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<document id="DDD063033C86A7BAC2F1CE1811D84593" ID-DOI="10.3897/zookeys.1218.132275" ID-ZooBank="85DC6275-A17C-4294-AAA8-9BF36A197A1B" ID-publisher-id="132275" URI-arpha="DCCCAA62-16AC-5E63-A3C4-DBEF580E0D4A" URI-zoobank="http://zoobank.org/85DC6275-A17C-4294-AAA8-9BF36A197A1B" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1732298777919" checkinUser="pensoft" docAuthor="Rázuri-Gonzales, Ernesto, Graf, Wolfram, Heckenhauer, Jacqueline, Schneider, Julio V. & Pauls, Steffen U." docDate="2024" docId="1717976C21BB506E8E9FB1D1BFE1B61E" docLanguage="en" docName="ZooKeys 1218: 295-314" docOrigin="ZooKeys 1218" docSource="https://zookeys.pensoft.net/article/132275/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Rhycophila lignumvallis Graf & Razuri-Gonzales 2024, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="314" masterDocId="DCCCAA6216AC5E63A3C4DBEF580E0D4A" masterDocTitle="A new species of Rhyacophila Pictet, 1834 (Trichoptera, Rhyacophilidae) from Corsica with the genomic characterization of the holotype" masterLastPageNumber="314" masterPageNumber="295" pageNumber="295" updateTime="1732303793471" updateUser="admin">
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<mods:titleInfo id="D9C635CE731D7FCB8637D3613FFC3BE4">
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<mods:title id="F64750DD678F2E1F247C191E9720CD4D">A new species of Rhyacophila Pictet, 1834 (Trichoptera, Rhyacophilidae) from Corsica with the genomic characterization of the holotype</mods:title>
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<mods:name id="A2DBA5928A2FE3C20BBFFD8F6807C24C" type="personal">
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<mods:roleTerm id="0332106EE4EC18BF9F4D0E0F3ADE1AC1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B78BC8204932655234734E7EBA831EC4">Rázuri-Gonzales, Ernesto</mods:namePart>
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<mods:nameIdentifier id="624EAAC8C58ADE068A1ABD80F3920836" type="ORCID">0000-0002-7554-0816</mods:nameIdentifier>
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<mods:affiliation id="708B82B85642D25E1312C32B3857EADB">Senckenberg Research Institute and Natural History Museum Frankfurt, Frankfurt, Germany</mods:affiliation>
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<mods:roleTerm id="09D61EABCD35FEF46D5667C7D845206D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BD3D6430F0F857189797356E54C4106A">Graf, Wolfram</mods:namePart>
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<mods:nameIdentifier id="78CC8A608D5212E9988E9FD927358912" type="ORCID">0000-0001-6559-0644</mods:nameIdentifier>
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<mods:affiliation id="DCC1901880750D431407AC9F5646D767">Institute of Hydrobiology and Aquatic Ecosystem Management (IHG), University of Natural Resources and Life Sciences, Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:name id="D8653FC138B84A335A6AABD54D8B72AC" type="personal">
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</mods:role>
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<mods:namePart id="857871C0F7DD6916BDFB4A6968BD83CD">Heckenhauer, Jacqueline</mods:namePart>
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<mods:nameIdentifier id="3A6B8C5B9D0655FA99418697690C9842" type="ORCID">0000-0001-8771-9154</mods:nameIdentifier>
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<mods:affiliation id="DCBAF299E79BE2101AB6C74096BCF952">Senckenberg Research Institute and Natural History Museum Frankfurt, Frankfurt, Germany & LOEWE Centre for Translational Biodiversity Genomics (LOEWE-TBG), Frankfurt, Germany</mods:affiliation>
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</mods:role>
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<mods:namePart id="EAB11252F43F53E656A7B7396772BD68">Schneider, Julio V.</mods:namePart>
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<mods:nameIdentifier id="A9EB5EF7DCA2E06254C068DBCF94DE6A" type="ORCID">0000-0002-9823-6569</mods:nameIdentifier>
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<mods:affiliation id="B2C8E662831AD37A443BD7FBCEAB9385">Senckenberg Research Institute and Natural History Museum Frankfurt, Frankfurt, Germany</mods:affiliation>
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<mods:namePart id="3DB4CA28A3BBFE9E80AE658BB51B55DD">Pauls, Steffen U.</mods:namePart>
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<mods:nameIdentifier id="E80846576424878F2F870CF95A58BC93" type="ORCID">0000-0002-6451-3425</mods:nameIdentifier>
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<mods:affiliation id="1F2CB1214654C391C1E52D90CFB94A22">Senckenberg Research Institute and Natural History Museum Frankfurt, Frankfurt, Germany & LOEWE Centre for Translational Biodiversity Genomics (LOEWE-TBG), Frankfurt, Germany & Institute of Insect Biotechnology, Justus-Liebig University, Gießen, Germany</mods:affiliation>
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<mods:title id="32ABFAF3468FC52626BB6CCFC9871499">ZooKeys</mods:title>
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<mods:part id="7B412181CE25C99006B7978886ED8EA9">
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<mods:date id="5561ED19DB3847D4F5D7088A2E8DAA9D">2024</mods:date>
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<mods:number id="CD8DB9EAAE2FAED11B023FF408644DF1">2024-11-22</mods:number>
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<mods:number id="A808FC9E41875CC96EC76B72FB4D97EB">1218</mods:number>
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<mods:start id="0C531F414C54907DF4C06B91ECEAC650">295</mods:start>
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<mods:classification id="AA1E2240B46FA39EF59AFCE61C2DDB10">journal article</mods:classification>
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<mods:identifier id="EC355521F1C3ED40AD9A21392DB38455" type="DOI">10.3897/zookeys.1218.132275</mods:identifier>
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<mods:identifier id="CA4B9B22C161F5FBF0EB026060B5598F" type="ZooBank">85DC6275-A17C-4294-AAA8-9BF36A197A1B</mods:identifier>
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</mods:mods>
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<treatment id="1717976C21BB506E8E9FB1D1BFE1B61E" ID-arpha="1717976C-21BB-506E-8E9F-B1D1BFE1B61E" ID-zoobank="https://zoobank.org/5A58EBFA-E945-4032-917A-877444F5DA2B" LSID="urn:lsid:plazi:treatment:1717976C21BB506E8E9FB1D1BFE1B61E" LSID-ZBK="urn:lsid:zoobank.org:act:5A58EBFA-E945-4032-917A-877444F5DA2B" httpUri="http://treatment.plazi.org/id/1717976C21BB506E8E9FB1D1BFE1B61E" scope_family="Rhyacophilidae" scope_order="Trichoptera">
|
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<subSubSection id="45FD3BC2EC845C741AE8F22FEBD0E731" type="nomenclature">
|
||||
<paragraph id="7C1A568D6F33C436CA88F7BB54D12014">
|
||||
<taxonomicName id="A92F1F5924A2CA37845FE11C5CEE0E6F" authority="Graf & Razuri-Gonzales" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhycophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
|
||||
<emphasis id="4811E0685FB7B62805A2933DCD1ED325" italics="true">Rhycophila lignumvallis</emphasis>
|
||||
Graf & Rázuri-Gonzales
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="5CE242521DFF3E5F78BB36B9369C0AEC" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EHYAI" type="Holotype">
|
||||
<paragraph id="68850CF13A7625D849537614D1EEEDA5">
|
||||
<heading id="8668130057F1F40BE1014D3156630A52" reason="title">
|
||||
<typeStatus id="B10E445EED57525395DA36DE0E5A0CB2" type="holotype">Holotype</typeStatus>
|
||||
.
