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G-W. -Chen -, Z-M. -Dong -, P-Z. -Wang +, coll. +G-W. Chen +, +Z-M. Dong +, +P-Z. Wang and co-workers, sagittal sections on 27 slides ; @@ -156,17 +156,19 @@ and 10, ibid., transverse sections on 44 and 10 slides respectively and 8, ibid., horizontal sections on 22 and 26 slides respectively ; - -RMNH -.VER.21595.1, ibid., sagittal sections on 13 slides + +RMNH.VER.21595.1 +, ibid., sagittal sections on 13 slides ; - -RMNH -.VER.21595.2, ibid., sagittal sections on 37 slides + +RMNH.VER.21595.2 +, ibid., sagittal sections on 37 slides . + + Habitat and reproduction. All animals were collected from a freshwater stream at Guanyin Mountain Protection Station in the central Qinling Mountains ( @@ -177,6 +179,8 @@ All animals were collected from a freshwater stream at Guanyin Mountain Protecti 1715 m a.s.l. (air temperature 9 ℃, water temperature 11 ℃). Many animals were sexually mature at collection, but no cocoons were found.After 11 months under laboratory conditions, the population failed to adapt to the laboratory environment and there were no surviving individuals. Furthermore, no cocoons were laid by the sexual animals. + + Karyotype. Each of the five, randomly selected specimens exhibited diploid chromosome complements. From a total of 104 metaphase plates examined, 86 chromosome complements were diploid with 2n = 2x = 40 = @@ -194,11 +198,9 @@ chromosomes were submetacentric, 10 chromosomes were subtelocentric and the remaining were metacentric ( Fig. 3 ). The chromosome complements of the remaining 18 plates could not be determined due to either lack of well-dispersed chromosomes or over-dispersed sets of chromosomes. Karyotype parameters, including relative length, arm ratio, and centromeric index are shown in -Table 1 +Table 1 . - - Description. In living sexual specimens, the body measured @@ -231,7 +233,7 @@ A pair of ventral ovaries are located at 1/3 – 1/5 of the distance between the 6 ). - + TABLE 1 Karyotype parameters (mean values and standard deviations) of @@ -242,7 +244,7 @@ Karyotype parameters (mean values and standard deviations) of - +
@@ -594,7 +596,6 @@ did not report on the chromosomes, the present study is the first account on the for which subtelocentric chromosomes have been described. - Presently known localities for P. asiatica @@ -625,9 +626,7 @@ a.s.l. 1715 m a.s.l. -) - -. +). The features of diff --git a/data/03/A2/87/03A287A3602B9D3DFF23FE64FD0BFC54.xml b/data/03/A2/87/03A287A3602B9D3DFF23FE64FD0BFC54.xml new file mode 100644 index 00000000000..c436c546cde --- /dev/null +++ b/data/03/A2/87/03A287A3602B9D3DFF23FE64FD0BFC54.xml @@ -0,0 +1,836 @@ + + + +Two species of Polycelis (Platyhelminthes, Tricladida, Planariidae) newly recorded for the Qinling Mountains and the Loess Plateau in China, with a comparative discussion on their karyotypes + + + +Author + +Sun, Xin-Xin +0000-0001-8167-1495 +College of Life Science, Henan Normal University, Xinxiang, 453007 Henan, China + + + +Author + +Wang, Lei +0000-0003-2328-1530 +College of Life Science, Henan Normal University, Xinxiang, 453007 Henan, China + + + +Author + +Li, Ning +0009-0003-1351-5089 +College of Life Science, Henan Normal University, Xinxiang, 453007 Henan, China +lining2021@htu.edu.cn + + + +Author + +Sluys, Ronald +0000-0002-9776-3471 +Naturalis Biodiversity Center, Leiden, The Netherlands +ronald.sluys@naturalis.nl + + + +Author + +Liu, De-Zeng +0000-0002-9354-2073 +College of Life Science, Henan Normal University, Xinxiang, 453007 Henan, China + + + +Author + +Dong, Zi-Mei +0000-0002-6416-590X +College of Life Science, Henan Normal University, Xinxiang, 453007 Henan, China + + + +Author + +Chen, Guang-Wen +0000-0001-8723-523X +College of Life Science, Henan Normal University, Xinxiang, 453007 Henan, China +chengw0183@sina.com + +text + + +Zootaxa + + +2024 + +2024-11-06 + + +5536 + + +4 + + +581 +599 + + + + +http://dx.doi.org/10.11646/zootaxa.5536.4.5 + +journal article +10.11646/zootaxa.5536.4.5 +1175-5326 +14239215 +AB30B22C-2FA0-494A-AE57-2C4778B6020C + + + + + + + +Polycelis eudendrocoeloides +( +Zabusova, 1929 +) + + + + + + + +Material examined. + +ZMHNU-JX8 +, +Wang Jiabian Town +( +38°14’50”N +110°14’24’’E +; alt. + +988 m +a.s.l. + +), +Jiaxian County +, +Shaanxi Province +, +China +, + +2 August 2017 + +, coll. +G-W. Chen +, +Z-M. Dong +, +Y-Q. Yu-wen +, and co-workers, sagittal sections on 28 slides + +; + +ZMHNU-JX2–7 +, ibid., sagittal sections on 13, 21, 9, 35, 9 and 9 slides, respectively + +; + +ZMHNU-JX9 +, ibid., horizontal sections on 19 slides + +; + +ZMHNU-JX10 +, ibid., transverse sections on 34 slides + +; + +RMNH. VER.21596.1 +, ibid., sagittal sections on 12 slides + +; + +RMNH.VER.21596.2 +, ibid., sagittal sections on 22 slides + +. + + + + +Habitat and reproduction. +All animals were collected from a stream on the Loess Plateau ( +Figs 1 +, +7A, B +) in northern +China +at +988 m +a.s.l. (air temperature 5 ℃, water temperature 15 ℃). Many animals were sexually mature at collection; however, no cocoons were collected. After 90 days under laboratory conditions, the sexual animals laid cocoons. Newly laid cocoons were yellow, but turned dark brown after 2 to 3 days. Cocoons were spherical in shape (< +1 mm +in diameter) and without a stalk. Since no young worms hatched from any of the cocoons, these were likely infertile. This population failed to adapt to the laboratory environment, and eventually there were no surviving individuals. + + + + + +Karyology. +Each of the five randomly selected specimens exhibited diploid chromosome complements. From a total of 109 metaphase plates examined, 90 chromosome complements were diploid with 2n = 2x = 36 ( +Fig. 8 +), while the chromosome complements of the remaining 19 plates failed to be determined, due to either lack of well-dispersed chromosomes or over-dispersed sets of chromosomes. All chromosomes were metacentric. Karyotype parameters, including relative length, arm ratio, and centromeric index, are shown in +Table 2 +. + + + + +
Chromosome Relative length
+ +FIGURE 7. +Habitat and external appearance of + +Polycelis eudendrocoeloides + +. +A & B +sampling site and habitat. +C +sexually mature, live individual. +D +Head with eyes, Scale bar: 500 μm. + + + +FIGURE 8. + +Polycelis eudendrocoeloides +. + +A +metaphase plate of diploid complement. +B +karyogram of diploid complement. +C +idiogram. Scale bar: 5 μm. + + + +TABLE 2 +Karyotype parameters (mean values and standard deviations) of + +Polycelis eudendrocoeloides + +; m: metacentric. + +
+ + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
ChromosomeRelative lengthArm ratioCentromere indexChromosome type
15.39±1.111.18±0.0746.06±0.01m
23.18±0.231.29±0.1143.79±0.02m
32.66±0.141.40±0.2142.41±0.03m
42.50±0.181.37±0.1542.43±0.03m
52.40±0.171.38±0.1442.23±0.02m
62.27±0.111.33±0.1743.68±0.02m
72.20±0.081.48±0.0540.72±001m
82.17±0.081.41±0.1041.68±0.02m
92.07±0.051.34±0.1943.13±0.03m
101.96±0.101.20±0.2746.23±0.05m
111.95±0.061.44±0.3241.69±0.05m
+ + +......continued on the next page + + + +TABLE 2. +(Continued) + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
ChromosomeRelative lengthArm ratioCentromere indexChromosome type
121.90±0.091.22±0.1545.49±0.03m
131.84±0.081.49±0.2340.77±0.03m
141.80±0.121.46±0.1341.17±0.02m
151.75±0.161.33±0.0543.18±0.01m
161.64±0.131.45±0.2341.95±0.03m
171.56±0.181.40±0.2142.00±0.04m
181.45±0.261.50±0.2140.78±0.04m
+ + +Description. +In living sexual specimens, the body measured +8–12 mm +in length and +4.3–4.7 mm +in width. The frontal margin of the body is slightly convex, and provided with a pair of bluntly pointed, laterally projecting auricles. Dorsal surface brownish, and the ventral surface is paler than the dorsal one, being slightly light brown ( +Fig. 7C +). About 49–80 small eyes are arranged into two bands that run from the frontal margin of the body along the lateral margins to a short distance posterior to the auricles ( +Fig. 7C +). + + +The pharynx, situated in the posterior half of the body, measures about 1/3–1/4 +th +of the body length; the mouth opening is located at the posterior end of the pharyngeal pocket ( +Fig. 7C +). Outer pharyngeal musculature is composed of a subepithelial layer of longitudinal muscles, followed by a layer of circular muscles. Inner pharyngeal musculature is composed of a thick subepithelial layer of circular muscles, followed by a thin layer of longitudinal muscles ( +Fig. 9A +). + + +The ventral, oval-shaped ovaries are located at 1/3–1/4 of the distance between the brain and the root of the pharynx, occupying about 1/3 of the dorso-ventral space ( +Fig. 9B +). The oviducts arise from the posterior wall of the ovaries and follow a ventral course in caudal direction. At the level of the male atrium, the two oviducts curve towards the dorsal body surface and dorsally to the atrium they unite to form a common oviduct, which opens into the dorsal portion of a very small and/or indistinct common atrium ( +Figs. 9C, D +, +10C +, +11 +). + + +The large, sac-shaped copulatory bursa occupies the entire dorso-ventral space and is lined by a vacuolated epithelium with basal nuclei ( +Figs. 9E +, +10E +). From the postero-dorsal wall of the copulatory bursa, the fairly long bursal canal runs in a caudal direction to the level of the gonopore and then curves ventrally to communicate with the common atrium and the gonopore ( +Figs. 10C, E +, +11 +). The bursal canal is lined with cylindrical, ciliated cells with basal nuclei and is surrounded by a subepithelial layer of circular muscles, followed by a thin layer of longitudinal muscles ( +Fig. 9C +). + + +The tall, ventrally located testes occupy almost the entire dorso-ventral space ( +Fig. 9F +). In fully mature individuals, there are about 20–30 testicular follicles extending from the level of the ovaries to the root of the pharynx. Mature spermatozoa are present in the testes and the vasa deferentia. The latter form spermiducal vesicles at the posterior level of the pharyngeal pocket in most specimens (RMNH.VER.21596.1, ZMHNU-JX2, 3, 4, 6, 7, 10, RMNH.VER.21596.2), except specimens ZMHNU-JX5, 8, and 9 ( +Fig. 10A +). At the level of the penis bulb, the vasa deferentia bend towards the dorsal body surface and then penetrate the antero-lateral or lateral wall of the penis bulb to open separately and symmetrically into the mid-lateral portion of the seminal vesicle. ( +Figs. 10B +, +11 +). + + +The large and well-developed penis bulb generally occupies most of the dorso-ventral space and is composed of intermingled muscle fibres. The intrabulbar seminal vesicle is very large in several specimens, except specimen ZMHNU-JX8, and then extends from near the ventral body surface to occupy most of both the penis bulb and the dorso-ventral space of the body ( +Figs. 10C–E +, +11 +). The seminal vesicle is lined by a flat, nucleated epithelium and is surrounded by a layer of irregularly crosswise arranged muscle fibres. Broad bands of erythrophil, granular secretion anterior to the penis bulb have their glandular ducts opening into the seminal vesicle. The secretion is present in the seminal vesicle in all specimens. + + +The penis papilla is a very short and blunt cone that projects into the seminal vesicle by means of an equally stubby kind of flagellum ( +Figs. 10C–E +, +11 +). The papilla is covered with an infranucleated epithelium, which is underlain with a subepithelial layer of circular muscles, followed by a layer of longitudinal muscle fibers. + + + +FIGURE 9. + +Polycelis eudendrocoeloides + +. Photomicrographs. +A +sagittal section of ZMHNU-JX8, showing pharyngeal musculature. +B +transverse section of ZMHNU-JX10, showing ovaries. +C +sagittal section of ZMHNU-JX8, showing common oviduct and longitudinal muscles and circular muscles around bursal canal. +D +transverse section of ZMHNU-JX10, showing longitudinal muscles and circular muscles around bursal canal. +E +sagittal section of ZMHNU-JX8, showing large copulatory bursa. +F +sagittal section of ZMHNU-JX8, showing testes. Scale bars: 100 μm. + + + +The elongated male atrium is lined with a tall, papillose epithelium. It is surrounded by a thick subepithelial layer of circular muscles, followed by a thin layer of longitudinal muscles. The male atrium communicates with a very small common atrium, which is not always well discernible ( +Figs. 10C, E, F +, +11 +). + + + + +Discussion. +The anatomy of the copulatory apparatus of our specimens from the Loess Plateau closely resembles that described by +Zabusova (1929) +for + +Polycelis eudendrocoeloides + +from Kamchatka. It is immediately obvious that both the +type +specimens and the Chinese animals exhibit a feature that is unique among species of + +Polycelis + +, viz., a highly reduced and blunt penis that is provided with a kind of flagellum that projects into the seminal vesicle. Although +Zabusova (1929) +also used the term flagellum and pointed to the resemblance with species of + +Dendrocoelum + +, we did not find any indication