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<figureCitation id="131C2A1FFF85B148FD53F917FCA81823" box="[739,828,1788,1815]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="45" pageNumber="46">Figs 5F</figureCitation>
<figureCitation id="131C2A1FFF85B148FD53F917FCA81823" box="[739,828,1788,1815]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="45" pageNumber="46">Figs 5F</figureCitation>
,
<figureCitation id="131C2A1FFF85B148FCF8F917FCF71822" box="[840,867,1788,1814]" captionStart="Fig" captionStartId="51.[227,270,1900,1926]" captionTargetBox="[291,1360,302,1875]" captionTargetId="figure@51.[291,1361,302,1875]" captionTargetPageId="51" captionText="Fig. 13. Male representatives of three subfamilies. A, C, E. Frontal view. B, D, F. Lateral view. — AB. Tatuidris tatusia, Agroecomyrmecinae (Panama, CASENT0178870, E. Prado). CD. Paraponera clavata, Paraponerinae (Guyana, CASENT0902407, R. Perry). EF. Pseudoponera stigma, Ponerinae (Paraguay, CASENT0178182, A. Nobile). Scale bars: A = 0.1 mm, B = 0.5 mm, C, F = 1.0 mm, D = 2.0 mm, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3780174" httpUri="https://zenodo.org/record/3780174/files/figure.png" pageId="45" pageNumber="46">13</figureCitation>
AB

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@ -52,7 +52,7 @@ Lepeletier de Saint-Fargeau, 1835
<paragraph id="8B98369AFF86B14BFDDAFB1AFB811A3F" blockId="46.[497,1166,1229,1292]" box="[618,1045,1265,1292]" pageId="46" pageNumber="47">
<figureCitation id="131C2A1FFF86B14BFDDAFB1AFD5C1A3F" box="[618,712,1265,1291]" captionStart="Fig" captionStartId="17.[227,270,1790,1816]" captionTargetBox="[349,1305,302,1757]" captionTargetId="figure@17.[348,1305,302,1757]" captionTargetPageId="17" captionText="Fig. 4. Male morphology. AC. Abdominal sternum IX in ventral view. B. Oblique. DE. Petiole in lateral view. F. Forewing in dorsal view. GH. Propodeum in lateral view. — A. Aenictogiton indet. (Zambia, CASENT0106126, M. Branstetter). B. Cerapachys “parasyscia” lineage (Kenya, B. Boudinot). C. Emeryopone buttelreepeni (Thailand, CASENT0278779, B. Boudinot). D. Paraponera clavata (?Panama, B. Boudinot). E. Cerapachys lividus (Madagascar, CASENT0138502, D. Raharinjanahary). F. Phaulomyrma indet. (Thailand, UCRENT150358, A. Nobile). G. Leptanillinae indet. (Thailand, CASENT0156249, B. Boudinot); arrow indicates dorsal margin of petiolar foramen. H. Adelomyrmex dentivagans; arrow indicates propodeal lobe. All scale bars = 0.2 mm, except DE = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3780158" httpUri="https://zenodo.org/record/3780158/files/figure.png" pageId="46" pageNumber="47">Figs 4C</figureCitation>
,
<figureCitation id="131C2A1FFF86B14BFD64FB1AFD6F1A38" box="[724,763,1265,1292]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="46" pageNumber="47">5H</figureCitation>
<figureCitation id="131C2A1FFF86B14BFD64FB1AFD6F1A38" box="[724,763,1265,1292]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="46" pageNumber="47">5H</figureCitation>
,
<figureCitation id="131C2A1FFF86B14BFCB7FB1AFCB81A3F" box="[775,812,1265,1291]" captionStart="Fig" captionStartId="19.[227,270,1755,1781]" captionTargetBox="[340,1306,302,1722]" captionTargetId="figure@19.[340,1306,302,1722]" captionTargetPageId="19" captionText="Fig. 6. Male morphology. AB, EG. Head capsule in frontal view. CD. Mesosoma, lateral view. HI. Petiole, ventrolateral view. — A. Probolomyrmex indet. (Madagascar, CASENT0080551, A. Nobile). B. Anochetus boltoni (Madagascar, CASENT0063847, A. Nobile). C. Leptanilloides gracilis (Guatemala, CASENT0234561, M. Borowiec). D. Odontomachus simillimus (Seychelles, CASENT0172666, A. Nobile). E. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). F. Protanilla indet. (Thailand, CASENT0119776, A. Nobile). G. Stigmatomma indet. (Madagascar, CASENT0007087, E. Prado). H. Nothomyrmecia macrops (Australia, B. Boudinot). I. Neoponera cf. apicalis (Honduras, B. Boudinot). Scale bars: A, FG= 0.1 mm, B = 0.2 mm, C = 0.25 mm, E, HI = 0.5 mm, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3780160" httpUri="https://zenodo.org/record/3780160/files/figure.png" pageId="46" pageNumber="47">6B</figureCitation>
, DE, I, 7D, 13EF
@ -99,7 +99,7 @@ is variable.