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||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="B0194E4518326E6B52D553AD345EC883">
|
||||
<materialsCitation id="6099A0D9A561D37B582094F85043B8F0" collectingDate="2019-07-25" collectionCode="SMF" collectorName="W. Graf" country="France" elevation="195" latitude="42.2275" location="bridge over the river Vecchio near the confluence with the river Tavignano" longLatPrecision="5" longitude="9.24306" municipality="river Tavignano" specimenCode="SMFTRI 00018634" specimenCount="1" specimenCount-male="1" stateProvince="Corsica" typeStatus="holotype">
|
||||
<collectingCountry id="8C03CBF88437C3244A6E2AB1FFC11547" name="France">France</collectingCountry>
|
||||
•
|
||||
<specimenCount id="76C5F417F6BC88FC9A5128D0B95930A0" count="1" type="male">♂</specimenCount>
|
||||
;
|
||||
<collectingRegion id="5C4589064F71FC33BDFB7ECDD3682D5C">Corsica</collectingRegion>
|
||||
,
|
||||
<location id="D119E089DCE516AB54AB23868AEB838A" LSID="urn:lsid:plazi:treatment:1717976C21BB506E8E9FB1D1BFE1B61E:D119E089DCE516AB54AB23868AEB838A" country="France" latitude="42.2275" longLatPrecision="5" longitude="9.24306" municipality="river Tavignano" name="bridge over the river Vecchio near the confluence with the river Tavignano" stateProvince="Corsica">bridge over the river Vecchio near the confluence with the river Tavignano</location>
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||||
;
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQYAI" specific-use="{"type":"Point","coordinates":[9.243060,42.227500]}">
|
||||
<geoCoordinate id="07AC4882D44A0A1BC0A3B012D532ACCF" degrees="42.2275" direction="north" orientation="latitude" precision="5" value="42.2275">42.2275 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="75F316028477760F8F18D16BFB8BA4C0" degrees="9.24306" direction="east" orientation="longitude" precision="1" value="9.24306">9.24306 ° E</geoCoordinate>
|
||||
</named-content>
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||||
;
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<elevation id="B1D9B6AEB03972398DE7AF120666D462" metricMagnitude="2" metricUnit="m" metricValue="1.95" unit="m" value="195.0">
|
||||
<quantity id="990EBC94AEA724BF43C03626FA8150B3" metricMagnitude="2" metricUnit="m" metricValue="1.95" unit="m" value="195.0">195 m</quantity>
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a. s. l.
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</elevation>
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;
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<date id="F4BAC052B39B4606A4ADD0E6356BBD8C" value="2019-07-25">
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<collectingDate id="228531F744E94B9E990FEA5AEEE69FB4" value="2019-07-25">25 Jul. 2019</collectingDate>
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</date>
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; col.
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<collectorName id="386ACC4F2F8C92126DBBB234D4EE577D">W. Graf</collectorName>
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leg.; in ethanol;
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<collectionCode id="3ED30E7699F528F62156A2B7FF2EE945">SMF</collectionCode>
|
||||
(
|
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<specimenCode id="CABFCC6465BBF361E71FCC7542B79B68" collectionCode="SMFTRI">SMFTRI 00018634</specimenCode>
|
||||
).
