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<mods:titleInfo id="8B707E7C5F5979351DBAFD3D3CE1F625">
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<mods:title id="FA24CECA4EA2D6BE92A214739824DC5F">The oldest false fairy wasp (Hymenoptera: Mymarommatidae) from Lower Cretaceous Lebanese amber</mods:title>
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<mods:namePart id="6A97F7A1D369CC4EE226745E06D4BC5C">Álvarez-Parra, Sergio</mods:namePart>
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<mods:affiliation id="FEFBECE4A426703AF9A3CE289CD6B458">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="0512AC1E9AD34F18825017A59BE607E8">Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation id="4D3FF4E76C8A80E4BFD0EBC374E5E799">Palaeontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="0F0F2A8DFD490F81078BE25709EB0191">Azar, Dany</mods:namePart>
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<mods:affiliation id="422FDCCF7FE142726C05649255CE8A02">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Lebanese University, Faculty of Science II, Natural Sciences Department, Fanar - El-Matn, P. O. Box 90656 Jdeidet, Lebanon * Corresponding author</mods:affiliation>
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<mods:date id="371B3F3D6B49E6E91BCD8A2CEDFBB87E">2024</mods:date>
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<heading id="D0BC81C2FFEBFFFADAFFFF0BC9AFFF70" bold="true" box="[151,974,151,178]" fontSize="11" level="1" pageId="6" pageNumber="51" reason="1">
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<emphasis id="B93FEABCFFEBFFFADAFFFF0BC9AFFF70" bold="true" box="[151,974,151,178]" pageId="6" pageNumber="51">
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<paragraph id="8BF436AEFFEBFFFADAFFFF0BC8A9FF73" blockId="6.[151,974,151,178]" box="[151,712,151,178]" pageId="6" pageNumber="51">
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Key to the species of the genus
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD872FF0BC8A9FF73" ID-CoL="ZY4" authority="Yoshimoto, 1975" authorityName="Yoshimoto" authorityYear="1975" box="[538,712,151,178]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEABCFFEBFFFAD872FF0BC8A9FF73" bold="true" box="[538,712,151,178]" italics="true" pageId="6" pageNumber="51">Archaeromma</emphasis>
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</taxonomicName>
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</paragraph>
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<subSubSection id="C3516525FFEBFFFAD8B8FF0BC9AFFF70" box="[720,974,151,177]" pageId="6" pageNumber="51" type="description">
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<paragraph id="8BF436AEFFEBFFFAD8B8FF0BC9AFFF70" blockId="6.[151,974,151,178]" box="[720,974,151,177]" pageId="6" pageNumber="51">
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(
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD8B0FF0BC9A7FF70" authorityName="Debauche" authorityYear="1948" box="[728,966,151,177]" class="Insecta" family="Mymarommatidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="family">Mymarommatidae</taxonomicName>
|
||||
)
|
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</paragraph>
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</emphasis>
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</heading>
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<subSubSection id="C3516525FFEBFFFADAFFFF40CFFDFB34" pageId="6" pageNumber="51" type="key">
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<paragraph id="8BF436AEFFEBFFFADAFFFF40CFFDFF30" blockId="6.[151,1436,220,1269]" box="[151,1436,220,242]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFF40CFFDFF30" box="[151,1436,220,242]" pageId="6" pageNumber="51">1. Clava 3-segmented.................................................................................... 2</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFF66CFFDFECE" blockId="6.[151,1436,220,1269]" box="[151,1436,249,271]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFF66CFFDFECE" box="[151,1436,249,271]" pageId="6" pageNumber="51">- Clava 4-segmented.................................................................................... 5</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFE8BCFFDFE88" pageId="6" pageNumber="51">
