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<document id="3248FC0D29058C2BFC981882BC8BAFD1" ID-DOI="10.1649/0010-065X(2001)055[0485:NSOPHC]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102835" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630156134743" checkinUser="tatiana" docAuthor="Morón, Miguel-Angel, Rivera-Cervantes, Luis Eugenio &amp; López-Vieyra, Mario" docDate="2001" docId="B25E87CDFFE3FFC2FE96FB6C971CFA26" docLanguage="en" docName="TheColeopBull.55.4.485-500.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docSource="http://dx.doi.org/10.1649/0010-065x(2001)055[0485:nsophc]2.0.co;2" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phytalus) dugesiana Moron, Rivera and Lopez, new species" docType="treatment" docVersion="1" lastPageNumber="497" masterDocId="4E67FFB5FFEAFFCEFF97FFD89305FFA0" masterDocTitle="New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México" masterLastPageNumber="500" masterPageNumber="485" pageNumber="494" updateTime="1730145459161" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="EC7B6153DB41662A0886BF691635D89D">New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México</mods:title>
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<mods:namePart id="6B3C4809CA3939E7934A3CB06ABE1FAF">Morón, Miguel-Angel</mods:namePart>
<mods:affiliation id="AEB4A7FD6A8C538D0F08B98DA201132A">Departamento de Entomología Instituto de Ecología, A. C. (Sector SEP­CONACYT) Apdo. Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
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<mods:namePart id="35D94253DF023A22E6E5BA9F68D649D4">Rivera-Cervantes, Luis Eugenio</mods:namePart>
<mods:affiliation id="4A8185B177FFD637B40D300037ACE3E9">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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<mods:namePart id="8C6BD3961D8164A5B9677CE6FFA24A97">López-Vieyra, Mario</mods:namePart>
<mods:affiliation id="274AF982FE54B150E0C9C28E999D2452">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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<mods:title id="728BDAFCAD79AD9DC7440C647F248B8F">The Coleopterists Bulletin</mods:title>
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<mods:date id="D769B356B1D7A6EA44390F7A8773A17F">2001</mods:date>
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<paragraph id="3A4836DBFFE3FFC7FE96FB6C9724FB69" blockId="9.[257,1057,1204,1252]" box="[257,1057,1204,1225]" pageId="9" pageNumber="494">
<taxonomicName id="FDF74D58FFE3FFC7FE96FB6C9097FB69" authority="Moron, Rivera and Lopez" authorityName="Moron, Rivera and Lopez" box="[257,914,1204,1225]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="dugesiana" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFE3FFC7FE96FB6C9174FB69" box="[257,625,1204,1225]" italics="true" pageId="9" pageNumber="494">Phyllophaga (Phytalus) dugesiana</emphasis>
Morón, Rivera and López
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,
<emphasis id="0883EAC9FFE3FFC7FC08FB6C9724FB69" bold="true" box="[927,1057,1204,1225]" pageId="9" pageNumber="494">
<taxonomicNameLabel id="13B057B2FFE3FFC7FC08FB6C9724FB69" box="[927,1057,1204,1225]" pageId="9" pageNumber="494" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="72ED6550FFE3FFC5FDC0FB17919BFCBE" lastPageId="11" lastPageNumber="496" pageId="9" pageNumber="494" type="description">
<paragraph id="3A4836DBFFE3FFC7FDC0FB1791D0FB44" blockId="9.[257,1057,1204,1252]" box="[599,725,1231,1252]" pageId="9" pageNumber="494">
<figureCitation id="A2CC2A5EFFE3FFC7FDC0FB1791D0FB44" box="[599,725,1231,1252]" captionStart="Figs" captionStartId="10.[272,315,1091,1110]" captionTargetBox="[253,1076,270,1070]" captionTargetId="figure-12@10.[250,1080,265,1070]" captionTargetPageId="10" captionText="Figs. 1822. Phyllophaga dugesiana. 18) head and pronotum, dorsal view; 19) tarsal claw, male; 20) genital capsule, lateral view; 21) paramera, distal view; 22) ventral genital plates, female. Scale lines: Figs. 19, 22 = 0.25 mm; Figs. 18, 2021 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102843" httpUri="https://zenodo.org/record/10102843/files/figure.png" pageId="9" pageNumber="494">Figs. 1822</figureCitation>
</paragraph>
<paragraph id="3A4836DBFFE3FFC5FE9AFB20913CFE41" blockId="9.[246,1080,1272,1652]" lastBlockId="11.[246,1079,270,481]" lastPageId="11" lastPageNumber="496" pageId="9" pageNumber="494">
<emphasis id="0883EAC9FFE3FFC7FE9AFB209262FAAB" bold="true" box="[269,359,1272,1291]" pageId="9" pageNumber="494">
<typeStatus id="E54C8879FFE3FFC7FE9AFB209262FAAB" box="[269,359,1272,1291]" pageId="9" pageNumber="494" type="holotype">Holotype</typeStatus>
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Male. Head shiny reddish brown; pronotum, elytra, pygidium, sternites and legs shiny yellowish testaceous. Clypeus short (
<figureCitation id="A2CC2A5EFFE3FFC7FD2AFAC99001FA84" box="[701,772,1297,1316]" captionStart="Figs" captionStartId="10.[272,315,1091,1110]" captionTargetBox="[253,1076,270,1070]" captionTargetId="figure-12@10.[250,1080,265,1070]" captionTargetPageId="10" captionText="Figs. 1822. Phyllophaga dugesiana. 18) head and pronotum, dorsal view; 19) tarsal claw, male; 20) genital capsule, lateral view; 21) paramera, distal view; 22) ventral genital plates, female. Scale lines: Figs. 19, 22 = 0.25 mm; Figs. 18, 2021 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102843" httpUri="https://zenodo.org/record/10102843/files/figure.png" pageId="9" pageNumber="494">Fig. 18</figureCitation>
), wider than long (3:1), anterior border widely sinuated with margins strongly elevated, disk surface nearly flat, with deep, round punctures, without setae. Fronto­clypeal suture slightly sinuated, finely impressed. Frons wider than long (1.8:1) convex, with round, deep punctures and without setae. Antenna 9 segmented, with 3 segmented club; lamellae longer than the preceeding four segments combined (1.8:1), segment 5
<emphasis id="0883EAC9FFE3FFC7FD0DFA5F91A4FA3B" bold="true" box="[666,673,1415,1435]" pageId="9" pageNumber="494">°</emphasis>
longer than segment 4
<emphasis id="0883EAC9FFE3FFC7FC14FA5F908FFA3B" bold="true" box="[899,906,1415,1435]" pageId="9" pageNumber="494">°</emphasis>
, segment 6
<emphasis id="0883EAC9FFE3FFC7FC6CFA5F9707FA3B" bold="true" box="[1019,1026,1415,1435]" pageId="9" pageNumber="494">°</emphasis>
very short, all without anterior projections. Frons 2.8
<emphasis id="0883EAC9FFE3FFC7FD28FA7891D4FA13" bold="true" box="[703,721,1440,1459]" pageId="9" pageNumber="494">×</emphasis>
wider than each eye dorsal diameter. Canthus short and narrowed, with 5 setae. Labrum widely bilobed, deeply sinuated, with slender setae at the borders. Mentum concave, with lateral bulks and slender setae, anterior border widely sinuated. Pronotum wider than long (1.7:1) and 2.2
<emphasis id="0883EAC9FFE3FFC7FC2EFA3090CEFA5B" bold="true" box="[953,971,1512,1531]" pageId="9" pageNumber="494">×</emphasis>
wider than frons. Pronotal disk glabrous, slightly convex, but with shallow depressions toward the posterior angles and lateral margins, irregularly punctate, round punctures separated by less of width of puncture to 13 punctures; anterior margin without setae; lateral borders widely rounded, marginal bead (
<figureCitation id="A2CC2A5EFFE3FFC7FDB0F991916EF9FC" box="[551,619,1609,1628]" captionStart="Figs" captionStartId="10.[272,315,1091,1110]" captionTargetBox="[253,1076,270,1070]" captionTargetId="figure-12@10.[250,1080,265,1070]" captionTargetPageId="10" captionText="Figs. 1822. Phyllophaga dugesiana. 18) head and pronotum, dorsal view; 19) tarsal claw, male; 20) genital capsule, lateral view; 21) paramera, distal view; 22) ventral genital plates, female. Scale lines: Figs. 19, 22 = 0.25 mm; Figs. 18, 2021 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102843" httpUri="https://zenodo.org/record/10102843/files/figure.png" pageId="9" pageNumber="494">Fig. 18</figureCitation>
) clear and widely crenulate, with scattered, short setae; anterior angles nearly straigth, prominent; posterior angles obtuse, prominent. Scu­ tellum 1.2
<emphasis id="0883EAC9FFE0FFC4FECDFB109269FB7B" bold="true" box="[346,364,1224,1243]" pageId="10" pageNumber="495">×</emphasis>
wider than long, with 30 shallow, small, irregular punctures. Each elytron 3.2
<emphasis id="0883EAC9FFE0FFC4FE81FB38922DFB53" bold="true" box="[278,296,1248,1267]" pageId="10" pageNumber="495">×</emphasis>
longer than wide, finely rugo­punctate, glabrous; epipleural border with the color of disk surface, narrowed, extended along the complete margin, without setae; humeral calla rounded, prominent; apical calla rounded. Metathoracic wings completely developed. Propygidium shiny, with fine, shallow punctures, glabrous. Pygidium slightly uniformly convex, shiny, shallow and finely punctate, glabrous, apical margin rounded, with 12 slender, short setae; basal margin widely interrupted at the middle. Pterosternon with medium size, moderately dense, yellowish vestiture. Visible abdominal sternites 2 to 4 slightly convex, with some minute setae at the midline; sternite 5 sligthly convex with group of medium size, erect setae at the middle; anal plate convex, with scattered shallow punctures and some slender setae. Protibiae shorter than protarsi (1:1.3), with external border nearly tridentate, preapical spur straight, acute, shorter than 2
<emphasis id="0883EAC9FFE0FFC4FC12FA179089FA43" bold="true" box="[901,908,1487,1507]" pageId="10" pageNumber="495">°</emphasis>
protarsomerus (1: 1.6). Mesotibiae with one oblique, strong, setiferous, transverse carina, preceeding by vague carina with 2 setae on external side; lower apical spur straight, narrow, slightly shorter than upper spur (1:1.2). Metatibiae nearly as long as metatarsi (1:1.2), with one oblique, poorly marked setiferous, transverse carina, and 35 setae on the external side; apical spurs articulated with the border, straight and narrowed with acute apex, upper spur shorter than 2
<emphasis id="0883EAC9FFE0FFC4FE3DF98792B4F9D3" bold="true" box="[426,433,1631,1651]" pageId="10" pageNumber="495">°</emphasis>
metatarsomere (1:1.3), and 1.2
<emphasis id="0883EAC9FFE0FFC4FD4FF9B891EFF9D3" bold="true" box="[728,746,1632,1651]" pageId="10" pageNumber="495">×</emphasis>
longer than lower spur. Tarsomeres semicylindrical, elongated, with enlarged apex, some setae around the apex and two lines of setae ventrally. Tarsal claws symmetrical, deep and narrowly cleft, with lower tooth clearly shorter and separated from the upper tooth (
<figureCitation id="A2CC2A5EFFE1FFC5FD74FEE6902FFEF1" box="[739,810,318,337]" captionStart="Figs" captionStartId="10.[272,315,1091,1110]" captionTargetBox="[253,1076,270,1070]" captionTargetId="figure-12@10.[250,1080,265,1070]" captionTargetPageId="10" captionText="Figs. 1822. Phyllophaga dugesiana. 18) head and pronotum, dorsal view; 19) tarsal claw, male; 20) genital capsule, lateral view; 21) paramera, distal view; 22) ventral genital plates, female. Scale lines: Figs. 19, 22 = 0.25 mm; Figs. 18, 2021 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102843" httpUri="https://zenodo.org/record/10102843/files/figure.png" pageId="11" pageNumber="496">Fig. 19</figureCitation>
). Genital capsule elongated, basal piece slightly shorter than paramera (1:1.1), tectum with transverse prominence near distal border, paramera elongate, fused at dorsal base, with apex narrowed, slightly curved downward, trema ojival (
<figureCitation id="A2CC2A5EFFE1FFC5FDA6FE5E91A2FE39" box="[561,679,390,409]" captionStart="Figs" captionStartId="10.[272,315,1091,1110]" captionTargetBox="[253,1076,270,1070]" captionTargetId="figure-12@10.[250,1080,265,1070]" captionTargetPageId="10" captionText="Figs. 1822. Phyllophaga dugesiana. 18) head and pronotum, dorsal view; 19) tarsal claw, male; 20) genital capsule, lateral view; 21) paramera, distal view; 22) ventral genital plates, female. Scale lines: Figs. 19, 22 = 0.25 mm; Figs. 18, 2021 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102843" httpUri="https://zenodo.org/record/10102843/files/figure.png" pageId="11" pageNumber="496">Figs. 2021</figureCitation>
). Aedeagus moderately long, with sclerotized, support structure, without outstanding dorsal or preapical sclerites. Length of genital capsule from apex of parameres to border of basal piece:
<quantity id="FD0F9B3EFFE1FFC5FCAFFE6E9087FE69" box="[824,898,438,457]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="496" unit="mm" value="1.9">1.9 mm</quantity>
. Total body length:
<quantity id="FD0F9B3EFFE1FFC5FF61FE169247FE41" box="[246,322,462,481]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="11" pageNumber="496" unit="mm" value="9.1">9.1 mm</quantity>
. Humeral width:
<quantity id="FD0F9B3EFFE1FFC5FE7FFE169131FE41" box="[488,564,462,481]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="11" pageNumber="496" unit="mm" value="3.3">3.3 mm</quantity>
.
</paragraph>
<caption id="6E886653FFE0FFC4FE87FB9B9734FB26" ID-DOI="http://doi.org/10.5281/zenodo.10102843" ID-Zenodo-Dep="10102843" httpUri="https://zenodo.org/record/10102843/files/figure.png" pageId="10" pageNumber="495" startId="10.[272,315,1091,1110]" targetBox="[253,1076,270,1070]" targetPageId="10">
<paragraph id="3A4836DBFFE0FFC4FE87FB9B9734FB26" blockId="10.[249,1081,1091,1158]" pageId="10" pageNumber="495">
<emphasis id="0883EAC9FFE0FFC4FE87FB9B9280FBF6" bold="true" box="[272,389,1091,1110]" pageId="10" pageNumber="495">Figs. 1822.</emphasis>
<emphasis id="0883EAC9FFE0FFC4FE0AFB9B9185FBF6" box="[413,640,1091,1110]" italics="true" pageId="10" pageNumber="495">
<taxonomicName id="FDF74D58FFE0FFC4FE0AFB9B9179FBF6" box="[413,636,1091,1110]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="495" phylum="Arthropoda" rank="species" species="dugesiana">Phyllophaga dugesiana</taxonomicName>
.
</emphasis>
<emphasis id="0883EAC9FFE0FFC4FD12FB9B91A1FBF6" bold="true" box="[645,676,1091,1110]" pageId="10" pageNumber="495">18)</emphasis>
head and pronotum, dorsal view;
<emphasis id="0883EAC9FFE0FFC4FC75FB9B9704FBF6" bold="true" box="[994,1025,1091,1110]" pageId="10" pageNumber="495">19)</emphasis>
tarsal claw, male;
<emphasis id="0883EAC9FFE0FFC4FEE3FB839296FBCE" bold="true" box="[372,403,1115,1134]" pageId="10" pageNumber="495">20)</emphasis>
genital capsule, lateral view;
<emphasis id="0883EAC9FFE0FFC4FD55FB8391E4FBCE" bold="true" box="[706,737,1115,1134]" pageId="10" pageNumber="495">21)</emphasis>
paramera, distal view;
<emphasis id="0883EAC9FFE0FFC4FC58FB8390EAFBCE" bold="true" box="[975,1007,1115,1134]" pageId="10" pageNumber="495">22)</emphasis>
ventral genital plates, female. Scale lines: Figs. 19, 22
<emphasis id="0883EAC9FFE0FFC4FD29FBAB91D5FB26" bold="true" box="[702,720,1139,1158]" pageId="10" pageNumber="495">=</emphasis>
0.25 mm; Figs. 18, 2021
<emphasis id="0883EAC9FFE0FFC4FC4FFBAB90EFFB26" bold="true" box="[984,1002,1139,1158]" pageId="10" pageNumber="495">=</emphasis>
1 mm.
</paragraph>
</caption>
<paragraph id="3A4836DBFFE1FFC5FE87FE269255FD14" blockId="11.[246,1079,510,1652]" pageId="11" pageNumber="496">
<emphasis id="0883EAC9FFE1FFC5FE87FE269270FDB3" bold="true" box="[272,373,510,531]" pageId="11" pageNumber="496">
<typeStatus id="E54C8879FFE1FFC5FE87FE269275FDB3" box="[272,368,510,531]" pageId="11" pageNumber="496" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Similar to male except as follows: antennal club slightly longer than preceeding four segments (1.2:1); pronotal punctures more deep; anal plate more convex, deep punctate; protibial external border with three acute teeth. Ventral genital plates slightly sclerotized, symmetrical, with apical border rounded without setae; dorsal genital plates rounded, thiny, with some setae at apical border (
<figureCitation id="A2CC2A5EFFE1FFC5FE67FD5C9146FD39" box="[496,579,644,665]" captionStart="Figs" captionStartId="10.[272,315,1091,1110]" captionTargetBox="[253,1076,270,1070]" captionTargetId="figure-12@10.[250,1080,265,1070]" captionTargetPageId="10" captionText="Figs. 1822. Phyllophaga dugesiana. 18) head and pronotum, dorsal view; 19) tarsal claw, male; 20) genital capsule, lateral view; 21) paramera, distal view; 22) ventral genital plates, female. Scale lines: Figs. 19, 22 = 0.25 mm; Figs. 18, 2021 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102843" httpUri="https://zenodo.org/record/10102843/files/figure.png" pageId="11" pageNumber="496">Fig. 22</figureCitation>
). Total body length:
<quantity id="FD0F9B3EFFE1FFC5FCBEFD5C9085FD39" box="[809,896,644,665]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="11" pageNumber="496" unit="mm" value="9.2">9.2 mm</quantity>
. Humeral width:
<quantity id="FD0F9B3EFFE1FFC5FF61FD479249FD14" box="[246,332,671,692]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="496" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
<paragraph id="3A4836DBFFE1FFC5FE87FD61919BFCBE" blockId="11.[246,1079,510,1652]" pageId="11" pageNumber="496">
<emphasis id="0883EAC9FFE1FFC5FE87FD6192F1FD6E" bold="true" box="[272,500,697,718]" pageId="11" pageNumber="496">
<typeStatus id="E54C8879FFE1FFC5FE87FD619273FD6E" box="[272,374,697,718]" pageId="11" pageNumber="496" type="paratype">Paratype</typeStatus>
Variation.
</emphasis>
<typeStatus id="E54C8879FFE1FFC5FE68FD619160FD6E" box="[511,613,697,718]" pageId="11" pageNumber="496" type="paratype">Paratypes</typeStatus>
of both sexes are similar to types except that some female specimens present darkened pronotum and elytra, or males are yellowish brown and females are reddish brown. Total body length:
<quantity id="FD0F9B3EFFE1FFC5FB9FFD37925CFCBE" metricMagnitude="-3" metricUnit="m" metricValue="8.85" metricValueMax="10.5" metricValueMin="7.2" pageId="11" pageNumber="496" unit="mm" value="8.85" valueMax="10.5" valueMin="7.2">7.2 10.5 mm</quantity>
. Humeral width: 3.0
<quantity id="FD0F9B3EFFE1FFC5FDD3FCD1919FFCBE" box="[580,666,777,798]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="496" unit="mm" value="3.9">3.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE1FFC5FE87FCFC9710FAEE" pageId="11" pageNumber="496" type="materials_examined">
<paragraph id="3A4836DBFFE1FFC5FE87FCFC9182FAB9" blockId="11.[246,1079,510,1652]" pageId="11" pageNumber="496">
<emphasis id="0883EAC9FFE1FFC5FE87FCFC9298FC99" bold="true" box="[272,413,804,825]" pageId="11" pageNumber="496">Type Series.</emphasis>
(Described from
<specimenCount id="2CF1FD52FFE1FFC5FDF5FCFC91D4FC99" box="[610,721,804,825]" pageId="11" pageNumber="496" type="male">202 males</specimenCount>
and
<specimenCount id="2CF1FD52FFE1FFC5FC9BFCFC9091FC99" box="[780,916,804,825]" pageId="11" pageNumber="496" type="female">179 females</specimenCount>
).
<materialsCitation id="8A9F3C86FFE1FFC5FC3EFCFC91F0FCF4" country="Mexico" location="Estacion Las Joyas" municipality="Autlan" pageId="11" pageNumber="496" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<typeStatus id="E54C8879FFE1FFC5FC3EFCFC9708FC99" box="[937,1037,804,825]" pageId="11" pageNumber="496" type="holotype">Holotype</typeStatus>
, ♂;
<collectingCountry id="42E0764BFFE1FFC5FF61FCE79264FCF4" box="[246,353,831,852]" name="Mexico" pageId="11" pageNumber="496">MEXICO</collectingCountry>
:
<emphasis id="0883EAC9FFE1FFC5FEF8FCE792BAFCF4" box="[367,447,831,852]" italics="true" pageId="11" pageNumber="496">
<collectingRegion id="F833F839FFE1FFC5FEF8FCE792B9FCF4" box="[367,444,831,852]" country="Mexico" name="Jalisco" pageId="11" pageNumber="496">Jalisco</collectingRegion>
,
</emphasis>
<collectingMunicipality id="DA2CACA1FFE1FFC5FE5DFCE79111FCF4" box="[458,532,831,852]" pageId="11" pageNumber="496">Autlán</collectingMunicipality>
,
<location id="3F286000FFE1FFC5FDB4FCE791F0FCF4" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FDB4FCE791F0FCF4" box="[547,757,831,852]" country="Mexico" municipality="Autlan" name="Estacion Las Joyas" pageId="11" pageNumber="496" stateProvince="Jalisco">Estación Las Joyas</location>
</materialsCitation>
,
<materialsCitation id="8A9F3C86FFE1FFC5FC94FCE791B0FCCE" collectingDate="1997-06-06" country="Mexico" elevation="1900" location="Lopez" pageId="11" pageNumber="496" specimenCount="1" typeStatus="holotype">
<date id="4E49101BFFE1FFC5FC94FCE79076FCF4" box="[771,883,831,852]" pageId="11" pageNumber="496" value="1997-06-06">
<collectingDate id="5E0DE9F3FFE1FFC5FC94FCE79076FCF4" box="[771,883,831,852]" pageId="11" pageNumber="496" value="1997-06-06">6­VI­1997</collectingDate>
</date>
,
<quantity id="FD0F9B3EFFE1FFC5FC15FCE790DAFCF4" box="[898,991,831,852]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="496" unit="m" value="1900.0">
<elevation id="B1DAD1E8FFE1FFC5FC15FCE790DAFCF4" box="[898,991,831,852]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="496" unit="m" value="1900.0">1,900 m</elevation>
</quantity>
, trampa luz, bosque mesófilo, M.
<location id="3F286000FFE1FFC5FD9EFC81914FFCCE" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FD9EFC81914FFCCE" box="[521,586,857,878]" country="Mexico" name="Lopez" pageId="11" pageNumber="496">López</location>
(
<collectionCode id="5CE6AE1EFFE1FFC5FDCCFC8191ABFCCE" box="[603,686,857,878]" pageId="11" pageNumber="496">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFE1FFC5FD56FC81923FFC29" collectingDate="1997-06-06" country="Mexico" elevation="1900" location="Lopez" pageId="11" pageNumber="496" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<typeStatus id="E54C8879FFE1FFC5FD56FC819019FCCE" box="[705,796,857,878]" pageId="11" pageNumber="496" type="allotype">Allotype</typeStatus>
<specimenCount id="2CF1FD52FFE1FFC5FCB1FC80903FFCCE" box="[806,826,856,878]" pageId="11" pageNumber="496" type="female"></specimenCount>
same data as holotype, except
</materialsCitation>
<materialsCitation id="8A9F3C86FFE1FFC5FED5FCAC9735FC29" box="[322,1072,883,905]" collectingDate="1997-05-06" country="Mexico" elevation="1900" location="Lopez" pageId="11" pageNumber="496" specimenCount="26" specimenCount-female="26" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FED5FCAC92A0FC29" box="[322,421,884,905]" pageId="11" pageNumber="496" value="1997-05-06">
<collectingDate id="5E0DE9F3FFE1FFC5FED5FCAC92A0FC29" box="[322,421,884,905]" pageId="11" pageNumber="496" value="1997-05-06">6­V­1997</collectingDate>
</date>
(
<collectionCode id="5CE6AE1EFFE1FFC5FE23FCAC9102FC29" box="[436,519,884,905]" pageId="11" pageNumber="496">MXAL</collectionCode>
).
<typeStatus id="E54C8879FFE1FFC5FD8EFCAC9186FC29" box="[537,643,884,905]" pageId="11" pageNumber="496" type="paratype">Paratypes</typeStatus>
: same data as holotype, (64 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FC4AFCAC9723FC29" box="[989,1062,883,905]" pageId="11" pageNumber="496" type="female">26 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFE1FFC5FF61FC5791B8FC04" box="[246,701,909,932]" collectingDate="1997-05-06" collectorName="C. Ibarra" country="Mexico" elevation="1900" location="Lopez" pageId="11" pageNumber="496" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
same data except
<collectorName id="9702530DFFE1FFC5FE20FC57911CFC04" box="[439,537,911,932]" pageId="11" pageNumber="496">C. Ibarra</collectorName>
, (4 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FDE2FC5791B6FC03" box="[629,691,909,932]" pageId="11" pageNumber="496" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
; D. Hernández (3 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FC2EFC5790E6FC03" box="[953,995,909,932]" pageId="11" pageNumber="496" type="female">1 ♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FC6BFC57913BFC1E" collectingDate="1997-06-04" country="Mexico" location="Lopez" pageId="11" pageNumber="496" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FC6BFC579228FC1E" pageId="11" pageNumber="496" value="1997-06-04">
<collectingDate id="5E0DE9F3FFE1FFC5FC6BFC579228FC1E" pageId="11" pageNumber="496" value="1997-06-04">4­VI­ 1997</collectingDate>
</date>
, M.
<location id="3F286000FFE1FFC5FEC8FC7192A5FC1E" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FEC8FC7192A5FC1E" box="[351,416,937,958]" country="Mexico" name="Lopez" pageId="11" pageNumber="496">López</location>
(7 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FE60FC719131FC1E" box="[503,564,936,958]" pageId="11" pageNumber="496" type="female">6 ♀♀</specimenCount>
)
</materialsCitation>
; C. Ibarra (2 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FC96FC71903BFC1E" box="[769,830,936,958]" pageId="11" pageNumber="496" type="female">3 ♀♀</specimenCount>
); D. Hernández (2 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FF61FC1C9227FC79" box="[246,290,963,985]" pageId="11" pageNumber="496" type="female">1 ♀</specimenCount>
);
<date id="4E49101BFFE1FFC5FEAAFC1C92ABFC79" box="[317,430,964,985]" pageId="11" pageNumber="496" value="1997-06-05">5­VI­1997</date>
, (2 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FD86FC1C9154FC79" box="[529,593,963,985]" pageId="11" pageNumber="496" type="female">2 ♀♀</specimenCount>
); M. López (2 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FCA2FC1C9064FC79" box="[821,865,963,985]" pageId="11" pageNumber="496" type="female">1 ♀</specimenCount>
); C. Ibarra (1 ♂
<specimenCount id="2CF1FD52FFE1FFC5FBBEFC1C921BFC53" pageId="11" pageNumber="496" type="female">2 ♀♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FEAFFC07917EFC54" box="[312,635,989,1012]" collectingDate="1997-07-03" country="Mexico" location="Garcia" pageId="11" pageNumber="496" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FEAFFC0792B4FC54" box="[312,433,991,1012]" pageId="11" pageNumber="496" value="1997-07-03">
<collectingDate id="5E0DE9F3FFE1FFC5FEAFFC0792B4FC54" box="[312,433,991,1012]" pageId="11" pageNumber="496" value="1997-07-03">3­VII­1997</collectingDate>
</date>
, J.
<location id="3F286000FFE1FFC5FE4DFC079125FC54" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FE4DFC079125FC54" box="[474,544,991,1012]" country="Mexico" name="Garcia" pageId="11" pageNumber="496">García</location>
(
<specimenCount id="2CF1FD52FFE1FFC5FDA5FC079174FC53" box="[562,625,989,1012]" pageId="11" pageNumber="496" type="female">3 ♀♀</specimenCount>
)
</materialsCitation>
; M. López (95 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FCCAFC0790ACFC53" box="[861,937,989,1012]" pageId="11" pageNumber="496" type="female">54 ♀♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FC22FC0792E7FBAE" collectingDate="1997-07-04" country="Mexico" location="Garcia" pageId="11" pageNumber="496" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FC22FC079737FC54" box="[949,1074,991,1012]" pageId="11" pageNumber="496" value="1997-07-04">
<collectingDate id="5E0DE9F3FFE1FFC5FC22FC079737FC54" box="[949,1074,991,1012]" pageId="11" pageNumber="496" value="1997-07-04">4­VII­1997</collectingDate>
</date>
, J.
<location id="3F286000FFE1FFC5FE87FC219253FBAE" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FE87FC219253FBAE" box="[272,342,1017,1038]" country="Mexico" name="Garcia" pageId="11" pageNumber="496">García</location>
(3 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FE39FC2192DDFBAE" box="[430,472,1016,1038]" pageId="11" pageNumber="496" type="female">1 ♀</specimenCount>
)
</materialsCitation>
; M. López (1 ♂
<specimenCount id="2CF1FD52FFE1FFC5FD09FC2191D9FBAE" box="[670,732,1016,1038]" pageId="11" pageNumber="496" type="female">3 ♀♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FD62FC21922EFB89" collectingDate="1997-08-05" country="Mexico" location="Nunez" pageId="11" pageNumber="496" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FD62FC219072FBAE" box="[757,887,1017,1038]" pageId="11" pageNumber="496" value="1997-08-05">
<collectingDate id="5E0DE9F3FFE1FFC5FD62FC219072FBAE" box="[757,887,1017,1038]" pageId="11" pageNumber="496" value="1997-08-05">5­VIII­1997</collectingDate>
</date>
, S.
<location id="3F286000FFE1FFC5FC35FC2190E3FBAE" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FC35FC2190E3FBAE" box="[930,998,1017,1038]" country="Mexico" name="Nunez" pageId="11" pageNumber="496">Núñez</location>
(2 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FF61FBCC9224FB89" box="[246,289,1043,1065]" pageId="11" pageNumber="496" type="female">1 ♀</specimenCount>
)
</materialsCitation>
; A. Vázquez (
<specimenCount id="2CF1FD52FFE1FFC5FE59FBCC910BFB89" box="[462,526,1043,1065]" pageId="11" pageNumber="496" type="female">2 ♀♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FDBFFBCC90C7FB89" box="[552,962,1043,1065]" collectingDate="1997-08-06" country="Mexico" location="Lopez" pageId="11" pageNumber="496" specimenCount="20" specimenCount-female="20" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FDBFFBCC91AFFB89" box="[552,682,1044,1065]" pageId="11" pageNumber="496" value="1997-08-06">
<collectingDate id="5E0DE9F3FFE1FFC5FDBFFBCC91AFFB89" box="[552,682,1044,1065]" pageId="11" pageNumber="496" value="1997-08-06">6­VIII­1997</collectingDate>
</date>
, M.
<location id="3F286000FFE1FFC5FD76FBCC9027FB89" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FD76FBCC9027FB89" box="[737,802,1044,1065]" country="Mexico" name="Lopez" pageId="11" pageNumber="496">López</location>
(1 ♂
<specimenCount id="2CF1FD52FFE1FFC5FCFCFBCC90BDFB89" box="[875,952,1043,1065]" pageId="11" pageNumber="496" type="female">20 ♀♀</specimenCount>
)
</materialsCitation>
; S. Núñez (
<specimenCount id="2CF1FD52FFE1FFC5FF68FBF59216FBE3" box="[255,275,1069,1091]" pageId="11" pageNumber="496" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FEBAFBF791D8FBE4" box="[301,733,1069,1092]" collectingDate="1997-08-07" country="Mexico" location="Lopez" pageId="11" pageNumber="496" specimenCount="12" specimenCount-female="12" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FEBAFBF792B5FBE4" box="[301,432,1071,1092]" pageId="11" pageNumber="496" value="1997-08-07">
<collectingDate id="5E0DE9F3FFE1FFC5FEBAFBF792B5FBE4" box="[301,432,1071,1092]" pageId="11" pageNumber="496" value="1997-08-07">7­VIII­1997</collectingDate>
</date>
, M.
<location id="3F286000FFE1FFC5FE70FBF7912DFBE4" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FE70FBF7912DFBE4" box="[487,552,1071,1092]" country="Mexico" name="Lopez" pageId="11" pageNumber="496">López</location>
(4 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FD11FBF791D6FBE3" box="[646,723,1069,1092]" pageId="11" pageNumber="496" type="female">12 ♀♀</specimenCount>
)
</materialsCitation>
; A. Partida (
<specimenCount id="2CF1FD52FFE1FFC5FCF8FBF59086FBE3" box="[879,899,1069,1091]" pageId="11" pageNumber="496" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FC0AFBF79268FBFE" collectingDate="1997-09-04" country="Mexico" location="Guerra" pageId="11" pageNumber="496" specimenCount="1" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FC0AFBF7970BFBE4" box="[925,1038,1071,1092]" pageId="11" pageNumber="496" value="1997-09-04">
<collectingDate id="5E0DE9F3FFE1FFC5FC0AFBF7970BFBE4" box="[925,1038,1071,1092]" pageId="11" pageNumber="496" value="1997-09-04">4­IX­1997</collectingDate>
</date>
, A.
