hbs.bishopmuseum.org/pubs-online/pdf/op24-5.pdf" volume="24" year="1970">Kondo (1970)</bibRefCitation>
. Note, Kondo found no difference between the male reproductive tracts of
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<emphasis italics="true" pageId="0" pageNumber="105">P. gibba</emphasis>
<taxonomicName id="5425A11771F7823A45104193958DFF73" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. gibba" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="gibba">
<emphasis id="CBE43BFC879054C1FB70EF090D8473F3" italics="true" pageId="0" pageNumber="105">P. gibba</emphasis>
</taxonomicName>
and
<taxonomicName class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
<taxonomicName id="991D7794D13AABD0C40D3F00F691B550" class="Gastropoda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. langfordi" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="langfordi">
<emphasis id="30FC99F9A1A5E7359C345CF440940349" italics="true" pageId="0" pageNumber="105">P. langfordi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="ecology">
<paragraph pageId="0" pageNumber="105">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="105">
<subSubSection id="BB2D47F50E6EE7D19F24D0B101E28131" pageId="0" pageNumber="105" type="ecology">
<paragraph id="A1A477DD1E742660B38CFB77B5BD55AD" pageId="0" pageNumber="105">Ecology.</paragraph>
<paragraph id="A4E566131DCF8B6DDF3723E1B69C1E25" pageId="0" pageNumber="105">
Type and paratype specimens were found on
<taxonomicName genus="Epiprenmum" lsidName="Epiprenmum aureum" pageId="0" pageNumber="105" rank="species" species="aureum">
<emphasis italics="true" pageId="0" pageNumber="105">Epiprenmum aureum</emphasis>
<taxonomicName id="5E7DF1F4831B927931918D9B14F3A0FD" genus="Epiprenmum" lsidName="Epiprenmum aureum" pageId="0" pageNumber="105" rank="species" species="aureum">
<emphasis id="E898597AD8EDEAC4170C6EBA4684C3AB" italics="true" pageId="0" pageNumber="105">Epiprenmum aureum</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Cav" authorityYear="1802" class="Gastropoda" family="Littorinidae" genus="Tectaria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tectaria crenata" order="Littorinimorpha" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="crenata">
<emphasis italics="true" pageId="0" pageNumber="105">Tectaria crenata</emphasis>
<taxonomicName id="26D22ECCD4BC38F4A645EEEA0CC0EC59" authorityName="Cav" authorityYear="1802" class="Gastropoda" family="Littorinidae" genus="Tectaria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tectaria crenata" order="Littorinimorpha" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="crenata">
<emphasis id="D6B1846B550370D2F2B839DDCFF8E2EA" italics="true" pageId="0" pageNumber="105">Tectaria crenata</emphasis>
</taxonomicName>
(J. Liske-Clark, Northern Mariana Department of Fish and Wildlife, personal communication).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="etymology">
<paragraph pageId="0" pageNumber="105">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="105">
<subSubSection id="933DEB4CDE9CD1DC1D3C62E850DC2840" pageId="0" pageNumber="105" type="etymology">
<paragraph id="85481811E2DCB5DB06005438FE772BDE" pageId="0" pageNumber="105">Etymology.</paragraph>
<paragraph id="BEDD8698DD5E30D3519EFAB55BBE7722" pageId="0" pageNumber="105">
The specific epithet
<taxonomicName authorityName="Sischo &amp; Hadfield" authorityYear="2021" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis italics="true" pageId="0" pageNumber="105">Partula lutaensis</emphasis>
<taxonomicName id="3DE5D016A216EB047B1F3ABDFDEEC5B0" authorityName="Sischo &amp; Hadfield" authorityYear="2021" class="Gastropoda" family="Partulidae" genus="Partula" kingdom="Animalia" lsidName="Partula lutaensis" order="Stylommatophora" pageId="0" pageNumber="105" phylum="Mollusca" rank="species" species="lutaensis">
<emphasis id="14E18B71CAF040CF76A88A084A175B31" italics="true" pageId="0" pageNumber="105">Partula lutaensis</emphasis>
</taxonomicName>
recognizes Luta, the indigenous Chamorro name for the island of Rota.
</paragraph>

View file

@ -1,118 +1,118 @@
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<treatment LSID="urn:lsid:plazi:treatment:6D5FA0B33A72574184E4D5AFAEB28B9E" httpUri="http://treatment.plazi.org/id/6D5FA0B33A72574184E4D5AFAEB28B9E" lastPageNumber="125" pageId="0" pageNumber="125">
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<treatment id="6D5FA0B33A72574184E4D5AFAEB28B9E" ID-GBIF-Taxon="197841589" LSID="urn:lsid:plazi:treatment:6D5FA0B33A72574184E4D5AFAEB28B9E" httpUri="http://treatment.plazi.org/id/6D5FA0B33A72574184E4D5AFAEB28B9E" lastPageNumber="125" pageId="0" pageNumber="125" scope_family="Heteropterygidae" scope_order="Phasmida" scope_subFamily="Dataminae">
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<paragraph id="961457D754CADA00CF5EE6402CE022C5" pageId="0" pageNumber="125">
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Pylaemenes
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, 1875
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<paragraph pageId="0" pageNumber="125">
<emphasis italics="true" pageId="0" pageNumber="125">Type species.</emphasis>
<subSubSection id="D0E66FE2022AF25E5ABF5DAD9355D0DC" pageId="0" pageNumber="125" type="type species">
<paragraph id="2013357FB7C4B05E0E8D3F3781143B3C" pageId="0" pageNumber="125">
<emphasis id="77093610E58547B025B513C8F569E9AD" italics="true" pageId="0" pageNumber="125">Type species.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="125">
<taxonomicName authorityName="Haan &amp; W. de" authorityYear="1842" class="Insecta" family="Phasmatidae" genus="Phasma" kingdom="Animalia" lsidName="Phasma coronatum" order="Phasmida" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="coronatum">
<emphasis italics="true" pageId="0" pageNumber="125">Phasma coronatum</emphasis>
<paragraph id="EE1EA6AAAEC0DDD3568B9ADC07D5F1EE" pageId="0" pageNumber="125">
<taxonomicName id="6A2FBD2DC0606F33AC728D0C155DC165" authorityName="Haan &amp; W. de" authorityYear="1842" class="Insecta" family="Phasmatidae" genus="Phasma" kingdom="Animalia" lsidName="Phasma coronatum" order="Phasmida" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="coronatum">
<emphasis id="18371C11EED63DF17D1F92E0EF03481E" italics="true" pageId="0" pageNumber="125">Phasma coronatum</emphasis>
</taxonomicName>
de Haan, 1842 (=
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<emphasis italics="true" pageId="0" pageNumber="125">Pylaemenes coronatus</emphasis>
<taxonomicName id="2CF76A1AD923B7BEF1640B6F2FF6B75D" baseAuthorityName="Haan &amp; W. de" baseAuthorityYear="1842" class="Insecta" family="Heteropterygidae" genus="Pylaemenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pylaemenes coronatus" order="Phasmatodea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="coronatus">
<emphasis id="A7A1A35261ADC3290BFF1D6EEFBB9B60" italics="true" pageId="0" pageNumber="125">Pylaemenes coronatus</emphasis>
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), by subsequent designation; authority:
<bibRefCitation author="Kirby, WF" journalOrPublisher="British Museum, London" pageId="0" pageNumber="125" refId="B11" refString="Kirby, WF, 1904. A synonymic catalogue of Orthoptera 1. OrthopteraEuplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). British Museum, London" title="A synonymic catalogue of Orthoptera 1. OrthopteraEuplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae)." year="1904">Kirby (1904)</bibRefCitation>
<bibRefCitation id="291A68A2232E33A9DF1620B5C8549823" author="Kirby, WF" journalOrPublisher="British Museum, London" pageId="0" pageNumber="125" refId="B11" refString="Kirby, WF, 1904. A synonymic catalogue of Orthoptera 1. OrthopteraEuplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). British Museum, London" title="A synonymic catalogue of Orthoptera 1. OrthopteraEuplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae)." year="1904">Kirby (1904)</bibRefCitation>
.
</paragraph>
</subSubSection>
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<paragraph pageId="0" pageNumber="125">
<emphasis italics="true" pageId="0" pageNumber="125">Distribution.</emphasis>
<subSubSection id="E9EA6D0FD93D5FDEFF3C14B71D690C6C" pageId="0" pageNumber="125" type="distribution">
<paragraph id="01894084089617FC428670ABA996EDE3" pageId="0" pageNumber="125">
<emphasis id="67CDAC7D925A51C3F9D45DA380F4DC88" italics="true" pageId="0" pageNumber="125">Distribution.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="125">China (Yunnan); Indonesia; Malaysia; Singapore; Thailand; Vietnam.</paragraph>
<paragraph id="319D880CEFEF12CA35137E7A16398BB9" pageId="0" pageNumber="125">China (Yunnan); Indonesia; Malaysia; Singapore; Thailand; Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="125" type="species included from china.">
<paragraph pageId="0" pageNumber="125">
<emphasis italics="true" pageId="0" pageNumber="125">Species included from China.</emphasis>
<subSubSection id="269A2EE478B969D828629A8E39647A7E" pageId="0" pageNumber="125" type="species included from china.">
<paragraph id="499AC95845BE15D5B94A5F786939EA01" pageId="0" pageNumber="125">
<emphasis id="2A101DB0DB31485A0A50BFD4ECBF16E8" italics="true" pageId="0" pageNumber="125">Species included from China.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="125">
<taxonomicName class="Insecta" family="Heteropterygidae" genus="Pylaemenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pylaemenes pui" order="Phasmatodea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="pui">
<emphasis italics="true" pageId="0" pageNumber="125">Pylaemenes pui</emphasis>
<paragraph id="B6C7789D0C09E928A15F5E7CF9FBE574" pageId="0" pageNumber="125">
<taxonomicName id="EDB7267921B92A5DF918EBC0C6C2FF47" class="Insecta" family="Heteropterygidae" genus="Pylaemenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pylaemenes pui" order="Phasmatodea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="pui">
<emphasis id="348C77BE20295F1C13D246CAC7858390" italics="true" pageId="0" pageNumber="125">Pylaemenes pui</emphasis>
</taxonomicName>
Ho, 2013 [China:
<normalizedToken originalValue="Puer">Pu'er</normalizedToken>
<normalizedToken id="C826DA261210F04DFA24E2256E5B83B5" originalValue="Puer">Pu'er</normalizedToken>
and Xishuangbanna] (Fig.
<figureCitation captionStart="Fig. 3" captionStartId="F3" captionText="Fig. 3. Living photos. A, B. Pylaemenes gulinqingensis sp. nov. (Gulinqing, Yunnan, China); C. Hainanphasma cristata Ho, 2013 (Mt. jianfengling, Hainan Island, China); D. P. pui Ho, 2013 (Mengla, Yunnan, China). Photo credit A-C. Pei-Lin Liao; D. Chao Wu." figureDoi="10.3897/jor.31.79783.fig3" httpUri="https://binary.pensoft.net/fig/718486" pageId="0" pageNumber="125">3D</figureCitation>
<figureCitation id="921BBB46795FD8130A1C114A9D316C1D" captionStart="Fig. 3" captionStartId="F3" captionText="Fig. 3. Living photos. A, B. Pylaemenes gulinqingensis sp. nov. (Gulinqing, Yunnan, China); C. Hainanphasma cristata Ho, 2013 (Mt. jianfengling, Hainan Island, China); D. P. pui Ho, 2013 (Mengla, Yunnan, China). Photo credit A-C. Pei-Lin Liao; D. Chao Wu." figureDoi="10.3897/jor.31.79783.fig3" httpUri="https://binary.pensoft.net/fig/718486" pageId="0" pageNumber="125">3D</figureCitation>
);
<taxonomicName authorityName="Gao &amp; Xie" authorityYear="2022" class="Insecta" family="Heteropterygidae" genus="Pylaemenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pylaemenes gulinqingensis" order="Phasmatodea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="gulinqingensis">
<emphasis italics="true" pageId="0" pageNumber="125">Pylaemenes gulinqingensis</emphasis>
<taxonomicName id="6268CBC398C2C90CBAB23C43ABA61306" authorityName="Gao &amp; Xie" authorityYear="2022" class="Insecta" family="Heteropterygidae" genus="Pylaemenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pylaemenes gulinqingensis" order="Phasmatodea" pageId="0" pageNumber="125" phylum="Arthropoda" rank="species" species="gulinqingensis">
<emphasis id="8CB071219B915A27D8EEFD7393C07A83" italics="true" pageId="0" pageNumber="125">Pylaemenes gulinqingensis</emphasis>
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sp. nov. [China: Wenshan].
</paragraph>
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<paragraph pageId="0" pageNumber="125">
<emphasis italics="true" pageId="0" pageNumber="125">Suggested common name (Chinese).</emphasis>
<subSubSection id="D573F05EC2D150C39645515E20B22ECA" pageId="0" pageNumber="125" type="suggested common name (chinese).">
<paragraph id="E55DD50299A8F9ED80A53DF303AE2499" pageId="0" pageNumber="125">
<emphasis id="563002948AB28313A2011F057D54280A" italics="true" pageId="0" pageNumber="125">Suggested common name (Chinese).</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="125">
<normalizedToken originalValue="瘤䗛属">瘤䗛属</normalizedToken>
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<normalizedToken id="A297017466738E84B1AE07FBD2F0E984" originalValue="瘤䗛属">瘤䗛属</normalizedToken>
</paragraph>
</subSubSection>
</treatment>

View file

@ -1,185 +1,185 @@
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<taxonomicName LSID="6D89FAA9-C7E6-5846-B30B-86D44DE894B4" authority="Sturany, 1905" authorityName="Sturany" authorityYear="1905" class="Bivalvia" family="Anatinellidae" genus="Raeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raeta jickelii" order="Venerida" pageId="0" pageNumber="571" phylum="Mollusca" rank="species" species="jickelii">Raeta jickelii Sturany, 1905</taxonomicName>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Raeta jickelii Sturany, 1905, Massaua, Eritrea. A, D. Original figures in Sturany, 1905; B, C, E. Illustrated syntype NHMW- Mollusca 39925: left valve outside view (B), left valve inside view (C), hinge detail (E). Scale bars: B, C: 10 mm, E: 2 mm." figureDoi="10.3897/zse.96.54707.figure3" httpUri="https://binary.pensoft.net/fig/448009" pageId="0" pageNumber="571">Figure 3</figureCitation>
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<figureCitation id="503FDF642EF446A061A63978CD2EA191" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Raeta jickelii Sturany, 1905, Massaua, Eritrea. A, D. Original figures in Sturany, 1905; B, C, E. Illustrated syntype NHMW- Mollusca 39925: left valve outside view (B), left valve inside view (C), hinge detail (E). Scale bars: B, C: 10 mm, E: 2 mm." figureDoi="10.3897/zse.96.54707.figure3" httpUri="https://binary.pensoft.net/fig/448009" pageId="0" pageNumber="571">Figure 3</figureCitation>
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<paragraph pageId="0" pageNumber="571">Sturany, 1905: 133-1134, with three figures.</paragraph>
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<paragraph id="07EBB6739E5D9B25961D4A4195E9CC3F" pageId="0" pageNumber="571">Sturany, 1905: 133-1134, with three figures.</paragraph>
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<paragraph pageId="0" pageNumber="571">Type material.</paragraph>
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<paragraph id="066C055302A54724DA8E04D9761D5B49" pageId="0" pageNumber="571">Type material.</paragraph>
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ERITREA • 1 complete left valve (of the original illustration), 2 fragmented left valves; Massaua; 8 Mar 1895; C.F. Jickeli coll.; syntypes; NHMW-
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39925; sizes: H = 31.7, L = 42.8 mm.