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<paragraph id="8BF436AEFFEBFFFADAFFFE8BCFFDFEED" blockId="6.[151,1436,220,1269]" box="[151,1436,279,301]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFE8BCFFDFEED" box="[151,1436,279,301]" pageId="6" pageNumber="51">2. Forewing posterobasal marginal seta present................................................................ 3</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFEA9CFFDFE88" blockId="6.[151,1436,220,1269]" box="[151,1436,308,330]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFEA9CFFDFE88" box="[151,1436,308,330]" pageId="6" pageNumber="51">- Forewing posterobasal marginal seta absent................................................................ 4</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFECECFFDFE7E" pageId="6" pageNumber="51">
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFECECFFDFE44" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFECECFFDFE44" pageId="6" pageNumber="51">
|
||||
3. Pronotum reaching tegula; forewing marginal fringe with 42 long, erect setae..............................................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD8D3FEF3CE16FE45" authority="(Kozlov & Rasnitsyn, 1979)" authorityName="Transferred" baseAuthorityName="Kozlov & Rasnitsyn" baseAuthorityYear="1979" box="[699,1143,367,389]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="mandibulatum">
|
||||
<emphasis id="B93FEABCFFEBFFFAD8D3FEF3C8ABFE45" box="[699,714,367,388]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD8BEFEF3C900FE45" box="[726,865,367,388]" italics="true" pageId="6" pageNumber="51">mandibulatum</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD905FEF3CE11FE45" author="Kozlov, M. A. & Rasnitsyn, A. P." box="[877,1136,367,389]" pageId="6" pageNumber="51" pagination="402 - 416" refId="ref8196" refString="Kozlov, M. A. & Rasnitsyn, A. P. (1979) On the limits of the family Serphitidae (Hymenoptera, Proctotrupoidea). Entomologischeskoe Obozrenie, 58, 402 - 416. [in Russian]." type="journal article" year="1979">Kozlov & Rasnitsyn, 1979</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADEEEFEF3CEF7FE45" box="[1158,1174,367,388]" count="1" pageId="6" pageNumber="51" type="female">♀</specimenCount>
|
||||
), Taimyr amber, Santonian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFE11CFFDFE7E" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFE11CFFDFE7E" pageId="6" pageNumber="51">
|
||||
- Pronotum not reaching tegula; forewing marginal fringe with 55 long, erect setae...............................................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD895FE36CE34FE7E" ID-CoL="C93K6" authority="Engel & Grimaldi, 2007" authorityName="Engel & Grimaldi" authorityYear="2007" box="[765,1109,425,447]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="carnifex">
|
||||
<emphasis id="B93FEABCFFEBFFFAD895FE36C96DFE7E" box="[765,780,426,447]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD97FFE35C907FE7F" box="[791,870,425,446]" italics="true" pageId="6" pageNumber="51">carnifex</emphasis>
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD904FE35CE34FE7E" author="Engel, M. S. & Grimaldi, D. A." box="[876,1109,425,447]" pageId="6" pageNumber="51" pagination="159 - 168" refId="ref7778" refString="Engel, M. S. & Grimaldi, D. A. (2007) New false fairy wasps in Cretaceous amber from New Jersey and Myanmar (Hymenoptera: Mymarommatoidea). Transactions of the Kansas Academy of Science, 110, 159 - 168. https: // doi. org / 10.1660 / 0022 - 8443 (2007) 110 [159: NFFWIC] 2.0. CO; 2" type="journal article" year="2007">Engel & Grimaldi, 2007</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE0BFE36CE16FE7E" box="[1123,1143,426,447]" count="1" pageId="6" pageNumber="51" type="male">♂</specimenCount>
|
||||
), New Jersey amber, Turonian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFE5BCFFDFDF4" pageId="6" pageNumber="51">
|
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<paragraph id="8BF436AEFFEBFFFADAFFFE5BCFFDFE3B" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFE5BCFFDFE3B" pageId="6" pageNumber="51">
|
||||