|
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</materialsCitation>
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||||
</paragraph>
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||||
</subSubSection>
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||||
<subSubSection id="SECID0EVYAI" type="diagnosis">
|
||||
<paragraph id="668AF9BF1C59208719229203833D40E0">
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<heading id="5B2D8DDDFE0858ABFA763053C62777B4" reason="title">Diagnosis.</heading>
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||||
</paragraph>
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||||
<paragraph id="AFED66C8F48A58E5C4828B0C03D1F9FC">
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||||
<taxonomicName id="128F429C84E3D81E3D058F50DAE2CF64" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
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<emphasis id="9221F759BCF8DFE940198BE118814B02" italics="true">Rhyacophila lignumvallis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A3E9A73AED803DF44048CF5D03B34AFD" rank="species">sp. nov.</taxonomicNameLabel>
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(Figs
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<figureCitation id="890B31C655EDDAFD88FC47471F19D5D1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4</figureCitation>
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,
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<figureCitation id="F05AB6F6CAC5CC93AEBD09ADE564C357" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Rhyacophila lignumvallis sp. nov., male genitalia, dorsal view (A), lateral view (B). R. pubescens, male genitalia, dorsal view (C), lateral view (D). R. tsurakiana, male genitalia, dorsal view (E), lateral view (F). Scale bars: 100 μm. Specimens were photographed by W. Graf." figureDoi="10.3897/zookeys.1218.132275.figure5" httpUri="https://binary.pensoft.net/fig/1180557">5 A, B</figureCitation>
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||||
) belongs to the
|
||||
<taxonomicName id="FA9407EECEDB7C60F8AF7EB64958D0EE" authorityName="Pictet" authorityYear="1834" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tristis">
|
||||
<emphasis id="58CD8BE0692E330F9D945BFE0E7B7B2A" italics="true">Rhyacophila tristis</emphasis>
|
||||
</taxonomicName>
|
||||
species group (
|
||||
<bibRefCitation id="491524C32F5F02A958AD30C0F283FD79" DOI="10.4039/entm10266fv" author="Schmid" firstAuthor="Schmid" journalOrPublisher="Memoires de la Société Entomologique du Canada" pagination="1-230" refId="B64" refString="Schmid F (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Société Entomologique du Canada 66: 1–230. https://doi.org/10.4039/entm10266fv" title="Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera)." volume="66" year="1970">Schmid 1970</bibRefCitation>
|
||||
). It is most similar to
|
||||
<taxonomicName id="DE9E8F20446C7F6C68E646A0F1460D34" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pubescens">
|
||||
<emphasis id="80AF3F6CF708CD1118B014BD4E09BC22" italics="true">R. pubescens</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="A5F22041731E96F6CB5B01CBFBAA2210" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Rhyacophila lignumvallis sp. nov., male genitalia, dorsal view (A), lateral view (B). R. pubescens, male genitalia, dorsal view (C), lateral view (D). R. tsurakiana, male genitalia, dorsal view (E), lateral view (F). Scale bars: 100 μm. Specimens were photographed by W. Graf." figureDoi="10.3897/zookeys.1218.132275.figure5" httpUri="https://binary.pensoft.net/fig/1180557">5 C, D</figureCitation>
|
||||
),
|
||||
<taxonomicName id="E6F196D918BDA0B556481BA5EE43A0C7" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="446247FC4F06BDC1C98210CF2CF47B0D" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
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<figureCitation id="82B922E99080B254BD5859313358E567" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Rhyacophila lignumvallis sp. nov., male genitalia, dorsal view (A), lateral view (B). R. pubescens, male genitalia, dorsal view (C), lateral view (D). R. tsurakiana, male genitalia, dorsal view (E), lateral view (F). Scale bars: 100 μm. Specimens were photographed by W. Graf." figureDoi="10.3897/zookeys.1218.132275.figure5" httpUri="https://binary.pensoft.net/fig/1180557">5 E, F</figureCitation>
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||||
),
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<taxonomicName id="47EE4E18007DF90C7E83111F927F8671" authority="Olah & Vincon, 2021" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="ligurica">
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||||
<emphasis id="6ECBA450377C1A3D1F35D88CCD0ADDCC" italics="true">R. ligurica</emphasis>
|
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<bibRefCitation id="95BDAC3F05727A93B6C05F0AD0F879D3" DOI="10.18348/opzool.2021.1.3" author="Oláh J & Vinçon G & Coppa G" journalOrPublisher="Opuscula Zoologica (Budapest)" pagination="3-67" refId="B53" refString="Olah J, Vincon G, Coppa G (2021) On the Trichoptera of Italy with delineation of incipient sibling species. Opuscula Zoologica (Budapest) 52: 03 - 67. https: // doi. org / 10.18348 / opzool. 2021.1. 3" title="On the Trichoptera of Italy with delineation of incipient sibling species." volume="52" year="2021">Oláh & Vinçon, 2021</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(in
|
||||
<bibRefCitation id="B1C202B57EC0224D3AE068F4E04982A7" DOI="10.18348/opzool.2021.1.3" author="Olah" etAl="et al." firstAuthor="Olah" journalOrPublisher="Opuscula Zoologica (Budapest)" pagination="3-67" refId="B53" refString="Oláh J, Vinçon G, Coppa G (2021) On the Trichoptera of Italy with delineation of incipient sibling species. Opuscula Zoologica (Budapest) 52: 03–67. https://doi.org/10.18348/opzool.2021.1.3" title="On the Trichoptera of Italy with delineation of incipient sibling species." volume="52" year="2021">Oláh et al. 2021</bibRefCitation>
|
||||
, figs 55–57 therein),
|
||||
<taxonomicName id="3D797507BD2E70A9665AE6C3C23C7E78" authority="Olah & Vincon, 2021" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="harmasa">
|
||||
<emphasis id="3DDA8C6A645D54A6D34DAEFEA1F1C7F6" italics="true">R. harmasa</emphasis>
|
||||
<bibRefCitation id="334E7D502C0AE59B4E72C4DEEBECC630" DOI="10.18348/opzool.2021.1.3" author="Oláh J & Vinçon G & Coppa G" journalOrPublisher="Opuscula Zoologica (Budapest)" pagination="3-67" refId="B53" refString="Olah J, Vincon G, Coppa G (2021) On the Trichoptera of Italy with delineation of incipient sibling species. Opuscula Zoologica (Budapest) 52: 03 - 67. https: // doi. org / 10.18348 / opzool. 2021.1. 3" title="On the Trichoptera of Italy with delineation of incipient sibling species." volume="52" year="2021">Oláh & Vinçon, 2021</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(in
|
||||
<bibRefCitation id="480496FD168A19013CB4902D3761B5B7" DOI="10.18348/opzool.2021.1.3" author="Olah" etAl="et al." firstAuthor="Olah" journalOrPublisher="Opuscula Zoologica (Budapest)" pagination="3-67" refId="B53" refString="Oláh J, Vinçon G, Coppa G (2021) On the Trichoptera of Italy with delineation of incipient sibling species. Opuscula Zoologica (Budapest) 52: 03–67. https://doi.org/10.18348/opzool.2021.1.3" title="On the Trichoptera of Italy with delineation of incipient sibling species." volume="52" year="2021">Oláh et al. 2021</bibRefCitation>
|
||||
, figs 52–54 therein), and
|
||||
<taxonomicName id="29EC07E0DE39BE6A7F4DC300E8BEC5E5" authority="Olah & Vincon, 2021" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="abruzzica">
|
||||
<emphasis id="00EC1E7F6BF5BC49A10EADE1F055BCDC" italics="true">R. abruzzica</emphasis>
|
||||
<bibRefCitation id="F04B4B86204CF9BC5C43C357972954A1" DOI="10.18348/opzool.2021.1.3" author="Oláh J & Vinçon G & Coppa G" journalOrPublisher="Opuscula Zoologica (Budapest)" pagination="3-67" refId="B53" refString="Olah J, Vincon G, Coppa G (2021) On the Trichoptera of Italy with delineation of incipient sibling species. Opuscula Zoologica (Budapest) 52: 03 - 67. https: // doi. org / 10.18348 / opzool. 2021.1. 3" title="On the Trichoptera of Italy with delineation of incipient sibling species." volume="52" year="2021">Oláh & Vinçon, 2021</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(in
|
||||
<bibRefCitation id="A3B97D849F81B3AB42714532F34EAB1C" DOI="10.18348/opzool.2021.1.3" author="Olah" etAl="et al." firstAuthor="Olah" journalOrPublisher="Opuscula Zoologica (Budapest)" pagination="3-67" refId="B53" refString="Oláh J, Vinçon G, Coppa G (2021) On the Trichoptera of Italy with delineation of incipient sibling species. Opuscula Zoologica (Budapest) 52: 03–67. https://doi.org/10.18348/opzool.2021.1.3" title="On the Trichoptera of Italy with delineation of incipient sibling species." volume="52" year="2021">Oláh et al. 2021</bibRefCitation>
|
||||
, figs 49–51 therein), but
|
||||
<taxonomicName id="7231C5AFDCC5D21EE5CF35A199BF87A6" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
|
||||
<emphasis id="4ECCD8E6E7D8719A5EAB7E33036CDC29" italics="true">R. lignumvallis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="23B86A0BC4138D5201E334A5E692953D" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is distinguishable from these species by the shape of tergum X, the dorsal arm of the phallic apparatus in lateral and ventral views, the shape of the aedeagus and parameres, and the second segment of the inferior appendages.