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<paragraph pageId="0" pageNumber="571">Original description.</paragraph>
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Diese Art wurde seinerzeit von Dr. Jickeli in Massaua in
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bloss die abgebildete linke Schale (43 mm lang, 31,5 mm hoch, 11,5 mm tief) gut erhalten und liegen
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nur noch die Fragmente von weiteren 2 linken Schalen (die bis 51 mm lang und 42 mm hoch gewesen sein
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eine rechte Valve gar nicht vertreten ist.
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Die
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Muschel hat vorne eine ziemlich starke
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in einen etwas nach aussen gekehrten Schnabel. Sie ist konzentrisch gefaltet, milchweiss
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das Schloss, welches eine Ligamentgrube (unter dem Wirbel), 2 davorstehende vertikale
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. Von den
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Arten sind
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<emphasis italics="true" pageId="0" pageNumber="571">Say und</emphasis>
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<emphasis id="3A8238BF84EE89139E9E9B8A012984F8" italics="true" pageId="0" pageNumber="571">Say und</emphasis>
B. abercrombiei
<emphasis italics="true" pageId="0" pageNumber="571">Melv.</emphasis>
<emphasis id="D8B97B76A6F0A245CE86C2F0ADF4E3F4" italics="true" pageId="0" pageNumber="571">Melv.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="571" type="translation">
<paragraph pageId="0" pageNumber="571">Translation.</paragraph>
<paragraph pageId="0" pageNumber="571">This species was discovered with incomplete specimens by Dr. Jickeli in Massaua. Only the figured left valve (43 mm long, 31.5 mm high, 11.5 mm deep) is in good condition and there are merely fragments of another two left valves (which were up to 51 mm long and 42 mm high), while a right valve is not present at all.</paragraph>
<paragraph pageId="0" pageNumber="571">
<subSubSection id="199D967808C94E90EDF5F2CB7CA8BD9C" pageId="0" pageNumber="571" type="translation">
<paragraph id="BE5CBA1C733388D8B3D04B0F1839F68F" pageId="0" pageNumber="571">Translation.</paragraph>
<paragraph id="96B51E37306D57774510BA7CA9E34B81" pageId="0" pageNumber="571">This species was discovered with incomplete specimens by Dr. Jickeli in Massaua. Only the figured left valve (43 mm long, 31.5 mm high, 11.5 mm deep) is in good condition and there are merely fragments of another two left valves (which were up to 51 mm long and 42 mm high), while a right valve is not present at all.</paragraph>
<paragraph id="BD6FD6B17BFF58A5456DCCB08EA1E41B" pageId="0" pageNumber="571">
The thin-shelled clam is rounded at the anterior end and has a slightly outwards turned posterior end. It is concentrically plicated, milky white and glossy. The beak lies centrally and dominates the hinge, which reveals a ligament pit (below the beak), two vertical teeth at the front and elongated front and back teeth running parallel to the upper margin. The anterior of the cardinal teeth is weak and simple, the posterior significantly stronger, but constricted from two sides, so that it has a star-shaped, three-pointed cross section. The muscle scars are indistinct, the pallial sinus extends to the centre. The closest related species are
<taxonomicName genus="R." lsidName="R. canaliculata" pageId="0" pageNumber="571" rank="species" species="canaliculata">
<emphasis italics="true" pageId="0" pageNumber="571">R. canaliculata</emphasis>
<taxonomicName id="AB7DC0620098678210753DB0F204E25E" lsidName="R. canaliculata" pageId="0" pageNumber="571" rank="species" species="canaliculata">
<emphasis id="4AC1C1301FE22F50791890E7514A12F4" italics="true" pageId="0" pageNumber="571">R. canaliculata</emphasis>
</taxonomicName>
Say and
<emphasis italics="true" pageId="0" pageNumber="571">B. abercrombiei</emphasis>
<emphasis id="2F1342AEA03446CD812C3AFA2402CBDE" italics="true" pageId="0" pageNumber="571">B. abercrombiei</emphasis>
Melv.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="571" type="comments">
<paragraph pageId="0" pageNumber="571">Comments.</paragraph>
<paragraph pageId="0" pageNumber="571">
<subSubSection id="5B65CDECAC387D68668ED49D99030DB1" pageId="0" pageNumber="571" type="comments">
<paragraph id="EA88197D73BAA77A12F7330C7B5ED5AD" pageId="0" pageNumber="571">Comments.</paragraph>
<paragraph id="EF108FB953D135B8C9AD6A145966D323" pageId="0" pageNumber="571">
Currently considered a junior synonym of
<taxonomicName baseAuthorityName="Reeve" baseAuthorityYear="1854" class="Bivalvia" family="Anatinellidae" genus="Raeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raeta pellicula" order="Venerida" pageId="0" pageNumber="571" phylum="Mollusca" rank="species" species="pellicula">
<emphasis italics="true" pageId="0" pageNumber="571">Raeta pellicula</emphasis>
<taxonomicName id="328B84C852584B2C6267FA73E904C362" baseAuthorityName="Reeve" baseAuthorityYear="1854" class="Bivalvia" family="Anatinellidae" genus="Raeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raeta pellicula" order="Venerida" pageId="0" pageNumber="571" phylum="Mollusca" rank="species" species="pellicula">
<emphasis id="2F1656C4145BE8FE3597FA50176DC015" italics="true" pageId="0" pageNumber="571">Raeta pellicula</emphasis>
</taxonomicName>
(Reeve, 1854) (
<bibRefCitation author="MolluscaBase" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="571" publicationUrl="http://www.molluscabase.org" refId="B15" refString="MolluscaBase, 2020. MolluscaBase. http://www.molluscabase.org" title="MolluscaBase." url="http://www.molluscabase.org" year="2020">MolluscaBase 2020</bibRefCitation>
<bibRefCitation id="8895CFB8D056283ECD1E5957AAD68B47" author="MolluscaBase" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="571" publicationUrl="http://www.molluscabase.org" refId="B15" refString="MolluscaBase, 2020. MolluscaBase. http://www.molluscabase.org" title="MolluscaBase." url="http://www.molluscabase.org" year="2020">MolluscaBase 2020</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zse.96.54707.figure3" httpUri="https://binary.pensoft.net/fig/448009" pageId="0" pageNumber="571" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="571">
<emphasis bold="true" pageId="0" pageNumber="571">Figure 3.</emphasis>
<taxonomicName authorityName="Sturany" authorityYear="1905" class="Bivalvia" family="Anatinellidae" genus="Raeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raeta jickelii" order="Venerida" pageId="0" pageNumber="571" phylum="Mollusca" rank="species" species="jickelii">
<emphasis italics="true" pageId="0" pageNumber="571">Raeta jickelii</emphasis>
<caption id="8A806E2707A767D8F4A59ABBCE740FAC" doi="10.3897/zse.96.54707.figure3" httpUri="https://binary.pensoft.net/fig/448009" pageId="0" pageNumber="571" start="Figure 3" startId="F3">
<paragraph id="AE49E868472C9CA9CC531648D55389BA" pageId="0" pageNumber="571">
<emphasis id="C8877E2DB5B113063A559279B47EAE07" bold="true" pageId="0" pageNumber="571">Figure 3.</emphasis>
<taxonomicName id="F0D9C67AD7AE0E5792C6E823E4AA5479" authorityName="Sturany" authorityYear="1905" class="Bivalvia" family="Anatinellidae" genus="Raeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raeta jickelii" order="Venerida" pageId="0" pageNumber="571" phylum="Mollusca" rank="species" species="jickelii">
<emphasis id="A464E5DEB21AFF7835CE52D8B1DDEFDE" italics="true" pageId="0" pageNumber="571">Raeta jickelii</emphasis>
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Sturany, 1905, Massaua, Eritrea.
<emphasis bold="true" pageId="0" pageNumber="571">A</emphasis>
<emphasis id="7523DC329327D94D5DD766330D3266CE" bold="true" pageId="0" pageNumber="571">A</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="571">D</emphasis>
<emphasis id="765C94C19B6A8FBC8D7D2A778E2783E1" bold="true" pageId="0" pageNumber="571">D</emphasis>
. Original figures in Sturany, 1905;
<emphasis bold="true" pageId="0" pageNumber="571">B</emphasis>
<emphasis id="718684A588977D5F4D1A3C844250B436" bold="true" pageId="0" pageNumber="571">B</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="571">C</emphasis>
<emphasis id="12B2610956D7D39C6C776289C49807E6" bold="true" pageId="0" pageNumber="571">C</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="571">E</emphasis>
<emphasis id="5F01F9F3EB13F44295735096A4435C5B" bold="true" pageId="0" pageNumber="571">E</emphasis>
. Illustrated syntype NHMW-
<taxonomicName authorityName="Gray" authorityYear="1853" class="Bivalvia" family="Anatinellidae" genus="Raeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Venerida" pageId="0" pageNumber="571" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
<taxonomicName id="72D743F2175462D0799B789DF73930C0" authorityName="Gray" authorityYear="1853" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" pageId="0" pageNumber="571" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
39925: left valve outside view (
<emphasis bold="true" pageId="0" pageNumber="571">B</emphasis>
<emphasis id="97CFD3B120399004EBC04682F0A7E898" bold="true" pageId="0" pageNumber="571">B</emphasis>
), left valve inside view (
<emphasis bold="true" pageId="0" pageNumber="571">C</emphasis>
<emphasis id="287B593C232623E689C4EB381FF099E5" bold="true" pageId="0" pageNumber="571">C</emphasis>
), hinge detail (
<emphasis bold="true" pageId="0" pageNumber="571">E</emphasis>
<emphasis id="5938878D319A186DAED1FA4E9294457A" bold="true" pageId="0" pageNumber="571">E</emphasis>
). Scale bars:
<emphasis bold="true" pageId="0" pageNumber="571">B, C</emphasis>
<emphasis id="2B53BAB8060797ADA6C6D4851A775ACB" bold="true" pageId="0" pageNumber="571">B, C</emphasis>
: 10 mm,
<emphasis bold="true" pageId="0" pageNumber="571">E</emphasis>
<emphasis id="9AC74D82125AF483809DEF7A6A06EA4F" bold="true" pageId="0" pageNumber="571">E</emphasis>
: 2 mm.
</paragraph>
</caption>

View file

@ -1,384 +1,384 @@
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<subSubSection id="E8683B356EBEAEE3D514354C3467D7C3" pageId="0" pageNumber="93" type="nomenclature">
<paragraph id="52660C7D8B55405E225CC995960B7CBD" pageId="0" pageNumber="93">
<taxonomicName id="5FE992B842E2E82EB086F2D7D24345DB" LSID="http://zoobank.org/16D79540-7339-434A-9662-29D1BB5F360B" authority="Song &amp; Qi" authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis" status="sp. nov.">Stenochironomus baishanzuensis Song &amp; Qi</taxonomicName>
<taxonomicNameLabel id="AE46FE88454E4BAED9577FF4DA7DD9D1" pageId="0" pageNumber="93">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="description">
<paragraph pageId="0" pageNumber="93">
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult (holotype, in dorsal view) of Stenochironomus baishanzuensis Song &amp; Qi sp. nov. male." figureDoi="10.3897/zookeys.1104.81403.figure6" httpUri="https://binary.pensoft.net/fig/693998" pageId="0" pageNumber="93">Figs 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A hypopygium in dorsal view B hypopygium in ventral view C superior volsella." figureDoi="10.3897/zookeys.1104.81403.figure8" httpUri="https://binary.pensoft.net/fig/694000" pageId="0" pageNumber="93">, 8</figureCitation>
<subSubSection id="187D379782286C324CF1A5000284A68B" pageId="0" pageNumber="93" type="description">
<paragraph id="3D5B3D380316CE6F889C93A87870437C" pageId="0" pageNumber="93">
<figureCitation id="E6C748466CD7CC5FD5C89C997DA0F41C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult (holotype, in dorsal view) of Stenochironomus baishanzuensis Song &amp; Qi sp. nov. male." figureDoi="10.3897/zookeys.1104.81403.figure6" httpUri="https://binary.pensoft.net/fig/693998" pageId="0" pageNumber="93">Figs 6</figureCitation>
<figureCitation id="925649D8E5B03E00C90555E31CEA9ED0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">, 7</figureCitation>
<figureCitation id="FDABA6D01CAA03BADF2778624C0C6432" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A hypopygium in dorsal view B hypopygium in ventral view C superior volsella." figureDoi="10.3897/zookeys.1104.81403.figure8" httpUri="https://binary.pensoft.net/fig/694000" pageId="0" pageNumber="93">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="type material">
<paragraph pageId="0" pageNumber="93">Type material.</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Holotype</emphasis>
<subSubSection id="998B8DEABA2D4760E1DC75A965D2FE4B" pageId="0" pageNumber="93" type="type material">
<paragraph id="BA4678D325926928992B2B2DDAEB3EE7" pageId="0" pageNumber="93">Type material.</paragraph>
<paragraph id="15909361FD2410B7F9B7B9FA714B621E" pageId="0" pageNumber="93">
<emphasis id="6E61F42FC2AC47553749626785D3D175" bold="true" italics="true" pageId="0" pageNumber="93">Holotype</emphasis>
(BOLD &amp; TZU sample ID: ZJCH226; Field ID: BSZ93) 1 male, China, Zhejiang Province, Lishui City, Qingyuan County, Baishanzu National Nature Reserve,
<geoCoordinate degrees="27.76" direction="north" orientation="latitude" precision="555" value="27.76">27.76°N</geoCoordinate>
<geoCoordinate id="96F25DDAEDFB1034B050761A87768C89" degrees="27.76" direction="north" orientation="latitude" precision="555" value="27.76">27.76°N</geoCoordinate>
,
<geoCoordinate degrees="119.31" direction="east" orientation="longitude" precision="555" value="119.31">119.31°E</geoCoordinate>
<geoCoordinate id="97078A02F20AC916B1225BF6E9325BBE" degrees="119.31" direction="east" orientation="longitude" precision="555" value="119.31">119.31°E</geoCoordinate>
, 11-12. VIII. 2020, Qi X. &amp; Song C., collected by light trap.