4. Pedicel shorter than scape; mesosoma narrower than head.......................................................................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD959FE78CE32FE3B" ID-CoL="C93KC" authority="(Yoshimoto, 1975)" authorityName="Transferred" baseAuthorityName="Yoshimoto" baseAuthorityYear="1975" box="[817,1107,484,506]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="masneri">
|
||||
<emphasis id="B93FEABCFFEBFFFAD959FE78C921FE38" box="[817,832,484,505]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD924FE79C9F8FE3B" box="[844,921,485,506]" italics="true" pageId="6" pageNumber="51">masneri</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD9CDFE78CE2DFE3B" author="Yoshimoto, C. M." box="[933,1100,484,506]" pageId="6" pageNumber="51" pagination="499 - 528" refId="ref8939" refString="Yoshimoto, C. M. (1975) Cretaceous chalcidoid fossils from Canadian amber. Canadian Entomologist, 107, 499 - 528. https: // doi. org / 10.4039 / Ent 107499 - 5" type="journal article" year="1975">Yoshimoto, 1975</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE0BFE78CE12FE38" box="[1123,1139,484,505]" count="1" pageId="6" pageNumber="51" type="female">♀</specimenCount>
|
||||
), Canadian amber, Campanian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFD9ECFFDFDF4" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFD9ECFFDFDF4" pageId="6" pageNumber="51">
|
||||
- Pedicel as long as scape; mesosoma broader than head......................................................................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD96DFD83CE34FDF5" ID-CoL="C93K7" authority="Engel & Grimaldi, 2007" authorityName="Engel & Grimaldi" authorityYear="2007" box="[773,1109,543,565]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="gibsoni">
|
||||
<emphasis id="B93FEABCFFEBFFFAD96DFD83C975FDF5" box="[773,788,543,564]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD977FD83C907FDF5" box="[799,870,543,564]" italics="true" pageId="6" pageNumber="51">gibsoni</emphasis>
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD904FD83CE34FDF5" author="Engel, M. S. & Grimaldi, D. A." box="[876,1109,543,565]" pageId="6" pageNumber="51" pagination="159 - 168" refId="ref7778" refString="Engel, M. S. & Grimaldi, D. A. (2007) New false fairy wasps in Cretaceous amber from New Jersey and Myanmar (Hymenoptera: Mymarommatoidea). Transactions of the Kansas Academy of Science, 110, 159 - 168. https: // doi. org / 10.1660 / 0022 - 8443 (2007) 110 [159: NFFWIC] 2.0. CO; 2" type="journal article" year="2007">Engel & Grimaldi, 2007</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE0BFD83CE16FDF5" box="[1123,1143,543,564]" count="1" pageId="6" pageNumber="51" type="male">♂</specimenCount>
|
||||
), New Jersey amber, Turonian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFDA0CFFDFD4C" pageId="6" pageNumber="51">
|
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<paragraph id="8BF436AEFFEBFFFADAFFFDA0CFFDFDAE" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFDA0CFFDFDAE" pageId="6" pageNumber="51">
|
||||
5. Notauli present; petiole with second segment longer than first segment......................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFADB35FDC6CE32FDAE" authority="Cockx, McKellar, Tappert, Vavrek & Muehlenbachs, 2020" authorityName="Cockx, McKellar, Tappert, Vavrek & Muehlenbachs" authorityYear="2020" box="[349,1107,601,623]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="trigonokephalion">
|
||||
<emphasis id="B93FEABCFFEBFFFADB35FDC6CB0DFDAE" box="[349,364,602,623]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFADB10FDC5C87CFDAF" box="[376,541,601,622]" italics="true" pageId="6" pageNumber="51">trigonokephalion</emphasis>
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD84BFDC5CE32FDAE" author="Cockx, P. & McKellar, R. & Tappert, R. & Vavrek, M. & Muehlenbachs, K." box="[547,1107,601,623]" pageId="6" pageNumber="51" pagination="378 - 389" refId="ref7295" refString="Cockx, P., McKellar, R., Tappert, R., Vavrek, M. & Muehlenbachs, K. (2020) Bonebed amber as a new source of paleontological data: The case of the Pipestone Creek deposit (Upper Cretaceous), Alberta, Canada. Gondwana Research, 81, 378 - 389. https: // doi. org / 10.1016 / j. gr. 2019.12.005" type="journal article" year="2020">Cockx, McKellar, Tappert, Vavrek & Muehlenbachs, 2020</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE09FDC6CE14FDAE" box="[1121,1141,602,623]" count="1" pageId="6" pageNumber="51" type="male">♂</specimenCount>
