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<treatment ID-GBIF-Taxon="159709676" LSID="urn:lsid:plazi:treatment:5804BA4F6A425552B3FD23F12459D9A2" httpUri="http://treatment.plazi.org/id/5804BA4F6A425552B3FD23F12459D9A2" lastPageNumber="43" pageId="0" pageNumber="43">
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<taxonomicName LSID="5804BA4F-6A42-5552-B3FD-23F12459D9A2" authority="Chou &amp; Lu, 1977" authorityName="Chou &amp; Lu" authorityYear="1977" class="Insecta" family="Ricaniidae" genus="Pochazia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pochazia shantungensis" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="shantungensis">Pochazia shantungensis (Chou &amp; Lu, 1977)</taxonomicName>
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<paragraph pageId="0" pageNumber="43">Description.</paragraph>
<paragraph pageId="0" pageNumber="43">
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<paragraph id="D3AEC7AA3E8AF5E7F1C4FB960A3C894B" pageId="0" pageNumber="43">Description.</paragraph>
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The mature spermatozoa of
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<emphasis italics="true" pageId="0" pageNumber="43">P. shantungensis</emphasis>
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are held together (totally 128 spermatozoa per spermatodesm) in the form of coiled sperm bundles in the seminal vesicles. Anterior ends of heads are embedded in a homogenous matrix that forms the spermatodesmata (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Fig. 1A, B, D</figureCitation>
<figureCitation id="E32483D7BB3F28E40B4330D63D50A2AF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Fig. 1A, B, D</figureCitation>
). The spermatozoon is long and filiform when it is separated from the bundles and exposed to a 0.9% saline solution (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Fig. 1C</figureCitation>
<figureCitation id="359DC8E1D395DE4F257F4F7E7F9634D4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Fig. 1C</figureCitation>
). It is approximately 127
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in mean length and has an elongate single head (about 16
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="4CD70542194CCDBDC1ACB30652D66E3C" originalValue="µm">µm</normalizedToken>
) and a conventional single flagellum (about 111
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).
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<emphasis bold="true" pageId="0" pageNumber="43">Figure 1.</emphasis>
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<emphasis id="5C141D7F448944FC4FE379335923AEDB" bold="true" pageId="0" pageNumber="43">Figure 1.</emphasis>
TEM and light micrographs of spermatozoa and spermatodesms of
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<emphasis italics="true" pageId="0" pageNumber="43">P. shantungensis</emphasis>
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.
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<emphasis id="97727A5C1E0CBEB68D6E9FB111450C35" bold="true" pageId="0" pageNumber="43">A, B</emphasis>
Light micrographs of spermatodesm and spermatozoa
<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
<emphasis id="3A69694EC2D90F6F238C7D423C6366AE" bold="true" pageId="0" pageNumber="43">C</emphasis>
light micrograph of a single spermatozoon with the head (
<emphasis bold="true" pageId="0" pageNumber="43">H</emphasis>
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, arrow) and wavy
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(
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<emphasis id="0D7E6E9F7C56C49856F2417FAFE535E7" bold="true" pageId="0" pageNumber="43">F</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="43">D, E</emphasis>
<emphasis id="AA5E4AB45B7F7372DC778A9D458563BE" bold="true" pageId="0" pageNumber="43">D, E</emphasis>
TEM micrographs of cross-sections of spermatozoa, showing the acrosome (
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<emphasis id="F62CCEA4028C431C006F2F60080923DC" bold="true" pageId="0" pageNumber="43">a</emphasis>
) and nucleus (
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<emphasis id="49A3C0388397DDB3D561ABB9A2E27FCA" bold="true" pageId="0" pageNumber="43">N</emphasis>
). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="06C1007DD09FC7E91CE6F64CA07CB73A" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="43">
<normalizedToken originalValue="AC">A-C</normalizedToken>
<emphasis id="1FC34C93142F170DFD03F7DA66262B50" bold="true" pageId="0" pageNumber="43">
<normalizedToken id="DC8416A6918FF00CA3C897E19E8E18A7" originalValue="AC">A-C</normalizedToken>
</emphasis>
); 2
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(
<emphasis bold="true" pageId="0" pageNumber="43">D</emphasis>
<emphasis id="D59DACE2595857524211B9935D3802C1" bold="true" pageId="0" pageNumber="43">D</emphasis>
); 0.5µm (
<emphasis bold="true" pageId="0" pageNumber="43">E</emphasis>
<emphasis id="907849E527358F09C55F76454B4C51D0" bold="true" pageId="0" pageNumber="43">E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="43">
<paragraph id="713391C2D466EF1D402185944E200B0C" pageId="0" pageNumber="43">
The head is formed by the nucleus and the acrosome. The acrosome has an irregular saccular acrosomal vesicle and a perforatorium, both located anterior to the nucleus (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Figs 1E</figureCitation>
<figureCitation id="FC59474AC90FBB6DF39603BBFF2A318F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Figs 1E</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">2A, B</figureCitation>
<figureCitation id="EAEFF8D6EF9FD49E628FA44FEB4D45FA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">2A, B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">3A, B</figureCitation>
<figureCitation id="9DA3AF50F9C03C442567A48F423CC733" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">3A, B</figureCitation>
). Between the base of the perforatorium and the anterior portion of the nucleus is a noticeable transition (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Figs 1E</figureCitation>
<figureCitation id="D1C3503A5DC47AB4B2A62D205A78F8D1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Figs 1E</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">2B</figureCitation>
<figureCitation id="8DA0AB2196619BB3B5DF663C924E3561" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">2B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
<figureCitation id="9CA110EE0D24D60AA0930B09397D974E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
3
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). The acrosome gradually invaginates posteriorly to form a subacrosomal space in which the anterior part of the elongated nucleus is inserted (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Figs 2B</figureCitation>
<figureCitation id="3F5E26CE4837D4CDE6C1C1966CC403FF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Figs 2B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">3C, D</figureCitation>
<figureCitation id="881B9143F19C976866E0FE598FBD8C68" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">3C, D</figureCitation>
). The nuclei, different in shape (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Figs 2B</figureCitation>
<figureCitation id="C8E0BE530950ECD1ED87F0186ECCBEC6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Figs 2B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
<figureCitation id="F21F301D3B65F9E908C33D40E84B5AA9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
3
<normalizedToken originalValue="BN">B-N</normalizedToken>
<normalizedToken id="6597E3B1A38D771FC9990E7A215FDD17" originalValue="BN">B-N</normalizedToken>
</figureCitation>
), are full of homogeneous condensed chromatin and are separated from each other by a cell membrane (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Fig. 1D</figureCitation>
<figureCitation id="4099C27B773D99AEAA45B8AEB6837A43" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. TEM and light micrographs of spermatozoa and spermatodesms of P. shantungensis. A, B Light micrographs of spermatodesm and spermatozoa C light micrograph of a single spermatozoon with the head (H, arrow) and wavy flagellum (F) D, E TEM micrographs of cross-sections of spermatozoa, showing the acrosome (a) and nucleus (N). Arrow shows head cluster, asterisk indicates the acrosome. Scale bars: 20 µm (A-C); 2 µm (D); 0.5 µm (E)." figureDoi="10.3897/zookeys.880.32810.figure1" httpUri="https://binary.pensoft.net/fig/346580" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure1">Fig. 1D</figureCitation>
). The ovoid nucleus measures approximately 0.94
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="4B34CDFE32E8547E098BAF6820985427" originalValue="µm">µm</normalizedToken>
in diameter in cross-section; in longitudinal-section, it turns into a cylinder-shape (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2A, B</figureCitation>
<figureCitation id="3E1C4E7DFE34F98AB493AE9426F6B77D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2A, B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Figure 2.</emphasis>
<caption id="6D841517BB6EEA70FE788D80B8044FD4" doi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" start="Figure 2" startId="F2">
<paragraph id="14BA65183277429974AE858B2334EDF9" pageId="0" pageNumber="43">
<emphasis id="C5001019A1DC9D4FC35AEDAA5961DE45" bold="true" pageId="0" pageNumber="43">Figure 2.</emphasis>
Longitudinal sections of spermatozoa of
<taxonomicName lsidName="P. shantungensis" pageId="0" pageNumber="43" rank="species" species="shantungensis">
<emphasis italics="true" pageId="0" pageNumber="43">P. shantungensis</emphasis>
<taxonomicName id="25F0E5F748E41E65CE53F5EBC488BCED" lsidName="P. shantungensis" pageId="0" pageNumber="43" rank="species" species="shantungensis">
<emphasis id="F1DF6D89AA5173FF5FC02C7C898B1F6F" italics="true" pageId="0" pageNumber="43">P. shantungensis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="43">A, B</emphasis>
<emphasis id="FB711FD27223AFBC38854FEAB3D1F049" bold="true" pageId="0" pageNumber="43">A, B</emphasis>
Spermatozoa, showing acrosome (
<emphasis bold="true" pageId="0" pageNumber="43">a</emphasis>
<emphasis id="D2304E765D5970F859CD42DCEEE24EEC" bold="true" pageId="0" pageNumber="43">a</emphasis>
), nucleus (
<emphasis bold="true" pageId="0" pageNumber="43">N</emphasis>
<emphasis id="FC554BF5271729ED248FD35C2F801834" bold="true" pageId="0" pageNumber="43">N</emphasis>
), arrow indicates connection area between acrosome and nucleus
<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
<emphasis id="1AB8939ACF4FD1A95DA39820C48248F9" bold="true" pageId="0" pageNumber="43">C</emphasis>
nucleus-flagellum transition, showing nucleus (
<emphasis bold="true" pageId="0" pageNumber="43">N</emphasis>
<emphasis id="B8292301F708BDE24CDEA491357A73BE" bold="true" pageId="0" pageNumber="43">N</emphasis>
), mitochondrial derivatives (
<emphasis bold="true" pageId="0" pageNumber="43">md</emphasis>
<emphasis id="C26F905D31E1369270B68FB40C626193" bold="true" pageId="0" pageNumber="43">md</emphasis>
), accessory body (
<emphasis bold="true" pageId="0" pageNumber="43">ab</emphasis>
<emphasis id="798D487C8C8628A66D70057A45F00CFF" bold="true" pageId="0" pageNumber="43">ab</emphasis>
), axoneme (
<emphasis bold="true" pageId="0" pageNumber="43">ax</emphasis>
<emphasis id="D2DD17C4FCFC07FB3C7CCE5D498790D0" bold="true" pageId="0" pageNumber="43">ax</emphasis>
), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct
<emphasis bold="true" pageId="0" pageNumber="43">D, E</emphasis>
<emphasis id="ADDB9C856BE0FE24C507283DA7FF45E5" bold="true" pageId="0" pageNumber="43">D, E</emphasis>
flagella of sperm, showing axoneme (
<emphasis bold="true" pageId="0" pageNumber="43">ax</emphasis>
<emphasis id="609EE457080AF8B472E2E2D1C6DFD454" bold="true" pageId="0" pageNumber="43">ax</emphasis>
), mitochondrial derivatives (
<emphasis bold="true" pageId="0" pageNumber="43">md</emphasis>
<emphasis id="126A4166A02006FF1536F1390BAC6D79" bold="true" pageId="0" pageNumber="43">md</emphasis>
) and cristae (arrow). Scale bars: 2
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="EADE8859E5CA2BF58689DEF0D1B4F9F4" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
<emphasis id="12D957D5D00DBC974AC0096F116FBDDA" bold="true" pageId="0" pageNumber="43">A</emphasis>
); 0.5
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="B74482D4F8F409F174E1DF2E445816E5" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="43">B, D</emphasis>
<emphasis id="DC7C7EAAEA74FAF6057479B4F09066EC" bold="true" pageId="0" pageNumber="43">B, D</emphasis>
); 0.1
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="3E258BC5CB3222DA4391733B6F4647F3" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="43">C, E</emphasis>
<emphasis id="B9FCD2E0A88D9A153BCD932DA7B0FA15" bold="true" pageId="0" pageNumber="43">C, E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="43">
<paragraph id="1508C69DD4F5E4E3B232E849421F7F8A" pageId="0" pageNumber="43">
The nucleus-flagellum transition region has a centriole and centriolar adjunct (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2B, C</figureCitation>
<figureCitation id="D02AF834D81AFC4EB27BD99DA1564630" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2B, C</figureCitation>
). The centriole starts near the terminal incurvation of the nucleus and terminates anterior of the axoneme (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2B, C</figureCitation>
<figureCitation id="C2E31428BC11D236C726EBBD0822F4F8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2B, C</figureCitation>
); it is parallel to the moderately electron-dense centriole adjunct (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2B</figureCitation>
<figureCitation id="133C5791C6DF6B7B3F033FD9A579CFC0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2B</figureCitation>
). The centriole adjunct contains dense granules between the accessory bodies and mitochondrial derivatives in longitudinal profile (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2B, C</figureCitation>
<figureCitation id="68B53D4F9D4D7616E28056BA058B2991" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2B, C</figureCitation>
); in cross-section the centriole adjunct arises near the end of the nucleus and terminates anterior to the mitochondrial derivatives (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
<figureCitation id="AE57D2CD0953868B9838C7419634EEB0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
Fig. 3
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<normalizedToken id="5336A8CBE122F8E2280DD860B2F9AA2C" originalValue="IK">I-K</normalizedToken>
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). The nucleus is wrapped by the centriolar adjunct (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Figs 2B, C</figureCitation>
<figureCitation id="23F8D1E3933958D33FB4D6C29CB8FC10" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Figs 2B, C</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
<figureCitation id="C8AB111E0C09AE6EA1DAE6A7DCAF2806" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
3
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). In several transverse profiles of spermatozoa, the posterior nucleus region overlaps the different regions of accessory bodies and mitochondrial derivatives (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
<figureCitation id="80B72CA8D7955655EDA12EA352A1791A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cross-sections of spermatozoa of P. shantungensis. A Acrosome, showing the dothideoid acrosome B-E serial cross-sections of head showing the dothideoid acrosome (a), the nucleus (N), and the plasma membrane (pm) F oval nucleus (N) G-N nucleus-flagellum transition region, showing the nucleus (N), mitochondrial derivatives (md), accessory bodies (ab), axoneme (ax). The asterisk indicates the centriolar adjunct (ca). Scale bars: 0.5 µm (B-D); 0.2 µm (A, K, L, N); 0.1 µm (E-J, M)." figureDoi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure3">
Fig. 3
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).
</paragraph>
<caption doi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Figure 3.</emphasis>
<caption id="9CFDCBC32EFB680061011B773DB03586" doi="10.3897/zookeys.880.32810.figure3" httpUri="https://binary.pensoft.net/fig/346582" pageId="0" pageNumber="43" start="Figure 3" startId="F3">
<paragraph id="D0B6D09B42CB434BBD4FE20664D5F43B" pageId="0" pageNumber="43">
<emphasis id="ABE2A712E6E916BD4F4C5A81BCD29BA5" bold="true" pageId="0" pageNumber="43">Figure 3.</emphasis>
Cross-sections of spermatozoa of
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<emphasis italics="true" pageId="0" pageNumber="43">P. shantungensis</emphasis>
<taxonomicName id="CC8639E3698CA9E91DD37A578CB0CDA5" lsidName="P. shantungensis" pageId="0" pageNumber="43" rank="species" species="shantungensis">
<emphasis id="71A5353F458C40C0E4E0C9F4ABA14C28" italics="true" pageId="0" pageNumber="43">P. shantungensis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
<emphasis id="3131FFFFEB370EE6E0D042EC3A1FD603" bold="true" pageId="0" pageNumber="43">A</emphasis>
Acrosome, showing the dothideoid acrosome
<emphasis bold="true" pageId="0" pageNumber="43">
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<emphasis id="9C5ED6C2C6A5A02058AEC29C1116B66B" bold="true" pageId="0" pageNumber="43">
<normalizedToken id="14B26D2AE50D1A1A22CCBD1BFBDC1CC1" originalValue="BE">B-E</normalizedToken>
</emphasis>
serial cross-sections of head showing the dothideoid acrosome (
<emphasis bold="true" pageId="0" pageNumber="43">a</emphasis>
<emphasis id="562B536A571BF0221E6C0C806F29F4ED" bold="true" pageId="0" pageNumber="43">a</emphasis>
), the nucleus (
<emphasis bold="true" pageId="0" pageNumber="43">N</emphasis>
<emphasis id="3DB572EA0FDEB977807DFC8D318EFE44" bold="true" pageId="0" pageNumber="43">N</emphasis>
), and the plasma membrane (
<emphasis bold="true" pageId="0" pageNumber="43">pm</emphasis>
<emphasis id="2C2228F101DE05156099EC5D823EAAB6" bold="true" pageId="0" pageNumber="43">pm</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="43">F</emphasis>
<emphasis id="C7D492F4F4C75FE66A8C7EFE256CF478" bold="true" pageId="0" pageNumber="43">F</emphasis>
oval nucleus (
<emphasis bold="true" pageId="0" pageNumber="43">N</emphasis>
<emphasis id="F1B23FA60246F9F48C2982A7D28F6F71" bold="true" pageId="0" pageNumber="43">N</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="43">
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<emphasis id="E9AF179B53DE04EB4A9155939D86399F" bold="true" pageId="0" pageNumber="43">
<normalizedToken id="7082098612EA1B4ED82E7E0E9AB3E365" originalValue="GN">G-N</normalizedToken>
</emphasis>
nucleus-flagellum transition region, showing the nucleus (
<emphasis bold="true" pageId="0" pageNumber="43">N</emphasis>
<emphasis id="17CD9EDE8DECE4F6253EDA92C29BECCD" bold="true" pageId="0" pageNumber="43">N</emphasis>
), mitochondrial derivatives (
<emphasis bold="true" pageId="0" pageNumber="43">md</emphasis>
<emphasis id="FF409AD064C66C211387511DFE66D371" bold="true" pageId="0" pageNumber="43">md</emphasis>
), accessory bodies (
<emphasis bold="true" pageId="0" pageNumber="43">ab</emphasis>
<emphasis id="416FA293D31365CF4D9353EBF8CC43BA" bold="true" pageId="0" pageNumber="43">ab</emphasis>
), axoneme (
<emphasis bold="true" pageId="0" pageNumber="43">ax</emphasis>
<emphasis id="BEFCC3BF6AB42CE8FDDBD86230A18D43" bold="true" pageId="0" pageNumber="43">ax</emphasis>
). The asterisk indicates the centriolar adjunct (
<emphasis bold="true" pageId="0" pageNumber="43">ca</emphasis>
<emphasis id="D511304843F1E4C82CCD485D43B556EE" bold="true" pageId="0" pageNumber="43">ca</emphasis>
). Scale bars: 0.5
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="9807F0D46A0B713A23797E852BD045FB" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="43">
<normalizedToken originalValue="BD">B-D</normalizedToken>
<emphasis id="6D8EDDBE20C03225341DB71A6542A3CF" bold="true" pageId="0" pageNumber="43">
<normalizedToken id="B050123E5AB3782D9DD2402A8B3D3C74" originalValue="BD">B-D</normalizedToken>
</emphasis>
); 0.2
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="07983581DA22A1E3AC494157D4FD5E2F" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="43">A, K, L, N</emphasis>
<emphasis id="18EA194F744CFE634BD1C15F5301F739" bold="true" pageId="0" pageNumber="43">A, K, L, N</emphasis>
); 0.1
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="B20C038FE0740114EB04F8C99E83045C" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="43">
<normalizedToken originalValue="EJ">E-J</normalizedToken>
<emphasis id="B0D25739211EE5144F34E0141F6C3950" bold="true" pageId="0" pageNumber="43">
<normalizedToken id="BC986F14F4738C712836EE6D38457D49" originalValue="EJ">E-J</normalizedToken>
, M
</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="43">
<paragraph id="12E6EF93EA89689AAC16D3E2B5F20EAE" pageId="0" pageNumber="43">
The flagellum region contains an axoneme, two mitochondrial derivatives and two accessory bodies (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">
<figureCitation id="FFCD70339B2FCDF4F299641F3AC470C7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">
Fig. 4
<normalizedToken originalValue="AD">A-D</normalizedToken>
<normalizedToken id="244F7D1F9352CE6634948A17817ADF8C" originalValue="AD">A-D</normalizedToken>
</figureCitation>
); they are parallel to each other throughout most of the length of the flagellum (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">
<figureCitation id="7096A359DDC38B305563474AFF6A270E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">
Fig. 2
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<normalizedToken id="414E894C5C5E63FF133FC02B563E4CFE" originalValue="BD">B-D</normalizedToken>
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). The axoneme arises from the centriole (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2C</figureCitation>
<figureCitation id="A2E086576CB5B95BA9BF2E3D6A9CE7A9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2C</figureCitation>
). It is composed of two innermost microtubules, nine outermost accessory microtubules, and nine doublets, showing the typical 9 + 9 + 2 microtubules arrangement in insects (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">Fig. 4D, G</figureCitation>
<figureCitation id="C8835874B6973FE76CB3B183498F4B2A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">Fig. 4D, G</figureCitation>
). The mitochondrial derivatives and accessory bodies are symmetrical in size and diameter in cross-section (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">
<figureCitation id="A5388899E0A104BF4F5E8EAF4C402F68" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">
Fig. 4
<normalizedToken originalValue="AD">A-D</normalizedToken>
<normalizedToken id="9C1764170B1AC2C09F52FB7AA6A59F08" originalValue="AD">A-D</normalizedToken>
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). Each mitochondrial derivative is made up of one serrated electron-dense area, one small oval electron-lucid portion and one mitochondrial cristae region (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">Fig. 4C</figureCitation>
<figureCitation id="A5B5F5D55F09C1B9194AA62D3225E039" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">Fig. 4C</figureCitation>
). In longitudinal-section, the mitochondrial derivatives are positioned lateral to the axoneme and are initiated near the extreme base of the centriole adjunct (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">
<figureCitation id="370DA4E0913A5A210B4A4940D50A2D8C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">
Fig. 2
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<normalizedToken id="34CE154D74533A60918182ECD37686BA" originalValue="BD">B-D</normalizedToken>
</figureCitation>
). The cristae are perpendicular to the longitudinal axis, bearing regular intervals (42 nm) between adjacent derivatives (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2E</figureCitation>
<figureCitation id="F7580F8922F6DD4DEBD34D3EE3176BC8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Longitudinal sections of spermatozoa of P. shantungensis. A, B Spermatozoa, showing acrosome (a), nucleus (N), arrow indicates connection area between acrosome and nucleus C nucleus-flagellum transition, showing nucleus (N), mitochondrial derivatives (md), accessory body (ab), axoneme (ax), arrow indicates centriole, triangular arrowhead indicates centriolar adjunct D, E flagella of sperm, showing axoneme (ax), mitochondrial derivatives (md) and cristae (arrow). Scale bars: 2 µm (A); 0.5 µm (B, D); 0.1 µm (C, E)." figureDoi="10.3897/zookeys.880.32810.figure2" httpUri="https://binary.pensoft.net/fig/346581" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure2">Fig. 2E</figureCitation>
). The accessory bodies are fishhook-shaped; they originate from the centriolar adjunct between the axoneme and mitochondrial derivatives (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">
<figureCitation id="9897731B3358A34D25E398F658EB9B8E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">
Fig. 4
<normalizedToken originalValue="BD">B-D</normalizedToken>
<normalizedToken id="1BCD9FBDED220C9EEC8720B2039824A0" originalValue="BD">B-D</normalizedToken>
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).
</paragraph>
<paragraph pageId="0" pageNumber="43">
<paragraph id="ED0FEF195CDB9D211ED00B1B190BD220" pageId="0" pageNumber="43">
Close to the posterior sperm tip, the axoneme becomes disorganized step by step, and the accessory bodies gradually taper to a cone-shape, while the mitochondrial derivatives disappear (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">Fig. 4E, F</figureCitation>
<figureCitation id="8EE4C2B882BC30AFA36121500CEC35C3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">Fig. 4E, F</figureCitation>
). At the terminal region of the flagellum, the doublet microtubules are the last to disappear (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">Fig. 4H</figureCitation>
<figureCitation id="E2E222A62889CE514C0A8A20DCAF58F4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cross-sections of the sperm flagellum of P. shantungensis. A-D Flagella, showing axoneme (ax), fishhook-shaped accessory bodies (ab), D-shaped mitochondrial derivatives (md), containing oval lucent region (1), serrated electron-dense region (2) and mitochondrial cristae region (3) E-F flagellum, mitochondrial derivatives slowly disappear, axonemes (ax) become disordered, accessory bodies (ab) become smaller G axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (am), nine doublet microtubules (dm) and two innermost central microtubules (cm) H Showing doublet microtubules finally disappearing. Scale bars: 2 µm (A); 0.5 µm (B); 0.1 µm (C-H)." figureDoi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" tableDoi="10.3897/zookeys.880.32810.figure4">Fig. 4H</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Figure 4.</emphasis>
<caption id="B535F1FE9751224DA9406A1AEEAF0EFB" doi="10.3897/zookeys.880.32810.figure4" httpUri="https://binary.pensoft.net/fig/346583" pageId="0" pageNumber="43" start="Figure 4" startId="F4">
<paragraph id="E27E21962AB619FD82692461BE27F797" pageId="0" pageNumber="43">
<emphasis id="F510C8A2B4435B89EFE61739C77942B0" bold="true" pageId="0" pageNumber="43">Figure 4.</emphasis>
Cross-sections of the sperm flagellum of
<taxonomicName lsidName="P. shantungensis" pageId="0" pageNumber="43" rank="species" species="shantungensis">
<emphasis italics="true" pageId="0" pageNumber="43">P. shantungensis</emphasis>
<taxonomicName id="E6EC6C2C582B12E8B5C29A39366EF654" lsidName="P. shantungensis" pageId="0" pageNumber="43" rank="species" species="shantungensis">
<emphasis id="8CE7DC7E83195CFEE5912A68EB2B0810" italics="true" pageId="0" pageNumber="43">P. shantungensis</emphasis>
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.
<emphasis bold="true" pageId="0" pageNumber="43">
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Flagella, showing axoneme (
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<emphasis id="F5F69B3E5E7DD9185E895F95480F527D" bold="true" pageId="0" pageNumber="43">ax</emphasis>
), fishhook-shaped accessory bodies (
<emphasis bold="true" pageId="0" pageNumber="43">ab</emphasis>
<emphasis id="2189C3BDEA86D8762A29DC73385A470A" bold="true" pageId="0" pageNumber="43">ab</emphasis>
), D-shaped mitochondrial derivatives (
<emphasis bold="true" pageId="0" pageNumber="43">md</emphasis>
<emphasis id="F8FB356F94AC30E9CE88DD47A1A3DAEF" bold="true" pageId="0" pageNumber="43">md</emphasis>
), containing oval lucent region (
<emphasis bold="true" pageId="0" pageNumber="43">1</emphasis>
<emphasis id="3F000D6A5801DDB21229102A6CE94AB7" bold="true" pageId="0" pageNumber="43">1</emphasis>
), serrated electron-dense region (
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<emphasis id="1B943B14847090293E5B233DDB5C4937" bold="true" pageId="0" pageNumber="43">2</emphasis>
) and mitochondrial cristae region (
<emphasis bold="true" pageId="0" pageNumber="43">3</emphasis>
<emphasis id="3540D6A9AA277FD9E09090FCA58EFFEA" bold="true" pageId="0" pageNumber="43">3</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="43">
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flagellum, mitochondrial derivatives slowly disappear, axonemes (
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<emphasis id="8E7B57819F65AEA7B6A7622DF57E71D8" bold="true" pageId="0" pageNumber="43">ax</emphasis>
) become disordered, accessory bodies (
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<emphasis id="D0E34251C71C05F8C2C83CCC4B44094E" bold="true" pageId="0" pageNumber="43">ab</emphasis>
) become smaller
<emphasis bold="true" pageId="0" pageNumber="43">G</emphasis>
<emphasis id="E066EE5EE72C6732964E65B60681C7BC" bold="true" pageId="0" pageNumber="43">G</emphasis>
axoneme, showing the typical 9 + 9 + 2 pattern, nine outermost accessory microtubules (
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<emphasis id="7E2E3BB0C634021C1EE310BEB26FEC3B" bold="true" pageId="0" pageNumber="43">am</emphasis>
), nine doublet microtubules (
<emphasis bold="true" pageId="0" pageNumber="43">dm</emphasis>
<emphasis id="3078C41AB13A1A6401933B2F79BCC542" bold="true" pageId="0" pageNumber="43">dm</emphasis>
) and two innermost central microtubules (
<emphasis bold="true" pageId="0" pageNumber="43">cm</emphasis>
<emphasis id="26F7F95B9E13C112CB4160B70992EF4B" bold="true" pageId="0" pageNumber="43">cm</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="43">H</emphasis>
<emphasis id="24ECB771E149EA08EF4CE9674F2076C3" bold="true" pageId="0" pageNumber="43">H</emphasis>
Showing doublet microtubules finally disappearing. Scale bars: 2
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(
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); 0.5
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(
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); 0.1
<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<emphasis id="C2ADDEC87881D438500D215A8C35D691" bold="true" pageId="0" pageNumber="43">
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</emphasis>
).
</paragraph>

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@ -1,93 +1,93 @@
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.220.96922" ID-Pensoft-Pub="1314-2003-220-83" ID-Pensoft-UUID="2DBED918C201533EBB75B147CD0BAE0E" ModsDocID="1314-2003-220-83" checkinTime="1677663952540" checkinUser="pensoft" docAuthor="Desjardins, Stuart D., Bailey, John P., Zhang, Baowei, Zhao, Kai &amp; Schwarzacher, Trude" docDate="2023" docId="583A4EAB4D5854D68DE57D616C7DBD41" docLanguage="en" docName="PhytoKeys 220: 83-108" docOrigin="PhytoKeys 220" docPubDate="2023-02-27" docSource="http://dx.doi.org/10.3897/phytokeys.220.96922" docTitle="Parogonum cynanchoides subsp. glabriusculum (A. J. Li) Desjardins &amp; J. P. Bailey, comb. et 2023, stat. nov." docType="treatment" docVersion="1" id="2DBED918C201533EBB75B147CD0BAE0E" lastPageNumber="83" masterDocId="2DBED918C201533EBB75B147CD0BAE0E" masterDocTitle="New insights into the phylogenetic relationships of Japanese knotweed (Reynoutria japonica) and allied taxa in subtribe Reynoutriinae (Polygonaceae)" masterLastPageNumber="108" masterPageNumber="83" pageNumber="83" updateTime="1677663952540" updateUser="pensoft">
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25: 104 (1998). Homotypic synonym.
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<taxonomicName authorityName="Linnaeus" authorityYear="1767" class="Insecta" family="Potamanthidae" genus="Ephemera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephemera luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">Ephemera luteus</taxonomicName>
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<taxonomicName id="100AD8AD3E8D84948780C671DA07FC37" authorityName="Linnaeus" authorityYear="1767" class="Insecta" family="Potamanthidae" genus="Ephemera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephemera luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">Ephemera luteus</taxonomicName>
Linnaeus, 1767: 906. Type: England.
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Ephemera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephemera reticulata" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="reticulata">Ephemera reticulata</taxonomicName>
<paragraph id="14CCD0E469131E3D793EE8AC628513C4" pageId="0" pageNumber="193">
<taxonomicName id="69446FD5506F2DDC7BF8CB6FEFD85B8A" class="Insecta" family="Potamanthidae" genus="Ephemera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephemera reticulata" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="reticulata">Ephemera reticulata</taxonomicName>
Fourcroy, 1785: 351. Synonymized by
<bibRefCitation author="Eaton, AE" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B6" refString="Eaton, AE, 1871. A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." title="A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." year="1871">Eaton (1871</bibRefCitation>
<bibRefCitation id="A0DD89AFF98E862D37A91D5DC8E5A53F" author="Eaton, AE" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B6" refString="Eaton, AE, 1871. A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." title="A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." year="1871">Eaton (1871</bibRefCitation>
: 76).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis mellea" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="mellea">Baetis mellea</taxonomicName>
<paragraph id="60F9DD2EDF66C30045EEC5C949977381" pageId="0" pageNumber="193">
<taxonomicName id="37C3B5FA08906B0FCCF3D8DBB6238752" class="Insecta" family="Potamanthidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis mellea" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="mellea">Baetis mellea</taxonomicName>
Curtis, 1834: 121. Types: subimago. Synonymized by
<bibRefCitation author="Eaton, AE" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B6" refString="Eaton, AE, 1871. A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." title="A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." year="1871">Eaton (1871</bibRefCitation>
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: 76).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis marginalis" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="marginalis">Baetis marginalis</taxonomicName>
<paragraph id="87F0B020D5A02F435DF55B6A2F0BF49B" pageId="0" pageNumber="193">
<taxonomicName id="97A6B20AB80B570EB4188C6F768DB0DD" class="Insecta" family="Potamanthidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis marginalis" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="marginalis">Baetis marginalis</taxonomicName>
Burmeister, 1839: 801. Types: male and female. Synonymized by
<bibRefCitation author="Eaton, AE" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B6" refString="Eaton, AE, 1871. A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." title="A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." year="1871">Eaton (1871</bibRefCitation>
<bibRefCitation id="3F6092B547A97C47F8D0CC5E792BEA00" author="Eaton, AE" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B6" refString="Eaton, AE, 1871. A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." title="A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." year="1871">Eaton (1871</bibRefCitation>
: 76).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Ephemera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephemera flavicans" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="flavicans">Ephemera flavicans</taxonomicName>
<paragraph id="5512A64310F79C9A24955AAD24FF3F78" pageId="0" pageNumber="193">
<taxonomicName id="2108CF7FB79579001F9E32309044A34D" class="Insecta" family="Potamanthidae" genus="Ephemera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephemera flavicans" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="flavicans">Ephemera flavicans</taxonomicName>
Rambur, 1842: 296. Types: male and female, from Paris, France. Synonymized by
<bibRefCitation author="Eaton, AE" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B6" refString="Eaton, AE, 1871. A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." title="A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." year="1871">Eaton (1871</bibRefCitation>
<bibRefCitation id="E85110204440EC4ADD69E33ED31DEE92" author="Eaton, AE" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B6" refString="Eaton, AE, 1871. A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." title="A monograph on the Ephemeridae. Transactions of the Entomological Society of London, 164 pp." year="1871">Eaton (1871</bibRefCitation>
: 76).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Ephemera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephemera chlorotica" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="chlorotica">Ephemera chlorotica</taxonomicName>
<paragraph id="ECE49D13B37B6ECC539FA8C508F60D30" pageId="0" pageNumber="193">
<taxonomicName id="32CB2A1AE15A24C7B811B2A14F3E2F0F" class="Insecta" family="Potamanthidae" genus="Ephemera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ephemera chlorotica" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="chlorotica">Ephemera chlorotica</taxonomicName>
Rambur, 1842: 296. Types: male and female subimagoes, from Paris, France. Synonymized by
<bibRefCitation author="Walker, F" journalOrPublisher=", Paris: Kessmann et Cherbuliez edit., aussi a Geneve" pageId="0" pageNumber="193" refId="B21" refString="Walker, F, 1853. List of the specimens of neuropterous insects in the collection of the British Museum, Part III (Termitidae - Ephemeridae): 533-585." title="List of the specimens of neuropterous insects in the collection of the British Museum, Part III (Termitidae - Ephemeridae): 533 - 585." year="1853">Walker (1853</bibRefCitation>
<bibRefCitation id="83925E399A583E1C384F5D3727518C8D" author="Walker, F" journalOrPublisher=", Paris: Kessmann et Cherbuliez edit., aussi a Geneve" pageId="0" pageNumber="193" refId="B21" refString="Walker, F, 1853. List of the specimens of neuropterous insects in the collection of the British Museum, Part III (Termitidae - Ephemeridae): 533-585." title="List of the specimens of neuropterous insects in the collection of the British Museum, Part III (Termitidae - Ephemeridae): 533 - 585." year="1853">Walker (1853</bibRefCitation>
: 539).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">Potamanthus luteus</taxonomicName>
<paragraph id="4B83A9842AA2C5036F60008E7C54FE7F" pageId="0" pageNumber="193">
<taxonomicName id="1A0641266BB43798B440030CBA85B209" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">Potamanthus luteus</taxonomicName>
(Linnaeus):
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.48625" author="Pictet, FJ" journalOrPublisher=", Paris: Kessmann et Cherbuliez edit., aussi a Geneve" pageId="0" pageNumber="193" refId="B18" refString="Pictet, FJ, 1843. Histoire naturelle generale et particulaire des Insectes Nevropteres-Famille des Ephemerines. Bailliere edit., Paris: Kessmann et Cherbuliez edit., aussi a Geneve, DOI: https://doi.org/10.5962/bhl.title.48625" title="Histoire naturelle generale et particulaire des Insectes Nevropteres-Famille des Ephemerines. Bailliere edit." url="https://doi.org/10.5962/bhl.title.48625" year="1843">Pictet 1843</bibRefCitation>
<bibRefCitation id="8EECFF9A590FEBC0E44F5C195BCD36E2" DOI="https://doi.org/10.5962/bhl.title.48625" author="Pictet, FJ" journalOrPublisher=", Paris: Kessmann et Cherbuliez edit., aussi a Geneve" pageId="0" pageNumber="193" refId="B18" refString="Pictet, FJ, 1843. Histoire naturelle generale et particulaire des Insectes Nevropteres-Famille des Ephemerines. Bailliere edit., Paris: Kessmann et Cherbuliez edit., aussi a Geneve, DOI: https://doi.org/10.5962/bhl.title.48625" title="Histoire naturelle generale et particulaire des Insectes Nevropteres-Famille des Ephemerines. Bailliere edit." url="https://doi.org/10.5962/bhl.title.48625" year="1843">Pictet 1843</bibRefCitation>
: 205; Eaton 1884: 79.
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus ferreri" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="ferreri">Potamanthus ferreri</taxonomicName>
<paragraph id="952E942F0B842D11303E5ECF04E32F41" pageId="0" pageNumber="193">
<taxonomicName id="4469B4A6DA3B366E47EF6E4C4C498EC3" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus ferreri" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="ferreri">Potamanthus ferreri</taxonomicName>
Pictet, 1843: 203. Types: male, from Italy. Synonymized by
<bibRefCitation author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991</bibRefCitation>
<bibRefCitation id="3CF61CD9FDBF09AFB3FCF0BEC907E211" author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991</bibRefCitation>
: 51).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Eucharidis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eucharidis reaumurii" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="reaumurii">Eucharidis reaumurii</taxonomicName>
<paragraph id="6BCF04E7E2B0A03E19A6DAE080DF731D" pageId="0" pageNumber="193">
<taxonomicName id="63B9C8541064B2158C120E1DD89DBF62" class="Insecta" family="Potamanthidae" genus="Eucharidis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eucharidis reaumurii" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="reaumurii">Eucharidis reaumurii</taxonomicName>
Joly &amp; Joly, 1876: 314. Types: nymph. Synonymized by Eaton (1884: 79).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">Potamanthus</taxonomicName>
<paragraph id="A967C189573B4A16066D38B3488A906D" pageId="0" pageNumber="193">
<taxonomicName id="8CA8B27427DA6E4B4A026CCAA873391D" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">Potamanthus</taxonomicName>
na Imanishi, 1940: 180, fig. 2 (nymph). Synonymized by
<bibRefCitation author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991</bibRefCitation>
<bibRefCitation id="8E34BB2CAE4C3A519163F1B3A3E733EC" author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991</bibRefCitation>
: 54).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">Potamanthus</taxonomicName>
<paragraph id="C5BBCD43A037CE6AA9A1D25B689BB45F" pageId="0" pageNumber="193">
<taxonomicName id="D08EAFE3532F0B00B89BBA3F33D7C47B" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">Potamanthus</taxonomicName>
naa Imanishi, 1940: 181 (nymph). Synonymized by
<bibRefCitation author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991</bibRefCitation>
<bibRefCitation id="BD3260B9E1B611F223FD519B385CE136" author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991</bibRefCitation>
: 54).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">Potamanthus luteus</taxonomicName>
<paragraph id="5A352DD92C100CE03CA3FF663AD43D76" pageId="0" pageNumber="193">
<taxonomicName id="778625C3CC5B1D0273B4EB1F18F4D46F" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">Potamanthus luteus</taxonomicName>
:
<bibRefCitation author="Wu, XY" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="193" pagination=": 336" refId="B22" refString="Wu, XY, 1987a. A new generic record of the Chinese Potamanthidae - Potamanthus Pictet. Acta Zootaxonomica Sinica 12 (3): 336" title="A new generic record of the Chinese Potamanthidae - Potamanthus Pictet." volume="12" year="1987 a">Wu 1987a</bibRefCitation>
<bibRefCitation id="28B7813F0E22DC94958E8227186676E9" author="Wu, XY" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="193" pagination=": 336" refId="B22" refString="Wu, XY, 1987a. A new generic record of the Chinese Potamanthidae - Potamanthus Pictet. Acta Zootaxonomica Sinica 12 (3): 336" title="A new generic record of the Chinese Potamanthidae - Potamanthus Pictet." volume="12" year="1987 a">Wu 1987a</bibRefCitation>
: 336 (female, first record from China);
<bibRefCitation author="You, DS" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="193" refId="B25" refString="You, DS, Gui, H, 1995. Ephemeroptera. Science Press, Beijing" title="Ephemeroptera." year="1995">You and Gui 1995</bibRefCitation>
<bibRefCitation id="72A9083BA8E9A5E560AEE04A258DACDC" author="You, DS" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="193" refId="B25" refString="You, DS, Gui, H, 1995. Ephemeroptera. Science Press, Beijing" title="Ephemeroptera." year="1995">You and Gui 1995</bibRefCitation>
: 115, fig. 122 (male);
<bibRefCitation DOI="https://doi.org/10.1163/9789004260887" author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B3" refString="Bauernfeind, E, Soldan, T, 2012. The Mayflies of Europe (Ephemeroptera). Apollo Books, Ollerup, DOI: https://doi.org/10.1163/9789004260887" title="The Mayflies of Europe (Ephemeroptera)." url="https://doi.org/10.1163/9789004260887" year="2012">
<bibRefCitation id="CCAC561C67141ECACC8550E3524D868F" DOI="https://doi.org/10.1163/9789004260887" author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B3" refString="Bauernfeind, E, Soldan, T, 2012. The Mayflies of Europe (Ephemeroptera). Apollo Books, Ollerup, DOI: https://doi.org/10.1163/9789004260887" title="The Mayflies of Europe (Ephemeroptera)." url="https://doi.org/10.1163/9789004260887" year="2012">
Bauernfeind and
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
<normalizedToken id="ADC091C8A39AABC46C06F25129092D12" originalValue="Soldán">Soldan</normalizedToken>
2012
</bibRefCitation>
: 634 (adult, nymph, egg).
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus (Patamanthus) luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subGenus="Patamanthus" subSpecies="oriens">Potamanthus (Patamanthus) luteus oriens</taxonomicName>
<paragraph id="8A0BBC6690B60788C257045FE5474370" pageId="0" pageNumber="193">
<taxonomicName id="A4284FCAB69D634966E8927E6DF3E97C" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus (Patamanthus) luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subGenus="Patamanthus" subSpecies="oriens">Potamanthus (Patamanthus) luteus oriens</taxonomicName>
:
<bibRefCitation author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty 1991</bibRefCitation>
<bibRefCitation id="FCCABC325200A2684E43217910C7DD03" author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty 1991</bibRefCitation>
: 54, fig. 4, 125 (subspecies established);
<bibRefCitation author="Bae, YJ" editor="Landolt, P" journalOrPublisher="Mauron + Tinguely &amp; Lachat SA, Moncor, Fribourg, Switzerland" pageId="0" pageNumber="193" pagination="405 - 417" refId="B1" refString="Bae, YJ, 1997. A historical review of Ephemeroptera systematics in northeast Asia. In: Landolt, P, Sartori, M, Eds., Ephemeroptera &amp; Plecoptera: Biology - Ecology - Systematics. Mauron + Tinguely &amp; Lachat SA, Moncor, Fribourg, Switzerland: 405 - 417" title="A historical review of Ephemeroptera systematics in northeast Asia." volumeTitle="Ephemeroptera &amp; Plecoptera: Biology - Ecology - Systematics." year="1997">Bae 1997</bibRefCitation>
<bibRefCitation id="C2E41CB6DA4368E7792DB1FACC750363" author="Bae, YJ" editor="Landolt, P" journalOrPublisher="Mauron + Tinguely &amp; Lachat SA, Moncor, Fribourg, Switzerland" pageId="0" pageNumber="193" pagination="405 - 417" refId="B1" refString="Bae, YJ, 1997. A historical review of Ephemeroptera systematics in northeast Asia. In: Landolt, P, Sartori, M, Eds., Ephemeroptera &amp; Plecoptera: Biology - Ecology - Systematics. Mauron + Tinguely &amp; Lachat SA, Moncor, Fribourg, Switzerland: 405 - 417" title="A historical review of Ephemeroptera systematics in northeast Asia." volumeTitle="Ephemeroptera &amp; Plecoptera: Biology - Ecology - Systematics." year="1997">Bae 1997</bibRefCitation>
: 408;
<bibRefCitation author="Zhou, CF" journalOrPublisher="BSA)," pageId="0" pageNumber="193" pagination="167 - 225" refId="B26" refString="Zhou, CF, 2013. A species list of Chinese mayflies (Insecta: Ephemeroptera). Inland Waters Biology, (Proc. 1st Symp. BSA), 6: 167 - 225" title="A species list of Chinese mayflies (Insecta: Ephemeroptera). Inland Waters Biology, (Proc. 1 st Symp." volume="6" year="2013">Zhou 2013</bibRefCitation>
<bibRefCitation id="646B731D7C1333FBCC16C52FE76852E2" author="Zhou, CF" journalOrPublisher="BSA)," pageId="0" pageNumber="193" pagination="167 - 225" refId="B26" refString="Zhou, CF, 2013. A species list of Chinese mayflies (Insecta: Ephemeroptera). Inland Waters Biology, (Proc. 1st Symp. BSA), 6: 167 - 225" title="A species list of Chinese mayflies (Insecta: Ephemeroptera). Inland Waters Biology, (Proc. 1 st Symp." volume="6" year="2013">Zhou 2013</bibRefCitation>
: 202;
<bibRefCitation author="Zhou, CF" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="193" refId="B27" refString="Zhou, CF, Su, CR, Gui, H, 2015. Outline of Chinese mayflies. Science Press, Beijing" title="Outline of Chinese mayflies." year="2015">Zhou et al. 2015</bibRefCitation>
<bibRefCitation id="6CFB0B76EA094482D04082ECABD568EA" author="Zhou, CF" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="193" refId="B27" refString="Zhou, CF, Su, CR, Gui, H, 2015. Outline of Chinese mayflies. Science Press, Beijing" title="Outline of Chinese mayflies." year="2015">Zhou et al. 2015</bibRefCitation>
: 252.
</paragraph>
<paragraph pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subSpecies="oriens">Potamanthus luteus oriens</taxonomicName>
<paragraph id="DF65CADB53ADFA6451E8F7CDCA3D1D22" pageId="0" pageNumber="193">
<taxonomicName id="A91518D08D343FA02D3C3B45D5BDC1ED" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subSpecies="oriens">Potamanthus luteus oriens</taxonomicName>
:
<bibRefCitation author="Quan, YT" journalOrPublisher=", Paris: Kessmann et Cherbuliez edit., aussi a Geneve" pageId="0" pageNumber="193" refId="B19" refString="Quan, YT, Bae, YJ, Jung, JC, Lee, JW, 2002. Ephemeroptera (Insecta) fauna of Northeast China. Insecta Koreana 19(3, 4): 241-269." title="Ephemeroptera (Insecta) fauna of Northeast China. Insecta Koreana 19 (3, 4): 241 - 269." year="2002">Quan et al. 2002</bibRefCitation>
<bibRefCitation id="1442E279FE160FF6256754715CA4BEAE" author="Quan, YT" journalOrPublisher=", Paris: Kessmann et Cherbuliez edit., aussi a Geneve" pageId="0" pageNumber="193" refId="B19" refString="Quan, YT, Bae, YJ, Jung, JC, Lee, JW, 2002. Ephemeroptera (Insecta) fauna of Northeast China. Insecta Koreana 19(3, 4): 241-269." title="Ephemeroptera (Insecta) fauna of Northeast China. Insecta Koreana 19 (3, 4): 241 - 269." year="2002">Quan et al. 2002</bibRefCitation>
: 257.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="distribution">
<paragraph pageId="0" pageNumber="193">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="193">China (Heilongjiang and Jilin Province); Palearctic and Oriental. From England east through Europe and Asia Minor, south to North Africa.</paragraph>
<subSubSection id="801197DBBBAABFFEBBC57DDDCF53356F" pageId="0" pageNumber="193" type="distribution">
<paragraph id="75158817FAFA91B4B89EEBB86DD11C31" pageId="0" pageNumber="193">Distribution.</paragraph>
<paragraph id="E11AACF5673D97A3AC14172C4AC65CED" pageId="0" pageNumber="193">China (Heilongjiang and Jilin Province); Palearctic and Oriental. From England east through Europe and Asia Minor, south to North Africa.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="description">
<paragraph pageId="0" pageNumber="193">Description.</paragraph>
<paragraph pageId="0" pageNumber="193">
<subSubSection id="6ACDBE0513B2D82E621FC970B3399FFD" pageId="0" pageNumber="193" type="description">
<paragraph id="0BD1746833560A33AB5934C842AEF824" pageId="0" pageNumber="193">Description.</paragraph>
<paragraph id="B68403AC5D36B348E53C31AEC809CBA8" pageId="0" pageNumber="193">
see
<bibRefCitation author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991)</bibRefCitation>
<bibRefCitation id="49FD4BE85F4B80CFDB7AFD08F938ED64" author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991)</bibRefCitation>
or
<bibRefCitation DOI="https://doi.org/10.1163/9789004260887" author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B3" refString="Bauernfeind, E, Soldan, T, 2012. The Mayflies of Europe (Ephemeroptera). Apollo Books, Ollerup, DOI: https://doi.org/10.1163/9789004260887" title="The Mayflies of Europe (Ephemeroptera)." url="https://doi.org/10.1163/9789004260887" year="2012">
<bibRefCitation id="1723E46E3BE1E2586071E375F1E899CB" DOI="https://doi.org/10.1163/9789004260887" author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B3" refString="Bauernfeind, E, Soldan, T, 2012. The Mayflies of Europe (Ephemeroptera). Apollo Books, Ollerup, DOI: https://doi.org/10.1163/9789004260887" title="The Mayflies of Europe (Ephemeroptera)." url="https://doi.org/10.1163/9789004260887" year="2012">
Bauernfeind and
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
<normalizedToken id="54FD0B1D60B65D9B0222ADCF7376BAE2" originalValue="Soldán">Soldan</normalizedToken>
(2012)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="diagnosis">
<paragraph pageId="0" pageNumber="193">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="193">
<subSubSection id="01DEE1DA9012811106D4FCECFFBDB55E" pageId="0" pageNumber="193" type="diagnosis">
<paragraph id="D9C3254C4BDB2D3BBB1D7B434CE33B0C" pageId="0" pageNumber="193">Diagnosis.</paragraph>
<paragraph id="DDC8160D56B83FD4975315658FDF4273" pageId="0" pageNumber="193">
see diagnosis of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="P. huoshanensis" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="huoshanensis">
<emphasis italics="true" pageId="0" pageNumber="193">P. huoshanensis</emphasis>
<taxonomicName id="5D0EC46E8BC2B10AC7107DC8A3C6A25E" class="Insecta" kingdom="Animalia" lsidName="P. huoshanensis" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="huoshanensis">
<emphasis id="7ECCE1FC75EA3C09303CC84405FC66FE" italics="true" pageId="0" pageNumber="193">P. huoshanensis</emphasis>
</taxonomicName>
. Males of this species can be identified by the more distinct color of the wings and penial lobes (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male head, female head, forewing and hindwing of two Potamanthus species: A, B, E, F P. huoshanensis and C, D, G, H P. luteus." figureDoi="10.3897/zookeys.1125.89219.figure5" httpUri="https://binary.pensoft.net/fig/762479" pageId="0" pageNumber="193">5E-H</figureCitation>
<figureCitation id="EA7D91F035E993138B6146782922F7CE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male head, female head, forewing and hindwing of two Potamanthus species: A, B, E, F P. huoshanensis and C, D, G, H P. luteus." figureDoi="10.3897/zookeys.1125.89219.figure5" httpUri="https://binary.pensoft.net/fig/762479" pageId="0" pageNumber="193">5E-H</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia (ventral and dorsal view) and penis (ventral, dorsal and lateral view) of two Potamanthus species: A-E P. huoshanensis and F-J P. luteus." figureDoi="10.3897/zookeys.1125.89219.figure6" httpUri="https://binary.pensoft.net/fig/762480" pageId="0" pageNumber="193">6</figureCitation>
<figureCitation id="BA1379CC46DBA8C33BD956D095B9A493" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia (ventral and dorsal view) and penis (ventral, dorsal and lateral view) of two Potamanthus species: A-E P. huoshanensis and F-J P. luteus." figureDoi="10.3897/zookeys.1125.89219.figure6" httpUri="https://binary.pensoft.net/fig/762480" pageId="0" pageNumber="193">6</figureCitation>
) and the foretibiae longer than the tarsi (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male and female adult morphology of two Potamanthus species: A, B P. huoshanensis and C, D P. luteus." figureDoi="10.3897/zookeys.1125.89219.figure4" httpUri="https://binary.pensoft.net/fig/762478" pageId="0" pageNumber="193">4A, C</figureCitation>
<figureCitation id="D77AB056991951E87447BC81B1EE5FC2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male and female adult morphology of two Potamanthus species: A, B P. huoshanensis and C, D P. luteus." figureDoi="10.3897/zookeys.1125.89219.figure4" httpUri="https://binary.pensoft.net/fig/762478" pageId="0" pageNumber="193">4A, C</figureCitation>
). The nymphs can be distinguished by the slightly larger mandibular tusks, longer foretibiae (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female head, thorax, foreleg claw, foreleg, midleg and hindleg of nymph of two Potamanthus species: A-D, I-K P. huoshanensis and E-H, L-N P. luteus." figureDoi="10.3897/zookeys.1125.89219.figure2" httpUri="https://binary.pensoft.net/fig/762476" pageId="0" pageNumber="193">2I, L</figureCitation>
<figureCitation id="4D20B4C692177C6AD4BD091A8CFC4E84" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female head, thorax, foreleg claw, foreleg, midleg and hindleg of nymph of two Potamanthus species: A-D, I-K P. huoshanensis and E-H, L-N P. luteus." figureDoi="10.3897/zookeys.1125.89219.figure2" httpUri="https://binary.pensoft.net/fig/762476" pageId="0" pageNumber="193">2I, L</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labrum, hypopharynx, left mandible, right mandible, maxilla and labium of nymph mouthparts of two Potamanthus species: A, C, E, G, I, K, M P. huoshanensis and B, D, F, H, J, L, N P. luteus. dd: distal dentisetae; dp: proximal dentisetae." figureDoi="10.3897/zookeys.1125.89219.figure3" httpUri="https://binary.pensoft.net/fig/762477" pageId="0" pageNumber="193">3E-H</figureCitation>
<figureCitation id="DF2B372FA130F7436DF48ED041EC551A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labrum, hypopharynx, left mandible, right mandible, maxilla and labium of nymph mouthparts of two Potamanthus species: A, C, E, G, I, K, M P. huoshanensis and B, D, F, H, J, L, N P. luteus. dd: distal dentisetae; dp: proximal dentisetae." figureDoi="10.3897/zookeys.1125.89219.figure3" httpUri="https://binary.pensoft.net/fig/762477" pageId="0" pageNumber="193">3E-H</figureCitation>
) and apical segment of the maxillary palpi (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labrum, hypopharynx, left mandible, right mandible, maxilla and labium of nymph mouthparts of two Potamanthus species: A, C, E, G, I, K, M P. huoshanensis and B, D, F, H, J, L, N P. luteus. dd: distal dentisetae; dp: proximal dentisetae." figureDoi="10.3897/zookeys.1125.89219.figure3" httpUri="https://binary.pensoft.net/fig/762477" pageId="0" pageNumber="193">3K, L</figureCitation>
<figureCitation id="00CB9CF4EF097810B867F7F334F18633" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Labrum, hypopharynx, left mandible, right mandible, maxilla and labium of nymph mouthparts of two Potamanthus species: A, C, E, G, I, K, M P. huoshanensis and B, D, F, H, J, L, N P. luteus. dd: distal dentisetae; dp: proximal dentisetae." figureDoi="10.3897/zookeys.1125.89219.figure3" httpUri="https://binary.pensoft.net/fig/762477" pageId="0" pageNumber="193">3K, L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="remarks">
<paragraph pageId="0" pageNumber="193">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="193">
<bibRefCitation author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991)</bibRefCitation>
<subSubSection id="9C6042F685AC4BE764047539AFA112EA" pageId="0" pageNumber="193" type="remarks">
<paragraph id="B4FE7592574820A39E1993B47E1A8259" pageId="0" pageNumber="193">Remarks.</paragraph>
<paragraph id="681D31ECA3A70115AF6D5A75AD4E38C5" pageId="0" pageNumber="193">
<bibRefCitation id="1E9107235ED0B3ED60CA872890F77FD6" author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991)</bibRefCitation>
mentioned that the nymphs of the subspecies
<taxonomicName class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subSpecies="oriens">
<emphasis italics="true" pageId="0" pageNumber="193">Potamanthus luteus oriens</emphasis>
<taxonomicName id="F507DBD27D9D8FF63E5E1A69FC3F27A1" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subSpecies="oriens">
<emphasis id="CC18D95C65612A436851A042C5FB5260" italics="true" pageId="0" pageNumber="193">Potamanthus luteus oriens</emphasis>
</taxonomicName>
have very pointed anterolateral projections of the pronotal and vestigial spine-row on the forefemora. In our material, the former character is distinct, and the transverse spine-row was not recognizable, which is consistent with the description of European
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="P. luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">
<emphasis italics="true" pageId="0" pageNumber="193">P. luteus</emphasis>
<taxonomicName id="56EAC8246E05BE6D9425EB2BD7B974F7" class="Insecta" kingdom="Animalia" lsidName="P. luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="185E8C4B19DC58849F4B860C2E754778" italics="true" pageId="0" pageNumber="193">P. luteus</emphasis>
</taxonomicName>
provided by
<bibRefCitation DOI="https://doi.org/10.1163/9789004260887" author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B3" refString="Bauernfeind, E, Soldan, T, 2012. The Mayflies of Europe (Ephemeroptera). Apollo Books, Ollerup, DOI: https://doi.org/10.1163/9789004260887" title="The Mayflies of Europe (Ephemeroptera)." url="https://doi.org/10.1163/9789004260887" year="2012">
<bibRefCitation id="33D09F2A6618F5657575343256B9CD3E" DOI="https://doi.org/10.1163/9789004260887" author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="193" refId="B3" refString="Bauernfeind, E, Soldan, T, 2012. The Mayflies of Europe (Ephemeroptera). Apollo Books, Ollerup, DOI: https://doi.org/10.1163/9789004260887" title="The Mayflies of Europe (Ephemeroptera)." url="https://doi.org/10.1163/9789004260887" year="2012">
Bauernfeind and
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<normalizedToken id="8F3B0D73C1F488C95CE3BBFC4BDD01B8" originalValue="Soldán">Soldan</normalizedToken>
(2012)
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. However, we do not know whether this variation is just at the population level or representative of different subspecies or geographical populations, because we have no material from abroad for comparison.
</paragraph>
<paragraph pageId="0" pageNumber="193">
<paragraph id="3B8BAF808D2330F3A0161B4B743BF781" pageId="0" pageNumber="193">
In the present comparison and photos, we can see clearly that
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<emphasis italics="true" pageId="0" pageNumber="193">P. huoshanensis</emphasis>
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<emphasis id="3EEED13159C4F1B87CC52F9C0ED8402C" italics="true" pageId="0" pageNumber="193">P. huoshanensis</emphasis>
</taxonomicName>
and
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<emphasis italics="true" pageId="0" pageNumber="193">P. luteus oriens</emphasis>
<taxonomicName id="860AB3DCAE9A9A2ABDBB0737C17E29F9" class="Insecta" kingdom="Animalia" lsidName="P. luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subSpecies="oriens">
<emphasis id="9D9228DE3D0C3C1CDCCC652DD492D78B" italics="true" pageId="0" pageNumber="193">P. luteus oriens</emphasis>
</taxonomicName>
are extremely similar in both nymphal and imaginal structures. The differences between them are very slight. Therefore, it is not surprising that
<bibRefCitation author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991)</bibRefCitation>
<bibRefCitation id="561E856BA73FC2C08F7EC68D2717582A" author="Bae, YJ" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="193" pagination="1 - 143" refId="B2" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3-4): 1 - 143" title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera)." volume="117" year="1991">Bae and McCafferty (1991)</bibRefCitation>
recognized Japanese materials of
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<emphasis italics="true" pageId="0" pageNumber="193">P. huoshanensis</emphasis>
<taxonomicName id="329AB5F21D06DA9FBB2AF3336DBEFCC7" class="Insecta" kingdom="Animalia" lsidName="P. huoshanensis" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="huoshanensis">
<emphasis id="4910CCC3B3C38D9BD2AE678FD35F69F4" italics="true" pageId="0" pageNumber="193">P. huoshanensis</emphasis>
</taxonomicName>
as
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="P. luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subSpecies="oriens">
<emphasis italics="true" pageId="0" pageNumber="193">P. luteus oriens</emphasis>
<taxonomicName id="C89F8561612FFEB29CAD78E6EA1696F5" class="Insecta" kingdom="Animalia" lsidName="P. luteus subsp. oriens" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="luteus" subSpecies="oriens">
<emphasis id="670880A9655676F48A8AE1E7FDBB2BFB" italics="true" pageId="0" pageNumber="193">P. luteus oriens</emphasis>
</taxonomicName>
, which was later corrected by
<bibRefCitation author="Ishiwata, SI" editor="Bae, YJ" journalOrPublisher="The Korean Society of Aquatic Entomology, Korea" pageId="0" pageNumber="193" pagination="55 - 84" refId="B12" refString="Ishiwata, SI, 2001. A checklist of Japanese Ephemeroptera. In: Bae, YJ, Ed., The 21st Century and Aquatic Entomology in East Asia. The Korean Society of Aquatic Entomology, Korea: 55 - 84" title="A checklist of Japanese Ephemeroptera." volumeTitle="The 21 st Century and Aquatic Entomology in East Asia." year="2001">Ishiwata (2001)</bibRefCitation>
<bibRefCitation id="9303546948F91E968CA5EF2B15E72751" author="Ishiwata, SI" editor="Bae, YJ" journalOrPublisher="The Korean Society of Aquatic Entomology, Korea" pageId="0" pageNumber="193" pagination="55 - 84" refId="B12" refString="Ishiwata, SI, 2001. A checklist of Japanese Ephemeroptera. In: Bae, YJ, Ed., The 21st Century and Aquatic Entomology in East Asia. The Korean Society of Aquatic Entomology, Korea: 55 - 84" title="A checklist of Japanese Ephemeroptera." volumeTitle="The 21 st Century and Aquatic Entomology in East Asia." year="2001">Ishiwata (2001)</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="193">
<paragraph id="03EF20A81B2B25FFBBF2333835F1F7D2" pageId="0" pageNumber="193">
The distribution of
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<emphasis italics="true" pageId="0" pageNumber="193">P. luteus</emphasis>
<taxonomicName id="2E90147DB6A99C022F4DD48C314BEDF6" class="Insecta" kingdom="Animalia" lsidName="P. luteus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="A82E17378BA7BED1E3FC4AE3F4308183" italics="true" pageId="0" pageNumber="193">P. luteus</emphasis>
</taxonomicName>
is wide, from Africa to Japan. In contrast,
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="P. huoshanensis" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="huoshanensis">
<emphasis italics="true" pageId="0" pageNumber="193">P. huoshanensis</emphasis>
<taxonomicName id="C79BAA613FF429F22B9FA9ABF370310B" class="Insecta" kingdom="Animalia" lsidName="P. huoshanensis" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="huoshanensis">
<emphasis id="F95328B6D065DC5D03ABB567290DE0F3" italics="true" pageId="0" pageNumber="193">P. huoshanensis</emphasis>
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was reported from three allopatric sites in Japan and China. Biogeographic and genetic studies at the population level are required for these species.
</paragraph>
<paragraph pageId="0" pageNumber="193">
<paragraph id="9E113CE2AFF238C4DF986B2F5187887F" pageId="0" pageNumber="193">
At the generic level, the definitions of the genera
<taxonomicName class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="193">Potamanthus</emphasis>
<taxonomicName id="FA9E366AA81DAE44E7A5101BB2CF88CD" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">
<emphasis id="87AA2A7D228DE34D4B90AF9DCEA2B4A9" italics="true" pageId="0" pageNumber="193">Potamanthus</emphasis>
</taxonomicName>
and
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<emphasis italics="true" pageId="0" pageNumber="193">Potamanthodes</emphasis>
<taxonomicName id="3878F44C9133266977CE3F78F6031C1E" authorityName="Ulmer" authorityYear="1920" class="Insecta" family="Potamanthidae" genus="Potamanthodes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Potamanthodes" order="Ephemeroptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">
<emphasis id="C59318AE17D39A2F63A8D3FDA36FE929" italics="true" pageId="0" pageNumber="193">Potamanthodes</emphasis>
</taxonomicName>
were updated by
<bibRefCitation DOI="https://doi.org/10.1080/01650424.2022.2074043" author="Li, WJ" journalOrPublisher="ZooKeys" pageId="0" pageNumber="193" refId="B16" refString="Li, WJ, Zhou, CF, 2022. The first description of the nymph of Potamanthodes macrophthalmus You, 1984 and reinstatements of the genera Potamanthodes Ulmer, 1920 and Stygifloris Bae, McCafferty and Edmunds, 1990 (Ephemeroptera: Potamanthidae). Aquatic Insects. https://doi.org/10.1080/01650424.2022.2074043" title="The first description of the nymph of Potamanthodes macrophthalmus You, 1984 and reinstatements of the genera Potamanthodes Ulmer, 1920 and Stygifloris Bae, McCafferty and Edmunds, 1990 (Ephemeroptera: Potamanthidae). Aquatic Insects." url="https://doi.org/10.1080/01650424.2022.2074043" year="2022">Li and Zhou (2022)</bibRefCitation>
<bibRefCitation id="C0BC03CD0D3402CA459A50795A5A9A12" DOI="https://doi.org/10.1080/01650424.2022.2074043" author="Li, WJ" journalOrPublisher="ZooKeys" pageId="0" pageNumber="193" refId="B16" refString="Li, WJ, Zhou, CF, 2022. The first description of the nymph of Potamanthodes macrophthalmus You, 1984 and reinstatements of the genera Potamanthodes Ulmer, 1920 and Stygifloris Bae, McCafferty and Edmunds, 1990 (Ephemeroptera: Potamanthidae). Aquatic Insects. https://doi.org/10.1080/01650424.2022.2074043" title="The first description of the nymph of Potamanthodes macrophthalmus You, 1984 and reinstatements of the genera Potamanthodes Ulmer, 1920 and Stygifloris Bae, McCafferty and Edmunds, 1990 (Ephemeroptera: Potamanthidae). Aquatic Insects." url="https://doi.org/10.1080/01650424.2022.2074043" year="2022">Li and Zhou (2022)</bibRefCitation>
and confirmed by the characters presented in this study.
</paragraph>
</subSubSection>