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Paratypes</emphasis>
<emphasis id="624AA5A786F34F1A72A2C90FF92631D8" bold="true" italics="true" pageId="0" pageNumber="93">Paratypes</emphasis>
: 2 males, same data as for holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="description">
<paragraph pageId="0" pageNumber="93">Diagnostic characters.</paragraph>
<paragraph pageId="0" pageNumber="93">
<subSubSection id="CEAAAE15455103BB110EFBC90A015CDB" pageId="0" pageNumber="93" type="description">
<paragraph id="9E3BDF9FEB657AD0E5274CBA180BC8D3" pageId="0" pageNumber="93">Diagnostic characters.</paragraph>
<paragraph id="30E64C14C9C9ECCCB2C70BBB235D0CE3" pageId="0" pageNumber="93">
Adult males of
<taxonomicName genus="S." lsidName="S. baishanzuensis" pageId="0" pageNumber="93" rank="species" species="baishanzuensis">
<emphasis italics="true" pageId="0" pageNumber="93">S. baishanzuensis</emphasis>
<taxonomicName id="345A947E720E0379899C9775E696ED4D" lsidName="S. baishanzuensis" pageId="0" pageNumber="93" rank="species" species="baishanzuensis">
<emphasis id="A060DE369A7D77567BE6FF6CEF52C472" italics="true" pageId="0" pageNumber="93">S. baishanzuensis</emphasis>
</taxonomicName>
sp. nov. can be distinguished from other related species by the following combination of characters: membrane of wing with large dark spots on median and apical parts; median vitta, lateral vitta, and postnotum with pigmentation; superior volsella short and broad with four setae; inferior volsella with four long bristles and one stout terminal spine, not overreaching apex of anal point.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="etymology">
<paragraph pageId="0" pageNumber="93">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="93">The specific name refers to the Baishanzu National Nature Reserve, where the holotype was collected.</paragraph>
<subSubSection id="483697BEDBBDF3BED8C4C99007F38F9A" pageId="0" pageNumber="93" type="etymology">
<paragraph id="69B2B192F7C004C1DBDA1187B3B294DC" pageId="0" pageNumber="93">Etymology.</paragraph>
<paragraph id="A7EEECA3369FC9482E5DAEE477DD70E9" pageId="0" pageNumber="93">The specific name refers to the Baishanzu National Nature Reserve, where the holotype was collected.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="description">
<paragraph pageId="0" pageNumber="93">Description.</paragraph>
<paragraph pageId="0" pageNumber="93">Male imago (N = 3). Total length 3.98-4.22, 4.07 mm. Wing length 1.90-2.13, 2.05 mm. Total length / wing length 1.87-2.22, 1.99. Wing length / length of pro-femur 1.63-1.65, 1.64.</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Coloration</emphasis>
<subSubSection id="F117C4D99CBA8CB2C282EFF5BB241F08" pageId="0" pageNumber="93" type="description">
<paragraph id="04E7C78158151BC0026FDD39CD7144C2" pageId="0" pageNumber="93">Description.</paragraph>
<paragraph id="E43792DDDC2FB8A9A89F03BFEC0DA707" pageId="0" pageNumber="93">Male imago (N = 3). Total length 3.98-4.22, 4.07 mm. Wing length 1.90-2.13, 2.05 mm. Total length / wing length 1.87-2.22, 1.99. Wing length / length of pro-femur 1.63-1.65, 1.64.</paragraph>
<paragraph id="CAF76EDA58D5AFA4CA124384E1B83990" pageId="0" pageNumber="93">
<emphasis id="E5B490A900C3C205DD2EDCA6FA281C9D" bold="true" italics="true" pageId="0" pageNumber="93">Coloration</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult (holotype, in dorsal view) of Stenochironomus baishanzuensis Song &amp; Qi sp. nov. male." figureDoi="10.3897/zookeys.1104.81403.figure6" httpUri="https://binary.pensoft.net/fig/693998" pageId="0" pageNumber="93">6</figureCitation>
<figureCitation id="07DE1A70D9B0105EB4EEABBF0467085F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult (holotype, in dorsal view) of Stenochironomus baishanzuensis Song &amp; Qi sp. nov. male." figureDoi="10.3897/zookeys.1104.81403.figure6" httpUri="https://binary.pensoft.net/fig/693998" pageId="0" pageNumber="93">6</figureCitation>
). Head. Antennal hairs dark; palpomeres dark. Thorax almost pale yellow, with posterior portion of media vitta, posterior 3/4 portion of postnotum, and almost the lateral vitta dark brown. Membrane with 2 dark spots located around RM and FCu areas (median spot is darker). Legs. P1: knee, tibia and Ta 1 dark brown, basal 3/4 of femur and Ta 2-5 pale yellowish; P2: yellowish with brownish knee; P3: yellowish with brownish knees and tibia. Abdomen almost light brown with T VI-VII dark brown.
</paragraph>
<caption doi="10.3897/zookeys.1104.81403.figure6" httpUri="https://binary.pensoft.net/fig/693998" pageId="0" pageNumber="93" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figure 6.</emphasis>
<caption id="92358672F8C87465107300584ECBDE3B" doi="10.3897/zookeys.1104.81403.figure6" httpUri="https://binary.pensoft.net/fig/693998" pageId="0" pageNumber="93" start="Figure 6" startId="F6">
<paragraph id="9DD79C7C52B1E0A3D7541FDC5D7E14A0" pageId="0" pageNumber="93">
<emphasis id="E89B10B3296EF95C65410F49B4825636" bold="true" pageId="0" pageNumber="93">Figure 6.</emphasis>
Male adult (holotype, in dorsal view) of
<taxonomicName authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis">
<emphasis italics="true" pageId="0" pageNumber="93">Stenochironomus baishanzuensis</emphasis>
<taxonomicName id="18D4C03A4E6262FDE7679CFA2716578B" authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis">
<emphasis id="7407E14FB5B111E4CC8CC29D60B34D68" italics="true" pageId="0" pageNumber="93">Stenochironomus baishanzuensis</emphasis>
</taxonomicName>
Song &amp; Qi sp. nov. male.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">
<emphasis italics="true" pageId="0" pageNumber="93">Head</emphasis>
<paragraph id="5576BAE3A44535DAF99D85752643F279" pageId="0" pageNumber="93">
<emphasis id="B1E2043132A68BDAE81C38C36DD63077" bold="true" pageId="0" pageNumber="93">
<emphasis id="3D2DBB562C0985525996774CEC72AE38" italics="true" pageId="0" pageNumber="93">Head</emphasis>
.
</emphasis>
AR 1.93, ultimate flagellomere 810
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="CBB07F298CB4CEE6CA7984045E6024B2" originalValue="µm">µm</normalizedToken>
long (n = 1); Temporals 12-15, 13 setae, including 5-7, 6 inner and 4-6, 5 outer verticals and 2-3, 3, postorbitals. Clypeus with 14-20, 18 setae. Tentorium 155-193, 170
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="6E0796280DFA65AE75F34588867225F2" originalValue="μm">μm</normalizedToken>
long, 35-53, 44
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="0070DD62BFF1CB0826A2B449C151E0F2" originalValue="μm">μm</normalizedToken>
wide. Palp 5-segmented, lengths (in
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="29DBE040F96357828109828EDF619901" originalValue="μm">μm</normalizedToken>
) of segments: 42-55, 47; 43-65, 53; 200-223, 214; 130-155, 146; 232-275, 253. Palpomere ratio (5th / 3rd) 1.13-1.26.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">
<emphasis italics="true" pageId="0" pageNumber="93">Thorax</emphasis>
<paragraph id="CB682667F86899E7590E109283F172BC" pageId="0" pageNumber="93">
<emphasis id="3D50D97038A9CE3B913D6EFE6AF42C9E" bold="true" pageId="0" pageNumber="93">
<emphasis id="58D9F2985A93BBD969AB2FBF7EEFF15F" italics="true" pageId="0" pageNumber="93">Thorax</emphasis>
.
</emphasis>
Dorsocentrals 15-16, 15; acrostichals 12-15, 14; prealars 6-7, 7. Scutellum with 8-9, 9 setae in 2 rows.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Wing</emphasis>
<paragraph id="BAEB9E51A32421C2B812489E5A379F2A" pageId="0" pageNumber="93">
<emphasis id="54D03D75676155EBE62482CD1CB93B86" bold="true" italics="true" pageId="0" pageNumber="93">Wing</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">7A</figureCitation>
<figureCitation id="18B23777FA5FC50DD98E67EE6CD89438" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">7A</figureCitation>
). VR 1.05-1.06, 1.06. Brachiolum with 2 setae. Distribution of setae on veins: R, 30-32, 31; R1, 22-26, 25; R4+5, 36-42, 39. Squama with 10-11, 11 setae. Anal lobe normally developed.
</paragraph>
<caption doi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figure 7.</emphasis>
<caption id="47C17779C40DCC53C824DDCD1385CE9A" doi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93" start="Figure 7" startId="F7">
<paragraph id="470B9DEEC10EB69023964233DF6D0D10" pageId="0" pageNumber="93">
<emphasis id="0789CA62F9FC8A7DEA0D055C21C2E438" bold="true" pageId="0" pageNumber="93">Figure 7.</emphasis>
Male adult (holotype) of
<taxonomicName authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis">
<emphasis italics="true" pageId="0" pageNumber="93">Stenochironomus baishanzuensis</emphasis>
<taxonomicName id="89F4748EFFA264BB609DC92DAAEA4BA9" authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis">
<emphasis id="DDBD2356B12C7A0432562C96B3B4F812" italics="true" pageId="0" pageNumber="93">Stenochironomus baishanzuensis</emphasis>
</taxonomicName>
Song &amp; Qi, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="93">A</emphasis>
<emphasis id="E0DE6EEFC7D3E5C3A08C3B1D5AC15739" bold="true" pageId="0" pageNumber="93">A</emphasis>
wing
<emphasis bold="true" pageId="0" pageNumber="93">B</emphasis>
<emphasis id="B03726DA30290D27F01353A53581CF65" bold="true" pageId="0" pageNumber="93">B</emphasis>
hypopygium in dorsal
<emphasis bold="true" pageId="0" pageNumber="93">C</emphasis>
<emphasis id="39DBAB3C39CE650A2DAA4FB320A2DCB2" bold="true" pageId="0" pageNumber="93">C</emphasis>
hypopygium in ventral view
<emphasis bold="true" pageId="0" pageNumber="93">D</emphasis>
<emphasis id="D751B70C6A9B4977A51745F9E1D33366" bold="true" pageId="0" pageNumber="93">D</emphasis>
legs.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Legs</emphasis>
<paragraph id="37C8BFE29CABACCF9C86A0D212C07657" pageId="0" pageNumber="93">
<emphasis id="0807ACB6D6690FDDCA736EBD03A95EC1" bold="true" italics="true" pageId="0" pageNumber="93">Legs</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">7D</figureCitation>
<figureCitation id="57AA4D170B9F3434D6296959780C1F6E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">7D</figureCitation>
). Fore leg: apex of tibia 55-61, 58
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="272F731BD4A79EC5197773AEF8B565E6" originalValue="μm">μm</normalizedToken>
width, tibia with pointed scale 35-38
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="2A6C04A112D56C61F15A3F2DA5D9CF9E" originalValue="µm">µm</normalizedToken>
long. Mid leg: apex of tibia 60-68, 65
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="3E6FC8CD6973C6F7C682988FACD7317F" originalValue="μm">μm</normalizedToken>
width; tibial spurs 48-55, 51 and 45-55, 49
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="AA4035E5EF0CD47CA4F5A98B55B969BB" originalValue="µm">µm</normalizedToken>
long, completely fused at midline part. Hind leg: apex of tibia 60-68, 64
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="74362832A28D509AE520AC6314E55455" originalValue="μm">μm</normalizedToken>
width, tibia with 2 apical spurs 37-50, 47 and 35-47, 41
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="B2612561F869A47661857C7ADCF592F2" originalValue="µm">µm</normalizedToken>
long. Lengths (in
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="CAF5A4BCEFCE95563843E37335102D27" originalValue="μm">μm</normalizedToken>
) and proportions of legs in Table
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Male adult of Stenochironomus baishanzuensis sp. nov. Lengths (in µm) and proportions of legs (N = 3, except where otherwise stated)." httpUri="http://table.plazi.org/id/26F317C58EC03E2E30E0B53D7B8712BF" pageId="0" pageNumber="93" tableUuid="26F317C58EC03E2E30E0B53D7B8712BF">5</tableCitation>
<tableCitation id="0A336E66BC531729F943E9C5D15D0C98" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Male adult of Stenochironomus baishanzuensis sp. nov. Lengths (in µm) and proportions of legs (N = 3, except where otherwise stated)." httpUri="http://table.plazi.org/id/26F317C58EC03E2E30E0B53D7B8712BF" pageId="0" pageNumber="93" tableUuid="26F317C58EC03E2E30E0B53D7B8712BF">5</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="26F317C58EC03E2E30E0B53D7B8712BF" httpUri="http://table.plazi.org/id/26F317C58EC03E2E30E0B53D7B8712BF" pageId="0" pageNumber="93" start="Table 5" startId="T5">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Table 5.</emphasis>
<caption id="26F317C58EC03E2E30E0B53D7B8712BF" ID-Table-UUID="26F317C58EC03E2E30E0B53D7B8712BF" httpUri="http://table.plazi.org/id/26F317C58EC03E2E30E0B53D7B8712BF" pageId="0" pageNumber="93" start="Table 5" startId="T5">
<paragraph id="E3B7FB2ED0AAD44DAFDC35B484299CAC" pageId="0" pageNumber="93">
<emphasis id="FDBA545798CC3C332E7A571DE2DF053B" bold="true" pageId="0" pageNumber="93">Table 5.</emphasis>
Male adult of
<taxonomicName authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis">
<emphasis italics="true" pageId="0" pageNumber="93">Stenochironomus baishanzuensis</emphasis>
<taxonomicName id="4131EF6DCCC49A8F8D32B9F48BBF6BE9" authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis">
<emphasis id="EC1A0BF92780435268432095699BE4CA" italics="true" pageId="0" pageNumber="93">Stenochironomus baishanzuensis</emphasis>
</taxonomicName>
sp. nov. Lengths (in
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="A4B3F5B87179EDE9DE0D217423CBA71D" originalValue="µm">µm</normalizedToken>