|
||||
), Canadian amber, Campanian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFDEBCFFDFD4C" blockId="6.[151,1436,220,1269]" box="[151,1436,631,653]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFDEBCFFDFD4C" box="[151,1436,631,653]" pageId="6" pageNumber="51">- Notauli absent; petiole with first segment longer or as long as second segment..................................... 6</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFD08CFFDFD06" pageId="6" pageNumber="51">
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFD08CFFDFD6B" blockId="6.[151,1436,220,1269]" box="[151,1436,660,682]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFD08CFFDFD6B" box="[151,1436,660,682]" pageId="6" pageNumber="51">6. Forewing posterobasal marginal seta present................................................................ 7</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFD2ECFFDFD06" blockId="6.[151,1436,220,1269]" box="[151,1436,689,711]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFD2ECFFDFD06" box="[151,1436,689,711]" pageId="6" pageNumber="51">- Forewing posterobasal marginal seta absent............................................................... 11</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFD53CFFDFCC0" pageId="6" pageNumber="51">
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFD53CFFDFD25" blockId="6.[151,1436,220,1269]" box="[151,1436,719,741]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFD53CFFDFD25" box="[151,1436,719,741]" pageId="6" pageNumber="51">7. Claval segments strongly compacted (sutures almost undistinguishable).......................................... 8</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFD71CFFDFCC0" blockId="6.[151,1436,220,1269]" box="[151,1436,748,770]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFD71CFFDFCC0" box="[151,1436,748,770]" pageId="6" pageNumber="51">- Claval segments not strongly compacted (sutures distinguishable)............................................... 9</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFC96CFFDFCB6" pageId="6" pageNumber="51">
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFC96CFFDFCFC" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFC96CFFDFCFC" pageId="6" pageNumber="51">
|
||||
8. Lengths of funicular segments increase towards apex; mesosoma longer than gaster...............................................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD963FCBBCE54FCFD" authority="(Brues, 1937)" authorityName="Transferred" baseAuthorityName="Brues" baseAuthorityYear="1937" box="[779,1077,807,828]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="minutissimum">
|
||||
<emphasis id="B93FEABCFFEBFFFAD963FCBBC97BFCFD" box="[779,794,807,828]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD94EFCBBC9CAFCFD" box="[806,939,807,828]" italics="true" pageId="6" pageNumber="51">minutissimum</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD9D0FCBBCE4FFCFD" author="Brues, C. T." box="[952,1070,807,828]" pageId="6" pageNumber="51" refId="ref7203" refString="Brues, C. T. (1937) Superfamilies Ichneumonoidea, Serphoidea, and Chalcidoidea. In: Carpenter, F. M., Folsom, J. W., Essig, E. O.," type="book" year="1937">Brues, 1937</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE2DFCBBCE37FCFD" box="[1093,1110,807,828]" count="1" pageId="6" pageNumber="51" type="female">♀</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE09FCBBCE14FCFD" box="[1121,1141,807,828]" count="1" pageId="6" pageNumber="51" type="male">♂</specimenCount>
|
||||
), Canadian amber, Campanian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFCD9CFFDFCB6" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFCD9CFFDFCB6" pageId="6" pageNumber="51">
|
||||
- Lengths of funicular segments similar; mesosoma shorter than gaster in females.................................................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD96EFCFECE54FCB6" authority="Yoshimoto, 1975" authorityName="Yoshimoto" authorityYear="1975" box="[774,1077,865,887]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="nearcticum">
|
||||
<emphasis id="B93FEABCFFEBFFFAD96EFCFEC974FCB6" box="[774,789,866,887]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD949FCFEC9EDFCB6" box="[801,908,866,887]" italics="true" pageId="6" pageNumber="51">nearcticum</emphasis>