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@ -1,225 +1,225 @@
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<taxonomicNameLabel pageId="3" pageNumber="4">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="062F019193902A9B379EDC69ECC919D4" pageId="3" pageNumber="4">sp. n.</taxonomicNameLabel>
Figs 3
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<normalizedToken id="04D9E217C7360A6860BD93ACE5981B3B" originalValue="AF">A-F</normalizedToken>
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<paragraph pageId="3" pageNumber="4">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alterosa caymmii" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
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<paragraph id="880B5E12CCAEA6FE10E6B49EB4068F0C" pageId="3" pageNumber="4">Remarks.</paragraph>
<paragraph id="00EBA778381AF7746D8640F7DB6C733B" pageId="3" pageNumber="4">
<taxonomicName id="37E6F0A0D2B23E5B4B86819F4A5363DC" genus="Alterosa" lsidName="Alterosa caymmii" pageId="3" pageNumber="4" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
sp. n. is very similar to
<taxonomicName class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alterosa amadoi" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="amadoi">Alterosa amadoi</taxonomicName>
<taxonomicName id="F95E6A9EF84133DAEE0C2752D32B2566" genus="Alterosa" lsidName="Alterosa amadoi" pageId="3" pageNumber="4" rank="species" species="amadoi">Alterosa amadoi</taxonomicName>
sp. n., as discussed under that species. It differs in that the lateral branch of each intermediate appendage is apically projecting in
<taxonomicName class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alterosa caymmii" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
<taxonomicName id="00703D5E3B2605777D9E67587B348175" genus="Alterosa" lsidName="Alterosa caymmii" pageId="3" pageNumber="4" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
sp. n., while in
<taxonomicName class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alterosa amadoi" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="amadoi">Alterosa amadoi</taxonomicName>
<taxonomicName id="A60B221C4E63B109C76F81BDFE5B402D" genus="Alterosa" lsidName="Alterosa amadoi" pageId="3" pageNumber="4" rank="species" species="amadoi">Alterosa amadoi</taxonomicName>
sp. n. it is broadly rounded.
<taxonomicName class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alterosa caymmii" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
<taxonomicName id="BDB4C4AAC4933FC3ACE494EC5292701F" genus="Alterosa" lsidName="Alterosa caymmii" pageId="3" pageNumber="4" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
sp. n. also differs from
<taxonomicName class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alterosa amadoi" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="amadoi">Alterosa amadoi</taxonomicName>
<taxonomicName id="7505488EC8E00E47B844BB8FEF9FC9D7" genus="Alterosa" lsidName="Alterosa amadoi" pageId="3" pageNumber="4" rank="species" species="amadoi">Alterosa amadoi</taxonomicName>
sp. n. in the shape of inferior appendages. Additionally, the phallus of
<taxonomicName class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alterosa caymmii" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
<taxonomicName id="4ADE8EF336E68ED98BABB1CFE0593BEE" genus="Alterosa" lsidName="Alterosa caymmii" pageId="3" pageNumber="4" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
sp. n. is more developed, with endotheca bearing several tracts of numerous large spines.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph pageId="3" pageNumber="4">Description.</paragraph>
<paragraph pageId="3" pageNumber="4">Adult. Color (in alcohol) brown; legs, palps, and antennae pale brown, forewing pattern not discernible. Male forewings 5.2-6.4 mm (n=10).</paragraph>
<paragraph pageId="4" pageNumber="5">
<pageBreakToken pageId="4" pageNumber="5" start="start">Male</pageBreakToken>
<subSubSection id="73471A6EB690329E1F217EDFE25BECB7" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="3D09AD95C0E066F502F0DFE1192C3E7F" pageId="3" pageNumber="4">Description.</paragraph>
<paragraph id="4DFD0170B2B4D3DB1B2A711C19DAA363" pageId="3" pageNumber="4">Adult. Color (in alcohol) brown; legs, palps, and antennae pale brown, forewing pattern not discernible. Male forewings 5.2-6.4 mm (n=10).</paragraph>
<paragraph id="CB0730B3527A486F8960D286540A4DB3" pageId="4" pageNumber="5">
<pageBreakToken id="C0D1B4DC6DDA0280B99844A61D7A09F2" pageId="4" pageNumber="5" start="start">Male</pageBreakToken>
genitalia. Tergum VIII with posteromesal margin not or scarcely emarginate. Sternum IX with anterolateral margins weakly rounded, subtruncate; posteroventral margin greatly produced, forming broadly rounded expansion (Fig. 3A). Tergum IX greatly reduced, membranous or fused to base of tergum X (Fig. 3B). Tergum X tapered from base in lateral view; apex sensillate, rounded as viewed dorsally, subtruncate and slightly enlarged in lateral view (Figs 3A, 3B). Intermediate appendages heavily sclerotized, branched subbasally; mesal branch moderately elongate, surpassing preanal appendages, slightly curved outward at midlength in dorsal view, rod-like, rounded apically, covered with spine-like setae; lateral branch short, flange-like, covered with spine-like setae in apical half, apex enlarged and rounded in lateral and dorsal views (Figs 3A, 3B). Preanal appendages elongate, club-like, covered with stiff, small setae; apex rounded with small subacuminate projection bearing small apical setae as viewed laterally and dorsally (Figs 3A, 3B). Inferior appendages elongate, linear, flattened on mesal surface, setose; each with 1st article, in lateral view, approximately 2 times as long as wide, tapering apically; 2nd article subequal in length to 1st article, relatively narrow, especially at midlength, apex rounded, with prominent pad of short, stiff apicomesal setae (Figs 3A, 3C). Phallobase tubular, very short, scarcely curved; endotheca elongate, with several tracts; 1 paired tract near base without spines, 1 apicolateral tract with several large spines, and 1 single tract at apex with 2 clusters of small spines; phallotremal sclerites indistinct (Figs 3D, 3E, 3F).
</paragraph>
<caption pageId="4" pageNumber="5">
<paragraph pageId="4" pageNumber="5">
<caption id="622C121174533E34C7657AC2E4FACF64" pageId="4" pageNumber="5">
<paragraph id="F1338EE733CC3AC3B0469F6A51D5D433" pageId="4" pageNumber="5">
Figure 3.
<taxonomicName class="Insecta" family="Philopotamidae" genus="Alterosa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alterosa caymmii" order="Trichoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
<taxonomicName id="681794EC1C182141082DC85154B33226" genus="Alterosa" lsidName="Alterosa caymmii" pageId="4" pageNumber="5" rank="species" species="caymmii">Alterosa caymmii</taxonomicName>
, new species, male genitalia. A left lateral view B dorsal view C left inferior appendage, dorsal view (setae of left lateral branch of intermediate appendages omitted) D phallic apparatus, left lateral view E phallic apparatus, dorsal view; 3F, phallic apparatus, ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="materials">
<paragraph pageId="4" pageNumber="5">Materials.</paragraph>
<paragraph pageId="4" pageNumber="5">
<subSubSection id="36A4970EAE757A86253C567B6EEA68EB" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="materials">
<paragraph id="A97C9AD5FDF0211EF7B8FFC2B5484AFA" pageId="4" pageNumber="5">Materials.</paragraph>
<paragraph id="CFFEC17065CFDC5503337EAC534A2208" pageId="4" pageNumber="5">
Holotype male: BRAZIL: Bahia: Varzedo, Serra da
<normalizedToken originalValue="Jibóia">Jiboia</normalizedToken>
<normalizedToken id="D7059D5C16F20234B8D08455F2A21AC9" originalValue="Jibóia">Jiboia</normalizedToken>
, Reserva
<normalizedToken originalValue="Jequitibá">Jequitiba</normalizedToken>
<normalizedToken id="F33A0C20687DC347FCB400DF86DD30DF" originalValue="Jequitibá">Jequitiba</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-12.872639">12°52'21.5&quot;S</geoCoordinate>
<geoCoordinate id="FA015D288419628A4B4C3BF4F8194130" direction="south" orientation="latitude" precision="1" value="-12.872639">12°52'21.5&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-39.48236">39°28'56.5&quot;W</geoCoordinate>
<geoCoordinate id="F4943579F2A7B329C8C5E2E4F0A29A37" direction="west" orientation="longitude" precision="1" value="-39.48236">39°28'56.5&quot;W</geoCoordinate>
, 07.xi.2010, ca 400 m, A. M. Silva-Neto &amp; M.
<normalizedToken originalValue="Araújo">Araujo</normalizedToken>
<normalizedToken id="E9AE567EFDFE85874FB9994902D973EF" originalValue="Araújo">Araujo</normalizedToken>
leg. (MZSP).
</paragraph>
<paragraph lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<paragraph id="05C2E3267CCE37585B26D229B44031E4" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Paratypes: BRAZIL: Bahia: same data as holotype, 7 males, 3 females (UFBA); Varzedo, Reserva
<normalizedToken originalValue="Gambá">Gamba</normalizedToken>
<normalizedToken id="77A5CE077E4492F14FF51CE08434D770" originalValue="Gambá">Gamba</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.953333">12°57'12&quot;S</geoCoordinate>
<geoCoordinate id="D713DF8344CE58E08C11626822E1CB97" direction="south" orientation="latitude" precision="15" value="-12.953333">12°57'12&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.475555">39°28'32&quot;W</geoCoordinate>
<geoCoordinate id="A39B7C37495944C04A56BDEB8B878417" direction="west" orientation="longitude" precision="15" value="-39.475555">39°28'32&quot;W</geoCoordinate>
, ca 510 m, 08.xi.2010, bandeja, A. R. Calor, F. Quinteiro, D.
<normalizedToken originalValue="França">Franca</normalizedToken>
<normalizedToken id="B01CF3A5E24DD0E944658567903BD230" originalValue="França">Franca</normalizedToken>
, R. Mariano &amp; A. Costa leg., 17 males, 1 female (UFBA); Santa Teresinha, Serra da
<normalizedToken originalValue="Jibóia">Jiboia</normalizedToken>
<normalizedToken id="E978AE64EF61250255D94CAB12E4C488" originalValue="Jibóia">Jiboia</normalizedToken>
, Riacho das Torres,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.849722">12°50'59&quot;S</geoCoordinate>
<geoCoordinate id="1CCF0EAE80797738A9866024C6A7CE32" direction="south" orientation="latitude" precision="15" value="-12.849722">12°50'59&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.47972">39°28'47&quot;W</geoCoordinate>
<geoCoordinate id="BA6EC1BF9F96146DF1AD0CF221DA0B5A" direction="west" orientation="longitude" precision="15" value="-39.47972">39°28'47&quot;W</geoCoordinate>
, ca 490 m, 06.x.2010, Luz, A. R. Calor, F. Quinteiro, D.
<normalizedToken originalValue="França">Franca</normalizedToken>
<normalizedToken id="AEE40F17BC2ADFDA0E0B956E533C9326" originalValue="França">Franca</normalizedToken>
, R. Mariano &amp; A. Costa leg., 37 males, 16 females (DZRJ 3792); Santa Teresinha, Distrito de Pedra Branca, Riacho das Torres,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.85">12°51'00&quot;S</geoCoordinate>
<geoCoordinate id="34A1056AE31079A0F77D5194C6DD7ECB" direction="south" orientation="latitude" precision="15" value="-12.85">12°51'00&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.48">39°28'48&quot;W</geoCoordinate>
<geoCoordinate id="BAFD211E92F2C0ADF506B03803A074F2" direction="west" orientation="longitude" precision="15" value="-39.48">39°28'48&quot;W</geoCoordinate>
, ca 490 m, 28.ix.2009, Luz, A. R. Calor &amp; A; Cruz leg., 1 male (UFBA); Santa Teresinha, Distrito de Pedra Branca, Riacho das Torres,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.85">12°51'00&quot;S</geoCoordinate>
<geoCoordinate id="2C3317C85F9B5E9E45A203D4F4C9546C" direction="south" orientation="latitude" precision="15" value="-12.85">12°51'00&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.48">39°28'48&quot;W</geoCoordinate>
<geoCoordinate id="E37CFAC37B60CA503A1AD95C17A9258F" direction="west" orientation="longitude" precision="15" value="-39.48">39°28'48&quot;W</geoCoordinate>
, ca 490 m, 10.vi.2010, Luz, A. R. Calor, D.
<normalizedToken originalValue="França">Franca</normalizedToken>
<normalizedToken id="0A697DCBE5BDDB1011EFA4A6224DC365" originalValue="França">Franca</normalizedToken>
&amp; F. B. Quinteiro leg., 1 male (UFBA); Santa Teresinha, Distrito de Pedra Branca, Riacho das Torres,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.850278">12°51'01&quot;S</geoCoordinate>
<geoCoordinate id="F0B4088B9619582C79FDBB4B88AC5D97" direction="south" orientation="latitude" precision="15" value="-12.850278">12°51'01&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.48">39°28'48&quot;W</geoCoordinate>
<geoCoordinate id="D02A34571FA7B7F6A729AC77F333FA57" direction="west" orientation="longitude" precision="15" value="-39.48">39°28'48&quot;W</geoCoordinate>
, ca 500 m, 07.viii.2009, A. R. Calor &amp; L. Lecci leg., 1 male (UFBA); Varzedo, Serra da
<normalizedToken originalValue="Jibóia">Jiboia</normalizedToken>
<normalizedToken id="BCB46E12599937454C366811C971A642" originalValue="Jibóia">Jiboia</normalizedToken>
, Fazenda Baixa da Areia Cai
<normalizedToken originalValue="Camarão">Camarao</normalizedToken>
<normalizedToken id="3792F71AD994E73AFD01071B7D215ED9" originalValue="Camarão">Camarao</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.9625">12°57'45&quot;S</geoCoordinate>
<geoCoordinate id="F724F37A766FC1AF38864F4877ED3B85" direction="south" orientation="latitude" precision="15" value="-12.9625">12°57'45&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.453335">39°27'12&quot;W</geoCoordinate>
<geoCoordinate id="CA1A6D99E6C47E8BC10176330E99869F" direction="west" orientation="longitude" precision="15" value="-39.453335">39°27'12&quot;W</geoCoordinate>
, ca 260 m, 27.iii.2012, Pan light trap, F. B. Quinteiro, T. Duarte &amp; I. Garcia leg., 2 males (UFBA); Camacan, Serra Bonita,
<normalizedToken originalValue="córrego">corrego</normalizedToken>
<normalizedToken id="7021ECA79EC674FD63D0BB185933D182" originalValue="córrego">corrego</normalizedToken>
1,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.391111">15°23'28&quot;S</geoCoordinate>
<geoCoordinate id="D19D6CBFA04E9AF617A71A9928444259" direction="south" orientation="latitude" precision="15" value="-15.391111">15°23'28&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.565556">39°33'56&quot;W</geoCoordinate>
<geoCoordinate id="42FFAA350658AEBC160AF6EA96F9B6A8" direction="west" orientation="longitude" precision="15" value="-39.565556">39°33'56&quot;W</geoCoordinate>
, ca 820 m, 31.vii.2008, Luz UV e branca, A. R. Calor, L. Lecci, L. C. Pinho &amp; R. Moretto leg., 2 males, 1 female (DZRJ 3793); Camacan, Serra Bonita, 1ª cachoeira,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.391111">15°23'28&quot;S</geoCoordinate>
<geoCoordinate id="319F5540001F082FE1C94127FFE33E5E" direction="south" orientation="latitude" precision="15" value="-15.391111">15°23'28&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.565834">39°33'57&quot;W</geoCoordinate>
<geoCoordinate id="FDF7F0A02E0EE776F65342B95CE45757" direction="west" orientation="longitude" precision="15" value="-39.565834">39°33'57&quot;W</geoCoordinate>
, ca 830 m, 04.xi.2009, A. R. Calor et al. leg., 2 males (UFBA); Camacan, Fazenda Waldemar da
<normalizedToken originalValue="farmácia">farmacia</normalizedToken>
<normalizedToken id="DDB355CDC05EAE27F89B6BD32B90C87D" originalValue="farmácia">farmacia</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.42139">15°25'17&quot;S</geoCoordinate>
<geoCoordinate id="D756E3F728EE56E263DFCA8F52A05E41" direction="south" orientation="latitude" precision="15" value="-15.42139">15°25'17&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.566944">39°34'01&quot;W</geoCoordinate>
<geoCoordinate id="65525DF2300C0D5DF895B6A1600002EF" direction="west" orientation="longitude" precision="15" value="-39.566944">39°34'01&quot;W</geoCoordinate>
, ca 310 m, 28.iii.2011, A. R. Calor, F. Quintero, D.
<normalizedToken originalValue="França">Franca</normalizedToken>
<normalizedToken id="3A695E94436F24376929CD729255DB2E" originalValue="França">Franca</normalizedToken>
&amp; H. Barreto leg., 1 male (DZRJ 3794); Camacan, RPPN Serra Bonita, riacho 1,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.394444">
<pageBreakToken pageId="5" pageNumber="6" start="start">15</pageBreakToken>
<geoCoordinate id="EDE96451EB5107ED48EE11BB2C20C24A" direction="south" orientation="latitude" precision="15" value="-15.394444">
<pageBreakToken id="0BF46CC4F9A734625E4E5BDAC3388182" pageId="5" pageNumber="6" start="start">15</pageBreakToken>
°23'40&quot;S
</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.56222">39°33'44&quot;W</geoCoordinate>
<geoCoordinate id="8D5E2EBE2654C747CC44829F77B9F2CB" direction="west" orientation="longitude" precision="15" value="-39.56222">39°33'44&quot;W</geoCoordinate>
, ca 720 m, 31.iii.2011, F. Quinteiro, D.
<normalizedToken originalValue="França">Franca</normalizedToken>
<normalizedToken id="DA7D215AF8B6ADF9CF95FCAD3274E4F3" originalValue="França">Franca</normalizedToken>
&amp; H. Barreto leg., 2 males (UFBA); Camacan, RPPN Serra Bonita, Malaise 1,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-15.391111">15°23'28&quot;S</geoCoordinate>
<geoCoordinate id="54C02980A9565E7C8299097D76FE3096" direction="south" orientation="latitude" precision="15" value="-15.391111">15°23'28&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.565556">39°33'56&quot;W</geoCoordinate>
<geoCoordinate id="642F3CF19538BFB6D89348C198E73619" direction="west" orientation="longitude" precision="15" value="-39.565556">39°33'56&quot;W</geoCoordinate>
, ca 260 m, iv.2011, A. R. Calor leg., 5 males, 2 females (DZRJ 3795); Wenceslau
<normalizedToken originalValue="Guimarães">Guimaraes</normalizedToken>
<normalizedToken id="AF44E51D8D4A5704D705D6560BC1188E" originalValue="Guimarães">Guimaraes</normalizedToken>
,
<normalizedToken originalValue="Estação">Estacao</normalizedToken>
<normalizedToken originalValue="Ecológica">Ecologica</normalizedToken>
<normalizedToken id="4AE1AD89734A6A4DC39E873B2F77FD1B" originalValue="Estação">Estacao</normalizedToken>
<normalizedToken id="79AD39E7877584CB92A311585E2CE1A7" originalValue="Ecológica">Ecologica</normalizedToken>
Estadual Wenceslau
<normalizedToken originalValue="Guimarães">Guimaraes</normalizedToken>
<normalizedToken id="4654CFD298C0DD34A7C9C35C5241944E" originalValue="Guimarães">Guimaraes</normalizedToken>
, Riacho Serra Grande, cachoeira em cima,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-13.595278">13°35'43&quot;S</geoCoordinate>
<geoCoordinate id="55CF36DCE767F406BEA9D62C00F96E52" direction="south" orientation="latitude" precision="15" value="-13.595278">13°35'43&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.72">39°43'12&quot;W</geoCoordinate>
<geoCoordinate id="8E6D72A40AB92B5028845149387ACD94" direction="west" orientation="longitude" precision="15" value="-39.72">39°43'12&quot;W</geoCoordinate>
, ca 580 m, 10.x.2010, A. R. Calor et al. leg., 2 males, 2 females (UFBA); Wenceslau
<normalizedToken originalValue="Guimarães">Guimaraes</normalizedToken>
<normalizedToken id="7DA66ED036D92DC4BCC6DF86BBBC69F5" originalValue="Guimarães">Guimaraes</normalizedToken>
,
<normalizedToken originalValue="Estação">Estacao</normalizedToken>
<normalizedToken originalValue="Ecológica">Ecologica</normalizedToken>
<normalizedToken id="071B49B36CA1ED29080FEED4329FE398" originalValue="Estação">Estacao</normalizedToken>
<normalizedToken id="A70AA2CE4B9AC36ECFC67D83A4B07E26" originalValue="Ecológica">Ecologica</normalizedToken>
Estadual Wenceslau
<normalizedToken originalValue="Guimarães">Guimaraes</normalizedToken>
<normalizedToken id="FF97680B6D47680FDDE33DE803B079F5" originalValue="Guimarães">Guimaraes</normalizedToken>
, Rio Patioba,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-13.580556">13°34'50&quot;S</geoCoordinate>
<geoCoordinate id="48E11D3C9A3CD14169983C36DC1DBDF3" direction="south" orientation="latitude" precision="15" value="-13.580556">13°34'50&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.704723">39°42'17&quot;W</geoCoordinate>
<geoCoordinate id="711AB1FEBAFC145E23EDD3BE4B232C3A" direction="west" orientation="longitude" precision="15" value="-39.704723">39°42'17&quot;W</geoCoordinate>
, ca 540 m, 09.x.2010, A. R. Calor et al. leg., 1 male (DZRJ 3796); Amargosa, Serra do
<normalizedToken originalValue="Timbó">Timbo</normalizedToken>
<normalizedToken id="E5522BC3CF80560F8A8FCE9B390EA6CE" originalValue="Timbó">Timbo</normalizedToken>
, Fazenda
<normalizedToken originalValue="Timbó">Timbo</normalizedToken>
<normalizedToken id="5C8D5777B0BF032C51B3A0F5A40A8184" originalValue="Timbó">Timbo</normalizedToken>
,
<normalizedToken originalValue="Córrego">Corrego</normalizedToken>
<normalizedToken id="D462B558515C872728B6BCE4A74A3028" originalValue="Córrego">Corrego</normalizedToken>
Santa Rita,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-13.106112">13°06'22&quot;S</geoCoordinate>
<geoCoordinate id="A3E38D4CB1BD8C24BE1C63C49CEB0274" direction="south" orientation="latitude" precision="15" value="-13.106112">13°06'22&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-39.66639">39°39'59&quot;W</geoCoordinate>
<geoCoordinate id="E85DB20A99AAEB419584D50C61FD9D0E" direction="west" orientation="longitude" precision="15" value="-39.66639">39°39'59&quot;W</geoCoordinate>
, ca. 710 m, 17.vii.2009, A. R. Calor &amp; L. Lecci leg.,1 male (UFBA).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="distribution">
<paragraph pageId="5" pageNumber="6">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="6">Brazil (Bahia).</paragraph>
<subSubSection id="D541DBAA24698FA66B7F08856DBAA150" pageId="5" pageNumber="6" type="distribution">
<paragraph id="835C902A9E1E2662A691A72E76BFA4D9" pageId="5" pageNumber="6">Distribution.</paragraph>
<paragraph id="482E141811CB0B1324AD7672A03B500B" pageId="5" pageNumber="6">Brazil (Bahia).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="etymology">
<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="6">
<subSubSection id="D6580165E1E55DFEECC3DEDE8941AFEE" pageId="5" pageNumber="6" type="etymology">
<paragraph id="256A7144BF94DFC45AB9932129529FFD" pageId="5" pageNumber="6">Etymology.</paragraph>
<paragraph id="5E04C5491E0641353844F5744204F0E8" pageId="5" pageNumber="6">
This species is named in memory of Dorival Caymmi, considered one of the most important Brazilian songwriters. Caymmi was born in Bahia state in 1914 and died at 2008. He became a national icon with his lyrics that evoked the charm of
<normalizedToken originalValue="Bahias">Bahia's</normalizedToken>
<normalizedToken id="7471ADA7BEBBE01D09785F0F0AEA29C9" originalValue="Bahias">Bahia's</normalizedToken>
fishing villages, beaches and beautiful women, like &quot;O Que
<normalizedToken originalValue="é">e</normalizedToken>
<normalizedToken id="A3AEA7BCA77916D6843D0596925AB6E2" originalValue="é">e</normalizedToken>
Que a Baiana Tem?&quot;,
<normalizedToken originalValue="“Marina”">&quot;Marina&quot;</normalizedToken>
<normalizedToken id="15F29DA0D7A9C49A02AB2CD39E5DB5C2" originalValue="“Marina”">&quot;Marina&quot;</normalizedToken>
, &quot;Rainha do Mar&quot;, &quot;Samba da Minha Terra&quot;, among others.
</paragraph>
</subSubSection>