in our specimens that this structure could be evaginated, as is the case in species of the last-mentioned genus. + + +Another noteworthy feature comparable between the +type +material of + +P. eudendrocoeloides + +and our specimens concerns the musculature of the male atrium. +Zabusova (1929) +mentioned that in the specimens from Kamchatka particularly the posterior portion of the male atrium was surrounded by a well-developed coat of circular and longitudinal muscles. Our Chinese specimens exhibited a relatively thick coat of circular and longitudinal muscles around the entire male atrium. Although this coat of muscles is well-developed, it does not yet reach the dimensions of the muscular coat surrounding the male atrium in species of + +Seidlia + +. Therefore, we agree with the classical taxonomic view that + +P. eudendrocoeloides + +belongs to the genus + +Polycelis + +s.s. + + + +FIGURE 10. + +Polycelis eudendrocoeloides + +. +A +sagittal section of ZMHNU-JX2, showing spermiducal vesicles. +B +horizontal section of ZMHNU-JX9, showing seminal vesicle, oviduct and vas deferens. +C +sagittal section of ZMHNU-JX8, showing seminal vesicle, bursal canal, male atrium and common oviduct. +D +sagittal section of ZMHNU-JX2, showing seminal vesicle, bursal canal and male atrium. +E +sagittal section of ZMHNU-JX4, showing copulatory bursa, bursal canal, male atrium and seminal vesicle. +F +sagittal section of ZMHNU-JX8, showing the thick layer of circular muscles and male atrium. Scale bars: 100 μm. + + + + +FIGURE 11. + +Polycelis eudendrocoeloides + +. Sagittal reconstruction of the copulatory apparatus of ZMHNU-JX8. Scale bar: 100 μm. + + +The dorsal surface of the animals from Kamchatka was described as pale yellow-brown. In the Chinese specimens, the dorsal surface is brownish, while the ventral surface is paler than the dorsal one, slightly light brown. + +The number of eyes in our Chinese animals ranged between 49–80 on either side of the head. +Zabusova (1929) +mentioned a considerably lower number for the +type +specimens, viz. 9–12 eyes on either side of the head. However, as already mentioned above, eye number in + +Polycelis + +is variable and generally increases during growth of an animal. The preserved specimens examined by +Zabusova (1929) +were about half the size of the live Chinese animals that we had at our disposal. This may, perhaps, signal that Zabusova’s animals were at a less advanced stage in their growth. + + +Our new finds of + +P. eudendrocoeloides + +are taxonomically and nomenclaturally interesting for several reasons. First of all, it falsifies +Kenk’s (1974) +conclusion that + +P. eudendrocoeloides + +would be a +species inquirenda +. The reasons for such a conclusion are difficult to apprehend, in view of the rather good description provided by +Zabusova (1929) +, in contrast to +Kenk’s (1953) +evaluation of it, while our new findings clearly support the species status of + +P. eudendrocoeloides + +. + + +Second, although +Kenk (1974) +considered + +P. eudendrocoeloides + +to be a +species inquirenda +, he did also designate it as the +type +species of the genus + +Sorocelides +Zabusova, 1929 + +, albeit he placed the latter under + +Polycelis + +s.l. +This taxonomic placement is supported by our examination of the new, Chinese specimens of + +P. eudendrocoeloides + +. + + +Third, according to +Dyganova & Porfirjeva (1990) +and +Kawakatsu & Mitchell (1998) +, + +P. eudendrocoeloides + +would be a junior synonym of + +Seidlia elongata +( +Zabusova, 1929 +) + +. Although the specific epithet + +eudendrocoeloides + +has page precedence (which does not imply priority), the act of +Dyganova & Porfirjeva (1990) +may be interpreted as being in accordance with the First Revisor principle ( +ICZN 1999 +, art. 24.2), so that the name + +elongata + +has priority over + +eudendrocoeloides + +. However, it is doubtful that the names are synonyms, as there are clear differences between the two species. + + +It is true that both species possess a very blunt penis papilla, which is provided with a kind of flagellum that projects into the seminal vesicle, a similarity that was already pointed out by +Zabusova-Zhdanova (1956) +. The latter doubted that this flagellum would be an eversible structure. However, +Dyganova & Porfirjeva (1990) +found that in + +S. elongata + +this flagellum-like structure that usually projects into the seminal vesicle, on occasions sits at the tip of the penis papilla as, what they called, a pseudo-flagellum (see +Dyganova & Porfirjeva 1990 +, fig. 35). Thus, apparently, it concerns an eversible flagellum. + + +However, + +S. elongata + +is different from + +P. eudendrocoeloides + +in that the former is an unpigmented species that also exhibits some notably different features in its copulatory apparatus. In particular, in + +S. elongata + +the posterior wall of the common atrium expands to form a distinct bulge towards the surrounding parenchyma (cf. +Zabusova 1929 +, fig. 9; +Dyganova & Porfirjeva 1990 +, fig. 35), which is absent in + +P. eudendrocoeloides + +. Furthermore, + +S. elongata + +is characterized by a very thick coat of muscles around the male atrium, as well as the bursal canal (cf. +Zabusova-Zhdanova 1956 +, fig. 4; + +Sluys +et al +., 2009b + +). The excessive musculature around the male atrium signals that + +S. elongata + +is indeed a member of the genus + +Seidlia + +and not of + +Polycelis + +s.s. +Although in + +P. eudendrocoeloides + +the musculature around the bursal canal is well developed, it does not reach the dimension of that in + +S. elongata +, + +in which it comprises ten layers of circular muscles, bounded by two layers of longitudinal muscles ( +Zabusova-Zhdanova 1956 +). + + +According to +Kawakatsu & Mitchell (1998) +, the following species would also be synonymous with + +S. elongata + +: + +Seidlia eurantron + +, + +S. relicta + +and + +Polycelis polyopsis +. + +That may well be true for + +P. eurantron + +as it also has a very stubby penis papilla and also shows the posterior extension at the base of the bursal canal, near its communication with the atrium. The latter feature was not specifically mentioned in the description but is apparent from the figures ( +Zabusova 1936 +, figs 7, 8). On the other hand, + +P. eurantron + +does not exhibit the flagellum-like structure pointing into the seminal vesicle that is characteristic for + +S. elongata + +. Furthermore, + +S. eurantron + +is a yellowish-brown species, whereas + +S. elongata + +is unpigmented or at least pale, while in + +S. eurantron + +the gonopore is displaced towards the posterior end of the body, in contrast to the standard position of the gonopore in + +S. elongata +( +Zabusova 1936 +) + +. + + +Although +Zabusova (1936) +considered + +S. eurantron + +to be very similar to + +S. relicta + +, the latter has a much more developed penis papilla, provided with a wide ejaculatory duct. + + +Finally, + +P. polyopsis + +is a member of the genus + +Polycelis + +s.s. +as +Zabusova (1936) +specifically described the musculature around the male atrium as being rather poorly developed, so that it cannot be synonymous with + +S. elongata + +. Furthermore, in + +P. polyopsis + +the penis papilla is conical, while its tip is curved towards the ventral body surface, in contrast to the condition in + +S. elongata + +. + + + + \ No newline at end of file diff --git a/data/03/A3/F0/03A3F02E90025A7C25B8FF5C290CF8A5.xml b/data/03/A3/F0/03A3F02E90025A7C25B8FF5C290CF8A5.xml new file mode 100644 index 00000000000..a3d2e173290 --- /dev/null +++ b/data/03/A3/F0/03A3F02E90025A7C25B8FF5C290CF8A5.xml @@ -0,0 +1,1654 @@ + + + +Review of the genus Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae) of Western Palaearctic region + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +451 +474 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.1 + +journal article +10.11646/zootaxa.5537.4.1 +1175-5326 +14240224 +46B2747F-200D-4C4A-9DAA-48CBC9FD6BBA + + + + + + + +Anthobium atrocephalum +( +Gyllenhal, 1827 +) + + + + + + + +( +Figs 1–3 +, +7–9 +) + + + + + + + +Omalium atrocephalum +Gyllenhal, 1827: 463 + + +. + + + + + + +Anthobium ruficolle +Stephens, 1834: 340 + + +. + + + + + + +Anthobium longipenne +Stephens, 1834: 342 + + +. + + + + + + +Lathrimaem cicatrix +Mulsant and Rey, 1880: 130 + + +. + + + + + + +Lathrimaeum concolor +Delahon, 1914: 621 + + +. + + + +Lathrimaem (Lathrimaeum) sardoum +Scheerpeltz 1961: 84 +, 92. + + + + +Lathrimaeum atrocephalum sardoum +: + + +Zanetti 1986: 96 + +. + + + + + +Anthobium atrocephalum + +: + +Khachikov 1998: 6 + +, + +Assing 2004: 676 + +, + +2011: 183 + +, + +Bordoni 2010: 52 + +, + +Assing & Schülke 2019: 98 + +, Khachikov +et al +. 2021: 249, + + +Salnitska +et al +. 2022: 57 + + +. + + + + + + +Lathrimaeum gracilicorne +Luze, 1905: 66 + + + +syn. n. + + + + + + +Lathrimaeum +( +Lathrimaeum +) +gracilicorne + +: + +Scheerpeltz, 1961: 83 + +. + + + + + + +Lathrimaeum abantense +Fagel, 1968: 121 + + + +syn. n. + + + + + + +Anthobium abantense + +: + +Assing 2013: 106 + +, + +2014: 1135 + +; + +Bordoni 2010: 52 + +. + + + + + + +Lathrimaeum urgelense +Fagel, 1968: 122 + + + +syn. n. + + + + +Other references see in +Shavrin (2020) +. + + + + +Type material examined. + +Holotype +of + +Lathrimaeum gracilicorne +Luze, 1905 + +( +Fig. 1 +; dissected; left side of the pronotum is slightly damaged): +‘longicorne Fv. +| Hammam R`iza | Africa bor. | leg. +Hauser’ +<handwritten>, ‘c. Eppelsh. | Steind. d.’ <printed>, ‘Type. gracili - | corne Luze’ <handwritten>, +‘atrocephalum +| Gyll. var.’ <handwritten>, ‘TYPUS’ <red, printed>, ‘ + +Anthobium + +| + +atrocephalum +( +Gyllenhal, 1827 +) + +| +Shavrin A.V. +det. 2024’ <printed> (NMW) + +. + + +Holotype +of + +Lathrimaeum abantense +Fagel, 1968 + +♂ +( +Fig. 2 +; dissected prior to the present study): [red round label], ‘Anatolie occid. | Abant Gölü | +1450 m +– +V.1967 +| G. Fagel’ <printed>, ‘TYPE’ <red printed label with black margin>, [blue round label], ‘G. Fagel det., [printed] | abantense | n. sp.’ <handwritten>, ‘ + +Anthobium + +| + +atrocephalum +( +Gyllenhal, 1827 +) + +| Shavrin A.V. det. 2024’ <printed> ( +IRSNB +). +Paratype +♂ +(dissected prior to the present study): [red round label], ‘Anatolie occid. | Abant Dagh. | +1400-1550 m +| +V.1967 +G.Fagel’ <printed>, ‘G. Fagel det. [printed] | sp. apud | atrocephalum | Gyll.’ <handwritten>, ‘G. Fagel det., [printed] | abantense | n. sp.’ <handwritten>, ‘ +Para- +| type’ <red printed label with black margin>, ‘ + +Anthobium + +| + +atrocephalum +( +Gyllenhal, 1827 +) + +| Shavrin A.V. det. 2024’ <printed> ( +IRSNB +). + + +Holotype +of + +Anthobium urgelense +Fagel, 1968 + +♂ +( +Fig. 3 +; dissected prior to the present study): ‘ +Catalogne +: Seo de | Urgel: Sierra del Cadi | +1000 m +, V-VI-1962 | G.Fagel 104 [handwritten]’ <printed>, [red small round label], ‘G.Fagel det., 1969 [69 - handwritten] | +L.atrocephalum +| ?? [handwritten] Gyll.’ <printed>, ‘G. Fagel det., | urgelense [handwritten] | n.sp [handwritten]’ <printed>, ‘TYPE’ <red printed label with black margin>, [blue small round label], ‘ + +Anthobium + +| + +atrocephalum +( +Gyllenhal, 1827 +) + +| Shavrin A.V. det. 2024’ <printed> ( +IRSNB +). +Paratypes +(all specimens dissected prior to the present study): +1 ♂ +: [red small round label], ‘ +Catalogne +: Seo de | Urgel: Sierra del Cadi | +1000 m +, V-VI-1962 | G.Fagel 109’ [handwritten] <printed>, ‘G. Fagel det., | urgelense [handwritten] | n.sp [handwritten]’ <printed>, ‘ +Para- +| type’ <red printed label with black margin>, [blue small round label] ( +IRSNB +); +1 ♂ +: [red small round label], ‘ +Espagne +| Castille | Puerto de Bejar | +800-1400 m +| +V – VI 1957 +G.Fagel’ <printed>, ‘G. Fagel det., | urgelense [handwritten] | n.sp [handwritten]’ <printed>, ‘ +Para- +| type’ <red printed label with black margin>, [blue small round label] ( +IRSNB +); +1 ♀ +: labels as previous except first printed label: ‘ +♀ +’ ( +IRSNB +). All +paratypes +with additional printed label: ‘ + +Anthobium + +| + +atrocephalum +( +Gyllenhal, 1827 +) + +| Shavrin A.V. det. 2024’. + + + +Additional material examined. +MOROCCO +: + +2 ♀♀ +: Azrou. 19- +20.05.1999 +. R. Milan leg. ( +NMPC +); + +ALGERIA +: + +21 ♂♂ +, +15 ♀♀ +: Gde Kabylie, Forêt d`Akfadou, +8 km +E Adekar. +1300 m +a.s.l. +15.05.1988 +. C. Besuchet, I. Löbl & D. Burckhardt leg. ( +MHNG +, cSh); +2 ♂♂ +, +1 ♀ +: same data, +1200 m +, +16.05.1988 +( +MHNG +); +1 ♂ +, +2 ♀♀ +: same data, +1050 m +a.s.l. ( +MHNG +); +1 ♀ +: same data, +9 km +W Adekar. +1300 m +a.s.l. +17.05.1988 +( +MHNG +); +1 ♀ +: ‘Massif des Mouzala’ ( +NMPC +); + +PORTUGAL +: + +1 ♂ +: Arrabães, Villa Real. +600 m +a.s.l. +27.07.1981 +. I. Löbl leg. ( +MHNG +); + +SPAIN +: HUESCA: + +1 ♂ +: Valle de Ordesa, +12.06.1966 +. C. Besuchet leg. ( +MHNG +); +TERUEL: +2 ♂♂ +, +3 ♀♀ +: Griegas. +1500 m +a.s.l. +21.05.1960 +. C. Besuchet leg. ( +MHNG +); +1 ♀ +: Entre Alcala de la Selva et Gudar. +1400 m +a.s.l. +22.05.1960 +. C. Besuchet leg. ( +MHNG +); +1 ♀ +: Noguera. +1550 m +a.s.l. +20.05.1960 +. C. Besuchet leg. ( +MHNG +); + +FRANCE +: ROUSSILON: + +1 ♂ +, +3 ♀♀ +: Montferrer-Corsavy. +14.04.1977 +. I. Löbl leg. ( +MHNG +); +1 ♂ +: Parcigoule. +15.4.1977 +. I. Löbl leg. ( +MNHG +); +2 ♂♂ +, +2 ♀♀ +: same data, Le Tech. ( +MHNG +); +FOIX: +1 ♀ +: Crouzette. 