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<treatment ID-GBIF-Taxon="135490007" LSID="urn:lsid:plazi:treatment:016660F62DF17563377579768AAF0440" httpUri="http://treatment.plazi.org/id/016660F62DF17563377579768AAF0440" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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<paragraph pageId="4" pageNumber="5">
<taxonomicName authority="Knudsen, 1970" authorityName="Knudsen" authorityYear="1970" class="Bivalvia" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vesicomya galatheae" order="Pholadomyoida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="galatheae">Vesicomya galatheae (Knudsen, 1970)</taxonomicName>
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<taxonomicName id="6EAFAA83F5E30DF160DB231555F797FE" authority="Knudsen, 1970" authorityName="Knudsen" authorityYear="1970" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Vesicomya galatheae" order="Pholadomyoida" pageId="4" pageNumber="5" rank="species" species="galatheae">Vesicomya galatheae (Knudsen, 1970)</taxonomicName>
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<subSubSection pageId="4" pageNumber="5" type="materials_examined">
<paragraph pageId="4" pageNumber="5">Material examined.</paragraph>
<paragraph pageId="4" pageNumber="5">NHM_143 NHMUK 20170040, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/c609ed0c-f881-44c9-a6a0-3e36f0934997</paragraph>
<paragraph pageId="4" pageNumber="5">NHM_255 NHMUK 20170041, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/314ef160-7cfa-4705-b091-640c3e69ad1a</paragraph>
<paragraph pageId="4" pageNumber="5">NHM_260 NHMUK 20170042.1-2, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/3add2560-71c1-4879-afb8-0a5ed1449c89</paragraph>
<subSubSection id="CD4F4C74786F9C6C9311C79C41DAFDE1" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="734604AEDB34D0BB6D6055873066047F" pageId="4" pageNumber="5">Material examined.</paragraph>
<paragraph id="E995AA1B6E963104748EC4FBD94F2E91" pageId="4" pageNumber="5">NHM_143 NHMUK 20170040, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/c609ed0c-f881-44c9-a6a0-3e36f0934997</paragraph>
<paragraph id="9950EB60ACB9809283D43A55AA294FFF" pageId="4" pageNumber="5">NHM_255 NHMUK 20170041, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/314ef160-7cfa-4705-b091-640c3e69ad1a</paragraph>
<paragraph id="C0552DAD45C58D2E7331F413DD20B22B" pageId="4" pageNumber="5">NHM_260 NHMUK 20170042.1-2, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/3add2560-71c1-4879-afb8-0a5ed1449c89</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="description">
<paragraph pageId="4" pageNumber="5">Description.</paragraph>
<paragraph pageId="4" pageNumber="5">Small, inflated sub-spherical. Sculpture of fine closely spaced low commarginal lamellae. Right valve with two cardinal teeth, posterior long, thin, anterior tooth small and short (Fig. 4). Specimen NHM_143 length 1.4 mm, height 1.2 mm.</paragraph>
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<paragraph pageId="4" pageNumber="5">
<subSubSection id="DAD3FFE7F382E923A657F33A8F2039F6" pageId="4" pageNumber="5" type="description">
<paragraph id="68A0D9C549B962B876EEE76E615C9ACD" pageId="4" pageNumber="5">Description.</paragraph>
<paragraph id="47E5AC48384C7CF27D2E5181B738255B" pageId="4" pageNumber="5">Small, inflated sub-spherical. Sculpture of fine closely spaced low commarginal lamellae. Right valve with two cardinal teeth, posterior long, thin, anterior tooth small and short (Fig. 4). Specimen NHM_143 length 1.4 mm, height 1.2 mm.</paragraph>
<caption id="53E032D6EEAC833293AB67EE3EEE0519" pageId="4" pageNumber="5">
<paragraph id="451F1B4541B976566FA8DE7FED8918B9" pageId="4" pageNumber="5">
Figure 4.
<taxonomicName class="Bivalvia" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vesicomya galatheae" order="Venerida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="galatheae">Vesicomya galatheae</taxonomicName>
<taxonomicName id="E007864FEE65C579805E7A51FC66258D" class="Bivalvia" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vesicomya galatheae" order="Venerida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="galatheae">Vesicomya galatheae</taxonomicName>
(Knudsen, 1970) A Live imaged specimens of NHM_260a,b,c habitus B Detail of NHM_143, probable juvenile, oil droplets arrowed C NHM_255 live imaged specimen
<normalizedToken originalValue="DE">D-E</normalizedToken>
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SEM detail of shell interior and hinge teeth of NHM_260a (right valve). Scale bars: 0.5 mm (B, E). Image attribution Glover, Taylor, Dahlgren &amp; Wiklund, 2017.
</paragraph>
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<paragraph pageId="4" pageNumber="5">Genetic data.</paragraph>
<paragraph pageId="4" pageNumber="5">GenBank NHM_143 18S-MF157474; NHM_255 16S-MF157460, 18S-MF157487, COI-MF157509; NHM_260 18S-MF157488, COI-MF157510.</paragraph>
<subSubSection id="FB3F7A55485963E95D660E02DB80F74C" pageId="4" pageNumber="5" type="genetic data">
<paragraph id="A199FADD776E4F1513051F84E1BEBDBE" pageId="4" pageNumber="5">Genetic data.</paragraph>
<paragraph id="0DE2AA687F43B2B8291C81283558A073" pageId="4" pageNumber="5">GenBank NHM_143 18S-MF157474; NHM_255 16S-MF157460, 18S-MF157487, COI-MF157509; NHM_260 18S-MF157488, COI-MF157510.</paragraph>
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<pageBreakToken pageId="5" pageNumber="6" start="start">Remarks</pageBreakToken>
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<pageBreakToken id="87C79790BD27DC0B2E1F41F342B39790" pageId="5" pageNumber="6" start="start">Remarks</pageBreakToken>
.
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<paragraph pageId="5" pageNumber="6">
<taxonomicName class="Bivalvia" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vesicomya galatheae" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="galatheae">Vesicomya galatheae</taxonomicName>
<paragraph id="EF0AE67DEBD2BBD0E67C90022853310F" pageId="5" pageNumber="6">
<taxonomicName id="862DDE604FD56D0056E50E9B2C5CDC59" class="Bivalvia" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vesicomya galatheae" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="galatheae">Vesicomya galatheae</taxonomicName>
was described from off Costa Rica and Panama at 2950- 3570 m. Morphologically similar to
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(Smith, 1885) holotype NHMUK 1887.2.9.2710-11 but
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regard this as a northern Pacific species distinguished from
<taxonomicName lsidName="V. galatheae" pageId="5" pageNumber="6" rank="species" species="galatheae">V. galatheae</taxonomicName>
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by the shape, hinge teeth and number of siphonal tentacles. When comparing sequences from our CCZ specimens with the
<taxonomicName class="Bivalvia" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vesicomya pacifica" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="pacifica">Vesicomya pacifica</taxonomicName>
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from
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<bibRefCitation id="326E953EE8BD5E7774243092E11FD60F" author="Krylova, EM" journalOrPublisher="Deep-Sea Research Part II: Topical Studies in Oceanography" pageId="30" pageNumber="31" pagination="198 - 209" title="Vesicomyinae (Bivalvia: Vesicomyidae) of the Kuril-Kamchatka Trench and adjacent abyssal regions." url="https://doi.org/10.1016/j.dsr2.2014.10.004" volume="111" year="2015">Krylova et al. (2015)</bibRefCitation>
, the K2P difference is 0.11. In the molecular tree (Fig. 5) it groups with a
<taxonomicName class="Bivalvia" family="Kelliellidae" genus="Kelliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelliella" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Kelliella</taxonomicName>
<taxonomicName id="1E524992FCA7B1A79FFE72B956F1DF9B" class="Bivalvia" family="Kelliellidae" genus="Kelliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelliella" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Kelliella</taxonomicName>
species from the northwestern Atlantic and these two species form a sister clade to
<taxonomicName class="Bivalvia" family="Vesicomyidae" genus="Calyptogena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calyptogena" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Calyptogena</taxonomicName>
<taxonomicName id="2CFC3728F2DFEA67235A02B3375D3A66" class="Bivalvia" family="Vesicomyidae" genus="Calyptogena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calyptogena" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Calyptogena</taxonomicName>
species.
<taxonomicName class="Bivalvia" family="Kelliellidae" genus="Kelliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelliella" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Kelliella</taxonomicName>
<taxonomicName id="468887993960C49FC998AEE80F2580A8" class="Bivalvia" family="Kelliellidae" genus="Kelliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kelliella" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Kelliella</taxonomicName>
species are very similar to
<taxonomicName class="Bivalvia" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vesicomya" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Vesicomya</taxonomicName>
<taxonomicName id="8C9437F3A9CA7483B4AC5077F6489967" class="Bivalvia" family="Vesicomyidae" genus="Vesicomya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vesicomya" order="Venerida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Vesicomya</taxonomicName>
and the relationships of species assigned to the two genera need clarification. Forms a unique monophyletic clade distinct from all other AB01 specimens. No genetic matches on GenBank.
</paragraph>
<caption pageId="5" pageNumber="6">
<paragraph pageId="5" pageNumber="6">
<caption id="B082746FA5D582804FFAD467E4D26027" pageId="5" pageNumber="6">
<paragraph id="5E8B321E6F0B6BCE5BD2E074AECD7B34" pageId="5" pageNumber="6">
Figure 5. Phylogenetic analysis of
<taxonomicName class="Bivalvia" lsidName="Heterodonta" pageId="5" pageNumber="6" rank="class">Bivalvia</taxonomicName>
<taxonomicName id="788EE0D49B86F7BEA5AD6E50CB511FB3" class="Bivalvia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterodonta" pageId="5" pageNumber="6" phylum="Mollusca" rank="class">Bivalvia</taxonomicName>
:
<taxonomicName lsidName="Heterodonta" pageId="5" pageNumber="6" rank="subClass" subClass="Heterodonta">Heterodonta</taxonomicName>
<taxonomicName id="35DBB9D68A5FF3A3C503C3F8CC4695D9" lsidName="Heterodonta" pageId="5" pageNumber="6" rank="subClass" subClass="Heterodonta">Heterodonta</taxonomicName>
. 50% majority rule consensus tree from the Bayesian analyses using 18S and COI. Asterisks denotes support values of 95 or above.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="ecology">
<paragraph pageId="5" pageNumber="6">Ecology.</paragraph>
<paragraph pageId="5" pageNumber="6">Found in polymetallic nodule province.</paragraph>
<subSubSection id="AB894F7D09932A56B9A491F916920033" pageId="5" pageNumber="6" type="ecology">
<paragraph id="2254675A87781A358E3C9E66536911DE" pageId="5" pageNumber="6">Ecology.</paragraph>
<paragraph id="E6C558824F35CC3B93F1495CE0F2B1E0" pageId="5" pageNumber="6">Found in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,113 +1,112 @@
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<taxonomicName authority="sp. (NHM _ 186)" class="Bivalvia" family="Cuspidariidae" genus="Myonera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myonera" order="Pholadomyoida" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">Myonera sp. (NHM_186)</taxonomicName>
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<taxonomicName id="18C2786D218D83FB3345B115B0045912" ID-CoL="7P9DY" authority="sp. (NHM _ 186)" family="Cuspidariidae" genus="Myonera" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Myonera" order="Pholadomyoida" pageId="2" pageNumber="3" rank="genus">Myonera sp. (NHM_186)</taxonomicName>
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<subSubSection pageId="2" pageNumber="3" type="materials_examined">
<paragraph pageId="2" pageNumber="3">Materials examined.</paragraph>
<paragraph pageId="2" pageNumber="3">NHM_186 NHMUK 20170037, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/45033e06-fb54-49d5-b632-767e63c1cfd3</paragraph>
<subSubSection id="4B61360E6ADFDD7BB3E1A3DD45B877FE" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="C3AB0778F7189C179D2840416239B14B" pageId="2" pageNumber="3">Materials examined.</paragraph>
<paragraph id="03E410E36ACDF24A1FEA7F72E98D1B19" pageId="2" pageNumber="3">NHM_186 NHMUK 20170037, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/45033e06-fb54-49d5-b632-767e63c1cfd3</paragraph>
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<subSubSection pageId="2" pageNumber="3" type="description">
<paragraph pageId="2" pageNumber="3">Description.</paragraph>
<paragraph pageId="2" pageNumber="3">Shell thin, translucent, sub-ovate tapering posteriorly. Postero-dorsal margin straight. Rostrum short, demarcated by single, carinate radial rib. Sculpture of a few strong, widely spaced, commarginal lamellae, reduced on rostrum. Shell surface minutely pustulose (Fig. 2). Maximum length 1.5 mm, maximum height 1 mm.</paragraph>
<caption pageId="2" pageNumber="3">
<paragraph pageId="2" pageNumber="3">
<subSubSection id="15EFAE5ACC7FAAE278F15A36FD77C618" pageId="2" pageNumber="3" type="description">
<paragraph id="6ECF09A6E857A30FCDF4CFFA37DAA208" pageId="2" pageNumber="3">Description.</paragraph>
<paragraph id="75344762094A17155B6CBC7FE0D7B68D" pageId="2" pageNumber="3">Shell thin, translucent, sub-ovate tapering posteriorly. Postero-dorsal margin straight. Rostrum short, demarcated by single, carinate radial rib. Sculpture of a few strong, widely spaced, commarginal lamellae, reduced on rostrum. Shell surface minutely pustulose (Fig. 2). Maximum length 1.5 mm, maximum height 1 mm.</paragraph>
<caption id="B134C0E7DE8C855238C0A3E8470F2062" pageId="2" pageNumber="3">
<paragraph id="A83B1B3EE79D5D836336379598FBFD05" pageId="2" pageNumber="3">
Figure 2.
<taxonomicName class="Bivalvia" family="Cuspidariidae" genus="Myonera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myonera" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">Myonera</taxonomicName>
<taxonomicName id="17F29AF8294762D6D5A3C99140D9137C" class="Bivalvia" family="Cuspidariidae" genus="Myonera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myonera" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">Myonera</taxonomicName>
sp. (NHM_186) A Live specimen imaged at sea, slightly broken shell with live animal B Detail of hinge C Detail of shell ornamentation. Scale bar: 0.5 mm (A). Image attribution Glover, Dahlgren and Wiklund, 2017.
</paragraph>
</caption>
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<subSubSection pageId="2" pageNumber="3" type="genetic data">
<paragraph pageId="2" pageNumber="3">Genetic data.</paragraph>
<paragraph pageId="2" pageNumber="3">GenBank NHM_186 18S-MF157481, COI-MF157508.</paragraph>
<subSubSection id="FA31EE2D03573BCA3661207AD61F2E65" pageId="2" pageNumber="3" type="genetic data">
<paragraph id="863E985E833546BAF4EA2FCF73124395" pageId="2" pageNumber="3">Genetic data.</paragraph>
<paragraph id="6338C1C3BDC3C6110A0D194C4D73ADA6" pageId="2" pageNumber="3">GenBank NHM_186 18S-MF157481, COI-MF157508.</paragraph>
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<subSubSection pageId="2" pageNumber="3" type="remarks">
<paragraph pageId="2" pageNumber="3">Remarks.</paragraph>
<paragraph pageId="2" pageNumber="3">
<subSubSection id="0E9083B2011559E907CD6C71D5C584F4" pageId="2" pageNumber="3" type="remarks">
<paragraph id="F586A43E81F807C69058367E9983000B" pageId="2" pageNumber="3">Remarks.</paragraph>
<paragraph id="0B75FABF46117D2758946220DBA01067" pageId="2" pageNumber="3">
The species resembles the supposedly cosmopolitan form
<taxonomicName class="Bivalvia" family="Cuspidariidae" genus="Myonera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myonera alleni" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="alleni">Myonera alleni</taxonomicName>
<taxonomicName id="BEDC5ACE14FF5ABD1DE785EF908BF861" class="Bivalvia" family="Cuspidariidae" genus="Myonera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myonera alleni" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="alleni">Myonera alleni</taxonomicName>
Poutiers &amp; Bernard, 1995, previously as
<taxonomicName class="Bivalvia" family="Cuspidariidae" genus="Myonera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myonera atlantica" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="atlantica">Myonera atlantica</taxonomicName>
<taxonomicName id="CD299D06FD062CF1CB4E2D70E5DA4228" class="Bivalvia" family="Cuspidariidae" genus="Myonera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myonera atlantica" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="atlantica">Myonera atlantica</taxonomicName>
(Allen &amp; Morgan, 1981). However, the type locality for this species is from the deep north Atlantic and no genetic data are available for comparison. No similar species is recorded from deep water of the eastern Pacific. Forms a unique monophyletic clade with two other cuspidariid species distinct from all other AB01 specimens (Fig. 5). No genetic matches on GenBank.
</paragraph>
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<paragraph pageId="2" pageNumber="3">Ecology.</paragraph>
<paragraph pageId="2" pageNumber="3">Found in polymetallic nodule province.</paragraph>
<subSubSection id="F9317DF2966AEAFE31818E17537BEE0C" pageId="2" pageNumber="3" type="ecology">
<paragraph id="3CA96BBF2134280647008492768C77D2" pageId="2" pageNumber="3">Ecology.</paragraph>
<paragraph id="4021F07F26E002889CAA2ED5E86F4FD1" pageId="2" pageNumber="3">Found in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
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@ -1,115 +1,114 @@
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<paragraph pageId="3" pageNumber="166">Taxon classification Animalia Pantopoda Phoxichilidiidae</paragraph>
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<paragraph id="25A5373E4A9067C1E88A51C9056B6B0A" pageId="3" pageNumber="166">Taxon classification Animalia Pantopoda Phoxichilidiidae</paragraph>
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<paragraph pageId="3" pageNumber="166">
<taxonomicName authority="Kroyer, 1844" authorityName="Kroyer" authorityYear="1844" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus petiolatus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="petiolatus">
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<paragraph id="516C0A5AF97FEE1F66275C511CE0B877" pageId="3" pageNumber="166">
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Anoplodactylus petiolatus (
<normalizedToken originalValue="Krøyer">Kroyer</normalizedToken>
<normalizedToken id="9367BB7B1B3128BCC75DC69705900D94" originalValue="Krøyer">Kroyer</normalizedToken>
, 1844)
</taxonomicName>
Figures 45, 48
</paragraph>
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<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus guyanensis" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="guyanensis">Anoplodactylus guyanensis</taxonomicName>
<subSubSection id="2A1281CA8CE606EA96CEECBBF31EE47E" pageId="3" pageNumber="166" type="reference_group">
<paragraph id="DCA346787A9D2DF8705344F59A07D107" pageId="3" pageNumber="166">
<taxonomicName id="2704B28E027BBF14DCA65B07F6EC03B1" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus guyanensis" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="guyanensis">Anoplodactylus guyanensis</taxonomicName>
Child, 1977
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus hedgpethi" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="hedgpethi">Anoplodactylus hedgpethi</taxonomicName>
<paragraph id="C003641B224A0E453B7DEE308957C6B3" pageId="3" pageNumber="166">
<taxonomicName id="76C8D9A97B41C1051B88354C9146F537" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus hedgpethi" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="hedgpethi">Anoplodactylus hedgpethi</taxonomicName>
Bacescu, 1959
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus longicollis" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="longicollis">Anoplodactylus longicollis</taxonomicName>
<paragraph id="EAEA4C3688EE14933C33C4181E4862C4" pageId="3" pageNumber="166">
<taxonomicName id="4B5A4204C773D9B07AC4EE97B09A5D74" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus longicollis" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="longicollis">Anoplodactylus longicollis</taxonomicName>
(Dohrn, 1881)
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus pygmaeus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="pygmaeus">Anoplodactylus pygmaeus</taxonomicName>
<paragraph id="A55298C6CA29A76A7378AF4A06A3ADB8" pageId="3" pageNumber="166">
<taxonomicName id="B2915A514E61510CB8EED03CCF333077" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus pygmaeus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="pygmaeus">Anoplodactylus pygmaeus</taxonomicName>
(Hoek, 1881)
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName authority="Johnston, 1837" authorityName="Johnston" authorityYear="1837" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pallene attenuata" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="attenuata">Pallene attenuata</taxonomicName>
<paragraph id="0884E032AFAD5E21F5685FD8348F5464" pageId="3" pageNumber="166">
<taxonomicName id="80257CBBD2FC6FEF9E41AC117B49C8B5" class="Pycnogonida" family="Phoxichilidiidae" genus="Pallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pallene attenuata" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="attenuata">Pallene attenuata</taxonomicName>
Hodge, 1864
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium attenuatus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="attenuatus">Phoxichilidium attenuatus</taxonomicName>
<paragraph id="695BC9897F85191F5B78A98434CF8835" pageId="3" pageNumber="166">
<taxonomicName id="C7ED9328521C284B7EE2C8B0BD3F574A" class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium attenuatus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="attenuatus">Phoxichilidium attenuatus</taxonomicName>
(Hodge, 1863)
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium longicolle" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="longicolle">Phoxichilidium longicolle</taxonomicName>
<paragraph id="027F60C80A4A5A1776B42002C5031F6D" pageId="3" pageNumber="166">
<taxonomicName id="9821FB7720BCFD71FD1E0DB4BDE79437" class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium longicolle" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="longicolle">Phoxichilidium longicolle</taxonomicName>
Dohrn, 1881
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium petiolatum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="petiolatum">Phoxichilidium petiolatum</taxonomicName>
<normalizedToken originalValue="Krøyer">Kroyer</normalizedToken>
<paragraph id="DEABAB13F96EDC4B4201F33BA9E6F564" pageId="3" pageNumber="166">
<taxonomicName id="F39BABC14A3C2C520BA96FFCACBCAEA4" class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium petiolatum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="petiolatum">Phoxichilidium petiolatum</taxonomicName>
<normalizedToken id="DDC628F1B0B0D360531FD4A113AFE51F" originalValue="Krøyer">Kroyer</normalizedToken>
, 1844
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="166" type="material">
<paragraph pageId="3" pageNumber="166">Material.</paragraph>
<paragraph pageId="3" pageNumber="166">ZSMA20071480: male; Banyuls-sur-mer, France; 07.2004. ZSMA20042384: male; Bayuls-sur-Mer, France; 06.2003.</paragraph>
<subSubSection id="02EB7AFA5D756CCB145C0B86C58F8AC9" pageId="3" pageNumber="166" type="material">
<paragraph id="0C12969C2E9E13BC2FE09DE6E6287D20" pageId="3" pageNumber="166">Material.</paragraph>
<paragraph id="EDED7D2CFE197E83FF5AF882AC879710" pageId="3" pageNumber="166">ZSMA20071480: male; Banyuls-sur-mer, France; 07.2004. ZSMA20042384: male; Bayuls-sur-Mer, France; 06.2003.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="166" type="remarks">
<paragraph pageId="3" pageNumber="166">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="166">
<subSubSection id="DA4DE5737B6DBA1E3DA21EC325DA6A85" pageId="3" pageNumber="166" type="remarks">
<paragraph id="D8D884B77C3902B7131A65F06DB3E5E0" pageId="3" pageNumber="166">Remarks.</paragraph>
<paragraph id="BC5F4F5560CCDC9A040AEB9559BFEDDE" pageId="3" pageNumber="166">
In literature there is diverse information about the articulation of the oviger, 7 articles in
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<bibRefCitation id="68854D38093C9B5673C3B851C282862B" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
and 6 articles in
<bibRefCitation author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
<bibRefCitation id="009FC33C840427FA75847008B212798E" author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
and
<bibRefCitation author="King, PE" journalOrPublisher="Field Studies" pageId="5" pageNumber="168" pagination="493 - 516" title="Sea spiders: a revised key to the adults of littoral Pycnogonida in the British Isles." volume="6" year="1986">King (1986)</bibRefCitation>
<bibRefCitation id="D8DEB26EDF26177AE429B07701E2B2C5" author="King, PE" journalOrPublisher="Field Studies" pageId="5" pageNumber="168" pagination="493 - 516" title="Sea spiders: a revised key to the adults of littoral Pycnogonida in the British Isles." volume="6" year="1986">King (1986)</bibRefCitation>
are reported. Here the ovigers have 6 articles.