|
||||
</paragraph>
|
||||
<paragraph id="288356111CF2067D569ED10003481D91">
|
||||
The dorsal surface of segment X is convex in all these species but narrower and higher in the new species,
|
||||
<taxonomicName id="0E8230D652B2206379CAC27B6E541BFD" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="C5B5F307F52E3A2AE2BB7E35545328B0" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="8622C321F34C75878B77A46DEDC7FCFA" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="harmasa">
|
||||
<emphasis id="BBCC4C1363B09D96AB4C1F9D309E5FFB" italics="true">R. harmasa</emphasis>
|
||||
</taxonomicName>
|
||||
. In dorsal view, however, the new species has a slightly membranous, mesally notched, and inflated segment X, while segment X in
|
||||
<taxonomicName id="70AACBE71587BE1A26DD656C96CD2E51" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="438C0FDC342F39BD33FE7B368D8F2BDE" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
appears flatter. Additionally, the dorsal branch of tergum X is rounded and broader in the new species, while it is narrower in
|
||||
<taxonomicName id="8607C1A489CBEBB859C4E408273DEB5A" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="54BAC86C9FA6CA721309569250B007DE" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="B2699AA5AAFF6AC6591CAA1FA27B2514" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="harmasa">
|
||||
<emphasis id="27A6D9AAD96DBDF395410A5D5F014D54" italics="true">R. harmasa</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="4F395A8E49C3EDE60C67912173D922C1">
|
||||
The dorsal appendix of the phallic apparatus in the new species is longer than the aedeagus and the parameres (Fig.
|
||||
<figureCitation id="B0867E7430AC7E5CF8758C1E0B5F4FCA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 D</figureCitation>
|
||||
). This also occurs in
|
||||
<taxonomicName id="6F1D4862235F9E637C3D3DAE408C6FC1" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="harmasa">
|
||||
<emphasis id="83D9308EA9BD60942AC205F4B7C2C4F1" italics="true">R. harmasa</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="759FAC994524EB63EF65FD454903772B" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="ligurica">
|
||||
<emphasis id="4B436C811EA8813D712E381004F2B2AC" italics="true">R. ligurica</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="370261356F4212E8BB0802A83072C29E" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pubescens">
|
||||
<emphasis id="43595150864C5FFAF1377A6D2B663481" italics="true">R. pubescens</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="A2364EEA835FDF2B024A03B669E7963C" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="F7281A8A3E5A0D57B515E8AF964F517C" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
but not in
|
||||
<taxonomicName id="873B3BEB553D5BCC8AA1045C7E9E2363" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="abruzzica">
|
||||
<emphasis id="D8B1541771EED335B826D7A775B64048" italics="true">R. abruzzica</emphasis>
|
||||
</taxonomicName>
|
||||
. However, the shape of the dorsal appendix in lateral view in the new species is digitate and slightly curved dorsad, whereas
|
||||
<taxonomicName id="0F004952D9C01FBB706CB3B6166A4EBE" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="harmasa">
|
||||
<emphasis id="32FC53ACE3D4C90EDBF8AB9588B30293" italics="true">R. harmasa</emphasis>
|
||||
</taxonomicName>
|
||||
has a slightly wider apical half,
|
||||
<taxonomicName id="A94AD377E74A49157E000142F6B60A0A" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="ligurica">
|
||||
<emphasis id="753CD83C713FE052316D87935CDC8F3E" italics="true">R. ligurica</emphasis>
|
||||
</taxonomicName>
|
||||
has a low bump mesally on its dorsal surface, and
|
||||
<taxonomicName id="FEAD96A0FDA8EA1F46D1FB705916E556" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="AE21D559B5A9F31A5BA909FD00304EDA" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
has a straight and flat dorsal appendix. In
|
||||
<taxonomicName id="D18B82A41C85101EFE1912B6DD45EDD4" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="abruzzica">
|
||||
<emphasis id="78039EFD98C7483E0E9223D184B07E5A" italics="true">R. abruzzica</emphasis>
|
||||
</taxonomicName>
|
||||
, the dorsal appendage is broad and medially widened in lateral view. In comparison to
|
||||
<taxonomicName id="2260AFF3B5A8F765D4E6937A2957EB7B" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
|
||||
<emphasis id="A0D17C0BCDC3366BFB2F7F9CC72DEBEA" italics="true">R. lignumvallis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A1AEFEC01E03341EDD037F9DCBE01224" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
(Figs
|
||||
<figureCitation id="D5FE590CAA015590EF4C7420547FF65E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 D</figureCitation>
|
||||
,
|
||||
<figureCitation id="D5B10A8CE27B928AC9DA35B5B546C568" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Rhyacophila lignumvallis sp. nov., male genitalia, dorsal view (A), lateral view (B). R. pubescens, male genitalia, dorsal view (C), lateral view (D). R. tsurakiana, male genitalia, dorsal view (E), lateral view (F). Scale bars: 100 μm. Specimens were photographed by W. Graf." figureDoi="10.3897/zookeys.1218.132275.figure5" httpUri="https://binary.pensoft.net/fig/1180557">5 A, B</figureCitation>
|
||||
),
|
||||
<taxonomicName id="F8587E1D8B6FB0A9134F045BFAAC714E" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="472E570992BCAFFCF12FA0DFC9DF32E5" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="8691B0158F6D51BE83E8A102EAD5F164" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="abruzzica">
|
||||
<emphasis id="3EAB4F1129352C6B7495C5F666813C4F" italics="true">R. abruzzica</emphasis>
|
||||
</taxonomicName>
|
||||
, the dorsal appendix of the phallic apparatus is much longer and clearly exceeds segment X in dorsal view in
|
||||
<taxonomicName id="9DA942496D3C407F2E9413AF9513E689" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="harmasa">
|
||||
<emphasis id="7D81E799CDEE7EE378A511B9B15163F8" italics="true">R. harmasa</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="B50F430DFCD89892D59826006C170387" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pubescens">
|
||||
<emphasis id="A2BEE6165C1BE260800300AC3488E80E" italics="true">R. pubescens</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4AA3B03DD89D8DBD3940E5A29CA94C4C" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="ligurica">
|
||||
<emphasis id="C3E7AE156D740AD147D2B4FEFF4183E9" italics="true">R. ligurica</emphasis>
|
||||
</taxonomicName>
|
||||
. In ventral view, the dorsal appendix is straight and rounded apically in the new species (Fig.