08.1967 ( +MHNG +); +DAUPHINÉ: +5 ♀♀ +: Vercors, Col de Limonches. +1000 m +a.s.l. +09.04.1980 +. I. Löbl leg. ( +MHNG +); +SAVOIE: +1 ♂ +: Col du Chat. +24.04.1975 +. I. Löbl leg. ( +MHNG +); +1 ♀ +: Le Bourget du Lac. +24.04.1975 +. I. Löbl leg. ( +MHNG +); +PROVENCE: +4 ♀♀ +: Uvernet. +1250 m +a.s.l. +05.06.1974 +. I. Löbl leg. ( +MHNG +); +3 ♀♀ +: Ubaye le Martinet. +03.06.1974 +. I. Löbl leg. ( +MHNG +); +1 ♂ +, +1 ♀ +: Cousson. +05.06.1970 +. Raffaldi leg. ( +MHNG +); +1 ♂ +: Gard, Mt. Aigonal. +1450 m +a.s.l. +06.05.1980 +. I. Löbl leg. ( +MHNG +); +2 ♀♀ +: Seranon. +22.07.1968 +( +MHNG +); +1 ♀ +: Col des Champs. 07.1965. ( +MHNG +); +NICE: +1 ♀ +: Sospel. 10.1962 ( +MHNG +); + +CORSICA +: + +1 ♂ +: Vizzavona. +1000 m +a.s.l. +18.07.1978 +. I. Löbl leg. ( +MHNG +); + +ITALY +: +PIEMONTE +: + +1 ♀ +: Crissolo. +11.07.1965 +( +MHNG +); + +VENETO +: + +1 ♂ +: Mt. Baldo Ferrara. +880 m +a.s.l. +31.05.1977 +. I. Löbl leg. ( +MHNG +); +1 ♂ +: Monte Grappa. +1800 m +a.s.l. 09.1969. Curti leg. ( +MHNG +); +2 ♂♂ +, +3 ♀♀ +: Fosse. +20.06.1968 +( +MHNG +); + +EMILIA-ROMAGNA +: + +2 ♂♂ +, +6 ♀♀ +: Grotte Santa Maria Madalena, Valestra. +12.06.1976 +. I. Löbl leg. ( +MHNG +); + +SARDEGNA +: + +1 ♀ +: Mt. Spada. +1100 m +a.s.l. +14.4.1977 +. S. Vit leg. ( +MHNG +); + +SERBIA +: + +10 ♂♂ +, +7 ♀♀ +: Krupanj. +450 m +a.s.l. +24.05.1984 +. C.Besuchet leg. ( +MHNG +); +6 ♂♂ +, +6 ♀♀ +: Rtanj Planina. +600 m +a.s.l. +27.05.1984 +. C.Besuchet leg. ( +MHNG +); +1 ♂ +, +2 ♀♀ +: Tara Planina, Mitrovac. +900 m +a.s.l. +23.05.1984 +. C.Besuchet leg. ( +MHNG +); +1 ♂ +: same data, +950 m +a.s.l. ( +MHNG +); +2 ♀♀ +: Stol Planina. +900 m +a.s.l. +26.05.1984 +. C.Besuchet leg. ( +MHNG +); + +UKRAINE +: +CRIMEA +: + +2 ex. +: Ay Petri ( +ZIN +); +1 ex. +: ‘Tauria, Пеньдикюль [Pendikyul`]. 11.4.14 V. Pliginski’ ( +ZIN +); 3 ex: Roman-Kosh. +22.05.1948 +. K. Arnol`di leg. ( +ZIN +); + +RUSSIA +: +STAVROPOL +TERRITORY: + +2 ♀♀ +: Zheleznovodsk, Razvalka Mt. +8.06.1985 +. A.B. Ryvkin leg. (cR); + +KRASNODAR +TERRITORY: + +1 ♂ +, +3 ♀♀ +: environs of Krasnodar. Floodplain forest, litter. +14.04.1995 +. A. Solodovnikov leg. ( +ZMUC +); +2 ♂♂ +, +1 ♀ +: Goryachiy Klyuch, gorge, north slope. +08.04.1952 +. K. Arnol`di leg. (cR., +ZMM +); + +ADYGEA +: + +1 ♂ +: Lagonaki. +1200 m +a.s.l. +14.06.1990 +. Ochrimenko leg. ( +SMNS +); +1 ♀ +: Abadzekhskaya. +25.09.1986 +. N.D. Silichenkov leg. (cSh); + +KARACHAY-CHERKESSIA +: + +1 ♂ +: Dombay. +2500 m +a.s.l. +10.10.1983 +. V. Biža leg. (cH); +2 ♀♀ +: Teberda, Dombay. 07.2010. E.A. Khachikov leg. (cKh); +2 ♂ +: Teberda State Nature Biosphere Reserve, Malaya Khatipara Mt. +1700 m +a.s.l. +30.05.1985 +. A.B. Ryvkin leg. (cR); +2 ♂♂ +, +4 ♀♀ +: same Reserve. +1.06.1985 +. A.B. Ryvkin leg. (cR, cSh); +1 ♂ +: same Reserve, Kizgich Canyon near Arkhyz. +1400 m +a.s.l., swamp with + +Carex +spp. + +, + +Juncus +sp. + +05.06.1985 +. A.B. Ryvkin leg. (cR); +5 ♂♂ +, +9 ♀♀ +: same Reserve, gorge of Gonachkhir River. +1700-1900 m +a.s.l., forest with + +Picea + +, + +Betula + +and + +Fagus + +, in litter. +01.06.1985 +. S.I. Golovatch & A.B. Ryvkin leg. (cR, cSh); + +NORTH OSSETIA-ALANIA +: + +2 ♀♀ +: Sunzhenskiy Mts., between Kardzhin and El`khotovo villages. +570 m +a.s.l., NW slope, + +Fagus + +forest with + +Rubus + +and + +Festuca + +. +03.10.1985 +. S.K. Alekseev leg. (cR); +2 ♂♂ +: Ardon River, Zintsar. +13.04.1984 +. Forest swamp. S.K. Alekseev leg. (cR); +1 ♂ +: valley of Ardok River. +26.08.1985 +. S.K. Alekseev leg. (cSh); + +DAGESTAN +: + +3 ♂♂ +, +1 ♀ +: Samur Valley, Garakh ca. +35 km +SW of Magaramkent. +700-800 m +a.s.l., + +Crataegus + +, + +Quercus + +, + +Acer +, +Rosa + +etc., scrub litter. +23.10.1987 +. S.I. Golovatch leg. (cSh); +3 ♂♂ +, +3 ♀♀ +: Dagestan. 1987. S.I. Golovatch leg. (cSh); + +ARMENIA +: +LORI +: + +11 ♂♂ +, +25 ♀♀ +: Stepanavan. +1500-1650 m +a.s.l., forest with + +Quercus + +, + +Fagus + +, + +Carpinus + +etc., litter and under bark. 21- +22.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, +ZMM +); +1 ♂ +, +1 ♀ +: Odzun W of Alaverdi. +1500-1550 m +a.s.l., forest with + +Quercus + +, + +Fagus + +, + +Carpinus + +, litter and under stones, 23- +24.07.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. ( +ZMM +); +2 ♂ +: Bazum Mt. Ridge, N slope Puskiri Pass. +1700 m +a.s.l., forest with + +Fagus + +and + +Pinus + +, litter. +22.05.1987 +. S.Golovatch & K.Yu. Es`kov leg. (cSh); + +TAVUSH +: + +5 ♂♂ +, +7 ♀♀ +: Dilijan National Park, Aknabat Valley near Salakh. +1400-1500 m +a.s.l., forest with + +Fagus + +, + +Taxus + +, + +Carpinus + +etc., in litter, under bark and stones. +17.04.1983 +. S.I. Golovatch leg. (cSh, +ZMM +); +3 ♀♀ +: same Park, Agartsyn. +1350-1400 m +a.s.l., forest with + +Fagus + +and + +Acer + +, litter. 28- +29.05.1987 +. S.I. Golovatch & K. Eskov leg. (cSh); +1 ♂ +, +3 ♀♀ +: W of Shamshadyn halfway between Idjevan and Berd. +1500-1600 m +a.s.l. 26- +27.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh); +5 ♂♂ +, +3 ♀ +: Idjevan District, Tsakhkavan. +850-900 m +, forest with + +Quercus + +, + +Acer + +, + +Carpinus + +etc., litter and tree hole. +25.07.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh); + +ARARAT +: + +19 ♂♂ +, +21 ♀♀ +: N Vartashen. +900 m +a.s.l., forest with + +Fagus + +and + +Quercus + +, in litter. +02.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, +ZMM +); + +SYUNIK +: + +2 ♂♂ +, +1 ♀ +: ca. +4 km +NNW Megri, village Lehvaz. +1000 m +a.s.l., + +Juglans + +and oak shrub with +Polyurus +and + +Rosa + +, litter and under stones. 24- +25.04.1983 +. S.I. Golovatch leg. (cSh, +ZMM +); +4 ♂♂ +, +1 ♀ +: Kapan District, Shikahogh State Reserve, vill. Shikahogh. +900-950 m +a.s.l., forest with + +Quercus + +, + +Fagus + +and + +Carpinus + +, litter and under stones. +28.04.1983 +. S.I. Golovatch leg. (cSh); +1 ♂ +: Nerkin Hand. Plane-tree grove. +30.04.1983 +. A.B. Ryvkin leg. (cR); +GEORGIA: ABKHAZIA: +1 ♂ +: W Gudauta, Myussera State Reserve. +100 m +a.s.l., in leaf litter on slope. +10.04.1983 +. S.I. Golovatch leg. ( +ZMM +); +1 ♂ +: Sukhumi, mouth of Kelasuri River. +16.09.1976 +. A.B. Ryvkin leg. (cR); +KAKHETI: +1 ♀ +: Lagodekh Nature Reserve. +500 m +a.s.l., wet forest litter near water. +06.05.1983 +. S.I. Golovatch leg. ( +ZMM +); +2 ♀♀ +: Batsara Strict State Reserve, ca +20 km +N of Akhmeta. +800-850 m +a.s.l., forest with + +Fagus + +and + +Castanea + +, litter. 05- +06.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh); +1 ♂ +: Magalakhari between Akhmeta and Tianeti. +1200 m +a.s.l., forest with + +Fagus + +and + +Carpinus + +, litter and under bark. 05.1987. S.I. Golovatch & K.Yu. Es`kov leg. ( +ZMM +); +1 ♂ +, +1 ♀ +: Mariandjvari Nature Reserve, NNE of Sagarejo. +1150-1250 m +a.s.l., forest with + +Fagus + +, + +Carpinus + +and + +Pinus + +. 13- +14.05.1987 +. SI. Golovatch & K.Yu. Es`kov leg. ( +ZMM +); +ADJARA: +1 ♀ +: +10 km +E Chakva. +07.05.1987 +. litter. S.A. Kurbatov leg. ( +RCPQ +); +MESKHETI: +1 ♀ +: +14 km +W Adigeni, flood-plain of Kvablani River, +1500 m +a.s.l., leaf litter on rill bank ( + +Alnus barbata + +,? + +Scirpus +sp. + +, + +Carex +sp. + +etc.). +15.05.1983 +. A.B. Ryvkin leg. (cR); +TBILISI: +9 ♂♂ +, +8 ♀♀ +: Tskneti. +1150-1300 m +a.s.l., forest with + +Fagus + +, + +Carpinus + +, + +Acer + +etc., litter. 16, +18.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cR); +KVEMO KARTLI: +2 ♀♀ +: Bolnisi District, Safario SSW of Patara Dinanisi. +1000-1050 m +a.s.l., forest with + +Quercus + +, + +Fagus + +and + +Acer + +, litter. +21.05.1987 +. S.I. Golovatch & K.Yu. Es1kov ( +ZMM +); +3 ♂♂ +, +5 ♀♀ +: Algeti National Park, W Manglisi. +1400-1500 m +a.s.l., forest with + +Fagus + +, + +Picea + +and + +Abies + +, litter and under bark. 16- +18.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, +ZMM +); +7 ♂♂ +, +10 ♀♀ +: Saguramo Reserve, NE of Etsketa, Zedazeni. +1100-1200 m +a.s.l., forest with + +Fagus + +, + +Carpinus + +, + +Acer + +etc., litter and under bark. +20.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, +ZMM +); + +REPUBLIC Of +SOUTH OSSETIA +: + +3 ♂♂ +, +2 ♀♀ +: Lower Roki, S of Roksky Pass. +1500 m +a.s.l., forest with + +Fagus + +, + +Betula + +etc., litter. +17.10.1987 +. S.I. Golovatch leg. (cSh); +AZERBAIJAN: NAKHCHIVAN: +1 ♂ +, +1 ♀ +: Aganos S of Lachin. +1100 m +, + +Quercus + +, + +Fagus + +, + +Carpinus + +forest, in litter. +30.04.1983 +. S.I. Golovatch (cSh, +ZMM +); +1 ♂ +: same data, Dashalty by Shusha. +1100-1300 m +a.s.l., forest with + +Quercus + +, + +Carpinus + +, litter, logs, under stones. +01.05.1983 +. (cSh); +EAST ZANGEZUR: +2 ♂♂ +: Chilisa ca. +7 km +N of Kelbadjar. +1450-1500 m +a.s.l., forest with + +Quercus + +, + +Carpinus + +, + +Acer + +, etc., litter and under bark. +31.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh); +SHAKI-ZAGATALA: +15 ♂♂ +, +21 ♀♀ +: NW above Bashlayski ca. +20 km +NNW of Sheki. +1250 m +a.s.l., forest with + +Fagus + +, + +Carpinus + +, + +Acer + +etc., litter. +03.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, +ZMM +); +9 ♂♂ +, +7 ♀♀ +: +5 km +N Kutkashen [=Gebele]. +1150-1200 m +a.s.l., forest with + +Fagus + +and + +Carpinus + +, in litter. +02.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, +ZMM +); +QUBA-KHACHMAZ: +1 ♂ +, +4 ♀♀ +: SW Kuba. +750 m +a.s.l., forest with + +Fagus + +, + +Quercus + +, + +Carpinus + +etc., litter and under bark. +23.04.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh); +MOUNTANOUS SHIRVAN: +15 ♂♂ +, +15 ♀♀ +: Shemakhi District, Pirkuli, near Observatorium. +1200-1500 m +a.s.l., forest with + +Quercus + +, + +Acer + +, + +Taxus + +etc. +30.04.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. (cSh, +ZMM +); + +IRAN +: +MAZANDARAN +: + +2 ♂♂ +, +1 ♀ +: E de Kiyasar, +36°14`N +53°33`E +. +1500 m +a.s.l. +11.07.1975 +. A. Senglet leg. ( +MHNG +); +1 ♀ +: Gol-e-Loveh. +37°18`N +55°43`E +. +1200 m +a.s.l. +21.08.1975 +. A. Senglet leg. ( +MHNG +); +CAUCASIAN REGION WITHOUT EXACT LOCALITIES: +1 ex. +: ‘Caucas’ ( +ZIN +); +1 ♀ +: ‘Kazbek’ ( +ZMM +). + + + + +Remarks. +Recently, +Shavrin (2020) +provided taxonomic and faunistic data for + +A. atrocephalum + +, widespread species in the Palaearctic region. It is known from North Africa and Europe to Eastern Siberia. The species is slightly varying in the body length, the coloration, and features of the punctation of the forebody. Some Caucasian specimens can be smaller, sometimes with paler head (usually darker in many specimens). The shape of the aedeagus is slightly variable also, especially in apical portion of the median lobe and parameres, which can be narrower ( +Figs 7–9 +) or somewhat broader ( + +Fig. +2 + +in +Shavrin (2020)) +. + + + +Lathrimaeum gracilicorne + +was originally described from “Nordafrika (Hammam R`iza…)” ( +Algeria +) based on the male +holotype +( +Fig. 1 +). I not found significant morphological differences with + +A. atrocephalum + +both in the shape of the body ( +Fig. 1 +) and the aedeagus ( +Fig. 7 +). Thus, + +A. gracilicorne + +was synonymized with it. Measurements of the +holotype +: HL: 0.40; HW: 0.65; AL: 1.24; OL: 0.17; PL: 0.57; PW: 0.90; ESL: 1.32; EW: 1.30; AW: 0.92; AedL: 0.55; TL: 2.65. + + + +Lathrimaeum abantense + +was originally described from “Anatolie occientale” based on +two specimens +. I studied both types from IRSNB. The aedeagus of the +paratype +as in +Fig. 8 +. Based on the external and internal morphological features, this species is undoubtedly belong to + +A. atrocephalum + +. Thus, I synonymized + +A. abantense + +with the latter species. Measurements of the +holotype +( +Fig. 2 +) and the +paratype +: HL: 0.41–0.45; HW: 0.66–0.72; AL: 1.35 ( +holotype +); OL: 0.25–0.30; PL: 0.52–0.60; PW: 0.96–1.07; ESL: 1.46–1.65; EW: 1.18–1.37; AW: 1.11–1.15; AedL: 0.62–0.70;TL: 3.16–3.65 ( +holotype +). + + + +Lathrimaeum urgelense + +was originally described from +Spain +based on +four specimens +. I studied all types from IRNSB. All males were dissected prior to the present study and placed to very dark Canadian balsam (?) on separate transparent plates. The shapes of the body and aedeagi are similar to that of other specimens of + +A. atrocephalum + +. Thus, I synonymized + +A. urgelense + +with the latter species. Measurements of the +holotype +( +Fig. 3 +) and +paratypes +: HL: 0.36–0.37; HW: 0.58–0.60; AL: 0.93 ( +holotype +); OL: 0.15; PL: 0.45–0.49; PW: 0.84–0.87; ESL: 1.23–1.25; EW: 1.10–1.15; AW: 0.91–0.96; AedL: 0.55–0.61;TL: 2.52–3.15 ( +holotype +: 2.67). + + + + +It is here recorded from +Morocco +, +Azerbaijan +and +Iran +for the first time. + + + + \ No newline at end of file diff --git a/data/03/A3/F0/03A3F02E90085A7125B8FC652EF2F8C1.xml b/data/03/A3/F0/03A3F02E90085A7125B8FC652EF2F8C1.xml new file mode 100644 index 00000000000..3041cfca351 --- /dev/null +++ b/data/03/A3/F0/03A3F02E90085A7125B8FC652EF2F8C1.xml @@ -0,0 +1,850 @@ + + + +Review of the genus Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae) of Western Palaearctic region + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +451 +474 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.1 + +journal article +10.11646/zootaxa.5537.4.1 +1175-5326 +14240224 +46B2747F-200D-4C4A-9DAA-48CBC9FD6BBA + + + + + + + +Anthobium melanocephalum +( +Illiger, 1794 +) + + + + + + + +( +Figs 4–5 +, +10–12 +) + + + + + + + +Silpha melanocephala +Illiger, 1794: 596 + + +. + + + + + +Lathrimaeum melanocephalum + +: + +Luze 1905: 58 + +, + +Lohse 1964: 52 + +, + +Tóth 1982: 80 + +, + +Zanetti 1987: 294 + + + + + + +Anthobium melanocephalum + +: + +Zanetti 2012: 55 + +. + + + + + + +Lathrimaeum kuehnelti +Scheerpeltz, 1963: 363 + + +. + + + + + + +Lathrimaeum hamatum +Luze, 1905: 59 + + + +syn. n. + + + + + + +Lathrimaeum hamatum + +: + +Bordoni, 1986: 43 + +, + +Khachikov 1998: 6 + +. + + + + + +Lathrimaeum +( +Lathrimaeum +) +hamatum + +: + +Scheerpeltz, 1961: 76 + +. + + + + + +Anthobium hamatum +: + + +Roubal 1913: 478 + +, + +Assing 2010: 76 + +, + +2011: 183 + +, + +2013: 106 + +, + +Bordoni 2010: 53 + +, + +Assing & Schülke 2019: 98 + +. + + + +Other references see in +Herman (2001) +. + + + + +Type material examined. + +Holotype +♂ +( +Fig. 4 +; dissected prior to the present study; abdomen slightly deformed): ‘Caucasus | Meskisches Geb.