) and proportions of legs (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="DCF32B1545C4024FF2661F47A13E377F" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 3, except where otherwise stated).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="93">
<table pageId="0" pageNumber="93">
<tr pageId="0" pageNumber="93">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="93" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="93" rowspan="1">P1</th>
<th colspan="1" pageId="0" pageNumber="93" rowspan="1">P2</th>
<th colspan="1" pageId="0" pageNumber="93" rowspan="1">P3</th>
<paragraph id="B66F9AAF0A2315A67630DC2AEDD0AB6D" pageId="0" pageNumber="93">
<table id="E48A782098E377D3805FADE4748CED97" pageId="0" pageNumber="93">
<tr id="6BE8E7E10558DCAAE5BFB1134336BE31" pageId="0" pageNumber="93">
<th id="22F96F2EE8D94EC3B48B1650B20ACBC7" colspan="1" isEmpty="true" pageId="0" pageNumber="93" rowspan="1">-</th>
<th id="C98AA918A5AD7EBEA5714D61971132BE" colspan="1" pageId="0" pageNumber="93" rowspan="1">P1</th>
<th id="8DDF293F49E86BB7E2C85A91E61B2988" colspan="1" pageId="0" pageNumber="93" rowspan="1">P2</th>
<th id="8114B1E2A7E6C070B8740539AF06FF61" colspan="1" pageId="0" pageNumber="93" rowspan="1">P3</th>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">Fe</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">1150-1300, 1250</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">970-1110, 1053</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">1140-1310, 1243</td>
<tr id="A65DCE1AC5A151846C03B6C86716BD80" pageId="0" pageNumber="93">
<td id="7FA7E298285DF846ED5943E289673569" colspan="1" pageId="0" pageNumber="93" rowspan="1">Fe</td>
<td id="2C48A40EC0F4D9DAC6450FCB0D59C215" colspan="1" pageId="0" pageNumber="93" rowspan="1">1150-1300, 1250</td>
<td id="4BBBD6AEBC53D83B6094F2D0653153BC" colspan="1" pageId="0" pageNumber="93" rowspan="1">970-1110, 1053</td>
<td id="5BA87686A2EF213A53820C67123CB878" colspan="1" pageId="0" pageNumber="93" rowspan="1">1140-1310, 1243</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">Ti</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">1130-1280, 1230</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">870-930, 907</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">1050-1250, 1160</td>
<tr id="35A0EC9301348DB8EA9F32295F477616" pageId="0" pageNumber="93">
<td id="A661C0CF4A44C9978185FCEE7D25F3BB" colspan="1" pageId="0" pageNumber="93" rowspan="1">Ti</td>
<td id="D5844D9EC58C0B2BA0F48D9C10A0243B" colspan="1" pageId="0" pageNumber="93" rowspan="1">1130-1280, 1230</td>
<td id="59E1555E1EF57727033603893C105C50" colspan="1" pageId="0" pageNumber="93" rowspan="1">870-930, 907</td>
<td id="6415FEF5A0BA2C4BC6DA467052688201" colspan="1" pageId="0" pageNumber="93" rowspan="1">1050-1250, 1160</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 1</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">
<tr id="5DEEAAA3353D5852202F3651AC884CEC" pageId="0" pageNumber="93">
<td id="BAB344800E4BA5F0CB068BDE8D12E147" colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 1</td>
<td id="7982365736AEC9C457A9B4269A1AA9C7" colspan="1" pageId="0" pageNumber="93" rowspan="1">
1250 (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="083CDED9620FF2A42F0FBAB23768F6C6" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 1)
</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">480-700, 606</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">820-950, 900</td>
<td id="CAC08050067667A7D3B7E550F735934B" colspan="1" pageId="0" pageNumber="93" rowspan="1">480-700, 606</td>
<td id="8764ED4829E4ACC675B7FD266A8232D6" colspan="1" pageId="0" pageNumber="93" rowspan="1">820-950, 900</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 2</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">
<tr id="5EF4D5E055EA1CFF69DD60E0D583551C" pageId="0" pageNumber="93">
<td id="C3A47CBAD22F07597507E407AFB9FF03" colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 2</td>
<td id="17B4C6AD9E3B022925F42FD862B3728E" colspan="1" pageId="0" pageNumber="93" rowspan="1">
800 (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="39C30CC3A20F5679A943D2236D90E195" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 1)
</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">370-400, 387</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">450-520, 477</td>
<td id="4A08D41734591B809C666715EEBBF173" colspan="1" pageId="0" pageNumber="93" rowspan="1">370-400, 387</td>
<td id="60E893A3643FD86F65AB52EE9887C7F8" colspan="1" pageId="0" pageNumber="93" rowspan="1">450-520, 477</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 3</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">
<tr id="C4AEFC196C1D492480BB50FF4089445D" pageId="0" pageNumber="93">
<td id="694ACA4951B1306B0016B57A8CEEF904" colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 3</td>
<td id="2A6F08B8FF90F8E83D809FF6BE297534" colspan="1" pageId="0" pageNumber="93" rowspan="1">
680 (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="3D6CC96512D743E1545E84A436A3702D" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 1)
</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">300-340, 317</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">380-410, 397</td>
<td id="CD8DCCEB2F7A6B618A7962912D637139" colspan="1" pageId="0" pageNumber="93" rowspan="1">300-340, 317</td>
<td id="92F6406FECA76B8E3CA1B592EBC9A298" colspan="1" pageId="0" pageNumber="93" rowspan="1">380-410, 397</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 4</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">
<tr id="7939B81052179D183E2EB29311CC976E" pageId="0" pageNumber="93">
<td id="1816CFC56E4B291CDB56A5709F6FEF10" colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 4</td>
<td id="D62B68E3C50724E6D7CC57A8ED43EA59" colspan="1" pageId="0" pageNumber="93" rowspan="1">
560 (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="0A834EB9348FD26CD4363F6C513139E1" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 1)
</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">180-210, 200</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">230-260, 240</td>
<td id="574728F27772986E7E53D6EFE7EE8639" colspan="1" pageId="0" pageNumber="93" rowspan="1">180-210, 200</td>
<td id="C50806B297F6E387D0F7E3FFDEC67599" colspan="1" pageId="0" pageNumber="93" rowspan="1">230-260, 240</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 5</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">
<tr id="DB56799F815A47BB7221A839A4A43C8F" pageId="0" pageNumber="93">
<td id="10A92C4A5A91E82AEEE0EA719C8719A8" colspan="1" pageId="0" pageNumber="93" rowspan="1">Ta 5</td>
<td id="70D01BA27D2B61082DCA8A1546CA761A" colspan="1" pageId="0" pageNumber="93" rowspan="1">
270 (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="465B1F72AA812E02F22BD5AA59E7C7EB" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 1)
</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">90-100, 95</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">90-110, 100</td>
<td id="E8C53A0B9873CDA6B3482DA5EEA9BC29" colspan="1" pageId="0" pageNumber="93" rowspan="1">90-100, 95</td>
<td id="8E513B990E2EB0835C419542B6E3A214" colspan="1" pageId="0" pageNumber="93" rowspan="1">90-110, 100</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">LR</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">
<tr id="25CFA1CF90DB5909C280B32631348A60" pageId="0" pageNumber="93">
<td id="C9D840D90F34EF68C1C16BD3EF600FF9" colspan="1" pageId="0" pageNumber="93" rowspan="1">LR</td>
<td id="541B2ED31248F2F5F614B051504449D3" colspan="1" pageId="0" pageNumber="93" rowspan="1">
0.98 (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="F4D19F23B904A0D2FD6853938181F7C9" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 1)
</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">0.52-0.75, 0.67</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">0.76-0.78, 0.77</td>
<td id="7787D77F2B381DD70391B96D3847968D" colspan="1" pageId="0" pageNumber="93" rowspan="1">0.52-0.75, 0.67</td>
<td id="0BB32C78403770111CC1FA2F53F3A923" colspan="1" pageId="0" pageNumber="93" rowspan="1">0.76-0.78, 0.77</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">BV</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">
<tr id="FCD7FFD4CC4EDA3A9BEEA63388633079" pageId="0" pageNumber="93">
<td id="C3290F1A679A3A0D23AEACF14129FC0E" colspan="1" pageId="0" pageNumber="93" rowspan="1">BV</td>
<td id="3AF6FD048818A062A95F0B6F2A0D38B1" colspan="1" pageId="0" pageNumber="93" rowspan="1">
2.35 (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="9E43E4CE77E4785503C8CE1C2C8964DB" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 1)
</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">2.46-2.64, 2.57</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">2.59-2.94, 2.72</td>
<td id="1155F2910801390EE778A2663D169C2A" colspan="1" pageId="0" pageNumber="93" rowspan="1">2.46-2.64, 2.57</td>
<td id="30377019DF2CCBCE648DAC9D0698FDE9" colspan="1" pageId="0" pageNumber="93" rowspan="1">2.59-2.94, 2.72</td>
</tr>
<tr pageId="0" pageNumber="93">
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">SV</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">
<tr id="105471EEEB81865E12AE97015D29CCD0" pageId="0" pageNumber="93">
<td id="96BE5480CD0AD80E1964D6F26511C849" colspan="1" pageId="0" pageNumber="93" rowspan="1">SV</td>
<td id="D8C785A306738DD62C1FF5BB85BE29F8" colspan="1" pageId="0" pageNumber="93" rowspan="1">
2.06 (
<emphasis italics="true" pageId="0" pageNumber="93">N</emphasis>
<emphasis id="79F8AAD66072E050627B425C93EFA449" italics="true" pageId="0" pageNumber="93">N</emphasis>
= 1)
</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">2.91-4.17, 3.31</td>
<td colspan="1" pageId="0" pageNumber="93" rowspan="1">2.66-2.67, 2.67</td>
<td id="9F8A10FB33B1548E8EBBB2551981B943" colspan="1" pageId="0" pageNumber="93" rowspan="1">2.91-4.17, 3.31</td>
<td id="1EF376823C3DD717556EDF554474C2E8" colspan="1" pageId="0" pageNumber="93" rowspan="1">2.66-2.67, 2.67</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Hypopygium</emphasis>
<paragraph id="022CAD2B30D15E81D2B8004BD23010DB" pageId="0" pageNumber="93">
<emphasis id="59BA44D4976DD1700AC874463F189121" bold="true" italics="true" pageId="0" pageNumber="93">Hypopygium</emphasis>
(Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">7B, C</figureCitation>
<figureCitation id="6C027F511E1E0ED46D5ED787BD795E2B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">7B, C</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A hypopygium in dorsal view B hypopygium in ventral view C superior volsella." figureDoi="10.3897/zookeys.1104.81403.figure8" httpUri="https://binary.pensoft.net/fig/694000" pageId="0" pageNumber="93">8</figureCitation>
<figureCitation id="F84B30444DEF004DB31BB40CAEE76395" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A hypopygium in dorsal view B hypopygium in ventral view C superior volsella." figureDoi="10.3897/zookeys.1104.81403.figure8" httpUri="https://binary.pensoft.net/fig/694000" pageId="0" pageNumber="93">8</figureCitation>
). Anal point 110-125, 120
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="0C865A7EA0ED7C3BA172AF531A1A46C0" originalValue="μm">μm</normalizedToken>
long, 30-45, 35
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="42D6306958213C8BFE8181D14CEB61C8" originalValue="µm">µm</normalizedToken>
wide at base, 10-13, 12
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="B83E8CA53CE78F924C31251E5758B944" originalValue="µm">µm</normalizedToken>
wide at apex. Tergite IX with 18-23, 20 long setae medially, and 3-4, 4 setae laterally. Posterior margin of tergite IX with 3-4, 4 spines and 5-7 long setae each side. Transverse sternapodeme 38-53, 44
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="7EA98768BE8FE8F55E29E04E92991991" originalValue="μm">μm</normalizedToken>
long; phallapodeme 95-113, 105
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="D9128C71D1A978DFAEADB07F4C2AAFD5" originalValue="μm">μm</normalizedToken>
long. Gonocoxite 183-188, 185
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="A1556BBE6ECFF21585E553CFE5B9C863" originalValue="μm">μm</normalizedToken>
long. Gonostylus 218-25, 221
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="F0D591B4B0E0B3FA5CC6CC71F5E6F3AC" originalValue="μm">μm</normalizedToken>
long. Superior volsella short and broad, 13-15, 14
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="95F2C222EE1F4E406B39BC68B3D8E19D" originalValue="μm">μm</normalizedToken>
long, 18-25, 21
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="6580EDD24F098A5CEA68BACF53389D8A" originalValue="μm">μm</normalizedToken>
wide, with 4 long setae (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">7C</figureCitation>
<figureCitation id="1D5F3A14382F06B5C872E19B51323D8D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male adult (holotype) of Stenochironomus baishanzuensis Song &amp; Qi, sp. nov. A wing B hypopygium in dorsal C hypopygium in ventral view D legs." figureDoi="10.3897/zookeys.1104.81403.figure7" httpUri="https://binary.pensoft.net/fig/693999" pageId="0" pageNumber="93">7C</figureCitation>
). Inferior volsella elongate, 183-192, 187
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="64C03EA234F3E4E515334BA879213714" originalValue="μm">μm</normalizedToken>
long, extending at most, to apex of anal point, with 4 setae and 1 strong terminal spine. HR 0.83-0.84,.084, HV 1.77-1.93.