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD9F9FCFDCE54FCB6" author="Yoshimoto, C. M." box="[913,1077,865,887]" pageId="6" pageNumber="51" pagination="499 - 528" refId="ref8939" refString="Yoshimoto, C. M. (1975) Cretaceous chalcidoid fossils from Canadian amber. Canadian Entomologist, 107, 499 - 528. https: // doi. org / 10.4039 / Ent 107499 - 5" type="journal article" year="1975">Yoshimoto, 1975</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE2DFCFECE37FCB6" box="[1093,1110,866,887]" count="1" pageId="6" pageNumber="51" type="female">♀</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE09FCFECE14FCB6" box="[1121,1141,866,887]" count="1" pageId="6" pageNumber="51" type="male">♂</specimenCount>
|
||||
), Canadian amber, Campanian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFCE3CFFDFC0E" pageId="6" pageNumber="51">
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFCE3CFFDFC73" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFCE3CFFDFC73" pageId="6" pageNumber="51">
|
||||
9. Mesoscutum without apparent sculpture; forewing marginal fringe with 31 long, erect setae..........................................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD975FC00CE26FC70" authority="Aiba & Inosa, 2024" authorityName="Aiba & Inosa" authorityYear="2024" box="[797,1095,924,946]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="chisatoi">
|
||||
<emphasis id="B93FEABCFFEBFFFAD975FC00C94DFC70" box="[797,812,924,945]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD95FFC00C9E2FC70" box="[823,899,924,945]" italics="true" pageId="6" pageNumber="51">chisatoi</emphasis>
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD9E1FC00CE26FC70" author="Aiba, H. & Inosa, H." box="[905,1095,924,946]" pageId="6" pageNumber="51" pagination="1 - 8" refId="ref6915" refString="Aiba, H. & Inosa, H. (2024) A new false fairy wasp (Hymenoptera: Mymarommatoidea: Mymarommatidae) in Late Cretaceous Iwaki amber from Futaba Group of Iwaki City, Fukushima Prefecture, Japan. Paleontological Research, 28 (3), 1 - 8. https: // doi. org / 10.2517 / PR 230020" type="journal article" year="2024">Aiba & Inosa, 2024</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE3FFC00CE09FC70" box="[1111,1128,924,945]" count="1" pageId="6" pageNumber="51" type="female">♀</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE1BFC00CEE6FC70" box="[1139,1159,924,945]" count="1" pageId="6" pageNumber="51" type="male">♂</specimenCount>
|
||||
), Japanese amber, Coniacian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFC26CFFDFC0E" blockId="6.[151,1436,220,1269]" box="[151,1436,953,975]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFC26CFFDFC0E" box="[151,1436,953,975]" pageId="6" pageNumber="51">- Mesoscutum with evident sculpture; forewing marginal fringe with 36–38 long, erect setae.......................... 10</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFC4BCFFDFB84" pageId="6" pageNumber="51">
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFC4BCFFDFBCB" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFC4BCFFDFBCB" pageId="6" pageNumber="51">
|
||||
10. Mesoscutum with transverse, striate sculpture; forewing posterobasal marginal seta offset by a series of nine short, posterior setae......................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD84DFC68CE04FBC8" ID-CoL="C93K9" authority="(Fursov, Shirota, Nomiya & Yamagishi, 2002)" baseAuthorityName="Fursov, Shirota, Nomiya & Yamagishi" baseAuthorityYear="2002" box="[549,1125,1012,1034]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="japonicum">
|
||||
<emphasis id="B93FEABCFFEBFFFAD84DFC68C855FBC8" box="[549,564,1012,1033]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD828FC69C8C4FBCB" box="[576,677,1013,1034]" italics="true" pageId="6" pageNumber="51">japonicum</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD8D9FC68CE01FBC8" author="Fursov, V. & Shirota, Y. & Nomiya, T. & Yamagishi, K." box="[689,1120,1012,1034]" pageId="6" pageNumber="51" pagination="51 - 54" refId="ref7852" refString="Fursov, V., Shirota, Y., Nomiya, T. & Yamagishi, K. (2002) New fossil mymarommatid species, Palaeomymar japonicum sp. nov. (Hymenoptera: Mymarommatidae), discovered in Cretaceous amber from Japan. Entomological Science, 5, 51 - 54." type="journal article" year="2002">Fursov, Shirota, Nomiya & Yamagishi, 2002</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE1CFC68CEE9FBC8" box="[1140,1160,1012,1033]" count="1" pageId="6" pageNumber="51" type="male">♂</specimenCount>