<location id="3F286000FFE1FFC5FF61FB91923AFBFE" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FF61FB91923AFBFE" box="[246,319,1097,1118]" country="Mexico" name="Guerra" pageId="11" pageNumber="496">Guerra</location>
(♂)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFE1FFC5FEEBFB91918AFBFE" box="[380,655,1096,1118]" collectingDate="1997-09-05" country="Mexico" location="Lopez" pageId="11" pageNumber="496" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FEEBFB9192E8FBFE" box="[380,493,1097,1118]" pageId="11" pageNumber="496" value="1997-09-05">
<collectingDate id="5E0DE9F3FFE1FFC5FEEBFB9192E8FBFE" box="[380,493,1097,1118]" pageId="11" pageNumber="496" value="1997-09-05">5­IX­1997</collectingDate>
</date>
, M.
<location id="3F286000FFE1FFC5FD88FB919165FBFE" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FD88FB919165FBFE" box="[543,608,1097,1118]" country="Mexico" name="Lopez" pageId="11" pageNumber="496">López</location>
(
<specimenCount id="2CF1FD52FFE1FFC5FDE6FB909180FBFE" box="[625,645,1096,1118]" pageId="11" pageNumber="496" type="female"></specimenCount>
)
</materialsCitation>
; A. Guerra (
<specimenCount id="2CF1FD52FFE1FFC5FC8EFB909028FBFE" box="[793,813,1096,1118]" pageId="11" pageNumber="496" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FCD2FB919218FBD9" collectingDate="1997-09-30" country="Mexico" location="Lopez" pageId="11" pageNumber="496" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FCD2FB9190C7FBFE" box="[837,962,1097,1118]" pageId="11" pageNumber="496" value="1997-09-30">
<collectingDate id="5E0DE9F3FFE1FFC5FCD2FB9190C7FBFE" box="[837,962,1097,1118]" pageId="11" pageNumber="496" value="1997-09-30">30­IX­1997</collectingDate>
</date>
, M.
<location id="3F286000FFE1FFC5FC62FB919733FBFE" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FC62FB919733FBFE" box="[1013,1078,1097,1118]" country="Mexico" name="Lopez" pageId="11" pageNumber="496">López</location>
(
<specimenCount id="2CF1FD52FFE1FFC5FF68FBBB9216FBD9" box="[255,275,1123,1145]" pageId="11" pageNumber="496" type="female"></specimenCount>
)
</materialsCitation>
; D. Pelayo (
<specimenCount id="2CF1FD52FFE1FFC5FE3DFBBB92BBFBD9" box="[426,446,1123,1145]" pageId="11" pageNumber="496" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FE40FBBC902BFBD9" box="[471,814,1123,1145]" collectingDate="1985-11-21" country="Mexico" location="Parada" pageId="11" pageNumber="496" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FE40FBBC9150FBD9" box="[471,597,1124,1145]" pageId="11" pageNumber="496" value="1985-11-21">
<collectingDate id="5E0DE9F3FFE1FFC5FE40FBBC9150FBD9" box="[471,597,1124,1145]" pageId="11" pageNumber="496" value="1985-11-21">21­XI­1985</collectingDate>
</date>
, G.
<location id="3F286000FFE1FFC5FD10FBBC91D4FBD9" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FD10FBBC91D4FBD9" box="[647,721,1124,1145]" country="Mexico" name="Parada" pageId="11" pageNumber="496">Parada</location>
, (
<specimenCount id="2CF1FD52FFE1FFC5FD70FBBC9023FBD9" box="[743,806,1123,1145]" pageId="11" pageNumber="496" type="female">3 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFE1FFC5FCA5FBBC9219FB34" collectingDate="1986-07-13" country="Mexico" location="Rivera" pageId="11" pageNumber="496" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FCA5FBBC90B9FBD9" box="[818,956,1124,1145]" pageId="11" pageNumber="496" value="1986-07-13">
<collectingDate id="5E0DE9F3FFE1FFC5FCA5FBBC90B9FBD9" box="[818,956,1124,1145]" pageId="11" pageNumber="496" value="1986-07-13">13­VII­1986</collectingDate>
</date>
, L.
<location id="3F286000FFE1FFC5FC7DFBBC9737FBD9" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FC7DFBBC9737FBD9" box="[1002,1074,1124,1145]" country="Mexico" name="Rivera" pageId="11" pageNumber="496">Rivera</location>
, (
<specimenCount id="2CF1FD52FFE1FFC5FF68FBA59216FB33" box="[255,275,1149,1171]" pageId="11" pageNumber="496" type="female"></specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFE1FFC5FEA3FBA792B7FB34" box="[308,434,1151,1172]" pageId="11" pageNumber="496" value="1986-06-15">15­VI­1986</date>
, (2 ♂♂
<specimenCount id="2CF1FD52FFE1FFC5FD99FBA79149FB33" box="[526,588,1149,1172]" pageId="11" pageNumber="496" type="female">8 ♀♀</specimenCount>
);
<date id="4E49101BFFE1FFC5FDF3FBA791E7FB34" box="[612,738,1151,1172]" pageId="11" pageNumber="496" value="1986-06-18">18­VI­1986</date>
, (♂);
<date id="4E49101BFFE1FFC5FCB3FBA790A7FB34" box="[804,930,1151,1172]" pageId="11" pageNumber="496" value="1986-06-19">19­VI­1986</date>
, (1 ♂
<specimenCount id="2CF1FD52FFE1FFC5FC7EFBA79723FB33" box="[1001,1062,1149,1172]" pageId="11" pageNumber="496" type="female">9 ♀♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FF61FB419208FB69" collectingDate="1986-07-11" collectorName="El Terrero" country="Mexico" elevation="2200" location="Minatitlan" pageId="11" pageNumber="496" specimenCount="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FF61FB419279FB0E" box="[246,380,1177,1198]" pageId="11" pageNumber="496" value="1986-07-11">
<collectingDate id="5E0DE9F3FFE1FFC5FF61FB419279FB0E" box="[246,380,1177,1198]" pageId="11" pageNumber="496" value="1986-07-11">11­VII­1986</collectingDate>
</date>
, (2 ♂♂).
<emphasis id="0883EAC9FFE1FFC5FE7EFB429138FB0F" box="[489,573,1178,1199]" italics="true" pageId="11" pageNumber="496">
<collectingRegion id="F833F839FFE1FFC5FE7EFB42913DFB0F" box="[489,568,1178,1199]" country="Mexico" name="Colima" pageId="11" pageNumber="496">Colima</collectingRegion>
,
</emphasis>
<location id="3F286000FFE1FFC5FDD1FB4191B3FB0E" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FDD1FB4191B3FB0E" box="[582,694,1177,1198]" country="Mexico" name="Minatitlan" pageId="11" pageNumber="496" stateProvince="Colima">Minatitlán</location>
,
<collectorName id="9702530DFFE1FFC5FD55FB419037FB0E" box="[706,818,1177,1198]" pageId="11" pageNumber="496">El Terrero</collectorName>
, bosque mesófilo,
<quantity id="FD0F9B3EFFE1FFC5FC6CFB419208FB69" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="496" unit="m" value="2200.0">
<elevation id="B1DAD1E8FFE1FFC5FC6CFB419208FB69" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="496" unit="m" value="2200.0">2,200 m</elevation>
</quantity>
</materialsCitation>
,
<materialsCitation id="8A9F3C86FFE1FFC5FE8EFB6C918CFB69" box="[281,649,1203,1225]" collectingDate="1998-06-27" country="Mexico" location="Garcia" pageId="11" pageNumber="496" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FE8EFB6C9292FB69" box="[281,407,1204,1225]" pageId="11" pageNumber="496" value="1998-06-27">
<collectingDate id="5E0DE9F3FFE1FFC5FE8EFB6C9292FB69" box="[281,407,1204,1225]" pageId="11" pageNumber="496" value="1998-06-27">27­VI­1998</collectingDate>
</date>
, luz, J.
<location id="3F286000FFE1FFC5FE7CFB6C9134FB69" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FE7CFB6C9134FB69" box="[491,561,1204,1225]" country="Mexico" name="Garcia" pageId="11" pageNumber="496">García</location>
(
<specimenCount id="2CF1FD52FFE1FFC5FDD5FB6C917AFB69" box="[578,639,1203,1225]" pageId="11" pageNumber="496" type="female">3 ♀♀</specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFE1FFC5FD00FB6C901EFB69" box="[663,795,1204,1225]" pageId="11" pageNumber="496" value="1998-07-23">23­VII­1998</date>
(
<specimenCount id="2CF1FD52FFE1FFC5FCBBFB6B9045FB69" box="[812,832,1203,1225]" pageId="11" pageNumber="496" type="female"></specimenCount>
);
<date id="4E49101BFFE1FFC5FCCFFB6C90D9FB69" box="[856,988,1204,1225]" pageId="11" pageNumber="496" value="1998-07-24">24­VII­1998</date>
(
<specimenCount id="2CF1FD52FFE1FFC5FC7AFB6B9704FB69" box="[1005,1025,1203,1225]" pageId="11" pageNumber="496" type="female"></specimenCount>
); M. López (
<specimenCount id="2CF1FD52FFE1FFC5FEDEFB159258FB43" box="[329,349,1229,1251]" pageId="11" pageNumber="496" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFE1FFC5FEE0FB1791B3FB44" box="[375,694,1229,1252]" collectingDate="1998-07-25" country="Mexico" location="Garcia" pageId="11" pageNumber="496" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="4E49101BFFE1FFC5FEE0FB1792F8FB44" box="[375,509,1231,1252]" pageId="11" pageNumber="496" value="1998-07-25">
<collectingDate id="5E0DE9F3FFE1FFC5FEE0FB1792F8FB44" box="[375,509,1231,1252]" pageId="11" pageNumber="496" value="1998-07-25">25­VII­1998</collectingDate>
</date>
, J.
<location id="3F286000FFE1FFC5FDB2FB17916EFB44" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FDB2FB17916EFB44" box="[549,619,1231,1252]" country="Mexico" name="Garcia" pageId="11" pageNumber="496">García</location>
(♂/
<specimenCount id="2CF1FD52FFE1FFC5FD00FB1591AEFB43" box="[663,683,1229,1251]" pageId="11" pageNumber="496" type="female"></specimenCount>
)
</materialsCitation>
.
<typeStatus id="E54C8879FFE1FFC5FD53FB17902FFB44" box="[708,810,1231,1252]" pageId="11" pageNumber="496" type="paratype">Paratypes</typeStatus>
are deposited at AAHC,
<collectionCode id="5CE6AE1EFFE1FFC5FF61FB31924CFB5E" box="[246,329,1257,1278]" pageId="11" pageNumber="496">AMNH</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FECCFB3192A7FB5E" box="[347,418,1257,1278]" pageId="11" pageNumber="496">CASC</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FE22FB3192E8FB5E" box="[437,493,1257,1278]" pageId="11" pageNumber="496">CNC</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FD96FB31914AFB5E" box="[513,591,1257,1278]" pageId="11" pageNumber="496">FMNH</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FDF5FB3191CAFB5E" box="[610,719,1257,1278]" pageId="11" pageNumber="496">IBUNAM</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FD75FB319026FB5E" box="[738,803,1257,1278]" pageId="11" pageNumber="496">IEXA</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FCA2FB319077FB5E" box="[821,882,1257,1278]" pageId="11" pageNumber="496">IMEJ</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FC12FB3190C8FB5E" box="[901,973,1257,1278]" pageId="11" pageNumber="496">GNPC</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FC77FB319736FB5E" box="[992,1075,1257,1278]" pageId="11" pageNumber="496">MNHN</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FF61FADC9243FAB9" box="[246,326,1284,1305]" pageId="11" pageNumber="496">MXAL</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FEC5FADC92A6FAB9" box="[338,419,1284,1305]" pageId="11" pageNumber="496">NHMC</collectionCode>
,
<collectionCode id="5CE6AE1EFFE1FFC5FE27FADC92FEFAB9" box="[432,507,1284,1305]" pageId="11" pageNumber="496">UNSM</collectionCode>
and
<collectionCode id="5CE6AE1EFFE1FFC5FDA4FADC9186FAB9" box="[563,643,1284,1305]" pageId="11" pageNumber="496">ZMHU</collectionCode>
.
</paragraph>
<paragraph id="3A4836DBFFE1FFC5FE87FAC69710FAEE" blockId="11.[246,1079,510,1652]" pageId="11" pageNumber="496">
<materialsCitation id="8A9F3C86FFE1FFC5FE87FAC69714FAEE" country="Mexico" county="Autlan" location="Las Joyas" municipality="Biosphere Reserve Sierra de Manantlan" pageId="11" pageNumber="496" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="0883EAC9FFE1FFC5FE87FAC692B1FA93" bold="true" box="[272,436,1310,1331]" pageId="11" pageNumber="496">
<typeStatus id="E54C8879FFE1FFC5FE87FAC6924CFA93" box="[272,329,1310,1331]" pageId="11" pageNumber="496">Type</typeStatus>
Locality.
</emphasis>
<location id="3F286000FFE1FFC5FE2AFAC7912DFA94" LSID="urn:lsid:plazi:treatment:B25E87CDFFE3FFC2FE96FB6C971CFA26:3F286000FFE1FFC5FE2AFAC7912DFA94" box="[445,552,1311,1332]" country="Mexico" county="Autlan" municipality="Biosphere Reserve Sierra de Manantlan" name="Las Joyas" pageId="11" pageNumber="496" stateProvince="Jalisco">Las Joyas</location>
,
<collectingMunicipality id="DA2CACA1FFE1FFC5FDA2FAC790E7FA94" box="[565,994,1311,1332]" pageId="11" pageNumber="496">Biosphere Reserve Sierra de Manantlán</collectingMunicipality>
,
<collectingCounty id="D3294E57FFE1FFC5FC78FAC79733FA94" box="[1007,1078,1311,1332]" pageId="11" pageNumber="496">Autlán</collectingCounty>
municipality, state of
<collectingRegion id="F833F839FFE1FFC5FE77FAE1912FFAEE" box="[480,554,1337,1358]" country="Mexico" name="Jalisco" pageId="11" pageNumber="496">Jalisco</collectingRegion>
,
<collectingCountry id="42E0764BFFE1FFC5FDA0FAE19182FAEE" box="[567,647,1337,1358]" name="Mexico" pageId="11" pageNumber="496">México</collectingCountry>
(approx. 19
<emphasis id="0883EAC9FFE1FFC5FC9DFAE19017FAEE" bold="true" box="[778,786,1337,1358]" pageId="11" pageNumber="496">°</emphasis>
35
<emphasis id="0883EAC9FFE1FFC5FCBAFAE09030FAEE" bold="true" box="[813,821,1336,1358]" pageId="11" pageNumber="496">'</emphasis>
13
<emphasis id="0883EAC9FFE1FFC5FCD8FAE09052FAEE" bold="true" box="[847,855,1336,1358]" pageId="11" pageNumber="496">&quot;</emphasis>
N; 104
<emphasis id="0883EAC9FFE1FFC5FC36FAE190ACFAEE" bold="true" box="[929,937,1337,1358]" pageId="11" pageNumber="496">°</emphasis>
16
<emphasis id="0883EAC9FFE1FFC5FC54FAE090CEFAEE" bold="true" box="[963,971,1336,1358]" pageId="11" pageNumber="496">'</emphasis>
48
<emphasis id="0883EAC9FFE1FFC5FC72FAE090E8FAEE" bold="true" box="[997,1005,1336,1358]" pageId="11" pageNumber="496">&quot;</emphasis>
W)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE1FFC5FE87FA8C927DF984" pageId="11" pageNumber="496" type="biology_ecology">
<paragraph id="3A4836DBFFE1FFC5FE87FA8C927DF984" blockId="11.[246,1079,510,1652]" pageId="11" pageNumber="496">
<emphasis id="0883EAC9FFE1FFC5FE87FA8C92C0FAC9" bold="true" box="[272,453,1364,1385]" pageId="11" pageNumber="496">Biological Data.</emphasis>
Specimens of
<taxonomicName id="FDF74D58FFE1FFC5FDFEFA8C91F0FAC9" box="[617,757,1364,1385]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="496" phylum="Arthropoda" rank="species" species="dugesiana" status="sp. nov.">
<emphasis id="0883EAC9FFE1FFC5FDFEFA8C91F0FAC9" box="[617,757,1364,1385]" italics="true" pageId="11" pageNumber="496">P. dugesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE1FFC5FD68FA8C9034FAC9" box="[767,817,1364,1385]" pageId="11" pageNumber="496" rank="species">n.sp.</taxonomicNameLabel>
were collected at lights in cloud forest located at
<quantity id="FD0F9B3EFFE1FFC5FD8EFAB79176FA24" box="[537,627,1391,1412]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="496" unit="m" value="1900.0">1,900 m</quantity>
of altitude. Phenology: June (158), July (167), August (45), September (5), November (3). This species represents nearly 50% of the samples of
<taxonomicName id="FDF74D58FFE1FFC5FD99FA7C9191FA19" box="[526,660,1444,1465]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="496" phylum="Arthropoda" rank="genus">
<emphasis id="0883EAC9FFE1FFC5FD99FA7C9191FA19" box="[526,660,1444,1465]" italics="true" pageId="11" pageNumber="496">Phyllophaga</emphasis>
</taxonomicName>
specimens in Las Joyas. Other species flying at the same time and place were
<taxonomicName id="FDF74D58FFE1FFC5FD24FA67923BFA4E" authority="Moron &amp; Rivera" authorityName="Moron &amp; Rivera-Cervantes" authorityYear="1992" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="496" phylum="Arthropoda" rank="species" species="mesophylla" subGenus="Phytalus">
<emphasis id="0883EAC9FFE1FFC5FD24FA6790C1FA74" box="[691,964,1471,1492]" italics="true" pageId="11" pageNumber="496">P. (Phytalus) mesophylla</emphasis>
Morón &amp; Rivera
</taxonomicName>
,
<taxonomicName id="FDF74D58FFE1FFC5FEC3FA02916BFA4F" box="[340,622,1498,1519]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="496" phylum="Arthropoda" rank="species" species="jalisciensis" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFE1FFC5FEC3FA02916BFA4F" box="[340,622,1498,1519]" italics="true" pageId="11" pageNumber="496">P. (Phytalus) jalisciensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE1FFC5FD17FA0191C0FA4E" box="[640,709,1497,1518]" pageId="11" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFE1FFC5FD4CFA029737FA4E" authority="Moron" authorityName="Moron" box="[731,1074,1497,1519]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="496" phylum="Arthropoda" rank="species" species="chamacayoca">
<emphasis id="0883EAC9FFE1FFC5FD4CFA0291F7FA4F" box="[731,754,1498,1519]" italics="true" pageId="11" pageNumber="496">P.</emphasis>
(
<emphasis id="0883EAC9FFE1FFC5FD6DFA029028FA4F" box="[762,813,1498,1519]" italics="true" pageId="11" pageNumber="496">s.str.</emphasis>
)
<emphasis id="0883EAC9FFE1FFC5FCD0FA0290D2FA4F" box="[839,983,1498,1519]" italics="true" pageId="11" pageNumber="496">chamacayoca</emphasis>
Morón
</taxonomicName>
,
<taxonomicName id="FDF74D58FFE1FFC5FF61FA2C92CFF9A9" box="[246,458,1524,1545]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="496" phylum="Arthropoda" rank="species" species="sayloriana" status="sp. nov.">
<emphasis id="0883EAC9FFE1FFC5FF61FA2C9208F9A9" box="[246,269,1524,1545]" italics="true" pageId="11" pageNumber="496">P.</emphasis>
(
<emphasis id="0883EAC9FFE1FFC5FE82FA2C924DF9A9" box="[277,328,1524,1545]" italics="true" pageId="11" pageNumber="496">s.str.</emphasis>
)
<emphasis id="0883EAC9FFE1FFC5FECDFA2C92CFF9A9" box="[346,458,1524,1545]" italics="true" pageId="11" pageNumber="496">sayloriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE1FFC5FE44FA2C9103F9A9" box="[467,518,1524,1545]" pageId="11" pageNumber="496" rank="species">n.sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFE1FFC5FD83FA2C91F7F9A9" box="[532,754,1524,1545]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="496" phylum="Arthropoda" rank="species" species="manantleca" status="sp. nov.">
<emphasis id="0883EAC9FFE1FFC5FD83FA2C912EF9A9" box="[532,555,1524,1545]" italics="true" pageId="11" pageNumber="496">P.</emphasis>
(
<emphasis id="0883EAC9FFE1FFC5FDA4FA2C9162F9A9" box="[563,615,1524,1545]" italics="true" pageId="11" pageNumber="496">s.str.</emphasis>
)
<emphasis id="0883EAC9FFE1FFC5FDEFFA2C91F7F9A9" box="[632,754,1524,1545]" italics="true" pageId="11" pageNumber="496">manantleca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE1FFC5FD6BFA2C9028F9A9" box="[764,813,1524,1545]" pageId="11" pageNumber="496" rank="species">n.sp.</taxonomicNameLabel>
and two other undescribed species.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE1FFC2FE87F9F1972AFA96" lastPageId="12" lastPageNumber="497" pageId="11" pageNumber="496" type="discussion">
<paragraph id="3A4836DBFFE1FFC2FE87F9F1972AFA96" blockId="11.[246,1079,510,1652]" lastBlockId="12.[249,1081,1260,1414]" lastPageId="12" lastPageNumber="497" pageId="11" pageNumber="496">
<emphasis id="0883EAC9FFE1FFC5FE87F9F19278F99E" bold="true" box="[272,381,1577,1598]" pageId="11" pageNumber="496">Remarks.</emphasis>
<taxonomicName id="FDF74D58FFE1FFC5FE10F9F29186F99F" box="[391,643,1578,1599]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="496" phylum="Arthropoda" rank="species" species="dugesiana" status="sp. nov.">
<emphasis id="0883EAC9FFE1FFC5FE10F9F29186F99F" box="[391,643,1578,1599]" italics="true" pageId="11" pageNumber="496">Phyllophaga dugesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE1FFC5FD19F9F191C5F99E" box="[654,704,1577,1598]" pageId="11" pageNumber="496" rank="species">n.sp.</taxonomicNameLabel>
presents the diagnostic characters of the species group
<emphasis id="0883EAC9FFE1FFC5FE7AF99C915FF9F9" box="[493,602,1604,1625]" italics="true" pageId="11" pageNumber="496">bucephala</emphasis>
but are clearly not related with any other species. The very elongated elytra (more than 3 times longer than wide), 9­ segmented antenna with elongated club, bilobed shape of clypeus, pronotum, elytra and pygidium glabrous, punctate pronotum, elytra moderately rugose, and shape of male genital capsule will aid in the recognition of this species.
</paragraph>
</subSubSection>
<caption id="6E886653FFE6FFC2FE87FBAB972AFB16" ID-DOI="http://doi.org/10.5281/zenodo.10102845" ID-Zenodo-Dep="10102845" httpUri="https://zenodo.org/record/10102845/files/figure.png" pageId="12" pageNumber="497" startId="12.[272,315,1139,1158]" targetBox="[249,1084,270,1114]" targetPageId="12">
<paragraph id="3A4836DBFFE6FFC2FE87FBAB972AFB16" blockId="12.[249,1082,1139,1206]" pageId="12" pageNumber="497">
<emphasis id="0883EAC9FFE6FFC2FE87FBAB928FFB26" bold="true" box="[272,394,1139,1158]" pageId="12" pageNumber="497">Figs. 2327.</emphasis>
<emphasis id="0883EAC9FFE6FFC2FE36FBAB9194FB26" box="[417,657,1139,1158]" italics="true" pageId="12" pageNumber="497">
<taxonomicName id="FDF74D58FFE6FFC2FE36FBAB9188FB26" box="[417,653,1139,1158]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="497" phylum="Arthropoda" rank="species" species="jalisciensis">Phyllophaga jalisciensis</taxonomicName>
.
</emphasis>
<emphasis id="0883EAC9FFE6FFC2FD0CFBAB91BFFB26" bold="true" box="[667,698,1139,1158]" pageId="12" pageNumber="497">23)</emphasis>
Head and pronotum, dorsal view;
<emphasis id="0883EAC9FFE6FFC2FB8DFBAB973FFB26" bold="true" box="[1050,1082,1139,1158]" pageId="12" pageNumber="497">24)</emphasis>
tarsal claw, male;
<emphasis id="0883EAC9FFE6FFC2FE33FB5392C6FB3E" bold="true" box="[420,451,1163,1182]" pageId="12" pageNumber="497">25)</emphasis>
paramera, distal view;
<emphasis id="0883EAC9FFE6FFC2FD09FB5391B8FB3E" bold="true" box="[670,701,1163,1182]" pageId="12" pageNumber="497">26)</emphasis>
genital capsule, lateral view;
<emphasis id="0883EAC9FFE6FFC2FC43FB5390F6FB3E" bold="true" box="[980,1011,1163,1182]" pageId="12" pageNumber="497">27)</emphasis>
ventral genital plates, female. Scale lines: Figs. 24, 27
<emphasis id="0883EAC9FFE6FFC2FD29FB7B91D5FB16" bold="true" box="[702,720,1187,1206]" pageId="12" pageNumber="497">=</emphasis>
0.25 mm; Figs. 23, 2526
<emphasis id="0883EAC9FFE6FFC2FC40FB7B90ECFB16" bold="true" box="[983,1001,1187,1206]" pageId="12" pageNumber="497">=</emphasis>
1 mm.
</paragraph>
</caption>
<subSubSection id="72ED6550FFE6FFC2FE84FAE4971CFA26" pageId="12" pageNumber="497" type="etymology">
<paragraph id="3A4836DBFFE6FFC2FE84FAE4971CFA26" blockId="12.[249,1081,1260,1414]" pageId="12" pageNumber="497">
<emphasis id="0883EAC9FFE6FFC2FE84FAE49296FAF1" bold="true" box="[275,403,1340,1361]" pageId="12" pageNumber="497">Etymology.</emphasis>
This new species is dedicated to the memory of the Mexican­ French coleopterist Eugenio D. Dugés (18331895), whose pioneering works with Mexican species of
<taxonomicName id="FDF74D58FFE6FFC2FD9FFAA9917EFA26" box="[520,635,1393,1414]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="497" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
have been recognized in recent years.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="F847A9387C95193D7730D7B0CBCBB3AD" ID-DOI="10.1649/0010-065X(2001)055[0485:NSOPHC]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102835" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630156134743" checkinUser="tatiana" docAuthor="Morón, Miguel-Angel, Rivera-Cervantes, Luis Eugenio &amp; López-Vieyra, Mario" docDate="2001" docId="B25E87CDFFE6FFC0FE97FA7F9124FC79" docLanguage="en" docName="TheColeopBull.55.4.485-500.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docSource="http://dx.doi.org/10.1649/0010-065x(2001)055[0485:nsophc]2.0.co;2" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phytalus) jalisciensis Moron, Rivera and Lopez, new species" docType="treatment" docVersion="1" lastPageNumber="499" masterDocId="4E67FFB5FFEAFFCEFF97FFD89305FFA0" masterDocTitle="New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México" masterLastPageNumber="500" masterPageNumber="485" pageNumber="497" updateTime="1730145459161" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="DA02061643B77B9EF7E3450AAC20FFA4">New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México</mods:title>
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<mods:namePart id="EC755A203266D666DA4A02F7DA21FD4D">Morón, Miguel-Angel</mods:namePart>
<mods:affiliation id="97F99BB795EC3A8FB20689506E0961DA">Departamento de Entomología Instituto de Ecología, A. C. (Sector SEP­CONACYT) Apdo. Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
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<mods:namePart id="85BEB7C4970F490A23D04F7D3C96F1BF">Rivera-Cervantes, Luis Eugenio</mods:namePart>
<mods:affiliation id="3A0E92D2FE20A88DDC849ECD5D958BC8">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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</mods:role>
<mods:namePart id="ED7114BD86CEC299049AC41C52F0FBDD">López-Vieyra, Mario</mods:namePart>
<mods:affiliation id="817339F0E1F290B384DEE7A58FFAC83A">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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<treatment id="B25E87CDFFE6FFC0FE97FA7F9124FC79" LSID="urn:lsid:plazi:treatment:B25E87CDFFE6FFC0FE97FA7F9124FC79" httpUri="http://treatment.plazi.org/id/B25E87CDFFE6FFC0FE97FA7F9124FC79" lastPageId="14" lastPageNumber="499" pageId="12" pageNumber="497">
<subSubSection id="72ED6550FFE6FFC2FE97FA7F972CFA1B" box="[256,1065,1446,1468]" pageId="12" pageNumber="497" type="nomenclature">
<paragraph id="3A4836DBFFE6FFC2FE97FA7F972CFA1B" blockId="12.[256,1065,1446,1494]" box="[256,1065,1446,1468]" pageId="12" pageNumber="497">
<taxonomicName id="FDF74D58FFE6FFC2FE97FA7F909CFA1C" authority="Moron, Rivera and Lopez" authorityName="Moron, Rivera and Lopez" box="[256,921,1447,1468]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="497" phylum="Arthropoda" rank="species" species="jalisciensis" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFE6FFC2FE97FA7F9283FA1C" box="[256,390,1447,1468]" italics="true" pageId="12" pageNumber="497">Phyllophaga</emphasis>
(
<emphasis id="0883EAC9FFE6FFC2FE0FFA7F92F6FA1C" box="[408,499,1447,1468]" italics="true" pageId="12" pageNumber="497">Phytalus</emphasis>
)
<emphasis id="0883EAC9FFE6FFC2FD93FA7F917DFA1C" box="[516,632,1447,1468]" italics="true" pageId="12" pageNumber="497">jalisciensis</emphasis>
Morón, Rivera and López
</taxonomicName>
,
<emphasis id="0883EAC9FFE6FFC2FC31FA7E972CFA1B" bold="true" box="[934,1065,1446,1467]" pageId="12" pageNumber="497">
<taxonomicNameLabel id="13B057B2FFE6FFC2FC31FA7E972CFA1B" box="[934,1065,1446,1467]" pageId="12" pageNumber="497" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE6FFC3FDCDFA1991C5FA74" lastPageId="13" lastPageNumber="498" pageId="12" pageNumber="497" type="description">
<paragraph id="3A4836DBFFE6FFC2FDCDFA1991DDFA76" blockId="12.[256,1065,1446,1494]" box="[602,728,1473,1494]" pageId="12" pageNumber="497">
<figureCitation id="A2CC2A5EFFE6FFC2FDCDFA1991DDFA76" box="[602,728,1473,1494]" captionStart="Figs" captionStartId="12.[272,315,1139,1158]" captionTargetBox="[249,1084,270,1114]" captionTargetId="figure-12@12.[244,1087,265,1118]" captionTargetPageId="12" captionText="Figs. 2327. Phyllophaga jalisciensis. 23) Head and pronotum, dorsal view; 24) tarsal claw, male; 25) paramera, distal view; 26) genital capsule, lateral view; 27) ventral genital plates, female. Scale lines: Figs. 24, 27 = 0.25 mm; Figs. 23, 2526 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102845" httpUri="https://zenodo.org/record/10102845/files/figure.png" pageId="12" pageNumber="497">Figs. 2325</figureCitation>
</paragraph>
<paragraph id="3A4836DBFFE6FFC3FE87FA3091EBFB11" blockId="12.[249,1082,1512,1652]" lastBlockId="13.[246,1079,270,1201]" lastPageId="13" lastPageNumber="498" pageId="12" pageNumber="497">
<emphasis id="0883EAC9FFE6FFC2FE87FA30926FFA5B" bold="true" box="[272,362,1512,1531]" pageId="12" pageNumber="497">
<typeStatus id="E54C8879FFE6FFC2FE87FA30926FFA5B" box="[272,362,1512,1531]" pageId="12" pageNumber="497" type="holotype">Holotype</typeStatus>
</emphasis>
Male. Frons, disk of pronotum and elytra, shiny dark brown; clypeus, lateral borders of pronotum, pygidium, sternites and legs shiny yellowish testaceous. Clypeus short (
<figureCitation id="A2CC2A5EFFE6FFC2FEAFF9C19284F98C" box="[312,385,1561,1580]" captionStart="Figs" captionStartId="12.[272,315,1139,1158]" captionTargetBox="[249,1084,270,1114]" captionTargetId="figure-12@12.[244,1087,265,1118]" captionTargetPageId="12" captionText="Figs. 2327. Phyllophaga jalisciensis. 23) Head and pronotum, dorsal view; 24) tarsal claw, male; 25) paramera, distal view; 26) genital capsule, lateral view; 27) ventral genital plates, female. Scale lines: Figs. 24, 27 = 0.25 mm; Figs. 23, 2526 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102845" httpUri="https://zenodo.org/record/10102845/files/figure.png" pageId="12" pageNumber="497">Fig. 23</figureCitation>
), wider than long (4.2:1), anterior border widely sinuated with margins strongly elevated, disk surface slightly concave, with wide, shallow, round punctures, without setae. Fronto­clypeal suture vaguely sinuated, finely impressed. Frons wider than long (2.5:1) convex, with sparce, round, shallow punctures and scarce, erect, slender setae. Antenna 9 segmented, with 3 segmented club; lamellae longer than the preceeding four segments combined (1.6:1), segment 5
<emphasis id="0883EAC9FFE7FFC3FD1AFEFD9191FE99" bold="true" box="[653,660,293,313]" pageId="13" pageNumber="498">°</emphasis>
as long as segment 4
<emphasis id="0883EAC9FFE7FFC3FCC9FEFD9060FE99" bold="true" box="[862,869,293,313]" pageId="13" pageNumber="498">°</emphasis>
, both without anterior projections; segment 6
<emphasis id="0883EAC9FFE7FFC3FE5BFEE592D6FEF1" bold="true" box="[460,467,317,337]" pageId="13" pageNumber="498">°</emphasis>
short, with acute projection directed forward. Frons 4.3
<emphasis id="0883EAC9FFE7FFC3FC7FFEE690FFFEF1" bold="true" box="[1000,1018,318,337]" pageId="13" pageNumber="498">×</emphasis>
wider than each eye dorsal diameter. Canthus much short and narrowed, with 4 setae. Labrum widely bilobed, deeply sinuated, with slender setae at the borders. Mentum concave, with lateral bulks and slender setae, anterior border wide and deeply sinuated. Pronotum wider than long (2:1) and 2
<emphasis id="0883EAC9FFE7FFC3FE56FE4692D6FE11" bold="true" box="[449,467,414,433]" pageId="13" pageNumber="498">×</emphasis>
wider than frons. Pronotal disk glabrous, slightly convex, irregularly punctate, round punctures separated by width of 12 punctures; anterior margin without setae; lateral borders wide and slightly rounded, marginal bead widely crenulate (
<figureCitation id="A2CC2A5EFFE7FFC3FF6AFE3E924DFE59" box="[253,328,486,505]" captionStart="Figs" captionStartId="12.[272,315,1139,1158]" captionTargetBox="[249,1084,270,1114]" captionTargetId="figure-12@12.[244,1087,265,1118]" captionTargetPageId="12" captionText="Figs. 2327. Phyllophaga jalisciensis. 23) Head and pronotum, dorsal view; 24) tarsal claw, male; 25) paramera, distal view; 26) genital capsule, lateral view; 27) ventral genital plates, female. Scale lines: Figs. 24, 27 = 0.25 mm; Figs. 23, 2526 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102845" httpUri="https://zenodo.org/record/10102845/files/figure.png" pageId="13" pageNumber="498">Fig. 23</figureCitation>
), with scattered, short setae; anterior angles nearly straigth, not prominent, rounded; posterior angles obtuse, not prominent, rounded. Scutellum 1.1
<emphasis id="0883EAC9FFE7FFC3FC22FE2690C2FDB1" bold="true" box="[949,967,510,529]" pageId="13" pageNumber="498">×</emphasis>
wider than long, with 18 shallow, small, irregular punctures. Each elytron 2.8
<emphasis id="0883EAC9FFE7FFC3FCE2FDCE9082FD89" bold="true" box="[885,903,534,553]" pageId="13" pageNumber="498">×</emphasis>
longer than wide, densely rugo­punctate, glabrous; epipleural border with color of disk surface, narrowed, extended along the complete margin, with scattered, short setae; humeral calla rounded, prominent; apical calla rounded. Metathoracic wings completely developed. Propygidium shiny, with fine, sparse punctures and 5 oblique striae, glabrous. Pygidium uniformly convex, shiny, regularly civered with round punctures, glabrous, apical margin rounded, with 13 slender, short setae; basal margin widely interrupted at the middle. Pterosternon with medium size, moderately dense to scarce, yellowish vestiture. Visible abdominal sternites 2 to 4 slightly convex, with short setae at the midline and toward sides; sternite 5 sligthly convex, with medium size, erect setae at middle; anal plate convex, with scattered shallow punctures and some slender, erect setae. Protibiae shorter than protarsi (1:1.2), with external border nearly tridentate, because basal tooth is vaguely indicated; preapical spur straight, acute, much shorter than 2
<emphasis id="0883EAC9FFE7FFC3FD75FCED91ECFCE9" bold="true" box="[738,745,821,841]" pageId="13" pageNumber="498">°</emphasis>
protarsomerus (1:2.8). Mesotibiae with one oblique, strong, setiferous, transverse carina, preceeding by vague carina with 3 setae on external side; lower apical spur straight, narrow, shorter than upper spur (1: 1.4). Metatibiae nearly as long as metatarsi (1:1.1), with one oblique, vague, setiferous, transverse carina, and 5 setae on external side; apical spurs articulated with border, straight and narrowed with rounded apex, upper spur much shorter than 2
<emphasis id="0883EAC9FFE7FFC3FC36FC7590ADFC61" bold="true" box="[929,936,941,961]" pageId="13" pageNumber="498">°</emphasis>
metatarsomere (1:2.3), and 1.2
<emphasis id="0883EAC9FFE7FFC3FE1BFC1E929BFC79" bold="true" box="[396,414,966,985]" pageId="13" pageNumber="498">×</emphasis>
longer than lower spur. Tarsomeres semicylindrical, elongated, with enlarged apex, some setae around the apex and many setae on ventrally slightly flattened side. Tarsal claws symmetrical, deep and narrowly cleft, with lower tooth clearly shorter and separated from the upper tooth (
<figureCitation id="A2CC2A5EFFE7FFC3FDF4FBD691A8FB81" box="[611,685,1038,1057]" captionStart="Figs" captionStartId="12.[272,315,1139,1158]" captionTargetBox="[249,1084,270,1114]" captionTargetId="figure-12@12.[244,1087,265,1118]" captionTargetPageId="12" captionText="Figs. 2327. Phyllophaga jalisciensis. 23) Head and pronotum, dorsal view; 24) tarsal claw, male; 25) paramera, distal view; 26) genital capsule, lateral view; 27) ventral genital plates, female. Scale lines: Figs. 24, 27 = 0.25 mm; Figs. 23, 2526 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102845" httpUri="https://zenodo.org/record/10102845/files/figure.png" pageId="13" pageNumber="498">Fig. 24</figureCitation>
). Genital capsule elongated, basal piece slightly shorter than paramera (1:1.1), tectum with 2 wide, lateral prominences near distal border, paramera elongate, fused at dorsal base, with distal half narrowed, slightly curved downward, apex nearly rounded, trema ojival (
<figureCitation id="A2CC2A5EFFE7FFC3FD25FB8E9020FBC9" box="[690,805,1110,1129]" captionStart="Figs" captionStartId="12.[272,315,1139,1158]" captionTargetBox="[249,1084,270,1114]" captionTargetId="figure-12@12.[244,1087,265,1118]" captionTargetPageId="12" captionText="Figs. 2327. Phyllophaga jalisciensis. 23) Head and pronotum, dorsal view; 24) tarsal claw, male; 25) paramera, distal view; 26) genital capsule, lateral view; 27) ventral genital plates, female. Scale lines: Figs. 24, 27 = 0.25 mm; Figs. 23, 2526 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102845" httpUri="https://zenodo.org/record/10102845/files/figure.png" pageId="13" pageNumber="498">Figs. 2526</figureCitation>
). Aedeagus moderately long, with poorly sclerotized support structure, without outstanding dorsal or preapical sclerites. Length of genital capsule from apex of parameres to border of basal piece:
<quantity id="FD0F9B3EFFE7FFC3FC7EFB5E9736FB39" box="[1001,1075,1158,1177]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="13" pageNumber="498" unit="mm" value="2.4">2.4 mm</quantity>
. Total body length:
<quantity id="FD0F9B3EFFE7FFC3FE3BFB4692F2FB11" box="[428,503,1182,1201]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="498" unit="mm" value="8.1">8.1 mm</quantity>
. Humeral width:
<quantity id="FD0F9B3EFFE7FFC3FD09FB4691EFFB11" box="[670,746,1182,1201]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="498" unit="mm" value="3.3">3.3 mm</quantity>
.