</paragraph>
</subSubSection>

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@ -1,76 +1,75 @@
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<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Endeidae" genus="Endeis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endeis charybdaea" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="charybdaea">Endeis charybdaea (Dohrn, 1881)</taxonomicName>
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Figures 38, 39, 40
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<taxonomicName authority="Stebbing, 1902" authorityName="Stebbing" authorityYear="1902" class="Pycnogonida" family="Endeidae" genus="Chilophoxus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chilophoxus charybdaeus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="charybdaeus">Chilophoxus charybdaeus</taxonomicName>
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<taxonomicName authority="Latreille, 1816" authorityName="Latreille" authorityYear="1816" class="Pycnogonida" family="Endeidae" genus="Phoxichilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phoxichilus charybdaeus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="charybdaeus">Phoxichilus charybdaeus</taxonomicName>
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<taxonomicName id="D471CC7B705EBE8112837C93287A0ED4" class="Pycnogonida" family="Endeidae" genus="Phoxichilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilus charybdaeus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="charybdaeus">Phoxichilus charybdaeus</taxonomicName>
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<paragraph pageId="3" pageNumber="166">Material.</paragraph>
<paragraph pageId="3" pageNumber="166">ZSMA20071497: male; Rovinj, Croatia; 04.09.2006; 30 m.</paragraph>
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<paragraph id="76014DFB28A63A0CCF5C0F2AE30E2663" pageId="3" pageNumber="166">ZSMA20071497: male; Rovinj, Croatia; 04.09.2006; 30 m.</paragraph>
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<taxonomicName authority="Dall, 1908" authorityName="Dall" authorityYear="1908" class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Pholadomyoida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar (Dall, 1908)</taxonomicName>
<treatment id="12D3DC37B39B5D35CC81BC44CBD55D57" ID-GBIF-Taxon="135490002" LSID="urn:lsid:plazi:treatment:12D3DC37B39B5D35CC81BC44CBD55D57" httpUri="http://treatment.plazi.org/id/12D3DC37B39B5D35CC81BC44CBD55D57" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" scope_class="Bivalvia" scope_order="Nuculanoida" scope_phylum="Mollusca" scope_subClass="Protobranchia">
<subSubSection id="16FD6F0F82D6D94829B7323B14BFA6B0" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="F321DE98291DED7D3123FFE338E44A86" pageId="6" pageNumber="7">
<taxonomicName id="44C66E94332B70864E24D525A89C2816" authority="Dall, 1908" authorityName="Dall" authorityYear="1908" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Pholadomyoida" pageId="6" pageNumber="7" rank="species" species="calcar">Bathyspinula calcar (Dall, 1908)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph pageId="6" pageNumber="7">Material examined.</paragraph>
<paragraph pageId="6" pageNumber="7">NHM_146 NHMUK 20170044, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/61f15e3c-f070-48a1-b484-780b37f7feb6</paragraph>
<paragraph pageId="7" pageNumber="8">
<pageBreakToken pageId="7" pageNumber="8" start="start">NHM</pageBreakToken>
<subSubSection id="0B975680D02C8CC955BF5CF23AFE1227" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="26D06DCFDC8846D37B984AE6DA562744" pageId="6" pageNumber="7">Material examined.</paragraph>
<paragraph id="0079A4C433A9998917851BCD9E7C236A" pageId="6" pageNumber="7">NHM_146 NHMUK 20170044, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/61f15e3c-f070-48a1-b484-780b37f7feb6</paragraph>
<paragraph id="1D011EB07CB5A600E7C80DAE964A8486" pageId="7" pageNumber="8">
<pageBreakToken id="1897EBAA7705BBDB694BE3CAFE14D297" pageId="7" pageNumber="8" start="start">NHM</pageBreakToken>
_149A NHMUK 20170045, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/c44da298-9b61-4d6d-a1cd-2d6c3bd70859
</paragraph>
<paragraph pageId="7" pageNumber="8">NHM_181 NHMUK 20170043, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/3ab74908-1a5d-465f-890c-49373a44906c</paragraph>
<paragraph pageId="7" pageNumber="8">NHM_284 NHMUK 20170046, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/ad2cb87b-1fce-415d-ab45-1619bbc4352b</paragraph>
<paragraph id="E52296877DE5A06A233A45CAF6642C84" pageId="7" pageNumber="8">NHM_181 NHMUK 20170043, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/3ab74908-1a5d-465f-890c-49373a44906c</paragraph>
<paragraph id="9D02AA1F16257E515D6C59A73CE649A9" pageId="7" pageNumber="8">NHM_284 NHMUK 20170046, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/ad2cb87b-1fce-415d-ab45-1619bbc4352b</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="description">
<paragraph pageId="7" pageNumber="8">Description.</paragraph>
<paragraph pageId="7" pageNumber="8">Shell sub-ovate, laterally compressed, with long, sharply pointed posterior rostrum. Periostracum shiny, medium brown. Posterior rostrum shorter, less defined in juveniles. Voucher specimen NHM_181 shell length 13.5 mm, width 7.6 mm (Fig. 6A).</paragraph>
<caption pageId="7" pageNumber="8">
<paragraph pageId="7" pageNumber="8">
<subSubSection id="3EB9F2EDA78885ED91CD12EE57600DA3" pageId="7" pageNumber="8" type="description">
<paragraph id="FD9FD83315861A1F6F464CB1DF13865A" pageId="7" pageNumber="8">Description.</paragraph>
<paragraph id="53F3663290805422687FEB133995073E" pageId="7" pageNumber="8">Shell sub-ovate, laterally compressed, with long, sharply pointed posterior rostrum. Periostracum shiny, medium brown. Posterior rostrum shorter, less defined in juveniles. Voucher specimen NHM_181 shell length 13.5 mm, width 7.6 mm (Fig. 6A).</paragraph>
<caption id="A91575BE8987E1B4302D394AC17A27C9" pageId="7" pageNumber="8">
<paragraph id="E41653D66FDB2059AAADAD9153419CB0" pageId="7" pageNumber="8">
Figure 6.
<taxonomicName class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
<taxonomicName id="7B009D8827C671090D0ECF9ECE116D41" class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
(Dall, 1908) A Specimen NHM_181, Image of live specimen after recovery, length 13.5 mm
<normalizedToken originalValue="BD">B-D</normalizedToken>
<normalizedToken id="417B10A93FF01AA96E1BB1CC7524F8C8" originalValue="BD">B-D</normalizedToken>
Specimen NHM_149A confirmed juvenile
<taxonomicName lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
<taxonomicName id="0DC07D715000481D232D117F5247F1BB" lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
using DNA evidence, total length of animal ~2mm. Scale bars: 5 mm (A); 1 mm (
<normalizedToken originalValue="BD">B-D</normalizedToken>
<normalizedToken id="71ED475BB01DDBC01C88C13D51996E23" originalValue="BD">B-D</normalizedToken>
). Image attribution Glover, Taylor, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="genetic data">
<paragraph pageId="7" pageNumber="8">Genetic data.</paragraph>
<paragraph pageId="7" pageNumber="8">GenBank NHM_146 18S-157475, COI-MF157505; NHM_149A COI-MF157506; NHM_181 18S-MF157479, COI-MF157507; NHM_284 COI-MF157514.</paragraph>
<subSubSection id="4434D29119574D203BEC8B09FC225B59" pageId="7" pageNumber="8" type="genetic data">
<paragraph id="DA2492B33543364FD353575272271C03" pageId="7" pageNumber="8">Genetic data.</paragraph>
<paragraph id="AA96A1C180A7B6B0B7A166924F6361B8" pageId="7" pageNumber="8">GenBank NHM_146 18S-157475, COI-MF157505; NHM_149A COI-MF157506; NHM_181 18S-MF157479, COI-MF157507; NHM_284 COI-MF157514.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="remarks">
<paragraph pageId="7" pageNumber="8">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="8">
<subSubSection id="E23AE99B24571AA4BFB7B04BAD60363F" pageId="7" pageNumber="8" type="remarks">
<paragraph id="FF6998F692A430061AF04F6E7F12B074" pageId="7" pageNumber="8">Remarks.</paragraph>
<paragraph id="0FD94190276034E9801228AAB4489A34" pageId="7" pageNumber="8">
Widely distributed in the eastern Pacific at depths of 400-5000 m (see
<bibRefCitation author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott 2012</bibRefCitation>
<bibRefCitation id="F70936D218EA28F63980F99D4A91A96F" author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott 2012</bibRefCitation>
). The holotype (USNM 110573) was collected 725 km west of Trujillo, Peru at 2370 fathoms (4334 m). Forms a unique monophyletic clade distinct from all other AB01 specimens. Genetic match in 18S to
<taxonomicName class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
<taxonomicName id="767A0DCD033E726BE7B7D1E0B4B65D5A" class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
(GenBank KC993875) from the north eastern Pacific (
<bibRefCitation author="Sharma, PP" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="31" pageNumber="32" pagination="188 - 204" title="Into the deep: A phylogenetic approach to the bivalve subclass Protobranchia." url="https://doi.org/10.1016/j.ympev.2013.05.018" volume="69" year="2013">Sharma et al. 2013</bibRefCitation>
<bibRefCitation id="398793AF67B623BCA691D8203B06D5A3" author="Sharma, PP" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="31" pageNumber="32" pagination="188 - 204" title="Into the deep: A phylogenetic approach to the bivalve subclass Protobranchia." url="https://doi.org/10.1016/j.ympev.2013.05.018" volume="69" year="2013">Sharma et al. 2013</bibRefCitation>
), but as the GenBank 18S sequence from
<taxonomicName lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
<taxonomicName id="37A5877AE54DB451CFECDD2E1E0987DC" lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
was only 289 bp long and as that specimen lacked COI, it was not included in the analyses. Some very small juvenile specimens (Fig. 6
<normalizedToken originalValue="BD">B-D</normalizedToken>
<normalizedToken id="98462F9B56CD1C155439F83F6090244C" originalValue="BD">B-D</normalizedToken>
) were recovered that superficially resemble
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
<taxonomicName id="3CBCE05B1B9162EFDFDF129F1F1B7EE5" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
sp. n. (Fig. 7) and may be easily confused. Genetic data confirmed these to be
<taxonomicName class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
<taxonomicName id="B45F79C6C7AFE9C29CC38B09791763B7" class="Bivalvia" family="Bathyspinulidae" genus="Bathyspinula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathyspinula calcar" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="calcar">Bathyspinula calcar</taxonomicName>
(Fig. 12). These may be distinguised from
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
<taxonomicName id="9C953CECD81A83F6171FBFC252E74D4A" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
by the shiny and iridescent nature of the shell surface of
<taxonomicName lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
<taxonomicName id="6191BD8E1684951A8EDB2A213DD47444" lsidName="B. calcar" pageId="7" pageNumber="8" rank="species" species="calcar">B. calcar</taxonomicName>
, which is preserved in the juveniles.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="ecology">
<paragraph pageId="7" pageNumber="8">Ecology.</paragraph>
<paragraph pageId="7" pageNumber="8">Relatively large bivalve recovered from epibenthic sledge tow in polymetallic nodule province.</paragraph>
<subSubSection id="8540BEAC3E411FF66F58FDBBB8D17522" pageId="7" pageNumber="8" type="ecology">
<paragraph id="80E0E73AC759F778E8E0C16EF23E13B6" pageId="7" pageNumber="8">Ecology.</paragraph>
<paragraph id="D3F97C4405A08FB6AF6F24613C9730F2" pageId="7" pageNumber="8">Relatively large bivalve recovered from epibenthic sledge tow in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph pageId="15" pageNumber="16">Material examined.</paragraph>
<paragraph pageId="15" pageNumber="16">NHM_453 NHMUK 20170069.1-2, collected 2013-10-21, 13.90165 -116.59, 4163 m. http://data.nhm.ac.uk/object/ce9cbed0-82cc-420d-baad-fdfff7cc0986</paragraph>
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<paragraph id="63D2F9E84F84595E4A541B013A02D521" pageId="15" pageNumber="16">Material examined.</paragraph>
<paragraph id="848CBCC892EE8D0D095013418F5274D3" pageId="15" pageNumber="16">NHM_453 NHMUK 20170069.1-2, collected 2013-10-21, 13.90165 -116.59, 4163 m. http://data.nhm.ac.uk/object/ce9cbed0-82cc-420d-baad-fdfff7cc0986</paragraph>
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<pageBreakToken pageId="16" pageNumber="17" start="start">Description</pageBreakToken>
<subSubSection id="6A85D0C29DAFE64A204BEF2733A1CF01" pageId="16" pageNumber="17" type="description">
<paragraph id="F4D741DA8F4C0BEDBD6DB5C1A1286E69" pageId="16" pageNumber="17">
<pageBreakToken id="3F3E1BEA713BBE4319CF168839EEE797" pageId="16" pageNumber="17" start="start">Description</pageBreakToken>
.
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<paragraph pageId="16" pageNumber="17">Subcircular to slightly oblique with slightly sinuous posterior margin. Periostracum with short, fine, bristles aligned in radial rows. Ligament small, triangular, set in shallow resilifer. Hinge teeth robust, 4 anterior and 5 posterior. Inner shell margin smooth. Voucher NHM_453 shell length 4.6 mm, height 4.3mm (Fig. 15).</paragraph>
<caption pageId="16" pageNumber="17">
<paragraph pageId="16" pageNumber="17">
<paragraph id="24002AF42847A8D56C4398A0292E3D03" pageId="16" pageNumber="17">Subcircular to slightly oblique with slightly sinuous posterior margin. Periostracum with short, fine, bristles aligned in radial rows. Ligament small, triangular, set in shallow resilifer. Hinge teeth robust, 4 anterior and 5 posterior. Inner shell margin smooth. Voucher NHM_453 shell length 4.6 mm, height 4.3mm (Fig. 15).</paragraph>
<caption id="CCCEEF1AAFB73C7E8CE4BF7E2CA334A3" pageId="16" pageNumber="17">
<paragraph id="D88A2FFD037B454CF023AA58261668F6" pageId="16" pageNumber="17">
Figure 15.
<taxonomicName class="Bivalvia" family="Limopsidae" genus="Limopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limopsis" order="Arcida" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Limopsis</taxonomicName>
<taxonomicName id="D1E358762353EAF9C719A46F6267F536" class="Bivalvia" family="Limopsidae" genus="Limopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limopsis" order="Arcida" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Limopsis</taxonomicName>
sp. (NHM_453) A Specimen NHM_453 live after recovery B Specimen NHM_453 after preservation
<normalizedToken originalValue="CD">C-D</normalizedToken>
<normalizedToken id="41873EC9D0352266B02EAEA2E40A7801" originalValue="CD">C-D</normalizedToken>
SEM of interior of right valve showing hinge teeth. Scale bars: 2mm B, 0.5mm D. Image attribution Glover, Taylor, Dahlgren &amp; Wiklund, 2017.
</paragraph>
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<subSubSection pageId="16" pageNumber="17" type="genetic data">
<paragraph pageId="16" pageNumber="17">Genetic data.</paragraph>
<paragraph pageId="16" pageNumber="17">GenBank NHM_453 18S-MF157498, COI-MF157524.</paragraph>
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<paragraph id="5E9351A405294E0079D206DFA2660099" pageId="16" pageNumber="17">Genetic data.</paragraph>
<paragraph id="79B83A187B1297288E94602B28B019D5" pageId="16" pageNumber="17">GenBank NHM_453 18S-MF157498, COI-MF157524.</paragraph>
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<subSubSection pageId="16" pageNumber="17" type="remarks">
<paragraph pageId="16" pageNumber="17">Remarks.</paragraph>
<paragraph pageId="16" pageNumber="17">
<subSubSection id="623EC4E6292B7B1884CCECE6BFABF277" pageId="16" pageNumber="17" type="remarks">
<paragraph id="1C6BCA9E4F92C3D7156986FAEDCB97E9" pageId="16" pageNumber="17">Remarks.</paragraph>
<paragraph id="DFC2882678D9D8785FC8537A646C598F" pageId="16" pageNumber="17">
Dissimilar in shape and periostracal bristle configuration to any recorded Eastern Pacific deep-water species (Coan &amp; Valentich-Scott 2012). However, shape and number of hinge teeth are known to change with age/size in
<taxonomicName class="Bivalvia" family="Limopsidae" genus="Limopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limopsis" order="Arcida" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Limopsis</taxonomicName>
<taxonomicName id="17B0E1938950F6C0B821FE0DB703B217" class="Bivalvia" family="Limopsidae" genus="Limopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limopsis" order="Arcida" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Limopsis</taxonomicName>
species. In molecular analysis (Fig. 17) forms part of a well supported monophyletic clade with other
<taxonomicName class="Bivalvia" family="Limopsidae" genus="Limopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limopsis" order="Arcida" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Limopsis</taxonomicName>
<taxonomicName id="5038C85D3CA53CEC47ABB7CE69F220E3" class="Bivalvia" family="Limopsidae" genus="Limopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limopsis" order="Arcida" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Limopsis</taxonomicName>
species and aligns closest to
<taxonomicName class="Bivalvia" family="Limopsidae" genus="Limopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limopsis marionensis" order="Arcida" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="marionensis">Limopsis marionensis</taxonomicName>
<taxonomicName id="42B8C4FA94226DE9706587BEC01B435F" class="Bivalvia" family="Limopsidae" genus="Limopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limopsis marionensis" order="Arcida" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="marionensis">Limopsis marionensis</taxonomicName>
Smith, 1885 from depths of 40-1000 m in the Southern Ocean. No genetic matches on GenBank.
</paragraph>
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<subSubSection pageId="16" pageNumber="17" type="ecology">
<paragraph pageId="16" pageNumber="17">Ecology.</paragraph>
<paragraph pageId="16" pageNumber="17">Found in polymetallic nodule province.</paragraph>
<subSubSection id="B218269FA6E92F62F8EDB9290FE2FC14" pageId="16" pageNumber="17" type="ecology">
<paragraph id="99F2EBF14D18513E2303E0833D7A040D" pageId="16" pageNumber="17">Ecology.</paragraph>
<paragraph id="4304E67E192B4A4463202D8B55755973" pageId="16" pageNumber="17">Found in polymetallic nodule province.</paragraph>
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<taxonomicName id="97574CF3A701A1C25C3E1465C0FE7C59" ID-CoL="7NFB8" authority="sp. (NHM _ 192)" class="Scaphopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" order="Gadilida" pageId="23" pageNumber="24" phylum="Mollusca" rank="order">Gadilida sp. (NHM_192)</taxonomicName>
</paragraph>
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<subSubSection pageId="23" pageNumber="24" type="materials_examined">
<paragraph pageId="23" pageNumber="24">Material examined.</paragraph>
<paragraph pageId="23" pageNumber="24">NHM_192 NHMUK 20170075, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/fc0e3ae8-9cce-46a0-bb8b-fafe0e2cb46b</paragraph>
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<paragraph id="54FAA35BF3811AE5FC48FD4AA63D6368" pageId="23" pageNumber="24">Material examined.</paragraph>
<paragraph id="13F7B6779FB7139F77B751F55FE288D1" pageId="23" pageNumber="24">NHM_192 NHMUK 20170075, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/fc0e3ae8-9cce-46a0-bb8b-fafe0e2cb46b</paragraph>
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<pageBreakToken pageId="24" pageNumber="25" start="start">Description</pageBreakToken>
<subSubSection id="E2F783FED4C11B37D41BBB1246F4CB46" pageId="24" pageNumber="25" type="description">
<paragraph id="7C5876E3A042F71574E06DD0EBC787FD" pageId="24" pageNumber="25">
<pageBreakToken id="2E901E6C832743BDF4C92E1B43CFAC33" pageId="24" pageNumber="25" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="24" pageNumber="25">Slender, smooth, transparent, annular growth increments, maximum diameter at mouth. Voucher NHM_192, length 4 mm, maximum width 0.5 mm (Fig. 23B).</paragraph>
<paragraph id="53EDEF61E6D2683AB08DE03FC7986A12" pageId="24" pageNumber="25">Slender, smooth, transparent, annular growth increments, maximum diameter at mouth. Voucher NHM_192, length 4 mm, maximum width 0.5 mm (Fig. 23B).</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="genetic data">
<paragraph pageId="24" pageNumber="25">Genetic data.</paragraph>
<paragraph pageId="24" pageNumber="25">GenBank NHM_192 16S-MF157459, 18S-MF157483.</paragraph>
<subSubSection id="AC9ADB4DF9DB56399033A85E6B95838F" pageId="24" pageNumber="25" type="genetic data">
<paragraph id="4BD393E2F3DB4C28D204B6619BD44C3C" pageId="24" pageNumber="25">Genetic data.</paragraph>
<paragraph id="76D79FB3233B9AF8BBC22BCEBE29FF3D" pageId="24" pageNumber="25">GenBank NHM_192 16S-MF157459, 18S-MF157483.</paragraph>
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<subSubSection pageId="24" pageNumber="25" type="remarks">
<paragraph pageId="24" pageNumber="25">Remarks.</paragraph>
<paragraph pageId="24" pageNumber="25">
<subSubSection id="6EA8615F4E06A0804B285C7BE3E9BA5D" pageId="24" pageNumber="25" type="remarks">
<paragraph id="5D8372182EC4678C3EA5EB2B9421AF4F" pageId="24" pageNumber="25">Remarks.</paragraph>
<paragraph id="E129F08D16529E880F68830BD7525C09" pageId="24" pageNumber="25">
Forms a unique monophyletic clade distinct from other AB01 specimens. No genetic matches on GenBank. Phylogenetic tree (Fig. 24) supports placement in order
<taxonomicName lsidName="Pteriomorphia" order="Gadilida" pageId="24" pageNumber="25" rank="order">Gadilida</taxonomicName>
<taxonomicName id="EAE4A540FBC3D5D1B5A42F1BEDB8DB4A" class="Scaphopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" order="Gadilida" pageId="24" pageNumber="25" phylum="Mollusca" rank="order">Gadilida</taxonomicName>
with NHM_345. Genetic and image data made available for future study when better specimens available.
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="ecology">
<paragraph pageId="24" pageNumber="25">Ecology.</paragraph>
<paragraph pageId="24" pageNumber="25">Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.</paragraph>
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<paragraph id="845106ACDD297512C9C38B7487F66645" pageId="24" pageNumber="25">Ecology.</paragraph>
<paragraph id="B19426A98D9C3D4FACFC42B7739C58C6" pageId="24" pageNumber="25">Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.</paragraph>
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<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella biunguiculata" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="biunguiculata">Ammothella biunguiculata (Dohrn, 1881)</taxonomicName>
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Figures 3, 15, 16
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<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea biunguiculata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="biunguiculata">
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<taxonomicName id="CD3038CD45AAC51BC4648D5D4F6327F6" class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea biunguiculata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="biunguiculata">
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biunguiculata
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Dohrn, 1881
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<paragraph pageId="2" pageNumber="165">ZSMA20071473: male; Cape Savudrija, Croatia; 08.10.2004: 0-1 m; on stone.</paragraph>
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<paragraph pageId="1" pageNumber="164">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
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<paragraph id="EF092F4FA382DE8D3C124DD4D622B05A" pageId="1" pageNumber="164">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
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<paragraph pageId="1" pageNumber="164">
<taxonomicName authority="Costa, 1861" authorityName="Costa" authorityYear="1861" class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia vulgaris" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="vulgaris">Achelia vulgaris (Costa, 1861)</taxonomicName>
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<paragraph id="9FDF08C14DD5849EC34B87E2D53364DF" pageId="1" pageNumber="164">
<taxonomicName id="7D2264355F8CAC1A5B3B58C796EB77A4" ID-CoL="96M7" authority="Costa, 1861" authorityName="Costa" authorityYear="1861" class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia vulgaris" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="vulgaris">Achelia vulgaris (Costa, 1861)</taxonomicName>
Figures 2, 11, 12
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<paragraph pageId="1" pageNumber="164">
<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia franciscana" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="franciscana">Achelia franciscana</taxonomicName>
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<paragraph id="C1748E9578038D0C44CFDBE0F04632E0" pageId="1" pageNumber="164">
<taxonomicName id="8C29FB3E250123963BCF155862C02FB9" class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia franciscana" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="franciscana">Achelia franciscana</taxonomicName>
(Dohrn, 1881)
</paragraph>
<paragraph pageId="1" pageNumber="164">
<taxonomicName authority="Deyrolle, 1864" authorityName="Deyrolle" authorityYear="1864" class="Insecta" family="Buprestidae" genus="Alcinous" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alcinous vulgaris" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="vulgaris">Alcinous vulgaris</taxonomicName>
<paragraph id="8D6606826B7B678DBA4DE6D9759DB9D5" pageId="1" pageNumber="164">
<taxonomicName id="8F02CC932B7A08006283EC93DE209A9F" class="Insecta" family="Buprestidae" genus="Alcinous" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alcinous vulgaris" order="Coleoptera" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="vulgaris">Alcinous vulgaris</taxonomicName>
Costa, 1861
</paragraph>
<paragraph pageId="1" pageNumber="164">
<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea fransciscana" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="fransciscana">Ammothea fransciscana</taxonomicName>
<paragraph id="1699C2E3CD200D557C978A32161B9071" pageId="1" pageNumber="164">
<taxonomicName id="318A9742589D359DA1407795FEE1FBB3" class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea fransciscana" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="fransciscana">Ammothea fransciscana</taxonomicName>
Dohrn, 1881
</paragraph>
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<subSubSection pageId="1" pageNumber="164" type="material">
<paragraph pageId="1" pageNumber="164">Material.</paragraph>
<paragraph pageId="1" pageNumber="164">ZSMA20071467: male; Bayuls-sur-Mer, France; 07.2004. ZSMA20071468: female; Giglio Island, Italy; 08.04.2005; 8-13 m; brown algae. ZSMA20071469: female; Giglio Island, Italy; 08.04.2005; 8-13 m; brown algae.</paragraph>
<subSubSection id="E3CEDA39E99952825218CC4F7D771E4D" pageId="1" pageNumber="164" type="material">
<paragraph id="7E26ED89170871822ACB9D8210831EA5" pageId="1" pageNumber="164">Material.</paragraph>
<paragraph id="72C3A6C774264DEB4E2C868CC226B50D" pageId="1" pageNumber="164">ZSMA20071467: male; Bayuls-sur-Mer, France; 07.2004. ZSMA20071468: female; Giglio Island, Italy; 08.04.2005; 8-13 m; brown algae. ZSMA20071469: female; Giglio Island, Italy; 08.04.2005; 8-13 m; brown algae.</paragraph>
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<subSubSection pageId="1" pageNumber="164" type="remarks">
<paragraph pageId="1" pageNumber="164">Remarks.</paragraph>
<paragraph pageId="1" pageNumber="164">
<subSubSection id="C7D91096768C12CD43C38DFB33851503" pageId="1" pageNumber="164" type="remarks">
<paragraph id="5032E78CD0C326205B79E72934E47D77" pageId="1" pageNumber="164">Remarks.</paragraph>
<paragraph id="503B1D3DCA1E33BF8AB2822093910BAB" pageId="1" pageNumber="164">
Adult males are differentiated by the character described by
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<bibRefCitation id="D1627D3265011473FC2995D882DA4ABA" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
and
<bibRefCitation author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
<bibRefCitation id="BCAFB4BBFCBFE22F8B8A85E76907F0C3" author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
: three protuberances with spine on each side of coxa 2. But Dohrn notes that there also exist varieties: two protuberances on each side of coxa 2 or two protuberances on one and three protuberances on the other side of coxa 2.
</paragraph>
<paragraph pageId="1" pageNumber="164">
<paragraph id="15A73EB143EECA6BA5D6C48A19517AB4" pageId="1" pageNumber="164">
Here, only coxa 2 of the first leg has three protuberances with spine on the one and two on the other side; the coxa 2 of the other legs has two protuberances with spine on each side (Fig. 11C, D). The other characters of the male, like lateral process not touching each other (Fig. 11B) correspond with the descriptions of
<bibRefCitation author="Costa, OG" journalOrPublisher="Stamperia dell'Iride, Napoli" pageId="5" pageNumber="168" title="Microdoride Mediterranea, o descrizione de'poco ben conosciuti od affatto ignoti viventi minuti e microscopici del Mediterraneo: I." year="1861">Costa (1861)</bibRefCitation>
<bibRefCitation id="EE70F40FCC466FA7478C9216C3E397C1" author="Costa, OG" journalOrPublisher="Stamperia dell'Iride, Napoli" pageId="5" pageNumber="168" title="Microdoride Mediterranea, o descrizione de'poco ben conosciuti od affatto ignoti viventi minuti e microscopici del Mediterraneo: I." year="1861">Costa (1861)</bibRefCitation>
,
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<bibRefCitation id="EE3842788104EB0A1193BAD224E19575" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
and
<bibRefCitation author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
<bibRefCitation id="82E3499CBCCEBF2EADE6B429BB6B3D70" author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
.