View file

@ -1,364 +1,364 @@
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<treatment ID-GBIF-Taxon="152032058" LSID="urn:lsid:zoobank.org:act:D953E83C-2809-4DE7-91F2-40451DEBFDEA" httpUri="http://treatment.plazi.org/id/58603F7E4C238B927F604A2986FDED1F" lastPageId="11" lastPageNumber="410" pageId="7" pageNumber="406">
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<paragraph pageId="7" pageNumber="406">
<taxonomicName LSID="58603F7E-4C23-8B92-7F60-4A2986FDED1F" authority="Erwin &amp; Ball" authorityName="Erwin &amp; Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Badister amazonus" order="Coleoptera" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="amazonus" status="sp. n.">Badister amazonus Erwin &amp; Ball</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="406">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="7" pageNumber="406">Figures 1</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="7" pageNumber="406">-7</figureCitation>
<treatment id="58603F7E4C238B927F604A2986FDED1F" ID-GBIF-Taxon="152032058" LSID="urn:lsid:zoobank.org:act:D953E83C-2809-4DE7-91F2-40451DEBFDEA" httpUri="http://treatment.plazi.org/id/58603F7E4C238B927F604A2986FDED1F" lastPageId="11" lastPageNumber="410" pageId="7" pageNumber="406" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Badister" scope_tribe="Licinini">
<subSubSection id="886BAF0C86F7846FA11F8C523ABAE82C" pageId="7" pageNumber="406" type="nomenclature">
<paragraph id="5BD0210BCAD81567672DB4B54522FD0F" pageId="7" pageNumber="406">
<taxonomicName id="E82B22581ED6CB32F278084C811AAA30" ID-CoL="KBR5" LSID="58603F7E-4C23-8B92-7F60-4A2986FDED1F" authority="Erwin &amp; Ball" authorityName="Erwin &amp; Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Badister amazonus" order="Coleoptera" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="amazonus" status="sp. n.">Badister amazonus Erwin &amp; Ball</taxonomicName>
<taxonomicNameLabel id="56EB9A427690A8863A1BA60E19FC3FC2" pageId="7" pageNumber="406">sp. n.</taxonomicNameLabel>
<figureCitation id="2FC0014F9E990E41474D8BA97AC9BDE3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="7" pageNumber="406">Figures 1</figureCitation>
<figureCitation id="C21F66330396F1E3157A029455BC601D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="7" pageNumber="406">-7</figureCitation>
</paragraph>
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<subSubSection pageId="7" pageNumber="406" type="holotype">
<paragraph pageId="7" pageNumber="406">Holotype.</paragraph>
<paragraph pageId="7" pageNumber="406">
<emphasis bold="true" pageId="7" pageNumber="406">
<normalizedToken originalValue="Perú">Peru</normalizedToken>
<subSubSection id="E0BF96F777839A3FF4154B34017F2730" pageId="7" pageNumber="406" type="holotype">
<paragraph id="01EB0DB705D27687E3C51F6FCCD0B135" pageId="7" pageNumber="406">Holotype.</paragraph>
<paragraph id="CEF32772E89B1327E883781D149EFAAC" pageId="7" pageNumber="406">
<emphasis id="BDC95FA5B6E696A1CFAE474443572369" bold="true" pageId="7" pageNumber="406">
<normalizedToken id="A03D24EF248B9447B27B5BEC1BA4233E" originalValue="Perú">Peru</normalizedToken>
</emphasis>
, Loreto, 1.0 km SW boca del Rio Samiria, Vigilante Post 1, 130m, &quot;
<geoCoordinate degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
<geoCoordinate id="97536DDA3F49E0DB90C5E4A646F9586E" degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
,
<geoCoordinate degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
<geoCoordinate id="2211F6CA043925FED2EC94A7AA4743DA" degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
&quot; 31 August 1991 (T.L. Erwin &amp; M.G. Pogue)(NMNH: ADP051824, male).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="407" type="derivation of specific epithet">
<paragraph pageId="8" pageNumber="407">
<pageBreakToken pageId="8" pageNumber="407" start="start">Derivation</pageBreakToken>
<subSubSection id="3564C75632AA61442CC2AE4581E69250" pageId="8" pageNumber="407" type="derivation of specific epithet">
<paragraph id="EBCD5DBFDE1933C7348306CD4A6E338B" pageId="8" pageNumber="407">
<pageBreakToken id="702377DFBCA9E946011346772260B883" pageId="8" pageNumber="407" start="start">Derivation</pageBreakToken>
of specific epithet.
</paragraph>
<paragraph pageId="8" pageNumber="407">
<paragraph id="0D0DD7C4786EA8CBA323B2AE64814BF4" pageId="8" pageNumber="407">
The epithet
<normalizedToken originalValue="“amazonus”">
<normalizedToken id="359F9398EED4C417E8EAC80C5A200965" originalValue="“amazonus”">
&quot;
<emphasis italics="true" pageId="8" pageNumber="407">amazonus</emphasis>
<emphasis id="A9AB100A982BD91A608AB2AFA703F114" italics="true" pageId="8" pageNumber="407">amazonus</emphasis>
&quot;
</normalizedToken>
is a singular Latinized masculine noun in apposition, based on the name of the area in which these beetles were found.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="407" type="proposed english vernacular name">
<paragraph pageId="8" pageNumber="407">Proposed English vernacular name.</paragraph>
<paragraph pageId="8" pageNumber="407">Amazon Fast-walking Beetle.</paragraph>
<subSubSection id="64239B0EF83573DF3F59730D24294A5D" pageId="8" pageNumber="407" type="proposed english vernacular name">
<paragraph id="68D0C4D8E8E756FE6983243582A3DFF5" pageId="8" pageNumber="407">Proposed English vernacular name.</paragraph>
<paragraph id="0D1E4CA9A41EDC5DFC45FE2A54385F82" pageId="8" pageNumber="407">Amazon Fast-walking Beetle.</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="408" pageId="8" pageNumber="407" type="diagnosis">
<paragraph pageId="8" pageNumber="407">Diagnosis.</paragraph>
<paragraph lastPageId="9" lastPageNumber="408" pageId="8" pageNumber="407">
<subSubSection id="057F1F8F28F1FE0572389B4F69F62EF8" lastPageId="9" lastPageNumber="408" pageId="8" pageNumber="407" type="diagnosis">
<paragraph id="4883C0321BEF1443057DACF3152866F8" pageId="8" pageNumber="407">Diagnosis.</paragraph>
<paragraph id="2F89731703567D691AC32A6BB9848219" lastPageId="9" lastPageNumber="408" pageId="8" pageNumber="407">
With the attributes of the subgenus
<taxonomicName authorityName="Clairville" authorityYear="1806" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="407">Badister</emphasis>
<taxonomicName id="1704FA604D63FA19AB406AE5FCAA0B61" authorityName="Clairville" authorityYear="1806" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="8396B08F32B60BA64B80FDF706A981D0" italics="true" pageId="8" pageNumber="407">Badister</emphasis>
</taxonomicName>
, as described by
<bibRefCitation author="Ball, GE" pageId="12" pageNumber="411" refId="B1" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1-258." title="A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1 - 258." year="1959">Ball (1959)</bibRefCitation>
<bibRefCitation id="CFD170B3274C07B02D3BC131948DE87E" author="Ball, GE" pageId="12" pageNumber="411" refId="B1" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1-258." title="A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1 - 258." year="1959">Ball (1959)</bibRefCitation>
and as noted above, and large-sized for the genus (mean SBL more than 6.1 mm for both males and females of
<taxonomicName authorityName="Erwin &amp; Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="407" phylum="Arthropoda" rank="species" species="amazonus">
<emphasis italics="true" pageId="8" pageNumber="407">Badister amazonus</emphasis>
<taxonomicName id="C151A5586F2B7315835DBBFDA33D7C18" authorityName="Erwin &amp; Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="407" phylum="Arthropoda" rank="species" species="amazonus">
<emphasis id="72B951E9DD2F20062A9A4A6ADB30DCE3" italics="true" pageId="8" pageNumber="407">Badister amazonus</emphasis>
</taxonomicName>
; mean SBL less than 5.9 mm for males and females of all other Western Hemisphere species samples). Adults with black head and elytra, the elytron with flavotestaceous base, sutural interneur, lateral margin, and epipleuron, and shiny throughout; prothorax and venter flavotestaceous; appendages testaceous. Microsculpture mesh pattern of head and pronotum isodiametric, sur
<pageBreakToken pageId="9" pageNumber="408" start="start">face</pageBreakToken>
<pageBreakToken id="3F18CA33887DEA76F6F41C3C39CF28B4" pageId="9" pageNumber="408" start="start">face</pageBreakToken>
luster somewhat dull; of elytron, very finely etched transverse lines, surface shiny iridescent. Labial palpomere 3 (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">Fig. 2</figureCitation>
<figureCitation id="AE847B81DC31BEA195D2018C65DEC82E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">Fig. 2</figureCitation>
,
<emphasis bold="true" pageId="9" pageNumber="408">lp3</emphasis>
<emphasis id="F99BCB461CA2E58214EEC8588ECC7E2A" bold="true" pageId="9" pageNumber="408">lp3</emphasis>
) swollen and triangulate (distal margin obliquely truncate) with a very small pointed knob at apex. Pronotum with lateral margin moderately explanate in anterior half, broadly so posteriorly. Male tarsomeres 1-3 (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="9" pageNumber="408">Fig. 3B, 3C</figureCitation>
<figureCitation id="020E04B82995B78A9171906FD59D019C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="9" pageNumber="408">Fig. 3B, 3C</figureCitation>
) slightly widened, with rows of adhesive vestiture on ventral surface narrow, confined to anterior half).
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="410" pageId="9" pageNumber="408" type="description">
<paragraph pageId="9" pageNumber="408">Description.</paragraph>
<paragraph pageId="9" pageNumber="408">
<subSubSection id="9A364F05FF79EC5E4F4627CF86A8CA5A" lastPageId="11" lastPageNumber="410" pageId="9" pageNumber="408" type="description">
<paragraph id="1663D16E8C106B8AFC2E19D707D16709" pageId="9" pageNumber="408">Description.</paragraph>
<paragraph id="7D9C994730C7A3559D245DFBABCA4F0D" pageId="9" pageNumber="408">
(
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="9" pageNumber="408">Fig. 1</figureCitation>
<figureCitation id="971E9CEA19B382E6132287FE683826AB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="9" pageNumber="408">Fig. 1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">2</figureCitation>
<figureCitation id="3B2D3FCD4BDB31C85B7E5CFEA9292B38" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">2</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="9" pageNumber="408">
<figureCitation id="9CAF0FA980AC892B60BDDF26E879E307" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="9" pageNumber="408">
3
<normalizedToken originalValue="AC">A-C</normalizedToken>
<normalizedToken id="7084049D1B6A51E479746B151FA9453F" originalValue="AC">A-C</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Badister amazonus sp. n., dorsal aspect, ADP 051543; type locality. Pygidial (defense) gland system, Legend: cc, collecting canal; ed, efferent duct; edbl, efferent duct, basal lobe; gldr, pygidial gland reservoir. Ovipositor sclerites, Legend: gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite" httpUri="https://binary.pensoft.net/fig/19733" pageId="9" pageNumber="408">4</figureCitation>
<figureCitation id="E5AF2355079DC97CCA7106B195484C82" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Badister amazonus sp. n., dorsal aspect, ADP 051543; type locality. Pygidial (defense) gland system, Legend: cc, collecting canal; ed, efferent duct; edbl, efferent duct, basal lobe; gldr, pygidial gland reservoir. Ovipositor sclerites, Legend: gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite" httpUri="https://binary.pensoft.net/fig/19733" pageId="9" pageNumber="408">4</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Badister amazonus sp. n., ADP 127149; type locality. Male genitalia, median lobe, and parameres, A - left lateral, B - dorsal, and C- right lateral aspects. Legend: a, apical area; bl, basal lobe; bo, basal orifice; lp, left paramere; ml, median lobe; om, ostial membrane; oo, ostial opening; par, preapical ridge; rp, right paramere; sh, shaft; ss, dorsal sclerotized strip." httpUri="https://binary.pensoft.net/fig/19734" pageId="9" pageNumber="408">
<figureCitation id="E43C01553D11B430DE905E1210AA64E8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Badister amazonus sp. n., ADP 127149; type locality. Male genitalia, median lobe, and parameres, A - left lateral, B - dorsal, and C- right lateral aspects. Legend: a, apical area; bl, basal lobe; bo, basal orifice; lp, left paramere; ml, median lobe; om, ostial membrane; oo, ostial opening; par, preapical ridge; rp, right paramere; sh, shaft; ss, dorsal sclerotized strip." httpUri="https://binary.pensoft.net/fig/19734" pageId="9" pageNumber="408">
5
<normalizedToken originalValue="AC">A-C</normalizedToken>
<normalizedToken id="989B13D828D54156F82195E37EAB5E25" originalValue="AC">A-C</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="9" pageNumber="408">
<figureCitation id="5509DA527EAF7A351863D1609389C18B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="9" pageNumber="408">
6
<normalizedToken originalValue="AD">A-D</normalizedToken>
<normalizedToken id="A3C93960DD04306279F0933443D82BBC" originalValue="AD">A-D</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="9" pageNumber="408">7</figureCitation>
<figureCitation id="B5BA8AC71DB957BEEC6B3B93879DF52E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="9" pageNumber="408">7</figureCitation>
).
<emphasis italics="true" pageId="9" pageNumber="408">Size</emphasis>
<emphasis id="C25081A046D1A5FE4B74A96DEAA94132" italics="true" pageId="9" pageNumber="408">Size</emphasis>
: Moderately large for the subgenus; ABL = 6.2-7.9mm, SBL = 5.36-6.73mm, EW (maximum width) 2.55-3.43mm, LP = 0.97-1.22mm, WP = 1.71-1.91 mm, LE = 3.86-4.84mm.
<emphasis italics="true" pageId="9" pageNumber="408">Color</emphasis>
<emphasis id="E89ADFD37025EBA2D239A42F40739683" italics="true" pageId="9" pageNumber="408">Color</emphasis>
: See Diagnosis above.
<emphasis italics="true" pageId="9" pageNumber="408">Luster</emphasis>
<emphasis id="CB9DE32F6008FD2216483884ED3CB681" italics="true" pageId="9" pageNumber="408">Luster</emphasis>
: See Diagnosis above.
<emphasis bold="true" pageId="9" pageNumber="408">Head</emphasis>
<emphasis id="684BF4EB47668E372C0DD83B8FA95843" bold="true" pageId="9" pageNumber="408">Head</emphasis>
(
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">Fig. 2</figureCitation>
<figureCitation id="8C0C82A5BC93AE01552FA7BE165C92FD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">Fig. 2</figureCitation>
): clypeus (
<emphasis bold="true" pageId="9" pageNumber="408">cl</emphasis>
<emphasis id="14EA1EF09C6887BEC1AA4257DAB67267" bold="true" pageId="9" pageNumber="408">cl</emphasis>
) medially depressed, unisetose laterally, no clear demarcation from frons. Frons markedly depressed, apically with depression extended to eye carina, posteriorly somewhat concave with slight swellings medially raised to occiput; occiput shallowly domed; neck broad. Eye moderately convex; gena short and flat. Antennae and mouthparts typical for genus (see above for details).
</paragraph>
<paragraph pageId="9" pageNumber="408">
<emphasis bold="true" pageId="9" pageNumber="408">Prothorax. Pronotum</emphasis>
<paragraph id="7C695CBE82B43B05C8BAF3B76E2B00E1" pageId="9" pageNumber="408">
<emphasis id="75297E21448587728E836751AF447372" bold="true" pageId="9" pageNumber="408">Prothorax. Pronotum</emphasis>
(
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="9" pageNumber="408">Fig. 1</figureCitation>
<figureCitation id="FB6FA16BDF6F197A0EA6BC7C9C2F0525" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="9" pageNumber="408">Fig. 1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">2</figureCitation>
<figureCitation id="EE68452EFB709362FF23D0694EF0096A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. Head, including eyes, antennomeres 1 - 3, mouthparts, and anterior part of pronotum. Legend: a 3, antennomere 3; cl, clypeus; lb, labrum; lp 3, labial palpomere 3; mb, mandibular boss; mn, mandibular notch; mp 4, maxillary palpomere 4." httpUri="https://binary.pensoft.net/fig/19731" pageId="9" pageNumber="408">2</figureCitation>
) markedly broad, about twice as long as head (mean LP/LH: 1.683 for males, 1.846 for females), cordiform, margin narrowly explanate with seta at anterior third; base deeply and bilaterally depressed; hind angle slightly obtusely produced and setose; moderately broader than long (mean W/L: 1.71 for males, 1.91 for females); surface medially smooth, slightly rugose laterally medial to explanation.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">
<pageBreakToken pageId="10" pageNumber="409" start="start">Pterothorax</pageBreakToken>
<paragraph id="EC1A423F9481EA83BFB7F5B31F2CFFEB" pageId="10" pageNumber="409">
<emphasis id="78062D70FD76E40FA91850FAFCAD2ECC" bold="true" pageId="10" pageNumber="409">
<pageBreakToken id="6FFC7077EFAAFA655DA7DDAF4BD31D16" pageId="10" pageNumber="409" start="start">Pterothorax</pageBreakToken>
.
</emphasis>
Normal for genus.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Elytra</emphasis>
<paragraph id="0A87D95F90E71C508BDCD1F4DBDF4E38" pageId="10" pageNumber="409">
<emphasis id="1F514D2BD8AAA65DB85BB1C09C6F74AC" bold="true" pageId="10" pageNumber="409">Elytra</emphasis>
. Elytron about same width as head across eyes (mean WH/WE: 0.998 for both sexes), moderately convex, intervals moderately convex and slightly more so laterally, interval 3 bisetose with each setigerous puncture adherent to interneur 2.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Hind wings</emphasis>
<paragraph id="8F732D661CD058A7BBEA89713A64714F" pageId="10" pageNumber="409">
<emphasis id="BAD8EAE470A94CC6B338655BDF4F05F2" bold="true" pageId="10" pageNumber="409">Hind wings</emphasis>
. Macropterous.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Legs.</emphasis>
<paragraph id="17D323A9C1B60E932E8E93531E1C9894" pageId="10" pageNumber="409">
<emphasis id="D03D68A64FD319A4589236595F8FF058" bold="true" pageId="10" pageNumber="409">Legs.</emphasis>
(
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="10" pageNumber="409">Fig. 1</figureCitation>
<figureCitation id="497B72C030928526D7DAD494723783F7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Badister amazonus sp. n., female, ADP 051544; type locality. Habitus, dorsal aspect. ABL = 7.13 mm." httpUri="https://binary.pensoft.net/fig/19730" pageId="10" pageNumber="409">Fig. 1</figureCitation>
). Overall, normal for subgenus. Male front tarsus (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3B, 3C</figureCitation>
<figureCitation id="DFDB0C374EA6DF1810F4D44A1185234B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3B, 3C</figureCitation>
) with tarsomeres 1-3 slightly dilated and each ventrally (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3C</figureCitation>
<figureCitation id="BE2E377D07DCBCC5A5D167FF98C87FB3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3C</figureCitation>
) with 2-4 rows of white articulo-setae.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Abdominal sterna</emphasis>
<paragraph id="DD1384457BA943610C0D0F46466BBC5B" pageId="10" pageNumber="409">
<emphasis id="92006169739860CD76294F72F9D59BB0" bold="true" pageId="10" pageNumber="409">Abdominal sterna</emphasis>
with normal setation for genus.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Male genitalia</emphasis>
<paragraph id="1FF3A1B0AFB9BE2A930B2339C215EFD3" pageId="10" pageNumber="409">
<emphasis id="3DDCD07278AAAE906254125386310814" bold="true" pageId="10" pageNumber="409">Male genitalia</emphasis>
(
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Badister amazonus sp. n., ADP 127149; type locality. Male genitalia, median lobe, and parameres, A - left lateral, B - dorsal, and C- right lateral aspects. Legend: a, apical area; bl, basal lobe; bo, basal orifice; lp, left paramere; ml, median lobe; om, ostial membrane; oo, ostial opening; par, preapical ridge; rp, right paramere; sh, shaft; ss, dorsal sclerotized strip." httpUri="https://binary.pensoft.net/fig/19734" pageId="10" pageNumber="409">
<figureCitation id="1BEADD4274F6F4A7A267304D86BAB8CC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Badister amazonus sp. n., ADP 127149; type locality. Male genitalia, median lobe, and parameres, A - left lateral, B - dorsal, and C- right lateral aspects. Legend: a, apical area; bl, basal lobe; bo, basal orifice; lp, left paramere; ml, median lobe; om, ostial membrane; oo, ostial opening; par, preapical ridge; rp, right paramere; sh, shaft; ss, dorsal sclerotized strip." httpUri="https://binary.pensoft.net/fig/19734" pageId="10" pageNumber="409">
Fig. 5
<normalizedToken originalValue="A">A-</normalizedToken>
<normalizedToken id="37B8F7F76626527FEFB7DDF210A442E7" originalValue="A">A-</normalizedToken>
5C
</figureCitation>
). Median lobe (
<emphasis bold="true" pageId="10" pageNumber="409">ml</emphasis>
<emphasis id="12745DFFD25452567C30817E6BB071C8" bold="true" pageId="10" pageNumber="409">ml</emphasis>
) with basal lobe (
<emphasis bold="true" pageId="10" pageNumber="409">bl</emphasis>
<emphasis id="04A3118E70F90F7B2601A5D146A0981D" bold="true" pageId="10" pageNumber="409">bl</emphasis>
) about half length of shaft (
<emphasis bold="true" pageId="10" pageNumber="409">sh</emphasis>
<emphasis id="9156646ED86DD1E334FC806DF9B74894" bold="true" pageId="10" pageNumber="409">sh</emphasis>
), basal opening (
<emphasis bold="true" pageId="10" pageNumber="409">bo</emphasis>
<emphasis id="6579116A9B416A42BD515E73F793F58D" bold="true" pageId="10" pageNumber="409">bo</emphasis>
) large. Shaft slender, curved ventrally, dorsally membranous (
<emphasis bold="true" pageId="10" pageNumber="409">om</emphasis>
<emphasis id="934D5DB297E0DCBBB0C3B3CC09600340" bold="true" pageId="10" pageNumber="409">om</emphasis>
) except for two long sclerotized strips (
<emphasis bold="true" pageId="10" pageNumber="409">ss</emphasis>
<emphasis id="7A105632B624F59FF5932DE4DA82EA07" bold="true" pageId="10" pageNumber="409">ss</emphasis>
) extended from basal lobe to ostial opening (
<emphasis bold="true" pageId="10" pageNumber="409">oo</emphasis>
<emphasis id="F0E4AA195A8C7B2D0B3C3670F635EAB7" bold="true" pageId="10" pageNumber="409">oo</emphasis>
); in ventral aspect tapered toward rather broadly rounded apex, preapically with prominent preapical ridge (
<emphasis bold="true" pageId="10" pageNumber="409">par</emphasis>
<emphasis id="36F77CDC4924955F95B862AD38EF2D1C" bold="true" pageId="10" pageNumber="409">par</emphasis>
), in lateral aspect, ridge a prominently projected point. Parameres (
<emphasis bold="true" pageId="10" pageNumber="409">lp, rp</emphasis>
<emphasis id="A22BD0331B765524D31DB5AC7475352D" bold="true" pageId="10" pageNumber="409">lp, rp</emphasis>
)broad, apices subtruncate left paramere
<emphasis bold="true" pageId="10" pageNumber="409">(lp</emphasis>
<emphasis id="24E2F4D2BE2F5C89D2D80E982ADDF78A" bold="true" pageId="10" pageNumber="409">(lp</emphasis>
) longer than right paramere (
<emphasis bold="true" pageId="10" pageNumber="409">rp</emphasis>
<emphasis id="8AA1C6DFA44289B540E19611D1F97A4D" bold="true" pageId="10" pageNumber="409">rp</emphasis>
) about three quarters length of shaft (measured in left lateral aspect). Internal sac with apical ring of rather densely distributed microtrichia, without preapical spines.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Female genitalia.</emphasis>
<paragraph id="00792E385368ACDDE2D0BBC694AE758A" pageId="10" pageNumber="409">
<emphasis id="66631A785114D07AEE06018C6EA6C967" bold="true" pageId="10" pageNumber="409">Female genitalia.</emphasis>
(
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="10" pageNumber="409">
<figureCitation id="4A8A671FAD459F9E3854ED04E243FA39" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="10" pageNumber="409">
Fig. 6
<normalizedToken originalValue="AD">A-D</normalizedToken>
<normalizedToken id="6A34AB4FDF6BEAE31C8D2817BB6444EF" originalValue="AD">A-D</normalizedToken>
</figureCitation>
). Ovipositor with broad laterotergite (
<emphasis bold="true" pageId="10" pageNumber="409">lt</emphasis>
<emphasis id="8D7FFE2FB7B4DDAA11BBFFEF7978EAA5" bold="true" pageId="10" pageNumber="409">lt</emphasis>
) and two gonocoxites (
<emphasis bold="true" pageId="10" pageNumber="409">gc 1, gc 2</emphasis>
<emphasis id="D934BD3059CE2A0B5AC3919DE32243F6" bold="true" pageId="10" pageNumber="409">gc 1, gc 2</emphasis>
); gonocoxite 1 asetose; gonocoxite 2 falcate, base (
<emphasis bold="true" pageId="10" pageNumber="409">b</emphasis>
<emphasis id="82899A7DAF1141F98804FAD83F189440" bold="true" pageId="10" pageNumber="409">b</emphasis>
) large, broad, blade (
<emphasis bold="true" pageId="10" pageNumber="409">bl</emphasis>
<emphasis id="711DDD2D4B9A16E3BDD7C19D2EC8A4B4" bold="true" pageId="10" pageNumber="409">bl</emphasis>
) rather short, with two dorsal ensiform setae (
<emphasis bold="true" pageId="10" pageNumber="409">des</emphasis>
<emphasis id="815629C274103F39284E4DFE97109CBE" bold="true" pageId="10" pageNumber="409">des</emphasis>
), and one ventral ensiform seta (
<emphasis bold="true" pageId="10" pageNumber="409">ves</emphasis>
<emphasis id="C99618F5BFF3AD993F21AF89C3FE4D3D" bold="true" pageId="10" pageNumber="409">ves</emphasis>
), all ensiform setae short; without ventral preapical nematiform setae. Reproductive tract (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="10" pageNumber="409">Fig. 6A, 6B</figureCitation>
<figureCitation id="82FAB405E8B84AC1307FC4E13C8C7330" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Badister amazonus sp. n. ADP 127151; type locality. Female genital tract. A. Ventral aspect, B. Dorsal aspect: Legend, b, base of gonocoxite 2; bl, blade of gonocoxite 2; des, dorsal ensiform seta; gc 1, gonocoxite 1; gc 2, gonocoxite 2; lt, laterotergite; ves, ventral ensiform setae. Ovipositor. C. Ventral aspect, D. Dorsal aspect: Legend, bc, bursa copulatrix; co, common oviduct; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca. dorsal aspect; vc, villous canal." httpUri="https://binary.pensoft.net/fig/19735" pageId="10" pageNumber="409">Fig. 6A, 6B</figureCitation>
) proximally with short, broad bursa copulatrix (
<emphasis bold="true" pageId="10" pageNumber="409">bc</emphasis>
<emphasis id="5527606CB94486CF60E7E95F99777D97" bold="true" pageId="10" pageNumber="409">bc</emphasis>
), continuous at its distal end with common oviduct (
<emphasis bold="true" pageId="10" pageNumber="409">co</emphasis>
<emphasis id="6BEB7AD82AA6369AC97226E31FB86677" bold="true" pageId="10" pageNumber="409">co</emphasis>
) and long spermatheca (
<emphasis bold="true" pageId="10" pageNumber="409">sp</emphasis>
<emphasis id="0CBB1FCFBC1F1A4A0142CC3BB4597B2C" bold="true" pageId="10" pageNumber="409">sp</emphasis>
), latter coiled distally; villous canal (
<emphasis bold="true" pageId="10" pageNumber="409">vc</emphasis>
<emphasis id="5966665EFDE23A6AE9EE32BBFCA2848A" bold="true" pageId="10" pageNumber="409">vc</emphasis>
) extended from near base of spermatheca well up common oviduct; spermathecal gland (
<emphasis bold="true" pageId="10" pageNumber="409">sg</emphasis>
<emphasis id="8C719FB01716E11B4A2A6FE722CC9B70" bold="true" pageId="10" pageNumber="409">sg</emphasis>
) bulbous; spermathecal gland duct (
<emphasis bold="true" pageId="10" pageNumber="409">sgd</emphasis>
<emphasis id="42FC047BC8FF55271D07AA59C4C31AC4" bold="true" pageId="10" pageNumber="409">sgd</emphasis>
) long, slender, attached to spermatheca at base of its coiled portion.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<paragraph id="6BCA9A89154FCF902604641DCFB850D7" pageId="10" pageNumber="409">
Markedly similar to homologous structures in
<taxonomicName authorityName="LeConte" authorityYear="1880" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="species" species="reflexus" subGenus="Baudia">
<emphasis italics="true" pageId="10" pageNumber="409">Badister (Baudia) reflexus</emphasis>
<taxonomicName id="85066A493BFA673022DD429374793108" authorityName="LeConte" authorityYear="1880" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="species" species="reflexus" subGenus="Baudia">
<emphasis id="655AEB211F91C092F0FF5020A80D202F" italics="true" pageId="10" pageNumber="409">Badister (Baudia) reflexus</emphasis>
</taxonomicName>
LeConte (
<bibRefCitation author="Will, KW" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="13" pageNumber="412" pagination="95 - 103" refId="B28" refString="Will, KW, 1998. A new species of Diplocheila Brulle from North America, with notes on female reproductive tract characters in selected Licinini and implications for evolution of the subgenus Isorembus Jeannel (Coleoptera: Carabidae: Licinini). Proceedings of the Entomological Society of Washington 100: 95 - 103" title="A new species of Diplocheila Brulle from North America, with notes on female reproductive tract characters in selected Licinini and implications for evolution of the subgenus Isorembus Jeannel (Coleoptera: Carabidae: Licinini)." volume="100" year="1998">Will 1998</bibRefCitation>
<bibRefCitation id="2E0F2BCE2FF6647D75C386BBB37FD36D" author="Will, KW" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="13" pageNumber="412" pagination="95 - 103" refId="B28" refString="Will, KW, 1998. A new species of Diplocheila Brulle from North America, with notes on female reproductive tract characters in selected Licinini and implications for evolution of the subgenus Isorembus Jeannel (Coleoptera: Carabidae: Licinini). Proceedings of the Entomological Society of Washington 100: 95 - 103" title="A new species of Diplocheila Brulle from North America, with notes on female reproductive tract characters in selected Licinini and implications for evolution of the subgenus Isorembus Jeannel (Coleoptera: Carabidae: Licinini)." volume="100" year="1998">Will 1998</bibRefCitation>
: 99, Fig. 8), which differ as follows: spermatheca with basal uncoiled portion more extensive; distal coiled portion short; spermathecal gland elongate digitiform.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Classification.</emphasis>
<bibRefCitation author="Ball, GE" pageId="12" pageNumber="411" refId="B1" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1-258." title="A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1 - 258." year="1959">Ball (1959</bibRefCitation>
<paragraph id="9547FFDFE610945AFB69D6155EE43936" pageId="10" pageNumber="409">
<emphasis id="89CADDF4E987CF7360C50353F3D0113E" bold="true" pageId="10" pageNumber="409">Classification.</emphasis>
<bibRefCitation id="ED40DE919ED3C67B2F578453A4374E22" author="Ball, GE" pageId="12" pageNumber="411" refId="B1" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1-258." title="A taxonomic study of the North American Licinini with notes on the Old World species of the Genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society, 16, iv + 1 - 258." year="1959">Ball (1959</bibRefCitation>
: 194-195) arranged the species of subgenus
<taxonomicName authorityName="Clairville" authorityYear="1806" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="409">Badister</emphasis>
<taxonomicName id="B3610636B02A2B5883E6A4306323DD05" authorityName="Clairville" authorityYear="1806" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="049733198FE5483069010699C45F3876" italics="true" pageId="10" pageNumber="409">Badister</emphasis>
</taxonomicName>
in two
<normalizedToken originalValue="“complexes”">&quot;complexes&quot;</normalizedToken>
<normalizedToken id="75636B8E7FD25B988CC51993FFA233F7" originalValue="“complexes”">&quot;complexes&quot;</normalizedToken>
characterized principally by features of the male genitalia and ovipositor gonocoxite 2. The combination of these features in
<taxonomicName authorityName="Erwin &amp; Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="species" species="amazonus">
<emphasis italics="true" pageId="10" pageNumber="409">Badister amazonus</emphasis>
<taxonomicName id="2A82415B4E426683F8BB3DDEB5C9EF20" authorityName="Erwin &amp; Ball" authorityYear="2011" class="Insecta" family="Carabidae" genus="Badister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="species" species="amazonus">
<emphasis id="CF9B6D5070E52F7333DD47FF3F7790C6" italics="true" pageId="10" pageNumber="409">Badister amazonus</emphasis>
</taxonomicName>
fits neither complex. Accordingly, we propose here the monobasic
<emphasis italics="true" pageId="10" pageNumber="409">amazonus</emphasis>
<emphasis id="C2CE417241982C9D442C69B5B4D710E9" italics="true" pageId="10" pageNumber="409">amazonus</emphasis>
complex, characterized as follows: median lobe of male genitalia dorsally with two long sclerotized strips, internal sac without sclerotized plates or spines; gonocoxite 2 of ovipositor relatively broad basally, ensiform setae relatively short; nematiform setae absent. Additional diagnostic features are: male fore tarsomeres 1-3 (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3B</figureCitation>
<figureCitation id="82EF0EE0947FE85E466D31612DA15B2B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3B</figureCitation>
) relatively slender, not much wider than tarsomeres 1-3 of middle and hind tarsi, fore tarsomere 1 longer than wide, ventral adhesive setae (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3C</figureCitation>
<figureCitation id="152C143CA7BF43B72299831D5853817B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Badister amazonus sp. n., dorsal aspect, female, ADP 051544; type locality. A. Scutellar region of elytra: (pss) parascutellar stria; B-C. Foreleg tarsomeres of male (ADP 051713); B. Dorsal aspect, C. ventral aspect; ats, articulo-seta; ta 1, ta 2, ta 3, tarsomeres 1 - 3." httpUri="https://binary.pensoft.net/fig/19732" pageId="10" pageNumber="409">Fig. 3C</figureCitation>
) concentrated in anterior half of each tarsomere.
</paragraph>
<paragraph pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Dispersal potential</emphasis>
<paragraph id="EE78F0E0889D124474A638037FCAF34F" pageId="10" pageNumber="409">
<emphasis id="13ECEE9A548C013CF3A653C813F5496B" bold="true" pageId="10" pageNumber="409">Dispersal potential</emphasis>
. These beetles are fully macropterous and are probably capable of flight; they are fast walkers. They also are climbers on grasses (
<taxonomicName class="Liliopsida" family="Poaceae" genus="Paspalum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="10" pageNumber="409" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="409">Paspalum</emphasis>
<taxonomicName id="C5940722F058B196A804C04E19074A09" class="Liliopsida" family="Poaceae" genus="Paspalum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="10" pageNumber="409" phylum="Tracheophyta" rank="genus">
<emphasis id="BAE5F5B82DF53C85F33103146A909035" italics="true" pageId="10" pageNumber="409">Paspalum</emphasis>
</taxonomicName>
spp.).
</paragraph>
<paragraph lastPageId="11" lastPageNumber="410" pageId="10" pageNumber="409">
<emphasis bold="true" pageId="10" pageNumber="409">Way of life</emphasis>
<paragraph id="151B61399CA44D6D7C8A409D69181333" lastPageId="11" lastPageNumber="410" pageId="10" pageNumber="409">
<emphasis id="D31D4B7099F94010475286B639AB4442" bold="true" pageId="10" pageNumber="409">Way of life</emphasis>
. Adults are associated with water: in open grassy marshes, on beaches of black and white water rivers, and in forest swamps at the edges of standing black
<pageBreakToken pageId="11" pageNumber="410" start="start">water</pageBreakToken>
<pageBreakToken id="9DB02B4782743A57578A2DBF08745ACE" pageId="11" pageNumber="410" start="start">water</pageBreakToken>
ponds in leaf litter on grey clay soil (Kaolin). They are active at night and take cover in the day among grass stems and under flood debris. They are active in May and
<normalizedToken originalValue="AugustSeptember">August-September</normalizedToken>
<normalizedToken id="C131F204E81C8B1C2E8B1743A84AA20B" originalValue="AugustSeptember">August-September</normalizedToken>
; no teneral adults were encountered in these months.
</paragraph>
<paragraph pageId="11" pageNumber="410">
<emphasis bold="true" pageId="11" pageNumber="410">Other specimens examined</emphasis>
<paragraph id="64AE3CE15462C370B3AA3CBC3C47016D" pageId="11" pageNumber="410">
<emphasis id="D13BFC8EE4BF5CF08CECB48151B6CD34" bold="true" pageId="11" pageNumber="410">Other specimens examined</emphasis>
.
<emphasis bold="true" pageId="11" pageNumber="410">
<normalizedToken originalValue="Perú">Peru</normalizedToken>
<emphasis id="C8C0052CB4ED67AA071294C2AD9D26F3" bold="true" pageId="11" pageNumber="410">
<normalizedToken id="8852B484D1479FFE33B8860C3B7A76C4" originalValue="Perú">Peru</normalizedToken>
,
</emphasis>
Loreto, SW bank of Rio Ucayali, 90m,
<geoCoordinate degrees="4.2089" direction="south" orientation="latitude" precision="5" value="-4.2089">4.2089°S</geoCoordinate>
<geoCoordinate id="5358F7007A8DACC2F3E970EEFE6AC63D" degrees="4.2089" direction="south" orientation="latitude" precision="5" value="-4.2089">4.2089°S</geoCoordinate>
,
<geoCoordinate degrees="73.3657" direction="west" orientation="longitude" precision="5" value="-73.3657">73.3657°W</geoCoordinate>
<geoCoordinate id="32BF2DD3D1C41A45E93362421EEC1CD9" degrees="73.3657" direction="west" orientation="longitude" precision="5" value="-73.3657">73.3657°W</geoCoordinate>
, 2 September 1991, (T.L. Erwin)(NMNH: ADP051862, female); Pacaya-Samiria National Reserve, Rio Samiria, Cocha Shinguito, 90m,
<geoCoordinate degrees="5.1775" direction="south" orientation="latitude" precision="5" value="-5.1775">5.1775°S</geoCoordinate>
<geoCoordinate id="794A9C11B3A9F34D55716B1506605486" degrees="5.1775" direction="south" orientation="latitude" precision="5" value="-5.1775">5.1775°S</geoCoordinate>
,
<geoCoordinate degrees="74.6556" direction="west" orientation="longitude" precision="5" value="-74.6556">74.6556°W</geoCoordinate>
<geoCoordinate id="600E914799BBA25BEBF44636EC424DD6" degrees="74.6556" direction="west" orientation="longitude" precision="5" value="-74.6556">74.6556°W</geoCoordinate>
, 21 May 1990 (T.L. Erwin)(NMNH: ADP093676, 128618, females); 1.0 km SW boca del Rio Samiria, Vigilante Post 1, 130m, &quot;
<geoCoordinate degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
<geoCoordinate id="99C0DF2288C57118E9A75A19DB8E4B05" degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
,
<geoCoordinate degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
<geoCoordinate id="1C46DFE227B0F212E8734DB2570324C5" degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
&quot; 31 August 1991-1 November 1991 (G.E. Ball &amp; D. Shpeley)(NMNH: ADP127151, female, 127149, male); 1.0 km SW boca del Rio Samiria, Vigilante Post 1, 130m, &quot;
<geoCoordinate degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
<geoCoordinate id="9F2D3CD9668097D19215CBE8E02582DA" degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
,
<geoCoordinate degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
<geoCoordinate id="F421A20D2A8F763FF76A429675EAB58B" degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
&quot; 31 August 1991 (T.L. Erwin &amp; M.G. Pogue)(NMNH: ADP051697, 051603, 051336, 051721, 051672, 051743, 051693, 051698, 051722, 051694, 051724, 051692, 051670, 051717, 051718, 051723, 050590, 050589, 050605, 051696, 051720 females and ADP051824, 051630, 051719, 051713, 051826, 051691, 051695 males); 1.0 km SW boca del Rio Samiria, Vigilante Post 1, 130m, &quot;
<geoCoordinate degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
<geoCoordinate id="9E4CCBE0B4F46B5D912EAA2260200CE6" degrees="04" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-4.675">04°40.5'S</geoCoordinate>
,
<geoCoordinate degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
<geoCoordinate id="4DB49BD5B8252C572F2FA2ADDC83CADA" degrees="074" direction="west" minutes="18.9" orientation="longitude" precision="92" value="-74.315">074°18.9'W</geoCoordinate>
&quot; 31 August 1991 (T.L. Erwin)(NMNH: ADP051592, 051563, 051544, 051583, 051589, 051559, 051590, 051564, 051567, 051569, 051561, 051581, 051585, 051570, 051584, 051565, 051543 females and ADP051562, 051541, 051591, 051542, 051566, 051588, 051560, 051568 males).
</paragraph>
<paragraph pageId="11" pageNumber="410">
<paragraph id="3D1BB027F0012D87238D26E21CC0C20B" pageId="11" pageNumber="410">
Two males and 12 females at UASM with the following label data: /PERU Dpto. Loreto/ 1 km SW Boca del Rio/Samiria
<geoCoordinate degrees="04" direction="south" minutes="40" orientation="latitude" precision="15" seconds="29" value="-4.674722">04°40'29&quot;S</geoCoordinate>
<geoCoordinate id="2387216551C5F3E75AF773CF828C0D22" degrees="04" direction="south" minutes="40" orientation="latitude" precision="15" seconds="29" value="-4.674722">04°40'29&quot;S</geoCoordinate>
,
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<geoCoordinate id="E45AC3E75DAECF07F2E2BBEA72F04C0A" degrees="74" direction="west" minutes="18" orientation="longitude" precision="15" seconds="55" value="-74.31528">74°18'55&quot;W</geoCoordinate>
130m/marsh, treading/31.VIII.&amp; 1.IX.91 30-91//T. L. ERWIN EXP./Res. Pacaya-Samiria/G.E. Ball &amp; D. Shpeley/collectors 1991//.
</paragraph>
<paragraph pageId="11" pageNumber="410">
<emphasis bold="true" pageId="11" pageNumber="410">Geographic distribution</emphasis>
<paragraph id="9274B1852FDCA6DB4619B04E49F8B48C" pageId="11" pageNumber="410">
<emphasis id="1500093A01C8ADB8AADE20A2C1B88873" bold="true" pageId="11" pageNumber="410">Geographic distribution</emphasis>
. (
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="11" pageNumber="410">Fig. 7</figureCitation>
<figureCitation id="43EC49F44650090520DF7F726090EB25" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution dot map for known localities of Badister amazonus sp. n .. along the rivers Samiria and Ucayali (★)." httpUri="https://binary.pensoft.net/fig/19736" pageId="11" pageNumber="410">Fig. 7</figureCitation>
). This species is currently known from only three localities: two along the
<normalizedToken originalValue="Río">Rio</normalizedToken>
<normalizedToken id="85E91F3C78F8920F12F9475853C7C0D6" originalValue="Río">Rio</normalizedToken>
Samiria, and one along the
<normalizedToken originalValue="Río">Rio</normalizedToken>
<normalizedToken id="E52B7B5E65B9C1663FCF9389AE9E84C3" originalValue="Río">Rio</normalizedToken>
Ucayali, Loreto Department,
<normalizedToken originalValue="Perú">Peru</normalizedToken>
<normalizedToken id="C781C527AD4E893AB8B652BB0BFDBABD" originalValue="Perú">Peru</normalizedToken>
.
</paragraph>
<paragraph pageId="11" pageNumber="410">
<emphasis bold="true" pageId="11" pageNumber="410">Notes</emphasis>
<paragraph id="093E61360A6C0DB9435CAFD10829AFEA" pageId="11" pageNumber="410">
<emphasis id="D6E30E5B474301600D94FBF98D316C1B" bold="true" pageId="11" pageNumber="410">Notes</emphasis>
. We are not presenting an analysis of variation in body proportions because this species can be separated from all other members of the genus without resorting to such attributes. In regard to the location (latitude/longitude) above for &quot;1.0 km SW del boca Rio Samiria,&quot; Google Earth allows us now to refine what is given on the labels. The actual site is
<geoCoordinate degrees="4.686779" direction="south" orientation="latitude" precision="1" value="-4.686779">4.686779°S</geoCoordinate>
<geoCoordinate id="819F3F883D5B57CD1E936D9BFAB397F4" degrees="4.686779" direction="south" orientation="latitude" precision="1" value="-4.686779">4.686779°S</geoCoordinate>
,
<geoCoordinate degrees="74.336711" direction="west" orientation="longitude" precision="1" value="-74.33671">74.336711°W</geoCoordinate>
<geoCoordinate id="83EE2B052FEF19BEC58C4DFEB503F6B0" degrees="74.336711" direction="west" orientation="longitude" precision="1" value="-74.33671">74.336711°W</geoCoordinate>
at an elevation of 101m. Also, the female specimen from the
<normalizedToken originalValue="Río">Rio</normalizedToken>
<normalizedToken id="1569617B5083464B59997E727DF49315" originalValue="Río">Rio</normalizedToken>
Ucayali was mistakenly labeled
<normalizedToken originalValue="“Río">&quot;Rio</normalizedToken>
<normalizedToken id="E7C0D955DDD77B03B81528FE7FF504F9" originalValue="“Río">&quot;Rio</normalizedToken>
Amazonas.&quot; The river changes names in the vicinity of Iquitos,
<normalizedToken originalValue="Perú">Peru</normalizedToken>
<normalizedToken id="69631CE8BCDCA98DC8B4D1C12671933B" originalValue="Perú">Peru</normalizedToken>
.
</paragraph>
</subSubSection>

View file

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Tshernyshev
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@ -63,7 +64,7 @@ Tshernyshev
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MATERIAL.
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,
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Figures 10-16
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<paragraph pageId="4" pageNumber="65">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="65">The specific name is derived from the long and dense hairs on the body.</paragraph>
<subSubSection id="3805A8CA490AA4B6BE5F9E01A5FD959B" pageId="4" pageNumber="65" type="etymology">
<paragraph id="1E2CED84A11AFF33E493043C84CADDB7" pageId="4" pageNumber="65">Etymology.</paragraph>
<paragraph id="687811C0ECA8B5D28B249CF866284425" pageId="4" pageNumber="65">The specific name is derived from the long and dense hairs on the body.</paragraph>
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<paragraph pageId="4" pageNumber="65">Types.</paragraph>
<paragraph pageId="4" pageNumber="65">Holotype, female, CHINA: Quannan County, 650m, Jiangxi Province, 29 June 2010, leg. Shi-Chang Li. Paratypes: 7 males, CHINA: Quannan County, 530 to 628m, Jiangxi Province, 12 May to 10 June 2008, leg. Shi-Chang Li. 3 males, CHINA: Quannan County, 628 to 700m, Jiangxi Province, 31 May to 18 June 2010, leg. Shi-Chang Li.</paragraph>
<subSubSection id="32ABB1B19718F914A9FDCE33AA6E9D84" pageId="4" pageNumber="65" type="types">
<paragraph id="E5C6DCF58B31B654A88B140AE4945DBD" pageId="4" pageNumber="65">Types.</paragraph>
<paragraph id="A3AD88E1F57E52DFFE3AA46C050FF125" pageId="4" pageNumber="65">Holotype, female, CHINA: Quannan County, 650m, Jiangxi Province, 29 June 2010, leg. Shi-Chang Li. Paratypes: 7 males, CHINA: Quannan County, 530 to 628m, Jiangxi Province, 12 May to 10 June 2008, leg. Shi-Chang Li. 3 males, CHINA: Quannan County, 628 to 700m, Jiangxi Province, 31 May to 18 June 2010, leg. Shi-Chang Li.</paragraph>
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<subSubSection pageId="4" pageNumber="65" type="diagnosis">
<paragraph pageId="4" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="65">Second tergum, hind femur and tibia black. Notaulus of male reaching beyond center of mesoscutum. Lateral sides of scutellum raised and median portion weakly concave in male. Scutellum with lateral carina extending 0.2 to 0.3 of its length.</paragraph>
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<paragraph pageId="4" pageNumber="65">
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<paragraph id="7B51D9CC369C5D873AB750E13175D581" pageId="4" pageNumber="65">Diagnosis.</paragraph>
<paragraph id="3546CDAC99D87E558F0369524B79F2F6" pageId="4" pageNumber="65">Second tergum, hind femur and tibia black. Notaulus of male reaching beyond center of mesoscutum. Lateral sides of scutellum raised and median portion weakly concave in male. Scutellum with lateral carina extending 0.2 to 0.3 of its length.</paragraph>
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<paragraph id="78D21CF126AD242B26DDCF2B4CBA34FC" pageId="4" pageNumber="65">
Figures 10-16.
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Carinityla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carinityla pilosa" order="Hymenoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="pilosa">Carinityla pilosa</taxonomicName>
<taxonomicName id="4C59D243A4C9AE3287C72F5525CE4ACF" genus="Carinityla" lsidName="Carinityla pilosa" pageId="4" pageNumber="65" rank="species" species="pilosa">Carinityla pilosa</taxonomicName>
Sheng &amp; Sun, sp. n. 10-13: Female. 10 Body, lateral view 11 Face 12 Mesopleuron 13 Propodeum 14-16: male 14 Body, lateral view 15 Median portion of flagellomeres and tyloids 16 Propodeum.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="65" type="description">
<paragraph pageId="4" pageNumber="65">Description.</paragraph>
<paragraph pageId="4" pageNumber="65">Female. Body length about 9.0 mm. Fore wing length about 7.2 mm. Ovipositor length about 2.8 mm. Head and mesosoma with long and dense yellowish brown hairs.</paragraph>
<subSubSection id="49C5E5F3CEEC054B713F8ADD73643091" pageId="4" pageNumber="65" type="description">
<paragraph id="F91C50BE189E36DB145DBAE5E7CA03F2" pageId="4" pageNumber="65">Description.</paragraph>
<paragraph id="08D87F349BEA79C933F3C9BAE44F8F75" pageId="4" pageNumber="65">Female. Body length about 9.0 mm. Fore wing length about 7.2 mm. Ovipositor length about 2.8 mm. Head and mesosoma with long and dense yellowish brown hairs.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="65" type="head">
<paragraph pageId="4" pageNumber="65">Head.</paragraph>
<paragraph pageId="4" pageNumber="65">
<subSubSection id="E80F29C56ACCE3E03C8CED490C329EE0" pageId="4" pageNumber="65" type="head">
<paragraph id="7D8E94EE58D6747BA3EA5834A2387F75" pageId="4" pageNumber="65">Head.</paragraph>
<paragraph id="20EE666BB204719054E9AE74C6034901" pageId="4" pageNumber="65">
Face (Figure 11) strongly convex medially, approximately 1.9 times as wide as long, with dense and irregular punctures. Clypeal suture vestigial between clypeal foveae. Clypeus slightly convex, basal portion with sparse and irregular punctures, distance between punctures 1.0 to 3.0 times diameter of puncture; subapical portion with shallow, transverse concavity; apical 0.2 smooth and impunctate, median section of apical margin distinctly raised. Mandible with indistinct longitudinal wrinkles and fine punctures; upper tooth longer than lower tooth. Cheek with elongate punctures. Subocular sulcus indistinct. Malar space approximately 0.5 times as long as basal width of mandible. Gena with dense punctures, distance between punctures 0.2 to 1.0 times diameter of puncture; in dorsal view approximately 0.6 times as long as width of eye. Vertex with irregular punctures, distance between punctures 0.2 to 1.5 times diameter of puncture. Postero-ocellar line about 0.6 times as long as ocular-ocellar line. Frons approximately flat, with regular and dense punctures, distance between punctures 0.2 to 1.0 times diameter of puncture. Antenna distinctly shorter than body in length, with 27 flagellomeres, ratio of length of flagellomeres 1:2:3:4:5 is 3.7:4.7:4.5:4.3:4.1. Flagellomeres 10 to 11 (12) of male with tyloids (Figure 15) similar to those of
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Carinityla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carinityla punctulata" order="Hymenoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="punctulata">Carinityla punctulata</taxonomicName>
<taxonomicName id="3A967C8A93C6540B2CE52A90E60FCCC6" genus="Carinityla" lsidName="Carinityla punctulata" pageId="4" pageNumber="65" rank="species" species="punctulata">Carinityla punctulata</taxonomicName>
. Tyloid on flagellomere 11 0.7 to 0.9 times as long as flagellomere. Occipital carina complete and strong, joining oral carina above base of mandible.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="67" pageId="4" pageNumber="65" type="mesosoma">
<paragraph pageId="4" pageNumber="65">Mesosoma.</paragraph>
<paragraph lastPageId="6" lastPageNumber="67" pageId="4" pageNumber="65">
<subSubSection id="2FB7748EDD42C07CCF8CC06DFB70472F" lastPageId="6" lastPageNumber="67" pageId="4" pageNumber="65" type="mesosoma">
<paragraph id="7F38CD1DEBC0491B5622CAA6643C315F" pageId="4" pageNumber="65">Mesosoma.</paragraph>
<paragraph id="3497D1D49857F61A9F400F8971ACCF92" lastPageId="6" lastPageNumber="67" pageId="4" pageNumber="65">
Anterior portion of pronotum with fine punctures, laterally concave and lower portion with dense, oblique transverse wrinkles; upper posterior portion with fine punctures, distance between punctures 0.2 to 1.0 times diameter of puncture; upper posterior margin slightly and narrowly raised. Mesoscutum with dense punctures, distance between punctures 0.2 to 0.5 times diameter of puncture; posterior median portion with irregular longitudinal wrinkles. Notaulus present anteriorly. Scutoscutellar groove with fine longitudinal wrinkles. Scutellum almost flat, with irregular punctures. Postscutellum smooth and shining, lateral portion strongly convex, anterior-lateral portion deeply concave. Subalar prominence strongly co
<pageBreakToken pageId="5" pageNumber="66" start="start">nvex</pageBreakToken>
<pageBreakToken id="15C0BCDD2BC3D15644022A97FEEE5952" pageId="5" pageNumber="66" start="start">nvex</pageBreakToken>
. Upper portion of mesopleuron (Figure 12) with dense punctures; lower portion, above sternaulus, with irregular punctures; lower posterior portion with transverse wrinkles. Speculum with dense punctures. Epicnemial carina strongly curved backward above sternaulus, upper end reaching to subalar prominence. Metapleuron with dense and irregular punctures. Juxtacoxal carina complete. Anterior section of submetapleural carina strongly projecting. Wings brownish hyaline. Fore wing with vein 1cu-a almost opposite 1-M. Vein 2-Cu approximately 2.0 times as long
<pageBreakToken pageId="6" pageNumber="67" start="start">as</pageBreakToken>
<pageBreakToken id="30D3BBA53D74F25477ABA7A4A5B1D87E" pageId="6" pageNumber="67" start="start">as</pageBreakToken>
2cu-a. Hind wing vein 1-cu about 3.0 times as long as cu-a. Legs robust, with long and dense brown hairs. Hind coxae and femora with distinct fine punctures. Spurs of hind tibia approximately half length of first tarsomere. Ratio of length of hind tarsomere 1:2:3:4:5 is 10.0:4.7:3.4:1.6:3.7. Propodeum (Figure 13) with long, brown hairs. Area superomedia hexagonal, approximately 1.15 times as wide as long, costula connecting slightly in front of its middle. Area basalis and area superomedia smooth and shining. Area externa with fine punctures. Area dentipara with indistinct wrinkles. Area spiracularis almost smooth. Area lateralis with oblique transverse wrinkles. Area petiolaris and area posteroexterna with transverse wrinkles. Propodeal apophysis short and compressed. Propodeal spiracle approximately 3.0 times as long as wide, distance to pleural carina approximately 1.3 times as long as distance to lateral longitudinal carina.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="67" type="metasoma">
<paragraph pageId="6" pageNumber="67">Metasoma.</paragraph>
<paragraph pageId="6" pageNumber="67">First to third terga smooth and shining. First tergum approximately 2.3 times as long as apical width. Hind section of dorsolateral carina, behind spiracle, indistinct. Lateral margins of petiole almost parallel, only posterior portion slightly broadened. Postpetiole weakly and evenly convex, anterior half with sparse and fine punctures. Spiracle circular, very small, located at about apical 0.3 of first tergum. Second tergum approximately 0.65 times as long as apical width. Remaining terga with short brown hairs and indistinct punctures. Ovipositor sheath approximately as long as hind tibia. Ovipositor compressed, with weak nodus.</paragraph>
<subSubSection id="AEE5C39FB2D3C79256B85C88655F1C28" pageId="6" pageNumber="67" type="metasoma">
<paragraph id="10AF642253B39A2E6A74CE5CD20F5C46" pageId="6" pageNumber="67">Metasoma.</paragraph>
<paragraph id="3A350250A90188E365EEAF5C20266B37" pageId="6" pageNumber="67">First to third terga smooth and shining. First tergum approximately 2.3 times as long as apical width. Hind section of dorsolateral carina, behind spiracle, indistinct. Lateral margins of petiole almost parallel, only posterior portion slightly broadened. Postpetiole weakly and evenly convex, anterior half with sparse and fine punctures. Spiracle circular, very small, located at about apical 0.3 of first tergum. Second tergum approximately 0.65 times as long as apical width. Remaining terga with short brown hairs and indistinct punctures. Ovipositor sheath approximately as long as hind tibia. Ovipositor compressed, with weak nodus.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="67" type="color">
<paragraph pageId="6" pageNumber="67">Color</paragraph>
<paragraph pageId="6" pageNumber="67">(Figure 10). Black, except the following. Ventral profiles of scape and pedicel dark brown. Flat portion of flagellomeres more or less brown. Dorsal profiles of eighth to fourteenth flagellomeres white. Maxillary and labial palpi buff except dark bases. Median portion of mandible crimson. All coxae and trochanters, inner profiles of front and mid femora yellowish brown. Remaining portion of fore legs and mid femora brown. Apices of mid femora, mid tibiae and tarsi puce. Apical portion of first tarsomere of hind tarsi, second to fourth tarsomeres, posterior median portions of sixth and seventh terga, main portion of eighth tergum white. Petiole of first tergum yellowish brown; postpetiole reddish brown. Stigma yellowish brown. veins brownish black.</paragraph>
<subSubSection id="F25D2C211A9E77566173C24AA1B2921B" pageId="6" pageNumber="67" type="color">
<paragraph id="BB80EF6F8176AEB91972B0718A787500" pageId="6" pageNumber="67">Color</paragraph>
<paragraph id="29E66227A2C12CC3F5D89F3B0FAF9BE5" pageId="6" pageNumber="67">(Figure 10). Black, except the following. Ventral profiles of scape and pedicel dark brown. Flat portion of flagellomeres more or less brown. Dorsal profiles of eighth to fourteenth flagellomeres white. Maxillary and labial palpi buff except dark bases. Median portion of mandible crimson. All coxae and trochanters, inner profiles of front and mid femora yellowish brown. Remaining portion of fore legs and mid femora brown. Apices of mid femora, mid tibiae and tarsi puce. Apical portion of first tarsomere of hind tarsi, second to fourth tarsomeres, posterior median portions of sixth and seventh terga, main portion of eighth tergum white. Petiole of first tergum yellowish brown; postpetiole reddish brown. Stigma yellowish brown. veins brownish black.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="67" type="male">
<paragraph pageId="6" pageNumber="67">Male</paragraph>
<paragraph pageId="6" pageNumber="67">(Figure 14). Body length 9.5 to 12.0 mm. Fore wing length 7.5 to 8.8 mm. Face 1.7 to 1.8 times as wide as long. Malar space 0.2 to 0.3 times as long as basal width of mandible. Antenna with 27 to 29 flagellomeres. Upper posterior portion of pronotum, in front of tegula, weakly convex. Notaulus long, reaching beyond center of mesoscutum. Lateral sides of scutellum raised, median portion weakly concave; basal 0.2 to 0.3 with lateral carina. Median portion of mesopleuron smooth and shining, impunctate. Area superomedia 1.5 to 1.6 times as wide as long, costula connecting at its anterior 0.3 (Figure 16). Propodeal spiracle 3.0 to 3.5 times as long as wide. First tergum 2.3 to 2.5 times as long as apical width. Dorsal profile of basal flagellomeres brownish black, ventral profile reddish brown; dorsal profiles of seventh to thirteen flagellomeres white; apical flagellomeres brownish black. Stigma and veins brownish black.</paragraph>
<subSubSection id="1D3FC885497FDDFADEDF9255387A7D89" pageId="6" pageNumber="67" type="male">
<paragraph id="820E0664F32A40502453E76DF40B291B" pageId="6" pageNumber="67">Male</paragraph>
<paragraph id="800248A444922B19FC4778824A85EBB3" pageId="6" pageNumber="67">(Figure 14). Body length 9.5 to 12.0 mm. Fore wing length 7.5 to 8.8 mm. Face 1.7 to 1.8 times as wide as long. Malar space 0.2 to 0.3 times as long as basal width of mandible. Antenna with 27 to 29 flagellomeres. Upper posterior portion of pronotum, in front of tegula, weakly convex. Notaulus long, reaching beyond center of mesoscutum. Lateral sides of scutellum raised, median portion weakly concave; basal 0.2 to 0.3 with lateral carina. Median portion of mesopleuron smooth and shining, impunctate. Area superomedia 1.5 to 1.6 times as wide as long, costula connecting at its anterior 0.3 (Figure 16). Propodeal spiracle 3.0 to 3.5 times as long as wide. First tergum 2.3 to 2.5 times as long as apical width. Dorsal profile of basal flagellomeres brownish black, ventral profile reddish brown; dorsal profiles of seventh to thirteen flagellomeres white; apical flagellomeres brownish black. Stigma and veins brownish black.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="67" type="host">
<paragraph pageId="6" pageNumber="67">Host.</paragraph>
<paragraph pageId="6" pageNumber="67">Unknown.</paragraph>
<subSubSection id="3A4913A7BBFAB9525E4331C6BD878EA2" pageId="6" pageNumber="67" type="host">
<paragraph id="86D523DB555D93363FDE28192D730B98" pageId="6" pageNumber="67">Host.</paragraph>
<paragraph id="4198CD5E15EF143477DA7D11D868D39F" pageId="6" pageNumber="67">Unknown.</paragraph>
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</treatment>
</document>