</paragraph>
<paragraph id="3A4836DBFFE7FFC3FE87FB169105FA24" blockId="13.[246,1079,1230,1652]" pageId="13" pageNumber="498">
<emphasis id="0883EAC9FFE7FFC3FE87FB169270FB43" bold="true" box="[272,373,1230,1251]" pageId="13" pageNumber="498">
<typeStatus id="E54C8879FFE7FFC3FE87FB169275FB43" box="[272,368,1230,1251]" pageId="13" pageNumber="498" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Similar to male except as follows: antennal club as long as preceeding four segments (1:1); frontal and pronotal punctures more deep; anal plate deep punctate; protibial external border with 3 teeth clearly marked. Ventral genital plates slightly sclerotized, symmetrical, with apical border widely rounded, without setae; dorsal genital plates rounded, slightly narrowed, with some erect setae at apical border (
<figureCitation id="A2CC2A5EFFE7FFC3FD32FA8C91FDFAC9" box="[677,760,1364,1385]" captionStart="Figs" captionStartId="12.[272,315,1139,1158]" captionTargetBox="[249,1084,270,1114]" captionTargetId="figure-12@12.[244,1087,265,1118]" captionTargetPageId="12" captionText="Figs. 2327. Phyllophaga jalisciensis. 23) Head and pronotum, dorsal view; 24) tarsal claw, male; 25) paramera, distal view; 26) genital capsule, lateral view; 27) ventral genital plates, female. Scale lines: Figs. 24, 27 = 0.25 mm; Figs. 23, 2526 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102845" httpUri="https://zenodo.org/record/10102845/files/figure.png" pageId="13" pageNumber="498">Fig. 27</figureCitation>
). Total body length:
<quantity id="FD0F9B3EFFE7FFC3FC4BFA8C9736FAC9" box="[988,1075,1364,1385]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="13" pageNumber="498" unit="mm" value="8.9">8.9 mm</quantity>
. Humeral width:
<quantity id="FD0F9B3EFFE7FFC3FE31FAB792F9FA24" box="[422,508,1391,1412]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="498" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
<paragraph id="3A4836DBFFE7FFC3FE87FA5191C5FA74" blockId="13.[246,1079,1230,1652]" pageId="13" pageNumber="498">
<emphasis id="0883EAC9FFE7FFC3FE87FA5192F1FA3E" bold="true" box="[272,500,1417,1438]" pageId="13" pageNumber="498">
<typeStatus id="E54C8879FFE7FFC3FE87FA519273FA3E" box="[272,374,1417,1438]" pageId="13" pageNumber="498" type="paratype">Paratype</typeStatus>
Variation.
</emphasis>
<typeStatus id="E54C8879FFE7FFC3FE68FA519160FA3E" box="[511,613,1417,1438]" pageId="13" pageNumber="498" type="paratype">Paratypes</typeStatus>
of both sexes are similar to types except that some specimens present pronotum completely darkned. Total body length:
<quantity id="FD0F9B3EFFE7FFC3FF61FA67927EFA74" box="[246,379,1471,1492]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.5" metricValueMin="7.9" pageId="13" pageNumber="498" unit="mm" value="8.7" valueMax="9.5" valueMin="7.9">7.99.5 mm</quantity>
. Humeral width: 3.0
<quantity id="FD0F9B3EFFE7FFC3FDF1FA6791B9FA74" box="[614,700,1471,1492]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="13" pageNumber="498" unit="mm" value="3.6">3.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE7FFC0FE87FA01971DFD8E" lastPageId="14" lastPageNumber="499" pageId="13" pageNumber="498" type="materials_examined">
<paragraph id="3A4836DBFFE7FFC0FE87FA01912BFE59" blockId="13.[246,1079,1230,1652]" lastBlockId="14.[249,1082,269,985]" lastPageId="14" lastPageNumber="499" pageId="13" pageNumber="498">
<emphasis id="0883EAC9FFE7FFC3FE87FA01929CFA4E" bold="true" box="[272,409,1497,1518]" pageId="13" pageNumber="498">Type Series.</emphasis>
(described from
<specimenCount id="2CF1FD52FFE7FFC3FDDBFA0191AFFA4E" box="[588,682,1497,1518]" pageId="13" pageNumber="498" type="male">42 males</specimenCount>
and
<specimenCount id="2CF1FD52FFE7FFC3FD4BFA019056FA4E" box="[732,851,1497,1518]" pageId="13" pageNumber="498" type="female">21 females</specimenCount>
).
<materialsCitation id="8A9F3C86FFE7FFC3FCF3FA019180F984" collectingDate="1997-07-04" collectorName="M. Lopez" elevation="1900" location="Estacion Las Joyas" municipality="Autlan" pageId="13" pageNumber="498" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<typeStatus id="E54C8879FFE7FFC3FCF3FA0190CDFA4E" box="[868,968,1497,1518]" pageId="13" pageNumber="498" type="holotype">Holotype</typeStatus>
, ♂; MEX­ ICO:
<emphasis id="0883EAC9FFE7FFC3FEA2FA2C9280F9A9" box="[309,389,1524,1545]" italics="true" pageId="13" pageNumber="498">
<collectingRegion id="F833F839FFE7FFC3FEA2FA2C9287F9A9" box="[309,386,1524,1545]" country="Mexico" name="Jalisco" pageId="13" pageNumber="498">Jalisco</collectingRegion>
,
</emphasis>
<collectingMunicipality id="DA2CACA1FFE7FFC3FE07FA2C92DFF9A9" box="[400,474,1524,1545]" pageId="13" pageNumber="498">Autlán</collectingMunicipality>
,
<location id="3F286000FFE7FFC3FE7EFA2C91BEF9A9" LSID="urn:lsid:plazi:treatment:B25E87CDFFE6FFC0FE97FA7F9124FC79:3F286000FFE7FFC3FE7EFA2C91BEF9A9" box="[489,699,1524,1545]" municipality="Autlan" name="Estacion Las Joyas" pageId="13" pageNumber="498" stateProvince="Jalisco">Estación Las Joyas</location>
,
<date id="4E49101BFFE7FFC3FD5EFA2C9046F9A9" box="[713,835,1524,1545]" pageId="13" pageNumber="498" value="1997-07-04">
<collectingDate id="5E0DE9F3FFE7FFC3FD5EFA2C9046F9A9" box="[713,835,1524,1545]" pageId="13" pageNumber="498" value="1997-07-04">4­VII­1997</collectingDate>
</date>
,
<quantity id="FD0F9B3EFFE7FFC3FCC6FA2C90ABF9A9" box="[849,942,1524,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="498" unit="m" value="1900.0">
<elevation id="B1DAD1E8FFE7FFC3FCC6FA2C90ABF9A9" box="[849,942,1524,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="498" unit="m" value="1900.0">1,900 m</elevation>
</quantity>
, trampa luz, bosque mesófilo,
<collectorName id="9702530DFFE7FFC3FE25F9D7911FF984" box="[434,538,1551,1572]" pageId="13" pageNumber="498">M. López</collectorName>
(
<collectionCode id="5CE6AE1EFFE7FFC3FDBCF9D7917BF984" box="[555,638,1551,1572]" pageId="13" pageNumber="498">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFE7FFC3FD06F9D79199F99E" collectingDate="1997-08-05" collectorName="Vazquez" elevation="1900" location="Estacion Las Joyas" municipality="Autlan" pageId="13" pageNumber="498" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="allotype">
<typeStatus id="E54C8879FFE7FFC3FD06F9D791EAF984" box="[657,751,1551,1572]" pageId="13" pageNumber="498" type="allotype">Allotype</typeStatus>
,
<specimenCount id="2CF1FD52FFE7FFC3FD6BF9D59015F983" box="[764,784,1549,1571]" pageId="13" pageNumber="498" type="female"></specimenCount>
same data as holotype, except
<date id="4E49101BFFE7FFC3FEBEF9F192AEF99E" box="[297,427,1577,1598]" pageId="13" pageNumber="498" value="1997-08-05">
<collectingDate id="5E0DE9F3FFE7FFC3FEBEF9F192AEF99E" box="[297,427,1577,1598]" pageId="13" pageNumber="498" value="1997-08-05">5­VIII­1997</collectingDate>
</date>
, A.
<collectorName id="9702530DFFE7FFC3FE40F9F19137F99E" box="[471,562,1577,1598]" pageId="13" pageNumber="498">Vázquez</collectorName>
(
<collectionCode id="5CE6AE1EFFE7FFC3FDD5F9F19190F99E" box="[578,661,1577,1598]" pageId="13" pageNumber="498">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFE7FFC3FD30F9F191D1F9F9" collectingDate="1997-07-04" collectorName="S. Nunez" elevation="1900" location="Estacion Las Joyas" municipality="Autlan" pageId="13" pageNumber="498" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
<typeStatus id="E54C8879FFE7FFC3FD30F9F19014F99E" box="[679,785,1577,1598]" pageId="13" pageNumber="498" type="paratype">Paratypes</typeStatus>
: same data as holotype (♂); same data as
<typeStatus id="E54C8879FFE7FFC3FE1BF99C92E6F9F9" box="[396,483,1604,1625]" pageId="13" pageNumber="498" type="allotype">allotype</typeStatus>
, except
<collectorName id="9702530DFFE7FFC3FDA8F99C91A6F9F9" box="[575,675,1604,1625]" pageId="13" pageNumber="498">S. Nuñez</collectorName>
(
<specimenCount id="2CF1FD52FFE7FFC3FD21F99B91CFF9F9" box="[694,714,1603,1625]" pageId="13" pageNumber="498" type="female"></specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFE7FFC3FD73F99C9058F9F9" box="[740,861,1604,1625]" pageId="13" pageNumber="498" value="1997-07-03">3­VII­1997</date>
, J. García (2 ♂♂
<specimenCount id="2CF1FD52FFE7FFC3FBBEF99C920FF9D3" pageId="13" pageNumber="498" type="female">1 ♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFE7FFC3FEB3F98792DFF9D4" box="[292,474,1629,1652]" location="Lopez" pageId="13" pageNumber="498" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
M.
<location id="3F286000FFE7FFC3FEDAF987928BF9D4" LSID="urn:lsid:plazi:treatment:B25E87CDFFE6FFC0FE97FA7F9124FC79:3F286000FFE7FFC3FEDAF987928BF9D4" box="[333,398,1631,1652]" name="Lopez" pageId="13" pageNumber="498" stateProvince="Jalisco">López</location>
(♂
<emphasis id="0883EAC9FFE7FFC3FE22F98692B9F9D3" bold="true" box="[437,444,1630,1651]" pageId="13" pageNumber="498">/</emphasis>
<specimenCount id="2CF1FD52FFE7FFC3FE2BF98592D5F9D3" box="[444,464,1629,1651]" pageId="13" pageNumber="498" type="female"></specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFE7FFC3FE7CF9879168F9D4" box="[491,621,1631,1652]" pageId="13" pageNumber="498" value="1997-08-05">5­VIII­1997</date>
, J. García (♂);
<date id="4E49101BFFE7FFC3FC89F9879098F9D4" box="[798,925,1631,1652]" pageId="13" pageNumber="498" value="1997-08-06">6­VIII­1997</date>
(3 ♂♂
<specimenCount id="2CF1FD52FFE7FFC3FC6CF9879723F9D3" box="[1019,1062,1629,1652]" pageId="13" pageNumber="498" type="female">1 ♀</specimenCount>
); A. Partida (6 ♂♂); M. López (
<specimenCount id="2CF1FD52FFE4FFC0FDC1FED5916FFE83" box="[598,618,269,291]" pageId="14" pageNumber="499" type="female"></specimenCount>
);
<date id="4E49101BFFE4FFC0FD12FED7900DFE84" box="[645,776,271,292]" pageId="14" pageNumber="499" value="1997-08-07">7­VIII­1997</date>
, J. García (7 ♂♂
<specimenCount id="2CF1FD52FFE4FFC0FC4FFED79700FE83" box="[984,1029,269,292]" pageId="14" pageNumber="499" type="female">1 ♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFE4FFC0FBB7FED792ECFE9E" location="Partida" pageId="14" pageNumber="499" specimenCount="3" specimenCount-female="3" stateProvince="Jalisco" typeStatus="paratype">
A.
<location id="3F286000FFE4FFC0FF6EFEF19246FE9E" LSID="urn:lsid:plazi:treatment:B25E87CDFFE6FFC0FE97FA7F9124FC79:3F286000FFE4FFC0FF6EFEF19246FE9E" box="[249,323,297,318]" name="Partida" pageId="14" pageNumber="499" stateProvince="Jalisco">Partida</location>
(3 ♂♂
<specimenCount id="2CF1FD52FFE4FFC0FE08FEF192DAFE9E" box="[415,479,296,318]" pageId="14" pageNumber="499" type="female">3 ♀♀</specimenCount>
)
</materialsCitation>
; S. Núñez (♂); M. López (
<specimenCount id="2CF1FD52FFE4FFC0FC8DFEF0902BFE9E" box="[794,814,296,318]" pageId="14" pageNumber="499" type="female"></specimenCount>
);
<date id="4E49101BFFE4FFC0FCDFFEF190BCFE9E" box="[840,953,297,318]" pageId="14" pageNumber="499" value="1997-09-04">4­IX­1997</date>
, A. Guerra, (2 ♂♂); M. López, (2 ♂♂
<specimenCount id="2CF1FD52FFE4FFC0FDB3FE9C914AFEF9" box="[548,591,323,345]" pageId="14" pageNumber="499" type="female">1 ♀</specimenCount>
);
<date id="4E49101BFFE4FFC0FDFEFE9C91DFFEF9" box="[617,730,324,345]" pageId="14" pageNumber="499" value="1997-09-05">5­IX­1997</date>
, M. López (♂);
<date id="4E49101BFFE4FFC0FC07FE9C9708FEF9" box="[912,1037,324,345]" pageId="14" pageNumber="499" value="1997-09-30">30­IX­1997</date>
, M. López (2 ♂♂);
<date id="4E49101BFFE4FFC0FE22FE87911EFED4" box="[437,539,351,372]" pageId="14" pageNumber="499" value="1997-10-01">1­X­1997</date>
(♂
<emphasis id="0883EAC9FFE4FFC0FDD5FE86914CFED3" bold="true" box="[578,585,350,371]" pageId="14" pageNumber="499">/</emphasis>
<specimenCount id="2CF1FD52FFE4FFC0FDDEFE859158FED3" box="[585,605,349,371]" pageId="14" pageNumber="499" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFE4FFC0FDEFFE8790BBFED4" box="[632,958,349,372]" collectingDate="1997-10-02" location="Pelayo" pageId="14" pageNumber="499" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
D.
<location id="3F286000FFE4FFC0FD0AFE8791E1FED4" LSID="urn:lsid:plazi:treatment:B25E87CDFFE6FFC0FE97FA7F9124FC79:3F286000FFE4FFC0FD0AFE8791E1FED4" box="[669,740,351,372]" name="Pelayo" pageId="14" pageNumber="499" stateProvince="Jalisco">Pelayo</location>
(♂);
<date id="4E49101BFFE4FFC0FCB0FE879089FED4" box="[807,908,351,372]" pageId="14" pageNumber="499" value="1997-10-02">
<collectingDate id="5E0DE9F3FFE4FFC0FCB0FE879089FED4" box="[807,908,351,372]" pageId="14" pageNumber="499" value="1997-10-02">2­X­1997</collectingDate>
</date>
(
<specimenCount id="2CF1FD52FFE4FFC0FC37FE8590B1FED3" box="[928,948,349,371]" pageId="14" pageNumber="499" type="female"></specimenCount>
)
</materialsCitation>
; M. López (♂);
<date id="4E49101BFFE4FFC0FEBBFEA192B7FE2E" box="[300,434,377,398]" pageId="14" pageNumber="499" value="1999-07-30">30­VII­1999</date>
, G. Mesa (♂
<emphasis id="0883EAC9FFE4FFC0FDAEFEA19145FE2E" bold="true" box="[569,576,377,398]" pageId="14" pageNumber="499">/</emphasis>
<specimenCount id="2CF1FD52FFE4FFC0FDD7FEA09151FE2E" box="[576,596,376,398]" pageId="14" pageNumber="499" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFE4FFC0FDFCFEA19015FE2E" box="[619,784,376,398]" location="Garcia" pageId="14" pageNumber="499" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
J.
<location id="3F286000FFE4FFC0FD15FEA191CDFE2E" LSID="urn:lsid:plazi:treatment:B25E87CDFFE6FFC0FE97FA7F9124FC79:3F286000FFE4FFC0FD15FEA191CDFE2E" box="[642,712,377,398]" name="Garcia" pageId="14" pageNumber="499" stateProvince="Jalisco">García</location>
(♂
<emphasis id="0883EAC9FFE4FFC0FD7CFEA191F7FE2E" bold="true" box="[747,754,377,398]" pageId="14" pageNumber="499">/</emphasis>
<specimenCount id="2CF1FD52FFE4FFC0FD65FEA09003FE2E" box="[754,774,376,398]" pageId="14" pageNumber="499" type="female"></specimenCount>
)
</materialsCitation>
; V. González (♂); M. López (
<specimenCount id="2CF1FD52FFE4FFC0FE95FE4B9213FE09" box="[258,278,403,425]" pageId="14" pageNumber="499" type="female"></specimenCount>
);
<date id="4E49101BFFE4FFC0FEA6FE4C92BDFE09" box="[305,440,404,425]" pageId="14" pageNumber="499" value="1999-07-31">31­VII­1999</date>
, V. González (♂);
<date id="4E49101BFFE4FFC0FD18FE4C9008FE09" box="[655,781,404,425]" pageId="14" pageNumber="499" value="1986-06-19">19­VI­1986</date>
, L. Rivera, (♂
<emphasis id="0883EAC9FFE4FFC0FC24FE4C90BFFE09" bold="true" box="[947,954,404,425]" pageId="14" pageNumber="499">/</emphasis>
<specimenCount id="2CF1FD52FFE4FFC0FC2DFE4B90CBFE09" box="[954,974,403,425]" pageId="14" pageNumber="499" type="female"></specimenCount>
);
<date id="4E49101BFFE4FFC0FC7EFE4C9235FE64" pageId="14" pageNumber="499" value="1986-07-12">12­VII­ 1986</date>
, barbechos, L. Rivera,(♂).
<typeStatus id="E54C8879FFE4FFC0FDC8FE7791C0FE64" box="[607,709,431,452]" pageId="14" pageNumber="499" type="paratype">Paratypes</typeStatus>
are deposited at AAHC,
<collectionCode id="5CE6AE1EFFE4FFC0FC74FE779733FE64" box="[995,1078,431,452]" pageId="14" pageNumber="499">AMNH</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FF6EFE119245FE7E" box="[249,320,457,478]" pageId="14" pageNumber="499">CASC</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FEC4FE11928EFE7E" box="[339,395,457,478]" pageId="14" pageNumber="499">CNC</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FE08FE1192E8FE7E" box="[415,493,457,478]" pageId="14" pageNumber="499">FMNH</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FD96FE11916BFE7E" box="[513,622,457,478]" pageId="14" pageNumber="499">IBUNAM</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FD16FE1191C7FE7E" box="[641,706,457,478]" pageId="14" pageNumber="499">IEXA</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FD42FE119017FE7E" box="[725,786,457,478]" pageId="14" pageNumber="499">IMEJ</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FCB2FE119068FE7E" box="[805,877,457,478]" pageId="14" pageNumber="499">GNPC</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FC17FE1190D6FE7E" box="[896,979,457,478]" pageId="14" pageNumber="499">MNHN</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FC71FE119733FE7E" box="[998,1078,457,478]" pageId="14" pageNumber="499">MXAL</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FF6EFE3C924FFE59" box="[249,330,484,505]" pageId="14" pageNumber="499">NHMC</collectionCode>
,
<collectionCode id="5CE6AE1EFFE4FFC0FEC0FE3C92A7FE59" box="[343,418,484,505]" pageId="14" pageNumber="499">UNSM</collectionCode>
and
<collectionCode id="5CE6AE1EFFE4FFC0FE4DFE3C912FFE59" box="[474,554,484,505]" pageId="14" pageNumber="499">ZMHU</collectionCode>
.
</paragraph>
<paragraph id="3A4836DBFFE4FFC0FE84FE26971DFD8E" blockId="14.[249,1082,269,985]" pageId="14" pageNumber="499">
<materialsCitation id="8A9F3C86FFE4FFC0FE84FE269710FD8E" country="Mexico" county="Autlan" location="Las Joyas" municipality="Biosphere Reserve Sierra de Manantlan" pageId="14" pageNumber="499" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="0883EAC9FFE4FFC0FE84FE2692B2FDB3" bold="true" box="[275,439,510,531]" pageId="14" pageNumber="499">
<typeStatus id="E54C8879FFE4FFC0FE84FE269249FDB3" box="[275,332,510,531]" pageId="14" pageNumber="499">Type</typeStatus>
Locality.
</emphasis>
<location id="3F286000FFE4FFC0FE57FE279129FDB4" LSID="urn:lsid:plazi:treatment:B25E87CDFFE6FFC0FE97FA7F9124FC79:3F286000FFE4FFC0FE57FE279129FDB4" box="[448,556,511,532]" country="Mexico" county="Autlan" municipality="Biosphere Reserve Sierra de Manantlan" name="Las Joyas" pageId="14" pageNumber="499" stateProvince="Jalisco">Las Joyas</location>
,
<collectingMunicipality id="DA2CACA1FFE4FFC0FDAFFE2790E0FDB4" box="[568,997,511,532]" pageId="14" pageNumber="499">Biosphere Reserve Sierra de Manantlán</collectingMunicipality>
,
<collectingCounty id="D3294E57FFE4FFC0FC65FE27973CFDB4" box="[1010,1081,511,532]" pageId="14" pageNumber="499">Autlán</collectingCounty>
municipality, state of
<collectingRegion id="F833F839FFE4FFC0FE74FDC19128FD8E" box="[483,557,537,558]" country="Mexico" name="Jalisco" pageId="14" pageNumber="499">Jalisco</collectingRegion>
,
<collectingCountry id="42E0764BFFE4FFC0FDADFDC1918FFD8E" box="[570,650,537,558]" name="Mexico" pageId="14" pageNumber="499">México</collectingCountry>
(approx. 19
<emphasis id="0883EAC9FFE4FFC0FC99FDC19013FD8E" bold="true" box="[782,790,537,558]" pageId="14" pageNumber="499">°</emphasis>
35
<emphasis id="0883EAC9FFE4FFC0FCA7FDC0903DFD8E" bold="true" box="[816,824,536,558]" pageId="14" pageNumber="499">'</emphasis>
13
<emphasis id="0883EAC9FFE4FFC0FCC5FDC0905FFD8E" bold="true" box="[850,858,536,558]" pageId="14" pageNumber="499">&quot;</emphasis>
N; 104
<emphasis id="0883EAC9FFE4FFC0FC33FDC190A9FD8E" bold="true" box="[932,940,537,558]" pageId="14" pageNumber="499">°</emphasis>
16
<emphasis id="0883EAC9FFE4FFC0FC51FDC090CBFD8E" bold="true" box="[966,974,536,558]" pageId="14" pageNumber="499">'</emphasis>
48
<emphasis id="0883EAC9FFE4FFC0FC7FFDC090F5FD8E" bold="true" box="[1000,1008,536,558]" pageId="14" pageNumber="499">&quot;</emphasis>
W)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE4FFC0FE84FDEC91A2FD6E" pageId="14" pageNumber="499" type="biology_ecology">
<paragraph id="3A4836DBFFE4FFC0FE84FDEC91A2FD6E" blockId="14.[249,1082,269,985]" pageId="14" pageNumber="499">
<emphasis id="0883EAC9FFE4FFC0FE84FDEC92C2FDE9" bold="true" box="[275,455,564,585]" pageId="14" pageNumber="499">Biological Data.</emphasis>
Specimens of
<taxonomicName id="FDF74D58FFE4FFC0FDFEFDEC91F8FDE9" box="[617,765,564,585]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="jalisciensis" status="sp. nov.">
<emphasis id="0883EAC9FFE4FFC0FDFEFDEC91F8FDE9" box="[617,765,564,585]" italics="true" pageId="14" pageNumber="499">P. jalisciensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE4FFC0FC91FDEC9032FDE9" box="[774,823,564,585]" pageId="14" pageNumber="499" rank="species">n.sp.</taxonomicNameLabel>
were collected at lights in cloud forest located at
<quantity id="FD0F9B3EFFE4FFC0FD98FD979163FDC4" box="[527,614,591,612]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="499" unit="m" value="1900.0">1,900 m</quantity>
of altitude. Phenology: June (2), July (15), August (32), September (9), October (5). Other species flying at the same time and place were
<taxonomicName id="FDF74D58FFE4FFC0FE32FD5C9076FD39" authority="Moron &amp; Rivera" authorityName="Moron &amp; Rivera-Cervantes" authorityYear="1992" box="[421,883,644,665]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="mesophylla" subGenus="Phytalus">
<emphasis id="0883EAC9FFE4FFC0FE32FD5C91B6FD39" box="[421,691,644,665]" italics="true" pageId="14" pageNumber="499">P. (Phytalus) mesophylla</emphasis>
Morón &amp; Rivera
</taxonomicName>
,
<taxonomicName id="FDF74D58FFE4FFC0FC17FD5C924FFD14" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="dugesiana" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFE4FFC0FC17FD5C924FFD14" italics="true" pageId="14" pageNumber="499">P. (Phytalus) dugesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE4FFC0FEC3FD479294FD14" box="[340,401,671,692]" pageId="14" pageNumber="499" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFE4FFC0FE37FD479171FD14" box="[416,628,671,692]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="sayloriana" status="sp. nov.">
<emphasis id="0883EAC9FFE4FFC0FE37FD4792B2FD14" box="[416,439,671,692]" italics="true" pageId="14" pageNumber="499">P.</emphasis>
(
<emphasis id="0883EAC9FFE4FFC0FE28FD4792F7FD14" box="[447,498,671,692]" italics="true" pageId="14" pageNumber="499">s.str.</emphasis>
)
<emphasis id="0883EAC9FFE4FFC0FD93FD479171FD14" box="[516,628,671,692]" italics="true" pageId="14" pageNumber="499">sayloriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE4FFC0FDE9FD4791B4FD14" box="[638,689,671,692]" pageId="14" pageNumber="499" rank="species">n.sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFE4FFC0FD57FD4790A4FD14" box="[704,929,671,692]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="manantleca">
<emphasis id="0883EAC9FFE4FFC0FD57FD4791D2FD14" box="[704,727,671,692]" italics="true" pageId="14" pageNumber="499">P.</emphasis>
(
<emphasis id="0883EAC9FFE4FFC0FD48FD479017FD14" box="[735,786,671,692]" italics="true" pageId="14" pageNumber="499">s.str.</emphasis>
)
<emphasis id="0883EAC9FFE4FFC0FCB3FD4790A4FD14" box="[804,929,671,692]" italics="true" pageId="14" pageNumber="499">manantleca</emphasis>
</taxonomicName>
<emphasis id="0883EAC9FFE4FFC0FC36FD4790A0FD14" box="[929,933,671,692]" italics="true" pageId="14" pageNumber="499">,</emphasis>
<taxonomicName id="FDF74D58FFE4FFC0FC39FD479229FD6F" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="joyana" status="sp. nov.">
<emphasis id="0883EAC9FFE4FFC0FC39FD4790C0FD14" box="[942,965,671,692]" italics="true" pageId="14" pageNumber="499">P.</emphasis>
(
<emphasis id="0883EAC9FFE4FFC0FC40FD47970FFD14" box="[983,1034,671,692]" italics="true" pageId="14" pageNumber="499">s.str.</emphasis>
)
<emphasis id="0883EAC9FFE4FFC0FB8BFD479229FD6F" italics="true" pageId="14" pageNumber="499">joyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE4FFC0FEA2FD619262FD6E" box="[309,359,697,718]" pageId="14" pageNumber="499" rank="species">n.sp.</taxonomicNameLabel>
and one undescribed species.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE4FFC0FE84FD0C92C5FCCF" pageId="14" pageNumber="499" type="discussion">
<paragraph id="3A4836DBFFE4FFC0FE84FD0C92C5FCCF" blockId="14.[249,1082,269,985]" pageId="14" pageNumber="499">
<emphasis id="0883EAC9FFE4FFC0FE84FD0C9285FD49" bold="true" box="[275,384,724,745]" pageId="14" pageNumber="499">Remarks.</emphasis>
<taxonomicName id="FDF74D58FFE4FFC0FE1EFD0C9189FD49" box="[393,652,724,745]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="jalisciensis" status="sp. nov.">
<emphasis id="0883EAC9FFE4FFC0FE1EFD0C9189FD49" box="[393,652,724,745]" italics="true" pageId="14" pageNumber="499">Phyllophaga jalisciensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE4FFC0FD01FD0C91CDFD49" box="[662,712,724,745]" pageId="14" pageNumber="499" rank="species">n.sp.</taxonomicNameLabel>
presents the diagnostic characters of the species group
<emphasis id="0883EAC9FFE4FFC0FE60FD379161FCA4" box="[503,612,751,772]" italics="true" pageId="14" pageNumber="499">bucephala</emphasis>
, with some similarity with
<taxonomicName id="FDF74D58FFE4FFC0FC38FD379236FCBE" authority="Bates" authorityName="Bates" authorityYear="1890" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="omiltemia">
<emphasis id="0883EAC9FFE4FFC0FC38FD37973FFCA4" box="[943,1082,751,772]" italics="true" pageId="14" pageNumber="499">P. omiltemia</emphasis>
Bates
</taxonomicName>
and
<taxonomicName id="FDF74D58FFE4FFC0FEFEFCD29038FCBE" authority="Moron &amp; Cervantes. The" authorityName="Moron &amp; Cervantes. The" box="[361,829,777,799]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="teosinteophaga">
<emphasis id="0883EAC9FFE4FFC0FEFEFCD2912DFCBF" box="[361,552,778,799]" italics="true" pageId="14" pageNumber="499">P. teosinteophaga</emphasis>
Morón &amp; Cervantes. The
</taxonomicName>
dense, ventral vestiture of the tarsal segments and male genital capsule are very different from the
<emphasis id="0883EAC9FFE4FFC0FBB4FCFC926CFCF4" italics="true" pageId="14" pageNumber="499">
<taxonomicName id="FDF74D58FFE4FFC0FBB4FCFC9260FCF4" authorityName="Bates" authorityYear="1890" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="omiltemia">P. omiltemia</taxonomicName>
.
