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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.701.13261" ID-GBIF-Dataset="a9b15909-ea61-40d7-ba04-325b8af91669" ID-PMC="PMC5674090" ID-Pensoft-Pub="1313-2970-701-1" ID-PubMed="29118596" ID-ZBK="23BDD7CE1C7E4D2092A8ED47267579FD" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-701-1" ModsDocOrigin="ZooKeys 701" ModsDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" checkinTime="1506095869158" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A. &amp; Dowling, Ashley P. G." docDate="2017" docId="02900DCF4969DD929E148EA2E89B4BF7" docLanguage="en" docName="ZooKeys 701: 1-496" docOrigin="ZooKeys 701" docSource="http://dx.doi.org/10.3897/zookeys.701.13261" docTitle="Torrenticola solisorta Fisher &amp; Dowling, sp. n." docType="treatment" docUuid="FDDDB599-3669-430E-B96E-09567B324EA4" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="3D440132FFABFFFC1505FFA0FFDDEA09" masterDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" masterLastPageNumber="496" masterPageNumber="1" pageNumber="1" updateTime="1668164828938" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, J. Ray</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Danielle M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skvarla, Michael J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nelson, Whitney A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>701</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>496</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.701.13261</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.701.13261</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-701-1</mods:identifier>
<mods:identifier type="ZBK">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
<mods:identifier type="ZooBank">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="134839650" LSID="urn:lsid:zoobank.org:act:FDDDB599-3669-430E-B96E-09567B324EA4" httpUri="http://treatment.plazi.org/id/02900DCF4969DD929E148EA2E89B4BF7" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/FDDDB599-3669-430E-B96E-09567B324EA4" authority="Fisher &amp; Dowling" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola solisorta" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solisorta">Torrenticola solisorta Fisher &amp; Dowling</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">HOLOTYPE (♀): from USA, Arkansas, Polk County, East Saline Creek, beside Forest Road 38, north of Shady Lake Recreation Area, 30 Jul 2011, by IM Smith, IMS 110041, DNA 1300.</paragraph>
<paragraph pageId="0" pageNumber="1">
PARATYPES (4 ♀; 6 ♂): Arkansas, USA: 2 ♂ from Montgomery County, Gaston, South Fork Ouachita River, 29 Jul 2011, by IM Smith, IMS110040
<normalizedToken originalValue="•">*</normalizedToken>
1 ♂ (ALLOTYPE) from Montgomery County, Ouachita National Forest, South Fork Ouachita River, 29 Jul 2011, by AJ Radwell, &amp; B Crump, AJR110302
<normalizedToken originalValue="•">*</normalizedToken>
4 ♀ and 3 ♂ from Polk County, East Saline Creek, beside Forest Road 38, north of Shady Lake Recreation Area, 30 Jul 2011, by IM Smith, IMS 110041.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type deposition">
<paragraph pageId="0" pageNumber="1">Type deposition.</paragraph>
<paragraph pageId="0" pageNumber="1">Holotype (♀), allotype (♂), and most paratypes (2 ♀; 4 ♂) deposited in the CNC; other paratypes (2 ♀; 2 ♂) deposited in ACUA.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola solisorta" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solisorta">Torrenticola solisorta</taxonomicName>
are similar to other members of the Nigroalba Group (
<taxonomicName lsidName="T. flangipalpa" pageId="0" pageNumber="1" rank="species" species="flangipalpa">T. flangipalpa</taxonomicName>
,
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
, and
<taxonomicName lsidName="T. dentirostra" pageId="0" pageNumber="1" rank="species" species="dentirostra">T. dentirostra</taxonomicName>
) in being small, slightly elongate, having purple dorsal coloration restricted posteriorly, and having distinct yet poorly-defined hind coxal margins.
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. flangipalpa" pageId="0" pageNumber="1" rank="species" species="flangipalpa">T. flangipalpa</taxonomicName>
by having tuberculate pedipalp femoral extension (flange-like and anteriorly-directed in
<taxonomicName lsidName="T. flangipalpa" pageId="0" pageNumber="1" rank="species" species="flangipalpa">T. flangipalpa</taxonomicName>
); a shorter anterior venter (192-225 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 235-265 in
<taxonomicName lsidName="T. flangipalpa" pageId="0" pageNumber="1" rank="species" species="flangipalpa">T. flangipalpa</taxonomicName>
); and more elongate pedipalp tibia (length/ width ♀ = 5.67-5.82 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 4.79-5.00 in
<taxonomicName lsidName="T. flangipalpa" pageId="0" pageNumber="1" rank="species" species="flangipalpa">T. flangipalpa</taxonomicName>
; ♂ = 5.08-5.17 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 4.40-4.86 in
<taxonomicName lsidName="T. flangipalpa" pageId="0" pageNumber="1" rank="species" species="flangipalpa">T. flangipalpa</taxonomicName>
).
