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<mods:title id="6DDD6996F018CF7D9A4CE4527DC13698">Re-evaluation of Zospeumschaufussi von Frauenfeld, 1862 and Z. suarezi Gittenberger, 1980, including the description of two new Iberian species using Computer Tomography (CT) (Eupulmonata, Ellobioidea, Carychiidae)</mods:title>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
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<mods:namePart>E. Prieto, Carlos</mods:namePart>
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<mods:namePart>Kampschulte, Marian</mods:namePart>
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<mods:namePart>Martels, Gunhild</mods:namePart>
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<mods:namePart>Ruthensteiner, Bernhard</mods:namePart>
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<treatment ID-GBIF-Taxon="154599944" LSID="urn:lsid:zoobank.org:act:3B97291A-8B05-41BA-96B1-8F6B50C22B69" httpUri="http://treatment.plazi.org/id/0410D4DC6B967E28E927F47701690A45" lastPageNumber="65" pageId="0" pageNumber="65">
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<taxonomicName LSID="http://zoobank.org/3B97291A-8B05-41BA-96B1-8F6B50C22B69" authority="Jochum, Prieto & De Winter" class="Gastropoda" family="Carychiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum praetermissum" order="Basommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="praetermissum">Zospeum praetermissum Jochum, Prieto & De Winter</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="65">sp. n.</taxonomicNameLabel>
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<treatment id="0410D4DC6B967E28E927F47701690A45" ID-GBIF-Taxon="154599944" LSID="urn:lsid:zoobank.org:act:3B97291A-8B05-41BA-96B1-8F6B50C22B69" httpUri="http://treatment.plazi.org/id/0410D4DC6B967E28E927F47701690A45" lastPageNumber="65" pageId="0" pageNumber="65" scope_family="Carychiidae" scope_superFamily="Ellobioidea" scope_superOrder="Eupulmonata">
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<subSubSection id="CA867938E7A8AAF6220CC91BF827622E" pageId="0" pageNumber="65" type="nomenclature">
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<paragraph id="F278A5A93C9F43A76DB81B841BC37A37" pageId="0" pageNumber="65">
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<taxonomicName id="DB6841699DA362C839D63023DBF86AAA" LSID="http://zoobank.org/3B97291A-8B05-41BA-96B1-8F6B50C22B69" authority="Jochum, Prieto & De Winter" family="Carychiidae" genus="Zospeum" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Zospeum praetermissum" order="Basommatophora" pageId="0" pageNumber="65" rank="species" species="praetermissum">Zospeum praetermissum Jochum, Prieto & De Winter</taxonomicName>
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<taxonomicNameLabel id="4EB7237FFE5AE9BFABC8CB6597B4A6A8" pageId="0" pageNumber="65">sp. n.</taxonomicNameLabel>
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Figures 5
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<normalizedToken originalValue="G–M">G-M</normalizedToken>
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<normalizedToken id="B636C62744A2ED46E365DBFD2EA6C96A" originalValue="G–M">G-M</normalizedToken>
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, 10, 11
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="reference_group">
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<paragraph pageId="0" pageNumber="65">
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<taxonomicName class="Gastropoda" family="Carychiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum suarezi" order="Basommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="suarezi">Zospeum suarezi</taxonomicName>
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<subSubSection id="975E3753BB2B54D0F2566800C48EB68B" pageId="0" pageNumber="65" type="reference_group">
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<paragraph id="163B9C095F3391433AD0250D3B589B12" pageId="0" pageNumber="65">
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<taxonomicName id="2DDE79B5BFBBC320BF3024D8066AF71E" family="Carychiidae" genus="Zospeum" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Zospeum suarezi" order="Basommatophora" pageId="0" pageNumber="65" rank="species" species="suarezi">Zospeum suarezi</taxonomicName>
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-
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<bibRefCitation author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger 1980</bibRefCitation>
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<bibRefCitation id="1BEEBCB2CAAC83C948DB2DFC34ADC82A" author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger 1980</bibRefCitation>
|
||||
: 203, Fig. 2 (only shells from Cueva de Inguanzo).