|
||||
<figureCitation id="CE056F675E341D5ACF0F8DD67D0F912C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 E</figureCitation>
|
||||
), slightly inflated on the apical half and rounded apically in
|
||||
<taxonomicName id="9820AFD462F49E365B2A3F3B83997CAB" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="harmasa">
|
||||
<emphasis id="9B01F5DE7009ADF5C28B1AF7AD5D3D90" italics="true">R. harmasa</emphasis>
|
||||
</taxonomicName>
|
||||
, almost straight and truncate apically in
|
||||
<taxonomicName id="B2B8932B7BD2A83B9F7305BCF96B3D5D" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="ligurica">
|
||||
<emphasis id="7F42F3766A6B3BC86C7555CA7990B369" italics="true">R. ligurica</emphasis>
|
||||
</taxonomicName>
|
||||
, constricted basally and truncate apically in
|
||||
<taxonomicName id="71E566D68AC9D0F363F9BA9A1AB3E3C5" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="0CE0AD39BDDC7E38B9ED28F34B81EAB5" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
, and rectangular in
|
||||
<taxonomicName id="E288BEC4CDA4A497C8195E18771189B6" authorityName="Olah & Vincon" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="abruzzica">
|
||||
<emphasis id="94A96ADBF8467F385E0DDA0AA7895F14" italics="true">R. abruzzica</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="05CCCF6DDD3FC254B6C33F8B2C3D6594">
|
||||
The aedeagus and parameres in
|
||||
<taxonomicName id="F56871116C14D1180B8EF1BE25F2AFB4" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
|
||||
<emphasis id="D1989B022415AF833CFDAE69B273D3E1" italics="true">R. lignumvallis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="1E72766905F271C1DDE3C9DC4B4D4357" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
are most similar to
|
||||
<taxonomicName id="5597B787622723E894E1712357E71C77" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pubescens">
|
||||
<emphasis id="F62D0D1E1D36ADDCD400826DDE0B00A8" italics="true">R. pubescens</emphasis>
|
||||
</taxonomicName>
|
||||
. However, in lateral view, the tip of the aedeagus in the new species is slenderer and slightly more curved apically than in
|
||||
<taxonomicName id="7E7619999BC8D28EB9185C6CFF230F3C" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pubescens">
|
||||
<emphasis id="17236CADAB877128D4FBDF818E1F5A62" italics="true">R. pubescens</emphasis>
|
||||
</taxonomicName>
|
||||
. In lateral view, the parameres in the new species are broader than in
|
||||
<taxonomicName id="890DF3037C2D1F1909FEFB2884F20299" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pubescens">
|
||||
<emphasis id="548126B9D7708CF8A18D5B62A521B362" italics="true">R. pubescens</emphasis>
|
||||
</taxonomicName>
|
||||
. In ventral view, the parameres in the new species are club-shaped and curved mesad, while in
|
||||
<taxonomicName id="CE9E44C8871543D0B5AB9EE2087027EF" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pubescens">
|
||||
<emphasis id="3DEC1B4847206B4A26962F4855EF137E" italics="true">R. pubescens</emphasis>
|
||||
</taxonomicName>
|
||||
, the parameres are digitate and directed posterad.
|
||||
</paragraph>
|
||||
<paragraph id="7C172F555DEBC5660F1D3B9AE8301BB8">
|
||||
The second segment of the inferior appendages in the new species is triangular, with a straight dorsal margin, while all the other species have a concave dorsal margin (Fig.