</paragraph>
</subSubSection>

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<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia langi" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="langi">Achelia langi (Dohrn, 1881)</taxonomicName>
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Figures 2, 9, 10
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<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea langi" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="langi">Ammothea langi</taxonomicName>
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<taxonomicName id="FB992CAD4D345F30880C25474F673DE2" class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea langi" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="langi">Ammothea langi</taxonomicName>
Dohrn, 1881
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<paragraph pageId="1" pageNumber="164">Material.</paragraph>
<paragraph pageId="1" pageNumber="164">ZSMA20071464: male; Cape Savudrija, Croatia; 08.10.2004: 0-1 m; under stones. ZSMA20071465: male; Cape Savudrija, Croatia; 22.05.2005: 0-1 m. ZSMA20071466: female; Cape Savudrija, Croatia; 08.08.2005; 0-1 m; on stone.</paragraph>
<subSubSection id="EC51E90363F24AAB62D3D85DE87FDE2A" pageId="1" pageNumber="164" type="material">
<paragraph id="3CA14A638CF6E20245D602D0EA2F73D7" pageId="1" pageNumber="164">Material.</paragraph>
<paragraph id="C2ADBEE19E2FB32A3F384B92E7BDF083" pageId="1" pageNumber="164">ZSMA20071464: male; Cape Savudrija, Croatia; 08.10.2004: 0-1 m; under stones. ZSMA20071465: male; Cape Savudrija, Croatia; 22.05.2005: 0-1 m. ZSMA20071466: female; Cape Savudrija, Croatia; 08.08.2005; 0-1 m; on stone.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="164" type="remarks">
<paragraph pageId="1" pageNumber="164">Remarks.</paragraph>
<paragraph pageId="1" pageNumber="164">
<subSubSection id="B61A41136FD36E1FA91017A776E04669" pageId="1" pageNumber="164" type="remarks">
<paragraph id="8E58EC36A81292C28659F54F2913FB47" pageId="1" pageNumber="164">Remarks.</paragraph>
<paragraph id="A5025B6B18EF6EB93AD2E1929514DFA7" pageId="1" pageNumber="164">
According to
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<bibRefCitation id="717F8DD5879E8A6F40A24B2AF66175D6" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
the lateral processes 1-3 are armed with one protuberance with spine. Here, in the male, the lateral process 1 has two protuberances with spine and the lateral process 2 and 3 have one (Fig. 9D, E). The lateral processes in the female correspond with
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description.
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@ -1,104 +1,103 @@
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<paragraph pageId="3" pageNumber="166">Taxon classification Animalia Pantopoda Phoxichilidiidae</paragraph>
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<paragraph id="D514A203A9DB5931EE602C5D70F4CB8C" pageId="3" pageNumber="166">Taxon classification Animalia Pantopoda Phoxichilidiidae</paragraph>
</subSubSection>
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<paragraph pageId="3" pageNumber="166">
<taxonomicName authority="Hodge, 1864" authorityName="Hodge" authorityYear="1864" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus pygmaeus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="pygmaeus">Anoplodactylus pygmaeus (Hodge, 1864)</taxonomicName>
<subSubSection id="7477B86DA17F3F10A04E264BC1C7A3A9" pageId="3" pageNumber="166" type="nomenclature">
<paragraph id="3B0234BFEE1F5A7A5B51BD16B32246B4" pageId="3" pageNumber="166">
<taxonomicName id="1BBBB9C93AAE74A92011C362D92BB842" ID-CoL="5V8RJ" authority="Hodge, 1864" authorityName="Hodge" authorityYear="1864" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus pygmaeus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="pygmaeus">Anoplodactylus pygmaeus (Hodge, 1864)</taxonomicName>
Figures 45, 49, 50, 51, 52
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="166" type="reference_group">
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus derjugini" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="derjugini">Anoplodactylus derjugini</taxonomicName>
<subSubSection id="8F8E406450EEE32273994794AE124901" pageId="3" pageNumber="166" type="reference_group">
<paragraph id="477159028BF4EDEEA954FF5A8331F29E" pageId="3" pageNumber="166">
<taxonomicName id="5F244AD1201AECEAE8AE2C13E978DA23" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus derjugini" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="derjugini">Anoplodactylus derjugini</taxonomicName>
(Losina-Losinsky, 1929)
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus exiguus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="exiguus">Anoplodactylus exiguus</taxonomicName>
<paragraph id="1F9D6BB969B8D34FE4C9AC68B5A7125C" pageId="3" pageNumber="166">
<taxonomicName id="2AC0E8E52C46FA2AE8ED32A4396884CA" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus exiguus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="exiguus">Anoplodactylus exiguus</taxonomicName>
(Dohrn, 1881)
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName authority="Johnston, 1837" authorityName="Johnston" authorityYear="1837" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pallene pygmaea" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="pygmaea">Pallene pygmaea</taxonomicName>
<paragraph id="8EF4BCEBE440503FD87149D425806AE6" pageId="3" pageNumber="166">
<taxonomicName id="1F7F6B21942D18B860CCCA38F1DC9E15" class="Pycnogonida" family="Phoxichilidiidae" genus="Pallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pallene pygmaea" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="pygmaea">Pallene pygmaea</taxonomicName>
Hodge, 1864
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium exiguum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="exiguum">Phoxichilidium exiguum</taxonomicName>
<paragraph id="43781C388696B888514F63C5FF47983F" pageId="3" pageNumber="166">
<taxonomicName id="05EEE867A1F60D16990A97BB00CD5A64" class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium exiguum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="exiguum">Phoxichilidium exiguum</taxonomicName>
Dohrn, 1881
</paragraph>
<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium pygmaeum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="pygmaeum">Phoxichilidium pygmaeum</taxonomicName>
<paragraph id="212E81DD898B4FF17A2ED007F9964FF4" pageId="3" pageNumber="166">
<taxonomicName id="5D29FB43471B87D53AC036CB79D175AC" class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium pygmaeum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="pygmaeum">Phoxichilidium pygmaeum</taxonomicName>
Hoek, 1881
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="166" type="material">
<paragraph pageId="3" pageNumber="166">Material.</paragraph>
<paragraph pageId="3" pageNumber="166">
<subSubSection id="DC57D2EED1F2127F7DE2BA47DB51F155" pageId="3" pageNumber="166" type="material">
<paragraph id="051F16292AAC7DD3019BD5F72C5B53BD" pageId="3" pageNumber="166">Material.</paragraph>
<paragraph id="0AA33555FBBE584FD28D6EBC6D0EBE4C" pageId="3" pageNumber="166">
ZSMA20071481: female; Elba, Italy; 20.10.2004; 3-5 m. ZSMA20071482: female; Elba, Italy; 21.10.2004; 3-5 m. ZSMA20071483: male; Elba, Italy; 20.10.2004; 0.5-4 m. ZSMA20071484: male; Cape Kamenjak, Croatia; 03.05.2007; 0-5 m;
<taxonomicName genus="Stypocaulon" lsidName="Stypocaulon scoparium" pageId="3" pageNumber="166" rank="species" species="scoparium">Stypocaulon scoparium</taxonomicName>
<taxonomicName id="6528011C42AA8EE597021FB6DECD5EC4" genus="Stypocaulon" lsidName="Stypocaulon scoparium" pageId="3" pageNumber="166" rank="species" species="scoparium">Stypocaulon scoparium</taxonomicName>
(Linnaeus)
<normalizedToken originalValue="Kützing">Kuetzing</normalizedToken>
<normalizedToken id="C1CA895FA4BEAA81EBF6D77CADB7FA66" originalValue="Kützing">Kuetzing</normalizedToken>
, 1843.
</paragraph>
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<subSubSection pageId="3" pageNumber="166" type="remarks">
<paragraph pageId="3" pageNumber="166">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="166">
<subSubSection id="F65CFD64C09AD734EB68D7232E220888" pageId="3" pageNumber="166" type="remarks">
<paragraph id="72125197789FCCBD20176F926E8809E3" pageId="3" pageNumber="166">Remarks.</paragraph>
<paragraph id="2AD0C56FD7FE13D64EBE6E63A7A0E0C3" pageId="3" pageNumber="166">
In the literature there is diverse information about the articulation of the oviger, 7 articles in
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<bibRefCitation id="687D1F04EAB3D61137FD171E6C94B018" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
and 6 articles in
<bibRefCitation author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
<bibRefCitation id="4F0E1734CD914CFF7C90E85BE1B1F1A4" author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
and
<bibRefCitation author="King, PE" journalOrPublisher="Field Studies" pageId="5" pageNumber="168" pagination="493 - 516" title="Sea spiders: a revised key to the adults of littoral Pycnogonida in the British Isles." volume="6" year="1986">King (1986)</bibRefCitation>
<bibRefCitation id="1DD977498812F57CA027FDB9CFCEA0B7" author="King, PE" journalOrPublisher="Field Studies" pageId="5" pageNumber="168" pagination="493 - 516" title="Sea spiders: a revised key to the adults of littoral Pycnogonida in the British Isles." volume="6" year="1986">King (1986)</bibRefCitation>
are reported. Here the ovigers have 6 articles (Fig. 50E, F).
</paragraph>
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<paragraph pageId="24" pageNumber="25">Material examined.</paragraph>
<paragraph pageId="24" pageNumber="25">NHM_132 NHMUK 20170077, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/6a1906d9-9ed1-4f6e-a0cf-2d53e2289a01</paragraph>
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<paragraph id="170549257949B7D94A956CB13B9D0572" pageId="24" pageNumber="25">Material examined.</paragraph>
<paragraph id="1C4DEB9C8700F06B05035711E2026C4D" pageId="24" pageNumber="25">NHM_132 NHMUK 20170077, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/6a1906d9-9ed1-4f6e-a0cf-2d53e2289a01</paragraph>
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<paragraph pageId="24" pageNumber="25">Shell slender, smooth, fairly transparent, increasing in diameter to a maximum about 2.5 mm from the anterior aperture, then decreasing towards the mouth. NHM_132 voucher specimen length 16.6 mm, max width 3 mm (Fig. 23D).</paragraph>
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<paragraph id="66BC104E0D113C024CE533BB2873C30A" pageId="24" pageNumber="25">Description.</paragraph>
<paragraph id="F14417AB5BEB9ED54CF401B07995297A" pageId="24" pageNumber="25">Shell slender, smooth, fairly transparent, increasing in diameter to a maximum about 2.5 mm from the anterior aperture, then decreasing towards the mouth. NHM_132 voucher specimen length 16.6 mm, max width 3 mm (Fig. 23D).</paragraph>
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<paragraph id="160570A48CD84190E9B990A68D03DEA7" pageId="24" pageNumber="25">Genetic data.</paragraph>
<paragraph id="33F147D48637F1CEC5C4A64E6D9C9222" pageId="24" pageNumber="25">GenBank NHM_132 16S-MF157456, 18S-MF157472.</paragraph>
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<paragraph pageId="24" pageNumber="25">Remarks.</paragraph>
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<paragraph id="145F8134F6ACB488D9091804ADF87191" pageId="24" pageNumber="25">Remarks.</paragraph>
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Forms a unique monophyletic clade distinct from other AB01 specimens. No genetic matches on GenBank. Phylogenetic tree (Fig. 24) supports place
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in order
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. Genetic and image data made available for future study when better specimens available.
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<paragraph pageId="25" pageNumber="26">Ecology.</paragraph>
<paragraph pageId="25" pageNumber="26">Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.</paragraph>
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<paragraph id="D6D862DBC3E2AEC565915A1962D12D7B" pageId="25" pageNumber="26">Ecology.</paragraph>
<paragraph id="EE02123C864541835B55AAAF351DAE0C" pageId="25" pageNumber="26">Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.</paragraph>
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<taxonomicName authority="sp. (NHM _ 261)" class="Scaphopoda" family="Dentaliidae" genus="Fissidentalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fissidentalium" order="Pholadomyoida" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">Fissidentalium sp. (NHM_261)</taxonomicName>
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<taxonomicName id="3B7A62BCFA384404DD68A2E833C253BE" authority="sp. (NHM _ 261)" family="Dentaliidae" genus="Fissidentalium" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Fissidentalium" order="Pholadomyoida" pageId="22" pageNumber="23" rank="genus">Fissidentalium sp. (NHM_261)</taxonomicName>
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<subSubSection pageId="22" pageNumber="23" type="materials_examined">
<paragraph pageId="22" pageNumber="23">Material examined.</paragraph>
<paragraph pageId="22" pageNumber="23">NHM_261 NHMUK 20170074, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/679fa0ca-d647-446d-87c5-e8d33949efe2</paragraph>
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<paragraph id="6AFA0E73E3ACC1AD9FA525F5A8D4F662" pageId="22" pageNumber="23">Material examined.</paragraph>
<paragraph id="C71C0F1D0FCECD8D9C7E498CD4A899B3" pageId="22" pageNumber="23">NHM_261 NHMUK 20170074, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/679fa0ca-d647-446d-87c5-e8d33949efe2</paragraph>
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<pageBreakToken pageId="23" pageNumber="24" start="start">Description</pageBreakToken>
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<paragraph id="88985484ABC7BB2CA6629CD98634472B" pageId="23" pageNumber="24">
<pageBreakToken id="12075EEE5E4ED9E2BF72A92B7F0FAD0B" pageId="23" pageNumber="24" start="start">Description</pageBreakToken>
.
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<paragraph pageId="23" pageNumber="24">
<paragraph id="3E6ACD24FAFDD83452918DDA4771425D" pageId="23" pageNumber="24">
A damaged shell with rib features and curvature similar to
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species (see Scarabino, 1995). Voucher NHM_261, poor preservation, length 21 mm, maximum width 3.1 mm (Fig. 23A).
</paragraph>
<caption pageId="23" pageNumber="24">
<paragraph pageId="23" pageNumber="24">
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<paragraph id="651C53426FE9A14A0DFF3515B060A9B6" pageId="23" pageNumber="24">
Figure 23.
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<taxonomicName id="CCFD9FDCB011DBE0B37713C12FB022EF" class="Scaphopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" pageId="23" pageNumber="24" phylum="Mollusca" rank="class">Scaphopoda</taxonomicName>
spp. A
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<taxonomicName id="6E5C5FB2562A6474D894483170B43435" class="Scaphopoda" family="Dentaliidae" genus="Fissidentalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fissidentalium" order="Dentaliida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">Fissidentalium</taxonomicName>
sp. (NHM_261) live specimen. B
<taxonomicName lsidName="Pteriomorphia" order="Gadilida" pageId="23" pageNumber="24" rank="order">Gadilida</taxonomicName>
<taxonomicName id="776DEF3D1FCC86C04726B09073733113" class="Scaphopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" order="Gadilida" pageId="23" pageNumber="24" phylum="Mollusca" rank="order">Gadilida</taxonomicName>
sp. (NHM_192) live specimen C
<taxonomicName class="Scaphopoda" family="Gadilidae" genus="Gadila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gadila" order="Gadilida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">Gadila</taxonomicName>
<taxonomicName id="398BAD69287D077B03B7814945F9E109" class="Scaphopoda" family="Gadilidae" genus="Gadila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gadila" order="Gadilida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">Gadila</taxonomicName>
sp. (NHM_345) live specimen D
<taxonomicName lsidName="Pteriomorphia" order="Gadilida" pageId="23" pageNumber="24" rank="order">Gadilida</taxonomicName>
<taxonomicName id="C3A5276D0CCF8ED7B25704163B91759A" class="Scaphopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" order="Gadilida" pageId="23" pageNumber="24" phylum="Mollusca" rank="order">Gadilida</taxonomicName>
sp. (NHM_132) live specimen. Scale bars: 5 mm (A, D); 1 mm (B); 2 mm (C). Image attribution Glover, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
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<subSubSection pageId="23" pageNumber="24" type="genetic data">
<paragraph pageId="23" pageNumber="24">Genetic data.</paragraph>
<paragraph pageId="23" pageNumber="24">GenBank NHM_261 16S-MF157461, 18S-MF157489, COI-MF157511.</paragraph>
<subSubSection id="B8559A31CF929E035CEA239E8638EE0F" pageId="23" pageNumber="24" type="genetic data">
<paragraph id="F372EEC70DA4B72685521F9674393A0F" pageId="23" pageNumber="24">Genetic data.</paragraph>
<paragraph id="94F0CCAB40056DAFCA7D701A710F843A" pageId="23" pageNumber="24">GenBank NHM_261 16S-MF157461, 18S-MF157489, COI-MF157511.</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="remarks">
<paragraph pageId="23" pageNumber="24">Remarks.</paragraph>
<paragraph pageId="23" pageNumber="24">
<subSubSection id="9E7BCD5D461C22B97696D04BAFAAEAE1" pageId="23" pageNumber="24" type="remarks">
<paragraph id="E999C347610A3589E7971CF8BBCDA235" pageId="23" pageNumber="24">Remarks.</paragraph>
<paragraph id="539C103A65F86E823472FB2537B15E70" pageId="23" pageNumber="24">
Forms a unique monophyletic clade distinct from other AB01 specimens. In the molecular analysis it groups with other
<taxonomicName class="Scaphopoda" family="Dentaliidae" genus="Fissidentalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fissidentalium" order="Dentaliida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">Fissidentalium</taxonomicName>
<taxonomicName id="0252CC4651575CC7BF8B777B9734E127" class="Scaphopoda" family="Dentaliidae" genus="Fissidentalium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fissidentalium" order="Dentaliida" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">Fissidentalium</taxonomicName>
species, but with very low support. No genetic matches on GenBank. Phylogenetic tree supports placement in order
<taxonomicName lsidName="Pteriomorphia" order="Dentaliida" pageId="23" pageNumber="24" rank="order">Dentaliida</taxonomicName>
<taxonomicName id="810F652034A613A086CB565EC27C9105" class="Scaphopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" order="Dentaliida" pageId="23" pageNumber="24" phylum="Mollusca" rank="order">Dentaliida</taxonomicName>
, family
<taxonomicName family="Dentaliidae" lsidName="" pageId="23" pageNumber="24" rank="family">Dentaliidae</taxonomicName>
<taxonomicName id="68C778E49DE9D5D63636027449CD4952" class="Scaphopoda" family="Dentaliidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Dentaliida" pageId="23" pageNumber="24" phylum="Mollusca" rank="family">Dentaliidae</taxonomicName>
(Fig. 24). Genetic data and imagery provided to facilitate future study.
</paragraph>
<caption pageId="23" pageNumber="24">
<paragraph pageId="23" pageNumber="24">
<caption id="1986779EFA7E56313770184E65AB02E7" pageId="23" pageNumber="24">
<paragraph id="5CF7BE7D1A98A34E8A21DD3980D933D9" pageId="23" pageNumber="24">
Figure 24. Phylogenetic analysis of
<taxonomicName class="Scaphopoda" lsidName="Pteriomorphia" pageId="23" pageNumber="24" rank="class">Scaphopoda</taxonomicName>
<taxonomicName id="BD82C6B87E5C5309A230E2D8FB6B3AAC" class="Scaphopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" pageId="23" pageNumber="24" phylum="Mollusca" rank="class">Scaphopoda</taxonomicName>
. 50% majority rule consensus tree from the Bayesian analyses using 18S. Asterisks denotes support values of 95 or above.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="ecology">
<paragraph pageId="23" pageNumber="24">Ecology.</paragraph>
<paragraph pageId="23" pageNumber="24">Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.</paragraph>
<subSubSection id="A7280A59114AD3BE8A93AF013EA59D63" pageId="23" pageNumber="24" type="ecology">
<paragraph id="2DF1216467689C76555A5198C51208E6" pageId="23" pageNumber="24">Ecology.</paragraph>
<paragraph id="2031C7CBBD0D4F42E06E9857F3A4BC63" pageId="23" pageNumber="24">Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.</paragraph>
</subSubSection>
</treatment>
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<paragraph pageId="1" pageNumber="164">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
<treatment id="6A34AF34A04A3F57FDDA869F0514C18D" ID-GBIF-Taxon="127870515" LSID="urn:lsid:plazi:treatment:6A34AF34A04A3F57FDDA869F0514C18D" httpUri="http://treatment.plazi.org/id/6A34AF34A04A3F57FDDA869F0514C18D" lastPageNumber="164" pageId="1" pageNumber="164" scope_family="Ammotheidae" scope_order="Pantopoda" scope_subOrder="Eupantopodida" scope_superFamily="Pycnogonoidea">
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<paragraph id="0F5908EFB06FD2129CF39E37B147EAA7" pageId="1" pageNumber="164">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
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<subSubSection pageId="1" pageNumber="164" type="nomenclature">
<paragraph pageId="1" pageNumber="164">
<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella appendiculata" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="appendiculata">Ammothella appendiculata (Dohrn, 1881)</taxonomicName>
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<paragraph id="A2BC626D63A1AE9468588C421136A481" pageId="1" pageNumber="164">
<taxonomicName id="08AE9BBDDA9A2A83A45987A724CBED09" ID-CoL="CVKX" authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella appendiculata" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="appendiculata">Ammothella appendiculata (Dohrn, 1881)</taxonomicName>
Figures 3, 13, 14
</paragraph>
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<subSubSection pageId="1" pageNumber="164" type="reference_group">
<paragraph pageId="1" pageNumber="164">
<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea appendiculata" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="appendiculata">Ammothea appendiculata</taxonomicName>
<subSubSection id="4D3E51235F3FC9AB3055A57EF02B2F98" pageId="1" pageNumber="164" type="reference_group">
<paragraph id="106A5480269015F0A745BB7DB736E89F" pageId="1" pageNumber="164">
<taxonomicName id="4EE8BFDEC5A85BC313A0D1B9114EB83D" class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea appendiculata" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="appendiculata">Ammothea appendiculata</taxonomicName>
Dohrn, 1881
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="164" type="material">
<paragraph pageId="1" pageNumber="164">Material.</paragraph>
<paragraph pageId="1" pageNumber="164">ZSMA20071470: female; Cape Savudrija, Croatia; 03.09.2004: 0-1 m. ZSMA20071471: female; Cape Savudrija, Croatia; 08.10.2004: 0-1 m; on stone. ZSMA20071472: female; Cape Savudrija, Croatia; 08.08.2005; 0-1 m; under stone.</paragraph>
<subSubSection id="DF39F0D7A2E117B77019B6DEB0A112B6" pageId="1" pageNumber="164" type="material">
<paragraph id="2E616746864680A5241ECFDC37F3408A" pageId="1" pageNumber="164">Material.</paragraph>
<paragraph id="DF7990AD1AB20EE4719E666896D0E188" pageId="1" pageNumber="164">ZSMA20071470: female; Cape Savudrija, Croatia; 03.09.2004: 0-1 m. ZSMA20071471: female; Cape Savudrija, Croatia; 08.10.2004: 0-1 m; on stone. ZSMA20071472: female; Cape Savudrija, Croatia; 08.08.2005; 0-1 m; under stone.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="164" type="remarks">
<paragraph pageId="1" pageNumber="164">Remarks.</paragraph>
<paragraph pageId="1" pageNumber="164">
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<subSubSection id="F72F4D76775A532D8A92B3550A43CC79" pageId="1" pageNumber="164" type="remarks">
<paragraph id="78B53E27A0D04E7351070C8D845080C0" pageId="1" pageNumber="164">Remarks.</paragraph>
<paragraph id="F1D3B2323F41EC0B022F9FAE2F5F1776" pageId="1" pageNumber="164">
<bibRefCitation id="6334169AB14695FD02304501DAA7EF26" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
described the palp with 10 articles; here the palp, as in other
<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">Ammothella</taxonomicName>
<taxonomicName id="1DFBA584A42EB46C81BB4BDF989E0330" class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="genus">Ammothella</taxonomicName>
, has 9 articles (Fig. 13E).