|Leder, (Reitter.)’ <printed label with black margins>, ‘c. Eppelsh.| Steind.d.’<printed>, ‘CO- [handwritten] | TYPUS’ <red, printed>, ‘ +Lathrimaeum +| +hamatum Luze +| det. BORDONI 198[printed]6 | extra xit et | defineavit’ <handwritten>, ‘HOLOTYPE’ <red, printed>, ‘ + +Anthobium + +| + +melanocephalum +( +Illiger, 1794 +) + +| +Shavrin A.V. +det. 2024’ <printed> ( +NMW +) + +. + + + +Additional material examined. +SPAIN +: +ÁVILA +: + +2 ♂♂ +: Sierra de Gredos. +15.06.1971 +. M. Curti leg. ( +MHNG +); + +FRANCE +: +AUVERGNE +: + +1 ♂ +: Le Mont-Dore. +25.09.1964 +( +MHNG +); + +LANGUEDOC +: + +1 ♀ +:Mt.Ventoux Vauce. +15.06.1963 +( +MHNG +); + +PROVENCE +: + +1 ♂ +: Mougins/Canne. +29.03.1986 +. C. Besuchet leg. ( +MHNG +); +2 ♀♀ +: Séranon. +01.05.1963 +( +MHNG +); +1 ♀ +: Lambruisse. B.A. +19.10.1966 +( +MHNG +); + +NICE +: + +1 ♂ +: Aopromonte Gambari +18.06.1976 +. M. Curti ( +MHNG +); + +ITALY +: +PIEMONTE +: + +1 ♂ +: Crissolo. 07.1965 ( +MHNG +); + +TURKEY +: +BURSA +: + +1 ♀ +: Uludað, +17 km +de Bursa. +1100 m +a.s.l. +13.05.1976 +. C. Besuchet & I. Löbl leg. ( +MHNG +); + +BOLU +: + +1 ♀ +: Ömerler. +800 m +a.s.l. +21.05.1976 +. C. Besuchet & I. Löbl leg. ( +MHNG +); +2 ♂♂ +: +9 km +Mengen. +750 m +a.s.l. +23.05.1976 +. C. Besuchet & I. Löbl leg. ( +MHNG +); +1 ♂ +: Abant. +1450 m +a.s.l. +25.05.1967 +. C. Besuchet leg. ( +MHNG +); +2 ♀♀ +: same data. +22.05.1976 +. +1500-1600 m +a.s.l. C. Besuchet & I. Löbl leg. ( +MHNG +); +1 ♂ +: Abant. +1450 m +a.s.l. +25.05.1967 +. C. Besuchet leg. ( +MHNG +); + +KASTAMONU +: + +1 ♂ +: +5 km +S Kürea. +1200 m +a.s.l. +18.05.1976 +. C. Besuchet & I. Löbl leg. ( +MHNG +); +1 ♂ +: Inebolu-Küre. +700 m +a.s.l. +18.05.1976 +. C. Besuchet & I. Löbl leg. ( +MHNG +); +1 ♀ +: Ilgardağ Diphan. +1300 m +a.s.l. 05.1976. C. Besuchet & I. Löbl leg. ( +MHNG +); +2 ♀♀ +: Inebolu-Küre. +700 m +a.s.l. +18.05.1976 +. C. Besuchet & I. Löbl leg. ( +MHNG +); +1 ♀ +: +13 km +E Aðli. +1200 m +a.s.l. +18.05.1976 +. C. Besuchet & I. Löbl leg. ( +MHNG +); + +SINOP +: + +1 ♀ +: Bürnük, +29 km +N Boyabat. +1250 m +a.s.l. +20.05.1976 +. C. Besuchet & I. Löbl leg. ( +MHNG +); + +ARTVIN +: + +1 ♀ +: SW Artvin. +1900 m +a.s.l. +09.06.1986 +. C. Besuchet, I. Löbl & D. Burckhardt leg. ( +MHNG +); +1 ♂ +: Pirnalli, Karkal Daði. +1250 m +a.s.l. +11.06.1986 +. C. Besuchet, I. Löbl & D. Burckhardt leg. ( +MHNG +); + +RUSSIA +: +ADYGEA +: + +1 ♀ +: Abadzekhskaya, “Lesnaya Skazka”. 05.2009. V. Bibin leg. (cKh); +2 ♂♂ +, +1 ♀ +: Abago Greenland, +11 km +Guzeripl`-Abago road. +06.10.2015 +. E.A. Khachikov leg. (cKh, cSh); + +KARACHAY-CHERKESSIA +: + +1 ♂ +: Teberda State Nature Biosphere Reserve, Malaya Khatipara Mt. +1900 m +a.s.l. +29.05.1985 +. A.B. Ryvkin leg. (cR); +1 ♂ +: +9 km +SW Teberda, Teberdinskiy Range, +43°22´40´´N +41°39´38´´E +. +2000 m +a.s.l. +27.07.2011 +. V. Assing leg. ( +NMW +); +1 ♀ +: +13 km +SW Teberda, +43°19´54´´N +41°39´58´´E +. +1450 m +a.s.l., spruce forest. +22.07.2011 +. V. Assing leg. ( +NMW +); + +NORTH OSSETIA-ALANIA +: + +2 ♀♀ +: Tseiskoye Gorge. Forest with + +Pinus + +and + +Carex + +. 1982. S.K. Alekseev leg. (cR); +1 ♂ +: North Ossetia Nature Reserve, basin of Ardon River. 1982. S.K. Alekseev leg. (cSh); + +DAGESTAN +: + +1 ♀ +: ‘ +♀ +’, ‘Daghestan Leder. Reitter.’, ‘Type. hamatum Luze’, ‘ex coll. Luze’, ‘TYPUS +Lathrimaeum hamatum +Luze’ ( +NMW +); + +ARMENIA +: + +1 ♀ +: ‘Caucasus. Armen. Geb. Leder. Reitter’, ‘coll. Reitter’, ‘Type. Hamatum Luze’, ‘ +Allotypus +1905 +Lathrimaeum hamatum +Luze’, ‘ +Lathrimaeum hamatum +det. Székessy’ ( +HNHM +); + +GEORGIA +: +SVANETI +: + +1 ♂ +: ‘Caucasus. Swanetien. Leder. Reitter.’, ‘coll. Reitter’, ‘Type. hamatum Luze’, ‘ +Paratypus +1905 +Lathrimaeum hamatum +Luze’ ( +HNHM +); +2 ♀♀ +: ‘Caucasus. Swanetien. Leder. Reitter.’, ‘ +Lathrimaeum hamatum Luze Coll. +Reitter’ ( +HNHM +); +3 ♂♂ +, +7 ♀♀ +: +30 km +NW Khaishi, near Kherkhvashi. +1750-1800 m +a.s.l., eutrophic-mesotrophic swamp ( + +Carex +spp. + +, + +Scirpus + +and mosses). +21.09.1986 +. A.B. Ryvkin leg. (cR, cSh); + +RACHA +: + +2 ♂♂ +, +1 ♀ +: Racha road E Ambrolauri [13], +43°31´04´´N +43°19´10´´E +. +1530 m +a.s.l., litter sifted. +27.07.2021 +. V. Assing leg. ( +NMW +); + +GURIA +: + +1 ♀ +: SE Ozurgeti [73], Gomismta, +41°49´57´´N +42°09´23´´E +. +1880 m +a.s.l., litter sifted. +07.08.2022 +. V. Assing leg. ( +NMW +); + +IMERETI +: + +1 ♂ +: SE Sairme [42], +41°50´15´´N +42°49´27´´E +. +2290 m +a.s.l., pass, litter sifted. +20.07.2019 +. V. Assing leg. ( +NMW +); +2 ♀♀ +: Imereti pass [69]. +25 km +SE Sachkhere, +42°09´54´´N +43°35´44´´E +. +1190 m +a.s.l., litter sifted. +23.10.2021 +. V. Assing leg. ( +NMW +); + +SAMTSKHE-JAVAKHETI +: + +2 ♀♀ +: ‘Caucasus Meskisches Geb. Leder, (Reitter)’, ‘ +Lathrimaeum hamatum Luze Coll. +Reitter’ ( +HNHM +); + +KVEMO KARTLI +: + +1 ♂ +: Algeti National Park, W Manglisi, +1400-1450 m +a.s.l., forest with + +Fagus + +, + +Picea + +, + +Abies + +etc., litter and under bark. 16- +18.05.1987 +. S.I. Golovatch & K.Yu. Es`kov leg. ( +ZMM +); + +TUSHETI +: + +1 ♀ +: ‘Caucasus Tbatani [18]79 Leder, (Reitter)’, ‘ +Lathrimaeum hamatum Luze Coll. +Reitter’ ( +HNHM +); + +AZERBAIJAN +: +LANKARAN-ASTARA +: + +1 ♂ +, +2 ♀♀ +: Yardimli District, Avash. +1200-1500 m +a.s.l. 14- +17.06.1996 +. S.I. Golovatch leg. (cSh); + +CAUCASIAN REGION WITHOUT EXACT LOCALITIES +: + +1 ♀ +: ‘Caucas.occ. Circassien Leder.Reitter’, ‘ +Lathrimaeum hamatum Luze Coll. +Reitter’ ( +HNHM +); + + +Besides that I studied +13 specimens +of “ + +A +. +hamatum + +” from NMW, label`s data of which were published by +Assing (2011 +, +2013 +) from +Turkey +and by +Assing & Schulke (2019) +from +Armenia +respectively. + + + + +Remarks. + +Anthobium melanocephalum + +is widely distributed in Europe. Besides that, it was recorded from +Turkey +by +Assing (2010 +, +2011 +). Turkish records of + +A melanocephalum + +by +Assing (2004) +refer to + +A. metallicum +( +Assing 2013 +) + +. +Eppelsheim (1881) +published the first record of + +A +. +melanocephalum + +from Caucasus (“Tbatani”). + + + +Lathrimaeum hamatum + +was originally described based on +two specimens +from “Meskisches Gebirge” (male) and “Armenisches Gebirge” (female). Later, it was recorded from many localities of Caucasus and +Turkey +. During the study of the type material ( +holotype +as in +Fig. 4 +) and additional specimens of + +A. melanocephalum + +from Europe, +Turkey +and Caucasus, I not found significant morphological differences between them. Some specimens from Caucasus and +Turkey +has paler (yellow-brown to brown) body and slightly vary in features of punctation of the head and the pronotum. Besides that, some specimens have more pronounced mediobasal impression on the pronotum. Specimens from the Middle Europe are usually larger, darker (reddish brown, with almost black head), and with more regular punctation on the pronotum. The shape of the aedeagus is slightly variable ( +Figs 10–12 +) if compared with aedeagi of specimens from Europe (Fig. 34f in +Zanetti (2012)) +. Thus, I synonymized + +A. hamatum + +with + +A. melanocephalum + +. Habitus as in +Figs 4–5 +. Measurements of specimens of + +A +. +melanocephalum + +from +Turkey +and Caucasus (n=50): HL: 0.41–0.44; HW: 0.66–0.70; AL: 1.20 ( +holotype +of + +A. hamatum + +); OL: 0.16–0.20; PL: 0.61– 0.70; PW: 1.16–1.20; ESL: 1.61–1.71; EW: 1.46–1.52; AW: 0.98–1.25; MTbL: 0.69 ( +holotype +of + +A. hamatum + +); MTrL ( +holotype +of + +A. hamatum + +): 0.35 (MTrL 1–4: 0.18; MTrL 5: 0.17); AedL: 0.50–0.77; TL: 3.20–4.07 ( +holotype +of + +A. hamatum + +: 4.04). It is here recorded from +Azerbaijan +for the first time. + + + + \ No newline at end of file diff --git a/data/03/A3/F0/03A3F02E900D5A6925B8FC662C46FE1D.xml b/data/03/A3/F0/03A3F02E900D5A6925B8FC662C46FE1D.xml new file mode 100644 index 00000000000..3daa070c2ba --- /dev/null +++ b/data/03/A3/F0/03A3F02E900D5A6925B8FC662C46FE1D.xml @@ -0,0 +1,493 @@ + + + +Review of the genus Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae) of Western Palaearctic region + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +451 +474 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.1 + +journal article +10.11646/zootaxa.5537.4.1 +1175-5326 +14240224 +46B2747F-200D-4C4A-9DAA-48CBC9FD6BBA + + + + + + + +Anthobium tenue +( +Eppelsheim, 1881 +) + + + + + + + +( +Figs 21 +, +26–27 +, +32 +) + + + + + + + +Lathrimaeum tenue +Eppelsheim, 1881: 507 + + +; + +Luze 1905: 60 + +, + +Bordoni 1986: 43 + +. + + + + + +Lathrimaeum +( +Lathrimaeum +) +tenue + +: + +Scheerpeltz 1961: 80 + +. + + + + + +Anthobium tenue + +: + +Assing & Schülke 2019: 98 + +. + + + + + +Type material examined. + +Lectotype +of + +Lathrimaeum tenue +Eppelsheim, 1881 + +( +here designated +) +♂ +( +Fig. 21 +; dissected prior to the present study): ‘ +Kaukas +| Leder | Suram [handwritten]’ <printed>, ‘c. Eppelsh. | Steind. d.’ <printed>, ‘TYPUS’ <red, printed>, +‘tenue Epp. +| det. [printed] Luze’ <handwritten>, ‘LECTOTYPE | + +Lathrimaeum + +| + +tenue +Eppelsheim, 1881 + +| Shavrin A.V. des. 2024’ <printed>, ‘ + +Anthobium + +| + +tenue +( +Eppelsheim, 1881 +) + +| +Shavrin A.V. +det. 2024’ <printed> ( +NMW +) + +. + +Paralectotype +♀ +: ‘Suram’ <handwritten>, ‘ +Kaukas +| Leder’ <printed>, ‘coll. Reitter’ <printed>, ‘Holotypus [printed in red] 1880 | +Lathrimaeum +| tenue | Eppelsheim’ <handwritten label with red margins>, ‘ +Lathrimaeum +| +tenue Epp. +| det. Székessy [printed]’ <handwritten>, ‘ + +Anthobium + +| + +tenue +( +Eppelsheim, 1881 +) + +| +Shavrin A.V. +det. 2024’ <printed> ( +HNHM +) + +. + + + +Additional material examined. +RUSSIA +: +KARACHAY-CHERKESSIA +: + +1 ♀ +: Teberda State Nature Biosphere Reserve, Malaya Khatipara Mt. +1700 m +a.s.l. +30.05.1985 +. A.B. Ryvkin leg. (cR); +1 ♀ +: same Reserve. +1.06.1985 +. A.B. Ryvkin leg. (cSh); + +GEORGIA +: SVANETI: + +1 ♀ +: ‘Caucasus Swanetien Leder Reitter’, ‘ +Lathrimaeum tenue Epp. Coll. +Reitter’ ( +HNHM +); +1 ♀ +: ‘Caucasus. Swanetien. Leder. Reitter’, ‘352’, +‘tenue Epp. +det. Luze’ ( +NMW +); +1 ♂ +: ‘ +♂ +’, ‘Caucasus. Swanetien. Leder. Reitter’, +‘tenue Epp. +det. Luze’, ‘ex. coll. Skalitzky’ ( +NMW +); +1 ♂ +: ‘ +♂ +’, ‘Caucasus. Swanetien. Leder. Reitter’, +‘tenue Epp. +det. Luze’, ‘ex. coll. Scheerpeltz’ ( +NMW +); + +SHIDA KARTLI +: + +2 ♂♂ +, +4 ♀♀ +: Surami (=Ricoti) pass, ca. +1000 m +a.s.l., forest with + +Fagus + +, + +Alnus + +, + +Castanea + +and + +Rhododendron + +, litter and under stones. 