<emphasis id="B953EA88FF86B14BFF53F8E9FEFE1828" bold="true" box="[227,362,1794,1820]" pageId="46" pageNumber="47">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8B98369AFF86B14BFF53F8DAFA0818EC" blockId="46.[227,1436,1841,2008]" pageId="46" pageNumber="47">
<paragraph id="8B98369AFF86B14AFF53F8DAFD621E5A" blockId="46.[227,1436,1841,2008]" lastBlockId="47.[227,1436,305,367]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
The
<taxonomicName id="4C274D19FF86B14BFEA7F8DAFE18187F" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[279,396,1841,1867]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subFamily" subFamily="Ponerinae">Ponerinae</taxonomicName>
is global in distribution and has recently been provided a molecular phylogeny (
@ -158,10 +158,7 @@ and
<taxonomicName id="4C274D19FF86B14BFD6FF855FCC018EC" box="[735,852,1982,2008]" class="Insecta" family="Formicidae" genus="Simopelta" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B953EA88FF86B14BFD6FF855FCC018EC" box="[735,852,1982,2008]" italics="true" pageId="46" pageNumber="47">Simopelta</emphasis>
</taxonomicName>
will be described in forthcoming publications (B.
</paragraph>
<paragraph id="8B98369AFF87B14AFF53FEDAFD621E5A" blockId="47.[227,1436,305,367]" pageId="47" pageNumber="48">
Boudinot in prep. and R.S. Probst
will be described in forthcoming publications (B. Boudinot in prep. and R.S. Probst
<emphasis id="B953EA88FF87B14AFDDFFED9FD351E7F" box="[623,673,305,331]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. in prep.). The male of
<taxonomicName id="4C274D19FF87B14AFC1FFEDAFBD11E7F" authorityName="Brown" authorityYear="1974" box="[943,1093,305,331]" class="Insecta" family="Formicidae" genus="Dolioponera" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">

View file

@ -1,42 +1,42 @@
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@ -54,7 +54,7 @@ Emery, 1901
,
<figureCitation id="131C2A1FFF86B14BFD47FD78FC8F1D99" box="[759,795,659,685]" captionStart="Fig" captionStartId="7.[227,270,1968,1994]" captionTargetBox="[322,1387,333,1918]" captionTargetId="figure@7.[227,1481,302,1946]" captionTargetPageId="7" captionText="Fig. 2. Pterothoracic venter morphology of representative hymenopterans. A. Tenthredinidae, female. B. Polistes (Vespidae), worker. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm. Abbreviations: see Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.3780156" httpUri="https://zenodo.org/record/3780156/files/figure.png" pageId="46" pageNumber="47">2C</figureCitation>
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<figureCitation id="131C2A1FFF86B14BFC97FD78FCD81D99" box="[807,844,659,685]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm." pageId="46" pageNumber="47">3C</figureCitation>
<figureCitation id="131C2A1FFF86B14BFC97FD78FCD81D99" box="[807,844,659,685]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm. Abbreviations: see Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.14900271" httpUri="https://zenodo.org/record/14900271/files/figure.png" pageId="46" pageNumber="47">3C</figureCitation>
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<figureCitation id="131C2A1FFF86B14BFCE8FD78FCE81D99" box="[856,892,659,685]" captionStart="Fig" captionStartId="17.[227,270,1790,1816]" captionTargetBox="[349,1305,302,1757]" captionTargetId="figure@17.[348,1305,302,1757]" captionTargetPageId="17" captionText="Fig. 4. Male morphology. AC. Abdominal sternum IX in ventral view. B. Oblique. DE. Petiole in lateral view. F. Forewing in dorsal view. GH. Propodeum in lateral view. — A. Aenictogiton indet. (Zambia, CASENT0106126, M. Branstetter). B. Cerapachys “parasyscia” lineage (Kenya, B. Boudinot). C. Emeryopone buttelreepeni (Thailand, CASENT0278779, B. Boudinot). D. Paraponera clavata (?Panama, B. Boudinot). E. Cerapachys lividus (Madagascar, CASENT0138502, D. Raharinjanahary). F. Phaulomyrma indet. (Thailand, UCRENT150358, A. Nobile). G. Leptanillinae indet. (Thailand, CASENT0156249, B. Boudinot); arrow indicates dorsal margin of petiolar foramen. H. Adelomyrmex dentivagans; arrow indicates propodeal lobe. All scale bars = 0.2 mm, except DE = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3780158" httpUri="https://zenodo.org/record/3780158/files/figure.png" pageId="46" pageNumber="47">4D</figureCitation>
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@ -1,42 +1,42 @@
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@ -50,7 +50,7 @@ Leach, 1815
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<figureCitation id="131C2A1FFF87B14AFD15F926FD7819DC" box="[677,748,1741,1768]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="47" pageNumber="48">Figs 5</figureCitation>
<figureCitation id="131C2A1FFF87B14AFD15F926FD7819DC" box="[677,748,1741,1768]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="47" pageNumber="48">Figs 5</figureCitation>
AB, E, 6C, 14CD
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Subfamily
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@ -46,14 +46,16 @@ Subfamily
M.R. Smith, 1952
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<figureCitation id="131C2A1FFF98B155FD7FFAF3FCDA1A07" box="[719,846,1304,1331]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="48" pageNumber="49">Figs 5B, E</figureCitation>