</paragraph>
</subSubSection>

View file

@ -1,133 +1,132 @@
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<taxonomicName authority="Dall, 1881" authorityName="Dall" authorityYear="1881" class="Bivalvia" family="Arcidae" genus="Bentharca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bentharca asperula" order="Pholadomyoida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="asperula">Bentharca cf. asperula (Dall, 1881)</taxonomicName>
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<taxonomicName id="3F39DCF95FEA9A98B647B670EABBB83C" authority="Dall, 1881" authorityName="Dall" authorityYear="1881" family="Arcidae" genus="Bentharca" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Bentharca asperula" order="Pholadomyoida" pageId="13" pageNumber="14" rank="species" species="asperula">Bentharca cf. asperula (Dall, 1881)</taxonomicName>
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<paragraph pageId="13" pageNumber="14">Material examined.</paragraph>
<paragraph pageId="13" pageNumber="14">NHM_108 NHMUK 20170061, collected 2013-10-11, 13.79335 -116.70308, 4081 m. http://data.nhm.ac.uk/object/9d29d7ec-55cd-4b41-929a-2379be221263</paragraph>
<paragraph pageId="13" pageNumber="14">NHM_150 NHMUK 20170062.1-2, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/96bfe548-f511-49c4-b2a3-0a9a45f9154b</paragraph>
<paragraph pageId="13" pageNumber="14">NHM_170 NHMUK 20170063, collected 2013-10-11, 13.7936 -116.70308, 4078 m. http://data.nhm.ac.uk/object/8d9beefd-2fbc-4204-9bf8-90551419ac1c</paragraph>
<paragraph pageId="13" pageNumber="14">NHM_282 NHMUK 20170064, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/ccdd114d-c8a8-47da-ba84-8b8ca5125a6a</paragraph>
<paragraph pageId="13" pageNumber="14">NHM_427 NHMUK 20170065, collected 2013-10-20, 13.86367 -116.54432, 4050 m. http://data.nhm.ac.uk/object/1d462c2a-bb98-4369-afc3-63a7c33a4bdd</paragraph>
<paragraph pageId="14" pageNumber="15">
<pageBreakToken pageId="14" pageNumber="15" start="start">NHM</pageBreakToken>
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<paragraph id="6954BCFF08E11DC06CC588E0D4D6492C" pageId="13" pageNumber="14">Material examined.</paragraph>
<paragraph id="7E2C371AA0CB412EB5A7F6C0D7077FA0" pageId="13" pageNumber="14">NHM_108 NHMUK 20170061, collected 2013-10-11, 13.79335 -116.70308, 4081 m. http://data.nhm.ac.uk/object/9d29d7ec-55cd-4b41-929a-2379be221263</paragraph>
<paragraph id="88C47FD114F56148FEAAA242B6D7DEDD" pageId="13" pageNumber="14">NHM_150 NHMUK 20170062.1-2, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/96bfe548-f511-49c4-b2a3-0a9a45f9154b</paragraph>
<paragraph id="AF7BF53999936F0E861D04C543637AD6" pageId="13" pageNumber="14">NHM_170 NHMUK 20170063, collected 2013-10-11, 13.7936 -116.70308, 4078 m. http://data.nhm.ac.uk/object/8d9beefd-2fbc-4204-9bf8-90551419ac1c</paragraph>
<paragraph id="97A3638FA0C3D4D7BA358D04E947E6D4" pageId="13" pageNumber="14">NHM_282 NHMUK 20170064, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/ccdd114d-c8a8-47da-ba84-8b8ca5125a6a</paragraph>
<paragraph id="16FA1BEB0AB134AD82DE3BF4C4C5A0C1" pageId="13" pageNumber="14">NHM_427 NHMUK 20170065, collected 2013-10-20, 13.86367 -116.54432, 4050 m. http://data.nhm.ac.uk/object/1d462c2a-bb98-4369-afc3-63a7c33a4bdd</paragraph>
<paragraph id="EF36971BC84817581D8D01E728B3DE64" pageId="14" pageNumber="15">
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_454 NHMUK 20170066, collected 2013-10-21, 13.90165 -116.59, 4163 m. http://data.nhm.ac.uk/object/a30eab51-5f52-4fec-89d7-d47152895c92
</paragraph>
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<subSubSection pageId="14" pageNumber="15" type="description">
<paragraph pageId="14" pageNumber="15">Description.</paragraph>
<paragraph pageId="14" pageNumber="15">Shell elongate, trapezoidal, strongly inequilateral, anteriorly attentuated and posteriorly expanded, umbones small, low, dorsal edge straight. Byssal sinus in ventral margin. Sculpture of irregular commarginal lamellae and low radial ribs but covered by a thick, shaggy, brown periostracum with projecting scales. Two pre- and post- umbonal hinge teeth with each tooth crossed by transverse grooves giving a lobate appearance (Fig. 13E, F). Inner shell margin smooth. DNA voucher NHM_150 shell length 3.2 mm shell width 1.9 mm.</paragraph>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
<subSubSection id="F14B158CBB64B00A1EB7FA842287DD25" pageId="14" pageNumber="15" type="description">
<paragraph id="89A280296D1742E148687EA236D121E7" pageId="14" pageNumber="15">Description.</paragraph>
<paragraph id="94AC9E7BA80D683CE8BABB7FC613D86C" pageId="14" pageNumber="15">Shell elongate, trapezoidal, strongly inequilateral, anteriorly attentuated and posteriorly expanded, umbones small, low, dorsal edge straight. Byssal sinus in ventral margin. Sculpture of irregular commarginal lamellae and low radial ribs but covered by a thick, shaggy, brown periostracum with projecting scales. Two pre- and post- umbonal hinge teeth with each tooth crossed by transverse grooves giving a lobate appearance (Fig. 13E, F). Inner shell margin smooth. DNA voucher NHM_150 shell length 3.2 mm shell width 1.9 mm.</paragraph>
<caption id="B153BE3BD02C83F6278D2702A41A600F" pageId="14" pageNumber="15">
<paragraph id="06823F9508EAD9F2B8F1AADEEB52F645" pageId="14" pageNumber="15">
Figure 13.
<taxonomicName class="Bivalvia" family="Arcidae" genus="Bentharca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bentharca asperula" order="Arcida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="asperula">Bentharca cf. asperula</taxonomicName>
<taxonomicName id="4B13D4500B0DB54EE54D92D554EA2C05" class="Bivalvia" family="Arcidae" genus="Bentharca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bentharca asperula" order="Arcida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="asperula">Bentharca cf. asperula</taxonomicName>
(Dall, 1881) A Voucher specimen NHM_150 live after recovery B Specimen NHM_150 after preservation and dissection for DNA sample showing valves C Specimen NHM_108 Live
<normalizedToken originalValue="DF">D-F</normalizedToken>
<normalizedToken id="898C084FFFF15A3612847E99C40B908C" originalValue="DF">D-F</normalizedToken>
Specimen NHM_150 SEM showing shell ornamentation and hinge teeth. Scale bars: 1 mm (A, E); 0.5 mm (B); 0.2 mm (F). Image attribution Glover, Taylor, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="genetic data">
<paragraph pageId="15" pageNumber="16">
<pageBreakToken pageId="15" pageNumber="16" start="start">Genetic</pageBreakToken>
<subSubSection id="E400E54B7E98E08113D4744B921B2DF6" pageId="15" pageNumber="16" type="genetic data">
<paragraph id="6ADCDF4E37F4F6D2A7C267A16E30E786" pageId="15" pageNumber="16">
<pageBreakToken id="1563AF8722F78EA4B6ACF1A96A105B92" pageId="15" pageNumber="16" start="start">Genetic</pageBreakToken>
data.
</paragraph>
<paragraph pageId="15" pageNumber="16">GenBank NHM_108 18S-MF157470, COI-MF157502; NHM_150 18S-MF157476; NHM_170 18S-MF157477; NHM_282 18S-MF157490, COI-MF157513; NHM_427 18S-MF157496; NHM_454 18S-MF157499.</paragraph>
<paragraph id="2D67A34D49E882F6BC229CB02F3DF855" pageId="15" pageNumber="16">GenBank NHM_108 18S-MF157470, COI-MF157502; NHM_150 18S-MF157476; NHM_170 18S-MF157477; NHM_282 18S-MF157490, COI-MF157513; NHM_427 18S-MF157496; NHM_454 18S-MF157499.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="remarks">
<paragraph pageId="15" pageNumber="16">Remarks.</paragraph>
<paragraph pageId="15" pageNumber="16">
<taxonomicName class="Bivalvia" family="Arcidae" genus="Bentharca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bentharca asperula" order="Arcida" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="asperula">Bentharca asperula</taxonomicName>
<subSubSection id="F3672D340AEE5AA192E4D3385997E7C0" pageId="15" pageNumber="16" type="remarks">
<paragraph id="E75DA9BCD8F0494E7668003F7B215318" pageId="15" pageNumber="16">Remarks.</paragraph>
<paragraph id="194EFB6A3C7D9A12C0AE8F70DC4F20CB" pageId="15" pageNumber="16">
<taxonomicName id="FA989F4A1DBB34A9FD558E2A21D1FD5C" class="Bivalvia" family="Arcidae" genus="Bentharca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bentharca asperula" order="Arcida" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="asperula">Bentharca asperula</taxonomicName>
has been regarded as a cosmopolitan deep-water species with a considerable recorded depth range of 430-5005 m (
<bibRefCitation author="Knudsen, J" journalOrPublisher="John Murray Expedition Scientific Reports" pageId="30" pageNumber="31" pagination="237 - 343" title="The deep-sea Bivalvia." volume="11" year="1967">Knudsen 1967</bibRefCitation>
<bibRefCitation id="61DB256AEF8DF6B866375A4FDAF5DC34" author="Knudsen, J" journalOrPublisher="John Murray Expedition Scientific Reports" pageId="30" pageNumber="31" pagination="237 - 343" title="The deep-sea Bivalvia." volume="11" year="1967">Knudsen 1967</bibRefCitation>
,
<bibRefCitation author="Knudsen, J" journalOrPublisher="Galathea Reports" pageId="30" pageNumber="31" pagination="1 - 241" title="The systematics and biology of abyssal and hadal Bivalvia." volume="11" year="1970">1970</bibRefCitation>
<bibRefCitation id="8694FDC358BA229137ACCA47A94A67AC" author="Knudsen, J" journalOrPublisher="Galathea Reports" pageId="30" pageNumber="31" pagination="1 - 241" title="The systematics and biology of abyssal and hadal Bivalvia." volume="11" year="1970">1970</bibRefCitation>
,
<bibRefCitation author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott 2012</bibRefCitation>
<bibRefCitation id="27C21DAE62596B29584FE7DB16181E77" author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott 2012</bibRefCitation>
) from Atlantic, Indian and Pacific Oceans. The lectotype and paralectotypes (USNM 63174, 887339, 94363) originated from the Gulf of Mexico, off Yucatan, 2868 m (Blake stn 33). Because of its epifaunal, byssate life habit
<taxonomicName lsidName="B. asperula" pageId="15" pageNumber="16" rank="species" species="asperula">B. asperula</taxonomicName>
<taxonomicName id="E390E372A93E442A0304F0D9EC335B2A" lsidName="B. asperula" pageId="15" pageNumber="16" rank="species" species="asperula">B. asperula</taxonomicName>
shows considerable shape variation and
<bibRefCitation author="Knudsen, J" journalOrPublisher="John Murray Expedition Scientific Reports" pageId="30" pageNumber="31" pagination="237 - 343" title="The deep-sea Bivalvia." volume="11" year="1967">Knudsen (1967)</bibRefCitation>
<bibRefCitation id="E79CF2A16E1AF581354A584E2E26FFB8" author="Knudsen, J" journalOrPublisher="John Murray Expedition Scientific Reports" pageId="30" pageNumber="31" pagination="237 - 343" title="The deep-sea Bivalvia." volume="11" year="1967">Knudsen (1967)</bibRefCitation>
synonymised several nominal species and described how the number of hinge teeth increases with shell size (age). Without supporting genetic evidence from samples from different oceans it is impossible to test whether the species is truly cosmopolitan. Perhaps significantly, no shell has been described with as few hinge teeth as the present sample and none with the transverse grooves (Fig. 13F). No genetic matches on GenBank.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="ecology">
<paragraph pageId="15" pageNumber="16">Ecology.</paragraph>
<paragraph pageId="15" pageNumber="16">Quite abundant. Found in polymetallic nodule province.</paragraph>
<subSubSection id="EBEED772A768D500EA47E56990AB6E47" pageId="15" pageNumber="16" type="ecology">
<paragraph id="53EE81A9A530A896D72E2BFC5258DDC3" pageId="15" pageNumber="16">Ecology.</paragraph>
<paragraph id="2C1E8DCC1F09EF04B4FFB9CFE4013904" pageId="15" pageNumber="16">Quite abundant. Found in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,145 +1,144 @@
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<taxonomicName authority="Smith, 1885" authorityName="Smith" authorityYear="1885" class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Pholadomyoida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum Smith, 1885</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="materials_examined">
<paragraph pageId="9" pageNumber="10">Material examined.</paragraph>
<paragraph pageId="9" pageNumber="10">NHM_141 NHMUK 20170051, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/f2133256-1cad-4255-a5cb-bd5331417127</paragraph>
<paragraph pageId="9" pageNumber="10">NHM_274A NHMUK 20170052, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/f96a470e-237e-46b4-ba85-4c6196106071</paragraph>
<paragraph pageId="9" pageNumber="10">NHM_378 NHMUK 20170053.1-2, collected 2013-10-19, 13.93307 -116.71628, 4182 m. http://data.nhm.ac.uk/object/65f8d1ed-dd6a-4265-90d2-daf07491cd76</paragraph>
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<paragraph id="2E4155D58D5F711D0A1CD0F66501AD13" pageId="9" pageNumber="10">Material examined.</paragraph>
<paragraph id="B8AA674F52BFADD82039192BCFADF7A2" pageId="9" pageNumber="10">NHM_141 NHMUK 20170051, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/f2133256-1cad-4255-a5cb-bd5331417127</paragraph>
<paragraph id="07E45D07D786DFFEA0742378440140EE" pageId="9" pageNumber="10">NHM_274A NHMUK 20170052, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/f96a470e-237e-46b4-ba85-4c6196106071</paragraph>
<paragraph id="049C9062FA411DCE6A8485DC5504281C" pageId="9" pageNumber="10">NHM_378 NHMUK 20170053.1-2, collected 2013-10-19, 13.93307 -116.71628, 4182 m. http://data.nhm.ac.uk/object/65f8d1ed-dd6a-4265-90d2-daf07491cd76</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="description">
<paragraph pageId="9" pageNumber="10">Description.</paragraph>
<paragraph pageId="9" pageNumber="10">Small, trigonal- subovate. Periostracum light brown, shiny. Sculpture of fine radial lirae. Resilifer small. Hinge teeth: 5 anterior, 4 posterior. Inner shell margin finely denticulate. Voucher NHM_274A width 2 mm, height 1.8 mm (Fig. 9).</paragraph>
<caption pageId="9" pageNumber="10">
<paragraph pageId="9" pageNumber="10">
<subSubSection id="2882D8A53A245939D91ADCA0DEC4B533" pageId="9" pageNumber="10" type="description">
<paragraph id="206E73FB3CB0212DD8BAF2092505B9EE" pageId="9" pageNumber="10">Description.</paragraph>
<paragraph id="838069D5182FCB407150A40F1998DE0E" pageId="9" pageNumber="10">Small, trigonal- subovate. Periostracum light brown, shiny. Sculpture of fine radial lirae. Resilifer small. Hinge teeth: 5 anterior, 4 posterior. Inner shell margin finely denticulate. Voucher NHM_274A width 2 mm, height 1.8 mm (Fig. 9).</paragraph>
<caption id="1DCA91766BBCBB9C8B636C6EE779A4BF" pageId="9" pageNumber="10">
<paragraph id="8ACC771685661A6EC1F80513DC853B22" pageId="9" pageNumber="10">
Figure 9.
<taxonomicName class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Nuculida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum</taxonomicName>
<taxonomicName id="F7264AD6CDAAE6D09C7B3E224F58916F" class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Nuculida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum</taxonomicName>
Smith, 1885 A Live specimen NHM_141 (for which 18S, CO1 and 16S sequences were obtained) B Live specimens NHM_274 (4 specimens from same sample) C Open shell from single individual NHM_274A with tissue sample taken for DNA sequencing
<normalizedToken originalValue="DE">D-E</normalizedToken>
<normalizedToken id="872D00E2F9D475C56CCF4212B9A423E0" originalValue="DE">D-E</normalizedToken>
SEM of NHM_378 valve showing hinge teeth. Scale bars: 1.5 mm (B); 0.5 mm (C). Image attribution Glover, Taylor, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="genetic data">
<paragraph pageId="9" pageNumber="10">Genetic data.</paragraph>
<paragraph pageId="9" pageNumber="10">GenBank NHM_141 16S-MF157457, 18S-MF157473, COI-MF157503; NHM_274A COI-MF157512; NHM_378 16S-MF157464, COI-MF157520.</paragraph>
<subSubSection id="5F080B541D8654552B63B17965FB9C49" pageId="9" pageNumber="10" type="genetic data">
<paragraph id="8D9933B6BD43AFD975952A04CFF79E15" pageId="9" pageNumber="10">Genetic data.</paragraph>
<paragraph id="867D87B24835C8885A427D8B181BD70D" pageId="9" pageNumber="10">GenBank NHM_141 16S-MF157457, 18S-MF157473, COI-MF157503; NHM_274A COI-MF157512; NHM_378 16S-MF157464, COI-MF157520.</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="remarks">
<paragraph pageId="10" pageNumber="11">
<pageBreakToken pageId="10" pageNumber="11" start="start">Remarks</pageBreakToken>
<subSubSection id="507BD47855D53EDF783657915FF33FCC" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="remarks">
<paragraph id="C4711EAA64FBD8DD236ECCD515665BF1" pageId="10" pageNumber="11">
<pageBreakToken id="411B20673B7AF60FB526847130B9B865" pageId="10" pageNumber="11" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<paragraph id="1D9FADC4ACC86C4D2F20AFE83616C2BF" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Morphologically matches
<taxonomicName class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Nuculida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum</taxonomicName>
<taxonomicName id="13DD60A590A03E0E8B355309E3948AD4" class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Nuculida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum</taxonomicName>
Smith, 1885 based on examination of the syntype specimens [NHMUK 1887.2.9.2919]. In the molecular analysis of nuculoid protobranchs (Fig. 12)
<taxonomicName class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Nuculida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum</taxonomicName>
<taxonomicName id="B248C008993C75EC5A718A75D7E595B4" class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Nuculida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum</taxonomicName>
and the Atlantic
<taxonomicName class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula atacellana" order="Nuculida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="atacellana">Nucula atacellana</taxonomicName>
<taxonomicName id="15CA82CFA63BCAFF513C209216EF869B" class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula atacellana" order="Nuculida" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="atacellana">Nucula atacellana</taxonomicName>
Schenck, 1939 are well supported sister species. However the
<taxonomicName lsidName="N. profundorum" pageId="10" pageNumber="11" rank="species" species="profundorum">N. profundorum</taxonomicName>
<taxonomicName id="13ED104E48AF5106B5A90F510FC5C247" lsidName="N. profundorum" pageId="10" pageNumber="11" rank="species" species="profundorum">N. profundorum</taxonomicName>
identified from the present samples differs genetically from the
<taxonomicName lsidName="N. profundorum" pageId="10" pageNumber="11" rank="species" species="profundorum">N. profundorum</taxonomicName>
<taxonomicName id="348FF4ACD7EE342ABAD1602B4918BA56" lsidName="N. profundorum" pageId="10" pageNumber="11" rank="species" species="profundorum">N. profundorum</taxonomicName>
record in GenBank (accession nr KJ950274;
<bibRefCitation author="Jennings, RM" journalOrPublisher="Polish Polar Research" pageId="30" pageNumber="31" pagination="261 - 278" title="Phylogeographic estimates of colonization of the deep Atlantic by the protobranch bivalve Nuculaatacellana." url="https://doi.org/10.2478/popore-2014-0017" volume="35" year="2014">Jennings and Etter 2014</bibRefCitation>
<bibRefCitation id="2AF4F129B2979D2F2E855079BB4350EB" author="Jennings, RM" journalOrPublisher="Polish Polar Research" pageId="30" pageNumber="31" pagination="261 - 278" title="Phylogeographic estimates of colonization of the deep Atlantic by the protobranch bivalve Nuculaatacellana." url="https://doi.org/10.2478/popore-2014-0017" volume="35" year="2014">Jennings and Etter 2014</bibRefCitation>
) which we believe may be misassigned. That sample came from 1045 m in the north eastern
<pageBreakToken pageId="11" pageNumber="12" start="start">Pacific</pageBreakToken>
<pageBreakToken id="3945AFC72D46FAA8EEED0EFD47878693" pageId="11" pageNumber="12" start="start">Pacific</pageBreakToken>
off San Diego (Figure 10). The shell illustrated by
<bibRefCitation author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott (2012</bibRefCitation>
<bibRefCitation id="59E53937BE39370E91F8AB9B7FDD7096" author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott (2012</bibRefCitation>
pl 12) as
<taxonomicName lsidName="N. profundorum" pageId="11" pageNumber="12" rank="species" species="profundorum">N. profundorum</taxonomicName>
<taxonomicName id="A7ED0424C6FDC5B200A122157A42E8E5" lsidName="N. profundorum" pageId="11" pageNumber="12" rank="species" species="profundorum">N. profundorum</taxonomicName>
has more hinge teeth. There may be a complex of morphologically similar species in the eastern Pacific. No genetic matches on GenBank.
</paragraph>
<caption pageId="11" pageNumber="12">
<paragraph pageId="11" pageNumber="12">
<caption id="F82F93E005F8176277062336B27B5D8A" pageId="11" pageNumber="12">
<paragraph id="03124256E4FBEF2A3B91D0955A8DCCA9" pageId="11" pageNumber="12">
Figure 10.
<taxonomicName class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Nuculida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum</taxonomicName>
<taxonomicName id="8D0051AFF3826431FB8E063D5F7EA152" class="Bivalvia" family="Nuculidae" genus="Nucula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nucula profundorum" order="Nuculida" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="profundorum">Nucula profundorum</taxonomicName>
Smith, 1885.
<normalizedToken originalValue="AD">A-D</normalizedToken>
<normalizedToken id="F2008B5426C6550E0EDB37E142264344" originalValue="AD">A-D</normalizedToken>
Syntype BMNH 1887.2.9.2919, scalebars 1mm E Type locality (red) of
<taxonomicName lsidName="N. profundorum" pageId="11" pageNumber="12" rank="species" species="profundorum">N. profundorum</taxonomicName>
<taxonomicName id="0081D4A310E5BAF08294619803204144" lsidName="N. profundorum" pageId="11" pageNumber="12" rank="species" species="profundorum">N. profundorum</taxonomicName>
from Challenger Expedition in relation to ABYSSLINE sampling location (green) and GenBank voucher specimen sampling location (white). Bathymetric data (D) from NOAA.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="ecology">
<paragraph pageId="11" pageNumber="12">Ecology.</paragraph>
<paragraph pageId="11" pageNumber="12">The most abundant bivalve mollusc recorded in the ABYSSLINE sampling programme, frequently found in epibenthic sledge and box core samples from region of sediment and polymetallic nodules.</paragraph>
<subSubSection id="80C0609AFF337A025BE0E1FF1E6623C3" pageId="11" pageNumber="12" type="ecology">
<paragraph id="BAD0E0E78A1BC5FC8A4D8835D775B6AC" pageId="11" pageNumber="12">Ecology.</paragraph>
<paragraph id="2215DDDFF35320949C122090C633E192" pageId="11" pageNumber="12">The most abundant bivalve mollusc recorded in the ABYSSLINE sampling programme, frequently found in epibenthic sledge and box core samples from region of sediment and polymetallic nodules.</paragraph>
</subSubSection>
</treatment>
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<paragraph pageId="3" pageNumber="166">Taxon classification Animalia Pantopoda Pycnogonidae</paragraph>
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<paragraph id="69A8BD0845EE9BE10B46E6E43631FCE1" pageId="3" pageNumber="166">Taxon classification Animalia Pantopoda Pycnogonidae</paragraph>
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<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnogonum nodulosum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="nodulosum">Pycnogonum nodulosum Dohrn, 1881</taxonomicName>
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<taxonomicName id="2078BA1EA75D9456E039DFA9A7F92F71" ID-CoL="4QMGP" authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Pycnogonidae" genus="Pycnogonum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnogonum nodulosum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="nodulosum">Pycnogonum nodulosum Dohrn, 1881</taxonomicName>
Figures 53, 54
</paragraph>
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<paragraph pageId="3" pageNumber="166">Material.</paragraph>
<paragraph pageId="3" pageNumber="166">ZSMA20071503: female; Banyuls-sur-mer, France; 07.2004; 30 m; red algae. ZSMA20030112: female; Tavolara Island, Sardinia, Italy; 2002.</paragraph>
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<paragraph id="6E87AAD3A6E677064CA86922F338A976" pageId="3" pageNumber="166">ZSMA20071503: female; Banyuls-sur-mer, France; 07.2004; 30 m; red algae. ZSMA20030112: female; Tavolara Island, Sardinia, Italy; 2002.</paragraph>
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<taxonomicName authority="sp. (NHM _ 190)" class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella" order="Pholadomyoida" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Yoldiella sp. (NHM_190)</taxonomicName>
<treatment id="7AAA55FEF36713D6978687AA33415C44" ID-GBIF-Taxon="135489990" LSID="urn:lsid:plazi:treatment:7AAA55FEF36713D6978687AA33415C44" httpUri="http://treatment.plazi.org/id/7AAA55FEF36713D6978687AA33415C44" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" scope_class="Bivalvia" scope_family="Yoldiidae" scope_order="Nuculanoida" scope_phylum="Mollusca" scope_subClass="Protobranchia">
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<taxonomicName id="ED9A24C8F0CE7A019BFE7150E51B6193" authority="sp. (NHM _ 190)" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Yoldiella" order="Pholadomyoida" pageId="11" pageNumber="12" rank="genus">Yoldiella sp. (NHM_190)</taxonomicName>
</paragraph>
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<subSubSection lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="materials_examined">
<paragraph pageId="11" pageNumber="12">Material examined.</paragraph>
<paragraph pageId="11" pageNumber="12">NHM_042 NHMUK 20170054, collected 2013-10-09, 13.8372 -116.55843, 4336 m. http://data.nhm.ac.uk/object/621deeed-8f8a-4d2e-9136-4e30794fc68e</paragraph>
<paragraph pageId="12" pageNumber="13">
<pageBreakToken pageId="12" pageNumber="13" start="start">NHM</pageBreakToken>
<subSubSection id="C1A92132646F273664427F7D9880F924" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="963C995266667D0015C51DDF4BB5D2C5" pageId="11" pageNumber="12">Material examined.</paragraph>
<paragraph id="90BDE97332E13CDA0B63D36B5CCBE1E4" pageId="11" pageNumber="12">NHM_042 NHMUK 20170054, collected 2013-10-09, 13.8372 -116.55843, 4336 m. http://data.nhm.ac.uk/object/621deeed-8f8a-4d2e-9136-4e30794fc68e</paragraph>
<paragraph id="1B49DA121F07647EFB74F8CC5A42F91D" pageId="12" pageNumber="13">
<pageBreakToken id="6CE6394AEF3029210EE8EFC6BE48AE52" pageId="12" pageNumber="13" start="start">NHM</pageBreakToken>
_185 NHMUK 20170055, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/6dfa8946-aa7a-448d-9f4f-703a3b2a10d9
</paragraph>
<paragraph pageId="12" pageNumber="13">NHM_190 NHMUK 20170056, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/b6e48ff4-2e02-42dc-b9ed-286d297d1459</paragraph>
<paragraph pageId="12" pageNumber="13">NHM_193 NHMUK 20170060, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/8923576e-4542-4fc7-9a89-016e8fb564cb</paragraph>
<paragraph pageId="12" pageNumber="13">NHM_194 NHMUK 20170057, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/7a6c76df-989b-4fcd-9e9c-a442d0a02443</paragraph>
<paragraph pageId="13" pageNumber="14">
<pageBreakToken pageId="13" pageNumber="14" start="start">NHM</pageBreakToken>
<paragraph id="E0E15BB0E49C911117A048DCF3AD2BD1" pageId="12" pageNumber="13">NHM_190 NHMUK 20170056, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/b6e48ff4-2e02-42dc-b9ed-286d297d1459</paragraph>
<paragraph id="04281175B34BF72149177F80A2267C5E" pageId="12" pageNumber="13">NHM_193 NHMUK 20170060, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/8923576e-4542-4fc7-9a89-016e8fb564cb</paragraph>
<paragraph id="08424F3DD484E185B71E77A1D998696A" pageId="12" pageNumber="13">NHM_194 NHMUK 20170057, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/7a6c76df-989b-4fcd-9e9c-a442d0a02443</paragraph>
<paragraph id="0252D86127AD57E79A270F0905582A83" pageId="13" pageNumber="14">
<pageBreakToken id="F895EA420AFC7E75DA86885FB68A5B08" pageId="13" pageNumber="14" start="start">NHM</pageBreakToken>
_246 NHMUK 20170058, collected 2013-10-16, 13.81166 -116.71, 4076 m. http://data.nhm.ac.uk/object/37b2493a-a725-4ec4-a720-cc9dd12fb49d
</paragraph>
<paragraph pageId="13" pageNumber="14">NHM_289 NHMUK 20170059, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/17d54bb4-9f38-4073-9bb6-17637773b058</paragraph>
<paragraph id="3843D57EA3D2EFE001C761209468F24C" pageId="13" pageNumber="14">NHM_289 NHMUK 20170059, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/17d54bb4-9f38-4073-9bb6-17637773b058</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="description">
<paragraph pageId="13" pageNumber="14">Description.</paragraph>
<paragraph pageId="13" pageNumber="14">Small, sub-ovate, longer than high, umbone at mid-line, dorsal margin horizontal to slightly curved, ventral margin deeply rounded, thin-shelled, shiny, semi-transparent, smooth except for growth increments. Internal features not investigated but 4-5 anterior and posterior chevron teeth. Hindgut visible though the shell forms a simple rounded loop on right side of body. DNA voucher NHM_190 shell length 1.6 mm, height 1 mm. Voucher specimen NHM_185 shell length 1.5 mm, height 1 mm (Fig. 11).</paragraph>
<caption pageId="13" pageNumber="14">
<paragraph pageId="13" pageNumber="14">
<subSubSection id="9CE97BB311660EA17057168CA9331D52" pageId="13" pageNumber="14" type="description">
<paragraph id="7278D4A4ACF9E29CD5C444157340BDB2" pageId="13" pageNumber="14">Description.</paragraph>
<paragraph id="1C0CCC9B736C92DAA6B39A7546AE24A4" pageId="13" pageNumber="14">Small, sub-ovate, longer than high, umbone at mid-line, dorsal margin horizontal to slightly curved, ventral margin deeply rounded, thin-shelled, shiny, semi-transparent, smooth except for growth increments. Internal features not investigated but 4-5 anterior and posterior chevron teeth. Hindgut visible though the shell forms a simple rounded loop on right side of body. DNA voucher NHM_190 shell length 1.6 mm, height 1 mm. Voucher specimen NHM_185 shell length 1.5 mm, height 1 mm (Fig. 11).</paragraph>
<caption id="F68B59D54C60C1F35DFF44D20B7DC6C6" pageId="13" pageNumber="14">
<paragraph id="E83BA0012F336C08008B640008B1E8F3" pageId="13" pageNumber="14">
Figure 11.