</paragraph>
<caption doi="10.3897/zookeys.1104.81403.figure8" httpUri="https://binary.pensoft.net/fig/694000" pageId="0" pageNumber="93" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figure 8.</emphasis>
<caption id="9A1987218AA12CAC71AFB0B97CB1E620" doi="10.3897/zookeys.1104.81403.figure8" httpUri="https://binary.pensoft.net/fig/694000" pageId="0" pageNumber="93" start="Figure 8" startId="F8">
<paragraph id="FF746F1075B094527E10885B42C8A182" pageId="0" pageNumber="93">
<emphasis id="BAB2A77544D2688742BCC85DE62AEB91" bold="true" pageId="0" pageNumber="93">Figure 8.</emphasis>
Male adult (holotype) of
<taxonomicName authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis">
<emphasis italics="true" pageId="0" pageNumber="93">Stenochironomus baishanzuensis</emphasis>
<taxonomicName id="2CBED505876CE4F1032558F439CF2B1C" authorityName="Song &amp; Qi" authorityYear="2022" class="Insecta" family="Chironomidae" genus="Stenochironomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenochironomus baishanzuensis" order="Diptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="baishanzuensis">
<emphasis id="E58E97CAC8A45C4C2C00169362FA8EF3" italics="true" pageId="0" pageNumber="93">Stenochironomus baishanzuensis</emphasis>
</taxonomicName>
Song &amp; Qi, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="93">A</emphasis>
<emphasis id="88F45809175F330E886231323635A527" bold="true" pageId="0" pageNumber="93">A</emphasis>
hypopygium in dorsal view
<emphasis bold="true" pageId="0" pageNumber="93">B</emphasis>
<emphasis id="28FD11D7ECDDAF6065CACE3F6A92306C" bold="true" pageId="0" pageNumber="93">B</emphasis>
hypopygium in ventral view
<emphasis bold="true" pageId="0" pageNumber="93">C</emphasis>
<emphasis id="F6ABEB54EED522896B61EA849F697C20" bold="true" pageId="0" pageNumber="93">C</emphasis>
superior volsella.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="93">Immature stages and female unknown.</paragraph>
<paragraph id="432849086730A5584430D7BF1460A296" pageId="0" pageNumber="93">Immature stages and female unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="remarks">
<paragraph pageId="0" pageNumber="93">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="93">
<subSubSection id="9C7BFE7DDF63A97F04C257198C1F618B" pageId="0" pageNumber="93" type="remarks">
<paragraph id="BCB90E609CEFC8745E2A635CEC3DF031" pageId="0" pageNumber="93">Remarks.</paragraph>
<paragraph id="2047F47746BC4E17B6D2D613D0367AF8" pageId="0" pageNumber="93">
The male adult of
<taxonomicName genus="S." lsidName="S. baishanzuensis" pageId="0" pageNumber="93" rank="species" species="baishanzuensis">
<emphasis italics="true" pageId="0" pageNumber="93">S. baishanzuensis</emphasis>
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<emphasis id="D6E67B988E8359ACFE629AD2155605BB" italics="true" pageId="0" pageNumber="93">S. baishanzuensis</emphasis>
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sp. nov. resembles that of
<taxonomicName genus="S." lsidName="S. gibbus" pageId="0" pageNumber="93" rank="species" species="gibbus">
<emphasis italics="true" pageId="0" pageNumber="93">S. gibbus</emphasis>
<taxonomicName id="5320AFBB773418ECBD85FB3EB3965402" lsidName="S. gibbus" pageId="0" pageNumber="93" rank="species" species="gibbus">
<emphasis id="273C1681E763746C28C18C9E5231EB1A" italics="true" pageId="0" pageNumber="93">S. gibbus</emphasis>
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(Fabricius, 1794) in the structure of the hypopygium and the wing patterns, but can be separated by the following characters: straight and parallel-sided anal point, and legs bearing different patterns (Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Main differences between S. annulus sp. nov., S. baishanzuensis sp. nov., S. gibbus, and S. xianjuensis." httpUri="http://table.plazi.org/id/75B996484767B660C9AD7B2025EBC620" pageId="0" pageNumber="93" tableUuid="75B996484767B660C9AD7B2025EBC620">4</tableCitation>
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).
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<subSubSection pageId="0" pageNumber="93" type="distribution">
<paragraph pageId="0" pageNumber="93">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="93">The species is currently known only from Zhejiang Province, Oriental China.</paragraph>
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<paragraph id="7959F3428D233C4C3A5A3B292F1DB790" pageId="0" pageNumber="93">Distribution.</paragraph>
<paragraph id="97425805524F80BDDD2F08BFB37D486E" pageId="0" pageNumber="93">The species is currently known only from Zhejiang Province, Oriental China.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,532 +1,532 @@
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<figureCitation id="F7F8F88938DC6A3DF204379136B85BB7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">, 5</figureCitation>
<figureCitation id="92946A2F1D7EAA97759958763FB7FC8A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">, 6</figureCitation>
<figureCitation id="B3B5607FBD08D97023FEFAF9647722EB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">, 7C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="materials_examined">
<paragraph pageId="0" pageNumber="105">Material.</paragraph>
<paragraph pageId="0" pageNumber="105">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Ukraine" county="It" location="Klesov" municipality="The" specimenCount="1" specimenCount-female="1" stateProvince="Rovno Region" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="105">
<typeStatus>Holotype</typeStatus>
<subSubSection id="8B39059FDEF0155A2FBF1A5A5446AE83" pageId="0" pageNumber="105" type="materials_examined">
<paragraph id="C854860A06797444D5842018305C02E7" pageId="0" pageNumber="105">Material.</paragraph>
<paragraph id="CD66C3E1ACD7A8D1C0D0A1520C8098C9" pageId="0" pageNumber="105">
<materialsCitation id="70597F5B72FA878F15EC35DDCF6DCD27" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Ukraine" county="It" location="Klesov" municipality="The" specimenCount="1" specimenCount-female="1" stateProvince="Rovno Region" typeStatus="Holotype">
<emphasis id="33941A822EA1EF3D373CBBD0BB9F45CD" bold="true" italics="true" pageId="0" pageNumber="105">
<typeStatus id="AE7DF9E4FF7C446D6695E241F99FC980">Holotype</typeStatus>
</emphasis>
, SIZK K-27005,
<specimenCount type="female">1 ♀</specimenCount>
<specimenCount id="671CC3F8452C8C86F39B787044B9058C" type="female">1 ♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:6DC9150112785681B544FBCC16A72905:26D55D3E6C90206EBEA5E31BE96DE9A4" country="Ukraine" county="It" municipality="The" name="Klesov" stateProvince="Rovno Region">Klesov</location>
<location id="26D55D3E6C90206EBEA5E31BE96DE9A4" LSID="urn:lsid:plazi:treatment:6DC9150112785681B544FBCC16A72905:26D55D3E6C90206EBEA5E31BE96DE9A4" country="Ukraine" county="It" municipality="The" name="Klesov" stateProvince="Rovno Region">Klesov</location>
,
<location LSID="urn:lsid:plazi:treatment:6DC9150112785681B544FBCC16A72905:B2077E599EA6E0B30650BF1CE5C2E4D7" country="Ukraine" county="It" municipality="The" name="Sarny District" stateProvince="Rovno Region">Sarny District</location>
<location id="B2077E599EA6E0B30650BF1CE5C2E4D7" LSID="urn:lsid:plazi:treatment:6DC9150112785681B544FBCC16A72905:B2077E599EA6E0B30650BF1CE5C2E4D7" country="Ukraine" county="It" municipality="The" name="Sarny District" stateProvince="Rovno Region">Sarny District</location>
,
<collectingRegion country="Ukraine" name="Rivne">Rovno Region</collectingRegion>
<collectingRegion id="3F91E67D3EE4BC99C35BB4D5CB9128A7" country="Ukraine" name="Rivne">Rovno Region</collectingRegion>
,
<collectingCountry name="Ukraine">Ukraine</collectingCountry>
<collectingCountry id="EC504DB4D4F2DEA3A1304A7B0693B5BA" name="Ukraine">Ukraine</collectingCountry>
;
<collectingRegion country="Ukraine" name="Rivne">Rovno</collectingRegion>
<collectingRegion id="2FB521ACAFFC3ACDA85262D441B27712" country="Ukraine" name="Rivne">Rovno</collectingRegion>
amber; late
<location LSID="urn:lsid:plazi:treatment:6DC9150112785681B544FBCC16A72905:91F58D04A057ACA677360D11CB7E8ACA" country="Ukraine" county="It" municipality="The" name="Eocene" stateProvince="Rovno Region">Eocene</location>
<location id="91F58D04A057ACA677360D11CB7E8ACA" LSID="urn:lsid:plazi:treatment:6DC9150112785681B544FBCC16A72905:91F58D04A057ACA677360D11CB7E8ACA" country="Ukraine" county="It" municipality="The" name="Eocene" stateProvince="Rovno Region">Eocene</location>
(deposited in SIZK).
<collectingMunicipality>The</collectingMunicipality>
<collectingMunicipality id="B69B58D124C3002ED16647265A6D43EB">The</collectingMunicipality>
inclusion is in a reddish-yellow and clear, parallelepiped piece of amber (ca. 7
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="DB0C6F3AC8800D527B6B7F744CC74688" originalValue="×">x</normalizedToken>
6.5
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
<normalizedToken id="75B33D8B7E87A5477C9D963CBC982E6C" originalValue="×">x</normalizedToken>
<quantity id="6B4673B303E15213680E4F567029A665" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
).
<collectingCounty>It</collectingCounty>
<collectingCounty id="41B272CD104EAD72565B0F9EEDF73B85">It</collectingCounty>
is well preserved, but its left side is obscured by a large air bubble (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1B</figureCitation>
<figureCitation id="D5FD4889E4204A95F3FA770E32718186" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1B</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2</figureCitation>
<figureCitation id="56E80A2A108945FCAD16A095E12DD5AE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2</figureCitation>
).
</materialsCitation>
</paragraph>
<caption doi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 1.</emphasis>
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
<caption id="7329B87F56008CF084A6BF68EAC3D460" doi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105" start="Figure 1" startId="F1">
<paragraph id="6AC840250F9279CA976EDC7ABB30776C" pageId="0" pageNumber="105">
<emphasis id="844D47197A774C803D9AB99EE609768E" bold="true" pageId="0" pageNumber="105">Figure 1.</emphasis>
<taxonomicName id="B5E7C263494595B7440658877DF50D7F" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="C5DB142C4960C81C38FD63E849C05B43" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
</taxonomicName>
gen. et sp. nov.,
<typeStatus>holotype</typeStatus>
<typeStatus id="3915DEFBC8A9A50AEFC7CC341A43C2E6">holotype</typeStatus>
female
<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
<emphasis id="56310F09B4DC1647FF664DBE7A6237CB" bold="true" pageId="0" pageNumber="105">A</emphasis>
body, lateral view
<emphasis bold="true" pageId="0" pageNumber="105">B</emphasis>
<emphasis id="763037CCD5A3B0126B051F1D1EAF7ABA" bold="true" pageId="0" pageNumber="105">B</emphasis>
body, anterodorsolateral view.
</paragraph>
</caption>
<caption doi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 2.</emphasis>
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
<caption id="1B5772B98D76858E15EB4A2AE3F50A53" doi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105" start="Figure 2" startId="F2">
<paragraph id="E5E1A4F1285AB8BE5A46E595579D4AB5" pageId="0" pageNumber="105">
<emphasis id="2B48992727B7CE012AD05B495006B0BA" bold="true" pageId="0" pageNumber="105">Figure 2.</emphasis>
<taxonomicName id="90A3D7A67B61D896661BF647942EF963" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="A7FE5712E7D52CF7B4846C1D6AFB850D" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
</taxonomicName>
gen. et sp. nov.,
<typeStatus>holotype</typeStatus>
<typeStatus id="CB1834D429137FAD181428B3A78EB0A5">holotype</typeStatus>
female
<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
<emphasis id="9A3C7E949E076328BF4FFE4DAF3220DA" bold="true" pageId="0" pageNumber="105">A</emphasis>
body, ventral (arrows indicate the metatibial pegs in inset)
<emphasis bold="true" pageId="0" pageNumber="105">B</emphasis>
<emphasis id="9FD74F0A1E479BF46998771F8750F718" bold="true" pageId="0" pageNumber="105">B</emphasis>
body, posterodorsal.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="syninclusions">
<paragraph pageId="0" pageNumber="105">Syninclusions.</paragraph>
<paragraph pageId="0" pageNumber="105">
<taxonomicName authorityName="Simutnik" authorityYear="2022" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" rank="subOrder" subOrder="Nematocera">Nematocera</taxonomicName>
<subSubSection id="6EA84D1A8605EB1B6D08CB4012A9B75C" pageId="0" pageNumber="105" type="syninclusions">
<paragraph id="CA5B29DFC88E7377D3B9AA58FCD82A7D" pageId="0" pageNumber="105">Syninclusions.</paragraph>
<paragraph id="46D5C8E2BC266CA74E6EAA851B4A7130" pageId="0" pageNumber="105">
<taxonomicName id="A4848E696FD728A9B164014D53C0455E" authorityName="Simutnik" authorityYear="2022" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="subOrder" subOrder="Nematocera">Nematocera</taxonomicName>
, spider web.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="etymology">
<paragraph pageId="0" pageNumber="105">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="105">The specific epithet means ancient in Latin.</paragraph>
<subSubSection id="C1D78F42A0A2D288F039B7157D8D6903" pageId="0" pageNumber="105" type="etymology">
<paragraph id="CC242E4FEFA74C50A7BECBD640B96D07" pageId="0" pageNumber="105">Etymology.</paragraph>
<paragraph id="AEAE8C3805BBFF0D3B4525CD782007BD" pageId="0" pageNumber="105">The specific epithet means ancient in Latin.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="description">
<paragraph pageId="0" pageNumber="105">Description.</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Female.</emphasis>
<subSubSection id="91EA076CE6E07603CEF7AC16837EBDD3" pageId="0" pageNumber="105" type="description">
<paragraph id="6479F30B68491B898D6534CF46970449" pageId="0" pageNumber="105">Description.</paragraph>
<paragraph id="876C33731C47E873A38210B62959A976" pageId="0" pageNumber="105">
<emphasis id="83FDE7DD0F7FEDDA65F2F5D320129B34" bold="true" pageId="0" pageNumber="105">Female.</emphasis>
Habitus as in Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1</figureCitation>
<figureCitation id="9687C7C9D7810626347D5295F9D00AA8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2</figureCitation>
<figureCitation id="DE932A57B0F0FB6AB8AB160C1556C368" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2</figureCitation>
. Body length 1.3 mm.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" italics="true" pageId="0" pageNumber="105">Coloration.</emphasis>
<paragraph id="5042A4BD31CC5A6FECEF25288747C8F1" pageId="0" pageNumber="105">
<emphasis id="1776CC7F628FFB0AF1E60832AE5DF2B5" bold="true" italics="true" pageId="0" pageNumber="105">Coloration.</emphasis>
Head, thorax, gaster dorsally, tegula, and all femora black; antenna uniformly dark brown; gaster ventrally dark brown; venation brown; mesotibial spur and tarsi pale yellow; surface of frontovertex, thorax, legs shiny due to presence of a thin layer of air, without metallic shine.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" italics="true" pageId="0" pageNumber="105">Sculpture.</emphasis>
<paragraph id="660990463F46AEA1A8050C8740C17388" pageId="0" pageNumber="105">
<emphasis id="5507DEF50AC91348F3C8B9E87DC60F81" bold="true" italics="true" pageId="0" pageNumber="105">Sculpture.</emphasis>
Head, pronotum and mesoscutum rough reticulate, but surface of head only sparsely punctate - frontovertex with 4 vertical rows of large but shallow piliferous punctures (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3A</figureCitation>
<figureCitation id="947D6E9CAF3F767156D6BAB946B46145" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3A</figureCitation>
), outer row separated from eye margin by a distance equal to or more than diameter of this punctures; scape, pedicel, scutellum, tegula, mesopleuron, coxae, legs, and gaster also relatively similarly but more smoothly reticulate; sculpture of face as in Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="03859C4484BF8989A44CFA1D09055BDA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
- horizontal row of three large, deep cells located under each torulus, lateral to clypeus, also intricately sculptured.