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@ -1,303 +1,303 @@
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<taxonomicName LSID="http://zoobank.org/8F185D84-D2A4-464B-97F9-A5FEDD791B47" authority="Sokolov, 2021" authorityName="Sokolov" authorityYear="2021" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus relictus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="relictus" status="sp. nov.">Anillinus relictus</taxonomicName>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">, 4A-C</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of the South of eastern United States, showing positions of locality records for the newly described species of Anillinus and the ranges of their presumed relatives (localities of the same color reflect supposed relatedness). A. felicianus, green circle; A. relictus, red circle. Green area with vertical line pattern - range of A. sinuaticollis. Black cross - type locality of A. sinuatus. Red areas with diagonal line pattern - ranges of the species of Anillinus whose males have a spine-like structure in the endophallus of the median lobe (after Sokolov et al. 2004; Sokolov and Watrous 2008; Sokolov 2011; Sokolov et al. 2014; Sokolov et al. 2017). Blue line - Mississippi River. State abbreviations follow Federal Information Processing Standards (https: // www. nlsinfo. org / content / cohorts / nlsy 79 / other-documentation / codebook-supplement / nlsy 79 - attachment- 102 - federal)." figureDoi="10.3897/zookeys.1016.61397.figure5" httpUri="https://binary.pensoft.net/fig/508297" pageId="0" pageNumber="63">, 5</figureCitation>
<treatment id="588729DA5AFB535A9D11D17A06E306A6" ID-GBIF-Taxon="177383248" LSID="urn:lsid:zoobank.org:act:8F185D84-D2A4-464B-97F9-A5FEDD791B47" httpUri="http://treatment.plazi.org/id/588729DA5AFB535A9D11D17A06E306A6" lastPageNumber="63" pageId="0" pageNumber="63" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Trechinae" scope_tribe="Anillini">
<subSubSection id="783D2876B663325B0C5040A78893E926" pageId="0" pageNumber="63" type="nomenclature">
<paragraph id="11E8E1FDB0D3B898998C5FA97A2C1D36" pageId="0" pageNumber="63">
<taxonomicName id="484882B29784A1D7EDC8D02904FECACB" ID-CoL="8MBJZ" LSID="http://zoobank.org/8F185D84-D2A4-464B-97F9-A5FEDD791B47" authority="Sokolov, 2021" authorityName="Sokolov" authorityYear="2021" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus relictus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="relictus" status="sp. nov.">Anillinus relictus</taxonomicName>
<taxonomicNameLabel id="C2B9E4294D56575D56B26641792635FE" pageId="0" pageNumber="63">sp. nov.</taxonomicNameLabel>
<figureCitation id="3F10A11B22E164BA8195E70CA305FAA8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">Figs 3</figureCitation>
<figureCitation id="B51CDB2928296E6F0A231F707199D9EB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">, 4A-C</figureCitation>
<figureCitation id="77BB36F238D9F00C03D4C6BB29CCC6AC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of the South of eastern United States, showing positions of locality records for the newly described species of Anillinus and the ranges of their presumed relatives (localities of the same color reflect supposed relatedness). A. felicianus, green circle; A. relictus, red circle. Green area with vertical line pattern - range of A. sinuaticollis. Black cross - type locality of A. sinuatus. Red areas with diagonal line pattern - ranges of the species of Anillinus whose males have a spine-like structure in the endophallus of the median lobe (after Sokolov et al. 2004; Sokolov and Watrous 2008; Sokolov 2011; Sokolov et al. 2014; Sokolov et al. 2017). Blue line - Mississippi River. State abbreviations follow Federal Information Processing Standards (https: // www. nlsinfo. org / content / cohorts / nlsy 79 / other-documentation / codebook-supplement / nlsy 79 - attachment- 102 - federal)." figureDoi="10.3897/zookeys.1016.61397.figure5" httpUri="https://binary.pensoft.net/fig/508297" pageId="0" pageNumber="63">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="type material">
<paragraph pageId="0" pageNumber="63">Type material.</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
<subSubSection id="B4C6B20FB7BF42DD4DB5FC5A463D3C2C" pageId="0" pageNumber="63" type="type material">
<paragraph id="4B8EF4B6FDAA0BCE75EDB6DCDF5BCD3A" pageId="0" pageNumber="63">Type material.</paragraph>
<paragraph id="C9A33B77C950BF021AD0F0F8C1B5F2AF" pageId="0" pageNumber="63">
<emphasis id="DE7085BC231A61AB6F033FCECF61F181" bold="true" italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
, one male (CMNH), dissected, labeled: \ ALABAMA: Blount Co., Tidwell Hollow Nature Trail east of Oneonta. T. N. King April 1 1972 \ 4/1/72 o [handwritten] \ THOMAS C. BARR COLLECTION 2011 Acc. No. 38,014 \.
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Paratype</emphasis>
<emphasis id="B1CB76083B5B4EC53AD8B07F71E735E8" bold="true" italics="true" pageId="0" pageNumber="63">Paratype</emphasis>
, one female, labeled as holotype (CMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="etymology">
<paragraph pageId="0" pageNumber="63">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="63">
<subSubSection id="CDD1048EC64FA339DE87B25CD09B00D0" pageId="0" pageNumber="63" type="etymology">
<paragraph id="1C0A2B06C2116400121F2D9DED932CE8" pageId="0" pageNumber="63">Etymology.</paragraph>
<paragraph id="1DD9462353718371FC7E053EE70C4C63" pageId="0" pageNumber="63">
The specific epithet is a Latin adjective,
<taxonomicName authorityName="Sokolov" authorityYear="2021" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus relictus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="relictus">
<emphasis italics="true" pageId="0" pageNumber="63">Anillinus relictus</emphasis>
<taxonomicName id="C8B22E29295A884FF0C34845BEC3AE7C" authorityName="Sokolov" authorityYear="2021" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus relictus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="75CA988AAFE5F24E4B84000B40CD166D" italics="true" pageId="0" pageNumber="63">Anillinus relictus</emphasis>
</taxonomicName>
(from Latin: abandoned, forsaken), in the masculine form, and refers to the geographical isolation of this species from its morphologically closest congeners, as it is believed to be the only remaining eastern representative of an ancestral group once more widespread.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="type locality">
<paragraph pageId="0" pageNumber="63">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="63">USA, Alabama, Blount County, the Oneonta area.</paragraph>
<subSubSection id="15F86ACCABD8B7C5226954193D8ECA81" pageId="0" pageNumber="63" type="type locality">
<paragraph id="719C7BADBA464A8DA796EF8931ECD80A" pageId="0" pageNumber="63">Type locality.</paragraph>
<paragraph id="BD8AAFAB69E37254C54C5FC30B43195E" pageId="0" pageNumber="63">USA, Alabama, Blount County, the Oneonta area.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="recognition">
<paragraph pageId="0" pageNumber="63">Recognition.</paragraph>
<paragraph pageId="0" pageNumber="63">
<subSubSection id="3DF4626C5CF115098817578D3DC231D0" pageId="0" pageNumber="63" type="recognition">
<paragraph id="E73B57E0AFF1220ECAB1E17EF7749180" pageId="0" pageNumber="63">Recognition.</paragraph>
<paragraph id="A6781B5C9E47667C4D2973ADD7170F17" pageId="0" pageNumber="63">
Adults of
<taxonomicName lsidName="A. relictus" pageId="0" pageNumber="63" rank="species" species="relictus">A. relictus</taxonomicName>
<taxonomicName id="9A982DCDC5DD8924F412F38E6938603B" lsidName="A. relictus" pageId="0" pageNumber="63" rank="species" species="relictus">A. relictus</taxonomicName>
can be distinguished from those of other members of eastern
<taxonomicName authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
<taxonomicName id="4F73B826F66040C0CF13C334021457E5" authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="5D46D46B2ABFED5F29C0942C3752EED2" italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
</taxonomicName>
by the combination of the large size, completely microsculptured head and pronotum, and, especially, by the long elytral vestiture equals to 0.5-0.7 of length of discal elytral setae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="description">
<paragraph pageId="0" pageNumber="63">Description.</paragraph>
<paragraph pageId="0" pageNumber="63">
<subSubSection id="DBF00D2A5B2BCED01BE07E0F8449B9FB" pageId="0" pageNumber="63" type="description">
<paragraph id="C1CB97C40740CC158A9197DCB7DFB67C" pageId="0" pageNumber="63">Description.</paragraph>
<paragraph id="03459AE701D57C236E0016017104E947" pageId="0" pageNumber="63">
Large-sized for genus (ABL 2.29-2.42 mm, mean 2.36
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="3D22CF57F0E4F925175A5D83EDC9EC35" originalValue="±">+/-</normalizedToken>
0.092 mm, n = 2).
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Habitus</emphasis>
<paragraph id="21DDB278F7A60474138DB7D7F550040E" pageId="0" pageNumber="63">
<emphasis id="4C713F04E7714486D22AAEBDCA6E587E" bold="true" italics="true" pageId="0" pageNumber="63">Habitus</emphasis>
: Body form (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">3A</figureCitation>
<figureCitation id="D685633EB3FF31F9D2E5B1AA98BD727B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">3A</figureCitation>
) moderately convex, ovoid (WE/ABL 0.39
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="CEA5618AC8CCA13B8BB01EE0464DA666" originalValue="±">+/-</normalizedToken>
0.012), head of average proportions for genus (WH/WPm 0.71
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="2B1F0BA30A448F202FC8A0FD3327C315" originalValue="±">+/-</normalizedToken>
0.023), pronotum moderately narrow in comparison to elytra (WPm/WE 0.77
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="E9DD3AA0183BE1B337094DA7E7B3A676" originalValue="±">+/-</normalizedToken>
0.007).
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Integument</emphasis>
<paragraph id="5214F4C6C6B38DD957B1AE47C5D8CFE3" pageId="0" pageNumber="63">
<emphasis id="2E09D90A38AA9FBF14C1E31DCAEB4D69" bold="true" italics="true" pageId="0" pageNumber="63">Integument</emphasis>
: Body color piceo-brunneus, appendages testaceous. Microsculpture (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">3B-D</figureCitation>
<figureCitation id="86E71962503B9EBD9B3F6AF9F7D5C083" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">3B-D</figureCitation>
) present across all head, pronotum, and elytra, where it is represented by isodiametric polygonal sculpticells. Body surface shiny, surface sparsely and finely punctate, covered with moderately dense, yellowish, long setae. Vestiture of elytra (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">3D</figureCitation>
<figureCitation id="F39B16F525DE652BB5D0FA0C6FDA8F5D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">3D</figureCitation>
) long (0.5-0.7 length of discal setae). Elytral chaetotaxy typical for
<taxonomicName authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
<taxonomicName id="699EC7411F6F31A27FAB6C9A1588CA54" authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="83D85DCDF2A978336F9095D1E0BCA968" italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
</taxonomicName>
, umbilicate series of type A (
<emphasis italics="true" pageId="0" pageNumber="63">sensu</emphasis>
<bibRefCitation author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie" pageId="0" pageNumber="63" pagination="33 - 204" refId="B13" refString="Jeannel, R, 1963a. Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]." volume="28" year="1963 a">Jeannel 1963a</bibRefCitation>
<emphasis id="35BB38E17B185602A689C8558E3BA39E" italics="true" pageId="0" pageNumber="63">sensu</emphasis>
<bibRefCitation id="32090A51ED2005EBDF6DB7ED603B0807" author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie" pageId="0" pageNumber="63" pagination="33 - 204" refId="B13" refString="Jeannel, R, 1963a. Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]." volume="28" year="1963 a">Jeannel 1963a</bibRefCitation>
and
<bibRefCitation author="Giachino, PM" journalOrPublisher="Biodiversity Journal, Monograph" pageId="0" pageNumber="63" pagination="1 - 112" refId="B9" refString="Giachino, PM, Vailati, D, 2011. Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini). Biodiversity Journal, Monograph 1: 1 - 112" title="Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini)." volume="1" year="2011">Giachino and Vailati 2011</bibRefCitation>
<bibRefCitation id="B596031FD1932004FED5D42FBC46B1ED" author="Giachino, PM" journalOrPublisher="Biodiversity Journal, Monograph" pageId="0" pageNumber="63" pagination="1 - 112" refId="B9" refString="Giachino, PM, Vailati, D, 2011. Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini). Biodiversity Journal, Monograph 1: 1 - 112" title="Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini)." volume="1" year="2011">Giachino and Vailati 2011</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 3.</emphasis>
<caption id="0F81B48F6AEB0644B87D57E7420A26D9" doi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63" start="Figure 3" startId="F3">
<paragraph id="06D44E4D56F72B7B12DB0405CB4626FD" pageId="0" pageNumber="63">
<emphasis id="4441409B8C6137DCCDA9FA2379F635E9" bold="true" pageId="0" pageNumber="63">Figure 3.</emphasis>
Digital images of external features of
<taxonomicName authorityName="Sokolov" authorityYear="2021" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus relictus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="relictus">
<emphasis italics="true" pageId="0" pageNumber="63">Anillinus relictus</emphasis>
<taxonomicName id="75898502E08A0A496DE684AE1DBA5186" authorityName="Sokolov" authorityYear="2021" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus relictus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="861C8963C68C924DE2F9C513FF14572C" italics="true" pageId="0" pageNumber="63">Anillinus relictus</emphasis>
</taxonomicName>
sp. nov. (male, E of Oneonta, Blount County, Alabama)
<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
<emphasis id="F13AFDE3552B02CEDE1E21FBE71D95A5" bold="true" pageId="0" pageNumber="63">A</emphasis>
habitus, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
<emphasis id="EE97A87475DB94D9FD1951287B869B86" bold="true" pageId="0" pageNumber="63">B</emphasis>
head, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="63">C</emphasis>
<emphasis id="815102B203841344B3797C88FFD0A93E" bold="true" pageId="0" pageNumber="63">C</emphasis>
pronotum, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="63">D</emphasis>
<emphasis id="967FE5DD5D32317AA562A1A30BD90FE8" bold="true" pageId="0" pageNumber="63">D</emphasis>
elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="63">A, D</emphasis>
<emphasis id="2E9C0FB7907D836AB2FA08DE96AFB24F" bold="true" pageId="0" pageNumber="63">A, D</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="63">B, C</emphasis>
<emphasis id="202015989D2DB3223A2EFA918B7AED6F" bold="true" pageId="0" pageNumber="63">B, C</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 4.</emphasis>
<caption id="FB4046BFB358DB180614CED68A3728D5" doi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63" start="Figure 4" startId="F4">
<paragraph id="802207888AE4957AD3A2225F3E6708F7" pageId="0" pageNumber="63">
<emphasis id="BD4671FFB1CCAC6AC3C34E6211571E00" bold="true" pageId="0" pageNumber="63">Figure 4.</emphasis>
Digital images and line drawings of male genitalia of
<taxonomicName authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
<taxonomicName id="4F8A5C735E169FC5C0E941E82BF2BDB9" authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="222F51A5FEE8F880A5B7E617D79641F7" italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
</taxonomicName>
species.
<taxonomicName lsidName="A. relictus" pageId="0" pageNumber="63" rank="species" species="relictus">
<emphasis italics="true" pageId="0" pageNumber="63">A. relictus</emphasis>
<taxonomicName id="C1AC5F2296E0E8D28F36A8A1906DE2D3" lsidName="A. relictus" pageId="0" pageNumber="63" rank="species" species="relictus">
<emphasis id="4A4501ADABA5005D71FFF640BD03FCE5" italics="true" pageId="0" pageNumber="63">A. relictus</emphasis>
</taxonomicName>
(E of Oneonta, Blount County, Alabama)
<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
<emphasis id="D85F985365D444EBCF5628B51C28F2FC" bold="true" pageId="0" pageNumber="63">A</emphasis>
median lobe, right lateral aspect
<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
<emphasis id="2C4F5FECAE8FD28E4E2B7F7AA9898A6E" bold="true" pageId="0" pageNumber="63">B</emphasis>
left paramere, left lateral aspect
<emphasis bold="true" pageId="0" pageNumber="63">C</emphasis>
<emphasis id="31A419CA5B0182BD57731E1D3C522321" bold="true" pageId="0" pageNumber="63">C</emphasis>
right paramere, right lateral aspect.
<taxonomicName lsidName="A. sinuatus" pageId="0" pageNumber="63" rank="species" species="sinuatus">
<emphasis italics="true" pageId="0" pageNumber="63">A. sinuatus</emphasis>
<taxonomicName id="A8C90BEA226D50B41F6624E640E0B592" lsidName="A. sinuatus" pageId="0" pageNumber="63" rank="species" species="sinuatus">
<emphasis id="C813D14591FECFBA41D3FA4334399F5D" italics="true" pageId="0" pageNumber="63">A. sinuatus</emphasis>
</taxonomicName>
(Bexar County, Texas)
<emphasis bold="true" pageId="0" pageNumber="63">D</emphasis>
<emphasis id="2908842ADD4E97E38E402A2953F91800" bold="true" pageId="0" pageNumber="63">D</emphasis>
median lobe, right lateral aspect
<emphasis bold="true" pageId="0" pageNumber="63">E</emphasis>
<emphasis id="64D6DB9B8BC8CE29AA94B04F7910E1B7" bold="true" pageId="0" pageNumber="63">E</emphasis>
left paramere, left lateral aspect
<emphasis bold="true" pageId="0" pageNumber="63">F</emphasis>
<emphasis id="AC9F46D563EFF0D03FE2482DB2C3188C" bold="true" pageId="0" pageNumber="63">F</emphasis>
right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1016.61397.figure5" httpUri="https://binary.pensoft.net/fig/508297" pageId="0" pageNumber="63" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" pageId="0" pageNumber="63">Figure 5.</emphasis>
<caption id="6823B7D7EB625F53FB8E1F551E9C49CB" doi="10.3897/zookeys.1016.61397.figure5" httpUri="https://binary.pensoft.net/fig/508297" pageId="0" pageNumber="63" start="Figure 5" startId="F5">
<paragraph id="8658E8DBCCE071BA647BD098DE2C3F3D" pageId="0" pageNumber="63">
<emphasis id="08A9098CEB86736561DE832AB38597BB" bold="true" pageId="0" pageNumber="63">Figure 5.</emphasis>
Map of the South of eastern United States, showing positions of locality records for the newly described species of
<taxonomicName authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
<taxonomicName id="3C09A8A96034863A23FD7765F3A853B1" authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="5B49F5C1E34088387945D28736305D0C" italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
</taxonomicName>
and the ranges of their presumed relatives (localities of the same color reflect supposed relatedness).
<taxonomicName lsidName="A. felicianus" pageId="0" pageNumber="63" rank="species" species="felicianus">
<emphasis italics="true" pageId="0" pageNumber="63">A. felicianus</emphasis>
<taxonomicName id="05FA88A91913A6061E835E81E83DEABA" lsidName="A. felicianus" pageId="0" pageNumber="63" rank="species" species="felicianus">
<emphasis id="0EE9A1A80A932F08D3AD31D91E195BA9" italics="true" pageId="0" pageNumber="63">A. felicianus</emphasis>
</taxonomicName>
, green circle;
<taxonomicName lsidName="A. relictus" pageId="0" pageNumber="63" rank="species" species="relictus">
<emphasis italics="true" pageId="0" pageNumber="63">A. relictus</emphasis>
<taxonomicName id="14A2C3BAF817B43E7A277C4B5FFDF882" lsidName="A. relictus" pageId="0" pageNumber="63" rank="species" species="relictus">
<emphasis id="5D706D693A27CA10F1A813517CD6BABF" italics="true" pageId="0" pageNumber="63">A. relictus</emphasis>
</taxonomicName>
, red circle. Green area with vertical line pattern - range of
<taxonomicName lsidName="A. sinuaticollis" pageId="0" pageNumber="63" rank="species" species="sinuaticollis">
<emphasis italics="true" pageId="0" pageNumber="63">A. sinuaticollis</emphasis>
<taxonomicName id="4CB1A7B31B353D1E7A2B9976EEC1C164" lsidName="A. sinuaticollis" pageId="0" pageNumber="63" rank="species" species="sinuaticollis">
<emphasis id="730154A205C77EB52782A484BA1735EA" italics="true" pageId="0" pageNumber="63">A. sinuaticollis</emphasis>
</taxonomicName>
. Black cross - type locality of
<taxonomicName lsidName="A. sinuatus" pageId="0" pageNumber="63" rank="species" species="sinuatus">
<emphasis italics="true" pageId="0" pageNumber="63">A. sinuatus</emphasis>
<taxonomicName id="23FF2EF23ECFD8350003449A2B7CE930" lsidName="A. sinuatus" pageId="0" pageNumber="63" rank="species" species="sinuatus">
<emphasis id="55F7EB4B0E4B043F42D3E8CB2FA7F13D" italics="true" pageId="0" pageNumber="63">A. sinuatus</emphasis>
</taxonomicName>
. Red areas with diagonal line pattern - ranges of the species of
<taxonomicName authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
<taxonomicName id="74C7577C8300AC7CA1B28E03D729C63C" authorityName="Casey" authorityYear="1918" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="C3F7AC498DDB8FE5ABE3474E28E7A86D" italics="true" pageId="0" pageNumber="63">Anillinus</emphasis>
</taxonomicName>
whose males have a spine-like structure in the endophallus of the median lobe (after
<bibRefCitation DOI="https://doi.org/10.1649/611" author="Sokolov, IM" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="63" pagination="185 - 233" refId="B22" refString="Sokolov, IM, Carlton, CE, Cornell, JF, 2004. Review of Anillinus with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Trechinae: Bembidiini). The Coleopterists Bulletin 58: 185 - 233, DOI: https://doi.org/10.1649/611" title="Review of Anillinus with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Trechinae: Bembidiini)." url="https://doi.org/10.1649/611" volume="58" year="2004">Sokolov et al. 2004</bibRefCitation>
<bibRefCitation id="11D7B3C622D6C0A89D4DE745E73EABE5" DOI="https://doi.org/10.1649/611" author="Sokolov, IM" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="63" pagination="185 - 233" refId="B22" refString="Sokolov, IM, Carlton, CE, Cornell, JF, 2004. Review of Anillinus with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Trechinae: Bembidiini). The Coleopterists Bulletin 58: 185 - 233, DOI: https://doi.org/10.1649/611" title="Review of Anillinus with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Trechinae: Bembidiini)." url="https://doi.org/10.1649/611" volume="58" year="2004">Sokolov et al. 2004</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1649/1114.1" author="Sokolov, IM" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="63" pagination="537 - 543" refId="B27" refString="Sokolov, IM, Watrous, LE, 2008. A new species and the first record of the genus Anillinus (Carabidae: Trechinae: Bembidiini) from the Ozark region. The Coleopterists Bulletin 62: 537 - 543, DOI: https://doi.org/10.1649/1114.1" title="A new species and the first record of the genus Anillinus (Carabidae: Trechinae: Bembidiini) from the Ozark region." url="https://doi.org/10.1649/1114.1" volume="62" year="2008">Sokolov and Watrous 2008</bibRefCitation>
<bibRefCitation id="22A2613704F54AA46F8065731DF1684D" DOI="https://doi.org/10.1649/1114.1" author="Sokolov, IM" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="63" pagination="537 - 543" refId="B27" refString="Sokolov, IM, Watrous, LE, 2008. A new species and the first record of the genus Anillinus (Carabidae: Trechinae: Bembidiini) from the Ozark region. The Coleopterists Bulletin 62: 537 - 543, DOI: https://doi.org/10.1649/1114.1" title="A new species and the first record of the genus Anillinus (Carabidae: Trechinae: Bembidiini) from the Ozark region." url="https://doi.org/10.1649/1114.1" volume="62" year="2008">Sokolov and Watrous 2008</bibRefCitation>
;
<bibRefCitation author="Sokolov, IM" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="63" pagination="1 - 14" refId="B20" refString="Sokolov, IM, 2011. Five new species of Anillinus Casey from the southern Appalachian Mountains and the Piedmont Plateau of eastern U.S.A. (Coleoptera: Carabidae: Trechinae: Bembidiini). Insecta Mundi 164: 1 - 14" title="Five new species of Anillinus Casey from the southern Appalachian Mountains and the Piedmont Plateau of eastern U. S. A. (Coleoptera: Carabidae: Trechinae: Bembidiini)." volume="164" year="2011">Sokolov 2011</bibRefCitation>
<bibRefCitation id="1295FA09F6E207715BE1055B9A291023" author="Sokolov, IM" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="63" pagination="1 - 14" refId="B20" refString="Sokolov, IM, 2011. Five new species of Anillinus Casey from the southern Appalachian Mountains and the Piedmont Plateau of eastern U.S.A. (Coleoptera: Carabidae: Trechinae: Bembidiini). Insecta Mundi 164: 1 - 14" title="Five new species of Anillinus Casey from the southern Appalachian Mountains and the Piedmont Plateau of eastern U. S. A. (Coleoptera: Carabidae: Trechinae: Bembidiini)." volume="164" year="2011">Sokolov 2011</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.417.7733" author="Sokolov, IM" journalOrPublisher="ZooKeys" pageId="0" pageNumber="63" pagination="71 - 101" refId="B26" refString="Sokolov, IM, Reddell, JR, Kavanaugh, DH, 2014. Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution. ZooKeys 417: 71 - 101, DOI: https://doi.org/10.3897/zookeys.417.7733" title="Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution." url="https://doi.org/10.3897/zookeys.417.7733" volume="417" year="2014">Sokolov et al. 2014</bibRefCitation>
<bibRefCitation id="86C03E3117DCC2D96021287877FB0B94" DOI="https://doi.org/10.3897/zookeys.417.7733" author="Sokolov, IM" journalOrPublisher="ZooKeys" pageId="0" pageNumber="63" pagination="71 - 101" refId="B26" refString="Sokolov, IM, Reddell, JR, Kavanaugh, DH, 2014. Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution. ZooKeys 417: 71 - 101, DOI: https://doi.org/10.3897/zookeys.417.7733" title="Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution." url="https://doi.org/10.3897/zookeys.417.7733" volume="417" year="2014">Sokolov et al. 2014</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1649/0010-065X-71.2.289" author="Sokolov, IM" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="63" pagination="289 - 297" refId="B23" refString="Sokolov, IM, Carlton, CE, Watrous, LE, Robison, HW, 2017. Anillinus alleni Sokolov and Carlton (Coleoptera: Carabidae: Trechinae: Bembidiini), a new species from the Ozark Interior Highlands of Arkansas, USA. The Coleopterists Bulletin 71: 289 - 297, DOI: https://doi.org/10.1649/0010-065X-71.2.289" title="Anillinus alleni Sokolov and Carlton (Coleoptera: Carabidae: Trechinae: Bembidiini), a new species from the Ozark Interior Highlands of Arkansas, USA." url="https://doi.org/10.1649/0010-065X-71.2.289" volume="71" year="2017">Sokolov et al. 2017</bibRefCitation>
<bibRefCitation id="4F5E2F91B3D27CE9356C9AA04A6BE079" DOI="https://doi.org/10.1649/0010-065X-71.2.289" author="Sokolov, IM" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="63" pagination="289 - 297" refId="B23" refString="Sokolov, IM, Carlton, CE, Watrous, LE, Robison, HW, 2017. Anillinus alleni Sokolov and Carlton (Coleoptera: Carabidae: Trechinae: Bembidiini), a new species from the Ozark Interior Highlands of Arkansas, USA. The Coleopterists Bulletin 71: 289 - 297, DOI: https://doi.org/10.1649/0010-065X-71.2.289" title="Anillinus alleni Sokolov and Carlton (Coleoptera: Carabidae: Trechinae: Bembidiini), a new species from the Ozark Interior Highlands of Arkansas, USA." url="https://doi.org/10.1649/0010-065X-71.2.289" volume="71" year="2017">Sokolov et al. 2017</bibRefCitation>
). Blue line - Mississippi River. State abbreviations follow Federal Information Processing Standards (https://www.nlsinfo.org/content/cohorts/nlsy79/other-documentation/codebook-supplement/nlsy79-attachment-102-federal).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Prothorax</emphasis>
<paragraph id="B14A3733906DBC1999D07E4127B48D44" pageId="0" pageNumber="63">
<emphasis id="C036EE0CECCB8931B4E7D4511AA5737B" bold="true" italics="true" pageId="0" pageNumber="63">Prothorax</emphasis>
: Pronotum (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">3C</figureCitation>
<figureCitation id="D4BB69E31205F906488E8DF8042AD0F0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Digital images of external features of Anillinus relictus sp. nov. (male, E of Oneonta, Blount County, Alabama) A habitus, dorsal aspect B head, dorsal aspect C pronotum, dorsal aspect D elytral vestiture, left lateral aspect. Scale bars: 0.5 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.1016.61397.figure3" httpUri="https://binary.pensoft.net/fig/508295" pageId="0" pageNumber="63">3C</figureCitation>
) moderately convex, of moderate size (LP/LE 0.40
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="550E391B3E7F757EE73E792B0801E57B" originalValue="±">+/-</normalizedToken>
0.003) and moderately transverse (WPm/LP 1.26
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="9BBD39B0CDD60D29E845F7B5915F3475" originalValue="±">+/-</normalizedToken>
0.028), with lateral margins almost rectilinearly and slightly constricted posteriorly (WPm/WPp 1.19
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="7AF9DBCBA81405841B47B33D5B3D052A" originalValue="±">+/-</normalizedToken>
0.013). Anterior angles slightly prominent, posterior angles nearly rectangular (95-100°). Width between posterior angles much greater than between anterior angles (WPa/WPp 0.87
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="75155BFE1991163C4DA939A45FE50D20" originalValue="±">+/-</normalizedToken>
0.038). Basal margin slightly concave in middle.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Scutellum</emphasis>
<paragraph id="7BA100A7686FAA5B6220B6D12D1B86C9" pageId="0" pageNumber="63">
<emphasis id="06AFFABF97946594D0E57648EBF38A8F" bold="true" italics="true" pageId="0" pageNumber="63">Scutellum</emphasis>
: Externally visible, triangular, with rounded apex.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Elytra</emphasis>
<paragraph id="4FF71B4D954F88AC478D6A761EF97246" pageId="0" pageNumber="63">
<emphasis id="8D6C24100882EED7A7B9EFB1492EDFDE" bold="true" italics="true" pageId="0" pageNumber="63">Elytra</emphasis>
: Narrowly depressed along suture, of average length (LE/ABL 0.59
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="DA35F8098365E995CBF6C5FBC6988A13" originalValue="±">+/-</normalizedToken>
0.005) and width (WE/LE 0.66
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="59970E3B884186C85152FB067AFD1FCD" originalValue="±">+/-</normalizedToken>
0.026) for genus, with traces of 6-7 striae. Humeri distinct, rounded, in outline forming obtuse angle with longitudinal axis of body. Lateral margins subparallel in middle, slightly convergent at basal fifth, evenly rounded to apex at apical fourth, with shallow subapical sinuation. Basal margination distinct.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Legs</emphasis>
<paragraph id="45C1C8E0859F6ADCF004DBFB76F810AC" pageId="0" pageNumber="63">
<emphasis id="159BCE1462D71448FB01DCD6A87851A5" bold="true" italics="true" pageId="0" pageNumber="63">Legs</emphasis>
: Protarsi of male with moderately dilated tarsomere 1. Profemora moderately swollen. Metafemora unmodified.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Male genitalia</emphasis>
<paragraph id="91EC6B34E83CF9D3068B522FFB7C7909" pageId="0" pageNumber="63">
<emphasis id="257F082C150E678D607CC718DDC0227C" bold="true" italics="true" pageId="0" pageNumber="63">Male genitalia</emphasis>
: Median lobe of aedeagus (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">4A</figureCitation>
<figureCitation id="B3F64AE46EC3F0F293C0E17D29D37C45" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">4A</figureCitation>
) anopic, slightly arcuate and slightly twisted. Basal orifice comparatively short for the genus. Shaft with long subparallel basal part, slightly dilating in apical third. Apical part with enlarged apex in form of rounded parallelogram. Dorsal margin slightly convex and strongly sclerotized at middle. Ventral margin curved near middle, where it is suddenly enlarged right before the apex. Endophallus with dorsal sclerite in form of a semicircular filament-like structure with short basal prolongations. Ventral sclerite located at apical orifice, in form of golf gap wedge plate. Dorsal scaly membranous field present at middle of dorsal sclerite. Enlarged apical area of median lobe with a dark spine-like structure (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">4A</figureCitation>
<figureCitation id="A444BBA0DBA16BE45FEF26534AAB6E21" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">4A</figureCitation>
, ss). Left paramere (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">4B</figureCitation>
<figureCitation id="D7B31123170DA1AE22E699D36AD9D397" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">4B</figureCitation>
) modified, with long, subparallel apical half of moderate width with one seta at angulate tip, basally with strong concave keel (bk), and thick basal processes of different length. Right paramere (Fig.
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<figureCitation id="0C6F4F929315DF71C0DFC5ED59DA7C69" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Digital images and line drawings of male genitalia of Anillinus species. A. relictus (E of Oneonta, Blount County, Alabama) A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect. A. sinuatus (Bexar County, Texas) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect. bk - basal keel, dp - dorsal process, ss - spine-like structure, vs - ventral sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1016.61397.figure4" httpUri="https://binary.pensoft.net/fig/508296" pageId="0" pageNumber="63">4C</figureCitation>
) of moderate length, with eight setiferous pores bearing only three long setae (several others might be broken), which are shorter than length of paramere.
</paragraph>
<paragraph pageId="0" pageNumber="63">
<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Female genitalia</emphasis>
<paragraph id="3FDEDAD122D5A3BA1209379EDE367195" pageId="0" pageNumber="63">
<emphasis id="BF7F7B26587BC234BAE6BC32DA5706CF" bold="true" italics="true" pageId="0" pageNumber="63">Female genitalia</emphasis>
: Spermatheca not investigated. Ovipositor sclerites standard for genus with falciform gonocoxite 2 bearing two ensiform setae. Laterotergite with 8-9 setae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="distribution">
<paragraph pageId="0" pageNumber="63">Geographic distribution.</paragraph>
<paragraph pageId="0" pageNumber="63">
<subSubSection id="D1357A35113D9303235FAA655681A310" pageId="0" pageNumber="63" type="distribution">
<paragraph id="2FEF764C69C2F3C4EDF5D650E42B1E64" pageId="0" pageNumber="63">Geographic distribution.</paragraph>
<paragraph id="A6DA46712603C2BE7079619A916CE3D3" pageId="0" pageNumber="63">
This species is known only from the type locality in Blount County, Alabama (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of the South of eastern United States, showing positions of locality records for the newly described species of Anillinus and the ranges of their presumed relatives (localities of the same color reflect supposed relatedness). A. felicianus, green circle; A. relictus, red circle. Green area with vertical line pattern - range of A. sinuaticollis. Black cross - type locality of A. sinuatus. Red areas with diagonal line pattern - ranges of the species of Anillinus whose males have a spine-like structure in the endophallus of the median lobe (after Sokolov et al. 2004; Sokolov and Watrous 2008; Sokolov 2011; Sokolov et al. 2014; Sokolov et al. 2017). Blue line - Mississippi River. State abbreviations follow Federal Information Processing Standards (https: // www. nlsinfo. org / content / cohorts / nlsy 79 / other-documentation / codebook-supplement / nlsy 79 - attachment- 102 - federal)." figureDoi="10.3897/zookeys.1016.61397.figure5" httpUri="https://binary.pensoft.net/fig/508297" pageId="0" pageNumber="63">5</figureCitation>
<figureCitation id="0AD36356F289A52D6414C9019FB7D890" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of the South of eastern United States, showing positions of locality records for the newly described species of Anillinus and the ranges of their presumed relatives (localities of the same color reflect supposed relatedness). A. felicianus, green circle; A. relictus, red circle. Green area with vertical line pattern - range of A. sinuaticollis. Black cross - type locality of A. sinuatus. Red areas with diagonal line pattern - ranges of the species of Anillinus whose males have a spine-like structure in the endophallus of the median lobe (after Sokolov et al. 2004; Sokolov and Watrous 2008; Sokolov 2011; Sokolov et al. 2014; Sokolov et al. 2017). Blue line - Mississippi River. State abbreviations follow Federal Information Processing Standards (https: // www. nlsinfo. org / content / cohorts / nlsy 79 / other-documentation / codebook-supplement / nlsy 79 - attachment- 102 - federal)." figureDoi="10.3897/zookeys.1016.61397.figure5" httpUri="https://binary.pensoft.net/fig/508297" pageId="0" pageNumber="63">5</figureCitation>
, red circle).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="habitat">
<paragraph pageId="0" pageNumber="63">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="63">The label does not contain any habitat information. Presumably, this species is not a cavernicolous species.</paragraph>
<subSubSection id="9F73E22445071EC78CA73191E42341B8" pageId="0" pageNumber="63" type="habitat">
<paragraph id="7ABA0D1582F312E9B205A3B422468588" pageId="0" pageNumber="63">Habitat.</paragraph>
<paragraph id="62712132988306CFB4E2E505AD4CB102" pageId="0" pageNumber="63">The label does not contain any habitat information. Presumably, this species is not a cavernicolous species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="63" type="relationships">
<paragraph pageId="0" pageNumber="63">Relationships.</paragraph>
<paragraph pageId="0" pageNumber="63">
<subSubSection id="F54B01449058DCF47E89779161AEA1BC" pageId="0" pageNumber="63" type="relationships">
<paragraph id="7DE3E541C24CEAF19AA0B071343130D3" pageId="0" pageNumber="63">Relationships.</paragraph>
<paragraph id="A20CECBCBE355C72229B6AE0C506A140" pageId="0" pageNumber="63">
Based on the structure of the median lobe,
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<emphasis italics="true" pageId="0" pageNumber="63">A. relictus</emphasis>
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<emphasis id="3D3502232D90CF26429E9845A031E0FC" italics="true" pageId="0" pageNumber="63">A. relictus</emphasis>
</taxonomicName>
is a sister species to the endogean
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<emphasis italics="true" pageId="0" pageNumber="63">A. sinuatus</emphasis>
<taxonomicName id="D52728BFDEF540204914369ABC586418" lsidName="A. sinuatus" pageId="0" pageNumber="63" rank="species" species="sinuatus">
<emphasis id="A3DB2C157153ED0BF682F02FC891CDE9" italics="true" pageId="0" pageNumber="63">A. sinuatus</emphasis>
</taxonomicName>
(Jeannel) (
<bibRefCitation author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie" pageId="0" pageNumber="63" pagination="33 - 204" refId="B13" refString="Jeannel, R, 1963a. Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]." volume="28" year="1963 a">Jeannel 1963a</bibRefCitation>
<bibRefCitation id="64D2D3FAACC64067DCC8413F8181BF30" author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie" pageId="0" pageNumber="63" pagination="33 - 204" refId="B13" refString="Jeannel, R, 1963a. Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]." volume="28" year="1963 a">Jeannel 1963a</bibRefCitation>
). The latter species is known to occur in Bexar County, Texas, where it was documented by a small series of three specimens extracted from the soil during surveys in peach orchards (
<bibRefCitation author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie" pageId="0" pageNumber="63" pagination="33 - 204" refId="B13" refString="Jeannel, R, 1963a. Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]." volume="28" year="1963 a">Jeannel 1963a</bibRefCitation>
<bibRefCitation id="CA7B9063F149236D05F2B928AE3CA195" author="Jeannel, R" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie" pageId="0" pageNumber="63" pagination="33 - 204" refId="B13" refString="Jeannel, R, 1963a. Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]. Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie 28: 33 - 204" title="Monographie des '' Anillini '', Bembidiides endoges [Coleoptera Trechidae]." volume="28" year="1963 a">Jeannel 1963a</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.417.7733" author="Sokolov, IM" journalOrPublisher="ZooKeys" pageId="0" pageNumber="63" pagination="71 - 101" refId="B26" refString="Sokolov, IM, Reddell, JR, Kavanaugh, DH, 2014. Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution. ZooKeys 417: 71 - 101, DOI: https://doi.org/10.3897/zookeys.417.7733" title="Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution." url="https://doi.org/10.3897/zookeys.417.7733" volume="417" year="2014">Sokolov et al. 2014</bibRefCitation>
<bibRefCitation id="CFD467F35F1BB2D18CDEC08F9AE413C6" DOI="https://doi.org/10.3897/zookeys.417.7733" author="Sokolov, IM" journalOrPublisher="ZooKeys" pageId="0" pageNumber="63" pagination="71 - 101" refId="B26" refString="Sokolov, IM, Reddell, JR, Kavanaugh, DH, 2014. Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution. ZooKeys 417: 71 - 101, DOI: https://doi.org/10.3897/zookeys.417.7733" title="Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution." url="https://doi.org/10.3897/zookeys.417.7733" volume="417" year="2014">Sokolov et al. 2014</bibRefCitation>
). The range of
<taxonomicName lsidName="A. sinuatus" pageId="0" pageNumber="63" rank="species" species="sinuatus">
<emphasis italics="true" pageId="0" pageNumber="63">A. sinuatus</emphasis>
<taxonomicName id="D12F93EDDCF84AE22F24194A0C135432" lsidName="A. sinuatus" pageId="0" pageNumber="63" rank="species" species="sinuatus">
<emphasis id="5594F070C2AC8DC80743D3FE09732945" italics="true" pageId="0" pageNumber="63">A. sinuatus</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of the South of eastern United States, showing positions of locality records for the newly described species of Anillinus and the ranges of their presumed relatives (localities of the same color reflect supposed relatedness). A. felicianus, green circle; A. relictus, red circle. Green area with vertical line pattern - range of A. sinuaticollis. Black cross - type locality of A. sinuatus. Red areas with diagonal line pattern - ranges of the species of Anillinus whose males have a spine-like structure in the endophallus of the median lobe (after Sokolov et al. 2004; Sokolov and Watrous 2008; Sokolov 2011; Sokolov et al. 2014; Sokolov et al. 2017). Blue line - Mississippi River. State abbreviations follow Federal Information Processing Standards (https: // www. nlsinfo. org / content / cohorts / nlsy 79 / other-documentation / codebook-supplement / nlsy 79 - attachment- 102 - federal)." figureDoi="10.3897/zookeys.1016.61397.figure5" httpUri="https://binary.pensoft.net/fig/508297" pageId="0" pageNumber="63">5</figureCitation>
<figureCitation id="FD5B373A7E947A6016AE57C722808C5C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of the South of eastern United States, showing positions of locality records for the newly described species of Anillinus and the ranges of their presumed relatives (localities of the same color reflect supposed relatedness). A. felicianus, green circle; A. relictus, red circle. Green area with vertical line pattern - range of A. sinuaticollis. Black cross - type locality of A. sinuatus. Red areas with diagonal line pattern - ranges of the species of Anillinus whose males have a spine-like structure in the endophallus of the median lobe (after Sokolov et al. 2004; Sokolov and Watrous 2008; Sokolov 2011; Sokolov et al. 2014; Sokolov et al. 2017). Blue line - Mississippi River. State abbreviations follow Federal Information Processing Standards (https: // www. nlsinfo. org / content / cohorts / nlsy 79 / other-documentation / codebook-supplement / nlsy 79 - attachment- 102 - federal)." figureDoi="10.3897/zookeys.1016.61397.figure5" httpUri="https://binary.pensoft.net/fig/508297" pageId="0" pageNumber="63">5</figureCitation>
, black cross) is situated approximately 770 miles southwest of the type locality of
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<emphasis italics="true" pageId="0" pageNumber="63">A. relictus</emphasis>
<taxonomicName id="EEE9496B0CC473AA92B257CAE70FC9C9" lsidName="A. relictus" pageId="0" pageNumber="63" rank="species" species="relictus">
<emphasis id="3EA52B3C735157FAC3D6B623D1BAD405" italics="true" pageId="0" pageNumber="63">A. relictus</emphasis>
</taxonomicName>
.
</paragraph>

View file

@ -1,194 +1,194 @@
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<subSection pageId="0" pageNumber="145" type="taxa of the agrotis fatidica (hübner, 1824) species-group and the history of their interpretation">
<treatment ID-GBIF-Taxon="182224749" LSID="urn:lsid:zoobank.org:act:C7C684FD-D230-479C-B13D-2E7F83EBAE8F" httpUri="http://treatment.plazi.org/id/58873912A176F4E7D8CE01EB585FF6A5" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="http://zoobank.org/C7C684FD-D230-479C-B13D-2E7F83EBAE8F" authority="Ronkay &amp; Huemer, 2018" authorityName="Ronkay &amp; Huemer" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis mazeli" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mazeli" status="sp. n.">Agrotis mazeli</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figures 3340" captionStartId="F8" captionText="Figures 33 - 40. Agrotis mazeli. 33 - 34 holotype male, France, 35 - 38 paratype males, France, 39 - 40 paratype female, France." figureDoi="10.3897/nl.41.23090.figures33-40" httpUri="https://binary.pensoft.net/fig/213652" pageId="0" pageNumber="145">Figs 33-40</figureCitation>
<figureCitation captionStart="Gen. figs 79" captionStartId="F7" captionText="Gen. figs 7 - 9. 7 Agrotis mayrorum sp. n., male genitalia, &quot; Corsica &quot;, 8 - 9 Agrotis mazeli sp. n., male genitalia, 8 holotype, 9 paratype." figureDoi="10.3897/nl.41.23090.genfigs7-9" httpUri="https://binary.pensoft.net/fig/213651" pageId="0" pageNumber="145">; Gen. figs 8-9</figureCitation>
<figureCitation captionStart="Gen. figs 1618" captionStartId="F13" captionText="Gen. figs 16 - 18. 16 Agrotis fatidica, female genitalia, Austria, 17 Agrotis mayrorum sp. n., paratype, female genitalia, France, 18 Agrotis mazeli sp. n., paratype, female genitalia, France." figureDoi="10.3897/nl.41.23090.genfigs16-18" httpUri="https://binary.pensoft.net/fig/213657" pageId="0" pageNumber="145">, 18</figureCitation>
<subSection id="286EDD4C1F7AF54EF1D666EAE8CC20F7" pageId="0" pageNumber="145" type="taxa of the agrotis fatidica (hübner, 1824) species-group and the history of their interpretation">
<treatment id="58873912A176F4E7D8CE01EB585FF6A5" ID-GBIF-Taxon="182224749" LSID="urn:lsid:zoobank.org:act:C7C684FD-D230-479C-B13D-2E7F83EBAE8F" httpUri="http://treatment.plazi.org/id/58873912A176F4E7D8CE01EB585FF6A5" lastPageNumber="145" pageId="0" pageNumber="145" scope_family="Noctuidae" scope_order="Lepidoptera">
<subSubSection id="5DC13A7404E773652A801F2A37E1003D" pageId="0" pageNumber="145" type="nomenclature">
<paragraph id="7513C407883979B7852BEBBB5B09790A" pageId="0" pageNumber="145">
<taxonomicName id="9F84B42454D7C7405F0A59B79D908AA8" LSID="http://zoobank.org/C7C684FD-D230-479C-B13D-2E7F83EBAE8F" authority="Ronkay &amp; Huemer, 2018" authorityName="Ronkay &amp; Huemer" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis mazeli" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mazeli" status="sp. n.">Agrotis mazeli</taxonomicName>
<taxonomicNameLabel id="5F1ED08257323442CADCF269D0F38EFE" pageId="0" pageNumber="145">sp. n.</taxonomicNameLabel>
<figureCitation id="42E8C7A490F4309F1609C07931EE9C6D" captionStart="Figures 3340" captionStartId="F8" captionText="Figures 33 - 40. Agrotis mazeli. 33 - 34 holotype male, France, 35 - 38 paratype males, France, 39 - 40 paratype female, France." figureDoi="10.3897/nl.41.23090.figures33-40" httpUri="https://binary.pensoft.net/fig/213652" pageId="0" pageNumber="145">Figs 33-40</figureCitation>
<figureCitation id="AA2B968CEA805B92D5D38D71890E7A2F" captionStart="Gen. figs 79" captionStartId="F7" captionText="Gen. figs 7 - 9. 7 Agrotis mayrorum sp. n., male genitalia, &quot; Corsica &quot;, 8 - 9 Agrotis mazeli sp. n., male genitalia, 8 holotype, 9 paratype." figureDoi="10.3897/nl.41.23090.genfigs7-9" httpUri="https://binary.pensoft.net/fig/213651" pageId="0" pageNumber="145">; Gen. figs 8-9</figureCitation>
<figureCitation id="5851EB13BBD7D4A09CA2CAF65091CA11" captionStart="Gen. figs 1618" captionStartId="F13" captionText="Gen. figs 16 - 18. 16 Agrotis fatidica, female genitalia, Austria, 17 Agrotis mayrorum sp. n., paratype, female genitalia, France, 18 Agrotis mazeli sp. n., paratype, female genitalia, France." figureDoi="10.3897/nl.41.23090.genfigs16-18" httpUri="https://binary.pensoft.net/fig/213657" pageId="0" pageNumber="145">, 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="holotype">
<paragraph pageId="0" pageNumber="145">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="145">Male, France, Canigou, Pla Guillem, 2280 m, 8.viii.1985, leg. G. Lutran; slide No. RL11749m (TLMF).</paragraph>
<subSubSection id="81C3EFF756232A6D4842A74E6E5831B1" pageId="0" pageNumber="145" type="holotype">
<paragraph id="64F35C1FC2F8FB813B8808073C30C42F" pageId="0" pageNumber="145">Holotype.</paragraph>
<paragraph id="561A1246EB575244C06C777767632136" pageId="0" pageNumber="145">Male, France, Canigou, Pla Guillem, 2280 m, 8.viii.1985, leg. G. Lutran; slide No. RL11749m (TLMF).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="paratypes">
<paragraph pageId="0" pageNumber="145">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">France</emphasis>
<subSubSection id="B330BEADC6400DC6AD25812C4195DA9C" pageId="0" pageNumber="145" type="paratypes">
<paragraph id="B1B74790A4B68239670894C6B6FEDF1B" pageId="0" pageNumber="145">Paratypes.</paragraph>
<paragraph id="BC66F469AFA13D2A265D49DCC0603DC5" pageId="0" pageNumber="145">
<emphasis id="22CBE75131EF82DEEE5CCAD9D165934A" bold="true" pageId="0" pageNumber="145">France</emphasis>
. 4 males, French Pyrenees orientales, Esquerdes de Rotja, 2200 m, 4.viii.1994, S. Peslier leg.; slide No. RL11311m, DNA Barcode ID TLMF Lep 14494 (TLMF); 1 male, Pyrenees orientales, Pla Guillem, 2.viii.1994, R. Mazel leg., DNA Barcode ID TLMF 14495 (TLMF); 1 female, Pyrenees orientales, Pla Guillem, 8.viii.1994, R. Mazel leg.; slide No. RL11312f (TLMF); 1 male, Pyrenees orientales, Pla de la Begude, 2300 m, 10.viii.1999, R. Mazel leg., DNA Barcode ID TLMF 14496 (TLMF); 1 male, &quot;la Vernet 1879,
<normalizedToken originalValue="Vallée">Vallee</normalizedToken>
<normalizedToken originalValue="Lipaudière">Lipaudiere</normalizedToken>
<normalizedToken id="F02454FFE4129FBFF892CB3D6BED8228" originalValue="Vallée">Vallee</normalizedToken>
<normalizedToken id="3D6771FB7FE5AFD71AE484CEC2346C43" originalValue="Lipaudière">Lipaudiere</normalizedToken>
, ex coll. Dr. Struve, Leipzig, coll.
<normalizedToken originalValue="Püngeler”">Puengeler&quot;</normalizedToken>
<normalizedToken id="8EE645B5DE74013EB45464C3E0453687" originalValue="Püngeler”">Puengeler&quot;</normalizedToken>
(see also
<bibRefCitation author="Struve, O" journalOrPublisher="Insect systematics &amp; evolution" pageId="0" pageNumber="145" refId="B52" refString="Struve, O, 1882. Drei Sommer in den Pyrenaeen. Entomologische Zeitung Stettin 1882: 393-405, 410-429." title="Drei Sommer in den Pyrenaeen. Entomologische Zeitung Stettin 1882: 393 - 405, 410 - 429." year="1882">Struve 1882</bibRefCitation>
<bibRefCitation id="316BE30A57C1AB6404F13FFE7644D735" author="Struve, O" journalOrPublisher="Insect systematics &amp; evolution" pageId="0" pageNumber="145" refId="B52" refString="Struve, O, 1882. Drei Sommer in den Pyrenaeen. Entomologische Zeitung Stettin 1882: 393-405, 410-429." title="Drei Sommer in den Pyrenaeen. Entomologische Zeitung Stettin 1882: 393 - 405, 410 - 429." year="1882">Struve 1882</bibRefCitation>
) (ZMHU).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="etymology">
<paragraph pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="145">Named in honour of Robert Mazel who collected most of the type-series and generously supported our work with this important material.</paragraph>
<subSubSection id="65095F6BA1A9FF42D6234EEB0DC90CA3" pageId="0" pageNumber="145" type="etymology">
<paragraph id="50049D54693F14C5171C26614F288F2F" pageId="0" pageNumber="145">Etymology.</paragraph>
<paragraph id="F505C25B7A7CF860AD33192015405205" pageId="0" pageNumber="145">Named in honour of Robert Mazel who collected most of the type-series and generously supported our work with this important material.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="CE6803C7F3B19BA855837E18BB266C5F" pageId="0" pageNumber="145" type="diagnosis">
<paragraph id="76158FA554171FD249E667413021BBE2" pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph id="E5C131FEDF3091BA285EC3DCCB773A95" pageId="0" pageNumber="145">
The new species is the allopatric sister species of
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
<taxonomicName id="44B54DCFB1995C0E90E9F6559738A5A2" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis id="93B7AB72418A96E7324AADF4D2B3E36C" italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
</taxonomicName>
, sharing the group features in the male antenna (longer and finer branches than in the
<emphasis italics="true" pageId="0" pageNumber="145">fatidica</emphasis>
<emphasis id="AC6556920AC5884834511E45ABF12E29" italics="true" pageId="0" pageNumber="145">fatidica</emphasis>
-line), and the white (or whitish) hindwing. The two species can be distinguished by a series of external morphological features which are recognisable in almost all specimens. These differences are ca as large as those between
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
<taxonomicName id="C8FCD2D8B60B2BC442EEEC9864F5A225" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis id="6EBE8726DA369F356398322D770ED6FF" italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
<taxonomicName id="DD94B15A6ED2E51C3A4E96A8EB4559CB" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis id="E87ED3AF2D7CE7C2FF3B911A10F4CB70" italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Ronkay &amp; Huemer" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis mazeli" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mazeli">
<emphasis italics="true" pageId="0" pageNumber="145">Agrotis mazeli</emphasis>
<paragraph id="7230A6FB5197982FB6844D078210B85B" pageId="0" pageNumber="145">
<taxonomicName id="93A4C047D776CFB2BBF6D52AED46A2B5" authorityName="Ronkay &amp; Huemer" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis mazeli" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mazeli">
<emphasis id="A245B460C117AB1A31B14B5A0A8A80DD" italics="true" pageId="0" pageNumber="145">Agrotis mazeli</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
<taxonomicName id="D6B64C64268A59E4E2AB7C9E30BA49A8" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis id="161BAE91238E35032C4B4D382A7B5BC1" italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
</taxonomicName>
by its somewhat narrower and more pointed forewings, which are also narrower and apically more pointed than those of
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
<taxonomicName id="3FA06903030A14CF9C478EE63C486B31" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis id="A31CA1D029CE5C6D7819732CBE9E8067" italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
</taxonomicName>
but broader than those of
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. proverai" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proverai">
<emphasis italics="true" pageId="0" pageNumber="145">A. proverai</emphasis>
<taxonomicName id="4C2BD1568E073772D6A5C0E1847089D1" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. proverai" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="proverai">
<emphasis id="083AC744EC96CD5004F66734B0A82E02" italics="true" pageId="0" pageNumber="145">A. proverai</emphasis>
</taxonomicName>
, a more sharply defined postmedial line with a clearly visible deep curve inwards at costal margin, on average smaller and more darkened orbicular and reniform stigmata, smaller and finer dark dots at terminal line, which are practically missing from the underside, and the smoothly, rather uniformly darkened forewing underside.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="2443645FBB10CB29ACA88205A3E309C5" pageId="0" pageNumber="145">
In the genitalia, the males of
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mazeli" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mazeli">
<emphasis italics="true" pageId="0" pageNumber="145">A. mazeli</emphasis>
<taxonomicName id="EB7A9999FEF3E6C4FB003802591172C3" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mazeli" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mazeli">
<emphasis id="D6E0852105B58FDB4038D89B010567FD" italics="true" pageId="0" pageNumber="145">A. mazeli</emphasis>
</taxonomicName>
can be distinguished from those of the closely related taxa by the short and basally broad valvae with the very typical, wide membranous ventral marginal section bordering the saccular part; the broad cucullus with finely pointed apex; the rather long and slender claspers; the comparatively short vinculum; the fine eversible bar of the ventral carina and the weak, dentated subbasal plate of the vesica; the rather short and obtusely conical dorso-lateral diverticulum of the inflated basal section of vesica; and the entire length of the vesica is somewhat shorter (in comparison with the total length of phallus) than in
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
<taxonomicName id="253F456CC09AF5C6C52013FD62D1A057" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis id="0F3C620F85E84681515E513733D06719" italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
<taxonomicName id="27F74BBEE7586BC1EF1182AB03A71CAC" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis id="72E429F5319D56A066B53BD62833EB70" italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<paragraph id="47D1762145753B8F6C6ADD82EBC74534" pageId="0" pageNumber="145">
In the female genitalia, the apophyses anteriores are longer and the appendix bursae is somewhat shorter than in
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
<taxonomicName id="DFE4C009BBD3BFD7B3E684D3C3EB1353" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis id="73961171AD9C0CD0D75823D1F2F2B158" italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
</taxonomicName>
; the ovipositor and the ductus bursae are proportionally shorter, and the appendix bursae is shorter than in
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
<taxonomicName id="BE7C3A871359087BBA897137147565D1" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis id="21937746B145776599180150F63F0911" italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">Description.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="7553C65FA6698CE3F1A37A45BE29C7F0" pageId="0" pageNumber="145" type="description">
<paragraph id="D53B55D41840102CD44155A234EC18B9" pageId="0" pageNumber="145">Description.</paragraph>
<paragraph id="0E11CF60DAB68E907475497F713E5CE2" pageId="0" pageNumber="145">
Wingspan: males 40-41 mm, female 32 mm. The main external features are typical of the
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
<taxonomicName id="8D9CA9CE93112F36570F0B3ADC3948A0" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis id="0946F241D1A73ED7ACAC6A2FDC60EFAB" italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
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species-complex. Forewing ground colour pale greyish-brown or slate-grey, with darker grey and brown irroration; costa and veins darker than ground colour and defined broadly with paler ochreous-grey intervenal stripes. Antemedial line variably but strongly marked, partly obsolete; median fascia absent; median area usually less variegated than in the other related species. Postmedial line recognisable though diffuse, pale ochreous-grey, and defined by a row of dark dots; its upper part curved inwards rather strongly around reniform stigma. Subterminal line obsolescent, ochreous-grey, defined by a row of conspicuous intervenal chevron-spots; terminal line fine, black(ish), with weak or indistinct dark dots; cilia ochreous-grey, with variably strong brown midline. Stigmata well-defined, medium-sized or relatively small, outlined by black or dark blackish-brown; claviform stigma long, dark, blackish-brown with only a slightly paler filling; orbicular stigma rather small, usually flattened-elliptical (drop-shaped) with pointed posterior edge and darker brown centre. Reniform stigma medium-sized, elliptical or lunulate, with more or less diffuse dark grey-brown outline and filling (which is stronger than in
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
<taxonomicName id="B70CB51F1401D36FBFC6734A51CE7EA0" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis id="F2C83CDC6543961E091497C3876A20DE" italics="true" pageId="0" pageNumber="145">A. mayrorum</emphasis>
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); intracellular dark patch between stigmata blackish-brown or blackish.
</paragraph>
<paragraph pageId="0" pageNumber="145">Hindwing milky-white or light whitish-grey with fine ochreous shade, costal area, termen and diffuse marginal stripe with ochreous-brown or greyish suffusion of variable strength; veins darkened. Discal spot rounded or somewhat lunulate, often shadow-like; terminal line more or less continuous, brownish; cilia whitish with some pale brownish shading. Forewing underside rather unicolorous grey, suffused with fuscous, discal dot and postmedial line obsolescent or missing. Hindwing underside whitish with darker veins, reduced transversal line and relatively well-defined discal spot; terminal line fine, brown, without or with only very small darker dots.</paragraph>
<paragraph id="6372BB484EC5DDFC6E62254DFB30C2E2" pageId="0" pageNumber="145">Hindwing milky-white or light whitish-grey with fine ochreous shade, costal area, termen and diffuse marginal stripe with ochreous-brown or greyish suffusion of variable strength; veins darkened. Discal spot rounded or somewhat lunulate, often shadow-like; terminal line more or less continuous, brownish; cilia whitish with some pale brownish shading. Forewing underside rather unicolorous grey, suffused with fuscous, discal dot and postmedial line obsolescent or missing. Hindwing underside whitish with darker veins, reduced transversal line and relatively well-defined discal spot; terminal line fine, brown, without or with only very small darker dots.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="molecular data">
<paragraph pageId="0" pageNumber="145">Molecular data.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="DF84F7E12A75695B9B19B65237F77D7B" pageId="0" pageNumber="145" type="molecular data">
<paragraph id="AC21C71ADBF17993EF6EE76F615E8FCD" pageId="0" pageNumber="145">Molecular data.</paragraph>
<paragraph id="A17DC59F3AB7ACF1D65FCC273D18A2E8" pageId="0" pageNumber="145">
BIN URI: BOLD: ACR9899. The intraspecific average and maximum distances of the barcode region are 0% (p-dist) (n=3), however intraspecific divergence may increase with additional samples from the Pyrenees. The minimum distance to the nearest neighbour
<taxonomicName authorityName="Ronkay &amp; Huemer" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis italics="true" pageId="0" pageNumber="145">Agrotis mayrorum</emphasis>
<taxonomicName id="AA9E1E2E12E418227AD17C15005B24FF" authorityName="Ronkay &amp; Huemer" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis mayrorum" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="mayrorum">
<emphasis id="9167E0ACE16562DBEBDEDB83AFF2E1BA" italics="true" pageId="0" pageNumber="145">Agrotis mayrorum</emphasis>
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is 1.12%, whereas the minimum distance to
<taxonomicName class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis italics="true" pageId="0" pageNumber="145">Agrotis fatidica</emphasis>
<taxonomicName id="A71F9E3AA4C54FDCEDC14C68CF1B727A" class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis id="EE7F8AA887927ABDAB9CB46DA8946021" italics="true" pageId="0" pageNumber="145">Agrotis fatidica</emphasis>
</taxonomicName>
is 2.18%.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="bionomics">
<paragraph pageId="0" pageNumber="145">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="145">Host-plant and early stages are unknown. The adults fly relatively late in the season, the specimens of the type series were collected in early August. Most of the specimens were obtained by light trapping using UV tubes and mercury vapour lamp. Adult records are restricted to alpine meadows and scree at high altitudes above the timberline (2200-2300 m a.s.l.).</paragraph>
<subSubSection id="AA7C5CA51D73DA8D25D3614CB3460885" pageId="0" pageNumber="145" type="bionomics">
<paragraph id="256772D6BB4EB02E0B5A285D6DAEF6F6" pageId="0" pageNumber="145">Bionomics.</paragraph>
<paragraph id="5C5FFEC67F505FB4FBB0ADDCAE6EAD91" pageId="0" pageNumber="145">Host-plant and early stages are unknown. The adults fly relatively late in the season, the specimens of the type series were collected in early August. Most of the specimens were obtained by light trapping using UV tubes and mercury vapour lamp. Adult records are restricted to alpine meadows and scree at high altitudes above the timberline (2200-2300 m a.s.l.).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
<subSubSection id="FDEE060730B8AED1376A54AD61ADFD4B" pageId="0" pageNumber="145" type="distribution">
<paragraph id="363A881BB466A081652A0D84A324A301" pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph id="2EE7E302273643E3303A006C56CBF26C" pageId="0" pageNumber="145">
The new species is known from the eastern Pyrenees in France and Spain. The type-series originates from the French side of the Pyrenees; the occurrence of the species on the Spanish side is documented by
<bibRefCitation author="Redondo, V" journalOrPublisher="Prames, Zaragoza" pageId="0" pageNumber="145" refId="B50" refString="Redondo, V, Gaston, J, Vicente, JC, 2010. Las Mariposas de Espana Peninsular. Manual Ilustrado de las Especies Diurnas y Nocturnas. Prames, Zaragoza" title="Las Mariposas de Espana Peninsular. Manual Ilustrado de las Especies Diurnas y Nocturnas." year="2010">Redondo et al. (2010</bibRefCitation>
<bibRefCitation id="B1A5EAE2AB1EC6B4941693E0597F9407" author="Redondo, V" journalOrPublisher="Prames, Zaragoza" pageId="0" pageNumber="145" refId="B50" refString="Redondo, V, Gaston, J, Vicente, JC, 2010. Las Mariposas de Espana Peninsular. Manual Ilustrado de las Especies Diurnas y Nocturnas. Prames, Zaragoza" title="Las Mariposas de Espana Peninsular. Manual Ilustrado de las Especies Diurnas y Nocturnas." year="2010">Redondo et al. (2010</bibRefCitation>
, under the name
<taxonomicName class="Insecta" genus="A." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
<taxonomicName id="47DD81DD3F859370DC0BDE8602D79056" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. fatidica" order="Lepidoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="fatidica">
<emphasis id="B60F148A2C00F95636C623D89CBD4756" italics="true" pageId="0" pageNumber="145">A. fatidica</emphasis>
</taxonomicName>
).
</paragraph>