<taxonomicName class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella" order="Nuculanida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">Yoldiella</taxonomicName>
<taxonomicName id="F5207CB1D4B3F6B1C8E3B8F89F6E5D5E" class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella" order="Nuculanida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">Yoldiella</taxonomicName>
sp. (NHM_190) A Voucher specimen NHM_190 B Live specimens NHM_185 C NHM_185 after preservation in ethanol for 3 months prior to DNA sequencing. Scale bar: 0.5 mm (C). Image attribution Glover, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
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<subSubSection pageId="13" pageNumber="14" type="genetic data">
<paragraph pageId="13" pageNumber="14">Genetic data.</paragraph>
<paragraph pageId="13" pageNumber="14">GenBank NHM_042 18S-MF157467; NHM_185 18S-MF157480; NHM_190 18S-MF157482; NHM_193 18S-MF157484; NHM_194 18S-MF157485; NHM_246 18S-MF157486; NHM_289 18S-MF157492, COI-MF157517.</paragraph>
<subSubSection id="C9E57361509B0A6380FDFB21FEE70220" pageId="13" pageNumber="14" type="genetic data">
<paragraph id="1E91CD022B8AB01C127573DC7BAA598D" pageId="13" pageNumber="14">Genetic data.</paragraph>
<paragraph id="2320B4F3D1BFAF1A263DA9AAF3D0DE9A" pageId="13" pageNumber="14">GenBank NHM_042 18S-MF157467; NHM_185 18S-MF157480; NHM_190 18S-MF157482; NHM_193 18S-MF157484; NHM_194 18S-MF157485; NHM_246 18S-MF157486; NHM_289 18S-MF157492, COI-MF157517.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="remarks">
<paragraph pageId="13" pageNumber="14">Remarks.</paragraph>
<paragraph pageId="13" pageNumber="14">
<subSubSection id="83A6F8EE5F929E6460D8F9B3B9A9EFC7" pageId="13" pageNumber="14" type="remarks">
<paragraph id="433D9237F971C46AAFC7B9DA6F10A889" pageId="13" pageNumber="14">Remarks.</paragraph>
<paragraph id="CEC79DBD05B0126763048A7BB025FD61" pageId="13" pageNumber="14">
Extremely small, semi-transparent bivalves typically about 1 mm in size.
<taxonomicName class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella" order="Nuculanida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">Yoldiella</taxonomicName>
<taxonomicName id="7365E7A8C263056EDEC6C3FF16824E18" class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella" order="Nuculanida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">Yoldiella</taxonomicName>
species are particularly difficult to identify (see
<bibRefCitation author="Killeen, IJ" journalOrPublisher="Journal of Conchology" pageId="30" pageNumber="31" pagination="733 - 778" title="Yoldiella and Portlandia (Bivalvia) from the Faroe-Shetland Channel and Rockall Trough, Northeast Atlantic." volume="39" year="2009">Killeen and Turner 2009</bibRefCitation>
<bibRefCitation id="270D684C0B57DFCB823F0890B3E7EB23" author="Killeen, IJ" journalOrPublisher="Journal of Conchology" pageId="30" pageNumber="31" pagination="733 - 778" title="Yoldiella and Portlandia (Bivalvia) from the Faroe-Shetland Channel and Rockall Trough, Northeast Atlantic." volume="39" year="2009">Killeen and Turner 2009</bibRefCitation>
). Forms a unique monophyletic clade distinct from all other AB01 specimens. No genetic matches on GenBank. In the molecular tree (Fig. 12) the genus
<taxonomicName class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella" order="Nuculanida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">Yoldiella</taxonomicName>
<taxonomicName id="04D4EEAE0B4688C81F0B91B2F9D0BD8E" class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella" order="Nuculanida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">Yoldiella</taxonomicName>
is not monophyletic, and the present species does not group with another Eastern Pacific bathyal species,
<taxonomicName class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella orcia" order="Nuculanida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="orcia">Yoldiella orcia</taxonomicName>
<taxonomicName id="63CB32E958F453C223BA5174E26A9F90" class="Bivalvia" family="Yoldiidae" genus="Yoldiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yoldiella orcia" order="Nuculanida" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="orcia">Yoldiella orcia</taxonomicName>
(Dall, 1916), which instead forms a well-supported subclade with two Atlantic species.
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Figure 12. Phylogenetic analysis of
<taxonomicName class="Bivalvia" lsidName="Protobranchia" pageId="13" pageNumber="14" rank="class">Bivalvia</taxonomicName>
<taxonomicName id="C22401836F9CBC8E17DC3DC4893810F7" class="Bivalvia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protobranchia" pageId="13" pageNumber="14" phylum="Mollusca" rank="class">Bivalvia</taxonomicName>
:
<taxonomicName lsidName="Protobranchia" pageId="13" pageNumber="14" rank="subClass" subClass="Protobranchia">Protobranchia</taxonomicName>
<taxonomicName id="6B7EF4185EEF636B7111E0A6C31D628E" lsidName="Protobranchia" pageId="13" pageNumber="14" rank="subClass" subClass="Protobranchia">Protobranchia</taxonomicName>
. 50% majority rule consensus tree from the Bayesian analyses using 18S and COI. Asterisks denotes support values of 95 or above.
</paragraph>
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<paragraph pageId="13" pageNumber="14">Ecology.</paragraph>
<paragraph pageId="13" pageNumber="14">Found in polymetallic nodule province.</paragraph>
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<paragraph id="B54FA67893954B7BED8158A54A8CA3A5" pageId="13" pageNumber="14">Ecology.</paragraph>
<paragraph id="150AC0FE20E00D26C25D6653E1F80069" pageId="13" pageNumber="14">Found in polymetallic nodule province.</paragraph>
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<paragraph pageId="1" pageNumber="164">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
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<taxonomicName authority="Hodge, 1864" authorityName="Hodge" authorityYear="1864" class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia echinata" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="echinata">Achelia echinata Hodge, 1864</taxonomicName>
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Figures 2, 5, 6, 7, 8
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<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea brevipes" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="brevipes">Ammothea brevipes</taxonomicName>
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Hodge, 1864
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Hodge, 1864
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<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea fibulifera" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="fibulifera">Ammothea fibulifera</taxonomicName>
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<taxonomicName id="139C1085037FF92E297D48B618FDBEF9" class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea fibulifera" order="Pantopoda" pageId="1" pageNumber="164" phylum="Arthropoda" rank="species" species="fibulifera">Ammothea fibulifera</taxonomicName>
(Dohrn, 1881)
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<paragraph pageId="1" pageNumber="164">Material.</paragraph>
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<paragraph id="7E6739354D0AEB69B11AF4C2E0FB4049" pageId="1" pageNumber="164">Material.</paragraph>
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ZSMA20071461: male; Giglio Island, Italy; 08.04.2005; 4-8 m; brown algae. ZSMA20071462: female; Giglio Island, Italy; 11.04.2005; 20 m. ZSMA20071463: female; Rovinj, Croatia; 05.07.2005; 0-1.5 m;
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, 1843.
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<paragraph pageId="2" pageNumber="165">Taxon classification Animalia Pantopoda Callipallenidae</paragraph>
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<paragraph id="32F4B1E9CBA9D53C5D81BC30160BDCEA" pageId="2" pageNumber="165">Taxon classification Animalia Pantopoda Callipallenidae</paragraph>
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<paragraph pageId="2" pageNumber="165">
<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene emaciata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="emaciata">Callipallene emaciata (Dohrn, 1881)</taxonomicName>
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<paragraph id="ABCA98203EB52E3F06AE693976E99904" pageId="2" pageNumber="165">
<taxonomicName id="4D32D115E56C7F03A89F483DE69BB7E0" ID-CoL="PS8X" authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene emaciata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="emaciata">Callipallene emaciata (Dohrn, 1881)</taxonomicName>
Figures 26, 28, 29
</paragraph>
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<subSubSection pageId="2" pageNumber="165" type="reference_group">
<paragraph pageId="2" pageNumber="165">
<taxonomicName class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene emaciata subsp. emaciata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="subspecies" species="emaciata" subspecies="emaciata">Callipallene emaciata emaciata</taxonomicName>
<subSubSection id="AE658ECE6ED7E3B49E292F08876BB3DA" pageId="2" pageNumber="165" type="reference_group">
<paragraph id="B0086C8907BE367E964638E677646142" pageId="2" pageNumber="165">
<taxonomicName id="164AECF6BA7A4065EB552CAF07FC0AE8" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene emaciata subsp. emaciata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="subspecies" species="emaciata" subspecies="emaciata">Callipallene emaciata emaciata</taxonomicName>
Stock, 1954
</paragraph>
<paragraph pageId="2" pageNumber="165">
<taxonomicName class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene emacinata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="emacinata">Callipallene emacinata</taxonomicName>
<paragraph id="F05901A073D454A1639CCE070EB459C1" pageId="2" pageNumber="165">
<taxonomicName id="3564366670881F63654E1F0CF7CFC5C7" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene emacinata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="emacinata">Callipallene emacinata</taxonomicName>
(Dohrn, 1881)
</paragraph>
<paragraph pageId="2" pageNumber="165">
<taxonomicName authority="Johnston, 1837" authorityName="Johnston" authorityYear="1837" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pallene emaciata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="emaciata">Pallene emaciata</taxonomicName>
<paragraph id="DD8FF286D5A694E6C3F30EEF80F05958" pageId="2" pageNumber="165">
<taxonomicName id="8140F69001FC9274F35402DFEF408A61" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pallene emaciata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="emaciata">Pallene emaciata</taxonomicName>
Dohrn, 1881
</paragraph>
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<subSubSection pageId="2" pageNumber="165" type="material">
<paragraph pageId="2" pageNumber="165">Material.</paragraph>
<paragraph pageId="2" pageNumber="165">
<subSubSection id="0F950BAB4EE5E0729FA7B538B80AFEAA" pageId="2" pageNumber="165" type="material">
<paragraph id="AD956407B6F78277A24F088B6CFED31E" pageId="2" pageNumber="165">Material.</paragraph>
<paragraph id="32226C5D0C21FFB1BC44CCDFE6190F3F" pageId="2" pageNumber="165">
ZSMA20071485: female; Giglio Island, Italy; 09.04.2005; 3-5 m;
<taxonomicName class="Phaeophyceae" family="Stypocaulaceae" genus="Halopteris" higherTaxonomySource="CoL" kingdom="Chromista" lsidName="Halopteris" order="Sphacelariales" pageId="2" pageNumber="165" phylum="Ochrophyta" rank="genus">Halopteris</taxonomicName>
<taxonomicName id="62A97E62DE8639D15CE673E2DE29466E" class="Hydrozoa" family="Halopterididae" genus="Halopteris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halopteris" order="Leptothecata" pageId="2" pageNumber="165" phylum="Cnidaria" rank="genus">Halopteris</taxonomicName>
sp.. ZSMA20071486: female; Giglio Island, Italy; 09.04.2005; 3-5 m;
<taxonomicName class="Phaeophyceae" family="Stypocaulaceae" genus="Halopteris" higherTaxonomySource="CoL" kingdom="Chromista" lsidName="Halopteris" order="Sphacelariales" pageId="2" pageNumber="165" phylum="Ochrophyta" rank="genus">Halopteris</taxonomicName>
<taxonomicName id="5E0DA4DF2C440518B78BD17BFDD21B95" class="Hydrozoa" family="Halopterididae" genus="Halopteris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halopteris" order="Leptothecata" pageId="2" pageNumber="165" phylum="Cnidaria" rank="genus">Halopteris</taxonomicName>
sp.. ZSMA20071487: male; Giglio Island, Italy; 09.04.2005; 3-5 m;
<taxonomicName class="Phaeophyceae" family="Stypocaulaceae" genus="Halopteris" higherTaxonomySource="CoL" kingdom="Chromista" lsidName="Halopteris" order="Sphacelariales" pageId="2" pageNumber="165" phylum="Ochrophyta" rank="genus">Halopteris</taxonomicName>
<taxonomicName id="1EA8196D8F128E73D9D0A01955857C35" class="Hydrozoa" family="Halopterididae" genus="Halopteris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halopteris" order="Leptothecata" pageId="2" pageNumber="165" phylum="Cnidaria" rank="genus">Halopteris</taxonomicName>
sp..
</paragraph>
</subSubSection>

View file

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<taxonomicName authority="sp. (NHM _ 381)" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Pholadomyoida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Ledella sp. (NHM_381)</taxonomicName>
<treatment id="8793970CF56119E8C21B32F1FC7E6A47" ID-GBIF-Taxon="135489987" LSID="urn:lsid:plazi:treatment:8793970CF56119E8C21B32F1FC7E6A47" httpUri="http://treatment.plazi.org/id/8793970CF56119E8C21B32F1FC7E6A47" lastPageNumber="9" pageId="8" pageNumber="9" scope_class="Bivalvia" scope_order="Nuculanoida" scope_phylum="Mollusca" scope_subClass="Protobranchia">
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<taxonomicName id="5CD984D6DE9A152668507E4A3B8D8D5A" authority="sp. (NHM _ 381)" family="Nuculanidae" genus="Ledella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ledella" order="Pholadomyoida" pageId="8" pageNumber="9" rank="genus">Ledella sp. (NHM_381)</taxonomicName>
</paragraph>
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<subSubSection pageId="8" pageNumber="9" type="materials_examined">
<paragraph pageId="8" pageNumber="9">Material examined.</paragraph>
<paragraph pageId="8" pageNumber="9">NHM_144 NHMUK 20170050, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/08d5c39f-b1e4-43d7-a8ea-2fe9abc05752</paragraph>
<paragraph pageId="8" pageNumber="9">NHM_381 NHMUK 20170049, collected 2013-10-19, 13.93307 -116.71628, 4182 m. http://data.nhm.ac.uk/object/8f077dac-baac-4fef-b6a1-7fd02d5f0070</paragraph>
<subSubSection id="A6EB410B57BB7DE9DA00607E24C9DDCC" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="36B9791C561DF7FBA9CB9769C92BA722" pageId="8" pageNumber="9">Material examined.</paragraph>
<paragraph id="38788726E0E10E826AA3B2B22E169325" pageId="8" pageNumber="9">NHM_144 NHMUK 20170050, collected 2013-10-11, 13.75833 -116.69852, 4080 m. http://data.nhm.ac.uk/object/08d5c39f-b1e4-43d7-a8ea-2fe9abc05752</paragraph>
<paragraph id="BE5AE10A24281EEDADD702040FDC5BE4" pageId="8" pageNumber="9">NHM_381 NHMUK 20170049, collected 2013-10-19, 13.93307 -116.71628, 4182 m. http://data.nhm.ac.uk/object/8f077dac-baac-4fef-b6a1-7fd02d5f0070</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="description">
<paragraph pageId="8" pageNumber="9">Description.</paragraph>
<paragraph pageId="8" pageNumber="9">Ovoid with short rostrum, shell shiny sub-translucent. Sculpture of fine closely spaced commarginal lamellae. Specimen NHM_381 length 2 mm (Fig. 8).</paragraph>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
<subSubSection id="F0DEB81653CA085DBFA76C2784603633" pageId="8" pageNumber="9" type="description">
<paragraph id="A0CA7C720C92867DF09BD250DF933E64" pageId="8" pageNumber="9">Description.</paragraph>
<paragraph id="96699E0E9C7C2C0A9D01AA0F237618A9" pageId="8" pageNumber="9">Ovoid with short rostrum, shell shiny sub-translucent. Sculpture of fine closely spaced commarginal lamellae. Specimen NHM_381 length 2 mm (Fig. 8).</paragraph>
<caption id="A39C8BA646CFD25EC5433B09C5234B0D" pageId="8" pageNumber="9">
<paragraph id="8506A5FA6E1FB8300588E3194F779589" pageId="8" pageNumber="9">
Figure 8.
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
<taxonomicName id="52C9C5EB0C1DEAFECD27B6090BCB3C1D" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
sp. (NHM_381). Scale bar: 1 mm. Image attribution Glover, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="genetic data">
<paragraph pageId="8" pageNumber="9">Genetic data.</paragraph>
<paragraph pageId="8" pageNumber="9">GenBank NHM_144 16S-MF157458, COI-MF157504; NHM_381 18S-MF157494, COI-MF157521.</paragraph>
<subSubSection id="34FD208B629D68335F917BD601188A26" pageId="8" pageNumber="9" type="genetic data">
<paragraph id="C4623281B6CA80F946240DD180C38482" pageId="8" pageNumber="9">Genetic data.</paragraph>
<paragraph id="D2D875AB3DBD96BEA5729492ADD768EE" pageId="8" pageNumber="9">GenBank NHM_144 16S-MF157458, COI-MF157504; NHM_381 18S-MF157494, COI-MF157521.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="remarks">
<paragraph pageId="8" pageNumber="9">Remarks.</paragraph>
<paragraph pageId="8" pageNumber="9">
<subSubSection id="CBA6E17F79B8463BC96A006BE1C3E0D4" pageId="8" pageNumber="9" type="remarks">
<paragraph id="140364AD5A64BCC8C6ADA29C46855A2B" pageId="8" pageNumber="9">Remarks.</paragraph>
<paragraph id="632CE7FD35C42E39A8E6A54287FDF185" pageId="8" pageNumber="9">
This species is morphologically very similar to the new
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
<taxonomicName id="8D3F0B951C28C76A2869A632BCC1494C" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
, its sister taxon in the molecular phylogenetic analyses (Fig. 12), and DNA might be required to properly identify the species. No genetic matches on GenBank.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="ecology">
<paragraph pageId="8" pageNumber="9">Ecology.</paragraph>
<paragraph pageId="8" pageNumber="9">Found in polymetallic nodule province.</paragraph>
<subSubSection id="9CA705A3FA52843A1D09794961733E48" pageId="8" pageNumber="9" type="ecology">
<paragraph id="47DDB98094EE71876155C92C09B5450B" pageId="8" pageNumber="9">Ecology.</paragraph>
<paragraph id="CF3682B0C1EED94F995B27E9083FB249" pageId="8" pageNumber="9">Found in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,87 +1,86 @@
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<paragraph pageId="2" pageNumber="165">Taxon classification Animalia Pantopoda Callipallenidae</paragraph>
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<paragraph id="4761D5E40626675B4754674FBBF388DD" pageId="2" pageNumber="165">Taxon classification Animalia Pantopoda Callipallenidae</paragraph>
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<subSubSection pageId="2" pageNumber="165" type="nomenclature">
<paragraph pageId="2" pageNumber="165">
<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene phantoma" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="phantoma">Callipallene phantoma (Dohrn, 1881)</taxonomicName>
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<paragraph id="00053588D6C4A15524BF927BD3149021" pageId="2" pageNumber="165">
<taxonomicName id="801BC94848BC23EB4BAA5422E437D0AB" ID-CoL="PS9J" authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene phantoma" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="phantoma">Callipallene phantoma (Dohrn, 1881)</taxonomicName>
Figures 26, 30, 31, 32, 33
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="165" type="reference_group">
<paragraph pageId="2" pageNumber="165">
<taxonomicName class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene phantoma subsp. crinita" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="subspecies" species="phantoma" subspecies="crinita">Callipallene phantoma crinita</taxonomicName>
<subSubSection id="41B95F6F326250961B8590F1045BCD64" pageId="2" pageNumber="165" type="reference_group">
<paragraph id="BF8016655F51CDF03AC0BCB36E868941" pageId="2" pageNumber="165">
<taxonomicName id="78C2CAA5603B3B171DAA91DBC7CA1373" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene phantoma subsp. crinita" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="subspecies" species="phantoma" subspecies="crinita">Callipallene phantoma crinita</taxonomicName>
Stock, 1952
</paragraph>
<paragraph pageId="2" pageNumber="165">
<taxonomicName class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene phantoma subsp. phantoma" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="subspecies" species="phantoma" subspecies="phantoma">Callipallene phantoma phantoma</taxonomicName>
<paragraph id="C70A23DF67725280232DAE75178D91E1" pageId="2" pageNumber="165">
<taxonomicName id="B08AE1D2E69AD0704ED7DE1B7AE3E6EF" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene phantoma subsp. phantoma" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="subspecies" species="phantoma" subspecies="phantoma">Callipallene phantoma phantoma</taxonomicName>
(Dohrn, 1881)
</paragraph>
<paragraph pageId="2" pageNumber="165">
<taxonomicName authority="Johnston, 1837" authorityName="Johnston" authorityYear="1837" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pallene phantoma" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="phantoma">Pallene phantoma</taxonomicName>
<paragraph id="50EA429AF768D3A7D663FE15B780F099" pageId="2" pageNumber="165">
<taxonomicName id="BB53B2FBB8505A926F0482BE13ECA563" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pallene phantoma" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="phantoma">Pallene phantoma</taxonomicName>
Dohrn, 1881
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="165" type="material">
<paragraph pageId="2" pageNumber="165">Material.</paragraph>
<paragraph pageId="2" pageNumber="165">ZSMA20071489: female; Brucoli, Sicily, Italy; 14.09.1988. ZSMA20071490: female; Banyuls-sur-mer, France; 07.2004. ZSMA20071491: female; Giglio Island, Italy; 11.04.2005; 20 m; red algae. ZSMA20071492: male; Giglio Island, Italy; 11.04.2005; 20 m; red algae. ZSMA20071493: male; Giglio Island, Italy; 11.04.2005; 20 m; red algae. ZSMA20071494: male; Rovinj, Croatia; 05.07.2005.</paragraph>
<subSubSection id="7B62D084BBCBC34BD757BF4C6A51B015" pageId="2" pageNumber="165" type="material">
<paragraph id="158E814C10CB021FE575CF77B2A2D216" pageId="2" pageNumber="165">Material.</paragraph>
<paragraph id="23776FFF2213BEF50A5E2A383630D669" pageId="2" pageNumber="165">ZSMA20071489: female; Brucoli, Sicily, Italy; 14.09.1988. ZSMA20071490: female; Banyuls-sur-mer, France; 07.2004. ZSMA20071491: female; Giglio Island, Italy; 11.04.2005; 20 m; red algae. ZSMA20071492: male; Giglio Island, Italy; 11.04.2005; 20 m; red algae. ZSMA20071493: male; Giglio Island, Italy; 11.04.2005; 20 m; red algae. ZSMA20071494: male; Rovinj, Croatia; 05.07.2005.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="165" type="remarks">
<paragraph pageId="2" pageNumber="165">Remarks.</paragraph>
<paragraph pageId="2" pageNumber="165">
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<subSubSection id="B760FE08C0A6CD46EC1B576751BCE8C1" pageId="2" pageNumber="165" type="remarks">
<paragraph id="521CA8F975328F7DA3C092C0F70297AC" pageId="2" pageNumber="165">Remarks.</paragraph>
<paragraph id="06777C62EB2B0DB792D45495A7F55D4B" pageId="2" pageNumber="165">
<bibRefCitation id="7ED9F9999E6E7CA99F78A8E50102F8A4" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
describes the auxiliary claws as half as long as the claw. According to
<bibRefCitation author="Stock, JH" journalOrPublisher="Beaufortia" pageId="6" pageNumber="169" pagination="1 - 15" title="Revision of the European representatives of the genus Callipallene Flynn, 1929 (Pycnogonida)." volume="13" year="1952">Stock (1952)</bibRefCitation>
<bibRefCitation id="FE35B65F746885A61059B5D29B5A089E" author="Stock, JH" journalOrPublisher="Beaufortia" pageId="6" pageNumber="169" pagination="1 - 15" title="Revision of the European representatives of the genus Callipallene Flynn, 1929 (Pycnogonida)." volume="13" year="1952">Stock (1952)</bibRefCitation>
the length of auxiliary claws can vary between specimens. Here all 6 specimens from 4 different locations have more or less rudimentary auxiliary claws (Fig. 31F; Fig. 33F, G).