|
||||
</paragraph>
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<paragraph pageId="0" pageNumber="65">
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||||
<taxonomicName class="Gastropoda" family="Carychiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum suarezi" order="Basommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="suarezi">Zospeum suarezi</taxonomicName>
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<paragraph id="F57362DD81622D9DB0E29FF64E44EBB3" pageId="0" pageNumber="65">
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<taxonomicName id="95D1A7112CCCC3B88A5C0F0B3BCAD38E" family="Carychiidae" genus="Zospeum" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Zospeum suarezi" order="Basommatophora" pageId="0" pageNumber="65" rank="species" species="suarezi">Zospeum suarezi</taxonomicName>
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||||
-
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<bibRefCitation pageId="0" pageNumber="65">
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<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
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<bibRefCitation id="0B069FD6F63B901BFB2B104D4C30C30B" pageId="0" pageNumber="65">
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<normalizedToken id="3DE25CA22FE93ABCF1CE242F4AC7E85B" originalValue="Gómez">Gomez</normalizedToken>
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and Prieto 1983
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</bibRefCitation>
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: 8, Fig. 1 (only the named shell).
|
||||
</paragraph>
|
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="type material">
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<paragraph pageId="0" pageNumber="65">Type Material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="65">Holotype. SPAIN Cueva del Puente de Inguanzo, Inguanzo, Concejo de Cabrales, Asturias, MGRS 30TUN4897097640; N43.315574, W4.860905; 230 m a.s.l.; 19 Feb 1979; G. Favre & R. Emery leg.; RMNH.MOL.55391 [55386]. Paratypes. SPAIN locus typicus: 2 shells; data as the holotype; RMNH.MOL.339954 [55386]. Other material. MHNG 96219/1 shell (now lost).</paragraph>
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<subSubSection id="83B0BFCA8F833AB90CF6F9EC13648ED5" pageId="0" pageNumber="65" type="type material">
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<paragraph id="967520830E48B0A1982D0906EDC86C77" pageId="0" pageNumber="65">Type Material.</paragraph>
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<paragraph id="4ED720613BF37A34DA5F36D842B2186C" pageId="0" pageNumber="65">Holotype. SPAIN Cueva del Puente de Inguanzo, Inguanzo, Concejo de Cabrales, Asturias, MGRS 30TUN4897097640; N43.315574, W4.860905; 230 m a.s.l.; 19 Feb 1979; G. Favre & R. Emery leg.; RMNH.MOL.55391 [55386]. Paratypes. SPAIN locus typicus: 2 shells; data as the holotype; RMNH.MOL.339954 [55386]. Other material. MHNG 96219/1 shell (now lost).</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="65" type="diagnosis">
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||||
<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="65">Shell ca. 1.1-1.2 mm, conical with a roundish and moderately thick peristome, lacking apparent apertural barriers but with a small distinct lamella (denticle) in the parietal-columellar corner; internally, columella robust with a central lamella, a low upper columellar bulge and a basal umbilical ridge.</paragraph>
|
||||
<subSubSection id="524C8BFEDA0B0898F803F10AD65FA48F" pageId="0" pageNumber="65" type="diagnosis">
|
||||
<paragraph id="FB2A9DFD5BDFAFEFD883A46EF4E97A69" pageId="0" pageNumber="65">Diagnosis.</paragraph>
|
||||
