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<document id="533A74808E343EA9FFF506957FEB806A" ID-DOI="10.11646/zootaxa.5487.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13210771" ID-ZooBank="50AC1672-FC20-4502-8D8A-4BCAE95F719B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722789583779" checkinUser="plazi" docAuthor="Zatwarnicki, Tadeusz &amp; Mathis, Wayne N." docDate="2024" docId="03D187B73A067B04FF4EFE525260F909" docLanguage="en" docName="zootaxa.5487.1.1.pdf" docOrigin="Zootaxa 5487 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5487.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Scatophila mystacina Zatwarnicki &amp; Mathis, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="88" masterDocId="FFE8FFCF3A537B53FFD9FFE4576AFFFA" masterDocTitle="Revision of the Nearctic Species of the Shore-Fly Genus Scatophila Becker (Diptera: Ephydridae)" masterLastPageNumber="100" masterPageNumber="1" pageNumber="86" updateTime="1722790937208" updateUser="ExternalLinkService">
<mods:mods id="BBC998A0CB7AE773CFD17723024614BF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="785D63D107D3B0C4970A776957FDCAAA">Revision of the Nearctic Species of the Shore-Fly Genus Scatophila Becker (Diptera: Ephydridae)</mods:title>
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<mods:namePart id="E4C5B1AA67560F6C4113C832DB51D2EF">Zatwarnicki, Tadeusz</mods:namePart>
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<mods:affiliation id="470FE7A8B8F080D3E289763B13B234E7">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
<mods:nameIdentifier id="8DB23E022F0146957F6CFED239441D63" type="email">zatwar@uni.opole.pl</mods:nameIdentifier>
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<mods:namePart id="B6DB02C751570F57DC8EB1DE0319C638">Mathis, Wayne N.</mods:namePart>
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<mods:affiliation id="858C6B96DB3F7BE3E24F8944D059E4DF">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143 &amp; Department of Entomology, PO BOX 37012, MRC 169, Smithsonian Institution, Washington, DC 20013 - 7012, USA mathisw @ si. edu; https: // orcid. org / 0000 - 0002 - 5012 - 1762 &amp; Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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<mods:title id="712E178D3DA15B14AC058048629AC435">Zootaxa</mods:title>
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<mods:date id="6C7257A88628B64963BDA037BE121B74">2024</mods:date>
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<mods:number id="04C44A172FFE3C5BFBC9784673E1D2C0">2024-07-30</mods:number>
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<subSubSection id="C362652A3A067B06FF4EFE525540FE2B" box="[151,554,438,465]" pageId="85" pageNumber="86" type="nomenclature">
<paragraph id="8BC736A13A067B06FF4EFE525540FE2B" blockId="85.[151,554,438,501]" box="[151,554,438,465]" pageId="85" pageNumber="86">
<emphasis id="B90CEAB33A067B06FF4EFE525540FE2B" bold="true" box="[151,554,438,465]" pageId="85" pageNumber="86">
23.
<taxonomicName id="4C784D223A067B06FF1AFE5256AFFE2B" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[195,453,438,465]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="mystacina" status="sp. nov.">
<emphasis id="B90CEAB33A067B06FF1AFE5256AFFE2B" bold="true" box="[195,453,438,465]" italics="true" pageId="85" pageNumber="86">Scatophila mystacina</emphasis>
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<taxonomicNameLabel id="A23F57C83A067B06FE14FE535540FE2B" box="[461,554,439,465]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C362652A3A067B04FF4EFE3F5260F909" lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86" type="description">
<paragraph id="8BC736A13A067B06FF4EFE3F56B4FE0F" blockId="85.[151,554,438,501]" box="[151,478,474,501]" pageId="85" pageNumber="86">
<figureCitation id="13432A243A067B06FF4EFE3F5668FE0F" box="[151,258,475,501]" captionStart="FIGURES 202–204" captionStartId="85.[151,264,1848,1872]" captionTargetBox="[151,1435,1094,1825]" captionTargetId="figure-325@85.[151,1435,1094,1825]" captionTargetPageId="85" captionText="FIGURES 202–204. Head, thorax, and wing of Scatophila. 202. S. hirsuta Sturtevant &amp; Wheeler, head and thorax, dorsal view. 203. S. mystacina sp. nov. male, wing, dorsal view. 204 S. picta Sturtevant &amp; Wheeler, wing, dorsal view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210935" httpUri="https://zenodo.org/record/13210935/files/figure.png" pageId="85" pageNumber="86">Figs. 203</figureCitation>
,
<figureCitation id="13432A243A067B06FED4FE3E5618FE0F" box="[269,370,474,501]" captionStart-0="FIGURES 205–207" captionStart-1="FIGURES 208–211" captionStartId-0="86.[151,264,1013,1037]" captionStartId-1="87.[151,264,1395,1419]" captionTargetBox-0="[159,1427,189,981]" captionTargetBox-1="[171,1435,181,1371]" captionTargetId-0="figure-30@86.[151,1435,181,989]" captionTargetId-1="figure-27@87.[151,1435,181,1371]" captionTargetPageId-0="86" captionTargetPageId-1="87" captionText-0="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." captionText-1="FIGURES 208–211. Head of Scatophila mystacina sp. nov. 208. Male, latero-oblique view. 209. Male, lateral view. 210. Female, latero-oblique view. 211. Female, lateral view. Scale bar = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13210939" figureDoi-1="http://doi.org/10.5281/zenodo.13210941" httpUri-0="https://zenodo.org/record/13210939/files/figure.png" httpUri-1="https://zenodo.org/record/13210941/files/figure.png" pageId="85" pageNumber="86">205–211</figureCitation>
,
<figureCitation id="13432A243A067B06FEA5FE3F56C3FE0F" box="[380,425,475,501]" captionStart="FIGURES 215–219" captionStartId="89.[151,264,1540,1564]" captionTargetBox="[167,1435,181,1516]" captionTargetId="figure-27@89.[151,1435,181,1517]" captionTargetPageId="89" captionText="FIGURES 215–219. Head, thorax, and wing of Scatophila. 215–216. Head of Scatophila picta Sturtevant &amp; Wheeler. 215. Latero-oblique view. 216. Lateral view. 217. Scatophila mystacina sp. nov., head and thorax, dorsal view. 218–219. Wing, dorsal view. 218. S. pulchra Sturtevant &amp; Wheeler. 219. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210947" httpUri="https://zenodo.org/record/13210947/files/figure.png" pageId="85" pageNumber="86">217</figureCitation>
,
<figureCitation id="13432A243A067B06FE6DFE3F56B4FE0F" box="[436,478,475,501]" captionStart="FIGURE 220" captionStartId="90.[151,250,1586,1610]" captionTargetBox="[151,1435,669,1563]" captionTargetId="figure-233@90.[151,1435,669,1563]" captionTargetPageId="90" captionText="FIGURE 220. Distribution map of Scatophila mystacina sp. nov. (circle), and S. rubribrunnea Sturtevant &amp; Wheeler (squares)." figureDoi="http://doi.org/10.5281/zenodo.13210951" httpUri="https://zenodo.org/record/13210951/files/figure.png" pageId="85" pageNumber="86">220</figureCitation>
</paragraph>
<paragraph id="8BC736A13A067B06FF4EFDF856A7FDA1" blockId="85.[151,1437,540,1071]" pageId="85" pageNumber="86">
Diagnosis.—This species is distinguished from congeners by the following combination of characters: Small shore flies, body length
<quantity id="4C809B443A067B06FEB8FDA456A2FDA1" box="[353,456,576,603]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="85" pageNumber="86" unit="mm" value="1.5">1.50 mm</quantity>
.
</paragraph>
<paragraph id="8BC736A13A067B06FF1EFD8056FFFCC8" blockId="85.[151,1437,540,1071]" pageId="85" pageNumber="86">
<emphasis id="B90CEAB33A067B06FF1EFD80566EFD84" box="[199,260,612,638]" italics="true" pageId="85" pageNumber="86">Head</emphasis>
(
<figureCitation id="13432A243A067B06FECAFD8056D0FD84" box="[275,442,612,638]" captionStart="FIGURES 208–211" captionStartId="87.[151,264,1395,1419]" captionTargetBox="[171,1435,181,1371]" captionTargetId="figure-27@87.[151,1435,181,1371]" captionTargetPageId="87" captionText="FIGURES 208–211. Head of Scatophila mystacina sp. nov. 208. Male, latero-oblique view. 209. Male, lateral view. 210. Female, latero-oblique view. 211. Female, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210941" httpUri="https://zenodo.org/record/13210941/files/figure.png" pageId="85" pageNumber="86">Figs. 208–211</figureCitation>
): Mesofrons and fronto-orbits gray, fronto-orbits slightly darker, brownish gray. Antenna blackish gray, basal flagellomere densely microtomentose. Face of
<specimenCount id="9D7EFD283A067B06FCAEFD6D54E6FD59" box="[887,908,649,675]" count="1" pageId="85" pageNumber="86" type="male">♂</specimenCount>
gray, ventral portion conspicuously protuberant (
<figureCitation id="13432A243A067B06FF46FD485651FD3C" box="[159,315,684,710]" captionStart="FIGURES 208–211" captionStartId="87.[151,264,1395,1419]" captionTargetBox="[171,1435,181,1371]" captionTargetId="figure-27@87.[151,1435,181,1371]" captionTargetPageId="87" captionText="FIGURES 208–211. Head of Scatophila mystacina sp. nov. 208. Male, latero-oblique view. 209. Male, lateral view. 210. Female, latero-oblique view. 211. Female, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210941" httpUri="https://zenodo.org/record/13210941/files/figure.png" pageId="85" pageNumber="86">Fig. 208–209</figureCitation>
), elongate, apex bearing numerous, slightly curved black setulae, these larger toward lateral margin of facial protuberance; face of
<specimenCount id="9D7EFD283A067B06FE2BFD35556DFD11" box="[498,519,721,747]" count="1" pageId="85" pageNumber="86" type="female">♀</specimenCount>
more shallowly protruded than male, more arched, bearing longer setulae toward ventral margin on each side (210–211). Gena moderately high, height slightly more than length of basal flagellomere; gena-to-eye ratio 0.40.