14.04, +17.05.1983 +. S.I. Golovatch leg. (cR, cSh, +ZMM +); + +SAMTSKHE-JAVAKHETI +: + +1 ♂ +, +1 ♀ +: ‘Caucasus Meskisches Geb. Leder, (Reitter)’, ‘ +Lathrimaeum tenue Epp. Coll. +Reitter’ ( +HNHM +); + +SAMTSKHE-JAVAKHETI +: + +1 ♂ +: ‘Caucasus Abastuman Leder, (Reitter) +Lathrimaeum tenue Epp. Coll. +Reitter’ ( +HNHM +); +CAUCASIAN REGION WITHOUT EXACT LOCALITIES: +1 ♂ +, +1 ♀ +: ‘Caucasus Circassien Leder. Reitter +Lathrimaeum tenue Epp. Coll. +Reitter’ ( +HNHM +); +2 ♂♂ +: ‘Caucasus Leder.Reitter’, ‘ +Lathrimaeum tenue Epp. Coll. +Reitter’ ( +HNHM +); +1 ♀ +: ‘Kaukas Leder,’, ‘c. Eppelsh. Steind. d.’, +‘tenue Epp. +det. Luze’, ‘TYPUS’ ( +NMW +); +1 ♀ +: ‘Kaukas Leder’, ‘c. Eppelsh. Steind. d.’, +‘tenue Eppelsh. Vrch. +zool. bot. Ges. Wien, 1880, p. 507’, ‘TYPUS’ ( +NMW +); +1 ♂ +, +1 ♀ +: ‘Kaukas Leder’ ( +HNHM +); +1 ♀ +: ‘ +♀ +’, ‘Kaukas Leder’, ‘Reitter’, +‘tenue Epp. +’, ‘355’, ‘ex. coll. Skalitzky’ ( +NMW +). + + + + +Redescription. +Measurements (n=20): HL: 0.37–0.45; HW: 0.64–0.72; AL: 1.45 ( +holotype +); OL: 0.11–0.12; PL: 0.61–0.63; PW: 1.14–1.27; ESL: 1.29–1.44; EW: 1.30–1.60; AW: 1.27–1.50; MTbL: 0.67 ( +holotype +); MTrL ( +holotype +): 0.34 (MTrL 1–4: 0.22; MTrL 5: 0.12); AedL: 0.70–0.72; TL: 2.60–3.70 ( +lectotype +: 3.35). + + + +FIGURES 19–23. +Habitus of + +Lathrimaeum + +(types) and + +Anthobium + +: 19– + +L. moczarskii + +(holotype), 20– + +L +. +metallicum + +(holotype), 21– + +L +. +tenue + +(lectotype), 22– + +L +. +laterale + +(holotype), 23– + +A +. +ganglbaueri + +(Turkey, Ulu Dað). Scale bar: 1.0 mm. + + + + +FIGURES 24–30. +Aedeagus of + +Anthobium + +(Figs 24–25: + +A. moczarskii + +(Spain, Pinilla del Valle), Figs 26–27: + +A. tenue + +(Georgia, Surami Pass), Fig. 28: + +A. ganglbaueri + +(Turkey, Bademli), Figs 29–30: + +A. ganglbaueri + +(Turkey, Kastamonu): 24, 26, 28–29– parameral view, 25, 27, 30–lateral view. Scale bar: 0.1 mm. + + + +Habitus as in +Fig. 21 +. Body yellow-brown to reddish-brown (some specimens with slightly darker head and abdomen); antennomeres 3–11 or 4–11 brownish (paler specimens with yellowish antennomeres); mouthparts, antennomeres 1–2 or 1–3 and legs yellow to yellow-brown. Head with transverse fine microreticulation in frontal portion, indistinct subdiagonal meshes in middle and dense, fine, transverse or subdiagonal microreticulation on laterobasal portions (some specimens without visible sculpture in middle). Head with moderately dense and irregular punctation, finer and sparser in middle, denser and sometimes deeper on infraorbital portion and between ocelli; neck with dense, regular and moderately fine punctation; pronotum with very dense, moderately fine punctation, with interspaces between punctures in middle about as long as diameters of one-two nearest punctures, usually finer and sparser in mediobasal portion; punctation of elytra about as that on pronotum, but sometimes slightly coarser and deeper in mediobasal part. + + + +FIGURE 31. +Distribution of + +Anthobium moczarskii + +in Iberian Peninsula. + + +Head 1.6–1.7 times as broad as long, with slightly or strongly elevated median portion; anteriomedian depressions moderately wide and deep, connected with relatively deep lateral depressions; postocular ridges acute, with interspace between posterior margin of eyes and ridge about as long as three nearest ommatidia. Ocelli moderately large, located at level or slightly below level of postocular ridges. Antennomere 4 distinctly shorter than 3, 5–7 slightly longer and indistinctly broader than 4, 8 slightly shorter and broader than 7, 9–10 slightly broader than 8. +Pronotum 1.8 times to twice as broad as long, from widest middle gradually narrowed both anteriad and posteriad; posterior angles obtuse; apical angles slightly protruded anteriad; middle portion usually without longitudinal elevation except of some specimens with shallow traces of it, mediobasal depression missing or indistinct. +Elytra 1.1 times as long as broad, slightly more than twice as long as pronotum, reaching apical margin of abdominal tergites V–VII, slightly broadened posteriad; middle surface of each elytron usually with irregular longitudinal elevations between punctures. + +Male. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII sinuate. Aedeagus with very wide basal portion, gradually narrowed toward wide truncate apex; parameres indistinctly exceeding apex of median lobe, slightly broadened in preapical portions, with four small apical setae; internal sac long and moderately narrow, spirally folded in basal portion, with wide field of acute spines in middle part ( +Fig. 26 +). Lateral aspect of the aedeagus as in +Fig. 27 +. + +Female. Apical margins of abdominal tergite VIII and sternite VIII truncate. + +Comparative notes. +Based on the general shape, the coloration and the length of the body, + +A. tenue + +is similar to widespread + +A. unicolor + +, from which can be distinguished by more transverse pronotum with significantly more protruded apical angles, the shape of the broader parameres and details of the structure of the internal sac. + + + + +Distribution. + +Anthobium tenue + +is known from several localities in Caucasus ( +Fig. 32 +). + + +Bionomics. +The detailed bionomical data for the most studied specimens is unknown. Some specimens were collected at elevations from +1000 to 1700 m +a.s.l. and were sifted from litter of deciduous forests. + + + + +Remarks. + +Lathrimaeum tenue + +was originally described from “Suramgebirge” based on “Wenige Stücke”. During the study of collections of NMW and HNHM I found +two specimens +collected in “Suram”. Thus, the male from NMW was designated as the +lectotype +in order to fix the indentity of the name. It is here recorded from +Russia +for the first time. + + + + \ No newline at end of file diff --git a/data/03/A3/F0/03A3F02E90115A6A25B8FD8D2E2DFE60.xml b/data/03/A3/F0/03A3F02E90115A6A25B8FD8D2E2DFE60.xml new file mode 100644 index 00000000000..7b07472e251 --- /dev/null +++ b/data/03/A3/F0/03A3F02E90115A6A25B8FD8D2E2DFE60.xml @@ -0,0 +1,636 @@ + + + +Review of the genus Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae) of Western Palaearctic region + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +451 +474 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.1 + +journal article +10.11646/zootaxa.5537.4.1 +1175-5326 +14240224 +46B2747F-200D-4C4A-9DAA-48CBC9FD6BBA + + + + + + + +Anthobium unicolor +( +Marsham, 1802 +) + + + + + + + +( +Fig. 22 +) + + + + + + + +Silpha unicolor +Marsham, 1802: 127 + + +. + + + + + +Anthobium unicolor + +: + +Stephens 1834: 343 + +; + +Assing 2004: 667 + +, + +2005: 57 + +; + +Assing & Schülke 2019: 98 + +. + + + + + +Lathrimaeum unicolor + +: + +Stephens 1839: 426 + +, +Fauvel 1871 +a: 119 [= 1872: 93], + +Ganglbauer 1895: 722 + +, + +Luze 1905: 61 + +, + +Reitter 1909: 188 + +, + +Johansen 1914: 596 + +, + +Portevin 1929: 441 + +, + +Palm 1948: 101 + +, + +Lohse 1964: 52 + +, + +Coiffait 1978: 275 + +, + +Tóth, 1982: 80 + +. + + + + + +Olophrum unicolor + +: + +Stephens, 1839: 427 + +. + + + + + +Lathrimaeum +( +Lathrimaeum +) +unicolor + +: + +Scheerpeltz, 1961: 81 + +. + + + + + + +Anthobium assimile +Stephens, 1834: 342 + + +. + + + + + + +Lathrimaeum luteum +Erichson, 1840: 869 + + +; G. + +Waterhouse, 1858: 31 + +. + + + + + + +Lathrimaeum diabolicum +Luze, 1905: 61 + + +. + + + + + + +Lathrimaeum roubali +Luze, 1911: 293 + + +; + +Roubal 1911: 2 + +. + + + + + + +Lathrimaeum laterale +Luze, 1911: 294 + + +. + + + + + + +Lathrimaeum deinekini +Roubal, 1913: 479 + + + +syn. n. + + + + +Other references see in +Herman (2001) +. + + + + +Type material examined. + +Holotype +of + +Lathrimaeum laterale +Luze, 1911 + +♂ +( +Fig. 22 +; left antennomeres 10–11 missing): ‘♂’ <printed>, ‘ +Caucasus +occ. | КраснаЯ ПолЯна | Roubal +VII. 1910 +’ <blue, printed>, ‘Type. | laterale m. | Luze’ <handwritten>, ‘ex. coll. | Luze’ <yellow, printed>, ‘TYPUS [printed] | +Lathrimaeum +| laterale | Luze.’ <red, handwritten>, ‘Holotypus | +Lathrimaeum +| +laterale Luze +| rev. V. +Assing 2004 +’ <printed>, ‘ +Anthobium +| unicolor | (Marsham) | det. + +V. +Assing 2004 + +’ ( +NMW +) + +. + + + +Paralectotype +of + +Lathrimaeum roubali +Luze, 1911 + +♂ +: ‘ +Caucasus +occ. | КраснаЯ ПолЯна [Krasnaya Polyana] Roubal + +VII. 1910 + +’ <blue, printed>, ‘Type | Lathri- | maeum Roubali Luze’ <red, printed> ( +SNMB +) + +. + + +Lectotype +( +here designated +) of + +Lathrimaeum deinekini +Roubal, 1913 + +♂ +: ‘Caucasus bor. | Teberda’ <printed>, ‘ +Lathrimaeum +| deineckini Roub.’ <handwritten>, ‘ +LECTOTYPE +| + +Lathrimaeum + +| + +deinekini +Roubal, 1913 + +| Shavrin A.V. des. 2014’ <red, printed>, ‘ +Anthobium +| + +unicolor ( +Marsham, 1802 +) + +| Shavrin A.V. det. 2024’ ( +NMPC +). +Paralectotypes +: +2 ♀♀ +: ‘Ca. b. Teberda | VI. 912 Roubal’ <yellow, printed>, ‘Cotypus’ <handwritten>, +‘deinekini +[handwritten] Roubal det’ <printed> ( +NMPC +); +1 ♂ +: ‘Ca. b. Teberda VI. 912 Roubal’ <printed>, ‘ +Lathrimaeum Deinekini +type Roubal det.’ <handwritten>, ‘Syntype’ <red, printed> ( +ZIN +); +1 ex. +: ‘Ca. b. Teberda VI. 912 Roubal’ <printed>, ‘TYPUS’ <yellow, printed>, ‘Deineckini Roub.’ <handwritten>, ‘Coll. Dr. Lgocki’ <printed> ( +ZIN +). +One paralectotype +deposited in the collection of MACN, Buenos Aires ( + +Bachmann +et al +. 2017 + +); photographs of the habitus and labels as in + +Figs +14–15 + +in + +Bachmann +et al +. (2017) + +. + + +Additional material examined. + + +SPAIN +: +HUESCA +: + +1 ♀ +: +Barranco de Izas +. + +1580 m + +a.s.l.. + +19.06.2005 + +. +A. Anichtchenko +leg. (cSh) + +; + + +RUSSIA +: +KRASNODAR +TERRITORY: + +2 ♂♂ +, +1 ♀ +: +Temnolesskoye +. + +04.05.1999 + +. +V. Grebennikov +leg. (cKh, cSh); 1 ex.: Aderbievka, Ostraya Mt. +20.04.1952 +. K. Arnol`di leg. (ZIN); + + + +ADYGEA +: + +1 ex. +: +Azovskaya +. + +11.04.1950 + +. +K. Arnol`di +leg. ( +ZIN +); + + +1 ♂ +: +Lagonaki +. +D. Fominykh +leg. (cSh); + + +4 ♂♂ +, +5 ♀♀ +: +Abadzekhskaya St. +, “Lesnaya Skazka”. 05.2009. +V. Bibin +leg. (cKh, cSh); + + + +KARACHAY-CHERKESSIA +: + +2 ♂ +: +Teberda State Nature Biosphere Reserve, Dombay +. + +1700 m + +, pebble bank of Amanaus River, near edge of forest. + +31.05.1985 + +. +P. K. Yeremin +leg. (cSh); + + + +GEORGIA +: +SHIDA KARTLI +: + +1 ♀ +: Surami (=Ricoti) pass, ca + +100 m +a.s.l. + +, forest with + +Fagus + +, + +Alnus + +, + +Castanea + +, + +Rhododendron + +, litter and under stones. 14.04, + +17.05.1983 + +. +S.I. Golovatch +leg. ( +ZMM +); + + + +KAKHETI +: + +1 ♀ +: +Mariandjvari State Reserve +, NNE Sagarejo. + +1150-1250 m +a.s.l. + +, forest with + +Fagus + +, + +Carpinus + +, + +Acer + +, + +Pinus + +etc. 13- + +14.05.1987 + +. +S.I. Golovatch +& +K.Yu. Es`kov +leg. (cSh) + +. + + + + +Remarks. + +Lathrimaeum roubali + +and + +L. laterale + +were originally described from “Kaukasus occid. (Krasnaja Poljana)”. +Assing (2004 +, +2005 +) synonymized both species with + +A. unicolor + +. The +holotype +of + +A. laterale + +as in +Fig. 22 +. Based on an unspecified number of +syntypes +, +Roubal (1913) +described + +L. deinekini + +from “Teberda”. The male from NMPC I designate here as the +lectotype +. During the study of specimens of + +A. deinekini + +, I didn`t find morphological differences between them and studied specimens of + +A. unicolor + +from different regions. Thus, + +A. deinekini + +was synonymized with the latter species. + +Anthobium unicolor + +is widely distributed in Europe ( +Herman 2001 +), including Caucasus. It is here recorded from +Georgia +for the first time. + + + + \ No newline at end of file diff --git a/data/03/A3/F0/03A3F02E90135A6B25B8FF5C2E6EFA90.xml b/data/03/A3/F0/03A3F02E90135A6B25B8FF5C2E6EFA90.xml new file mode 100644 index 00000000000..a21c0bbb172 --- /dev/null +++ b/data/03/A3/F0/03A3F02E90135A6B25B8FF5C2E6EFA90.xml @@ -0,0 +1,636 @@ + + + +Review of the genus Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae) of Western Palaearctic region + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +451 +474 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.1 + +journal article +10.11646/zootaxa.5537.4.1 +1175-5326 +14240224 +46B2747F-200D-4C4A-9DAA-48CBC9FD6BBA + + + + + + + +Anthobium ganglbaueri +( +Luze, 1905 +) + + + + + + + +( +Figs 23 +, +28–30 +) + + + + + + + +Lathrimaeum ganglbaueri +Luze, 1905: 65 + + +; + +Rambousek 1909: 5 + +; + +Tóth, 1982: 81 + +. + + + + + +Lathrimaeum +( +Lathrimaeum +) +ganglbaueri + +: + +Scheerpeltz, 1961: 83 + +. + + + + + +Anthobium ganglbaueri +: + + +Assing & Wunderle 2001: 105 + +, + +Smetana 2004: 238 + +. + + + + + + +Lathrimaeum knappei +Scheerpeltz, 1961: 89 + + +. + + + + + + +Lathrimaeum anatolicum +Fagel, 1968: 118 + + +; + +Bordoni 1986: 44 + +. + + + + + +Anthobium anatolicum + +: + +Assing 2004: 676 + +, + +2009: 121 + +, + +2013: 106 + +, + +2015: 16 + +; + +Bordoni 2010: 52 + +. + + + +Other references see in +Herman (2001) +. + + + + +Material examined. + + +SERBIA +: + +1 ♂ +, +2 ♀♀ +: +Majdanpek +, +44°25´45´´N +21°57´08´´E +. + +07.05.2015 + +. +V. Zieris +leg. (cSh) + +; + + +ROMANIA +: + +1 ♀ +: +Mehedinţi +, Motru Sec., +Lazului. +09- + +10.05.2015 + +. +V. Zieris +leg. (cH) + +; + + +GEORGIA +: +ADJARA +: + +1 ♂ +: NE Khulo, +41°42´17´´N +42°21´49´´E +. + +1120 m +a.s.l. + +, E slope with + +Corylus + +. + +12.07.2019 + +. +V. Assing +leg. ( +NMW +) + +; + + +TURKEY +: +BURSA +: + +1 ♀ +: +Ulu Dað +, +20 km +Bursa. + +1300 m +a.s.l. + + +12.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); 1 ♂: Ulu Dað, 1300 m a.s.l. +02.05.1995 +. H. Enghoff leg. (ZMUC); + + + +BOLU +: + +2 ♂♂ +, +1 ♀ +: Ömenter pres Bolu. + +800 m +a.s.l. + + +21.05.1976 + +. +C. Besuchet +leg. ( +MHNG +); + + +4 ♂♂ +, +5 ♀♀ +: +Abant +. + +1500-1600 m +a.s.l. + + +22.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); + + +3 ♂♂ +, +3 ♀♀ +: +9 km N Mengen. + +750 m +a.s.l. + + +23.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); + + + +ZONGULDAK +: + +4 ♂♂ +, 8 f ##: +17 km N Safranbolu. + +1000 m +a.s.l. + + +16.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); + + + +KASTAMONU +: + +10 ♂♂ +, +14 ♀♀ +: Küre 5 km N. + +600 m +a.s.l. + + +18.