</paragraph>
<caption doi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 3.</emphasis>
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
<caption id="BCEAA3BB5EC1C4B8544476EBE8A1B52F" doi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105" start="Figure 3" startId="F3">
<paragraph id="185324CDD45E55490EE6689583209BFD" pageId="0" pageNumber="105">
<emphasis id="E8A6E9152ACE34CF8444F5E5E696664A" bold="true" pageId="0" pageNumber="105">Figure 3.</emphasis>
<taxonomicName id="0F6E81677EFA82944FD1F4D9DB8CC648" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="DCAA65540216B5BD84871EB2602A3692" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
</taxonomicName>
gen. et sp. nov., holotype female
<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
<emphasis id="7F43925A12D2A46B3A438321ABB42EE0" bold="true" pageId="0" pageNumber="105">A</emphasis>
head, antennae, part of mesosoma, anterodorsolateral view
<emphasis bold="true" pageId="0" pageNumber="105">B</emphasis>
<emphasis id="61BCB7DFB820C0C5874E6A952D0B2286" bold="true" pageId="0" pageNumber="105">B</emphasis>
head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression).
</paragraph>
</caption>
<caption doi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 4.</emphasis>
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
<caption id="1FF80F08BA80C9A5074AF90CA5E90CFA" doi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105" start="Figure 4" startId="F4">
<paragraph id="85B192DFFAB7B7ED179929E3140E52B5" pageId="0" pageNumber="105">
<emphasis id="AD793761A1C3B5B8EEDC211FFF4363EC" bold="true" pageId="0" pageNumber="105">Figure 4.</emphasis>
<taxonomicName id="F956EFC0622B856D462BA460491579F7" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="4930BFD7ED257185FAF49909AC4DA959" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
</taxonomicName>
gen. et sp. nov., holotype female, antennae, head anteroventral.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" italics="true" pageId="0" pageNumber="105">Head.</emphasis>
<paragraph id="93C24F01D6F5002B4BBC6B016292851A" pageId="0" pageNumber="105">
<emphasis id="C2A111377D7CA2C669CF958C80939471" bold="true" italics="true" pageId="0" pageNumber="105">Head.</emphasis>
Lenticular, slightly wider than thorax in dorsal view; occipital margin sharp, but not carinate (Fig.
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); frontovertex curved to posterior ocelli; broadly rounded in frontal view, not vaulted above eyes; eyes bare, without visible setae, with inner orbits parallel over much of height but ventrally divergent (Figs
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<figureCitation id="C5DCEFA40ED5844E0D929BFB0C8B0415" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3B</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="7D8C86B06F81746F9A4ED7396F07700A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
); frontovertex as long as broad, minimum distance between eyes about 0.4
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head width; ocelli forming a right angled triangle, posterior ones closer to eye than occipital margin; OOL about equal to posterior ocellar diameter; OOL:POL:LOL:OCL about 3:11:8:4; eye reaching occipital margin (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A</figureCitation>
<figureCitation id="DBDE9E87777A9C098B97EAA2B622EDD9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3</figureCitation>
<figureCitation id="00CC379E200E012A2CB524CD2D633D03" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3</figureCitation>
); antennal scrobes as in Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3</figureCitation>
<figureCitation id="5DD1242F2C20B9648FEC4A55123F2FC0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="D4F9D5A3250150ECBBC1882855280869" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
, v- or u-shaped and meeting dorsally, not extended to anterior ocellus, in dorsal view anterior ocellus about equidistant from occipital margin and from upper margin of scrobal depression; interantennal prominence presents (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="B0A68EC46B61F81B4A5EA5B0C80E9E3A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
); antennal toruli located about at level of lower eye margin, separated from mouth margin by distance equal to their own width (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="2D0225A7ABA3826E913E087F7BC9AC06" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
); distance between toruli equal to distance between torulus and eye, about 1.5
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width of torulus, about 0.5
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<normalizedToken id="73A26157D38427396335DEB158A96991" originalValue="×">x</normalizedToken>
length of malar space; malar space with complete malar sulcus (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="B84917BAEB11CB5E5755E67517EA88F5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
), about 2
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shorter than height of eye; clypeus strongly emarginate, with short lateral margins; mandible 3-dentate, with middle tooth longest (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="C367F162D1BA86A9D867F6760AFED073" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Antenna</emphasis>
<paragraph id="F891F208CAF0DAE254480B238B412BC9" pageId="0" pageNumber="105">
<emphasis id="F1D1649114B1666FB120464E823C852F" bold="true" pageId="0" pageNumber="105">
<emphasis id="15B85B96DC80783CC804FEF3A7E16930" italics="true" pageId="0" pageNumber="105">Antenna</emphasis>
.
</emphasis>
Geniculate, 11-segmented, without differentiated anelli, with 6 funicular segments and 3-segmented clava; radicle short, about 2
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<normalizedToken id="0F3E9B5E6CAA75B07BB211E05F9778C6" originalValue="×">x</normalizedToken>
as long as broad (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2B</figureCitation>
<figureCitation id="14A55FDD165F56D0CB770702440F40C0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2B</figureCitation>
); antennal scape ~4
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as long as broad; pedicel conical, little longer than first two funicular segments combined, longer than any segment of funicle; F1 subquadrate, slightly longer than broad, F2 and F3 distinctly longer than broad, F4-F6 subquadrate, F3-F6 slightly flattened; width of flagellomeres slightly increases toward apex; F5 and F6 noticeably wider than F2-F4 (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="9D5326E2D1056993D7A7B94F47A05B58" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
); all segments of funicle, at least F2-F6, and two basal segments of clava with mps; clava a little shorter than F3-F6 combined, with small oblique truncation at apical segment only (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3</figureCitation>
<figureCitation id="43A2967A6878F95B246B1B56B05975F1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
<figureCitation id="40CF31D51EA206AA816C69C2757188C5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Electronoyesella antiqua gen. et sp. nov., holotype female, antennae, head anteroventral." figureDoi="10.3897/jhr.94.94773.figure4" httpUri="https://binary.pensoft.net/fig/784450" pageId="0" pageNumber="105">4</figureCitation>
), wider than F6; flagellum and clava clothed in very short setae.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Mesosoma</emphasis>
<paragraph id="A8828B734F9B6529BCF199DD8C782B1B" pageId="0" pageNumber="105">
<emphasis id="EA757ECAADA0E5395633B553F0DAC98C" bold="true" pageId="0" pageNumber="105">
<emphasis id="FC98C3B1AF0FD5C7644278F4FA49408B" italics="true" pageId="0" pageNumber="105">Mesosoma</emphasis>
.
</emphasis>
Pronotum short, not conspicuously narrower than mesoscutum, almost vertical (in lateral view), with posterior edge only slightly emarginate (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3B</figureCitation>
<figureCitation id="55ECE4D29ED2919CEAD04A5CB8C72A1C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3B</figureCitation>
); mesoscutum as broad as long or nearly to (in dorsal view); mesothoracic spiracle open, not concealed beneath pronotum (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A</figureCitation>
<figureCitation id="F67FC1B58E141CEDEE6E6BAF7F1552F7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A</figureCitation>
); notauli present as depressions at extreme antero-lateral margin of mesoscutum (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3A, B</figureCitation>
<figureCitation id="15468C24BC2510D67A1D4FCE3ECD589D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Electronoyesella antiqua gen. et sp. nov., holotype female A head, antennae, part of mesosoma, anterodorsolateral view B head, antennae, part of mesosoma, anterodorsal view (arrow indicates the notaular depression)." figureDoi="10.3897/jhr.94.94773.figure3" httpUri="https://binary.pensoft.net/fig/784449" pageId="0" pageNumber="105">3A, B</figureCitation>
: arrow); meeting of axillae not visible; scutellum flat, apically pointed, with several long, vertical setae at own apex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2B</figureCitation>
<figureCitation id="B0BFA000E7EA3409702C14B7DB626704" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2B</figureCitation>
); mesopleuron enlarged posteriorly; metapleuron triangular, narrow, without visible setation (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A</figureCitation>
<figureCitation id="2B8DABC0644C28E670D174ED23CA69E8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A</figureCitation>
); propodeum bare, with relatively large lateral parts, touching hind coxa (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A</figureCitation>
<figureCitation id="52835F36B76C8AA558F6F423B4A0CAEA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Wings</emphasis>
<paragraph id="A5D13D50029AAC75DC1DA6829EE6030B" pageId="0" pageNumber="105">
<emphasis id="83443CF36F5BEE958465B54151C8DB22" bold="true" pageId="0" pageNumber="105">
<emphasis id="DBC39C2F63F446D4F406AE79F0033466" italics="true" pageId="0" pageNumber="105">Wings</emphasis>
.
</emphasis>
Fully developed, hyaline. Forewing 2.5
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<normalizedToken id="22E8B8527892AE8649DE059D82F632C4" originalValue="×">x</normalizedToken>
as long as broad; linea calva not interrupted but closed on posterior margin, without filum spinosum, with well-developed line of long setae alongside basal margin of dorsal surface (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5A, B</figureCitation>
<figureCitation id="728BA85250442B059BD0C151EBD2972B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5A, B</figureCitation>
); parastigma thickened, hyaline break (unpigmented area) present; marginal vein 5
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<normalizedToken id="2D78231E1A98ED4224F675B022C3EB18" originalValue="×">x</normalizedToken>
as long as broad; stigmal vein as long as marginal, with long narrow uncus consisting row of 4 uncal sensilla (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5B</figureCitation>
<figureCitation id="40CB44B179EF854B3A8A42FF7E83935C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5B</figureCitation>
); postmarginal vein 1.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B2F9000B0F317B3F65BFBAE76F737494" originalValue="×">x</normalizedToken>
as long as marginal vein, enlarged seta marking apex of postmarginal vein of forewing absent (as long as others on this vein); setae of marginal fringe short. Hindwing relatively wide; membrane of costal cell along submarginal vein with line of long setae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
<figureCitation id="7CA2A9284D12BB5261323C3953143366" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
: ls1); spur vein originating from submarginal vein visible as differentiated hyaline process, alongside this a row of 5 thickened setae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
<figureCitation id="EEA528975D131AC7E8864A08857E5F94" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Electronoyesella antiqua gen. et sp. nov., holotype female A wings B forewing venation C hind wing venation (ls 1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (A); 0.1 mm (B, C)." figureDoi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105">5C</figureCitation>
: spv, ls2); apex of marginal vein with 3 hamuli; marginal fringe ~ 0.2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="5D60F41BCC800BF2EBDEA5138FC0ABE2" originalValue="×">x</normalizedToken>
as long as wing width.
</paragraph>
<caption doi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 5.</emphasis>
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
<caption id="52854DBA9F3B9ACDA16456526E55C0D3" doi="10.3897/jhr.94.94773.figure5" httpUri="https://binary.pensoft.net/fig/784451" pageId="0" pageNumber="105" start="Figure 5" startId="F5">
<paragraph id="B55DCEF76F3D2EBFBC52D028EA891505" pageId="0" pageNumber="105">
<emphasis id="E1D6FC92EBA119F7A331ED3039F2E508" bold="true" pageId="0" pageNumber="105">Figure 5.</emphasis>
<taxonomicName id="AA3BA57358FAB36536B861D697E6A757" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="A5BCABE63F1E28CFA683A2D33AAA6A69" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
</taxonomicName>
gen. et sp. nov., holotype female
<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
<emphasis id="E87C82049877BE446BD5F367AD3D6115" bold="true" pageId="0" pageNumber="105">A</emphasis>
wings
<emphasis bold="true" pageId="0" pageNumber="105">B</emphasis>
<emphasis id="59907E394523713FDFF0CE35CF71280C" bold="true" pageId="0" pageNumber="105">B</emphasis>
forewing venation
<emphasis bold="true" pageId="0" pageNumber="105">C</emphasis>
<emphasis id="9A40C907E36CE68BC43E9BF93545DF59" bold="true" pageId="0" pageNumber="105">C</emphasis>
hind wing venation (ls1, 2 - lines of setae, spv - spur vein, pst - parastigma). Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
<emphasis id="446B2C50300306019FF019787D3801A0" bold="true" pageId="0" pageNumber="105">A</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="105">B, C</emphasis>
<emphasis id="51B52D618E2BBB5F8E8E09D634136635" bold="true" pageId="0" pageNumber="105">B, C</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Legs</emphasis>
<paragraph id="CEABB8AECC675E241872D3346D3C9E94" pageId="0" pageNumber="105">
<emphasis id="DF7982CFAAA08AE0B34724BCFAD86731" bold="true" pageId="0" pageNumber="105">
<emphasis id="1910CBE1CAF3CB0DEA91D9C1010C0056" italics="true" pageId="0" pageNumber="105">Legs</emphasis>
.