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<figureCitation id="131C2A1FFF98B155FD7FFAF3FCDA1A07" box="[719,846,1304,1331]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="48" pageNumber="49">Figs 5B, E</figureCitation>
,
<figureCitation id="131C2A1FFF98B155FCEBFAF3FCE21A07" box="[859,886,1304,1331]" captionStart="Fig" captionStartId="53.[227,270,1898,1924]" captionTargetBox="[227,1436,510,1849]" captionTargetId="figure@53.[227,1436,510,1849]" captionTargetPageId="53" captionText="Fig. 15. Male representatives of three subfamilies. A, D. Frontal view. BC, E. Lateral view. — AB. Pseudomyrmex holmgreni, Pseudomyrmecinae (Paraguay, CASENT0173758, A. Nobile). C. Aneuretus simoni, Aneuretinae, used with permission from Wilson et al. (1956). DE. Technomyrmex difficilis, Dolichoderinae (Madagascar, CASENT0049968, A. Nobile). Scale bars: A, D = 0.2 mm, B = 1.0 mm, E = 0.5 mm, no scale available for C." figureDoi="http://doi.org/10.5281/zenodo.3780178" httpUri="https://zenodo.org/record/3780178/files/figure.png" pageId="48" pageNumber="49">15</figureCitation>
AB
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@ -50,7 +50,7 @@ Emery, 1877
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<figureCitation id="131C2A1FFF98B155FD08FD36FD6B1DCC" box="[696,767,733,760]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="48" pageNumber="49">Figs 5</figureCitation>
<figureCitation id="131C2A1FFF98B155FD08FD36FD6B1DCC" box="[696,767,733,760]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="48" pageNumber="49">Figs 5</figureCitation>
CD, 6H, 14EF
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@ -50,7 +50,7 @@ Latreille, 1809
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,
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@ -52,7 +52,7 @@ Lepeletier de Saint-Fargeau, 1835
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<figureCitation id="131C2A1FFF9FB152FD05FDC9FC811D08" box="[693,789,546,572]" captionStart="Fig" captionStartId="17.[227,270,1790,1816]" captionTargetBox="[349,1305,302,1757]" captionTargetId="figure@17.[348,1305,302,1757]" captionTargetPageId="17" captionText="Fig. 4. Male morphology. AC. Abdominal sternum IX in ventral view. B. Oblique. DE. Petiole in lateral view. F. Forewing in dorsal view. GH. Propodeum in lateral view. — A. Aenictogiton indet. (Zambia, CASENT0106126, M. Branstetter). B. Cerapachys “parasyscia” lineage (Kenya, B. Boudinot). C. Emeryopone buttelreepeni (Thailand, CASENT0278779, B. Boudinot). D. Paraponera clavata (?Panama, B. Boudinot). E. Cerapachys lividus (Madagascar, CASENT0138502, D. Raharinjanahary). F. Phaulomyrma indet. (Thailand, UCRENT150358, A. Nobile). G. Leptanillinae indet. (Thailand, CASENT0156249, B. Boudinot); arrow indicates dorsal margin of petiolar foramen. H. Adelomyrmex dentivagans; arrow indicates propodeal lobe. All scale bars = 0.2 mm, except DE = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3780158" httpUri="https://zenodo.org/record/3780158/files/figure.png" pageId="55" pageNumber="56">Figs 4H</figureCitation>
,
<figureCitation id="131C2A1FFF9FB152FC91FDC9FCFF1D08" box="[801,875,546,573]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="55" pageNumber="56">5A, G</figureCitation>
<figureCitation id="131C2A1FFF9FB152FC91FDC9FCFF1D08" box="[801,875,546,573]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="55" pageNumber="56">5A, G</figureCitation>
,
<figureCitation id="131C2A1FFF9FB152FCC7FDC9FC001D08" box="[887,916,546,572]" captionStart="Fig" captionStartId="54.[227,270,1932,1958]" captionTargetBox="[227,1436,334,1894]" captionTargetId="figure@54.[227,1436,333,1895]" captionTargetPageId="54" captionText="Fig. 16. Male representatives of three subfamilies. AB. Formica wheeleri, Formicinae (U.S.A., CASENT0173024, A. Nobile). CD. Rhytidoponera, Ectatomminae, ectaheteromorph clade (Australia, CASENT0004610, A. Nobile). EF. Pogonomyrmex rastratus (Argentina, CASENT0172673, A." figureDoi="http://doi.org/10.5281/zenodo.3780180" httpUri="https://zenodo.org/record/3780180/files/figure.png" pageId="55" pageNumber="56">16</figureCitation>
DE

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@ -66,12 +66,12 @@ with the following apomorphies:
<paragraph id="8B98369AFFA3B16EFF53FA00FB5B1931" blockId="11.[227,1436,1374,2034]" box="[227,1231,1515,1542]" pageId="11" pageNumber="12">5. Antenna geniculate between long scape and funiculus (worker, gyne) (notes 1, 3).</paragraph>
<paragraph id="8B98369AFFA3B16EFF53F9E5FA08197F" blockId="11.[227,1436,1374,2034]" pageId="11" pageNumber="12">
6. Disticoxal foramen directed laterally and completely enclosing protrochanteral base, including protrochanteral condyles, such that all disticoxal membrane concealed (all castes,
<figureCitation id="131C2A1FFFA3B16EFB60F9DAFABD197F" box="[1232,1321,1585,1611]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm." pageId="11" pageNumber="12">Fig. 3C</figureCitation>
<figureCitation id="131C2A1FFFA3B16EFB60F9DAFABD197F" box="[1232,1321,1585,1611]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm. Abbreviations: see Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.14900271" httpUri="https://zenodo.org/record/14900271/files/figure.png" pageId="11" pageNumber="12">Fig. 3C</figureCitation>
) (note 4).