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); + + +3 ♀♀ +: +13 km E Aðli. + +1200 m +a.s.l. + + +18.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); + + +1 ♂ +: +Ilgardağ Diphan +. + +1300 m +a.s.l. + + +17.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); + + + +SINOP +: + +1 ♂ +: Bürnük, +29 km N Boyabat. + +1250 m +a.s.l. + + +20.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); + + +5 ♂♂ +, +1 ♀ +: +Bektras +, +23 km N Boyabat. + +1100 m +a.s.l. + + +20.05.1976 + +. +C. Besuchet +& +I. Löbl +leg. ( +MHNG +); + + + +SAMSUN +: + +2 ♂♂ +, +2 ♀♀ +: +Samsun-Kavak +. + +20.05.1967 + +. +C. Besuchet +leg. ( +MHNG +); + + + +ARTVIN +: + +2 ♂♂ +: +Pirnalli +, Karkal Daðli. + +1250 m +a.s.l. + + +11.06.1986 + +. +C. Besuchet +, +I. Löbl +& +D. Burckhardt +leg. ( +MHNG +); + + + +ANTALYA +: + +1 ♂ +: +Bademli +, Bademli geçidi (ca + +30 km +N of Akseki + +). 02- + +03.05.2001 + +. +R. Lohaj +leg. (cSh) + +. + + +Besides that I studied +111 specimens +( +NMW +) data of which were published by +Assing (2004 +, +2009 +, 213) and +Assing & Wunderle (2001) +from +Turkey +and +Greece +respectively. + + + + +Remarks. + +Lathrimaeum ganglbaueri + +was originally described from “Südungarn (Herkulesbad)”. + +Lathrimaeum knappei + +was originally described from “…dem Nordwestfuss des Berguges Galičica gelegenen Dorfe Ramne südöstlich von +Ochrid +in +Mazedonien +in etwa +1000 bis 1200 m +..”. +Zerche (2001) +designated the +lectotype +of + +L. knappei + +and synonymized it with + +A. ganglbaueri + +. + +Lathrimaeum anatolicum + +was originally described from “Anatolie occidentale:Abant Dagh…”. +Assing (2015) +synonymized this species with + +A. ganglbaueri + +. The habitus as in +Fig. 23 +; the aedeagus as in +Figs 29–30 +. + +Anthobium ganglbaueri + +is known from south Europe ( +Bulgaria +, +Greece +, +Macedonia +, +Serbia +, +Romania +) and +Turkey +. The map of distribution of the species in +Turkey +see in +Assing (2009) +. It is here recorded from the Caucasian region and +Georgia +for the first time. + + + + \ No newline at end of file diff --git a/data/03/A3/F0/03A3F02E90135A6C25B8FA01288DFC84.xml b/data/03/A3/F0/03A3F02E90135A6C25B8FA01288DFC84.xml new file mode 100644 index 00000000000..2fe8ab06e36 --- /dev/null +++ b/data/03/A3/F0/03A3F02E90135A6C25B8FA01288DFC84.xml @@ -0,0 +1,317 @@ + + + +Review of the genus Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae) of Western Palaearctic region + + + +Author + +Shavrin, Alexey V. + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +451 +474 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.1 + +journal article +10.11646/zootaxa.5537.4.1 +1175-5326 +14240224 +46B2747F-200D-4C4A-9DAA-48CBC9FD6BBA + + + + + + +Key to Western Palaearctic species of + +Anthobium + + + + + + + + + +1 Body elongate. Pronotum relatively narrow, about one and a half as broad as long. Elytra distinctly longer than broad. Median lobe and apical portions of parameres significantly broadened ( + +Fig. +23 + +in +Shavrin (2022a)) +. Habitus as in + +Fig. +1 + +in +Shavrin (2022a) +. Body reddish-brown to dark brown. Body large: +4.30–5.50 mm +. Middle Europe....................... + +algidum + + + + +- Body wide, somewhat shortened. Pronotum transverse, about twice as broad as long or slightly narrower. Median lobe and apical portions of parameres narrower. Body smaller......................................................... 2 + + + + + +2 Forebody less elevated, somewhat flattened. Middle potion of pronotum evenly elevated, sometimes with indistinct longitudinal impression. Aedeagus as in Fig. +62 in +Shavrin (2022a) +. Habitus as in + +Fig. +2 + +in +Shavrin (2022a) +. Body brown to reddish-brown. Body length: +2.70–4.50 mm +. Transpalaearctic species.................................................... + +tectum + + + + +- Forebody distinctly elevated, convex. Middle portion strongly elevated, usually with distinct longitudinal and/or mediobasal impressions......................................................................................... 3 + + + + +3 Lateral edges of pronotum with distinct crenulation, sometimes forming indistinct mediolateral angles................. 4 + + +- Lateral edges of pronotum without crenulation.............................................................. 5 + + + + + +4 Pronotum about twice as broad as long, with anterior angles relatively strongly protruded anteriad, from widest middle slightly more narrowed anteriad than posteriad, with slightly concave latero-apical margin and lateral edges, sometimes forming indistinct mediolateral angles; median impressions usually indistinct. Aedeagus as in +Figs. 29–30 +. Habitus as in +Fig. 24 +. Body paler, yellow-brown to brown. Body length: +3.05–4.10 mm +. South-eastern Europe, +Turkey +, Caucasus......... + +ganglbaueri + + + + + +- Pronotum smaller, distinctly less transverse, with anterior angles slightly protruded anteriad, from widest middle relatively evenly narrowed both anteriad and posteriad, without concave latero-apical margin and mediolateral angles; median impressions deep. Aedeagus as in Figs. +38–43 in +Shavrin & Smetana (2018) +. Habitus as in + +Fig. +1 + +in +Shavrin & Smetana (2018) +. Body darker, brown to black. Body length: +2.10–3.40 mm +. Palaearctic species....................................... + +fusculum + + + + + + +5 Anterior angles of pronotum strongly protruded anteriad...................................................... 6 + + +- Anterior angles of pronotum slightly protruded anteriad...................................................... 7 + + + + + +6 Laterobasal margins of pronotum strongly narrowed posteriad; median pronotal impressions deep. Apical part of median lobe moderately narrow; parameres significantly longer than apex of median lobe ( +Figs 10–11 +). Habitus as in +Figs 4–5 +. Body brown to reddish brown. Body length: +3.20–4.07 mm +. Europe, +Turkey +, Caucasus........................... + +melanocephalum + + + + + +- +Laterobasal margins of pronotum slightly narrowed posteriad; median pronotal impressions shallow.Apical part of median lobe wide; parameres indistinctly exceeding apex of median lobe ( +Figs 14, 16 +). Habitus as in +Fig. 6 +. Body reddish-brown. Body length: +3.55–4.10 mm +. North Africa.............................................................. + +longicorne + + + + + + + +7 Body yellowish or yellow-brown. Aedeagus as in Fig. 34h in +Zanetti (2012) +. Body length: 3.00– +3.50 mm +. Europe, Caucasus.............................................................................................. + +unicolor + + + + + +- +Body darker......................................................................................... 8 + + + + + +8 Pronotum without or with indistinct longitudinal impressions. Apical part of median lobe wide....................... 9 + + + +- +Pronotum with distinct longitudinal impression. Apical part of median lobe moderately narrow...................... 10 + + + + + + +9 Punctation of pronotum and elytra moderately fine. Parameres slightly exceeding apex of median lobe, with distinctly broadened preapical parts ( +Fig. 27 +). Habitus as in +Fig. 22 +. Body length: +2.60–3.70 mm +. Caucasus........................... + +tenue + + + + + +- +Punctation of pronotum and elytra large and deep. Parameres significantly longer than apex of median lobe, with narrow apical parts ( +Fig. 25 +). Habitus as in +Fig. 20 +. Body length: +3.50–4.90 mm +. +Portugal +, +Spain +......................... + +moczarskii + + + + + + + +10 Apical angles of pronotum not or slightly protruded anteriad. Aedeagus elongate, with relatively narrow apical part of median lobe ( +Figs. 7–9 +and + +Figs 2, +4–6 + +in +Shavrin (2020)) +. Habitus as in +Figs 1–3 +and + +Fig. +1 + +in +Shavrin (2020) +. Body length: +2.40–3.60 mm +. Palaearctic species...................................................................... + +atrocephalum + + + + + +- +Apical angles of pronotum distinctly protruded anteriad. Aedeagus short, with slightly broadened apical part of median lobe ( +Fig. 17 +). Habitus as in +Fig. 21 +. Body length: +3.20–4.15 mm +. +Syria +, +Lebanon +, +Turkey +....................... + +metallicum + + + + + + + \ No newline at end of file diff --git a/data/03/B8/87/03B88790FF9AFF9A9EE7C8A09D544AC6.xml b/data/03/B8/87/03B88790FF9AFF9A9EE7C8A09D544AC6.xml new file mode 100644 index 00000000000..51ec17ded7b --- /dev/null +++ b/data/03/B8/87/03B88790FF9AFF9A9EE7C8A09D544AC6.xml @@ -0,0 +1,859 @@ + + + +Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China + + + +Author + +Wu, Jingyu +0000-0001-6677-7726 +Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659 +wujingyu176@163.com + + + +Author + +Liu, Xingyue +0000-0001-6677-7726 +Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659 +wujingyu176@163.com + +text + + +Zootaxa + + +2024 + +2024-11-22 + + +5540 + + +1 + + +1 +169 + + + + +http://dx.doi.org/10.11646/zootaxa.5540.1.1 + +journal article +10.11646/zootaxa.5540.1.1 +1175-5334 +14240496 +4DBD3A92-F14B-4C5B-95B3-2B430EC197DD + + + + + +[24] + +Chrysopidia +( +Chrysotropia +) +orientalis +( +Hölzel, 1973 +) + + + + + + +( +Figs 32 +, +82 +) + + + + + + + +Chrysotropia orientalis +Hölzel, 1973: 355 + + +. Type locality: +Nepal +(Kathmandu Valley). +Holotype +in ZSM. + + + + + +Diagnosis. +This species is characterized by the presence of the head without spots, the maxillary palpi black on outer side, the thorax without markings, the short forewing with triangular cell +im +, the 1st rs-psm crossvein located at the end of +im +, the male sterna VIII +IV slightly exceeded the apex of ectoproct, the quadrate entoprocessus with an invagination on top. + + + + +FIGURE 32. + +Chrysopidia +( +Chrysotropia +) +orientalis +( +Hölzel, 1973 +) + +. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcal complex, dorsal view; G. Gonarcal complex, lateral view; H. Female genitalia, lateral view; I. Subgenitale; J. Spermatheca. Scale bar: 5 mm (A), 1 mm (B–D). + + + + +Description. +Male. Body length +9.2–10.3 mm +, forewing length +11.9–13.1 mm +, hindwing length 10.1–11.0 mm. + +Head without spots, vertex raised. Compound eyes black and prominent. Antennae yellow, scape and pedicel wider than flagellum. Frons, clypeus and gena without spots. Maxillary palpi black on outer side, labial palpi brownish yellow. +Pronotum anterior margin shorter than posterior margin. Mesonotum and metanotum sometimes with pale reddish spots. +Legs with dense setae, without spots. Pretarsal claws brown, with basal dilation. + +Forewing short and narrow. Pterostigma indistinct. Costal area with 19–23 crossveins; 10–13 radial crossveins present. 1st rs-psm crossvein located at end of +im +; +im +narrow and triangular; +m1 +smaller than +m2 +; 9 crossveins present between Psm and Psc. +c1 +smaller than +c2 +, +dcc +opened. A1 forked, A2 and A3 simple. Inner gradate series with 6–7 crossveins, outer gradate series with 7–8 crossveins. + +Hindwing narrow, with black suffusion on apical posterior margin. Pterostigma indistinct. Costal area with 16– 18 crossveins; 10–13 radial crossveins and 8 psm-psc crossveins present. Inner gradate series with 6–7 crossveins, outer gradate series with 6–8 crossveins. +Abdomen with dense setae. Tergum VII and sternum VII subquadrate in lateral view; tergum VIII obviously shorter than tergum VII; sternum VIII fused with sternum IX, about 2.0 times as long as tergum VIII; tergum IX fused with ectoproct. Callus cerci ovoid. Gonarcus arcuate, lateral arms narrow at base and wide at tip; entoprocessus quadrate, with an invagination on top; arcessus short, wide and curved. + +Female. Body length 10.2–11.0 mm, forewing length 14.0– +14.8 mm +, hindwing length +11.3–12.3 mm +. + +Sternum VII slightly longer than tergum VII; tergum VII subquadrate in lateral view; tergum VIII shorter than tergum VII; tergum IX fused with ectoproct. Callus cerci ovoid. Subgenitale bilobed apically; spermatheca thick; vela short; ventral impression absent; duct long and coiled. + + + +Material. + +1♂ +1♀ +, +China +, +Guangxi +, +Baise +, +Cenwanglaoshan National Nature Reserve +, +Dalongping +, + +1260 m + +, + +2016.V.19 + +, +Feiyang Liang +( +CAU +) + +; + +1♂ +, +China +, +Chongqing +, +Nanchuan District +, +Jinfoshan National Nature Reserve +, + +2021.VIII.15 + +, +Xingyue Liu +( +CAU +) + +; + +2♀ +, +China +, +Guizhou +, +Leishan +, +Leigongshan National Nature Reserve +, + +1528 m + +, + +2014.VII.19 + +, +Yuting Dai +( +CAU +) + +; + +1♀ +, +China +, +Guizhou +, +Zunyi +, +Suiyang +, +Kuankuoshui Reservoir +, + +2010.VI.04 + +, +Dan Zhou +( +CAU +) + +; + +1♂ +, +China +, +Yunnan +, +Yingjiang +, +Xima +, + +1800–2100 m + +, + +2017.V.31 + +, +Xingyue Liu +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Yingjiang +, +Xima +, + +1000 m + +, 2019.VIII, +Weizong Yang +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Yingjiang +, +Xima +, 2020. V–VI, +Weizong Yang +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Yingjiang +, +Xima +, + +1000 m + +, 