</paragraph>
<paragraph id="8BC736A13A067B06FF1EFCD85656FBD4" blockId="85.[151,1437,540,1071]" pageId="85" pageNumber="86">
<emphasis id="B90CEAB33A067B06FF1EFCD85672FCAC" box="[199,280,828,854]" italics="true" pageId="85" pageNumber="86">Thorax</emphasis>
(
<figureCitation id="13432A243A067B06FEF3FCD856F0FCAC" box="[298,410,828,854]" captionStart="FIGURES 202–204" captionStartId="85.[151,264,1848,1872]" captionTargetBox="[151,1435,1094,1825]" captionTargetId="figure-325@85.[151,1435,1094,1825]" captionTargetPageId="85" captionText="FIGURES 202–204. Head, thorax, and wing of Scatophila. 202. S. hirsuta Sturtevant &amp; Wheeler, head and thorax, dorsal view. 203. S. mystacina sp. nov. male, wing, dorsal view. 204 S. picta Sturtevant &amp; Wheeler, wing, dorsal view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210935" httpUri="https://zenodo.org/record/13210935/files/figure.png" pageId="85" pageNumber="86">Figs. 203</figureCitation>
,
<figureCitation id="13432A243A067B06FE71FCD856BDFCAC" box="[424,471,828,854]" captionStart="FIGURES 215–219" captionStartId="89.[151,264,1540,1564]" captionTargetBox="[167,1435,181,1516]" captionTargetId="figure-27@89.[151,1435,181,1517]" captionTargetPageId="89" captionText="FIGURES 215–219. Head, thorax, and wing of Scatophila. 215–216. Head of Scatophila picta Sturtevant &amp; Wheeler. 215. Latero-oblique view. 216. Lateral view. 217. Scatophila mystacina sp. nov., head and thorax, dorsal view. 218–219. Wing, dorsal view. 218. S. pulchra Sturtevant &amp; Wheeler. 219. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210947" httpUri="https://zenodo.org/record/13210947/files/figure.png" pageId="85" pageNumber="86">217</figureCitation>
): Mesonotum (
<figureCitation id="13432A243A067B06FD5CFCD8558DFCAC" box="[645,743,828,854]" captionStart="FIGURES 215–219" captionStartId="89.[151,264,1540,1564]" captionTargetBox="[167,1435,181,1516]" captionTargetId="figure-27@89.[151,1435,181,1517]" captionTargetPageId="89" captionText="FIGURES 215–219. Head, thorax, and wing of Scatophila. 215–216. Head of Scatophila picta Sturtevant &amp; Wheeler. 215. Latero-oblique view. 216. Lateral view. 217. Scatophila mystacina sp. nov., head and thorax, dorsal view. 218–219. Wing, dorsal view. 218. S. pulchra Sturtevant &amp; Wheeler. 219. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210947" httpUri="https://zenodo.org/record/13210947/files/figure.png" pageId="85" pageNumber="86">Fig. 217</figureCitation>
) mostly grayish tan, stripes faint or lacking; scutum mostly uniformly gray, microtomentose, dull, lacking stripes, but lighter in coloration between acrostichal tracks; disc of scutellum whitish gray; acrostichal setae in 2 rows on anterior third; 2 larger postsutural setae, smaller setae anterior of larger setae. Wing (
<figureCitation id="13432A243A067B06FE55FC4C5680FC38" box="[396,490,936,962]" captionStart="FIGURES 202–204" captionStartId="85.[151,264,1848,1872]" captionTargetBox="[151,1435,1094,1825]" captionTargetId="figure-325@85.[151,1435,1094,1825]" captionTargetPageId="85" captionText="FIGURES 202–204. Head, thorax, and wing of Scatophila. 202. S. hirsuta Sturtevant &amp; Wheeler, head and thorax, dorsal view. 203. S. mystacina sp. nov. male, wing, dorsal view. 204 S. picta Sturtevant &amp; Wheeler, wing, dorsal view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210935" httpUri="https://zenodo.org/record/13210935/files/figure.png" pageId="85" pageNumber="86">Fig. 203</figureCitation>
) with faint pattern of white spots, veins and crossveins slightly darker; veins R
<subScript id="17FC34E43A067B06FA8DFC5E5207FC32" attach="left" box="[1364,1389,954,968]" fontSize="6" pageId="85" pageNumber="86">2+3</subScript>
and R
<subScript id="17FC34E43A067B06FF73FC3A57AEFC16" attach="left" box="[170,196,990,1004]" fontSize="6" pageId="85" pageNumber="86">4+5</subScript>
shallowly undulate near white spots; costal vein ratio 0.27; M
<subScript id="17FC34E43A067B06FCAAFC3A5411FC16" attach="left" box="[883,891,990,1004]" fontSize="6" pageId="85" pageNumber="86">1</subScript>
vein ratio 0.78. Legs with femora and tibiae dark gray, tibiae slightly yellowish at apices; tarsi pale, yellowish, especially basal tarsomeres, apical tarsomeres darker, gray to brown.
</paragraph>
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<paragraph id="8BC736A13A067B06FF4EF8DC52F0F88E" blockId="85.[151,1437,1848,1908]" pageId="85" pageNumber="86">
<emphasis id="B90CEAB33A067B06FF4EF8DC560CF8AA" bold="true" box="[151,358,1848,1872]" pageId="85" pageNumber="86">FIGURES 202–204</emphasis>
. Head, thorax, and wing of
<taxonomicName id="4C784D223A067B06FDA7F8DC5580F8AA" box="[638,746,1848,1872]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB33A067B06FDA7F8DC5580F8AA" box="[638,746,1848,1872]" italics="true" pageId="85" pageNumber="86">Scatophila</emphasis>
</taxonomicName>
. 202.
<taxonomicName id="4C784D223A067B06FCF1F8DD5306F8AA" authority="Sturtevant &amp; Wheeler" authorityName="Sturtevant &amp; Wheeler" box="[808,1132,1848,1872]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B90CEAB33A067B06FCF1F8DD54E3F8AA" box="[808,905,1848,1872]" italics="true" pageId="85" pageNumber="86">S. hirsuta</emphasis>
Sturtevant &amp; Wheeler
</taxonomicName>
, head and thorax, dorsal view. 203.
<taxonomicName id="4C784D223A067B06FF13F8B95620F88E" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[202,330,1885,1908]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="mystacina" status="sp. nov.">
<emphasis id="B90CEAB33A067B06FF13F8B95620F88E" box="[202,330,1885,1908]" italics="true" pageId="85" pageNumber="86">S. mystacina</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A23F57C83A067B06FE89F8B856CBF88E" box="[336,417,1884,1908]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male, wing, dorsal view. 204
<taxonomicName id="4C784D223A067B06FD0BF8B95369F88E" authority="Sturtevant &amp; Wheeler" authorityName="Sturtevant &amp; Wheeler" box="[722,1027,1884,1908]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B90CEAB33A067B06FD0BF8B95474F88E" box="[722,798,1885,1908]" italics="true" pageId="85" pageNumber="86">S. picta</emphasis>
Sturtevant &amp; Wheeler
</taxonomicName>
, wing, dorsal view. Scale bar = 0.2 mm.
</paragraph>
</caption>
<caption id="DF0766293A057B05FF4EFC1152F2FBCB" ID-DOI="http://doi.org/10.5281/zenodo.13210939" ID-Zenodo-Dep="13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87" startId="86.[151,264,1013,1037]" targetBox="[159,1427,189,981]" targetPageId="86" targetType="figure">
<paragraph id="8BC736A13A057B05FF4EFC1152F2FBCB" blockId="86.[151,1436,1013,1073]" pageId="86" pageNumber="87">
<emphasis id="B90CEAB33A057B05FF4EFC11560DFBF7" bold="true" box="[151,359,1013,1037]" pageId="86" pageNumber="87">FIGURES 205–207</emphasis>
. Male terminalia of
<taxonomicName id="4C784D223A057B05FDEFFC115465FBF7" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[566,783,1013,1037]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="mystacina" status="sp. nov.">
<emphasis id="B90CEAB33A057B05FDEFFC115465FBF7" box="[566,783,1013,1037]" italics="true" pageId="86" pageNumber="87">Scatophila mystacina</emphasis>
</taxonomicName>
<emphasis id="B90CEAB33A057B05FCCCFC11540CFBF7" bold="true" box="[789,870,1013,1037]" pageId="86" pageNumber="87">
<taxonomicNameLabel id="A23F57C83A057B05FCCCFC11540CFBF7" box="[789,870,1013,1037]" pageId="86" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A057B05FF1EFB675699F8DF" blockId="86.[151,1437,1155,2010]" pageId="86" pageNumber="87">
<emphasis id="B90CEAB33A057B05FF1EFB67565BFB67" box="[199,305,1155,1181]" italics="true" pageId="86" pageNumber="87">Abdomen</emphasis>
(
<figureCitation id="13432A243A057B05FEE7FB675688FB64" box="[318,482,1155,1182]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Figs. 205–207</figureCitation>
): Tergites uniformly gray, concolorous with coloration of thorax. Male terminalia (
<figureCitation id="13432A243A057B05FABDFB675795FB38" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Figs. 205–207</figureCitation>
): Epandrium in posterior view (
<figureCitation id="13432A243A057B05FDB8FB4355AAFB38" box="[609,704,1191,1218]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 205</figureCitation>
) generally squatly oval except for broadly stepwise emargination at ventral margin, width almost equal to height, lateral margins shallowly curved with short, papilla-like projection at ventrolateral corner; dorsal margin of epandrium in posterior view (
<figureCitation id="13432A243A057B05FC7BFB0B536BFAF0" box="[930,1025,1263,1290]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 205</figureCitation>
) evenly rounded; length of epandrial setulae variable, those along ventral margin shorter, sparse along lateral margins; cercal cavity nearly round, only slightly higher than wide; cerci in posterior view (
<figureCitation id="13432A243A057B05FCC4FAD354EAFAA8" box="[797,896,1335,1362]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 205</figureCitation>
) hemispherical, medial margin nearly straight; aedeagus in lateral view (
<figureCitation id="13432A243A057B05FE76FABF5567FA8F" box="[431,525,1371,1397]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 207</figureCitation>
) broadly rounded basally, irregularly rectangular, dorsal margin deeply excavated, excavation V-shaped, depth of cavity longer than width of opening, posterior margin broadly truncate, in ventral view (
<figureCitation id="13432A243A057B05FF04FA475654FA47" box="[221,318,1443,1469]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 206</figureCitation>
) more or less pentagonal, widest sub-basally, posterior margin tapered toward apical margin, apical margin truncate, basal margin concave; ventral aedeagal process in lateral view (
<figureCitation id="13432A243A057B05FBC0FA235312FA1B" box="[1049,1144,1479,1505]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 207</figureCitation>
) C-shaped; phallapodeme in lateral view (
<figureCitation id="13432A243A057B05FE93FA0F56C0F9FF" box="[330,426,1515,1541]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 207</figureCitation>
) very elongate, narrow, curved, keel evident as a medial thumb-like protuberance, dorsal length about equal to ventral length, in ventral view (
<figureCitation id="13432A243A057B05FD3BF9EB542BF9D3" box="[738,833,1551,1577]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 206</figureCitation>
) T-shaped, basal crossbar slightly less than half length of medial process, crossbar extensions slightly tapered, especially apically, curved posteriorly; postgonite in lateral view (
<figureCitation id="13432A243A057B05FF06F9B3562AF98B" box="[223,320,1623,1649]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 207</figureCitation>
) regularly, widely, and shallowly V-shaped, dorsal arm subrectangular, wider basally and anterior margin emarginate, ventral extension elongate, rod-like, parallel-sided, posterior apex tapered, thumb-like, narrow apex shallowly bifurcate, anteroventral corner distinctly produced, apex shallowly and narrowly bifurcate, bearing setulae on dorsal surface, in ventral view (
<figureCitation id="13432A243A057B05FDACF92755BCF927" box="[629,726,1731,1757]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 206</figureCitation>
) with base wide and excavated, irregularly V-shaped, posterior extensions oriented medially, narrow, thumb-like; neohypandrium in lateral view (
<figureCitation id="13432A243A057B05FBEBF90353F8F8FB" box="[1074,1170,1767,1793]" captionStart="FIGURES 205–207" captionStartId="86.[151,264,1013,1037]" captionTargetBox="[159,1427,189,981]" captionTargetId="figure-30@86.[151,1435,181,989]" captionTargetPageId="86" captionText="FIGURES 205–207. Male terminalia of Scatophila mystacina sp. nov. 205. Epandrium and cerci, posterior view. 206. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 207. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210939" httpUri="https://zenodo.org/record/13210939/files/figure.png" pageId="86" pageNumber="87">Fig. 207</figureCitation>
) narrow, parallel-sided, irregularly and deeply sinuous.