|
||||
), Japanese amber, Santonian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFB8ECFFDFB84" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFB8ECFFDFB84" pageId="6" pageNumber="51">
|
||||
- Mesoscutum with reticulate sculpture; forewing posterobasal marginal seta offset by a series of four–five short, posterior setae......................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD8B7FBB3CE16FB85" authority="(Kozlov & Rasnitsyn, 1979)" authorityName="Transferred" baseAuthorityName="Kozlov & Rasnitsyn" baseAuthorityYear="1979" box="[735,1143,1071,1093]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="senonicum">
|
||||
<emphasis id="B93FEABCFFEBFFFAD8B7FBB3C88FFB85" box="[735,750,1071,1092]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD892FBB3C900FB85" box="[762,865,1071,1092]" italics="true" pageId="6" pageNumber="51">senonicum</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD905FBB3CE11FB85" author="Kozlov, M. A. & Rasnitsyn, A. P." box="[877,1136,1071,1093]" pageId="6" pageNumber="51" pagination="402 - 416" refId="ref8196" refString="Kozlov, M. A. & Rasnitsyn, A. P. (1979) On the limits of the family Serphitidae (Hymenoptera, Proctotrupoidea). Entomologischeskoe Obozrenie, 58, 402 - 416. [in Russian]." type="journal article" year="1979">Kozlov & Rasnitsyn, 1979</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADEEEFBB3CEF7FB85" box="[1158,1174,1071,1092]" count="1" pageId="6" pageNumber="51" type="female">♀</specimenCount>
|
||||
), Taimyr amber, Santonian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30BF2E0BFFEBFFFADAFFFBD0CFFDFB34" pageId="6" pageNumber="51">
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFBD0CFFDFB5C" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFBD0CFFDFB5C" pageId="6" pageNumber="51">
|
||||
11. Forewing marginal fringe with 42–43 long, erect setae; forewing without mid-sized posterior setae; mesotibiae and metatibiae without pseudospurs; metatibiae with four distinct setae forming a comb on posterior distal margin.............................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD867FB1BCE10FB5D" ID-CoL="C93K8" authority="Ortega-Blanco, Penalver, Delclos & Engel, 2011" authorityName="Ortega-Blanco, Penalver, Delclos & Engel" authorityYear="2011" box="[527,1137,1159,1181]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="hispanicum">
|
||||
<emphasis id="B93FEABCFFEBFFFAD867FB1BC87FFB5D" box="[527,542,1159,1180]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD841FB1BC8F9FB5D" box="[553,664,1159,1180]" italics="true" pageId="6" pageNumber="51">hispanicum</emphasis>
|
||||
<bibRefCitation id="EFDA4B5FFFEBFFFAD8F6FB1BCE10FB5D" author="Ortega-Blanco, J. & Penalver, E. & Delclos, X. & Engel, M. S." box="[670,1137,1159,1181]" pageId="6" pageNumber="51" pagination="511 - 523" refId="ref8489" refString="Ortega-Blanco, J., Penalver, E., Delclos, X. & Engel, M. S. (2011) False fairy wasps in early Cretaceous amber from Spain (Hymenoptera: Mymarommatoidea). Palaeontology, 54 (3), 511 - 523. https: // doi. org / 10.1111 / j. 1475 - 4983.2011.01049. x" type="journal article" year="2011">Ortega-Blanco, Peñalver, Delclòs & Engel, 2011</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADEE9FB1BCEF3FB5D" box="[1153,1170,1159,1180]" count="1" pageId="6" pageNumber="51" type="female">♀</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D4DFD27FFEBFFFADEF5FB1BCED0FB5D" box="[1181,1201,1159,1180]" count="1" pageId="6" pageNumber="51" type="male">♂</specimenCount>
|
||||
), Spanish amber, Albian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BF436AEFFEBFFFADAFFFB39CFFDFB34" blockId="6.[151,1436,220,1269]" pageId="6" pageNumber="51">
|
||||
<keyLead id="30BA959BFFEBFFFADAFFFB39CFFDFB34" pageId="6" pageNumber="51">
|
||||
- Forewing marginal fringe with 38 long, erect setae; forewing with four mid-sized posterior setae; mesotibiae with two pseudospurs and metatibiae with four pseudospurs; metatibiae without setae forming a comb on posterior distal margin............................................................