2020.X, +Weizong Yang +( +CAU +) + +; + +4♀ +, +China +, +Yunnan +, +Baoshan +, +Longyang District +, +Baihualing +, +Guxingzhai +, + +1800–2000 m + +, + +2020.VI.08–10 + +, +Yuchen Zheng +& +Jiazhi Zhang +( +CAU +) + +; + +3♂ +5♀ +, +China +, +Yunnan +, +Baoshan +, +Gaoligong National Nature Reserve +, +Nankang Station +, light, + +2048 m + +, + +2015.VII.16–20 + +, +Yunlan Jiang +& +Lu Yue +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Baoshan +, +Gaoligong National Nature Reserve +, +Baihualing +, + +1500 m + +, + +2015.VII.25 + +, +Lu Yue +( +CAU +) + +; + +1♀ +, +China +, +Zhaotong +, +Yiliang +, +Xiaocaoba +, + +2022. VII.18 + +, +Xingyue Liu +( +CAU +) + +; + +1♂ +1♀ +, +China +, +Yunnan +, +Lvchun +, +Huanglianshan National Nature Reserve +, +Bamawudu +, + +1700–2100 m + +, + +2021.V.29–VI.02 + +, +Yuchen Zheng +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Lushui +, + +1900 m + +, + +2012.VII.25 + +, +Junchao Wang +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Lushui +, light, + +2016.VII.13–14 + +, +Liang Wang +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Lushui +, +Pianma +, light, + +2018.V.01 + +, +Yunlong Ma +( +CAU +) + +; + +1♂ +, +China +, +Yunnan +, +Lvchun +, +Huanglianshan National Nature Reserve +, + +1090 m + +, +Xingyue Liu +( +CAU +) + +; + +1♂ +, +China +, +Yunnan +, +Lvchun +, +Huanglianshan National Nature Reserve +, light, + +2013.VI.05 + +, +Jie Zhang +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Jinping +, +Maandi +, + +1225 m + +, + +2016.VII.12 + +, +Ya’nan Lv +( +CAU +) + +; + +1♂ +, +China +, +Yunnan +, +Puer +, +Wuliangshan National Nature Reserve +, light, + +2221 m + +, + +2016.VII.16 + +( +CAU +) + +; + +1♀ +, +China +, +Yunnan +, +Maguan +, +Jinchang +, +Xiajinchang +, + +1430 m + +, + +2016.VII.29 + +, +Yunlan Jiang +( +CAU +) + +; + +3♂ +, +China +, +Xizang +, +Mêdog +, 80K, light, + +2115 m + +, + +2018.VII.31 + +, +Xingyue Liu +( +CAU +) + +; + +4♂ +3♀ +, +China +, +Xizang +, +Mêdog +, 80K, + +2115 m + +, + +2020.VIII.20– 27 + +, +Yuchen Zheng +( +CAU +) + +; + +1♂ +1♀ +, +China +, +Xizang +, +Nyingchi +, +Mêdog +, +Gyêling +, + +1500 m + +, + +2020.VIII.22 + +, +Yuchen Zheng +( +CAU +) + +; + +1♀ +, +China +, +Xizang +, +Shigatse +, +Gyirong +, + +2950 m + +, + +2020.VII.26 + +, +Yuchen Zheng +( +CAU +) + +; + +1♀ +, +China +, +Xizang +, +Nyingchi +, +Bomê +, +Tangmai +, + +2170 m + +, + +2020.VIII.07 + +, +Yuchen Zheng +( +CAU +) + +; + +1♀ +, +China +, +Xizang +, +Shigatse +, +Zhêntang +, + +2200 m + +, + +2022.VII.12 + +, +Yuezheng Tu +( +CAU +) + +. + + + + +Distribution. +China +( +Guangxi +, +Guizhou +, +Xizang +, +Yunnan +); +Nepal +(Kathmandu). + + + + +Remarks. +Hölzel (1973) +described + +C. orientalis + +based on a female specimen from Kathmandu Valley, +Nepal +, and no further records of this species has been published since the original description. We examined many specimens that fit the original description of this species, especially the head without spots, the thorax without markings, the 1st rs-psm crossvein located at the end of +im +, etc. Here, we identify our material as + +C. orientalis + +, which represents the new record of this species from +China +. + + + + \ No newline at end of file diff --git a/data/03/B8/87/03B88790FFF0FFFD9EE7CF249ED84EDE.xml b/data/03/B8/87/03B88790FFF0FFFD9EE7CF249ED84EDE.xml index 61c4e4cea38..cd697af181d 100644 --- a/data/03/B8/87/03B88790FFF0FFFD9EE7CF249ED84EDE.xml +++ b/data/03/B8/87/03B88790FFF0FFFD9EE7CF249ED84EDE.xml @@ -1,56 +1,56 @@ - - - -Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China + + + +Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China - - -Author + + +Author -Wu, Jingyu -0000-0001-6677-7726 -Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659 -wujingyu176@163.com +Wu, Jingyu +0000-0001-6677-7726 +Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659 +wujingyu176@163.com - - -Author + + +Author -Liu, Xingyue -0000-0001-6677-7726 -Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659 -wujingyu176@163.com +Liu, Xingyue +0000-0001-6677-7726 +Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659 +wujingyu176@163.com -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-22 + +2024 + +2024-11-22 - -5540 + +5540 - -1 + +1 - -1 -169 + +1 +169 - -http://dx.doi.org/10.11646/zootaxa.5540.1.1 + +http://dx.doi.org/10.11646/zootaxa.5540.1.1 -journal article -10.11646/zootaxa.5540.1.1 -1175-5334 -14240496 -4DBD3A92-F14B-4C5B-95B3-2B430EC197DD +journal article +10.11646/zootaxa.5540.1.1 +1175-5334 +14240496 +4DBD3A92-F14B-4C5B-95B3-2B430EC197DD @@ -73,15 +73,13 @@ ) - + Tumeochrysa praeclara - Hölzel, 1973: 376 - . Type locality: @@ -91,14 +89,11 @@ in ZSM. - - + Tumeochrysa hui - Yang, 1987: 209 - . Type locality: @@ -110,11 +105,8 @@ in ZSM. ). Holotype in CAU. -syn. nov. Diagnosis. -This species is characterized by the presence of the frons, clypeus and gena without spots, the vertex with reddish triangular markings, the scape with small tubercle on the outer side, the wings without shadings, and the median arch of male gonarcus with short wide projections. - - - + +syn. nov. FIGURE 44. @@ -129,6 +121,12 @@ This species is characterized by the presence of the frons, clypeus and gena wit . A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Thorax, dorsal view; F. Male genitalia, lateral view; G. Gonarcal complex, dorsal view; H. Gonarcal complex, lateral view. Scale bar: 5 mm (A), 1 mm (B–D). + +Diagnosis. This species is characterized by the presence of the frons, clypeus and gena without spots, the vertex with reddish triangular markings, the scape with small tubercle on the outer side, the wings without shadings, and the median arch of male gonarcus with short wide projections. + + + + Description. @@ -160,7 +158,6 @@ closed. Gradates in four rows, basal crossvein of inner gradate series not meeti Material. - Holotype of @@ -185,7 +182,7 @@ of , Shengchang Hu ( -CAU +CAU ). 1♂ , @@ -209,7 +206,7 @@ of , Yuchen Zheng ( -CAU +CAU ); 1♂ , @@ -235,9 +232,8 @@ of , Yuchen Zheng ( -CAU +CAU ). - diff --git a/data/9D/49/87/9D4987B60611FFC6C0C9FA048CCF981E.xml b/data/9D/49/87/9D4987B60611FFC6C0C9FA048CCF981E.xml new file mode 100644 index 00000000000..fa3b95606c1 --- /dev/null +++ b/data/9D/49/87/9D4987B60611FFC6C0C9FA048CCF981E.xml @@ -0,0 +1,73 @@ + + + +A new Potamocypridini genus of Cypridopsinae (Crustacea, Ostracoda) from Thailand, with a discussion on taxonomic characters and morphological evolution in the tribe + + + +Author + +Savatenalinton, Sukonthip + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +475 +492 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.2 + +journal article +10.11646/zootaxa.5537.4.2 +1175-5326 +14240272 +FCB0EA5F-ED89-45E8-9105-9F4942933A2D + + + + + + +Tribe + +Potamocypridini +Ghetti & McKenzie, 1981 + + + + + + + + +Genera included. +Potamocypris + +, + +Neopotamocypris + + +gen. nov. + + + + + \ No newline at end of file diff --git a/data/9D/49/87/9D4987B60612FFC0C0C9FF348F7D9F7F.xml b/data/9D/49/87/9D4987B60612FFC0C0C9FF348F7D9F7F.xml new file mode 100644 index 00000000000..6ba9afc2ad5 --- /dev/null +++ b/data/9D/49/87/9D4987B60612FFC0C0C9FF348F7D9F7F.xml @@ -0,0 +1,549 @@ + + + +A new Potamocypridini genus of Cypridopsinae (Crustacea, Ostracoda) from Thailand, with a discussion on taxonomic characters and morphological evolution in the tribe + + + +Author + +Savatenalinton, Sukonthip + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +475 +492 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.2 + +journal article +10.11646/zootaxa.5537.4.2 +1175-5326 +14240272 +FCB0EA5F-ED89-45E8-9105-9F4942933A2D + + + + + + + +Neopotamocypris indivisa + +gen. nov. +sp. nov. + + + + + +urn:lsid:zoobank.org:act: +ACBAFC94-E72D-4B01-888D-B6B042426CDB + + + + + +( +Figs 1–5 +) + + + + + + +Potamocypris +sp. + +: + +Savatenalinton & Suttajit, 2016: 8 + +. + + + + + +Etymology. +The new species is named with regard to the outstanding character of the T +2 in +which the undivided penultimate segment is apparent. The specific epithet +“indivisa +” means “undivided” in Latin. + + + + +FIGURE 1. + +Neopotamocypris indivisa + + +gen. nov. +sp. nov. + +A. CpL from left side (MSU-ZOC.412). B. CpD (MSU-ZOC.411). C. Detail of surface of LVe (MSU-ZOC.412). D. LVi (MSU-ZOC.409). E. RVi (MSU-ZOC.409). F. Detail of muscle scars of RVi (MSU-ZOC.409). Scale bars: A–B, E–F = 100 µm, C = 10 µm, D = 20 µm. + + + + +Diagnosis. +RV overlapping LV ventrally, Cp in dorsal view narrow, anterior and posterior extremities pointed with subequal LV and RV margins, Cp in lateral view subtriangular, posterior margin narrowly rounded, valve surface set sparsely with long stiff, unrimmed-pore setae and long, thin, rimmed-pore setae. LV with anterior and posterior inwardly displaced selvage. RV with submarginal anterior selvage. A1 seven-segmented, Rome organ small, Wouters organ not observed. A2 natatory setae long; aesthetasc Y four-segmented with large and striated last section, A2 terminal claws well-developed. Terminal segment of Md-palp with robust claws set with (strong) setules, α seta with flagellum-like tip, γ seta markedly long with sparse setules and flagellum-like ending. Mx1-palp with spatula-shaped terminal segment bearing three large claws, a tiny seta and a robust seta, lateral subapical seta on basal segment robust with very long apical setules, lateral setae on third endite robust with very long apical setules. Apical setae on T1-endite with few setae longer than others; a, b, c, d setae absent. T2 with undivided penultimate segment, d2 seta present (d1 seta absent). T3 with pincer organ, d1 seta absent, F seta present. CR reduced with triangular base and considerably long flagellum-like apical seta, basal seta long, with thread-like tip. Female genital hook stubby with strongly curved tip. + + + + +Type materials: + + +Holotype +. + +Female +, soft parts dissected and preserved in glycerine on a slide, and valves stored dry in a micropalaeontological slide ( +MSU-ZOC.405 +). + + + + + +Paratypes +. + +Four dissected +females +( +MSU-ZOC.406 +, 409, 410, 413) stored like the holotype and five undissected females (MSU-ZOC.407, 408, 411, 412, 414) stored dry in micropalaeontological slides. Whole specimens (ca. 20 individuals) kept in 70% ethanol (MSU-ZOC.415, 416) + +. + + +Type locality. +Sap Pradu Reservoir +, +Sikhio District +, +Nakhon Ratchasima Province +; coordinates: +14° 51' 09" N +, +101° 41' 52" E +, collected on + +6 February 2006 +, +5 October 2007 + +. + + +Other localities. +See +Table 1 +. + + + + +Measurements (μm). +Cp (n=5): L = 533–558, H = 282–301, W = 179–189, LV (n=3): L = 533–555, H = 262–287, RV (n=3): L = 528–542, H = 291–304. + + +Description + + +Cp in lateral view ( +Fig 1A +) subtriangular, anterior margin rounded, posterior margin more pointed than anterior margin, greatest height situated ca. at mid-length. RV overlapping LV ventrally, dorsal margin of RV higher than that of LV. Valve surface sparsely set with long stiff, unrimmed-pore setae and with long, thin, rimmed-pore setae ( +Fig 1C +). + + +Cp in dorsal view ( +Fig 1B +) narrowly elongated, anterior and posterior extremities pointed with subequal LV and RV margins, greatest width situated ca. at mid-length. + + + +FIGURE 2. + +Neopotamocypris indivisa + + +gen. nov. +sp. nov. + +(MSU-ZOC.409). A. Detail of the posterior part of LVi. B. Detail of the anterior part of LVi. C. Detail of the anterior part of RVi. D. Detail of the posterior part of RVi. Scale bars: A–D = 50 µm. + + + + +TABLE 1. +List of localities of + +Neopotamocypris indivisa + +n. gen. +n. sp. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
NumberLocalityDistrict, ProvinceSampled dateCollecting depth (m)SedimentAquatic plant
1Sap Pradu ReservoirSikhio, Nakhon Ratchasima6 Feb 2006, 5 Oct 20070.5Sand, organic detritus (small amount of death plant material) +No aquatic +plants +
2Pasak Cholasit DamPhatthana Nikhom, Lopburi6 Feb 20060.6Sand, organic detritus (small amount of death plant material) +No aquatic +plants +
3Lopburi RiverMuang, Lopburi5 Oct 20070.9Sand, mud, organic detritus (small amount of death plant material) + +Hydrilla verticillata + +(L.f.) Royle, + +Eichhornia crassipes +(Mart.) Solms + +
4Nong Naree (swamp)Muang, Phetchabun9 Oct 20070.5Mud, organic detritus (plenty of death plant material) + +Lemna perpusilla +Torr. + +, + +Nymphaea pubescens +Willd. + +, + +Ipomoea aquatic +Forsk. + +, + +Cyperus involucratus +Roxb. + +
5Huai Pao (stream)Kasetsomboon, Chaiyaphum15 Nov 20060.6Sand, mud, organic detritus (moderate amount of death plant material) +No aquatic +plants +
6Nong Krang ReservoirBo Phloi, Kanchanaburi12 Mar 20190.6Mud, organic detritus (plenty of death plant material) + +Hydrilla verticillata + +(L.f.) Royle, + +Ottelia alismoides + +(L.) Pers., + +Typha angustifolia + +L. +
7A swampMuang, Kanchanaburi18 Oct 20180.6Mud, organic detritus (small amount of death plant material) + +Nymphaea pubescens +Willd. + +, + +Ipomoea aquatic +Forsk. + +, + +Cyperus involucratus +Roxb. + +, + +Colocasia esculenta +var. +aquatilis +Hassk + +