|
||||
<figureCitation id="E10C990E3D08F8E9D5EF454B83B93F24" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 A</figureCitation>
|
||||
,
|
||||
<figureCitation id="2EAC90695B3D8B7CB79E2A7165FBD271" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Rhyacophila lignumvallis sp. nov., male genitalia, dorsal view (A), lateral view (B). R. pubescens, male genitalia, dorsal view (C), lateral view (D). R. tsurakiana, male genitalia, dorsal view (E), lateral view (F). Scale bars: 100 μm. Specimens were photographed by W. Graf." figureDoi="10.3897/zookeys.1218.132275.figure5" httpUri="https://binary.pensoft.net/fig/1180557">5 B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ESHBI" type="description">
|
||||
<paragraph id="503E5EF841DCA1958A8830753441BBB3">
|
||||
<heading id="8EED433F3AEAF287043422CD818635D2" reason="title">Description.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="933902702C2FA7698865535AB0AD4FD3">
|
||||
<emphasis id="8E0591AD500064B01AE991EABC624FA2" bold="true">
|
||||
<emphasis id="6A68F7A9A91D0410CC489E418A2DC347" bold="true" italics="true">Adult male</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Specimen in ethanol, mostly denuded; dorsally brown, ventrally light brown. Legs light brown with slightly darker tibial spurs. Head with frontal setal wart triangular; antennal setal wart subtriangular and smaller than frontal setal wart; posterior setal warts oval and connected to ocellar setal warts via a raised cuticular “ bridge ” (see
|
||||
<bibRefCitation id="7837DAC405074B39B5F627960425C027" DOI="10.4039/entm10266fv" author="Schmid F" journalOrPublisher="Memoires de la Société Entomologique du Canada" pagination="1-230" refId="B64" refString="Schmid F (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada 66: 1 - 230. https: // doi. org / 10.4039 / entm 10266 fv" title="Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera)." volume="66" year="1970">Schmid, 1970</bibRefCitation>
|
||||
; pl. I, fig. 1). Forewing length (
|
||||
<quantity id="CF79DBCC56032EE418371AF978D71016" metricMagnitude="-3" metricUnit="m" metricValue="8.8" unit="mm" value="8.8">8.8 mm</quantity>
|
||||
,
|
||||
<emphasis id="A9F5E49265B69536FC705816A187C4FC" italics="true">N</emphasis>
|
||||
= 1) mostly denuded, with sparse, very short light brown setae and golden brown microtrichia. Hind wings also mostly denuded, with slightly longer light brown setae. Forewing (Fig.
|
||||
<figureCitation id="7A13A4F71C6C0D03A423DED79261F693" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhyacophila lignumvallis sp. nov., wing venation. C, costal vein; Sc, subcostal vein; R 1 – R 5, first to fifth branches of the radial vein; M 1 – M 4, first to fourth branches of the medial vein; M 3 + 4, medial vein 3 + 4 (hindwing); Cu 1 a, anterior branch of first cubital vein; Cu 1 b, posterior branch of first cubital vein; Cu 2, second cubital vein; 1 A – 3 A, first to third anal veins; r, radial crossvein; r-m, radiomedial crossvein; m-cu, mediocubital crossvein; I – V, first to fifth wing forks; TC, thyridial cell. Scale bar: 1 mm. Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure3" httpUri="https://binary.pensoft.net/fig/1180555">3 A</figureCitation>
|
||||
) with crossveins connecting costal (C) and subcostal (Sc) veins; subcostal (Sc) and first radial (R
|
||||
<subScript id="83AC60341E59995718DB9754CAB08FE7">1</subScript>
|
||||
) veins, first (R
|
||||
<subScript id="7B9710BD9F27DBC9C010C7B9F3F221FC">1</subScript>
|
||||
) and second radial (R
|
||||
<subScript id="494B7D5937EE74B70B91D32C28B5434C">2</subScript>
|
||||
) veins (
|
||||
<emphasis id="93B2370EF796A4B2F9FFED8C51863A74" italics="true">r</emphasis>
|
||||
), fifth radial (R
|
||||
<subScript id="845A07D39E0933B58BF8B41B9DCE15C5">5</subScript>
|
||||
) and first medial (M) veins (
|
||||
<emphasis id="5A7783AC4D8C52E43C73EB8794931704" italics="true">r-m</emphasis>
|
||||
), first medial (M) and first cubital (Cu 1) veins (
|
||||
<emphasis id="6C2CAEB12EEE490C1EB1DF15CE5ECAFB" italics="true">m-cu</emphasis>
|
||||
), and first cubital (Cu 1) and second cubital (Cu 2) veins. Hind wing (Fig.
|
||||
<figureCitation id="C3BF3F880F611A85299FF95FC9C8E288" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhyacophila lignumvallis sp. nov., wing venation. C, costal vein; Sc, subcostal vein; R 1 – R 5, first to fifth branches of the radial vein; M 1 – M 4, first to fourth branches of the medial vein; M 3 + 4, medial vein 3 + 4 (hindwing); Cu 1 a, anterior branch of first cubital vein; Cu 1 b, posterior branch of first cubital vein; Cu 2, second cubital vein; 1 A – 3 A, first to third anal veins; r, radial crossvein; r-m, radiomedial crossvein; m-cu, mediocubital crossvein; I – V, first to fifth wing forks; TC, thyridial cell. Scale bar: 1 mm. Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure3" httpUri="https://binary.pensoft.net/fig/1180555">3 B</figureCitation>
|
||||
) with crossveins connecting subcostal (Sc) and first radial (R
|
||||
<subScript id="2320C2C9EC1DFF960780AAFA6BAAF0F8">1</subScript>
|
||||
) veins, fifth radial (R
|
||||
<subScript id="29692C56C55BDC33B97891B44E9C2CB4">5</subScript>
|
||||
) and first medial (M) veins (
|
||||
<emphasis id="D6B8C2FFB9263450BD9658091037330A" italics="true">r-m</emphasis>
|
||||
), and M
|
||||
<subScript id="5AC566742B1F1D6A0CECE7CD30CA1716">3 + 4</subScript>
|
||||
and first cubital (Cu 1) present.
|
||||
</paragraph>
|
||||
<caption id="AA2F5EC821FA7EA2188CE306DFB2814D" ID-DOI="10.3897/zookeys.1218.132275.figure3" ID-arpha="F68DAFA5-0A9F-5DB7-95B5-561B0F46FEF7" httpUri="https://binary.pensoft.net/fig/1180555" startId="F3">
|
||||
<paragraph id="E9690AAEF551D465AF5881327F0AC867">
|
||||
<label id="58BE4BA6D81C735FAFCD65F8559D6FCE">Figure 3.</label>
|
||||
</paragraph>
|
||||
<paragraph id="F33CC45DCE9E033A508B4A2F9ED31DB0">
|
||||
<taxonomicName id="58BC326495B3FA2CA92B0D6E020819C2" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
|
||||
<emphasis id="353F5EFFAB151285FABA8FCDAA23E384" italics="true">Rhyacophila lignumvallis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="ABA3A0164B0EA93C78D2DA6ED847FEEB" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, wing venation.