<paragraph id="380DC0F1A8FCE1F25345AA5A7A77C660" pageId="0" pageNumber="65">Shell ca. 1.1-1.2 mm, conical with a roundish and moderately thick peristome, lacking apparent apertural barriers but with a small distinct lamella (denticle) in the parietal-columellar corner; internally, columella robust with a central lamella, a low upper columellar bulge and a basal umbilical ridge.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="65" type="description">
|
||||
<paragraph pageId="0" pageNumber="65">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="65">Measurements of holotype and paratypes are presented in Table 1. Shell minute, fresh shells transparent, variable in shape (SH:SD ratio 1.41-1.57) with ca. 5 whorls, regularly coiled, suture deep, whorls convex, more or less strongly shouldered; teleoconch sculpture consists of occasional blunt growth lines; robust columella with an inclinate, central lamella (Fig. 11F); an upper lamellar bulge swells from the base of the inner penultimate whorl above the central lamella (Fig. 11G, I, K); a distinct, short, ridge projects from the base of the columella above the umbilical indentation (Fig. 11G, I, K); aperture more or less circular; peristome adhering to spire, reflected, moderately thickened, roundish; umbilicus closed; apertural barriers absent apart from the low lamella that appears externally as a small, but distinct denticle on the parietal-columellar corner visible in oblique apertural view (Figs 5M, 10D).</paragraph>
|
||||
<caption pageId="0" pageNumber="65">
|
||||
<paragraph pageId="0" pageNumber="65">
|
||||
<subSubSection id="58C7E46EA15E89F4F6BE89AC91BA4CCA" pageId="0" pageNumber="65" type="description">
|
||||
<paragraph id="31CF76425C72AB46F281BC6E41A4E9E2" pageId="0" pageNumber="65">Description.</paragraph>
|
||||
<paragraph id="3F3850C45BCFA301ED5B6B23236C3DA6" pageId="0" pageNumber="65">Measurements of holotype and paratypes are presented in Table 1. Shell minute, fresh shells transparent, variable in shape (SH:SD ratio 1.41-1.57) with ca. 5 whorls, regularly coiled, suture deep, whorls convex, more or less strongly shouldered; teleoconch sculpture consists of occasional blunt growth lines; robust columella with an inclinate, central lamella (Fig. 11F); an upper lamellar bulge swells from the base of the inner penultimate whorl above the central lamella (Fig. 11G, I, K); a distinct, short, ridge projects from the base of the columella above the umbilical indentation (Fig. 11G, I, K); aperture more or less circular; peristome adhering to spire, reflected, moderately thickened, roundish; umbilicus closed; apertural barriers absent apart from the low lamella that appears externally as a small, but distinct denticle on the parietal-columellar corner visible in oblique apertural view (Figs 5M, 10D).</paragraph>
|
||||
<caption id="B02B2F582ABAAE34D06BCEB3D9D54495" pageId="0" pageNumber="65">
|
||||
<paragraph id="83E37A0C236F9571D04629FE22B1DE50" pageId="0" pageNumber="65">
|
||||
Figure 4.
|
||||
<normalizedToken originalValue="A–C">A-C</normalizedToken>
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum schaufussi" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="schaufussi">Zospeum schaufussi</taxonomicName>
|
||||
<normalizedToken id="35432E5C456F650F406BCFA5838AF19B" originalValue="A–C">A-C</normalizedToken>
|
||||
<taxonomicName id="212F795B53269EC83D158C129BB18324" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum schaufussi" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="schaufussi">Zospeum schaufussi</taxonomicName>
|
||||
von Frauenfeld, 1862, Cueva del
|
||||
<normalizedToken originalValue="Búho">Buho</normalizedToken>
|
||||
<normalizedToken id="EA54E54860A34357ED31C0D0F0410EE2" originalValue="Búho">Buho</normalizedToken>
|
||||
, Puente Viesgo, Santander, (holotype of
|
||||
<taxonomicName lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
<taxonomicName id="A072F53983E90E2E2C0A6C93F2ADC241" lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
, RMNH.MOL.55383)
|
||||
<normalizedToken originalValue="D–E">D-E</normalizedToken>