</paragraph>
<paragraph id="8B98369AFFA3B16EFF53F9BEFC7219A7" blockId="11.[227,1436,1374,2034]" pageId="11" pageNumber="12">
7. All meso- and metacoxal cavities small, circular, monocondylic, ventrally-directed, and disticoxae strongly produced laterally (all adult castes,
<figureCitation id="131C2A1FFFA3B16EFCA9F993FCE519A6" box="[793,881,1656,1682]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm." pageId="11" pageNumber="12">Fig. 3C</figureCitation>
<figureCitation id="131C2A1FFFA3B16EFCA9F993FCE519A6" box="[793,881,1656,1682]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm. Abbreviations: see Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.14900271" httpUri="https://zenodo.org/record/14900271/files/figure.png" pageId="11" pageNumber="12">Fig. 3C</figureCitation>
) (note 5).
</paragraph>
<paragraph id="8B98369AFFA3B16EFF53F970FC091982" blockId="11.[227,1436,1374,2034]" box="[227,925,1691,1718]" pageId="11" pageNumber="12">8. Metapleural gland present (adult castes, but see note 6).</paragraph>
@ -127,13 +127,13 @@ is to release large quantities of sexuals. The wording has been specifically rep
4. The procoxa of
<taxonomicName id="4C274D19FFA4B169FE74FC53FDDE1CE6" authorityName="Latreille" authorityYear="1809" box="[452,586,952,978]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
is characteristically modified. The trochanteral foramen (situated apically on the procoxa) is directed laterally and entirely enclosed, revealing no membrane in undamaged specimens (
<figureCitation id="131C2A1FFFA4B169FE2BFC15FE6C1B2D" box="[411,504,1022,1049]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm." pageId="12" pageNumber="13">Fig. 3C</figureCitation>
<figureCitation id="131C2A1FFFA4B169FE2BFC15FE6C1B2D" box="[411,504,1022,1049]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm. Abbreviations: see Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.14900271" httpUri="https://zenodo.org/record/14900271/files/figure.png" pageId="12" pageNumber="13">Fig. 3C</figureCitation>
, left column, top row). Medially, the foramen is closed by an unfused seam of the anterior and posterior apical coxal lobes, which completely surround the anterior trochanteral process. The axis of coxal-trochanteral articulation, rather than being lateromedial as in Symphyta (
<figureCitation id="131C2A1FFFA4B169FEA9FB83FEE21BB6" box="[281,374,1128,1154]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm." pageId="12" pageNumber="13">Fig. 3A</figureCitation>
<figureCitation id="131C2A1FFFA4B169FEA9FB83FEE21BB6" box="[281,374,1128,1154]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm. Abbreviations: see Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.14900271" httpUri="https://zenodo.org/record/14900271/files/figure.png" pageId="12" pageNumber="13">Fig. 3A</figureCitation>
), or rotated obliquely as in many
<taxonomicName id="4C274D19FFA4B169FD4AFB83FCF61BB6" authorityName="Latreille" authorityYear="1802" box="[762,866,1128,1154]" class="Insecta" higherTaxonomySource="GBIF,CoL" infraOrder="Aculeata" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="infraOrder">Aculeata</taxonomicName>
(
<figureCitation id="131C2A1FFFA4B169FCC1FB83FC581BB6" box="[881,972,1128,1154]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm." pageId="12" pageNumber="13">Fig. 3B</figureCitation>
<figureCitation id="131C2A1FFFA4B169FCC1FB83FC581BB6" box="[881,972,1128,1154]" captionStart="Fig" captionStartId="8.[227,269,1899,1925]" captionTargetBox="[349,1285,302,1860]" captionTargetId="figure@8.[317,1335,302,1869]" captionTargetPageId="8" captionText="Fig. 3. Coxal morphology of representative hymenopterans: right procoxae, left column; right mesocoxae, middle column; right metacoxae, right column; top procoxa lateral view, bottom procoxa medial view; top meso- and metacoxae anterior view, bottom meso- and metacoxae posterior view. A. Tenthredinidae. B. Polistes sp., Vespidae. C. Paraponera clavata gyne (Paraponerinae, Formicidae). Scale bars = 1.0 mm. Abbreviations: see Material and Methods." figureDoi="http://doi.org/10.5281/zenodo.14900271" httpUri="https://zenodo.org/record/14900271/files/figure.png" pageId="12" pageNumber="13">Fig. 3B</figureCitation>
), is almost entirely anteroposterior. Leg adduction and abduction occurs along this anteroposterior axis in more-or-less one plane of motion, with the trochanter rotating within the closed disticoxal foramen. The coxae and their articulations with the mesosoma and trochanters are poorly studied and show promise for valuable systematic characters. Previous work on hymenopteran coxae include
<bibRefCitation id="EFB64B6BFFA4B169FC0FFB1EFBE31A3B" author="Johnson N. R." box="[959,1143,1269,1295]" pageId="12" pageNumber="13" pagination="870 - 881" refId="ref31538" refString="Johnson N. R. 1988. Midcoxal articulations and phylogeny of the order Hymenoptera. Annals of the Entomological Society of America 81: 870 - 881." type="journal article" year="1988">Johnson (1988)</bibRefCitation>
, which solely focused on the basicoxite and its musculature,
@ -385,7 +385,7 @@ LaPolla
).