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. dentirostra" pageId="0" pageNumber="1" rank="species" species="dentirostra">T. dentirostra</taxonomicName>
by having a smooth rostrum (
<taxonomicName lsidName="T. dentirostra" pageId="0" pageNumber="1" rank="species" species="dentirostra">T. dentirostra</taxonomicName>
has a dentate bump midway on the dorsal edge of the rostrum) and more elongate pedipalpal tibiae (length/width ♀ = 5.6-5.9 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 4.5-5.0 in
<taxonomicName lsidName="T. dentirostra" pageId="0" pageNumber="1" rank="species" species="dentirostra">T. dentirostra</taxonomicName>
; ♂ = 5.0-5.2 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 4.5-4.7 in
<taxonomicName lsidName="T. dentirostra" pageId="0" pageNumber="1" rank="species" species="dentirostra">T. dentirostra</taxonomicName>
).
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
by having orangish coloration immediately anterior to the purple dorsal coloration. Additionally, female
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
can be differentiated from female
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
by having a slightly shorter dorsum (475-500 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 500-530 in
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
); a thicker gnathosomal bay (length/width = 1.3-1.5 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 1.25-1.55 in
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
); and a slightly thinner subcapitulum (3.14-3.30 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 3.00-3.14 in
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
). Male
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
also can be differentiated from
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
by anterior venter/medial suture (2.87-3.26 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 2.54-2.77 in
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
) and having a wider dorsum (305-320 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 290-300 in
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
). Other
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Torrenticola</taxonomicName>
with purple dorsal coloration restricted posteriorly can be confused with
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, such as
<taxonomicName lsidName="T. tahoei" pageId="0" pageNumber="1" rank="species" species="tahoei">T. tahoei</taxonomicName>
and
<taxonomicName lsidName="T. oregonensis" pageId="0" pageNumber="1" rank="species" species="oregonensis">T. oregonensis</taxonomicName>
. Both of these species are larger (dorsum length ♀ = 475-500 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 600-840 in others; ♂ = 425-460 in
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
, 560-820 in others) and distributed in the west (
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
is known only from the Ouachita Mountains in Arkansas).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female (Figure 247) (n = 5) (holotype measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsum - (475-500 (500) long; 325-360 (350) wide) ovoid with bluish-purple to purple coloration restricted posteriorly with orange immediately anterior and fading anteriorly. Anterio-medial platelets (102.5-112.5 (112.5) long; 42.5-46.25 (45) wide). Anterio-lateral platelets (138.75-150 (147.5) long; 47.5-50 (47.5) wide) free from dorsal plate. Dgl-4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 220-240 (240)). Dorsal plate proportions: dorsum length/width 1.38-1.46 (1.43); dorsal width/distance between Dgl-4 1.45-1.50 (1.46); anterio-medial platelet length/width 2.22-2.53 (2.50); anterio-lateral platelet length/width 2.78-3.16 (3.11); anterio-lateral/anterio-medial length 1.31-1.43 (1.31).</paragraph>
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (280-300 (300) long (ventral); 199-217 (217) long (dorsal); 85-92.5 (92.5) tall) elongate and colorless. Rostrum (105-110 (110) long; 35-37.5 (36.25) wide). Chelicerae (256-281 (278) long) with curved fangs (34-44 (43) long). Subcapitular proportions: ventral length/height 3.14-3.29 (3.24); rostrum length/width 2.87-3.03 (3.03). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua ending broadly and dentate. Palpomeres: trochanter (27.5-30 (30) long); femur (86.25-93.75 (93.75) long); genu (52.5-55 (55) long); tibia (80-85 (85) long; 13.75-15 (15) wide); tarsus (12.5-15 (12.5) long). Palpomere proportions: femur/genu 1.64-1.70 (1.70); tibia/femur 0.91-0.99 (0.91); tibia length/width 5.67-5.82 (5.67).</paragraph>