</emphasis>
The bicolored pronotum and shape of parameres are different from
<emphasis id="0883EAC9FFE4FFC0FF6EFC8292C5FCCF" box="[249,448,858,879]" italics="true" pageId="14" pageNumber="499">
<taxonomicName id="FDF74D58FFE4FFC0FF6EFC8292B9FCCF" box="[249,444,858,879]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="teosinteophaga">P. teosinteophaga</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE4FFC0FE84FCAC9124FC79" pageId="14" pageNumber="499" type="etymology">
<paragraph id="3A4836DBFFE4FFC0FE84FCAC9124FC79" blockId="14.[249,1082,269,985]" pageId="14" pageNumber="499">
<emphasis id="0883EAC9FFE4FFC0FE84FCAC9296FC29" bold="true" box="[275,403,884,905]" pageId="14" pageNumber="499">Etymology.</emphasis>
Derived from the name of state of
<collectingRegion id="F833F839FFE4FFC0FC83FCAC905EFC29" box="[788,859,884,905]" country="Mexico" name="Jalisco" pageId="14" pageNumber="499">Jalisco</collectingRegion>
or Xallisco, old Nahuatl language name meaning
<emphasis id="0883EAC9FFE4FFC0FDD5FC57917EFC04" box="[578,635,911,932]" italics="true" pageId="14" pageNumber="499">Xalli,</emphasis>
sand,
<emphasis id="0883EAC9FFE4FFC0FD75FC5791FEFC04" box="[738,763,911,932]" italics="true" pageId="14" pageNumber="499">co</emphasis>
in, in sandy land (
<bibRefCitation id="5E664B2AFFE4FFC0FB92FC579264FC1E" author="Simeon, R." bookContentInfo="Siglo XXI Editores, Mexico. 782 pp." pageId="14" pageNumber="499" refId="ref10587" refString="Simeon, R. 1988. Diccionario de la lengua nahuatl o mexicana. Siglo XXI Editores, Mexico. 782 pp." title="Diccionario de la lengua nahuatl o mexicana" type="book" year="1988">Simeón 1988</bibRefCitation>
) and Latin suffix ­
<emphasis id="0883EAC9FFE4FFC0FDAFFC729176FC1F" box="[568,627,938,959]" italics="true" pageId="14" pageNumber="499">ensis,</emphasis>
belonging (
<bibRefCitation id="5E664B2AFFE4FFC0FCB2FC7190A8FC1E" author="Jaeger, E. C." bookContentInfo="Charles C. Thomas Publisher. Spingfield, Illinois. 323 pp." box="[805,941,937,958]" pageId="14" pageNumber="499" refId="ref10411" refString="Jaeger, E. C. 1978. A source book of biological names and terms. (Third edition). Charles C. Thomas Publisher. Spingfield, Illinois. 323 pp." title="A source book of biological names and terms. (Third edition)" type="book" year="1978">Jaeger 1978</bibRefCitation>
);
<taxonomicName id="FDF74D58FFE4FFC0FC52FC72973CFC1F" box="[965,1081,938,959]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="499" phylum="Arthropoda" rank="species" species="jalisciensis">
<emphasis id="0883EAC9FFE4FFC0FC52FC72973CFC1F" box="[965,1081,938,959]" italics="true" pageId="14" pageNumber="499">jalisciensis</emphasis>
</taxonomicName>
belonging to sandy land.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,815 @@
<document id="8EC542E76E10F915CCAFCAAFD62EA73E" ID-DOI="10.1649/0010-065X(2001)055[0485:NSOPHC]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102835" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630156134743" checkinUser="tatiana" docAuthor="Morón, Miguel-Angel, Rivera-Cervantes, Luis Eugenio &amp; López-Vieyra, Mario" docDate="2001" docId="B25E87CDFFEBFFCAFF60FB939703FAA4" docLanguage="en" docName="TheColeopBull.55.4.485-500.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docSource="http://dx.doi.org/10.1649/0010-065x(2001)055[0485:nsophc]2.0.co;2" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) sayloriana Moron, Rivera y Lopez, new species" docType="treatment" docVersion="1" lastPageNumber="489" masterDocId="4E67FFB5FFEAFFCEFF97FFD89305FFA0" masterDocTitle="New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México" masterLastPageNumber="500" masterPageNumber="485" pageNumber="486" updateTime="1730145459161" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="FB9FACDFB5150B8F7A7FC9F389102A3B">New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México</mods:title>
</mods:titleInfo>
<mods:name id="36567F8D219E111F8F772994698A9623" type="personal">
<mods:role id="0F4CD21159FAE442B93C2BC76176D5D3">
<mods:roleTerm id="CE257165FDDD46F3E446A6C3091340DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6E861735E5BC7722E11029B64BB5641">Morón, Miguel-Angel</mods:namePart>
<mods:affiliation id="A5D65C7CE2CC3200A1CCEBC44F245C53">Departamento de Entomología Instituto de Ecología, A. C. (Sector SEP­CONACYT) Apdo. Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="101AB5E1C1B052F6E7F087077B5C4134">Rivera-Cervantes, Luis Eugenio</mods:namePart>
<mods:affiliation id="491B0F0E49451BB462667C5463A176CC">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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</mods:role>
<mods:namePart id="88EB247D96A72D76C840252B42FA3C89">López-Vieyra, Mario</mods:namePart>
<mods:affiliation id="E739BC58851D60624CE8C52D90132C09">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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<mods:date id="24FC2DBCF0B9F80D3C14AC96AD36D3CB">2001</mods:date>
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<treatment id="B25E87CDFFEBFFCAFF60FB939703FAA4" LSID="urn:lsid:plazi:treatment:B25E87CDFFEBFFCAFF60FB939703FAA4" httpUri="http://treatment.plazi.org/id/B25E87CDFFEBFFCAFF60FB939703FAA4" lastPageId="4" lastPageNumber="489" pageId="1" pageNumber="486">
<subSubSection id="72ED6550FFEBFFCFFF60FB939729FBFF" box="[247,1068,1098,1120]" pageId="1" pageNumber="486" type="nomenclature">
<paragraph id="3A4836DBFFEBFFCFFF60FB939729FBFF" blockId="1.[247,1068,1098,1146]" box="[247,1068,1098,1120]" pageId="1" pageNumber="486">
<taxonomicName id="FDF74D58FFEBFFCFFF60FB939098FBC0" authority="Moron, Rivera y Lopez" authorityName="Moron, Rivera y Lopez" box="[247,925,1099,1120]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="486" phylum="Arthropoda" rank="species" species="sayloriana" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="0883EAC9FFEBFFCFFF60FB939190FBC0" box="[247,661,1099,1120]" italics="true" pageId="1" pageNumber="486">Phyllophaga (Phyllophaga) sayloriana</emphasis>
Morón, Rivera y López
</taxonomicName>
,
<emphasis id="0883EAC9FFEBFFCFFC3EFB929729FBFF" bold="true" box="[937,1068,1098,1119]" pageId="1" pageNumber="486">
<taxonomicNameLabel id="13B057B2FFEBFFCFFC3EFB929729FBFF" box="[937,1068,1098,1119]" pageId="1" pageNumber="486" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFEBFFCDFDF3FBBD9255FB41" lastPageId="3" lastPageNumber="488" pageId="1" pageNumber="486" type="description">
<paragraph id="3A4836DBFFEBFFCFFDF3FBBD91CDFBDA" blockId="1.[247,1068,1098,1146]" box="[612,712,1125,1146]" pageId="1" pageNumber="486">
<figureCitation id="A2CC2A5EFFEBFFCFFDF3FBBD91CDFBDA" box="[612,712,1125,1146]" captionStart="Figs" captionStartId="2.[272,315,1339,1358]" captionTargetBox="[256,1071,268,1318]" captionTargetId="figure-12@2.[254,1075,265,1318]" captionTargetPageId="2" captionText="Figs. 18. Phyllophaga sayloriana. 1) male mesotarsomeres, latero­ventral view; 2) male metatarsomeres, latero­ventral view; 3) tarsal claw, male; 4) genital capsule, lateral view; 5) paramera, distal view; 6) apex of paramera, distal­ventral view; 7) distal half of genital capsule, dorsal view; 8) ventral genital plates, female. Scale lines: Figs. 13 = 0.5 mm; Figs. 48 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102837" httpUri="https://zenodo.org/record/10102837/files/figure.png" pageId="1" pageNumber="486">Figs. 18</figureCitation>
</paragraph>
<paragraph id="3A4836DBFFEBFFCDFE9AFB4B9189FD49" blockId="1.[246,1079,1171,1652]" lastBlockId="3.[246,1079,270,745]" lastPageId="3" lastPageNumber="488" pageId="1" pageNumber="486">
<emphasis id="0883EAC9FFEBFFCFFE9AFB4B92F5FB06" bold="true" box="[269,496,1171,1190]" pageId="1" pageNumber="486">
Description.
<typeStatus id="E54C8879FFEBFFCFFE07FB4B92E9FB06" box="[400,492,1171,1190]" pageId="1" pageNumber="486" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Head, pronotum, elytra and pygidium shiny reddish dark brown, sterna, and legs shiny reddish light brown. Clypeus bilobed, wider than long (4:1), anterior border widely sinuated with margins moderately elevated, surface slightly convex, densely rugo­punctate, with erect, very long, slender setae. Fronto­clypeal suture slightly sinuated and vaguely impressed, obscured by dense punctuation. Frons wider than long (2.1:1) convex, coasely rugo­punctate, with erect, very long, slender setae. Antenna 10 segmented, with 3 segmented club; lamellae slightly longer than length of five preceeding segments combined (1.2:1), segment 4
<emphasis id="0883EAC9FFEBFFCFFC97FAE49002FAF0" bold="true" box="[768,775,1340,1360]" pageId="1" pageNumber="486">°</emphasis>
as long as segment 3, segment 5
<emphasis id="0883EAC9FFEBFFCFFE96FA8C920DFAC8" bold="true" box="[257,264,1364,1384]" pageId="1" pageNumber="486">°</emphasis>
with rounded anterior tubercle, segments 6
<emphasis id="0883EAC9FFEBFFCFFD27FA8C91B2FAC8" bold="true" box="[688,695,1364,1384]" pageId="1" pageNumber="486">°</emphasis>
and 7
<emphasis id="0883EAC9FFEBFFCFFD60FA8C91FBFAC8" bold="true" box="[759,766,1364,1384]" pageId="1" pageNumber="486">°</emphasis>
wider than long, each one with conical or acute process directed forward. Frons 6.5
<emphasis id="0883EAC9FFEBFFCFFD65FAB69001FA21" bold="true" box="[754,772,1390,1409]" pageId="1" pageNumber="486">×</emphasis>
wider than each eye dorsal diameter. Canthus curved and rounded, with 79 setae. Labrum deeply bilobed, with curved slender setae on the borders. Mentum widely concave, polished, with lateral slender setae, anterior border widely cleft. Pronotum wider than long (1.8:1) and 1.9
<emphasis id="0883EAC9FFEBFFCFFBB3FA6E9733FA69" bold="true" box="[1060,1078,1462,1481]" pageId="1" pageNumber="486">×</emphasis>
wider than frons. Pronotal disk densely rugo­punctate, with many erect, very long, slen­ der setae; lateral borders widely angulated, marginal bead widely crenulate, with long, slender setae; anterior angles slightly obtuse, rounded, scarcely prominent; posterior angles obtuse, scarcely prominent. Scutellum 1.5
<emphasis id="0883EAC9FFEBFFCFFD2CF9CF91C8F98A" bold="true" box="[699,717,1559,1578]" pageId="1" pageNumber="486">×</emphasis>
wider than long, with more than 15 deep and shallow, small, setiferous punctures. Elytron 2.3
<emphasis id="0883EAC9FFEBFFCFFCB3F9E89033F9E3" bold="true" box="[804,822,1584,1603]" pageId="1" pageNumber="486">×</emphasis>
longer than wide, densely rugo­punctate, with many erect, long, slender setae uniformly distributed; epipleural bor­ der narrowed, extended along the complete margin, provided with a fringe of long setae; humeral calla rounded, prominent; apical calla rounded. Metathoracic wings completely developed. Propygidium shiny, finely punctate with scattered, short setae. Pygidium convex, shiny, markedly rugose, with erect, long, slender setae, mainly toward the basal border; apical margin with 20 slender setae; basal margin effaced at the middle. Pterosternon with long, dense, yellowish vestiture. Visible abdominal sternites 2
<emphasis id="0883EAC9FFE8FFCCFC5EF99F90D5F9FB" bold="true" box="[969,976,1607,1627]" pageId="2" pageNumber="487">°</emphasis>
to 4
<emphasis id="0883EAC9FFE8FFCCFB97F99F9702F9FB" bold="true" box="[1024,1031,1607,1627]" pageId="2" pageNumber="487">°</emphasis>
with wide and deep mesial sulcus; sternite 5
<emphasis id="0883EAC9FFE8FFCCFDFAF9879171F9D3" bold="true" box="[621,628,1631,1651]" pageId="2" pageNumber="487">°</emphasis>
convex, with scattered setiferous punctures and anterior border slightly furrowed; anal plate widely concave, finely granulose­punctate, anterior and posterior borders thickened, with scattered erect, long setae. Protibiae as long as protarsi, with two large teeth and one small tooth on external border, preapical spur long, straight, acute, slightly longer than 2
<emphasis id="0883EAC9FFE9FFCDFD2EFE8D91C5FEC9" bold="true" box="[697,704,341,361]" pageId="3" pageNumber="488">°</emphasis>
protarsomerus (1.2:1). Mesotibiae with very long, slender setae, and one oblique, short, setiferous, transverse carina; upper apical spur straight, narrow, as long as lower spur. Metatibiae shorter than metatarsi (1:1.2), with very long, slender setae, and one oblique, short setiferous transverse carina; apical spurs articulated with the border, upper spur lanceolate, slightly curved, with nearly rounded apex, nearly as long as 2
<emphasis id="0883EAC9FFE9FFCDFDDEFE159155FE41" bold="true" box="[585,592,461,481]" pageId="3" pageNumber="488">°</emphasis>
metatarsomerus, and 1.5 longer than lower spur. Tarsomeres semicylindrical, elongated, with enlarged apex, and many setae around the apex; mesotarsomeres 2
<emphasis id="0883EAC9FFE9FFCDFE4CFE2592E7FDB1" bold="true" box="[475,482,509,529]" pageId="3" pageNumber="488">°</emphasis>
to 5
<emphasis id="0883EAC9FFE9FFCDFD86FE25911DFDB1" bold="true" box="[529,536,509,529]" pageId="3" pageNumber="488">°</emphasis>
ventrally present one line of slender setae (
<figureCitation id="A2CC2A5EFFE9FFCDFC55FE2690FEFDB1" box="[962,1019,510,529]" captionStart="Figs" captionStartId="2.[272,315,1339,1358]" captionTargetBox="[256,1071,268,1318]" captionTargetId="figure-12@2.[254,1075,265,1318]" captionTargetPageId="2" captionText="Figs. 18. Phyllophaga sayloriana. 1) male mesotarsomeres, latero­ventral view; 2) male metatarsomeres, latero­ventral view; 3) tarsal claw, male; 4) genital capsule, lateral view; 5) paramera, distal view; 6) apex of paramera, distal­ventral view; 7) distal half of genital capsule, dorsal view; 8) ventral genital plates, female. Scale lines: Figs. 13 = 0.5 mm; Figs. 48 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102837" httpUri="https://zenodo.org/record/10102837/files/figure.png" pageId="3" pageNumber="488">Fig. 1</figureCitation>
); metatarsomeres 2
<emphasis id="0883EAC9FFE9FFCDFEE9FDCD9280FD89" bold="true" box="[382,389,533,553]" pageId="3" pageNumber="488">°</emphasis>
to 4
<emphasis id="0883EAC9FFE9FFCDFE26FDCD92BDFD89" bold="true" box="[433,440,533,553]" pageId="3" pageNumber="488">°</emphasis>
ventrally present one line of stout setae, nearly comb­like structure (
<figureCitation id="A2CC2A5EFFE9FFCDFF6AFDF6923CFDE1" box="[253,313,558,577]" captionStart="Figs" captionStartId="2.[272,315,1339,1358]" captionTargetBox="[256,1071,268,1318]" captionTargetId="figure-12@2.[254,1075,265,1318]" captionTargetPageId="2" captionText="Figs. 18. Phyllophaga sayloriana. 1) male mesotarsomeres, latero­ventral view; 2) male metatarsomeres, latero­ventral view; 3) tarsal claw, male; 4) genital capsule, lateral view; 5) paramera, distal view; 6) apex of paramera, distal­ventral view; 7) distal half of genital capsule, dorsal view; 8) ventral genital plates, female. Scale lines: Figs. 13 = 0.5 mm; Figs. 48 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102837" httpUri="https://zenodo.org/record/10102837/files/figure.png" pageId="3" pageNumber="488">Fig. 2</figureCitation>
). Tarsal claws symmetrical, similar on all legs, with median acute­angled tooth located near basal dilatation, but also near apical tooth (
<figureCitation id="A2CC2A5EFFE9FFCDFC89FD9E905FFDF9" box="[798,858,582,601]" captionStart="Figs" captionStartId="2.[272,315,1339,1358]" captionTargetBox="[256,1071,268,1318]" captionTargetId="figure-12@2.[254,1075,265,1318]" captionTargetPageId="2" captionText="Figs. 18. Phyllophaga sayloriana. 1) male mesotarsomeres, latero­ventral view; 2) male metatarsomeres, latero­ventral view; 3) tarsal claw, male; 4) genital capsule, lateral view; 5) paramera, distal view; 6) apex of paramera, distal­ventral view; 7) distal half of genital capsule, dorsal view; 8) ventral genital plates, female. Scale lines: Figs. 13 = 0.5 mm; Figs. 48 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102837" httpUri="https://zenodo.org/record/10102837/files/figure.png" pageId="3" pageNumber="488">Fig. 3</figureCitation>
). Genital capsule with long, narrowed, angled paramera, fused dorsally at their basis, apex angled, with preapical long projections ended in flattened, blade­like structures (
<figureCitation id="A2CC2A5EFFE9FFCDFCA9FDAE909CFD29" box="[830,921,630,649]" captionStart="Figs" captionStartId="2.[272,315,1339,1358]" captionTargetBox="[256,1071,268,1318]" captionTargetId="figure-12@2.[254,1075,265,1318]" captionTargetPageId="2" captionText="Figs. 18. Phyllophaga sayloriana. 1) male mesotarsomeres, latero­ventral view; 2) male metatarsomeres, latero­ventral view; 3) tarsal claw, male; 4) genital capsule, lateral view; 5) paramera, distal view; 6) apex of paramera, distal­ventral view; 7) distal half of genital capsule, dorsal view; 8) ventral genital plates, female. Scale lines: Figs. 13 = 0.5 mm; Figs. 48 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102837" httpUri="https://zenodo.org/record/10102837/files/figure.png" pageId="3" pageNumber="488">Figs. 47</figureCitation>
). Tectum convex with widely sinuated distal border. Aedeagus long, wide, with sclerotized tube­like support and preapical, dorsal, sclerotized plates and blades (
<figureCitation id="A2CC2A5EFFE9FFCDFC89FD7E9079FD19" box="[798,892,678,697]" captionStart="Figs" captionStartId="2.[272,315,1339,1358]" captionTargetBox="[256,1071,268,1318]" captionTargetId="figure-12@2.[254,1075,265,1318]" captionTargetPageId="2" captionText="Figs. 18. Phyllophaga sayloriana. 1) male mesotarsomeres, latero­ventral view; 2) male metatarsomeres, latero­ventral view; 3) tarsal claw, male; 4) genital capsule, lateral view; 5) paramera, distal view; 6) apex of paramera, distal­ventral view; 7) distal half of genital capsule, dorsal view; 8) ventral genital plates, female. Scale lines: Figs. 13 = 0.5 mm; Figs. 48 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102837" httpUri="https://zenodo.org/record/10102837/files/figure.png" pageId="3" pageNumber="488">Figs. 47</figureCitation>
). Length of genital capsule from the apex of parameres to the border of basal piece:
<quantity id="FD0F9B3EFFE9FFCDFCE1FD6690C6FD71" box="[886,963,702,721]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="488" unit="mm" value="3.3">3.3 mm</quantity>
. Total body length:
<quantity id="FD0F9B3EFFE9FFCDFEA9FD0E9293FD49" box="[318,406,726,745]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="3" pageNumber="488" unit="mm" value="13.2">13.2 mm</quantity>
. Humeral width:
<quantity id="FD0F9B3EFFE9FFCDFDABFD0E918DFD49" box="[572,648,726,745]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="488" unit="mm" value="5.3">5.3 mm</quantity>
.
</paragraph>
<caption id="6E886653FFE8FFCCFE87FAE39121FA0E" ID-DOI="http://doi.org/10.5281/zenodo.10102837" ID-Zenodo-Dep="10102837" httpUri="https://zenodo.org/record/10102837/files/figure.png" pageId="2" pageNumber="487" startId="2.[272,315,1339,1358]" targetBox="[256,1071,268,1318]" targetPageId="2">
<paragraph id="3A4836DBFFE8FFCCFE87FAE39121FA0E" blockId="2.[249,1081,1339,1454]" pageId="2" pageNumber="487">
<emphasis id="0883EAC9FFE8FFCCFE87FAE39275FAEE" bold="true" box="[272,368,1339,1358]" pageId="2" pageNumber="487">Figs. 18.</emphasis>
<emphasis id="0883EAC9FFE8FFCCFE10FAE3916AFAEE" box="[391,623,1339,1358]" italics="true" pageId="2" pageNumber="487">
<taxonomicName id="FDF74D58FFE8FFCCFE10FAE3916EFAEE" box="[391,619,1339,1358]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="487" phylum="Arthropoda" rank="species" species="sayloriana">Phyllophaga sayloriana</taxonomicName>
.
</emphasis>
<emphasis id="0883EAC9FFE8FFCCFDE1FAE3918CFAEE" bold="true" box="[630,649,1339,1358]" pageId="2" pageNumber="487">1)</emphasis>
male mesotarsomeres, latero­ventral view;
<emphasis id="0883EAC9FFE8FFCCFBB1FAE3973CFAEE" bold="true" box="[1062,1081,1339,1358]" pageId="2" pageNumber="487">2)</emphasis>
male metatarsomeres, latero­ventral view;
<emphasis id="0883EAC9FFE8FFCCFD1DFA8B9198FAC6" bold="true" box="[650,669,1363,1382]" pageId="2" pageNumber="487">3)</emphasis>
tarsal claw, male;
<emphasis id="0883EAC9FFE8FFCCFCD9FA8B9067FAC6" bold="true" box="[846,866,1363,1382]" pageId="2" pageNumber="487">4)</emphasis>
genital capsule, lateral view;
<emphasis id="0883EAC9FFE8FFCCFEA2FAB3924CFADE" bold="true" box="[309,329,1387,1406]" pageId="2" pageNumber="487">5)</emphasis>
paramera, distal view;
<emphasis id="0883EAC9FFE8FFCCFDB9FAB39144FADE" bold="true" box="[558,577,1387,1406]" pageId="2" pageNumber="487">6)</emphasis>
apex of paramera, distal­ventral view;
<emphasis id="0883EAC9FFE8FFCCFC28FAB390D7FADE" bold="true" box="[959,978,1387,1406]" pageId="2" pageNumber="487">7)</emphasis>
distal half of genital capsule, dorsal view;
<emphasis id="0883EAC9FFE8FFCCFDB9FA5B9144FA36" bold="true" box="[558,577,1411,1430]" pageId="2" pageNumber="487">8)</emphasis>
ventral genital plates, female. Scale lines: Figs. 13
<emphasis id="0883EAC9FFE8FFCCFF6EFA43920EFA0E" bold="true" box="[249,267,1435,1454]" pageId="2" pageNumber="487">=</emphasis>
0.5 mm; Figs. 48
<emphasis id="0883EAC9FFE8FFCCFE5BFA4392DBFA0E" bold="true" box="[460,478,1435,1454]" pageId="2" pageNumber="487">=</emphasis>
1 mm.
</paragraph>
</caption>
<paragraph id="3A4836DBFFE9FFCDFE87FCDE9065FB86" blockId="3.[246,1079,774,1652]" pageId="3" pageNumber="488">
<emphasis id="0883EAC9FFE9FFCDFE87FCDE9270FCBB" bold="true" box="[272,373,774,795]" pageId="3" pageNumber="488">
<typeStatus id="E54C8879FFE9FFCDFE87FCDE9275FCBB" box="[272,368,774,795]" pageId="3" pageNumber="488" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Similar to the male except as follows: antennal club 0.8 shorter than the five preceeding segments combined; pygidium with apical fifth concave, polished, without punctures or setae, apex slightly upturned; visible abdominal sternites 2
<emphasis id="0883EAC9FFE9FFCDFE48FC8E92E2FCCB" bold="true" box="[479,487,854,875]" pageId="3" pageNumber="488">°</emphasis>
to 4
<emphasis id="0883EAC9FFE9FFCDFDB7FC8E912DFCCB" bold="true" box="[544,552,854,875]" pageId="3" pageNumber="488">°</emphasis>
with mesial narrow, shallow sulcus; sternite 5
<emphasis id="0883EAC9FFE9FFCDFBB9FC8E9733FCCB" bold="true" box="[1070,1078,854,875]" pageId="3" pageNumber="488">°</emphasis>
coarsely rugo­punctate, with scattered short setae; anal plate convex, rugopunctate, with basal border slightly uplift, and lines of 1020 slender setae along basal and apical borders. Protibia with external border clearly tridentate. Apical spurs of metatibiae wide, slightly curvated with rounded apexes. Ventral genital plates ovate, strongly sclerotized, nearly symmetrical, convex, with some short setae; dorsal genital plates ovate, with borders scarcely setose (
<figureCitation id="A2CC2A5EFFE9FFCDFB9BFC2F9200FB86" captionStart="Figs" captionStartId="2.[272,315,1339,1358]" captionTargetBox="[256,1071,268,1318]" captionTargetId="figure-12@2.[254,1075,265,1318]" captionTargetPageId="2" captionText="Figs. 18. Phyllophaga sayloriana. 1) male mesotarsomeres, latero­ventral view; 2) male metatarsomeres, latero­ventral view; 3) tarsal claw, male; 4) genital capsule, lateral view; 5) paramera, distal view; 6) apex of paramera, distal­ventral view; 7) distal half of genital capsule, dorsal view; 8) ventral genital plates, female. Scale lines: Figs. 13 = 0.5 mm; Figs. 48 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10102837" httpUri="https://zenodo.org/record/10102837/files/figure.png" pageId="3" pageNumber="488">Fig. 8</figureCitation>
). Total body length:
<quantity id="FD0F9B3EFFE9FFCDFE7FFBC9914EFB86" box="[488,587,1041,1062]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="3" pageNumber="488" unit="mm" value="13.9">13.9 mm</quantity>
. Humeral width: 6.0 mm.
</paragraph>
<paragraph id="3A4836DBFFE9FFCDFE87FBF49255FB41" blockId="3.[246,1079,774,1652]" pageId="3" pageNumber="488">
<emphasis id="0883EAC9FFE9FFCDFE87FBF492EAFBE1" bold="true" box="[272,495,1068,1089]" pageId="3" pageNumber="488">
<typeStatus id="E54C8879FFE9FFCDFE87FBF49273FBE1" box="[272,374,1068,1089]" pageId="3" pageNumber="488" type="paratype">Paratype</typeStatus>
Variation.
</emphasis>
Males. Similar to the
<typeStatus id="E54C8879FFE9FFCDFD4CFBF49033FBE1" box="[731,822,1068,1089]" pageId="3" pageNumber="488" type="holotype">holotype</typeStatus>
except as follows: body color dark reddish brown to light reddish brown; abdominal sternites with mesial furrow less deep; total body length:
<quantity id="FD0F9B3EFFE9FFCDFD2AFBB99052FBD6" box="[701,855,1121,1142]" metricMagnitude="-2" metricUnit="m" metricValue="1.265" metricValueMax="1.3699999999999999" metricValueMin="1.16" pageId="3" pageNumber="488" unit="mm" value="12.649999999999999" valueMax="13.7" valueMin="11.6">11.613.7 mm</quantity>
, humeral width:
<quantity id="FD0F9B3EFFE9FFCDFB9FFBB99249FB31" metricMagnitude="-3" metricUnit="m" metricValue="5.199999999999999" metricValueMax="5.6" metricValueMin="4.8" pageId="3" pageNumber="488" unit="mm" value="5.199999999999999" valueMax="5.6" valueMin="4.8">4.8 5.6 mm</quantity>
. Female
<typeStatus id="E54C8879FFE9FFCDFE39FBA49117FB31" box="[430,530,1148,1169]" pageId="3" pageNumber="488" type="paratype">paratypes</typeStatus>
are similar to the
<typeStatus id="E54C8879FFE9FFCDFD4CFBA4902AFB31" box="[731,815,1148,1169]" pageId="3" pageNumber="488" type="allotype">allotype</typeStatus>
except as follows: body color dark reddish brown to dark yellowish brown; pygidial preapical fifth more or less concave; total body length:
<quantity id="FD0F9B3EFFE9FFCDFD26FB69904AFB66" box="[689,847,1201,1222]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" metricValueMax="1.49" metricValueMin="1.35" pageId="3" pageNumber="488" unit="mm" value="14.2" valueMax="14.9" valueMin="13.5">13.514.9 mm</quantity>
; humeral width:
<quantity id="FD0F9B3EFFE9FFCDFB90FB699249FB41" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.2" metricValueMin="5.9" pageId="3" pageNumber="488" unit="mm" value="6.05" valueMax="6.2" valueMin="5.9">5.9 6.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE9FFCAFE87FB3E9724FD8E" lastPageId="4" lastPageNumber="489" pageId="3" pageNumber="488" type="materials_examined">
<paragraph id="3A4836DBFFE9FFCAFE87FB3E903AFE59" blockId="3.[246,1079,774,1652]" lastBlockId="4.[249,1082,269,1284]" lastPageId="4" lastPageNumber="489" pageId="3" pageNumber="488">
<emphasis id="0883EAC9FFE9FFCDFE87FB3E929AFB5B" bold="true" box="[272,415,1254,1275]" pageId="3" pageNumber="488">Type Series.</emphasis>
(Described from
<specimenCount id="2CF1FD52FFE9FFCDFDFDFB3F91CBFB5C" box="[618,718,1255,1276]" pageId="3" pageNumber="488" type="male">50 males</specimenCount>
and
<specimenCount id="2CF1FD52FFE9FFCDFC9AFB3F908FFB5C" box="[781,906,1255,1276]" pageId="3" pageNumber="488" type="female">92 females</specimenCount>
).