</paragraph>
</subSubSection>
<subSubSection id="C33D6511FFA5B168FF53F985FCF519BE" box="[227,865,1645,1674]" pageId="13" pageNumber="14" type="nomenclature">
<subSubSection id="C33D6511FFA5B168FF53F985FCF519BE" box="[227,865,1645,1674]" pageId="13" pageNumber="14" type="multiple">
<paragraph id="8B98369AFFA5B168FF53F985FCF519BE" blockId="13.[227,865,1645,1674]" box="[227,865,1645,1674]" pageId="13" pageNumber="14">
<heading id="D0D081F6FFA5B168FF53F985FCF519BE" box="[227,865,1645,1674]" pageId="13" pageNumber="14">
<emphasis id="B953EA88FFA5B168FF53F985FCF519BE" bold="true" box="[227,865,1645,1674]" italics="true" pageId="13" pageNumber="14">Key to global subfamilies, based on extant males</emphasis>
@ -396,7 +396,7 @@ LaPolla
<paragraph id="8B98369AFFA5B168FF53F949FB6C1812" blockId="13.[227,1437,1698,1830]" pageId="13" pageNumber="14">Notes: It may be expected that some taxa will fail this key, as males of at least 70 genera are undescribed and/or uncollected (see subfamily diagnoses for specific genera). This key treats alate males only, because ergatoid males are identifiable similarly to the workers. Ergatoid males generally bear shorter scapes and more antennomeres than the female, and are known for about a dozen genera.</paragraph>
</subSubSection>
<subSubSection id="C33D6511FFA5B175FF53F8A6FA081CEA" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14" type="key">
<key id="FFB661ABFFA5B16AFF53F8A6FA0818CB" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<key id="FFB661ABFFA5B175FF53F8A6FA081CEA" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<keyStep id="30D32E3FFFA5B168FF53F8A6FA0918BE" pageId="13" pageNumber="14">
<paragraph id="8B98369AFFA5B168FF53F8A6FA091853" blockId="13.[227,1437,1869,1930]" box="[227,1437,1869,1896]" pageId="13" pageNumber="14">
<keyLead id="30D695AFFFA5B168FF53F8A6FA091853" box="[227,1437,1869,1896]" pageId="13" pageNumber="14">
@ -476,7 +476,7 @@ propodeal lobes conspicuous and present (
<paragraph id="8B98369AFFA6B16BFF53FCFFFA091C41" blockId="14.[227,1437,788,1026]" pageId="14" pageNumber="15">
<keyLead id="30D695AFFFA6B16BFF53FCFFFA091C41" pageId="14" pageNumber="15">
4 Abdominal segment III strongly reduced and differentiated from abdominal segment IV (
<figureCitation id="131C2A1FFFA6B16BFADDFCFFFEDE1C66" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5A, B</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFADDFCFFFEDE1C66" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5A, B</figureCitation>
)
<emphasis id="B953EA88FFA6B16BFEEDFCDCFE181C65" bold="true" box="[349,396,823,849]" pageId="14" pageNumber="15">and</emphasis>
antennal toruli separated from anterior clypeal margin by at least one antennal socket diameter
@ -488,7 +488,7 @@ terminal abdominal tergum never produced as spine…………………………
<paragraph id="8B98369AFFA6B16BFF53FC95FA091B35" blockId="14.[227,1437,788,1026]" pageId="14" pageNumber="15">
<keyLead id="30D695AFFFA6B16BFF53FC95FA091B35" pageId="14" pageNumber="15">
Abdominal segment III not reduced relative to abdominal segment IV or somewhat reduced, but not differentiated from abdominal segment IV (
<figureCitation id="131C2A1FFFA6B16BFC38FC4AFC7C1C88" box="[904,1000,929,956]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5C</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFC38FC4AFC7C1C88" box="[904,1000,929,956]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5C</figureCitation>
)
<emphasis id="B953EA88FFA6B16BFC4FFC4AFB8E1C8F" bold="true" box="[1023,1050,929,955]" pageId="14" pageNumber="15">or</emphasis>
antennal toruli separated from anterior clypeal margin by less than one antennal socket diameter
@ -512,7 +512,7 @@ terminal abdominal tergum posteriorly produced as spine……………………
<paragraph id="8B98369AFFA6B16BFF53FB66FA091BFE" blockId="14.[227,1437,1165,1297]" pageId="14" pageNumber="15">
<keyLead id="30D695AFFFA6B16BFF53FB66FA091BFE" pageId="14" pageNumber="15">
6 Jugal lobe present. Frontal carinae usually robust and conspicuous (
<figureCitation id="131C2A1FFFA6B16BFB92FB66FBE91B9C" box="[1058,1149,1165,1192]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5D</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFB92FB66FBE91B9C" box="[1058,1149,1165,1192]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5D</figureCitation>
). Cuticle very thick and usually coarsely sculptured.