|
||||
<taxonomicName lsidName="Z. praetermissum" pageId="0" pageNumber="65" rank="species" species="praetermissum">Z. praetermissum</taxonomicName>
|
||||
<normalizedToken id="A13061C9748FA380F6D869BFC6789F0C" originalValue="D–E">D-E</normalizedToken>
|
||||
<taxonomicName id="A873D52C0B3EC62B7B7FCD9A68CAE689" lsidName="Z. praetermissum" pageId="0" pageNumber="65" rank="species" species="praetermissum">Z. praetermissum</taxonomicName>
|
||||
sp. n., Cueva del Puente de Inguanzo (RMNH.MOL55389). Scale bar: 500
|
||||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||||
<normalizedToken id="EE82A30AB2E3D0896AE996A1D7FE07B7" originalValue="μm">μm</normalizedToken>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="65" type="differential diagnosis">
|
||||
<paragraph pageId="0" pageNumber="65">Differential diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="65">
|
||||
<subSubSection id="55C28F182FC3DA0F645ADCA5FED1078E" pageId="0" pageNumber="65" type="differential diagnosis">
|
||||
<paragraph id="053A68D5D7E2898026D1B241F357A6FD" pageId="0" pageNumber="65">Differential diagnosis.</paragraph>
|
||||
<paragraph id="A15E65B461905F1FAE4524ACAA547A47" pageId="0" pageNumber="65">
|
||||
Differs from
|
||||
<taxonomicName lsidName="Z. schaufussi" pageId="0" pageNumber="65" rank="species" species="schaufussi">Z. schaufussi</taxonomicName>
|
||||
<taxonomicName id="A61CD14524E8469777748FC267E9B214" lsidName="Z. schaufussi" pageId="0" pageNumber="65" rank="species" species="schaufussi">Z. schaufussi</taxonomicName>
|
||||
externally by its more conical shell with less tightly coiled whorls - adult shells having around 5 rather than 6 whorls of the same size - and a more pronounced lamella/denticle visible in the parietal-columellar corner in oblique view; internally by its robust columella and by its pronounced basal columellar ridge (above umbilical indentation).
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum praetermissum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="praetermissum">Zospeum praetermissum</taxonomicName>
|
||||
<taxonomicName id="D3605537C36C90C5215AED98F13B5978" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum praetermissum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="praetermissum">Zospeum praetermissum</taxonomicName>
|
||||
is easily distinguished from
|
||||
<taxonomicName lsidName="Z. biscaiense" pageId="0" pageNumber="65" rank="species" species="biscaiense">Z. biscaiense</taxonomicName>
|
||||
<taxonomicName id="9C652CFCAA02837674641C71323E7C64" lsidName="Z. biscaiense" pageId="0" pageNumber="65" rank="species" species="biscaiense">Z. biscaiense</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="Z. zaldivarae" pageId="0" pageNumber="65" rank="species" species="zaldivarae">Z. zaldivarae</taxonomicName>
|
||||
<taxonomicName id="FEB64612791776C51C39534F98A347C3" lsidName="Z. zaldivarae" pageId="0" pageNumber="65" rank="species" species="zaldivarae">Z. zaldivarae</taxonomicName>
|
||||
in shell and peristome shape and apertural characters, whereas
|
||||
<taxonomicName lsidName="Z. percostulatum" pageId="0" pageNumber="65" rank="species" species="percostulatum">Z. percostulatum</taxonomicName>
|
||||
<taxonomicName id="24311CBE1CBACC2A9DB6080122E1C2F7" lsidName="Z. percostulatum" pageId="0" pageNumber="65" rank="species" species="percostulatum">Z. percostulatum</taxonomicName>
|
||||
is distinctly ribbed.
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum gittenbergeri" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="gittenbergeri">Zospeum gittenbergeri</taxonomicName>
|
||||
<taxonomicName id="878FDAF257B08A5F4636F53ADC71D056" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum gittenbergeri" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="gittenbergeri">Zospeum gittenbergeri</taxonomicName>
|
||||
differs by its broad, angular peristome and barely developed lamella on the columella.