View file

@ -1,189 +1,189 @@
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<paragraph pageId="0" pageNumber="551">
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<paragraph id="8AB17E94821A6BC470CBDF672CA2406E" pageId="0" pageNumber="551">Reference.</paragraph>
<paragraph id="C476F3C877576A1689E6D18B2676DC00" pageId="0" pageNumber="551">
<bibRefCitation id="56B15DA6FB0832553B2E6A019A2F278B" author="Grichanov, IYa" journalOrPublisher="Nature Conservation Research" pageId="0" pageNumber="551" pagination="23 - 31" refId="B5" refString="Grichanov, IYa, Khruleva, OA, 2018. Fauna and ecology of Dolichopodidae (Diptera) from Wrangel Island Nature Reserve (Chukotka AD, Russia). Nature Conservation Research 3 (3): 23 - 31" title="Fauna and ecology of Dolichopodidae (Diptera) from Wrangel Island Nature Reserve (Chukotka AD, Russia)." volume="3" year="2018">Grichanov and Khruleva 2018</bibRefCitation>
: 40, Fig.
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<figureCitation id="2BA50845D86A497255293A35193904B3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Rogers Bay environs. Middle part of gravel-loamy foothill ridge with spotted willow-sedge-moss cover; a site where Hydrophorus alpinus was collected. Photo by O. A. Khruleva." httpUri="https://binary.pensoft.net/fig/474276" pageId="0" pageNumber="551">5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="551" type="materials_examined">
<paragraph pageId="0" pageNumber="551">Material.</paragraph>
<paragraph pageId="0" pageNumber="551">
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<specimenCount type="male">1♂</specimenCount>
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<paragraph id="1D11EC13D486A35B1D455E86B7F6E112" pageId="0" pageNumber="551">Material.</paragraph>
<paragraph id="8367AEA41D31AAB2D17C5371ED8480ED" pageId="0" pageNumber="551">
<materialsCitation id="E8838097BB1A3991104BBF61CEF198A9" collectingDate="2016-07-31" collectorName="L. F. Volkova" latitude="70.95" location="Thomas Mt." longLatPrecision="973" longitude="178.71666" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
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,
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,
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,
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178°43'
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178°43'
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, gravel southern slope with spotted grassy-forb and herb-dwarf willow-moss cover, 11-
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, gravel southern slope with spotted grassy-forb and herb-dwarf willow-moss cover, 11-
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,
<collectorName>L.F. Volkova</collectorName>
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</materialsCitation>
;
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<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="3A54BF6F644651BDDF72C021BDF945D1" collectingDate="2016-07-17" collectorName="L. F. Volkova" latitude="71.28253" location="Krasnyi Flag River" longLatPrecision="1" longitude="-178.8359" specimenCount="1" specimenCount-male="1">
<specimenCount id="4A7640D1C5C3B713BC45EC134BE7C427" type="male">1♂</specimenCount>
, middle-upper reaches of
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<location id="C159703826F33B4661E299467CD64CCC" LSID="urn:lsid:plazi:treatment:58A02AFFACD15820BA3ADF3D5FFCD815:C159703826F33B4661E299467CD64CCC" latitude="71.28253" longLatPrecision="1" longitude="-178.8359" name="Krasnyi Flag River">Krasnyi Flag River</location>
,
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, wet plot in a river valley with continuous grass-dryad-sedge-moss hummocks, 4-
<collectingDate value="2016-07-17">17.07.2016</collectingDate>
<geoCoordinate id="AF553013FF6397B906E7371DA646B171" degrees="71" direction="north" minutes="16.952" orientation="latitude" precision="1" value="71.28253">71°16.952'N</geoCoordinate>
<geoCoordinate id="C48E4CBECA21EB27B94CBA39676E6642" degrees="178" direction="west" minutes="50.154" orientation="longitude" precision="1" value="-178.8359">178°50.154'W</geoCoordinate>
, wet plot in a river valley with continuous grass-dryad-sedge-moss hummocks, 4-
<collectingDate id="65CFEB54ECA9A6F6A8F0BB9B47F38D75" value="2016-07-17">17.07.2016</collectingDate>
,
<collectorName>L.F. Volkova</collectorName>
<collectorName id="3060DAE05A768EC91408E886F44AF97B">L.F. Volkova</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2019-07-09" collectingMethod="yellow pan traps" collectorName="U. V. Babiy" latitude="71.29776" location="Tundrovaya Mt." longLatPrecision="1" longitude="-179.79578" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="4D731918FEC9A5B871A90C67602AC350" collectingDate="2019-07-09" collectingMethod="yellow pan traps" collectorName="U. V. Babiy" latitude="71.29776" location="Tundrovaya Mt." longLatPrecision="1" longitude="-179.79578" specimenCount="1" specimenCount-male="1">
<specimenCount id="F6C58530D0BF5D9C6F1DFAEAAA2D3295" type="male">1♂</specimenCount>
, environs of
<location LSID="urn:lsid:plazi:treatment:58A02AFFACD15820BA3ADF3D5FFCD815:616DAF0A28521A5270225DC29229826D" latitude="71.29776" longLatPrecision="1" longitude="-179.79578" name="Tundrovaya Mt.">Tundrovaya Mt.</location>
<location id="616DAF0A28521A5270225DC29229826D" LSID="urn:lsid:plazi:treatment:58A02AFFACD15820BA3ADF3D5FFCD815:616DAF0A28521A5270225DC29229826D" latitude="71.29776" longLatPrecision="1" longitude="-179.79578" name="Tundrovaya Mt.">Tundrovaya Mt.</location>
,
<geoCoordinate degrees="71.29776" direction="north" orientation="latitude" precision="1" value="71.29776">71.29776N</geoCoordinate>
<geoCoordinate id="D3486CE54D8DD14A895FBA8CE6EAAA69" degrees="71.29776" direction="north" orientation="latitude" precision="1" value="71.29776">71.29776N</geoCoordinate>
,
<geoCoordinate degrees="179.79578" direction="west" orientation="longitude" precision="1" value="-179.79578">179.79578W</geoCoordinate>
<geoCoordinate id="A1168C96A5E02979CD2386E91FD38D51" degrees="179.79578" direction="west" orientation="longitude" precision="1" value="-179.79578">179.79578W</geoCoordinate>
, loamy high shore with a sparse grass and wormwood cover,
<collectingMethod>yellow pan traps</collectingMethod>
, 6-
<collectingDate value="2019-07-09">9.07.2019</collectingDate>
<collectingMethod id="0E7BBC3C78ABBAE93432EC7602F0A28E">yellow pan traps</collectingMethod>
, 6-
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,
<collectorName>U.V. Babiy</collectorName>
<collectorName id="8068867C81DA4A947D74F1450E4113D6">U.V. Babiy</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2019-07-26" collectorName="O. A. Khruleva" latitude="70.98783" location="Chertov Ovrag" longLatPrecision="1" longitude="-179.84161" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1♀</specimenCount>
<materialsCitation id="5936D48234837EAE2817D84FB53A1EB9" collectingDate="2019-07-26" collectorName="O. A. Khruleva" latitude="70.98783" location="Chertov Ovrag" longLatPrecision="1" longitude="-179.84161" specimenCount="1" specimenCount-female="1">
<specimenCount id="9BABAFFB559FDD00ED9C3D536ADDCA63" type="female">1♀</specimenCount>
,
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<location id="4CF34D8DB7491B83650C3F7C590373EF" LSID="urn:lsid:plazi:treatment:58A02AFFACD15820BA3ADF3D5FFCD815:4CF34D8DB7491B83650C3F7C590373EF" latitude="70.98783" longLatPrecision="1" longitude="-179.84161" name="Chertov Ovrag">Chertov Ovrag</location>
,
<geoCoordinate degrees="70.987833" direction="north" orientation="latitude" precision="1" value="70.98783">70.987833°N</geoCoordinate>
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,
<geoCoordinate degrees="179.841611" direction="west" orientation="longitude" precision="1" value="-179.84161">179.841611°W</geoCoordinate>
<geoCoordinate id="E292EA3161444C7E2B9F8E53738F1927" degrees="179.841611" direction="west" orientation="longitude" precision="1" value="-179.84161">179.841611°W</geoCoordinate>
, the lower part of the loamy-gravelly slope of the southern exposure with a spotted forb-moss-willow-dryad cover, sweeping,
<collectingDate value="2019-07-26">26.07.2019</collectingDate>
<collectingDate id="B502FC48B759606386F018D78B9DC4CB" value="2019-07-26">26.07.2019</collectingDate>
,
<collectorName>O.A. Khruleva</collectorName>
<collectorName id="8707B778165EB3F1A6E92801F94C9070">O.A. Khruleva</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2019-07-24" collectorName="O. A. Khruleva" latitude="71.00389" location="Vyuchnyi" longLatPrecision="1" longitude="-179.70367" specimenCount="7" specimenCount-female="4" specimenCount-male="3">
<specimenCount type="male">3♂♂</specimenCount>
<materialsCitation id="A5AE93555AF2516FEADBFFB7A8EAC8EF" collectingDate="2019-07-24" collectorName="O. A. Khruleva" latitude="71.00389" location="Vyuchnyi" longLatPrecision="1" longitude="-179.70367" specimenCount="7" specimenCount-female="4" specimenCount-male="3">
<specimenCount id="FC35C97EF2F1C4245BE87E96B0446180" type="male">3♂♂</specimenCount>
,
<specimenCount type="female">4♀♀</specimenCount>
<specimenCount id="A989A5C45AE68A27F05D3F456ACFB435" type="female">4♀♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:58A02AFFACD15820BA3ADF3D5FFCD815:15544FE41B2F6E543D293E1F2985F3F1" latitude="71.00389" longLatPrecision="1" longitude="-179.70367" name="Vyuchnyi">Vyuchnyi</location>
<location id="15544FE41B2F6E543D293E1F2985F3F1" LSID="urn:lsid:plazi:treatment:58A02AFFACD15820BA3ADF3D5FFCD815:15544FE41B2F6E543D293E1F2985F3F1" latitude="71.00389" longLatPrecision="1" longitude="-179.70367" name="Vyuchnyi">Vyuchnyi</location>
brook,
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<geoCoordinate id="BCD563F40599675889CBB8DD91FFAFBC" degrees="71.003889" direction="north" orientation="latitude" precision="1" value="71.00389">71.003889°N</geoCoordinate>
,
<geoCoordinate degrees="179.703667" direction="west" orientation="longitude" precision="1" value="-179.70367">179.703667°W</geoCoordinate>
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, base and southern slope of a hill with willow-sedge-moss-dryad hummocky cover and spotted lichen-moss-herb-sedge cover,
<collectingDate value="2019-07-24">24.07.2019</collectingDate>
<collectingDate id="B437B4981A98978A6F695C5BE70C9F2E" value="2019-07-24">24.07.2019</collectingDate>
, sweeping and hand collection,
<collectorName>O.A. Khruleva</collectorName>
<collectorName id="C7C1224C0DC4F9162FA88E31BF9426F2">O.A. Khruleva</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2019-07-11" collectorName="O. A. Khruleva" latitude="70.98478" location="Rogers Bay" longLatPrecision="1" longitude="-178.49414" specimenCount="2" specimenCount-male="2">
<specimenCount type="male">2♂♂</specimenCount>
<materialsCitation id="D5A358A1AFCD7ECEA6309F59E092DA49" collectingDate="2019-07-11" collectorName="O. A. Khruleva" latitude="70.98478" location="Rogers Bay" longLatPrecision="1" longitude="-178.49414" specimenCount="2" specimenCount-male="2">
<specimenCount id="E316A5D34DEDE98D69CB155CE99F4B8D" type="male">2♂♂</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:58A02AFFACD15820BA3ADF3D5FFCD815:A52509BF74FD2492727B100A4EAF74C0" latitude="70.98478" longLatPrecision="1" longitude="-178.49414" name="Rogers Bay">Rogers Bay</location>
<location id="A52509BF74FD2492727B100A4EAF74C0" LSID="urn:lsid:plazi:treatment:58A02AFFACD15820BA3ADF3D5FFCD815:A52509BF74FD2492727B100A4EAF74C0" latitude="70.98478" longLatPrecision="1" longitude="-178.49414" name="Rogers Bay">Rogers Bay</location>
env.,
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<geoCoordinate id="F31CCEE508BD0576CB876E42264F9925" degrees="70.984778" direction="north" orientation="latitude" precision="1" value="70.98478">70.984778°N</geoCoordinate>
,
<geoCoordinate degrees="178.494139" direction="west" orientation="longitude" precision="1" value="-178.49414">178.494139°W</geoCoordinate>
<geoCoordinate id="CF87FC863757DA8C8C868A6AED1A7657" degrees="178.494139" direction="west" orientation="longitude" precision="1" value="-178.49414">178.494139°W</geoCoordinate>
, moderately moistened middle part of gravel-loamy foothill ridge with spotted willow-sedge-moss cover, sweeping,
<collectingDate value="2019-07-11">11.07.2019</collectingDate>
<collectingDate id="C571F0A083E252C308DB980CDDD877A9" value="2019-07-11">11.07.2019</collectingDate>
,
<collectorName>O.A. Khruleva</collectorName>
<collectorName id="216DBF9FEDC08DE153BB3E425A4B2022">O.A. Khruleva</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="551" type="notes. hydrophorus alpinus">
<paragraph pageId="0" pageNumber="551">
<subSubSection id="0B820741A9263EED4439FE72ECA1ACE1" pageId="0" pageNumber="551" type="notes. hydrophorus alpinus">
<paragraph id="3ADBA16635705053B1D1DEDDE57BC7F4" pageId="0" pageNumber="551">
Notes.
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<emphasis italics="true" pageId="0" pageNumber="551">Hydrophorus alpinus</emphasis>
<taxonomicName id="9F4728CBF7D7C54F15D913B8705DF610" authorityName="Wahlberg" authorityYear="1844" class="Insecta" family="Dolichopodidae" genus="Hydrophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrophorus alpinus" order="Diptera" pageId="0" pageNumber="551" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="3599E1260906C7635FAB201BC9F8B7D4" italics="true" pageId="0" pageNumber="551">Hydrophorus alpinus</emphasis>
</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="551">
<paragraph id="B20734BA6CF4B3B45FECF4A139322010" pageId="0" pageNumber="551">
is the commonest species on the Island, but previously only one specimen has been collected outside the warmest central part (
<bibRefCitation author="Grichanov, IYa" journalOrPublisher="Nature Conservation Research" pageId="0" pageNumber="551" pagination="23 - 31" refId="B5" refString="Grichanov, IYa, Khruleva, OA, 2018. Fauna and ecology of Dolichopodidae (Diptera) from Wrangel Island Nature Reserve (Chukotka AD, Russia). Nature Conservation Research 3 (3): 23 - 31" title="Fauna and ecology of Dolichopodidae (Diptera) from Wrangel Island Nature Reserve (Chukotka AD, Russia)." volume="3" year="2018">Grichanov and Khruleva 2018</bibRefCitation>
<bibRefCitation id="52C7C19DBAF6EB8C6B6D073DAA2E1537" author="Grichanov, IYa" journalOrPublisher="Nature Conservation Research" pageId="0" pageNumber="551" pagination="23 - 31" refId="B5" refString="Grichanov, IYa, Khruleva, OA, 2018. Fauna and ecology of Dolichopodidae (Diptera) from Wrangel Island Nature Reserve (Chukotka AD, Russia). Nature Conservation Research 3 (3): 23 - 31" title="Fauna and ecology of Dolichopodidae (Diptera) from Wrangel Island Nature Reserve (Chukotka AD, Russia)." volume="3" year="2018">Grichanov and Khruleva 2018</bibRefCitation>
). In 2016 and 2019, it was discovered in six new plots (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Vyuchnyi brook. Southern gravelly slope of a hill with a &quot; tundra-steppe &quot; plant community; a site where Dolichopus terminasianae and Hydrophorus alpinus were collected. Photo by O. A. Khruleva." httpUri="https://binary.pensoft.net/fig/474274" pageId="0" pageNumber="551">3</figureCitation>
<figureCitation id="BDE49C1BE9F60E5CDAE78B6E8AE4C91F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Vyuchnyi brook. Southern gravelly slope of a hill with a &quot; tundra-steppe &quot; plant community; a site where Dolichopus terminasianae and Hydrophorus alpinus were collected. Photo by O. A. Khruleva." httpUri="https://binary.pensoft.net/fig/474274" pageId="0" pageNumber="551">3</figureCitation>
-
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Rogers Bay environs. Middle part of gravel-loamy foothill ridge with spotted willow-sedge-moss cover; a site where Hydrophorus alpinus was collected. Photo by O. A. Khruleva." httpUri="https://binary.pensoft.net/fig/474276" pageId="0" pageNumber="551">5</figureCitation>
<figureCitation id="C550689CB3671FE857A7E8DB6D089841" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Rogers Bay environs. Middle part of gravel-loamy foothill ridge with spotted willow-sedge-moss cover; a site where Hydrophorus alpinus was collected. Photo by O. A. Khruleva." httpUri="https://binary.pensoft.net/fig/474276" pageId="0" pageNumber="551">5</figureCitation>
) located mainly outside the warmest inland areas, including the coast of the Island.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="551" type="distribution">
<paragraph pageId="0" pageNumber="551">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="551">Type locality: Sweden: &quot;Walli et Snjerack prope Quickjock&quot;. Palaearctic: Finland, Norway, Russia (Arkhangelsk, Chukotka mainland, Wrangel Island, Leningrad, Murmansk, Nenetsia, Yakutia, Yamalia), Sweden; Nearctic: Canada, USA.</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/474276" pageId="0" pageNumber="551" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="551">
<emphasis bold="true" pageId="0" pageNumber="551">Figure 5.</emphasis>
<subSubSection id="8782C24124AB4B522E09F0B02782FB05" pageId="0" pageNumber="551" type="distribution">
<paragraph id="06FCFA43D5CA0C52C87BC2C5401C49D0" pageId="0" pageNumber="551">Distribution.</paragraph>
<paragraph id="96D9E7E8AC155102A052B8ED68894423" pageId="0" pageNumber="551">Type locality: Sweden: &quot;Walli et Snjerack prope Quickjock&quot;. Palaearctic: Finland, Norway, Russia (Arkhangelsk, Chukotka mainland, Wrangel Island, Leningrad, Murmansk, Nenetsia, Yakutia, Yamalia), Sweden; Nearctic: Canada, USA.</paragraph>
<caption id="D55F3B93B64FDDB062A834FD577548C2" httpUri="https://binary.pensoft.net/fig/474276" pageId="0" pageNumber="551" start="Figure 5" startId="F5">
<paragraph id="7D5F8386E3DBCBA33038A6C7F9B85042" pageId="0" pageNumber="551">
<emphasis id="1C3B30DCAA85A851525E48890924B93C" bold="true" pageId="0" pageNumber="551">Figure 5.</emphasis>
Rogers Bay environs. Middle part of gravel-loamy foothill ridge with spotted willow-sedge-moss cover; a site where
<taxonomicName authorityName="Wahlberg" authorityYear="1844" class="Insecta" family="Dolichopodidae" genus="Hydrophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrophorus alpinus" order="Diptera" pageId="0" pageNumber="551" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis italics="true" pageId="0" pageNumber="551">Hydrophorus alpinus</emphasis>
<taxonomicName id="D5FE9156DC69E2B37F0AB3FADED63B4D" authorityName="Wahlberg" authorityYear="1844" class="Insecta" family="Dolichopodidae" genus="Hydrophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrophorus alpinus" order="Diptera" pageId="0" pageNumber="551" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="D7D4B6E491A3216F832139F6977504FD" italics="true" pageId="0" pageNumber="551">Hydrophorus alpinus</emphasis>
</taxonomicName>
was collected. Photo by O.A. Khruleva.
</paragraph>

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<mods:title>Two new Marquesan species of the southeastern Polynesian genus Oparanthus (Asteraceae, Coreopsidinae)</mods:title>
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<mods:title id="8290BD1E03B8D6D3E1D628A005DCC5F4">Two new Marquesan species of the southeastern Polynesian genus Oparanthus (Asteraceae, Coreopsidinae)</mods:title>
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<mods:namePart>Wagner, Warren L.</mods:namePart>
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<mods:namePart>Lorence, David H.</mods:namePart>
<mods:namePart id="1E892873CA09C8A7D66492F903F19D19">Lorence, David H.</mods:namePart>
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<treatment ID-GBIF-Taxon="152025417" LSID="urn:lsid:plazi:treatment:58AB36715F1CE7D5A9723FEE44876BF0" httpUri="http://treatment.plazi.org/id/58AB36715F1CE7D5A9723FEE44876BF0" lastPageId="5" lastPageNumber="144" pageId="3" pageNumber="142">
<subSubSection pageId="3" pageNumber="142" type="nomenclature">
<paragraph pageId="3" pageNumber="142">
<taxonomicName LSID="58AB3671-5F1C-E7D5-A972-3FEE44876BF0" authority="W. L. Wagner &amp; Lorence" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Oparanthus woodii" order="Asterales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="woodii" status="sp. nov.">Oparanthus woodii W. L. Wagner &amp; Lorence</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="142">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oparanthus woodii W. L. Wagner &amp; Lorence. A-N drawn from holotype Wood 6375, PTBG, except B from US isotype. A flowering branch B abaxial leaf surface showing naked domatia in axils of primary veins C lower leaf D head E receptacular bract of ray floret F receptacular bract of disk floret G-H ray corolla and functional style I disk corolla with non-functional style and fertile stamens J longitudinal view of disk corolla with non-functional style and fertile stamens K sterile disk achene L-M ray achenes N fertile stamen from disk floret." pageId="3" pageNumber="142">Fig. 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Field images of Oparanthus. A Oparanthus hivaoanus, (Hiva Oa, Price et al 201, photo D. Lorence) B Oparanthus teikiteetinii (Nuku Hiva, Lorence 6078, photo D. Lorence) C Oparanthus tiva (Tahuata, Wood 6523, photo K. Wood) D Oparanthus woodii (Nuku Hiva, Wood 6338, photo K. Wood)." pageId="3" pageNumber="142">3D</figureCitation>
<treatment id="58AB36715F1CE7D5A9723FEE44876BF0" ID-GBIF-Taxon="152025417" LSID="urn:lsid:plazi:treatment:58AB36715F1CE7D5A9723FEE44876BF0" httpUri="http://treatment.plazi.org/id/58AB36715F1CE7D5A9723FEE44876BF0" lastPageId="5" lastPageNumber="144" pageId="3" pageNumber="142" scope_family="Asteraceae">
<subSubSection id="8EF1FF711A1AA707F24A3DD9D3A49105" pageId="3" pageNumber="142" type="nomenclature">
<paragraph id="BA81A6778FE7E4789A2D394776012FBB" pageId="3" pageNumber="142">
<taxonomicName id="8907E303093BFA5E06A24F4D4E974440" ID-CoL="49TJ3" LSID="58AB3671-5F1C-E7D5-A972-3FEE44876BF0" authority="W. L. Wagner &amp; Lorence" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Oparanthus woodii" order="Asterales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="woodii" status="sp. nov.">Oparanthus woodii W. L. Wagner &amp; Lorence</taxonomicName>
<taxonomicNameLabel id="3F232E052C2ABDFE5D07F3F1D6F7203D" pageId="3" pageNumber="142">sp. nov.</taxonomicNameLabel>
<figureCitation id="3EDF53DFF073D2A8D186E8D2CC9D5B31" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oparanthus woodii W. L. Wagner &amp; Lorence. A-N drawn from holotype Wood 6375, PTBG, except B from US isotype. A flowering branch B abaxial leaf surface showing naked domatia in axils of primary veins C lower leaf D head E receptacular bract of ray floret F receptacular bract of disk floret G-H ray corolla and functional style I disk corolla with non-functional style and fertile stamens J longitudinal view of disk corolla with non-functional style and fertile stamens K sterile disk achene L-M ray achenes N fertile stamen from disk floret." pageId="3" pageNumber="142">Fig. 2</figureCitation>
<figureCitation id="28ABD3D42C65C43318B802221B660D99" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Field images of Oparanthus. A Oparanthus hivaoanus, (Hiva Oa, Price et al 201, photo D. Lorence) B Oparanthus teikiteetinii (Nuku Hiva, Lorence 6078, photo D. Lorence) C Oparanthus tiva (Tahuata, Wood 6523, photo K. Wood) D Oparanthus woodii (Nuku Hiva, Wood 6338, photo K. Wood)." pageId="3" pageNumber="142">3D</figureCitation>
</paragraph>
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<subSubSection pageId="3" pageNumber="142" type="latin">
<paragraph pageId="3" pageNumber="142">Latin.</paragraph>
<paragraph pageId="3" pageNumber="142">
<emphasis italics="true" pageId="3" pageNumber="142">Ab O. teikiteetinii fere glabra arbore, domatiis apilosis in foliis abaxialibus in axilibus nervis secundariis presentibus, foliis basi connatis, flosculis radiorum tubis corollarum 7.5-8 mm differt.</emphasis>
<subSubSection id="AD58E9F997AD0EECC9DAF9FDA3153F83" pageId="3" pageNumber="142" type="latin">
<paragraph id="538C685F912B062534A1C14DEFE86FD0" pageId="3" pageNumber="142">Latin.</paragraph>
<paragraph id="C24E9B94D2EE9E6F9C5059F3F28EE564" pageId="3" pageNumber="142">
<emphasis id="401A1D1441EEB994DCF8CFA1986D75C4" italics="true" pageId="3" pageNumber="142">Ab O. teikiteetinii fere glabra arbore, domatiis apilosis in foliis abaxialibus in axilibus nervis secundariis presentibus, foliis basi connatis, flosculis radiorum tubis corollarum 7.5-8 mm differt.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="142" type="type">
<paragraph pageId="3" pageNumber="142">Type.</paragraph>
<paragraph pageId="3" pageNumber="142">
<emphasis bold="true" pageId="3" pageNumber="142">Marquesas Islands</emphasis>
<subSubSection id="3B4B7935F4049E655A82C67A23EC0FC9" pageId="3" pageNumber="142" type="type">
<paragraph id="CC7C0E7B4333069BD58873B0061DF80A" pageId="3" pageNumber="142">Type.</paragraph>
<paragraph id="DA509EA88C1F971D0BE168F86697CCB2" pageId="3" pageNumber="142">
<emphasis id="875B6DE7149FE19A20217A5BD527735C" bold="true" pageId="3" pageNumber="142">Marquesas Islands</emphasis>
: Nuku Hiva: Ooumu region, top of Tapueahu Valley off new Hwy, [08 51'53S, 140 10'63W], 1067-1128 m, 23 June 1997, K. R. Wood 6375 (Holotype: PTBG-025565!; isotypes: BISH!, P!, PAP!, US!).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142" type="description">
<paragraph pageId="3" pageNumber="142">Description.</paragraph>
<paragraph lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142">
<emphasis italics="true" pageId="3" pageNumber="142">Trees</emphasis>
<subSubSection id="BDC57889BBB0E9393E347E3AB200B8A8" lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142" type="description">
<paragraph id="5B70700D989DDAA63CB624602EBF61DD" pageId="3" pageNumber="142">Description.</paragraph>
<paragraph id="D88428EA053646F81D018D6C3339ACC7" lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142">
<emphasis id="99C16CF73F5B765EA528604449FFBA49" italics="true" pageId="3" pageNumber="142">Trees</emphasis>
2-5 m tall, glabrous, functionally monoecious, moderately to diffusely branched, the trunk often with multiple prop roots, bark brown, wood cream-colored.
<emphasis italics="true" pageId="3" pageNumber="142">Leaves</emphasis>
<emphasis id="E5D1DE6B65D6E7D3CF689FB874E18AB9" italics="true" pageId="3" pageNumber="142">Leaves</emphasis>
thinly coriaceous, the blade broadly elliptic to broadly elliptic-obovate, 11-23.1
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="8B4F9D5FBD325B552D85E5CE5F6749DA" originalValue="×">x</normalizedToken>
3.5-12.7 cm, secondary veins 4-12 mm apart, conspicuously arching upward from midrib, then spreading to margin, abaxial axils often with domatia, these without associated hairs, margins entire, apex rounded to truncate, base cuneate, often oblique; petioles 3.5-6 cm long, the base conspicuously connate around stem with paired leaf petiole.
<emphasis italics="true" pageId="3" pageNumber="142">Inflorescences</emphasis>
<emphasis id="9EB8DFC4CC2B6D9D465F79C38BCDF532" italics="true" pageId="3" pageNumber="142">Inflorescences</emphasis>
terminal, heads solitary, 12-16 mm in diameter, 22-35 mm high, peduncles 3-15 mm long, stout; involucre campanulate; involucral bracts 8, in 2 series, the external ones 8-13 mm long, connate at the base, thick and broadly triangular, becoming lignified in fruit, the internal ones usually longer (up to 15 mm) and narrower, triangular to elliptic; receptacular bracts of the ray florets 13-14 mm long, those of the disk florets 12-13 mm long;
<emphasis italics="true" pageId="3" pageNumber="142">ray florets</emphasis>
<emphasis id="32B3C7F72C3EE0154158ACE6A26101EB" italics="true" pageId="3" pageNumber="142">ray florets</emphasis>
ca. 18-22, in 2 series, corolla tube and throat 7.5-8 mm long, limb 4.2-4.5 mm long, 2-3-lobed, the lobes usually divided to near corolla throat, occasionally only shallowly so;
<emphasis italics="true" pageId="3" pageNumber="142">disk florets</emphasis>
<emphasis id="590B355A05C825E1A463F660D0039CA2" italics="true" pageId="3" pageNumber="142">disk florets</emphasis>
ca. 40-50 or perhaps more, corolla tube and throat 6.3-6.6 mm long, the lobes 4.4-5.4
<pageBreakToken pageId="4" pageNumber="143" start="start">mm</pageBreakToken>
<pageBreakToken id="D2C69F627F5F4C6E5D68411EACC7A1BF" pageId="4" pageNumber="143" start="start">mm</pageBreakToken>
long.
<emphasis italics="true" pageId="4" pageNumber="143">Ray achenes</emphasis>
<emphasis id="D7C678E6A128D2547BA86039D544A107" italics="true" pageId="4" pageNumber="143">Ray achenes</emphasis>
elliptic, 5-6 mm long, distinctly winged, the wings ca.1 mm wide, extending slightly beyond the achene apex; disk achenes sterile, linear, 10-11 mm long, with 2 awns.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="143" type="etymology">
<paragraph pageId="4" pageNumber="143">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="143">This new species is named for Kenneth R. Wood, who first collected it and who has contributed greatly to our knowledge of the flora of the Marquesas and Hawaii through his collections and field observations.</paragraph>
<subSubSection id="C0749C10D3BC990DBABA6B318C5D9120" pageId="4" pageNumber="143" type="etymology">
<paragraph id="5D24FFCF39157D143D75702BEDE8D34B" pageId="4" pageNumber="143">Etymology.</paragraph>
<paragraph id="489814B248EBBFC5437795BE0F5DC176" pageId="4" pageNumber="143">This new species is named for Kenneth R. Wood, who first collected it and who has contributed greatly to our knowledge of the flora of the Marquesas and Hawaii through his collections and field observations.</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="144" pageId="4" pageNumber="143" type="distribution">
<paragraph pageId="4" pageNumber="143">Distribution.</paragraph>
<paragraph lastPageId="5" lastPageNumber="144" pageId="4" pageNumber="143">
<subSubSection id="3C7C51178A43737E8B1BCBC36C7CC44E" lastPageId="5" lastPageNumber="144" pageId="4" pageNumber="143" type="distribution">
<paragraph id="7E2739C383A5462684C9E2C8944D9573" pageId="4" pageNumber="143">Distribution.</paragraph>
<paragraph id="7146501B894D0AF57004BD278D4610C4" lastPageId="5" lastPageNumber="144" pageId="4" pageNumber="143">
Endemic to Nuku Hiva, Marquesas Islands, and apparently restricted to the Ooumu region, in gulches near the top of Tapueahu Valley, from 1060 to 1130 m elevation
<pageBreakToken pageId="5" pageNumber="144" start="start">.</pageBreakToken>
<pageBreakToken id="7079396CBC937A8508DF47C5DD32327E" pageId="5" pageNumber="144" start="start">.</pageBreakToken>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="ecology">
<paragraph pageId="5" pageNumber="144">Ecology.</paragraph>
<paragraph pageId="5" pageNumber="144">
<subSubSection id="50EE8C4DF1826641CAEF475089C01F34" pageId="5" pageNumber="144" type="ecology">
<paragraph id="35E8F1096FACDC210EB14721A6CC5295" pageId="5" pageNumber="144">Ecology.</paragraph>
<paragraph id="993C822DC325F7BF865C36AA4760520C" pageId="5" pageNumber="144">
Occurring in montane mesic to wet forest, ravines and steep slopes, with
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="5" pageNumber="144" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Metrosideros</emphasis>
<taxonomicName id="ADC49A1E12C05DBDDF2E8279E65D802C" class="Magnoliopsida" family="Myrtaceae" genus="Metrosideros" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Myrtales" pageId="5" pageNumber="144" phylum="Magnoliophyta" rank="genus">
<emphasis id="3ACFFEA1CFEAF5FECC5E134A80C8A586" italics="true" pageId="5" pageNumber="144">Metrosideros</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Oxalidales" pageId="5" pageNumber="144" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Weinmannia</emphasis>
<taxonomicName id="80C17AC6B0759E7F8C30B5D19BE5EDFA" class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Oxalidales" pageId="5" pageNumber="144" phylum="Magnoliophyta" rank="genus">
<emphasis id="866204673C6A662E2FAE3DD5508F3FED" italics="true" pageId="5" pageNumber="144">Weinmannia</emphasis>
</taxonomicName>
dominant and a diverse understory of
<taxonomicName class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="5" pageNumber="144" phylum="Polypodiopsida" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Asplenium</emphasis>
<taxonomicName id="7F387C57D903FD3BFF62F1D5B4734702" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="5" pageNumber="144" phylum="Polypodiopsida" rank="genus">
<emphasis id="E520A6E634B45AFF23C6FC39A9B0CEE0" italics="true" pageId="5" pageNumber="144">Asplenium</emphasis>
</taxonomicName>
,
<taxonomicName genus="Blechmum" lsidName="" pageId="5" pageNumber="144" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Blechmum</emphasis>
<taxonomicName id="443CB021D7BC585AD668C7497EA23545" genus="Blechmum" lsidName="" pageId="5" pageNumber="144" rank="genus">
<emphasis id="1DFFD913B9891973F02A1A175E4A5142" italics="true" pageId="5" pageNumber="144">Blechmum</emphasis>
</taxonomicName>
,
<taxonomicName class="Dicotyledoneae" family="Gesneriaceae" genus="Cyrtandra" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Tubiflorae" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Cyrtandra</emphasis>
<taxonomicName id="D533247C03933033BFA13FE464B5D9FB" class="Dicotyledoneae" family="Gesneriaceae" genus="Cyrtandra" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Tubiflorae" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis id="ADAE884CF23150CCF1F6B35BCC808814" italics="true" pageId="5" pageNumber="144">Cyrtandra</emphasis>
</taxonomicName>
,
<taxonomicName class="Filicopsida" family="Dennstaedtiaceae" genus="Hypolepis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="5" pageNumber="144" phylum="Pteridophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Hypolepis</emphasis>
<taxonomicName id="B10836C44A5309101CBA2439A6C53826" class="Filicopsida" family="Dennstaedtiaceae" genus="Hypolepis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Polypodiales" pageId="5" pageNumber="144" phylum="Pteridophyta" rank="genus">
<emphasis id="5E5B21391481246C31CCF9F0F29D3CF8" italics="true" pageId="5" pageNumber="144">Hypolepis</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Aquifoliales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Ilex</emphasis>
<taxonomicName id="5756A06F3F0AA591A7B1CB334AACADFD" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Aquifoliales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="989A40F7A4ADAC28A137190E133A7265" italics="true" pageId="5" pageNumber="144">Ilex</emphasis>
</taxonomicName>
, and
<taxonomicName class="Dicotyledoneae" family="Rutaceae" genus="Melicope" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rutales" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Melicope</emphasis>
<taxonomicName id="2C842C2CE0B0D65E9F47002416A2DA27" class="Dicotyledoneae" family="Rutaceae" genus="Melicope" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rutales" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis id="FE724F9B85DC7010A05EC31AF2685298" italics="true" pageId="5" pageNumber="144">Melicope</emphasis>
</taxonomicName>
, with stands of
<taxonomicName class="Monocotyledoneae" family="Pandanaceae" genus="Freycinetia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Pandanales" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Freycinetia</emphasis>
<taxonomicName id="DFD940258DFDDC2D3FF4CC68AB8B89CB" class="Monocotyledoneae" family="Pandanaceae" genus="Freycinetia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Pandanales" pageId="5" pageNumber="144" phylum="Angiospermae" rank="genus">
<emphasis id="218C010A9478A4A8AD2E371AF4E3F232" italics="true" pageId="5" pageNumber="144">Freycinetia</emphasis>
</taxonomicName>
nearby. Known to flower in June, but probably for some months after that.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="conservation status">
<paragraph pageId="5" pageNumber="144">Conservation status.</paragraph>
<paragraph pageId="5" pageNumber="144">
<subSubSection id="B87FAA45DB71991D8E3E31A113FF8C76" pageId="5" pageNumber="144" type="conservation status">
<paragraph id="6E7B5BA040910E64B794217E5CFD8AD7" pageId="5" pageNumber="144">Conservation status.</paragraph>
<paragraph id="749FFFE4D160E3568E8C401C0D5B3D46" pageId="5" pageNumber="144">
Based on the IUCN criteria and categories this species is assigned a preliminary Red List status of
<emphasis bold="true" pageId="5" pageNumber="144">Critically</emphasis>
<emphasis bold="true" pageId="5" pageNumber="144">Endangered</emphasis>
<emphasis id="946B8C341F38F8E40D213E3EE51A99EC" bold="true" pageId="5" pageNumber="144">Critically</emphasis>
<emphasis id="435DF50906CB003B97CA9D1CF4BC2A74" bold="true" pageId="5" pageNumber="144">Endangered</emphasis>
(CR) B2a, B2b (i-iii); D: B2: total area of occupancy less than 10 km2 (ca. 5 km2). B2a, a single population known; b (
<normalizedToken originalValue="iiii">i-iii</normalizedToken>
<normalizedToken id="9B3DE4F8BA94BAFE835DEC2898CC87AE" originalValue="iiii">i-iii</normalizedToken>
), habitat continuing decline inferred; D, population estimated to number fewer than 250 individuals. The suitable habitat for
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
<taxonomicName id="D8669DE878FDF2B4E7A83DFB90E89F27" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="E00BA507C91C9BF982646C9859E40D1A" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
on Nuku Hiva (
<emphasis italics="true" pageId="5" pageNumber="144">c.</emphasis>
<emphasis id="C0519505E1B701C6A03A7307D699C709" italics="true" pageId="5" pageNumber="144">c.</emphasis>
340 km2) is indicated as an endangered environment, threatened by human activity (deforestation and fire), feral animals, and invasive plants, thus reducing the extent of the forest.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="discussion">
<paragraph pageId="5" pageNumber="144">Discussion.</paragraph>
<paragraph pageId="5" pageNumber="144">
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
<subSubSection id="1CCEAE436A841B60DB80D686EC3BFC22" pageId="5" pageNumber="144" type="discussion">
<paragraph id="B51D5506BD18D1553366C02E4E40F427" pageId="5" pageNumber="144">Discussion.</paragraph>
<paragraph id="04A1AB7A68272BFB857C1B195471D34D" pageId="5" pageNumber="144">
<taxonomicName id="086346FF14E16ED2C5432B6816CADA85" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="3085DA5CAA573B53DBB283829878C74F" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
appears to be closely related to
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
<taxonomicName id="0FC769F879E149664D976C0B50695619" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis id="D55A1FE347E14C8925AFEC260415E156" italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
</taxonomicName>
, which grows at lower elevations, but approaches the known range of
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
<taxonomicName id="292A8BCC360C4BA5A925473239E99F0C" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="A3817B1BC65A045835B899760D61E740" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
within a few hundred meters. While
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
<taxonomicName id="276D24FE5FA24B7C9E4F014AEA6F6442" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis id="114F2DB00C87C2625C51DFF13607F751" italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
</taxonomicName>
is generally distinctive in the genus for its large size, attaining heights of up to 12 m, and for its large, often solitary capitula,
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
<taxonomicName id="6577EC543C88B8D1438AA38E5468D880" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="3A6344D0F92AC7D6A0A16F4F1B12C1A9" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
is a smaller tree up to 5 m tall and has large solitary heads on much shorter and stouter peduncles up to 15 mm long. The corollas are similar in these two species, but the ray corollas of
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
<taxonomicName id="ACAAF219D8BC1CCC212DF33732FEA246" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="97D02720BA855447D370517A3628DEDA" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
are also distinctive in that they are deeply divided to near the corolla throat. Likewise the leaves of
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
<taxonomicName id="E34F4669700B4179A5B3C22D7D42AA72" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="5FF4DB965029D96CA00DDB9BA94D38AF" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
are distinctive in that they are conspicuously connate at the petiole bases, have secondary veins that arch upwards, and havenaked domatia in the abaxial vein axils. These are not always present and they lack the tufts of hairs in the always present domatia of
<taxonomicName authorityName="R. K. Shannon &amp; W. L. Wagner" authorityYear="1997" baseAuthorityName="O. Deg. &amp; Sherff" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="hivoanus">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus hivoanus</emphasis>
<taxonomicName id="67ED179BDFEDBEB67172265F340626C2" authorityName="R. K. Shannon &amp; W. L. Wagner" authorityYear="1997" baseAuthorityName="O. Deg. &amp; Sherff" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="hivoanus">
<emphasis id="BEE9E6164FFAEADEDC113E3CBE287A31" italics="true" pageId="5" pageNumber="144">Oparanthus hivoanus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="tiva">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus tiva</emphasis>
<taxonomicName id="3EA175BC969EE97D3ECA64A69F6F7D20" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="tiva">
<emphasis id="58A5D6EF4786CEC2970BA17F01CBBBD4" italics="true" pageId="5" pageNumber="144">Oparanthus tiva</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="specimens examined">
<paragraph pageId="5" pageNumber="144">Specimens examined.</paragraph>
<paragraph pageId="5" pageNumber="144">
<emphasis bold="true" pageId="5" pageNumber="144">Marquesas Islands:</emphasis>
<subSubSection id="75602818B46862806BC09C375138DCCB" pageId="5" pageNumber="144" type="specimens examined">
<paragraph id="643141E05422C1146A137FB680ABAA4C" pageId="5" pageNumber="144">Specimens examined.</paragraph>
<paragraph id="AE5962DC2FE117779AC4A8D2954F1B46" pageId="5" pageNumber="144">
<emphasis id="C10690C3A236771494AFCC7FB5843F46" bold="true" pageId="5" pageNumber="144">Marquesas Islands:</emphasis>
MARQUESAS ISLANDS: Nuku Hiva, Ooumu region, top of Tapueahu Valley off new Hwy, [08 51'53S, 140 10'63W], Wood et. al. 6338 (P, PTBG, US); Wood 6376 (PTBG, US); Wood 6377 (BISH, K, P, PAP, PTBG, US).
</paragraph>
<caption pageId="5" pageNumber="144" start="Figure 2" startId="F2">
<paragraph pageId="5" pageNumber="144">
<emphasis bold="true" pageId="5" pageNumber="144">Figure 2.</emphasis>
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
<caption id="297FD13A16E04D645A105378F5449F16" pageId="5" pageNumber="144" start="Figure 2" startId="F2">
<paragraph id="0FBED967E9978E0240E08D9E6C5BF145" pageId="5" pageNumber="144">
<emphasis id="14937A6445F188F9B8DCAB8408F926A7" bold="true" pageId="5" pageNumber="144">Figure 2.</emphasis>
<taxonomicName id="21B32103110D461EE0AD289CC0DEF2D6" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="2966CDB8ECD688BA3D30760E2D35F68A" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
W. L. Wagner &amp; Lorence.
<emphasis bold="true" pageId="5" pageNumber="144">
<normalizedToken originalValue="AN">A-N</normalizedToken>
<emphasis id="B469A31642FCB32CC92E0261992AA45D" bold="true" pageId="5" pageNumber="144">
<normalizedToken id="FF04D0BF9E673FC9341E6AE446586EDA" originalValue="AN">A-N</normalizedToken>
</emphasis>
drawn from holotype Wood 6375, PTBG, except
<emphasis bold="true" pageId="5" pageNumber="144">B</emphasis>
<emphasis id="EF1A7C2228B4BE0FE41B2F3DE2725C96" bold="true" pageId="5" pageNumber="144">B</emphasis>
from US isotype.
<emphasis bold="true" pageId="5" pageNumber="144">A</emphasis>
<emphasis id="3FD0E0211300EA9AEDB7903B66C2D607" bold="true" pageId="5" pageNumber="144">A</emphasis>
flowering branch
<emphasis bold="true" pageId="5" pageNumber="144">B</emphasis>
<emphasis id="E178AC75B7B457FC946D947C38E66695" bold="true" pageId="5" pageNumber="144">B</emphasis>
abaxial leaf surface showing naked domatia in axils of primary veins
<emphasis bold="true" pageId="5" pageNumber="144">C</emphasis>
<emphasis id="2157DB02FC98E36333B12401C4EC965C" bold="true" pageId="5" pageNumber="144">C</emphasis>
lower leaf
<emphasis bold="true" pageId="5" pageNumber="144">D</emphasis>
<emphasis id="8A5A8264E3E1D52533080FB142A6E239" bold="true" pageId="5" pageNumber="144">D</emphasis>
head
<emphasis bold="true" pageId="5" pageNumber="144">E</emphasis>
<emphasis id="7B7FA78E99D750C0998903AF58D1C10E" bold="true" pageId="5" pageNumber="144">E</emphasis>
receptacular bract of ray floret
<emphasis bold="true" pageId="5" pageNumber="144">F</emphasis>
<emphasis id="589E4BD999AB79627731BEA9F61A0E26" bold="true" pageId="5" pageNumber="144">F</emphasis>
receptacular bract of disk floret
<emphasis bold="true" pageId="5" pageNumber="144">
<normalizedToken originalValue="GH">G-H</normalizedToken>
<emphasis id="6B1CABFDCFE507DFCFBA2E2D196D3A2A" bold="true" pageId="5" pageNumber="144">
<normalizedToken id="CF0A8A5C7CADB77F21F6B1DC8B712FD3" originalValue="GH">G-H</normalizedToken>
</emphasis>
ray corolla and functional style
<emphasis bold="true" pageId="5" pageNumber="144">I</emphasis>
<emphasis id="F639156958EC0246C4CD552E3F508B8B" bold="true" pageId="5" pageNumber="144">I</emphasis>
disk corolla with non-functional style and fertile stamens
<emphasis bold="true" pageId="5" pageNumber="144">J</emphasis>
<emphasis id="F90B10F5FA7B6072D7B08F386E1E464E" bold="true" pageId="5" pageNumber="144">J</emphasis>
longitudinal view of disk corolla with non-functional style and fertile stamens
<emphasis bold="true" pageId="5" pageNumber="144">K</emphasis>
<emphasis id="965FE2EEBF25AC4F3CDBD8186776CA9A" bold="true" pageId="5" pageNumber="144">K</emphasis>
sterile disk achene
<emphasis bold="true" pageId="5" pageNumber="144">
<normalizedToken originalValue="LM">L-M</normalizedToken>
<emphasis id="210789E92B1F17B4660E3CDB5939988E" bold="true" pageId="5" pageNumber="144">
<normalizedToken id="9AEF2F07E7BC766822E1D9377F44E682" originalValue="LM">L-M</normalizedToken>
</emphasis>
ray achenes
<emphasis bold="true" pageId="5" pageNumber="144">N</emphasis>
<emphasis id="594F76EC59FD14FF19CD807684D0AF21" bold="true" pageId="5" pageNumber="144">N</emphasis>
fertile stamen from disk floret.
</paragraph>
</caption>
<caption pageId="5" pageNumber="144" start="Figure 3" startId="F3">
<paragraph pageId="5" pageNumber="144">
<emphasis bold="true" pageId="5" pageNumber="144">Figure 3.</emphasis>
<caption id="9D74D9E7A65A6FC07ED7F91E4B0AA75F" pageId="5" pageNumber="144" start="Figure 3" startId="F3">
<paragraph id="2F87023DBF5A2815946D32A6A443F2F9" pageId="5" pageNumber="144">
<emphasis id="718680157F7BDE14562B0597DE99377B" bold="true" pageId="5" pageNumber="144">Figure 3.</emphasis>
Field images of
<taxonomicName authorityName="Sherff" authorityYear="1937" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus</emphasis>
<taxonomicName id="42C41EB1959ACCA21772EBA3C2F3F60A" authorityName="Sherff" authorityYear="1937" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="1FF1720505C93CCDF0C63E3D5D1A8736" italics="true" pageId="5" pageNumber="144">Oparanthus</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="5" pageNumber="144">A</emphasis>
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="hivaoanus">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus hivaoanus</emphasis>
<emphasis id="4A62EDCCF108C868189F3C37A1064511" bold="true" pageId="5" pageNumber="144">A</emphasis>
<taxonomicName id="54311BB17C771CDCD28D1722182923D7" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="hivaoanus">
<emphasis id="4BEBDCD5ADC16077D22B62C3FF4B78D9" italics="true" pageId="5" pageNumber="144">Oparanthus hivaoanus</emphasis>
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, (Hiva Oa, Price et al 201, photo D. Lorence)
<emphasis bold="true" pageId="5" pageNumber="144">B</emphasis>
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
<emphasis id="A3A084B3262F4B1D344FD39F1F31B89A" bold="true" pageId="5" pageNumber="144">B</emphasis>
<taxonomicName id="2FCB42BFDB6F1D035030810683E070F6" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="teikiteetinii">
<emphasis id="B30028A5DB01D8DA06F8FE7FC86DCFAE" italics="true" pageId="5" pageNumber="144">Oparanthus teikiteetinii</emphasis>
</taxonomicName>
(Nuku Hiva, Lorence 6078, photo D. Lorence)
<emphasis bold="true" pageId="5" pageNumber="144">C</emphasis>
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="tiva">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus tiva</emphasis>
<emphasis id="1EC9223B34F1EEC5952D979D52759141" bold="true" pageId="5" pageNumber="144">C</emphasis>
<taxonomicName id="3B4C70E4A1A6B75E1E3699E434FF368D" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="tiva">
<emphasis id="43D2D41459FFA4CADA35B99F705858D4" italics="true" pageId="5" pageNumber="144">Oparanthus tiva</emphasis>
</taxonomicName>
(Tahuata, Wood 6523, photo K. Wood)
<emphasis bold="true" pageId="5" pageNumber="144">D</emphasis>
<taxonomicName authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
<emphasis id="A99BAB202B4F427E6995C2697001AA10" bold="true" pageId="5" pageNumber="144">D</emphasis>
<taxonomicName id="33DEFB23E7A258C36EED1F64CA14A767" authorityName="W. L. Wagner &amp; Lorence" authorityYear="2011" class="Magnoliopsida" family="Asteraceae" genus="Oparanthus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="144" phylum="Tracheophyta" rank="species" species="woodii">
<emphasis id="7151D0F41CC3754DFC0E7D96A3DC4947" italics="true" pageId="5" pageNumber="144">Oparanthus woodii</emphasis>
</taxonomicName>
(Nuku Hiva, Wood 6338, photo K. Wood).
</paragraph>

View file

@ -1,97 +1,97 @@
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<mods:identifier id="9376FD868842DEFB835EA17AF18B9BDE" type="DOI">http://dx.doi.org/10.3897/mycokeys.101.111882</mods:identifier>
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<treatment id="58AEB2484ADD5D01A8522E2CBBBB5703" LSID="urn:lsid:plazi:treatment:58AEB2484ADD5D01A8522E2CBBBB5703" httpUri="http://treatment.plazi.org/id/58AEB2484ADD5D01A8522E2CBBBB5703" lastPageNumber="329" pageId="0" pageNumber="329">
<treatment id="58AEB2484ADD5D01A8522E2CBBBB5703" ID-GBIF-Taxon="221055194" LSID="urn:lsid:plazi:treatment:58AEB2484ADD5D01A8522E2CBBBB5703" httpUri="http://treatment.plazi.org/id/58AEB2484ADD5D01A8522E2CBBBB5703" lastPageNumber="329" pageId="0" pageNumber="329" scope_family="Cordycipitaceae" scope_order="Hypocreales">
<subSubSection id="CB707BB8DF6C68AC24BD8024BEBC736B" pageId="0" pageNumber="329" type="nomenclature">
<paragraph id="9298DE867EFC92846CAAE5791FE6288B" pageId="0" pageNumber="329">
<taxonomicName id="8130372F7F3E5061FCC13D7478677AAD" LSID="58AEB248-4ADD-5D01-A852-2E2CBBBB5703" authority="T. Wang, Ming J. Chen &amp; B. Huang" authorityName="T. Wang, Ming J. Chen &amp; B. Huang" authorityYear="2024" class="Insecta" family="Cordycipitaceae" genus="Samsoniella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Samsoniella aranea" order="Hypocreales" pageId="0" pageNumber="329" phylum="Ascomycota" rank="species" species="aranea" status="sp. nov.">Samsoniella aranea T. Wang, Ming J. Chen &amp; B. Huang</taxonomicName>
<taxonomicName id="8130372F7F3E5061FCC13D7478677AAD" LSID="58AEB248-4ADD-5D01-A852-2E2CBBBB5703" authority="T. Wang, Ming J. Chen &amp; B. Huang" authorityName="T. Wang, Ming J. Chen &amp; B. Huang" authorityYear="2024" class="Insecta" family="Cordycipitaceae" genus="Samsoniella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Samsoniella aranea" order="Hypocreales" pageId="0" pageNumber="329" phylum="Arthropoda" rank="species" species="aranea" status="sp. nov.">Samsoniella aranea T. Wang, Ming J. Chen &amp; B. Huang</taxonomicName>
<taxonomicNameLabel id="90843DB9D4E5CCDE08DCCE0B9C8126B4" pageId="0" pageNumber="329">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
@ -104,7 +104,7 @@
<paragraph id="8FC3615870C92D47FD6FE83B492ABCBD" pageId="0" pageNumber="329">Etymology.</paragraph>
<paragraph id="B594CF1268281F3691D383C5F3E5A859" pageId="0" pageNumber="329">
Referring to its host, spider, family
<taxonomicName id="224D4E62E3233452778D216F6AA34D96" authorityName="Mongkols., Noisrip., Thanakitp., Spatafora &amp; Luangsa-ard" authorityYear="2018" class="Insecta" kingdom="Fungi" lsidName="" order="Araneae" pageId="0" pageNumber="329" rank="order">Araneae</taxonomicName>
<taxonomicName id="224D4E62E3233452778D216F6AA34D96" authorityName="Mongkols., Noisrip., Thanakitp., Spatafora &amp; Luangsa-ard" authorityYear="2018" class="Insecta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Araneae" pageId="0" pageNumber="329" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
.
</paragraph>
</subSubSection>
@ -161,7 +161,7 @@ diam hyphae (Fig.
<caption id="6EB3501D520D73332CCF276E6B9C4DC2" doi="10.3897/mycokeys.101.111882.figure3" httpUri="https://binary.pensoft.net/fig/978126" pageId="0" pageNumber="329" start="Figure 3" startId="F3">
<paragraph id="F7445FD7F024F03EE308C7A871EE0D3F" pageId="0" pageNumber="329">
<emphasis id="CBF51DE26723A78D48D6369B015FFBF5" bold="true" pageId="0" pageNumber="329">Figure 3.</emphasis>
<taxonomicName id="DCBED19BC6D03E39570961D13A769A19" authorityName="T. Wang, Ming J. Chen &amp; B. Huang" authorityYear="2024" class="Insecta" family="Cordycipitaceae" genus="Samsoniella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Samsoniella aranea" order="Hypocreales" pageId="0" pageNumber="329" phylum="Ascomycota" rank="species" species="aranea">
<taxonomicName id="DCBED19BC6D03E39570961D13A769A19" authorityName="T. Wang, Ming J. Chen &amp; B. Huang" authorityYear="2024" class="Insecta" family="Cordycipitaceae" genus="Samsoniella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Samsoniella aranea" order="Hypocreales" pageId="0" pageNumber="329" phylum="Arthropoda" rank="species" species="aranea">
<emphasis id="712C901E6375515EAD8071D83995A594" italics="true" pageId="0" pageNumber="329">Samsoniella aranea</emphasis>
</taxonomicName>
<emphasis id="B11CB4ABFEC0B4E62FB1298786B79D68" bold="true" pageId="0" pageNumber="329">A</emphasis>
@ -237,11 +237,11 @@ wide at the base, and 0.5-0.9μm wide at the apex (Fig.
<subSubSection id="8F757677AF6485D43ED9498A6CEEF9CC" pageId="0" pageNumber="329" type="notes">
<paragraph id="4727CD3F42FD3F4F73947C32D75ACBB4" pageId="0" pageNumber="329">Notes.</paragraph>
<paragraph id="36B2F43AF86DB67224B8F6FB0A75A7F0" pageId="0" pageNumber="329">
<taxonomicName id="573D41FF05790C7C34AF945CC4F001DA" authorityName="T. Wang, Ming J. Chen &amp; B. Huang" authorityYear="2024" class="Insecta" family="Cordycipitaceae" genus="Samsoniella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Samsoniella aranea" order="Hypocreales" pageId="0" pageNumber="329" phylum="Ascomycota" rank="species" species="aranea">
<taxonomicName id="573D41FF05790C7C34AF945CC4F001DA" authorityName="T. Wang, Ming J. Chen &amp; B. Huang" authorityYear="2024" class="Insecta" family="Cordycipitaceae" genus="Samsoniella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Samsoniella aranea" order="Hypocreales" pageId="0" pageNumber="329" phylum="Arthropoda" rank="species" species="aranea">
<emphasis id="28D7210650DE8D112E27E9B760DE53C6" italics="true" pageId="0" pageNumber="329">Samsoniella aranea</emphasis>
</taxonomicName>
was readily classified within the genus
<taxonomicName id="A4380BA7292E786A4540F9ECEB50FE2A" authorityName="Mongkols., Noisrip., Thanakitp., Spatafora &amp; Luangsa-ard" authorityYear="2018" class="Insecta" family="Cordycipitaceae" genus="Samsoniella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Samsoniella" order="Hypocreales" pageId="0" pageNumber="329" phylum="Ascomycota" rank="genus">
<taxonomicName id="A4380BA7292E786A4540F9ECEB50FE2A" authorityName="Mongkols., Noisrip., Thanakitp., Spatafora &amp; Luangsa-ard" authorityYear="2018" class="Insecta" family="Cordycipitaceae" genus="Samsoniella" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Samsoniella" order="Hypocreales" pageId="0" pageNumber="329" phylum="Arthropoda" rank="genus">
<emphasis id="6813607F54BDC865D9BD63EB9D2AFA3C" italics="true" pageId="0" pageNumber="329">Samsoniella</emphasis>
</taxonomicName>
through phylogenetic analyses (Fig.