</paragraph>
<paragraph id="8BC736A13A057B05FF1EF8D4548EF823" blockId="86.[151,1437,1155,2010]" pageId="86" pageNumber="87">
Type Material.—
<materialsCitation id="3B103CFC3A057B05FE51F8CB548BF823" collectingDate="2007-07-30" collectionCode="UT, USNM" collectorName="D. &amp; W. N. Mathis" country="United States of America" elevation="1225" latitude="39.116665" location="The" longLatPrecision="928" longitude="-110.11" pageId="86" pageNumber="87" specimenCode="ENT 00250329" specimenCount="7" specimenCount-female="2" specimenCount-male="2" typeStatus="holotype">
<location id="8EA7607A3A057B05FE51F8CB56DEF8B3" LSID="urn:lsid:plazi:treatment:03D187B73A067B04FF4EFE525260F909:8EA7607A3A057B05FE51F8CB56DEF8B3" box="[392,436,1839,1865]" country="United States of America" latitude="39.116665" longLatPrecision="928" longitude="-110.11" name="The" pageId="86" pageNumber="87">The</location>
<typeStatus id="54C388033A057B05FE65F8CB5574F8B3" box="[444,542,1839,1865]" pageId="86" pageNumber="87" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C784D223A057B05FD5AF8CB541EF8B3" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[643,884,1839,1865]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="mystacina" status="sp. nov.">
<emphasis id="B90CEAB33A057B05FD5AF8CB541EF8B3" box="[643,884,1839,1865]" italics="true" pageId="86" pageNumber="87">Scatophila mystacina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57C83A057B05FCA2F8D45369F8B3" box="[891,1027,1839,1866]" pageId="86" pageNumber="87" rank="species">new species</taxonomicNameLabel>
is labeled “
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.
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. Grand: Green River (
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N;
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,
<geoCoordinate id="EE4C50663A057B05FE10F8B75554F894" box="[457,574,1875,1902]" degrees="110" direction="west" minutes="6.6" orientation="longitude" pageId="86" pageNumber="87" precision="92" value="-110.11">110°6.6'W</geoCoordinate>
;
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</quantity>
),
<date id="FFC610613A057B05FD63F8B7542AF897" box="[698,832,1875,1901]" pageId="86" pageNumber="87" value="2007-07-30">
<collectingDate id="EF82E9893A057B05FD63F8B7542AF897" box="[698,832,1875,1901]" pageId="86" pageNumber="87" value="2007-07-30">30 Jul 2007</collectingDate>
</date>
,
<collectorName id="268D53773A057B05FC92F8B75360F897" box="[843,1034,1875,1902]" pageId="86" pageNumber="87">D.&amp;W.N. Mathis</collectorName>
/
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<specimenCode id="DBDE9EDA3A057B05FBB2F8B05272F894" box="[1131,1304,1875,1902]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="86" pageNumber="87" type="Museum">ENT 00250329</specimenCode>
[plastic bar code label]/
<typeStatus id="54C388033A057B05FEC2F89356D8F86B" box="[283,434,1911,1937]" pageId="86" pageNumber="87" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D7EFD283A057B05FE65F89C56BBF868" box="[444,465,1912,1938]" count="1" pageId="86" pageNumber="87" type="male">♂</specimenCount>
<taxonomicName id="4C784D223A057B05FE02F8935359F86B" authority="Zatwarnicki &amp; Mathis USNM" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[475,1075,1911,1937]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="mystacina">
<emphasis id="B90CEAB33A057B05FE02F89355A5F86B" box="[475,719,1911,1937]" italics="true" pageId="86" pageNumber="87">Scatophila mystacina</emphasis>
Zatwarnicki &amp; Mathis USNM
</taxonomicName>
[red].” The
<typeStatus id="54C388033A057B05FB1EF8935243F86B" box="[1223,1321,1911,1937]" pageId="86" pageNumber="87" type="holotype">holotype</typeStatus>
is double mounted (minute nadel in block of plastic elastomere), is in excellent condition, and is deposited in the
<collectionCode id="ED69AE643A057B05FA9DF87F52F3F84F" box="[1348,1433,1947,1973]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="86" pageNumber="87" type="Museum">USNM</collectionCode>
.
<specimenCount id="9D7EFD283A057B05FF4EF85B5620F820" box="[151,330,1983,2010]" count="3" pageId="86" pageNumber="87" type="generic" typeStatus="paratype">Three paratypes</specimenCount>
(
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,
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;
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) bear the same label data as the holotype
</materialsCitation>
.
</paragraph>
<caption id="DF0766293A047B04FF4EFA9754FDFA55" ID-DOI="http://doi.org/10.5281/zenodo.13210941" ID-Zenodo-Dep="13210941" httpUri="https://zenodo.org/record/13210941/files/figure.png" pageId="87" pageNumber="88" startId="87.[151,264,1395,1419]" targetBox="[171,1435,181,1371]" targetPageId="87" targetType="figure">
<paragraph id="8BC736A13A047B04FF4EFA9754FDFA55" blockId="87.[151,1437,1395,1455]" pageId="87" pageNumber="88">
<emphasis id="B90CEAB33A047B04FF4EFA975603FA71" bold="true" box="[151,361,1395,1419]" pageId="87" pageNumber="88">FIGURES 208–211</emphasis>
. Head of
<taxonomicName id="4C784D223A047B04FE0EFA9755DEFA71" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[471,692,1395,1419]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="mystacina" status="sp. nov.">
<emphasis id="B90CEAB33A047B04FE0EFA9755DEFA71" box="[471,692,1395,1419]" italics="true" pageId="87" pageNumber="88">Scatophila mystacina</emphasis>
</taxonomicName>
<emphasis id="B90CEAB33A047B04FD64FA97547BFA71" bold="true" box="[701,785,1395,1419]" pageId="87" pageNumber="88">
<taxonomicNameLabel id="A23F57C83A047B04FD64FA97547BFA71" box="[701,785,1395,1419]" pageId="87" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
208. Male, latero-oblique view. 209. Male, lateral view. 210. Female, latero-oblique view. 211. Female, lateral view. Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A047B04FF1EF9E6524DF9E1" blockId="87.[151,1436,1537,1780]" box="[199,1319,1537,1564]" pageId="87" pageNumber="88">
<materialsCitation id="3B103CFC3A047B04FF1EF9E6524EF9E1" box="[199,1316,1537,1564]" country="United States of America" elevation="1225" latitude="39.116665" location="Green River" longLatPrecision="928" longitude="-110.11" municipality="Grand" pageId="87" pageNumber="88" specimenCount="1" stateProvince="Utah" typeStatus="holotype">
<typeStatus id="54C388033A047B04FF1EF9E65795F9E6" box="[199,255,1538,1564]" pageId="87" pageNumber="88">Type</typeStatus>
Locality.—
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.
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(
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N;
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,
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.
</paragraph>
<paragraph id="8BC736A13A047B04FF1EF9C1542CF9C5" blockId="87.[151,1436,1537,1780]" box="[199,838,1573,1599]" pageId="87" pageNumber="88">
Distribution (
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).—
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<taxonomicName id="4C784D223A047B04FE30F9C25521F9C5" box="[489,587,1574,1599]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
:
</emphasis>
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(
<collectingRegion id="49BCF8433A047B04FD26F9C15450F9C5" box="[767,826,1573,1599]" country="United States of America" name="Utah" pageId="87" pageNumber="88">Utah</collectingRegion>
).
</paragraph>
<paragraph id="8BC736A13A047B04FF1EF9AD54C8F97D" blockId="87.[151,1436,1537,1780]" pageId="87" pageNumber="88">
Etymology.—The species epithet,
<emphasis id="B90CEAB33A047B04FD95F9AE55AFF999" box="[588,709,1610,1635]" italics="true" pageId="87" pageNumber="88">
<taxonomicName id="4C784D223A047B04FD95F9AE55AAF999" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[588,704,1610,1635]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="mystacina">mystacina</taxonomicName>
,
</emphasis>
is of Greek derivation and means “mustache” and has reference to the dense patch of setae at the apex of the ventral facial protrusion.
</paragraph>
<paragraph id="8BC736A13A047B04FF1EF9755260F909" blockId="87.[151,1436,1537,1780]" pageId="87" pageNumber="88">
Remarks.—This species is placed in the
<emphasis id="B90CEAB33A047B04FD5EF975544BF951" box="[647,801,1681,1707]" italics="true" pageId="87" pageNumber="88">quadriguttata</emphasis>
species group, where it is similar and closely related to
<emphasis id="B90CEAB33A047B04FA5EF9765791F935" italics="true" pageId="87" pageNumber="88">
<taxonomicName id="4C784D223A047B04FA5EF976579DF935" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="conifera">S. conifera</taxonomicName>
,
</emphasis>
differing by the more pronounced ventral facial protrusion of the male that bears a dense patch of curved, black setulae, and by structures of the male terminalia as noted in the key and respective descriptions.
</paragraph>
</subSubSection>
</treatment>
</document>

630
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<mods:title id="23C9EC12DB512CBACCD139E22370A876">Revision of the Nearctic Species of the Shore-Fly Genus Scatophila Becker (Diptera: Ephydridae)</mods:title>
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<mods:affiliation id="7EA7E07409AC124EF626480301094B76">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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<emphasis id="B90CEAB33A637B63FF4EFC12556DFBEB" bold="true" box="[151,519,1014,1041]" pageId="48" pageNumber="49">
11.