<collectingCountry id="F330760AFFA6B16BFDF3FB5BFD3B1BFE" box="[579,687,1200,1226]" name="Australia" pageId="14" pageNumber="15">Australia</collectingCountry>
…………………
@ -531,7 +531,7 @@ terminal abdominal tergum posteriorly produced as spine……………………
<paragraph id="8B98369AFFA6B16BFF53FB3FFA081A25" blockId="14.[227,1437,1165,1297]" pageId="14" pageNumber="15">
<keyLead id="30D695AFFFA6B16BFF53FB3FFA081A25" pageId="14" pageNumber="15">
Jugal lobe absent. Frontal carinae fine, inconspicuous, or absent (
<figureCitation id="131C2A1FFFA6B16BFBBDFB3FFBFD1BDA" box="[1037,1129,1235,1262]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5E</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFBBDFB3FFBFD1BDA" box="[1037,1129,1235,1262]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5E</figureCitation>
). Cuticle thin and usually finely to not sculptured. New World; African, Asian, Australian…………………
<taxonomicName id="4C274D19FFA6B16BFB18FB1CFA081A25" ID-CoL="K88" authorityName="M. R. Smith" authorityYear="1952" box="[1192,1436,1271,1297]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subFamily" subFamily="Pseudomyrmecinae">
<emphasis id="B953EA88FFA6B16BFB18FB1CFA081A25" bold="true" box="[1192,1436,1271,1297]" pageId="14" pageNumber="15">Pseudomyrmecinae</emphasis>
@ -543,9 +543,9 @@ terminal abdominal tergum posteriorly produced as spine……………………
<paragraph id="8B98369AFFA6B16BFF53FAD2FA091936" blockId="14.[227,1437,1336,1750]" pageId="14" pageNumber="15">
<keyLead id="30D695AFFFA6B16BFF53FAD2FA091936" pageId="14" pageNumber="15">
7 Abdominal tergum IV strongly and evenly convex in profile view and much longer than abdominal sternum IV (
<figureCitation id="131C2A1FFFA6B16BFE27FAB0FE7A1A42" box="[407,494,1371,1398]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5F</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFE27FAB0FE7A1A42" box="[407,494,1371,1398]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5F</figureCitation>
, black lines). Helcium supraaxial: Anteroposterior axis of helcium situated well above anteroposterior axis of abdominal segment III postsclerites, such that poststernite with very dorsoventrally tall anterior face relative to anterior face of posttergite (
<figureCitation id="131C2A1FFFA6B16BFBDCFA49FB531A89" box="[1132,1223,1442,1469]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5F</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFBDCFA49FB531A89" box="[1132,1223,1442,1469]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5F</figureCitation>
, dark green lines along anterior faces of abdominal segment III). Spiracle of abdominal tergum IV located in extreme anteroventral corner(within at least five spiracular diameters)………
<taxonomicName id="4C274D19FFA6B16BFBA7FA03FA091936" authority="(Tatuidris)" authorityName="Carpenter" authorityYear="1930" baseAuthorityName="Tatuidris" box="[1047,1437,1512,1539]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subFamily" subFamily="Agroecomyrmecinae">
<emphasis id="B953EA88FFA6B16BFBA7FA03FA091936" bold="true" box="[1047,1437,1512,1539]" pageId="14" pageNumber="15">
@ -559,9 +559,9 @@ Agroecomyrmecinae (
<paragraph id="8B98369AFFA6B16BFF53F9E7FA0819E1" blockId="14.[227,1437,1336,1750]" pageId="14" pageNumber="15">
<keyLead id="30D695AFFFA6B16BFF53F9E7FA0819E1" pageId="14" pageNumber="15">
Abdominal tergum IV weakly or unevenly convex in profile view and about as long as abdominal sternum IV (
<figureCitation id="131C2A1FFFA6B16BFD9FF9C4FD00197E" box="[559,660,1583,1610]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5G</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFD9FF9C4FD00197E" box="[559,660,1583,1610]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5G</figureCitation>
, black lines). Helcium axial: Anteroposterior axis of helcium situated at about midheight of abdominal segment III postsclerites, such that anterior faces of postsclerites roughly equivalent, or anterior face of posttergite somewhat longer than that of poststernite (
<figureCitation id="131C2A1FFFA6B16BFE79F973FDBB1987" box="[457,559,1688,1715]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5G</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFE79F973FDBB1987" box="[457,559,1688,1715]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5G</figureCitation>
, green lines). Spiracle of abdominal tergum IV located distant from anteroventral corner (distant by at least ten spiracular diameters)……………………