View file

@ -1,222 +1,222 @@
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<taxonomicNameLabel id="40FBB4B343A14E53C5886FA44D7CFA7D" pageId="6" pageNumber="31">sp. n.</taxonomicNameLabel>
Figs 1235
</paragraph>
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<subSubSection pageId="6" pageNumber="31" type="type material">
<paragraph pageId="6" pageNumber="31">Type material.</paragraph>
<paragraph pageId="6" pageNumber="31">
<subSubSection id="944F6B3DFE1CE5477DE9A54FA28CFD56" pageId="6" pageNumber="31" type="type material">
<paragraph id="E6590896696C0CDD07EF7C157D0E8965" pageId="6" pageNumber="31">Type material.</paragraph>
<paragraph id="A86CAA66799E5C21D2C065666C3532B8" pageId="6" pageNumber="31">
Holotype ♀ (Fig. 5A) (in Museo Nacional de Ciencias Naturales, Madrid, Spain (MNCN), card-mounted. Cat.
<normalizedToken originalValue="nº"></normalizedToken>
<normalizedToken id="3796641FAA0042DFCC0AFF4DE0E23CCD" originalValue="nº"></normalizedToken>
2315). PANAMA,
<normalizedToken originalValue="Chiriquí">Chiriqui</normalizedToken>
<normalizedToken id="BD4FFE0EB0A071F7A077D6571AEA0D22" originalValue="Chiriquí">Chiriqui</normalizedToken>
,
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
<normalizedToken originalValue="Barú">Baru</normalizedToken>
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.776889">8°46'36.8&quot;N</geoCoordinate>
<normalizedToken id="1FB131E8B9E2D2E9BFAD64E9E78E6820" originalValue="Volcán">Volcan</normalizedToken>
<normalizedToken id="09DA6021BDE2EE5961E0D72410689A88" originalValue="Barú">Baru</normalizedToken>
<geoCoordinate id="1377F2C708049317A16E122B45E6FCB2" direction="north" orientation="latitude" precision="1" value="8.776889">8°46'36.8&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-82.52759">82°31'39.3&quot;W</geoCoordinate>
<geoCoordinate id="D315CF072D6E4ACE9FD638CC832C6582" direction="west" orientation="longitude" precision="1" value="-82.52759">82°31'39.3&quot;W</geoCoordinate>
, 2515-3045 m; from galls on leaves of
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus bumelioides" order="Fagales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="bumelioides">Quercus bumelioides</taxonomicName>
<taxonomicName id="04A1BD8DB2C122CCBDEC8E2C71990DA9" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus bumelioides" order="Fagales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="bumelioides">Quercus bumelioides</taxonomicName>
Liebm. (
<taxonomicName genus="Fagaceae" lsidName="Fagaceae" pageId="6" pageNumber="31" rank="genus">Fagaceae</taxonomicName>
<taxonomicName id="8CD4BCC7BC97ACFD6B249639858E1D2F" genus="Fagaceae" lsidName="Fagaceae" pageId="6" pageNumber="31" rank="genus">Fagaceae</taxonomicName>
), gall collected 16.vi.2008, insect emerged vii.08, E. Medianero leg. Paratypes: 1♀ same data as holotype, 2♀ same data as holotype but collected 15.iii.2008, insect emerged iv.08, 1♀ same data as holotype but collected 24 i.2009, insect emerged ii.09. Three paratypes in MNCN, one paratype in
<normalizedToken originalValue="Maestría">Maestria</normalizedToken>
<normalizedToken id="B7EAC6E34D0C38A719045B5CE639EBEB" originalValue="Maestría">Maestria</normalizedToken>
en
<normalizedToken originalValue="Entomología">Entomologia</normalizedToken>
<normalizedToken id="7DB028B3D8BA2FD9083A7AB2D94AEF1D" originalValue="Entomología">Entomologia</normalizedToken>
, Universidad de
<normalizedToken originalValue="Panamá">Panama</normalizedToken>
<normalizedToken id="97C1DE552C9E9DD04FAC3A5FE469761D" originalValue="Panamá">Panama</normalizedToken>
(MEUP).
</paragraph>
<paragraph pageId="6" pageNumber="31">Additionally, 1♀ paratype of the type series was dissected for SEM observation (in MNCN).</paragraph>
<paragraph id="6EBD95E4CA48CED3B0592EBA01FC207F" pageId="6" pageNumber="31">Additionally, 1♀ paratype of the type series was dissected for SEM observation (in MNCN).</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="31" type="etymology">
<paragraph pageId="6" pageNumber="31">Etymology.</paragraph>
<paragraph pageId="6" pageNumber="31">Named after the country where the new species was collected.</paragraph>
<subSubSection id="39A7E0102A377D2A8DAEAF06288E8B8C" pageId="6" pageNumber="31" type="etymology">
<paragraph id="2D659EA5B9D8502FF1E02A11D2C8E141" pageId="6" pageNumber="31">Etymology.</paragraph>
<paragraph id="3489155F2920704D853E22F097D4FB5D" pageId="6" pageNumber="31">Named after the country where the new species was collected.</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="34" pageId="6" pageNumber="31" type="description">
<paragraph pageId="6" pageNumber="31">Description.</paragraph>
<paragraph lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31">
<subSubSection id="1D545C6D33E5EC3E55CDA7B132B749E6" lastPageId="9" lastPageNumber="34" pageId="6" pageNumber="31" type="description">
<paragraph id="265051503634E52619A00F822E031735" pageId="6" pageNumber="31">Description.</paragraph>
<paragraph id="33AAC193C09A3DC2FC6BD93519FC1D8C" lastPageId="7" lastPageNumber="32" pageId="6" pageNumber="31">
Asexual female. Body length 2.9 mm (range 2.59-3.42; N = 4). Head and mesosoma black. Metasoma, flagellomeres and tarsi brownish; hypopygium
<pageBreakToken pageId="7" pageNumber="32" start="start">yellowish</pageBreakToken>
<pageBreakToken id="4239EC4C1162B9B32D42670DD22BFB4A" pageId="7" pageNumber="32" start="start">yellowish</pageBreakToken>
. Mandibles yellowish with black teeth. Antennal scape and pedicel yellowish in part. Half basal of coxae black; coxae apically, femora and tibiae ventrally yellowish. Forewing hyaline, slightly and uniformly darkened; veins dark brown.
</paragraph>
<paragraph pageId="7" pageNumber="32">Head alutaceous-reticulate, moderately pubescent, with relatively long white setae, except on vertex, upper frons and gena, in dorsal view about 2.7 times wider than long. POL 1.8 times longer than OOL, posterior ocellus separated from inner orbit of eye by 2.0 times its longest diameter. Head in anterior view (Fig. 1A) 1.2 times wider than high. Genae slightly expanded behind eyes, strongly alutaceus-reticulate (Fig. 1B). Clypeus more or less trapezoidal, 1.7 times wider than high moderately pubescent, ventral margin sinuate, moderately projecting over mandibles. Anterior tentorial pits visible; epistomal sulcus not visible, clypeo-pleurostomal lines visible. Malar space 0.2 times height of compound eye, without malar sulcus strongly alutaceus-reticulate, without irradiating striae from clypeus (Fig. 1B). Toruli situated slightly above mid-height of compound eye; distance between antennal rim and compound eye 0.8 times width of antennal socket including rim. Ocellar plate slightly raised. Head, posterior view (Fig. 1C) without occipital carina. Gula short; distance between occipital and oral foramina ashigh as occipital foramen (Fig. 1C). Hypostomal sulci well separate at oral fossa.</paragraph>
<paragraph pageId="7" pageNumber="32">Mouthparts (Figs 1D): mandibles exposed; with dense setae in base, right mandible with three teeth; left with two teeth. Cardo of maxilla visible, maxillary stipes long, about 2.3 times longer than wide. Maxillary palp five-segmented. Labial palp three-segmented, both moderately pubescent (Fig. 1D).</paragraph>
<paragraph pageId="7" pageNumber="32">
<paragraph id="ED38B78397C5B9D5A18F94EA976F52B0" pageId="7" pageNumber="32">Head alutaceous-reticulate, moderately pubescent, with relatively long white setae, except on vertex, upper frons and gena, in dorsal view about 2.7 times wider than long. POL 1.8 times longer than OOL, posterior ocellus separated from inner orbit of eye by 2.0 times its longest diameter. Head in anterior view (Fig. 1A) 1.2 times wider than high. Genae slightly expanded behind eyes, strongly alutaceus-reticulate (Fig. 1B). Clypeus more or less trapezoidal, 1.7 times wider than high moderately pubescent, ventral margin sinuate, moderately projecting over mandibles. Anterior tentorial pits visible; epistomal sulcus not visible, clypeo-pleurostomal lines visible. Malar space 0.2 times height of compound eye, without malar sulcus strongly alutaceus-reticulate, without irradiating striae from clypeus (Fig. 1B). Toruli situated slightly above mid-height of compound eye; distance between antennal rim and compound eye 0.8 times width of antennal socket including rim. Ocellar plate slightly raised. Head, posterior view (Fig. 1C) without occipital carina. Gula short; distance between occipital and oral foramina ashigh as occipital foramen (Fig. 1C). Hypostomal sulci well separate at oral fossa.</paragraph>
<paragraph id="1BB8365CE61002ABC24065C30377A233" pageId="7" pageNumber="32">Mouthparts (Figs 1D): mandibles exposed; with dense setae in base, right mandible with three teeth; left with two teeth. Cardo of maxilla visible, maxillary stipes long, about 2.3 times longer than wide. Maxillary palp five-segmented. Labial palp three-segmented, both moderately pubescent (Fig. 1D).</paragraph>
<paragraph id="2AC55D1CB7BE39E6F820456F1AB7226D" pageId="7" pageNumber="32">
Antenna (Fig. 1
<normalizedToken originalValue="EG">E-G</normalizedToken>
<normalizedToken id="A79C61E85BA6491690B3F9DF5DD1AA8C" originalValue="EG">E-G</normalizedToken>
) of moderate length, as long as 1/2 body length, with 16 antennomeres; flagellum not broadening towards apex; with relatively long, erect setae, and visible elongate placodeal sensilla (Fig. 1F). Relative lengths of antennal segments: 19:12:32:25:20:21:17:18:16:15:13:12:13:12:11:16. Pedicel, globose, 0.6 as long as scape (Fig. 1G). F1 1.3 times as long as F2. F14 2.1 times longer than wide, 1.5 times as long as F13 (Fig. 1F). Placodeal sensillae on F2-F14 disposed in one row of 8-9 sensillae in half dorsal area of each flagellomere.
</paragraph>
<paragraph pageId="7" pageNumber="32">Mesosoma. Uniformlyalutaceus, moderately pubescent, in lateral view 1.2 times as long as high, slightly convex dorsally (Fig. 2B). Pronotum, densely pubescent; lateral surface of pronotum with longitudinal wrinkles; with long and dense white setae (Fig. 2A-B). Pronotum short medially, ratio of length of pronotum medially/laterally = 0.2. Pronotal plate indistinct dorsally (Fig. 2A).</paragraph>
<paragraph lastPageId="8" lastPageNumber="33" pageId="7" pageNumber="32">
<paragraph id="293D0006F08407F54C6F9BD101D2369C" pageId="7" pageNumber="32">Mesosoma. Uniformlyalutaceus, moderately pubescent, in lateral view 1.2 times as long as high, slightly convex dorsally (Fig. 2B). Pronotum, densely pubescent; lateral surface of pronotum with longitudinal wrinkles; with long and dense white setae (Fig. 2A-B). Pronotum short medially, ratio of length of pronotum medially/laterally = 0.2. Pronotal plate indistinct dorsally (Fig. 2A).</paragraph>
<paragraph id="FA5B7A03B9540AB492A4AE471B723CB3" lastPageId="8" lastPageNumber="33" pageId="7" pageNumber="32">
Mesonotum (Fig. 2C). Mesoscutum coriaceous-alutaceus, only slightly pubescent medially and along notauli, slightly broader than long in dorsal view. Notauli percurrent, smooth, well impressed along entire length, reaching pronotum, not quite convergent, well separated posteriorly, median mesoscutal impression not visible. Anteroadmedian and parapsidal signa visible. Transscutal fissure narrow, well-visible, deeply impressed, slightly sinuate. Scutellar foveae not well differentiated, shallow, confluent medially, with some longitudinal striae and indistinctly margined posteriorly. Mesoscutellum (Fig. 2D) rounded from above moderately pubescent, about 0.6 as long as mesoscutum, strongly alutaceus-reticulate, in lateral view extending posteriorly slightly
<pageBreakToken pageId="8" pageNumber="33" start="start">over</pageBreakToken>
<pageBreakToken id="73FD982720500625D0670C7A9398CA05" pageId="8" pageNumber="33" start="start">over</pageBreakToken>
the dorsellum. Axillula slightly pubescent, their anterior margins marked and posterior indistinctly. Mesopleuron alutaceus, slightly pubescent with mesopleural triangle densely pubescent (Fig. 2B).
</paragraph>
<paragraph pageId="9" pageNumber="34">
<pageBreakToken pageId="9" pageNumber="34" start="start">Metanotum</pageBreakToken>
<paragraph id="CD92A702E0B50046D8C4057A9D61A17F" pageId="9" pageNumber="34">
<pageBreakToken id="F4C3BECC37B934F54F89E231E83AF07A" pageId="9" pageNumber="34" start="start">Metanotum</pageBreakToken>
(Fig. 2E). Metapectal-propodeal complex. Metapleural sulcus reaching posterior margin of mesopectus at about two thirds height (Fig. 2B). Lateral propodeal carinae poorly defined, fragmented, slightly divergent posteriorly and reaching nucha. Median propodeal area narrow, bare, with a median longitudinal carina fragmented but well visible (Fig. 2E); lateral propodeal area densely pubescent, with relatively long white setae, nucha rugose.
</paragraph>
<paragraph pageId="9" pageNumber="34">Legs (Fig. 2F). Densely pubescent, metatarsal claws with an acute basal lobe or short tooth (Fig. 2G).</paragraph>
<paragraph pageId="9" pageNumber="34">Forewing (Fig. 5B) 1.2 times as long as body, without smoky spots or stripes and densely pubescent; basal cell pubescent; radial cell 4.2 times longer than wide, open along anterior margin; areolet large, triangular, closed. R1, Rs and M nearly straight not reaching wing margin. R1 forming a quite acute angle with anterior margin of wing; Rs+M not reaching basalis. Basalis slightly curved, 2r well pigmented. Apical margin of wing with moderately long hair fringe.</paragraph>
<paragraph pageId="9" pageNumber="34">
<paragraph id="9CD5AB60EB590EE789DEE22A11919F1B" pageId="9" pageNumber="34">Legs (Fig. 2F). Densely pubescent, metatarsal claws with an acute basal lobe or short tooth (Fig. 2G).</paragraph>
<paragraph id="1DB30D0DE82392BD010F37AE17ECC14D" pageId="9" pageNumber="34">Forewing (Fig. 5B) 1.2 times as long as body, without smoky spots or stripes and densely pubescent; basal cell pubescent; radial cell 4.2 times longer than wide, open along anterior margin; areolet large, triangular, closed. R1, Rs and M nearly straight not reaching wing margin. R1 forming a quite acute angle with anterior margin of wing; Rs+M not reaching basalis. Basalis slightly curved, 2r well pigmented. Apical margin of wing with moderately long hair fringe.</paragraph>
<paragraph id="77B48F3D1AECE91E4C89803014847550" pageId="9" pageNumber="34">
Metasoma (Fig. 3A) large, as long as head and mesosoma combined, in lateral view as wide as high. Second metasomal tergite covering about two thirds of metasoma, with a patch of setae in its lateral anteromedial area. Projecting part of hypopygial spine long, lanceolate, tapering from the base to apex, at least 4.0 times as long as broad, with dense long basal setae arising in the base of the projected part which reach the apex of the spine forming a tuft (Fig. 3
<normalizedToken originalValue="BC">B-C</normalizedToken>
<normalizedToken id="DCF98FD6DDA5530C5092927557452B7D" originalValue="BC">B-C</normalizedToken>
).
</paragraph>
<paragraph pageId="9" pageNumber="34">
<paragraph id="111551E86B7DF4F2D8107611C04ABCD2" pageId="9" pageNumber="34">
Gall (Fig. 5
<normalizedToken originalValue="CG">C-G</normalizedToken>
<normalizedToken id="BD5CC34C7CAC925C3D712182785C2A77" originalValue="CG">C-G</normalizedToken>
). Irregular small formations (up to 4-9 mm) arising from crevices on the stems and on the petiole and midribs of leaves. The gall is displayed as a dense mass covered with light brown hairs, solitary, containing a single larval cell or more frequently forming clusters, and then appearing as polythalamous. Inside, the gall has a highly lignified core enclosing the larva (Fig. 5E).
</paragraph>
<paragraph pageId="9" pageNumber="34">
<paragraph id="0225E2E8FD3622A9CB21D8B5895F1DA8" pageId="9" pageNumber="34">
Host plant.
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus bumelioides" order="Fagales" pageId="9" pageNumber="34" phylum="Tracheophyta" rank="species" species="bumelioides">Quercus bumelioides</taxonomicName>
<taxonomicName id="052364161AA5DD9AD68FC7F33E081714" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus bumelioides" order="Fagales" pageId="9" pageNumber="34" phylum="Tracheophyta" rank="species" species="bumelioides">Quercus bumelioides</taxonomicName>
Liebm. (section Quercus of
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="9" pageNumber="34" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
<taxonomicName id="5ED5BB3E710A5875DD510FCD920EB8A2" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="9" pageNumber="34" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
; white oaks (
<taxonomicName genus="Fagaceae" lsidName="Fagaceae" pageId="9" pageNumber="34" rank="genus">Fagaceae</taxonomicName>
<taxonomicName id="54D0E8EC80D995C9AD8321870398E796" genus="Fagaceae" lsidName="Fagaceae" pageId="9" pageNumber="34" rank="genus">Fagaceae</taxonomicName>
), a species distributed from Mexico to Panama (
<bibRefCitation pageId="9" pageNumber="34">Breedlove 2001</bibRefCitation>
<bibRefCitation id="0670E3026829DD9D3E9EB70EE4741FF1" pageId="9" pageNumber="34">Breedlove 2001</bibRefCitation>
). The gall resembles that of the species of the Nubila complex, established by Kinsey within the subgenus
<taxonomicName class="Insecta" family="Cynipidae" genus="Acraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acraspis" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="genus">Acraspis</taxonomicName>
<taxonomicName id="BD91902F6C1B0D1E29E05DB08CD7BE08" class="Insecta" family="Cynipidae" genus="Acraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acraspis" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="genus">Acraspis</taxonomicName>
of
<taxonomicName class="Insecta" family="Cynipidae" genus="Cynips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cynips" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="genus">Cynips</taxonomicName>
<taxonomicName id="0BE9C133E6326D6051EE652EFC901DB0" class="Insecta" family="Cynipidae" genus="Cynips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cynips" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="genus">Cynips</taxonomicName>
(
<bibRefCitation pageId="9" pageNumber="34">Kinsey 1936</bibRefCitation>
<bibRefCitation id="4A2944F47AA0FE35EEA2F01C6FB21C00" pageId="9" pageNumber="34">Kinsey 1936</bibRefCitation>
), all known from Mexico. These galls were described as a mass of coarse hairs containing a spherical hard core, attached to mid-veins, on or under the surfaces of leaves. However, the insects are quite different in important characteristics such as the number of antennal segments and the shape of the hypopygial spine.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="34" type="distribution">
<paragraph pageId="9" pageNumber="34">Distribution.</paragraph>
<paragraph pageId="9" pageNumber="34">
<taxonomicName class="Insecta" family="Cynipidae" genus="Barucynips" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Barucynips panamensis" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="panamensis">Barucynips panamensis</taxonomicName>
<subSubSection id="4DC5746DBC8962197215229AABABC8A2" pageId="9" pageNumber="34" type="distribution">
<paragraph id="6A75D372A537D82AFBD8AC5E09D653BA" pageId="9" pageNumber="34">Distribution.</paragraph>
<paragraph id="1467A33F8343F1404F5F2B12847E8567" pageId="9" pageNumber="34">
<taxonomicName id="B82F83E546B3ED0D82AB22084B554BCC" genus="Barucynips" lsidName="Barucynips panamensis" pageId="9" pageNumber="34" rank="species" species="panamensis">Barucynips panamensis</taxonomicName>
was found between 2515-3045 m a.s.l. at
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
<normalizedToken originalValue="Barú">Baru</normalizedToken>
<normalizedToken id="953B98B8215F84BCFD03DE68CD6DCC72" originalValue="Volcán">Volcan</normalizedToken>
<normalizedToken id="5A976B1251249A80CC1B8A75C687A903" originalValue="Barú">Baru</normalizedToken>
, Chiriqui, Panama. Although currently known only at this locality, it is a species that is relatively abundant at the higher elevations of
<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
<normalizedToken originalValue="Barú">Baru</normalizedToken>
<normalizedToken id="E102723AC9A54CED457250E9C4190133" originalValue="Volcán">Volcan</normalizedToken>
<normalizedToken id="68F8C2CE22500B36485CE2990D0BB1CD" originalValue="Barú">Baru</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="34" type="biology">
<paragraph pageId="9" pageNumber="34">Biology.</paragraph>
<paragraph pageId="9" pageNumber="34">
<subSubSection id="9D5881844F0CEC5CDA188440A4602CBA" pageId="9" pageNumber="34" type="biology">
<paragraph id="E8C02A602056655D866C0DC51E55A7E4" pageId="9" pageNumber="34">Biology.</paragraph>
<paragraph id="13918AC1A41687053A16E958596FD6AA" pageId="9" pageNumber="34">
Only the asexual generation is known, inducing galls on
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus bumelioides" order="Fagales" pageId="9" pageNumber="34" phylum="Tracheophyta" rank="species" species="bumelioides">Quercus bumelioides</taxonomicName>
<taxonomicName id="6C042BAEDB3B7885CA9B5F74EBB5D67E" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus bumelioides" order="Fagales" pageId="9" pageNumber="34" phylum="Tracheophyta" rank="species" species="bumelioides">Quercus bumelioides</taxonomicName>
Liebm (section Quercus). The galls are common and can be found at every time of year in different grades of maturation on stems and leaves. The galls frequently are found growing together with galls of the new species, also described in this paper,
<taxonomicName class="Insecta" family="Cynipidae" genus="Coffeikokkos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coffeikokkos korytkowskii" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="korytkowskii">Coffeikokkos korytkowskii</taxonomicName>
<taxonomicName id="74C6717A25FFA1D553458BA925B6CAD2" genus="Coffeikokkos" lsidName="Coffeikokkos korytkowskii" pageId="9" pageNumber="34" rank="species" species="korytkowskii">Coffeikokkos korytkowskii</taxonomicName>
. The insects studied emerged from January to July.
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34" type="inquiline and parasitoid associated community">
<paragraph pageId="9" pageNumber="34">Inquiline and parasitoid associated community.</paragraph>
<paragraph lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34">
<subSubSection id="3819F72586BF6A518DF35541BCB983A2" lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34" type="inquiline and parasitoid associated community">
<paragraph id="E3AEC38F84BF1A6B573028419F5E1593" pageId="9" pageNumber="34">Inquiline and parasitoid associated community.</paragraph>
<paragraph id="4225758593EDCD6E92702BBD79B6F09D" lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34">
From the gall of
<taxonomicName class="Insecta" family="Cynipidae" genus="Barucynips" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Barucynips panamensis" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="panamensis">Barucynips panamensis</taxonomicName>
<taxonomicName id="E750C399268EA05EE38BF27EADD04252" genus="Barucynips" lsidName="Barucynips panamensis" pageId="9" pageNumber="34" rank="species" species="panamensis">Barucynips panamensis</taxonomicName>
, two species of inquilines were reared,
<taxonomicName class="Insecta" family="Cynipidae" genus="Synergus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synergus elegans" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="elegans">Synergus elegans</taxonomicName>
<taxonomicName id="2421ECD3CDE58ADFF260E2FDC5B8B0B8" class="Insecta" family="Cynipidae" genus="Synergus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synergus elegans" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="elegans">Synergus elegans</taxonomicName>
Nieves-Aldrey &amp; Medianero, and one indeterminate species similar to
<taxonomicName class="Insecta" family="Cynipidae" genus="Synergus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synergus luteus" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="luteus">Synergus luteus</taxonomicName>
<taxonomicName id="174CF73B0ECF8F3C2DBCD3B0A3E8BAAF" class="Insecta" family="Cynipidae" genus="Synergus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synergus luteus" order="Hymenoptera" pageId="9" pageNumber="34" phylum="Arthropoda" rank="species" species="luteus">Synergus luteus</taxonomicName>
Nieves-Aldrey &amp; Medi
<pageBreakToken pageId="10" pageNumber="35" start="start">anero</pageBreakToken>
<pageBreakToken id="6FA4D92DCF4A2E79426698AFF7314AF6" pageId="10" pageNumber="35" start="start">anero</pageBreakToken>
or
<taxonomicName class="Insecta" family="Cynipidae" genus="Synergus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synergus gabrieli" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="gabrieli">Synergus gabrieli</taxonomicName>
<taxonomicName id="B523EA8781FD4D80BB687FAAB7174ABD" class="Insecta" family="Cynipidae" genus="Synergus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synergus gabrieli" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="gabrieli">Synergus gabrieli</taxonomicName>
Nieves-Aldrey &amp; Medianero (
<bibRefCitation author="Nieves-Aldrey, JL" journalOrPublisher="Zootaxa" pageId="17" pageNumber="42" pagination="1 - 47" title="Taxonomy of inquilines of oak gall wasps of Panama, with description of eight new species of Synergus Hartig (Hymenoptera, Cynipidae, Synergini)." volume="2744" year="2011">Nieves-Aldrey and Medianero 2011</bibRefCitation>
<bibRefCitation id="2DAD3AAA74A8F85D8088F736AD9E40CC" author="Nieves-Aldrey, JL" journalOrPublisher="Zootaxa" pageId="17" pageNumber="42" pagination="1 - 47" title="Taxonomy of inquilines of oak gall wasps of Panama, with description of eight new species of Synergus Hartig (Hymenoptera, Cynipidae, Synergini)." volume="2744" year="2011">Nieves-Aldrey and Medianero 2011</bibRefCitation>
). Additionally, the parasitoid species
<taxonomicName class="Insecta" family="Ormyridae" genus="Ormyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormyrus venustus" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="venustus">Ormyrus venustus</taxonomicName>
<taxonomicName id="B283A84EE691B08F27F8DE6753904A41" class="Insecta" family="Ormyridae" genus="Ormyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormyrus venustus" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="venustus">Ormyrus venustus</taxonomicName>
Hanson 1992 (
<taxonomicName genus="Chalcidoidea" lsidName="Chalcidoidea" pageId="10" pageNumber="35" rank="genus">Chalcidoidea</taxonomicName>
<taxonomicName id="0B6CD82FCBE03371503739C68433F9DC" genus="Chalcidoidea" lsidName="Chalcidoidea" pageId="10" pageNumber="35" rank="genus">Chalcidoidea</taxonomicName>
,
<taxonomicName family="Ormyridae" lsidName="" pageId="10" pageNumber="35" rank="family">Ormyridae</taxonomicName>
<taxonomicName id="A490678262EEE5B077294A99CD73EAE7" class="Insecta" family="Ormyridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="family">Ormyridae</taxonomicName>
) was also reared.
</paragraph>
<caption pageId="10" pageNumber="35">
<paragraph pageId="10" pageNumber="35">
<caption id="B40BF74E009050DC035D457B618614AC" pageId="10" pageNumber="35">
<paragraph id="242FED8CB84B60087491254CEEF82B81" pageId="10" pageNumber="35">
Figure 4. Detail of ventral spine of hypopygium of females of species of the Cynips group: A
<taxonomicName class="Insecta" family="Cynipidae" genus="Cynips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cynips divisa" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="divisa">Cynips divisa</taxonomicName>
<taxonomicName id="D77C091EC5834880F0605773E4399C66" class="Insecta" family="Cynipidae" genus="Cynips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cynips divisa" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="divisa">Cynips divisa</taxonomicName>
, asexual generation B
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, sexual generation C
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, asexual generation D
<taxonomicName class="Insecta" family="Cynipidae" genus="Acraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acraspis erinacei" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="erinacei">Acraspis erinacei</taxonomicName>
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, sexual generation E
<taxonomicName class="Insecta" family="Cynipidae" genus="Cynips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cynips" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="genus">Cynips</taxonomicName>
<taxonomicName id="4A2F62D752986D6C075862564378C3B2" class="Insecta" family="Cynipidae" genus="Cynips" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cynips" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="genus">Cynips</taxonomicName>
=
<taxonomicName class="Insecta" family="Cynipidae" genus="Antron" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Antron douglasi" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="douglasi">Antron douglasi</taxonomicName>
<taxonomicName id="F7C2151A0E2117356E19DD77E82CA5FC" genus="Antron" lsidName="Antron douglasi" pageId="10" pageNumber="35" rank="species" species="douglasi">Antron douglasi</taxonomicName>
F
<taxonomicName class="Insecta" family="Cynipidae" genus="Atrusca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrusca emergens" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="emergens">Atrusca emergens</taxonomicName>
<taxonomicName id="C447E02DAED1566130F7D2785D780240" class="Insecta" family="Cynipidae" genus="Atrusca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrusca emergens" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="emergens">Atrusca emergens</taxonomicName>
G
<taxonomicName class="Insecta" family="Cynipidae" genus="Philonix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonix gigas" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="gigas">Philonix gigas</taxonomicName>
<taxonomicName id="F3C1982FB22B3C419D3781578FCFBA3A" class="Insecta" family="Cynipidae" genus="Philonix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonix gigas" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="gigas">Philonix gigas</taxonomicName>
H
<taxonomicName class="Insecta" family="Cynipidae" genus="Biorhiza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biorhiza" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="genus">Biorhiza</taxonomicName>
<taxonomicName id="D773C7C0014D34DA5ADAA6803C149A22" class="Insecta" family="Cynipidae" genus="Biorhiza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biorhiza" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="genus">Biorhiza</taxonomicName>
=
<taxonomicName class="Insecta" family="Cynipidae" genus="Sphaeroteras" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sphaeroteras mellea" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="mellea">Sphaeroteras mellea</taxonomicName>
<taxonomicName id="AA2F0671A3ADE557C82E44049B2ECC7D" genus="Sphaeroteras" lsidName="Sphaeroteras mellea" pageId="10" pageNumber="35" rank="species" species="mellea">Sphaeroteras mellea</taxonomicName>
. [after
<bibRefCitation author="Liljeblad, J" journalOrPublisher="Zootaxa" pageId="17" pageNumber="42" pagination="1 - 73" title="A fully web-illustrated morphological phylogenetic study of relationships among oak gall wasps and their closest relatives (Hymenoptera: Cynipidae)." volume="1796" year="2008">Liljeblad et al. 2008</bibRefCitation>
<bibRefCitation id="58540403E4D5B87F837F66236B1C2A87" author="Liljeblad, J" journalOrPublisher="Zootaxa" pageId="17" pageNumber="42" pagination="1 - 73" title="A fully web-illustrated morphological phylogenetic study of relationships among oak gall wasps and their closest relatives (Hymenoptera: Cynipidae)." volume="1796" year="2008">Liljeblad et al. 2008</bibRefCitation>
http://www.morphbank.net/].
</paragraph>
</caption>
<caption pageId="10" pageNumber="35">
<paragraph pageId="10" pageNumber="35">
<caption id="888525799206C64B180A602D9A77B62D" pageId="10" pageNumber="35">
<paragraph id="7A0FD10834D9038C031E3EB41B67CA9D" pageId="10" pageNumber="35">
Figure 5. Habitus, forewings and galls of
<taxonomicName class="Insecta" family="Cynipidae" genus="Barucynips" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Barucynips panamensis" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="panamensis">Barucynips panamensis</taxonomicName>
<taxonomicName id="8E05C5CC6FCFE6A286756C10A312D069" genus="Barucynips" lsidName="Barucynips panamensis" pageId="10" pageNumber="35" rank="species" species="panamensis">Barucynips panamensis</taxonomicName>
A Habitus, female B Forewing of female
<normalizedToken originalValue="CG">C-G</normalizedToken>
<normalizedToken id="F1A54CE864CD75B6708476FC08151760" originalValue="CG">C-G</normalizedToken>
Mature galls C Galls on stems D Cluster of galls on midrib of leaves E Section of a gall showing larval cells F Galls on stems G Detail of a single gall.
</paragraph>
</caption>
<caption pageId="10" pageNumber="35">
<paragraph pageId="10" pageNumber="35">
<caption id="90121C5EE1D6EE4EFCB8C43F5A7B8244" pageId="10" pageNumber="35">
<paragraph id="13FECD3635C3B790A5459510C5D10F45" pageId="10" pageNumber="35">
Figure 6.
<taxonomicName class="Insecta" family="Cynipidae" genus="Coffeikokkos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Coffeikokkos korytkowskii" order="Hymenoptera" pageId="10" pageNumber="35" phylum="Arthropoda" rank="species" species="korytkowskii">Coffeikokkos korytkowskii</taxonomicName>
<taxonomicName id="7CE2D9E742C18F9E1C1C47A5673CA7CF" genus="Coffeikokkos" lsidName="Coffeikokkos korytkowskii" pageId="10" pageNumber="35" rank="species" species="korytkowskii">Coffeikokkos korytkowskii</taxonomicName>
: A Head anterior view B Head posterior view C Pronotum antero-dorsal view D Mesosoma lateral view E Mesosoma dorsal view D Propodeum.
</paragraph>
</caption>

View file

@ -1,216 +1,216 @@
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<figureCitation captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Gnamptogenys dentihumera holotype worker (No. G 150067). 1 head in full-face view 2 petiole in lateral view 3 body in dorsal view 4 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures1-4" httpUri="https://binary.pensoft.net/fig/124100" pageId="2" pageNumber="95">Figs 1</figureCitation>
<figureCitation captionStart="Figures 510" captionStartId="F3" captionText="Figures 5 - 10. Gnamptogenys dentihumera paratype queen. 5 head in full-face view 6 forewing in dorsal view 7 hindwing in dorsal view 8 gaster in dorsal view 9 body in dorsal view 10 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures5-10" httpUri="https://binary.pensoft.net/fig/124102" pageId="2" pageNumber="95">, 10</figureCitation>
<figureCitation captionStart="Map 1" captionStartId="F2" captionText="Map 1. Distribution of ten known species of the genus Gnamptogenys in China. The data came from collection specimens and literatures." figureDoi="10.3897/jhr.54.10816.map1" httpUri="https://binary.pensoft.net/fig/124101" pageId="2" pageNumber="95">; Map 1</figureCitation>
<treatment id="58C0C0C7595BEFD78B6D4D4E9B14B7FE" ID-GBIF-Taxon="154473319" LSID="urn:lsid:zoobank.org:act:9307A812-9B3F-4C5A-A4E8-D36B3752C682" httpUri="http://treatment.plazi.org/id/58C0C0C7595BEFD78B6D4D4E9B14B7FE" lastPageId="4" lastPageNumber="96" pageId="2" pageNumber="95" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="37ED4720C58C2703E5897AC6E9A193AF" pageId="2" pageNumber="95" type="nomenclature">
<paragraph id="00140B1BDE9C059A6ADC77594EDA7B27" pageId="2" pageNumber="95">
<taxonomicName id="B01574C57472F7669CF7049B431AACA5" LSID="http://zoobank.org/9307A812-9B3F-4C5A-A4E8-D36B3752C682" authority="Chen, Lattke &amp; Zhou" authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="2" pageNumber="95" phylum="Arthropoda" rank="species" species="dentihumera" status="sp. n.">Gnamptogenys dentihumera Chen, Lattke &amp; Zhou</taxonomicName>
<taxonomicNameLabel id="8AECC42D92AAB2F566D0857EAD8C84B9" pageId="2" pageNumber="95">sp. n.</taxonomicNameLabel>
<figureCitation id="418BD158D5228C6EBE8BC7CC89CF0F73" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Gnamptogenys dentihumera holotype worker (No. G 150067). 1 head in full-face view 2 petiole in lateral view 3 body in dorsal view 4 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures1-4" httpUri="https://binary.pensoft.net/fig/124100" pageId="2" pageNumber="95">Figs 1</figureCitation>
<figureCitation id="BF781E2E5EE1834A633EB9361676EA41" captionStart="Figures 510" captionStartId="F3" captionText="Figures 5 - 10. Gnamptogenys dentihumera paratype queen. 5 head in full-face view 6 forewing in dorsal view 7 hindwing in dorsal view 8 gaster in dorsal view 9 body in dorsal view 10 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures5-10" httpUri="https://binary.pensoft.net/fig/124102" pageId="2" pageNumber="95">, 10</figureCitation>
<figureCitation id="71CB2202782132A0DC952F422B9CDC86" captionStart="Map 1" captionStartId="F2" captionText="Map 1. Distribution of ten known species of the genus Gnamptogenys in China. The data came from collection specimens and literatures." figureDoi="10.3897/jhr.54.10816.map1" httpUri="https://binary.pensoft.net/fig/124101" pageId="2" pageNumber="95">; Map 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="95" type="type material">
<paragraph pageId="2" pageNumber="95">Type material.</paragraph>
<paragraph pageId="2" pageNumber="95">
<subSubSection id="AC6706EEEE09B2F569624B453AFEE315" pageId="2" pageNumber="95" type="type material">
<paragraph id="DC68335C9AA4593624C7261F042453A0" pageId="2" pageNumber="95">Type material.</paragraph>
<paragraph id="D9A113265C693EB7675EBB4E509EB954" pageId="2" pageNumber="95">
Holotype. Worker. CHINA: Guangxi, Huanjiang County, Jiuwan Mountain,
<geoCoordinate degrees="25" direction="north" minutes="11" orientation="latitude" precision="15" seconds="00" value="25.183332">25°11'00&quot;N</geoCoordinate>
<geoCoordinate id="79E6A73405D34D19BA78B7F0059977C5" degrees="25" direction="north" minutes="11" orientation="latitude" precision="15" seconds="00" value="25.183332">25°11'00&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="36" orientation="longitude" precision="15" seconds="57" value="108.61583">108°36'57&quot;E</geoCoordinate>
<geoCoordinate id="47E34BA21315774156967CABBFB17AC3" degrees="108" direction="east" minutes="36" orientation="longitude" precision="15" seconds="57" value="108.61583">108°36'57&quot;E</geoCoordinate>
, 925m, 8.VIII.2015, leg. Zhilin Chen, No. G150067. Paratypes. 12 workers and 1 queen, data the same as holotype. [1 worker holotype, 8 workers pratypes and 1 queen paratype specimens are deposited in the Insect Collection, Guangxi Normal University, Guilin, China (GXNU); 2 workers paratypes will be deposited in the Insect Collection, Southwest Forestry University, Kunming, Yunnan Province, China (SWFU); 2 workers paratypes will be deposited in the Institute of Zoology, Chinese Academy of Sciences, Beijing, China (IZCAS)].
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="96" pageId="2" pageNumber="95" type="holotype worker">
<paragraph pageId="2" pageNumber="95">Holotype worker</paragraph>
<paragraph pageId="2" pageNumber="95">
<subSubSection id="1AC3FCEC6567FF0296747EFE5E02DF9C" lastPageId="3" lastPageNumber="96" pageId="2" pageNumber="95" type="holotype worker">
<paragraph id="7D669EEE94441D7180E92C2CE4E2C21D" pageId="2" pageNumber="95">Holotype worker</paragraph>
<paragraph id="6C02356F8C1A0A4AEFDFF2544CDE84F9" pageId="2" pageNumber="95">
(Figs
<figureCitation captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Gnamptogenys dentihumera holotype worker (No. G 150067). 1 head in full-face view 2 petiole in lateral view 3 body in dorsal view 4 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures1-4" httpUri="https://binary.pensoft.net/fig/124100" pageId="2" pageNumber="95">1-4</figureCitation>
<figureCitation id="258A6BCAAEDFFD366D9624C94C857A42" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Gnamptogenys dentihumera holotype worker (No. G 150067). 1 head in full-face view 2 petiole in lateral view 3 body in dorsal view 4 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures1-4" httpUri="https://binary.pensoft.net/fig/124100" pageId="2" pageNumber="95">1-4</figureCitation>
). TL 4.61, HL 1.45, HW 1.22, CI 84, SL 1.05, SI 86, ED 0.14, PW 0.96, MSL 1.71, PL 0.41, PH 0.95, DPW 0.58, LPI 231, DPI 141. In full-face view head rectangular, longer than broad; posterior margin almost straight, posterior corner bluntly angular, lateral margin weakly convex. Mandible triangular, masticatory margin crenulate. Anterior margin of clypeus with convex to bluntly pointed median lobe. Antennal scape just slightly surpassing posterior corner of head, flagellar segments longer than broad. Eye small, with 10 ommatidia along maximum diameter, situated on midpoint of cephalic lateral margin.
</paragraph>
<paragraph pageId="2" pageNumber="95">In lateral view occipital lamella of head forming a blunt triangular denticle. Dorsum of mesosoma strongly convex, promesonotal suture slightly impressed. Propodeum with short triangular denticle; propodeal declivity weakly concave close to tooth then relatively straight. Petiole roughly trapezoidal, about 2.1 times higher than long, dorsal and anterior margins weakly convex, posterior margin straight, subpetiolar process broadly triangular. Sting extruding.</paragraph>
<paragraph pageId="2" pageNumber="95">In dorsal view anterolateral margin of pronotum with strong constriction, forming pointed denticle on humeral area. Metanotal groove ambiguously evident, evidenced by a change in sculpturing. Petiolar anterior corner rounded, lateral margin weakly convex in dorsal view.</paragraph>
<paragraph lastPageId="3" lastPageNumber="96" pageId="2" pageNumber="95">
<paragraph id="4488CC07383731D5D11118C3E3DF471F" pageId="2" pageNumber="95">In lateral view occipital lamella of head forming a blunt triangular denticle. Dorsum of mesosoma strongly convex, promesonotal suture slightly impressed. Propodeum with short triangular denticle; propodeal declivity weakly concave close to tooth then relatively straight. Petiole roughly trapezoidal, about 2.1 times higher than long, dorsal and anterior margins weakly convex, posterior margin straight, subpetiolar process broadly triangular. Sting extruding.</paragraph>
<paragraph id="6006F17404430EF19DCD83CCEB77BAF2" pageId="2" pageNumber="95">In dorsal view anterolateral margin of pronotum with strong constriction, forming pointed denticle on humeral area. Metanotal groove ambiguously evident, evidenced by a change in sculpturing. Petiolar anterior corner rounded, lateral margin weakly convex in dorsal view.</paragraph>
<paragraph id="C030667E63C524B73F53C08AB9D93065" lastPageId="3" lastPageNumber="96" pageId="2" pageNumber="95">
Mandibular dorsum and clypeus with longitudinal fine striae; cephalic dorsum mostly longitudinally rugulose-foveolate; side of pronotum, propodeum, petiole and third abdominal tergite and sternite rugulose-foveolate; posteromediand pronotum and mesonotum with longitudinal striae; propleuron irregularly rugulose; meso- and
<pageBreakToken pageId="3" pageNumber="96" start="start">metapleuron</pageBreakToken>
<pageBreakToken id="79C687C68FC7EE56F679B6063F5CA7D3" pageId="3" pageNumber="96" start="start">metapleuron</pageBreakToken>
longitudinally striate; fourth abdominal tergite finely irregularly rugulose, sternite rugulae denser than on tergite. Propodeal declivity transversely striate. Head, mesosoma, petiole, and gaster with abundant suberect hairs and sparse decumbent pubescence. Scape and tibiae with sparse sub-decumbent hairs and dense decumbent pubescence. Body color black; mandible, antenna and leg reddish-brown.
</paragraph>
<caption ID-Zenodo-Dep="987899" doi="10.3897/jhr.54.10816.figures1-4" httpUri="https://binary.pensoft.net/fig/124100" pageId="3" pageNumber="96" start="Figures 14" startId="F1">
<paragraph pageId="3" pageNumber="96">
<emphasis bold="true" pageId="3" pageNumber="96">Figures 1-4.</emphasis>
<taxonomicName authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis italics="true" pageId="3" pageNumber="96">Gnamptogenys dentihumera</emphasis>
<caption id="B8C017962782BA23C3AC40E43D9B8287" ID-Zenodo-Dep="987899" doi="10.3897/jhr.54.10816.figures1-4" httpUri="https://binary.pensoft.net/fig/124100" pageId="3" pageNumber="96" start="Figures 14" startId="F1">
<paragraph id="7B624C99A129A4DA0C453E72BD21A5F6" pageId="3" pageNumber="96">
<emphasis id="584061384E78EF46A922F1C38322267E" bold="true" pageId="3" pageNumber="96">Figures 1-4.</emphasis>
<taxonomicName id="610E4A92B7BE2A137A2BCC388EB026B3" authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis id="02719D7A64E07EA08783CE66CC3BDF2D" italics="true" pageId="3" pageNumber="96">Gnamptogenys dentihumera</emphasis>
</taxonomicName>
holotype worker (No. G150067).
<emphasis bold="true" pageId="3" pageNumber="96">1</emphasis>
<emphasis id="AB1C9EF21396454528B7C9AADBFCEE0E" bold="true" pageId="3" pageNumber="96">1</emphasis>
head in full-face view
<emphasis bold="true" pageId="3" pageNumber="96">2</emphasis>
<emphasis id="701A4836A018AFC039715F0842E2586C" bold="true" pageId="3" pageNumber="96">2</emphasis>
petiole in lateral view
<emphasis bold="true" pageId="3" pageNumber="96">3</emphasis>
<emphasis id="3EE7000F2A1A6D7D272D626B08AC649A" bold="true" pageId="3" pageNumber="96">3</emphasis>
body in dorsal view
<emphasis bold="true" pageId="3" pageNumber="96">4</emphasis>
<emphasis id="74A961E9E63A2DB2095FEE208609F708" bold="true" pageId="3" pageNumber="96">4</emphasis>
body in lateral view.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="987913" doi="10.3897/jhr.54.10816.map1" httpUri="https://binary.pensoft.net/fig/124101" pageId="3" pageNumber="96" start="Map 1" startId="F2">
<paragraph pageId="3" pageNumber="96">
<emphasis bold="true" pageId="3" pageNumber="96">Map 1.</emphasis>
<caption id="0695353844849B16D923A8A3C50A7A70" ID-Zenodo-Dep="987913" doi="10.3897/jhr.54.10816.map1" httpUri="https://binary.pensoft.net/fig/124101" pageId="3" pageNumber="96" start="Map 1" startId="F2">
<paragraph id="6BA013CCA2738F1E34D6707B2325DE7A" pageId="3" pageNumber="96">
<emphasis id="1F4D27DC72EC0C310BDB4EEAF9F78709" bold="true" pageId="3" pageNumber="96">Map 1.</emphasis>
Distribution of ten known species of the genus
<taxonomicName authorityName="Roger" authorityYear="1863" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="96">Gnamptogenys</emphasis>
<taxonomicName id="7BE081482AE2B8FE98A79D4EDEF4195D" authorityName="Roger" authorityYear="1863" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="1DB959D94D09C78959F1DB559369E3C5" italics="true" pageId="3" pageNumber="96">Gnamptogenys</emphasis>
</taxonomicName>
in China. The data came from collection specimens and literatures.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="96" type="paratype workers">
<paragraph pageId="3" pageNumber="96">Paratype workers</paragraph>
<paragraph pageId="3" pageNumber="96">(n = 12). TL 4.56-4.65, HL 1.43-1.45, HW 1.20-1.23, CI 83-85, SL 1.03-1.05, SI 84-87, ED 0.13-0.15, PW 0.94-0.95, MSL 1.70-1.72, PL 0.40-0.43, PH 0.93-0.96, DPW 0.56-0.58, LPI 230-233, DPI 137-143. General appearance as in holotype, with some specimens reddish brown to yellowish brown in color. We believe they are callow workers.</paragraph>
<subSubSection id="01246E3867DD557B59611648F4BF144D" pageId="3" pageNumber="96" type="paratype workers">
<paragraph id="5621402929159D439F3E1C1922476BA3" pageId="3" pageNumber="96">Paratype workers</paragraph>
<paragraph id="B212359355321398AAFE8A0338FA0638" pageId="3" pageNumber="96">(n = 12). TL 4.56-4.65, HL 1.43-1.45, HW 1.20-1.23, CI 83-85, SL 1.03-1.05, SI 84-87, ED 0.13-0.15, PW 0.94-0.95, MSL 1.70-1.72, PL 0.40-0.43, PH 0.93-0.96, DPW 0.56-0.58, LPI 230-233, DPI 137-143. General appearance as in holotype, with some specimens reddish brown to yellowish brown in color. We believe they are callow workers.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="96" type="paratype queen">
<paragraph pageId="3" pageNumber="96">Paratype queen</paragraph>
<paragraph pageId="3" pageNumber="96">
<subSubSection id="BA5987CD9D5D71D120397165A3724F08" pageId="3" pageNumber="96" type="paratype queen">
<paragraph id="DBA073CC751972A072EBBABC6494EAE2" pageId="3" pageNumber="96">Paratype queen</paragraph>
<paragraph id="5CA6F041130506F9990BE09D99C025CE" pageId="3" pageNumber="96">
(Figs
<figureCitation captionStart="Figures 510" captionStartId="F3" captionText="Figures 5 - 10. Gnamptogenys dentihumera paratype queen. 5 head in full-face view 6 forewing in dorsal view 7 hindwing in dorsal view 8 gaster in dorsal view 9 body in dorsal view 10 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures5-10" httpUri="https://binary.pensoft.net/fig/124102" pageId="3" pageNumber="96">5-10</figureCitation>
<figureCitation id="E9B474DCBD6844E84A306FFCA39D5A38" captionStart="Figures 510" captionStartId="F3" captionText="Figures 5 - 10. Gnamptogenys dentihumera paratype queen. 5 head in full-face view 6 forewing in dorsal view 7 hindwing in dorsal view 8 gaster in dorsal view 9 body in dorsal view 10 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures5-10" httpUri="https://binary.pensoft.net/fig/124102" pageId="3" pageNumber="96">5-10</figureCitation>
). TL 6.42, HL 1.55, HW 1.21, CI 78, SL 1.11, SI 91, ED 0.25, PW 1.21, MSL 2.31, PL 0.37, PH 1.12, DPW 0.71, LPI 302, DPI 191. In full-face view head rectangular, longer than broad, posterior margin weakly concave, posterior corner bluntly angular, lateral margin almost parallel. Three ocelli on vertex (two lateral and one median) bordered by dark patches. Mandible triangular, masticatory margin crenulate. Anterior margin of clypeus with convex to bluntly pointed median lobe. Antennal scape just slightly surpassing posterior corner of head. Eye situated on midpoint of lateral cephalic margin.
</paragraph>
<paragraph pageId="3" pageNumber="96">In lateral view occipital lamella of head forming blunt point. Dorsum of mesosoma convex. Promesonotal suture obvious. Petiole relatively thin, anterior margin convex, posterior margin straight, dorsal and anterior margin with no obvious boundary, subpetiolar process broadly triangular. Sting extruding.</paragraph>
<paragraph pageId="3" pageNumber="96">In dorsal view anterior margin of pronotum rounded; metanotal groove distinct. Metanotum oval, very narrow, bordered by mesoscutellum and propodeum. Petiole anterior weakly rounded.</paragraph>
<paragraph pageId="3" pageNumber="96">
<paragraph id="FDE06EAC77BED1298F75A0243198B921" pageId="3" pageNumber="96">In lateral view occipital lamella of head forming blunt point. Dorsum of mesosoma convex. Promesonotal suture obvious. Petiole relatively thin, anterior margin convex, posterior margin straight, dorsal and anterior margin with no obvious boundary, subpetiolar process broadly triangular. Sting extruding.</paragraph>
<paragraph id="02ED4C6B27D0DB292F28E3DC245E7F42" pageId="3" pageNumber="96">In dorsal view anterior margin of pronotum rounded; metanotal groove distinct. Metanotum oval, very narrow, bordered by mesoscutellum and propodeum. Petiole anterior weakly rounded.</paragraph>
<paragraph id="C2E0D8B485BBBD1C0FDD6E0DBAE3EA2B" pageId="3" pageNumber="96">
Forewing and hindwing of queen shown in Figs
<figureCitation captionStart="Figures 510" captionStartId="F3" captionText="Figures 5 - 10. Gnamptogenys dentihumera paratype queen. 5 head in full-face view 6 forewing in dorsal view 7 hindwing in dorsal view 8 gaster in dorsal view 9 body in dorsal view 10 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures5-10" httpUri="https://binary.pensoft.net/fig/124102" pageId="3" pageNumber="96">6-7</figureCitation>
<figureCitation id="FA02947C12490C0A9A50BF235C24930A" captionStart="Figures 510" captionStartId="F3" captionText="Figures 5 - 10. Gnamptogenys dentihumera paratype queen. 5 head in full-face view 6 forewing in dorsal view 7 hindwing in dorsal view 8 gaster in dorsal view 9 body in dorsal view 10 body in lateral view." figureDoi="10.3897/jhr.54.10816.figures5-10" httpUri="https://binary.pensoft.net/fig/124102" pageId="3" pageNumber="96">6-7</figureCitation>
.
</paragraph>
<paragraph pageId="3" pageNumber="96">Mandibular dorsum, anterolateral cephalic surface and middle part of clypeus with fine longitudinal striae. Head, pronotum, propodeum and petiole coarsely rugulose; middle part of mesonotum longitudinally striate; propleuron irregularly rugulose; mesopleuron and metapleuron longitudinally rugulose; in lateral view, third abdominal tergite with semicircular rugulae; fourth abdominal tergite with fine short rugulae; propodeal declivity transversely striate. Head, mesosoma, petiole, and gaster with abundant sub-erect hairs and sparse decumbent pubescence; scape and tibiae with sparse sub-decumbent hairs and dense decumbent pubescence. Whole body reddish-brown.</paragraph>
<caption ID-Zenodo-Dep="987915" doi="10.3897/jhr.54.10816.figures5-10" httpUri="https://binary.pensoft.net/fig/124102" pageId="3" pageNumber="96" start="Figures 510" startId="F3">
<paragraph pageId="3" pageNumber="96">
<emphasis bold="true" pageId="3" pageNumber="96">Figures 5-10.</emphasis>
<taxonomicName authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis italics="true" pageId="3" pageNumber="96">Gnamptogenys dentihumera</emphasis>
<paragraph id="1C591486F90DA90874E80792B10A5195" pageId="3" pageNumber="96">Mandibular dorsum, anterolateral cephalic surface and middle part of clypeus with fine longitudinal striae. Head, pronotum, propodeum and petiole coarsely rugulose; middle part of mesonotum longitudinally striate; propleuron irregularly rugulose; mesopleuron and metapleuron longitudinally rugulose; in lateral view, third abdominal tergite with semicircular rugulae; fourth abdominal tergite with fine short rugulae; propodeal declivity transversely striate. Head, mesosoma, petiole, and gaster with abundant sub-erect hairs and sparse decumbent pubescence; scape and tibiae with sparse sub-decumbent hairs and dense decumbent pubescence. Whole body reddish-brown.</paragraph>
<caption id="6A7E91904F9110BF8178381140E26DDF" ID-Zenodo-Dep="987915" doi="10.3897/jhr.54.10816.figures5-10" httpUri="https://binary.pensoft.net/fig/124102" pageId="3" pageNumber="96" start="Figures 510" startId="F3">
<paragraph id="D7B51507058FDB003121CCB0634E94A4" pageId="3" pageNumber="96">
<emphasis id="6392253A5A51969907AD3A43E3B2B62B" bold="true" pageId="3" pageNumber="96">Figures 5-10.</emphasis>
<taxonomicName id="2B46E37962235CC3D1C4B23DD3498C0C" authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis id="E37332C3ECBAA1D21C3B3FCA2C055DC0" italics="true" pageId="3" pageNumber="96">Gnamptogenys dentihumera</emphasis>
</taxonomicName>
paratype queen.
<emphasis bold="true" pageId="3" pageNumber="96">5</emphasis>
<emphasis id="7AAED1694C279B95C932CEEF9BADCD1C" bold="true" pageId="3" pageNumber="96">5</emphasis>
head in full-face view
<emphasis bold="true" pageId="3" pageNumber="96">6</emphasis>
<emphasis id="471D83A6D673F5844A2D34A5B86220BB" bold="true" pageId="3" pageNumber="96">6</emphasis>
forewing in dorsal view
<emphasis bold="true" pageId="3" pageNumber="96">7</emphasis>
<emphasis id="F84E6C1EAC70A1F0ADCA229021F259CE" bold="true" pageId="3" pageNumber="96">7</emphasis>
hindwing in dorsal view
<emphasis bold="true" pageId="3" pageNumber="96">8</emphasis>
<emphasis id="5E1511A0DFB988E969B029B848DC2624" bold="true" pageId="3" pageNumber="96">8</emphasis>
gaster in dorsal view
<emphasis bold="true" pageId="3" pageNumber="96">9</emphasis>
<emphasis id="A45D3B16DD4454EC3C253C8E1534834C" bold="true" pageId="3" pageNumber="96">9</emphasis>
body in dorsal view
<emphasis bold="true" pageId="3" pageNumber="96">10</emphasis>
<emphasis id="01F2FED53BA9F75C89CCA8251433A522" bold="true" pageId="3" pageNumber="96">10</emphasis>
body in lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="96" type="male">
<paragraph pageId="3" pageNumber="96">Male.</paragraph>
<paragraph pageId="3" pageNumber="96">Unknown.</paragraph>
<subSubSection id="269B5AB6222193EB2F685D4D4216B091" pageId="3" pageNumber="96" type="male">
<paragraph id="9BFE6BB0C0CE47EA04DFF882A5EB3190" pageId="3" pageNumber="96">Male.</paragraph>
<paragraph id="85025A324F3A245CB003FA9AE9B068A8" pageId="3" pageNumber="96">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="96" type="habitat">
<paragraph pageId="3" pageNumber="96">Habitat.</paragraph>
<paragraph pageId="3" pageNumber="96">This species was found nesting in deadwood of a branch on the ground in a subtropical evergreen broad-leaved forest, at the altitude of 925 m. All ants came from the same nest.</paragraph>
<subSubSection id="580E4A0132E188CDBC6CAAEE8F82E1DD" pageId="3" pageNumber="96" type="habitat">
<paragraph id="3AC40F86D89C6DF20663AC3FFE804083" pageId="3" pageNumber="96">Habitat.</paragraph>
<paragraph id="BA3CF739C597F4A3D35E26826EB0BC28" pageId="3" pageNumber="96">This species was found nesting in deadwood of a branch on the ground in a subtropical evergreen broad-leaved forest, at the altitude of 925 m. All ants came from the same nest.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="96" type="etymology">
<paragraph pageId="3" pageNumber="96">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="96">
<subSubSection id="430F8307115AD5FD50DB3117CC6EF139" pageId="3" pageNumber="96" type="etymology">
<paragraph id="F638829EB5E469BBB68E827FA0795A52" pageId="3" pageNumber="96">Etymology.</paragraph>
<paragraph id="B7CC2BC262E337C988D9AA78498A074B" pageId="3" pageNumber="96">
This species is named from the Latin words
<normalizedToken originalValue="“dent”">&quot;dent&quot;</normalizedToken>
<normalizedToken id="0D1F562C969624FDB166708F501B0636" originalValue="“dent”">&quot;dent&quot;</normalizedToken>
(denticle) and
<normalizedToken originalValue="“humer”">&quot;humer&quot;</normalizedToken>
<normalizedToken id="171181EDC84A807508FDADCEAF867E2D" originalValue="“humer”">&quot;humer&quot;</normalizedToken>
(humeral), referring to the pronotum humeral area forming a pair of pointed denticles.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="97" pageId="3" pageNumber="96" type="differential diagnosis">
<paragraph pageId="3" pageNumber="96">Differential diagnosis.</paragraph>
<paragraph lastPageId="4" lastPageNumber="97" pageId="3" pageNumber="96">
<subSubSection id="947D1781C0F64A03B8A25B9A148666EE" lastPageId="4" lastPageNumber="97" pageId="3" pageNumber="96" type="differential diagnosis">
<paragraph id="DAFB49675207B32E9D7B895C0044386D" pageId="3" pageNumber="96">Differential diagnosis.</paragraph>
<paragraph id="8625A5A85BB7001C071C5F9D39DFF985" lastPageId="4" lastPageNumber="97" pageId="3" pageNumber="96">
The workers of
<taxonomicName authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis italics="true" pageId="3" pageNumber="96">Gnamptogenys dentihumera</emphasis>
<taxonomicName id="D50A7B3188E68814E331588C9EA86652" authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis id="C64D9E708B8487DAA402676585060B44" italics="true" pageId="3" pageNumber="96">Gnamptogenys dentihumera</emphasis>
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are similar to workers of
<taxonomicName baseAuthorityName="Brown" baseAuthorityYear="1948" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys panda" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="panda">
<emphasis italics="true" pageId="3" pageNumber="96">Gnamptogenys panda</emphasis>
<taxonomicName id="DD5620B25A577CDBD75A860720C15460" baseAuthorityName="Brown" baseAuthorityYear="1948" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys panda" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="panda">
<emphasis id="09F7F857C0CEC5E34E103488FBD6CBF7" italics="true" pageId="3" pageNumber="96">Gnamptogenys panda</emphasis>
</taxonomicName>
, but
<taxonomicName authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis italics="true" pageId="3" pageNumber="96">Gnamptogenys dentihumera</emphasis>
<taxonomicName id="4AB968170EB1EE5F8A531837962467EB" authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="3" pageNumber="96" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis id="A34B6EF378783A1849EA1AE32A3EFCFE" italics="true" pageId="3" pageNumber="96">Gnamptogenys dentihumera</emphasis>
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can be distinguished from the latter by propodeum
<pageBreakToken pageId="4" pageNumber="97" start="start">with</pageBreakToken>
<pageBreakToken id="73C8F666ECBAADFAF8FDEEC1B74E8A5D" pageId="4" pageNumber="97" start="start">with</pageBreakToken>
short triangular denticle and triangular subpetiolar process. The workers of
<taxonomicName authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis italics="true" pageId="4" pageNumber="97">Gnamptogenys dentihumera</emphasis>
<taxonomicName id="026DD8AE4A13293506314B52AA3C4732" authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis id="1E2AE255BB5FB116BA0B71C94F69A4FF" italics="true" pageId="4" pageNumber="97">Gnamptogenys dentihumera</emphasis>
</taxonomicName>
also resemble
<taxonomicName authorityName="Wu &amp; Xiao" authorityYear="1987" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys sinensis" order="Hymenoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis italics="true" pageId="4" pageNumber="97">Gnamptogenys sinensis</emphasis>
<taxonomicName id="5278BE7989277C6024BBF38B50E0B9A9" authorityName="Wu &amp; Xiao" authorityYear="1987" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys sinensis" order="Hymenoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="39AD2D5E98C486F168226EA9833155F0" italics="true" pageId="4" pageNumber="97">Gnamptogenys sinensis</emphasis>
</taxonomicName>
, but differ from the latter by the occipital lamella of head shaped as a blunt triangular denticle, and the pronotal humeral denticle of the workers of
<taxonomicName authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis italics="true" pageId="4" pageNumber="97">Gnamptogenys dentihumera</emphasis>
<taxonomicName id="0739743796C96A572AA1C43FC0FD25C2" authorityName="Chen, Lattke &amp; Zhou" authorityYear="2017" class="Insecta" family="Formicidae" genus="Gnamptogenys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnamptogenys dentihumera" order="Hymenoptera" pageId="4" pageNumber="97" phylum="Arthropoda" rank="species" species="dentihumera">
<emphasis id="A5C9CB8AD1C5D0011729FBB2956F5F20" italics="true" pageId="4" pageNumber="97">Gnamptogenys dentihumera</emphasis>
</taxonomicName>
is short, pointing laterally, unsurpassing the anterior margin of pronotum.
</paragraph>