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum vasconicum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="vasconicum">Zospeum vasconicum</taxonomicName>
|
||||
<taxonomicName id="92C55DB402A2440F6D40D3B0EEA08CEE" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum vasconicum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="vasconicum">Zospeum vasconicum</taxonomicName>
|
||||
has a more rounded aperture with an almost uniformly-thickened peristome.
|
||||
<taxonomicName lsidName="Z. bellesi" pageId="0" pageNumber="65" rank="species" species="bellesi">Z. bellesi</taxonomicName>
|
||||
<taxonomicName id="3A54DC0DB274B440EB99E13A5911DA81" lsidName="Z. bellesi" pageId="0" pageNumber="65" rank="species" species="bellesi">Z. bellesi</taxonomicName>
|
||||
has no apertural barriers nor a columellar lamella.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="65" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="65">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="65">
|
||||
<subSubSection id="587A2A70B8C4BB87EAB990FCA4D91ED6" pageId="0" pageNumber="65" type="etymology">
|
||||
<paragraph id="72B661DD92D092A058B664E6C123B9C8" pageId="0" pageNumber="65">Etymology.</paragraph>
|
||||
<paragraph id="5BB8430D2E6FA6F166021A7B2CD2A7F3" pageId="0" pageNumber="65">
|
||||
The name,
|
||||
<taxonomicName lsidName="praetermissum" pageId="0" pageNumber="65" rank="species" species="praetermissum">praetermissum</taxonomicName>
|
||||
<taxonomicName id="CB41D5B4B185A8FC6FA7FF8825488340" lsidName="praetermissum" pageId="0" pageNumber="65" rank="species" species="praetermissum">praetermissum</taxonomicName>
|
||||
, refers to the situation that the holotype shell was originally not recognized as distinct from
|
||||
<taxonomicName lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
<taxonomicName id="BDBD68A8E66439A59ABED267A605FE57" lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="65" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="65">Only known from the type locality.</paragraph>
|
||||
<subSubSection id="95547EE4950099F3FAD60022890C5CC0" pageId="0" pageNumber="65" type="distribution">
|
||||
<paragraph id="9A97AE726381FB91219A0799408EBA83" pageId="0" pageNumber="65">Distribution.</paragraph>
|
||||
<paragraph id="F6C4A39DB564BA99119998F4B7CEDDE0" pageId="0" pageNumber="65">Only known from the type locality.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="65" type="ecology">
|
||||
<paragraph pageId="0" pageNumber="65">Ecology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="65">
|
||||
<subSubSection id="EFAF2DE029354F55AC3CB4BC8D846680" pageId="0" pageNumber="65" type="ecology">
|
||||
<paragraph id="796DA74936DAF8DB3F8DD329B7038D6A" pageId="0" pageNumber="65">Ecology.</paragraph>
|
||||
<paragraph id="F213B01FB821512A3F20C6C076071D2F" pageId="0" pageNumber="65">
|
||||
According to the records of the speleologist
|
||||
<normalizedToken originalValue="Gérald">Gerald</normalizedToken>
|
||||
<normalizedToken id="8658D156DD051DFB2E0528A223E9E676" originalValue="Gérald">Gerald</normalizedToken>
|
||||
Favre (pers. comm. 2017), the collection site was located ca. 200 m from the cave entrance (also noted on the data label NHMG-Moll 96219 (978.363)). His field notes document that the collection area consisted of sandy substrate, some chestnut fragments and that the humidity level was low. The total length of the cave is 1500 m.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="65" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="65">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="65">
|
||||
<subSubSection id="9A0A64AEF21759A445FFB98DA3CA0AF7" pageId="0" pageNumber="65" type="remarks">
|
||||
<paragraph id="F1D5DE39D817E0F0AA9636082A5FB176" pageId="0" pageNumber="65">Remarks.</paragraph>
|
||||
<paragraph id="6662100A81AB633F2DAA6C99C54BEEBF" pageId="0" pageNumber="65">
|
||||
The label to the shell material
|
||||