</paragraph>
</subSubSection>

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@ -1,104 +1,103 @@
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<taxonomicName authority="sp. (NHM _ 345)" class="Scaphopoda" family="Gadilidae" genus="Gadila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gadila" order="Pholadomyoida" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">Gadila sp. (NHM_345)</taxonomicName>
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<paragraph pageId="24" pageNumber="25">Material examined.</paragraph>
<paragraph pageId="24" pageNumber="25">NHM_345 NHMUK 20170076, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/c301a72f-54cb-435e-8aae-17cf4d37675f</paragraph>
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<paragraph id="FE0A4C42A882E6799408DF9D30C537FD" pageId="24" pageNumber="25">Material examined.</paragraph>
<paragraph id="217242CC31FD128C58FBC1D4704AF26B" pageId="24" pageNumber="25">NHM_345 NHMUK 20170076, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/c301a72f-54cb-435e-8aae-17cf4d37675f</paragraph>
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<paragraph pageId="24" pageNumber="25">Short, glossy, transparent, maximum diameter near centre, ventral side curved, dorsal side near straight. Mouth simple, oblique. NHM_345 voucher specimen length 6 mm, width 1.4 mm (Fig. 23C).</paragraph>
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<paragraph id="E0E61AD33563A0911E611514D4F18FE8" pageId="24" pageNumber="25">Description.</paragraph>
<paragraph id="BD3844FA55764879864E1A7CCD9645D5" pageId="24" pageNumber="25">Short, glossy, transparent, maximum diameter near centre, ventral side curved, dorsal side near straight. Mouth simple, oblique. NHM_345 voucher specimen length 6 mm, width 1.4 mm (Fig. 23C).</paragraph>
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<paragraph pageId="24" pageNumber="25">Remarks.</paragraph>
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Forms a unique monophyletic clade distinct from other AB01 specimens. No genetic matches on GenBank. Phylogenetic tree supports placement in order
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(Figure 24). Genetic and image data made available for future study when better specimens available.
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<paragraph pageId="24" pageNumber="25">Ecology.</paragraph>
<paragraph pageId="24" pageNumber="25">Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.</paragraph>
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<paragraph id="2CE9E1F3BEF8EA4064BC664E75174B89" pageId="24" pageNumber="25">Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.</paragraph>
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<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene tiberi" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="tiberi">Callipallene tiberi (Dohrn, 1881)</taxonomicName>
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<taxonomicName id="EE328CDDE56F597A83AEAEB02BB24421" authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene tiberi" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="tiberi">Callipallene tiberi (Dohrn, 1881)</taxonomicName>
Figures 27, 37
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<taxonomicName class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene emaciata subsp. tiberii" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="subspecies" species="emaciata" subspecies="tiberii">Callipallene emaciata tiberii</taxonomicName>
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<taxonomicName id="758DD17DA46A89195E30F76B6CAAFAD6" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene emaciata subsp. tiberii" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="subspecies" species="emaciata" subspecies="tiberii">Callipallene emaciata tiberii</taxonomicName>
(Dohrn, 1881)
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<paragraph pageId="3" pageNumber="166">
<taxonomicName authority="Johnston, 1837" authorityName="Johnston" authorityYear="1837" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pallene tiberi" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="tiberi">Pallene tiberi</taxonomicName>
<paragraph id="FE2B94F21B735742CA05E113D1BF1CFC" pageId="3" pageNumber="166">
<taxonomicName id="FC87228DCD6C0C9FD98E2EA621E4E361" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pallene tiberi" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="tiberi">Pallene tiberi</taxonomicName>
Dohrn, 1881
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<paragraph pageId="3" pageNumber="166">Material.</paragraph>
<paragraph pageId="3" pageNumber="166">
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ZSMA20071488: female; Giglio Island, Italy; 12.04.2005; 1 m;
<taxonomicName class="Phaeophyceae" family="Stypocaulaceae" genus="Halopteris" higherTaxonomySource="CoL" kingdom="Chromista" lsidName="Halopteris" order="Sphacelariales" pageId="3" pageNumber="166" phylum="Ochrophyta" rank="genus">Halopteris</taxonomicName>
<taxonomicName id="519E4F8669544B727EBA07E9153BD4D7" class="Hydrozoa" family="Halopterididae" genus="Halopteris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halopteris" order="Leptothecata" pageId="3" pageNumber="166" phylum="Cnidaria" rank="genus">Halopteris</taxonomicName>
sp..
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<taxonomicName authority="Sars, 1888" authorityName="Sars" authorityYear="1888" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene producta" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="producta">Callipallene producta (Sars, 1888)</taxonomicName>
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Figures 26, 34, 35
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<taxonomicName class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene brevirostris subsp. producta" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="subspecies" species="brevirostris" subspecies="producta">Callipallene brevirostris producta</taxonomicName>
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<taxonomicName id="D7BB2CC830A172F1B70E79DA7E966317" class="Pycnogonida" family="Callipallenidae" genus="Callipallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Callipallene brevirostris subsp. producta" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="subspecies" species="brevirostris" subspecies="producta">Callipallene brevirostris producta</taxonomicName>
(Sars, 1888)
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<taxonomicName authority="Johnston, 1837" authorityName="Johnston" authorityYear="1837" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pallene producta" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="producta">Pallene producta</taxonomicName>
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<taxonomicName id="758FF38E489309386787A157121922A8" class="Pycnogonida" family="Callipallenidae" genus="Pallene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pallene producta" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="producta">Pallene producta</taxonomicName>
Sars, 1888
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<paragraph pageId="2" pageNumber="165">Material.</paragraph>
<paragraph pageId="2" pageNumber="165">ZSMA20071495: female; Giglio Island, Italy; 08.04.2005; 13-18 m; on brown algae. ZSMA20071496: female; Giglio Island, Italy; 08.04.2005; 13-18 m; on brown algae.</paragraph>
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<paragraph id="29FADA9E5519EF90AA4E7640E5D7D212" pageId="2" pageNumber="165">ZSMA20071495: female; Giglio Island, Italy; 08.04.2005; 13-18 m; on brown algae. ZSMA20071496: female; Giglio Island, Italy; 08.04.2005; 13-18 m; on brown algae.</paragraph>
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<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus angulatus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="angulatus">Anoplodactylus angulatus (Dohrn, 1881)</taxonomicName>
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Figures 45, 46, 47
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<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus angulirostris" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="angulirostris">Anoplodactylus angulirostris</taxonomicName>
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<taxonomicName id="36100F6733F6EAA2F54669109CE5AB2C" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodactylus angulirostris" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="angulirostris">Anoplodactylus angulirostris</taxonomicName>
(Dohrn, 1881)
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<paragraph pageId="3" pageNumber="166">
<taxonomicName class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium angulatum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="angulatum">Phoxichilidium angulatum</taxonomicName>
<paragraph id="CFAAA5D1397DEC3CFA47BB22D51F5B9A" pageId="3" pageNumber="166">
<taxonomicName id="DAEDC572D7E18443E520490F3BD43A11" class="Pycnogonida" family="Phoxichilidiidae" genus="Phoxichilidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilidium angulatum" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="angulatum">Phoxichilidium angulatum</taxonomicName>
Dohrn, 1881
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<paragraph pageId="3" pageNumber="166">Material.</paragraph>
<paragraph pageId="3" pageNumber="166">ZSMA20071478: female; Banyuls-sur-mer, France; 07.2004. ZSMA20071479: female; Giglio Island, Italy; 14.4.2005; 20 m; red algae.</paragraph>
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<paragraph id="5B05C4C7A705B1C751ECC768465C3992" pageId="3" pageNumber="166">ZSMA20071478: female; Banyuls-sur-mer, France; 07.2004. ZSMA20071479: female; Giglio Island, Italy; 14.4.2005; 20 m; red algae.</paragraph>
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@ -1,125 +1,124 @@
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<paragraph pageId="2" pageNumber="3">Material examined.</paragraph>
<paragraph pageId="2" pageNumber="3">NHM_051 NHMUK 20170038, collected 2013-10-09, 13.8372 -116.55843, 4336 m. http://data.nhm.ac.uk/object/49b2f599-bda4-4177-932f-59effe8a3320</paragraph>
<paragraph pageId="2" pageNumber="3">NHM_180 NHMUK 20170039, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/b84e470d-73bc-413b-88f9-3d702509a37a</paragraph>
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<paragraph id="D5269507578B039F1F7D77169DD8B56F" pageId="2" pageNumber="3">NHM_051 NHMUK 20170038, collected 2013-10-09, 13.8372 -116.55843, 4336 m. http://data.nhm.ac.uk/object/49b2f599-bda4-4177-932f-59effe8a3320</paragraph>
<paragraph id="9CB6D2167C5800024F5605C544E3FFFF" pageId="2" pageNumber="3">NHM_180 NHMUK 20170039, collected 2013-10-13, 13.93482 -116.55018, 4082 m. http://data.nhm.ac.uk/object/b84e470d-73bc-413b-88f9-3d702509a37a</paragraph>
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<subSubSection pageId="2" pageNumber="3" type="description">
<paragraph pageId="2" pageNumber="3">Description.</paragraph>
<paragraph pageId="2" pageNumber="3">Minute, thin-shelled, translucent, anteriorly extended, longer than high, umbones posterior of mid-line, posteriorly angulate, antero-dorsal margin long, evenly curved, shell surface smooth. Gill with single demibranch of about 10 widely spaced filaments, ventral edge of the gill does not cover the body pouches. Foot relatively large with distal bulb (Fig. 3). NHM_180 length 1.1 mm.</paragraph>
<caption pageId="2" pageNumber="3">
<paragraph pageId="2" pageNumber="3">
<subSubSection id="A54DD9F520F6FEAF239E474C6BD15395" pageId="2" pageNumber="3" type="description">
<paragraph id="A6F6142C76A7AA8C272D0BB2148064B4" pageId="2" pageNumber="3">Description.</paragraph>
<paragraph id="67CF03B3E3C53A339A0661C99CFFD852" pageId="2" pageNumber="3">Minute, thin-shelled, translucent, anteriorly extended, longer than high, umbones posterior of mid-line, posteriorly angulate, antero-dorsal margin long, evenly curved, shell surface smooth. Gill with single demibranch of about 10 widely spaced filaments, ventral edge of the gill does not cover the body pouches. Foot relatively large with distal bulb (Fig. 3). NHM_180 length 1.1 mm.</paragraph>
<caption id="8E1086F08E1FD590DD41EE16EA77EC32" pageId="2" pageNumber="3">
<paragraph id="C5F2343FABD93A178ADBB31A1D8D9ABB" pageId="2" pageNumber="3">
Figure 3.
<taxonomicName class="Bivalvia" family="Thyasiridae" genus="Thyasira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thyasira" order="Lucinida" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">Thyasira</taxonomicName>
<taxonomicName id="07C5D136CE160ECBD58AAD2390EC8AAF" class="Bivalvia" family="Thyasiridae" genus="Thyasira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thyasira" order="Lucinida" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">Thyasira</taxonomicName>
sp. (NHM_180) A Preserved specimen (NHM_180) with pieces of polymetallic nodule adhered to shell margin B Additional small specimen (live imaged at sea) NHM_051. Scale bar: 0.5 mm (A). Image attribution Glover, Dahlgren and Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="genetic data">
<paragraph pageId="3" pageNumber="4">
<pageBreakToken pageId="3" pageNumber="4" start="start">Genetic</pageBreakToken>
<subSubSection id="08C08AAC42CAFEF17475B7835B43EBAE" pageId="3" pageNumber="4" type="genetic data">
<paragraph id="8A59D90730C8950A85EE5C299DE6451E" pageId="3" pageNumber="4">
<pageBreakToken id="8C4893F4CEC9D315A37ADC4D1BE27E1C" pageId="3" pageNumber="4" start="start">Genetic</pageBreakToken>
data.
</paragraph>
<paragraph pageId="3" pageNumber="4">GenBank NHM_051 18S-MF157468, COI-MF157501; NHM_180 18S-MF157478.</paragraph>
<paragraph id="DF31F55BD68AF620A72E0B81635B2261" pageId="3" pageNumber="4">GenBank NHM_051 18S-MF157468, COI-MF157501; NHM_180 18S-MF157478.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="remarks">
<paragraph pageId="3" pageNumber="4">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="4">
<subSubSection id="3C52B9FF0221742AA96938AFEFA75A11" pageId="3" pageNumber="4" type="remarks">
<paragraph id="1CF1CF7121C4B6A441829C28BA6C1369" pageId="3" pageNumber="4">Remarks.</paragraph>
<paragraph id="8A91B2757296A78893BF9E8DAA572F8E" pageId="3" pageNumber="4">
Forms a monophyletic clade with four other thyasirid species (Fig. 5) and distinct from all other AB01 specimens. No genetic matches on GenBank. Morphologically the species is similar in shape to abyssal thyasirid species (
<taxonomicName class="Bivalvia" family="Thyasiridae" genus="Thyasira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thyasira inflata" order="Lucinida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="inflata">Thyasira inflata</taxonomicName>
<taxonomicName id="A0E55EBF7F760220058BF9A7484BC385" class="Bivalvia" family="Thyasiridae" genus="Thyasira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thyasira inflata" order="Lucinida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="inflata">Thyasira inflata</taxonomicName>
,
<taxonomicName lsidName="T. transversa" pageId="3" pageNumber="4" rank="species" species="transversa">T. transversa</taxonomicName>
<taxonomicName id="5CAAD61B16914A779BDA0C9BA7C85655" lsidName="T. transversa" pageId="3" pageNumber="4" rank="species" species="transversa">T. transversa</taxonomicName>
) from the south Atlantic described and placed in
<taxonomicName class="Bivalvia" family="Thyasiridae" genus="Thyasira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thyasira (Mendicula)" order="Lucinida" pageId="3" pageNumber="4" phylum="Mollusca" rank="subGenus" subGenus="Mendicula">Thyasira (Mendicula)</taxonomicName>
<taxonomicName id="D7B3B14AD489B334160B5970413FD010" class="Bivalvia" family="Thyasiridae" genus="Thyasira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thyasira (Mendicula)" order="Lucinida" pageId="3" pageNumber="4" phylum="Mollusca" rank="subGenus" subGenus="Mendicula">Thyasira (Mendicula)</taxonomicName>
by Payne &amp; Allen (1991) but not similar to the type species of
<taxonomicName class="Bivalvia" family="Thyasiridae" genus="Mendicula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mendicula (Lucina) induta" order="Lucinida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="induta" subGenus="Lucina">Mendicula (Lucina) induta</taxonomicName>
<taxonomicName id="89C692D47CF79AB97CE50B76413F7A58" class="Bivalvia" family="Thyasiridae" genus="Mendicula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mendicula (Lucina) induta" order="Lucinida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="induta" subGenus="Lucina">Mendicula (Lucina) induta</taxonomicName>
Hedley, 1907 =
<taxonomicName lsidName="M. memorata" pageId="3" pageNumber="4" rank="species" species="memorata">M. memorata</taxonomicName>
<taxonomicName id="40CBB614CC943FD8CB5B80C660140F60" lsidName="M. memorata" pageId="3" pageNumber="4" rank="species" species="memorata">M. memorata</taxonomicName>
Iredale, 1924) or the widespread
<taxonomicName class="Bivalvia" family="Thyasiridae" genus="Mendicula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mendicula ferruginosa" order="Lucinida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="ferruginosa">Mendicula ferruginosa</taxonomicName>
<taxonomicName id="3F7F5EA915F1215D880C4873AE617C7D" class="Bivalvia" family="Thyasiridae" genus="Mendicula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mendicula ferruginosa" order="Lucinida" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="ferruginosa">Mendicula ferruginosa</taxonomicName>
(Forbes, 1844). No similarly shaped species has been recorded from the abyssal eastern Pacific.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="ecology">
<paragraph pageId="3" pageNumber="4">Ecology.</paragraph>
<paragraph pageId="3" pageNumber="4">Found in polymetallic nodule province.</paragraph>
<subSubSection id="7CE065F95CECC51DC710FC972AFDD2F5" pageId="3" pageNumber="4" type="ecology">
<paragraph id="66C9A1C5E3A609A6D474B09E60463BF5" pageId="3" pageNumber="4">Ecology.</paragraph>
<paragraph id="A22A16F57F35A87BACC57C8C6704A313" pageId="3" pageNumber="4">Found in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,84 +1,83 @@
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<paragraph pageId="2" pageNumber="165">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
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<paragraph id="70213500925D09099B48B8B3DC8BEFB4" pageId="2" pageNumber="165">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
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<paragraph pageId="2" pageNumber="165">
<taxonomicName authority="Hodge, 1864" authorityName="Hodge" authorityYear="1864" class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella longipes" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="longipes">Ammothella longipes (Hodge, 1864)</taxonomicName>
<subSubSection id="126C9FB400010D8B217F036300100C87" pageId="2" pageNumber="165" type="nomenclature">
<paragraph id="170C87A88640C6E3B01B2CD1D1B3691E" pageId="2" pageNumber="165">
<taxonomicName id="04246B9874A0C879F1691049D83F9306" ID-CoL="CVLF" authority="Hodge, 1864" authorityName="Hodge" authorityYear="1864" class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella longipes" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="longipes">Ammothella longipes (Hodge, 1864)</taxonomicName>
Figures 3, 17
</paragraph>
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<paragraph pageId="2" pageNumber="165">
<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia longipes" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="longipes">Achelia longipes</taxonomicName>
<subSubSection id="FB65D57D33A08658E944CEB502E7672A" pageId="2" pageNumber="165" type="reference_group">
<paragraph id="2829F0D71E4758F07D1560075E50009C" pageId="2" pageNumber="165">
<taxonomicName id="87B3BBAD04D4673FD118340777DCDB1D" class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia longipes" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="longipes">Achelia longipes</taxonomicName>
(Hodge, 1864)
</paragraph>
<paragraph pageId="2" pageNumber="165">
<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia magnirostris" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="magnirostris">Achelia magnirostris</taxonomicName>
<paragraph id="B5AC2F98BE0DB4F32FD4D37512DF7BE3" pageId="2" pageNumber="165">
<taxonomicName id="9FED5632CE1911E833BC1420D456F024" class="Pycnogonida" family="Ammotheidae" genus="Achelia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achelia magnirostris" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="magnirostris">Achelia magnirostris</taxonomicName>
(Dohrn, 1881)
</paragraph>
<paragraph pageId="2" pageNumber="165">
<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea longipes" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="longipes">Ammothea longipes</taxonomicName>
<paragraph id="15916B611B05D51C6DB4DD82B73565E4" pageId="2" pageNumber="165">
<taxonomicName id="8CAD42BC1869D407DE81E00CED713D7E" class="Pycnogonida" family="Ammotheidae" genus="Ammothea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothea longipes" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="longipes">Ammothea longipes</taxonomicName>
Hodge, 1864
</paragraph>
<paragraph pageId="2" pageNumber="165">
<taxonomicName class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella magnirostris" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="magnirostris">Ammothella magnirostris</taxonomicName>
<paragraph id="389A5346E31704F42A949144249A75A0" pageId="2" pageNumber="165">
<taxonomicName id="08F6BA1744015BD82FB9DD1E801A4B83" class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella magnirostris" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="magnirostris">Ammothella magnirostris</taxonomicName>
(Dohrn, 1881)
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<paragraph pageId="2" pageNumber="165">Material.</paragraph>
<paragraph pageId="2" pageNumber="165">ZSMA20071477: female; Bayuls-sur-Mer, France; 07.2004.</paragraph>
<subSubSection id="38691CD5B2D1DBE326FC85573B3F7381" pageId="2" pageNumber="165" type="material">
<paragraph id="F13D0E9B91525784CCF0B93BE3B9D9FE" pageId="2" pageNumber="165">Material.</paragraph>
<paragraph id="A9123659991CFCAAECE030D8C1D844F2" pageId="2" pageNumber="165">ZSMA20071477: female; Bayuls-sur-Mer, France; 07.2004.</paragraph>
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<treatment ID-GBIF-Taxon="135489967" LSID="urn:lsid:plazi:treatment:C0CE17664142D67ACC3322871333FD62" httpUri="http://treatment.plazi.org/id/C0CE17664142D67ACC3322871333FD62" lastPageNumber="22" pageId="21" pageNumber="22">
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<paragraph pageId="21" pageNumber="22">
<taxonomicName authority="Sirenko, 2015" authorityName="Sirenko" authorityYear="2015" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochiton macleani" order="Pholadomyoida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="macleani">Leptochiton macleani Sirenko, 2015</taxonomicName>
<treatment id="C0CE17664142D67ACC3322871333FD62" ID-GBIF-Taxon="135489967" LSID="urn:lsid:plazi:treatment:C0CE17664142D67ACC3322871333FD62" httpUri="http://treatment.plazi.org/id/C0CE17664142D67ACC3322871333FD62" lastPageNumber="22" pageId="21" pageNumber="22" scope_class="Polyplacophora" scope_family="Leptochitonidae" scope_phylum="Mollusca">
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<paragraph id="C87DE0B6731143AD0343EF995758F4AC" pageId="21" pageNumber="22">
<taxonomicName id="8BE3DE8E63B730F3090350CEAC36D078" authority="Sirenko, 2015" authorityName="Sirenko" authorityYear="2015" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Leptochiton macleani" order="Pholadomyoida" pageId="21" pageNumber="22" rank="species" species="macleani">Leptochiton macleani Sirenko, 2015</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="materials_examined">
<paragraph pageId="21" pageNumber="22">Material examined.</paragraph>
<paragraph pageId="21" pageNumber="22">NHM_446 NHMUK 20170073.1-2, collected 2013-10-20, 13.86367 -116.54432, 4050 m. http://data.nhm.ac.uk/object/d69b581d-8a79-4c4d-8f70-88b2ec07d86e</paragraph>
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<paragraph id="11EA63A3B82817EE8C1260193CE4D7A4" pageId="21" pageNumber="22">Material examined.</paragraph>
<paragraph id="897D0D75773EE894CA1039FB34CD0393" pageId="21" pageNumber="22">NHM_446 NHMUK 20170073.1-2, collected 2013-10-20, 13.86367 -116.54432, 4050 m. http://data.nhm.ac.uk/object/d69b581d-8a79-4c4d-8f70-88b2ec07d86e</paragraph>
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<subSubSection pageId="21" pageNumber="22" type="description">
<paragraph pageId="21" pageNumber="22">Description.</paragraph>
<paragraph pageId="21" pageNumber="22">
<subSubSection id="9EC177ED6283940CEE03E6AB186FFE45" pageId="21" pageNumber="22" type="description">
<paragraph id="C1D37E42DE750C6C39B86247978F92BD" pageId="21" pageNumber="22">Description.</paragraph>
<paragraph id="BC7E92FB0ACE594EE832E0495E1665DC" pageId="21" pageNumber="22">
The form and pattern of tegmental granules together with the three aesthete pores are most similar to the images of
<taxonomicName class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochiton macleani" order="Lepidopleurida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="macleani">Leptochiton macleani</taxonomicName>
<taxonomicName id="4039CF9F3D6F156F3D8E741A42275950" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochiton macleani" order="Lepidopleurida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="macleani">Leptochiton macleani</taxonomicName>
(Sirenko, 2015: figs 34-36). Voucher NHM_446 length approx 10 mm, width 3.2 mm (Fig. 21).
</paragraph>
<caption pageId="21" pageNumber="22">
<paragraph pageId="21" pageNumber="22">
<caption id="42FBEBEADE8ED1E57DF5535A4F1C5179" pageId="21" pageNumber="22">
<paragraph id="6550C3F030F793511119809B3439E786" pageId="21" pageNumber="22">
Figure 21.