|
||||
<taxonomicName id="4C4B4D2DFFEBFFFAD916FB43CE68FB35" authorityName="Álvarez-Parra & Rasnitsyn & Azar" authorityYear="2024" box="[894,1033,1247,1268]" class="Insecta" family="Mymarommatidae" genus="Archaeromma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="phoenicium" status="sp. nov.">
|
||||
<emphasis id="B93FEABCFFEBFFFAD916FB43C9ECFB35" box="[894,909,1247,1268]" italics="true" pageId="6" pageNumber="51">A</emphasis>
|
||||
.
|
||||
<emphasis id="B93FEABCFFEBFFFAD9F1FB43CE68FB35" box="[921,1033,1247,1268]" italics="true" pageId="6" pageNumber="51">phoenicium</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B93FEABCFFEBFFFADE67FB43CE3AFB35" bold="true" box="[1039,1115,1247,1268]" pageId="6" pageNumber="51">
|
||||
<taxonomicNameLabel id="A20C57C7FFEBFFFADE67FB43CE3AFB35" box="[1039,1115,1247,1268]" pageId="6" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(
|
||||
<specimenCount id="9D4DFD27FFEBFFFADE03FB43CE1AFB35" box="[1131,1147,1247,1268]" count="1" pageId="6" pageNumber="51" type="female">♀</specimenCount>
|
||||
), Lebanese amber, Barremian
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,62 +1,62 @@
|
|||
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<mods:mods id="9255592BE8EBF98E297D21C9D39F6965" xmlns:mods="http://www.loc.gov/mods/v3">
|
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(
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<figureCitation id="13702A2BFFEFFFFEDAF7F9E3CB63F958" box="[159,258,1663,1689]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1903,1927]" captionStartId-1="4.[151,250,815,839]" captionStartId-2="5.[151,250,1316,1340]" captionTargetBox-0="[151,1436,522,1856]" captionTargetBox-1="[151,1426,181,791]" captionTargetBox-2="[151,1436,181,1291]" captionTargetId-0="figure-166@3.[151,1436,522,1877]" captionTargetId-1="figure-20@4.[151,1436,181,791]" captionTargetId-2="figure-19@5.[151,1436,181,1291]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Archaeromma phoenicium sp. nov., holotype, female. A, Habitus, right lateral view. B, Habitus, left lateral view. C, Drawing of the specimen from left lateral view, head from ventral view. D, Interpretative drawing of antenna. Abbreviations: sc—scape, p—pedicel, f1–f7—funicular segments, cl1–cl4—claval segments." captionText-1="FIGURE 2. Archaeromma phoenicium sp. nov., holotype, female. A, Forewings in right lateral view. B, Detail of stigmal complex of left forewing indicating macrochaeta (red arrowhead) and hindwings (black arrowheads). C, Drawing of right forewing. D, Interpretative drawing of stigmal complex of left forewing. E, Interpretative drawing of hind wing. Abbreviations: cc—costal cell, ham—hamulus, mb—hind wing membrane, mv—marginal vein, op—opposite projection, ret—retinaculum, smv—submarginal vein, stv—stigmal vein, tv—hind wing tubular vein." captionText-2="FIGURE 3. Archaeromma phoenicium sp. nov., holotype, female. A, Head in dorsal view and mesosoma in right lateral view. B, Head in ventral view and mesosoma in left lateral view. C, Protarsus and interpretative drawing of protibial calcar and basitarsus. D, Mesotarsus with two pseudospurs (red arrowheads). E, Metatarsus with four pseudospurs (red arrowheads). F, Metasoma in right lateral view. G, Metasoma in left lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.14595754" figureDoi-1="http://doi.org/10.5281/zenodo.14595756" figureDoi-2="http://doi.org/10.5281/zenodo.14595758" httpUri-0="https://zenodo.org/record/14595754/files/figure.png" httpUri-1="https://zenodo.org/record/14595756/files/figure.png" httpUri-2="https://zenodo.org/record/14595758/files/figure.png" pageId="2" pageNumber="47">Figs 1–3</figureCitation>
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<emphasis id="B93FEABCFFEFFFFEDAFFF95BCB28F920" bold="true" box="[151,329,1735,1761]" pageId="2" pageNumber="47">Type material.</emphasis>