View file

@ -1,81 +1,81 @@
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<taxonomicName LSID="58CF53A1-471E-576A-B53E-46F60A0EA5BC" authority="Singh" authorityName="Singh" authorityYear="1931" class="Insecta" family="Aleyrodidae" genus="Aleuroclava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleuroclava" order="Hemiptera" pageId="0" pageNumber="121" phylum="Arthropoda" rank="genus">Aleuroclava Singh</taxonomicName>
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<paragraph pageId="0" pageNumber="121">Diagnosis.</paragraph>
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<paragraph id="EF5FA86F1D4EE7F8016620998D4A0A28" pageId="0" pageNumber="121">Diagnosis.</paragraph>
<paragraph id="96578DDA47EB9CED9EADA17FCB6A8884" pageId="0" pageNumber="121">
Puparia small in size, elliptical or subelliptical. Margin with one row of teeth. Submarginal area not separated from dorsal disc, with papillae-like markings in some species, dorsum generally with tubercles. Thoracic tracheal folds and pores not discernible; caudal furrow and pore distinct.
<taxonomicName genus="Vasiform" lsidName="Vasiform orifice" pageId="0" pageNumber="121" rank="species" species="orifice">Vasiform orifice</taxonomicName>
<taxonomicName id="7C9AB92900C271211472F97C4D29AC77" genus="Vasiform" lsidName="Vasiform orifice" pageId="0" pageNumber="121" rank="species" species="orifice">Vasiform orifice</taxonomicName>
generally notched posteriorly; operculum cordate, nearly filling
<taxonomicName genus="Vasiform" lsidName="orifice" pageId="0" pageNumber="121" rank="species" species="orifice">orifice</taxonomicName>
<taxonomicName id="BEE76CE065A15EA28987E57C0A3510AD" genus="Vasiform" lsidName="orifice" pageId="0" pageNumber="121" rank="species" species="orifice">orifice</taxonomicName>
; lingual hidden.
</paragraph>
<caption doi="10.3897/zookeys.991.47725.figures1-2" httpUri="https://binary.pensoft.net/fig/474325" pageId="0" pageNumber="121" start="Figures 12" startId="F1">
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<emphasis bold="true" pageId="0" pageNumber="121">Figures 1-2.</emphasis>
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<emphasis id="1BC184DCD03B3F2E0C0EC995C3847D27" bold="true" pageId="0" pageNumber="121">Figures 1-2.</emphasis>
Leaves of host plant
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<emphasis italics="true" pageId="0" pageNumber="121">Schima superba</emphasis>
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<emphasis id="6C1975F7E639BB21CB4CA0A5242DD5A3" italics="true" pageId="0" pageNumber="121">Schima superba</emphasis>
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infested by
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<emphasis italics="true" pageId="0" pageNumber="121">A. schimae</emphasis>
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<emphasis id="EB635323D790A2CA76ADAF315F867EE0" italics="true" pageId="0" pageNumber="121">A. schimae</emphasis>
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sp. nov.
</paragraph>

View file

@ -1,92 +1,92 @@
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@ -109,7 +109,7 @@ Desf.,
. Mus. Hist. Nat. 4: 5 (1818).
<taxonomicName id="A28F061B856FCA6F61EE9B6DAF54BD6C" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Vangueria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vangueria rubiginosa" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="rubiginosa" status="comb. nov.">Vangueria rubiginosa</taxonomicName>
(Desf.) Lantz, Pl. Syst. Evol. 253: 181 (2005), non
<taxonomicName id="C5F755C6055667FD4911AF577B734DC0" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" family="Rubiaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="V. rubiginosa" order="Gentianales" pageId="0" pageNumber="145" rank="species" species="rubiginosa" status="comb. nov.">V. rubiginosa</taxonomicName>
<taxonomicName id="C5F755C6055667FD4911AF577B734DC0" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="V. rubiginosa" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="rubiginosa" status="comb. nov.">V. rubiginosa</taxonomicName>
K.Schum., Bot. Jahrb. Syst. 23: 457 (1897) [=
<taxonomicName id="6B05B64C1910D6D54A87336CB25CAC0D" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Rytigynia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rytigynia rubiginosa" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="rubiginosa" status="comb. nov.">Rytigynia rubiginosa</taxonomicName>
(K.Schum.) Robyns, Bull. Jard. Bot.
@ -127,7 +127,7 @@ Welw., J. Trav. Nat. Hist. 1: 29 (1868), nomen nudum.
<paragraph id="2131D92291EF304F2DC8F70D0815ED6B" pageId="0" pageNumber="145">
<taxonomicName id="973AF88C2C6C709D52ECAE81F9EAE456" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Ancylanthos" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ancylanthos bainesii" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="bainesii" status="comb. nov.">Ancylanthos bainesii</taxonomicName>
Hiern in Oliv. (ed.), Fl. Trop. Afr. 3: 160 (1877). Type:
<taxonomicName id="F472AACB2C776203BCB6EDC807629F7A" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" family="Rubiaceae" genus="Baines" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Baines" order="Gentianales" pageId="0" pageNumber="145" rank="genus" status="comb. nov.">Baines</taxonomicName>
<taxonomicName id="F472AACB2C776203BCB6EDC807629F7A" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Baines" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Baines" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus" status="comb. nov.">Baines</taxonomicName>
s.n. (lectotype: K (K000412071) designated by
<bibRefCitation id="0D7AA31BC211D82F7287401FA26B0C1E" DOI="https://doi.org/10.2307/3666476" author="Robyns, W" journalOrPublisher="Bulletin du Jardin Botanique de l'Etat Bruxelles" pageId="0" pageNumber="145" pagination="1 - 359" refId="B49" refString="Robyns, W, 1928. Tentamen monographiae Vangueriae generis affinium. Bulletin du Jardin Botanique de l'Etat Bruxelles 11 (1): 1 - 359, DOI: https://doi.org/10.2307/3666476" title="Tentamen monographiae Vangueriae generis affinium." url="https://doi.org/10.2307/3666476" volume="11" year="1928">Robyns 1928</bibRefCitation>
: 329).
@ -149,7 +149,7 @@ is a small woody species with conspicuous orange flowers. It can form single-ste
</paragraph>
<paragraph id="EAF9E298CB4070355C13F4019F1A8AB1" pageId="0" pageNumber="145">
Molecular studies of the tribe
<taxonomicName id="2C6BD3670A91E3C97042F6A8FEE8C209" family="Rubiaceae" kingdom="Plantae" lsidName="" order="Gentianales" pageId="0" pageNumber="145" rank="tribe" tribe="Vanguerieae">Vanguerieae</taxonomicName>
<taxonomicName id="2C6BD3670A91E3C97042F6A8FEE8C209" class="Magnoliopsida" family="Rubiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="tribe" tribe="Vanguerieae">Vanguerieae</taxonomicName>
A.Rich. ex Dumort. (
<bibRefCitation id="BC29648AE28DC91D33D18F07667DC8FD" DOI="https://doi.org/10.1111/j.1095-8339.2004.00338.x" author="Lantz, H" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="145" pagination="257 - 283" refId="B37" refString="Lantz, H, Bremer, B, 2004. Phylogeny inferred from morphology and DNA data: Characterising well-supported groups in Vanguerieae (Rubiaceae). Botanical Journal of the Linnean Society 146 (3): 257 - 283, DOI: https://doi.org/10.1111/j.1095-8339.2004.00338.x" title="Phylogeny inferred from morphology and DNA data: Characterising well-supported groups in Vanguerieae (Rubiaceae)." url="https://doi.org/10.1111/j.1095-8339.2004.00338.x" volume="146" year="2004">Lantz and Bremer 2004</bibRefCitation>
;
@ -171,7 +171,7 @@ Juss., and combinations under the latter were made in
<emphasis id="36E03BF285CD54CF908355A530DA4AE7" italics="true" pageId="0" pageNumber="145">Vangueria rubiginosa</emphasis>
</taxonomicName>
(Desf.) Lantz is a later homonym of
<taxonomicName id="172723066D532FEFE1B9DB8DCAF3172D" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" family="Rubiaceae" kingdom="Plantae" lsidName="V. rubiginosa" order="Gentianales" pageId="0" pageNumber="145" rank="species" species="rubiginosa">
<taxonomicName id="172723066D532FEFE1B9DB8DCAF3172D" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="V. rubiginosa" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="rubiginosa">
<emphasis id="EB350AF445B095AF300147CF35E16B13" italics="true" pageId="0" pageNumber="145">V. rubiginosa</emphasis>
</taxonomicName>
K.Schum. (
@ -215,7 +215,7 @@ were published simultaneously and the epithets are both available under
<emphasis id="CFCED510B9DBFD4BD320F31C9379A129" italics="true" pageId="0" pageNumber="145">Vangueria</emphasis>
</taxonomicName>
, so either could be used as a basionym for a new combination to replace the illegitimate
<taxonomicName id="F0B7BD539005200E36C6A3750D6FFDE8" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" family="Rubiaceae" kingdom="Plantae" lsidName="V. rubiginosa" order="Gentianales" pageId="0" pageNumber="145" rank="species" species="rubiginosa">
<taxonomicName id="F0B7BD539005200E36C6A3750D6FFDE8" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="V. rubiginosa" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="rubiginosa">
<emphasis id="52E2DEFC88D35552E55EE9527838AD54" italics="true" pageId="0" pageNumber="145">V. rubiginosa</emphasis>
</taxonomicName>
(Desf.) Lantz - we have chosen to use
@ -253,17 +253,17 @@ cited two specimens in his description of
<emphasis id="34F5E7A35B2CF4D8907EA584A964CCF1" italics="true" pageId="0" pageNumber="145">Ancylanthos bainesii</emphasis>
</taxonomicName>
. One, without a locality, was just cited as &quot;T.
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<taxonomicName id="EDA4F0EB8F0B23C41309B9356ED0CDA6" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Baines" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Baines" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">Baines</taxonomicName>
!&quot;. The second is the Chapman &amp;
<taxonomicName id="18DE0428CEE8B092574C603C99C8E2DF" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" family="Rubiaceae" genus="Baines" kingdom="Plantae" lsidName="Baines" order="Gentianales" pageId="0" pageNumber="145" rank="genus">Baines</taxonomicName>
<taxonomicName id="18DE0428CEE8B092574C603C99C8E2DF" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Baines" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Baines" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">Baines</taxonomicName>
collection labelled &quot;lat. 23°&quot;.
<bibRefCitation id="A5A25BFDD96C5EB1D3781970F50C19B2" DOI="https://doi.org/10.1007/s12225-016-9656-3" author="Goyder, DJ" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="145" pagination=": 46" refId="B25" refString="Goyder, DJ, 2016b. Localities of plants collected by Thomas Baines on the Baines -Chapman expedition to Lake Ngami and Victoria Falls 1861-1863. Kew Bulletin 71 (3): 46, DOI: https://doi.org/10.1007/s12225-016-9656-3" title="Localities of plants collected by Thomas Baines on the Baines - Chapman expedition to Lake Ngami and Victoria Falls 1861 - 1863." url="https://doi.org/10.1007/s12225-016-9656-3" volume="71" year="2016 b">Goyder (2016b)</bibRefCitation>
drew attention to problems localising Chapman &amp;
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<taxonomicName id="E3B9E648634688335299E425A1F38B22" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Baines" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Baines" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">Baines</taxonomicName>
' collections particularly those labelled lat. 23° - if taken literally, this would imply they were collected near the Namibian coast E of Walvis Bay, but as this is desert and therefore unsuitable habitat for most of the species bearing these labels, it is much more likely that, given the well-documented route of the expedition, they originated in northern Botswana.
<bibRefCitation id="E6AE5FE8A6EDCB419FA9EB5D3A5192BF" DOI="https://doi.org/10.2307/3666476" author="Robyns, W" journalOrPublisher="Bulletin du Jardin Botanique de l'Etat Bruxelles" pageId="0" pageNumber="145" pagination="1 - 359" refId="B49" refString="Robyns, W, 1928. Tentamen monographiae Vangueriae generis affinium. Bulletin du Jardin Botanique de l'Etat Bruxelles 11 (1): 1 - 359, DOI: https://doi.org/10.2307/3666476" title="Tentamen monographiae Vangueriae generis affinium." url="https://doi.org/10.2307/3666476" volume="11" year="1928">Robyns (1928</bibRefCitation>
: 329) effectively lectotypified the name by citing the unlocalised &quot;T.
<taxonomicName id="4A38152C3D03C61D5B18A54955D323E5" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" family="Rubiaceae" genus="Baines" kingdom="Plantae" lsidName="Baines" order="Gentianales" pageId="0" pageNumber="145" rank="genus">Baines</taxonomicName>
<taxonomicName id="4A38152C3D03C61D5B18A54955D323E5" authorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" authorityYear="2023" baseAuthorityName="Goyder &amp; Davies &amp; Finckh &amp; Gomes &amp; Gonçalves &amp; Meller &amp; Paton" baseAuthorityYear="2023" class="Magnoliopsida" family="Rubiaceae" genus="Baines" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Baines" order="Gentianales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">Baines</taxonomicName>
&quot; collection as the type.
</paragraph>
</subSubSection>

View file

@ -1,236 +1,236 @@
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<treatment ID-GBIF-Taxon="159365013" LSID="urn:lsid:plazi:treatment:58DFEF222735FD2714C1FCFF8E4D5FFE" httpUri="http://treatment.plazi.org/id/58DFEF222735FD2714C1FCFF8E4D5FFE" lastPageId="7" lastPageNumber="67" pageId="4" pageNumber="65">
<subSubSection pageId="4" pageNumber="65" type="nomenclature">
<paragraph pageId="4" pageNumber="65">
<taxonomicName LSID="http://www.species-id.net/wiki/Laccoporus_nigritulus" authority="Gschwendtner, 1936" authorityName="Gschwendtner" authorityYear="1936" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus (Gschwendtner, 1936)</taxonomicName>
<treatment id="58DFEF222735FD2714C1FCFF8E4D5FFE" ID-GBIF-Taxon="159365013" LSID="urn:lsid:plazi:treatment:58DFEF222735FD2714C1FCFF8E4D5FFE" httpUri="http://treatment.plazi.org/id/58DFEF222735FD2714C1FCFF8E4D5FFE" lastPageId="7" lastPageNumber="67" pageId="4" pageNumber="65" scope_family="Dytiscidae" scope_order="Coleoptera">
<subSubSection id="B10BB3530417B8B7581D0AA0C04D7FEE" pageId="4" pageNumber="65" type="nomenclature">
<paragraph id="2B44716F85545C4DFA3B2C26D2B06011" pageId="4" pageNumber="65">
<taxonomicName id="6B6EEAB184C6C5A2DB66E8F245035082" ID-CoL="3RMD5" LSID="http://www.species-id.net/wiki/Laccoporus_nigritulus" authority="Gschwendtner, 1936" authorityName="Gschwendtner" authorityYear="1936" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus (Gschwendtner, 1936)</taxonomicName>
Figs 10-15
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="65" type="reference_group">
<paragraph pageId="4" pageNumber="65">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
<subSubSection id="749FA532F8DDB2E21A8C191F572E7356" pageId="4" pageNumber="65" type="reference_group">
<paragraph id="3B72DC2A81464907FC1F8F28F3ADA778" pageId="4" pageNumber="65">
<taxonomicName id="F0048FBFEF2155C12D9329A53A8118F4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
var. nigritulus
<bibRefCitation author="Gschwendtner, L" journalOrPublisher="Records of the Indian Museum" pageId="7" pageNumber="68" pagination="365 - 374" title="Interessante und neue Schwimmkaefer des Indischen Museums in Calcutta" volume="37" year="1936">Gschwendtner 1936</bibRefCitation>
<bibRefCitation id="CD8A708D89E3F2E581839E5B53ECCE49" author="Gschwendtner, L" journalOrPublisher="Records of the Indian Museum" pageId="7" pageNumber="68" pagination="365 - 374" title="Interessante und neue Schwimmkaefer des Indischen Museums in Calcutta" volume="37" year="1936">Gschwendtner 1936</bibRefCitation>
: 367.
</paragraph>
<paragraph pageId="4" pageNumber="65">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
<paragraph id="5179063E35A4BAB41B314ABF04580BE4" pageId="4" pageNumber="65">
<taxonomicName id="21AA89F35451C955D600A04C8A76DC24" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
(Gschwendtner):
<bibRefCitation author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani 1970</bibRefCitation>
<bibRefCitation id="067277C9860752A3CB6896728F1CE235" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani 1970</bibRefCitation>
: 563-564 (new combination, new status);
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson 2001</bibRefCitation>
<bibRefCitation id="DDB2C46D7DA1894F7C5FAF12FBCFD4B9" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson 2001</bibRefCitation>
: 253.
</paragraph>
<paragraph pageId="4" pageNumber="65">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="The Annals and Magazine of Natural History (11)" pageId="7" pageNumber="68" pagination="97 - 114" title="A contribution to the study of the Dytiscidae. - I. (Coleoptera, Adephaga)." volume="3" year="1939">Balfour-Browne 1939</bibRefCitation>
<paragraph id="ABD7E3C75E87D45B63305A96CD013D4D" pageId="4" pageNumber="65">
<taxonomicName id="2FCE0DB958A28FE0F2D2C897350859DB" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<bibRefCitation id="9EF06DEC778BD62F48A37216A4C73CC5" author="Balfour-Browne, J" journalOrPublisher="The Annals and Magazine of Natural History (11)" pageId="7" pageNumber="68" pagination="97 - 114" title="A contribution to the study of the Dytiscidae. - I. (Coleoptera, Adephaga)." volume="3" year="1939">Balfour-Browne 1939</bibRefCitation>
: 104;
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson 2001</bibRefCitation>
<bibRefCitation id="6DDD2BBBD9F10E4315A6EA508F0888DD" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson 2001</bibRefCitation>
: 253; new synonymy.
</paragraph>
<paragraph pageId="4" pageNumber="65">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
<paragraph id="4B4F772AC6293C8A8429C321FB8D663E" pageId="4" pageNumber="65">
<taxonomicName id="B1DE07F6510052FD1EE891FB3ED78CE7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
sp.:
<bibRefCitation author="Nilsson, AN" editor="Jaech, MA" journalOrPublisher="Zoologisch-Botanische Gesellschaft in Oesterreich and Wiener Coleopterologenverein, Wien" pageId="7" pageNumber="68" pagination="35 - 96" title="Noteridae and Dytiscidae: Annotated checklist of the Noteridae and Dytiscidae of China (Coleoptera)" volumeTitle="Water beetles of China, Vol. 1" year="1995">Nilsson 1995</bibRefCitation>
<bibRefCitation id="B5BABBB48BB2FD90FE9E626F6B98823E" author="Nilsson, AN" editor="Jaech, MA" journalOrPublisher="Zoologisch-Botanische Gesellschaft in Oesterreich and Wiener Coleopterologenverein, Wien" pageId="7" pageNumber="68" pagination="35 - 96" title="Noteridae and Dytiscidae: Annotated checklist of the Noteridae and Dytiscidae of China (Coleoptera)" volumeTitle="Water beetles of China, Vol. 1" year="1995">Nilsson 1995</bibRefCitation>
: 71.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="65" type="type localities">
<paragraph pageId="4" pageNumber="65">Type localities:</paragraph>
<paragraph pageId="4" pageNumber="65">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
<subSubSection id="F35840086E22A7F5C30F235352E66654" pageId="4" pageNumber="65" type="type localities">
<paragraph id="C414F5F0F23DB9B31C7FDB712819EE0C" pageId="4" pageNumber="65">Type localities:</paragraph>
<paragraph id="1DC57B2015944591EAA8B2D01D5C670B" pageId="4" pageNumber="65">
<taxonomicName id="C2478677B62EA95236151D918DFDE284" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
var. nigritulus: China: Tibet, Te-ring Gompa;
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<taxonomicName id="A3FB047F7F86DACC8901B55D8B5FE3D5" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
: China: Tibet, Gyangtse.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65" type="type material">
<paragraph pageId="4" pageNumber="65">Type material:</paragraph>
<paragraph pageId="4" pageNumber="65">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
<subSubSection id="A432A10D5CC45F4C6C9B8E6BE1B42FF7" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65" type="type material">
<paragraph id="10982AEB25262A530FCE1515E95B9AC0" pageId="4" pageNumber="65">Type material:</paragraph>
<paragraph id="848B7E9B779FE0A25C6BC105B9FCEF02" pageId="4" pageNumber="65">
<taxonomicName id="136AE52CFAAB3C9EFFD82AF894514E70" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
var. nigritulus: 10 syntypes, Te-ring Gompa. 14,000 ft. Tibet (F.H. Stewart) in ZSIC [not studied].
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson (2001)</bibRefCitation>
<bibRefCitation id="E0D21427754F910A2C27520C97A9A667" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson (2001)</bibRefCitation>
mentioned a lectotype designation by
<bibRefCitation author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
<bibRefCitation id="8BD93B87765CA70B16A61DAB181C167D" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
, but Vazirani mentioned that he studied the holotype and one paratype, the type status is therefore rather unclear and we could not access types from ZSIC to clarify the situation.
</paragraph>
<paragraph pageId="5" pageNumber="66">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="viator">
<pageBreakToken pageId="5" pageNumber="66" start="start">Laccoporus</pageBreakToken>
<paragraph id="1234CED17ED7748E78E369092A7265EC" pageId="5" pageNumber="66">
<taxonomicName id="1054F635E49E67F5992425303AC0E4CE" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="viator">
<pageBreakToken id="8379D4CCE23E46239F505EA3C2B9562F" pageId="5" pageNumber="66" start="start">Laccoporus</pageBreakToken>
viator
</taxonomicName>
: Holotype,
<normalizedToken originalValue="„allotype”">&quot;allotype&quot;</normalizedToken>
<normalizedToken id="72B7C3D2FADC3B62BFFFD9A16EBD3822" originalValue="„allotype”">&quot;allotype&quot;</normalizedToken>
and 10 other paratypes in BMNH. We studied 2 ♂♂ paratypes labelled:
<normalizedToken originalValue="„Co-type”">&quot;Co-type&quot;</normalizedToken>
<normalizedToken id="C546F7E52FBECE133452875CAB823230" originalValue="„Co-type”">&quot;Co-type&quot;</normalizedToken>
, &quot;Gyangtse. / 13,000 ft. / June 1904. / Tibet Expedit. / H.J. Walton. / 1903
<normalizedToken originalValue="172.”">-172.&quot;</normalizedToken>
<normalizedToken id="A0F46FA38F5F80A8ADE16321A817A99A" originalValue="172.”">-172.&quot;</normalizedToken>
, &quot;Laccoporus / viator. B-B. / Co-type.&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="66" type="other material examined">
<paragraph pageId="5" pageNumber="66">Other material examined:</paragraph>
<paragraph pageId="5" pageNumber="66">
<subSubSection id="72B610E288FCD7D33B51339C74B0BFF0" pageId="5" pageNumber="66" type="other material examined">
<paragraph id="23EB4B598A15E346E1E407D170543D82" pageId="5" pageNumber="66">Other material examined:</paragraph>
<paragraph id="C1B373200C2F2F4B24E682D756193272" pageId="5" pageNumber="66">
20 exs (SYSU): 2 ♂♂CHINA: Xizang, Dingri (Tering), 4300 m, 2.vi.1974, leg. Xuezhong Zhang, Academia Sinica; 1 ♀ the same data as male; 1 ♀ Xizang, suburb of Rikaze, 3826 m, 20-23.vii.1986, leg. Geqiu Liang; 5 ♂♂, 3 ♀♀ Xizang, Rikaze, 3862 m, 4.viii.1986, leg. Geqiu Liang; 1 ♂ the same data as the former, with a label &quot;Laccophilus indicus?&quot;; 4 ♂♂, 3 ♀♀ Xizang, Mozhugongka County, 7.viii.1986, leg. Geqiu Liang; 8 exs (MTD, NMPC, CHF) &quot;Tibet, Yamtso-ufer bei / Nagartze, N28°58'31,9 / E90°24'6,0; 4450mNN; / 29.VII. 1998; leg. O.
<normalizedToken originalValue="Jäger”">Jaeger&quot;</normalizedToken>
<normalizedToken id="ABC8A2AF240831BC75CE51D12C7B1F42" originalValue="Jäger”">Jaeger&quot;</normalizedToken>
, &quot;Laccoporus nigritulus / (Gschwendtner) / Fery det. 1999&quot;; 11 exs (MTD, CHF) &quot;Tibet, Tingriebene / N28°34'39,7/E86°36'52,7; 4400 m / 3.-5.VIII. 1998, lg.
<normalizedToken originalValue="Jäger“">Jaeger&quot;</normalizedToken>
<normalizedToken id="CA5038E5BB6A6915D52E7CA66665B267" originalValue="Jäger“">Jaeger&quot;</normalizedToken>
, &quot;Laccoporus nigritulus / (Gschwendtner) / Fery det. 1999&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="66" type="redescription">
<paragraph pageId="5" pageNumber="66">Redescription.</paragraph>
<paragraph pageId="5" pageNumber="66">
<subSubSection id="974421BC02018C91EC0726184E5B79C8" pageId="5" pageNumber="66" type="redescription">
<paragraph id="47456DE7B72C225493028754592E6F3F" pageId="5" pageNumber="66">Redescription.</paragraph>
<paragraph id="A295E13E88487B5B323A7CC47BEDEF4B" pageId="5" pageNumber="66">
Measurements. Length 4.5-5.0 mm, width 2.5-2.6 mm (paratypes of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<taxonomicName id="040E7E0DFB2A104B96D4C7C7A16FAAF7" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
: length 4.5-4.8 mm, width 2.3-2.6 mm).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="colouration">
<paragraph pageId="5" pageNumber="66">Colouration</paragraph>
<paragraph lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
<subSubSection id="56300221AC574C4EAF420B2541FE6C97" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="colouration">
<paragraph id="27CAD3BFC6AF490C714219E627F52A4B" pageId="5" pageNumber="66">Colouration</paragraph>
<paragraph id="7492459CFA4EBF007E52EBC2E61F0357" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
(Figs 10, 11, 15). Dorsum uniformly yellow brown. Head yellowish brown, with one short dark brown line near antennae. Antenna with antennomeres 1-4 and base of antennomeres 5-6 yellow brown, antennomeres 7-11 andapical parts of antennomeres 5-6 dark brown. Maxillary palpomeres yellow brown, apical palpomere apically dark. Labial palpomeres yellow brown, apical palpomere dark. Pronotum sometimes paler with darker median, anterior and posterior transverse bands. Elytra clearly darker than pronotum, but with the same color at base as pronotum. Ventral surface of head and thorax black. Abdomen black, ventrites 3-6 yellow brown along posterior margins (e.g. Fig. 15). Legs yellow brown with coxae black, posterior half of metacoxal process yellowstructures(see http://www.species-id.net/wiki/Laccoporus_nigritulus for high resolution images of surfaces). Head with surface sculpture consisting of small irregular polygonal meshes, with some sparse coarse and shallow punctures in posterior half. Meshes somewhat stronger on posterior portion. Clypeus with transverse series of large punctures present anteriorly. Labrum with very deep excavation, surface with short transverse meshes, anterior notch with dense and short white setae. Antenna filiform, not dilated. Pronotum more or less depressed laterally, without lateral bead, with small irregular polygonal meshes, sometimes very tiny punctures detectable. Posterior corner almost rectangular. Elytra with base as wide as posterior portion of pronotum, surface with irregular polygonal meshes in size and shape, sculpture more or less stronger than that on pronotum, tiny punctures very sparse and clearly detectable. Prosternal process thin and long,bisinuate in lateral view, almost reaching posterior margin of mesocoxae. Metaventrite medially with some coarse and strong punctures behind mesocoxae. Metacoxae and metasternum strongly and irregularly scratched, with dense longitudinal sculptures and sparse tiny punctures. Pro- and mesofemora with tuft of long setae at basal portion posteriorly. Metafemora with fine and dense sculpture and tiny punctures. Metatibia with inner spur longer than outer one. Pro- and mesotarsi smooth. Metatarsi smooth, with fine sculpture dorsally. First tarsomere of metatarsi almost as long as following two tarsomeres combined. Claws
<pageBreakToken pageId="6" pageNumber="67" start="start">normally</pageBreakToken>
<pageBreakToken id="06B41EFC279AE7110B96C41CD4BC8554" pageId="6" pageNumber="67" start="start">normally</pageBreakToken>
curved, protarsal claws longer than metatarsal claws. Metatarsomere 1 almost as long as metatarsomeres 2-3 combined. Claws regularly curved, proclaws longer than metaclaws.
</paragraph>
<caption pageId="6" pageNumber="67">
<paragraph pageId="6" pageNumber="67">
<caption id="965094D4338C4A92D236589AAA48D62A" pageId="6" pageNumber="67">
<paragraph id="0BF3210FBA2BBE870D29044A04EB9FEE" pageId="6" pageNumber="67">
Figures 10-15.
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
<taxonomicName id="8AFDE4D894360747C97D545C79A4DB82" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
male 10 habitus, dorsal view 11 habitus, ventral view 12 left paramere 13 right paramere 14 median lobe of aedeagus, lateral view 15 ventral view of 6th ventrite.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="6" pageNumber="67" type="male">
<paragraph pageId="6" pageNumber="67">Male.</paragraph>
<paragraph pageId="6" pageNumber="67">Protarsus not dilated. Median lobe and parameres as in Figs 12-14.</paragraph>
<subSubSection id="29028309650CE16C0E097B79FE22D715" pageId="6" pageNumber="67" type="male">
<paragraph id="77179F9AA5C06D1C9F9F789FC2581578" pageId="6" pageNumber="67">Male.</paragraph>
<paragraph id="421A713AE26138A466A6FE9CCEBFE33B" pageId="6" pageNumber="67">Protarsus not dilated. Median lobe and parameres as in Figs 12-14.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="67" type="distribution">
<paragraph pageId="6" pageNumber="67">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="67">China: Xizang.</paragraph>
<subSubSection id="2EED11E81E7B6765BB2F691CFC4EBEAD" pageId="6" pageNumber="67" type="distribution">
<paragraph id="179F7BAD95CAB2191C7C4C451B2AEFA4" pageId="6" pageNumber="67">Distribution.</paragraph>
<paragraph id="4E8F5E664E1D3D6818FC6BEC5D02EC20" pageId="6" pageNumber="67">China: Xizang.</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67" type="remarks">
<paragraph pageId="6" pageNumber="67">Remarks.</paragraph>
<paragraph pageId="6" pageNumber="67">
<subSubSection id="2C853BA12F7383D26BBA2760582366BF" lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67" type="remarks">
<paragraph id="8EB4758C974ACFF19441A6BF38462407" pageId="6" pageNumber="67">Remarks.</paragraph>
<paragraph id="29BD3E009C1D040B6DF156EDF205F381" pageId="6" pageNumber="67">
Genus
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">Laccoporus</taxonomicName>
<taxonomicName id="D6A292997620ABF48BCD81148E1169AB" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">Laccoporus</taxonomicName>
was erected by
<bibRefCitation author="Balfour-Browne, J" journalOrPublisher="The Annals and Magazine of Natural History (11)" pageId="7" pageNumber="68" pagination="97 - 114" title="A contribution to the study of the Dytiscidae. - I. (Coleoptera, Adephaga)." volume="3" year="1939">Balfour-Browne (1939)</bibRefCitation>
<bibRefCitation id="3EA2B2E0C22596525DB80ABCE11537BE" author="Balfour-Browne, J" journalOrPublisher="The Annals and Magazine of Natural History (11)" pageId="7" pageNumber="68" pagination="97 - 114" title="A contribution to the study of the Dytiscidae. - I. (Coleoptera, Adephaga)." volume="3" year="1939">Balfour-Browne (1939)</bibRefCitation>
with
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<taxonomicName id="8B1DBD9393BE7221642CAB78454A3E43" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
Balfour-Browne, 1939 as the type species. This author had no knowledge about
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicornis" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="apicornis">Laccophilus apicornis</taxonomicName>
<taxonomicName id="FE5B61F68B902DA1F75D25242AC9B785" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicornis" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="apicornis">Laccophilus apicornis</taxonomicName>
var. nigritulus Gschwendtner, 1936. The latter taxon was transferred to
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">Laccoporus</taxonomicName>
<taxonomicName id="9A50D23F66D3138166D64BBB938B16CC" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">Laccoporus</taxonomicName>
and raised to species status by
<bibRefCitation author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
<bibRefCitation id="3A4E6413BD15E3767689E9BED0177978" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
who studied two syntypes housed in ZSIC.
<bibRefCitation author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
<bibRefCitation id="C4B28C19E122630E7E50CE2F11F2C6E3" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
had doubts that
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<taxonomicName id="5982219F1A2923EEB0EB9121A8B7DC25" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
and
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
<taxonomicName id="A2F81753AA117C504F60DBDDFEE8609F" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
were different species, but he could not study
<normalizedToken originalValue="Balfour-Brownes">Balfour-Browne's</normalizedToken>
<normalizedToken id="9483515D32EFDFFDB6D0A7FE489ED3FB" originalValue="Balfour-Brownes">Balfour-Browne's</normalizedToken>
types. We have no access to specimens in ZSIC, but
<bibRefCitation author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
<bibRefCitation id="AD0F16125605EEE1CDE54198AB732064" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
provided an illustration of the ventral side of the median lobe of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
<taxonomicName id="E8F7A6C866307EF22F2EF5C12CEC096E" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
, which agrees with our specimens.
</paragraph>
<paragraph pageId="6" pageNumber="67">
<paragraph id="4185D86B51B2ADF81B35ECE0237228AD" pageId="6" pageNumber="67">
The types of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<taxonomicName id="65938EFD8BC31F62767535081E5B43E9" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
are from Gyangtse (Jiangzi). The types of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
<taxonomicName id="AD57E8B127131FC97E9C92E1527DED74" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
were collected from
<normalizedToken originalValue="“Te-ring”">&quot;Te-ring&quot;</normalizedToken>
<normalizedToken id="02C306FAEF7C15FD2EB2489D4F4FF4E0" originalValue="“Te-ring”">&quot;Te-ring&quot;</normalizedToken>
(Dingri) and
<normalizedToken originalValue="“Gompa”">&quot;Gompa&quot;</normalizedToken>
<normalizedToken id="83E8F623831719D3E37983E4593631AD" originalValue="“Gompa”">&quot;Gompa&quot;</normalizedToken>
(Gangba). The distance between these sites is only about 100-120 km (Fig. 16). The two latter sites are not far (about 90-120 km respectively) from Rikaze where some of our newly studied specimens originate from. The altiude of these three sites is 3900-4300 m.
</paragraph>
<caption pageId="6" pageNumber="67">
<paragraph pageId="6" pageNumber="67">
<caption id="3FA9661644508C67B1AAB042CE3D2EB5" pageId="6" pageNumber="67">
<paragraph id="5C44C26580F0D7F2048C6964004A2683" pageId="6" pageNumber="67">
Figure 16. Xizang Autonomous Region, collecting localities of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus fengi" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="fengi">Rhantus fengi</taxonomicName>
<taxonomicName id="EFF5A8D3DB2229BC2E2505A3A60D3858" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus fengi" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="fengi">Rhantus fengi</taxonomicName>
and
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
<taxonomicName id="D7C3B402BFFED4320BCE68D576CBA953" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
and notable geographic localities for orientation. Dingri = Te-ring (type locality of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
<taxonomicName id="7803648738DB9FAA644C8A392083C814" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
); Gangba = Gompa; Jiangzi = Gyangtse (type locality of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<taxonomicName id="8C6AFB766CEAA6848E31BDF46B73FC66" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
).
</paragraph>
</caption>
<paragraph lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67">
<bibRefCitation author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
<paragraph id="560F6615EBF6BFA2BEBF9E27995E5BC3" lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67">
<bibRefCitation id="8C5CA7977763B4350630A096557F9FA4" author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
described &quot;
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus thibetanus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="thibetanus">Laccophilus thibetanus</taxonomicName>
<taxonomicName id="B5F96DEB3E58BDD83E7FA5E2CA309AAE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus thibetanus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="thibetanus">Laccophilus thibetanus</taxonomicName>
sp. n.&quot; from Rikaze, Xizang in her PhD thesis (unpublished, in Chinese) (in the corrected thesis version, the name was
<pageBreakToken pageId="7" pageNumber="68" start="start">changed</pageBreakToken>
<pageBreakToken id="CEADBEA4C2250A05719C0F5CB14789AF" pageId="7" pageNumber="68" start="start">changed</pageBreakToken>
to
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus xizangensis" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="xizangensis">Laccoporus xizangensis</taxonomicName>
<taxonomicName id="5F2AE8F5D7D63FDB7C1928667426DD6B" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus xizangensis" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="xizangensis">Laccoporus xizangensis</taxonomicName>
sp. n. by herself, possibly acknowledging that Xizang is a more precise reference to the type locality than
<normalizedToken originalValue="“Tibet”">&quot;Tibet&quot;</normalizedToken>
<normalizedToken id="48594C74D16F59371A88A4E61E7DF95E" originalValue="“Tibet”">&quot;Tibet&quot;</normalizedToken>
).
<bibRefCitation author="Nilsson, AN" editor="Jaech, MA" journalOrPublisher="Zoologisch-Botanische Gesellschaft in Oesterreich and Wiener Coleopterologenverein, Wien" pageId="7" pageNumber="68" pagination="35 - 96" title="Noteridae and Dytiscidae: Annotated checklist of the Noteridae and Dytiscidae of China (Coleoptera)" volumeTitle="Water beetles of China, Vol. 1" year="1995">Nilsson (1995: 71)</bibRefCitation>
<bibRefCitation id="6BD529BB7C004539F149F06CB0805A44" author="Nilsson, AN" editor="Jaech, MA" journalOrPublisher="Zoologisch-Botanische Gesellschaft in Oesterreich and Wiener Coleopterologenverein, Wien" pageId="7" pageNumber="68" pagination="35 - 96" title="Noteridae and Dytiscidae: Annotated checklist of the Noteridae and Dytiscidae of China (Coleoptera)" volumeTitle="Water beetles of China, Vol. 1" year="1995">Nilsson (1995: 71)</bibRefCitation>
recorded a &quot;
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
<taxonomicName id="DF7EAEA16DBFC03085D29272B6D1AAB2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
sp.&quot; and stated that it is based on &quot;
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus tibetanus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="tibetanus">Laccophilus tibetanus</taxonomicName>
<taxonomicName id="570FE554C5E611BE85B5172ED78BC18F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus tibetanus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="tibetanus">Laccophilus tibetanus</taxonomicName>
Zeng, 1989: 4 (nom. nud., Tibet)&quot;. We found the specimens that have the same locality data as recorded by
<bibRefCitation author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
<bibRefCitation id="732D43CFFF68C6ED1ACC1EC393F4A93A" author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
in SYSU. Although there is no specific name and no type designation attached to these specimens, we are confident that
<bibRefCitation author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
<bibRefCitation id="14CDF6B925F1BD3968B4AF0307BC9C89" author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
based her unpublished
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus xizangensis" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="xizangensis">Laccophilus xizangensis</taxonomicName>
<taxonomicName id="29BB22D41FB57751C336634E20488695" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus xizangensis" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="xizangensis">Laccophilus xizangensis</taxonomicName>
on these specimens. All the specimens are conspecific with
<taxonomicName class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
<taxonomicName id="EE1CA68547715F2984198B0176A84651" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
.
</paragraph>
</subSubSection>