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<emphasis id="B90CEAB33A637B63FF1BFC1256C8FBEB" bold="true" box="[194,418,1014,1041]" italics="true" pageId="48" pageNumber="49">Scatophila bolwigi</emphasis>
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<paragraph id="8BC736A13A637B63FF4EFBFF56DEFBCF" blockId="48.[151,519,1014,1077]" box="[151,436,1051,1077]" pageId="48" pageNumber="49">
<figureCitation id="13432A243A637B63FF4EFBFF579EFBCF" box="[151,244,1051,1077]" captionStart="FIGURES 89–91" captionStartId="46.[151,264,1944,1968]" captionTargetBox="[151,1433,1140,1920]" captionTargetId="figure-70@46.[151,1435,1140,1920]" captionTargetPageId="46" captionText="FIGURES 89–91. Head, thorax and wing of Scatophila. 89. Scatophila viridella Sturtevant &amp; Wheeler, head and thorax, dorsal view. 90–91. Wing, dorsal view. 90. S. bolwigi sp. nov. 91. S. hirtirostris Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210835" httpUri="https://zenodo.org/record/13210835/files/figure.png" pageId="48" pageNumber="49">Figs. 90</figureCitation>
,
<figureCitation id="13432A243A637B63FF26FBFF5622FBCF" box="[255,328,1051,1077]" captionStart-0="FIGURES 93–96" captionStart-1="FIGURES 97–99" captionStartId-0="49.[151,264,1642,1666]" captionStartId-1="50.[151,264,1195,1219]" captionTargetBox-0="[171,1435,387,1618]" captionTargetBox-1="[159,1427,425,1164]" captionTargetId-0="figure-109@49.[151,1435,381,1618]" captionTargetId-1="figure-153@50.[151,1435,417,1172]" captionTargetPageId-0="49" captionTargetPageId-1="50" captionText-0="FIGURES 93–96. Head of Scatophila bolwigi sp. nov. 93. Male, latero-oblique view. 94. Male, lateral view. 95. Female, laterooblique view. 96. Female, lateral view. Scale bar = 0.2 mm." captionText-1="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13210839" figureDoi-1="http://doi.org/10.5281/zenodo.13210841" httpUri-0="https://zenodo.org/record/13210839/files/figure.png" httpUri-1="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">93–99</figureCitation>
,
<figureCitation id="13432A243A637B63FE8AFBFF5615FBCF" box="[339,383,1051,1077]" captionStart="FIGURES 107–109" captionStartId="53.[151,264,947,971]" captionTargetBox="[151,1435,181,923]" captionTargetId="figure-27@53.[151,1435,181,923]" captionTargetPageId="53" captionText="FIGURES 107–109. Head, thorax and wing of Scatophila. 107. Scatophila bolwigi sp. nov., head and thorax, dorsal view. 108–109. Wing, dorsal view. 108. S. tuberculosa Cresson. 109. S. unicornis Czerny. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210847" httpUri="https://zenodo.org/record/13210847/files/figure.png" pageId="48" pageNumber="49">107</figureCitation>
,
<figureCitation id="13432A243A637B63FE52FBFF56DEFBCF" box="[395,436,1051,1077]" captionStart="FIGURE 110" captionStartId="54.[151,250,1228,1252]" captionTargetBox="[151,1435,584,1204]" captionTargetId="figure-199@54.[151,1435,584,1204]" captionTargetPageId="54" captionText="FIGURE 110. Distribution map of Scatophila bolwigi sp. nov. (squares), S. tuberculosa Cresson (triangles), and S. variofacialis Sturtevant &amp; Wheeler (circles)." figureDoi="http://doi.org/10.5281/zenodo.13210849" httpUri="https://zenodo.org/record/13210849/files/figure.png" pageId="48" pageNumber="49">110</figureCitation>
</paragraph>
<paragraph id="8BC736A13A637B63FF4EFB875566FB5B" blockId="48.[151,1437,1123,2013]" pageId="48" pageNumber="49">
Diagnosis.—This species is distinguished from congeners by the following combination of characters: Small shore flies, body length
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.
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<emphasis id="B90CEAB33A637B63FF1EFB4F566EFB3F" box="[199,260,1195,1221]" italics="true" pageId="48" pageNumber="49">Head</emphasis>
(
<figureCitation id="13432A243A637B63FECDFB4F56CAFB3F" box="[276,416,1195,1221]" captionStart="FIGURES 93–96" captionStartId="49.[151,264,1642,1666]" captionTargetBox="[171,1435,387,1618]" captionTargetId="figure-109@49.[151,1435,381,1618]" captionTargetPageId="49" captionText="FIGURES 93–96. Head of Scatophila bolwigi sp. nov. 93. Male, latero-oblique view. 94. Male, lateral view. 95. Female, laterooblique view. 96. Female, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210839" httpUri="https://zenodo.org/record/13210839/files/figure.png" pageId="48" pageNumber="49">Figs. 93–96</figureCitation>
): Mesofrons and fronto-orbits tan to brown. Face of
<specimenCount id="9D7EFD283A637B63FC21FB4F5367FB3F" box="[1016,1037,1195,1221]" count="1" pageId="48" pageNumber="49" type="male">♂</specimenCount>
brown, ventral portion protuberant with an elongate, medial depression bordered by dense curly setulae, especially laterally; face of
<specimenCount id="9D7EFD283A637B63FADFFB2B5271FB13" box="[1286,1307,1231,1257]" count="1" pageId="48" pageNumber="49" type="female">♀</specimenCount>
with some subshiny luster. Gena moderately short, slightly more than half height of basal flagellomere; gena-to-eye ratio 0.25–0.27.
</paragraph>
<paragraph id="8BC736A13A637B63FF1EFADF5347F9F3" blockId="48.[151,1437,1123,2013]" pageId="48" pageNumber="49">
<emphasis id="B90CEAB33A637B63FF1EFADF5672FAAF" box="[199,280,1339,1365]" italics="true" pageId="48" pageNumber="49">Thorax</emphasis>
(
<figureCitation id="13432A243A637B63FEFDFADF5615FAAF" box="[292,383,1339,1365]" captionStart="FIGURES 89–91" captionStartId="46.[151,264,1944,1968]" captionTargetBox="[151,1433,1140,1920]" captionTargetId="figure-70@46.[151,1435,1140,1920]" captionTargetPageId="46" captionText="FIGURES 89–91. Head, thorax and wing of Scatophila. 89. Scatophila viridella Sturtevant &amp; Wheeler, head and thorax, dorsal view. 90–91. Wing, dorsal view. 90. S. bolwigi sp. nov. 91. S. hirtirostris Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210835" httpUri="https://zenodo.org/record/13210835/files/figure.png" pageId="48" pageNumber="49">Figs.90</figureCitation>
,
<figureCitation id="13432A243A637B63FE5EFADF56DFFAAF" box="[391,437,1339,1365]" captionStart="FIGURES 107–109" captionStartId="53.[151,264,947,971]" captionTargetBox="[151,1435,181,923]" captionTargetId="figure-27@53.[151,1435,181,923]" captionTargetPageId="53" captionText="FIGURES 107–109. Head, thorax and wing of Scatophila. 107. Scatophila bolwigi sp. nov., head and thorax, dorsal view. 108–109. Wing, dorsal view. 108. S. tuberculosa Cresson. 109. S. unicornis Czerny. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210847" httpUri="https://zenodo.org/record/13210847/files/figure.png" pageId="48" pageNumber="49">107</figureCitation>
): Mesonotum (
<figureCitation id="13432A243A637B63FD81FADF55DEFAAF" box="[600,692,1339,1365]" captionStart="FIGURES 107–109" captionStartId="53.[151,264,947,971]" captionTargetBox="[151,1435,181,923]" captionTargetId="figure-27@53.[151,1435,181,923]" captionTargetPageId="53" captionText="FIGURES 107–109. Head, thorax and wing of Scatophila. 107. Scatophila bolwigi sp. nov., head and thorax, dorsal view. 108–109. Wing, dorsal view. 108. S. tuberculosa Cresson. 109. S. unicornis Czerny. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210847" httpUri="https://zenodo.org/record/13210847/files/figure.png" pageId="48" pageNumber="49">Fig.107</figureCitation>
) mostly brown with grayish stripes; scutum mostly microtomentose, dull to subshiny brown with pattern of linear gray stripes anterior, between acrostichal tracks and laterad of acrostichal tracks, fainter linear spots laterad of dorsocentral tracks; presutural dorsocentral setae lacking. Wing (
<figureCitation id="13432A243A637B63FF79FA435799FA3B" box="[160,243,1447,1473]" captionStart="FIGURES 89–91" captionStartId="46.[151,264,1944,1968]" captionTargetBox="[151,1433,1140,1920]" captionTargetId="figure-70@46.[151,1435,1140,1920]" captionTargetPageId="46" captionText="FIGURES 89–91. Head, thorax and wing of Scatophila. 89. Scatophila viridella Sturtevant &amp; Wheeler, head and thorax, dorsal view. 90–91. Wing, dorsal view. 90. S. bolwigi sp. nov. 91. S. hirtirostris Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210835" httpUri="https://zenodo.org/record/13210835/files/figure.png" pageId="48" pageNumber="49">Fig. 90</figureCitation>
) generally faint with moderate pattern of white spots; veins R
<subScript id="17FC34E43A637B63FC64FA5D54BCFA3D" attach="left" box="[957,982,1465,1479]" fontSize="6" pageId="48" pageNumber="49">2+3</subScript>
and R
<subScript id="17FC34E43A637B63FBFAFA5D5357FA3D" attach="left" box="[1059,1085,1465,1479]" fontSize="6" pageId="48" pageNumber="49">4+5</subScript>
shallowly undulate near white spots; costal vein ratio 0.29–0.30; M
<subScript id="17FC34E43A637B63FDE5FA39552EFA11" attach="left" box="[572,580,1501,1515]" fontSize="6" pageId="48" pageNumber="49">1</subScript>
vein ratio 0.56–0.66. Legs dark; male midfemur lacking row of spine-like setulae along posteroventral margin Tarsi often yellowish to brownish red basally.