<paragraph pageId="0" pageNumber="1">Venter - (580-640 (630) long; 356-439 (380) wide) with bluish-purple or purple coloration. Gnathosomal bay (97.5-110 (110) long; 70-77.5 (77.5) wide). Cxgl-4 far from apex. Medial suture (47.5-50 (47.5) long). Genital plates (132.5-140 (140) long; 117.5-122.5 (122.5) wide). Additional measurements: Cx-1 (228-249 (249) long (total); 88-121 (121) long (medial)); Cx-3 (236.5-259.75 (252) wide); anterior venter (192.5-207.5 (205) long). Ventral proportions: gnathosomal bay length/width 1.30-1.50 (1.42); anterior venter/genital field length 1.45-1.54 (1.46); anterior venter length/genital field width 1.60-1.77 (1.67); anterior venter/medial suture 3.85-4.37 (4.32).</paragraph>
<paragraph pageId="0" pageNumber="1">Male (Figure 248) (n = 6) (allotypic measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsum - (425-460 (435) long; 305-320 (305) wide) ovoid with bluish-purple to purple coloration restricted posteriorly with orange immediately anterior and fading anteriorly. Anterio-medial platelets (97.5-105 (100) long; 40-45 (42.5) wide). Anterio-lateral platelets (130-145 (145) long; 45-50 (45) wide) free from dorsal plate. Dgl-4 closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 205-220 (215)). Dorsal plate proportions: dorsum length/width 1.37-1.48 (1.43); dorsal width/distance between Dgl-4 1.42-1.51 (1.42); anterio-medial platelet length/width 2.33-2.44 (2.35); anterio-lateral platelet length/width 2.85-3.22 (3.22); anterio-lateral/anterio-medial length 1.32-1.45 (1.45).</paragraph>
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (252.5-265 (255) long (ventral); 181-196 (190) long (dorsal); 80-87.5 (80) tall) elongate and colorless. Rostrum (92.5-100 (100) long; 32.5-35 (35) wide). Chelicerae (240-246 (240) long) with curved fangs (33-45 (45) long). Subcapitular proportions: ventral length/height 2.97-3.19 (3.19); rostrum length/width 2.81-2.92 (2.86). Pedipalps elongate (especially tibiae) with tuberculate ventral extensions on femora and genua ending broadly and dentate. Palpomeres: trochanter (27.5-30 (27.5) long); femur (80-82.5 (80) long); genu (48.75-51.25 (48.75) long); tibia (77.5-82.5 (77.5) long; 15-16.25 (15) wide); tarsus (12.5-15 (12.5) long). Palpomere proportions: femur/genu 1.59-1.65 (1.64); tibia/femur 0.94-1.00 (0.97); tibia length/width 5.08-5.17 (5.17).</paragraph>
<paragraph pageId="0" pageNumber="1">Venter - (520-570 (545) long; 335-382 (335) wide) with bluish-purple or purple coloration. Gnathosomal bay (85-107.5 (102.5) long; 60-72.5 (67.5) wide). Cxgl-4 far from apex. Medial suture (65-77.5 (75) long). Genital plates (105-115 (105) long; 90-95 (92.5) wide). Additional measurements: Cx-1 (208-231 (230) long (total); 91-130 (130) long (medial)); Cx-3 (240-281 (240) wide); anterior venter (210-225 (215) long). Ventral proportions: gnathosomal bay length/width 1.17-1.52 (1.52); anterior venter/genital field length 1.87-2.10 (2.05); anterior venter length/genital field width 2.27-2.50 (2.32); anterior venter/medial suture 2.87-3.26 (2.87).</paragraph>
<paragraph pageId="0" pageNumber="1">Immatures unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Specific epithet (
<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea solisorta" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solisorta">solisorta</taxonomicName>
) refers to the dorsal coloration of this species, which is the easiest way to differentiate it from its sister species (
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
). If one imagines the posterior purple color as the landscape and horizon, then the orangish coloration resembles the sun beginning to rise (solis ortus, L. sunrise).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Ouachita Mountains (Figure 246).</paragraph>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 246.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola solisorta" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solisorta">Torrenticola solisorta</taxonomicName>
sp. n. distribution.