</emphasis>
Mid tibial spur about as long as basal mid tarsal segment or slightly shorter, both relatively long; mesotibia with row of pegs along anteroapical edge (Figs
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<figureCitation id="DDFF097FDCB931D6608E58DEEA15610F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1B</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
<figureCitation id="62687AF47F7BC558D00CE1401302BB5F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
); ventral surface of mesobasitarsus and each next tarsomere with differentiated setation along anteroventral edge (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
<figureCitation id="36C8D25791E92549F7F89382E34AB637" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
); apex of metatibia with one peg originating from round, deep pit and row of spine-like setae (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
<figureCitation id="BD7AECB65EF86103E1CB86CD638A16DB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Electronoyesella antiqua gen. et sp. nov., holotype female A body, ventral (arrows indicate the metatibial pegs in inset) B body, posterodorsal." figureDoi="10.3897/jhr.94.94773.figure2" httpUri="https://binary.pensoft.net/fig/784448" pageId="0" pageNumber="105">2A</figureCitation>
: arrows); tarsi 5-segmented.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Metasoma</emphasis>
<paragraph id="BB330A7961D058734414E614D21650CD" pageId="0" pageNumber="105">
<emphasis id="46E718B027225462DAB9E4AF264B9858" bold="true" pageId="0" pageNumber="105">
<emphasis id="B9BF62EE2909BC046EC454ABEDABA14B" italics="true" pageId="0" pageNumber="105">Metasoma</emphasis>
.
</emphasis>
As long as head and mesosoma together; cerci situated in apical third of metasoma, with long vertical setae (or bristles) (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A, B</figureCitation>
<figureCitation id="E41C74D460ABEEEEEFCECC7C703B2894" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Electronoyesella antiqua gen. et sp. nov., holotype female A body, lateral view B body, anterodorsolateral view." figureDoi="10.3897/jhr.94.94773.figure1" httpUri="https://binary.pensoft.net/fig/784447" pageId="0" pageNumber="105">1A, B</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
<figureCitation id="B8DB0991AF1653B4714E950A93B9C4B7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
); syntergum (Mt8 + Mt9) v-shaped, no longer than 1/3 of metasoma; posterior margins of Mt2-Mt5 straight, parallel; Mt6 and Mt7 slightly produced, M-shaped, U-like between cerci; apex of hypopygium with developed mucro, reaching apex of syntergum (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
<figureCitation id="D43CA6D92D279CE716700C2A878E3E5B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
); lateral margin of hypopygium bare, without row of setae; ovipositor stylet thick; ovipositor sheaths (gonostyli or third valvulae) visible in dorsal view (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6A, B</figureCitation>
<figureCitation id="C038E829E8771F32F739C30ADDA9DF69" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6A, B</figureCitation>
: osh (v3)), as long as mesobasitarsus, connect at some angle with second valvifers (inner plates of the ovipositor, Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
<figureCitation id="9C188E54918ED4C5D20D631A568DBC76" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
: vr2), apparently, movably fused; outer plates of ovipositor not visible but (as J. S. Noyes, pers. comm. 2022, suggested) appear very slender, appear to have filamentous connection to paratergite (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
<figureCitation id="D9A01EB625E3535EECEC4F7F514FF96B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
: fc, ptrg?) running forwards to outside of cercal plate, similar to that of
<taxonomicName class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
<taxonomicName id="B24CEDF4C1A2A0E53310AE924DF1E962" class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="16243C7EAE688B86E1F80C1782526FB4" italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7</figureCitation>
<figureCitation id="85C387111C0B851B2FB58A8616BC1B1A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7</figureCitation>
).
</paragraph>
<caption doi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 6.</emphasis>
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
<caption id="BB3E7598F82EE243363F7FFF9B973E95" doi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105" start="Figure 6" startId="F6">
<paragraph id="1AB16091B5CF53B064BA5F0076523797" pageId="0" pageNumber="105">
<emphasis id="02380B03F7AA41C9965E8ACBC5E3EB3C" bold="true" pageId="0" pageNumber="105">Figure 6.</emphasis>
<taxonomicName id="820324B3205D5082278E0304B99CBE9A" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="DE89988E89985C13FF15A8B0615A2CBE" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
</taxonomicName>
gen. et sp. nov., holotype female
<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
<emphasis id="B2B04267BF92AD86F6AFB93440D71459" bold="true" pageId="0" pageNumber="105">A</emphasis>
metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr2 - second valvifer)
<emphasis bold="true" pageId="0" pageNumber="105">B</emphasis>
<emphasis id="CEF76CFBAA235AA27E95343833B5E42C" bold="true" pageId="0" pageNumber="105">B</emphasis>
metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh(v3) - ovipositor sheaths).
</paragraph>
</caption>
<caption doi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Figure 7.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="105">A, B</emphasis>
<taxonomicName class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
<caption id="BAE6BF2338443A8A0B0A96024B69EC45" doi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105" start="Figure 7" startId="F7">
<paragraph id="E75BA3A33DB1158BD5229467827E4842" pageId="0" pageNumber="105">
<emphasis id="93DF9AAAA7C0226642F4141F91194C2B" bold="true" pageId="0" pageNumber="105">Figure 7.</emphasis>
<emphasis id="F8D5E4C2A44B13124405A41B31BD1D16" bold="true" pageId="0" pageNumber="105">A, B</emphasis>
<taxonomicName id="D20E4C124FDF7646F74FBDC4ECFAD47F" class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="C97352CB106330FF32987BC1C6450CCB" italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
</taxonomicName>
sp., female
<emphasis bold="true" pageId="0" pageNumber="105">A</emphasis>
<emphasis id="BE0A206DD417A31463FDADF6F38A16ED" bold="true" pageId="0" pageNumber="105">A</emphasis>
metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite)
<emphasis bold="true" pageId="0" pageNumber="105">B</emphasis>
<emphasis id="CD276A6579738C69D21124160239891F" bold="true" pageId="0" pageNumber="105">B</emphasis>
apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt7, v3 - ovipositor sheaths)
<emphasis bold="true" pageId="0" pageNumber="105">C</emphasis>
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
<emphasis id="508D62D811AFBEBAFD3B25E00F758835" bold="true" pageId="0" pageNumber="105">C</emphasis>
<taxonomicName id="37D36C36FAE270FAB95A084E71D3A092" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="FA381D1E6F0A394D159A296BFAEB250A" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
</taxonomicName>
gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="105">
<emphasis bold="true" pageId="0" pageNumber="105">Male.</emphasis>
<paragraph id="0047E57BD93AF41D05F093773054E0DE" pageId="0" pageNumber="105">
<emphasis id="2EF3C7F687264A08BB817347DD4B57AB" bold="true" pageId="0" pageNumber="105">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="genus composition">
<paragraph pageId="0" pageNumber="105">Genus composition.</paragraph>
<paragraph pageId="0" pageNumber="105">Type species only.</paragraph>
<subSubSection id="8C8DE259C7F419CD7B91D1F8F1576D3E" pageId="0" pageNumber="105" type="genus composition">
<paragraph id="6331435139D03689A53C71BE6D167681" pageId="0" pageNumber="105">Genus composition.</paragraph>
<paragraph id="270E0EB04A453A1FEAFEEA9430F707EA" pageId="0" pageNumber="105">Type species only.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="remarks">
<paragraph pageId="0" pageNumber="105">Remark.</paragraph>
<paragraph pageId="0" pageNumber="105">
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" rank="species" species="antiqua">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
<subSubSection id="B1648FF0EE7981487D3AE41656C89B18" pageId="0" pageNumber="105" type="remarks">
<paragraph id="D46665F5E3384FECFA191ABEF76F8C9C" pageId="0" pageNumber="105">Remark.</paragraph>
<paragraph id="D6F610FD8A1076EF6ED3C001442F02B9" pageId="0" pageNumber="105">
<taxonomicName id="6908529A309383687C7DAD65605F4274" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella antiqua" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="antiqua">
<emphasis id="937683141F061DC766B6505DA510EB10" italics="true" pageId="0" pageNumber="105">Electronoyesella antiqua</emphasis>
</taxonomicName>
gen. et sp. nov. from Rovno amber is the 23th non-ant hymenopteran genus (from 57, 40.4%) and 50th non-ant hymenopteran species (from 73, 68.5%), unknown from Baltic amber (Olmi 2022a, b).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="discussion">
<paragraph pageId="0" pageNumber="105">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="105">
<subSubSection id="62FFB613089E27D41040F93D69611FCF" pageId="0" pageNumber="105" type="discussion">
<paragraph id="335C5368034C5F92C34324BD878E9A8C" pageId="0" pageNumber="105">Discussion.</paragraph>
<paragraph id="D1E48EA0F19593CAD5DEAF4776D4058A" pageId="0" pageNumber="105">
Late Eocene and extant encyrtid faunas differ (
<bibRefCitation DOI="https://doi.org/10.1080/08912963.2020.1835887" author="Simutnik, SA" journalOrPublisher="Entomological Review" pageId="0" pageNumber="105" refId="B14" refString="Simutnik, SA, 2021. https://doi.org/10.1080/08912963.2020.1835887" url="https://doi.org/10.1080/08912963.2020.1835887" year="2021">Simutnik 2021</bibRefCitation>
<bibRefCitation id="49EA6875B00BD2985B88FA29BB5CE6EB" DOI="https://doi.org/10.1080/08912963.2020.1835887" author="Simutnik, SA" journalOrPublisher="Entomological Review" pageId="0" pageNumber="105" refId="B14" refString="Simutnik, SA, 2021. https://doi.org/10.1080/08912963.2020.1835887" url="https://doi.org/10.1080/08912963.2020.1835887" year="2021">Simutnik 2021</bibRefCitation>
). For example, fossil
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
<taxonomicName id="A8318CB60B3D1BF0858CD3977813B47A" baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
with reduced or shortened wings are unknown. The presence of the filum spinosum (FS), one of the main features of the subfamily
<taxonomicName lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
<taxonomicName id="A1F691212CFC75420DA22ED862E9980F" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
, have been found in about one third (22) of the 64 examined Eocene encyrtid specimens (males and females). In extant
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
<taxonomicName id="880E532C05DF1245ADE60BEFEA940973" baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
, the number of species of
<taxonomicName lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
<taxonomicName id="E9DCEC0589552578E94EB532422DD45A" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
(with
<emphasis italics="true" pageId="0" pageNumber="105">FS</emphasis>
<emphasis id="F8853F554311661BAD86EE16947FCF66" italics="true" pageId="0" pageNumber="105">FS</emphasis>
) is approximately three times more than the number of
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
<taxonomicName id="3D07AE9DBDE9332169E05F42112CE716" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
species (without
<emphasis italics="true" pageId="0" pageNumber="105">FS</emphasis>
<emphasis id="20B5C2CA8C9020C661304AA4204B7842" italics="true" pageId="0" pageNumber="105">FS</emphasis>
). The venation of the forewings and the structure of the linea calva in the fossil described here, as well as in other known Eocene
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
<taxonomicName id="9FE1004D92B1F96E0A22A353B779AA83" baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
without
<emphasis italics="true" pageId="0" pageNumber="105">FS</emphasis>
<emphasis id="4217D3C7E42361FB1BDE3D8529F0A01A" italics="true" pageId="0" pageNumber="105">FS</emphasis>
, most closely resemble those of
<taxonomicName authorityName="Tryapitsyn" authorityYear="1979" class="Insecta" family="Encyrtidae" genus="Savzdargia" kingdom="Animalia" lsidName="Savzdargia" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Savzdargia</emphasis>
<taxonomicName id="5542DAF46CB14D8B80814D8D5017FAB6" authorityName="Tryapitsyn" authorityYear="1979" class="Insecta" family="Encyrtidae" genus="Savzdargia" kingdom="Animalia" lsidName="Savzdargia" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="08082BC1CFD6DAB7CBDB2E4BC3811C95" italics="true" pageId="0" pageNumber="105">Savzdargia</emphasis>
</taxonomicName>
Trjapitzin, 1979;
<taxonomicName authorityName="Hoffer" authorityYear="1954" class="Insecta" family="Encyrtidae" genus="Moraviella" kingdom="Animalia" lsidName="Moraviella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Moraviella</emphasis>
<taxonomicName id="E282126D0B0FFC0A28A2BC73DE34B839" authorityName="Hoffer" authorityYear="1954" class="Insecta" family="Encyrtidae" genus="Moraviella" kingdom="Animalia" lsidName="Moraviella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="5703CAC7DB6DD248FBF3F758030FF0B7" italics="true" pageId="0" pageNumber="105">Moraviella</emphasis>
</taxonomicName>
Hoffer, 1954;
<taxonomicName class="Insecta" family="Encyrtidae" genus="Monodiscodes" kingdom="Animalia" lsidName="Monodiscodes" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Monodiscodes</emphasis>
<taxonomicName id="4FD3E1845CD65488BF79B7F95C63E7DD" class="Insecta" family="Encyrtidae" genus="Monodiscodes" kingdom="Animalia" lsidName="Monodiscodes" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="F4853C1D8D4C9288DEC7C2CBC626B3A2" italics="true" pageId="0" pageNumber="105">Monodiscodes</emphasis>
</taxonomicName>
Hoffer, 1954; possibly some species of
<taxonomicName class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
<taxonomicName id="282333701322C74C3CD5903860EC5D8F" class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="A43D63E3F78DC084996111B57783D921" italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
</taxonomicName>
; and some other extant
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
<taxonomicName id="D0522FE41054CCA83CFE9C42EAAF938E" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
. At the same time, distinct paratergites (the presence of which is one of the main features of the
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
<taxonomicName id="818D48D8F02C2121AF1ACB4B5940E4C8" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
:
<bibRefCitation author="Trjapitzin, VA" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="105" refId="B22" refString="Trjapitzin, VA, 1968. The problems of morphological evolution and the classification of the family Encyrtidae (Hymenoptera, Chalcidoidea). Readings in Memory of N.A. Kholodkovsky (1967), 44-62. [in Russian]" title="The problems of morphological evolution and the classification of the family Encyrtidae (Hymenoptera, Chalcidoidea). Readings in Memory of N. A. Kholodkovsky (1967), 44 - 62. [in Russian]" year="1968">Trjapitzin 1968</bibRefCitation>
<bibRefCitation id="F88C52313E0BDED8603E5572001252A3" author="Trjapitzin, VA" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="105" refId="B22" refString="Trjapitzin, VA, 1968. The problems of morphological evolution and the classification of the family Encyrtidae (Hymenoptera, Chalcidoidea). Readings in Memory of N.A. Kholodkovsky (1967), 44-62. [in Russian]" title="The problems of morphological evolution and the classification of the family Encyrtidae (Hymenoptera, Chalcidoidea). Readings in Memory of N. A. Kholodkovsky (1967), 44 - 62. [in Russian]" year="1968">Trjapitzin 1968</bibRefCitation>
) have not been previously found in fossil females. Therefore, there is not enough morphological evidence to classify them as members of the
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
<taxonomicName id="1E90A40524904D8D6E52A1879A8BE870" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4441.3.8" author="Simutnik, SA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="105" pagination="543 - 548" refId="B15" refString="Simutnik, SA, Perkovsky, EE, 2018. Archaeocercus gen. nov. (Hymenoptera, Chalcidoidea, En-cyrtidae) from late Eocene Rovno amber. Zootaxa 4441 (3): 543 - 548, DOI: https://doi.org/10.11646/zootaxa.4441.3.8" title="Archaeocercus gen. nov. (Hymenoptera, Chalcidoidea, En-cyrtidae) from late Eocene Rovno amber." url="https://doi.org/10.11646/zootaxa.4441.3.8" volume="4441" year="2018">Simutnik and Perkovsky 2018</bibRefCitation>
<bibRefCitation id="E063000A12CAA4504CA7B8FA78C5658E" DOI="https://doi.org/10.11646/zootaxa.4441.3.8" author="Simutnik, SA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="105" pagination="543 - 548" refId="B15" refString="Simutnik, SA, Perkovsky, EE, 2018. Archaeocercus gen. nov. (Hymenoptera, Chalcidoidea, En-cyrtidae) from late Eocene Rovno amber. Zootaxa 4441 (3): 543 - 548, DOI: https://doi.org/10.11646/zootaxa.4441.3.8" title="Archaeocercus gen. nov. (Hymenoptera, Chalcidoidea, En-cyrtidae) from late Eocene Rovno amber." url="https://doi.org/10.11646/zootaxa.4441.3.8" volume="4441" year="2018">Simutnik and Perkovsky 2018</bibRefCitation>
;
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<bibRefCitation id="1F1698C9850D09E560913D0D4A6A7AC3" DOI="https://doi.org/10.3897/jhr.89.79180" author="Simutnik, SA" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="105" pagination="47 - 60" refId="B20" refString="Simutnik, SA, Perkovsky, EE, Khomych, MR, Vasilenko, DV, 2022a. Two new genera of Encyrtidae (Hymenoptera, Chalcidoidea) with reduced ovipositor sheaths. Journal of Hymenoptera Research 89: 47 - 60, DOI: https://doi.org/10.3897/jhr.89.79180" title="Two new genera of Encyrtidae (Hymenoptera, Chalcidoidea) with reduced ovipositor sheaths." url="https://doi.org/10.3897/jhr.89.79180" volume="89" year="2022 a">Simutnik et al. 2022a</bibRefCitation>
).