</paragraph>
<paragraph id="8BC736A13A637B62FF1EF9F75373FE9F" blockId="48.[151,1437,1123,2013]" lastBlockId="49.[151,1437,151,357]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="B90CEAB33A637B63FF1EF9F7565BF9D7" box="[199,305,1555,1581]" italics="true" pageId="48" pageNumber="49">Abdomen</emphasis>
(
<figureCitation id="13432A243A637B63FE9AF9F756BBF9D7" box="[323,465,1555,1581]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Figs. 97–99</figureCitation>
): Mostly blackish brown to black, subshiny to shiny, with some blackish gray areas basally on tergites 2–4; subshiny, male tergite 5 truncate posteriorly. Male terminalia (
<figureCitation id="13432A243A637B63FBBAF9D35385F9AB" box="[1123,1263,1591,1617]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Figs. 97–99</figureCitation>
): Epandrium in posterior view (
<figureCitation id="13432A243A637B63FE95F9BF56F5F98F" box="[332,415,1627,1653]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Fig. 97</figureCitation>
) generally squatly oval except for broadly stepped emargination at ventral margin, width almost equal to height, lateral margins shallowly curved with short, papilla-like projection at ventrolateral corner; dorsal margin of epandrium in posterior view (
<figureCitation id="13432A243A637B63FD7EF9475593F947" box="[679,761,1699,1725]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Fig. 97</figureCitation>
) evenly rounded; length of epandrial setulae variable, those along ventral margin shorter, sparse along lateral margins; cercal cavity nearly round, only slightly higher than wide; cerci in posterior view (
<figureCitation id="13432A243A637B63FE30F90F5551F8FF" box="[489,571,1771,1797]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Fig. 97</figureCitation>
) hemispherical, medial margin nearly straight; aedeagus in lateral view (
<figureCitation id="13432A243A637B63FAB6F90F57DEF8D3" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Fig. 99</figureCitation>
) broadly rounded basally, irregularly rectangular, dorsal margin deeply excavated, excavation V-shaped, depth of cavity longer than width of opening, posterior margin broadly truncate, in ventral view (
<figureCitation id="13432A243A637B63FB72F8D75394F8B7" box="[1195,1278,1843,1869]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Fig. 98</figureCitation>
) more or less pentagonal, widest sub-basally, posterior margin tapered toward apical margin, apical margin truncate, basal margin concave; ventral aedeagal process in lateral view (
<figureCitation id="13432A243A637B63FD0BF89F544FF86F" box="[722,805,1915,1941]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Fig. 99</figureCitation>
) C-shaped; phallapodeme in lateral view (
<figureCitation id="13432A243A637B63FADEF89F5230F86F" box="[1287,1370,1915,1941]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Fig. 99</figureCitation>
) very elongate, narrow, curved, keel evident as a medial thumb-like protuberance, dorsal length about equal to ventral length, in ventral view (
<figureCitation id="13432A243A637B63FE7FF8275692F827" box="[422,504,1987,2013]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="48" pageNumber="49">Fig. 98</figureCitation>
) T-shaped, basal crossbar slightly less than half length of medial process, crossbar extensions slightly tapered, especially apically, curved posteriorly; postgonite in lateral view (
<figureCitation id="13432A243A627B62FB1BFF73527FFF4B" box="[1218,1301,151,177]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="49" pageNumber="50">Fig. 99</figureCitation>
) irregularly and widely V-shaped, dorsal arm subrectangular, wider basally and anterior margin emarginate, ventral extension an elongate, rod-like, parallel-sided projection, posterior apex a tapered, thumb-like projection, narrow apex shallowly bifurcate, posterodorsal corner distinctly produced, apex shallowly and narrowly bifurcate, bearing setulae on dorsal surface, in ventral view (
<figureCitation id="13432A243A627B62FE69FEC35568FEBB" box="[432,514,295,321]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="49" pageNumber="50">Fig. 98</figureCitation>
) with posterior extensions oriented medially, narrow, thumb-like; neohypandrium in lateral view (
<figureCitation id="13432A243A627B62FE9EFEAF56F2FE9F" box="[327,408,331,357]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="49" pageNumber="50">Fig. 99</figureCitation>
) narrow, parallel-sided, irregularly and deeply C-shaped.
</paragraph>
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<paragraph id="8BC736A13A627B62FF4EF98E5598F95C" blockId="49.[151,1436,1642,1702]" pageId="49" pageNumber="50">
<emphasis id="B90CEAB33A627B62FF4EF98E5626F978" bold="true" box="[151,332,1642,1666]" pageId="49" pageNumber="50">FIGURES 93–96</emphasis>
. Head of
<taxonomicName id="4C784D223A627B62FE77F98E5501F978" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[430,619,1642,1666]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="bolwigi" status="sp. nov.">
<emphasis id="B90CEAB33A627B62FE77F98E5501F978" box="[430,619,1642,1666]" italics="true" pageId="49" pageNumber="50">Scatophila bolwigi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A23F57C83A627B62FDA8F98E55AAF978" box="[625,704,1642,1666]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
93. Male, latero-oblique view. 94. Male, lateral view. 95. Female, laterooblique view. 96. Female, lateral view. Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A627B61FF1EF91D5332FE73" blockId="49.[151,1437,1784,2027]" lastBlockId="50.[151,1437,150,393]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
Type Material.—
<materialsCitation id="3B103CFC3A627B61FE5FF91C53BDFF4B" collectingDate="1917-09-25" collectingDateMax="2011-08-07" collectingDateMin="1917-09-25" collectionCode="USNM" collectorName="The" country="United States of America" county="Gulkana River" lastPageId="50" lastPageNumber="51" latitude="60.791668" location="Twitchell" longLatPrecision="102" longitude="-161.755" municipality="Census Area" pageId="49" pageNumber="50" specimenCode="ENT 00250327" specimenCount="15" specimenCount-female="1" specimenCount-male="10" stateProvince="D" typeStatus="holotype">
<collectorName id="268D53773A627B62FE5FF91C56D8F8E8" box="[390,434,1784,1810]" pageId="49" pageNumber="50">The</collectorName>
<typeStatus id="54C388033A627B62FE61F91C5570F8E8" box="[440,538,1784,1810]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C784D223A627B62FDA0F91C5420F8E8" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[633,842,1784,1810]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="bolwigi" status="sp. nov.">
<emphasis id="B90CEAB33A627B62FDA0F91C5420F8E8" box="[633,842,1784,1810]" italics="true" pageId="49" pageNumber="50">Scatophila bolwigi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57C83A627B62FC89F91D54BFF8E8" box="[848,981,1784,1811]" pageId="49" pageNumber="50" rank="species">new species</taxonomicNameLabel>
is labeled “
<collectingCountry id="F36F76313A627B62FB8EF91D53F9F8E9" box="[1111,1171,1785,1811]" name="United States of America" pageId="49" pageNumber="50">USA</collectingCountry>
. A[LAS]K[A].
<collectingCounty id="62A64E2D3A627B62FAE4F91C57BFF8CC" pageId="49" pageNumber="50">Gulkana River</collectingCounty>
(
<locationDeviation id="9A752F833A627B62FF3EF8F956B3F8CC" box="[231,473,1820,1847]" location="Glenallen" pageId="49" pageNumber="50">
<quantity id="4C809B443A627B62FF3EF8F9562CF8CC" box="[231,326,1820,1847]" metricMagnitude="4" metricUnit="m" metricValue="1.93" pageId="49" pageNumber="50" unit="km" value="19.3">19.3 km</quantity>
N Glenallen
</locationDeviation>
[should be Glennallen];
<geoCoordinate id="EE4C50663A627B62FD21F8F85406F8CC" box="[760,876,1820,1846]" degrees="62" direction="north" minutes="16.1" orientation="latitude" pageId="49" pageNumber="50" precision="92" value="62.268333">62°16.1'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A627B62FCA0F8F85494F8CD" box="[889,1022,1820,1847]" degrees="145" direction="west" minutes="23.1" orientation="longitude" pageId="49" pageNumber="50" precision="92" value="-145.385">145°23.1'W</geoCoordinate>
)[,]
<date id="FFC610613A627B62FBF2F8F953DCF8CD" box="[1067,1206,1821,1847]" pageId="49" pageNumber="50" value="2011-08-07">
<collectingDate id="EF82E9893A627B62FBF2F8F953DCF8CD" box="[1067,1206,1821,1847]" pageId="49" pageNumber="50" value="2011-08-07">7 Aug 2011</collectingDate>
</date>
,
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. &amp;
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.
<collectingRegion id="49BCF8433A627B62FAFBF8F95253F8CD" box="[1314,1337,1821,1847]" country="United States of America" name="N" pageId="49" pageNumber="50">N</collectingRegion>
. Mathis/ Sp. photographed
<collectingRegion id="49BCF8433A627B62FEBCF8A5561DF8A1" box="[357,375,1857,1883]" country="United States of America" name="T" pageId="49" pageNumber="50">T</collectingRegion>
. Zatwarnicki,
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. Cielniak, 2018/
<collectionCode id="ED69AE643A627B62FD36F8A5542AF8A1" box="[751,832,1857,1883]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="49" pageNumber="50" type="Museum">USNM</collectionCode>
<specimenCode id="DBDE9EDA3A627B62FC91F8A5549CF8A1" box="[840,1014,1856,1883]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="49" pageNumber="50" type="Museum">ENT 00250327</specimenCode>
[plastic bar code label]/
<typeStatus id="54C388033A627B62FAD0F8A552F7F8A1" box="[1289,1437,1857,1883]" pageId="49" pageNumber="50" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D7EFD283A627B62FF4EF88157C6F885" box="[151,172,1893,1919]" count="1" pageId="49" pageNumber="50" type="male">♂</specimenCount>
<taxonomicName id="4C784D223A627B62FF6EF880559DF885" authority="Zatwarnicki &amp; Mathis USNM" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[183,759,1892,1919]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="bolwigi">
<emphasis id="B90CEAB33A627B62FF6EF88056E4F884" box="[183,398,1892,1918]" italics="true" pageId="49" pageNumber="50">Scatophila bolwigi</emphasis>
Zatwarnicki &amp; Mathis USNM
</taxonomicName>
[red].” The
<typeStatus id="54C388033A627B62FC54F8805485F884" box="[909,1007,1892,1918]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
is double mounted (minute nadel in block of plastic elastomere), is in excellent condition, and is deposited in the
<collectingRegion id="49BCF8433A627B62FBC4F86D5318F859" box="[1053,1138,1929,1955]" country="United States of America" name="USNM" pageId="49" pageNumber="50">USNM</collectingRegion>
.
<specimenCount id="9D7EFD283A627B62FBA6F86D5242F859" box="[1151,1320,1929,1955]" count="4" pageId="49" pageNumber="50" type="generic" typeStatus="paratype">Four paratypes</specimenCount>
(
<specimenCount id="9D7EFD283A627B62FAE5F86D520BF859" box="[1340,1377,1929,1955]" count="3" pageId="49" pageNumber="50" type="male">3♂</specimenCount>
,
<specimenCount id="9D7EFD283A627B62FAA9F86D52FBF859" box="[1392,1425,1929,1955]" count="1" pageId="49" pageNumber="50" type="female">1♀</specimenCount>
) bear the same label data as the holotype and five other
<typeStatus id="54C388033A627B62FCDCF849541AF83D" box="[773,880,1965,1991]" pageId="49" pageNumber="50" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D7EFD283A627B62FC58F84854CFF83D" box="[897,933,1964,1991]" count="5" pageId="49" pageNumber="50" type="male">5♂</specimenCount>
) bear the same locality label, but the date of collection is
<collectingDate id="EF82E9893A627B62FEF3F835550FF811" box="[298,613,2000,2027]" isEnumeration="true" pageId="49" pageNumber="50">
<date id="FFC610613A627B62FEF3F83556C8F810" box="[298,418,2000,2027]" pageId="49" pageNumber="50" value="2006-07-09">9 Jul 2006</date>
and
<date id="FFC610613A627B62FE02F835550FF811" box="[475,613,2001,2027]" pageId="49" pageNumber="50" value="2012-08-07">7 Aug 2012</date>
</collectingDate>
. Other
<typeStatus id="54C388033A627B62FD60F835544EF811" box="[697,804,2001,2027]" pageId="49" pageNumber="50" type="paratype">paratypes</typeStatus>
are as follows:
<emphasis id="B90CEAB33A627B62FC04F8355220F811" box="[989,1354,2000,2027]" italics="true" pageId="49" pageNumber="50">
<collectingRegion id="49BCF8433A627B62FC04F835532EF811" box="[989,1092,2001,2027]" country="United States of America" name="Alaska" pageId="49" pageNumber="50">ALASKA</collectingRegion>
. Bethel (
<collectingMunicipality id="6BA3ACDB3A627B62FB70F8355256F811" box="[1193,1340,2001,2027]" pageId="49" pageNumber="50">Census Area</collectingMunicipality>
):
</emphasis>
Bethel (
<geoCoordinate id="EE4C50663A617B61FF46FF72567EFF4B" box="[159,276,150,177]" degrees="60" direction="north" minutes="47.5" orientation="latitude" pageId="50" pageNumber="51" precision="92" value="60.791668">60°47.5'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A617B61FEF9FF7256C3FF4B" box="[288,425,150,177]" degrees="161" direction="west" minutes="45.3" orientation="longitude" pageId="50" pageNumber="51" precision="92" value="-161.755">161°45.3'W</geoCoordinate>
),
<date id="FFC610613A617B61FE64FF725521FF4B" box="[445,587,150,177]" pageId="50" pageNumber="51" value="1917-09-25">
<collectingDate id="EF82E9893A617B61FE64FF725521FF4B" box="[445,587,150,177]" pageId="50" pageNumber="51" value="1917-09-25">25 Sep 1917</collectingDate>
</date>
,
<collectingRegion id="49BCF8433A617B61FD8CFF735506FF4B" box="[597,620,151,177]" country="United States of America" name="A" pageId="50" pageNumber="51">A</collectingRegion>
.
<collectingRegion id="49BCF8433A617B61FDAEFF7355E4FF4B" box="[631,654,151,177]" country="United States of America" name="H" pageId="50" pageNumber="51">H</collectingRegion>
.
<location id="8EA7607A3A617B61FD41FF73546BFF4B" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FD41FF73546BFF4B" box="[664,769,151,177]" country="United States of America" county="Gulkana River" latitude="60.791668" longLatPrecision="102" longitude="-161.755" municipality="Census Area" name="Twitchell" pageId="50" pageNumber="51" stateProvince="D">Twitchell</location>
(
<specimenCount id="9D7EFD283A617B61FCCBFF73545FFF4B" box="[786,821,151,177]" count="1" pageId="50" pageNumber="51" type="male">1♂</specimenCount>
<typeStatus id="54C388033A617B61FCE2FF7354F1FF4B" box="[827,923,151,177]" pageId="50" pageNumber="51" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C784D223A617B61FC18FF735301FF4B" box="[961,1131,151,177]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="variofacialis">
<emphasis id="B90CEAB33A617B61FC18FF735301FF4B" box="[961,1131,151,177]" italics="true" pageId="50" pageNumber="51">S. variofacialis</emphasis>
</taxonomicName>
;
<collectionCode id="ED69AE643A617B61FBA0FF7353A5FF4B" box="[1145,1231,151,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="50" pageNumber="51" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103CFC3A617B61FB38FF7352F3FF2F" collectingDate="2003-08-10" collectionCode="USNM" country="United States of America" county="North Star" latitude="64.793335" location="Mathis" longLatPrecision="100" longitude="-147.19" municipality="Chena Lake Recreation Area" pageId="50" pageNumber="51" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="D" typeStatus="holotype">
<emphasis id="B90CEAB33A617B61FB38FF7357A5FF2F" italics="true" pageId="50" pageNumber="51">
Fairbanks
<collectingCounty id="62A64E2D3A617B61FA82FF7357A3FF2F" pageId="50" pageNumber="51">North Star</collectingCounty>
:
</emphasis>
<collectingMunicipality id="6BA3ACDB3A617B61FF0FFF5F5572FF2F" box="[214,536,187,213]" pageId="50" pageNumber="51">Chena Lake Recreation Area</collectingMunicipality>
(
<geoCoordinate id="EE4C50663A617B61FDFEFF5F55F7FF2F" box="[551,669,187,213]" degrees="64" direction="north" minutes="47.6" orientation="latitude" pageId="50" pageNumber="51" precision="92" value="64.793335">64°47.6'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A617B61FD71FF5F545AFF2F" box="[680,816,187,213]" degrees="147" direction="west" minutes="11.4" orientation="longitude" pageId="50" pageNumber="51" precision="92" value="-147.19">147°11.4'W</geoCoordinate>
),
<date id="FFC610613A617B61FC9DFF5F54BDFF2F" box="[836,983,187,213]" pageId="50" pageNumber="51" value="2003-08-10">
<collectingDate id="EF82E9893A617B61FC9DFF5F54BDFF2F" box="[836,983,187,213]" pageId="50" pageNumber="51" value="2003-08-10">10 Aug 2003</collectingDate>
</date>
,
<collectingRegion id="49BCF8433A617B61FC3BFF5F5493FF2F" box="[994,1017,187,213]" country="United States of America" name="D" pageId="50" pageNumber="51">D</collectingRegion>
. and
<collectingRegion id="49BCF8433A617B61FBEAFF5F5325FF2F" box="[1075,1103,187,213]" country="United States of America" name="W" pageId="50" pageNumber="51">W</collectingRegion>
.
<collectingRegion id="49BCF8433A617B61FB81FF5F5305FF2F" box="[1112,1135,187,213]" country="United States of America" name="N" pageId="50" pageNumber="51">N</collectingRegion>
.
<location id="8EA7607A3A617B61FBA3FF5F53A2FF2F" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FBA3FF5F53A2FF2F" box="[1146,1224,187,213]" country="United States of America" county="North Star" latitude="64.793335" longLatPrecision="100" longitude="-147.19" municipality="Chena Lake Recreation Area" name="Mathis" pageId="50" pageNumber="51" stateProvince="D">Mathis</location>
(
<specimenCount id="9D7EFD283A617B61FB01FF5F5394FF2F" box="[1240,1278,187,213]" count="4" pageId="50" pageNumber="51" type="male">4♂</specimenCount>
,
<specimenCount id="9D7EFD283A617B61FAD0FF5F5244FF2F" box="[1289,1326,187,213]" count="2" pageId="50" pageNumber="51" type="female">2♀</specimenCount>
;
<collectionCode id="ED69AE643A617B61FAE2FF5F52FBFF2F" box="[1339,1425,187,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="50" pageNumber="51" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103CFC3A617B61FF4EFF3B5261FF03" box="[151,1291,222,249]" collectingDate="2002-08-02" collectionCode="USNM" collectorName="Kenai Peninsula &amp; Homer" country="United States of America" latitude="59.646667" location="Mathis" longLatPrecision="103" longitude="-151.525" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="D" typeStatus="holotype">
<emphasis id="B90CEAB33A617B61FF4EFF3B5636FF03" box="[151,348,223,249]" italics="true" pageId="50" pageNumber="51">
<collectorName id="268D53773A617B61FF4EFF3B563CFF03" box="[151,342,223,249]" pageId="50" pageNumber="51">Kenai Peninsula</collectorName>
:
</emphasis>
<collectorName id="268D53773A617B61FEBCFF3B56D9FF03" box="[357,435,223,249]" pageId="50" pageNumber="51">Homer</collectorName>
(
<geoCoordinate id="EE4C50663A617B61FE1AFF3A5552FF03" box="[451,568,222,249]" degrees="59" direction="north" minutes="38.8" orientation="latitude" pageId="50" pageNumber="51" precision="92" value="59.646667">59°38.8'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A617B61FD9CFF3A55A5FF03" box="[581,719,222,249]" degrees="151" direction="west" minutes="31.5" orientation="longitude" pageId="50" pageNumber="51" precision="92" value="-151.525">151°31.5'W</geoCoordinate>
),
<date id="FFC610613A617B61FD3DFF3B5407FF03" box="[740,877,223,249]" pageId="50" pageNumber="51" value="2002-08-02">
<collectingDate id="EF82E9893A617B61FD3DFF3B5407FF03" box="[740,877,223,249]" pageId="50" pageNumber="51" value="2002-08-02">2 Aug 2002</collectingDate>
</date>
,
<collectingRegion id="49BCF8433A617B61FCA3FF3B54FBFF03" box="[890,913,223,249]" country="United States of America" name="D" pageId="50" pageNumber="51">D</collectingRegion>
. and
<collectingRegion id="49BCF8433A617B61FC17FF3B5480FF03" box="[974,1002,223,249]" country="United States of America" name="W" pageId="50" pageNumber="51">W</collectingRegion>
.
<collectingRegion id="49BCF8433A617B61FC2FFF3B5367FF03" box="[1014,1037,223,249]" country="United States of America" name="N" pageId="50" pageNumber="51">N</collectingRegion>
.
<location id="8EA7607A3A617B61FBC0FF3B530DFF03" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FBC0FF3B530DFF03" box="[1049,1127,223,249]" country="United States of America" latitude="59.646667" longLatPrecision="103" longitude="-151.525" name="Mathis" pageId="50" pageNumber="51" stateProvince="D">Mathis</location>
(
<specimenCount id="9D7EFD283A617B61FBA3FF3B53F4FF03" box="[1146,1182,223,249]" count="1" pageId="50" pageNumber="51" type="male">1♂</specimenCount>
;
<collectionCode id="ED69AE643A617B61FB74FF3B5269FF03" box="[1197,1283,223,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="50" pageNumber="51" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103CFC3A617B61FACFFF3B5223FEE7" collectingDate="2002-08-07" collectionCode="USNM" collectorName="Matanuska-Susitna &amp; Eklutna &amp; Knik Arm" country="United States of America" latitude="61.47" location="Mathis" longLatPrecision="102" longitude="-149.35667" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="D" typeStatus="holotype">
<emphasis id="B90CEAB33A617B61FACFFF3B5799FEE7" italics="true" pageId="50" pageNumber="51">
<collectorName id="268D53773A617B61FACFFF3B5786FEE7" pageId="50" pageNumber="51">Matanuska-Susitna</collectorName>
:
</emphasis>
<collectorName id="268D53773A617B61FF24FEE7563EFEE7" box="[253,340,259,285]" pageId="50" pageNumber="51">Eklutna</collectorName>
(
<collectorName id="268D53773A617B61FEBFFEE7568AFEE7" box="[358,480,259,285]" pageId="50" pageNumber="51">Knik Arm</collectorName>
;
<geoCoordinate id="EE4C50663A617B61FE36FEE75509FEE7" box="[495,611,259,285]" degrees="61" direction="north" minutes="28.2" orientation="latitude" pageId="50" pageNumber="51" precision="92" value="61.47">61°28.2'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A617B61FDABFEE75591FEE7" box="[626,763,259,285]" degrees="149" direction="west" minutes="21.4" orientation="longitude" pageId="50" pageNumber="51" precision="92" value="-149.35667">149°21.4'W</geoCoordinate>
),
<date id="FFC610613A617B61FCCBFEE754F5FEE7" box="[786,927,259,285]" pageId="50" pageNumber="51" value="2002-08-07">
<collectingDate id="EF82E9893A617B61FCCBFEE754F5FEE7" box="[786,927,259,285]" pageId="50" pageNumber="51" value="2002-08-07">7 Aug 2002</collectingDate>
</date>
,
<collectingRegion id="49BCF8433A617B61FC74FEE754AEFEE7" box="[941,964,259,285]" country="United States of America" name="D" pageId="50" pageNumber="51">D</collectingRegion>
. and
<collectingRegion id="49BCF8433A617B61FBDCFEE7534BFEE7" box="[1029,1057,259,285]" country="United States of America" name="W" pageId="50" pageNumber="51">W</collectingRegion>
.
<collectingRegion id="49BCF8433A617B61FBF6FEE7532CFEE7" box="[1071,1094,259,285]" country="United States of America" name="N" pageId="50" pageNumber="51">N</collectingRegion>
.
<location id="8EA7607A3A617B61FB8DFEE753C8FEE7" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FB8DFEE753C8FEE7" box="[1108,1186,259,285]" country="United States of America" latitude="61.47" longLatPrecision="102" longitude="-149.35667" name="Mathis" pageId="50" pageNumber="51" stateProvince="D">Mathis</location>
(
<specimenCount id="9D7EFD283A617B61FB6EFEE753B1FEE7" box="[1207,1243,259,285]" count="1" pageId="50" pageNumber="51" type="male">1♂</specimenCount>
;
<collectionCode id="ED69AE643A617B61FB32FEE7522BFEE7" box="[1259,1345,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="50" pageNumber="51" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103CFC3A617B61FA81FEE75300FEBB" collectingDate="2011-08-10" collectionCode="USNM" country="United States of America" elevation="55" latitude="61.97167" location="Mathis" longLatPrecision="445" longitude="-150.08333" municipality="Sheep Creek" pageId="50" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="D" typeStatus="holotype">
<collectingMunicipality id="6BA3ACDB3A617B61FA81FEE757B0FEBB" pageId="50" pageNumber="51">Sheep Creek</collectingMunicipality>
(
<geoCoordinate id="EE4C50663A617B61FF33FEC25635FEBB" box="[234,351,294,321]" degrees="61" direction="north" minutes="58.3" orientation="latitude" pageId="50" pageNumber="51" precision="92" value="61.97167">61°58.3'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A617B61FEB5FEC256B5FEBB" box="[364,479,294,321]" degrees="150" direction="west" minutes="05" orientation="longitude" pageId="50" pageNumber="51" precision="925" value="-150.08333">150°05'W</geoCoordinate>
;
<quantity id="4C809B443A617B61FE32FEC25542FEBB" box="[491,552,294,321]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="50" pageNumber="51" unit="m" value="55.0">
<elevation id="0055D1923A617B61FE32FEC25542FEBB" box="[491,552,294,321]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="50" pageNumber="51" unit="m" value="55.0">55 m</elevation>
</quantity>
),
<date id="FFC610613A617B61FDE4FEC355BBFEBB" box="[573,721,295,321]" pageId="50" pageNumber="51" value="2011-08-10">
<collectingDate id="EF82E9893A617B61FDE4FEC355BBFEBB" box="[573,721,295,321]" pageId="50" pageNumber="51" value="2011-08-10">10 Aug 2011</collectingDate>
</date>
,
<collectingRegion id="49BCF8433A617B61FD04FEC3559EFEBB" box="[733,756,295,321]" country="United States of America" name="D" pageId="50" pageNumber="51">D</collectingRegion>
. and
<collectingRegion id="49BCF8433A617B61FCE9FEC35426FEBB" box="[816,844,295,321]" country="United States of America" name="W" pageId="50" pageNumber="51">W</collectingRegion>
.
<collectingRegion id="49BCF8433A617B61FC8EFEC35404FEBB" box="[855,878,295,321]" country="United States of America" name="N" pageId="50" pageNumber="51">N</collectingRegion>
.
<location id="8EA7607A3A617B61FCA3FEC354A2FEBB" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FCA3FEC354A2FEBB" box="[890,968,295,321]" country="United States of America" latitude="61.97167" longLatPrecision="445" longitude="-150.08333" municipality="Sheep Creek" name="Mathis" pageId="50" pageNumber="51" stateProvince="D">Mathis</location>
(
<specimenCount id="9D7EFD283A617B61FC05FEC35497FEBB" box="[988,1021,295,321]" count="1" pageId="50" pageNumber="51" type="female">1♀</specimenCount>
;
<collectionCode id="ED69AE643A617B61FBD5FEC35308FEBB" box="[1036,1122,295,321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="50" pageNumber="51" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B103CFC3A617B61FBACFEC354D7FE9F" collectingDate="2011-07-26" collectionCode="USNM" country="United States of America" elevation="50" latitude="61.768333" location="Mathis" longLatPrecision="102" longitude="-150.07" municipality="Willow Creek" pageId="50" pageNumber="51" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="D" typeStatus="holotype">
<collectingMunicipality id="6BA3ACDB3A617B61FBACFEC35279FEBB" box="[1141,1299,295,321]" pageId="50" pageNumber="51">Willow Creek</collectingMunicipality>
(
<geoCoordinate id="EE4C50663A617B61FAFAFEC352F2FEBB" box="[1315,1432,295,321]" degrees="61" direction="north" minutes="46.1" orientation="latitude" pageId="50" pageNumber="51" precision="92" value="61.768333">61°46.1'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A617B61FF4EFEAE5675FE9F" box="[151,287,330,357]" degrees="150" direction="west" minutes="04.2" orientation="longitude" pageId="50" pageNumber="51" precision="92" value="-150.07">150°04.2'W</geoCoordinate>
;
<quantity id="4C809B443A617B61FEF3FEAE560CFE9F" box="[298,358,330,357]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="m" value="50.0">
<elevation id="0055D1923A617B61FEF3FEAE560CFE9F" box="[298,358,330,357]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="m" value="50.0">50 m</elevation>
</quantity>
),
<date id="FFC610613A617B61FEA0FEAF5697FE9F" box="[377,509,331,357]" pageId="50" pageNumber="51" value="2011-07-26">
<collectingDate id="EF82E9893A617B61FEA0FEAF5697FE9F" box="[377,509,331,357]" pageId="50" pageNumber="51" value="2011-07-26">26 Jul 2011</collectingDate>
</date>
,
<collectingRegion id="49BCF8433A617B61FDD1FEAF5575FE9F" box="[520,543,331,357]" country="United States of America" name="D" pageId="50" pageNumber="51">D</collectingRegion>
. and
<collectingRegion id="49BCF8433A617B61FD81FEAF551EFE9F" box="[600,628,331,357]" country="United States of America" name="W" pageId="50" pageNumber="51">W</collectingRegion>
.
<collectingRegion id="49BCF8433A617B61FDA7FEAF55FFFE9F" box="[638,661,331,357]" country="United States of America" name="N" pageId="50" pageNumber="51">N</collectingRegion>
.
<location id="8EA7607A3A617B61FD46FEAF5587FE9F" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FD46FEAF5587FE9F" box="[671,749,331,357]" country="United States of America" latitude="61.768333" longLatPrecision="102" longitude="-150.07" municipality="Willow Creek" name="Mathis" pageId="50" pageNumber="51" stateProvince="D">Mathis</location>
(
<specimenCount id="9D7EFD283A617B61FD24FEAE5448FE9F" box="[765,802,330,357]" count="5" pageId="50" pageNumber="51" type="male">5♂</specimenCount>
,
<specimenCount id="9D7EFD283A617B61FCF7FEAF5439FE9F" box="[814,851,331,357]" count="4" pageId="50" pageNumber="51" type="female">4♀</specimenCount>
;
<collectionCode id="ED69AE643A617B61FC86FEAF54DFFE9F" box="[863,949,331,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="50" pageNumber="51" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103CFC3A617B61FC1EFEAF533EFE73" collectingDate="2012-08-02" collectionCode="USNM" country="United States of America" county="Nome" latitude="64.565" location="Snake River" longLatPrecision="100" longitude="-165.51" municipality="Census Area" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="D" typeStatus="holotype">
<emphasis id="B90CEAB33A617B61FC1EFEAF53DDFE9F" box="[967,1207,331,357]" italics="true" pageId="50" pageNumber="51">
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NW Nome
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(
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<caption id="DF0766293A617B61FF4EFB4F523EFB1D" ID-DOI="http://doi.org/10.5281/zenodo.13210841" ID-Zenodo-Dep="13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="50" pageNumber="51" startId="50.[151,264,1195,1219]" targetBox="[159,1427,425,1164]" targetPageId="50" targetType="figure">
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<emphasis id="B90CEAB33A617B61FF4EFB4F563FFB39" bold="true" box="[151,341,1195,1219]" pageId="50" pageNumber="51">FIGURES 97–99.</emphasis>
Male terminalia of
<taxonomicName id="4C784D223A617B61FDFAFB4F5589FB39" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[547,739,1195,1219]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="bolwigi" status="sp. nov.">
<emphasis id="B90CEAB33A617B61FDFAFB4F5589FB39" box="[547,739,1195,1219]" italics="true" pageId="50" pageNumber="51">Scatophila bolwigi</emphasis>
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<taxonomicNameLabel id="A23F57C83A617B61FD33FB4F5456FB39" box="[746,828,1195,1219]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A617B61FF1EFADD56FFFA45" blockId="50.[151,1437,1337,1976]" pageId="50" pageNumber="51">
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.—
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.
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Fairbanks
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:
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<location id="8EA7607A3A617B61FB79FADD5264FAA9" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FB79FADD5264FAA9" box="[1184,1294,1337,1363]" country="United States of America" county="Fairbanks North Star" latitude="64.83833" longLatPrecision="404" longitude="-147.71666" municipality="North Star" name="Fairbanks" pageId="50" pageNumber="51" stateProvince="Alaska">Fairbanks</location>
(
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,
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),
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,
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.
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(
<specimenCount id="9D7EFD283A617B61FD1DFAB9558DFA8D" box="[708,743,1373,1399]" count="2" pageId="50" pageNumber="51" type="female">2♀</specimenCount>
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of
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<emphasis id="B90CEAB33A617B61FC55FABA5351FA8D" box="[908,1083,1373,1400]" italics="true" pageId="50" pageNumber="51">S. variofacialis</emphasis>
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;
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).
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<collectingMunicipality id="6BA3ACDB3A617B61FB62FAB952FCFA8D" box="[1211,1430,1373,1399]" pageId="50" pageNumber="51">Matanuska-Susitna</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A617B61FF4EFA655678FA61" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FF4EFA655678FA61" box="[151,274,1409,1435]" country="United States of America" county="Aldrich" latitude="61.541668" longLatPrecision="102" longitude="-149.23" municipality="Matanuska-Susitna" name="Matanuska" pageId="50" pageNumber="51" stateProvince="J">Matanuska</location>
(
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,
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;
<collectingMethod id="52394EB63A617B61FDE1FA6655D9FA61" box="[568,691,1409,1436]" pageId="50" pageNumber="51">rotary trap</collectingMethod>
),
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<collectingDate id="EF82E9893A617B61FD1EFA6554D0FA61" box="[711,954,1409,1436]" pageId="50" pageNumber="51" value="1944-05-19" valueMax="1944-06-20" valueMin="1944-05-19">19 May–20 Jun 1944</collectingDate>
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,
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.
<collectingRegion id="49BCF8433A617B61FC39FA65549CFA61" box="[992,1014,1409,1435]" country="United States of America" name="C" pageId="50" pageNumber="51">C</collectingRegion>
.
<location id="8EA7607A3A617B61FBDBFA6553ECFA61" LSID="urn:lsid:plazi:treatment:03D187B73A637B61FF4EFC1254FEF842:8EA7607A3A617B61FBDBFA6553ECFA61" box="[1026,1158,1409,1435]" country="United States of America" name="Chamberlin" pageId="50" pageNumber="51" stateProvince="J">Chamberlin</location>
(
<specimenCount id="9D7EFD283A617B61FB41FA6553D7FA61" box="[1176,1213,1409,1435]" count="3" pageId="50" pageNumber="51" type="male">3♂</specimenCount>
,
<specimenCount id="9D7EFD283A617B61FB13FA655387FA61" box="[1226,1261,1409,1435]" count="4" pageId="50" pageNumber="51" type="female">4♀</specimenCount>
<typeStatus id="54C388033A617B61FB2CFA66520AFA66" box="[1269,1376,1410,1436]" pageId="50" pageNumber="51" type="paratype">paratypes</typeStatus>
of
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<emphasis id="B90CEAB33A617B61FA5EFA66564FFA45" italics="true" pageId="50" pageNumber="51">S. variofacialis</emphasis>
</taxonomicName>
;
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)
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.
</paragraph>
<paragraph id="8BC736A13A617B61FF1EFA2E56C7F9FD" blockId="50.[151,1437,1337,1976]" pageId="50" pageNumber="51">
<materialsCitation id="3B103CFC3A617B61FF1EFA2E56CDF9FD" country="United States of America" county="Valdez-Cordova" latitude="62.268333" location="Gulkana River" longLatPrecision="102" longitude="-145.385" municipality="Census Area" pageId="50" pageNumber="51" specimenCount="1" stateProvince="Alaska" typeStatus="holotype">
<typeStatus id="54C388033A617B61FF1EFA2E5795FA1E" box="[199,255,1482,1508]" pageId="50" pageNumber="51">Type</typeStatus>
Locality.—
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.
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.
<collectingCounty id="62A64E2D3A617B61FD59FA2D545CFA19" box="[640,822,1481,1507]" pageId="50" pageNumber="51">Valdez-Cordova</collectingCounty>
(
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):
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(
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N Glennallen
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;
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,
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)
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.
</paragraph>
<paragraph id="8BC736A13A617B61FF1EF9F55430F9D1" blockId="50.[151,1437,1337,1976]" box="[199,858,1553,1579]" pageId="50" pageNumber="51">
Distribution (
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).—
<taxonomicName id="4C784D223A617B61FE3CF9F55521F9D1" box="[485,587,1553,1579]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="97" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
:
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(
<collectingRegion id="49BCF8433A617B61FD23F9F55424F9D1" box="[762,846,1553,1579]" country="United States of America" name="Alaska" pageId="50" pageNumber="51">Alaska</collectingRegion>
).
</paragraph>
<paragraph id="8BC736A13A617B61FF1EF9D15211F96D" blockId="50.[151,1437,1337,1976]" pageId="50" pageNumber="51">
Etymology.—The species epithet,
<emphasis id="B90CEAB33A617B61FD96F9D155C0F9B5" box="[591,682,1589,1615]" italics="true" pageId="50" pageNumber="51">
<taxonomicName id="4C784D223A617B61FD96F9D155CCF9B5" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[591,678,1589,1615]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="bolwigi">bolwigi</taxonomicName>
,
</emphasis>
is genitive patronym to honor Dr. Niels Bolwig (
<date id="FFC610613A617B61FB3BF9D15218F9B5" box="[1250,1394,1589,1615]" pageId="50" pageNumber="51" value="1911-09-26">26 Sep 1911</date>
–
<date id="FFC610613A617B61FA5BF9D1566FF989" pageId="50" pageNumber="51" value="2004-09-10">10 Sep 2004</date>
), who was a Danish zoologist, behavioral biologist, entomologist, lecturer, and professor Dr. N. Bolwig published three papers on the biology and natural history of
<taxonomicName id="4C784D223A617B61FCE3F9995349F96D" box="[826,1059,1661,1687]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B90CEAB33A617B61FCE3F9995349F96D" box="[826,1059,1661,1687]" italics="true" pageId="50" pageNumber="51">Scatophila unicornis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE94B503A617B61FBE8F99953BEF96D" author="Bolwig, N." box="[1073,1236,1661,1687]" pageId="50" pageNumber="51" pagination="129 - 137" refId="ref69079" refString="Bolwig, N. (1940 a) The description of Scatophila unicornis Czerny, 1900 (Ephydridae, Diptera). The Proceedings of the Royal Entomological Society of London, Series B, 9, 129 - 137. https: // doi. org / 10.1111 / j. 1365 - 3113.1940. tb 00363. x" type="journal article" year="1940">Bolwig 1940a</bibRefCitation>
,
<bibRefCitation id="EFE94B503A617B61FB06F999524DF96D" author="Bolwig, N." box="[1247,1319,1661,1687]" pageId="50" pageNumber="51" pagination="97 - 102" refId="ref69139" refString="Bolwig, N. (1940 b) The reproductive organs of Scatophila unicornis Czerny (Diptera). The Proceedings of the Royal Entomological Society of London, Series A, 15, 97 - 102. https: // doi. org / 10.1111 / j. 1365 - 3032.1940. tb 00577. x" type="journal article" year="1940">1940b</bibRefCitation>
,
<bibRefCitation id="EFE94B503A617B61FAEAF9995205F96D" author="Bolwig, N." box="[1331,1391,1661,1687]" pageId="50" pageNumber="51" pagination="1 - 10" refId="ref69196" refString="Bolwig, N. (1941) The head of Scatophila unicornis Czerny (Diptera). The Proceedings of the Royal Entomological Society of London, Series A, 16 (1 - 3), 1 - 10. https: // doi. org / 10.1111 / j. 1365 - 3032.1941. tb 01213. x" type="journal article" year="1941">1941</bibRefCitation>
).
</paragraph>
<paragraph id="8BC736A13A617B61FF1EF94552FEF869" blockId="50.[151,1437,1337,1976]" pageId="50" pageNumber="51">
Remarks.—
<bibRefCitation id="EFE94B503A617B61FE94F945559EF941" author="Sturtevant, A. H. &amp; Wheeler, M. R." box="[333,756,1697,1724]" pageId="50" pageNumber="51" pagination="151 - 257" refId="ref70834" refString="Sturtevant, A. H. &amp; Wheeler, M. R. (1954) Synopses of Nearctic Ephydridae (Diptera). Transactions of the American Entomological Society, 79, 151 - 257." type="journal article" year="1954">Sturtevant and Wheeler (1954: 200)</bibRefCitation>
noted that “the mesonotal glossiness is scarcely evident” in specimens from
<collectingRegion id="49BCF8433A617B61FEACF92156A9F925" box="[373,451,1733,1759]" country="United States of America" name="Alaska" pageId="50" pageNumber="51">Alaska</collectingRegion>
that they had identified as
<emphasis id="B90CEAB33A617B61FCDDF92254DDF925" box="[772,951,1733,1760]" italics="true" pageId="50" pageNumber="51">
<taxonomicName id="4C784D223A617B61FCDDF92254D9F925" box="[772,947,1733,1760]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="variofacialis">S. variofacialis</taxonomicName>
,
</emphasis>
a new species they were then describing. We concur with their observation and have also discovered that structures of the male terminalia of the Alaskan populations are consistently quite different. These differences include much smaller ventrolateral projections of the epandrium (
<figureCitation id="13432A243A617B61FE95F8D556C8F8B1" box="[332,418,1841,1867]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="50" pageNumber="51">Fig. 97</figureCitation>
), the pentagonal shape of the aedeagus in ventral view (
<figureCitation id="13432A243A617B61FBF4F8D553E9F8B1" box="[1069,1155,1841,1867]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="50" pageNumber="51">Fig. 98</figureCitation>
), the smaller keel of the phallapodeme in lateral view (
<figureCitation id="13432A243A617B61FE30F8B15556F895" box="[489,572,1877,1903]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="50" pageNumber="51">Fig. 99</figureCitation>
), and the shape and orientation of the paired postgonites in ventral and lateral views (
<figureCitation id="13432A243A617B61FF31F89D5618F869" box="[232,370,1913,1939]" captionStart="FIGURES 97–99" captionStartId="50.[151,264,1195,1219]" captionTargetBox="[159,1427,425,1164]" captionTargetId="figure-153@50.[151,1435,417,1172]" captionTargetPageId="50" captionText="FIGURES 97–99. Male terminalia of Scatophila bolwigi sp. nov. 97. Epandrium and cerci, posterior view. 98. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 99. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210841" httpUri="https://zenodo.org/record/13210841/files/figure.png" pageId="50" pageNumber="51">Figs. 98–99</figureCitation>
). Thus, we are recognizing the populations from
<collectingRegion id="49BCF8433A617B61FC4CF89D5489F869" box="[917,995,1913,1939]" country="United States of America" name="Alaska" pageId="50" pageNumber="51">Alaska</collectingRegion>
as a separate and undescribed species.
</paragraph>
<paragraph id="8BC736A13A617B61FF1EF87954FEF842" blockId="50.[151,1437,1337,1976]" box="[199,916,1949,1976]" pageId="50" pageNumber="51">
Like
<emphasis id="B90CEAB33A617B61FED8F87A56D8F84D" box="[257,434,1949,1976]" italics="true" pageId="50" pageNumber="51">
<taxonomicName id="4C784D223A617B61FED8F87A56C7F84D" box="[257,429,1949,1976]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="variofacialis">S. variofacialis</taxonomicName>
,
</emphasis>
this species is placed in the
<emphasis id="B90CEAB33A617B61FD29F879542EF84D" box="[752,836,1949,1975]" italics="true" pageId="50" pageNumber="51">noctula</emphasis>
group.
</paragraph>
</subSubSection>
</treatment>
</document>