<materialsCitation id="8A9F3C86FFE9FFCDFC36FB3F91B3FA91" collectingDate="1997-08-07" collectingMethod="trampa luz" collectionCode="MXAL" collectorName="M. Lopez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Est. Las Joyas" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="holotype">
<typeStatus id="E54C8879FFE9FFCDFC36FB3F9707FB5C" box="[929,1026,1255,1276]" pageId="3" pageNumber="488" type="holotype">Holotype</typeStatus>
<specimenCount id="2CF1FD52FFE9FFCDFB99FB3D9727FB5B" box="[1038,1058,1253,1275]" count="1" pageId="3" pageNumber="488" type="male"></specimenCount>
;
<collectingCountry id="42E0764BFFE9FFCDFF61FAD99264FAB6" box="[246,353,1281,1302]" name="Mexico" pageId="3" pageNumber="488">MEXICO</collectingCountry>
:
<emphasis id="0883EAC9FFE9FFCDFEF9FADA92BBFAB7" box="[366,446,1282,1303]" italics="true" pageId="3" pageNumber="488">
<collectingRegion id="F833F839FFE9FFCDFEF9FADA92BEFAB7" box="[366,443,1282,1303]" country="Mexico" name="Jalisco" pageId="3" pageNumber="488">Jalisco</collectingRegion>
,
</emphasis>
<collectingCounty id="D3294E57FFE9FFCDFE5FFAD99117FAB6" box="[456,530,1281,1302]" pageId="3" pageNumber="488">Autlán</collectingCounty>
,
<location id="3F286000FFE9FFCDFD88FAD991B9FAB6" LSID="urn:lsid:plazi:treatment:B25E87CDFFEBFFCAFF60FB939703FAA4:3F286000FFE9FFCDFD88FAD991B9FAB6" box="[543,700,1281,1302]" country="Mexico" county="Autlan" municipality="Est. Las Joyas" name="Est. Las Joyas" pageId="3" pageNumber="488" stateProvince="Jalisco">Est. Las Joyas</location>
, bosque mesófilo, Alt.
<quantity id="FD0F9B3EFFE9FFCDFC2FFAD99716FAB6" box="[952,1043,1281,1302]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="488" unit="m" value="1900.0">
<elevation id="B1DAD1E8FFE9FFCDFC2FFAD99716FAB6" box="[952,1043,1281,1302]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="488" unit="m" value="1900.0">1,900 m</elevation>
</quantity>
,
<date id="4E49101BFFE9FFCDFBB7FAD99266FA91" pageId="3" pageNumber="488" value="1997-08-07">
<collectingDate id="5E0DE9F3FFE9FFCDFBB7FAD99266FA91" pageId="3" pageNumber="488" value="1997-08-07">7­ VIII­1997</collectingDate>
</date>
,
<collectingMethod id="E3B64ECCFFE9FFCDFEFAFAC492DBFA91" box="[365,478,1308,1329]" pageId="3" pageNumber="488">trampa luz</collectingMethod>
,
<collectorName id="9702530DFFE9FFCDFE7EFAC4914BFA91" box="[489,590,1308,1329]" pageId="3" pageNumber="488">M. López</collectorName>
(
<collectionCode id="5CE6AE1EFFE9FFCDFDCBFAC491AAFA91" box="[604,687,1308,1329]" pageId="3" pageNumber="488">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFE9FFCDFD56FAC49122FAEC" collectingDate="1997-08-07" collectingMethod="trampa luz" collectionCode="MXAL" collectorName="A. Vazquez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="allotype">
<typeStatus id="E54C8879FFE9FFCDFD56FAC49019FA91" box="[705,796,1308,1329]" pageId="3" pageNumber="488" type="allotype">Allotype</typeStatus>
<specimenCount id="2CF1FD52FFE9FFCDFCB5FAC39033FA91" box="[802,822,1307,1329]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
; same data as holotype, except
<collectorName id="9702530DFFE9FFCDFED5FAEF92BBFAEC" box="[322,446,1335,1356]" pageId="3" pageNumber="488">A. Vázquez</collectorName>
(
<collectionCode id="5CE6AE1EFFE9FFCDFE5AFAEF9125FAEC" box="[461,544,1335,1356]" pageId="3" pageNumber="488">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFE9FFCDFDA5FAEF9283FAC6" collectingDate="1997-08-07" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Vazquez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<typeStatus id="E54C8879FFE9FFCDFDA5FAEF9199FAEC" box="[562,668,1335,1356]" pageId="3" pageNumber="488" type="paratype">Paratypes</typeStatus>
: same data as holotype except
<collectorName id="9702530DFFE9FFCDFC74FAEF9222FAC6" pageId="3" pageNumber="488">A. Vázquez</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFEAFFA899273FAC6" box="[312,374,1360,1382]" count="2" pageId="3" pageNumber="488" type="male">2 ♂♂</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFE19FA8990C2FAC6" box="[398,967,1360,1382]" collectingDate="1997-07-03" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
same data except
<date id="4E49101BFFE9FFCDFDDAFA8991C3FAC6" box="[589,710,1361,1382]" pageId="3" pageNumber="488" value="1997-07-03">
<collectingDate id="5E0DE9F3FFE9FFCDFDDAFA8991C3FAC6" box="[589,710,1361,1382]" pageId="3" pageNumber="488" value="1997-07-03">3­VII­1997</collectingDate>
</date>
,
<collectorName id="9702530DFFE9FFCDFD45FA899036FAC6" box="[722,819,1361,1382]" pageId="3" pageNumber="488">J. García</collectorName>
, (
<specimenCount id="2CF1FD52FFE9FFCDFCDDFA8990B2FAC6" box="[842,951,1360,1382]" count="1" pageId="3" pageNumber="488" type="female">1 ♂ 2 ♀♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFC58FA899293FA21" collectingDate="1997-08-07" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="8" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFC58FA899733FAC6" box="[975,1078,1361,1382]" pageId="3" pageNumber="488">M. López</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFF69FAB49238FA21" box="[254,317,1387,1409]" pageId="3" pageNumber="488">3 ♂♂</specimenCount>
<specimenCount id="2CF1FD52FFE9FFCDFED0FAB49283FA21" box="[327,390,1387,1409]" pageId="3" pageNumber="488">5 ♀♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFE37FAB4901BFA21" box="[416,798,1387,1409]" collectingDate="1997-07-04" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="4" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFE37FAB4911FFA21" box="[416,538,1388,1409]" pageId="3" pageNumber="488" value="1997-07-04">
<collectingDate id="5E0DE9F3FFE9FFCDFE37FAB4911FFA21" box="[416,538,1388,1409]" pageId="3" pageNumber="488" value="1997-07-04">4­VII­1997</collectingDate>
</date>
,
<collectorName id="9702530DFFE9FFCDFDB0FAB4918DFA21" box="[551,648,1388,1409]" pageId="3" pageNumber="488">J. García</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFD0DFAB491DCFA21" box="[666,729,1387,1409]" pageId="3" pageNumber="488">3 ♂♂</specimenCount>
<specimenCount id="2CF1FD52FFE9FFCDFD74FAB4900BFA21" box="[739,782,1387,1409]" pageId="3" pageNumber="488">1 ♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFCBFFAB49248FA3C" collectingDate="1997-08-05" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Vazquez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-female="2" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFCBFFAB490AFFA21" box="[808,938,1388,1409]" pageId="3" pageNumber="488" value="1997-08-05">
<collectingDate id="5E0DE9F3FFE9FFCDFCBFFAB490AFFA21" box="[808,938,1388,1409]" pageId="3" pageNumber="488" value="1997-08-05">5­VIII­1997</collectingDate>
</date>
,
<collectorName id="9702530DFFE9FFCDFC2FFAB49733FA21" box="[952,1078,1388,1409]" pageId="3" pageNumber="488">A. Vázquez</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFF69FA5F9238FA3C" box="[254,317,1414,1436]" count="2" pageId="3" pageNumber="488" type="female">2 ♀♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFEC1FA5F9109FA3C" box="[342,524,1414,1436]" collectingDate="1997-08-06" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Vazquez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFEC1FA5F92D0FA3C" box="[342,469,1415,1436]" pageId="3" pageNumber="488" value="1997-08-06">
<collectingDate id="5E0DE9F3FFE9FFCDFEC1FA5F92D0FA3C" box="[342,469,1415,1436]" pageId="3" pageNumber="488" value="1997-08-06">6­VIII­1997</collectingDate>
</date>
(
<specimenCount id="2CF1FD52FFE9FFCDFE7FFA5E92F9FA3C" box="[488,508,1414,1436]" count="1" pageId="3" pageNumber="488" type="male"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFD82FA5F91DBFA3C" box="[533,734,1414,1436]" collectingDate="1997-08-06" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="4" specimenCount-female="4" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFD82FA5F9178FA3C" box="[533,637,1415,1436]" pageId="3" pageNumber="488">M. López</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFD18FA5F91CBFA3C" box="[655,718,1414,1436]" count="4" pageId="3" pageNumber="488" type="female">4 ♀♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFD70FA5F90A2FA3C" box="[743,935,1414,1436]" collectingDate="1997-08-06" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-female="2" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFD70FA5F9042FA3C" box="[743,839,1415,1436]" pageId="3" pageNumber="488">J. García</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFCCFFA5F9092FA3C" box="[856,919,1414,1436]" count="2" pageId="3" pageNumber="488" type="female">2 ♀♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFC27FA5F92BFFA17" collectingDate="1997-08-07" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="S. Nunez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-female="2" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFC27FA5F9737FA3C" box="[944,1074,1415,1436]" pageId="3" pageNumber="488" value="1997-08-07">
<collectingDate id="5E0DE9F3FFE9FFCDFC27FA5F9737FA3C" box="[944,1074,1415,1436]" pageId="3" pageNumber="488" value="1997-08-07">7­VIII­1997</collectingDate>
</date>
,
<collectorName id="9702530DFFE9FFCDFF61FA7A925CFA17" box="[246,345,1442,1463]" pageId="3" pageNumber="488">S. Núñez</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFEFDFA7A92AFFA17" box="[362,426,1441,1463]" count="2" pageId="3" pageNumber="488" type="male">2 ♂♂</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFE54FA7A91A6FA17" box="[451,675,1441,1463]" collectingDate="1997-08-07" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Vazquez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-female="2" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFE54FA7A9144FA17" box="[451,577,1442,1463]" pageId="3" pageNumber="488">A. Vázquez</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFDC4FA7A9197FA17" box="[595,658,1441,1463]" count="2" pageId="3" pageNumber="488" type="female">2 ♀♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFD3BFA7A9086FA17" box="[684,899,1441,1463]" collectingDate="1997-08-07" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="11" specimenCount-female="11" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFD3BFA7A9011FA17" box="[684,788,1442,1463]" pageId="3" pageNumber="488">M. López</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFCB0FA7A9076FA17" box="[807,883,1441,1463]" count="11" pageId="3" pageNumber="488" type="female">11 ♀♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFC1BFA7A9243FA72" collectingDate="1997-08-07" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Partida" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFC1BFA7A90FCFA17" box="[908,1017,1442,1463]" pageId="3" pageNumber="488">A. Partida</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFB9AFA7A9233FA72" count="1" pageId="3" pageNumber="488" type="female">1 ♂ 2 ♀♀</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFEC6FA65916CFA72" box="[337,617,1468,1490]" collectingDate="1997-09-05" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-female="2" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFEC6FA6592B6FA72" box="[337,435,1469,1490]" pageId="3" pageNumber="488">J. García</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFE51FA6492DFFA72" box="[454,474,1468,1490]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
<date id="4E49101BFFE9FFCDFE62FA659163FA72" box="[501,614,1469,1490]" pageId="3" pageNumber="488" value="1997-09-05">
<collectingDate id="5E0DE9F3FFE9FFCDFE62FA659163FA72" box="[501,614,1469,1490]" pageId="3" pageNumber="488" value="1997-09-05">5­IX­1997</collectingDate>
</date>
,
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFDE2FA65901CFA72" box="[629,793,1468,1490]" collectingDate="1997-09-05" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Guerra" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFDE2FA6591E7FA72" box="[629,738,1469,1490]" pageId="3" pageNumber="488">A. Guerra</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFD62FA64900CFA72" box="[757,777,1468,1490]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFCB3FA6590C3FA72" box="[804,966,1468,1490]" collectingDate="1997-09-05" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFCB3FA65908BFA72" box="[804,910,1469,1490]" pageId="3" pageNumber="488">M. López</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFC35FA6490B3FA72" box="[930,950,1468,1490]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFC47FA65923BFA4D" collectingDate="1997-10-01" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Vazquez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFC47FA659733FA72" box="[976,1078,1469,1490]" pageId="3" pageNumber="488" value="1997-10-01">
<collectingDate id="5E0DE9F3FFE9FFCDFC47FA659733FA72" box="[976,1078,1469,1490]" pageId="3" pageNumber="488" value="1997-10-01">1­X­1997</collectingDate>
</date>
(
<specimenCount id="2CF1FD52FFE9FFCDFF68FA0F9216FA4D" box="[255,275,1495,1517]" count="1" pageId="3" pageNumber="488" type="male"></specimenCount>
<emphasis id="0883EAC9FFE9FFCDFE84FA00921FFA4D" bold="true" box="[275,282,1496,1517]" pageId="3" pageNumber="488">/</emphasis>
<specimenCount id="2CF1FD52FFE9FFCDFE8DFA0F922BFA4D" box="[282,302,1495,1517]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFEDCFA0091D5FA4D" box="[331,720,1495,1517]" collectingDate="1999-07-30" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="V. Gonzalez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-female="1" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFEDCFA0092D4FA4D" box="[331,465,1496,1517]" pageId="3" pageNumber="488" value="1999-07-30">
<collectingDate id="5E0DE9F3FFE9FFCDFEDCFA0092D4FA4D" box="[331,465,1496,1517]" pageId="3" pageNumber="488" value="1999-07-30">30­VII­1999</collectingDate>
</date>
,
<collectorName id="9702530DFFE9FFCDFE75FA00916DFA4D" box="[482,616,1496,1517]" pageId="3" pageNumber="488">V. González</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFDEBFA0091BAFA4D" box="[636,703,1495,1517]" count="2" pageId="3" pageNumber="488" type="male">2 ♂♂</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFD4BFA009070FA4D" box="[732,885,1495,1517]" collectingDate="1999-07-30" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="G. Mesa" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFD4BFA00903EFA4D" box="[732,827,1496,1517]" pageId="3" pageNumber="488">G. Mesa</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFCC7FA0F9061FA4D" box="[848,868,1495,1517]" count="1" pageId="3" pageNumber="488" type="male"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFC16FA00928CF9A8" collectingDate="1999-07-31" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFC16FA009702FA4D" box="[897,1031,1496,1517]" pageId="3" pageNumber="488" value="1999-07-31">
<collectingDate id="5E0DE9F3FFE9FFCDFC16FA009702FA4D" box="[897,1031,1496,1517]" pageId="3" pageNumber="488" value="1999-07-31">31­VII­1999</collectingDate>
</date>
,
<collectorName id="9702530DFFE9FFCDFB8FFA009232F9A8" pageId="3" pageNumber="488">M. López</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFEDEFA2A9258F9A8" box="[329,349,1522,1544]" count="1" pageId="3" pageNumber="488" type="male"></specimenCount>
<emphasis id="0883EAC9FFE9FFCDFECAFA2B9261F9A8" bold="true" box="[349,356,1523,1544]" pageId="3" pageNumber="488">/</emphasis>
<specimenCount id="2CF1FD52FFE9FFCDFEF2FA2A927CF9A8" box="[357,377,1522,1544]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFE05FA2B912DF9A8" box="[402,552,1522,1544]" collectingDate="1997-09-05" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFE05FA2B92F7F9A8" box="[402,498,1523,1544]" pageId="3" pageNumber="488">J. García</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFD93FA2A911DF9A8" box="[516,536,1522,1544]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFDA6FA2B91EFF9A8" box="[561,746,1522,1544]" collectingDate="1997-09-05" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="V. Gonzalez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
<collectorName id="9702530DFFE9FFCDFDA6FA2B91B1F9A8" box="[561,692,1523,1544]" pageId="3" pageNumber="488">V. González</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFD52FA2A91DCF9A8" box="[709,729,1522,1544]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFD65FA2B914BF983" collectingDate="1985-11-21" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="G. Parada" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
same data except
<date id="4E49101BFFE9FFCDFC23FA2B9737F9A8" box="[948,1074,1523,1544]" pageId="3" pageNumber="488" value="1985-11-21">
<collectingDate id="5E0DE9F3FFE9FFCDFC23FA2B9737F9A8" box="[948,1074,1523,1544]" pageId="3" pageNumber="488" value="1985-11-21">21­XI­1985</collectingDate>
</date>
,
<collectorName id="9702530DFFE9FFCDFF61F9D69266F983" box="[246,355,1550,1571]" pageId="3" pageNumber="488">G. Parada</collectorName>
, veg. secundaria (
<specimenCount id="2CF1FD52FFE9FFCDFDBDF9D5913BF983" box="[554,574,1549,1571]" count="1" pageId="3" pageNumber="488" type="male"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFDCFF9D69733F983" box="[600,1078,1549,1571]" collectingDate="1986-06-17" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="L. Rivera" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFDCFF9D691D0F983" box="[600,725,1550,1571]" pageId="3" pageNumber="488" value="1986-06-17">
<collectingDate id="5E0DE9F3FFE9FFCDFDCFF9D691D0F983" box="[600,725,1550,1571]" pageId="3" pageNumber="488" value="1986-06-17">17­VI­1986</collectingDate>
</date>
,
<collectorName id="9702530DFFE9FFCDFD74F9D69049F983" box="[739,844,1550,1571]" pageId="3" pageNumber="488">L. Rivera</collectorName>
, veg. secundaria (
<specimenCount id="2CF1FD52FFE9FFCDFB85F9D59723F983" box="[1042,1062,1549,1571]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFF61F9F192DDF99E" box="[246,472,1576,1598]" collectingDate="1986-06-19" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Vazquez" country="Mexico" county="Autlan" location="Alt" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="2" specimenCount-male="2" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFF61F9F19271F99E" box="[246,372,1577,1598]" pageId="3" pageNumber="488" value="1986-06-19">
<collectingDate id="5E0DE9F3FFE9FFCDFF61F9F19271F99E" box="[246,372,1577,1598]" pageId="3" pageNumber="488" value="1986-06-19">19­VI­1986</collectingDate>
</date>
, (
<specimenCount id="2CF1FD52FFE9FFCDFE1EF9F192CDF99E" box="[393,456,1576,1598]" count="2" pageId="3" pageNumber="488" type="male">2 ♂♂</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFE76F9F19011F99E" box="[481,788,1576,1598]" collectingDate="1985-07" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="A. Vazquez" country="Mexico" county="Autlan" elevation="1900" latitude="19.586945" location="Alt" longitude="-104.28001" municipality="Est. Las Joyas" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFE76F9F19140F99E" box="[481,581,1577,1598]" pageId="3" pageNumber="488" value="1985-07">
<collectingDate id="5E0DE9F3FFE9FFCDFE76F9F19140F99E" box="[481,581,1577,1598]" pageId="3" pageNumber="488" value="1985-07">VII­1985</collectingDate>
</date>
, bosque pinos, (
<specimenCount id="2CF1FD52FFE9FFCDFD61F9F0900FF99E" box="[758,778,1576,1598]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFE9FFCDFCB4F9F190DFF99E" box="[803,986,1576,1598]" collectingDate="1987-09-10" collectingMethod="trampa luz" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" country="Mexico" county="Autlan" elevation="2300" latitude="19.586945" location="El Terrero" longitude="-104.28001" municipality="Minatitlan" pageId="3" pageNumber="488" specimenCount="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFCB4F9F190A5F99E" box="[803,928,1577,1598]" pageId="3" pageNumber="488" value="1987-09-10">
<collectingDate id="5E0DE9F3FFE9FFCDFCB4F9F190A5F99E" box="[803,928,1577,1598]" pageId="3" pageNumber="488" value="1987-09-10">10­IX­1987</collectingDate>
</date>
, (
<specimenCount id="2CF1FD52FFE9FFCDFC20F9F090CEF99E" box="[951,971,1576,1598]" count="1" pageId="3" pageNumber="488" type="male"></specimenCount>
).
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFC74F9F192B5F9D4" collectingDate="1987-09-10" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" elevation="2300" location="El Terrero" municipality="Minatitlan" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<emphasis id="0883EAC9FFE9FFCDFC74F9F19733F99E" box="[995,1078,1577,1598]" italics="true" pageId="3" pageNumber="488">
<collectingRegion id="F833F839FFE9FFCDFC74F9F19737F99E" box="[995,1074,1577,1598]" country="Mexico" name="Colima" pageId="3" pageNumber="488">Colima</collectingRegion>
,
</emphasis>
<collectingMunicipality id="DA2CACA1FFE9FFCDFF61F99C9263F9F9" box="[246,358,1604,1625]" pageId="3" pageNumber="488">Minatitlán</collectingMunicipality>
,
<location id="3F286000FFE9FFCDFEE6F99C92DAF9F9" LSID="urn:lsid:plazi:treatment:B25E87CDFFEBFFCAFF60FB939703FAA4:3F286000FFE9FFCDFEE6F99C92DAF9F9" box="[369,479,1604,1625]" country="Mexico" municipality="Minatitlan" name="El Terrero" pageId="3" pageNumber="488" stateProvince="Colima">El Terrero</location>
, bosque mesófilo, Alt.
<quantity id="FD0F9B3EFFE9FFCDFD44F99C9029F9F9" box="[723,812,1604,1625]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="488" unit="m" value="2300.0">
<elevation id="B1DAD1E8FFE9FFCDFD44F99C9029F9F9" box="[723,812,1604,1625]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="488" unit="m" value="2300.0">2,300 m</elevation>
</quantity>
,
<date id="4E49101BFFE9FFCDFCA0F99C90B0F9F9" box="[823,949,1604,1625]" pageId="3" pageNumber="488" value="1998-06-25">
<collectingDate id="5E0DE9F3FFE9FFCDFCA0F99C90B0F9F9" box="[823,949,1604,1625]" pageId="3" pageNumber="488" value="1998-06-25">25­VI­1998</collectingDate>
</date>
, trampa luz,
<collectorName id="9702530DFFE9FFCDFF61F9879251F9D4" box="[246,340,1631,1652]" pageId="3" pageNumber="488">J. García</collectorName>
(
<specimenCount id="2CF1FD52FFE9FFCDFEF3F98792A5F9D3" box="[356,416,1629,1652]" count="3" pageId="3" pageNumber="488" type="male">3 ♂♂</specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCDFE20F98790ABF9D4" box="[439,942,1629,1652]" collectingDate="1998-06-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="C. Hernandez" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="3" pageNumber="488" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
same data except
<collectorName id="9702530DFFE9FFCDFDE5F9879000F9D4" box="[626,773,1631,1652]" pageId="3" pageNumber="488">C. Hernández</collectorName>
, trampa uv (
<specimenCount id="2CF1FD52FFE9FFCDFC1EF9859098F9D3" box="[905,925,1629,1651]" count="1" pageId="3" pageNumber="488" type="female"></specimenCount>
);
</materialsCitation>
<materialsCitation id="8A9F3C86FFE9FFCAFC23F987928DFE84" collectingDate="1998-06-26" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="C. Hernandez" country="Mexico" elevation="2300" lastPageId="4" lastPageNumber="489" location="Colima" municipality="Minatitlan" pageId="3" pageNumber="488" specimenCount="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFE9FFCDFC23F9879737F9D4" box="[948,1074,1631,1652]" pageId="3" pageNumber="488" value="1998-06-26">
<collectingDate id="5E0DE9F3FFE9FFCDFC23F9879737F9D4" box="[948,1074,1631,1652]" pageId="3" pageNumber="488" value="1998-06-26">26­VI­1998</collectingDate>
</date>
, luz fluor. (
<specimenCount id="2CF1FD52FFEEFFCAFEFDFED5927BFE83" box="[362,382,269,291]" count="1" pageId="4" pageNumber="489" type="male"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFE00FED79032FE84" box="[407,823,269,292]" collectingDate="1998-06-26" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="C. Hernandez" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFE00FED79113FE84" box="[407,534,271,292]" pageId="4" pageNumber="489" value="1998-06-27">27­VI­1998</date>
, M. López, trampa uv (
<specimenCount id="2CF1FD52FFEEFFCAFC8EFED59028FE83" box="[793,813,269,291]" count="1" pageId="4" pageNumber="489" type="female"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFCD1FED79296FE9E" collectingDate="1998-07-23" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="C. Hernandez" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="16" specimenCount-female="7" specimenCount-male="9" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFCD1FED790C9FE84" box="[838,972,271,292]" pageId="4" pageNumber="489" value="1998-07-23">23­VII­1998</date>
, luz fluor. (
<specimenCount id="2CF1FD52FFEEFFCAFE96FEF1928CFE9E" box="[257,393,296,318]" count="9" pageId="4" pageNumber="489" type="female">9 ♂♂ 7 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFE35FEF191CAFE9E" box="[418,719,296,318]" collectingDate="1998-07-24" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFE35FEF1912CFE9E" box="[418,553,297,318]" pageId="4" pageNumber="489" value="1998-07-24">24­VII­1998</date>
, M. López (
<specimenCount id="2CF1FD52FFEEFFCAFD26FEF091C0FE9E" box="[689,709,296,318]" count="1" pageId="4" pageNumber="489" type="female"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFD49FEF1922EFEF9" collectingDate="1998-07-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="11" specimenCount-female="3" specimenCount-male="8" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFD49FEF19061FE9E" box="[734,868,297,318]" pageId="4" pageNumber="489" value="1998-07-25">25­VII­1998</date>
, J. García (
<specimenCount id="2CF1FD52FFEEFFCAFC73FEF19224FEF9" count="8" pageId="4" pageNumber="489" type="female">8 ♂♂ 3 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFEABFE9C9100FEF9" box="[316,517,323,345]" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="2" specimenCount-male="2" stateProvince="Colima" typeStatus="paratype">
M. López (
<specimenCount id="2CF1FD52FFEEFFCAFE2DFE9C92FEFEF9" box="[442,507,323,345]" count="2" pageId="4" pageNumber="489" type="male">2 ♂♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFD81FE9C90D5FEF9" box="[534,976,323,345]" collectingDate="1998-08-20" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFD81FE9C91A0FEF9" box="[534,677,324,345]" pageId="4" pageNumber="489" value="1998-08-20">20­VIII­1998</date>
, M. López, trampa uv (
<specimenCount id="2CF1FD52FFEEFFCAFC25FE9B90C3FEF9" box="[946,966,323,345]" count="1" pageId="4" pageNumber="489" type="male"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFC76FE9C92FFFED4" collectingDate="1998-08-21" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="3" specimenCount-female="3" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFC76FE9C9235FED4" pageId="4" pageNumber="489" value="1998-08-21">21­VIII­ 1998</date>
, J. García (
<specimenCount id="2CF1FD52FFEEFFCAFE26FE8792F5FED3" box="[433,496,349,372]" count="3" pageId="4" pageNumber="489" type="female">3 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFD9DFE8791FFFED4" box="[522,762,349,372]" collectingDate="1998-07-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
luz fluor. (
<specimenCount id="2CF1FD52FFEEFFCAFDEAFE8791F5FED3" box="[637,752,349,372]" count="1" pageId="4" pageNumber="489" type="female">1 ♂ 5 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFC9EFE879736FED4" box="[777,1075,349,372]" collectingDate="1998-08-22" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="17" specimenCount-female="16" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFC9EFE879093FED4" box="[777,918,351,372]" pageId="4" pageNumber="489" value="1998-08-22">22­VIII­1998</date>
(
<specimenCount id="2CF1FD52FFEEFFCAFC3DFE87972CFED3" box="[938,1065,349,372]" count="1" pageId="4" pageNumber="489" type="female">1 ♂ 16 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFF6EFEA192C0FE2E" box="[249,453,376,398]" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="6" specimenCount-female="6" stateProvince="Colima" typeStatus="paratype">
trampa uv (
<specimenCount id="2CF1FD52FFEEFFCAFEEDFEA192BEFE2E" box="[378,443,376,398]" count="6" pageId="4" pageNumber="489" type="female">6 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFE41FEA19029FE2E" box="[470,812,376,398]" collectingDate="1998-08-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFE41FEA19160FE2E" box="[470,613,377,398]" pageId="4" pageNumber="489" value="1998-08-25">25­VIII­1998</date>
, M. López (
<specimenCount id="2CF1FD52FFEEFFCAFD64FEA09002FE2E" box="[755,775,376,398]" count="1" pageId="4" pageNumber="489" type="male"></specimenCount>
<emphasis id="0883EAC9FFEEFFCAFC90FEA1900BFE2E" bold="true" box="[775,782,377,398]" pageId="4" pageNumber="489">/</emphasis>
<specimenCount id="2CF1FD52FFEEFFCAFC99FEA09027FE2E" box="[782,802,376,398]" count="1" pageId="4" pageNumber="489" type="female"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFCAAFEA190D5FE2E" box="[829,976,376,398]" collectingDate="1998-08-21" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
J. García (
<specimenCount id="2CF1FD52FFEEFFCAFC25FEA090C3FE2E" box="[946,966,376,398]" count="1" pageId="4" pageNumber="489" type="male"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFC76FEA192D9FE09" collectingDate="1998-08-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="M. Lopez" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="1" specimenCount-female="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFC76FEA19235FE09" pageId="4" pageNumber="489" value="1998-08-27">27­VIII­ 1998</date>
, M. López (
<specimenCount id="2CF1FD52FFEEFFCAFE29FE4B92D7FE09" box="[446,466,403,425]" count="1" pageId="4" pageNumber="489" type="female"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFE7AFE4C90A8FE09" box="[493,941,403,425]" collectingDate="1998-08-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" collectorName="J. Garcia" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="4" specimenCount-female="4" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFE7AFE4C916FFE09" box="[493,618,404,425]" pageId="4" pageNumber="489" value="1998-09-24">24­IX­1998</date>
, J. García, luz fluor. (
<specimenCount id="2CF1FD52FFEEFFCAFCF5FE4C90A6FE09" box="[866,931,403,425]" count="4" pageId="4" pageNumber="489" type="female">4 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFC29FE4C9225FE64" collectingDate="1998-08-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="1" specimenCount-female="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFC29FE4C973FFE09" box="[958,1082,404,425]" pageId="4" pageNumber="489" value="1998-09-25">25­IX­1998</date>
(
<specimenCount id="2CF1FD52FFEEFFCAFE95FE759213FE63" box="[258,278,429,451]" count="1" pageId="4" pageNumber="489" type="female"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFEB8FE779149FE64" box="[303,588,429,452]" collectingDate="1998-08-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="1" specimenCount-female="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFEB8FE7792A1FE64" box="[303,420,431,452]" pageId="4" pageNumber="489" value="1998-10-24">24­X­1998</date>
, trampa uv (
<specimenCount id="2CF1FD52FFEEFFCAFDB9FE759147FE63" box="[558,578,429,451]" count="1" pageId="4" pageNumber="489" type="female"></specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFEEFFCAFDCCFE77907CFE64" box="[603,889,429,452]" collectingDate="1998-08-25" collectionCode="AAHC, AMNH, CASC, CNC, FMNH, IBUNAM, IEXA, IMEJ, GNPC, MNHN, MXAL, NHMC, UNSM, ZMHU" country="Mexico" elevation="2300" location="Colima" municipality="Minatitlan" pageId="4" pageNumber="489" specimenCount="1" specimenCount-male="1" stateProvince="Colima" typeStatus="paratype">
<date id="4E49101BFFEEFFCAFDCCFE7791DDFE64" box="[603,728,431,452]" pageId="4" pageNumber="489" value="1998-11-24">24­XI­1998</date>
, luz fluor. (
<specimenCount id="2CF1FD52FFEEFFCAFCC2FE75906CFE63" box="[853,873,429,451]" count="1" pageId="4" pageNumber="489" type="male"></specimenCount>
).
</materialsCitation>
<typeStatus id="E54C8879FFEEFFCAFC16FE7790E2FE64" box="[897,999,431,452]" pageId="4" pageNumber="489" type="paratype">Paratypes</typeStatus>
are deposited at
<collectionCode id="5CE6AE1EFFEEFFCAFEE7FE1192B8FE7E" box="[368,445,457,478]" collectionName="AAHC" pageId="4" pageNumber="489">AAHC</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFE5BFE11911AFE7E" box="[460,543,457,478]" collectionName="USA, New York, New York, American Museum of Natural History" pageId="4" pageNumber="489">AMNH</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFDB8FE119173FE7E" box="[559,630,457,478]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="4" pageNumber="489">CASC</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFD11FE1191BBFE7E" box="[646,702,457,478]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="4" pageNumber="489">CNC</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFD59FE119019FE7E" box="[718,796,457,478]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="4" pageNumber="489">FMNH</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFCBBFE11909CFE7E" box="[812,921,457,478]" pageId="4" pageNumber="489">IBUNAM</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFC3FFE1190ECFE7E" box="[936,1001,457,478]" pageId="4" pageNumber="489">IEXA</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFC6EFE119733FE7E" box="[1017,1078,457,478]" collectionName="IMEJ" pageId="4" pageNumber="489">IMEJ</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFF6EFE3C9244FE59" box="[249,321,484,505]" collectionName="GNPC" pageId="4" pageNumber="489">GNPC</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFED9FE3C92A4FE59" box="[334,417,484,505]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="4" pageNumber="489">MNHN</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFE3AFE3C92F8FE59" box="[429,509,484,505]" pageId="4" pageNumber="489">MXAL</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFD9DFE3C915EFE59" box="[522,603,484,505]" collectionName="Myanmar, Rangoon, Natural History Museum" pageId="4" pageNumber="489">NHMC</collectionCode>
,
<collectionCode id="5CE6AE1EFFEEFFCAFDF0FE3C91B7FE59" box="[615,690,484,505]" collectionName="USA, Nebraska, Lincoln, University of Nebraska State Museum" pageId="4" pageNumber="489">UNSM</collectionCode>
and
<collectionCode id="5CE6AE1EFFEEFFCAFD7DFE3C903FFE59" box="[746,826,484,505]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="4" pageNumber="489">ZMHU</collectionCode>
.