|
||||
<emphasis id="F0F6954AEC5C336F4107A792361D633E" bold="true">C</emphasis>
|
||||
, costal vein;
|
||||
<emphasis id="93CA76F9873D3F657B7B969E426DA706" bold="true">Sc</emphasis>
|
||||
, subcostal vein;
|
||||
<emphasis id="97DCC0F01020CDAF1573E04CE6687CCF" bold="true">R 1 – R 5</emphasis>
|
||||
, first to fifth branches of the radial vein;
|
||||
<emphasis id="D5C61EBA682DD0AE82A9E94E1CF0973D" bold="true">M 1 – M 4</emphasis>
|
||||
, first to fourth branches of the medial vein;
|
||||
<emphasis id="80E777D3D1D0493DEE0C26A10A22F6B7" bold="true">M 3 + 4</emphasis>
|
||||
, medial vein 3 + 4 (hindwing);
|
||||
<emphasis id="3DB1547B0F49F0273743111422943135" bold="true">Cu 1 a</emphasis>
|
||||
, anterior branch of first cubital vein;
|
||||
<emphasis id="73557F5F2F89EBDE92133B5F35A0F4A5" bold="true">Cu 1 b</emphasis>
|
||||
, posterior branch of first cubital vein;
|
||||
<emphasis id="CF686CD72D6B6AB385D72CF9AF9E1BFE" bold="true">Cu 2</emphasis>
|
||||
, second cubital vein;
|
||||
<emphasis id="B1F5553B2305AE18E8E53B21E3320974" bold="true">1 A – 3 A</emphasis>
|
||||
, first to third anal veins;
|
||||
<emphasis id="451BF0613D68D55047152F39E47E2E29" bold="true">r</emphasis>
|
||||
, radial crossvein;
|
||||
<emphasis id="DB2E3086CBBBBCF75F16916EFCEDA19F" bold="true">r-m</emphasis>
|
||||
, radiomedial crossvein;
|
||||
<emphasis id="0C5EC3403C491036E2D66C5D6A66C918" bold="true">m-cu</emphasis>
|
||||
, mediocubital crossvein;
|
||||
<emphasis id="E5C6591D10D32A68DD6F522AEDBD08AE" bold="true">I – V</emphasis>
|
||||
, first to fifth wing forks;
|
||||
<emphasis id="3975152FE99D01BF32F22A46504874D0" bold="true">TC</emphasis>
|
||||
, thyridial cell. Scale bar: 1 mm. Illustrations were produced by Ernesto Rázuri-Gonzales.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="2E2CFF5A917CCA4034D054B2C19579B8">
|
||||
<emphasis id="BBC27D948904BA38450611EB6FADADD5" bold="true">
|
||||
<emphasis id="E2B24BF2E92CD536BCC2A64BF3EE7EB9" bold="true" italics="true">Male genitalia</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Segment IX longitudinally short in lateral view (Fig.
|
||||
<figureCitation id="FF58B541279CD0A9FD72D25905E377AE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 A</figureCitation>
|
||||
), anterior and posterior margins slightly concave, dorsal half slightly longer than ventral. Dorsal surface of segment X membranous, slightly inflated, shallowly notched mesally in dorsal view (Fig.
|
||||
<figureCitation id="601A07F3501D2958C953AA7E97792050" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 C</figureCitation>
|
||||
). Dorsal branch of segment X short and rounded in lateral view. Anal sclerites partially fused to each other basally and to segment X, in lateral view, directed ventrad. First article of inferior appendages (Fig.
|
||||
<figureCitation id="825ED1A1844AFCDA5DE3ED896493911D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 A</figureCitation>
|
||||
) rectangular in lateral view, slightly broader basally than apically; in ventral view (Fig.
|
||||
<figureCitation id="E7AA3A431D5FA6F3423A35C6ACE461D5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 B</figureCitation>
|
||||
), slightly broader apically than basally, with a small setose bump basally on mesal surface. Second article of inferior appendages (Fig.
|
||||
<figureCitation id="D124BFAFC81107F090D483B9D3E930C4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 A</figureCitation>
|
||||
) quadrangular in lateral view, dorsal and ventral margins slightly diverging, posterodorsal margin straight, at a 130 ° angle to dorsal margin; in ventral view (Fig.
|
||||
<figureCitation id="A77173832345D193353E98E954878AC4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 B</figureCitation>
|
||||
), mitton-shaped, mesal margin with very short, peg-like setae basally and longer setae apically. Phallic apparatus (Fig.
|
||||
<figureCitation id="DD071D693032773D89AFDBC962D131F2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 D</figureCitation>
|
||||
) with dorsal appendix straight in lateral view, slightly curved dorsad, rounded apically, longer than parameres and aedeagus; in ventral view, straight, lateral margins slightly sinuous, rounded apically. Parameres in lateral view (Fig.
|
||||
<figureCitation id="2803701BC1E42A21820E7321A9BEA040" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 D</figureCitation>
|
||||
) broader than aedeagus, slightly curved posterodorsad, ventral margin straight, dorsal margin slightly sinuous, rounded apically; in ventral view (Fig.
|
||||
<figureCitation id="8F7ED6CE16E1256E0BAD81A0BADDFB14" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 E</figureCitation>
|
||||
), club-shaped, directed mesad. Aedeagus in lateral view (Fig.
|
||||
<figureCitation id="A7C75C2888A85CA06281C98EB298F183" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Rhyacophila lignumvallis sp. nov., male genitalia, lateral view (A), ventral view (B), segment X in dorsal view (C), phallic apparatus in lateral view (D), and phallic apparatus in ventral view (E). Illustrations were produced by Ernesto Rázuri-Gonzales." figureDoi="10.3897/zookeys.1218.132275.figure4" httpUri="https://binary.pensoft.net/fig/1180556">4 D</figureCitation>
|
||||
) slender, slightly sinuous, tapering towards its apex; in ventral view, slender and straight.