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<materialsCitation id="3B233CF3FFEFFFFEDB3BF95BCF37F8E8" collectorName="Science II" country="Lebanon" location="Faculty of Science" pageId="2" pageNumber="47" specimenCount="2" typeStatus="holotype">
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<materialsCitation id="3B233CF3FFEFFFFEDB3BF95BCF37F8E8" collectorName="Science II" country="Lebanon" location="Faculty of Science" pageId="2" pageNumber="47" specimenCode="1701-A" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54F0880CFFEFFFFEDB3BF95BCBDAF920" box="[339,443,1735,1761]" pageId="2" pageNumber="47" type="holotype">Holotype</typeStatus>
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1701-A; a complete female specimen; left antenna and left forewing are affected by fractures in the amber. Syninclusion with two spiders (Araneae) and two springtails (Collembola). The
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is deposited at the Natural History Museum of the Lebanese University,
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Faculty of
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II, Fanar,
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<collectingCountry id="F35C763EFFEFFFFEDE84F893CF33F8E8" box="[1260,1362,1807,1833]" name="Lebanon" pageId="2" pageNumber="47">Lebanon</collectingCountry>
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.
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<specimenCode id="DBED9ED5FFEFFFFEDBAEF95BC841F920" box="[454,544,1735,1761]" pageId="2" pageNumber="47">1701-A</specimenCode>
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; a complete
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<specimenCount id="9D4DFD27FFEFFFFED8DFF95BC962F920" box="[695,771,1735,1761]" pageId="2" pageNumber="47" type="female">female</specimenCount>
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specimen; left antenna and left forewing are affected by fractures in the amber. Syninclusion with two spiders (Araneae) and two springtails (Collembola). The holotype is deposited at the Natural History Museum of the Lebanese University, Faculty of Science II, Fanar, Lebanon.
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|
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@ -107,11 +101,13 @@ After Phoenicia, ancient name of the region of modern-day
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.
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|
||||
Female. Compound eyes composed by about 30–40 ommatidia; no ocelli; mandibles exodont, bidentate; toruli not contiguous, inserted above compound eyes; antenna with funicle 7-segmented and clava 4- segmented; scape 2.8× longer than wide; pedicel longer than cumulative lengths of f1 and f2; clava slightly longer than cumulative lengths of scape and pedicel; forewing: submarginal and marginal veins separated by a bulla, stigmal vein present as a bulbous area, marginal fringe composed by: six short anterior setae, eight mid-sized anterior setae, 38 long erect setae deeply inserted in membrane, and four mid-sized posterior setae, no long posterobasal marginal seta; hind wing: stalk-like, with posterior membrane slender, distal pincer-like structure present; protibial calcar curved and simple; mesotibiae with two pseudospurs; metatibiae with four pseudospurs; basitarsi as long as cumulative lengths of second and third tarsomeres; petiole bi-segmented with first segment 1.2× longer than second; gaster flattened; dorsal part of gaster setose.
|
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3516525FFEEFFFFDAAFFEF2CFE9FE6C" pageId="3" pageNumber="48" type="distribution">
|
||||
<paragraph id="8BF436AEFFEEFFFFDAAFFEF2CFE9FE6C" blockId="3.[151,1437,151,501]" pageId="3" pageNumber="48">
|
||||
<emphasis id="B93FEABCFFEEFFFFDAAFFEF2CBA7FE49" bold="true" box="[199,454,366,393]" pageId="3" pageNumber="48">Locality and horizon.</emphasis>
|
||||
Hammana-Mdeyrij amber-bearing outcrop, Caza Baabda, Mouhafazet Mount
|
||||
|
|
|
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