</paragraph>
</caption>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 247.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola solisorta" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solisorta">Torrenticola solisorta</taxonomicName>
sp. n. female: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 248.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola solisorta" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solisorta">Torrenticola solisorta</taxonomicName>
sp. n. male: A dorsal plates, note the faint orange coloration, which is a rare character state (most specimens are much brighter), not a male character B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola solisorta" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solisorta">Torrenticola solisorta</taxonomicName>
groups with other members of the Raptor Complex with high support in all analyses and specimens are less than 1% different in COI sequence from each other. In all analyses,
<taxonomicName lsidName="T. solisorta" pageId="0" pageNumber="1" rank="species" species="solisorta">T. solisorta</taxonomicName>
groups with two other morphologically similar species:
<taxonomicName lsidName="T. flangipalpa" pageId="0" pageNumber="1" rank="species" species="flangipalpa">T. flangipalpa</taxonomicName>
and
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola solisorta" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solisorta">Torrenticola solisorta</taxonomicName>
is greater than 4% different in COI from its sister species (
<taxonomicName lsidName="T. nigroalba" pageId="0" pageNumber="1" rank="species" species="nigroalba">T. nigroalba</taxonomicName>
). That clade of three species corresponds to an identification group, the Nigroalba Group, the members of which are easily differentiated by their size, coloration, long medial suture in females, and overall appearance.
</paragraph>
<paragraph pageId="0" pageNumber="1">This species hypothesis is supported by low COI divergence within the species (0-2%) and high divergence between species (3-15%), and by the morphological characters outlined in the diagnosis.</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="D76B8B4FAC36441EBBCFEB05D8ABCE2C" pageNumber="1571" type="nomenclature">
<paragraph id="64B32C2366BE3A24910D2889AA74C3AD" pageNumber="1571">
<taxonomicName id="546BA1CCBFB7805A028F0AD1133C3DC5" ID-CoL="9FZP" authority="(Waterhouse 1841)" baseAuthorityName="Waterhouse" baseAuthorityYear="1841" class="Mammalia" family="Octodontidae" genus="Aconaemys" kingdom="Animalia" order="Rodentia" pageNumber="1571" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="51A5AD7C9684832B4BED91BFB3D69001" italics="true" pageNumber="1571">Aconaemys fuscus</emphasis>
(Waterhouse 1841)
</taxonomicName>
</paragraph>
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<subSubSection id="2D0ABA62C8828CD5751E936684DB8EF2" pageNumber="1571" type="reference_group">
<paragraph id="9898740D3DE65DBBE841F90C92D7B652" pageNumber="1571">
<treatmentCitation id="A840C5880FA7C2ADB2F87D468511B510" author="Waterhouse" authority="Waterhouse 1842" authorityName="Waterhouse" authorityYear="1842" class="Mammalia" family="Octodontidae" genus="Schizodon" journal="Proc. Zool. Soc. Lond." kingdom="Animalia" order="Rodentia" page="91" pageNumber="1571" phylum="Chordata" rank="species" species="fuscus" title="Schizodon fuscus" volume="1841" year="1841">
<taxonomicName id="985E74D25F4108E09F5FBC588449D3E2" ID-CoL="C4HLK" authority="Waterhouse 1842" authorityName="Waterhouse" authorityYear="1842" class="Mammalia" family="Octodontidae" genus="Schizodon" kingdom="Animalia" order="Rodentia" pageNumber="1571" phylum="Chordata" rank="species" species="fuscus">
<emphasis id="55A4E2A13BC23787F95D9BCCBB787FA7" italics="true" pageNumber="1571">[Schizodon] fuscus</emphasis>
<bibRefCitation id="A4F0169C55A4A2F65933F39C9332AA1E" author="Waterhouse, G. R." refId="ref383282" refString="Waterhouse, G. R. 1842 [1843]. Descriptions of a new species of quadrupeds collected by Mr. Fraser at Fernando Po. Proceedings of the Zoological Society of London, 1842: 124 - 130." year="1842">Waterhouse 1842</bibRefCitation>
</taxonomicName>
,