View file

@ -1,285 +1,285 @@
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<mods:title>A new genus and two new species of oonopid spiders from Tibet, China (Araneae, Oonopidae)</mods:title>
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<mods:title id="314786782EB03C0A6BADC07465F8A24E">A new genus and two new species of oonopid spiders from Tibet, China (Araneae, Oonopidae)</mods:title>
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<mods:namePart>Cheng, Weihua</mods:namePart>
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<mods:namePart>Bian, Dongju</mods:namePart>
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<mods:namePart>Tong, Yanfeng</mods:namePart>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paramolotra pome sp. nov., male left palp, SEM A, B, D prolateral, retrolateral, and dorsal views C, E, F distal part of cymbiobulbus, prolateral, retrolateral, and dorsal views. Abbreviations: ap = anterior protrusion; bp = basal protrusion; dp = distal protrusion; pp = posterior protrusion. Scale bars: 0.2 mm (A, B, D); 0.02 mm (C, E, F)." figureDoi="10.3897/zookeys.1052.66402.figure2" httpUri="https://binary.pensoft.net/fig/571978" pageId="0" pageNumber="55">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">, 3</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paramolotra pome sp. nov. A, B, E, F, male holotype (SYNU- 435); Paramolotra metok sp. nov. C, D, G, H male holotype (SYNU- 441) A, C left palp, prolateral view B, D left palp, retrolateral view E, G left chelicerae, anterior view F, H left chelicerae, lateral view. Scale bars: 0.2 mm (A-D); 0.1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66402.figure7" httpUri="https://binary.pensoft.net/fig/571983" pageId="0" pageNumber="55">, 7A, B, E, F</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Paramolotra pome sp. nov. A-F male holotype (SYNU- 435); Paramolotra metok sp. nov. G-L male holotype (SYNU- 441) A, G labium and endite, ventral view B, H left endite, ventral view C, I, E, K, F, L left chelicerae, lateral, posterior, and anterior views D, J left chelicerae, posterior magnified views. Abbreviations: ami = anterior median indentation; csp = cone-shaped protuberance; lt = large tooth; ss = slit sensillum. Scale bars: 0.1 mm (A, C, E-G, I, K, L); 0.04 mm (B, D, H, J)." figureDoi="10.3897/zookeys.1052.66402.figure8" httpUri="https://binary.pensoft.net/fig/571984" pageId="0" pageNumber="55">, 8A-F</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">, 9A, B</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Paramolotra species from Tibet, China 1. P. pome sp. nov. 2. P. metok sp. nov." figureDoi="10.3897/zookeys.1052.66402.figure10" httpUri="https://binary.pensoft.net/fig/571986" pageId="0" pageNumber="55">, 10</figureCitation>
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<figureCitation id="17480B5C2D8E760145E7FAD234274314" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">Figures 1</figureCitation>
<figureCitation id="54ED556E50F18703B91D27ED11B76734" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paramolotra pome sp. nov., male left palp, SEM A, B, D prolateral, retrolateral, and dorsal views C, E, F distal part of cymbiobulbus, prolateral, retrolateral, and dorsal views. Abbreviations: ap = anterior protrusion; bp = basal protrusion; dp = distal protrusion; pp = posterior protrusion. Scale bars: 0.2 mm (A, B, D); 0.02 mm (C, E, F)." figureDoi="10.3897/zookeys.1052.66402.figure2" httpUri="https://binary.pensoft.net/fig/571978" pageId="0" pageNumber="55">, 2</figureCitation>
<figureCitation id="A1741449F2BAD95AE73BAF64BF500B03" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">, 3</figureCitation>
<figureCitation id="B7F7BA894C9D39A13A1A25647A26ABA5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paramolotra pome sp. nov. A, B, E, F, male holotype (SYNU- 435); Paramolotra metok sp. nov. C, D, G, H male holotype (SYNU- 441) A, C left palp, prolateral view B, D left palp, retrolateral view E, G left chelicerae, anterior view F, H left chelicerae, lateral view. Scale bars: 0.2 mm (A-D); 0.1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66402.figure7" httpUri="https://binary.pensoft.net/fig/571983" pageId="0" pageNumber="55">, 7A, B, E, F</figureCitation>
<figureCitation id="DB5AD53654B05068CAD0D8C0BF41DDEF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Paramolotra pome sp. nov. A-F male holotype (SYNU- 435); Paramolotra metok sp. nov. G-L male holotype (SYNU- 441) A, G labium and endite, ventral view B, H left endite, ventral view C, I, E, K, F, L left chelicerae, lateral, posterior, and anterior views D, J left chelicerae, posterior magnified views. Abbreviations: ami = anterior median indentation; csp = cone-shaped protuberance; lt = large tooth; ss = slit sensillum. Scale bars: 0.1 mm (A, C, E-G, I, K, L); 0.04 mm (B, D, H, J)." figureDoi="10.3897/zookeys.1052.66402.figure8" httpUri="https://binary.pensoft.net/fig/571984" pageId="0" pageNumber="55">, 8A-F</figureCitation>
<figureCitation id="25416D596A9479B2DFD6B3638EB8D227" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">, 9A, B</figureCitation>
<figureCitation id="993E46F89B5C5AF142C3F7A65BF8428C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Paramolotra species from Tibet, China 1. P. pome sp. nov. 2. P. metok sp. nov." figureDoi="10.3897/zookeys.1052.66402.figure10" httpUri="https://binary.pensoft.net/fig/571986" pageId="0" pageNumber="55">, 10</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="55">Type materials.</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Holotype</emphasis>
<subSubSection id="66E4573D3349EF539159C6F29025E816" pageId="0" pageNumber="55" type="type material">
<paragraph id="A9B57ADF729E237E95FFC159460FC620" pageId="0" pageNumber="55">Type materials.</paragraph>
<paragraph id="221CCD01E0F6631363F69907B1513B08" pageId="0" pageNumber="55">
<emphasis id="D6E6FE2AE900AB906BE427CA747AC8F1" bold="true" italics="true" pageId="0" pageNumber="55">Holotype</emphasis>
♂: China, Tibet, Nyingchi, Pome County, road to Metok County, 80 K;
<geoCoordinate degrees="29" direction="north" minutes="39.897" orientation="latitude" precision="1" value="29.66495">29°39.897'N</geoCoordinate>
<geoCoordinate id="DDEDAC17BF10AF9631F7F6E15C7814AA" degrees="29" direction="north" minutes="39.897" orientation="latitude" precision="1" value="29.66495">29°39.897'N</geoCoordinate>
,
<geoCoordinate degrees="95" direction="east" minutes="29.963" orientation="longitude" precision="1" value="95.49938">95°29.963'E</geoCoordinate>
<geoCoordinate id="8F8EAF1FD4BD5AE4B21124F1ABA40D23" degrees="95" direction="east" minutes="29.963" orientation="longitude" precision="1" value="95.49938">95°29.963'E</geoCoordinate>
; 2140 m a.s.l.; 10.VIII.2013; Y. Lin leg. (SYNU-435).
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Paratypes</emphasis>
<emphasis id="0323FAB8FB9592A615FA079A174D4B05" bold="true" italics="true" pageId="0" pageNumber="55">Paratypes</emphasis>
2♂3♀: same data as for holotype (SYNU-436-440).
</paragraph>
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<paragraph pageId="0" pageNumber="55">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="55">
<subSubSection id="CE46C2F1D0AA53CD92C71C56E0829E4B" pageId="0" pageNumber="55" type="diagnosis">
<paragraph id="F44FDFACDEAC337E86814ACC38A05FDC" pageId="0" pageNumber="55">Diagnosis.</paragraph>
<paragraph id="427F6DEE9E79DE54F2A4DF55DD5FA7A0" pageId="0" pageNumber="55">
This new species is similar to
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
<taxonomicName id="A06E84323F61EACB052257A982BF6F63" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis id="919EFF4AEA49D0E6E4F9DB87ACAF89E2" italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
</taxonomicName>
sp. nov., but can be distinguished by the narrow anterior protrusion (ap) and broad posterior protrusion (pp) of embolar region (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paramolotra pome sp. nov., male left palp, SEM A, B, D prolateral, retrolateral, and dorsal views C, E, F distal part of cymbiobulbus, prolateral, retrolateral, and dorsal views. Abbreviations: ap = anterior protrusion; bp = basal protrusion; dp = distal protrusion; pp = posterior protrusion. Scale bars: 0.2 mm (A, B, D); 0.02 mm (C, E, F)." figureDoi="10.3897/zookeys.1052.66402.figure2" httpUri="https://binary.pensoft.net/fig/571978" pageId="0" pageNumber="55">2</figureCitation>
<figureCitation id="F7C7BB4EA306898CA71BAC6E32659822" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paramolotra pome sp. nov., male left palp, SEM A, B, D prolateral, retrolateral, and dorsal views C, E, F distal part of cymbiobulbus, prolateral, retrolateral, and dorsal views. Abbreviations: ap = anterior protrusion; bp = basal protrusion; dp = distal protrusion; pp = posterior protrusion. Scale bars: 0.2 mm (A, B, D); 0.02 mm (C, E, F)." figureDoi="10.3897/zookeys.1052.66402.figure2" httpUri="https://binary.pensoft.net/fig/571978" pageId="0" pageNumber="55">2</figureCitation>
), and the long, straight arms of T-shaped sclerite of endogyne (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">9B</figureCitation>
<figureCitation id="D3DB1E5A3BE27F8272D6E0469C82244A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">9B</figureCitation>
).
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
<taxonomicName id="633A8660D384B03FBCD3B8809B5B86BF" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis id="F2918C354D88CB8EE7FA31CC5FAC27ED" italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
</taxonomicName>
sp. nov. males have hook like anterior protrusion (ap) and basally broad, distally finger like posterior protrusion (pp) of embolar region (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paramolotra metok sp. nov., male left palp, SEM A, B, D prolateral, retrolateral, and dorsal views C, E, F distal part of cymbiobulbus, prolateral, retrolateral, and dorsal views. Abbreviations: am = anterior membrane; ap = anterior protrusion; bp = basal protrusion; pm = posterior membrane; pp = posterior protrusion. Scale bars: 0.2 mm (A, B, D); 0.02 mm (C, E, F)." figureDoi="10.3897/zookeys.1052.66402.figure5" httpUri="https://binary.pensoft.net/fig/571981" pageId="0" pageNumber="55">5</figureCitation>
<figureCitation id="0131BFCC39D282AE69A0DAC75D9428A2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paramolotra metok sp. nov., male left palp, SEM A, B, D prolateral, retrolateral, and dorsal views C, E, F distal part of cymbiobulbus, prolateral, retrolateral, and dorsal views. Abbreviations: am = anterior membrane; ap = anterior protrusion; bp = basal protrusion; pm = posterior membrane; pp = posterior protrusion. Scale bars: 0.2 mm (A, B, D); 0.02 mm (C, E, F)." figureDoi="10.3897/zookeys.1052.66402.figure5" httpUri="https://binary.pensoft.net/fig/571981" pageId="0" pageNumber="55">5</figureCitation>
), and females have very short anterior arm of T-shaped sclerite of endogyne (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">9D</figureCitation>
<figureCitation id="739A72E66F4BD1BF8D6CA836951BDDFD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">9D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="description">
<paragraph pageId="0" pageNumber="55">Description.</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
<subSubSection id="A7D3FF34585A92F0E20B90933B0FF828" pageId="0" pageNumber="55" type="description">
<paragraph id="F2858A528574596EA7946A8000F15297" pageId="0" pageNumber="55">Description.</paragraph>
<paragraph id="B375C05073777913F1F42591509FBA23" pageId="0" pageNumber="55">
<emphasis id="20365F92631F2211C8F836F4736016BF" bold="true" pageId="0" pageNumber="55">
Male (holotype).
<emphasis italics="true" pageId="0" pageNumber="55">Body</emphasis>
<emphasis id="E50818AED603CF6C59522B14A82C81CE" italics="true" pageId="0" pageNumber="55">Body</emphasis>
</emphasis>
: carapace brown, abdomen light brown; habitus as in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">1A, C, E</figureCitation>
<figureCitation id="4D9E57128711F323117533A41D55C784" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">1A, C, E</figureCitation>
; body length 2.15.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Carapace</emphasis>
<emphasis id="6BAF9679B2DA8C1C4D4D4CB663EE6712" bold="true" italics="true" pageId="0" pageNumber="55">Carapace</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">1B, H</figureCitation>
<figureCitation id="2B3E6CC2B781FDD944A0D6DF57A69512" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">1B, H</figureCitation>
): 1.04 long, 0.80 wide.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Clypeus</emphasis>
<emphasis id="CC9965B077122EC16984E975C07011BE" bold="true" italics="true" pageId="0" pageNumber="55">Clypeus</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">1I</figureCitation>
<figureCitation id="025FFF0E918FD2056E8E571F7C6EB1CD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">1I</figureCitation>
): ALE separated from edge of carapace by 1.1 times their diameter.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Abdomen</emphasis>
<emphasis id="F6A9792F6B129AC1B431FBF4685A1782" bold="true" italics="true" pageId="0" pageNumber="55">Abdomen</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">1F, G</figureCitation>
<figureCitation id="F7FA0058554524EA48DA9761921E6A13" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Paramolotra pome sp. nov., male holotype (SYNU- 435) A, C, E habitus in dorsal, ventral, and lateral views B, D, H, I prosoma in dorsal, ventral, lateral, and anterior views F, G abdomen in lateral and ventral views. Scale bars: 0.8 mm (A, C, E); 0.4 mm (B, D, F-I)." figureDoi="10.3897/zookeys.1052.66402.figure1" httpUri="https://binary.pensoft.net/fig/571977" pageId="0" pageNumber="55">1F, G</figureCitation>
): 1.06 long, 0.82 wide.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Palp</emphasis>
<emphasis id="2B003D9FF67D8E3AFAC14725518C2A8E" bold="true" italics="true" pageId="0" pageNumber="55">Palp</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paramolotra pome sp. nov., male left palp, SEM A, B, D prolateral, retrolateral, and dorsal views C, E, F distal part of cymbiobulbus, prolateral, retrolateral, and dorsal views. Abbreviations: ap = anterior protrusion; bp = basal protrusion; dp = distal protrusion; pp = posterior protrusion. Scale bars: 0.2 mm (A, B, D); 0.02 mm (C, E, F)." figureDoi="10.3897/zookeys.1052.66402.figure2" httpUri="https://binary.pensoft.net/fig/571978" pageId="0" pageNumber="55">2</figureCitation>
<figureCitation id="5A227E8B0BDA661985E36787BFB1018F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Paramolotra pome sp. nov., male left palp, SEM A, B, D prolateral, retrolateral, and dorsal views C, E, F distal part of cymbiobulbus, prolateral, retrolateral, and dorsal views. Abbreviations: ap = anterior protrusion; bp = basal protrusion; dp = distal protrusion; pp = posterior protrusion. Scale bars: 0.2 mm (A, B, D); 0.02 mm (C, E, F)." figureDoi="10.3897/zookeys.1052.66402.figure2" httpUri="https://binary.pensoft.net/fig/571978" pageId="0" pageNumber="55">2</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paramolotra pome sp. nov. A, B, E, F, male holotype (SYNU- 435); Paramolotra metok sp. nov. C, D, G, H male holotype (SYNU- 441) A, C left palp, prolateral view B, D left palp, retrolateral view E, G left chelicerae, anterior view F, H left chelicerae, lateral view. Scale bars: 0.2 mm (A-D); 0.1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66402.figure7" httpUri="https://binary.pensoft.net/fig/571983" pageId="0" pageNumber="55">7A, B</figureCitation>
<figureCitation id="1FF3B91E316C34D2BE1EA6E00771DCA5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paramolotra pome sp. nov. A, B, E, F, male holotype (SYNU- 435); Paramolotra metok sp. nov. C, D, G, H male holotype (SYNU- 441) A, C left palp, prolateral view B, D left palp, retrolateral view E, G left chelicerae, anterior view F, H left chelicerae, lateral view. Scale bars: 0.2 mm (A-D); 0.1 mm (E-H)." figureDoi="10.3897/zookeys.1052.66402.figure7" httpUri="https://binary.pensoft.net/fig/571983" pageId="0" pageNumber="55">7A, B</figureCitation>
): femur 0.25 long, patella 0.18 long, tibia 0.12 long, cymbiobulbus 0.57 long, 0.33 wide, length/maximal width = 1.58; embolar region with a broad basal protrusion (bp), a narrow anterior protrusion (ap), a broad posterior protrusion (pp) and a distal dorsal protrusion (dp).
</paragraph>
<caption doi="10.3897/zookeys.1052.66402.figure7" httpUri="https://binary.pensoft.net/fig/571983" pageId="0" pageNumber="55" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figure 7.</emphasis>
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra pome" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="pome">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra pome</emphasis>
<caption id="CBF213E8C0E2B4005734ACCE8DDA6C2D" doi="10.3897/zookeys.1052.66402.figure7" httpUri="https://binary.pensoft.net/fig/571983" pageId="0" pageNumber="55" start="Figure 7" startId="F7">
<paragraph id="BDE827EC0C9BB69A385BD4B6A8A3C52E" pageId="0" pageNumber="55">
<emphasis id="5CF7BDA9D9F3D5DB99F83C9A13A6EE1E" bold="true" pageId="0" pageNumber="55">Figure 7.</emphasis>
<taxonomicName id="7115B0AB0DF8F2C61E57B0A8E1364D63" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra pome" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="pome">
<emphasis id="5718E13ADF97E156442B9F7E482C969C" italics="true" pageId="0" pageNumber="55">Paramolotra pome</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="55">A, B, E, F</emphasis>
<emphasis id="B32C83C2AE8B1A559EA6D125666B02B4" bold="true" pageId="0" pageNumber="55">A, B, E, F</emphasis>
, male holotype (SYNU-435);
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
<taxonomicName id="4F5866247C724BC37D72E5D68E270404" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis id="B0859C1433C361CE06AC3FEC5184087A" italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="55">C, D, G, H</emphasis>
<emphasis id="45A03769D80D45C61ADE01BB82664094" bold="true" pageId="0" pageNumber="55">C, D, G, H</emphasis>
male holotype (SYNU-441)
<emphasis bold="true" pageId="0" pageNumber="55">A, C</emphasis>
<emphasis id="967B625685A686E3A90EE1E4FB31CC2F" bold="true" pageId="0" pageNumber="55">A, C</emphasis>
left palp, prolateral view
<emphasis bold="true" pageId="0" pageNumber="55">B, D</emphasis>
<emphasis id="F1C2B4D2940B080E97FBB5D6341A324A" bold="true" pageId="0" pageNumber="55">B, D</emphasis>
left palp, retrolateral view
<emphasis bold="true" pageId="0" pageNumber="55">E, G</emphasis>
<emphasis id="04C4D45DED6F3A61D8E1DC298A047CF1" bold="true" pageId="0" pageNumber="55">E, G</emphasis>
left chelicerae, anterior view
<emphasis bold="true" pageId="0" pageNumber="55">F, H</emphasis>
<emphasis id="9FB19D7FA08A8463719552BC0435F4DF" bold="true" pageId="0" pageNumber="55">F, H</emphasis>
left chelicerae, lateral view. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="55">A-D</emphasis>
<emphasis id="EF5F27E0E647D04CB292362E8C694A3A" bold="true" pageId="0" pageNumber="55">A-D</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="55">E-H</emphasis>
<emphasis id="B92E40694731C73837115F02B2954B9E" bold="true" pageId="0" pageNumber="55">E-H</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1052.66402.figure8" httpUri="https://binary.pensoft.net/fig/571984" pageId="0" pageNumber="55" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figure 8.</emphasis>
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra pome" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="pome">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra pome</emphasis>
<caption id="3D54084D32557C935D280D315CA92ED1" doi="10.3897/zookeys.1052.66402.figure8" httpUri="https://binary.pensoft.net/fig/571984" pageId="0" pageNumber="55" start="Figure 8" startId="F8">
<paragraph id="16F86573FCF10D59DA4C9F125A996736" pageId="0" pageNumber="55">
<emphasis id="2A7D36B4D29BEC01CFF8E038D32A7D43" bold="true" pageId="0" pageNumber="55">Figure 8.</emphasis>
<taxonomicName id="72E1488BE06A4259B11D84EFCF6C60CE" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra pome" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="pome">
<emphasis id="B8B2D5E82F1A8C60F40795D98B41A86F" italics="true" pageId="0" pageNumber="55">Paramolotra pome</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="55">A-F</emphasis>
<emphasis id="16CD7A46D7B545BE5C4107C343A1A959" bold="true" pageId="0" pageNumber="55">A-F</emphasis>
male holotype (SYNU-435);
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
<taxonomicName id="EA09B3C038C5C826952E6780B5900F40" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis id="E1978A4F44B2CE88F4466BDD3268C9FC" italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="55">G-L</emphasis>
<emphasis id="F4A642CC13C7AAFA30256C932EEC636C" bold="true" pageId="0" pageNumber="55">G-L</emphasis>
male holotype (SYNU-441)
<emphasis bold="true" pageId="0" pageNumber="55">A, G</emphasis>
<emphasis id="C5F197C4C46F5FF321F1E75B47470117" bold="true" pageId="0" pageNumber="55">A, G</emphasis>
labium and endite, ventral view
<emphasis bold="true" pageId="0" pageNumber="55">B, H</emphasis>
<emphasis id="391936F1B9F4508C48630B5FAD338D03" bold="true" pageId="0" pageNumber="55">B, H</emphasis>
left endite, ventral view
<emphasis bold="true" pageId="0" pageNumber="55">C, I, E, K, F, L</emphasis>
<emphasis id="623A8DBA3ED7149426AD988852265EC4" bold="true" pageId="0" pageNumber="55">C, I, E, K, F, L</emphasis>
left chelicerae, lateral, posterior, and anterior views
<emphasis bold="true" pageId="0" pageNumber="55">D, J</emphasis>
<emphasis id="BF415BBC5E4A0D05C0D77B4491717238" bold="true" pageId="0" pageNumber="55">D, J</emphasis>
left chelicerae, posterior magnified views. Abbreviations: ami = anterior median indentation; csp = cone-shaped protuberance; lt = large tooth; ss = slit sensillum. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="55">A, C, E-G, I, K, L</emphasis>
<emphasis id="38FCC5556DF2385BB853DC015CEE3CBC" bold="true" pageId="0" pageNumber="55">A, C, E-G, I, K, L</emphasis>
); 0.04 mm (
<emphasis bold="true" pageId="0" pageNumber="55">B, D, H, J</emphasis>
<emphasis id="35F310B04510278CC7B68A9DF74A5137" bold="true" pageId="0" pageNumber="55">B, D, H, J</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Female (paratype, SYNU-438).</emphasis>
<paragraph id="3DDCEB1E0C1AEF6A1570764C951E5280" pageId="0" pageNumber="55">
<emphasis id="107243CD2E06D39C3D88C58F802868D9" bold="true" pageId="0" pageNumber="55">Female (paratype, SYNU-438).</emphasis>
As in male except as noted.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Body</emphasis>
<emphasis id="BF0EA01A1AC6317522B72D2DA25D8C70" bold="true" italics="true" pageId="0" pageNumber="55">Body</emphasis>
: habitus as in Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">3A, B, E</figureCitation>
<figureCitation id="9197F911990CE6626FC08F4887312543" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">3A, B, E</figureCitation>
; body length 2.24.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Carapace</emphasis>
<emphasis id="AE4C0EEFC16DA9AAF4C64D66AC9E9526" bold="true" italics="true" pageId="0" pageNumber="55">Carapace</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">3C, F, H</figureCitation>
<figureCitation id="3F47EFEAA7D03E4668C90CC2D3CBF33D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">3C, F, H</figureCitation>
): 1.06 long, 0.82 wide.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Abdomen</emphasis>
<emphasis id="DBE8F0250D07A83E2A337BF770FFEB07" bold="true" italics="true" pageId="0" pageNumber="55">Abdomen</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">3G</figureCitation>
<figureCitation id="754C1728C5AD71E6BB3D0D1DF52E466E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">3G</figureCitation>
): 1.24 long, 0.97 wide.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Epigastric area</emphasis>
<emphasis id="9BABF6ECF012A39046A387445C33FD6A" bold="true" italics="true" pageId="0" pageNumber="55">Epigastric area</emphasis>
(Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">3G</figureCitation>
<figureCitation id="25A895247865ECCC42F98F9C369345CF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Paramolotra pome sp. nov., female paratype (SYNU- 438) A, B, E habitus in dorsal, ventral, and lateral views C, D, F, H prosoma in dorsal, ventral, lateral, and anterior views G abdomen in ventral view. Scale bars: 0.8 mm (A, B, E); 0.4 mm (C, D, F-H)." figureDoi="10.3897/zookeys.1052.66402.figure3" httpUri="https://binary.pensoft.net/fig/571979" pageId="0" pageNumber="55">3G</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">9A</figureCitation>
<figureCitation id="3102B8AD216FB400BE06D2EED3232852" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">9A</figureCitation>
): genital atrium relatively wide, broadly oval.
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Endogyne</emphasis>
<emphasis id="25B3FAC4ABFFFC269C7A91A915F96DEB" bold="true" italics="true" pageId="0" pageNumber="55">Endogyne</emphasis>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">9B</figureCitation>
<figureCitation id="8144CAF4F5BDC6B658D63BBB0CA81274" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Paramolotra pome sp. nov. A, B female paratype (SYNU- 438); Paramolotra metok sp. nov. C, D female paratype (SYNU- 443) A, C copulatory organ, ventral view B, D copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55">9B</figureCitation>
): with a large T-shaped sclerite; the anterior arm of the T-shaped sclerite long and strong.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="etymology">
<paragraph pageId="0" pageNumber="55">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="55">The specific name is a noun in apposition taken from the type locality.</paragraph>
<subSubSection id="9AB911F13C5EDC8A9C42D0B486CA2CEB" pageId="0" pageNumber="55" type="etymology">
<paragraph id="07AFFCBBCDEAFF8A862A80AC2736939C" pageId="0" pageNumber="55">Etymology.</paragraph>
<paragraph id="868A66B1E40D099226791A2D3DD07A4F" pageId="0" pageNumber="55">The specific name is a noun in apposition taken from the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="distribution">
<paragraph pageId="0" pageNumber="55">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="55">
<subSubSection id="AB2314E04052C17816B5E9BB43ED244B" pageId="0" pageNumber="55" type="distribution">
<paragraph id="4DC4C4C6903E76CD7AA29DBC432B5FED" pageId="0" pageNumber="55">Distribution.</paragraph>
<paragraph id="41B7E18663FB73AB40C15157D21A6D7C" pageId="0" pageNumber="55">
Known only from the type locality (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Paramolotra species from Tibet, China 1. P. pome sp. nov. 2. P. metok sp. nov." figureDoi="10.3897/zookeys.1052.66402.figure10" httpUri="https://binary.pensoft.net/fig/571986" pageId="0" pageNumber="55">10</figureCitation>
<figureCitation id="D80A4B6018A75E608073DCA2D5E0EEDA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Paramolotra species from Tibet, China 1. P. pome sp. nov. 2. P. metok sp. nov." figureDoi="10.3897/zookeys.1052.66402.figure10" httpUri="https://binary.pensoft.net/fig/571986" pageId="0" pageNumber="55">10</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figure 9.</emphasis>
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra pome" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="pome">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra pome</emphasis>
<caption id="FEAA6D0AA18F9EEBBBCD2ABEDB640AD2" doi="10.3897/zookeys.1052.66402.figure9" httpUri="https://binary.pensoft.net/fig/571985" pageId="0" pageNumber="55" start="Figure 9" startId="F9">
<paragraph id="D7758C7275854619FBDC2BE43A38F2A2" pageId="0" pageNumber="55">
<emphasis id="DA651912852BE2BF7451C36FF008A426" bold="true" pageId="0" pageNumber="55">Figure 9.</emphasis>
<taxonomicName id="A9A5A32EE96A30B58DE10B536F844CFE" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra pome" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="pome">
<emphasis id="B509EF6BA20AE618E0AE8D9E5AFC80A0" italics="true" pageId="0" pageNumber="55">Paramolotra pome</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="55">A, B</emphasis>
<emphasis id="B96C7637AE2AD0B102E7D0D24BBAF09F" bold="true" pageId="0" pageNumber="55">A, B</emphasis>
female paratype (SYNU-438);
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
<taxonomicName id="0BF50DD40BB49C10776C4C8C4090AF68" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra metok" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="metok">
<emphasis id="76813DDF414FE5D34BF64DD9869A2C18" italics="true" pageId="0" pageNumber="55">Paramolotra metok</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="55">C, D</emphasis>
<emphasis id="87753CE0F73A1CC3B343F8F03EAA87D7" bold="true" pageId="0" pageNumber="55">C, D</emphasis>
female paratype (SYNU-443)
<emphasis bold="true" pageId="0" pageNumber="55">A, C</emphasis>
<emphasis id="9587BB316E0484FE512DB0473A2044F1" bold="true" pageId="0" pageNumber="55">A, C</emphasis>
copulatory organ, ventral view
<emphasis bold="true" pageId="0" pageNumber="55">B, D</emphasis>
<emphasis id="9A1DE75582DBAEC7A5209F97AC00FE0C" bold="true" pageId="0" pageNumber="55">B, D</emphasis>
copulatory organ, dorsal view. Abbreviations: apo = apodemes; pr = posterior receptaculum; tls = tube-like structure; tsc = T-shaped sclerite. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1052.66402.figure10" httpUri="https://binary.pensoft.net/fig/571986" pageId="0" pageNumber="55" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figure 10.</emphasis>
<caption id="DEB5F167C66DDEF7D431919BE5F6B248" doi="10.3897/zookeys.1052.66402.figure10" httpUri="https://binary.pensoft.net/fig/571986" pageId="0" pageNumber="55" start="Figure 10" startId="F10">
<paragraph id="E763E5096BBDE99746BEB04739363C12" pageId="0" pageNumber="55">
<emphasis id="873663C00D41EFC5F6773B3099856584" bold="true" pageId="0" pageNumber="55">Figure 10.</emphasis>
Distribution records of
<taxonomicName authorityName="Tong &amp; Li" authorityYear="2021" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Paramolotra</emphasis>
<taxonomicName id="D111865E1A252A57EA7D82897400247A" authorityName="Tong &amp; Li" authorityYear="2021" class="Arachnida" family="Oonopidae" genus="Paramolotra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramolotra" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="515C721EC62F3A0F7970402831F5B1C1" italics="true" pageId="0" pageNumber="55">Paramolotra</emphasis>
</taxonomicName>
species from Tibet, China
<emphasis bold="true" pageId="0" pageNumber="55">1</emphasis>
<emphasis id="8951F4B5B3F138C6625B277A4A8BF5DC" bold="true" pageId="0" pageNumber="55">1</emphasis>
.
<taxonomicName lsidName="P. pome" pageId="0" pageNumber="55" rank="species" species="pome">
<emphasis italics="true" pageId="0" pageNumber="55">P. pome</emphasis>
<taxonomicName id="75396024ECBADE1031A1D35AB24DEF7B" lsidName="P. pome" pageId="0" pageNumber="55" rank="species" species="pome">
<emphasis id="57D8E627293ECE47921A746EB1C6F714" italics="true" pageId="0" pageNumber="55">P. pome</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="55">2</emphasis>
<emphasis id="11B1923C6127122EDF8A0F3F21EBFF49" bold="true" pageId="0" pageNumber="55">2</emphasis>
.
<taxonomicName lsidName="P. metok" pageId="0" pageNumber="55" rank="species" species="metok">
<emphasis italics="true" pageId="0" pageNumber="55">P. metok</emphasis>
<taxonomicName id="A861A41D516455F2F1A682CF8ACF8908" lsidName="P. metok" pageId="0" pageNumber="55" rank="species" species="metok">
<emphasis id="30BCB183A90BA8DB79F7B60306DEA867" italics="true" pageId="0" pageNumber="55">P. metok</emphasis>
</taxonomicName>
sp. nov.
</paragraph>

View file

@ -1,242 +1,242 @@
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<mods:title id="A14A04E55B47E925939B916EABC658A5">ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-03</mods:number>
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<mods:date id="F46110EC4BAB3C5D5AD7EAD808FFFB2D">2022</mods:date>
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<mods:number id="E82E4DA0CDBEAC47AF28B62ECBBC19A9">2022-06-03</mods:number>
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<mods:detail type="volume">
<mods:number>1103</mods:number>
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<mods:number id="BB124B845E2225D7966FE7A627308543">1103</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1103.83952</mods:url>
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<mods:url id="79A4DD6CC50D79D6C752E5217E4AE483">http://dx.doi.org/10.3897/zookeys.1103.83952</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1103.83952</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1103-189</mods:identifier>
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<mods:identifier id="616808F00E987D55EAF822E3BE264EDA" type="DOI">http://dx.doi.org/10.3897/zookeys.1103.83952</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:5C8F64C1-7008-4356-8CAC-A775D3F05D12" httpUri="http://treatment.plazi.org/id/58E3D93AEB8A56F38AC18292CCD29472" lastPageNumber="189" pageId="0" pageNumber="189">
<subSubSection pageId="0" pageNumber="189" type="nomenclature">
<paragraph pageId="0" pageNumber="189">
<taxonomicName LSID="http://zoobank.org/5C8F64C1-7008-4356-8CAC-A775D3F05D12" authority="Huemer, 2022" authorityName="Huemer" authorityYear="2022" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum herwigvanstaai" order="Lepidoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="herwigvanstaai" status="sp. nov.">Caryocolum herwigvanstaai</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="189">sp. nov.</taxonomicNameLabel>
<treatment id="58E3D93AEB8A56F38AC18292CCD29472" ID-GBIF-Taxon="195738904" LSID="urn:lsid:zoobank.org:act:5C8F64C1-7008-4356-8CAC-A775D3F05D12" httpUri="http://treatment.plazi.org/id/58E3D93AEB8A56F38AC18292CCD29472" lastPageNumber="189" pageId="0" pageNumber="189" scope_family="Gelechiidae" scope_order="Lepidoptera">
<subSubSection id="E66507CCABEB8FD476AA07AC45B851FD" pageId="0" pageNumber="189" type="nomenclature">
<paragraph id="2315EF31203363F1F84D882DAFBE32D4" pageId="0" pageNumber="189">
<taxonomicName id="C95BCBF78E490FFB8AF096BDA2AC6E09" ID-CoL="999BC" LSID="http://zoobank.org/5C8F64C1-7008-4356-8CAC-A775D3F05D12" authority="Huemer, 2022" authorityName="Huemer" authorityYear="2022" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum herwigvanstaai" order="Lepidoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="herwigvanstaai" status="sp. nov.">Caryocolum herwigvanstaai</taxonomicName>
<taxonomicNameLabel id="3F2C04944C83491AFFFB0F2FD51F5A5E" pageId="0" pageNumber="189">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="type material">
<paragraph pageId="0" pageNumber="189">Type material.</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">
<emphasis italics="true" pageId="0" pageNumber="189">Holotype</emphasis>
<subSubSection id="722C5C86C59AC76A1537A39CFDFA85C9" pageId="0" pageNumber="189" type="type material">
<paragraph id="37F7F1F21167449B0B2EB3FBC458CE50" pageId="0" pageNumber="189">Type material.</paragraph>
<paragraph id="D6C0960D81330E71782AE3A424778180" pageId="0" pageNumber="189">
<emphasis id="550BAF2590AE8C02AC5D50130CB98250" bold="true" pageId="0" pageNumber="189">
<emphasis id="C8C79D1203DEBA64B7FEAAC1898F2655" italics="true" pageId="0" pageNumber="189">Holotype</emphasis>
.
</emphasis>
[Italy] • ♂;
<normalizedToken originalValue="LAquila">L'Aquila</normalizedToken>
<normalizedToken id="3893CC2BC4077FF7AFC84548B4890524" originalValue="LAquila">L'Aquila</normalizedToken>
, NP Gran Sasso, ex Miniera di Lignite; 1750 m; 14-15 Jul 2010; [genitalia slide number] GEL 1153♂, P. Huemer; P. Huemer leg; TLMF.
</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">
<emphasis italics="true" pageId="0" pageNumber="189">Paratypes</emphasis>
<paragraph id="5B8804122A3CE7607F930552724460E5" pageId="0" pageNumber="189">
<emphasis id="124AC3EAFB374E701A8143264158A26F" bold="true" pageId="0" pageNumber="189">
<emphasis id="FC826D987742C282613EB398AE4F7CC0" italics="true" pageId="0" pageNumber="189">Paratypes</emphasis>
.
</emphasis>
[Italy] • 5 ♂, 5 ♀; same collection data as for holotype; [genitalia slide number] GEL 1155♀, P. Huemer; [1 ♂, 1 ♀ genitalia in glycerin capsule]; [DNA barcode ids] BC TLMF Lep 01600; all TLMF; • 10 ♂, 3 ♀; same collection data as for holotype; 1750 m; 15 Jul 2010; T. Mayr leg.; RCTM; • 1 ♂; same collection data as for holotype; 1750 m; 14 Jul 2010; T. Mayr leg.; RCTM; • 1 ♀; Rieti, Monte Terminillo; 1730-1780 m; 11 Jul 2010; P. Huemer leg.; [DNA barcode ids] BC TLMF Lep 01601; • 6 ♂; Rieti, Monte Terminillo; 1700 m; 17 Jul 2011; T. Mayr leg.; RCTM; • 1 ♀; Chieti, PN della Majella, Taranta Peligna, Pian di Valle; 770 m; 20 Jul 2011; P. Huemer leg.; BC TLMF Lep 05038; all TLMF.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="diagnosis">
<paragraph pageId="0" pageNumber="189">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="189">
<taxonomicName authorityName="Huemer" authorityYear="2022" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum herwigvanstaai" order="Lepidoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="herwigvanstaai">
<emphasis italics="true" pageId="0" pageNumber="189">Caryocolum herwigvanstaai</emphasis>
<subSubSection id="6B745DF118129D5121B7B092B226CF09" pageId="0" pageNumber="189" type="diagnosis">
<paragraph id="38053FED21C3C347E9CC3D2ADAD3F6CB" pageId="0" pageNumber="189">Diagnosis.</paragraph>
<paragraph id="EA849D8855F9A929B8F9887535784512" pageId="0" pageNumber="189">
<taxonomicName id="8F4E280F5A5952EA22E55C5817E317B2" authorityName="Huemer" authorityYear="2022" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum herwigvanstaai" order="Lepidoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="herwigvanstaai">
<emphasis id="CD2D5276E146EF263389E8D7A23CC198" italics="true" pageId="0" pageNumber="189">Caryocolum herwigvanstaai</emphasis>
</taxonomicName>
differs from
<taxonomicName genus="C." lsidName="C. tricolorella" pageId="0" pageNumber="189" rank="species" species="tricolorella">
<emphasis italics="true" pageId="0" pageNumber="189">C. tricolorella</emphasis>
<taxonomicName id="877BECA366C1F0392EF8B10FEBB8B2B5" lsidName="C. tricolorella" pageId="0" pageNumber="189" rank="species" species="tricolorella">
<emphasis id="606B778B99C54EF8D0EADDCB63951673" italics="true" pageId="0" pageNumber="189">C. tricolorella</emphasis>
</taxonomicName>
by its distinctly smaller size and the less extensive ochreous-orange markings, and from
<taxonomicName genus="C." lsidName="C. fibigerium" pageId="0" pageNumber="189" rank="species" species="fibigerium">
<emphasis italics="true" pageId="0" pageNumber="189">C. fibigerium</emphasis>
<taxonomicName id="716575E2050FE99E5227973549D3282B" lsidName="C. fibigerium" pageId="0" pageNumber="189" rank="species" species="fibigerium">
<emphasis id="D859125F7656303C3C6168E95F5BCF85" italics="true" pageId="0" pageNumber="189">C. fibigerium</emphasis>
</taxonomicName>
by the extended white forewing markings which are, however, less pronounced at the inner margin compared to
<taxonomicName genus="C." lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
<taxonomicName id="85F718AEBD8CE8D01C49B7A21107CA06" lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis id="6BD0A9A284A75233DFE523B1309F297A" italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
</taxonomicName>
. The male genitalia differ from
<taxonomicName genus="C." lsidName="C. tricolorella" pageId="0" pageNumber="189" rank="species" species="tricolorella">
<emphasis italics="true" pageId="0" pageNumber="189">C. tricolorella</emphasis>
<taxonomicName id="8FD6274C9ACACBDB00B12757B7366134" lsidName="C. tricolorella" pageId="0" pageNumber="189" rank="species" species="tricolorella">
<emphasis id="775AEBA1C5074DDF72F6F64039AF3C56" italics="true" pageId="0" pageNumber="189">C. tricolorella</emphasis>
</taxonomicName>
by the shorter valva and sacculus and the additional, although moderately low, humps of the posterior margin of the vinculum. From
<emphasis italics="true" pageId="0" pageNumber="189">
<taxonomicName genus="C." lsidName="C. fibigerium" pageId="0" pageNumber="189" rank="species" species="fibigerium">C. fibigerium</taxonomicName>
<taxonomicName genus="C." lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">C. herwigvanstaai</taxonomicName>
<emphasis id="932DA5B7F1F187C6363BD3DF1A3F075B" italics="true" pageId="0" pageNumber="189">
<taxonomicName id="0698A51198714D29AB03644E9AB913BE" lsidName="C. fibigerium" pageId="0" pageNumber="189" rank="species" species="fibigerium">C. fibigerium</taxonomicName>
<taxonomicName id="2E275CBFD4932AAD041A289B829315E8" lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">C. herwigvanstaai</taxonomicName>
</emphasis>
differs in particular by the more slender sacculus and the distinct lateral humps of the posterior margin of the vinculum, and from
<taxonomicName genus="C." lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
<taxonomicName id="2DC60687A65CAB63B65577D0E3A93D5E" lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis id="2864C1349DAF25C1435476AACEE2F449" italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
</taxonomicName>
by the apically slightly dilated valva and the slender sacculus. The antrum of the female genitalia is much larger than in
<taxonomicName genus="C." lsidName="C. tricolorella" pageId="0" pageNumber="189" rank="species" species="tricolorella">
<emphasis italics="true" pageId="0" pageNumber="189">C. tricolorella</emphasis>
<taxonomicName id="13256441FF83557D60A0ABC17C63F3FE" lsidName="C. tricolorella" pageId="0" pageNumber="189" rank="species" species="tricolorella">
<emphasis id="712492555EEFA2A61C149C3B316DE3CB" italics="true" pageId="0" pageNumber="189">C. tricolorella</emphasis>
</taxonomicName>
but smaller than in
<taxonomicName genus="C." lsidName="C. fibigerium" pageId="0" pageNumber="189" rank="species" species="fibigerium">
<emphasis italics="true" pageId="0" pageNumber="189">C. fibigerium</emphasis>
<taxonomicName id="24F17DF5D291611BE040962743B4C0CE" lsidName="C. fibigerium" pageId="0" pageNumber="189" rank="species" species="fibigerium">
<emphasis id="C7F97A90ECB2D68666990DA380C6D75F" italics="true" pageId="0" pageNumber="189">C. fibigerium</emphasis>
</taxonomicName>
, not extending the length of apophysis anterior. The anterior margin of the antrum is concave in
<taxonomicName genus="C." lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">
<emphasis italics="true" pageId="0" pageNumber="189">C. herwigvanstaai</emphasis>
<taxonomicName id="1E67BEF3F4EE11A7409F21C22FC3447A" lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">
<emphasis id="7EBDA39B3814B0CAC621C0BE4777A343" italics="true" pageId="0" pageNumber="189">C. herwigvanstaai</emphasis>
</taxonomicName>
but convex in
<taxonomicName genus="C." lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
<taxonomicName id="E34E3347B4675D9C74EAB4F05A1D5B37" lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis id="8760467110FE9E1FBC4A42C6AAA85D70" italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="description">
<paragraph pageId="0" pageNumber="189">Description.</paragraph>
<paragraph pageId="0" pageNumber="189">
<subSubSection id="AF9B3586E9712AFBA67473A4D80D6634" pageId="0" pageNumber="189" type="description">
<paragraph id="B8ED6627F9BC9F41E6AA453780D7D7A3" pageId="0" pageNumber="189">Description.</paragraph>
<paragraph id="FAA17ECEC70641AEEE6A95974A6FDDB9" pageId="0" pageNumber="189">
Adult (Fig.
<figureCitation captionStart="Figures 25" captionStartId="F2" captionText="Figures 2 - 5. Adults 2 Caryocolum tricolorella, male, Germany 3 C. fibigerium, male, paratype, Spain 4 C. herwigvanstaai sp. nov., male, holotype, Italy 5 C. olekarsholti, male, holotype, Greece." figureDoi="10.3897/zookeys.1103.83952.figures2-5" httpUri="https://binary.pensoft.net/fig/691658" pageId="0" pageNumber="189">4</figureCitation>
<figureCitation id="BCA84EF3D9D0C250FFA4FF34ECD302FF" captionStart="Figures 25" captionStartId="F2" captionText="Figures 2 - 5. Adults 2 Caryocolum tricolorella, male, Germany 3 C. fibigerium, male, paratype, Spain 4 C. herwigvanstaai sp. nov., male, holotype, Italy 5 C. olekarsholti, male, holotype, Greece." figureDoi="10.3897/zookeys.1103.83952.figures2-5" httpUri="https://binary.pensoft.net/fig/691658" pageId="0" pageNumber="189">4</figureCitation>
). Forewing length. ♂ 4.9-5.5 mm (
<normalizedToken originalValue="ø">o</normalizedToken>
<normalizedToken id="11F44FAE5E23A3B8783BE697EC77FB98" originalValue="ø">o</normalizedToken>
= 5.25 mm,
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
<emphasis id="41AF67F605CD0F1B70135244B0CA69DE" italics="true" pageId="0" pageNumber="189">n</emphasis>
= 4), ♀ 5.1-5.7 mm (
<normalizedToken originalValue="ø">o</normalizedToken>
<normalizedToken id="0A8896533EB4D7D6A6693D75FD4C1EC5" originalValue="ø">o</normalizedToken>
= 5.40 mm,
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
<emphasis id="840A61697825CE7359609BDF64C156A9" italics="true" pageId="0" pageNumber="189">n</emphasis>
= 4). Head with fuscous vertex, frons cream-white; second segment of labial palpus cream-white on inner and upper surface, predominantly grey-brown on outer surface, third segment dark brown with a few white scales particularly at apex; antenna black, weakly ringed whitish. Thorax and tegula dark brown with a few intermixed ochreous scales. Abdomen dorsally grey, ventrally whitish, pale grey at margins. Forewing predominantly fuscous in costal and terminal area, dorsum mixed ochreous-whitish with scattered fuscous scales, extensive white mottling from dorsum to costa at 1/5 and 1/2, large white costal and tornal spots nearly fused, separated by few fuscous scales, irregularly shaped black patch from fold to costa at about 1/3 interrupted by ochreous scales, black plical and discal spot; cilia light grey with fuscous ciliary line, buff-whitish beyond line. Hindwing light grey, cilia greyish buff.
</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Variation</emphasis>
<paragraph id="4C93AA3B361FA0BAAC27A84E007A69CA" pageId="0" pageNumber="189">
<emphasis id="2FB4B77FD3D3B5D97BC8EA6131FCA557" bold="true" pageId="0" pageNumber="189">Variation</emphasis>
: the extent of ochreous scales, particularly along the dorsum, is slightly variable.
</paragraph>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Male genitalia</emphasis>
<paragraph id="8A1F05BA9A1B22087A1059E43164DB56" pageId="0" pageNumber="189">
<emphasis id="8BE34DF169E3C35FA31B4B7F8487AC60" bold="true" italics="true" pageId="0" pageNumber="189">Male genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 8, 9" captionStartId="F4" captionText="Figures 8, 9. Male genitalia 8 C. herwigvanstaai sp. nov., holotype, Italy, slide GEL 1153 P. Huemer 9 C. olekarsholti, paratype, Greece, slide GEL 1213 P. Huemer;" figureDoi="10.3897/zookeys.1103.83952.figures8-9" httpUri="https://binary.pensoft.net/fig/691660" pageId="0" pageNumber="189">8</figureCitation>
<figureCitation id="32D0AF14F7E5DA27985D8DF462F8FE7D" captionStart="Figures 8, 9" captionStartId="F4" captionText="Figures 8, 9. Male genitalia 8 C. herwigvanstaai sp. nov., holotype, Italy, slide GEL 1153 P. Huemer 9 C. olekarsholti, paratype, Greece, slide GEL 1213 P. Huemer;" figureDoi="10.3897/zookeys.1103.83952.figures8-9" httpUri="https://binary.pensoft.net/fig/691660" pageId="0" pageNumber="189">8</figureCitation>
). Uncus long, suboval, posterior edges rounded; gnathos with large mesial sclerite, culcitula small; posterior 1/3 of tegumen slender, anterior part strongly widened towards broadly rounded pedunculi of about twice size of uncus, anterior margin with deep concave emargination; transtilla membranous with few microtrichia; valva basally curved ventrad, moderately short, slender, apical part slightly dilated, obliquely pointed apex with group of stiff setae; sacculus moderately long, more slender and shorter than valva, apex rounded, with dorsally pointed projection; vinculum wide and short, posterior margin moderately sclerotized, with shallow medial incision and distinctly rounded lateromedial projections, lateral projections distinct, anterior margin with strongly sclerotized concave ridge; saccus slender, basally weakly widened, gradually narrowing towards pointed apex, slightly exceeding length of apex of valva to anterior margin of vinculum; anellus with pair of needle-shaped sclerites; phallus stout, distal part weakly curved and contorted, coecum weakly inflated, longitudinal ridge from about middle to apex, two small sclerotized hooklets at apex.
</paragraph>
<caption doi="10.3897/zookeys.1103.83952.figures8-9" httpUri="https://binary.pensoft.net/fig/691660" pageId="0" pageNumber="189" start="Figures 8, 9" startId="F4">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figures 8, 9.</emphasis>
<caption id="4223352859C2024CBA14D691B4E269F3" doi="10.3897/zookeys.1103.83952.figures8-9" httpUri="https://binary.pensoft.net/fig/691660" pageId="0" pageNumber="189" start="Figures 8, 9" startId="F4">
<paragraph id="C26474D9545BB89B371CF06B16A767C7" pageId="0" pageNumber="189">
<emphasis id="AD17F10DC41DF18C30DC17539183C858" bold="true" pageId="0" pageNumber="189">Figures 8, 9.</emphasis>
Male genitalia
<emphasis bold="true" pageId="0" pageNumber="189">8</emphasis>
<taxonomicName genus="C." lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">
<emphasis italics="true" pageId="0" pageNumber="189">C. herwigvanstaai</emphasis>
<emphasis id="9072CD36B7BDD3C1F25B0D1EECA47B24" bold="true" pageId="0" pageNumber="189">8</emphasis>
<taxonomicName id="8FE3636A79A990549749870315BEFC7C" lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">
<emphasis id="E0B0E799D3CBEB7AFD0F65B73F8FE33C" italics="true" pageId="0" pageNumber="189">C. herwigvanstaai</emphasis>
</taxonomicName>
sp. nov., holotype, Italy, slide GEL 1153 P. Huemer
<emphasis bold="true" pageId="0" pageNumber="189">9</emphasis>
<taxonomicName genus="C." lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
<emphasis id="CC9FEBB18D30469C8B4203EE4C352E43" bold="true" pageId="0" pageNumber="189">9</emphasis>
<taxonomicName id="42EDB591761549FC5FCD44A8C0F49435" lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis id="27A0EB780621512F6E1676CAE465E4EB" italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
</taxonomicName>
, paratype, Greece, slide GEL 1213 P. Huemer;
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1103.83952.figures10-11" httpUri="https://binary.pensoft.net/fig/691661" pageId="0" pageNumber="189" start="Figures 10, 11" startId="F5">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figures 10, 11.</emphasis>
<caption id="14C871EB6B3636BD93659D5BC3E8938D" doi="10.3897/zookeys.1103.83952.figures10-11" httpUri="https://binary.pensoft.net/fig/691661" pageId="0" pageNumber="189" start="Figures 10, 11" startId="F5">
<paragraph id="5C35CAEF36E26DEFB8CF4974AAAA88E4" pageId="0" pageNumber="189">
<emphasis id="55884302BD061733073C63C2994D57FD" bold="true" pageId="0" pageNumber="189">Figures 10, 11.</emphasis>
Female genitalia
<emphasis bold="true" pageId="0" pageNumber="189">10</emphasis>
<taxonomicName baseAuthorityName="Haworth" baseAuthorityYear="1812" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum tricolorella" order="Lepidoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tricolorella">
<emphasis italics="true" pageId="0" pageNumber="189">Caryocolum tricolorella</emphasis>
<emphasis id="62680A330158E22C2660A01ABB2824A3" bold="true" pageId="0" pageNumber="189">10</emphasis>
<taxonomicName id="5878CE3BDBA504EC6393CD60BA8049EB" baseAuthorityName="Haworth" baseAuthorityYear="1812" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum tricolorella" order="Lepidoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="tricolorella">
<emphasis id="FCD72FB23C5944C59A5916B74B121CA4" italics="true" pageId="0" pageNumber="189">Caryocolum tricolorella</emphasis>
</taxonomicName>
, Germany, slide GEL 1092 P. Huemer
<emphasis bold="true" pageId="0" pageNumber="189">11</emphasis>
<taxonomicName genus="C." lsidName="C. fibigerium" pageId="0" pageNumber="189" rank="species" species="fibigerium">
<emphasis italics="true" pageId="0" pageNumber="189">C. fibigerium</emphasis>
<emphasis id="B60D5E62CA3461C014C7060A69FE25E5" bold="true" pageId="0" pageNumber="189">11</emphasis>
<taxonomicName id="277D4CB52D089A2BC90BE7A110F28AAF" lsidName="C. fibigerium" pageId="0" pageNumber="189" rank="species" species="fibigerium">
<emphasis id="B306115F59FA012E572223D9D3D999F5" italics="true" pageId="0" pageNumber="189">C. fibigerium</emphasis>
</taxonomicName>
, Spain, slide GEL 1095 P. Huemer.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" italics="true" pageId="0" pageNumber="189">Female genitalia</emphasis>
<paragraph id="D20FF66B3E879CA1A2D22DFFA6276D90" pageId="0" pageNumber="189">
<emphasis id="44C5EB05B3FF7A411BB8CCCFE859DCC7" bold="true" italics="true" pageId="0" pageNumber="189">Female genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 12, 13" captionStartId="F6" captionText="Figures 12, 13. Female genitalia 12 C. herwigvanstaai sp. nov., paratype, Italy, slide GEL 1155 P. Huemer 13 C. olekarsholti, paratype, Greece, slide GEL 1231 P. Huemer." figureDoi="10.3897/zookeys.1103.83952.figures12-13" httpUri="https://binary.pensoft.net/fig/691662" pageId="0" pageNumber="189">12</figureCitation>
<figureCitation id="5FCE6B69F8519753530DB3C7802E8771" captionStart="Figures 12, 13" captionStartId="F6" captionText="Figures 12, 13. Female genitalia 12 C. herwigvanstaai sp. nov., paratype, Italy, slide GEL 1155 P. Huemer 13 C. olekarsholti, paratype, Greece, slide GEL 1231 P. Huemer." figureDoi="10.3897/zookeys.1103.83952.figures12-13" httpUri="https://binary.pensoft.net/fig/691662" pageId="0" pageNumber="189">12</figureCitation>
). Apophysis posterior about 4.5 times length of apophysis anterior; segment VIII with suboval sclerotized dorsolateral zones, with small dorsolateral flaps, posterior and inner edge strongly sclerotized, membranous ventromedial part with numerous microtrichia; apophysis anterior about length of segment VIII; antrum moderately large, funnel-shaped, shorter than apophysis anterior and segment VIII, basally about 1/2 width of segment VIII between bases of apophyses anteriores, posterior edge weakly concave; ductus bursae about twice length of apophysis anterior; corpus bursae semi-oval, signum a crescent-shaped basal plate with moderately long and stout hook.
</paragraph>
<caption doi="10.3897/zookeys.1103.83952.figures12-13" httpUri="https://binary.pensoft.net/fig/691662" pageId="0" pageNumber="189" start="Figures 12, 13" startId="F6">
<paragraph pageId="0" pageNumber="189">
<emphasis bold="true" pageId="0" pageNumber="189">Figures 12, 13.</emphasis>
<caption id="384466DFDDC29C88F8A8631B782C7C22" doi="10.3897/zookeys.1103.83952.figures12-13" httpUri="https://binary.pensoft.net/fig/691662" pageId="0" pageNumber="189" start="Figures 12, 13" startId="F6">
<paragraph id="FBFB31561319CDE2433094FB4D22E4FD" pageId="0" pageNumber="189">
<emphasis id="DEB8B60DB9FB592EFC413C56DC2BA8FB" bold="true" pageId="0" pageNumber="189">Figures 12, 13.</emphasis>
Female genitalia
<emphasis bold="true" pageId="0" pageNumber="189">12</emphasis>
<taxonomicName genus="C." lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">
<emphasis italics="true" pageId="0" pageNumber="189">C. herwigvanstaai</emphasis>
<emphasis id="AACF2CB2250FEA3D085035EB51FB7A81" bold="true" pageId="0" pageNumber="189">12</emphasis>
<taxonomicName id="2005C2AA47125187EF32509B5D286E44" lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">
<emphasis id="453E34AFABC91B080B744CB60570BF01" italics="true" pageId="0" pageNumber="189">C. herwigvanstaai</emphasis>
</taxonomicName>
sp. nov., paratype, Italy, slide GEL 1155 P. Huemer
<emphasis bold="true" pageId="0" pageNumber="189">13</emphasis>
<taxonomicName genus="C." lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
<emphasis id="EA88B84D6CC5CCAF295793A7C50C7A02" bold="true" pageId="0" pageNumber="189">13</emphasis>
<taxonomicName id="0507829A6675DA70BFD58E6739889CFA" lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis id="01A974E7F83D2E6FB7727D94F607248A" italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
</taxonomicName>
, paratype, Greece, slide GEL 1231 P. Huemer.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="molecular data">
<paragraph pageId="0" pageNumber="189">Molecular data.</paragraph>
<paragraph pageId="0" pageNumber="189">
<subSubSection id="B6E4B4292729A75A2C5BED87FC213881" pageId="0" pageNumber="189" type="molecular data">
<paragraph id="B54DE52F374D495534F47D1C5494115F" pageId="0" pageNumber="189">Molecular data.</paragraph>
<paragraph id="39F1A1200B87256DEF3E5F243F474F9B" pageId="0" pageNumber="189">
BINs: BOLD:AAO2674, BOLD:ADK9243. A genetically variable species splitting into two BINs which, however, require re-evaluation from additional material. The distance between both BINs is 2.1% (
<emphasis italics="true" pageId="0" pageNumber="189">n</emphasis>
<emphasis id="25A0E949A6689D752B89F3CE4E492089" italics="true" pageId="0" pageNumber="189">n</emphasis>
= 3). The minimum distance to the nearest neighbour,
<taxonomicName genus="C." lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
<taxonomicName id="84F2E1B750BF35619280C1A70163956C" lsidName="C. olekarsholti" pageId="0" pageNumber="189" rank="species" species="olekarsholti">
<emphasis id="9EF5A404C660F2925ADA894AB9210151" italics="true" pageId="0" pageNumber="189">C. olekarsholti</emphasis>
</taxonomicName>
, is 4.12%.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="etymology">
<paragraph pageId="0" pageNumber="189">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="189">The species is dedicated to DDr Herwig van Staa (Innsbruck, Austria), former governor of the province of Tyrol on his 80th birthday on the 10 June 2022, and in recognition of his tremendous support of the Tyrolean Federal State Museums and the Alpenzoo Innsbruck, resulting in a joint Natural History Museum.</paragraph>
<subSubSection id="9521E2FE6115B4CE25E65A1459A48218" pageId="0" pageNumber="189" type="etymology">
<paragraph id="A9467CB275812C8E79821A81CA93751B" pageId="0" pageNumber="189">Etymology.</paragraph>
<paragraph id="AA3AD8319193DD44BFE7D1F0DD2B2A85" pageId="0" pageNumber="189">The species is dedicated to DDr Herwig van Staa (Innsbruck, Austria), former governor of the province of Tyrol on his 80th birthday on the 10 June 2022, and in recognition of his tremendous support of the Tyrolean Federal State Museums and the Alpenzoo Innsbruck, resulting in a joint Natural History Museum.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="distribution">
<paragraph pageId="0" pageNumber="189">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="189">
<subSubSection id="378C6A1F91E56057271FE4B4F6363683" pageId="0" pageNumber="189" type="distribution">
<paragraph id="AC2FF985C432F674FFA3B594DCEB3150" pageId="0" pageNumber="189">Distribution.</paragraph>
<paragraph id="87BD1A67274B684266B9324604D6AA84" pageId="0" pageNumber="189">
The species is currently only known from Central Italy but may have a wider distribution on the Italian Peninsula.
<bibRefCitation author="Mariani, M" journalOrPublisher="Memoria" pageId="0" pageNumber="189" pagination="1 - 237" refId="B27" refString="Mariani, M, 1943. Fauna Lepidopterorum Italiae. Parte I. Catalogo ragionato dei Lepidotteri d'Italia. Giornale di Scienze naturali ed economiche, Palermo, 42 (1940-41). Memoria 3: 1 - 237" title="Fauna Lepidopterorum Italiae. Parte I. Catalogo ragionato dei Lepidotteri d'Italia. Giornale di Scienze naturali ed economiche, Palermo, 42 (1940 - 41)." volume="3" year="1943">Mariani (1943)</bibRefCitation>
<bibRefCitation id="E138555255515FD6B5AD2E0CA9EA1C3D" author="Mariani, M" journalOrPublisher="Memoria" pageId="0" pageNumber="189" pagination="1 - 237" refId="B27" refString="Mariani, M, 1943. Fauna Lepidopterorum Italiae. Parte I. Catalogo ragionato dei Lepidotteri d'Italia. Giornale di Scienze naturali ed economiche, Palermo, 42 (1940-41). Memoria 3: 1 - 237" title="Fauna Lepidopterorum Italiae. Parte I. Catalogo ragionato dei Lepidotteri d'Italia. Giornale di Scienze naturali ed economiche, Palermo, 42 (1940 - 41)." volume="3" year="1943">Mariani (1943)</bibRefCitation>
had published a record of
<taxonomicName genus="C." lsidName="C. tricolorella" pageId="0" pageNumber="189" rank="species" species="tricolorella">
<emphasis italics="true" pageId="0" pageNumber="189">C. tricolorella</emphasis>
<taxonomicName id="2EDC29DAF463E1E13775DAFD21831932" lsidName="C. tricolorella" pageId="0" pageNumber="189" rank="species" species="tricolorella">
<emphasis id="9A3C804BE8CF01A1A270A401CDE90EF4" italics="true" pageId="0" pageNumber="189">C. tricolorella</emphasis>
</taxonomicName>
from Sicily, which possibly is
<taxonomicName genus="C." lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">
<emphasis italics="true" pageId="0" pageNumber="189">C. herwigvanstaai</emphasis>
<taxonomicName id="655C210BF11ACAA45D20811F8EEBD114" lsidName="C. herwigvanstaai" pageId="0" pageNumber="189" rank="species" species="herwigvanstaai">
<emphasis id="052D89AFFCD40600601F5B2D821CC48C" italics="true" pageId="0" pageNumber="189">C. herwigvanstaai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="189" type="bionomics">
<paragraph pageId="0" pageNumber="189">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="189">
<subSubSection id="D135329588C10FC8D74EE19EC9C166DC" pageId="0" pageNumber="189" type="bionomics">
<paragraph id="9FCE08E0C4A1FDC438DDD574DC45CFCF" pageId="0" pageNumber="189">Bionomics.</paragraph>
<paragraph id="141A62B5BF183D46102D61E8415473E4" pageId="0" pageNumber="189">
Host-plant and early stages are undescribed but it seems most likely that the species shows a similar behaviour as related taxa with the potential host-plant among
<taxonomicName class="Magnoliopsida" family="Caryophyllaceae" genus="Cerastium" kingdom="Plantae" lsidName="Cerastium" order="Caryophyllales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="189">Cerastium</emphasis>
<taxonomicName id="8537C62459C5DFDF6C75BBDD6C70D228" class="Magnoliopsida" family="Caryophyllaceae" genus="Cerastium" kingdom="Plantae" lsidName="Cerastium" order="Caryophyllales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">
<emphasis id="AE6C350E9F0064E72A80E32E22EABB28" italics="true" pageId="0" pageNumber="189">Cerastium</emphasis>
</taxonomicName>
or related genera of
<taxonomicName class="Magnoliopsida" family="Caryophyllaceae" genus="Cerastium" kingdom="Plantae" lsidName="" order="Caryophyllales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="family">Caryophyllaceae</taxonomicName>
<taxonomicName id="962BE1CD845268010A4D694E7E56C065" class="Magnoliopsida" family="Caryophyllaceae" kingdom="Plantae" lsidName="" order="Caryophyllales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="family">Caryophyllaceae</taxonomicName>
. The adults have been found in mid-July at artificial light sources near rock and scree on calcareous soil at altitudes of about 1700-1800 m.
</paragraph>
</subSubSection>