<taxonomicName class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochiton macleani" order="Lepidopleurida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="macleani">Leptochiton macleani</taxonomicName>
<taxonomicName id="AE1CF639343E8D21D2B22FA9DB73F837" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochiton macleani" order="Lepidopleurida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="macleani">Leptochiton macleani</taxonomicName>
Sirenko, 2015. NHM_446 voucher specimen. A Live specimen (lateral view) after recovery from the ROV scoop sample B Preserved specimen (ventro-lateral view) following DNA extraction C Dorsal view D surface detail E SEM of tegmentum surface and pores. Scale bars: 4 mm (A); 0.5 mm (D); 0.3 mm (E). Image attribution Glover, Taylor, Ikebe, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="genetic data">
<paragraph pageId="21" pageNumber="22">Genetic data.</paragraph>
<paragraph pageId="21" pageNumber="22">GenBank NHM_446 16S-MF157466, 18S-MF157497, COI-MF157523.</paragraph>
<subSubSection id="290B168D269BE42D89EA7827C3EB5E4B" pageId="21" pageNumber="22" type="genetic data">
<paragraph id="4243C98AF39E93869A5A83DB7480FB0E" pageId="21" pageNumber="22">Genetic data.</paragraph>
<paragraph id="EAFF695CDDFDB5F608F00ABB6B35603F" pageId="21" pageNumber="22">GenBank NHM_446 16S-MF157466, 18S-MF157497, COI-MF157523.</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="remarks">
<paragraph pageId="21" pageNumber="22">Remarks.</paragraph>
<paragraph pageId="21" pageNumber="22">
<bibRefCitation author="Sirenko, B" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="151 - 202" title="Shallow and deep-sea chitons of the genus Leptochiton Gray, 1847 (Mollusca: Polyplacophora: Lepidopleurida) from Peruvian and Chilean waters." url="https://doi.org/10.11646/zootaxa.4033.2.1" volume="4033" year="2015">Sirenko (2015)</bibRefCitation>
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<paragraph id="A7B9D3BF14C82871EB301405CFF94732" pageId="21" pageNumber="22">Remarks.</paragraph>
<paragraph id="956EA8E18D59151A5BBF70C09A027B9B" pageId="21" pageNumber="22">
<bibRefCitation id="EF83EA9BB6F3E8FF5EB08A973ACA418F" author="Sirenko, B" journalOrPublisher="Zootaxa" pageId="31" pageNumber="32" pagination="151 - 202" title="Shallow and deep-sea chitons of the genus Leptochiton Gray, 1847 (Mollusca: Polyplacophora: Lepidopleurida) from Peruvian and Chilean waters." url="https://doi.org/10.11646/zootaxa.4033.2.1" volume="4033" year="2015">Sirenko (2015)</bibRefCitation>
has recently reviewed
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of the southeastern Pacific Ocean and described several new species that had been previously confounded with
<taxonomicName class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochiton belknapi" order="Lepidopleurida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="belknapi">Leptochiton belknapi</taxonomicName>
<taxonomicName id="F71B82770BBE507A84F870DBB7F46B89" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochiton belknapi" order="Lepidopleurida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="belknapi">Leptochiton belknapi</taxonomicName>
Dall, 1878. The specimen morphologically matches
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, type locality Peru-Chile Trench, East Pacific, 4600 m depth. Forms a unique monophyletic clade distinct from other AB01 specimens. No genetic matches on GenBank. In the molecular analyses based on the 18S and COI genes, it falls with strong support as sister taxon to two other
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species, but in the phylogenetic tree the genus
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<taxonomicName id="6DA3B02F92CD3D73A4E021CD2A79E337" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptochiton" order="Lepidopleurida" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">Leptochiton</taxonomicName>
is not monophyletic (Fig. 22).
</paragraph>
<caption pageId="21" pageNumber="22">
<paragraph pageId="21" pageNumber="22">
<caption id="0AC1CFBAABF6C6BA933D2A2DD632411A" pageId="21" pageNumber="22">
<paragraph id="8CCDEE052C36337E8888275A8643415C" pageId="21" pageNumber="22">
Figure 22. Phylogenetic analysis of
<taxonomicName family="Leptochitonidae" lsidName="" pageId="21" pageNumber="22" rank="family">Leptochitonidae</taxonomicName>
<taxonomicName id="11E6986FB5D52B681412117CDF289A16" class="Polyplacophora" family="Leptochitonidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidopleurida" pageId="21" pageNumber="22" phylum="Mollusca" rank="family">Leptochitonidae</taxonomicName>
,
<taxonomicName class="Polyplacophora" lsidName="Pteriomorphia" pageId="21" pageNumber="22" rank="class">Polyplacophora</taxonomicName>
<taxonomicName id="99C38C07D660A059705D61A0EB63CE12" class="Polyplacophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" pageId="21" pageNumber="22" phylum="Mollusca" rank="class">Polyplacophora</taxonomicName>
. 50% majority rule consensus tree from the Bayesian analyses, combining 18S and COI. Asterisks denotes posterior probability values of 95 or above.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="ecology">
<paragraph pageId="21" pageNumber="22">Ecology.</paragraph>
<paragraph pageId="21" pageNumber="22">Specimen collected from an ROV scoop in region of sediment and polymetallic nodules, presumed living associated or on the nodule surface, but not directly observed doing so.</paragraph>
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<paragraph id="6421CC441EA6567288330A151D541F8F" pageId="21" pageNumber="22">Ecology.</paragraph>
<paragraph id="39BF7FD82B26FC3265A96478520D7367" pageId="21" pageNumber="22">Specimen collected from an ROV scoop in region of sediment and polymetallic nodules, presumed living associated or on the nodule surface, but not directly observed doing so.</paragraph>
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</treatment>
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@ -1,82 +1,81 @@
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<paragraph pageId="2" pageNumber="165">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
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<paragraph id="5233DF89A80A6DDE29551137C9F32803" pageId="2" pageNumber="165">Taxon classification Animalia Pantopoda Ammotheidae</paragraph>
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<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Ammotheidae" genus="Trygaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trygaeus communis" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="communis">Trygaeus communis Dohrn, 1881</taxonomicName>
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Figures 4, 22, 23, 24, 25
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<paragraph pageId="2" pageNumber="165">Material.</paragraph>
<paragraph pageId="2" pageNumber="165">
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<paragraph id="D18C666CF4861AD0E1F641E23466DEBA" pageId="2" pageNumber="165">Material.</paragraph>
<paragraph id="69D24ECBB605CCCC24D54CC4DD35071D" pageId="2" pageNumber="165">
ZSMA20071509: female; Giglio Island, Italy; 12.04.2005. ZSMA20071510: male; Giglio Island, Italy; 12.04.2005. ZSMA20071511: male; Giglio Island, Italy; 12.04.2005. ZSMA20071512: male; Giglio Island, Italy; 12.04.2005. ZSMA20071513: male; Cape Kamenjak, Croatia; 03.05.2007; 0-5 m;
<taxonomicName genus="Stypocaulon" lsidName="Stypocaulon scoparium" pageId="2" pageNumber="165" rank="species" species="scoparium">Stypocaulon scoparium</taxonomicName>
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(Linnaeus)
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, 1843.
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<paragraph pageId="2" pageNumber="165">Remarks.</paragraph>
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<paragraph id="0711F6E3C6D42A319B9BB1C491322BF1" pageId="2" pageNumber="165">Remarks.</paragraph>
<paragraph id="B3986F7D2579BC689B5C9566D57CEDDB" pageId="2" pageNumber="165">
According to
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<bibRefCitation id="34C49A4CFDD50B46C80766C73A4C90F7" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
,
<bibRefCitation author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
<bibRefCitation id="DC92C404F668FF174355FFA886AB34F2" author="Bouvier, EL" journalOrPublisher="Paris" pageId="5" pageNumber="168" title="Faune de France - Pycnogonides" year="1923">Bouvier (1923)</bibRefCitation>
and
<bibRefCitation author="Stock, JH" journalOrPublisher="Vie et Milieu" pageId="6" pageNumber="169" pagination="407 - 417" title="Sur quelques Pycnogonides de la region de Banyuls (3 e note)." volume="17" year="1966">Stock (1966)</bibRefCitation>
<bibRefCitation id="93FD5DA1532959FF3EF96B12406137EE" author="Stock, JH" journalOrPublisher="Vie et Milieu" pageId="6" pageNumber="169" pagination="407 - 417" title="Sur quelques Pycnogonides de la region de Banyuls (3 e note)." volume="17" year="1966">Stock (1966)</bibRefCitation>
the palps vary in the number of articles from 4 to 7 and the oviger from 6 to 10. Here the palp has 7 articles (Fig. 23A) and the male and female oviger have 9 articles (Fig. 23C, D; Fig. 25D). Other specimens not depicted have four palp articles.
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<treatment ID-GBIF-Taxon="135489988" LSID="urn:lsid:zoobank.org:act:66E692B5-7C61-4ADC-9539-EFC085424147" httpUri="http://treatment.plazi.org/id/CC5F0ABB15F6989E849383880CD39530" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection pageId="7" pageNumber="8" type="nomenclature">
<paragraph pageId="7" pageNumber="8">
<taxonomicName LSID="http://zoobank.org/66E692B5-7C61-4ADC-9539-EFC085424147" authority="Taylor &amp; Wiklund" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Pholadomyoida" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni Taylor &amp; Wiklund</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="8">sp. n.</taxonomicNameLabel>
<treatment id="CC5F0ABB15F6989E849383880CD39530" ID-GBIF-Taxon="135489988" LSID="urn:lsid:zoobank.org:act:66E692B5-7C61-4ADC-9539-EFC085424147" httpUri="http://treatment.plazi.org/id/CC5F0ABB15F6989E849383880CD39530" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" scope_class="Bivalvia" scope_order="Nuculanoida" scope_phylum="Mollusca" scope_subClass="Protobranchia">
<subSubSection id="6B7309B8D136DED9F697BA520B201C70" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="5D6EF44B8BB9354FE1B67388DBC863C2" pageId="7" pageNumber="8">
<taxonomicName id="30683AEC4529E8C08F4F22DDC01AA274" LSID="http://zoobank.org/66E692B5-7C61-4ADC-9539-EFC085424147" authority="Taylor &amp; Wiklund" family="Nuculanidae" genus="Ledella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ledella knudseni" order="Pholadomyoida" pageId="7" pageNumber="8" rank="species" species="knudseni">Ledella knudseni Taylor &amp; Wiklund</taxonomicName>
<taxonomicNameLabel id="02FF18AF3A4CF515DACFB0C168552E23" pageId="7" pageNumber="8">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="materials_examined">
<paragraph pageId="7" pageNumber="8">Material examined.</paragraph>
<paragraph pageId="7" pageNumber="8">Paratype NHM_288A NHMUK 20170047.1-2, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/8aec47f4-dcec-4668-8398-9e4b0c28ecb8</paragraph>
<paragraph pageId="7" pageNumber="8">Holotype NHM_288C NHMUK 20170048, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/f1886d78-22bf-403e-bdb2-784b91c0eb12</paragraph>
<subSubSection id="42C8FDD5E455D26E4C3AF604F40BBF37" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="664B8DD1DF0DB184921BEA8CD25FFA10" pageId="7" pageNumber="8">Material examined.</paragraph>
<paragraph id="73BF21513025096871C08B72C4394957" pageId="7" pageNumber="8">Paratype NHM_288A NHMUK 20170047.1-2, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/8aec47f4-dcec-4668-8398-9e4b0c28ecb8</paragraph>
<paragraph id="D9B3CDC4EDD106787CE8B45912BBBC1E" pageId="7" pageNumber="8">Holotype NHM_288C NHMUK 20170048, collected 2013-10-17, 13.75583 -116.48667, 4076 m. http://data.nhm.ac.uk/object/f1886d78-22bf-403e-bdb2-784b91c0eb12</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph pageId="7" pageNumber="8">Description.</paragraph>
<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="8BFE55E602D09AE508AD6F12D7AF7A92" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="AF963D49C19E2130F05708FCCF900015" pageId="7" pageNumber="8">Description.</paragraph>
<paragraph id="5D3D08B2E12413BA75D886D96E93CDA4" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Shell relatively thick, robust. Ovoid with short rostrum, umbones broad, prominent; postero ventral margin sinuous; broad, shallow sulcus extending from umbones to posteroventral margin. Sculpture of low, relatively broad, closely
<pageBreakToken pageId="8" pageNumber="9" start="start">spaced</pageBreakToken>
<pageBreakToken id="1B20E545193C9D713A550EBF11060D76" pageId="8" pageNumber="9" start="start">spaced</pageBreakToken>
, commarginal lamellae; fine radial striations on rostrum and juvenile shell. Ligament internal, situated on broad resilium beneath umbones. Hinge robust, with 8-9 chevron shaped, blunt teeth to either side of ligament. Inner shell margin smooth. Prodissoconch large, ellipsoidal 0.3 mm long, with sharp rim, surface irregularly pitted. Holotype NHM_288C shell length 2.2 mm, width 1.5 mm; paratype NHM_288A shell length 2.1 mm, height 1.5 mm. (Figure 7).
</paragraph>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
<caption id="0BE2D147EF0AB5BFDBD88DF182351E6B" pageId="8" pageNumber="9">
<paragraph id="B1CEA5742ECD08A856440AD5EE1E7BD5" pageId="8" pageNumber="9">
Figure 7.
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
<taxonomicName id="23CA6FC9AFAB6286CA8F90C0D9A1796F" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
sp. n. A Holotype, specimen NHM_288c B Paratype, specimen NHM_288a C Specimen NHM 288a dissected prior to DNA sequencing and SEM
<normalizedToken originalValue="DG">D-G</normalizedToken>
<normalizedToken id="463F528E1D57E2CAE5C059DDADCD8148" originalValue="DG">D-G</normalizedToken>
SEM of valve, hinge teeth and protoconch. Scale bars: 1 mm (
<normalizedToken originalValue="BC">B-C</normalizedToken>
<normalizedToken id="FFF160641FE5F79C0EA98F1BE8A88C9E" originalValue="BC">B-C</normalizedToken>
); 0.5 mm (
<normalizedToken originalValue="DE">D-E</normalizedToken>
<normalizedToken id="0B1E492E591FD63FCE113BE2671E8DE9" originalValue="DE">D-E</normalizedToken>
); 0.1 mm (
<normalizedToken originalValue="FG">F-G</normalizedToken>
<normalizedToken id="EF4B7E1BC6D46C59BE72A4CD3E59A681" originalValue="FG">F-G</normalizedToken>
). Image attribution Glover, Taylor, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="genetic data">
<paragraph pageId="8" pageNumber="9">Genetic data.</paragraph>
<paragraph pageId="8" pageNumber="9">GenBank NHM_288A COI-MF157515; NHM_288C 18S-MF157491, COI-MF157516.</paragraph>
<subSubSection id="EA177AD5706BDBA9C373A592162B69D3" pageId="8" pageNumber="9" type="genetic data">
<paragraph id="E24ADC95218C6F751DE65FB2F207C3F6" pageId="8" pageNumber="9">Genetic data.</paragraph>
<paragraph id="B59E80AD5B95C5B1C46FF5F7FFE8A150" pageId="8" pageNumber="9">GenBank NHM_288A COI-MF157515; NHM_288C 18S-MF157491, COI-MF157516.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="remarks">
<paragraph pageId="8" pageNumber="9">Remarks.</paragraph>
<paragraph pageId="8" pageNumber="9">
<subSubSection id="3FC6B2396601ADA33AF7119750E598F8" pageId="8" pageNumber="9" type="remarks">
<paragraph id="0942A7CA7A52032C80747C71C2C0860A" pageId="8" pageNumber="9">Remarks.</paragraph>
<paragraph id="0E720D441BFD1BA48C44FFFEF03A48BB" pageId="8" pageNumber="9">
Similar in form to
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella ultima" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ultima">Ledella ultima</taxonomicName>
<taxonomicName id="B97408288B5F6F6712CA272918E83492" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella ultima" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ultima">Ledella ultima</taxonomicName>
(Smith, 1885) widespread in the abyssal Atlantic (
<bibRefCitation author="Allen, JA" journalOrPublisher="Malacologia" pageId="29" pageNumber="30" pagination="57 - 103" title="Bivalvia of the deep Atlantic." url="https://doi.org/10.4002/0076-2997-50.1.57" volume="50" year="2008">Allen 2008</bibRefCitation>
<bibRefCitation id="8101B0E6277D3CF955E250DCB5459291" author="Allen, JA" journalOrPublisher="Malacologia" pageId="29" pageNumber="30" pagination="57 - 103" title="Bivalvia of the deep Atlantic." url="https://doi.org/10.4002/0076-2997-50.1.57" volume="50" year="2008">Allen 2008</bibRefCitation>
), but has a less massive hinge with more teeth, 8-9 compared with 6-8 in
<taxonomicName lsidName="L. ultima" pageId="8" pageNumber="9" rank="species" species="ultima">L. ultima</taxonomicName>
<taxonomicName id="72F627C4A7B072BA439706E8721DEE2D" lsidName="L. ultima" pageId="8" pageNumber="9" rank="species" species="ultima">L. ultima</taxonomicName>
. Also similar is the species identified by
<bibRefCitation author="Knudsen, J" journalOrPublisher="Galathea Reports" pageId="30" pageNumber="31" pagination="1 - 241" title="The systematics and biology of abyssal and hadal Bivalvia." volume="11" year="1970">Knudsen (1970)</bibRefCitation>
<bibRefCitation id="57FDB3AE36176E5EC90898B01906B3D7" author="Knudsen, J" journalOrPublisher="Galathea Reports" pageId="30" pageNumber="31" pagination="1 - 241" title="The systematics and biology of abyssal and hadal Bivalvia." volume="11" year="1970">Knudsen (1970)</bibRefCitation>
as
<taxonomicName lsidName="L. ultima" pageId="8" pageNumber="9" rank="species" species="ultima">L. ultima</taxonomicName>
<taxonomicName id="C457FA68C37087E19CBB980CBCD465AE" lsidName="L. ultima" pageId="8" pageNumber="9" rank="species" species="ultima">L. ultima</taxonomicName>
from the Sunda Trench in Indian Ocean at 3810 m. The only species recorded from the deep eastern Pacific is
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella dicella" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="dicella">Ledella dicella</taxonomicName>
<taxonomicName id="18F6CB30A9A620046C4D5948786F1EBC" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella dicella" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="dicella">Ledella dicella</taxonomicName>
(Dall, 1908) from 734-1200 m off Ecuador but this lacks the short rostrum and has 12-13 hinge teeth on each side of the ligament (
<bibRefCitation author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott 2012</bibRefCitation>
<bibRefCitation id="E67C49E05FE00DB4C6B47E3C9816336B" author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott 2012</bibRefCitation>
pl. 26). No genetic matches on GenBank.
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
<taxonomicName id="134242A39A0EE7A368DC641C83C8BD7D" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella knudseni" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="knudseni">Ledella knudseni</taxonomicName>
groups in a small subclade with but is distinct from the Atlantic species
<taxonomicName lsidName="L. ultima" pageId="8" pageNumber="9" rank="species" species="ultima">L. ultima</taxonomicName>
<taxonomicName id="6F4EE6D3F285F1DC4B9C5A04514DECA8" lsidName="L. ultima" pageId="8" pageNumber="9" rank="species" species="ultima">L. ultima</taxonomicName>
and
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella jamesi" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="jamesi">Ledella jamesi</taxonomicName>
<taxonomicName id="62FB9194C0E32B897D8276CCA1DEF487" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella jamesi" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="jamesi">Ledella jamesi</taxonomicName>
Allen &amp; Hannah, 1989, as well as another
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
<taxonomicName id="150C3002D7EB9D0634A24B625889DE67" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
species from this study in the Pacific,
<taxonomicName class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
<taxonomicName id="E942378A92E0EF55D675E0C911CD8FE0" class="Bivalvia" family="Nuculanidae" genus="Ledella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ledella" order="Nuculanida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Ledella</taxonomicName>
sp. (NHM_381) (Figure 12). The new species can be confused with juveniles of
<taxonomicName lsidName="B. calcar" pageId="8" pageNumber="9" rank="species" species="calcar">B. calcar</taxonomicName>
<taxonomicName id="CECD739474E54407C965419B8D8AF8D7" lsidName="B. calcar" pageId="8" pageNumber="9" rank="species" species="calcar">B. calcar</taxonomicName>
(see above), but shell is less shiny and iridescent, and ribs are more pronounced. DNA may be required to confirm identification.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="etymology">
<paragraph pageId="8" pageNumber="9">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="9">
<subSubSection id="80EDE35B5BB5437F86D8255EB09543C2" pageId="8" pageNumber="9" type="etymology">
<paragraph id="305A4FABAC196CF50F0B6B10E618CC7B" pageId="8" pageNumber="9">Etymology.</paragraph>
<paragraph id="C901AFE7700A140B8C8E2C71CBC6E927" pageId="8" pageNumber="9">
Named for
<normalizedToken originalValue="Jørgen">Jorgen</normalizedToken>
<normalizedToken id="5F8E3C1B50AB93284DC3A6DED9C57756" originalValue="Jørgen">Jorgen</normalizedToken>
Knudsen (1918-2009), deep-sea bivalve systematist and author of the Galathea Report on abyssal and hadal
<taxonomicName class="Bivalvia" lsidName="Protobranchia" pageId="8" pageNumber="9" rank="class">Bivalvia</taxonomicName>
<taxonomicName id="769BE6998B5B94E49FAA087F3D32F77F" class="Bivalvia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protobranchia" pageId="8" pageNumber="9" phylum="Mollusca" rank="class">Bivalvia</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="ecology">
<paragraph pageId="8" pageNumber="9">Ecology.</paragraph>
<paragraph pageId="8" pageNumber="9">Found in polymetallic nodule province.</paragraph>
<subSubSection id="B1D07D8C4336DCA18CDC6D31B13D0D33" pageId="8" pageNumber="9" type="ecology">
<paragraph id="2F20FB4A37BBB5694D4E6807306F2325" pageId="8" pageNumber="9">Ecology.</paragraph>
<paragraph id="B7ED524E87A9C6D39E38C2668BD881E5" pageId="8" pageNumber="9">Found in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="F1EDCE29FFA08FAFC76512E299CD85CE">Abyssal fauna of the UK- 1 polymetallic nodule exploration area, Clarion-Clipperton Zone, central Pacific Ocean: Mollusca</mods:title>
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<mods:namePart>Taylor, John D.</mods:namePart>
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<mods:namePart>Dahlgren, Thomas G.</mods:namePart>
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<mods:namePart>Todt, Christiane</mods:namePart>
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<mods:namePart>Ikebe, Chiho</mods:namePart>
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<mods:namePart>Rabone, Muriel</mods:namePart>
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<taxonomicName authority="Rokop, 1972" authorityName="Rokop" authorityYear="1972" class="Monoplacophora" family="Neopilinidae" genus="Veleropilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Veleropilina oligotropha" order="Pholadomyoida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="oligotropha">Veleropilina oligotropha (Rokop, 1972)</taxonomicName>
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<taxonomicName id="8BD11C45A3651BA827B7040EFC8E037F" authority="Rokop, 1972" authorityName="Rokop" authorityYear="1972" family="Neopilinidae" genus="Veleropilina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Veleropilina oligotropha" order="Pholadomyoida" pageId="19" pageNumber="20" rank="species" species="oligotropha">Veleropilina oligotropha (Rokop, 1972)</taxonomicName>
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<subSubSection pageId="19" pageNumber="20" type="materials_examined">
<paragraph pageId="19" pageNumber="20">Material examined.</paragraph>
<paragraph pageId="19" pageNumber="20">NHM_405 NHMUK 20170072, collected 2013-10-20, 13.86328 -116.54885, 4050 m. http://data.nhm.ac.uk/object/bf968b01-1991-43b7-87e4-25da4d5a9dc5</paragraph>
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<paragraph id="EC4842E08D75BD5DEFC2C9AF54A88EA9" pageId="19" pageNumber="20">Material examined.</paragraph>
<paragraph id="08D046C755E08B1496E0AFB0CA40D7F7" pageId="19" pageNumber="20">NHM_405 NHMUK 20170072, collected 2013-10-20, 13.86328 -116.54885, 4050 m. http://data.nhm.ac.uk/object/bf968b01-1991-43b7-87e4-25da4d5a9dc5</paragraph>
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<pageBreakToken pageId="20" pageNumber="21" start="start">Description</pageBreakToken>
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<pageBreakToken id="5162F3B43D4981812D320566AA578681" pageId="20" pageNumber="21" start="start">Description</pageBreakToken>
.
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<paragraph pageId="20" pageNumber="21">Shell transparent, sculpture is reticulate, reticulation not covering the smooth apical area. Voucher specimen NHM_405, specimen length 2.2 mm, specimen width 1 mm (Fig. 19).</paragraph>
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<paragraph pageId="20" pageNumber="21">
<paragraph id="FEF6A12EBC4A652CDBF824DA1E6BA6B9" pageId="20" pageNumber="21">Shell transparent, sculpture is reticulate, reticulation not covering the smooth apical area. Voucher specimen NHM_405, specimen length 2.2 mm, specimen width 1 mm (Fig. 19).</paragraph>
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<paragraph id="FFA7E8F074708FFEFE4664AA19EFA3D3" pageId="20" pageNumber="21">
Figure 19.
<taxonomicName class="Monoplacophora" family="Neopilinidae" genus="Veleropilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Veleropilina oligotropha" order="Tryblidiida" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oligotropha">Veleropilina oligotropha</taxonomicName>
<taxonomicName id="F6960534221FB68CA695C9C60786A918" class="Monoplacophora" family="Neopilinidae" genus="Veleropilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Veleropilina oligotropha" order="Neopilinida" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oligotropha">Veleropilina oligotropha</taxonomicName>
(Rokop, 1972) Specimen NHM_405. A Dorsal view of living specimen B Ventral view of living specimen. C Lateral view of ethanol-preserved specimen D Dorsal shell sculpture detail, just below apex E Ventral view of mouth and shell margin. Scale bars: 1mm. Image attribution Glover, Dahlgren &amp; Wiklund, 2017.
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<subSubSection pageId="20" pageNumber="21" type="genetic data">
<paragraph pageId="20" pageNumber="21">Genetic data.</paragraph>
<paragraph pageId="20" pageNumber="21">GenBank NHM_405 16S-MF157465, 18S-MF157495, COI-MF157522.</paragraph>
<subSubSection id="5097B39ACB6D5B109E05EDDEA199DA16" pageId="20" pageNumber="21" type="genetic data">
<paragraph id="F466024FD8C3E0ECBEF79867B88DB028" pageId="20" pageNumber="21">Genetic data.</paragraph>
<paragraph id="891C5749A999BBE4582E285FF3FADF0B" pageId="20" pageNumber="21">GenBank NHM_405 16S-MF157465, 18S-MF157495, COI-MF157522.</paragraph>
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<subSubSection pageId="20" pageNumber="21" type="remarks">
<paragraph pageId="20" pageNumber="21">Remarks.</paragraph>
<paragraph pageId="20" pageNumber="21">
<subSubSection id="A9EAF138E252D042F6B5F708C7DE8F08" pageId="20" pageNumber="21" type="remarks">
<paragraph id="A6E448703BCF0DB842B8BC65F8B37382" pageId="20" pageNumber="21">Remarks.</paragraph>
<paragraph id="44C84F6032D0EF8F5F02EA3BE4B13CD9" pageId="20" pageNumber="21">
Morphologically agrees with description of
<taxonomicName class="Monoplacophora" family="Neopilinidae" genus="Veleropilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Veleropilina oligotropha" order="Tryblidiida" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oligotropha">Veleropilina oligotropha</taxonomicName>
<taxonomicName id="327EE859854FA47872CA04F4D78B8268" class="Monoplacophora" family="Neopilinidae" genus="Veleropilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Veleropilina oligotropha" order="Neopilinida" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="oligotropha">Veleropilina oligotropha</taxonomicName>
(Rokop, 1972) described from ~6000 m water depth in the central North Pacific.