</paragraph>
<paragraph id="3A4836DBFFEEFFCAFE84FE269724FD8E" blockId="4.[249,1082,269,1284]" pageId="4" pageNumber="489">
<emphasis id="0883EAC9FFEEFFCAFE84FE2692B2FDB3" bold="true" box="[275,439,510,531]" pageId="4" pageNumber="489">Type Locality.</emphasis>
<location id="3F286000FFEEFFCAFE57FE279129FDB4" LSID="urn:lsid:plazi:treatment:B25E87CDFFEBFFCAFF60FB939703FAA4:3F286000FFEEFFCAFE57FE279129FDB4" box="[448,556,511,532]" country="Mexico" county="Autlan" municipality="Biosphere Reserve Sierra de Manantlan" name="Las Joyas" pageId="4" pageNumber="489" stateProvince="Jalisco">Las Joyas</location>
,
<collectingMunicipality id="DA2CACA1FFEEFFCAFDAFFE2790E0FDB4" box="[568,997,511,532]" pageId="4" pageNumber="489">Biosphere Reserve Sierra de Manantlán</collectingMunicipality>
,
<collectingCounty id="D3294E57FFEEFFCAFC65FE27973CFDB4" box="[1010,1081,511,532]" pageId="4" pageNumber="489">Autlán</collectingCounty>
municipality, state of
<collectingRegion id="F833F839FFEEFFCAFE74FDC19128FD8E" box="[483,557,537,558]" country="Mexico" name="Jalisco" pageId="4" pageNumber="489">Jalisco</collectingRegion>
,
<collectingCountry id="42E0764BFFEEFFCAFDADFDC1918FFD8E" box="[570,650,537,558]" name="Mexico" pageId="4" pageNumber="489">
<collectingRegion id="F833F839FFEEFFCAFDADFDC1918FFD8E" box="[570,650,537,558]" country="Mexico" name="Mexico" pageId="4" pageNumber="489">México</collectingRegion>
</collectingCountry>
(approx.
<geoCoordinate id="5FC3501CFFEEFFCAFD63FDC19075FD8E" box="[756,880,536,558]" degrees="19" direction="north" minutes="35" orientation="latitude" pageId="4" pageNumber="489" precision="15" seconds="13" value="19.586945">
19
<emphasis id="0883EAC9FFEEFFCAFC99FDC19013FD8E" bold="true" box="[782,790,537,558]" pageId="4" pageNumber="489">°</emphasis>
35
<emphasis id="0883EAC9FFEEFFCAFCA7FDC0903DFD8E" bold="true" box="[816,824,536,558]" pageId="4" pageNumber="489">'</emphasis>
13
<emphasis id="0883EAC9FFEEFFCAFCC5FDC0905FFD8E" bold="true" box="[850,858,536,558]" pageId="4" pageNumber="489">&quot;</emphasis>
N
</geoCoordinate>
;
<geoCoordinate id="5FC3501CFFEEFFCAFCEAFDC19713FD8E" box="[893,1046,536,558]" degrees="104" direction="west" minutes="16" orientation="longitude" pageId="4" pageNumber="489" precision="15" seconds="48" value="-104.28001">
104
<emphasis id="0883EAC9FFEEFFCAFC33FDC190A9FD8E" bold="true" box="[932,940,537,558]" pageId="4" pageNumber="489">°</emphasis>
16
<emphasis id="0883EAC9FFEEFFCAFC51FDC090CBFD8E" bold="true" box="[966,974,536,558]" pageId="4" pageNumber="489">'</emphasis>
48
<emphasis id="0883EAC9FFEEFFCAFC66FDC090FCFD8E" bold="true" box="[1009,1017,536,558]" pageId="4" pageNumber="489">&quot;</emphasis>
W
</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFEEFFCAFE84FDEC92E5FD4A" pageId="4" pageNumber="489" type="biology_ecology">
<paragraph id="3A4836DBFFEEFFCAFE84FDEC92E5FD4A" blockId="4.[249,1082,269,1284]" pageId="4" pageNumber="489">
<emphasis id="0883EAC9FFEEFFCAFE84FDEC92CDFDE9" bold="true" box="[275,456,564,585]" pageId="4" pageNumber="489">Biological Data.</emphasis>
Specimens of
<taxonomicName id="FDF74D58FFEEFFCAFDFCFDEC91FEFDE9" box="[619,763,564,585]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="sayloriana" status="sp. nov.">
<emphasis id="0883EAC9FFEEFFCAFDFCFDEC91FEFDE9" box="[619,763,564,585]" italics="true" pageId="4" pageNumber="489">P. sayloriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEEFFCAFC93FDEC9033FDE9" box="[772,822,564,585]" pageId="4" pageNumber="489" rank="species">n.sp.</taxonomicNameLabel>
were collected at white fluorescent and UV lights in montane cloud forest, pine forest and secondary vegetation located at
<quantity id="FD0F9B3EFFEEFFCAFE76FDB19187FDDE" box="[481,642,617,638]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.3" metricValueMin="1.9" pageId="4" pageNumber="489" unit="m" value="2100.0" valueMax="2300.0" valueMin="1900.0">
<geoCoordinate id="5FC3501CFFEEFFCAFE76FDB1911EFDDE" box="[481,539,617,638]" degrees="1,900" direction="north" orientation="latitude" pageId="4" pageNumber="489" precision="55" value="1.9">1,900</geoCoordinate>
<geoCoordinate id="5FC3501CFFEEFFCAFD8CFDB19161FDDE" box="[539,612,617,638]" degrees="2,300" direction="west" orientation="longitude" pageId="4" pageNumber="489" precision="55" value="-2.3">2,300</geoCoordinate>
m
</quantity>
altitude. Phenology: June (9), July (53), August (67), September (8), October (3), November (2). Other species of
<taxonomicName id="FDF74D58FFEEFFCAFF6EFD47927AFD14" box="[249,383,671,692]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="0883EAC9FFEEFFCAFF6EFD47927AFD14" box="[249,383,671,692]" italics="true" pageId="4" pageNumber="489">Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and place were
<taxonomicName id="FDF74D58FFEEFFCAFCAFFD47973CFD14" box="[824,1081,671,692]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="dugesiana" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFEEFFCAFCAFFD47973CFD14" box="[824,1081,671,692]" italics="true" pageId="4" pageNumber="489">P. (Phytalus) dugesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEEFFCAFF6EFD629229FD6F" box="[249,300,698,719]" pageId="4" pageNumber="489" rank="species">n.sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFEEFFCAFEAEFD62913AFD6F" box="[313,575,698,719]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="jalisciensis" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFEEFFCAFEAEFD62913AFD6F" box="[313,575,698,719]" italics="true" pageId="4" pageNumber="489">P. (Phytalus) jalisciensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEEFFCAFDD0FD62917FFD6F" box="[583,634,698,719]" pageId="4" pageNumber="489" rank="species">n.sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFEEFFCAFD10FD62906FFD6F" box="[647,874,698,719]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="manantleca" status="sp. nov.">
<emphasis id="0883EAC9FFEEFFCAFD10FD62919BFD6F" box="[647,670,698,719]" italics="true" pageId="4" pageNumber="489">P.</emphasis>
(
<emphasis id="0883EAC9FFEEFFCAFD3AFD6291E4FD6F" box="[685,737,698,719]" italics="true" pageId="4" pageNumber="489">s.str.</emphasis>
)
<emphasis id="0883EAC9FFEEFFCAFD67FD62906FFD6F" box="[752,874,698,719]" italics="true" pageId="4" pageNumber="489">manantleca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEEFFCAFCE5FD6290A6FD6F" box="[882,931,698,719]" pageId="4" pageNumber="489" rank="species">n.sp.</taxonomicNameLabel>
and
<taxonomicName id="FDF74D58FFEEFFCAFC40FD6292E5FD4A" authority="Moron." authorityName="Moron." class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="chamacayoca">
<emphasis id="0883EAC9FFEEFFCAFC40FD6290EBFD6F" box="[983,1006,698,719]" italics="true" pageId="4" pageNumber="489">P.</emphasis>
(
<emphasis id="0883EAC9FFEEFFCAFC69FD629734FD6F" box="[1022,1073,698,719]" italics="true" pageId="4" pageNumber="489">s.str.</emphasis>
)
<emphasis id="0883EAC9FFEEFFCAFF6EFD0D928CFD4A" box="[249,393,725,746]" italics="true" pageId="4" pageNumber="489">chamacayoca</emphasis>
Morón.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFEEFFCAFE84FD37905BFB14" pageId="4" pageNumber="489" type="discussion">
<paragraph id="3A4836DBFFEEFFCAFE84FD37905BFB14" blockId="4.[249,1082,269,1284]" pageId="4" pageNumber="489">
<emphasis id="0883EAC9FFEEFFCAFE84FD379285FCA4" bold="true" box="[275,384,751,772]" pageId="4" pageNumber="489">Remarks.</emphasis>
<taxonomicName id="FDF74D58FFEEFFCAFE1FFD289180FCA5" box="[392,645,752,773]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="sayloriana" status="sp. nov.">
<emphasis id="0883EAC9FFEEFFCAFE1FFD289180FCA5" box="[392,645,752,773]" italics="true" pageId="4" pageNumber="489">Phyllophaga sayloriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEEFFCAFD1AFD2891BAFCA5" box="[653,703,752,773]" pageId="4" pageNumber="489" rank="species">n.sp.</taxonomicNameLabel>
belong to the species group
<taxonomicName id="FDF74D58FFEEFFCAFB95FD289238FC80" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="0883EAC9FFEEFFCAFB95FD289238FC80" italics="true" pageId="4" pageNumber="489">blanchardi</emphasis>
</taxonomicName>
, complex
<taxonomicName id="FDF74D58FFEEFFCAFE45FCD39145FC80" box="[466,576,779,800]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="pubicauda">
<emphasis id="0883EAC9FFEEFFCAFE45FCD39145FC80" box="[466,576,779,800]" italics="true" pageId="4" pageNumber="489">pubicauda</emphasis>
</taxonomicName>
(
<emphasis id="0883EAC9FFEEFFCAFDF4FCD39198FC80" box="[611,669,779,800]" italics="true" pageId="4" pageNumber="489">sensu</emphasis>
<bibRefCitation id="5E664B2AFFEEFFCAFD30FCD39034FC80" author="Moron, M. A." bookContentInfo="Publ. 20, Instituto de Ecologia, Mexico. 341 pp." box="[679,817,779,800]" pageId="4" pageNumber="489" refId="ref10446" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica. (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico. 341 pp." title="El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica. (Insecta: Coleoptera)" type="book" year="1986">Morón 1986</bibRefCitation>
). The diagnostic combination of characters for both sexes include: anterior border of clypeus deep and widely sinuated; frons, pronotum and elytra with dense vestiture of erect, very long setae; abdominal sternites 2
<emphasis id="0883EAC9FFEEFFCAFD1FFC839195FCD0" bold="true" box="[648,656,859,880]" pageId="4" pageNumber="489">°</emphasis>
to 4
<emphasis id="0883EAC9FFEEFFCAFD28FC8391C2FCD0" bold="true" box="[703,711,859,880]" pageId="4" pageNumber="489">°</emphasis>
with deep mesial sulcus; pygidium noticeably rugose; and metatarsal segments 2
<emphasis id="0883EAC9FFEEFFCAFD7AFCAE91F0FC2B" bold="true" box="[749,757,886,907]" pageId="4" pageNumber="489">°</emphasis>
to 4
<emphasis id="0883EAC9FFEEFFCAFCBAFCAE9030FC2B" bold="true" box="[813,821,886,907]" pageId="4" pageNumber="489">°</emphasis>
with comb­like line of setae in ventral sides. Shape of paramera is similar to
<taxonomicName id="FDF74D58FFEEFFCAFCD4FC499712FC06" authority="Bates" authorityName="Bates" box="[835,1047,913,934]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="pubicauda">
<emphasis id="0883EAC9FFEEFFCAFCD4FC4990D4FC06" box="[835,977,913,934]" italics="true" pageId="4" pageNumber="489">P. pubicauda</emphasis>
Bates
</taxonomicName>
,
<taxonomicName id="FDF74D58FFEEFFCAFBB4FC4992BEFC61" authority="Arrow" authorityName="Arrow" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="0883EAC9FFEEFFCAFBB4FC499269FC61" italics="true" pageId="4" pageNumber="489">P. blanchardi</emphasis>
Arrow
</taxonomicName>
,
<taxonomicName id="FDF74D58FFEEFFCAFE5FFC7491A9FC61" authority="Arrow" authorityName="Arrow" box="[456,684,940,961]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="microcerus">
<emphasis id="0883EAC9FFEEFFCAFE5FFC749158FC61" box="[456,605,940,961]" italics="true" pageId="4" pageNumber="489">P. microcerus</emphasis>
Arrow
</taxonomicName>
,
<taxonomicName id="FDF74D58FFEEFFCAFD2EFC749088FC61" authority="Moser" authorityName="Moser" box="[697,909,940,961]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="rufithorax">
<emphasis id="0883EAC9FFEEFFCAFD2EFC749047FC61" box="[697,834,940,961]" italics="true" pageId="4" pageNumber="489">P. rufithorax</emphasis>
Moser
</taxonomicName>
,
<taxonomicName id="FDF74D58FFEEFFCAFC0DFC74973CFC61" authority="Saylor" authorityName="Saylor" authorityYear="1938" box="[922,1081,940,961]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="moei">
<emphasis id="0883EAC9FFEEFFCAFC0DFC7490E8FC61" box="[922,1005,940,961]" italics="true" pageId="4" pageNumber="489">P. moei</emphasis>
Saylor
</taxonomicName>
and
<taxonomicName id="FDF74D58FFEEFFCAFEB0FC1F910CFC7C" authority="Saylor" authorityName="Saylor" authorityYear="1943" box="[295,521,967,988]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="hidalgoana">
<emphasis id="0883EAC9FFEEFFCAFEB0FC1F92B9FC7C" box="[295,444,967,988]" italics="true" pageId="4" pageNumber="489">P. hidalgoana</emphasis>
Saylor
</taxonomicName>
, but the proportions of each part of the paramera are different, and in all of these species the pygidium is not rugose and the sternites are not deeply sulcated. The dorsal vestiture and punctuation, body color and mesial sulcus on sternites, suggests some relation with
<taxonomicName id="FDF74D58FFEEFFCAFCF7FBC09730FB8D" authority="Moser" authorityName="Moser" box="[864,1077,1048,1069]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="hirticollis">
<emphasis id="0883EAC9FFEEFFCAFCF7FBC090E2FB8D" box="[864,999,1048,1069]" italics="true" pageId="4" pageNumber="489">P. hirticollis</emphasis>
Moser
</taxonomicName>
, known only from Buena Vista,
<collectingCountry id="42E0764BFFEEFFCAFDF6FBEB91BCFBE8" box="[609,697,1075,1096]" name="Mexico" pageId="4" pageNumber="489">Mexico</collectingCountry>
, but the details and proportions of male genital capsule, pygidial sculpture, and metatarsi vestiture, as well as body size, are different. Dorsal aspect of
<taxonomicName id="FDF74D58FFEEFFCAFD56FBB19054FBDE" box="[705,849,1129,1150]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="sayloriana">
<emphasis id="0883EAC9FFEEFFCAFD56FBB19054FBDE" box="[705,849,1129,1150]" italics="true" pageId="4" pageNumber="489">P. sayloriana</emphasis>
</taxonomicName>
also resembles some species in the group
<emphasis id="0883EAC9FFEEFFCAFE63FB5C9170FB39" box="[500,629,1156,1177]" italics="true" pageId="4" pageNumber="489">heteronycha</emphasis>
, but the shape of abdominal segments, tarsal claws and male genital capsule are much different.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFEEFFCAFE84FB619703FAA4" pageId="4" pageNumber="489" type="etymology">
<paragraph id="3A4836DBFFEEFFCAFE84FB619703FAA4" blockId="4.[249,1082,269,1284]" pageId="4" pageNumber="489">
<emphasis id="0883EAC9FFEEFFCAFE84FB619296FB6E" bold="true" box="[275,403,1209,1230]" pageId="4" pageNumber="489">Etymology.</emphasis>
This new species is dedicated to Lawrence W. Saylor, whose numerous descriptions of species of the genus
<taxonomicName id="FDF74D58FFEEFFCAFD6CFB0D9084FB4A" box="[763,897,1237,1258]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="0883EAC9FFEEFFCAFD6CFB0D9084FB4A" box="[763,897,1237,1258]" italics="true" pageId="4" pageNumber="489">Phyllophaga</emphasis>
</taxonomicName>
provided an important step for the study of these beetles in
<collectingCountry id="42E0764BFFEEFFCAFD4AFB379028FAA4" box="[733,813,1263,1284]" name="Mexico" pageId="4" pageNumber="489">Mexico</collectingCountry>
and Latin America.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,373 @@
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<mods:title id="3D71BC86AB6E2F4ED42ACDC4B169F30B">New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México</mods:title>
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<mods:namePart id="BEDD396496D089CE290D72E544A10B97">Morón, Miguel-Angel</mods:namePart>
<mods:affiliation id="3A62FC0839FDC5258CBD120B98C197E0">Departamento de Entomología Instituto de Ecología, A. C. (Sector SEP­CONACYT) Apdo. Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
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<mods:namePart id="8E0440C78AA975D024752CE3C5184593">Rivera-Cervantes, Luis Eugenio</mods:namePart>
<mods:affiliation id="733B84229ACE3C3A60D5D0BCAC6DEF76">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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<mods:namePart id="60D279021625D88C42D936A414865D5E">López-Vieyra, Mario</mods:namePart>
<mods:affiliation id="E805BE0C3E18BD663CEF870E38607140">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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<treatment id="B25E87CDFFEDFFC7FE9CFC8290B5FB34" LSID="urn:lsid:plazi:treatment:B25E87CDFFEDFFC7FE9CFC8290B5FB34" httpUri="http://treatment.plazi.org/id/B25E87CDFFEDFFC7FE9CFC8290B5FB34" lastPageId="9" lastPageNumber="494" pageId="7" pageNumber="492">
<subSubSection id="72ED6550FFEDFFC9FE9CFC82971DFCCE" box="[267,1048,857,879]" pageId="7" pageNumber="492" type="nomenclature">
<paragraph id="3A4836DBFFEDFFC9FE9CFC82971DFCCE" blockId="7.[267,1048,857,905]" box="[267,1048,857,879]" pageId="7" pageNumber="492">
<taxonomicName id="FDF74D58FFEDFFC9FE9CFC82908DFCCE" authority="Moron, Rivera y Lopez" authorityName="Moron, Rivera y Lopez" box="[267,904,857,879]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="joyana" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="0883EAC9FFEDFFC9FE9CFC829294FCCF" box="[267,401,858,879]" italics="true" pageId="7" pageNumber="492">Phyllophaga</emphasis>
(
<emphasis id="0883EAC9FFEDFFC9FE35FC829122FCCF" box="[418,551,858,879]" italics="true" pageId="7" pageNumber="492">Phyllophaga</emphasis>
)
<emphasis id="0883EAC9FFEDFFC9FDAEFC829185FCCF" box="[569,640,858,879]" italics="true" pageId="7" pageNumber="492">joyana</emphasis>
Morón, Rivera y López
</taxonomicName>
,
<emphasis id="0883EAC9FFEDFFC9FC02FC81971DFCCE" bold="true" box="[917,1048,857,878]" pageId="7" pageNumber="492">
<taxonomicNameLabel id="13B057B2FFEDFFC9FC02FC81971DFCCE" box="[917,1048,857,878]" pageId="7" pageNumber="492" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFEDFFC7FDC1FCAC971DFDB4" lastPageId="9" lastPageNumber="494" pageId="7" pageNumber="492" type="description">
<paragraph id="3A4836DBFFEDFFC9FDC1FCAC91D0FC29" blockId="7.[267,1048,857,905]" box="[598,725,884,905]" pageId="7" pageNumber="492">
<figureCitation id="A2CC2A5EFFEDFFC9FDC1FCAC91D0FC29" box="[598,725,884,905]" captionStart="Figs" captionStartId="8.[272,315,990,1009]" captionTargetBox="[249,1081,268,965]" captionTargetId="figure-12@8.[246,1084,265,969]" captionTargetPageId="8" captionText="Figs. 1417. Phyllophaga joyana. 14) tarsal claw, male; 15) genital capsule, lateral view; 16) paramera, distal view; 17) distal half of genital capsule, dorsal view. Scale lines = 1 mm, except Fig. 14 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102841" httpUri="https://zenodo.org/record/10102841/files/figure.png" pageId="7" pageNumber="492">Figs. 1417</figureCitation>
</paragraph>
<paragraph id="3A4836DBFFEDFFC6FE9AFC7A9289FA59" blockId="7.[246,1079,930,1652]" lastBlockId="8.[249,1082,1125,1529]" lastPageId="8" lastPageNumber="493" pageId="7" pageNumber="492">
<emphasis id="0883EAC9FFEDFFC9FE9AFC7A9262FC15" bold="true" box="[269,359,930,949]" pageId="7" pageNumber="492">
<typeStatus id="E54C8879FFEDFFC9FE9AFC7A9262FC15" box="[269,359,930,949]" pageId="7" pageNumber="492" type="holotype">Holotype</typeStatus>
</emphasis>
Male. Head, pronotum and elytra shiny yellowish dark brown; pygidium, sterna, and legs shiny yellowish testaceous. Clypeus clearly bilobed, wider than long (4.8:1), anterior border deeply sinuated, with margins scarcely elevated, surface convex, with large, round punctures, nearly confluent, with long, slender setae. Fronto­clypeal suture slightly sinuose, finely impressed. Frons wider than long (2.2:1) convex, irregularly rugo­punctate, with long, slender setae. Antenna 10 segmented, with 3 segmented club; lamellae as long as the length of five preceeding segments combined (1:1), segment 4
<emphasis id="0883EAC9FFEDFFC9FE96FB91920DFBFD" bold="true" box="[257,264,1097,1117]" pageId="7" pageNumber="492">°</emphasis>
nearly as long as segment 3, segment 5
<emphasis id="0883EAC9FFEDFFC9FD35FB9191ACFBFD" bold="true" box="[674,681,1097,1117]" pageId="7" pageNumber="492">°</emphasis>
with small anterior, rounded projection, segments 6
<emphasis id="0883EAC9FFEDFFC9FEC9FBBA9260FBD6" bold="true" box="[350,357,1122,1142]" pageId="7" pageNumber="492">°</emphasis>
and 7
<emphasis id="0883EAC9FFEDFFC9FE37FBBA92A2FBD6" bold="true" box="[416,423,1122,1142]" pageId="7" pageNumber="492">°</emphasis>
wider than long, each one with acute process directed forward. Frons 6.2
<emphasis id="0883EAC9FFEDFFC9FE84FBA39220FB2E" bold="true" box="[275,293,1147,1166]" pageId="7" pageNumber="492">×</emphasis>
wider than each eye dorsal diameter. Canthus curved and rounded, with 57 setae. Labrum deeply bilobed, with curved, slender setae near borders. Mentum widely concave, polished, with slender setae at center and near lateral borders, anterior border clearly sinuated. Pronotum wider than long (1.9:1) and 1.9
<emphasis id="0883EAC9FFEDFFC9FCB1FB1C903DFB77" bold="true" box="[806,824,1220,1239]" pageId="7" pageNumber="492">×</emphasis>
wider than frons. Pronotal disk with heterogeneous, deep, round punctures separated by 13 diameters, with many erect, very long, slender setae; lateral borders widely angled, marginal bead vaguely crenulate, with long, slender setae; anterior angles obtuse, scarcely prominent; posterior angles widely obtuse, not prominent. Scutellum 1.4
<emphasis id="0883EAC9FFEDFFC9FD73FAFD91F3FA98" bold="true" box="[740,758,1317,1336]" pageId="7" pageNumber="492">×</emphasis>
wider than long, with 9 shallow, small punctures. Elytron 2.6
<emphasis id="0883EAC9FFEDFFC9FD91FAE5911DFAF0" bold="true" box="[518,536,1341,1360]" pageId="7" pageNumber="492">×</emphasis>
longer than wide, densely rugo­punctate, without setae; epipleural border narrowed, extended along complete margin, provided with a fringe of setae progressvely shorter toward the apex of elytra; humeral calla rounded, prominent; apical calla rounded. Metathoracic wings completely developed. Propygidium shiny, fine and shallowly punctate, without setae. Pygidium convex, slightly prominent toward base, shiny, with scattered, shallow punctures, without setae on disk; apical margin with 14 slender setae; basal margin effaced at middle. Pterosternon with long, dense, yellowish vestiture. Visible abdominal sternites 2
<emphasis id="0883EAC9FFEDFFC9FDF7FA3E9162FA5A" bold="true" box="[608,615,1510,1530]" pageId="7" pageNumber="492">°</emphasis>
to 4
<emphasis id="0883EAC9FFEDFFC9FD07FA3E9192FA5A" bold="true" box="[656,663,1510,1530]" pageId="7" pageNumber="492">°</emphasis>
with shallow, wide, longitudinal depression; sternite 5
<emphasis id="0883EAC9FFEDFFC9FED9FA269250F9B2" bold="true" box="[334,341,1534,1554]" pageId="7" pageNumber="492">°</emphasis>
convex, scattered punctures and some slender setae; posterior border slightly excavated toward sides; anal plate wide and shallowly concave, with vague punctures, without setae; posterior border thickened, scarcely notched at middle, with 10 erect, long setae. Protibiae slightly shorter than protarsi (0.9:1), with two large teeth and one acute, small tooth on external border, preapical spur long, straight, acute, slightly shorter than 2
<emphasis id="0883EAC9FFE2FFC6FE92FBBD9209FBD9" bold="true" box="[261,268,1125,1145]" pageId="8" pageNumber="493">°</emphasis>
protarsomerus (0.8:1). Mesotibiae with some long, slender setae, and one oblique, short, setiferous, transverse carina; upper apical spur straight, narrow, slightly shorter than lower spur (0.8:1). Metatibiae shorter than metatarsi (0.9:1), with some long, slender setae, and one oblique, short setiferous transverse carina; apical spurs articulated with the border, upper spur lanceolate, slightly curved, with rounded apex, nearly as long as 2
<emphasis id="0883EAC9FFE2FFC6FE92FB059209FB51" bold="true" box="[261,268,1245,1265]" pageId="8" pageNumber="493">°</emphasis>
metatarsomerus, and 1.2
<emphasis id="0883EAC9FFE2FFC6FE6BFB06910BFB51" bold="true" box="[508,526,1246,1265]" pageId="8" pageNumber="493">×</emphasis>
longer than lower spur. Tarsomeres semicylindrical, elongated, with enlarged apex, and many setae around apex; mesotarsomeres 2
<emphasis id="0883EAC9FFE2FFC6FC5DFB2D90D4FAA9" bold="true" box="[970,977,1269,1289]" pageId="8" pageNumber="493">°</emphasis>
to 5
<emphasis id="0883EAC9FFE2FFC6FB97FB2D9702FAA9" bold="true" box="[1024,1031,1269,1289]" pageId="8" pageNumber="493">°</emphasis>
ventrally present two lines of scattered, slender setae; metatarsomeres 2
<emphasis id="0883EAC9FFE2FFC6FC0BFAD590A6FA81" bold="true" box="[924,931,1293,1313]" pageId="8" pageNumber="493">°</emphasis>
to 4
<emphasis id="0883EAC9FFE2FFC6FC42FAD590D9FA81" bold="true" box="[981,988,1293,1313]" pageId="8" pageNumber="493">°</emphasis>
ventrally present two lines of setae. Tarsal claws symmetrical, similar on all legs, with median acute­angled tooth located near basal dilatation, but also near apical tooth (
<figureCitation id="A2CC2A5EFFE2FFC6FC72FAE6972AFAF1" box="[997,1071,1342,1361]" captionStart="Figs" captionStartId="8.[272,315,990,1009]" captionTargetBox="[249,1081,268,965]" captionTargetId="figure-12@8.[246,1084,265,969]" captionTargetPageId="8" captionText="Figs. 1417. Phyllophaga joyana. 14) tarsal claw, male; 15) genital capsule, lateral view; 16) paramera, distal view; 17) distal half of genital capsule, dorsal view. Scale lines = 1 mm, except Fig. 14 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102841" httpUri="https://zenodo.org/record/10102841/files/figure.png" pageId="8" pageNumber="493">Fig. 14</figureCitation>
). Genital capsule with long, narrowed, curved paramera, fused dorsally at their basis, and ventrally at their apex, with lateral, rounded laminar projections and preapical short, tooth­like, curved projections (
<figureCitation id="A2CC2A5EFFE2FFC6FD88FA5E9197FA39" box="[543,658,1414,1433]" captionStart="Figs" captionStartId="8.[272,315,990,1009]" captionTargetBox="[249,1081,268,965]" captionTargetId="figure-12@8.[246,1084,265,969]" captionTargetPageId="8" captionText="Figs. 1417. Phyllophaga joyana. 14) tarsal claw, male; 15) genital capsule, lateral view; 16) paramera, distal view; 17) distal half of genital capsule, dorsal view. Scale lines = 1 mm, except Fig. 14 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102841" httpUri="https://zenodo.org/record/10102841/files/figure.png" pageId="8" pageNumber="493">Figs. 1517</figureCitation>
). Tectum convex with widely sinuated distal border. Aedeagus long, wide, with sclerotized tube­like support and preapical, dorsal, bilobed, excavated, sclerotized plate (
<figureCitation id="A2CC2A5EFFE2FFC6FDC9FA6E91D4FA69" box="[606,721,1462,1481]" captionStart="Figs" captionStartId="8.[272,315,990,1009]" captionTargetBox="[249,1081,268,965]" captionTargetId="figure-12@8.[246,1084,265,969]" captionTargetPageId="8" captionText="Figs. 1417. Phyllophaga joyana. 14) tarsal claw, male; 15) genital capsule, lateral view; 16) paramera, distal view; 17) distal half of genital capsule, dorsal view. Scale lines = 1 mm, except Fig. 14 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102841" httpUri="https://zenodo.org/record/10102841/files/figure.png" pageId="8" pageNumber="493">Figs. 1517</figureCitation>
). Length of genital capsule from apex of parameres to border of basal piece:
<quantity id="FD0F9B3EFFE2FFC6FDE4FA1691C5FA41" box="[627,704,1486,1505]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="493" unit="mm" value="3.4">3.4 mm</quantity>
. Total body length: 13.0 mm. Humeral width:
<quantity id="FD0F9B3EFFE2FFC6FEABFA3E928DFA59" box="[316,392,1510,1529]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="493" unit="mm" value="5.5">5.5 mm</quantity>
.
</paragraph>
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<paragraph id="3A4836DBFFE2FFC6FE87FC069183FB81" blockId="8.[249,1081,990,1057]" pageId="8" pageNumber="493">
<emphasis id="0883EAC9FFE2FFC6FE87FC069282FC51" bold="true" box="[272,391,990,1009]" pageId="8" pageNumber="493">Figs. 1417.</emphasis>
<emphasis id="0883EAC9FFE2FFC6FE09FC069166FC51" box="[414,611,990,1009]" italics="true" pageId="8" pageNumber="493">
<taxonomicName id="FDF74D58FFE2FFC6FE09FC06915AFC51" box="[414,607,990,1009]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="493" phylum="Arthropoda" rank="species" species="joyana">Phyllophaga joyana</taxonomicName>
.
</emphasis>
<emphasis id="0883EAC9FFE2FFC6FDFDFC06918CFC51" bold="true" box="[618,649,990,1009]" pageId="8" pageNumber="493">14)</emphasis>
tarsal claw, male;
<emphasis id="0883EAC9FFE2FFC6FCA8FC06905BFC51" bold="true" box="[831,862,990,1009]" pageId="8" pageNumber="493">15)</emphasis>
genital capsule, lateral view;
<emphasis id="0883EAC9FFE2FFC6FEA1FC2E9250FBA9" bold="true" box="[310,341,1014,1033]" pageId="8" pageNumber="493">16)</emphasis>
paramera, distal view;
<emphasis id="0883EAC9FFE2FFC6FDAAFC2E9159FBA9" bold="true" box="[573,604,1014,1033]" pageId="8" pageNumber="493">17)</emphasis>
distal half of genital capsule, dorsal view. Scale lines
<emphasis id="0883EAC9FFE2FFC6FEBAFBD6923AFB81" bold="true" box="[301,319,1038,1057]" pageId="8" pageNumber="493">=</emphasis>
1 mm, except Fig. 14
<emphasis id="0883EAC9FFE2FFC6FD8BFBD6912BFB81" bold="true" box="[540,558,1038,1057]" pageId="8" pageNumber="493">=</emphasis>
0.5 mm.
</paragraph>
</caption>
<paragraph id="3A4836DBFFE2FFC7FE84F9D6929EFE64" blockId="8.[249,1081,1550,1652]" lastBlockId="9.[246,1079,271,1172]" lastPageId="9" lastPageNumber="494" pageId="8" pageNumber="493">
<emphasis id="0883EAC9FFE2FFC6FE84F9D6927CF983" bold="true" box="[275,377,1550,1571]" pageId="8" pageNumber="493">
<typeStatus id="E54C8879FFE2FFC6FE84F9D69276F983" box="[275,371,1550,1571]" pageId="8" pageNumber="493" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Similar to the male except as follows: antennal club as long as the four preceeding segments combined; pygidium transversally concave at middle with preapical prominence briefly bilobed; visible abdominal sternites 2
<emphasis id="0883EAC9FFE2FFC6FEFDF9869277F9D3" bold="true" box="[362,370,1630,1651]" pageId="8" pageNumber="493">°</emphasis>
to 4
<emphasis id="0883EAC9FFE2FFC6FE3BF98692B1F9D3" bold="true" box="[428,436,1630,1651]" pageId="8" pageNumber="493">°</emphasis>
with mesial sulcus vaguely indicated; sternite 5
<emphasis id="0883EAC9FFE2FFC6FC44F98690DEF9D3" bold="true" box="[979,987,1630,1651]" pageId="8" pageNumber="493">°</emphasis>
convex, punctate, with scattered short setae; anal plate convex, punctate, with scattered, short and medium size setae. Protibia with external border clearly tridentate. Apical spurs of metatibiae wide, slightly curvated with rounded apexes. Ventral genital plates widely ovate, moderately sclerotized, nearly symmetrical, poorly convex, without setae; dorsal genital plates nearly rectangular, fused mesially, with scarce setae on distal borders. Total body length: 13.0 mm. Humeral width:
<quantity id="FD0F9B3EFFE3FFC7FED5FE779292FE64" box="[322,407,431,452]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="9" pageNumber="494" unit="mm" value="5.6">5.6 mm</quantity>
.