<bibRefCitation author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger (1980)</bibRefCitation>
|
||||
<bibRefCitation id="F4E851AE0C28C5F68D5226C56266A607" author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger (1980)</bibRefCitation>
|
||||
assigned as
|
||||
<taxonomicName lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
<taxonomicName id="AF35274F61118FB904EB18F8BC02663D" lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
paratypes (NHMG-Moll.96219/1 shell), bore
|
||||
<normalizedToken originalValue="Gittenberger’s">Gittenberger's</normalizedToken>
|
||||
<normalizedToken id="9D3ADE7BF96DD31FF616F17B21CB036E" originalValue="Gittenberger’s">Gittenberger's</normalizedToken>
|
||||
apparent ambivalence "Zospeum schaufussi Frfld Gitt. det. 1979" (Fig. 5L). Although currently lost, this shell was most likely conspecific with
|
||||
<taxonomicName lsidName="Z. praetermissum" pageId="0" pageNumber="65" rank="species" species="praetermissum">Z. praetermissum</taxonomicName>
|
||||
<taxonomicName id="39EEC680C39F2480949D4378229499C2" lsidName="Z. praetermissum" pageId="0" pageNumber="65" rank="species" species="praetermissum">Z. praetermissum</taxonomicName>
|
||||
. We consider the paratype
|
||||
<taxonomicName lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
<taxonomicName id="BDA5E339C739113AC22E200BDE45DDDA" lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
shell (NHMW-MO 75000-E-48815) from Cueva Hernialde, Guipuzcoa to be Z.
|
||||
<taxonomicName lsidName="vasconicum" pageId="0" pageNumber="65" rank="species" species="vasconicum">cf. vasconicum</taxonomicName>
|
||||
<taxonomicName id="AA7EAB702BAFCDA56A8B1C7D92B735D8" lsidName="vasconicum" pageId="0" pageNumber="65" rank="species" species="vasconicum">cf. vasconicum</taxonomicName>
|
||||
(Fig. 6).
|
||||
</paragraph>
|
||||
<caption pageId="0" pageNumber="65">
|
||||
<paragraph pageId="0" pageNumber="65">
|
||||
<caption id="7430A14264105921A76781345980477F" pageId="0" pageNumber="65">
|
||||
<paragraph id="66F8827761C41629C3066FD7E2C85431" pageId="0" pageNumber="65">
|
||||
Figure 5.
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">Zospeum</taxonomicName>
|
||||
<taxonomicName id="FA5F76A92BA240D4491E958282AF96B6" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">Zospeum</taxonomicName>
|
||||
material assessed in
|
||||
<bibRefCitation author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger (1980)</bibRefCitation>
|
||||
<bibRefCitation id="61D2BE3AFA68BAE2892AA6052DD8489E" author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger (1980)</bibRefCitation>
|
||||
.
|
||||
<normalizedToken originalValue="A–F">A-F</normalizedToken>
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum praetermissum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="praetermissum">Zospeum praetermissum</taxonomicName>
|
||||
<normalizedToken id="85010DAFD5FBEB7A3980E78825086066" originalValue="A–F">A-F</normalizedToken>
|
||||
<taxonomicName id="9C36342F7D276DD8D56D81E8C0B45D77" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum praetermissum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="praetermissum">Zospeum praetermissum</taxonomicName>
|
||||
sp. n. (paratype of
|
||||
<taxonomicName lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
<taxonomicName id="4270F716C7E300E961DFE8A88FA5501F" lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
, MHNG-Mol 96220/1 shell, now lost), Cueva Los Quesos (showing ambivalent label,
|
||||
<taxonomicName lsidName="Z. schaufussi" pageId="0" pageNumber="65" rank="species" species="schaufussi">Z. schaufussi</taxonomicName>
|
||||
<taxonomicName id="6509F8D36882E229BDCC45018D889CDA" lsidName="Z. schaufussi" pageId="0" pageNumber="65" rank="species" species="schaufussi">Z. schaufussi</taxonomicName>
|
||||
Frfld)
|
||||
<normalizedToken originalValue="G–M">G-M</normalizedToken>
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum praetermissum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="praetermissum">Zospeum praetermissum</taxonomicName>