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella</emphasis>
<taxonomicName id="B235454A86B5D573CB7A2EB3B646C80B" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="BE59B5E2A36B5130805342A52E67C1AF" italics="true" pageId="0" pageNumber="105">Electronoyesella</emphasis>
</taxonomicName>
has a sclerotized, ribbon-like, separated part of Mt8 along the lateral edge of the syntergum (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
<figureCitation id="70718B39E9A60551A728CE0644A9775B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Electronoyesella antiqua gen. et sp. nov., holotype female A metasoma, part of mesosoma, posterodorsal (cers - cercal seta, hyp - hypopygium, osh - ovipositor sheaths, ost - ovipositor stylet, ptrg? - paratergite?, spr - spiracle, syn - syntergum, vr 2 - second valvifer) B metasoma, part of mesosoma, posterolateral (cer - cercus, fc - filamentous connection, muc - mucro, opo? - possible, outer plate of the ovipositor, spv - hyaline spur vein; osh (v 3) - ovipositor sheaths)." figureDoi="10.3897/jhr.94.94773.figure6" httpUri="https://binary.pensoft.net/fig/784452" pageId="0" pageNumber="105">6B</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7C</figureCitation>
<figureCitation id="CED86C084E6566AD730AC0819859B8C7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7C</figureCitation>
: ptrg?). This structure closely resembles the paratergites of extant
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
<taxonomicName id="AD404F8073AC9386A30D7FA63943126E" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7A, B</figureCitation>
<figureCitation id="7518F07D6B3C7241188664D0B28CD7E6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7A, B</figureCitation>
: ptrg).
</paragraph>
<paragraph pageId="0" pageNumber="105">
<paragraph id="5D5686D1F736FF59F4AB85FB8E315A2A" pageId="0" pageNumber="105">
One of the most intriguing things in the evolution of encyrtids is what happens to the connection of tergite IX of the abdomen (Mt8) and the outer plates of the ovipositor (
<emphasis italics="true" pageId="0" pageNumber="105">OPO</emphasis>
<emphasis id="1C174EB8A5CA528C7267154EC9A842C1" italics="true" pageId="0" pageNumber="105">OPO</emphasis>
) (J. S. Noyes, pers. comm. 2022). The
<emphasis italics="true" pageId="0" pageNumber="105">OPO</emphasis>
<emphasis id="818CE71595C289B2C018EC0B3341BE85" italics="true" pageId="0" pageNumber="105">OPO</emphasis>
are completely separated from the syntergum behind the cercal plate in
<taxonomicName lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
<taxonomicName id="05B1B44E20CECAE0D9D3EF50C33D9AE4" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
. In
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<taxonomicName id="EE5AB013165D7A7F0151624C5222FBB6" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
they are connected by the paratergite. Both features almost certainly result from advancement of cerci (
<bibRefCitation author="Noyes, JS" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="105" pagination="1 - 459" refId="B5" refString="Noyes, JS, 2004. Encyrtidae of Costa Rica (Hymenoptera: Chalcidoidea), 2. Metaphycus and related genera, parasitoids of scale insects (Coccoidea) and whiteflies (Aleyrodidae). Memoirs of the American Entomological Institute 73: 1 - 459" title="Encyrtidae of Costa Rica (Hymenoptera: Chalcidoidea), 2. Metaphycus and related genera, parasitoids of scale insects (Coccoidea) and whiteflies (Aleyrodidae)." volume="73" year="2004">Noyes 2004</bibRefCitation>
<bibRefCitation id="40403967BE7906B1272BD9BF2CF87DD5" author="Noyes, JS" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="105" pagination="1 - 459" refId="B5" refString="Noyes, JS, 2004. Encyrtidae of Costa Rica (Hymenoptera: Chalcidoidea), 2. Metaphycus and related genera, parasitoids of scale insects (Coccoidea) and whiteflies (Aleyrodidae). Memoirs of the American Entomological Institute 73: 1 - 459" title="Encyrtidae of Costa Rica (Hymenoptera: Chalcidoidea), 2. Metaphycus and related genera, parasitoids of scale insects (Coccoidea) and whiteflies (Aleyrodidae)." volume="73" year="2004">Noyes 2004</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="105">
<paragraph id="0CB903AB87A44DD1D224F12860A55A99" pageId="0" pageNumber="105">
Cerci of the earliest known middle Eocene
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
<taxonomicName id="3BFFB1BC2DF30EC5A8102CEF77A170DC" baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
from Sakhalinian amber are close to each other, located at the very top of the gaster, similar to the ground plan state for
<taxonomicName authorityName="Simutnik" authorityYear="2022" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" rank="superFamily" superFamily="Chalcidoidea">Chalcidoidea</taxonomicName>
<taxonomicName id="FE5A98D303FC73F1BAF4B89BC1A7D237" authorityName="Simutnik" authorityYear="2022" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="superFamily" superFamily="Chalcidoidea">Chalcidoidea</taxonomicName>
. Then, tentatively in late Eocene, they began to shift towards the base of the gaster and the distance between them began to increase (figs 12-14 in
<bibRefCitation DOI="https://doi.org/10.1080/08912963.2020.1835887" author="Simutnik, SA" journalOrPublisher="Entomological Review" pageId="0" pageNumber="105" refId="B14" refString="Simutnik, SA, 2021. https://doi.org/10.1080/08912963.2020.1835887" url="https://doi.org/10.1080/08912963.2020.1835887" year="2021">Simutnik 2021</bibRefCitation>
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). This process was most likely the result of adaptation to parasitization of the host, probably by
<taxonomicName authorityName="Simutnik" authorityYear="2022" infraOrder="Coccinea" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" rank="infraOrder">Coccinea</taxonomicName>
<taxonomicName id="5930D824DE41EDABDE7BF95C4F443523" authorityName="Simutnik" authorityYear="2022" class="Insecta" higherTaxonomySource="CoL" infraOrder="Coccinea" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="infraOrder">Coccinea</taxonomicName>
(for example,
<taxonomicName authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Archaeocercoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archaeocercoides puchkovi" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="puchkovi">
<emphasis italics="true" pageId="0" pageNumber="105">Archaeocercoides puchkovi</emphasis>
<taxonomicName id="1C1F33893B9FDEFE0093463A1076A429" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Archaeocercoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archaeocercoides puchkovi" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="puchkovi">
<emphasis id="EF42DF9FEFD5819B80219FEA34FE06AA" italics="true" pageId="0" pageNumber="105">Archaeocercoides puchkovi</emphasis>
</taxonomicName>
Simutnik, 2022 from Rovno amber was fossilized near an undescribed crawler, see fig. 1C, D in
<bibRefCitation DOI="https://doi.org/10.3897/jhr.89.79180" author="Simutnik, SA" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="105" pagination="47 - 60" refId="B20" refString="Simutnik, SA, Perkovsky, EE, Khomych, MR, Vasilenko, DV, 2022a. Two new genera of Encyrtidae (Hymenoptera, Chalcidoidea) with reduced ovipositor sheaths. Journal of Hymenoptera Research 89: 47 - 60, DOI: https://doi.org/10.3897/jhr.89.79180" title="Two new genera of Encyrtidae (Hymenoptera, Chalcidoidea) with reduced ovipositor sheaths." url="https://doi.org/10.3897/jhr.89.79180" volume="89" year="2022 a">Simutnik et al. 2022a</bibRefCitation>
<bibRefCitation id="2F9ADAA6B2E447AACF2A453C9219EC75" DOI="https://doi.org/10.3897/jhr.89.79180" author="Simutnik, SA" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="105" pagination="47 - 60" refId="B20" refString="Simutnik, SA, Perkovsky, EE, Khomych, MR, Vasilenko, DV, 2022a. Two new genera of Encyrtidae (Hymenoptera, Chalcidoidea) with reduced ovipositor sheaths. Journal of Hymenoptera Research 89: 47 - 60, DOI: https://doi.org/10.3897/jhr.89.79180" title="Two new genera of Encyrtidae (Hymenoptera, Chalcidoidea) with reduced ovipositor sheaths." url="https://doi.org/10.3897/jhr.89.79180" volume="89" year="2022 a">Simutnik et al. 2022a</bibRefCitation>
). As the cerci advanced forward, Mt8 (or syntergum Mt8+Mt9) and then the previous terga began to separate into a dorsal part and lateral lobes (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7</figureCitation>
<figureCitation id="097CFA4BA719766F28C270971820BA0F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="105">
<paragraph id="0D34BE245ED8D295C95F93B81890ACCC" pageId="0" pageNumber="105">
The process of cercus advancement in both
<taxonomicName lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
<taxonomicName id="99339B207BF3A78C20D9D49CC15A570A" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Encyrtinae">Encyrtinae</taxonomicName>
and
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
<taxonomicName id="3A5C9B291762C1E1AA4031D904B7A396" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
occurred independently and in parallel, and it began precisely in the late Eocene, since the cerci still remain in the apical or subapical position in most encyrtids from European ambers. Cerci extremely advanced to the base of the metasoma, as in many extant taxa, have not been found in known Eocene
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
<taxonomicName id="ED4E1B81DA70FE47CC2E0A22291965CC" baseAuthorityName="Forster" baseAuthorityYear="1856" class="Insecta" family="Encyrtidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="family">Encyrtidae</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="105">
<paragraph id="28D52453F25DCB89A11EC4E4236DBCC8" pageId="0" pageNumber="105">
The elongate sclerotized strip (paratergite) in
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
<taxonomicName id="A923C0B23EE103505AE7ECC952C7DE27" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
apparently separated from the lateral margin of the syntergum when the latter became long enough and the cercal plates advanced significantly towards the base of the gaster. The true paratergite must directly maintain a connection to tergite IX (Mt8). This connection is traceable in many species of extant
<taxonomicName class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
<taxonomicName id="AE817ABE6637EB21B7D7DDC4D97E2A6B" class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="07FC72B6254A52D2D074A6D507C52230" italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
</taxonomicName>
around the side and anterior of the cercal plate (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7A, B</figureCitation>
<figureCitation id="4DA2F5F5597B76224932A27DFE556F66" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A, B Ericydnus sp., female A metasoma, dorsal (opo - outer plates of ovipositor, ptrg - paratergite) B apex of metasoma, lateral (spr - spiracle on the lateral lobe of the Mt 7, v 3 - ovipositor sheaths) C Electronoyesella antiqua gen. et sp. nov., holotype female, apex of metasoma, lateral (fc - filamentous connection). Scale bars: 0.2 mm." figureDoi="10.3897/jhr.94.94773.figure7" httpUri="https://binary.pensoft.net/fig/784453" pageId="0" pageNumber="105">7A, B</figureCitation>
) but not in any other genera where the connection, at most is at best tenuous (J. S. Noyes, pers. comm. 2022). What is possibly the paratergite of
<taxonomicName authorityName="Simutnik" authorityYear="2022" family="Encyrtidae" genus="Electronoyesella" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Electronoyesella</emphasis>
<taxonomicName id="712C8DEEF26BE7C26D2210187B55FF77" authorityName="Simutnik" authorityYear="2022" class="Insecta" family="Encyrtidae" genus="Electronoyesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Electronoyesella" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="CAD94C9C6EC07F67D7CABA561E3A831F" italics="true" pageId="0" pageNumber="105">Electronoyesella</emphasis>
</taxonomicName>
running anteriorly to the outside of the cercal plate is very similar to the paratergite of
<taxonomicName class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
<taxonomicName id="CDA2652412E409AA2D9705E950F0908B" class="Arachnida" family="Encyrtidae" genus="Ericydnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericydnus" order="Hymenoptera" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="BEE17D41237F312A0C051B6916B010CD" italics="true" pageId="0" pageNumber="105">Ericydnus</emphasis>
</taxonomicName>
and may provide morphological evidence for the existence of the subfamily
<taxonomicName authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
<taxonomicName id="BAD332CAB801DD61DDCA47AA79840C09" authorityName="Howard" authorityYear="1892" lsidName="" pageId="0" pageNumber="105" rank="subFamily" subFamily="Tetracneminae">Tetracneminae</taxonomicName>
in the late Eocene.
</paragraph>
</subSubSection>