</paragraph>
<paragraph id="3A4836DBFFE3FFC7FE87FE11971DFDB4" blockId="9.[246,1079,271,1172]" pageId="9" pageNumber="494">
<materialsCitation id="8A9F3C86FFE3FFC7FE87FE119711FDB4" location="Minatitlan" pageId="9" pageNumber="494" specimenCount="1" stateProvince="Colima" typeStatus="paratype">
<emphasis id="0883EAC9FFE3FFC7FE87FE1192F4FE7E" bold="true" box="[272,497,457,478]" pageId="9" pageNumber="494">
<typeStatus id="E54C8879FFE3FFC7FE87FE119273FE7E" box="[272,374,457,478]" pageId="9" pageNumber="494" type="paratype">Paratype</typeStatus>
Variation.
</emphasis>
Males. Specimens from
<location id="3F286000FFE3FFC7FD68FE11906AFE7E" LSID="urn:lsid:plazi:treatment:B25E87CDFFEDFFC7FE9CFC8290B5FB34:3F286000FFE3FFC7FD68FE11906AFE7E" box="[767,879,457,478]" name="Minatitlan" pageId="9" pageNumber="494" stateProvince="Colima">Minatitlán</location>
,
<collectingRegion id="F833F839FFE3FFC7FCECFE1190CDFE7E" box="[891,968,457,478]" country="Mexico" name="Colima" pageId="9" pageNumber="494">Colima</collectingRegion>
have pronotal punctures closely approximated, wider and denser than specimens from Las Joyas. Total body length: 13.0
<quantity id="FD0F9B3EFFE3FFC7FDE0FE2791DCFDB4" box="[631,729,511,532]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="9" pageNumber="494" unit="mm" value="13.2">13.2 mm</quantity>
, humeral width:
<quantity id="FD0F9B3EFFE3FFC7FC07FE279711FDB4" box="[912,1044,511,532]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="9" pageNumber="494" unit="mm" value="5.55" valueMax="5.6" valueMin="5.5">5.55.6 mm</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE3FFC7FE87FDC19710FD49" pageId="9" pageNumber="494" type="materials_examined">
<paragraph id="3A4836DBFFE3FFC7FE87FDC192F5FD14" blockId="9.[246,1079,271,1172]" pageId="9" pageNumber="494">
<emphasis id="0883EAC9FFE3FFC7FE87FDC1929EFD8E" bold="true" box="[272,411,537,558]" pageId="9" pageNumber="494">Type Series.</emphasis>
(Described from
<specimenCount id="2CF1FD52FFE3FFC7FDCCFDC191AAFD8E" box="[603,687,537,558]" pageId="9" pageNumber="494" type="male">4 males</specimenCount>
and
<specimenCount id="2CF1FD52FFE3FFC7FD70FDC1904CFD8E" box="[743,841,537,558]" pageId="9" pageNumber="494" type="female">1 female</specimenCount>
).
<materialsCitation id="8A9F3C86FFE3FFC7FCCBFDC1919AFDC4" collectingDate="1997-08-05" collectorName="M. Lopez" elevation="1900" location="Autlan" pageId="9" pageNumber="494" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<typeStatus id="E54C8879FFE3FFC7FCCBFDC190B8FD8E" box="[860,957,537,558]" pageId="9" pageNumber="494" type="holotype">Holotype</typeStatus>
♂; MEXI­ CO:
<emphasis id="0883EAC9FFE3FFC7FEBBFDEC9278FDE9" box="[300,381,564,585]" italics="true" pageId="9" pageNumber="494">
<collectingRegion id="F833F839FFE3FFC7FEBBFDEC927CFDE9" box="[300,377,564,585]" country="Mexico" name="Jalisco" pageId="9" pageNumber="494">Jalisco</collectingRegion>
,
</emphasis>
<location id="3F286000FFE3FFC7FE10FDEC92D4FDE9" LSID="urn:lsid:plazi:treatment:B25E87CDFFEDFFC7FE9CFC8290B5FB34:3F286000FFE3FFC7FE10FDEC92D4FDE9" box="[391,465,564,585]" name="Autlan" pageId="9" pageNumber="494" stateProvince="Jalisco">Autlán</location>
, Est. Las Joyas, bosque mesófilo, Alt.
<quantity id="FD0F9B3EFFE3FFC7FC17FDEC90D9FDE9" box="[896,988,564,585]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="494" unit="m" value="1900.0">
<elevation id="B1DAD1E8FFE3FFC7FC17FDEC90D9FDE9" box="[896,988,564,585]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="494" unit="m" value="1900.0">1,900 m</elevation>
</quantity>
,
<date id="4E49101BFFE3FFC7FC7CFDEC9228FDC4" pageId="9" pageNumber="494" value="1997-08-05">
<collectingDate id="5E0DE9F3FFE3FFC7FC7CFDEC9228FDC4" pageId="9" pageNumber="494" value="1997-08-05">5­VIII­ 1997</collectingDate>
</date>
, trampa luz,
<collectorName id="9702530DFFE3FFC7FE51FD979134FDC4" box="[454,561,591,612]" pageId="9" pageNumber="494">M. López</collectorName>
(
<collectionCode id="5CE6AE1EFFE3FFC7FDD2FD97919DFDC4" box="[581,664,591,612]" pageId="9" pageNumber="494">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFE3FFC7FD27FD979252FD39" collectingDate="1998-07-25" collectorName="El Terrero &amp; J. Garcia" elevation="2200" location="El Terrero" municipality="Minatitlan" pageId="9" pageNumber="494" specimenCount="1" specimenCount-female="1" stateProvince="Colima" typeStatus="allotype">
<typeStatus id="E54C8879FFE3FFC7FD27FD97900EFDC4" box="[688,779,591,612]" pageId="9" pageNumber="494" type="allotype">Allotype</typeStatus>
<specimenCount id="2CF1FD52FFE3FFC7FC8FFD959029FDC3" box="[792,812,589,611]" pageId="9" pageNumber="494" type="female"></specimenCount>
:
<emphasis id="0883EAC9FFE3FFC7FCA8FD979096FDC4" box="[831,915,591,612]" italics="true" pageId="9" pageNumber="494">
<collectingRegion id="F833F839FFE3FFC7FCA8FD97908BFDC4" box="[831,910,591,612]" country="Mexico" name="Colima" pageId="9" pageNumber="494">Colima</collectingRegion>
,
</emphasis>
<collectingMunicipality id="DA2CACA1FFE3FFC7FC08FD97970AFDC4" box="[927,1039,591,612]" pageId="9" pageNumber="494">Minatitlán</collectingMunicipality>
,
<collectorName id="9702530DFFE3FFC7FB88FD979240FDDE" pageId="9" pageNumber="494">
<location id="3F286000FFE3FFC7FB88FD979240FDDE" LSID="urn:lsid:plazi:treatment:B25E87CDFFEDFFC7FE9CFC8290B5FB34:3F286000FFE3FFC7FB88FD979240FDDE" municipality="Minatitlan" name="El Terrero" pageId="9" pageNumber="494" stateProvince="Colima">El Terrero</location>
</collectorName>
, bosque mesófilo, Alt.
<quantity id="FD0F9B3EFFE3FFC7FDAAFDB19192FDDE" box="[573,663,617,638]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="494" unit="m" value="2200.0">
<elevation id="B1DAD1E8FFE3FFC7FDAAFDB19192FDDE" box="[573,663,617,638]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="494" unit="m" value="2200.0">2,200 m</elevation>
</quantity>
,
<date id="4E49101BFFE3FFC7FD34FDB1902CFDDE" box="[675,809,617,638]" pageId="9" pageNumber="494" value="1998-07-25">
<collectingDate id="5E0DE9F3FFE3FFC7FD34FDB1902CFDDE" box="[675,809,617,638]" pageId="9" pageNumber="494" value="1998-07-25">25­VII­1998</collectingDate>
</date>
, trampa luz UV,
<collectorName id="9702530DFFE3FFC7FC77FDB19733FDDE" box="[992,1078,617,638]" pageId="9" pageNumber="494">J.García</collectorName>
(
<collectionCode id="5CE6AE1EFFE3FFC7FF6AFD5C9255FD39" box="[253,336,644,665]" pageId="9" pageNumber="494">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFE3FFC7FEF4FD5C92E9FD14" collectingDate="1997-08-05" collectorName="M. Lopez" elevation="1900" location="Autlan" pageId="9" pageNumber="494" specimenCount="2" stateProvince="Jalisco" typeStatus="paratype">
<typeStatus id="E54C8879FFE3FFC7FEF4FD5C92C8FD39" box="[355,461,644,665]" pageId="9" pageNumber="494" type="paratype">Paratypes</typeStatus>
: same data as holotype (♂) (
<collectionCode id="5CE6AE1EFFE3FFC7FC97FD5C9045FD39" box="[768,832,644,665]" pageId="9" pageNumber="494">IMEJ</collectionCode>
); same data as
<typeStatus id="E54C8879FFE3FFC7FC75FD5C9733FD39" box="[994,1078,644,665]" pageId="9" pageNumber="494" type="allotype">allotype</typeStatus>
(2 ♂♂)(
<collectionCode id="5CE6AE1EFFE3FFC7FEDCFD47928BFD14" box="[331,398,671,692]" pageId="9" pageNumber="494">IEXA</collectionCode>
,
<collectionCode id="5CE6AE1EFFE3FFC7FE05FD4792E0FD14" box="[402,485,671,692]" pageId="9" pageNumber="494">MXAL</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3A4836DBFFE3FFC7FE87FD619710FD49" blockId="9.[246,1079,271,1172]" pageId="9" pageNumber="494">
<materialsCitation id="8A9F3C86FFE3FFC7FE87FD619714FD49" country="Mexico" county="Autlan" location="Las Joyas" municipality="Biosphere Reserve Sierra de Manantlan" pageId="9" pageNumber="494" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="0883EAC9FFE3FFC7FE87FD6192B1FD6E" bold="true" box="[272,436,697,718]" pageId="9" pageNumber="494">
<typeStatus id="E54C8879FFE3FFC7FE87FD61924CFD6E" box="[272,329,697,718]" pageId="9" pageNumber="494">Type</typeStatus>
Locality.
</emphasis>
<location id="3F286000FFE3FFC7FE2AFD61912DFD6E" LSID="urn:lsid:plazi:treatment:B25E87CDFFEDFFC7FE9CFC8290B5FB34:3F286000FFE3FFC7FE2AFD61912DFD6E" box="[445,552,697,718]" country="Mexico" county="Autlan" municipality="Biosphere Reserve Sierra de Manantlan" name="Las Joyas" pageId="9" pageNumber="494" stateProvince="Jalisco">Las Joyas</location>
,
<collectingMunicipality id="DA2CACA1FFE3FFC7FDA2FD6190E7FD6E" box="[565,994,697,718]" pageId="9" pageNumber="494">Biosphere Reserve Sierra de Manantlán</collectingMunicipality>
,
<collectingCounty id="D3294E57FFE3FFC7FC78FD619733FD6E" box="[1007,1078,697,718]" pageId="9" pageNumber="494">Autlán</collectingCounty>
municipality, state of
<collectingRegion id="F833F839FFE3FFC7FE77FD0C912FFD49" box="[480,554,724,745]" country="Mexico" name="Jalisco" pageId="9" pageNumber="494">Jalisco</collectingRegion>
,
<collectingCountry id="42E0764BFFE3FFC7FDA0FD0C9182FD49" box="[567,647,724,745]" name="Mexico" pageId="9" pageNumber="494">México</collectingCountry>
(approx. 19
<emphasis id="0883EAC9FFE3FFC7FC9DFD0C9017FD49" bold="true" box="[778,786,724,745]" pageId="9" pageNumber="494">°</emphasis>
35
<emphasis id="0883EAC9FFE3FFC7FCBAFD0B9030FD49" bold="true" box="[813,821,723,745]" pageId="9" pageNumber="494">'</emphasis>
13
<emphasis id="0883EAC9FFE3FFC7FCD8FD0B9052FD49" bold="true" box="[847,855,723,745]" pageId="9" pageNumber="494">&quot;</emphasis>
N; 104
<emphasis id="0883EAC9FFE3FFC7FC36FD0C90ACFD49" bold="true" box="[929,937,724,745]" pageId="9" pageNumber="494">°</emphasis>
16
<emphasis id="0883EAC9FFE3FFC7FC54FD0B90CEFD49" bold="true" box="[963,971,723,745]" pageId="9" pageNumber="494">'</emphasis>
48
<emphasis id="0883EAC9FFE3FFC7FC72FD0B90E8FD49" bold="true" box="[997,1005,723,745]" pageId="9" pageNumber="494">&quot;</emphasis>
W)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE3FFC7FE87FD3691F8FCCE" pageId="9" pageNumber="494" type="biology_ecology">
<paragraph id="3A4836DBFFE3FFC7FE87FD3691F8FCCE" blockId="9.[246,1079,271,1172]" pageId="9" pageNumber="494">
<emphasis id="0883EAC9FFE3FFC7FE87FD3692C6FCA3" bold="true" box="[272,451,750,771]" pageId="9" pageNumber="494">Biological Data.</emphasis>
Specimens of
<taxonomicName id="FDF74D58FFE3FFC7FDF6FD3791C2FCA4" box="[609,711,751,772]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="joyana" status="sp. nov.">
<emphasis id="0883EAC9FFE3FFC7FDF6FD3791C2FCA4" box="[609,711,751,772]" italics="true" pageId="9" pageNumber="494">P. joyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE3FFC7FD58FD379004FCA4" box="[719,769,751,772]" pageId="9" pageNumber="494" rank="species">n.sp.</taxonomicNameLabel>
were collected at fluorescent white lights or UV lights in montane cloud forest located at
<quantity id="FD0F9B3EFFE3FFC7FC04FCD19733FCBE" box="[915,1078,777,798]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="9" pageNumber="494" unit="m" value="2050.0" valueMax="2200.0" valueMin="1900.0">
<geoCoordinate id="5FC3501CFFE3FFC7FC04FCD190C8FCBE" box="[915,973,777,798]" degrees="1,900" direction="north" orientation="latitude" pageId="9" pageNumber="494" precision="55" value="1.9">1,900</geoCoordinate>
<geoCoordinate id="5FC3501CFFE3FFC7FC5AFCD19712FCBE" box="[973,1047,777,798]" degrees="2,200" direction="west" orientation="longitude" pageId="9" pageNumber="494" precision="55" value="-2.2">2,200</geoCoordinate>
m
</quantity>
altitude, between 20:00 and 20:30 h. Phenology: July (3), August (2). Other species of
<taxonomicName id="FDF74D58FFE3FFC7FEFDFCE792F5FCF4" box="[362,496,831,852]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="0883EAC9FFE3FFC7FEFDFCE792F5FCF4" box="[362,496,831,852]" italics="true" pageId="9" pageNumber="494">Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and place were
<taxonomicName id="FDF74D58FFE3FFC7FC3EFCE7926FFCCF" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="jalisciensis" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFE3FFC7FC3EFCE7926FFCCF" italics="true" pageId="9" pageNumber="494">P. (Phytalus) jalisciensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE3FFC7FEE4FC8192A1FCCE" box="[371,420,857,878]" pageId="9" pageNumber="494" rank="species">n.sp.</taxonomicNameLabel>
and
<taxonomicName id="FDF74D58FFE3FFC7FE4BFC8291C7FCCF" box="[476,706,857,879]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="manantleca" status="sp. nov.">
<emphasis id="0883EAC9FFE3FFC7FE4BFC8292F6FCCF" box="[476,499,858,879]" italics="true" pageId="9" pageNumber="494">P.</emphasis>
(
<emphasis id="0883EAC9FFE3FFC7FD93FC829132FCCF" box="[516,567,858,879]" italics="true" pageId="9" pageNumber="494">s.str.</emphasis>
)
<emphasis id="0883EAC9FFE3FFC7FDDFFC8291C7FCCF" box="[584,706,858,879]" italics="true" pageId="9" pageNumber="494">manantleca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE3FFC7FD5CFC8191F8FCCE" box="[715,765,857,878]" pageId="9" pageNumber="494" rank="species">n.sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE3FFC7FE87FCAC91B1FBFE" pageId="9" pageNumber="494" type="discussion">
<paragraph id="3A4836DBFFE3FFC7FE87FCAC91B1FBFE" blockId="9.[246,1079,271,1172]" pageId="9" pageNumber="494">
<emphasis id="0883EAC9FFE3FFC7FE87FCAC9278FC29" bold="true" box="[272,381,884,905]" pageId="9" pageNumber="494">Remarks.</emphasis>
<taxonomicName id="FDF74D58FFE3FFC7FE1FFCAC9165FC29" box="[392,608,884,905]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="joyana" status="sp. nov.">
<emphasis id="0883EAC9FFE3FFC7FE1FFCAC9165FC29" box="[392,608,884,905]" italics="true" pageId="9" pageNumber="494">Phyllophaga joyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFE3FFC7FDFCFCAC9198FC29" box="[619,669,884,905]" pageId="9" pageNumber="494" rank="species">n.sp.</taxonomicNameLabel>
belongs to the species group
<emphasis id="0883EAC9FFE3FFC7FC78FCAC923FFC04" italics="true" pageId="9" pageNumber="494">
<taxonomicName id="FDF74D58FFE3FFC7FC6DFCAC923FFC04" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="blanchardi">blanchardi</taxonomicName>
</emphasis>
, complex
<taxonomicName id="FDF74D58FFE3FFC7FE5CFC579114FC04" box="[459,529,911,932]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="xanthe">
<emphasis id="0883EAC9FFE3FFC7FE5CFC579114FC04" box="[459,529,911,932]" italics="true" pageId="9" pageNumber="494">xanthe</emphasis>
</taxonomicName>
(
<emphasis id="0883EAC9FFE3FFC7FDA6FC57916EFC04" box="[561,619,911,932]" italics="true" pageId="9" pageNumber="494">sensu</emphasis>
<bibRefCitation id="5E664B2AFFE3FFC7FDE3FC5791FEFC04" author="Moron, M. A." bookContentInfo="Publ. 20, Instituto de Ecologia, Mexico. 341 pp." box="[628,763,911,932]" pageId="9" pageNumber="494" refId="ref10446" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica. (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico. 341 pp." title="El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica. (Insecta: Coleoptera)" type="book" year="1986">Morón 1986</bibRefCitation>
). The diagnostic combination of characters for male include: anterior border of clypeus deeply sinuated; frons and pronotum with dense vestiture of erect, long setae; abdominal sternites 2
<emphasis id="0883EAC9FFE3FFC7FEABFC069241FC53" bold="true" box="[316,324,990,1011]" pageId="9" pageNumber="494">°</emphasis>
to 4
<emphasis id="0883EAC9FFE3FFC7FEE1FC06927BFC53" bold="true" box="[374,382,990,1011]" pageId="9" pageNumber="494">°</emphasis>
with shallow mesial concavity; abdominal sternite 5
<emphasis id="0883EAC9FFE3FFC7FC38FC0690B2FC53" bold="true" box="[943,951,990,1011]" pageId="9" pageNumber="494">°</emphasis>
with scarce punctuation or vestiture; and pigydium vaguely punctate, without setae on disk. The shape of the paramera and aedeagus have some similarities with
<taxonomicName id="FDF74D58FFE3FFC7FC44FBCC9236FBE4" authority="Bates" authorityName="Bates" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="xanthe">
<emphasis id="0883EAC9FFE3FFC7FC44FBCC9733FB89" box="[979,1078,1044,1065]" italics="true" pageId="9" pageNumber="494">P. xanthe</emphasis>
Bates
</taxonomicName>
,
<taxonomicName id="FDF74D58FFE3FFC7FED6FBF79144FBE4" authority="Moser" authorityName="Moser" box="[321,577,1071,1092]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="verruciventris">
<emphasis id="0883EAC9FFE3FFC7FED6FBF792F1FBE4" box="[321,500,1071,1092]" italics="true" pageId="9" pageNumber="494">P. verruciventris</emphasis>
Moser
</taxonomicName>
and
<taxonomicName id="FDF74D58FFE3FFC7FDECFBF7903CFBE4" authority="Moser" authorityName="Moser" box="[635,825,1071,1092]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="494" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="0883EAC9FFE3FFC7FDECFBF791E9FBE4" box="[635,748,1071,1092]" italics="true" pageId="9" pageNumber="494">P. collaris</emphasis>
Moser
</taxonomicName>
, but the proportions or positions of each part are much different.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFE3FFC7FE87FBBC90B5FB34" pageId="9" pageNumber="494" type="etymology">
<paragraph id="3A4836DBFFE3FFC7FE87FBBC90B5FB34" blockId="9.[246,1079,271,1172]" pageId="9" pageNumber="494">
<emphasis id="0883EAC9FFE3FFC7FE87FBBC928AFBD9" bold="true" box="[272,399,1124,1145]" pageId="9" pageNumber="494">Etymology.</emphasis>
Derived from the name of Las Joyas creek, that flow near the middle of the Biological Station Las Joyas, Sierra de Manantlán.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,685 @@
<document id="4F093929A8080318382818D4E280D1E0" ID-DOI="10.1649/0010-065X(2001)055[0485:NSOPHC]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102835" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630156134743" checkinUser="tatiana" docAuthor="Morón, Miguel-Angel, Rivera-Cervantes, Luis Eugenio &amp; López-Vieyra, Mario" docDate="2001" docId="B25E87CDFFEEFFC9FF6EFAFD91BAFC99" docLanguage="en" docName="TheColeopBull.55.4.485-500.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docSource="http://dx.doi.org/10.1649/0010-065x(2001)055[0485:nsophc]2.0.co;2" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) manantleca Moron, Rivera y Lopez, new species" docType="treatment" docVersion="1" lastPageNumber="492" masterDocId="4E67FFB5FFEAFFCEFF97FFD89305FFA0" masterDocTitle="New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México" masterLastPageNumber="500" masterPageNumber="485" pageNumber="489" updateTime="1730145459161" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="F0E49E3528425B6910BD5F6280F8FFB1">New Species Of Phyllophaga Harris (Coleoptera: Melolonthidae, Melolonthinae) From Biosphere Reserve Of Manantlán, Jalisco-Colima, México</mods:title>
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<mods:name id="D568367587366DD4D70F4F48543A7427" type="personal">
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<mods:namePart id="19B34047558A2E226BC358D137A36A34">Morón, Miguel-Angel</mods:namePart>
<mods:affiliation id="BF07156DAD2DDCADAEA94BFE2A779778">Departamento de Entomología Instituto de Ecología, A. C. (Sector SEP­CONACYT) Apdo. Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
</mods:name>
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<mods:namePart id="D941B3B2C93D7AA654E978B82CAC66BF">Rivera-Cervantes, Luis Eugenio</mods:namePart>
<mods:affiliation id="47D763335519501A2AC1930D8F0E0F6C">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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</mods:role>
<mods:namePart id="570086470BBC389E3A4848D4A2B2D73E">López-Vieyra, Mario</mods:namePart>
<mods:affiliation id="4969925AA40426460EDBB8BF019A6692">Instituto Manantlán de Ecología y Conservación de la Biodiversidad Avenida Independencia Nacional 151 Autlán, Jalisco 48900, MÉXICO</mods:affiliation>
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<mods:date id="BE9532CE526EFACF4FE0985EF18DDCC9">2001</mods:date>
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<treatment id="B25E87CDFFEEFFC9FF6EFAFD91BAFC99" LSID="urn:lsid:plazi:treatment:B25E87CDFFEEFFC9FF6EFAFD91BAFC99" httpUri="http://treatment.plazi.org/id/B25E87CDFFEEFFC9FF6EFAFD91BAFC99" lastPageId="7" lastPageNumber="492" pageId="4" pageNumber="489">
<subSubSection id="72ED6550FFEEFFCAFF6EFAFD973CFA9A" box="[249,1081,1317,1338]" pageId="4" pageNumber="489" type="nomenclature">
<paragraph id="3A4836DBFFEEFFCAFF6EFAFD973CFA9A" blockId="4.[249,1081,1317,1365]" box="[249,1081,1317,1338]" pageId="4" pageNumber="489">
<taxonomicName id="FDF74D58FFEEFFCAFF6EFAFD90ACFA9A" authority="Moron, Rivera y Lopez" authorityName="Moron, Rivera y Lopez" box="[249,937,1317,1338]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="manantleca" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="0883EAC9FFEEFFCAFF6EFAFD91A4FA9A" box="[249,673,1317,1338]" italics="true" pageId="4" pageNumber="489">Phyllophaga (Phyllophaga) manantleca</emphasis>
Morón, Rivera y López
</taxonomicName>
,
<emphasis id="0883EAC9FFEEFFCAFC21FAFD973CFA9A" bold="true" box="[950,1081,1317,1338]" pageId="4" pageNumber="489">
<taxonomicNameLabel id="13B057B2FFEEFFCAFC21FAFD973CFA9A" box="[950,1081,1317,1338]" pageId="4" pageNumber="489" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="72ED6550FFEEFFC8FDF7FA989193FC29" lastPageId="6" lastPageNumber="491" pageId="4" pageNumber="489" type="description">
<paragraph id="3A4836DBFFEEFFCAFDF7FA9891D7FAF5" blockId="4.[249,1081,1317,1365]" box="[608,722,1344,1365]" pageId="4" pageNumber="489">
<figureCitation id="A2CC2A5EFFEEFFCAFDF7FA9891D7FAF5" box="[608,722,1344,1365]" captionStart="Figs" captionStartId="5.[269,312,891,910]" captionTargetBox="[252,1072,268,868]" captionTargetId="figure-12@5.[251,1073,265,870]" captionTargetPageId="5" captionText="Figs. 913. Phyllophaga manantleca. 9) tarsal claw, male; 10) genital capsule, lateral view; 11) paramera, distal view; 12) distal half of genital capsule, dorsal view; 13) ventral genital plates, female. Scale lines = 1 mm, except Fig. 9 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102839" httpUri="https://zenodo.org/record/10102839/files/figure.png" pageId="4" pageNumber="489">Figs. 913</figureCitation>
</paragraph>
<paragraph id="3A4836DBFFEEFFC8FE87FAB691F8FE69" blockId="4.[249,1083,1390,1652]" lastBlockId="6.[249,1082,270,457]" lastPageId="6" lastPageNumber="491" pageId="4" pageNumber="489">
<emphasis id="0883EAC9FFEEFFCAFE87FAB6926FFA21" bold="true" box="[272,362,1390,1409]" pageId="4" pageNumber="489">
<typeStatus id="E54C8879FFEEFFCAFE87FAB6926FFA21" box="[272,362,1390,1409]" pageId="4" pageNumber="489" type="holotype">Holotype</typeStatus>
</emphasis>
Male. Head, pronotum and elytra shiny reddish dark brown; pygidium, sterna, and legs shiny reddish light brown. Clypeus moderately bilobed, wider than long (4: 1), anterior border shallow and widely sinuated, with margins scarcely elevated, surface convex, with large, round punctures, nearly confluent, with erect, long, slender setae. Fronto­clypeal suture nearly straight and scarcely impressed. Frons wider than long (2.1: 1) convex, with large, round punctures and erect, long, slender setae. Antenna 10 segmented, with 3 segmented club; lamellae slightly longer than length of five preceeding segments combined (1.2:1), segment 4
<emphasis id="0883EAC9FFEEFFCAFDF3F9CF916EF98B" bold="true" box="[612,619,1559,1579]" pageId="4" pageNumber="489">°</emphasis>
nearly as long as segment 3, segment 5
<emphasis id="0883EAC9FFEEFFCAFC72F9CF90E9F98B" bold="true" box="[997,1004,1559,1579]" pageId="4" pageNumber="489">°</emphasis>
without anterior projections, segments 6
<emphasis id="0883EAC9FFEEFFCAFDB1F9F79128F9E3" bold="true" box="[550,557,1583,1603]" pageId="4" pageNumber="489">°</emphasis>
and 7
<emphasis id="0883EAC9FFEEFFCAFDFEF9F79175F9E3" bold="true" box="[617,624,1583,1603]" pageId="4" pageNumber="489">°</emphasis>
wider than long, each one with conical or acute process directed forward. Frons 6.2
<emphasis id="0883EAC9FFEEFFCAFDCFF990916FF9FB" bold="true" box="[600,618,1608,1627]" pageId="4" pageNumber="489">×</emphasis>
wider than each eye dorsal diameter. Canthus curved and rounded, with 9 setae. Labrum deeply bilobed, with curved, slender setae on all surface. Mentum widely concave, polished, with lateral slender setae, anterior border deeply cleft. Pronotum wider than long (1.8:1) and 2
<emphasis id="0883EAC9FFEFFFCBFD64FBD09000FBBB" bold="true" box="[755,773,1032,1051]" pageId="5" pageNumber="490">×</emphasis>
wider than frons. Pronotal disk rugo­punctate, punctures separated by one diameter or less, with many erect, long, slen­ der setae; lateral borders widely angulated, marginal bead vaguely crenulate, with long, slender setae; anterior angles slightly obtuse, rounded, scarcely prominent; posterior angles widely obtuse, not prominent. Scutellum 1.6
<emphasis id="0883EAC9FFEFFFCBFD5DFBB091D9FBDB" bold="true" box="[714,732,1128,1147]" pageId="5" pageNumber="490">×</emphasis>
wider than long, without punctures. Elytron 2.6
<emphasis id="0883EAC9FFEFFFCBFEF7FB589277FB33" bold="true" box="[352,370,1152,1171]" pageId="5" pageNumber="490">×</emphasis>
longer than wide, densely rugo­punctate, with some, scattered, short setae at basal third; epipleural border narrowed, extended along the complete margin, provided with a fringe of medium size setae; humeral calla rounded, prominent; apical calla round­ ed. Metathoracic wings completely developed. Propygidium shiny, finely punctate with abundant short setae. Pygidium convex, shiny, moderately punctate, with erect, medium size, slender setae near center of disk and short setae toward basal border; preapical area without setae; apical margin with 20 slender setae; basal margin effaced at middle. Pterosternon with long, dense, yellowish vestiture. Visible abdominal sternites 2
<emphasis id="0883EAC9FFEFFFCBFC6EFAFF9705FA9B" bold="true" box="[1017,1024,1319,1339]" pageId="5" pageNumber="490">°</emphasis>
to 4
<emphasis id="0883EAC9FFEFFFCBFBB8FAFF9733FA9B" bold="true" box="[1071,1078,1319,1339]" pageId="5" pageNumber="490">°</emphasis>
with narrow and deep mesial sulcus; sternite 5
<emphasis id="0883EAC9FFEFFFCBFD3EFAE791B5FAF3" bold="true" box="[681,688,1343,1363]" pageId="5" pageNumber="490">°</emphasis>
convex, with patch of setiferous granules and posterior border slightly excavated transversally; anal plate wide and shallowly concave, finely granulose­setose, anterior and posterior borders thickened, with scattered erect, long setae on posterior border. Protibiae slightly shorter than protarsi (0.84:1), with two large teeth and one small tooth on external border, preapical spur long, straight, acute, shorter than 2
<emphasis id="0883EAC9FFEFFFCBFE23FA6F92BEFA6B" bold="true" box="[436,443,1463,1483]" pageId="5" pageNumber="490">°</emphasis>
protarsomerus (1.6:2). Mesotibiae with long, slender setae, and one oblique, short, setiferous, transverse carina; upper apical spur straight, narrow, nearly as long as lower spur. Metatibiae shorter than metatarsi (1:1.2), with long, slender setae, and one oblique, short setiferous transverse carina; apical spurs articulated with border, upper spur lanceolate, slightly curved, with rounded apex, nearly as long as 2
<emphasis id="0883EAC9FFEFFFCBFC4CF9CF90E7F98B" bold="true" box="[987,994,1559,1579]" pageId="5" pageNumber="490">°</emphasis>
metatarsomerus, and 1.2
<emphasis id="0883EAC9FFEFFFCBFE0FF9E892AFF9E3" bold="true" box="[408,426,1584,1603]" pageId="5" pageNumber="490">×</emphasis>
longer than lower spur. Tarsomeres semicylindrical, elongated, with enlarged apex, and many setae around the apex; mesotarsomeres 2
<emphasis id="0883EAC9FFEFFFCBFCC8F99F9063F9FB" bold="true" box="[863,870,1607,1627]" pageId="5" pageNumber="490">°</emphasis>
to 5
<emphasis id="0883EAC9FFEFFFCBFC18F99F9093F9FB" bold="true" box="[911,918,1607,1627]" pageId="5" pageNumber="490">°</emphasis>
ventrally present one line of scattered, slender setae; metatarsomeres 2
<emphasis id="0883EAC9FFEFFFCBFD62F98791F9F9D3" bold="true" box="[757,764,1631,1651]" pageId="5" pageNumber="490">°</emphasis>
to 4
<emphasis id="0883EAC9FFEFFFCBFCBDF9879034F9D3" bold="true" box="[810,817,1631,1651]" pageId="5" pageNumber="490">°</emphasis>
ventrally present two lines of setae. Tarsal claws symmetrical, similar on all legs, with median acute­angled tooth located near basal dilatation, but also near apical tooth (
<figureCitation id="A2CC2A5EFFECFFC8FCB6FEFE905AFE99" box="[801,863,294,313]" captionStart="Figs" captionStartId="5.[269,312,891,910]" captionTargetBox="[252,1072,268,868]" captionTargetId="figure-12@5.[251,1073,265,870]" captionTargetPageId="5" captionText="Figs. 913. Phyllophaga manantleca. 9) tarsal claw, male; 10) genital capsule, lateral view; 11) paramera, distal view; 12) distal half of genital capsule, dorsal view; 13) ventral genital plates, female. Scale lines = 1 mm, except Fig. 9 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102839" httpUri="https://zenodo.org/record/10102839/files/figure.png" pageId="6" pageNumber="491">Fig. 9</figureCitation>
). Genital capsule with long, narrowed, angled paramera, fused dorsally at their basis, apex angled, with preapical short projections ended in slightly elbowed, ovate structure (
<figureCitation id="A2CC2A5EFFECFFC8FCF8FE8E90E6FEC9" box="[879,995,342,361]" captionStart="Figs" captionStartId="5.[269,312,891,910]" captionTargetBox="[252,1072,268,868]" captionTargetId="figure-12@5.[251,1073,265,870]" captionTargetPageId="5" captionText="Figs. 913. Phyllophaga manantleca. 9) tarsal claw, male; 10) genital capsule, lateral view; 11) paramera, distal view; 12) distal half of genital capsule, dorsal view; 13) ventral genital plates, female. Scale lines = 1 mm, except Fig. 9 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102839" httpUri="https://zenodo.org/record/10102839/files/figure.png" pageId="6" pageNumber="491">Figs. 1012</figureCitation>
). Tectum convex with widely sinuated distal border. Aedeagus long, wide, with sclerotized tubelike support and preapical, dorsal, curved, sharply pointed, sclerotized plate (
<figureCitation id="A2CC2A5EFFECFFC8FC49FE5E9216FE11" captionStart="Figs" captionStartId="5.[269,312,891,910]" captionTargetBox="[252,1072,268,868]" captionTargetId="figure-12@5.[251,1073,265,870]" captionTargetPageId="5" captionText="Figs. 913. Phyllophaga manantleca. 9) tarsal claw, male; 10) genital capsule, lateral view; 11) paramera, distal view; 12) distal half of genital capsule, dorsal view; 13) ventral genital plates, female. Scale lines = 1 mm, except Fig. 9 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102839" httpUri="https://zenodo.org/record/10102839/files/figure.png" pageId="6" pageNumber="491">Figs. 10 12</figureCitation>
). Length of genital capsule from apex of parameres to border of basal piece:
<quantity id="FD0F9B3EFFECFFC8FC7BFE469733FE11" box="[1004,1078,414,433]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="491" unit="mm" value="2.6">2.6 mm</quantity>
. Total body length:
<quantity id="FD0F9B3EFFECFFC8FE38FE6E9103FE69" box="[431,518,438,457]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="6" pageNumber="491" unit="mm" value="13.1">13.1 mm</quantity>
. Humeral width:
<quantity id="FD0F9B3EFFECFFC8FD3AFE6E91FCFE69" box="[685,761,438,457]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="6" pageNumber="491" unit="mm" value="5.6">5.6 mm</quantity>
.