|
||||
<normalizedToken id="1BC455C74B1BABEAE18669B800E0ECC3" originalValue="G–M">G-M</normalizedToken>
|
||||
<taxonomicName id="236CBC93E4F0D13B814F2DDD12366B5B" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum praetermissum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="praetermissum">Zospeum praetermissum</taxonomicName>
|
||||
sp. n (figured paratype of
|
||||
<taxonomicName lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
<taxonomicName id="E5A18BBC1E0198C96D32CD83862DC900" lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
, MHNG-Moll 96219, now lost), Cueva del Puente de Inguanzo L,
|
||||
<normalizedToken originalValue="N–R">N-R</normalizedToken>
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum gittenbergeri" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="gittenbergeri">Zospeum gittenbergeri</taxonomicName>
|
||||
<normalizedToken id="FC5681700A54EF574917A95480985FE0" originalValue="N–R">N-R</normalizedToken>
|
||||
<taxonomicName id="AF49D0F3ED4904D1553C045B06096C30" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum gittenbergeri" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="gittenbergeri">Zospeum gittenbergeri</taxonomicName>
|
||||
sp. n. (figured shell of
|
||||
<taxonomicName lsidName="Z. schaufussi" pageId="0" pageNumber="65" rank="species" species="schaufussi">Z. schaufussi</taxonomicName>
|
||||
<taxonomicName id="A7940CA6E20C06AF77E051AA65A219FA" lsidName="Z. schaufussi" pageId="0" pageNumber="65" rank="species" species="schaufussi">Z. schaufussi</taxonomicName>
|
||||
sensu
|
||||
<bibRefCitation author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger (1980)</bibRefCitation>
|
||||
<bibRefCitation id="990A5E20105FB1F06653DD5BE5CBB907" author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger (1980)</bibRefCitation>
|
||||
, MHNG 96219, now lost).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption pageId="0" pageNumber="65">
|
||||
<paragraph pageId="0" pageNumber="65">
|
||||
<caption id="D7A1F213CE072338DDFDA823197E6B09" pageId="0" pageNumber="65">
|
||||
<paragraph id="6A8A683DBF104FB16C75E6B0F73D9730" pageId="0" pageNumber="65">
|
||||
Figure 6.
|
||||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum vasconicum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="vasconicum">Zospeum cf. vasconicum</taxonomicName>
|
||||
<taxonomicName id="6C42DC182565FCEAE0083DC32FB95CBC" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum vasconicum" order="Ellobiida" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="vasconicum">Zospeum cf. vasconicum</taxonomicName>
|
||||
Prieto, De Winter, Weigand,
|
||||
<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
|
||||
<normalizedToken id="4AFE89039DBA0656350DC4FF4A910786" originalValue="Gómez">Gomez</normalizedToken>
|
||||
& Jochum, 2015. Ex.
|
||||
<taxonomicName lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
<taxonomicName id="587C0D4429D79C8C250C8A5654F79AD0" lsidName="Z. suarezi" pageId="0" pageNumber="65" rank="species" species="suarezi">Z. suarezi</taxonomicName>
|
||||
paratype, NHMW-MO 75000-E-48815, Cueva Hernialde, Guipuzcoa, assessed in
|
||||
<bibRefCitation author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger (1980)</bibRefCitation>
|
||||
<bibRefCitation id="C795D6DD24C69C2E07DDACF0888A4849" author="Gittenberger, E" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="65" pagination="201 - 213" title="Three Notes on Iberian Terrestrial Gastropods." volume="55" year="1980">Gittenberger (1980)</bibRefCitation>
|
||||
("NWW-Coll. Edlauer 48815, ex. Coll.
|
||||
<normalizedToken originalValue="Robić/1shell”">Robic/1shell"</normalizedToken>
|
||||
<normalizedToken id="8D0F7D0E2EE3AA66A5D898357D10808A" originalValue="Robić/1shell”">Robic/1shell"</normalizedToken>
|
||||
).
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
|
|||
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