|
||||
</paragraph>
|
||||
<caption id="C8CBB993D6EA842B0BA81F302DBEF48F" ID-DOI="10.3897/zookeys.1218.132275.figure4" ID-arpha="B3AAB607-F236-5298-8412-50EB9AD782E0" httpUri="https://binary.pensoft.net/fig/1180556" startId="F4">
|
||||
<paragraph id="C99B04FB940BE715667F6BAD9B87CDD5">
|
||||
<label id="79D2B4DD2A680E5DCBB16E155C8E2313">Figure 4.</label>
|
||||
</paragraph>
|
||||
<paragraph id="E3BCABDAF3E2E81AB37027087389B9F6">
|
||||
<taxonomicName id="496BA4EA24DDE88F1F3A4F6C588E73E0" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
|
||||
<emphasis id="04E866356AA2216F5BB5AB80C696F5AF" italics="true">Rhyacophila lignumvallis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="8047EEA3FB59C576BFD7A0A4546FDFDF" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, male genitalia, lateral view (
|
||||
<emphasis id="B931E6ECDC3DF963BD37DFB9D9C3D54D" bold="true">A</emphasis>
|
||||
), ventral view (
|
||||
<emphasis id="6216C675B028A0E9229645EC3FB169D9" bold="true">B</emphasis>
|
||||
), segment X in dorsal view (
|
||||
<emphasis id="212E73D887CD2EC1EBB5AFCC135FB381" bold="true">C</emphasis>
|
||||
), phallic apparatus in lateral view (
|
||||
<emphasis id="8C70A3616CA595CC35E1E147F2B7318A" bold="true">D</emphasis>
|
||||
), and phallic apparatus in ventral view (
|
||||
<emphasis id="8B16C9B8B355BDDFCC5891A38DBBA072" bold="true">E</emphasis>
|
||||
). Illustrations were produced by Ernesto Rázuri-Gonzales.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="7038B1989A8206F19284BF0AD496C0B9" ID-DOI="10.3897/zookeys.1218.132275.figure5" ID-arpha="A7F42EAC-73B0-55E9-81AE-3B87279C4458" httpUri="https://binary.pensoft.net/fig/1180557" startId="F5">
|
||||
<paragraph id="910FC687A38D7BA0B1AE9E47926C5E0A">
|
||||
<label id="46507BFA42C23582D92F7CEA2BAA7318">Figure 5.</label>
|
||||
</paragraph>
|
||||
<paragraph id="A33629BDA63FCED5E1AF5F78BBC6137B">
|
||||
<taxonomicName id="59B74B902A639241CCF47BCFD702C9AD" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
|
||||
<emphasis id="93B547B2BD8FBB2CC01F72C3A73AD4E9" italics="true">Rhyacophila lignumvallis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="484381A974925911E941E0F5471BD49B" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
, male genitalia, dorsal view (
|
||||
<emphasis id="5E886760155FD73F62B27EE33D136EF8" bold="true">A</emphasis>
|
||||
), lateral view (
|
||||
<emphasis id="E5927E36F25DCE06330C5CB0B12E2F7E" bold="true">B</emphasis>
|
||||
).
|
||||
<taxonomicName id="69D7A8FEF179EEFBBB80B9DC2BFA9876" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="pubescens">
|
||||
<emphasis id="40B12587DACB95960AD53563D71CABC1" italics="true">R. pubescens</emphasis>
|
||||
</taxonomicName>
|
||||
, male genitalia, dorsal view (
|
||||
<emphasis id="82568F9F3D4E6D729FAF8AFD2ABB1CE4" bold="true">C</emphasis>
|
||||
), lateral view (
|
||||
<emphasis id="39357A7FE35AF561112F2EC41364EF8D" bold="true">D</emphasis>
|
||||
).
|
||||
<taxonomicName id="1AD58687064DCCEF4B7491AD7FEC79AF" authorityName="Malicky" authorityYear="1984" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="tsurakiana">
|
||||
<emphasis id="190A7052C12479DE0152B487B47C18B9" italics="true">R. tsurakiana</emphasis>
|
||||
</taxonomicName>
|
||||
, male genitalia, dorsal view (
|
||||
<emphasis id="225D2F3C4AC53D1EB94C509234466458" bold="true">E</emphasis>
|
||||
), lateral view (
|
||||
<emphasis id="21C2B2FD89059588E7D50268D07A4F01" bold="true">F</emphasis>
|
||||
). Scale bars: 100 μm. Specimens were photographed by W. Graf.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E3OBI" type="etymology">
|
||||
<paragraph id="2AFC29CBA9FAC272C2FC0269A0D0C193">
|
||||
<heading id="1B4D5C47502E6A7C0EDF58B0450818A8" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="9024759FFC7E3A96470B0DC5CDDBF3C2">
|
||||
We dedicate this species to Dr Ralph W. Holzenthal to honor his contributions to caddisfly taxonomy and systematics.
|
||||
<emphasis id="CBE267CCADAC8CA56909943E599A13F1" italics="true">Lignumvallis</emphasis>
|
||||
, wood valley, is derived from the Latin translation of Ralph’s last name.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EJPBI" type="habitat">
|
||||
<paragraph id="89CD1C2E0FB1C6ADCE7C0DD363F78C3B">
|
||||
<heading id="FFBEB1FAF7B5046A772EB411E894A1EA" reason="title">Habitat.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="A9BDB625BD560AA90E6AC93F7BC95714">
|
||||
The river Vecchio is a crystal-clear, slow-flowing stream with a heterogeneous bottom substrate that varies from sandy patches to gravel to boulders. Stabile substrates were densely covered by
|
||||
<taxonomicName id="194074BD4831DE10CFFD4B351D920B1C" authorityName="Mosely" authorityYear="1930" class="Insecta" family="Glossosomatidae" genus="Agapetus" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="cyrnensis">
|
||||
<emphasis id="C351AB74E4309BCB584F98199A8581E0" italics="true">Agapetus cyrnensis</emphasis>
|
||||
</taxonomicName>
|
||||
pupae. As many spring trickles enter the river on its left margin and the specimen was collected by sweeping the vegetation, the habitat of
|
||||
<taxonomicName id="699213F81F9FCF46FCDACCF0BA1FCE9C" authorityName="Graf & Razuri-Gonzales" authorityYear="2024" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" phylum="Arthropoda" rank="species" species="lignumvallis" status="sp. nov.">
|
||||
<emphasis id="E855B709BC147EA30AA894A2F9F607CC" italics="true">R. lignumvallis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="AE2428DDB0F7C5C78AEF23369026A9DE" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
remains unknown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue