Add updates up until 2024-12-17 19:45:09

2024-12-17 19:50:13 +00:00 · 2024-12-17 19:50:13 +00:00 · 45d463932f
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 (from 
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 Gardner 
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 . 2008
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 ): Relatively small to moderate-sized skinks (adult SVL 
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 ); head and body squarish in cross-section; midbody scales in 34‒52 rows, the dorsals smooth (keeled in 
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 ); nasal scale without postnarial groove; subocular scale row incomplete; eyes relatively large, the eyelids usually with distinct cream margins, easily differentiated from adjacent scales. Terrestrial; mainly diurnal with some species crepuscular to nocturnal; viviparous. For content, see 
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 and 
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 .
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 provided no derivation of name, but it is presumably from the original Greek 
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 latinised to 
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 meaning smooth or flat and from the Greek 
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 meaning scale of a reptile.
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 <mods:role id="D702F89E1611541146AB92C3159726A0">
 <mods:roleTerm id="B25C0294F9B9D4E4431DDE6E18DB3349">Author</mods:roleTerm>
 </mods:role>
 <mods:namePart id="86E36138FA49D85FEA83B82999F821FD">Donnellan, Stephen C.</mods:namePart>
 <mods:affiliation id="05F58E364C28373F0EBCD7D9573C6AAE">South Australian Museum, North Terrace, Adelaide SA, 5000, AUSTRALIA. &amp; School of Biological Sciences, The University of Adelaide, Adelaide SA, 5005, AUSTRALIA. &amp; Australian Museum Research Institute, Australian Museum, Sydney NSW, 2010, AUSTRALIA.</mods:affiliation>
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 <mods:typeOfResource id="87B359475C41A89346246BAFD7B41A27">text</mods:typeOfResource>
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 <mods:title id="34D6FD243BC0CAA3BD71049D9C13ADAD">Zootaxa</mods:title>
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 <mods:part id="4A5AD58B430F8064C2540C5A63A36903">
 <mods:date id="AA82025543CBA0964871AD007AAD367E">2024</mods:date>
 <mods:detail id="A0472E0BA150851414622A3767CBBD8A" type="pubDate">
 <mods:number id="90288EAA78C0A920A0E99192DF26477C">2024-12-12</mods:number>
 </mods:detail>
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 <mods:number id="C7D51B72BBD764C92C0D4B13E51E28E7">5551</mods:number>
 </mods:detail>
 <mods:detail id="B78D83E1AC26E6CF1F781A8BC2D6AB60" type="issue">
 <mods:number id="307F3DEB383BF26E8750166B6D791925">2</mods:number>
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 <mods:start id="D78BAFEE3DBE1A941B3EDF5DB9FF04B4">299</mods:start>
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 <mods:url id="4D7CF43E0ADF690D994743499CA19037">https://doi.org/10.11646/zootaxa.5551.2.4</mods:url>
 </mods:location>
 <mods:classification id="0699453860B4C51DA433CF187AA18311">journal article</mods:classification>
 <mods:identifier id="E0D7849389A3A57D9EDF9ECBDA46ED00" type="DOI">10.11646/zootaxa.5551.2.4</mods:identifier>
 <mods:identifier id="F0DAD5D6D5266B81081602B726E2DB0E" type="ISSN">1175-5326</mods:identifier>
 <mods:identifier id="5D852A80546B5463F36662B37E4BE094" type="Zenodo-Dep">14421211</mods:identifier>
 <mods:identifier id="A7B67A656412075F5C63D642D0971DAE" type="ZooBank">43648902-1550-47C3-84F0-A7EA392CF27D</mods:identifier>
 </mods:mods>
 <treatment id="EC6387B48F134125FF0E9EED72AED51F" LSID="urn:lsid:plazi:treatment:EC6387B48F134125FF0E9EED72AED51F" httpUri="http://treatment.plazi.org/id/EC6387B48F134125FF0E9EED72AED51F" lastPageId="23" lastPageNumber="322" pageId="17" pageNumber="316">
 <subSubSection id="2CD065298F134123FF0E9EED70ABD0D5" box="[151,442,973,1000]" pageId="17" pageNumber="316" type="nomenclature">
 <paragraph id="647536A28F134123FF0E9EED70ABD0D5" blockId="17.[151,442,973,1072]" box="[151,442,973,1000]" pageId="17" pageNumber="316">
 <heading id="3F3D81CE8F134123FF0E9EED70ABD0D5" bold="true" box="[151,442,973,1000]" fontSize="11" level="1" pageId="17" pageNumber="316" reason="1">
 <emphasis id="56BEEAB08F134123FF0E9EED70ABD0D5" bold="true" box="[151,442,973,1000]" pageId="17" pageNumber="316">
 <taxonomicName id="A3CA4D218F134123FF0E9EED7044D0DA" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[151,341,973,1000]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F134123FF0E9EED7044D0DA" bold="true" box="[151,341,973,1000]" italics="true" pageId="17" pageNumber="316">Liopholis parva</emphasis>
 </taxonomicName>
 <taxonomicNameLabel id="4D8D57CB8F134123FEC59EED70ABD0D5" box="[348,442,973,999]" pageId="17" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 </heading>
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F134123FF0E9ED170B3D739" box="[151,418,1009,1035]" pageId="17" pageNumber="316" type="vernacular_names">
 <paragraph id="647536A28F134123FF0E9ED170B3D739" blockId="17.[151,442,973,1072]" box="[151,418,1009,1035]" pageId="17" pageNumber="316">North-West Cape Skink</paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F134123FF0E99357036D71D" box="[151,295,1045,1072]" pageId="17" pageNumber="316" type="description">
 <paragraph id="647536A28F134123FF0E99357036D71D" blockId="17.[151,442,973,1072]" box="[151,295,1045,1072]" pageId="17" pageNumber="316">
 (
 <figureCitation id="FCF12A278F134123FF06993571F9D71D" box="[159,232,1045,1071]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="17" pageNumber="316">Figs 6</figureCitation>
 , 
 <figureCitation id="FCF12A278F134123FF6D99367015D702" box="[244,260,1046,1072]" captionStart="FIGURE 7" captionStartId="19.[151,250,1462,1486]" captionTargetBox="[151,1436,181,1438]" captionTargetId="figure-17@19.[151,1436,181,1438]" captionTargetPageId="19" captionText="FIGURE 7. Liopholis parva sp. nov. A) Preserved holotype WAM R156452 in dorsal, ventral and lateral views, B) adult from the Learmonth Air Weapons Range, North-West Cape, Western Australia, and C) juvenile born in captivity, photographed in life (photos B. Maryan)." figureDoi="http://doi.org/10.5281/zenodo.14421236" httpUri="https://zenodo.org/record/14421236/files/figure.png" pageId="17" pageNumber="316">7</figureCitation>
 , 
 <figureCitation id="FCF12A278F134123FE899935700FD71D" box="[272,286,1045,1071]" captionStart="FIGURE 8" captionStartId="20.[151,250,1477,1501]" captionTargetBox="[165,1435,453,1439]" captionTargetId="figure-152@20.[151,1435,453,1454]" captionTargetPageId="20" captionText="FIGURE 8. Lateral and dorsal views of head scalation of A‒B) L. parva sp. nov. holotype WAM R156452, and C‒D) L. purnululu sp. nov. holotype WAM R103112." figureDoi="http://doi.org/10.5281/zenodo.14421238" httpUri="https://zenodo.org/record/14421238/files/figure.png" pageId="17" pageNumber="316">8</figureCitation>
 )
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F134123FF0E997C75EBD67C" pageId="17" pageNumber="316" type="materials_examined">
 <paragraph id="647536A28F134123FF0E997C7374D78C" blockId="17.[151,1437,1116,2042]" pageId="17" pageNumber="316">
 <materialsCitation id="D4A23CFF8F134123FF0E997C7374D78C" collectingDate="2005-03-17" collectionCode="WAM, SAMA" collectorName="M. Bamford" country="Australia" location="Western Australia" pageId="17" pageNumber="316" specimenCode="R156452, ABTC 101420" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
 <emphasis id="56BEEAB08F134123FF0E997C701AD744" bold="true" box="[151,267,1116,1142]" pageId="17" pageNumber="316">
 <typeStatus id="BB7188008F134123FF0E997C7017D744" box="[151,262,1116,1142]" pageId="17" pageNumber="316" type="holotype">Holotype</typeStatus>
 .
 </emphasis>
 <collectionCode id="02DBAE678F134123FE8F997C704BD744" box="[278,346,1116,1142]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="316" type="Museum">WAM</collectionCode>
 <specimenCode id="346C9ED98F134123FEFC997C70DFD745" box="[357,462,1116,1143]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="316">R156452</specimenCode>
 , male, Learmonth Air Weapons Range, North-West Cape (-22.45, 113.83), 
 <collectingRegion id="A60EF8408F134123FAD8997D7010D7A8" country="Australia" name="Western Australia" pageId="17" pageNumber="316">Western Australia</collectingRegion>
 , 
 <collectingCountry id="1CDD76328F134123FE9399A07065D7A8" box="[266,372,1152,1178]" name="Australia" pageId="17" pageNumber="316">Australia</collectingCountry>
 , collected by 
 <collectorName id="C93F53748F134123FD9499A17388D7A8" box="[525,665,1152,1179]" pageId="17" pageNumber="316">M. Bamford</collectorName>
 , 
 <date id="107410628F134123FD3D99A07242D7A9" box="[676,851,1152,1179]" pageId="17" pageNumber="316" value="2005-03-17">
 <collectingDate id="0030E98A8F134123FD3D99A07242D7A9" box="[676,851,1152,1179]" pageId="17" pageNumber="316" value="2005-03-17">17 March 2005</collectingDate>
 </date>
 . Fixed in 10% formalin, stored in 70% ethanol, liver tissue stored at 
 <collectionCode id="02DBAE678F134123FEDC99847085D78C" box="[325,404,1188,1214]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="17" pageNumber="316" type="Museum">SAMA</collectionCode>
 as 
 <specimenCode id="346C9ED98F134123FE2199847370D78C" box="[440,609,1188,1214]" collectionCode="ABTC" country="Australia" httpUri="http://grbio.org/cool/y82c-h1ev" name="Australian Biological Tissue Collection, South Australian Museum" pageId="17" pageNumber="316" type="Museum">ABTC 101420</specimenCode>
 .
 </materialsCitation>
 </paragraph>
 <paragraph id="647536A28F134123FF5E99E87554D634" blockId="17.[151,1437,1116,2042]" pageId="17" pageNumber="316">
 <materialsCitation id="D4A23CFF8F134123FF5E99E87550D634" collectionCode="WAM" location="Bullara Station" municipality="Two" pageId="17" pageNumber="316" specimenCode="R153831, R156453" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
 <emphasis id="56BEEAB08F134123FF5E99E87057D7D0" bold="true" box="[199,326,1224,1250]" pageId="17" pageNumber="316">
 <typeStatus id="BB7188008F134123FF5E99E87050D7D0" box="[199,321,1224,1250]" pageId="17" pageNumber="316" type="paratype">Paratypes</typeStatus>
 .
 </emphasis>
 <collectingMunicipality id="8411ACD88F134123FEC899E97093D7D1" box="[337,386,1225,1251]" pageId="17" pageNumber="316">Two</collectingMunicipality>
 from 
 <collectingRegion id="A60EF8408F134123FE5699E9738FD7D0" box="[463,670,1224,1251]" country="Australia" name="Western Australia" pageId="17" pageNumber="316">Western Australia</collectingRegion>
 . 
 <collectionCode id="02DBAE678F134123FD3599E873E1D7D0" box="[684,752,1224,1250]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="316" type="Museum">WAM</collectionCode>
 <specimenCode id="346C9ED98F134123FD6599E87277D7D1" box="[764,870,1224,1251]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="316">R153831</specimenCode>
 , female, 
 <location id="611560798F134123FC4D99E87591D7D0" LSID="urn:lsid:plazi:treatment:EC6387B48F134125FF0E9EED72AED51F:611560798F134123FC4D99E87591D7D0" box="[980,1152,1224,1250]" municipality="Two" name="Bullara Station" pageId="17" pageNumber="316" stateProvince="Western Australia">Bullara Station</location>
 (-22.89, 113.92); 
 <collectionCode id="02DBAE678F134123FAC199E8748DD7D0" box="[1368,1436,1224,1250]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="316" type="Museum">WAM</collectionCode>
 <specimenCode id="346C9ED98F134123FF0E99CC7010D635" box="[151,257,1260,1287]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="316">R156453</specimenCode>
 , male, Learmonth Air Weapons Range, North-West Cape (-22.45, 113.83)
 </materialsCitation>
 .
 </paragraph>
 <paragraph id="647536A28F134123FF5E983075EBD67C" blockId="17.[151,1437,1116,2042]" pageId="17" pageNumber="316">
 <emphasis id="56BEEAB08F134123FF5E983070A0D618" bold="true" box="[199,433,1296,1322]" pageId="17" pageNumber="316">Referred specimen.</emphasis>
 <collectionCode id="02DBAE678F134123FE20983070ECD618" box="[441,509,1296,1322]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="17" pageNumber="316" type="Museum">WAM</collectionCode>
 <collectionCode id="02DBAE678F134123FD9E9830730AD618" box="[519,539,1296,1322]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="316">R</collectionCode>
 8601, male, Merlinleigh Outstation, Kennedy Range (-24.32, 115.18), 
 <collectingRegion id="A60EF8408F134123FAD898317010D67C" country="Australia" name="Western Australia" pageId="17" pageNumber="316">Western Australia</collectingRegion>
 , 
 <collectingCountry id="1CDD76328F134123FE9398147065D67C" box="[266,372,1332,1358]" name="Australia" pageId="17" pageNumber="316">Australia</collectingCountry>
 . This specimen has no specific collection date but was accessioned during 1943.
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F134123FF5E98787589D474" pageId="17" pageNumber="316" type="diagnosis">
 <paragraph id="647536A28F134123FF5E987870C3D6ED" blockId="17.[151,1437,1116,2042]" pageId="17" pageNumber="316">
 <emphasis id="56BEEAB08F134123FF5E98787053D640" bold="true" box="[199,322,1368,1394]" pageId="17" pageNumber="316">Diagnosis.</emphasis>
 A moderately robust, small-sized member of the 
 <taxonomicName id="A3CA4D218F134123FC1198797518D640" baseAuthorityName="Rosen" baseAuthorityYear="1905" box="[904,1033,1369,1394]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F134123FC1198797518D640" box="[904,1033,1369,1394]" italics="true" pageId="17" pageNumber="316">L. inornata</emphasis>
 </taxonomicName>
 species group (maximum SVL 
 <quantity id="A3329B478F134123FA19987871D6D6A4" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="17" pageNumber="316" unit="mm" value="66.0">66 mm</quantity>
 ), with round pupil, smooth dorsal and caudal scales, a proportionally short neck and head, proportionally short forelimbs, a relatively short tail, up to 34 midbody scale rows, ≤ 24 fourth toe lamellae and a plain orange-brown colour with obscure pattern.
 </paragraph>
 <paragraph id="647536A28F134123FF5E98C87589D474" blockId="17.[151,1437,1116,2042]" pageId="17" pageNumber="316">
 <emphasis id="56BEEAB08F134123FF5E98C8735CD530" bold="true" box="[199,589,1512,1538]" pageId="17" pageNumber="316">Comparisons with other species.</emphasis>
 <taxonomicName id="A3CA4D218F134123FDCC98C87219D530" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[597,776,1512,1538]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F134123FDCC98C87219D530" box="[597,776,1512,1538]" italics="true" pageId="17" pageNumber="316">Liopholis parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F134123FC8898C8727DD530" bold="true" box="[785,876,1512,1538]" pageId="17" pageNumber="316">
 <taxonomicNameLabel id="4D8D57CB8F134123FC8898C8727DD530" box="[785,876,1512,1538]" pageId="17" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 differs from nominate 
 <taxonomicName id="A3CA4D218F134123FBEC98C975E5D530" baseAuthorityName="Rosen" baseAuthorityYear="1905" box="[1141,1268,1513,1538]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F134123FBEC98C975E5D530" box="[1141,1268,1513,1538]" italics="true" pageId="17" pageNumber="316">L. inornata</emphasis>
 </taxonomicName>
 from southern semiarid to arid 
 <collectingRegion id="A60EF8408F134123FEC99B2D730AD514" box="[336,539,1548,1575]" country="Australia" name="Western Australia" pageId="17" pageNumber="316">Western Australia</collectingRegion>
 , all known populations of 
 <emphasis id="56BEEAB08F134123FCD19B2D72D9D514" box="[840,968,1548,1574]" italics="true" pageId="17" pageNumber="316">
 L. s. 
 <taxonomicName id="A3CA4D218F134123FC199B2C72D9D514" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[896,968,1548,1574]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="slateri">slateri</taxonomicName>
 </emphasis>
 , and 
 <emphasis id="56BEEAB08F134123FB9E9B2D759ED514" box="[1031,1167,1549,1574]" italics="true" pageId="17" pageNumber="316">L. s. virgata</emphasis>
 in its smaller adult size (SVL to 
 <quantity id="A3329B478F134123FF639B10705BD579" box="[250,330,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="17" pageNumber="316" unit="mm" value="66.0">66 mm</quantity>
 <emphasis id="56BEEAB08F134123FECB9B11708BD578" box="[338,410,1585,1610]" italics="true" pageId="17" pageNumber="316">versus</emphasis>
 to 
 <quantity id="A3329B478F134123FE269B107300D579" box="[447,529,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="17" pageNumber="316" unit="mm" value="84.0">84 mm</quantity>
 , 
 <quantity id="A3329B478F134123FD849B10737CD579" box="[541,621,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="9.3" pageId="17" pageNumber="316" unit="mm" value="93.0">93 mm</quantity>
 and 
 <quantity id="A3329B478F134123FD3C9B1073E6D579" box="[677,759,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="17" pageNumber="316" unit="mm" value="94.0">94 mm</quantity>
 , respectively). 
 <taxonomicName id="A3CA4D218F134123FC3D9B107547D578" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[932,1110,1584,1610]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F134123FC3D9B107547D578" box="[932,1110,1584,1610]" italics="true" pageId="17" pageNumber="316">Liopholis parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F134123FBC69B1075A8D578" bold="true" box="[1119,1209,1584,1610]" pageId="17" pageNumber="316">
 <taxonomicNameLabel id="4D8D57CB8F134123FBC69B1075A8D578" box="[1119,1209,1584,1610]" pageId="17" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 further differs from 
 <taxonomicName id="A3CA4D218F134123FF0E9B757007D55C" baseAuthorityName="Rosen" baseAuthorityYear="1905" box="[151,278,1621,1646]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F134123FF0E9B757007D55C" box="[151,278,1621,1646]" italics="true" pageId="17" pageNumber="316">L. inornata</emphasis>
 </taxonomicName>
 and 
 <emphasis id="56BEEAB08F134123FED69B7570C1D55C" box="[335,464,1620,1646]" italics="true" pageId="17" pageNumber="316">
 L. s. 
 <taxonomicName id="A3CA4D218F134123FE119B7470C1D55C" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[392,464,1620,1646]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="slateri">slateri</taxonomicName>
 </emphasis>
 by having a shorter head, modally lower midbody and paravertebral scale counts and subdigital lamellae counts (
 <tableCitation id="294803198F134123FE549B587334D5A1" box="[461,549,1656,1683]" captionStart="TABLE 5" captionStartId="13.[151,239,1685,1709]" captionTargetPageId="13" captionText="TABLE 5. Summary statistics for measurements (mm) and meristic counts for various taxa and populations of the Liopholis inornata species group. N = sample size. NA = not available. Upper row is the range and the lower row is the mean ± SD. See Table 2 for abbreviations of measurements and counts. 1 AMS R.105730. 2 Sample size 4, except where indicated in parentheses." pageId="17" pageNumber="316">Table 5</tableCitation>
 ). 
 <taxonomicName id="A3CA4D218F134123FDA29B5873FFD5A0" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[571,750,1656,1682]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F134123FDA29B5873FFD5A0" box="[571,750,1656,1682]" italics="true" pageId="17" pageNumber="316">Liopholis parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F134123FD6F9B587240D5A0" bold="true" box="[758,849,1656,1682]" pageId="17" pageNumber="316">
 <taxonomicNameLabel id="4D8D57CB8F134123FD6F9B587240D5A0" box="[758,849,1656,1682]" pageId="17" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 further differs from 
 <emphasis id="56BEEAB08F134123FBA59B5975D4D5A0" box="[1084,1221,1657,1682]" italics="true" pageId="17" pageNumber="316">L. s. virgata</emphasis>
 by lacking a black vertebral stripe (
 <figureCitation id="FCF12A278F134123FECD9BBC708AD584" box="[340,411,1692,1718]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="17" pageNumber="316">Fig. 6</figureCitation>
 ). 
 <taxonomicName id="A3CA4D218F134123FE2A9BBC737ED584" box="[435,623,1692,1718]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="striata">
 <emphasis id="56BEEAB08F134123FE2A9BBC737ED584" box="[435,623,1692,1718]" italics="true" pageId="17" pageNumber="316">Liopholis striata</emphasis>
 </taxonomicName>
 attains a much larger size (SVL to 
 <quantity id="A3329B478F134123FB969BBC7560D584" box="[1039,1137,1692,1718]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="17" pageNumber="316" unit="mm" value="112.0">112 mm</quantity>
 ), has higher midbody and paravertebral scale counts, and has a vertically ovate rather than rounded pupil. 
 <taxonomicName id="A3CA4D218F134123FBAC9BE07422D5E8" baseAuthorityName="Mitchell &amp; Behrndt" baseAuthorityYear="1949" box="[1077,1331,1728,1754]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="multiscutata">
 <emphasis id="56BEEAB08F134123FBAC9BE07422D5E8" box="[1077,1331,1728,1754]" italics="true" pageId="17" pageNumber="316">Liopholis multiscutata</emphasis>
 </taxonomicName>
 attains a much larger size (SVL to 
 <quantity id="A3329B478F134123FE279BC47301D5CC" box="[446,528,1764,1790]" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="17" pageNumber="316" unit="mm" value="96.0">96 mm</quantity>
 ) and has a predominately greyish or brownish colour (often with dorsal stripes) and bicarinate subdigital lamellae. 
 <taxonomicName id="A3CA4D218F134123FDB99A2873C3D410" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[544,722,1800,1826]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F134123FDB99A2873C3D410" box="[544,722,1800,1826]" italics="true" pageId="17" pageNumber="316">Liopholis parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F134123FD439A287225D410" bold="true" box="[730,820,1800,1826]" pageId="17" pageNumber="316">
 <taxonomicNameLabel id="4D8D57CB8F134123FD439A287225D410" box="[730,820,1800,1826]" pageId="17" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 differs from 
 <taxonomicName id="A3CA4D218F134123FC509A297544D410" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[969,1109,1800,1826]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="17" pageNumber="316" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F134123FC509A297544D410" box="[969,1109,1800,1826]" italics="true" pageId="17" pageNumber="316">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F134123FBC49A2875A6D410" bold="true" box="[1117,1207,1800,1826]" pageId="17" pageNumber="316">
 <taxonomicNameLabel id="4D8D57CB8F134123FBC49A2875A6D410" box="[1117,1207,1800,1826]" pageId="17" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 by having a smaller size (SVL to 
 <quantity id="A3329B478F134123FEB29A0C706BD475" box="[299,378,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="17" pageNumber="316" unit="mm" value="66.0">66 mm</quantity>
 <emphasis id="56BEEAB08F134123FE199A0D70D9D474" box="[384,456,1837,1862]" italics="true" pageId="17" pageNumber="316">versus</emphasis>
 to 
 <quantity id="A3329B478F134123FE759A0D732DD474" box="[492,572,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="17" pageNumber="316" unit="mm" value="77.0">77 mm</quantity>
 ) and orange-brown (
 <emphasis id="56BEEAB08F134123FCBE9A0D727ED474" box="[807,879,1837,1862]" italics="true" pageId="17" pageNumber="316">versus</emphasis>
 brownish) colour (
 <figureCitation id="FCF12A278F134123FBDC9A0C759BD474" box="[1093,1162,1836,1862]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="17" pageNumber="316">Fig. 6</figureCitation>
 ).
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F134127FF5E9A7075CBD22F" lastPageId="21" lastPageNumber="320" pageId="17" pageNumber="316" type="description">
 <paragraph id="647536A28F134123FF5E9A7071CED4C8" blockId="17.[151,1437,1116,2042]" pageId="17" pageNumber="316">
 <emphasis id="56BEEAB08F134123FF5E9A7070A9D458" bold="true" box="[199,440,1872,1898]" pageId="17" pageNumber="316">
 Details of 
 <typeStatus id="BB7188008F134123FEDD9A7070A2D458" box="[324,435,1872,1898]" pageId="17" pageNumber="316" type="holotype">Holotype</typeStatus>
 .
 </emphasis>
 (
 <figureCitation id="FCF12A278F134123FE539A707336D458" box="[458,551,1872,1898]" captionStart="FIGURE 7" captionStartId="19.[151,250,1462,1486]" captionTargetBox="[151,1436,181,1438]" captionTargetId="figure-17@19.[151,1436,181,1438]" captionTargetPageId="19" captionText="FIGURE 7. Liopholis parva sp. nov. A) Preserved holotype WAM R156452 in dorsal, ventral and lateral views, B) adult from the Learmonth Air Weapons Range, North-West Cape, Western Australia, and C) juvenile born in captivity, photographed in life (photos B. Maryan)." figureDoi="http://doi.org/10.5281/zenodo.14421236" httpUri="https://zenodo.org/record/14421236/files/figure.png" pageId="17" pageNumber="316">Fig. 7A</figureCitation>
 ). SVL 57.9; TL 71; HD 7.9; HL 13.1; HW 10.5; HNL 19.5; FLL 16.6; HLL 23.8; AGL 28.7; MBS 34; PV 50; SDL 23; nasals widely separated; prefrontals in broad contact; supraciliaries 8/8; nuchals 1/1; upper palpebrals 10/10; primary temporal single; enlarged upper secondary temporals 1/1, each with one adjacent smaller temporal bordering the parietal; supralabials 7/7; ear lobules 3/4, uppermost lobule largest in series.
 </paragraph>
 <caption id="30B5662A8F104120FF0E9BC9738ED4EB" ID-DOI="http://doi.org/10.5281/zenodo.14421232" ID-Zenodo-Dep="14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="18" pageNumber="317" startId="18.[151,250,1769,1793]" targetBox="[151,1436,181,1745]" targetPageId="18" targetType="figure">
 <paragraph id="647536A28F104120FF0E9BC9738ED4EB" blockId="18.[151,1436,1769,2009]" pageId="18" pageNumber="317">
 <emphasis id="56BEEAB08F104120FF0E9BC97003D433" bold="true" box="[151,274,1769,1793]" pageId="18" pageNumber="317">FIGURE 6.</emphasis>
 Selection of 
 <taxonomicName id="A3CA4D218F104120FE0E9BC97345D433" baseAuthorityName="Rosen" baseAuthorityYear="1905" box="[407,596,1769,1793]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="18" pageNumber="317" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F104120FE0E9BC97345D433" box="[407,596,1769,1793]" italics="true" pageId="18" pageNumber="317">Liopholis inornata</emphasis>
 </taxonomicName>
 species group specimens illustrating variation in colour and pattern. 
 <emphasis id="56BEEAB08F104120FA939BC9740DD433" bold="true" box="[1290,1308,1769,1793]" pageId="18" pageNumber="317">A</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F104120FAB29BCA71FCD417" authority="WAM R" authorityName="WAM R" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="18" pageNumber="317" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F104120FAB29BCA748DD433" box="[1323,1436,1770,1793]" italics="true" pageId="18" pageNumber="317">L. inornata</emphasis>
 WAM R
 </taxonomicName>
 166716, Tropicana Mine, Western Australia, 
 <emphasis id="56BEEAB08F104120FD2E9A2D73D9D417" bold="true" box="[695,712,1805,1829]" pageId="18" pageNumber="317">B</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F104120FD419A2E72B9D417" authority="WAM R" authorityName="WAM R" box="[728,936,1805,1829]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="18" pageNumber="317" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F104120FD419A2E725BD417" box="[728,842,1806,1829]" italics="true" pageId="18" pageNumber="317">L. inornata</emphasis>
 WAM R
 </taxonomicName>
 163368, Neale Junction Nature Reserve, Western Australia, 
 <emphasis id="56BEEAB08F104120FE989A117002D47B" bold="true" box="[257,275,1841,1865]" pageId="18" pageNumber="317">C</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F104120FEBB9A1270E1D47B" authority="WAM R" authorityName="WAM R" box="[290,496,1841,1865]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="18" pageNumber="317" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F104120FEBB9A127085D47B" box="[290,404,1842,1865]" italics="true" pageId="18" pageNumber="317">L. inornata</emphasis>
 WAM R
 </taxonomicName>
 78798, 19 km ENE of Toomey Hills, Western Australia, 
 <emphasis id="56BEEAB08F104120FBB79A117551D47B" bold="true" box="[1070,1088,1841,1865]" pageId="18" pageNumber="317">D</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F104120FBD69A12740CD47B" authority="WAM R" authorityName="WAM R" box="[1103,1309,1841,1865]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="18" pageNumber="317" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F104120FBD69A1275D0D47B" box="[1103,1217,1842,1865]" italics="true" pageId="18" pageNumber="317">L. inornata</emphasis>
 WAM R
 </taxonomicName>
 65835, 3 km NE of Comet Vale, Western Australia, 
 <emphasis id="56BEEAB08F104120FD869A757321D45F" bold="true" box="[543,560,1877,1901]" pageId="18" pageNumber="317">E</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F104120FDA79A767385D45F" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[574,660,1878,1901]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="18" pageNumber="317" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F104120FDA79A767385D45F" box="[574,660,1878,1901]" italics="true" pageId="18" pageNumber="317">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F104120FD029A7573FAD45F" bold="true" box="[667,747,1877,1901]" pageId="18" pageNumber="317">
 <taxonomicNameLabel id="4D8D57CB8F104120FD029A7573FAD45F" box="[667,747,1877,1901]" pageId="18" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, 
 <emphasis id="56BEEAB08F104120FE9D9A597002D4A3" bold="true" box="[260,275,1913,1937]" pageId="18" pageNumber="317">F</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F104120FEBF9A5A70B8D4A3" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[294,425,1913,1937]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="18" pageNumber="317" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F104120FEBF9A5A70B8D4A3" box="[294,425,1913,1937]" italics="true" pageId="18" pageNumber="317">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F104120FE2A9A597316D4A3" bold="true" box="[435,519,1913,1937]" pageId="18" pageNumber="317">
 <taxonomicNameLabel id="4D8D57CB8F104120FE2A9A597316D4A3" box="[435,519,1913,1937]" pageId="18" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 paratype WAM R156716, Purnululu National Park, Western Australia, 
 <emphasis id="56BEEAB08F104120FB649A597400D4A3" bold="true" box="[1277,1297,1913,1937]" pageId="18" pageNumber="317">G</emphasis>
 ) 
 <emphasis id="56BEEAB08F104120FABA9A5A748DD4A3" box="[1315,1436,1913,1937]" italics="true" pageId="18" pageNumber="317">
 L. s. 
 <taxonomicName id="A3CA4D218F104120FAC29A59748DD4A3" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[1371,1436,1913,1937]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="18" pageNumber="317" phylum="Chordata" rank="species" species="slateri">slateri</taxonomicName>
 </emphasis>
 holotype WAM R29130, Alice Springs, Northern Territory, and 
 <emphasis id="56BEEAB08F104120FCAC9ABD7258D487" bold="true" box="[821,841,1949,1973]" pageId="18" pageNumber="317">H</emphasis>
 ) 
 <emphasis id="56BEEAB08F104120FCC39ABE72C8D487" box="[858,985,1950,1973]" italics="true" pageId="18" pageNumber="317">L. s. virgata</emphasis>
 SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach).
 </paragraph>
 </caption>
 <caption id="30B5662A8F114121FF0E98967077D524" ID-DOI="http://doi.org/10.5281/zenodo.14421236" ID-Zenodo-Dep="14421236" httpUri="https://zenodo.org/record/14421236/files/figure.png" pageId="19" pageNumber="318" startId="19.[151,250,1462,1486]" targetBox="[151,1436,181,1438]" targetPageId="19" targetType="figure">
 <paragraph id="647536A28F114121FF0E98967077D524" blockId="19.[151,1436,1462,1558]" pageId="19" pageNumber="318">
 <emphasis id="56BEEAB08F114121FF0E98967002D6FC" bold="true" box="[151,275,1462,1486]" pageId="19" pageNumber="318">FIGURE 7.</emphasis>
 <taxonomicName id="A3CA4D218F114121FE80989670ABD6FC" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[281,442,1462,1486]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="19" pageNumber="318" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F114121FE80989670ABD6FC" box="[281,442,1462,1486]" italics="true" pageId="19" pageNumber="318">Liopholis parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F114121FE589896733BD6FC" bold="true" box="[449,554,1462,1486]" pageId="19" pageNumber="318">
 <taxonomicNameLabel id="4D8D57CB8F114121FE5898967300D6FC" box="[449,529,1462,1486]" pageId="19" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
 A
 </emphasis>
 ) Preserved holotype WAM R156452 in dorsal, ventral and lateral views, 
 <emphasis id="56BEEAB08F114121FA8B98967432D6FC" bold="true" box="[1298,1315,1462,1486]" pageId="19" pageNumber="318">B</emphasis>
 ) adult from the Learmonth Air Weapons Range, North-West Cape, Western Australia, and 
 <emphasis id="56BEEAB08F114121FC3098FA72AAD6C0" bold="true" box="[937,955,1498,1522]" pageId="19" pageNumber="318">C</emphasis>
 ) juvenile born in captivity, photographed in life (photos B. Maryan).
 </paragraph>
 </caption>
 <paragraph id="647536A28F114121FF5E9B6473F6D5F8" blockId="19.[151,1437,1604,2027]" pageId="19" pageNumber="318">
 <emphasis id="56BEEAB08F114121FF5E9B647040D56C" bold="true" box="[199,337,1604,1630]" pageId="19" pageNumber="318">Description</emphasis>
 . See 
 <tableCitation id="294803198F114121FE179B6470F1D56D" box="[398,480,1604,1631]" captionStart="TABLE 5" captionStartId="13.[151,239,1685,1709]" captionTargetPageId="13" captionText="TABLE 5. Summary statistics for measurements (mm) and meristic counts for various taxa and populations of the Liopholis inornata species group. N = sample size. NA = not available. Upper row is the range and the lower row is the mean ± SD. See Table 2 for abbreviations of measurements and counts. 1 AMS R.105730. 2 Sample size 4, except where indicated in parentheses." pageId="19" pageNumber="318">Table 5</tableCitation>
 for summary statistics of mensural and meristic data and Supplementary 
 <tableCitation id="294803198F114121FA8A9B647465D56C" box="[1299,1396,1604,1630]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionTargetPageId="2" captionText="TABLE 1. Locality and sample information for all individuals of Liopholis genetically and morphologically examined (prefixes omitted). Under the registration# column: H indicates holotype. Abbreviations are: ABTC Australian Biological Tissue Collection,Adelaide;AMSAustralian Museum, Sydney;ANWCAustralian National Wildlife Collection, Canberra; MAGNT Museum and Art Gallery of the Northern Territory, Darwin; NMV National Museum of Victoria, Melbourne; SAMA South Australian Museum, Adelaide; WAM Western Australian Museum, Perth; NSW New South Wales; NT Northern Territory; SA South Australia; VIC Victoria; WA Western Australia; CP Conservation Park; HS Homestead; NP National Park; NR Nature Reserve. Under the analysis column specimens examined as follows: M morphology, S SNPs." pageId="19" pageNumber="318">Table S1</tableCitation>
 for measurements and counts of individual specimens. 
 <figureCitation id="FCF12A278F114121FD539B487218D5B0" box="[714,777,1640,1666]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="19" pageNumber="318">Fig. 6</figureCitation>
 shows the 
 <typeStatus id="BB7188008F114121FCE69B4872F0D5B0" box="[895,993,1640,1666]" pageId="19" pageNumber="318" type="holotype">holotype</typeStatus>
 WAM R156452 compared to a selection of other 
 <taxonomicName id="A3CA4D218F114121FF6D9BAC70ADD594" box="[244,444,1676,1702]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="19" pageNumber="318" phylum="Chordata" rank="species" species="undetermined">
 <emphasis id="56BEEAB08F114121FF6D9BAC704FD594" box="[244,350,1676,1702]" italics="true" pageId="19" pageNumber="318">Liopholis</emphasis>
 species.
 </taxonomicName>
 <figureCitation id="FCF12A278F114121FE589BAC7305D595" box="[449,532,1676,1703]" captionStart="FIGURE 7" captionStartId="19.[151,250,1462,1486]" captionTargetBox="[151,1436,181,1438]" captionTargetId="figure-17@19.[151,1436,181,1438]" captionTargetPageId="19" captionText="FIGURE 7. Liopholis parva sp. nov. A) Preserved holotype WAM R156452 in dorsal, ventral and lateral views, B) adult from the Learmonth Air Weapons Range, North-West Cape, Western Australia, and C) juvenile born in captivity, photographed in life (photos B. Maryan)." figureDoi="http://doi.org/10.5281/zenodo.14421236" httpUri="https://zenodo.org/record/14421236/files/figure.png" pageId="19" pageNumber="318">Fig. 7B</figureCitation>
 shows an adult and 
 <figureCitation id="FCF12A278F114121FD689BAC7247D595" box="[753,854,1676,1703]" captionStart="FIGURE 7" captionStartId="19.[151,250,1462,1486]" captionTargetBox="[151,1436,181,1438]" captionTargetId="figure-17@19.[151,1436,181,1438]" captionTargetPageId="19" captionText="FIGURE 7. Liopholis parva sp. nov. A) Preserved holotype WAM R156452 in dorsal, ventral and lateral views, B) adult from the Learmonth Air Weapons Range, North-West Cape, Western Australia, and C) juvenile born in captivity, photographed in life (photos B. Maryan)." figureDoi="http://doi.org/10.5281/zenodo.14421236" httpUri="https://zenodo.org/record/14421236/files/figure.png" pageId="19" pageNumber="318">Fig. 7C a</figureCitation>
 juvenile born in captivity, both photographed in life. Head and details of scalation are illustrated in 
 <figureCitation id="FCF12A278F114121FD069B9073F3D5F8" box="[671,738,1712,1738]" captionStart="FIGURE 8" captionStartId="20.[151,250,1477,1501]" captionTargetBox="[165,1435,453,1439]" captionTargetId="figure-152@20.[151,1435,453,1454]" captionTargetPageId="20" captionText="FIGURE 8. Lateral and dorsal views of head scalation of A‒B) L. parva sp. nov. holotype WAM R156452, and C‒D) L. purnululu sp. nov. holotype WAM R103112." figureDoi="http://doi.org/10.5281/zenodo.14421238" httpUri="https://zenodo.org/record/14421238/files/figure.png" pageId="19" pageNumber="318">Fig. 8</figureCitation>
 .
 </paragraph>
 <paragraph id="647536A28F114121FF5E9BF475B7D468" blockId="19.[151,1437,1604,2027]" pageId="19" pageNumber="318">Head moderately deep, barely distinct from neck; external ear opening prominent, narrow and much higher than wide; snout very short, rounded in profile and slightly angular in lateral view; body moderately robust, round in cross-section with flat venter and well-developed, thickset pentadactyl limbs; digits moderately long and slender; claws strong with long sharp tip; tail round in cross-section gradually tapering to pointed tip.</paragraph>
 <paragraph id="647536A28F114126FF5E9A44730CD29F" blockId="19.[151,1437,1604,2027]" lastBlockId="20.[151,1437,151,429]" lastPageId="20" lastPageNumber="319" pageId="19" pageNumber="318">Nasals widely separated; groove representing nasal-postnasal suture absent; two loreals, first slightly higher than wide and second mostly wider than high; two presuboculars, first much higher than second; prefrontals in broad or point contact (e.g. WAM R153831); frontal much wider anteriorly, tapering gradually to a rounded point, equal or only slightly wider than interparietal; frontoparietals paired; parietals bordered posteriorly by an enlarged nuchal scale and laterally by two temporal scales; 3‒4 ear lobules, small and obtusely rounded, uppermost lobule largest and with slight decrease in size downwards; 7‒8 supraciliaries, first and third largest; five supraoculars, second much larger and fifth smallest with anterior two in contact with frontal; 9‒10 upper palpebrals; one nuchal on each side; 6‒7 supralabials, the fifth and sixth or fourth and fifth largest and subocular, subocular supralabials separated from granular eyelid scales by a row of small scales; first pair of chin shields in broad contact, second pair separated medially by one scale; third pair widely separated medially by three scales; 34 midbody scale rows; 49‒50 paravertebral scales; 23‒24 lamellae under fourth toe, with moderately strong calli, outer edges denticulate, palmar and plantar scales small and ovate.</paragraph>
 <caption id="30B5662A8F164126FF0E98E5734ED533" ID-DOI="http://doi.org/10.5281/zenodo.14421238" ID-Zenodo-Dep="14421238" httpUri="https://zenodo.org/record/14421238/files/figure.png" pageId="20" pageNumber="319" startId="20.[151,250,1477,1501]" targetBox="[165,1435,453,1439]" targetPageId="20" targetType="figure">
 <paragraph id="647536A28F164126FF0E98E5734ED533" blockId="20.[151,1437,1477,1537]" pageId="20" pageNumber="319">
 <emphasis id="56BEEAB08F164126FF0E98E57007D6EF" bold="true" box="[151,278,1477,1501]" pageId="20" pageNumber="319">FIGURE 8.</emphasis>
 Lateral and dorsal views of head scalation of 
 <emphasis id="56BEEAB08F164126FC9B98E57223D6EF" bold="true" box="[770,818,1477,1501]" pageId="20" pageNumber="319">A‒B</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F164126FCDA98E6728DD6EF" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[835,924,1478,1501]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="20" pageNumber="319" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F164126FCDA98E6728DD6EF" box="[835,924,1478,1501]" italics="true" pageId="20" pageNumber="319">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F164126FC3C98E572E8D6EF" bold="true" box="[933,1017,1477,1501]" pageId="20" pageNumber="319">
 <taxonomicNameLabel id="4D8D57CB8F164126FC3C98E572E8D6EF" box="[933,1017,1477,1501]" pageId="20" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 holotype WAM R156452, and 
 <emphasis id="56BEEAB08F164126FADC98E57467D6EF" bold="true" box="[1349,1398,1477,1501]" pageId="20" pageNumber="319">C‒D</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F164126FA1198E671EAD533" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="20" pageNumber="319" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F164126FA1198E671EAD533" italics="true" pageId="20" pageNumber="319">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F164126FE9898C97043D533" bold="true" box="[257,338,1513,1537]" pageId="20" pageNumber="319">
 <taxonomicNameLabel id="4D8D57CB8F164126FE9898C97043D533" box="[257,338,1513,1537]" pageId="20" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 holotype WAM R103112.
 </paragraph>
 </caption>
 <paragraph id="647536A28F164126FF5E9B0F7027D483" blockId="20.[151,1437,1583,1970]" pageId="20" pageNumber="319">
 <emphasis id="56BEEAB08F164126FF5E9B0F7079D57B" bold="true" box="[199,360,1583,1609]" pageId="20" pageNumber="319">Colour in life</emphasis>
 . Colour in life is based on illustrated figures and field observations of live individuals. Dorsal surface pale orange-brown, grading to greyish on head above eyes. Dorsal scales immaculate (e.g. WAM R153831) or spotted on posterior edge with cream (e.g. WAM R156452‒3), the spots tending to be transversely elongate and forming very indistinct longitudinal lines that continue to dorsolateral surface of tail. Lateral surface with creamy white base colour. Some upper lateral scales bearing pale brown edges, forming an irregular reticulum that commences behind the ear aperture and terminates on proximal part of tail. Tail concolorous with dorsal surface, except for presence of brownish smudges, most prominent dorsolaterally and aggregated on the dorsal surface to form narrow transverse bands. Scales in interspaces between bands on tail with cream posterior edges, forming a central series of narrow pale bars. Upper surface of limbs pale brown, with no discernible markings. Posterior supralabials and temporals with diffuse brown smudges. Ventral surface creamy white. Claws with dark culmen on upper surface.
 </paragraph>
 <paragraph id="647536A28F174127FF5E9DB771D9D3E7" blockId="21.[151,1437,151,1905]" pageId="21" pageNumber="320">
 Juveniles are a rich bright orange with yellowish faces, dark crowns and prominent bands on the tail (
 <figureCitation id="FCF12A278F174127FAF69DB771AAD3E7" captionStart="FIGURE 7" captionStartId="19.[151,250,1462,1486]" captionTargetBox="[151,1436,181,1438]" captionTargetId="figure-17@19.[151,1436,181,1438]" captionTargetPageId="19" captionText="FIGURE 7. Liopholis parva sp. nov. A) Preserved holotype WAM R156452 in dorsal, ventral and lateral views, B) adult from the Learmonth Air Weapons Range, North-West Cape, Western Australia, and C) juvenile born in captivity, photographed in life (photos B. Maryan)." figureDoi="http://doi.org/10.5281/zenodo.14421236" httpUri="https://zenodo.org/record/14421236/files/figure.png" pageId="21" pageNumber="320">Fig. 7C</figureCitation>
 ).
 </paragraph>
 <paragraph id="647536A28F174127FF5E9DFF75CBD22F" blockId="21.[151,1437,151,1905]" pageId="21" pageNumber="320">The referred specimen WAM R8601 has been in preservative for about 80 years and the ground colour has faded to a creamy white, with no discernible spots on the dorsum and only faint bands on the tail.</paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F174127FF5E9C077350D257" pageId="21" pageNumber="320" type="etymology">
 <paragraph id="647536A28F174127FF5E9C077350D257" blockId="21.[151,1437,151,1905]" pageId="21" pageNumber="320">
 <emphasis id="56BEEAB08F174127FF5E9C077059D273" bold="true" box="[199,328,295,321]" pageId="21" pageNumber="320">Etymology</emphasis>
 . From the Latin 
 <emphasis id="56BEEAB08F174127FD989C07735CD272" box="[513,589,295,320]" italics="true" pageId="21" pageNumber="320">parvus</emphasis>
 , small, little, minute, in specific reference to the distinctive small size of this species. Used as a noun in apposition.
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F174127FF5E9C4E70F5D06F" pageId="21" pageNumber="320" type="distribution">
 <paragraph id="647536A28F174127FF5E9C4E75AFD19B" blockId="21.[151,1437,151,1905]" pageId="21" pageNumber="320">
 <emphasis id="56BEEAB08F174127FF5E9C4E7048D2BA" bold="true" box="[199,345,366,392]" pageId="21" pageNumber="320">Distribution</emphasis>
 . Known from two disjunct populations in 
 <collectingRegion id="A60EF8408F174127FCC99C4F750ED2BB" box="[848,1055,367,393]" country="Australia" name="Western Australia" pageId="21" pageNumber="320">Western Australia</collectingRegion>
 , one in the dune systems on the North-West Cape, the other in similar habitat on the Kennedy Range (
 <figureCitation id="FCF12A278F174127FC009CB372E1D29F" box="[921,1008,403,429]" captionStart="FIGURE 1" captionStartId="7.[151,250,1832,1856]" captionTargetBox="[151,1436,150,1825]" captionTargetId="figure-17@7.[151,1436,150,1825]" captionTargetPageId="7" captionText="FIGURE 1. A) Map of Australia showing distribution and sample locations of the Liopholis inornata species group used in the molecular genetic analysis. Purple star L. parva sp. nov., pink triangle L. purnululu sp. nov., green circles L. inornata, blue crosses L. s. slateri. B) showing sample locations in central Australia. C) locations for L. parva sp. nov. a—Merlinleigh Outstation, Kennedy Range, b—Bullara Station, c—Learmonth Air Weapons Range, d—20 km W of Learmonth, e—8 km N of Exmouth. The location numbering scheme is used to identify individuals in plots and terminals on the trees in Figs 2–3. Locality details are provided in Table 1. Voucher collection records are based on the Atlas of Living Australia (2024)." figureDoi="http://doi.org/10.5281/zenodo.14421213" httpUri="https://zenodo.org/record/14421213/files/figure.png" pageId="21" pageNumber="320">Fig. 1C</figureCitation>
 ). On the North-West Cape, it is known from four separate locations associated with two main dune systems: Cape Range National Park, Learmonth Air Weapons Range, Bullara Station and immediately north of Exmouth. Merlinleigh Outstation on the Kennedy Range has not been intensively surveyed and the status of 
 <taxonomicName id="A3CA4D218F174127FD799CDF7251D12A" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[736,832,511,536]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F174127FD799CDF7251D12A" box="[736,832,511,536]" italics="true" pageId="21" pageNumber="320">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F174127FCD19CDF72B2D12B" bold="true" box="[840,931,511,537]" pageId="21" pageNumber="320">
 <taxonomicNameLabel id="4D8D57CB8F174127FCD19CDF72B2D12B" box="[840,931,511,537]" pageId="21" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 at this location is not known. Herpetofaunal assessments on the North-West Cape, the Kennedy Range National Park and adjacent regions have not detected any populations of 
 <taxonomicName id="A3CA4D218F174127FEEF9F6770C9D152" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[374,472,583,608]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F174127FEEF9F6770C9D152" box="[374,472,583,608]" italics="true" pageId="21" pageNumber="320">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F174127FE7B9F67732FD153" bold="true" box="[482,574,583,609]" pageId="21" pageNumber="320">
 <taxonomicNameLabel id="4D8D57CB8F174127FE7B9F67732FD153" box="[482,574,583,609]" pageId="21" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 or any other 
 <taxonomicName id="A3CA4D218F174127FD479F6772B3D153" box="[734,930,583,609]" pageId="21" pageNumber="320">
 <emphasis id="56BEEAB08F174127FD479F677259D153" box="[734,840,583,609]" italics="true" pageId="21" pageNumber="320">Liopholis</emphasis>
 species
 </taxonomicName>
 (e.g. 
 <bibRefCitation id="005B4B538F174127FC7E9F6775CED153" author="Storr, G. M. &amp; Hanlon, T. M. S." box="[999,1247,583,609]" pageId="21" pageNumber="320" pagination="423 - 439" refId="ref22521" refString="Storr, G. M. &amp; Hanlon, T. M. S. (1980) Herpetofauna of the Exmouth Region, Western Australia. Records of the Western Australian Museum, 8, 423 - 439." type="journal article" year="1980">Storr &amp; Hanlon 1980</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F174127FB749F6771C3D1B7" author="Storr, G. M. &amp; Harold, G." pageId="21" pageNumber="320" pagination="173 - 189" refId="ref22561" refString="Storr, G. M. &amp; Harold, G. (1984) Herpetofauna of the Lake MacLeod Region, Western Australia. Records of the Western Australian Museum, 11, 173 - 189." type="journal article" year="1984">Storr &amp; Harold 1984</bibRefCitation>
 , 
 <bibRefCitation id="005B4B538F174127FF479F4A7008D1B7" author="Storr, G. M. &amp; Harold, G." box="[222,281,618,645]" pageId="21" pageNumber="320" pagination="277 - 291" refId="ref22598" refString="Storr, G. M. &amp; Harold, G. (1985) Herpetofauna of the Onslow Region, Western Australia. Records of the Western Australian Museum, 12, 277 - 291." type="journal article" year="1985">1985</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F174127FEBC9F4B70DED1B7" author="Kendrick, P. G." box="[293,463,619,645]" pageId="21" pageNumber="320" pagination="193 - 206" refId="ref20534" refString="Kendrick, P. G. (1993) Biogeography of the vertebrates of the Cape Range peninsula, Western Australia. In: W. F. Humphreys (Ed.) The Biogeography of Cape Range, Western Australia. Records of the Western Australian Museum, 45 (Supplement), 193 - 206." type="journal article" year="1993">Kendrick 1993</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F174127FE459F4B73C5D1B7" author="McKenzie, N. L. &amp; Rolfe, J. K. &amp; Aplin, K. P. &amp; Cowan, M. A. &amp; Smith, L. A." box="[476,724,619,645]" pageId="21" pageNumber="320" pagination="335 - 360" refId="ref21182" refString="McKenzie, N. L., Rolfe, J. K., Aplin, K. P., Cowan, M. A. &amp; Smith, L. A. (2000) Herpetofauna of the southern Carnarvon Basin, Western Australia. In: A. H. Burbidge, M. S. Harvey and N. L. McKenzie (Eds.) Biodiversity of the southern Carnarvon Basin. Records of the Western Australian Museum, 61 (Supplement), 335 - 360." type="journal article" year="2000">
 McKenzie 
 <emphasis id="56BEEAB08F174127FDC19F4B7398D1B7" box="[600,649,619,645]" italics="true" pageId="21" pageNumber="320">et al</emphasis>
 . 2000
 </bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F174127FD469F4B72AED1B7" author="Beranek, C. &amp; Mahony, S. &amp; Scott, S." box="[735,959,619,645]" pageId="21" pageNumber="320" pagination="146 - 155" refId="ref18667" refString="Beranek, C., Mahony, S. &amp; Scott, S. (2021) A significant range extension for the Western Soil-Crevice Skink Proablepharus reginae (Glauert 1960) and an updated reptile species list of Cape Range, Western Australia. Australian Zoologist, 42, 146 - 155. https: // doi. org / 10.7882 / AZ. 2021.029" type="journal article" year="2021">
 Beranek 
 <emphasis id="56BEEAB08F174127FCDD9F4B7264D1B7" box="[836,885,619,645]" italics="true" pageId="21" pageNumber="320">et al</emphasis>
 . 2021
 </bibRefCitation>
 ). However, recent biological surveys have recorded this species for the first time in Cape Range National Park (L. Clausen, pers. comm.).
 </paragraph>
 <paragraph id="647536A28F174127FF5E9F9370F5D06F" blockId="21.[151,1437,151,1905]" pageId="21" pageNumber="320">
 Based on collection records (
 <bibRefCitation id="005B4B538F174127FD889F937263D1FF" author="Atlas of Living Australia" box="[529,882,691,717]" pageId="21" pageNumber="320" refId="ref18479" refString="Atlas of Living Australia (2024) Available from: Atlas of Living Australia - Open access to Australia's biodiversity data (ala. org. au) (accessed 12 July 2024)" type="book" year="2024">
 Atlas of Living 
 <collectingCountry id="1CDD76328F174127FD5E9F93723FD1FF" box="[711,814,691,717]" name="Australia" pageId="21" pageNumber="320">Australia</collectingCountry>
 2024
 </bibRefCitation>
 ), 
 <taxonomicName id="A3CA4D218F174127FC1F9F93752BD1FE" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[902,1082,691,717]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F174127FC1F9F93752BD1FE" box="[902,1082,691,717]" italics="true" pageId="21" pageNumber="320">Liopholis parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F174127FBDB9F93758CD1FF" bold="true" box="[1090,1181,691,717]" pageId="21" pageNumber="320">
 <taxonomicNameLabel id="4D8D57CB8F174127FBDB9F93758CD1FF" box="[1090,1181,691,717]" pageId="21" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 is widely allopatric to any other 
 <taxonomicName id="A3CA4D218F174127FE9E9FF770C1D1C3" box="[263,464,727,753]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="undetermined">
 <emphasis id="56BEEAB08F174127FE9E9FF77060D1C3" box="[263,369,727,753]" italics="true" pageId="21" pageNumber="320">Liopholis</emphasis>
 species.
 </taxonomicName>
 The geographically nearest congeners consist of single records of 
 <taxonomicName id="A3CA4D218F174127FB2F9FF77422D1C2" box="[1206,1331,727,752]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F174127FB2F9FF77422D1C2" box="[1206,1331,727,752]" italics="true" pageId="21" pageNumber="320">L. inornata</emphasis>
 </taxonomicName>
 recorded approximately 
 <quantity id="A3329B478F174127FED89FDB7086D027" box="[321,407,763,789]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="320" unit="km" value="200.0">200 km</quantity>
 to the southwest on the Peron Peninsula and Hamelin Station Reserve at Shark Bay (
 <bibRefCitation id="005B4B538F174127FAFD9FDB7029D00B" author="Shea, G. M. &amp; Wells, R. W." pageId="21" pageNumber="320" refId="ref22133" refString="Shea, G. M. &amp; Wells, R. W. (1985) Egernia inornata, an additional reptile record for the Shark Bay region. Western Australian Naturalist, 16, 91. https: // doi. org / 10.3853 / j. 0067 - 1975.37.1985.337" type="journal volume" year="1985">Shea &amp; Wells 1985</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F174127FEDC9E3F7330D00B" author="Maryan, B. &amp; Gaikhorst, G. &amp; Parkhurst, B." box="[325,545,799,825]" pageId="21" pageNumber="320" pagination="230 - 251" refId="ref21102" refString="Maryan, B., Gaikhorst, G. &amp; Parkhurst, B. (2024) The terrestrial herpetofauna of the Zuytdorp coast and hinterland of Western Australia: Exceptional richness in a global biodiversity hotspot. The Western Australian Naturalist, 33, 230 - 251." type="journal article" year="2024">
 Maryan 
 <emphasis id="56BEEAB08F174127FE3C9E3F70CED00B" box="[421,479,799,825]" italics="true" pageId="21" pageNumber="320">et al.</emphasis>
 2024
 </bibRefCitation>
 ) and a disjunct population of 
 <taxonomicName id="A3CA4D218F174127FCE09E3F7533D00B" baseAuthorityName="Mitchell &amp; Behrndt" baseAuthorityYear="1949" box="[889,1058,799,825]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="multiscutata">
 <emphasis id="56BEEAB08F174127FCE09E3F7533D00B" box="[889,1058,799,825]" italics="true" pageId="21" pageNumber="320">L. multiscutata</emphasis>
 </taxonomicName>
 on Bernier Island, offshore from Carnarvon (
 <bibRefCitation id="005B4B538F174127FE859E6370C9D06F" author="Storr, G. M. &amp; Smith, L. A. &amp; Johnstone, R. E." box="[284,472,835,861]" pageId="21" pageNumber="320" refId="ref22708" refString="Storr, G. M., Smith, L. A. &amp; Johnstone, R. E. (1999) Lizards of Western Australia. I Skinks. Revised Edition. Western Australian Museum, 291 pp." type="book" year="1999">
 Storr 
 <emphasis id="56BEEAB08F174127FEC49E637084D06F" box="[349,405,835,861]" italics="true" pageId="21" pageNumber="320">et al.</emphasis>
 1999
 </bibRefCitation>
 ).
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F174125FF5E9E467564D153" lastPageId="23" lastPageNumber="322" pageId="21" pageNumber="320" type="biology_ecology">
 <paragraph id="647536A28F174127FF5E9E4670DED7DB" blockId="21.[151,1437,151,1905]" pageId="21" pageNumber="320">
 <emphasis id="56BEEAB08F174127FF5E9E467344D0B2" bold="true" box="[199,597,870,897]" pageId="21" pageNumber="320">Habitat, ecology and conservation</emphasis>
 . The 
 <typeStatus id="BB7188008F174127FD099E4773E3D0B3" box="[656,754,871,897]" pageId="21" pageNumber="320" type="holotype">holotype</typeStatus>
 and WAM R156453 were pit or funnel-trapped on a sandplain. The vegetation comprised a low, mixed shrubland of 
 <taxonomicName id="A3CA4D218F174127FD6B9EAB729CD097" authority="Miller" authorityName="Miller" box="[754,909,907,933]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="320" phylum="Tracheophyta" rank="genus">
 <emphasis id="56BEEAB08F174127FD6B9EAB7251D097" box="[754,832,907,933]" italics="true" pageId="21" pageNumber="320">Acacia</emphasis>
 Miller
 </taxonomicName>
 with emergent 
 <taxonomicName id="A3CA4D218F174127FBA79EAB7472D097" authority="Hill &amp; Johnson" authorityName="Hill &amp; Johnson" box="[1086,1379,907,933]" class="Insecta" family="Cerambycidae" genus="Corymbia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="320" phylum="Arthropoda" rank="genus">
 <emphasis id="56BEEAB08F174127FBA79EAB75BFD097" box="[1086,1198,907,933]" italics="true" pageId="21" pageNumber="320">Corymbia</emphasis>
 Hill &amp; Johnson
 </taxonomicName>
 over a mosaic groundcover of low shrubs and 
 <taxonomicName id="A3CA4D218F174127FDE99E8F7206D0FB" authority="Brown" authorityName="Brown" box="[624,791,943,969]" class="Insecta" family="Hepialidae" genus="Triodia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="320" phylum="Arthropoda" rank="genus">
 <emphasis id="56BEEAB08F174127FDE99E8F73D3D0FB" box="[624,706,943,969]" italics="true" pageId="21" pageNumber="320">Triodia</emphasis>
 Brown
 </taxonomicName>
 hummocks, with patches of exposed limestone (
 <bibRefCitation id="005B4B538F174127FADA9E8F7073D0DF" author="Metcalf, B. M. &amp; Bamford, M. J." pageId="21" pageNumber="320" refId="ref21458" refString="Metcalf, B. M. &amp; Bamford, M. J. (2005) Fauna assessment of Exmouth unallocated crown land. Unpublished report for Department of Conservation and Land Management." type="book" year="2005">Metcalf &amp; Bamford 2005</bibRefCitation>
 ; HLA‒Envirosciences, 
 <bibRefCitation id="005B4B538F174127FDEC9EF37284D0DF" author="Ecoscape Australia" box="[629,917,978,1005]" pageId="21" pageNumber="320" refId="ref20141" refString="HLA - Envirosciences, Ecoscape Australia (2005) Learmonth Air Weapons Range ecological survey interim report. Unpublished report for Department of Defence Corporate Services and Infrastructure." type="book" year="2005">
 Ecoscape 
 <collectingCountry id="1CDD76328F174127FD719EF3725ED0DF" box="[744,847,979,1005]" name="Australia" pageId="21" pageNumber="320">Australia</collectingCountry>
 2005
 </bibRefCitation>
 , 
 <figureCitation id="FCF12A278F174127FC3D9EF37510D0DF" box="[932,1025,979,1005]" captionStart="FIGURE 9" captionStartId="22.[151,250,1473,1497]" captionTargetBox="[152,1435,166,1450]" captionTargetId="figure-23@22.[152,1435,166,1450]" captionTargetPageId="22" captionText="FIGURE 9. A) The habitat of L. parva sp. nov., dense Triodia sandplain in swale with sand dunes vegetated with low Acacia and emergent Corymbia on red sand, 8 km N of Exmouth, North-West Cape, Western Australia, B) aerial view of dunefield habitat on North-West Cape, and C) active burrow on crest of a red dune (photos A: B. Maryan, B: L. Clausen, C: S. Wilson)." figureDoi="http://doi.org/10.5281/zenodo.14421242" httpUri="https://zenodo.org/record/14421242/files/figure.png" pageId="21" pageNumber="320">Fig. 9A</figureCitation>
 ). Nearby were stands of low, open woodland of 
 <taxonomicName id="A3CA4D218F174127FEB69ED77335D723" authority="L'Heritier" authorityName="L'Heritier" box="[303,548,1015,1041]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="21" pageNumber="320" phylum="Tracheophyta" rank="genus">
 <emphasis id="56BEEAB08F174127FEB69ED770BAD723" box="[303,427,1015,1041]" italics="true" pageId="21" pageNumber="320">Eucalyptus</emphasis>
 L’Héritier
 </taxonomicName>
 and 
 <taxonomicName id="A3CA4D218F174127FDC79ED772F0D723" authority="Schott &amp; Endlicher." authorityName="Schott &amp; Endlicher." box="[606,993,1015,1041]" class="Magnoliopsida" family="Malvaceae" genus="Brachychiton" kingdom="Plantae" order="Malvales" pageId="21" pageNumber="320" phylum="Tracheophyta" rank="genus">
 <emphasis id="56BEEAB08F174127FDC79ED773E4D723" box="[606,757,1015,1041]" italics="true" pageId="21" pageNumber="320">Brachychiton</emphasis>
 Schott &amp; Endlicher.
 </taxonomicName>
 The associated dune habitat supported predominately 
 <taxonomicName id="A3CA4D218F174127FEA6993B7080D707" box="[319,401,1051,1077]" class="Insecta" family="Hepialidae" genus="Triodia" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="320" phylum="Arthropoda" rank="genus">
 <emphasis id="56BEEAB08F174127FEA6993B7080D707" box="[319,401,1051,1077]" italics="true" pageId="21" pageNumber="320">Triodia</emphasis>
 </taxonomicName>
 with scattered 
 <taxonomicName id="A3CA4D218F174127FDA1993B727FD707" authority="Baker. Field" authorityName="Baker. Field" box="[568,878,1051,1077]" class="Magnoliopsida" family="Proteaceae" genus="Banksia" kingdom="Plantae" order="Proteales" pageId="21" pageNumber="320" phylum="Tracheophyta" rank="species" species="ashbyi">
 <emphasis id="56BEEAB08F174127FDA1993B73F1D707" box="[568,736,1051,1077]" italics="true" pageId="21" pageNumber="320">Banksia ashbyi</emphasis>
 Baker. Field
 </taxonomicName>
 observations of other individuals were pit-trapped on red dunes (L. Clausen, pers. comm., 
 <figureCitation id="FCF12A278F174127FDCD991F73BAD76B" box="[596,683,1087,1113]" captionStart="FIGURE 9" captionStartId="22.[151,250,1473,1497]" captionTargetBox="[152,1435,166,1450]" captionTargetId="figure-23@22.[152,1435,166,1450]" captionTargetPageId="22" captionText="FIGURE 9. A) The habitat of L. parva sp. nov., dense Triodia sandplain in swale with sand dunes vegetated with low Acacia and emergent Corymbia on red sand, 8 km N of Exmouth, North-West Cape, Western Australia, B) aerial view of dunefield habitat on North-West Cape, and C) active burrow on crest of a red dune (photos A: B. Maryan, B: L. Clausen, C: S. Wilson)." figureDoi="http://doi.org/10.5281/zenodo.14421242" httpUri="https://zenodo.org/record/14421242/files/figure.png" pageId="21" pageNumber="320">Fig. 9B</figureCitation>
 ). 
 <specimenCount id="72CCFD2B8F174127FD27991F724FD76B" box="[702,862,1087,1113]" count="1" pageId="21" pageNumber="320" type="generic">One specimen</specimenCount>
 (WAM R153831) was excavated from a burrow on a thinly vegetated crest of a red dune (
 <figureCitation id="FCF12A278F174127FDD6994373B9D74F" box="[591,680,1123,1149]" captionStart="FIGURE 9" captionStartId="22.[151,250,1473,1497]" captionTargetBox="[152,1435,166,1450]" captionTargetId="figure-23@22.[152,1435,166,1450]" captionTargetPageId="22" captionText="FIGURE 9. A) The habitat of L. parva sp. nov., dense Triodia sandplain in swale with sand dunes vegetated with low Acacia and emergent Corymbia on red sand, 8 km N of Exmouth, North-West Cape, Western Australia, B) aerial view of dunefield habitat on North-West Cape, and C) active burrow on crest of a red dune (photos A: B. Maryan, B: L. Clausen, C: S. Wilson)." figureDoi="http://doi.org/10.5281/zenodo.14421242" httpUri="https://zenodo.org/record/14421242/files/figure.png" pageId="21" pageNumber="320">Fig. 9C</figureCitation>
 ). The vegetation of the two main dune systems on the North-West Cape, the northern and southern red sand dunes, have been mapped by 
 <bibRefCitation id="005B4B538F174127FC2499A77547D793" author="Beard, J. S." box="[957,1110,1158,1185]" pageId="21" pageNumber="320" refId="ref18623" refString="Beard, J. S. (1975) Vegetation Surveys of Western Australia. Pilbara 1: 1 000 000 Vegetation Series. Explanatory Notes to Sheet 5. Vegetation of the Pilbara Region. University of Western Australia Press, Perth." type="journal volume" year="1975">Beard (1975)</bibRefCitation>
 . The referred specimen from Merlinleigh Outstation on the Kennedy Range (WAM R8601) has no habitat details but this area is predominantly dunefield/sandplain habitat.
 </paragraph>
 <paragraph id="647536A28F174127FF5E99D37409D6D7" blockId="21.[151,1437,151,1905]" pageId="21" pageNumber="320">
 The ecology of this species is largely unknown. The limited field observations of 
 <taxonomicName id="A3CA4D218F174127FBCA99D375A0D63E" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[1107,1201,1267,1292]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F174127FBCA99D375A0D63E" box="[1107,1201,1267,1292]" italics="true" pageId="21" pageNumber="320">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F174127FB2199D37400D63F" bold="true" box="[1208,1297,1267,1293]" pageId="21" pageNumber="320">
 <taxonomicNameLabel id="4D8D57CB8F174127FB2199D37400D63F" box="[1208,1297,1267,1293]" pageId="21" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 suggest that it is numerous at two localities on the North-West Cape based on captures in traps, appearing to favour the lower slopes of dunes where sand is more compacted (M. Bamford &amp; R. Browne-Cooper, pers. obs.). Typical of other burrow-digging members of 
 <taxonomicName id="A3CA4D218F174127FE40987F7352D64B" authorityName="Fitzinger" authorityYear="1843" box="[473,579,1375,1401]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="genus">
 <emphasis id="56BEEAB08F174127FE40987F7352D64B" box="[473,579,1375,1401]" italics="true" pageId="21" pageNumber="320">Liopholis</emphasis>
 </taxonomicName>
 , their presence can often be indicated by excavated multi-entranced burrows (
 <figureCitation id="FCF12A278F174127FF0698A371E6D6AF" box="[159,247,1411,1437]" captionStart="FIGURE 9" captionStartId="22.[151,250,1473,1497]" captionTargetBox="[152,1435,166,1450]" captionTargetId="figure-23@22.[152,1435,166,1450]" captionTargetPageId="22" captionText="FIGURE 9. A) The habitat of L. parva sp. nov., dense Triodia sandplain in swale with sand dunes vegetated with low Acacia and emergent Corymbia on red sand, 8 km N of Exmouth, North-West Cape, Western Australia, B) aerial view of dunefield habitat on North-West Cape, and C) active burrow on crest of a red dune (photos A: B. Maryan, B: L. Clausen, C: S. Wilson)." figureDoi="http://doi.org/10.5281/zenodo.14421242" httpUri="https://zenodo.org/record/14421242/files/figure.png" pageId="21" pageNumber="320">Fig. 9C</figureCitation>
 ). 
 <bibRefCitation id="005B4B538F174127FE9098A37353D6AF" author="Bamford, M. J. &amp; Bamford, A. R." box="[265,578,1411,1437]" pageId="21" pageNumber="320" pagination="107 - 111" refId="ref18577" refString="Bamford, M. J. &amp; Bamford, A. R. (2012) Behavioural observations on an unnamed species of skink Liopholis (formerly Egernia) &quot; Cape Range &quot; from Cape Range. Western Australian Naturalist 28, 107 - 111." type="journal article" year="2012">Bamford &amp; Bamford (2012)</bibRefCitation>
 provide details of behavioural observations and captive breeding for 
 <taxonomicName id="A3CA4D218F174127FAA698A3748DD6AE" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[1343,1436,1411,1436]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F174127FAA698A3748DD6AE" box="[1343,1436,1411,1436]" italics="true" pageId="21" pageNumber="320">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F174127FF0E988771E1D6F3" bold="true" box="[151,240,1447,1473]" pageId="21" pageNumber="320">
 <taxonomicNameLabel id="4D8D57CB8F174127FF0E988771E1D6F3" box="[151,240,1447,1473]" pageId="21" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 listed as 
 <taxonomicName id="A3CA4D218F174127FECE988770D0D6F3" authority="Cape Range" authorityName="Fitzinger" authorityYear="1843" box="[343,449,1447,1473]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="genus">
 <emphasis id="56BEEAB08F174127FECE988770D0D6F3" box="[343,449,1447,1473]" italics="true" pageId="21" pageNumber="320">Liopholis</emphasis>
 </taxonomicName>
 “Cape Range”. Their information also included the SVL measurements (in mm) of 
 <quantity id="A3329B478F174127FAFD9887748DD6F3" box="[1380,1436,1447,1473]" metricMagnitude="0" metricUnit="m" metricValue="1.5748" pageId="21" pageNumber="320" unit="in" value="62.0">62 in</quantity>
 <specimenCount id="72CCFD2B8F174127FF0E98EB71EED6D7" box="[151,255,1483,1509]" count="1" pageId="21" pageNumber="320" type="adult">one adult</specimenCount>
 male and 56 and 
 <quantity id="A3329B478F174127FE5C98EA70EFD6D7" box="[453,510,1482,1509]" metricMagnitude="0" metricUnit="m" metricValue="1.4478" pageId="21" pageNumber="320" unit="in" value="57.0">57 in</quantity>
 <specimenCount id="72CCFD2B8F174127FD9C98EB737FD6D7" box="[517,622,1483,1509]" count="2" pageId="21" pageNumber="320" type="adult">two adult</specimenCount>
 females that displayed no growth after six years in captivity.
 </paragraph>
 <paragraph id="647536A28F174127FF5E98CF70DDD443" blockId="21.[151,1437,151,1905]" pageId="21" pageNumber="320">
 The current land use of the locations where 
 <taxonomicName id="A3CA4D218F174127FD2D98CF7202D53A" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[692,787,1519,1544]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F174127FD2D98CF7202D53A" box="[692,787,1519,1544]" italics="true" pageId="21" pageNumber="320">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F174127FC8398CF7265D53B" bold="true" box="[794,884,1519,1545]" pageId="21" pageNumber="320">
 <taxonomicNameLabel id="4D8D57CB8F174127FC8398CF7265D53B" box="[794,884,1519,1545]" pageId="21" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 are known from comprise a mixture of protected gazetted conservation, Commonwealth Defence and non-protected areas. Currently, within the protected areas, 
 <taxonomicName id="A3CA4D218F174127FF0E9B1771E9D562" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[151,248,1591,1616]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F174127FF0E9B1771E9D562" box="[151,248,1591,1616]" italics="true" pageId="21" pageNumber="320">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F174127FE9B9B17704FD563" bold="true" box="[258,350,1591,1617]" pageId="21" pageNumber="320">
 <taxonomicNameLabel id="4D8D57CB8F174127FE9B9B17704FD563" box="[258,350,1591,1617]" pageId="21" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 has been recorded from single localities in both the Cape Range National Park (
 <quantity id="A3329B478F174127FA999B177448D563" box="[1280,1369,1591,1617]" metricMagnitude="5" metricUnit="m" metricValue="4.76" pageId="21" pageNumber="320" unit="km" value="476.0">476 km</quantity>
 <superScript id="93BF9BEA8F174127FAC09B167470D576" attach="left" box="[1369,1377,1590,1604]" fontSize="6" pageId="21" pageNumber="320">2</superScript>
 ) and Learmonth Air Weapons Range (
 <quantity id="A3329B478F174127FD939B7B7370D547" box="[522,609,1627,1653]" metricMagnitude="5" metricUnit="m" metricValue="1.89" pageId="21" pageNumber="320" unit="km" value="189.0">189 km</quantity>
 <superScript id="93BF9BEA8F174127FDF89B7A7378D55A" attach="left" box="[609,617,1626,1640]" fontSize="6" pageId="21" pageNumber="320">2</superScript>
 ). Commonwealth Defence land is not routinely considered conservation estate, but it does fall under heritage management objectives that includes natural heritage in compliance with the Australian Government’s Environment Protection and Biodiversity Act (EPBC Act). The non-protected areas include both an active pastoral station and unallocated Crown Land (e.g. 
 <bibRefCitation id="005B4B538F174127FC489BE7756AD5D3" author="Meissner, R. A." box="[977,1147,1735,1761]" pageId="21" pageNumber="320" refId="ref21332" refString="Meissner, R. A. (2011) Biodiversity values of unallocated Crown Land on Cape Range peninsula, Western Australia. Department of Environment and Conservation, Perth, 45 pp." type="book" year="2011">Meissner 2011</bibRefCitation>
 ). This mosaic of different land tenures is noted by 
 <bibRefCitation id="005B4B538F174127FE3C9BCB7389D437" author="Doughty, P. &amp; Oliver, P. &amp; Adams, M." box="[421,664,1771,1797]" pageId="21" pageNumber="320" pagination="247 - 265" refId="ref19354" refString="Doughty, P., Oliver, P. &amp; Adams, M. (2008) Systematics of stone geckos in the genus Diplodactylus (Reptilia: Diplodactylidae) from northwestern Australia, with a description of a new species from the Northwest Cape, Western Australia. Records of the Western Australian Museum, 24, 247 - 265. https: // doi. org / 10.18195 / issn. 0312 - 3162.24 (3). 2008.247 - 265" type="journal article" year="2008">
 Doughty 
 <emphasis id="56BEEAB08F174127FD979BCB7354D437" box="[526,581,1771,1797]" italics="true" pageId="21" pageNumber="320">et al.</emphasis>
 (2008)
 </bibRefCitation>
 , whereby only the Cape Range is currently protected within the Cape Range National Park, excluding much of the North-West Cape from the same statutory protection. The single record of 
 <taxonomicName id="A3CA4D218F174127FF2E9A137009D47E" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[183,280,1843,1868]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="21" pageNumber="320" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F174127FF2E9A137009D47E" box="[183,280,1843,1868]" italics="true" pageId="21" pageNumber="320">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F174127FEB89A13706CD47F" bold="true" box="[289,381,1843,1869]" pageId="21" pageNumber="320">
 <taxonomicNameLabel id="4D8D57CB8F174127FEB89A13706CD47F" box="[289,381,1843,1869]" pageId="21" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 from Merlinleigh Outstation on the Kennedy Range is protected within the Kennedy Range National Park (
 <quantity id="A3329B478F174127FEDD9A7770A2D443" box="[324,435,1879,1905]" metricMagnitude="6" metricUnit="m" metricValue="1.416" pageId="21" pageNumber="320" unit="km" value="1416.0">1, 416 km</quantity>
 <superScript id="93BF9BEA8F174127FE2A9A7670AAD456" attach="left" box="[435,443,1878,1892]" fontSize="6" pageId="21" pageNumber="320">2</superScript>
 ).
 </paragraph>
 <caption id="30B5662A8F144124FF0E98E17480D510" ID-DOI="http://doi.org/10.5281/zenodo.14421242" ID-Zenodo-Dep="14421242" httpUri="https://zenodo.org/record/14421242/files/figure.png" pageId="22" pageNumber="321" startId="22.[151,250,1473,1497]" targetBox="[152,1435,166,1450]" targetPageId="22" targetType="figure">
 <paragraph id="647536A28F144124FF0E98E17480D510" blockId="22.[151,1437,1473,1570]" pageId="22" pageNumber="321">
 <emphasis id="56BEEAB08F144124FF0E98E1703AD6EB" bold="true" box="[151,299,1473,1498]" pageId="22" pageNumber="321">FIGURE 9. A</emphasis>
 ) The habitat of 
 <taxonomicName id="A3CA4D218F144124FE4998E27336D6EB" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[464,551,1474,1497]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="22" pageNumber="321" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F144124FE4998E27336D6EB" box="[464,551,1474,1497]" italics="true" pageId="22" pageNumber="321">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F144124FDB798E2736ED6E8" bold="true" box="[558,639,1474,1498]" pageId="22" pageNumber="321">
 <taxonomicNameLabel id="4D8D57CB8F144124FDB798E2736ED6E8" box="[558,639,1474,1498]" pageId="22" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 , dense 
 <taxonomicName id="A3CA4D218F144124FD5498E27209D6E8" box="[717,792,1474,1498]" class="Insecta" family="Hepialidae" genus="Triodia" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="321" phylum="Arthropoda" rank="genus">
 <emphasis id="56BEEAB08F144124FD5498E27209D6E8" box="[717,792,1474,1498]" italics="true" pageId="22" pageNumber="321">Triodia</emphasis>
 </taxonomicName>
 sandplain in swale with sand dunes vegetated with low 
 <taxonomicName id="A3CA4D218F144124FACF98E2748DD6EB" box="[1366,1436,1474,1497]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="22" pageNumber="321" phylum="Tracheophyta" rank="genus">
 <emphasis id="56BEEAB08F144124FACF98E2748DD6EB" box="[1366,1436,1474,1497]" italics="true" pageId="22" pageNumber="321">Acacia</emphasis>
 </taxonomicName>
 and emergent 
 <taxonomicName id="A3CA4D218F144124FEB398C6709ED6CC" box="[298,399,1510,1534]" class="Insecta" family="Cerambycidae" genus="Corymbia" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="321" phylum="Arthropoda" rank="genus">
 <emphasis id="56BEEAB08F144124FEB398C6709ED6CC" box="[298,399,1510,1534]" italics="true" pageId="22" pageNumber="321">Corymbia</emphasis>
 </taxonomicName>
 on red sand, 8 km N of Exmouth, North-West Cape, Western Australia, 
 <emphasis id="56BEEAB08F144124FB1D98C67584D6CC" bold="true" box="[1156,1173,1510,1534]" pageId="22" pageNumber="321">B</emphasis>
 ) aerial view of dunefield habitat on North-West Cape, and 
 <emphasis id="56BEEAB08F144124FE739B2970EDD513" bold="true" box="[490,508,1545,1569]" pageId="22" pageNumber="321">C</emphasis>
 ) active burrow on crest of a red dune (photos A: B. Maryan, B: L. Clausen, C: S. Wilson).
 </paragraph>
 </caption>
 <paragraph id="647536A28F144124FF5E9B70708CD4C4" blockId="22.[151,1436,1616,2038]" pageId="22" pageNumber="321">
 Similarly to other relatively small 
 <taxonomicName id="A3CA4D218F144124FDDC9B707217D558" box="[581,774,1616,1642]" pageId="22" pageNumber="321">
 <emphasis id="56BEEAB08F144124FDDC9B7073BED558" box="[581,687,1616,1642]" italics="true" pageId="22" pageNumber="321">Liopholis</emphasis>
 species
 </taxonomicName>
 with restricted distributions (
 <bibRefCitation id="005B4B538F144124FBD59B70743AD558" author="Chapple, D. G. &amp; Tingley, R. &amp; Mitchell, N. J. &amp; Macdonald, S. L. &amp; Keogh, J. S. &amp; Shea, G. M. &amp; Bowles, P. &amp; Cox, N. A. &amp; Woinarski, J. C. Z." box="[1100,1323,1616,1642]" pageId="22" pageNumber="321" refId="ref19115" refString="Chapple, D. G., Tingley, R., Mitchell, N. J., Macdonald, S. L., Keogh, J. S., Shea, G. M., Bowles, P., Cox, N. A. &amp; Woinarski, J. C. Z. (2019) The Action Plan for Australian Lizards and Snakes 2017. CSIRO Publishing, Melbourne, 663 pp." type="book" year="2019">
 Chapple 
 <emphasis id="56BEEAB08F144124FB2B9B7175F8D558" box="[1202,1257,1616,1642]" italics="true" pageId="22" pageNumber="321">et al.</emphasis>
 2019
 </bibRefCitation>
 ), 
 <taxonomicName id="A3CA4D218F144124FAA79B71748DD558" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[1342,1436,1616,1642]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="22" pageNumber="321" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F144124FAA79B71745FD558" box="[1342,1358,1617,1642]" italics="true" pageId="22" pageNumber="321">L</emphasis>
 . 
 <emphasis id="56BEEAB08F144124FAC29B71748DD558" box="[1371,1436,1617,1642]" italics="true" pageId="22" pageNumber="321">parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F144124FF0E9B5471E2D5BC" bold="true" box="[151,243,1652,1678]" pageId="22" pageNumber="321">
 <taxonomicNameLabel id="4D8D57CB8F144124FF0E9B5471E2D5BC" box="[151,243,1652,1678]" pageId="22" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 faces potential threats that warrant conservation attention. As highlighted for 
 <emphasis id="56BEEAB08F144124FBE99B5575E3D5BC" box="[1136,1266,1652,1678]" italics="true" pageId="22" pageNumber="321">
 L. s. 
 <taxonomicName id="A3CA4D218F144124FB339B5475E3D5BC" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[1194,1266,1652,1678]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="22" pageNumber="321" phylum="Chordata" rank="species" species="slateri">slateri</taxonomicName>
 </emphasis>
 populations in central 
 <collectingCountry id="1CDD76328F144124FF739BB87040D580" box="[234,337,1688,1714]" name="Australia" pageId="22" pageNumber="321">Australia</collectingCountry>
 (
 <bibRefCitation id="005B4B538F144124FEF99BB870FBD580" author="Pavey, C." box="[352,490,1688,1714]" pageId="22" pageNumber="321" refId="ref21885" refString="Pavey, C. (2004) Recovery plan for Slater's Skink, Egernia slateri, 2005 - 2010. Northern Territory Department of Infrastructure, Planning and Environment, Darwin." type="book" year="2004">Pavey 2004</bibRefCitation>
 , 
 <bibRefCitation id="005B4B538F144124FE6F9BB87320D580" author="Pavey, C." box="[502,561,1688,1714]" pageId="22" pageNumber="321" pagination="176 - 177" refId="ref21917" refString="Pavey, C. (2007) Slater's Skink Egernia slateri. In: J. Woinarski, C. Pavey, R. Kerrigan, I. Cowie and S. Ward (Eds.), Lost from Our Landscape: Threatened species of the Northern Territory, Northern Territory Government, Darwin, Northern Territory, Australia, pp. 176 - 177." type="book chapter" year="2007">2007</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F144124FDA79BB87219D580" author="Pavey, C. R. &amp; Burwell, C. J. &amp; Nano, C. E." box="[574,776,1688,1714]" pageId="22" pageNumber="321" pagination="563 - 571" refId="ref21984" refString="Pavey, C. R., Burwell, C. J. &amp; Nano, C. E. (2010) Foraging ecology and habitat use of Slater's skink (Egernia slateri): An endangered Australian desert lizard. Journal of Herpetology, 44, 563 - 571." type="journal article" year="2010">
 Pavey 
 <emphasis id="56BEEAB08F144124FD129BB973ACD580" box="[651,701,1688,1714]" italics="true" pageId="22" pageNumber="321">et al</emphasis>
 . 2010
 </bibRefCitation>
 ), the species is potentially impacted by the degradation of existing habitat caused by the invasion of the introduced pasture plant buffel grass 
 <taxonomicName id="A3CA4D218F144124FBBF9B9C7442D5E4" authority="Linnaeus" authorityName="Linnaeus" box="[1062,1363,1724,1750]" class="Liliopsida" family="Poaceae" genus="Cenchrus" kingdom="Plantae" order="Poales" pageId="22" pageNumber="321" phylum="Tracheophyta" rank="species" species="ciliaris">
 <emphasis id="56BEEAB08F144124FBBF9B9C75F7D5E4" box="[1062,1254,1724,1750]" italics="true" pageId="22" pageNumber="321">Cenchrus ciliaris</emphasis>
 Linnaeus
 </taxonomicName>
 on the North-West Cape (
 <bibRefCitation id="005B4B538F144124FEF39BC0739DD5C8" author="Keighery, G. &amp; Gibson, N." box="[362,652,1760,1786]" pageId="22" pageNumber="321" pagination="51 - 85" refId="ref20471" refString="Keighery, G. &amp; Gibson, N. (1993) Biogeography and composition of the flora of the Cape Range peninsula, Western Australia. In: W. F. Humphreys (Ed.) The Biogeography of Cape Range, Western Australia. Records of the Western Australian Museum, 45 (Supplement), 51 - 85." type="journal article" year="1993">Keighery &amp; Gibson 1993</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F144124FD009BC07252D5C8" author="Meissner, R. A." box="[665,835,1760,1786]" pageId="22" pageNumber="321" refId="ref21332" refString="Meissner, R. A. (2011) Biodiversity values of unallocated Crown Land on Cape Range peninsula, Western Australia. Department of Environment and Conservation, Perth, 45 pp." type="book" year="2011">Meissner 2011</bibRefCitation>
 ). The success of buffel grass as an invasive weed not only impacts on the vegetation communities but also on the fauna that use them for habitat and refuge (
 <bibRefCitation id="005B4B538F144124FA8F9A2471C2D470" author="Dixon, I. R. &amp; Dixon, K. W. &amp; Barret, M." pageId="22" pageNumber="321" pagination="92 - 101" refId="ref19265" refString="Dixon, I. R., Dixon, K. W. &amp; Barret, M. (2001) Eradication of buffel grass (Cenchrus ciliaris) on Airlie Island, Pilbara coast, Western Australia. In: C. R. Veitch and M. N. Clout (Eds.) Turning the Tide in the Eradication of Invasive Species. IUCN SSC Invasive Species Specialist Group, IUCN, Gland, Switzerland and Cambridge, UK, pp. 92 - 101." type="book chapter" year="2001">
 Dixon 
 <emphasis id="56BEEAB08F144124FAFD9A257484D42C" box="[1380,1429,1796,1822]" italics="true" pageId="22" pageNumber="321">et al</emphasis>
 . 2001
 </bibRefCitation>
 ). For example, it has been further hypothesised that the foraging behaviour of 
 <emphasis id="56BEEAB08F144124FBA69A0975ACD470" box="[1087,1213,1832,1858]" italics="true" pageId="22" pageNumber="321">
 L. s. 
 <taxonomicName id="A3CA4D218F144124FBEC9A0875ACD470" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[1141,1213,1832,1858]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="22" pageNumber="321" phylum="Chordata" rank="species" species="slateri">slateri</taxonomicName>
 </emphasis>
 , which rely on high visibility outside their burrow entrance to ambush prey, makes them particularly sensitive to the impacts of buffel grass (
 <bibRefCitation id="005B4B538F144124FF799A5070CED4B8" author="McKinney, M. A. &amp; Schlesinger, C. A. &amp; Pavey, C. R." box="[224,479,1904,1930]" pageId="22" pageNumber="321" pagination="477 - 482" refId="ref21275" refString="McKinney, M. A., Schlesinger, C. A. &amp; Pavey, C. R. (2014) Foraging behaviour of the endangered Australian skink (Liopholis slateri). Australian Journal of Zoology, 62, 477 - 482. https: // doi. org / 10.1071 / ZO 14071" type="journal article" year="2014">
 McKinney 
 <emphasis id="56BEEAB08F144124FEF89A51708BD4B8" box="[353,410,1904,1930]" italics="true" pageId="22" pageNumber="321">et al.</emphasis>
 2014
 </bibRefCitation>
 ). The trampling by feral herbivores and cattle in addition to altered fire regimes may also impact the long-term survival of the habitat occupied by 
 <taxonomicName id="A3CA4D218F144124FCCB9AB572A3D49C" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[850,946,1941,1966]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="22" pageNumber="321" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F144124FCCB9AB572A3D49C" box="[850,946,1941,1966]" italics="true" pageId="22" pageNumber="321">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F144124FC239AB47504D49C" bold="true" box="[954,1045,1940,1966]" pageId="22" pageNumber="321">
 <taxonomicNameLabel id="4D8D57CB8F144124FC239AB47504D49C" box="[954,1045,1940,1966]" pageId="22" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 Another significant consideration is the predation by cats and foxes that are both known to take small lizard prey; thus, juveniles are likely to be particularly vulnerable.
 </paragraph>
 <paragraph id="647536A28F154125FF5E9DB77564D153" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="322">
 Available knowledge indicates this species occurs in fragmented populations on the North-West Cape associated with two main dune systems, the northern sand dunes that occur within unallocated Crown Land, and the southern sand dunes that are partially protected within the Cape Range National Park (
 <bibRefCitation id="005B4B538F154125FB969DFF75ACD3CB" author="Meissner, R. A." box="[1039,1213,223,249]" pageId="23" pageNumber="322" refId="ref21332" refString="Meissner, R. A. (2011) Biodiversity values of unallocated Crown Land on Cape Range peninsula, Western Australia. Department of Environment and Conservation, Perth, 45 pp." type="book" year="2011">Meissner 2011</bibRefCitation>
 ). Based on existing collection records, including the Kennedy Range National Park (N = 4, 
 <bibRefCitation id="005B4B538F154125FC219C23741ED22F" author="Atlas of Living Australia" box="[952,1295,259,285]" pageId="23" pageNumber="322" refId="ref18479" refString="Atlas of Living Australia (2024) Available from: Atlas of Living Australia - Open access to Australia's biodiversity data (ala. org. au) (accessed 12 July 2024)" type="book" year="2024">
 Atlas of Living 
 <collectingCountry id="1CDD76328F154125FBF19C2375DED22F" box="[1128,1231,259,285]" name="Australia" pageId="23" pageNumber="322">Australia</collectingCountry>
 2024
 </bibRefCitation>
 ) and limited field observations, we calculated the extent of occurrence(EOO; minimum convex hull) and area of occupancy(AOO; 2 x 
 <quantity id="A3329B478F154125FF269C6B71E6D257" box="[191,247,331,357]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="322" unit="km" value="2.0">2 km</quantity>
 grid cells) of 
 <taxonomicName id="A3CA4D218F154125FE089C6B70FFD256" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[401,494,331,356]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F154125FE089C6B70FFD256" box="[401,494,331,356]" italics="true" pageId="23" pageNumber="322">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F154125FE6C9C6B735FD257" bold="true" box="[501,590,331,357]" pageId="23" pageNumber="322">
 <taxonomicNameLabel id="4D8D57CB8F154125FE6C9C6B735FD257" box="[501,590,331,357]" pageId="23" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 in GeoCat (https;//geocat.kew.org; 
 <bibRefCitation id="005B4B538F154125FC409C6B75D3D257" author="Bachman, S. &amp; Moat, J. &amp; Hill, A. W. &amp; de la Torre, J. &amp; Scott, B." box="[985,1218,331,357]" pageId="23" pageNumber="322" pagination="117 - 126" refId="ref18513" refString="Bachman, S., Moat, J., Hill, A. W., de la Torre, J. &amp; Scott, B. (2011) Supporting Red List threat assessments with GeoCAT: geospatial conservation assessment tool. ZooKeys, 150, 117 - 126. https: // doi. org / 10.3897 / zookeys. 150.2109" type="journal article" year="2011">
 Bachman 
 <emphasis id="56BEEAB08F154125FBD09C6B7591D257" box="[1097,1152,331,357]" italics="true" pageId="23" pageNumber="322">et al.</emphasis>
 2011
 </bibRefCitation>
 ). The species’ EOO is 
 <quantity id="A3329B478F154125FF289C4F703ED2BB" box="[177,303,367,393]" metricMagnitude="7" metricUnit="m" metricValue="1.0076" pageId="23" pageNumber="322" unit="km" value="10076.0">10, 076 km</quantity>
 <superScript id="93BF9BEA8F154125FEB69C4E7026D24E" attach="left" box="[303,311,366,380]" fontSize="6" pageId="23" pageNumber="322">2</superScript>
 and the AOO is considerably smaller at 
 <quantity id="A3329B478F154125FD619C4F722ED2BB" box="[760,831,367,393]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="23" pageNumber="322" unit="km" value="24.0">24 km</quantity>
 <superScript id="93BF9BEA8F154125FCA79C4E7257D24E" attach="left" box="[830,838,366,380]" fontSize="6" pageId="23" pageNumber="322">2</superScript>
 . Targeted biological surveys on the North-West Cape and Kennedy Range in suitable habitat for this species would improve the issue. Current distributional knowledge of 
 <taxonomicName id="A3CA4D218F154125FF2C9C977003D2E2" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[181,274,439,464]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F154125FF2C9C977003D2E2" box="[181,274,439,464]" italics="true" pageId="23" pageNumber="322">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F154125FE809C977060D2E3" bold="true" box="[281,369,439,465]" pageId="23" pageNumber="322">
 <taxonomicNameLabel id="4D8D57CB8F154125FE809C977060D2E3" box="[281,369,439,465]" pageId="23" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 allows for an IUCN listing assessment under criteria B2 of geographic range (AOO &lt;
 <quantity id="A3329B478F154125FAA99C967495D2E3" box="[1328,1412,438,465]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="322" unit="km" value="500.0">500 km</quantity>
 <superScript id="93BF9BEA8F154125FA1D9C96749DD2F6" attach="left" box="[1412,1420,438,452]" fontSize="6" pageId="23" pageNumber="322">2</superScript>
 ), B(a) number of locations (equal to 5) and possibly B(biii) reduction in quality of habitat due to buffel grass invasion and altered fire regimes, that qualifies this species for Endangered, under IUCN guidelines and criteria (
 <bibRefCitation id="005B4B538F154125FACF9CDF71C5D10F" author="IUCN" pageId="23" pageNumber="322" refId="ref20318" refString="IUCN (2022) The Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1 (July 2022). Available from: https: // www. iucnredlist. org / resources / redlistguidelines (accessed 12 th July 2024)" type="journal volume" year="2022">IUCN 2022</bibRefCitation>
 ). Further consideration is that without any detailed knowledge of the species’ ecology, life history, distribution and threats to effectively assess the species, it may also be considered as Data Deficient.
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F154125FF5E9F4B72AED51F" pageId="23" pageNumber="322" type="discussion">
 <paragraph id="647536A28F154125FF5E9F4B72FFD00B" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="322">
 <emphasis id="56BEEAB08F154125FF5E9F4B7025D1B7" bold="true" box="[199,308,619,645]" pageId="23" pageNumber="322">Remarks</emphasis>
 . 
 <specimenCount id="72CCFD2B8F154125FEDD9F4B70F7D1B7" box="[324,486,619,645]" count="1" pageId="23" pageNumber="322" type="generic">One specimen</specimenCount>
 (WAM R8601) from Merlinleigh Outstation on the Kennedy Range dunefield was included in 
 <taxonomicName id="A3CA4D218F154125FEB89FAF70B0D19A" box="[289,417,655,680]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F154125FEB89FAF70B0D19A" box="[289,417,655,680]" italics="true" pageId="23" pageNumber="322">L. inornata</emphasis>
 </taxonomicName>
 by 
 <bibRefCitation id="005B4B538F154125FE489FAF739AD19B" author="Storr, G. M." box="[465,651,654,681]" pageId="23" pageNumber="322" pagination="51 - 62" refId="ref22410" refString="Storr, G. M. (1968) Revision of the Egernia whitei species-group (Lacertilia: Scincidae). Journal of the Royal Society of Western Austustralia, 51, 51 - 62." type="journal article" year="1968">Storr (1968: 59)</bibRefCitation>
 and mapped as this taxon by 
 <bibRefCitation id="005B4B538F154125FC729FAF75FDD19B" author="Storr, G. M. &amp; Smith, L. A. &amp; Johnstone, R. E." box="[1003,1260,655,681]" pageId="23" pageNumber="322" refId="ref22446" refString="Storr, G. M., Smith, L. A. &amp; Johnstone, R. E. (1981) Lizards of Western Australia. I Skinks. Western Australian Museum, 200 pp." type="book" year="1981">
 Storr 
 <emphasis id="56BEEAB08F154125FBB59FAF7579D19B" box="[1068,1128,655,681]" italics="true" pageId="23" pageNumber="322">et al.</emphasis>
 (1981: 89)
 </bibRefCitation>
 . However, in a later revision 
 <bibRefCitation id="005B4B538F154125FEAB9F937325D1FF" author="Storr, G. M. &amp; Smith, L. A. &amp; Johnstone, R. E." box="[306,564,691,717]" pageId="23" pageNumber="322" refId="ref22708" refString="Storr, G. M., Smith, L. A. &amp; Johnstone, R. E. (1999) Lizards of Western Australia. I Skinks. Revised Edition. Western Australian Museum, 291 pp." type="book" year="1999">
 Storr 
 <emphasis id="56BEEAB08F154125FEE89F9370BBD1FF" box="[369,426,691,717]" italics="true" pageId="23" pageNumber="322">et al.</emphasis>
 (1999: 112)
 </bibRefCitation>
 omitted this record from the species map, presumably due to emerging doubt over its true affinity. Despite the faded condition of the specimen which was accessioned over 80 years ago, the morphological evidence supports its inclusion within 
 <taxonomicName id="A3CA4D218F154125FD759FDB725BD026" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[748,842,763,788]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F154125FD759FDB725BD026" box="[748,842,763,788]" italics="true" pageId="23" pageNumber="322">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F154125FCC89FDB72B8D027" bold="true" box="[849,937,763,789]" pageId="23" pageNumber="322">
 <taxonomicNameLabel id="4D8D57CB8F154125FCC89FDB72B8D027" box="[849,937,763,789]" pageId="23" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 Further material from the Kennedy Range is required to clarify the genetic affinities of this significant outlier population.
 </paragraph>
 <paragraph id="647536A28F154125FF5E9E6372B9D707" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="322">
 <bibRefCitation id="005B4B538F154125FF5E9E6370A6D06F" author="Shea, G. M. &amp; Wells, R. W." box="[199,439,834,861]" pageId="23" pageNumber="322" refId="ref22133" refString="Shea, G. M. &amp; Wells, R. W. (1985) Egernia inornata, an additional reptile record for the Shark Bay region. Western Australian Naturalist, 16, 91. https: // doi. org / 10.3853 / j. 0067 - 1975.37.1985.337" type="journal volume" year="1985">Shea &amp; Wells (1985)</bibRefCitation>
 referred a 
 <taxonomicName id="A3CA4D218F154125FDAF9E6373B1D06F" authorityName="Fitzinger" authorityYear="1843" box="[566,672,835,861]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="genus">
 <emphasis id="56BEEAB08F154125FDAF9E6373B1D06F" box="[566,672,835,861]" italics="true" pageId="23" pageNumber="322">Liopholis</emphasis>
 </taxonomicName>
 specimen (AMS R.105730) from Peron Peninsula at Shark Bay to 
 <taxonomicName id="A3CA4D218F154125FF0E9E477004D0B2" box="[151,277,871,896]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F154125FF0E9E477004D0B2" box="[151,277,871,896]" italics="true" pageId="23" pageNumber="322">L. inornata</emphasis>
 </taxonomicName>
 and this record was included on the updated map of 
 <bibRefCitation id="005B4B538F154125FCF49E47752BD0B3" author="Storr, G. M. &amp; Smith, L. A. &amp; Johnstone, R. E." box="[877,1082,871,897]" pageId="23" pageNumber="322" refId="ref22708" refString="Storr, G. M., Smith, L. A. &amp; Johnstone, R. E. (1999) Lizards of Western Australia. I Skinks. Revised Edition. Western Australian Museum, 291 pp." type="book" year="1999">
 Storr 
 <emphasis id="56BEEAB08F154125FC349E4772CFD0B3" box="[941,990,871,897]" italics="true" pageId="23" pageNumber="322">et al</emphasis>
 . (1999)
 </bibRefCitation>
 . We agree with this assessment based on its colour pattern and the following characteristics and measurements (in mm): SVL 71.7; tail broken; HD 8.9; HL 16.4; HW 13.5; HNL 28.1; FLL 13.6; HLL 18.2; AGL 37.2; MBS 38; PV 50; SDL 22; nasals widely separated; prefrontals in broad contact; supraciliaries 8/8; nuchals 1/1; upper palpebrals 10/10; primary temporal single; enlarged upper secondary temporals 1/1, each with one adjacent smaller temporal bordering the parietal; supralabials 7/7; ear lobules 3/4, first from top lobule largest in series.
 </paragraph>
 <paragraph id="647536A28F154125FF5E991F72ABD63F" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="322">
 The fact that only a few specimens of 
 <taxonomicName id="A3CA4D218F154125FDFB991F73DDD76B" authorityName="Fitzinger" authorityYear="1843" box="[610,716,1087,1113]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="genus">
 <emphasis id="56BEEAB08F154125FDFB991F73DDD76B" box="[610,716,1087,1113]" italics="true" pageId="23" pageNumber="322">Liopholis</emphasis>
 </taxonomicName>
 have been collected in each of the Kennedy Range and Shark Bay regions, despite combined biological survey effort and numerous visits by field herpetologists in both areas (
 <bibRefCitation id="005B4B538F154125FAFB99437052D793" author="Storr, G. M. &amp; Harold, G." pageId="23" pageNumber="322" pagination="449 - 467" refId="ref22484" refString="Storr, G. M. &amp; Harold, G. (1978) Herpetofauna of the Shark Bay Region, Western Australia. Records of the Western Australian Museum, 6, 449 - 467." type="journal article" year="1978">Storr &amp; Harold 1978</bibRefCitation>
 , 
 <bibRefCitation id="005B4B538F154125FED799A77098D793" author="Storr, G. M. &amp; Harold, G." box="[334,393,1159,1185]" pageId="23" pageNumber="322" pagination="279 - 285" refId="ref22634" refString="Storr, G. M. &amp; Harold, G. (1990) Amphibians and Reptiles of the Shark Bay area, Western Australia. In: P. F. Berry, S. D. Bradshaw and B. R. Wilson (Eds.) Research in Shark Bay: Report of the France-Australe Bicentenary Expedition Committee. Western Australian Museum, Perth, pp. 279 - 285." type="book chapter" year="1990">1990</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F154125FE0D99A7737FD793" author="Shea, G. M. &amp; Wells, R. W." box="[404,622,1158,1185]" pageId="23" pageNumber="322" refId="ref22133" refString="Shea, G. M. &amp; Wells, R. W. (1985) Egernia inornata, an additional reptile record for the Shark Bay region. Western Australian Naturalist, 16, 91. https: // doi. org / 10.3853 / j. 0067 - 1975.37.1985.337" type="journal volume" year="1985">Shea &amp; Wells 1985</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F154125FDE099A7727ED793" author="McKenzie, N. L. &amp; Rolfe, J. K. &amp; Aplin, K. P. &amp; Cowan, M. A. &amp; Smith, L. A." box="[633,879,1159,1185]" pageId="23" pageNumber="322" pagination="335 - 360" refId="ref21182" refString="McKenzie, N. L., Rolfe, J. K., Aplin, K. P., Cowan, M. A. &amp; Smith, L. A. (2000) Herpetofauna of the southern Carnarvon Basin, Western Australia. In: A. H. Burbidge, M. S. Harvey and N. L. McKenzie (Eds.) Biodiversity of the southern Carnarvon Basin. Records of the Western Australian Museum, 61 (Supplement), 335 - 360." type="journal article" year="2000">
 McKenzie 
 <emphasis id="56BEEAB08F154125FD6D99A7723DD793" box="[756,812,1159,1185]" italics="true" pageId="23" pageNumber="322">et al.</emphasis>
 2000
 </bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F154125FCE099A77540D793" author="Maryan, B. &amp; Gaikhorst, G. &amp; Parkhurst, B." box="[889,1105,1159,1185]" pageId="23" pageNumber="322" pagination="230 - 251" refId="ref21102" refString="Maryan, B., Gaikhorst, G. &amp; Parkhurst, B. (2024) The terrestrial herpetofauna of the Zuytdorp coast and hinterland of Western Australia: Exceptional richness in a global biodiversity hotspot. The Western Australian Naturalist, 33, 230 - 251." type="journal article" year="2024">
 Maryan 
 <emphasis id="56BEEAB08F154125FC4E99A7751ED793" box="[983,1039,1159,1185]" italics="true" pageId="23" pageNumber="322">et al.</emphasis>
 2024
 </bibRefCitation>
 ), highlights the serendipity in locating populations of this genus in arid sandy habitats. For example, during a series of biological surveys by GHD Pty Ltd. in the Zuytdorp coast and hinterland, a total of 1,386 trap nights from 28 sites using buckets, Elliott and funnel traps captured only 
 <specimenCount id="72CCFD2B8F154125FE5A99D37318D63F" box="[451,521,1267,1293]" count="1" pageId="23" pageNumber="322" type="generic">
 one 
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 <emphasis id="56BEEAB08F154125FE6A99D37312D63E" box="[499,515,1267,1292]" italics="true" pageId="23" pageNumber="322">L</emphasis>
 .
 </taxonomicName>
 </specimenCount>
 <emphasis id="56BEEAB08F154125FD8999D37361D63E" box="[528,624,1267,1292]" italics="true" pageId="23" pageNumber="322">inornata</emphasis>
 (G. Gaikhorst, pers. comm.).
 </paragraph>
 <paragraph id="647536A28F154125FF5E983772AED51F" blockId="23.[151,1437,151,1581]" pageId="23" pageNumber="322">
 <taxonomicName id="A3CA4D218F154125FF5E9837706DD602" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[199,380,1303,1329]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F154125FF5E9837706DD602" box="[199,380,1303,1329]" italics="true" pageId="23" pageNumber="322">Liopholis parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F154125FE1E983770F5D603" bold="true" box="[391,484,1303,1329]" pageId="23" pageNumber="322">
 <taxonomicNameLabel id="4D8D57CB8F154125FE1E983770F5D603" box="[391,484,1303,1329]" pageId="23" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 is the only reptile (or vertebrate) species that is known exclusively from disjunct populations on the North-West Cape and the Kennedy Range. Six reptile species are endemic to North-West Cape: the geckos 
 <taxonomicName id="A3CA4D218F154125FE8C987F724FD64B" authority="Doughty, Ellis &amp; Oliver" authorityName="Doughty, Ellis &amp; Oliver" authorityYear="2016" box="[277,862,1375,1401]" class="Squamata" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="tuberculatus">
 <emphasis id="56BEEAB08F154125FE8C987F7358D64B" box="[277,585,1375,1401]" italics="true" pageId="23" pageNumber="322">Crenadactylus tuberculatus</emphasis>
 Doughty, Ellis &amp; Oliver
 </taxonomicName>
 , 
 <taxonomicName id="A3CA4D218F154125FCF3987F748DD64B" authority="Doughty, Oliver &amp; Adams" authorityName="Doughty, Oliver &amp; Adams" authorityYear="2008" box="[874,1436,1375,1401]" class="Squamata" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="capensis">
 <emphasis id="56BEEAB08F154125FCF3987F757FD64A" box="[874,1134,1375,1401]" italics="true" pageId="23" pageNumber="322">Diplodactylus capensis</emphasis>
 Doughty, Oliver &amp; Adams
 </taxonomicName>
 and 
 <taxonomicName id="A3CA4D218F154125FF5198A372DED6AF" authority="Kealley, Doughty, Pepper, Keogh, Hillyer &amp; Huey" authorityName="Kealley, Doughty, Pepper, Keogh, Hillyer &amp; Huey" authorityYear="2018" box="[200,975,1411,1437]" class="Squamata" family="Gekkonidae" genus="Gehyra" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="capensis">
 <emphasis id="56BEEAB08F154125FF5198A37094D6AE" box="[200,389,1411,1437]" italics="true" pageId="23" pageNumber="322">Gehyra capensis</emphasis>
 Kealley, Doughty, Pepper, Keogh, Hillyer &amp; Huey
 </taxonomicName>
 , the legless lizard 
 <taxonomicName id="A3CA4D218F154125FB3D98A3705CD6F3" authority="Maryan, Aplin &amp; Adams" authorityName="Maryan, Aplin &amp; Adams" authorityYear="2007" class="Squamata" family="Pygopodidae" genus="Delma" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="tealei">
 <emphasis id="56BEEAB08F154125FB3D98A37424D6AF" box="[1188,1333,1411,1437]" italics="true" pageId="23" pageNumber="322">Delma tealei</emphasis>
 Maryan, Aplin &amp; Adams
 </taxonomicName>
 , the skink 
 <taxonomicName id="A3CA4D218F154125FE5E988773E5D6F3" authority="Kendrick" authorityName="Kendrick" authorityYear="1989" box="[455,756,1447,1473]" class="Squamata" family="Scincidae" genus="Lerista" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="allochira">
 <emphasis id="56BEEAB08F154125FE5E98877394D6F3" box="[455,645,1447,1473]" italics="true" pageId="23" pageNumber="322">Lerista allochira</emphasis>
 Kendrick
 </taxonomicName>
 and the blind snake 
 <taxonomicName id="A3CA4D218F154125FC469887741AD6F3" authority="(Aplin)" baseAuthorityName="Aplin" baseAuthorityYear="1998" box="[991,1291,1447,1473]" class="Squamata" family="Typhlopidae" genus="Anilios" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="splendidus">
 <emphasis id="56BEEAB08F154125FC46988775BFD6F3" box="[991,1198,1447,1473]" italics="true" pageId="23" pageNumber="322">Anilios splendidus</emphasis>
 (Aplin)
 </taxonomicName>
 , with further endemics awaiting description. The only known endemic species on the Kennedy Range is the skink 
 <taxonomicName id="A3CA4D218F154125FAD498EB708ED53B" authority="Kendrick" authorityName="Kendrick" authorityYear="1989" class="Squamata" family="Scincidae" genus="Lerista" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="kennedyensis">
 <emphasis id="56BEEAB08F154125FAD498EB703AD53B" italics="true" pageId="23" pageNumber="322">Lerista kennedyensis</emphasis>
 Kendrick
 </taxonomicName>
 , though 
 <bibRefCitation id="005B4B538F154125FD9B98CF7256D53B" author="Smith, L. A. &amp; Adams, M." box="[514,839,1519,1545]" pageId="23" pageNumber="322" pagination="309 - 357" refId="ref22241" refString="Smith, L. A. &amp; Adams, M. (2007) Revision of the Lerista muelleri species-group (Lacertilia: Scincidae) in Western Australia, with a redescription of L. muelleri (Fischer, 1881) and the description of nine new species. Records of the Western Australian Museum, 23, 309 - 357. https: // doi. org / 10.18195 / issn. 0312 - 3162.23 (4). 2007.309 - 357" type="journal article" year="2007">Smith &amp; Adams (2007: 323)</bibRefCitation>
 mention another morphologically and allozymically distinctive population with similarities to 
 <taxonomicName id="A3CA4D218F154125FDF29B3372AED51F" authority="Smith &amp; Adams." authorityName="Smith &amp; Adams." box="[619,959,1555,1581]" class="Squamata" family="Scincidae" genus="Lerista" kingdom="Animalia" pageId="23" pageNumber="322" phylum="Chordata" rank="species" species="clara">
 <emphasis id="56BEEAB08F154125FDF29B3373EDD51F" box="[619,764,1555,1581]" italics="true" pageId="23" pageNumber="322">Lerista clara</emphasis>
 Smith &amp; Adams.
 </taxonomicName>
 </paragraph>
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 <emphasis id="56BEEAB08F154125FF0E9B5E709CD5AB" bold="true" box="[151,397,1662,1689]" italics="true" pageId="23" pageNumber="322">Liopholis purnululu</emphasis>
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 <paragraph id="647536A28F154125FF0E9BE7704FD5D3" blockId="23.[151,497,1662,1761]" box="[151,350,1735,1761]" pageId="23" pageNumber="322">
 (
 <figureCitation id="FCF12A278F154125FF069BE771F9D5D3" box="[159,232,1735,1761]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="23" pageNumber="322">Figs 6</figureCitation>
 , 
 <figureCitation id="FCF12A278F154125FF6D9BE77012D5D3" box="[244,259,1735,1761]" captionStart="FIGURE 8" captionStartId="20.[151,250,1477,1501]" captionTargetBox="[165,1435,453,1439]" captionTargetId="figure-152@20.[151,1435,453,1454]" captionTargetPageId="20" captionText="FIGURE 8. Lateral and dorsal views of head scalation of A‒B) L. parva sp. nov. holotype WAM R156452, and C‒D) L. purnululu sp. nov. holotype WAM R103112." figureDoi="http://doi.org/10.5281/zenodo.14421238" httpUri="https://zenodo.org/record/14421238/files/figure.png" pageId="23" pageNumber="322">8</figureCitation>
 , 
 <figureCitation id="FCF12A278F154125FE899BE7703FD5D3" box="[272,302,1735,1761]" captionStart="FIGURE 10" captionStartId="25.[151,250,1892,1916]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-17@25.[151,1436,181,1868]" captionTargetPageId="25" captionText="FIGURE 10. Liopholis purnululu sp. nov. A) Preserved holotype WAM R103112 in dorsal, ventral and lateral views, and B) adult paratype WAM R156716 from the Bungle Bungle Range, Purnululu National Park, Western Australia, photographed in life (photos B. Maryan)." figureDoi="http://doi.org/10.5281/zenodo.14421246" httpUri="https://zenodo.org/record/14421246/files/figure.png" pageId="23" pageNumber="322">10</figureCitation>
 , 
 <figureCitation id="FCF12A278F154125FEA39BE77044D5D3" box="[314,341,1735,1761]" captionStart="FIGURE 11" captionStartId="27.[151,250,1859,1883]" captionTargetBox="[152,1436,181,1835]" captionTargetId="figure-17@27.[152,1436,181,1835]" captionTargetPageId="27" captionText="FIGURE 11. A) The habitat of L. purnululu sp. nov., dense Triodia on shallow sandy soils along edge of ephemeral watercourse, Bungle Bungle Range, Purnululu National Park, Western Australia, and B) L. purnululu sp. nov. photographed in situ outside burrow (photos A: B. Maryan, B: N. Gambold)." figureDoi="http://doi.org/10.5281/zenodo.14421248" httpUri="https://zenodo.org/record/14421248/files/figure.png" pageId="23" pageNumber="322">11</figureCitation>
 )
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F154125FF0E9A2F737DD48B" pageId="23" pageNumber="322" type="materials_examined">
 <paragraph id="647536A28F154125FF0E9A2F7451D47F" blockId="23.[151,1436,1807,1977]" pageId="23" pageNumber="322">
 <materialsCitation id="D4A23CFF8F154125FF0E9A2F7451D47F" collectingDate="1989-07-05" collectionCode="WAM" collectorName="N. Gambold" country="Australia" location="Purnululu National Park" pageId="23" pageNumber="322" specimenCode="R103112" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
 <emphasis id="56BEEAB08F154125FF0E9A2F7015D41B" bold="true" box="[151,260,1807,1833]" pageId="23" pageNumber="322">
 <typeStatus id="BB7188008F154125FF0E9A2F7015D41B" box="[151,260,1807,1833]" pageId="23" pageNumber="322" type="holotype">Holotype</typeStatus>
 </emphasis>
 . 
 <collectionCode id="02DBAE678F154125FE8E9A2F704AD41B" box="[279,347,1807,1833]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="23" pageNumber="322" type="Museum">WAM</collectionCode>
 <specimenCode id="346C9ED98F154125FEF19A2F70C0D41B" box="[360,465,1807,1833]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="322">R103112</specimenCode>
 , female, Bungle Bungle Range, 
 <location id="611560798F154125FCC19A2F7562D41B" LSID="urn:lsid:plazi:treatment:EC6387B48F15412EFF0E9B5E73F1D2BB:611560798F154125FCC19A2F7562D41B" box="[856,1139,1807,1833]" country="Australia" name="Purnululu National Park" pageId="23" pageNumber="322" stateProvince="Western Australia">Purnululu National Park</location>
 (-17.43, 128.4), 
 <collectingRegion id="A60EF8408F154125FAD89A2F7010D47F" country="Australia" name="Western Australia" pageId="23" pageNumber="322">Western Australia</collectingRegion>
 , 
 <collectingCountry id="1CDD76328F154125FE939A137065D47F" box="[266,372,1843,1869]" name="Australia" pageId="23" pageNumber="322">Australia</collectingCountry>
 , collected by 
 <collectorName id="C93F53748F154125FD979A13738AD47F" box="[526,667,1843,1869]" pageId="23" pageNumber="322">N. Gambold</collectorName>
 , 
 <date id="107410628F154125FD3F9A12723DD47F" box="[678,812,1842,1869]" pageId="23" pageNumber="322" value="1989-07-05">
 <collectingDate id="0030E98A8F154125FD3F9A12723DD47F" box="[678,812,1842,1869]" pageId="23" pageNumber="322" value="1989-07-05">5 July 1989</collectingDate>
 </date>
 . Fixed in 10% formalin, stored in 70% ethanol.
 </materialsCitation>
 </paragraph>
 <paragraph id="647536A28F154125FF5E9A77737DD48B" blockId="23.[151,1436,1807,1977]" pageId="23" pageNumber="322">
 <materialsCitation id="D4A23CFF8F154125FF5E9A777379D48B" collectionCode="WAM" county="Five" location="Purnululu National Park" municipality="Bungle Bungle Range" pageId="23" pageNumber="322" specimenCode="R103113-114, R156715-716, R156745" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
 <emphasis id="56BEEAB08F154125FF5E9A77702ED443" bold="true" box="[199,319,1879,1905]" pageId="23" pageNumber="322">
 <typeStatus id="BB7188008F154125FF5E9A77702ED443" box="[199,319,1879,1905]" pageId="23" pageNumber="322" type="paratype">Paratypes</typeStatus>
 </emphasis>
 . 
 <collectingCounty id="8D144E2E8F154125FED49A77706ED443" box="[333,383,1879,1905]" pageId="23" pageNumber="322">Five</collectingCounty>
 from 
 <collectingRegion id="A60EF8408F154125FE5B9A77739DD443" box="[450,652,1879,1905]" country="Australia" name="Western Australia" pageId="23" pageNumber="322">Western Australia</collectingRegion>
 . 
 <collectionCode id="02DBAE678F154125FD0F9A7773CBD443" box="[662,730,1879,1905]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="23" pageNumber="322" type="Museum">WAM</collectionCode>
 <specimenCode id="346C9ED98F154125FD789A777290D443" box="[737,897,1879,1905]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="322">R103113–114</specimenCode>
 , male, female, 
 <collectingMunicipality id="8411ACD88F154125FBB09A777432D443" box="[1065,1315,1879,1905]" pageId="23" pageNumber="322">Bungle Bungle Range</collectingMunicipality>
 , 
 <location id="611560798F154125FAB79A777020D4A7" LSID="urn:lsid:plazi:treatment:EC6387B48F15412EFF0E9B5E73F1D2BB:611560798F154125FAB79A777020D4A7" county="Five" municipality="Bungle Bungle Range" name="Purnululu National Park" pageId="23" pageNumber="322" stateProvince="Western Australia">Purnululu National Park</location>
 (-17.43, 128.4); 
 <collectionCode id="02DBAE678F154125FE769A5B7322D4A7" box="[495,563,1915,1941]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="23" pageNumber="322" type="Museum">WAM</collectionCode>
 <specimenCode id="346C9ED98F154125FDA39A5A73CAD4A7" box="[570,731,1914,1941]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="322">R156715–716</specimenCode>
 , 
 <collectionCode id="02DBAE678F154125FD7F9A5B723BD4A7" box="[742,810,1915,1941]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="23" pageNumber="322" type="Museum">WAM</collectionCode>
 <specimenCode id="346C9ED98F154125FCA89A5A728AD4A7" box="[817,923,1914,1941]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="322">R156745</specimenCode>
 , female, female, male, 
 <location id="611560798F154125FB079A5B7489D4A7" LSID="urn:lsid:plazi:treatment:EC6387B48F15412EFF0E9B5E73F1D2BB:611560798F154125FB079A5B7489D4A7" box="[1182,1432,1915,1941]" county="Five" municipality="Bungle Bungle Range" name="Bungle Bungle Range" pageId="23" pageNumber="322" stateProvince="Western Australia">Bungle Bungle Range</location>
 , 
 <location id="611560798F154125FF0E9ABF70B6D48B" LSID="urn:lsid:plazi:treatment:EC6387B48F15412EFF0E9B5E73F1D2BB:611560798F154125FF0E9ABF70B6D48B" box="[151,423,1951,1977]" county="Five" municipality="Bungle Bungle Range" name="Purnululu National Park" pageId="23" pageNumber="322" stateProvince="Western Australia">Purnululu National Park</location>
 (-17.35, 128.62)
 </materialsCitation>
 .
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F1A412AFF5E9DB67438D19B" pageId="24" pageNumber="323" type="diagnosis">
 <paragraph id="647536A28F1A412AFF5E9DB6737AD3CB" blockId="24.[151,1437,150,1977]" pageId="24" pageNumber="323">
 <emphasis id="56BEEAB08F1A412AFF5E9DB6702BD382" bold="true" box="[199,314,150,176]" pageId="24" pageNumber="323">Diagnosis</emphasis>
 . A moderately robust, small-sized 
 <taxonomicName id="A3CA4D218F1A412AFD4B9DB7722DD383" authorityName="Fitzinger" authorityYear="1843" box="[722,828,151,177]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="genus">
 <emphasis id="56BEEAB08F1A412AFD4B9DB7722DD383" box="[722,828,151,177]" italics="true" pageId="24" pageNumber="323">Liopholis</emphasis>
 </taxonomicName>
 (maximum SVL 
 <quantity id="A3329B478F1A412AFB939DB77570D383" box="[1034,1121,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="24" pageNumber="323" unit="mm" value="77.0">77 mm</quantity>
 ), with round pupil, smooth dorsal and caudal scales, relatively short tail, up to 39 midbody scale rows, ≤ 20 fourth toe lamellae and a plain brownish coloration with obscure pattern.
 </paragraph>
 <paragraph id="647536A28F1A412AFF5E9C227438D19B" blockId="24.[151,1437,150,1977]" pageId="24" pageNumber="323">
 <emphasis id="56BEEAB08F1A412AFF5E9C227352D22E" bold="true" box="[199,579,258,285]" pageId="24" pageNumber="323">Comparisons with other species.</emphasis>
 <taxonomicName id="A3CA4D218F1A412AFDD19C237234D22F" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[584,805,259,285]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F1A412AFDD19C237234D22F" box="[584,805,259,285]" italics="true" pageId="24" pageNumber="323">Liopholis purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F1A412AFCB29C237292D22F" bold="true" box="[811,899,259,285]" pageId="24" pageNumber="323">
 <taxonomicNameLabel id="4D8D57CB8F1A412AFCB29C237292D22F" box="[811,899,259,285]" pageId="24" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 differs from nominate 
 <taxonomicName id="A3CA4D218F1A412AFBE69C2375EBD22E" box="[1151,1274,259,284]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F1A412AFBE69C2375EBD22E" box="[1151,1274,259,284]" italics="true" pageId="24" pageNumber="323">L. inornata</emphasis>
 </taxonomicName>
 from southern semiarid to arid 
 <collectingRegion id="A60EF8408F1A412AFED29C077305D273" box="[331,532,295,321]" country="Australia" name="Western Australia" pageId="24" pageNumber="323">Western Australia</collectingRegion>
 , all populations of 
 <emphasis id="56BEEAB08F1A412AFD7F9C077272D273" box="[742,867,295,321]" italics="true" pageId="24" pageNumber="323">
 L. s. 
 <taxonomicName id="A3CA4D218F1A412AFC829C077272D273" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[795,867,295,321]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="slateri">slateri</taxonomicName>
 </emphasis>
 , and 
 <emphasis id="56BEEAB08F1A412AFC079C077532D272" box="[926,1059,295,320]" italics="true" pageId="24" pageNumber="323">L. s. virgata</emphasis>
 in its smaller size (SVL to 
 <quantity id="A3329B478F1A412AFAD69C07748CD273" box="[1359,1437,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="24" pageNumber="323" unit="mm" value="77.0">77 mm</quantity>
 <emphasis id="56BEEAB08F1A412AFF0E9C6B71CED256" box="[151,223,331,356]" italics="true" pageId="24" pageNumber="323">versus</emphasis>
 to 
 <quantity id="A3329B478F1A412AFE9C9C6B7049D257" box="[261,344,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="24" pageNumber="323" unit="mm" value="84.0">84 mm</quantity>
 , 
 <quantity id="A3329B478F1A412AFEFD9C6B70A4D257" box="[356,437,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="9.3" pageId="24" pageNumber="323" unit="mm" value="93.0">93 mm</quantity>
 and 
 <quantity id="A3329B478F1A412AFE749C6B7351D257" box="[493,576,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="24" pageNumber="323" unit="mm" value="94.0">94 mm</quantity>
 , respectively). 
 <taxonomicName id="A3CA4D218F1A412AFD749C6B72DCD257" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[749,973,331,357]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F1A412AFD749C6B72DCD257" box="[749,973,331,357]" italics="true" pageId="24" pageNumber="323">Liopholis purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F1A412AFC4F9C6B7521D257" bold="true" box="[982,1072,331,357]" pageId="24" pageNumber="323">
 <taxonomicNameLabel id="4D8D57CB8F1A412AFC4F9C6B7521D257" box="[982,1072,331,357]" pageId="24" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 further differs from 
 <taxonomicName id="A3CA4D218F1A412AFA849C6B748DD256" box="[1309,1436,331,356]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F1A412AFA849C6B748DD256" box="[1309,1436,331,356]" italics="true" pageId="24" pageNumber="323">L. inornata</emphasis>
 </taxonomicName>
 and 
 <emphasis id="56BEEAB08F1A412AFF5F9C4F7055D2BB" box="[198,324,367,393]" italics="true" pageId="24" pageNumber="323">
 L. s. 
 <taxonomicName id="A3CA4D218F1A412AFF659C4F7055D2BB" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[252,324,367,393]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="slateri">slateri</taxonomicName>
 </emphasis>
 by having lower paravertebral scale and subdigital lamellae counts (
 <tableCitation id="294803198F1A412AFBA69C4F7582D2BB" box="[1087,1171,366,393]" captionStart="TABLE 5" captionStartId="13.[151,239,1685,1709]" captionTargetPageId="13" captionText="TABLE 5. Summary statistics for measurements (mm) and meristic counts for various taxa and populations of the Liopholis inornata species group. N = sample size. NA = not available. Upper row is the range and the lower row is the mean ± SD. See Table 2 for abbreviations of measurements and counts. 1 AMS R.105730. 2 Sample size 4, except where indicated in parentheses." pageId="24" pageNumber="323">Table 5</tableCitation>
 ) and having less dense, dark pigmentation on the flanks and back (
 <figureCitation id="FCF12A278F1A412AFD1B9CB373D9D29F" box="[642,712,403,429]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="24" pageNumber="323">Fig. 6</figureCitation>
 ). 
 <taxonomicName id="A3CA4D218F1A412AFD469CB372D1D29F" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[735,960,403,429]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F1A412AFD469CB372D1D29F" box="[735,960,403,429]" italics="true" pageId="24" pageNumber="323">Liopholis purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F1A412AFC509CB37535D29F" bold="true" box="[969,1060,403,429]" pageId="24" pageNumber="323">
 <taxonomicNameLabel id="4D8D57CB8F1A412AFC509CB37535D29F" box="[969,1060,403,429]" pageId="24" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 further differs from 
 <emphasis id="56BEEAB08F1A412AFA8B9CB3748DD29E" box="[1298,1436,403,428]" italics="true" pageId="24" pageNumber="323">L. s. virgata</emphasis>
 by lacking a black vertebral stripe (
 <figureCitation id="FCF12A278F1A412AFDBC9C97737BD2E3" box="[549,618,439,465]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="24" pageNumber="323">Fig. 6</figureCitation>
 ). Dorsal colour in 
 <taxonomicName id="A3CA4D218F1A412AFCA79C9772ADD2E2" box="[830,956,439,465]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="inornata">
 <emphasis id="56BEEAB08F1A412AFCA79C97725FD2E2" box="[830,846,439,464]" italics="true" pageId="24" pageNumber="323">L</emphasis>
 . 
 <emphasis id="56BEEAB08F1A412AFCC59C9772ADD2E2" box="[860,956,439,464]" italics="true" pageId="24" pageNumber="323">inornata</emphasis>
 </taxonomicName>
 and 
 <taxonomicName id="A3CA4D218F1A412AFC6D9C977542D2E2" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[1012,1107,439,464]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F1A412AFC6D9C977542D2E2" box="[1012,1107,439,464]" italics="true" pageId="24" pageNumber="323">L. parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F1A412AFBC39C9775A5D2E3" bold="true" box="[1114,1204,439,465]" pageId="24" pageNumber="323">
 <taxonomicNameLabel id="4D8D57CB8F1A412AFBC39C9775A5D2E3" box="[1114,1204,439,465]" pageId="24" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 (see above) tends to be yellowish to reddish or orange-brown rather than brownish as in 
 <taxonomicName id="A3CA4D218F1A412AFC0B9CFB750FD2C7" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[914,1054,475,501]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F1A412AFC0B9CFB750FD2C7" box="[914,1054,475,501]" italics="true" pageId="24" pageNumber="323">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F1A412AFBBF9CFB7591D2C7" bold="true" box="[1062,1152,475,501]" pageId="24" pageNumber="323">
 <taxonomicNameLabel id="4D8D57CB8F1A412AFBBF9CFB7591D2C7" box="[1062,1152,475,501]" pageId="24" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 (
 <figureCitation id="FCF12A278F1A412AFB169CFB75C5D2C7" box="[1167,1236,475,501]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="24" pageNumber="323">Fig. 6</figureCitation>
 ). 
 <taxonomicName id="A3CA4D218F1A412AFB739CFB748DD2C6" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[1258,1436,475,501]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="parva" status="sp. nov.">
 <emphasis id="56BEEAB08F1A412AFB739CFB748DD2C6" box="[1258,1436,475,501]" italics="true" pageId="24" pageNumber="323">Liopholis parva</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F1A412AFF0E9CDF71E0D12B" bold="true" box="[151,241,511,537]" pageId="24" pageNumber="323">
 <taxonomicNameLabel id="4D8D57CB8F1A412AFF0E9CDF71E0D12B" box="[151,241,511,537]" pageId="24" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 further differs by having an even smaller size (SVL to 
 <quantity id="A3329B478F1A412AFCC09CDF72B9D12B" box="[857,936,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="24" pageNumber="323" unit="mm" value="66.0">66 mm</quantity>
 <emphasis id="56BEEAB08F1A412AFC379CDF72E7D12A" box="[942,1014,511,536]" italics="true" pageId="24" pageNumber="323">versus</emphasis>
 to 
 <quantity id="A3329B478F1A412AFB839CDF7578D12B" box="[1050,1129,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="24" pageNumber="323" unit="mm" value="77.0">77 mm</quantity>
 ) and proportionally shorter neck and head (the latter caused by a short snout). The geographically closest congener to 
 <taxonomicName id="A3CA4D218F1A412AFB0D9F037431D10F" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[1172,1312,547,573]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F1A412AFB0D9F037431D10F" box="[1172,1312,547,573]" italics="true" pageId="24" pageNumber="323">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F1A412AFAB19F037493D10F" bold="true" box="[1320,1410,547,573]" pageId="24" pageNumber="323">
 <taxonomicNameLabel id="4D8D57CB8F1A412AFAB19F037493D10F" box="[1320,1410,547,573]" pageId="24" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 is 
 <taxonomicName id="A3CA4D218F1A412AFF0E9F6771ECD152" box="[151,253,583,609]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="24" pageNumber="323" phylum="Chordata" rank="species" species="striata">
 <emphasis id="56BEEAB08F1A412AFF0E9F6771B6D152" box="[151,167,583,608]" italics="true" pageId="24" pageNumber="323">L</emphasis>
 . 
 <emphasis id="56BEEAB08F1A412AFF2D9F6771ECD152" box="[180,253,583,608]" italics="true" pageId="24" pageNumber="323">striata</emphasis>
 </taxonomicName>
 which attains a much larger size (SVL to 
 <quantity id="A3329B478F1A412AFD499F677221D153" box="[720,816,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="24" pageNumber="323" unit="mm" value="112.0">112 mm</quantity>
 ), has a predominately reddish-brown (
 <emphasis id="56BEEAB08F1A412AFB449F677434D152" box="[1245,1317,583,608]" italics="true" pageId="24" pageNumber="323">versus</emphasis>
 brownish) colour, fewer subdigital lamellae (16‒19 
 <emphasis id="56BEEAB08F1A412AFDF09F4B73A0D1B6" box="[617,689,619,644]" italics="true" pageId="24" pageNumber="323">versus</emphasis>
 19‒20), higher paravertebral scale counts (55 or more 
 <emphasis id="56BEEAB08F1A412AFABA9F4B747AD1B6" box="[1315,1387,619,644]" italics="true" pageId="24" pageNumber="323">versus</emphasis>
 46‒ 48), higher midbody scale counts (40‒43 
 <emphasis id="56BEEAB08F1A412AFDF09FAF73A0D19A" box="[617,689,655,680]" italics="true" pageId="24" pageNumber="323">versus</emphasis>
 38‒39) and a vertically ovate rather than rounded pupil.
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F1A412AFF5E9F9270A2D48B" pageId="24" pageNumber="323" type="description">
 <paragraph id="647536A28F1A412AFF5E9F9273B1D06F" blockId="24.[151,1437,150,1977]" pageId="24" pageNumber="323">
 <emphasis id="56BEEAB08F1A412AFF5E9F9270B8D1FF" bold="true" box="[199,425,690,717]" pageId="24" pageNumber="323">
 Details of 
 <typeStatus id="BB7188008F1A412AFEDB9F9370B8D1FF" box="[322,425,691,717]" pageId="24" pageNumber="323" type="holotype">holotype</typeStatus>
 </emphasis>
 . (
 <figureCitation id="FCF12A278F1A412AFE599F93733BD1FF" box="[448,554,691,717]" captionStart="FIGURE 10" captionStartId="25.[151,250,1892,1916]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-17@25.[151,1436,181,1868]" captionTargetPageId="25" captionText="FIGURE 10. Liopholis purnululu sp. nov. A) Preserved holotype WAM R103112 in dorsal, ventral and lateral views, and B) adult paratype WAM R156716 from the Bungle Bungle Range, Purnululu National Park, Western Australia, photographed in life (photos B. Maryan)." figureDoi="http://doi.org/10.5281/zenodo.14421246" httpUri="https://zenodo.org/record/14421246/files/figure.png" pageId="24" pageNumber="323">Fig. 10A</figureCitation>
 ). SVL 77.1; TL 90; HD 10.9; HL 18.0; HW 13.8; HNL 29.5; FLL 22.6; HLL 28.1; AGL 36.2; MBS 38; PV 46; SDL 19; nasals widely separated; prefrontals in broad contact; supraciliaries 6/7, fused to second loreal on left side; nuchals 1/1; upper palpebrals 11/11; primary temporal single; enlarged upper secondary temporals 1/1, each with one adjacent smaller temporal bordering the parietal; supralabials 7/7; ear lobules 4/4, uppermost lobule largest in series.
 </paragraph>
 <paragraph id="647536A28F1A412AFF5E9E467065D0DF" blockId="24.[151,1437,150,1977]" pageId="24" pageNumber="323">
 <emphasis id="56BEEAB08F1A412AFF5E9E467040D0B2" bold="true" box="[199,337,870,896]" pageId="24" pageNumber="323">Description</emphasis>
 . See 
 <tableCitation id="294803198F1A412AFE179E4770F1D0B3" box="[398,480,870,897]" captionStart="TABLE 5" captionStartId="13.[151,239,1685,1709]" captionTargetPageId="13" captionText="TABLE 5. Summary statistics for measurements (mm) and meristic counts for various taxa and populations of the Liopholis inornata species group. N = sample size. NA = not available. Upper row is the range and the lower row is the mean ± SD. See Table 2 for abbreviations of measurements and counts. 1 AMS R.105730. 2 Sample size 4, except where indicated in parentheses." pageId="24" pageNumber="323">Table 5</tableCitation>
 for summary statistics of mensural and meristic data and Supplementary 
 <tableCitation id="294803198F1A412AFA8A9E477465D0B3" box="[1299,1396,871,897]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionTargetPageId="2" captionText="TABLE 1. Locality and sample information for all individuals of Liopholis genetically and morphologically examined (prefixes omitted). Under the registration# column: H indicates holotype. Abbreviations are: ABTC Australian Biological Tissue Collection,Adelaide;AMSAustralian Museum, Sydney;ANWCAustralian National Wildlife Collection, Canberra; MAGNT Museum and Art Gallery of the Northern Territory, Darwin; NMV National Museum of Victoria, Melbourne; SAMA South Australian Museum, Adelaide; WAM Western Australian Museum, Perth; NSW New South Wales; NT Northern Territory; SA South Australia; VIC Victoria; WA Western Australia; CP Conservation Park; HS Homestead; NP National Park; NR Nature Reserve. Under the analysis column specimens examined as follows: M morphology, S SNPs." pageId="24" pageNumber="323">Table S1</tableCitation>
 for measurements and counts of individual specimens. 
 <figureCitation id="FCF12A278F1A412AFD449EAB720ED097" box="[733,799,907,933]" captionStart="FIGURE 6" captionStartId="18.[151,250,1769,1793]" captionTargetBox="[151,1436,181,1745]" captionTargetId="figure-23@18.[151,1436,181,1745]" captionTargetPageId="18" captionText="FIGURE 6. Selection of Liopholis inornata species group specimens illustrating variation in colour and pattern. A) L. inornata WAM R166716, Tropicana Mine, Western Australia, B) L. inornata WAM R163368, Neale Junction Nature Reserve, Western Australia, C) L. inornata WAM R78798, 19 km ENE of Toomey Hills, Western Australia, D) L. inornata WAM R65835, 3 km NE of Comet Vale, Western Australia, E) L. parva sp. nov. holotype WAM R156452, Learmonth Air Weapons Range, Western Australia, F) L. purnululu sp. nov. paratype WAM R156716, Purnululu National Park, Western Australia, G) L. s. slateri holotype WAM R29130, Alice Springs, Northern Territory, and H) L. s. virgata SAM R48873, Neales Camp No. 1, South Australia (photos A‒G: B. Maryan, H: C. Kovach)." figureDoi="http://doi.org/10.5281/zenodo.14421232" httpUri="https://zenodo.org/record/14421232/files/figure.png" pageId="24" pageNumber="323">Fig. 6</figureCitation>
 shows WAM R156716 compared to a selection of other 
 <taxonomicName id="A3CA4D218F1A412AFF0E9E8F704BD0FB" box="[151,346,943,969]" pageId="24" pageNumber="323">
 <emphasis id="56BEEAB08F1A412AFF0E9E8F7010D0FB" box="[151,257,943,969]" italics="true" pageId="24" pageNumber="323">Liopholis</emphasis>
 species
 </taxonomicName>
 and 
 <figureCitation id="FCF12A278F1A412AFE0A9E8F70E6D0FB" box="[403,503,943,969]" captionStart="FIGURE 10" captionStartId="25.[151,250,1892,1916]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure-17@25.[151,1436,181,1868]" captionTargetPageId="25" captionText="FIGURE 10. Liopholis purnululu sp. nov. A) Preserved holotype WAM R103112 in dorsal, ventral and lateral views, and B) adult paratype WAM R156716 from the Bungle Bungle Range, Purnululu National Park, Western Australia, photographed in life (photos B. Maryan)." figureDoi="http://doi.org/10.5281/zenodo.14421246" httpUri="https://zenodo.org/record/14421246/files/figure.png" pageId="24" pageNumber="323">Fig. 10B</figureCitation>
 shows the same individual photographed in life. Head and details of scalation are illustrated in 
 <figureCitation id="FCF12A278F1A412AFEB29EF3707FD0DF" box="[299,366,979,1005]" captionStart="FIGURE 8" captionStartId="20.[151,250,1477,1501]" captionTargetBox="[165,1435,453,1439]" captionTargetId="figure-152@20.[151,1435,453,1454]" captionTargetPageId="20" captionText="FIGURE 8. Lateral and dorsal views of head scalation of A‒B) L. parva sp. nov. holotype WAM R156452, and C‒D) L. purnululu sp. nov. holotype WAM R103112." figureDoi="http://doi.org/10.5281/zenodo.14421238" httpUri="https://zenodo.org/record/14421238/files/figure.png" pageId="24" pageNumber="323">Fig. 8</figureCitation>
 .
 </paragraph>
 <paragraph id="647536A28F1A412AFF5E9ED772D1D563" blockId="24.[151,1437,150,1977]" pageId="24" pageNumber="323">Head moderately deep, barely distinct from neck; external ear opening prominent, narrow and much higher than wide; snout very short, rounded in profile and slightly angular in lateral view; body moderately robust, round in cross-section with flat venter and well-developed, thickset pentadactyl limbs; digits moderately long and slender; claws strong with long sharp tip; tail round in cross-section gradually tapering to pointed tip. Nasals widely separated; groove representing nasal-postnasal suture discernible but weak; two loreals, mostly equal or second usually a little higher than wide, fused to first supraciliary on left side (e.g. WAM R103112, WAM R156715); two presuboculars, first higher than second; prefrontals in broad contact; frontal much wider anteriorly, tapering gradually to a rounded point, equal or only slightly wider than interparietal; frontoparietals paired; parietals bordered posteriorly by an enlarged nuchal scale and laterally by two temporal scales; 3‒5 ear lobules, small and obtuse, mostly equal in size or decreasing slightly in size downwards; 6‒8 (mostly 7) supraciliaries, first three larger and decreasing in size to slightly larger last scale; five supraoculars, second much larger and fifth smallest with anterior two in contact with frontal; 9‒12 upper palpebrals; one nuchal on each side; seven supralabials with the fifth and sixth subocular, subocular supralabials separated from granular eyelid scales by a row of small scales; first pair of chin shields in broad contact, second pair typically separated medially by one scale or three scales (e.g. WAM R103114), third pair longitudinally divided, in contact (e.g. WAM R156716) or separated medially by one or two scales; 38‒39 midbody scale rows; 46‒48 paravertebral scales; 19‒20 lamellae under fourth toe, with an apical series of weak or strong calli, outer edges denticulate, palmar and plantar scales small and ovate.</paragraph>
 <paragraph id="647536A28F1A412AFF5E9B7A70A2D48B" blockId="24.[151,1437,150,1977]" pageId="24" pageNumber="323">
 <emphasis id="56BEEAB08F1A412AFF5E9B7A7079D546" bold="true" box="[199,360,1626,1652]" pageId="24" pageNumber="323">Colour in life</emphasis>
 . Colour in life is based on illustrated figures and field observations of live individuals. Dorsal surface pale brown, head slightly paler. Many dorsal scales with outer edge bearing a faint darker brown smudge, forming very indistinct longitudinal lines, more pronounced in one darker specimen (e.g. WAM R156745). Lateral surface uniform pale grey or with some faint indication of cream posterior edging becoming more distinct above hindlimb. Some specimens (e.g. WAM R156715, WAM R156745) have some dark lateral edges of scales forming obscure streaks that are more distinct between ear and forelimb. Tail concolorous with dorsal surface, with cream posterior edge on many scales forming an obscure transverse alignment of narrow bands. Upper surface of limbs pale brown with no discernible markings. Posterior supralabials and sometimes infralabials with diffuse darker brown smudge on anterior edge of fourth to seventh supralabial. Ventral surface creamy white. Claws with dark culmen on upper surface.
 </paragraph>
 </subSubSection>
 <caption id="30B5662A8F1B412BFF0E9A44709FD4F7" ID-DOI="http://doi.org/10.5281/zenodo.14421246" ID-Zenodo-Dep="14421246" httpUri="https://zenodo.org/record/14421246/files/figure.png" pageId="25" pageNumber="324" startId="25.[151,250,1892,1916]" targetBox="[151,1436,181,1868]" targetPageId="25" targetType="figure">
 <paragraph id="647536A28F1B412BFF0E9A44709FD4F7" blockId="25.[151,1436,1892,1989]" pageId="25" pageNumber="324">
 <emphasis id="56BEEAB08F1B412BFF0E9A447030D44E" bold="true" box="[151,289,1892,1916]" pageId="25" pageNumber="324">FIGURE 10.</emphasis>
 <taxonomicName id="A3CA4D218F1B412BFEB19A4470E2D44E" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[296,499,1892,1916]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="25" pageNumber="324" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F1B412BFEB19A4470E2D44E" box="[296,499,1892,1916]" italics="true" pageId="25" pageNumber="324">Liopholis purnululu</emphasis>
 </taxonomicName>
 <taxonomicNameLabel id="4D8D57CB8F1B412BFE639A45735CD44E" box="[506,589,1893,1917]" pageId="25" pageNumber="324" rank="species">
 <emphasis id="56BEEAB08F1B412BFE639A457357D44F" bold="true" box="[506,582,1893,1917]" pageId="25" pageNumber="324">sp. nov</emphasis>
 .
 </taxonomicNameLabel>
 <emphasis id="56BEEAB08F1B412BFDCD9A447377D44E" bold="true" box="[596,614,1892,1916]" pageId="25" pageNumber="324">A</emphasis>
 ) Preserved holotype WAM R103112 in dorsal, ventral and lateral views, and 
 <emphasis id="56BEEAB08F1B412BFA1A9A457485D44F" bold="true" box="[1411,1428,1893,1917]" pageId="25" pageNumber="324">B</emphasis>
 ) adult paratype WAM R156716 from the Bungle Bungle Range, Purnululu National Park, Western Australia, photographed in life (photos B. Maryan).
 </paragraph>
 </caption>
 <subSubSection id="2CD065298F184128FF5E9DB77239D3CB" pageId="26" pageNumber="325" type="etymology">
 <paragraph id="647536A28F184128FF5E9DB77239D3CB" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="325">
 <emphasis id="56BEEAB08F184128FF5E9DB77059D383" bold="true" box="[199,328,151,177]" pageId="26" pageNumber="325">Etymology</emphasis>
 . After the regional name in the Aboriginal Kija language for the sandstone of the Bungle Bungle Range, and with the aim of drawing attention to the biological endemism of this significant landform in the southeast Kimberley (
 <bibRefCitation id="005B4B538F184128FE849DFF70DBD3CB" author="Gambold, N." box="[285,458,223,249]" pageId="26" pageNumber="325" pagination="160 - 192" refId="ref19747" refString="Gambold, N. (1992) Herpetofauna of the Bungle Bungle area. In: Woinarski, J. C. Z. (Ed.) A survey of the wildlife and vegetation of Purnululu (Bungle Bungle) National Park and adjacent area. Research Bulletin No. 6, Department of Conservation and Land Management, Perth, pp. 160 - 192." type="book chapter" year="1992">Gambold 1992</bibRefCitation>
 ). Used as a noun in apposition.
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F184128FF5E9C227303D10F" pageId="26" pageNumber="325" type="distribution">
 <paragraph id="647536A28F184128FF5E9C227384D2BB" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="325">
 <emphasis id="56BEEAB08F184128FF5E9C227048D22E" bold="true" box="[199,345,258,284]" pageId="26" pageNumber="325">Distribution</emphasis>
 . Apparently restricted to the Bungle Bungle Range in the southeast Kimberley, a heavily dissected sandstone plateau, partly bounded by cliffs and cut by numerous gorges, which is surrounded by an extensive sandplain (
 <bibRefCitation id="005B4B538F184128FE8A9C6B70EBD257" author="Hoatson, D. &amp; Blake, D. &amp; Mory, A. &amp; Tyler, I. &amp; Pittavino, M. &amp; Allen, B. &amp; Kamprad, J. &amp; Oswald-Jacobs, I." box="[275,506,331,357]" pageId="26" pageNumber="325" refId="ref20170" refString="Hoatson, D., Blake, D., Mory, A., Tyler, I., Pittavino, M., Allen, B., Kamprad, J. &amp; Oswald-Jacobs, I. (1997) Bungle Bungle Range, Purnululu National Park, East Kimberley, Western Australia: a guide to the rocks, landforms, plants, animals, and human impact. Australian Geological Survey Organisation, Canberra, 82 pp." type="book" year="1997">
 Hoatson 
 <emphasis id="56BEEAB08F184128FEE29C6B70A4D257" box="[379,437,331,357]" italics="true" pageId="26" pageNumber="325">et al.</emphasis>
 1997
 </bibRefCitation>
 ). It is possible that the plateau of the Bungle Bungle Range within the Purnululu National Park comprises its total distribution.
 </paragraph>
 <paragraph id="647536A28F184128FF5E9CB37303D10F" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="325">
 Biological surveys conducted in the Purnululu National Park to date have not recorded any other species of 
 <taxonomicName id="A3CA4D218F184128FF0E9C977010D2E3" authorityName="Fitzinger" authorityYear="1843" box="[151,257,439,465]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="genus">
 <emphasis id="56BEEAB08F184128FF0E9C977010D2E3" box="[151,257,439,465]" italics="true" pageId="26" pageNumber="325">Liopholis</emphasis>
 </taxonomicName>
 (
 <bibRefCitation id="005B4B538F184128FE969C9770ABD2E3" author="Gambold, N." box="[271,442,439,465]" pageId="26" pageNumber="325" pagination="160 - 192" refId="ref19747" refString="Gambold, N. (1992) Herpetofauna of the Bungle Bungle area. In: Woinarski, J. C. Z. (Ed.) A survey of the wildlife and vegetation of Purnululu (Bungle Bungle) National Park and adjacent area. Research Bulletin No. 6, Department of Conservation and Land Management, Perth, pp. 160 - 192." type="book chapter" year="1992">Gambold 1992</bibRefCitation>
 , T. Partridge, pers. comm.). Based on collection records (
 <bibRefCitation id="005B4B538F184128FBAE9C977481D2E3" author="Atlas of Living Australia" box="[1079,1424,439,465]" pageId="26" pageNumber="325" refId="ref18479" refString="Atlas of Living Australia (2024) Available from: Atlas of Living Australia - Open access to Australia's biodiversity data (ala. org. au) (accessed 12 July 2024)" type="book" year="2024">
 Atlas of Living 
 <collectingCountry id="1CDD76328F184128FB7E9C97745FD2E3" box="[1255,1358,439,465]" name="Australia" pageId="26" pageNumber="325">Australia</collectingCountry>
 2024
 </bibRefCitation>
 ), 
 <taxonomicName id="A3CA4D218F184128FF0E9CFB7035D2C7" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[151,292,475,501]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F184128FF0E9CFB7035D2C7" box="[151,292,475,501]" italics="true" pageId="26" pageNumber="325">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F184128FEB49CFB7099D2C7" bold="true" box="[301,392,475,501]" pageId="26" pageNumber="325">
 <taxonomicNameLabel id="4D8D57CB8F184128FEB49CFB7099D2C7" box="[301,392,475,501]" pageId="26" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 is widely allopatric to any other 
 <taxonomicName id="A3CA4D218F184128FC9D9CFB72DED2C7" box="[772,975,475,501]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="species" species="undetermined">
 <emphasis id="56BEEAB08F184128FC9D9CFB727FD2C7" box="[772,878,475,501]" italics="true" pageId="26" pageNumber="325">Liopholis</emphasis>
 species.
 </taxonomicName>
 The geographically nearest congener, 
 <taxonomicName id="A3CA4D218F184128FA1F9CFB71F1D12A" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="species" species="striata">
 <emphasis id="56BEEAB08F184128FA1F9CFB71F1D12A" italics="true" pageId="26" pageNumber="325">L. striata</emphasis>
 </taxonomicName>
 , has been recorded approximately 
 <quantity id="A3329B478F184128FDF19CDF73ACD12B" box="[616,701,511,537]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="325" unit="km" value="300.0">300 km</quantity>
 to the south in the vicinity of Billiluna on the western edge of the Tanami Desert (
 <bibRefCitation id="005B4B538F184128FED09F037314D10F" author="Storr, G. M. &amp; Smith, L. A. &amp; Johnstone, R. E." box="[329,517,547,573]" pageId="26" pageNumber="325" refId="ref22708" refString="Storr, G. M., Smith, L. A. &amp; Johnstone, R. E. (1999) Lizards of Western Australia. I Skinks. Revised Edition. Western Australian Museum, 291 pp." type="book" year="1999">
 Storr 
 <emphasis id="56BEEAB08F184128FE139F0370D3D10F" box="[394,450,547,573]" italics="true" pageId="26" pageNumber="325">et al.</emphasis>
 1999
 </bibRefCitation>
 ).
 </paragraph>
 </subSubSection>
 <subSubSection id="2CD065298F184128FF5E9F667554D48B" pageId="26" pageNumber="325" type="biology_ecology">
 <paragraph id="647536A28F184128FF5E9F6673D3D0FB" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="325">
 <emphasis id="56BEEAB08F184128FF5E9F66734ED152" bold="true" box="[199,607,582,609]" pageId="26" pageNumber="325">Habitat, ecology and conservation</emphasis>
 . 
 <bibRefCitation id="005B4B538F184128FDF79F6772B8D153" author="Menkhorst, K. &amp; Cowie, I." box="[622,937,583,609]" pageId="26" pageNumber="325" pagination="23 - 94" refId="ref21366" refString="Menkhorst, K. &amp; Cowie, I. (1992) Flora of the Bungle Bungle area. In: Woinarski, J. C. Z. (Ed.) A survey of the wildlife and vegetation of Purnululu (Bungle Bungle) National Park and adjacent area. Research Bulletin No. 6, Department of Conservation and Land Management, Perth, pp. 23 - 94. https: // doi. org / 10.18195 / issn. 0313 - 122 x. 61.2000.335 - 360" type="book chapter" year="1992">Menkhorst &amp; Cowie (1992)</bibRefCitation>
 provide a detailed description of the habitat where individuals were either pit-trapped or excavated from burrows on sandy loams of the Elder Uplands of the Bungle Bungle Range plateau (
 <figureCitation id="FCF12A278F184128FDB69FAF7392D19B" box="[559,643,655,681]" captionStart="FIGURE 11" captionStartId="27.[151,250,1859,1883]" captionTargetBox="[152,1436,181,1835]" captionTargetId="figure-17@27.[152,1436,181,1835]" captionTargetPageId="27" captionText="FIGURE 11. A) The habitat of L. purnululu sp. nov., dense Triodia on shallow sandy soils along edge of ephemeral watercourse, Bungle Bungle Range, Purnululu National Park, Western Australia, and B) L. purnululu sp. nov. photographed in situ outside burrow (photos A: B. Maryan, B: N. Gambold)." figureDoi="http://doi.org/10.5281/zenodo.14421248" httpUri="https://zenodo.org/record/14421248/files/figure.png" pageId="26" pageNumber="325">Fig. 11</figureCitation>
 ). Their account describes a low and sparse tree layer, dominated by 
 <taxonomicName id="A3CA4D218F184128FF0E9F937305D1FF" authority="(Fitzgerald)" baseAuthorityName="Fitzgerald" box="[151,532,691,717]" class="Insecta" family="Cerambycidae" genus="Corymbia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="325" phylum="Arthropoda" rank="species" species="cliftoniana">
 <emphasis id="56BEEAB08F184128FF0E9F937096D1FF" box="[151,391,691,717]" italics="true" pageId="26" pageNumber="325">Corymbia cliftoniana</emphasis>
 (Fitzgerald)
 </taxonomicName>
 with occasional 
 <taxonomicName id="A3CA4D218F184128FD489F9372DAD1FF" authority="(Fitzgerald)" baseAuthorityName="Fitzgerald" box="[721,971,691,717]" class="Insecta" family="Cerambycidae" genus="Corymbia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="325" phylum="Arthropoda" rank="species" species="collina">
 <emphasis id="56BEEAB08F184128FD489F937251D1FF" box="[721,832,691,717]" italics="true" pageId="26" pageNumber="325">C. collina</emphasis>
 (Fitzgerald)
 </taxonomicName>
 , and with a canopy height of 4–8 metres. 
 <taxonomicName id="A3CA4D218F184128FF0E9FF77324D1C3" authority="(Mueller)" baseAuthorityName="Mueller" box="[151,565,727,753]" class="Insecta" family="Cerambycidae" genus="Corymbia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="325" phylum="Arthropoda" rank="species" species="dichromophloia">
 <emphasis id="56BEEAB08F184128FF0E9FF770D0D1C3" box="[151,449,727,753]" italics="true" pageId="26" pageNumber="325">Corymbia dichromophloia</emphasis>
 (Mueller)
 </taxonomicName>
 occurs on rocky slopes and 
 <taxonomicName id="A3CA4D218F184128FCE19FF775DBD1C3" authority="Gardner" authorityName="Gardner" box="[888,1226,727,753]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="26" pageNumber="325" phylum="Tracheophyta" rank="species" species="cupularis">
 <emphasis id="56BEEAB08F184128FCE19FF77577D1C3" box="[888,1126,727,753]" italics="true" pageId="26" pageNumber="325">Eucalyptus cupularis</emphasis>
 Gardner
 </taxonomicName>
 is restricted to the sandy ephemeral watercourses. The shrub layer is well developed, particularly along drainage lines and includes 
 <taxonomicName id="A3CA4D218F184128FF0E9E3F70DBD00B" authority="Mueller" authorityName="Mueller" box="[151,458,799,825]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="26" pageNumber="325" phylum="Tracheophyta" rank="species" species="gonocarpa">
 <emphasis id="56BEEAB08F184128FF0E9E3F7076D00A" box="[151,359,799,825]" italics="true" pageId="26" pageNumber="325">Acacia gonocarpa</emphasis>
 Mueller
 </taxonomicName>
 , 
 <taxonomicName id="A3CA4D218F184128FE419E3F73C4D00B" authority="Mueller" authorityName="Mueller" box="[472,725,799,825]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="26" pageNumber="325" phylum="Tracheophyta" rank="species" species="stipuligera">
 <emphasis id="56BEEAB08F184128FE419E3F7363D00B" box="[472,626,799,825]" italics="true" pageId="26" pageNumber="325">A. stipuligera</emphasis>
 Mueller
 </taxonomicName>
 , 
 <taxonomicName id="A3CA4D218F184128FD7B9E3F72A5D00B" authority="Mueller" authorityName="Mueller" box="[738,948,799,825]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="26" pageNumber="325" phylum="Tracheophyta" rank="species" species="tumida">
 <emphasis id="56BEEAB08F184128FD7B9E3F7240D00B" box="[738,849,799,825]" italics="true" pageId="26" pageNumber="325">A. tumida</emphasis>
 Mueller
 </taxonomicName>
 , 
 <taxonomicName id="A3CA4D218F184128FC589E3F75D9D00B" authority="Hook" authorityName="Hook" box="[961,1224,799,825]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="26" pageNumber="325" phylum="Tracheophyta" rank="species" species="lycopodiifolia">
 <emphasis id="56BEEAB08F184128FC589E3F756FD00B" box="[961,1150,799,825]" italics="true" pageId="26" pageNumber="325">A. lycopodiifolia</emphasis>
 Hook
 </taxonomicName>
 , 
 <taxonomicName id="A3CA4D218F184128FB4C9E3F701AD06F" authority="Fitzgerald" authorityName="Fitzgerald" class="Magnoliopsida" family="Proteaceae" genus="Grevillea" kingdom="Plantae" order="Proteales" pageId="26" pageNumber="325" phylum="Tracheophyta" rank="species" species="miniata">
 <emphasis id="56BEEAB08F184128FB4C9E3F748CD00A" box="[1237,1437,799,825]" italics="true" pageId="26" pageNumber="325">Grevillea miniata</emphasis>
 Fitzgerald
 </taxonomicName>
 , 
 <taxonomicName id="A3CA4D218F184128FE8E9E6370F4D06F" authority="Brown" authorityName="Brown" box="[279,485,835,861]" class="Magnoliopsida" family="Proteaceae" genus="Grevillea" kingdom="Plantae" order="Proteales" pageId="26" pageNumber="325" phylum="Tracheophyta" rank="species" species="refracta">
 <emphasis id="56BEEAB08F184128FE8E9E637083D06F" box="[279,402,835,861]" italics="true" pageId="26" pageNumber="325">G. refracta</emphasis>
 Brown
 </taxonomicName>
 and 
 <taxonomicName id="A3CA4D218F184128FD829E637295D06F" authority="Mueller." authorityName="Mueller." box="[539,900,835,861]" class="Insecta" family="Erebidae" genus="Jacksonia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="325" phylum="Arthropoda" rank="species" species="odontoclada">
 <emphasis id="56BEEAB08F184128FD829E63720FD06F" box="[539,798,835,861]" italics="true" pageId="26" pageNumber="325">Jacksonia odontoclada</emphasis>
 Mueller.
 </taxonomicName>
 <taxonomicName id="A3CA4D218F184128FC139E6375B2D06F" authority="Burbidge" authorityName="Burbidge" box="[906,1187,835,861]" class="Insecta" family="Hepialidae" genus="Triodia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="325" phylum="Arthropoda" rank="species" species="spicata">
 <emphasis id="56BEEAB08F184128FC139E637524D06E" box="[906,1077,835,861]" italics="true" pageId="26" pageNumber="325">Triodia spicata</emphasis>
 Burbidge
 </taxonomicName>
 forms a dense ground layer with 
 <taxonomicName id="A3CA4D218F184128FE8B9E47730BD0B3" authority="Brown" authorityName="Brown" box="[274,538,871,897]" class="Insecta" family="Hepialidae" genus="Triodia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="325" phylum="Arthropoda" rank="species" species="microstachya">
 <emphasis id="56BEEAB08F184128FE8B9E4770D4D0B3" box="[274,453,871,897]" italics="true" pageId="26" pageNumber="325">T. microstachya</emphasis>
 Brown
 </taxonomicName>
 forming large hummocks on the sandy banks of ephemeral watercourses. This vegetation community recorded the highest percentage cover of shrubs and hummock grasses, and no introduced plants were detected (
 <bibRefCitation id="005B4B538F184128FE139E8F73A4D0FB" author="Menkhorst, K. &amp; Cowie, I." box="[394,693,943,969]" pageId="26" pageNumber="325" pagination="23 - 94" refId="ref21366" refString="Menkhorst, K. &amp; Cowie, I. (1992) Flora of the Bungle Bungle area. In: Woinarski, J. C. Z. (Ed.) A survey of the wildlife and vegetation of Purnululu (Bungle Bungle) National Park and adjacent area. Research Bulletin No. 6, Department of Conservation and Land Management, Perth, pp. 23 - 94. https: // doi. org / 10.18195 / issn. 0313 - 122 x. 61.2000.335 - 360" type="book chapter" year="1992">Menkhorst &amp; Cowie 1992</bibRefCitation>
 ).
 </paragraph>
 <paragraph id="647536A28F184128FF5E9EF37341D76B" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="325">
 The ecology of this species is largely unknown. The limited field observations of 
 <taxonomicName id="A3CA4D218F184128FBEC9EF37413D0DF" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[1141,1282,979,1005]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F184128FBEC9EF37413D0DF" box="[1141,1282,979,1005]" italics="true" pageId="26" pageNumber="325">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F184128FA959EF37479D0DF" bold="true" box="[1292,1384,979,1005]" pageId="26" pageNumber="325">
 <taxonomicNameLabel id="4D8D57CB8F184128FA959EF37479D0DF" box="[1292,1384,979,1005]" pageId="26" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 in a small area on the Bungle Bungle Range plateau indicate that this species is numerous, with conspicuous multi-entranced burrows positioned at the base of 
 <taxonomicName id="A3CA4D218F184128FD12993B73CCD707" box="[651,733,1051,1077]" class="Insecta" family="Hepialidae" genus="Triodia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="325" phylum="Arthropoda" rank="genus">
 <emphasis id="56BEEAB08F184128FD12993B73CCD707" box="[651,733,1051,1077]" italics="true" pageId="26" pageNumber="325">Triodia</emphasis>
 </taxonomicName>
 clumps on shallow sandy soils (
 <bibRefCitation id="005B4B538F184128FBD4993B75EBD707" author="Gambold, N." box="[1101,1274,1051,1077]" pageId="26" pageNumber="325" pagination="160 - 192" refId="ref19747" refString="Gambold, N. (1992) Herpetofauna of the Bungle Bungle area. In: Woinarski, J. C. Z. (Ed.) A survey of the wildlife and vegetation of Purnululu (Bungle Bungle) National Park and adjacent area. Research Bulletin No. 6, Department of Conservation and Land Management, Perth, pp. 160 - 192." type="book chapter" year="1992">Gambold 1992</bibRefCitation>
 , B. Maryan &amp; R. Browne-Cooper, pers. obs., 
 <figureCitation id="FCF12A278F184128FE6B991F7353D76B" box="[498,578,1087,1113]" captionStart="FIGURE 11" captionStartId="27.[151,250,1859,1883]" captionTargetBox="[152,1436,181,1835]" captionTargetId="figure-17@27.[152,1436,181,1835]" captionTargetPageId="27" captionText="FIGURE 11. A) The habitat of L. purnululu sp. nov., dense Triodia on shallow sandy soils along edge of ephemeral watercourse, Bungle Bungle Range, Purnululu National Park, Western Australia, and B) L. purnululu sp. nov. photographed in situ outside burrow (photos A: B. Maryan, B: N. Gambold)." figureDoi="http://doi.org/10.5281/zenodo.14421248" httpUri="https://zenodo.org/record/14421248/files/figure.png" pageId="26" pageNumber="325">Fig. 11</figureCitation>
 ).
 </paragraph>
 <paragraph id="647536A28F184128FF5E9943743ED6F3" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="325">
 The species has a potentially restricted geographic distribution on the largely inaccessible Bungle Bungle Range plateau (
 <quantity id="A3329B478F184128FF6399A67042D793" box="[250,339,1158,1185]" metricMagnitude="5" metricUnit="m" metricValue="4.5" pageId="26" pageNumber="325" unit="km" value="450.0">450 km</quantity>
 <superScript id="93BF9BEA8F184128FECA99A6704AD7A6" attach="left" box="[339,347,1158,1172]" fontSize="6" pageId="26" pageNumber="325">2</superScript>
 ) that is protected within the Purnululu National Park (
 <quantity id="A3329B478F184128FC4F99A77554D793" box="[982,1093,1159,1185]" metricMagnitude="6" metricUnit="m" metricValue="2.397" pageId="26" pageNumber="325" unit="km" value="2397.0">2,397 km</quantity>
 <superScript id="93BF9BEA8F184128FBDC99A6755CD7A6" attach="left" box="[1093,1101,1158,1172]" fontSize="6" pageId="26" pageNumber="325">2</superScript>
 ). The national park is also a UNESCO World Heritage Site recognised for its universal value criteria of containing areas of natural and geological significance. Despite this level of protection, caution is certainly warranted considering the conservation status of the closely related 
 <taxonomicName id="A3CA4D218F184128FEF799D370F2D63F" baseAuthorityName="Stirling &amp; Zietz" baseAuthorityYear="1893" box="[366,483,1267,1293]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="species" species="kintorei">
 <emphasis id="56BEEAB08F184128FEF799D370F2D63F" box="[366,483,1267,1293]" italics="true" pageId="26" pageNumber="325">L. kintorei</emphasis>
 </taxonomicName>
 and 
 <emphasis id="56BEEAB08F184128FD8599D3738CD63F" box="[540,669,1267,1293]" italics="true" pageId="26" pageNumber="325">
 L. s. 
 <taxonomicName id="A3CA4D218F184128FDCC99D3738CD63F" baseAuthorityName="Storr" baseAuthorityYear="1968" box="[597,669,1267,1293]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="species" species="slateri">slateri</taxonomicName>
 </emphasis>
 in which populations of both species have declined significantly in recent decades (
 <bibRefCitation id="005B4B538F184128FED4983770E9D603" author="McAlpin, S." box="[333,504,1303,1329]" pageId="26" pageNumber="325" refId="ref21151" refString="McAlpin, S. (2001) A recovery plan for the Great Desert Skink (Egernia kintorei) 2001 - 2011. Arid Lands Environment Centre, Alice Springs." type="book" year="2001">McAlpin 2001</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F184128FD9F9837739FD603" author="Pavey, C." box="[518,654,1303,1329]" pageId="26" pageNumber="325" refId="ref21885" refString="Pavey, C. (2004) Recovery plan for Slater's Skink, Egernia slateri, 2005 - 2010. Northern Territory Department of Infrastructure, Planning and Environment, Darwin." type="book" year="2004">Pavey 2004</bibRefCitation>
 , 
 <bibRefCitation id="005B4B538F184128FD02983773C7D603" author="Pavey, C." box="[667,726,1303,1329]" pageId="26" pageNumber="325" pagination="176 - 177" refId="ref21917" refString="Pavey, C. (2007) Slater's Skink Egernia slateri. In: J. Woinarski, C. Pavey, R. Kerrigan, I. Cowie and S. Ward (Eds.), Lost from Our Landscape: Threatened species of the Northern Territory, Northern Territory Government, Darwin, Northern Territory, Australia, pp. 176 - 177." type="book chapter" year="2007">2007</bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F184128FD7D983772A6D603" author="Moore, D. &amp; Kearney, M. R. &amp; Paltridge, R. &amp; McAlpin, S. &amp; Stow, A." box="[740,951,1302,1329]" pageId="26" pageNumber="325" pagination="207 - 216" refId="ref21492" refString="Moore, D., Kearney, M. R., Paltridge, R., McAlpin, S. &amp; Stow, A. (2015) Is fire a threatening process for Liopholis kintorei, a nationally threatened skink? Wildlife Research, 42, 207 - 216. https: // doi. org / 10.1071 / WR 14227" type="journal article" year="2015">
 Moore 
 <emphasis id="56BEEAB08F184128FCAE9837727BD603" box="[823,874,1303,1329]" italics="true" pageId="26" pageNumber="325">et al</emphasis>
 . 2015
 </bibRefCitation>
 ). Furthermore, as highlighted by 
 <bibRefCitation id="005B4B538F184128FAAD983771F5D667" author="Gambold, N." pageId="26" pageNumber="325" pagination="160 - 192" refId="ref19747" refString="Gambold, N. (1992) Herpetofauna of the Bungle Bungle area. In: Woinarski, J. C. Z. (Ed.) A survey of the wildlife and vegetation of Purnululu (Bungle Bungle) National Park and adjacent area. Research Bulletin No. 6, Department of Conservation and Land Management, Perth, pp. 160 - 192." type="book chapter" year="1992">Gambold (1992)</bibRefCitation>
 , optimum fire regimes for this species have not been defined, and further survey effort is required to refine distributional knowledge including the extensive sandplain surrounding the Bungle Bungle Range. Importantly, no feral herbivores or predators, particularly cats have been recorded on the plateau, the latter tending to be uncommon but widespread, and occasionally trapped and removed by rangers around camps (
 <bibRefCitation id="005B4B538F184128FBB598877432D6F3" author="Woinarski, J. &amp; Menkhorst, K. &amp; Gambold, N. &amp; Braithwaite, R." box="[1068,1315,1447,1473]" pageId="26" pageNumber="325" pagination="99 - 117" refId="ref22955" refString="Woinarski, J., Menkhorst, K., Gambold, N. &amp; Braithwaite, R. (1992) Mammals of the Bungle Bungle area. In: Woinarski, J. C. Z. (Ed.) A survey of the wildlife and vegetation of Purnululu (Bungle Bungle) National Park and adjacent area. Research Bulletin No. 6, Department of Conservation and Land Management, Perth, pp. 99 - 117." type="book chapter" year="1992">
 Woinarski 
 <emphasis id="56BEEAB08F184128FB31988775F1D6F3" box="[1192,1248,1447,1473]" italics="true" pageId="26" pageNumber="325">et al.</emphasis>
 1992
 </bibRefCitation>
 ).
 </paragraph>
 <paragraph id="647536A28F184128FF5E98EB7554D48B" blockId="26.[151,1437,151,1977]" pageId="26" pageNumber="325">
 Available knowledge indicates this species is restricted to the Bungle Bungle Range plateau. The extent of rocky terrain affects the amount of suitable habitat for this burrow-digging species that to date has only been found on sandy loams of the Elder Uplands. Based on existing collection records (N = 6, 
 <bibRefCitation id="005B4B538F184128FB969B337477D51F" author="Atlas of Living Australia" box="[1039,1382,1555,1581]" pageId="26" pageNumber="325" refId="ref18479" refString="Atlas of Living Australia (2024) Available from: Atlas of Living Australia - Open access to Australia's biodiversity data (ala. org. au) (accessed 12 July 2024)" type="book" year="2024">
 Atlas of Living 
 <collectingCountry id="1CDD76328F184128FB279B337434D51F" box="[1214,1317,1555,1581]" name="Australia" pageId="26" pageNumber="325">Australia</collectingCountry>
 2024
 </bibRefCitation>
 ) and limited field observations from only two locations, we calculated the extent of occurrence (EOO; minimum convex hull) and area of occupancy (AOO; 2 x 
 <quantity id="A3329B478F184128FDFF9B7B73B0D547" box="[614,673,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="325" unit="km" value="2.0">2 km</quantity>
 grid cells) of 
 <taxonomicName id="A3CA4D218F184128FCDE9B7B72C4D547" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[839,981,1627,1653]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F184128FCDE9B7B72C4D547" box="[839,981,1627,1653]" italics="true" pageId="26" pageNumber="325">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F184128FC479B7B752BD547" bold="true" box="[990,1082,1627,1653]" pageId="26" pageNumber="325">
 <taxonomicNameLabel id="4D8D57CB8F184128FC479B7B752BD547" box="[990,1082,1627,1653]" pageId="26" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 in GeoCat (https;//geocat.kew. org; 
 <bibRefCitation id="005B4B538F184128FF559B5F70AAD5AB" author="Bachman, S. &amp; Moat, J. &amp; Hill, A. W. &amp; de la Torre, J. &amp; Scott, B." box="[204,443,1663,1689]" pageId="26" pageNumber="325" pagination="117 - 126" refId="ref18513" refString="Bachman, S., Moat, J., Hill, A. W., de la Torre, J. &amp; Scott, B. (2011) Supporting Red List threat assessments with GeoCAT: geospatial conservation assessment tool. ZooKeys, 150, 117 - 126. https: // doi. org / 10.3897 / zookeys. 150.2109" type="journal article" year="2011">
 Bachman 
 <emphasis id="56BEEAB08F184128FEA79B5F7066D5AB" box="[318,375,1663,1689]" italics="true" pageId="26" pageNumber="325">et al.</emphasis>
 2011
 </bibRefCitation>
 ). Due to only two occurrence location points, GeoCat cannot create a minimum convex polygon, and hence the EOO, as per IUCN guidelines, must be made identical to the AOO value, which is very small at 
 <quantity id="A3329B478F184128FF289BE771F8D5D3" box="[177,233,1735,1761]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="325" unit="km" value="8.0">8 km</quantity>
 <superScript id="93BF9BEA8F184128FF709BE671E0D5E6" attach="left" box="[233,241,1734,1748]" fontSize="6" pageId="26" pageNumber="325">2</superScript>
 . Targeted biological surveys on the Bungle Bungle Range in suitable habitat for this species would improve the accuracy of these estimates. Current distributional knowledge of 
 <taxonomicName id="A3CA4D218F184128FC2C9BCB7555D437" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[949,1092,1771,1797]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="26" pageNumber="325" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F184128FC2C9BCB7555D437" box="[949,1092,1771,1797]" italics="true" pageId="26" pageNumber="325">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F184128FBD79BCB75BAD437" bold="true" box="[1102,1195,1771,1797]" pageId="26" pageNumber="325">
 <taxonomicNameLabel id="4D8D57CB8F184128FBD79BCB75BAD437" box="[1102,1195,1771,1797]" pageId="26" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 allows for an IUCN listing assessment under criterion B2 of geographic range (AOO &lt;
 <quantity id="A3329B478F184128FC1F9A2F72DCD41B" box="[902,973,1807,1833]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="325" unit="km" value="10.0">10 km</quantity>
 <superScript id="93BF9BEA8F184128FC549A2E72C4D42E" attach="left" box="[973,981,1806,1820]" fontSize="6" pageId="26" pageNumber="325">2</superScript>
 ) and B(a) number of locations (&lt;5) that qualifies this species for Critically Endangered or at the very least Endangered, under IUCN guidelines and criteria (
 <bibRefCitation id="005B4B538F184128FF069A77703CD443" author="IUCN" box="[159,301,1879,1905]" pageId="26" pageNumber="325" refId="ref20318" refString="IUCN (2022) The Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1 (July 2022). Available from: https: // www. iucnredlist. org / resources / redlistguidelines (accessed 12 th July 2024)" type="journal volume" year="2022">IUCN 2022</bibRefCitation>
 ). Ascertaining further conditions under criterion B in relation to population decline are unknown. Further consideration is that without any detailed knowledge of the species’ ecology, life history, distribution and threats to effectively assess the species, it may also be considered as Data Deficient.
 </paragraph>
 </subSubSection>
 <caption id="30B5662A8F194129FF0E9A63736DD491" ID-DOI="http://doi.org/10.5281/zenodo.14421248" ID-Zenodo-Dep="14421248" httpUri="https://zenodo.org/record/14421248/files/figure.png" pageId="27" pageNumber="326" startId="27.[151,250,1859,1883]" targetBox="[152,1436,181,1835]" targetPageId="27" targetType="figure">
 <paragraph id="647536A28F194129FF0E9A63736DD491" blockId="27.[151,1437,1859,1956]" pageId="27" pageNumber="326">
 <emphasis id="56BEEAB08F194129FF0E9A637025D469" bold="true" box="[151,308,1859,1883]" pageId="27" pageNumber="326">FIGURE 11. A</emphasis>
 ) The habitat of 
 <taxonomicName id="A3CA4D218F194129FE489A64735FD469" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[465,590,1859,1883]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="27" pageNumber="326" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F194129FE489A64735FD469" box="[465,590,1859,1883]" italics="true" pageId="27" pageNumber="326">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F194129FDCA9A6473B3D46E" bold="true" box="[595,674,1860,1884]" pageId="27" pageNumber="326">
 <taxonomicNameLabel id="4D8D57CB8F194129FDCA9A6473B3D46E" box="[595,674,1860,1884]" pageId="27" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 , dense 
 <taxonomicName id="A3CA4D218F194129FD759A637226D469" box="[748,823,1859,1883]" class="Insecta" family="Hepialidae" genus="Triodia" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="326" phylum="Arthropoda" rank="genus">
 <emphasis id="56BEEAB08F194129FD759A637226D469" box="[748,823,1859,1883]" italics="true" pageId="27" pageNumber="326">Triodia</emphasis>
 </taxonomicName>
 on shallow sandy soils along edge of ephemeral watercourse, Bungle Bungle Range, Purnululu National Park, Western Australia, and 
 <emphasis id="56BEEAB08F194129FCE39A48729AD4B2" bold="true" box="[890,907,1896,1920]" pageId="27" pageNumber="326">B</emphasis>
 ) 
 <taxonomicName id="A3CA4D218F194129FC039A48750BD44D" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[922,1050,1895,1919]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="27" pageNumber="326" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F194129FC039A48750BD44D" box="[922,1050,1895,1919]" italics="true" pageId="27" pageNumber="326">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F194129FBB89A487563D4B2" bold="true" box="[1057,1138,1896,1920]" pageId="27" pageNumber="326">
 <taxonomicNameLabel id="4D8D57CB8F194129FBB89A487563D4B2" box="[1057,1138,1896,1920]" pageId="27" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 photographed 
 <emphasis id="56BEEAB08F194129FA959A48745DD44D" box="[1292,1356,1896,1919]" italics="true" pageId="27" pageNumber="326">in situ</emphasis>
 outside burrow (photos A: B. Maryan, B: N. Gambold).
 </paragraph>
 </caption>
 <subSubSection id="2CD065298F1E412EFF5E9DB773F1D2BB" pageId="28" pageNumber="327" type="discussion">
 <paragraph id="647536A28F1E412EFF5E9DB773F1D2BB" blockId="28.[151,1436,151,393]" pageId="28" pageNumber="327">
 <emphasis id="56BEEAB08F1E412EFF5E9DB77025D383" bold="true" box="[199,308,151,177]" pageId="28" pageNumber="327">Remarks</emphasis>
 . The description of 
 <taxonomicName id="A3CA4D218F1E412EFD839DB773B6D383" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[538,679,151,177]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="28" pageNumber="327" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F1E412EFD839DB773B6D383" box="[538,679,151,177]" italics="true" pageId="28" pageNumber="327">L. purnululu</emphasis>
 </taxonomicName>
 <emphasis id="56BEEAB08F1E412EFD369DB7721BD383" bold="true" box="[687,778,151,177]" pageId="28" pageNumber="327">
 <taxonomicNameLabel id="4D8D57CB8F1E412EFD369DB7721BD383" box="[687,778,151,177]" pageId="28" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
 </emphasis>
 raises the number of reptile species endemic to Purnululu National Park and the adjacent Osmond Range to three. The other endemics are the gecko, 
 <taxonomicName id="A3CA4D218F1E412EFB2E9D9B748DD3E7" authority="Horner" authorityName="Horner" authorityYear="2005" box="[1207,1436,187,213]" class="Squamata" family="Gekkonidae" genus="Gehyra" kingdom="Animalia" pageId="28" pageNumber="327" phylum="Chordata" rank="species" species="ipsa">
 <emphasis id="56BEEAB08F1E412EFB2E9D9B7455D3E6" box="[1207,1348,187,213]" italics="true" pageId="28" pageNumber="327">Gehyra ipsa</emphasis>
 Horner
 </taxonomicName>
 and the skink, 
 <taxonomicName id="A3CA4D218F1E412EFEA39DFF7378D3CB" authority="Storr" authorityName="Storr" authorityYear="1991" box="[314,617,223,249]" class="Squamata" family="Scincidae" genus="Lerista" kingdom="Animalia" pageId="28" pageNumber="327" phylum="Chordata" rank="species" species="bunglebungle">
 <emphasis id="56BEEAB08F1E412EFEA39DFF7339D3CB" box="[314,552,223,249]" italics="true" pageId="28" pageNumber="327">Lerista bunglebungle</emphasis>
 Storr
 </taxonomicName>
 , which is only known from 
 <specimenCount id="72CCFD2B8F1E412EFC3B9DFF7547D3CB" box="[930,1110,223,249]" count="3" pageId="28" pageNumber="327" type="generic">three specimens</specimenCount>
 (e.g. 
 <bibRefCitation id="005B4B538F1E412EFB0C9DFF740DD3CB" author="Lloyd, R." box="[1173,1308,222,249]" pageId="28" pageNumber="327" pagination="36 - 42" refId="ref20990" refString="Lloyd, R. (2015) Twenty-five years buried: Rediscovery of the Bungle Bungle Robust Slider, Lerista bunglebungle (Lacertilia: Sphenomorphidae). Western Australian Naturalist, 30, 36 - 42." type="journal article" year="2015">Lloyd 2015</bibRefCitation>
 ). Purnululu National Park is also the 
 <typeStatus id="BB7188008F1E412EFE309C2370C8D22F" box="[425,473,259,285]" pageId="28" pageNumber="327">type</typeStatus>
 locality for the skink 
 <taxonomicName id="A3CA4D218F1E412EFD509C2372E9D22F" authority="Horner. All" authorityName="Horner. All" box="[713,1016,259,285]" class="Squamata" family="Scincidae" genus="Ctenotus" kingdom="Animalia" pageId="28" pageNumber="327" phylum="Chordata" rank="species" species="vagus">
 <emphasis id="56BEEAB08F1E412EFD509C237265D22E" box="[713,884,259,285]" italics="true" pageId="28" pageNumber="327">Ctenotus vagus</emphasis>
 Horner. All
 </taxonomicName>
 the above taxa including 
 <taxonomicName id="A3CA4D218F1E412EFA8B9C23748DD22F" authorityName="Aplin &amp; Maryan &amp; Armstrong &amp; Pavey &amp; Donnellan" authorityYear="2024" box="[1298,1436,259,285]" class="Squamata" family="Scincidae" genus="Liopholis" kingdom="Animalia" pageId="28" pageNumber="327" phylum="Chordata" rank="species" species="purnululu" status="sp. nov.">
 <emphasis id="56BEEAB08F1E412EFA8B9C23748DD22F" box="[1298,1436,259,285]" italics="true" pageId="28" pageNumber="327">L. purnululu</emphasis>
 </taxonomicName>
 <taxonomicNameLabel id="4D8D57CB8F1E412EFF0E9C0771E4D273" box="[151,245,295,321]" pageId="28" pageNumber="327" rank="species">
 <emphasis id="56BEEAB08F1E412EFF0E9C0771A0D273" bold="true" box="[151,177,295,321]" pageId="28" pageNumber="327">sp</emphasis>
 . 
 <emphasis id="56BEEAB08F1E412EFF5A9C0771FED273" bold="true" box="[195,239,295,321]" pageId="28" pageNumber="327">nov</emphasis>
 .
 </taxonomicNameLabel>
 were collected during a single biological survey (
 <bibRefCitation id="005B4B538F1E412EFCA39C0772FBD273" author="Gambold, N." box="[826,1002,295,321]" pageId="28" pageNumber="327" pagination="160 - 192" refId="ref19747" refString="Gambold, N. (1992) Herpetofauna of the Bungle Bungle area. In: Woinarski, J. C. Z. (Ed.) A survey of the wildlife and vegetation of Purnululu (Bungle Bungle) National Park and adjacent area. Research Bulletin No. 6, Department of Conservation and Land Management, Perth, pp. 160 - 192." type="book chapter" year="1992">Gambold 1992</bibRefCitation>
 ). An emerging pattern in areas of the Kimberley with distinctive underlying geology is that they support several endemic reptile species, particularly geckos (
 <bibRefCitation id="005B4B538F1E412EFF6A9C4F70A9D2BB" author="Laver, R. J. &amp; Nielsen, S. V. &amp; Rosauer, D. F. &amp; Oliver, P. M." box="[243,440,367,393]" pageId="28" pageNumber="327" pagination="62 - 70" refId="ref20796" refString="Laver, R. J., Nielsen, S. V., Rosauer, D. F. &amp; Oliver, P. M. (2017) Trans-biome diversity in Australian grass-specialist lizards (Diplodactylidae: Strophurus). Molecular Phylogenetics and Evolution, 115, 62 - 70. https: // doi. org / 10.1016 / j. ympev. 2017.07.015" type="journal article" year="2017">
 Laver 
 <emphasis id="56BEEAB08F1E412EFEA79C4F707ED2BB" box="[318,367,367,393]" italics="true" pageId="28" pageNumber="327">et al</emphasis>
 . 2017
 </bibRefCitation>
 ; 
 <bibRefCitation id="005B4B538F1E412EFE5A9C4F739DD2BB" author="Oliver, P. M. &amp; Laver, R. J. &amp; Martins Mello, F. D. &amp; Pratt, R. C. &amp; Hunjan, S. &amp; Moritz, C." box="[451,652,367,393]" pageId="28" pageNumber="327" pagination="53 - 66" refId="ref21622" refString="Oliver, P. M., Laver, R. J., Martins Mello, F. D., Pratt, R. C., Hunjan, S. &amp; Moritz, C. (2017) A novel hotspot of vertebrate endemism and an evolutionary refugium in tropical Australia. Diversity and Distributions, 23, 53 - 66. https: // doi. org / 10.1111 / ddi. 12506" type="journal article" year="2017">
 Oliver 
 <emphasis id="56BEEAB08F1E412EFD8B9C4F7352D2BB" box="[530,579,367,393]" italics="true" pageId="28" pageNumber="327">et al</emphasis>
 . 2017
 </bibRefCitation>
 , 
 <bibRefCitation id="005B4B538F1E412EFD0E9C4F73C5D2BB" author="Oliver, P. M. &amp; Prasetya, A. M. &amp; Tedeschi, L. G. &amp; Fenker, J. &amp; Ellis, R. J. &amp; Doughty, P. &amp; Moritz, C." box="[663,724,367,393]" pageId="28" pageNumber="327" refId="ref21699" refString="Oliver, P. M., Prasetya, A. M., Tedeschi, L. G., Fenker, J., Ellis, R. J., Doughty, P. &amp; Moritz, C. (2020) Crypsis and convergence: integrative taxonomic revision of the Gehyra australis group (Squamata: Gekkonidae) from northern Australia. PeerJ. 8, e 7971. https: // doi. org / 10.7717 / peerj. 7971." type="journal volume" year="2020">2020</bibRefCitation>
 ).
 </paragraph>
 </subSubSection>
 </treatment>
 </document>