<taxonomicName id="4C274D19FFA6B16BFAB3F950FA0819E1" ID-CoL="JY2" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[1283,1436,1723,1749]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="60" phylum="Arthropoda" rank="subFamily" subFamily="Myrmicinae">
<emphasis id="B953EA88FFA6B16BFAB3F950FA0819E1" bold="true" box="[1283,1436,1723,1749]" pageId="14" pageNumber="15">Myrmicinae</emphasis>
@ -573,7 +573,7 @@ Agroecomyrmecinae (
<paragraph id="8B98369AFFA6B16BFF53F916FA09180E" blockId="14.[227,1437,1789,1921]" pageId="14" pageNumber="15">
<keyLead id="30D695AFFFA6B16BFF53F916FA09180E" pageId="14" pageNumber="15">
8 Abdominal segment IV with cinctus (constriction) between the pre- and postsclerite (
<figureCitation id="131C2A1FFFA6B16BFABEF916FAF9182C" box="[1294,1389,1789,1816]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5H</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFABEF916FAF9182C" box="[1294,1389,1789,1816]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5H</figureCitation>
)
<emphasis id="B953EA88FFA6B16BFA31F916FA081823" bold="true" box="[1409,1436,1789,1815]" pageId="14" pageNumber="15">or</emphasis>
jugal lobe present
@ -585,7 +585,7 @@ oblique mesopleural sulcus absent or indistinct……………………………
<paragraph id="8B98369AFFA6B16BFF53F8AFFA0818B4" blockId="14.[227,1437,1789,1921]" pageId="14" pageNumber="15">
<keyLead id="30D695AFFFA6B16BFF53F8AFFA0818B4" pageId="14" pageNumber="15">
Abdominal segment IV without a cinctus (
<figureCitation id="131C2A1FFFA6B16BFCF7F8A8FC31186A" box="[839,933,1859,1886]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="14" pageNumber="15">Fig. 5I</figureCitation>
<figureCitation id="131C2A1FFFA6B16BFCF7F8A8FC31186A" box="[839,933,1859,1886]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="14" pageNumber="15">Fig. 5I</figureCitation>
)
<emphasis id="B953EA88FFA6B16BFC73F8AFFC66186A" bold="true" box="[963,1010,1860,1886]" pageId="14" pageNumber="15">and</emphasis>
jugal lobe absent
@ -690,7 +690,7 @@ part,
<paragraph id="8B98369AFFA7B16AFF53FCBAFA081C85" blockId="15.[227,1437,848,1051]" pageId="15" pageNumber="16">
<keyLead id="30D695AFFFA7B16AFF53FCBAFA081C85" pageId="15" pageNumber="16">
13 Abdominal segment IV without cinctus between pre- and postsclerites (
<figureCitation id="131C2A1FFFA7B16AFB0BFCBBFAB61C5F" box="[1211,1314,848,875]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="15" pageNumber="16">Fig. 5C</figureCitation>
<figureCitation id="131C2A1FFFA7B16AFB0BFCBBFAB61C5F" box="[1211,1314,848,875]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="15" pageNumber="16">Fig. 5C</figureCitation>
). Petiolar tergum and sternum completely fused in anterior third, without visible suture (
<figureCitation id="131C2A1FFFA7B16AFA96FC9FFA1B1CBB" box="[1318,1423,884,911]" captionStart="Fig" captionStartId="19.[227,270,1755,1781]" captionTargetBox="[340,1306,302,1722]" captionTargetId="figure@19.[340,1306,302,1722]" captionTargetPageId="19" captionText="Fig. 6. Male morphology. AB, EG. Head capsule in frontal view. CD. Mesosoma, lateral view. HI. Petiole, ventrolateral view. — A. Probolomyrmex indet. (Madagascar, CASENT0080551, A. Nobile). B. Anochetus boltoni (Madagascar, CASENT0063847, A. Nobile). C. Leptanilloides gracilis (Guatemala, CASENT0234561, M. Borowiec). D. Odontomachus simillimus (Seychelles, CASENT0172666, A. Nobile). E. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). F. Protanilla indet. (Thailand, CASENT0119776, A. Nobile). G. Stigmatomma indet. (Madagascar, CASENT0007087, E. Prado). H. Nothomyrmecia macrops (Australia, B. Boudinot). I. Neoponera cf. apicalis (Honduras, B. Boudinot). Scale bars: A, FG= 0.1 mm, B = 0.2 mm, C = 0.25 mm, E, HI = 0.5 mm, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3780160" httpUri="https://zenodo.org/record/3780160/files/figure.png" pageId="15" pageNumber="16">Fig. 6H</figureCitation>
). Mandibular teeth robust…………
@ -709,7 +709,7 @@ part,
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Abdominal segment III with cinctus between pre- and postsclerites (
<figureCitation id="131C2A1FFFA7B16AFBD5FC51FB5D1CE1" box="[1125,1225,954,981]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A." pageId="15" pageNumber="16">Fig. 5H</figureCitation>