<bibCitation id="7692F0C0A268F8660D5843B05DDA7DEE" author="Waterhouse" journal="Proc. Zool. Soc. Lond." pageNumber="1571" pagination="91" title="Schizodon fuscus" volume="1841" year="1841">Proc. Zool. Soc. Lond., 1841: 91</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CD860F25809611ECCAC91D0D610732F6" pageNumber="1571" type="materials_examined">
<paragraph id="ABC6453FB959F1F271D17CD3A092DAC6" pageNumber="1571">
<emphasis id="486374DE8AD93E17988F0FD58CF61903" inLineHeading="true" pageNumber="1571">Type Locality:</emphasis>
<materialsCitation id="751F5D14EE499B9B6C64D09192D79801" ID-GBIF-Occurrence="4867356305" country="Argentina" pageNumber="1571" stateProvince="Mendoza">
<collectingCountry id="A1FD0EF2BFD84425486F9AED5168CC00" name="Argentina">Argentina</collectingCountry>
,
<collectingRegion id="DBF7A9087BAE32F20CE4A8CB2CB4D7A1" country="Argentina" name="Mendoza">Mendoza Prov.</collectingRegion>
, Valle de las Cuevas; see comments.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9CE904F80756383E8ED767B1F0F990F4" pageNumber="1571" type="vernacular_names">
<paragraph id="4CF4D3C36D17B0A7C9E7E94A072AE751" pageNumber="1571">
<emphasis id="11471EED78F745F7A426582C6EAFEAC1" inLineHeading="true" pageNumber="1571">Vernacular Names:</emphasis>
<vernacularName id="BE4CA22A322B11A2F0EB77BFFAE5243F" language="eng" pageNumber="1571">Chilean Rock Rat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="58BE287FDEAB0BAAD5663C451729CC34" pageNumber="1571" type="distribution">
<paragraph id="ADF2A8CEBCAF44C25F72987FD53AA8B7" pageNumber="1571">
<emphasis id="56723B90EA3AAB1EB7747F10659F1C3F" inLineHeading="true" pageNumber="1571">Distribution:</emphasis>
High Andes of
<collectingCountry id="BF909B95CDED3FD68D104F09DDC3D317" name="Chile">Chile</collectingCountry>
and
<collectingCountry id="7E92C2738EAC6F03F7476620B7C417CD" name="Argentina">Argentina</collectingCountry>
(between 33
<sup id="4D489A920A300DD32A97AD09A1D51E6A" pageNumber="1571">o</sup>
and 41
<sup id="7E874EFBC8F5D816C7333A95C63FB1AB" pageNumber="1571">o</sup>
S).
</paragraph>
</subSubSection>
<subSubSection id="AFE64CE8FCA52AC60943FD9E78222A03" pageNumber="1571" type="conservation">
<paragraph id="EE7B7F5FBF4F9004211E81DDBA294213" pageNumber="1571">
<emphasis id="3CF3B2758BD781CAB2E7B8C980378D2E" inLineHeading="true" pageNumber="1571">Conservation:</emphasis>
<collectionCode id="B10139F238469EF1A5E47BCC2CE16838">IUCN</collectionCode>
– Lower Risk (lc). Locally abundant.
</paragraph>
</subSubSection>
<subSubSection id="95C75CD9185761CA89668B3329884E38" pageNumber="1571" type="discussion">
<paragraph id="754F0528632E4DFB9B37259F46621C77" pageNumber="1571">
<emphasis id="07C362B72A16729CED8C2F9CB4D608A8" inLineHeading="true" pageNumber="1571">Discussion:</emphasis>
Waterhouse never designated a type specimen, which was finally designated by
<bibRefCitation id="6DBC10989BD23FF478B00B01F036046E" author="Thomas, O." refId="ref354263" refString="Thomas, O. 1917 a. A new species of Aconaemys from southern Chile. Annals and Magazine of Natural History, ser. 8, 19: 281 - 282." year="1917">
Thomas (1917
<emphasis id="C772E4C2CCE317A753E71DAB4E5B25F9" italics="true" pageNumber="1571">a</emphasis>
)
</bibRefCitation>
. Waterhouse’s original type locality was &quot;from
<collectingCountry id="4DDEC4996E4AC115484656197849AB5E" name="Chile">Chile</collectingCountry>
&quot;, but in 1848 he added that the locality was from the Valle de las Cuevas near the Volcano of Peteroa, which would make the locality in
<collectingCountry id="C6C9BE57F7573498FCF22A41532158FA" name="Argentina">Argentina</collectingCountry>
(
<bibRefCitation id="CB9F7BDC6E738D96DBFF562BEDC7D9EA" author="Pearson, O. P." refId="ref278495" refString="Pearson, O. P. 1984. Taxonomy and natural history of some fossorial rodents of Patagonia, southern Argentina. Journal of Zoology, London, 202: 225 - 237." year="1984">Pearson, 1984</bibRefCitation>
). Karyotype has 2n=56 and FN=108 (
<bibRefCitation id="F9C14187D99B72E13111C487D8984467" author="Gallardo, M. H. &amp; D. Reise" refId="ref111470" refString="Gallardo, M. H., and D. Reise. 1992. Systematics of Aconaemys (Rodentia, Octodontidae). Journal of Mammalogy, 73 (4): 779 - 788." year="1992">Gallardo and Reise, 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>