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Forms a unique monophyletic clade distinct from all other AB01 specimens. No genetic matches on GenBank. In the molecular analyses based on the 16S gene, the
<taxonomicName class="Monoplacophora" lsidName="Pteriomorphia" pageId="20" pageNumber="21" rank="class">Monoplacophora</taxonomicName>
<taxonomicName id="07ED7E2EA34DD3C011E90E048776AFD1" class="Monoplacophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" pageId="20" pageNumber="21" phylum="Mollusca" rank="class">Monoplacophora</taxonomicName>
clade is strongly supported, but internal branches are unresolved or, when clades are present, they have low support (Fig. 20).
</paragraph>
<caption pageId="20" pageNumber="21">
<paragraph pageId="20" pageNumber="21">
<caption id="A71F3AF16D81CB7BF783C13590B3A793" pageId="20" pageNumber="21">
<paragraph id="E86766EE8F484FEA1601A7899378D0FF" pageId="20" pageNumber="21">
Figure 20. Phylogenetic analysis of
<taxonomicName class="Monoplacophora" lsidName="Pteriomorphia" pageId="20" pageNumber="21" rank="class">Monoplacophora</taxonomicName>
<taxonomicName id="9D0418E756D1B38D6E6247950944D5C5" class="Monoplacophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" pageId="20" pageNumber="21" phylum="Mollusca" rank="class">Monoplacophora</taxonomicName>
. 50% majority rule consensus tree from the Bayesian analyses using 16S. Asterisks denotes support values of 95 or above.
</paragraph>
</caption>
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<subSubSection pageId="20" pageNumber="21" type="ecology">
<paragraph pageId="20" pageNumber="21">Ecology.</paragraph>
<paragraph pageId="20" pageNumber="21">
<subSubSection id="70D62BE8C6DD5E4AF3D7E49C70384546" pageId="20" pageNumber="21" type="ecology">
<paragraph id="49A5AAFD4F599ACD4A4F3DEAE77918BF" pageId="20" pageNumber="21">Ecology.</paragraph>
<paragraph id="D8A9B3C44AA510B99D1A9AD807504852" pageId="20" pageNumber="21">
Specimen collected from an epibenthic sledge tow across region of sediment and polymetallic nodules.
<bibRefCitation author="Rokop, FJ" journalOrPublisher="Veliger" pageId="30" pageNumber="31" pagination="91 - 95" title="A new species of monoplacophoran from the abyssal north Pacific." volume="15" year="1972">Rokop (1972)</bibRefCitation>
<bibRefCitation id="C351B9567CDC8337220367211D3BB9BF" author="Rokop, FJ" journalOrPublisher="Veliger" pageId="30" pageNumber="31" pagination="91 - 95" title="A new species of monoplacophoran from the abyssal north Pacific." volume="15" year="1972">Rokop (1972)</bibRefCitation>
did not observe the species directly on nodules, they were just recovered from the epibenthic sledge sample, as was the case in this study. The importance of the nodules as a habitat for the species remains uncertain until they are directly observed live on the seafloor.
</paragraph>
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Figures 4, 19, 20, 21
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Dohrn, 1881
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<paragraph pageId="2" pageNumber="165">ZSMA20071505: female; Bayuls-sur-Mer, France; 07.2004. ZSMA20071506: female; Bayuls-sur-Mer, France; 07.2004. ZSMA20071507: male; Bayuls-sur-Mer, France; 07.2004. ZSMA20071508: male; Bayuls-sur-Mer, France; 07.2004.</paragraph>
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<paragraph id="9FCC52015443AEA20F97E4CFDC8AEEAA" pageId="2" pageNumber="165">ZSMA20071505: female; Bayuls-sur-Mer, France; 07.2004. ZSMA20071506: female; Bayuls-sur-Mer, France; 07.2004. ZSMA20071507: male; Bayuls-sur-Mer, France; 07.2004. ZSMA20071508: male; Bayuls-sur-Mer, France; 07.2004.</paragraph>
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<taxonomicName authority="Dohrn, 1881" authorityName="Dohrn" authorityYear="1881" class="Pycnogonida" family="Ammotheidae" genus="Ammothella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammothella uniunguiculata" order="Pantopoda" pageId="2" pageNumber="165" phylum="Arthropoda" rank="species" species="uniunguiculata">Ammothella uniunguiculata (Dohrn, 1881)</taxonomicName>
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Figures 4, 18
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Dohrn, 1881
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<paragraph pageId="3" pageNumber="166">Taxon classification Animalia Pantopoda Endeidae</paragraph>
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<taxonomicName authority="Montagu, 1808" authorityName="Montagu" authorityYear="1808" class="Pycnogonida" family="Endeidae" genus="Endeis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endeis spinosa" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="spinosa">Endeis spinosa (Montagu, 1808)</taxonomicName>
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Figures 38, 41, 42, 43, 44
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<taxonomicName authority="Stebbing, 1902" authorityName="Stebbing" authorityYear="1902" class="Pycnogonida" family="Endeidae" genus="Chilophoxus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chilophoxus spinosus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="spinosus">Chilophoxus spinosus</taxonomicName>
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<taxonomicName id="C6F7986269936EAFFCF16CCC2410A553" class="Pycnogonida" family="Endeidae" genus="Chilophoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chilophoxus spinosus" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="spinosus">Chilophoxus spinosus</taxonomicName>
Montagu
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<taxonomicName class="Pycnogonida" family="Endeidae" genus="Endeis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endeis gracilis" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="gracilis">Endeis gracilis</taxonomicName>
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<taxonomicName id="1F3638324B6E3A53341C5EF507CBFD39" class="Pycnogonida" family="Endeidae" genus="Endeis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endeis gracilis" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="gracilis">Endeis gracilis</taxonomicName>
Philippi, 1843
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<taxonomicName class="Pycnogonida" family="Endeidae" genus="Endeis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endeis laevis" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="laevis">Endeis laevis</taxonomicName>
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<taxonomicName id="9B107CB50D91D318CAC1CEFE2D88CB2B" class="Pycnogonida" family="Endeidae" genus="Endeis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endeis laevis" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="laevis">Endeis laevis</taxonomicName>
(Grube, 1871)
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<taxonomicName class="Pycnogonida" family="Endeidae" genus="Endeis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endeis vulgaris" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="vulgaris">Endeis vulgaris</taxonomicName>
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<taxonomicName id="C39BC73052F063C8BAEF3E3C11565458" class="Pycnogonida" family="Endeidae" genus="Endeis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endeis vulgaris" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="vulgaris">Endeis vulgaris</taxonomicName>
(Dohrn, 1881)
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<taxonomicName genus="Phallangium" lsidName="Phallangium spinosum" pageId="3" pageNumber="166" rank="species" species="spinosum">Phallangium spinosum</taxonomicName>
<paragraph id="CED48BAB5294328CE09126705D990FBE" pageId="3" pageNumber="166">
<taxonomicName id="3ABAC8D0AC125379C94D38F823207DC5" genus="Phallangium" lsidName="Phallangium spinosum" pageId="3" pageNumber="166" rank="species" species="spinosum">Phallangium spinosum</taxonomicName>
Montagu, 1808
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<paragraph pageId="3" pageNumber="166">
<taxonomicName authority="Latreille, 1816" authorityName="Latreille" authorityYear="1816" class="Pycnogonida" family="Endeidae" genus="Phoxichilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phoxichilus vulgaris" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="vulgaris">Phoxichilus vulgaris</taxonomicName>
<paragraph id="B2BD08BEA0FF766B0A120B4744CA9298" pageId="3" pageNumber="166">
<taxonomicName id="A56416EEE551DDD0775B32AB9CEA2C8D" class="Pycnogonida" family="Endeidae" genus="Phoxichilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phoxichilus vulgaris" order="Pantopoda" pageId="3" pageNumber="166" phylum="Arthropoda" rank="species" species="vulgaris">Phoxichilus vulgaris</taxonomicName>
Dohrn, 1881
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<paragraph pageId="3" pageNumber="166">Material.</paragraph>
<paragraph pageId="3" pageNumber="166">
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<paragraph id="D5DDFF4C951EEB4035B2A2419919E623" pageId="3" pageNumber="166">Material.</paragraph>
<paragraph id="63086A7D5F821F0DE52EA976F216932A" pageId="3" pageNumber="166">
ZSMA20071498: female; Rovinj, Croatia; 04.2004. ZSMA20071499: male; Rovinj, Croatia; 05.05.2007; 0-2 m;
<taxonomicName family="Corallinaceae" genus="Corallina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corallina elongata" order="Cryptonemiales" pageId="3" pageNumber="166" phylum="Rhodophyta" rank="species" species="elongata">Corallina elongata</taxonomicName>
<taxonomicName id="41376D39CE30532EC95701414698BFE5" class="Polychaeta" family="Sabellidae" genus="Corallina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corallina elongata" order="Sabellida" pageId="3" pageNumber="166" phylum="Annelida" rank="species" species="elongata">Corallina elongata</taxonomicName>
J.Ellis &amp; Solander, 1786.
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<subSubSection pageId="3" pageNumber="166" type="remarks">
<paragraph pageId="3" pageNumber="166">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="166">
<subSubSection id="2D2933DC26CEDEA074F1FF5673133C84" pageId="3" pageNumber="166" type="remarks">
<paragraph id="4704DFC22D40848DAB2602F8C4A2743D" pageId="3" pageNumber="166">Remarks.</paragraph>
<paragraph id="1052FDCFF5880CF8F39263AEBD81B549" pageId="3" pageNumber="166">
According to
<bibRefCitation author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
<bibRefCitation id="71672967BA0562C23E933987297C32AA" author="Dohrn, A" journalOrPublisher="Monographie der Fauna und Flora des golfes von Neapel" pageId="5" pageNumber="168" pagination="1 - 252" title="Die Pantopoden des Golfes von Neapel und der angrenzenden Meeresabschnitte." volume="3" year="1881">Dohrn (1881)</bibRefCitation>
and
<bibRefCitation author="Bamber, RN" journalOrPublisher="Linnean Society of London by Field Studies Council, London, Shrewsbury" pageId="5" pageNumber="168" title="Sea-spiders (Pycnogonida) of the north-east Atlantic: Keys and Notes for Identification the Species." year="2010">Bamber (2010)</bibRefCitation>
<bibRefCitation id="8D53827212E59E153462B0F3AC7CC40A" author="Bamber, RN" journalOrPublisher="Linnean Society of London by Field Studies Council, London, Shrewsbury" pageId="5" pageNumber="168" title="Sea-spiders (Pycnogonida) of the north-east Atlantic: Keys and Notes for Identification the Species." year="2010">Bamber (2010)</bibRefCitation>
each lateral process and coxa 1 is armed with one protuberance. Here, in the males the lateral process 1 and the coxa 1 of the first leg has two protuberances (Fig. 42D, F), the remaining legs correspond with previous observations (e.g. Fig. 42E, G). The females are without such protuberances (Fig. 44C, D).
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Figures 27, 36
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emaciata spectrum
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(Dohrn, 1881)
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Dohrn, 1881
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<paragraph pageId="3" pageNumber="166">Material.</paragraph>
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ZSMA20140004: male; Banjole, Rovinj, Croatia; 07.1968; 5 m; on
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<taxonomicName id="CD74C6C4BC9DC328C56DB17B98FB9088" class="Phaeophyceae" family="Sargassaceae" genus="Cystoseira" higherTaxonomySource="CoL" kingdom="Chromista" lsidName="Cystoseira" order="Fucales" pageId="3" pageNumber="166" phylum="Ochrophyta" rank="genus">Cystoseira</taxonomicName>
sp..
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Figures 53, 55
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<paragraph pageId="4" pageNumber="167">ZSMA20071504: male; Brucoli, Sicily, Italy; 14.09.1988.</paragraph>
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<taxonomicName authority="Knudsen, 1970" authorityName="Knudsen" authorityYear="1970" class="Bivalvia" family="Mytilidae" genus="Dacrydium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dacrydium panamensis" order="Pholadomyoida" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="panamensis">Dacrydium panamensis Knudsen, 1970</taxonomicName>
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<subSubSection pageId="15" pageNumber="16" type="materials_examined">
<paragraph pageId="15" pageNumber="16">Material examined.</paragraph>
<paragraph pageId="15" pageNumber="16">NHM_117 NHMUK 20170067, collected 2013-10-11, 13.79335 -116.70308, 4081 m. http://data.nhm.ac.uk/object/180e485f-f1c2-41e1-b858-f02ba537804b</paragraph>
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<paragraph id="443CC0428B0A348307A3BD7A9831DF42" pageId="15" pageNumber="16">Material examined.</paragraph>
<paragraph id="CB1D5634755EFEBDDF04C0B915F9CA46" pageId="15" pageNumber="16">NHM_117 NHMUK 20170067, collected 2013-10-11, 13.79335 -116.70308, 4081 m. http://data.nhm.ac.uk/object/180e485f-f1c2-41e1-b858-f02ba537804b</paragraph>
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<paragraph pageId="15" pageNumber="16">Description.</paragraph>
<paragraph pageId="15" pageNumber="16">Shell small, subovate, translucent, anterior-ventral margin slightly produced, highest point near mid-line. Voucher NHM_117 Shell length 1.7 mm, shell height 2.5 mm (Fig. 14).</paragraph>
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<paragraph id="A18D8C38A670B8B629DEEB1DB829E170" pageId="15" pageNumber="16">Description.</paragraph>
<paragraph id="047370DC480132CF8A4FD66FFD813AC0" pageId="15" pageNumber="16">Shell small, subovate, translucent, anterior-ventral margin slightly produced, highest point near mid-line. Voucher NHM_117 Shell length 1.7 mm, shell height 2.5 mm (Fig. 14).</paragraph>
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Figure 14.
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Knudsen, 1970 Specimen NHM_117. Scale bar: 0.5 mm. Image attribution Glover, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
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<paragraph pageId="15" pageNumber="16">Genetic data.</paragraph>
<paragraph pageId="15" pageNumber="16">GenBank NHM_117 18S-MF157471.</paragraph>
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<paragraph id="C8635E48E822B5791EDAE40B0688CE98" pageId="15" pageNumber="16">Genetic data.</paragraph>
<paragraph id="4E8DAA6710334CFA557C744FC194493D" pageId="15" pageNumber="16">GenBank NHM_117 18S-MF157471.</paragraph>
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<paragraph pageId="15" pageNumber="16">Remarks.</paragraph>
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<paragraph id="3530ED0E5BDABC01852563D41A242441" pageId="15" pageNumber="16">Remarks.</paragraph>
<paragraph id="243B4BE118947BFFB880E5C50FC7AA02" pageId="15" pageNumber="16">
Identified from figures in
<bibRefCitation author="Knudsen, J" journalOrPublisher="Galathea Reports" pageId="30" pageNumber="31" pagination="1 - 241" title="The systematics and biology of abyssal and hadal Bivalvia." volume="11" year="1970">Knudsen (1970)</bibRefCitation>
<bibRefCitation id="941C054EEDC304BF69A066F22BF52F2C" author="Knudsen, J" journalOrPublisher="Galathea Reports" pageId="30" pageNumber="31" pagination="1 - 241" title="The systematics and biology of abyssal and hadal Bivalvia." volume="11" year="1970">Knudsen (1970)</bibRefCitation>
and Coan &amp; Valentich-Scott (2012). The holotype of
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was collected on the Galathea expedition (stn 726) at 3670-3270 m depth in Gulf of Panama. In the molecular analysis (Fig. 17) it aligns as a sister species to many shallow water
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. No genetic matches on GenBank.
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<paragraph pageId="15" pageNumber="16">Ecology.</paragraph>
<paragraph pageId="15" pageNumber="16">Found in polymetallic nodule province.</paragraph>
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<paragraph id="E3FF1BBC1872153DF0A259E09D8C0996" pageId="15" pageNumber="16">Ecology.</paragraph>
<paragraph id="A4DD4C2AB8EF4C510D2418D019C3E6A9" pageId="15" pageNumber="16">Found in polymetallic nodule province.</paragraph>
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<mods:url>http://dx.doi.org/10.3897/zookeys.707.13042</mods:url>
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<mods:url id="4BD811992006F26DB3775251F16C95DB">http://dx.doi.org/10.3897/zookeys.707.13042</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.707.13042</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-707-1</mods:identifier>
<mods:identifier type="ZBK">6A0BCD992749450C874B440D0C0A232C</mods:identifier>
<mods:identifier type="ZooBank">6A0BCD992749450C874B440D0C0A232C</mods:identifier>
<mods:classification id="53E410C79F0D40536F0A8CE669DD89A8">journal article</mods:classification>
<mods:identifier id="9D0BF6356F2011F363F587945905BACD" type="DOI">http://dx.doi.org/10.3897/zookeys.707.13042</mods:identifier>
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<treatment ID-GBIF-Taxon="135489983" LSID="urn:lsid:plazi:treatment:FC3B76110BDF6C6AF88EE68461190B54" httpUri="http://treatment.plazi.org/id/FC3B76110BDF6C6AF88EE68461190B54" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection pageId="17" pageNumber="18" type="nomenclature">
<paragraph pageId="17" pageNumber="18">
<taxonomicName authority="sp. (NHM _ 105)" class="Bivalvia" family="Propeamussiidae" genus="Catillopecten" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catillopecten" order="Pholadomyoida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Catillopecten sp. (NHM_105)</taxonomicName>
<treatment id="FC3B76110BDF6C6AF88EE68461190B54" ID-GBIF-Taxon="135489983" LSID="urn:lsid:plazi:treatment:FC3B76110BDF6C6AF88EE68461190B54" httpUri="http://treatment.plazi.org/id/FC3B76110BDF6C6AF88EE68461190B54" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" scope_class="Bivalvia" scope_family="Propeamussiidae" scope_order="Pectinoida" scope_phylum="Mollusca" scope_subClass="Pteriomorphia">
<subSubSection id="FBF5A78203F74694F7823A80DB3FB925" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="272B38264C1FE25D3174986F41EB7DFB" pageId="17" pageNumber="18">
<taxonomicName id="1E46F51B7A9D0DDF8D331F7CB4F77FD1" authority="sp. (NHM _ 105)" family="Propeamussiidae" genus="Catillopecten" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Catillopecten" order="Pholadomyoida" pageId="17" pageNumber="18" rank="genus">Catillopecten sp. (NHM_105)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="18" type="materials_examined">
<paragraph pageId="17" pageNumber="18">Material examined.</paragraph>
<paragraph pageId="17" pageNumber="18">NHM_105 NHMUK 20170070, collected 2013-10-11, 13.79335 -116.70308, 4081 m. http://data.nhm.ac.uk/object/24f5c5bb-e419-48ef-baaa-4a6493f691d9</paragraph>
<subSubSection id="16CB03547C3C511AAD1C4148386E7FAD" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="E1EFA560FC5189CD2853E8821A3DF287" pageId="17" pageNumber="18">Material examined.</paragraph>
<paragraph id="EC375156D0BB48C9E52A95C3EDA3D802" pageId="17" pageNumber="18">NHM_105 NHMUK 20170070, collected 2013-10-11, 13.79335 -116.70308, 4081 m. http://data.nhm.ac.uk/object/24f5c5bb-e419-48ef-baaa-4a6493f691d9</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="18" type="description">
<paragraph pageId="17" pageNumber="18">Description.</paragraph>
<paragraph pageId="17" pageNumber="18">Small, thin-shelled, subcircular. Right valve flat, left valve slightly convex. Both valves with commarginal undulations that become stronger towards the margin, fine radial striations on both valves. Well defined anterior auricle and byssal notch. Voucher NHM_105 1.8 mm shell length, height 1.5 mm (Fig. 16).</paragraph>
<caption pageId="17" pageNumber="18">
<paragraph pageId="17" pageNumber="18">
<subSubSection id="9FB7FC604BAFD75F733E42BF7E89F222" pageId="17" pageNumber="18" type="description">
<paragraph id="982481CE91B12069C1097194E7420C93" pageId="17" pageNumber="18">Description.</paragraph>
<paragraph id="B249ADF3B49522783DBEEE610DE374FF" pageId="17" pageNumber="18">Small, thin-shelled, subcircular. Right valve flat, left valve slightly convex. Both valves with commarginal undulations that become stronger towards the margin, fine radial striations on both valves. Well defined anterior auricle and byssal notch. Voucher NHM_105 1.8 mm shell length, height 1.5 mm (Fig. 16).</paragraph>
<caption id="ADA5C35449215BE74DD05BDC0634D02E" pageId="17" pageNumber="18">
<paragraph id="D268FFF7FF081BF333D81E94DE2588BA" pageId="17" pageNumber="18">
Figure 16.
<taxonomicName class="Bivalvia" family="Propeamussiidae" genus="Catillopecten" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catillopecten" order="Pectinida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Catillopecten</taxonomicName>
<taxonomicName id="37B10852BEC35569469AB276A58C6373" class="Bivalvia" family="Propeamussiidae" genus="Catillopecten" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catillopecten" order="Pectinida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Catillopecten</taxonomicName>
sp. (NHM_105) live after recovery. Scale bar: 1 mm. Image attribution Glover, Dahlgren &amp; Wiklund, 2017.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="17" pageNumber="18" type="genetic data">
<paragraph pageId="17" pageNumber="18">Genetic data.</paragraph>
<paragraph pageId="17" pageNumber="18">GenBank NHM_105 18S-MF157469.</paragraph>
<subSubSection id="21FA868457C0482E9737456ACCA1C9F5" pageId="17" pageNumber="18" type="genetic data">
<paragraph id="1F01B944A4C5D200B3591B0F245C51C1" pageId="17" pageNumber="18">Genetic data.</paragraph>
<paragraph id="327B462431F90A1ECDACF43E45BBDE7B" pageId="17" pageNumber="18">GenBank NHM_105 18S-MF157469.</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="remarks">
<paragraph pageId="17" pageNumber="18">Remarks.</paragraph>
<paragraph lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="A0F40558C9212BC51E607D4C693C498A" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="remarks">
<paragraph id="F7897B49A13F5950F8851197CF840EC0" pageId="17" pageNumber="18">Remarks.</paragraph>
<paragraph id="D78B8E377B90295854E277450FBEEEB1" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Holotype (ZMUC) from Gulf of Panama, 3270-3670 m Galathea stn 726, figured by
<bibRefCitation author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott (2012</bibRefCitation>
<bibRefCitation id="26E2384D2D82128B16243FDDEF792039" author="Coan, EV" journalOrPublisher="Santa Barbara Museum of Natural History Monographs" pageId="29" pageNumber="30" pagination="1 - 1258" title="Bivalve seashells of tropical west America: Marine bivalves from Baja California to northern Peru." volume="6" year="2012">Coan and Valentich-Scott (2012</bibRefCitation>
pl. 100). In the molecular tree it
<pageBreakToken pageId="18" pageNumber="19" start="start">groups</pageBreakToken>
<pageBreakToken id="5E32170EE0B6A717DF3ECED02D05221D" pageId="18" pageNumber="19" start="start">groups</pageBreakToken>
with two other species of
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<taxonomicName id="EDC4F92D9204D0C6951846DAE55D7DD4" family="Propeamussidae" lsidName="" pageId="18" pageNumber="19" rank="family">Propeamussidae</taxonomicName>
on a long branch and distinct from other
<taxonomicName lsidName="Pteriomorphia" order="Pectinoida" pageId="18" pageNumber="19" rank="order">Pectinoida</taxonomicName>
<taxonomicName id="B1FC87E93D8A75AC656A625F3FAF8EBC" lsidName="Pteriomorphia" order="Pectinoida" pageId="18" pageNumber="19" rank="order">Pectinoida</taxonomicName>
, but a GenBank species (VLG_2013) identified as
<taxonomicName class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Propeamussium" order="Pectinida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">Propeamussium</taxonomicName>
<taxonomicName id="C3F7B9ECD38F1B1D1AF089E291139618" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Propeamussium" order="Pectinida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">Propeamussium</taxonomicName>
sp. is distinct from these (Fig. 17). Henk H. Dijkstra (Naturalis Biodiversity Center in Leiden, Netherlands) advised on identification of this species. Forms a unique monophyletic clade distinct from all other AB01 specimens. No genetic matches on GenBank.
</paragraph>
<caption pageId="18" pageNumber="19">
<paragraph pageId="18" pageNumber="19">
<caption id="81D786672E77EDB2A0A33127C6CD9BD1" pageId="18" pageNumber="19">
<paragraph id="A056CBF52DF6C99D2E34A55E505E4AB5" pageId="18" pageNumber="19">
Figure 17. Phylogenetic analysis of
<taxonomicName class="Bivalvia" lsidName="Pteriomorphia" pageId="18" pageNumber="19" rank="class">Bivalvia</taxonomicName>
<taxonomicName id="6B84F5B862BF703BFCEB01DAA02EDE51" class="Bivalvia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteriomorphia" pageId="18" pageNumber="19" phylum="Mollusca" rank="class">Bivalvia</taxonomicName>
:
<taxonomicName lsidName="Pteriomorphia" order="Pteriomorpha" pageId="18" pageNumber="19" rank="order">Pteriomorpha</taxonomicName>
<taxonomicName id="A7BC69C8432EA846A6F60E35C7084D44" lsidName="Pteriomorphia" order="Pteriomorpha" pageId="18" pageNumber="19" rank="order">Pteriomorpha</taxonomicName>
. 50% majority rule consensus tree from the Bayesian analyses using 18S and COI. Asterisks denotes support values of 95 or above.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="ecology">
<paragraph pageId="18" pageNumber="19">Ecology.</paragraph>
<paragraph pageId="18" pageNumber="19">Found in polymetallic nodule province.</paragraph>
<subSubSection id="7A27BF70E60DB39BED07FCA4091D8D5F" pageId="18" pageNumber="19" type="ecology">
<paragraph id="BAB823BA9AA128BCDDEB04A23525B477" pageId="18" pageNumber="19">Ecology.</paragraph>
<paragraph id="2A69C8DDAB5046C4EBCAAC633297051D" pageId="18" pageNumber="19">Found in polymetallic nodule province.</paragraph>
</subSubSection>
</treatment>
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