8Samnak Mai Teng ReservoirMuang, Ratchaburi13 Mar 20190.5Sand, mud, organic detritus (small amount of death plant material) + +Ceratophyllum demersum + +L. +
9Ban Wang Hin ReservoirNong Ya Plong, Phetchaburi14 Mar 20191Mud, organic detritus (small amount of death plant material) +No aquatic +plants +
10Yang Chum DamKui Buri, Prachuap Khiri Khan15 Mar 20190.6Sand, organic detritus (small amount of death plant material) + +Ottelia alismoides + +(L.) Pers. +
+ + +LV in interior view ( +Figs 1D +, +2A–B +) with anterior and posterior inwardly displaced selvages, posterior selvage with postero-ventral angulate corner, calcified inner lamella with one inner list anteriorly and posteriorly, ventral margin slightly sinuate at mid-length, greatest height situated in front of mid-length. + + +RV in interior view ( +Figs 1E +, +2C–D +) with marginal anterior selvage, calcified inner lamella with weak inner lists anteriorly and posteriorly, dorsal margin strongly arched, greatest height situated at mid-length, strongly sloping down to posterior margin. + + +A1 ( +Fig 3A +) seven-segmented. First segment with one long subapical dorsal seta (reaching mid-length of next segment) and two long apical ventral setae (subequal in length), Wouters organ not seen. Second segment slightly wider than long, with one long apical dorsal seta (reaching slightly beyond mid-length of next segment) and small Rome organ. Third segment with one long apical dorsal setae (reaching slightly beyond tip of next segment) and one short apical ventral seta. Fourth segment with two long dorsal setae and two short ventral setae (longer one reaching slightly beyond tip of next segment). Fifth segment with two long dorsal setae and two shorter ventral setae (longer one reaching mid-length of terminal segment, shorter one reaching tip of next segment). Sixth segment with four long apical setae. Terminal segment elongated with three (two long, one shorter) apical setae and aesthetasc ya, length of aesthetasc ya ca. 2/3 of shorter seta. + + + +FIGURE 3. + +Neopotamocypris indivisa + + +gen. nov. +sp. nov. + +A. A1 (MSU-ZOC.405). B. A2 (MSU-ZOC.405). C. Aesthetasc Y of A2 (MSU-ZOC.405). D. Md coxa (MSU-ZOC.406). E. Rake-like organ ((MSU-ZOC.409). Scale bars: A–B, D = 50 µm, C = 20 µm. + + + +A2 ( +Fig 3B–C +): Basal segment with two slender proximal setae and one long slender ventro-apical seta. Exopodite with three (one long, two short) setae, the long seta reaching tip of penultimate segment. First endopodal segment with five long (reaching beyond tip of terminal claws) and one shorter accompanying setae, length of latter seta ca. 3/4 of penultimate segment, aesthetasc Y slender, four-sectioned, fourth section striated and larger than other sections, apical ventral seta long, reaching far beyond tip of terminal segment. Penultimate segment undivided, medially with two long dorsal setae (length of shorter one ca. 2/3 of that of long one) and four ventral t-setae, t2 seta longer and larger than others; this segment distally with three large, serrated claws G1, G2 and G3, claw G2 stronger serrated than claws G1 and G3, z1–z3 setae long, z1 and z2 not reaching tip of terminal claws, aesthetasc y2 short (reaching mid-length of terminal segment). Terminal segment slightly elongated with large, serrated claw GM, shorter and slender claw Gm (length ca. 2/3 of that of claw GM), and short aesthetasc y3, length of the latter ca. half of claw Gm and ca. 4/5 of that of accompanying seta, g-seta not seen. + + +Rake-like organ ( +Fig 3E +) T-shaped, apically with c.17 small blunt teeth. + + +Md-coxa ( +Fig 3D +) elongated, apically with rows of teeth (ca. 12 teeth) and small setae, largest tooth on dorsal margin and other teeth gradually decreasing in size towards ventral margin, and with cone-shaped, pointed tip subapical dorsal seta. + + +Md-palp ( +Fig 4A–C +): first segment with two large, hirsute S1 and S2 setae (S1seta shorter than S2 seta), one long and slender seta and a smooth, thin α-seta with flagellum-like tip. Second segment dorsally with three subequal long apical setae, one of them robust with long setules; ventrally with a group of three long, hirsute setae, one shorter seta and β-seta, the latter small, cone-shaped and slightly plumose. Penultimate segment with three groups of setae: dorsally with a group of four subequal, long, subapical setae, one of them with short setules, shortest seta thin (reaching ca. half of long ones), two long seta reaching beyond tip of terminal claws; laterally with apical γ-seta and two further apical setae; the former long, slender with slightly wide base and flagellum-like ending and with sparsely few setules, the latter smooth, thin and short; ventrally with one long apical seta (length ca. 4 times of that of terminal segment) and one short and thin seta. Terminal segment bearing three long, large, serrated claws (one of them shorter but stronger serrated than other claws) and two short setae. + + +Mx1 ( +Fig 4D–F +). Palp two-segmented. Basal segment of palp subquadrate with a group of five apical setae and one subapical seta; the former large, subequal and sparsely set with long setules, the latter long bearing short setules; medially of this segment with robust lateral subapical seta distally set with very long setules (reaching slightly beyond tip of terminal segment). Terminal segment spatula-shaped, apically with four (three robust, one small) claws and two (one tiny and one large) setae, large seta sparsely set with long setules. Three endites stout, third endite laterally with a group of six large setae, two of them with short setules, four setae stubby with very long setules. + + +T1 ( +Figs 4G +): protopodite without a, b, c, and d-setae, distally with ca. 10 hirsute apical setae, two of them obviously longer than other setae. Endopodite elongated, weakly built palp with three (two long, one short) setae, length of longest seta more than twice that of endopodite length, length of shortest seta ca. 1/5 of that of longest one. + + +T2 ( +Fig 5A +) with tiny d2 seta, d1 seta absent. Second segment with short e-seta (length ca. 1/3 of that of penultimate segment). Penultimate segment undivided, medially with short seta (not reaching tip of its segment), distally with a pair of apical setae (short g-seta and one tiny seta). Terminal segment dorsally bearing one short apical seta (h1), ventrally bearing one apical spine-like seta (h3) and medially with a very long and curved serrated claw seta (h2), length of h1 less than 1/7 of that of h2, length of h2 ca. that of last three segments. + + +T3 ( +Fig 5B–C +) with pincer organ. First segment with short d2 seta and long dp setae (length of d2 ca. 2/3 of that of dp seta), d1 seta absent, and with curved, weak sclerotized distal part. Second segment with short apical e-seta (length ca. 1/3 of that of next segment). Third segment with medially short f-seta (not reaching tip of segment) and with short subapical F seta (not reaching tip of segment). Terminal segment bearing three setae (h1-h3 setae), h1 seta short, h2 seta long, claw-like and h3 reflexed, hirsute subapical seta, h3 seta slightly longer than that of h2 seta, apical part of segment (close to base of h2 seta) with small projection. + + +CR ( +Fig. 5D +) reduced with triangular base, markedly long, flagellum-like apical seta and short subapical seta, the latter with thread-like tip. + +Female reproductive organ with stubby genital hook ending with strongly curved tip. +Male unknown. + + + +Ecology. +Most samples of + +N. indivisa + + +gen. nov. +sp. nov. + +were collected at the depth of ca. +40–50 cm +except for two localities, which were taken at ca. +90–100 cm +depth. ( +Table 1 +). The sediments were sand, mud and a small amount of organic detritus, which mostly was dead plant material, but two localities (number 4 and 6) had plenty of organic matter. No macrophyte was found in four localities (number 1, 2, 5, 9). Submerged plants found were + +Hydrilla verticillata + +(L.f.) Royle, + +Ottelia alismoides + +(L.) Pers. and + +Ceratophyllum demersum + +L. (locality number 3, 6, 8, 10), while floating plants found were + +Eichhornia crassipes +(Mart.) Solms + +, + +Lemna perpusilla +Torr. + +, + +Nymphaea pubescens +Willd. + +and + +Ipomoea aquatica +Forsk. + +(locality number 3, 4, 7). Emerged plants were found in three localities (number 4, 6, 7): + +Cyperus involucratus +Roxb. + +, + +Typha angustifolia + +L. and + +Colocasia esculenta +var. +aquatilis +Hassk. + + + + + \ No newline at end of file diff --git a/data/9D/49/87/9D4987B6061DFFC9C0C9FBC18AF09C27.xml b/data/9D/49/87/9D4987B6061DFFC9C0C9FBC18AF09C27.xml new file mode 100644 index 00000000000..caffb7eddaf --- /dev/null +++ b/data/9D/49/87/9D4987B6061DFFC9C0C9FBC18AF09C27.xml @@ -0,0 +1,249 @@ + + + +A new Potamocypridini genus of Cypridopsinae (Crustacea, Ostracoda) from Thailand, with a discussion on taxonomic characters and morphological evolution in the tribe + + + +Author + +Savatenalinton, Sukonthip + +text + + +Zootaxa + + +2024 + +2024-11-12 + + +5537 + + +4 + + +475 +492 + + + + +http://dx.doi.org/10.11646/zootaxa.5537.4.2 + +journal article +10.11646/zootaxa.5537.4.2 +1175-5326 +14240272 +FCB0EA5F-ED89-45E8-9105-9F4942933A2D + + + + + + +Origin of + +Neopotamocypris + +gen. nov. + + + + + + + +Neopotamocypris + + +gen. nov. + +is very similar to + +Potamocypris + +in external morphology. For example, the general shape of the Cp in lateral view is subtriangular, the RV overlaps the LV ventrally, and the valve surface is usually set with tiny pits and setae. In addition, the similarity also exists in the valve structure, especially on the LV, which has inwardly displaced selvages on both, the anterior and posterior parts. The discrimination between these two genera thus mainly relies on the soft parts morphology, especially the T2, T3, Md-palp, Mx1 and genital hook of the female. As both genera belong to the same tribe ( +Potamocypridini +), such differences existing in the new genus could be constituted via divergent evolution from the + +Potamocypris + +ancestor and the new genus might be of a more recent origin evidenced by its derived characters (see discussion above). + + + +Neopotamocypris indivisa + + +gen. nov. +sp. nov. + +is the only species of the tribe +Potamocypridini +recorded in Southeast Asia and +Thailand +. It generally resembles + +P. acuminata +, + +which was described based on Pleistocene fossil specimens ( +Fuhrmann & Pietrzeniuk 1990 +; Fuhrman & Goth 2011). However, recently, the first living specimens of this species were reported from +Japan +and the description of soft body parts was provided for the first time ( + +Smith +et al +. 2024 + +). Thus far, + +P. acuminata + +has been recorded from the Palaearctic region. Nonetheless, + +Smith +et al +. (2021) + +reported on exotic species, from Japanese pet shops, including + +Pseudostrandesia tenebrarum + +which is very similar to the endemic Thai + +P. phetchabunensis + +(see +Savatenalinton & Martens 2010 +; + +Smith +et al +. 2021 + +), and also proposed that its origin is most likely in Southeast Asia, with transportation occurring via the pet trade. Moreover, + +Smith +et al +. (2024) + +recently presented six more exotic species of +Japan +, including + +P. acuminata + +and + +Siamopsis sudzukii +(Okubo, 1992) + +, collected from pet shops and hobbyist’s aquaria. The record of the latter species in +Japan +is remarkable in that + +Siamopsis + +is no longer endemic to +Thailand +. Such evidence most likely suggests the possibility of transportation of ostracod faunas between these two regions. Hence, it could be assumed that + +N. indivisa + + +gen. nov. +sp. nov. + +evolved from an ancestor in + +Potamocypris + +, like + +P. acuminata + +. + + +Another possibility is that + +N. indivisa + + +gen. nov. +sp. nov. + +evolved from an Oriental + +Potamocypris + +ancestor. In the Oriental region, only one identified + +Potamocypris +species + +has been recorded, the Indian + +P. narayanani + +(see +George & Martens 2002 +). Further findings of + +Potamocypris + +, either living or fossil, in +Thailand +or Southeast Asia would provide more explanation on this phenomenon. + + + + +Distribution + + +In the last two decades, extensive surveys of freshwater ostracods in +Thailand +have been carried out mainly in three parts of the country: northern, northeast and western parts. About 410 water bodies, ranging from various +types +of habitats in 37 provinces (from a total of 77 provinces) were sampled and ca. 620 samples were obtained. +The +examination of such a large number of sampled localities unveils that + +N. indivisa + + +gen. nov. +sp. nov. + +is very rare as it has so far been found only in 10 localities in eight provinces, namely +Nakhon Ratchasima +, +Chaiyaphume +, +Lopburi +, +Phetchabun +, +Kanchanaburi +, +Ratchaburi +, +Phetchaburi +and +Prachuap Khiri Khan +( +Table 1 +, +Fig 6 +). + + + + \ No newline at end of file