<figureCitation id="131C2A1FFFA7B16AFBD5FC51FB5D1CE1" box="[1125,1225,954,981]" captionStart="Fig" captionStartId="18.[227,270,1756,1782]" captionTargetBox="[314,1363,302,1734]" captionTargetId="figure@18.[314,1363,302,1734]" captionTargetPageId="18" captionText="Fig. 5. Male morphology. AC, FI. Metasoma, lateral view. DE. Head capsule, frontal view. — A. Myrmica latifrons (U.S.A., CASENT0104816, A. Nobile). B. Pseudomyrmex indet. (Mexico, CASENT0103327, A. Nobile). C. Nothomyrmecia macrops (Australia, CASENT0902784, Z. Lieberman); note that abdominal segment III is reduced relative to, but not differentiated from segment IV. D. Myrmecia pilosula (Australia, CASENT0902800, Z. Lieberman). E. Tetraponera indet. (Madagascar, CASENT0053316, A. Nobile). F. Tatzuidris tatusia (Panama, CASENT0178870, E. Prado). G. Acromyrmex volcanus (Costa Rica, INBIOCRI001283114, E. Ortega). H. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). I. Formica pallidefulva (U.S.A., CASENT0172882, A. Nobile). Scale bars: AF = 0.5 mm, GI = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14900306" httpUri="https://zenodo.org/record/14900306/files/figure.png" pageId="15" pageNumber="16">Fig. 5H</figureCitation>
). Petiolar tergum and sternum not insensibly fused in anterior third; if fused, suture visible along entire petiolar length (
<figureCitation id="131C2A1FFFA7B16AFEC7FBEBFE5B1B2F" box="[375,463,1024,1051]" captionStart="Fig" captionStartId="19.[227,270,1755,1781]" captionTargetBox="[340,1306,302,1722]" captionTargetId="figure@19.[340,1306,302,1722]" captionTargetPageId="19" captionText="Fig. 6. Male morphology. AB, EG. Head capsule in frontal view. CD. Mesosoma, lateral view. HI. Petiole, ventrolateral view. — A. Probolomyrmex indet. (Madagascar, CASENT0080551, A. Nobile). B. Anochetus boltoni (Madagascar, CASENT0063847, A. Nobile). C. Leptanilloides gracilis (Guatemala, CASENT0234561, M. Borowiec). D. Odontomachus simillimus (Seychelles, CASENT0172666, A. Nobile). E. Platythyrea arthuri (Mayotte, CASENT0132466, E. Prado). F. Protanilla indet. (Thailand, CASENT0119776, A. Nobile). G. Stigmatomma indet. (Madagascar, CASENT0007087, E. Prado). H. Nothomyrmecia macrops (Australia, B. Boudinot). I. Neoponera cf. apicalis (Honduras, B. Boudinot). Scale bars: A, FG= 0.1 mm, B = 0.2 mm, C = 0.25 mm, E, HI = 0.5 mm, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3780160" httpUri="https://zenodo.org/record/3780160/files/figure.png" pageId="15" pageNumber="16">Fig. 6I</figureCitation>
). Mandibular teeth absent, fine, or robust……………………………………
@ -824,7 +824,7 @@ Martialinae (
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18 Mesosoma anteriorly elongated: Mesonotum almost twice as long as broad in dorsal view, and lateral pronotal face longer in profile view than head in full-face view (
@ -840,13 +840,9 @@ Martialinae (
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<keyLead id="30D695AFFFA7B16AFF53F829FA0818CB" pageId="15" pageNumber="16"> Mesosoma not anteriorly elongated: Mesonotum much less than twice as long as broad in dorsal view, and lateral pronotal face in profile view shorter than to as</keyLead>
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long as head length in profile view (
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Mesosoma not anteriorly elongated: Mesonotum much less than twice as long as broad in dorsal view, and lateral pronotal face in profile view shorter than to as long as head length in profile view (
<figureCitation id="131C2A1FFFB8B175FCADFEDAFC1D1E78" box="[797,905,305,332]" captionStart="Fig" captionStartId="20.[227,270,1860,1886]" captionTargetBox="[319,1356,302,1826]" captionTargetId="figure@20.[317,1356,302,1830]" captionTargetPageId="20" captionText="Fig. 7. AB. Forewing. A. Ventral view, gyne. B. Dorsal view, male. CD. Mesosoma, lateral view, male. EF. Paramere, lateral view, male. — A. Aneuretus simoni (Sri Lanka, CASENT0172259, A. Nobile). B. Nylanderia vividula (U.S.A., CASENT0058918, A. Nobile). C. Apomyrma stygia (Central African Republic, CASENT0086073, E. Prado). D. Anochetus boltoni (Madagascar, CASENT0063847, A. Nobile). E. Dolichoderus validus (Costa Rica, INB0003662427, B. Boudinot). F. Formica pacifica (U.S.A., JTLC000006350, B. Boudinot). Scale bars: AC, EF = 0.5 mm, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3780162" httpUri="https://zenodo.org/record/3780162/files/figure.png" pageId="16" pageNumber="17">Fig. 7D</figureCitation>
). Pterostigma present or absent. Petiole very narrowly attached to abdominal segment III, node strong, except
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@ -860,8 +856,9 @@ long as head length in profile view (
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part)
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