</paragraph>
<caption id="6E886653FFEFFFCBFE9AFCA3908DFC1E" ID-DOI="http://doi.org/10.5281/zenodo.10102839" ID-Zenodo-Dep="10102839" httpUri="https://zenodo.org/record/10102839/files/figure.png" pageId="5" pageNumber="490" startId="5.[269,312,891,910]" targetBox="[252,1072,268,868]" targetPageId="5">
<paragraph id="3A4836DBFFEFFFCBFE9AFCA3908DFC1E" blockId="5.[246,1078,891,958]" pageId="5" pageNumber="490">
<emphasis id="0883EAC9FFEFFFCBFE9AFCA39273FC2E" bold="true" box="[269,374,891,910]" pageId="5" pageNumber="490">Figs. 913.</emphasis>
<emphasis id="0883EAC9FFEFFFCBFE1AFCA39178FC2E" box="[397,637,891,910]" italics="true" pageId="5" pageNumber="490">
<taxonomicName id="FDF74D58FFEFFFCBFE1AFCA3917CFC2E" box="[397,633,891,910]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="490" phylum="Arthropoda" rank="species" species="manantleca">Phyllophaga manantleca</taxonomicName>
.
</emphasis>
<emphasis id="0883EAC9FFEFFFCBFD15FCA39190FC2E" bold="true" box="[642,661,891,910]" pageId="5" pageNumber="490">9)</emphasis>
tarsal claw, male;
<emphasis id="0883EAC9FFEFFFCBFCD4FCA39067FC2E" bold="true" box="[835,866,891,910]" pageId="5" pageNumber="490">10)</emphasis>
genital capsule, lateral view;
<emphasis id="0883EAC9FFEFFFCBFEB8FC4B924BFC06" bold="true" box="[303,334,915,934]" pageId="5" pageNumber="490">11)</emphasis>
paramera, distal view;
<emphasis id="0883EAC9FFEFFFCBFDB0FC4B9143FC06" bold="true" box="[551,582,915,934]" pageId="5" pageNumber="490">12)</emphasis>
distal half of genital capsule, dorsal view;
<emphasis id="0883EAC9FFEFFFCBFC45FC4B90F4FC06" bold="true" box="[978,1009,915,934]" pageId="5" pageNumber="490">13)</emphasis>
ventral genital plates, female. Scale lines
<emphasis id="0883EAC9FFEFFFCBFDACFC739148FC1E" bold="true" box="[571,589,939,958]" pageId="5" pageNumber="490">=</emphasis>
1 mm, except Fig. 9
<emphasis id="0883EAC9FFEFFFCBFC89FC739035FC1E" bold="true" box="[798,816,939,958]" pageId="5" pageNumber="490">=</emphasis>
0.5 mm.
</paragraph>
</caption>
<paragraph id="3A4836DBFFECFFC8FE84FE3C913EFCA4" blockId="6.[249,1082,484,1652]" pageId="6" pageNumber="491">
<emphasis id="0883EAC9FFECFFC8FE84FE3C927CFE59" bold="true" box="[275,377,484,505]" pageId="6" pageNumber="491">
<typeStatus id="E54C8879FFECFFC8FE84FE3C9276FE59" box="[275,371,484,505]" pageId="6" pageNumber="491" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Similar to the male except as follows: antennal club 0.9 shorter than the five preceeding segments combined; punctuation of frons and pronotum slightly more coarse; apex of pygidium slightly prominent; visible abdominal sternites 2
<emphasis id="0883EAC9FFECFFC8FE48FDEC92E2FDE9" bold="true" box="[479,487,564,585]" pageId="6" pageNumber="491">°</emphasis>
to 4
<emphasis id="0883EAC9FFECFFC8FD8AFDEC9120FDE9" bold="true" box="[541,549,564,585]" pageId="6" pageNumber="491">°</emphasis>
with mesial sulcus vaguely indicated; sternite 5
<emphasis id="0883EAC9FFECFFC8FBA6FDEC973CFDE9" bold="true" box="[1073,1081,564,585]" pageId="6" pageNumber="491">°</emphasis>
convex, punctate, with scattered short setae; anal plate convex, punctate, with scattered, short and medium size setae. Protibia with external border clearly tridentate. Apical spurs of metatibiae wide, slightly curvated with rounded apexes. Ventral genital plates ovate, moderately sclerotized, nearly symmetrical, convex, with some short setae near acute, apical border; dorsal genital plates ovate, with borders scarcely setose (
<figureCitation id="A2CC2A5EFFECFFC8FD47FD0C9027FD49" box="[720,802,724,745]" captionStart="Figs" captionStartId="5.[269,312,891,910]" captionTargetBox="[252,1072,268,868]" captionTargetId="figure-12@5.[251,1073,265,870]" captionTargetPageId="5" captionText="Figs. 913. Phyllophaga manantleca. 9) tarsal claw, male; 10) genital capsule, lateral view; 11) paramera, distal view; 12) distal half of genital capsule, dorsal view; 13) ventral genital plates, female. Scale lines = 1 mm, except Fig. 9 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10102839" httpUri="https://zenodo.org/record/10102839/files/figure.png" pageId="6" pageNumber="491">Fig. 13</figureCitation>
). Total body length:
<quantity id="FD0F9B3EFFECFFC8FB9CFD0C9220FCA4" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="6" pageNumber="491" unit="mm" value="14.1">14.1 mm</quantity>
. Humeral width:
<quantity id="FD0F9B3EFFECFFC8FE76FD379132FCA4" box="[481,567,751,772]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="491" unit="mm" value="6.1">6.1 mm</quantity>
.
</paragraph>
<paragraph id="3A4836DBFFECFFC8FE84FCD19193FC29" blockId="6.[249,1082,484,1652]" pageId="6" pageNumber="491">
<emphasis id="0883EAC9FFECFFC8FE84FCD192F1FCBE" bold="true" box="[275,500,777,798]" pageId="6" pageNumber="491">
<typeStatus id="E54C8879FFECFFC8FE84FCD1927CFCBE" box="[275,377,777,798]" pageId="6" pageNumber="491" type="paratype">Paratype</typeStatus>
Variation.
</emphasis>
Males are similar to the
<typeStatus id="E54C8879FFECFFC8FC95FCD19058FCBE" box="[770,861,777,798]" pageId="6" pageNumber="491" type="holotype">holotype</typeStatus>
except in total body length:
<quantity id="FD0F9B3EFFECFFC8FEDEFCFC92E1FC99" box="[329,484,804,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.265" metricValueMax="1.35" metricValueMin="1.18" pageId="6" pageNumber="491" unit="mm" value="12.65" valueMax="13.5" valueMin="11.8">11.813.5 mm</quantity>
, humeral width:
<quantity id="FD0F9B3EFFECFFC8FD0FFCFC901FFC99" box="[664,794,804,825]" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="5.8" metricValueMin="5.1" pageId="6" pageNumber="491" unit="mm" value="5.449999999999999" valueMax="5.8" valueMin="5.1">5.15.8 mm</quantity>
. Female
<typeStatus id="E54C8879FFECFFC8FCEEFCFC90D8FC99" box="[889,989,804,825]" pageId="6" pageNumber="491" type="paratype">paratypes</typeStatus>
are similar to the
<typeStatus id="E54C8879FFECFFC8FEFAFCE792C4FCF4" box="[365,449,831,852]" pageId="6" pageNumber="491" type="allotype">allotype</typeStatus>
except as follows: body color dark reddish brown to dark yellowish brown; pygidium more or less setiferous; total body length:
<quantity id="FD0F9B3EFFECFFC8FC69FC819259FC29" metricMagnitude="-2" metricUnit="m" metricValue="1.4149999999999998" metricValueMax="1.51" metricValueMin="1.3199999999999998" pageId="6" pageNumber="491" unit="mm" value="14.149999999999999" valueMax="15.1" valueMin="13.2">13.2 15.1 mm</quantity>
; humeral width: 6.0
<quantity id="FD0F9B3EFFECFFC8FDD4FCAC9193FC29" box="[579,662,884,905]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="491" unit="mm" value="6.5">6.5 mm</quantity>
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFECFFC8FE84FC569724F9D4" pageId="6" pageNumber="491" type="materials_examined">
<paragraph id="3A4836DBFFECFFC8FE84FC5690D3F99E" blockId="6.[249,1082,484,1652]" pageId="6" pageNumber="491">
<emphasis id="0883EAC9FFECFFC8FE84FC5692A4FC03" bold="true" box="[275,417,910,931]" pageId="6" pageNumber="491">Type Series.</emphasis>
(Described from
<specimenCount id="2CF1FD52FFECFFC8FDFFFC5791D2FC04" box="[616,727,911,932]" pageId="6" pageNumber="491" type="male">189 males</specimenCount>
and
<specimenCount id="2CF1FD52FFECFFC8FC83FC579099FC04" box="[788,924,911,932]" pageId="6" pageNumber="491" type="female">243 females</specimenCount>
).
<materialsCitation id="8A9F3C86FFECFFC8FC25FC5791B7FC79" collectingDate="1997-10-02" collectorName="D. Pelayo" country="Mexico" county="Autlan" elevation="1900" location="Alt." municipality="Est. Las Joyas" pageId="6" pageNumber="491" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<typeStatus id="E54C8879FFECFFC8FC25FC579716FC04" box="[946,1043,911,932]" pageId="6" pageNumber="491" type="holotype">Holotype</typeStatus>
♂;
<collectingCountry id="42E0764BFFECFFC8FF6EFC719261FC1E" box="[249,356,937,958]" name="Mexico" pageId="6" pageNumber="491">MEXICO</collectingCountry>
:
<emphasis id="0883EAC9FFECFFC8FEE6FC7292C4FC1F" box="[369,449,938,959]" italics="true" pageId="6" pageNumber="491">
<collectingRegion id="F833F839FFECFFC8FEE6FC7292BBFC1F" box="[369,446,938,959]" country="Mexico" name="Jalisco" pageId="6" pageNumber="491">Jalisco</collectingRegion>
,
</emphasis>
<collectingCounty id="D3294E57FFECFFC8FE5CFC719110FC1E" box="[459,533,937,958]" pageId="6" pageNumber="491">Autlán</collectingCounty>
,
<collectingMunicipality id="DA2CACA1FFECFFC8FDB5FC7191BAFC1E" box="[546,703,937,958]" pageId="6" pageNumber="491">Est. Las Joyas</collectingMunicipality>
, bosque mesófilo,
<location id="3F286000FFECFFC8FC1DFC7190B7FC1E" LSID="urn:lsid:plazi:treatment:B25E87CDFFEEFFC9FF6EFAFD91BAFC99:3F286000FFECFFC8FC1DFC7190B7FC1E" box="[906,946,937,958]" country="Mexico" county="Autlan" municipality="Est. Las Joyas" name="Alt." pageId="6" pageNumber="491" stateProvince="Jalisco">Alt.</location>
<quantity id="FD0F9B3EFFECFFC8FC2CFC719713FC1E" box="[955,1046,937,958]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="491" unit="m" value="1900.0">
<elevation id="B1DAD1E8FFECFFC8FC2CFC719713FC1E" box="[955,1046,937,958]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="491" unit="m" value="1900.0">1,900 m</elevation>
</quantity>
,
<date id="4E49101BFFECFFC8FBB4FC719249FC79" pageId="6" pageNumber="491" value="1997-10-02">
<collectingDate id="5E0DE9F3FFECFFC8FBB4FC719249FC79" pageId="6" pageNumber="491" value="1997-10-02">2­ X­1997</collectingDate>
</date>
, trampa luz,
<collectorName id="9702530DFFECFFC8FE4BFC1C9143FC79" box="[476,582,964,985]" pageId="6" pageNumber="491">D. Pelayo</collectorName>
(
<collectionCode id="5CE6AE1EFFECFFC8FDCFFC1C91AEFC79" box="[600,683,964,985]" pageId="6" pageNumber="491">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFECFFC8FD28FC1C925FFC54" collectingDate="1997-10-02" collectorName="D. Pelayo" country="Mexico" county="Autlan" elevation="1900" location="Alt." municipality="Est. Las Joyas" pageId="6" pageNumber="491" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="allotype">
<typeStatus id="E54C8879FFECFFC8FD28FC1C901FFC79" box="[703,794,964,985]" pageId="6" pageNumber="491" type="allotype">Allotype</typeStatus>
<specimenCount id="2CF1FD52FFECFFC8FCB3FC1B903DFC79" box="[804,824,963,985]" pageId="6" pageNumber="491" type="female"></specimenCount>
; same data as holotype (
<collectionCode id="5CE6AE1EFFECFFC8FE97FC079256FC54" box="[256,339,991,1012]" pageId="6" pageNumber="491">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8A9F3C86FFECFFC8FEFFFC079074FC54" box="[360,881,989,1012]" collectingDate="1997-10-02" collectorName="D. Pelayo" country="Mexico" county="Autlan" elevation="1900" location="Alt." municipality="Est. Las Joyas" pageId="6" pageNumber="491" specimenCount="3" specimenCount-female="3" stateProvince="Jalisco" typeStatus="paratype">
<typeStatus id="E54C8879FFECFFC8FEFFFC0792D7FC54" box="[360,466,991,1012]" pageId="6" pageNumber="491" type="paratype">Paratypes</typeStatus>
: same data as holotype (4 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FCBFFC079062FC53" box="[808,871,989,1012]" pageId="6" pageNumber="491" type="female">3 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFECFFC8FC17FC0790ACFBAE" collectingDate="1997-08-05" collectorName="Nunez &amp; Vazquez &amp; J. Garcia" country="Mexico" county="Autlan" elevation="1900" location="Alt." municipality="Est. Las Joyas" pageId="6" pageNumber="491" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
same data except
<date id="4E49101BFFECFFC8FF6EFC21927EFBAE" box="[249,379,1017,1038]" pageId="6" pageNumber="491" value="1997-08-05">
<collectingDate id="5E0DE9F3FFECFFC8FF6EFC21927EFBAE" box="[249,379,1017,1038]" pageId="6" pageNumber="491" value="1997-08-05">5­VIII­1997</collectingDate>
</date>
, S.
<collectorName id="9702530DFFECFFC8FE3DFC2192F4FBAE" box="[426,497,1017,1038]" pageId="6" pageNumber="491">Núñez</collectorName>
, (♂); A.
<collectorName id="9702530DFFECFFC8FDCBFC2191B2FBAE" box="[604,695,1017,1038]" pageId="6" pageNumber="491">Vázquez</collectorName>
(♂);
<collectorName id="9702530DFFECFFC8FD6DFC219059FBAE" box="[762,860,1017,1038]" pageId="6" pageNumber="491">J. García</collectorName>
(♂
<emphasis id="0883EAC9FFECFFC8FC13FC21908EFBAE" bold="true" box="[900,907,1017,1038]" pageId="6" pageNumber="491">/</emphasis>
<specimenCount id="2CF1FD52FFECFFC8FC1CFC20909AFBAE" box="[907,927,1016,1038]" pageId="6" pageNumber="491" type="female"></specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFECFFC8FC2DFC21973CFBAE" box="[954,1081,1017,1038]" pageId="6" pageNumber="491" value="1997-08-06">6­VIII­1997</date>
(♂
<emphasis id="0883EAC9FFECFFC8FE81FBCC9218FB89" bold="true" box="[278,285,1044,1065]" pageId="6" pageNumber="491">/</emphasis>
<specimenCount id="2CF1FD52FFECFFC8FE8AFBCB9234FB89" box="[285,305,1043,1065]" pageId="6" pageNumber="491" type="female"></specimenCount>
); A. Partida (3 ♂♂);
<date id="4E49101BFFECFFC8FD84FBCC9193FB89" box="[531,662,1044,1065]" pageId="6" pageNumber="491" value="1997-08-07">7­VIII­1997</date>
, S. Núñez (
<specimenCount id="2CF1FD52FFECFFC8FC98FBCB9026FB89" box="[783,803,1043,1065]" pageId="6" pageNumber="491" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFECFFC8FCADFBCC9262FBFE" collectingDate="1997-09-04" collectorName="A. Partida &amp; J. Garcia &amp; Guerra &amp; Lopez" country="Mexico" location="Lopez" municipality="Vazquez" pageId="6" pageNumber="491" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectorName id="9702530DFFECFFC8FCADFBCC90A1FB89" box="[826,932,1044,1065]" pageId="6" pageNumber="491">A. Partida</collectorName>
(♂);
<collectorName id="9702530DFFECFFC8FC4AFBCC973FFB89" box="[989,1082,1044,1065]" pageId="6" pageNumber="491">J. García</collectorName>
(♂); A.
<collectingMunicipality id="DA2CACA1FFECFFC8FED8FBF792AFFBE4" box="[335,426,1071,1092]" pageId="6" pageNumber="491">Vázquez</collectingMunicipality>
(♂); M.
<location id="3F286000FFECFFC8FD9CFBF79149FBE4" LSID="urn:lsid:plazi:treatment:B25E87CDFFEEFFC9FF6EFAFD91BAFC99:3F286000FFECFFC8FD9CFBF79149FBE4" box="[523,588,1071,1092]" country="Mexico" municipality="Vazquez" name="Lopez" pageId="6" pageNumber="491">López</location>
(8 ♂♂);
<date id="4E49101BFFECFFC8FD26FBF79027FBE4" box="[689,802,1071,1092]" pageId="6" pageNumber="491" value="1997-09-04">
<collectingDate id="5E0DE9F3FFECFFC8FD26FBF79027FBE4" box="[689,802,1071,1092]" pageId="6" pageNumber="491" value="1997-09-04">4­IX­1997</collectingDate>
</date>
, A.
<collectorName id="9702530DFFECFFC8FCD8FBF7909DFBE4" box="[847,920,1071,1092]" pageId="6" pageNumber="491">Guerra</collectorName>
(♂); M.
<collectorName id="9702530DFFECFFC8FC6FFBF7973CFBE4" box="[1016,1081,1071,1092]" pageId="6" pageNumber="491">López</collectorName>
(2 ♂
<specimenCount id="2CF1FD52FFECFFC8FEA3FB919258FBFE" box="[308,349,1096,1118]" pageId="6" pageNumber="491" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFECFFC8FEE1FB9192E2FBFE" box="[374,487,1097,1118]" pageId="6" pageNumber="491" value="1997-09-05">5­IX­1997</date>
, A.Guerra (♂
<emphasis id="0883EAC9FFECFFC8FDEDFB919184FBFE" bold="true" box="[634,641,1097,1118]" pageId="6" pageNumber="491">/</emphasis>
<specimenCount id="2CF1FD52FFECFFC8FD15FB909193FBFE" box="[642,662,1096,1118]" pageId="6" pageNumber="491" type="female"></specimenCount>
); M. López (1 ♂
<specimenCount id="2CF1FD52FFECFFC8FCCFFB919090FBFE" box="[856,917,1096,1118]" pageId="6" pageNumber="491" type="female">2 ♀♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFECFFC8FC39FB91926AFBD9" country="Mexico" location="Lopez" pageId="6" pageNumber="491" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
M.
<location id="3F286000FFECFFC8FC44FB919712FBFE" LSID="urn:lsid:plazi:treatment:B25E87CDFFEEFFC9FF6EFAFD91BAFC99:3F286000FFECFFC8FC44FB919712FBFE" box="[979,1047,1097,1118]" country="Mexico" name="Lopez" pageId="6" pageNumber="491">López</location>
, (3 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FEBFFBBC9260FBD9" box="[296,357,1123,1145]" pageId="6" pageNumber="491" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFECFFC8FEEBFBBC92E7FBD9" box="[380,482,1124,1145]" pageId="6" pageNumber="491" value="1997-10-01">1­X­1997</date>
(8 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FDA2FBBC9174FBD9" box="[565,625,1123,1145]" pageId="6" pageNumber="491" type="female">2 ♀♀</specimenCount>
); D. Pelayo (
<specimenCount id="2CF1FD52FFECFFC8FC97FBBB9011FBD9" box="[768,788,1123,1145]" pageId="6" pageNumber="491" type="female"></specimenCount>
);
<date id="4E49101BFFECFFC8FCBDFBBC909CFBD9" box="[810,921,1124,1145]" pageId="6" pageNumber="491" value="1997-11-04">4­XI­1997</date>
(♂); M. López (♂);
<date id="4E49101BFFECFFC8FEB9FBA7929BFB34" box="[302,414,1151,1172]" pageId="6" pageNumber="491" value="1997-11-06">6­XI­1997</date>
, D. Pelayo (♂
<emphasis id="0883EAC9FFECFFC8FDAFFBA6913AFB33" bold="true" box="[568,575,1150,1171]" pageId="6" pageNumber="491">/</emphasis>
<specimenCount id="2CF1FD52FFECFFC8FDA8FBA59156FB33" box="[575,595,1149,1171]" pageId="6" pageNumber="491" type="female"></specimenCount>
); M. López (♂);
<date id="4E49101BFFECFFC8FC99FBA79080FB34" box="[782,901,1151,1172]" pageId="6" pageNumber="491" value="1997-12-04">4­XII­1997</date>
(2 ♂♂);
<date id="4E49101BFFECFFC8FC7EFBA79235FB0E" pageId="6" pageNumber="491" value="1999-07-29">29­VII­ 1999</date>
, J. García (♂);
<date id="4E49101BFFECFFC8FE76FB419162FB0E" box="[481,615,1177,1198]" pageId="6" pageNumber="491" value="1999-07-30">30­VII­1999</date>
, V. González (♂); J. García (♂); G. Mesa (
<specimenCount id="2CF1FD52FFECFFC8FE95FB6B9213FB69" box="[258,278,1203,1225]" pageId="6" pageNumber="491" type="female"></specimenCount>
); M. López (
<specimenCount id="2CF1FD52FFECFFC8FE3FFB6B92B9FB69" box="[424,444,1203,1225]" pageId="6" pageNumber="491" type="female"></specimenCount>
);
<date id="4E49101BFFECFFC8FE42FB6C915CFB69" box="[469,601,1204,1225]" pageId="6" pageNumber="491" value="1999-07-31">31­VII­1999</date>
(
<specimenCount id="2CF1FD52FFECFFC8FDFCFB6B917AFB69" box="[619,639,1203,1225]" pageId="6" pageNumber="491" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFECFFC8FD00FB6C9156FB44" collectingDate="1986-06-18" collectorName="V. Gonzalez &amp; L. Rivera" country="Mexico" location="Lopez" pageId="6" pageNumber="491" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectorName id="9702530DFFECFFC8FD00FB6C901CFB69" box="[663,793,1204,1225]" pageId="6" pageNumber="491">V. González</collectorName>
(♂); same data except
<date id="4E49101BFFECFFC8FB81FB6C9250FB44" pageId="6" pageNumber="491" value="1986-06-18">
<collectingDate id="5E0DE9F3FFECFFC8FB81FB6C9250FB44" pageId="6" pageNumber="491" value="1986-06-18">18­ VI­1986</collectingDate>
</date>
,
<collectorName id="9702530DFFECFFC8FEF4FB1792C8FB44" box="[355,461,1231,1252]" pageId="6" pageNumber="491">L. Rivera</collectorName>
, (1 ♂
<specimenCount id="2CF1FD52FFECFFC8FD8AFB17914CFB43" box="[541,585,1229,1252]" pageId="6" pageNumber="491" type="female">2 ♀</specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFECFFC8FDF3FB1791D4FB44" box="[612,721,1231,1252]" pageId="6" pageNumber="491" value="1987-08">VIII­1987</date>
, (
<specimenCount id="2CF1FD52FFECFFC8FD7EFB1591F8FB43" box="[745,765,1229,1251]" pageId="6" pageNumber="491" type="female"></specimenCount>
);
<date id="4E49101BFFECFFC8FC8FFB179090FB44" box="[792,917,1231,1252]" pageId="6" pageNumber="491" value="1985-11-17">17­XI­1985</date>
, bosque pinos, (♂
<emphasis id="0883EAC9FFECFFC8FE81FB319218FB5E" bold="true" box="[278,285,1257,1278]" pageId="6" pageNumber="491">/</emphasis>
<specimenCount id="2CF1FD52FFECFFC8FE8AFB309234FB5E" box="[285,305,1256,1278]" pageId="6" pageNumber="491" type="female"></specimenCount>
);
<materialsCitation id="8A9F3C86FFECFFC8FEDCFB31928BFAB9" collectingDate="1994-09-12" collectorName="Nogueira" country="Mexico" elevation="1650" location="Alt." municipality="Sierra de Manantlan" pageId="6" pageNumber="491" specimenCount="2" specimenCount-female="2" stateProvince="Jalisco" typeStatus="paratype">
<collectingRegion id="F833F839FFECFFC8FEDCFB319290FB5E" box="[331,405,1257,1278]" country="Mexico" name="Jalisco" pageId="6" pageNumber="491">Jalisco</collectingRegion>
,
<collectingMunicipality id="DA2CACA1FFECFFC8FE35FB319185FB5E" box="[418,640,1257,1278]" pageId="6" pageNumber="491">Sierra de Manantlán</collectingMunicipality>
,
<location id="3F286000FFECFFC8FD19FB3191B3FB5E" LSID="urn:lsid:plazi:treatment:B25E87CDFFEEFFC9FF6EFAFD91BAFC99:3F286000FFECFFC8FD19FB3191B3FB5E" box="[654,694,1257,1278]" country="Mexico" municipality="Sierra de Manantlan" name="Alt." pageId="6" pageNumber="491" stateProvince="Jalisco">Alt.</location>
<quantity id="FD0F9B3EFFECFFC8FD57FB31901EFB5E" box="[704,795,1257,1278]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="491" unit="m" value="1650.0">
<elevation id="B1DAD1E8FFECFFC8FD57FB31901EFB5E" box="[704,795,1257,1278]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="491" unit="m" value="1650.0">1,650 m</elevation>
</quantity>
,
<date id="4E49101BFFECFFC8FCBEFB3190A2FB5E" box="[809,935,1257,1278]" pageId="6" pageNumber="491" value="1994-09-12">
<collectingDate id="5E0DE9F3FFECFFC8FCBEFB3190A2FB5E" box="[809,935,1257,1278]" pageId="6" pageNumber="491" value="1994-09-12">12­IX­1994</collectingDate>
</date>
, G.
<collectorName id="9702530DFFECFFC8FC4FFB31973CFB5E" box="[984,1081,1257,1278]" pageId="6" pageNumber="491">Nogueira</collectorName>
(3 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FED0FADC9281FAB9" box="[327,388,1283,1305]" pageId="6" pageNumber="491" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFECFFC8FE0BFADC90E5FAB9" box="[412,992,1283,1305]" collectingDate="1993-07-17" collectorName="Nogueira" country="Mexico" elevation="1500" location="Alt." municipality="Sierra de Manantlan" pageId="6" pageNumber="491" specimenCount="2" specimenCount-female="2" stateProvince="Jalisco" typeStatus="paratype">
same data except
<date id="4E49101BFFECFFC8FDCDFADC91E4FAB9" box="[602,737,1284,1305]" pageId="6" pageNumber="491" value="1993-07-17">
<collectingDate id="5E0DE9F3FFECFFC8FDCDFADC91E4FAB9" box="[602,737,1284,1305]" pageId="6" pageNumber="491" value="1993-07-17">17­VII­1993</collectingDate>
</date>
,
<quantity id="FD0F9B3EFFECFFC8FD7AFADC9046FAB9" box="[749,835,1284,1305]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="491" unit="m" value="1500.0">
<elevation id="B1DAD1E8FFECFFC8FD7AFADC9046FAB9" box="[749,835,1284,1305]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="491" unit="m" value="1500.0">1,500 m</elevation>
</quantity>
(3 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FC0EFADC90D3FAB9" box="[921,982,1283,1305]" pageId="6" pageNumber="491" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFECFFC8FC79FADC9118FA94" collectingDate="1994-08-02" collectorName="Nogueira" country="Mexico" elevation="1470" isTailEnumeration="true" location="Alt." municipality="Sierra de Manantlan" pageId="6" pageNumber="491" specimenCount="3" specimenCount-female="3" stateProvince="Jalisco" typeStatus="paratype">
<date id="4E49101BFFECFFC8FC79FADC9235FA94" pageId="6" pageNumber="491" value="1994-08-02">
<collectingDate id="5E0DE9F3FFECFFC8FC79FADC9235FA94" pageId="6" pageNumber="491" value="1994-08-02">2­VIII­ 1994</collectingDate>
</date>
,
<quantity id="FD0F9B3EFFECFFC8FEABFAC79296FA94" box="[316,403,1311,1332]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="6" pageNumber="491" unit="m" value="1470.0">
<elevation id="B1DAD1E8FFECFFC8FEABFAC79296FA94" box="[316,403,1311,1332]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="6" pageNumber="491" unit="m" value="1470.0">1,470 m</elevation>
</quantity>
(1 ♂
<specimenCount id="2CF1FD52FFECFFC8FE41FAC79116FA93" box="[470,531,1309,1332]" pageId="6" pageNumber="491" type="female">3 ♀♀</specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFECFFC8FDBCFAC791A3FA94" box="[555,678,1311,1332]" pageId="6" pageNumber="491" value="1996-06-15">15­VI­1996</date>
(18 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FC9DFAC79051FA93" box="[778,852,1309,1332]" pageId="6" pageNumber="491" type="female">10 ♀♀</specimenCount>
).
<materialsCitation id="8A9F3C86FFECFFC8FCFCFAC7924FFAC9" collectingDate="1998-07-23" collectorName="El Terrero &amp; J. Garcia" country="Mexico" elevation="2200" location="Alt." municipality="Minatitlan" pageId="6" pageNumber="491" specimenCount="2" specimenCount-female="2" stateProvince="Colima" typeStatus="paratype">
<emphasis id="0883EAC9FFECFFC8FCFCFAC790BBFA94" box="[875,958,1311,1332]" italics="true" pageId="6" pageNumber="491">
<collectingRegion id="F833F839FFECFFC8FCFCFAC790BFFA94" box="[875,954,1311,1332]" country="Mexico" name="Colima" pageId="6" pageNumber="491">Colima</collectingRegion>
,
</emphasis>
<collectingMunicipality id="DA2CACA1FFECFFC8FC51FAC79733FA94" box="[966,1078,1311,1332]" pageId="6" pageNumber="491">Minatitlán</collectingMunicipality>
,
<collectorName id="9702530DFFECFFC8FF6EFAE1926DFAEE" box="[249,360,1337,1358]" pageId="6" pageNumber="491">El Terrero</collectorName>
, bosque mesófilo,
<location id="3F286000FFECFFC8FDA7FAE1915CFAEE" LSID="urn:lsid:plazi:treatment:B25E87CDFFEEFFC9FF6EFAFD91BAFC99:3F286000FFECFFC8FDA7FAE1915CFAEE" box="[560,601,1337,1358]" country="Mexico" municipality="Minatitlan" name="Alt." pageId="6" pageNumber="491" stateProvince="Colima">Alt.</location>
<quantity id="FD0F9B3EFFECFFC8FDF6FAE191BEFAEE" box="[609,699,1337,1358]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="491" unit="m" value="2200.0">
<elevation id="B1DAD1E8FFECFFC8FDF6FAE191BEFAEE" box="[609,699,1337,1358]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="491" unit="m" value="2200.0">2,200 m</elevation>
</quantity>
,
<date id="4E49101BFFECFFC8FD5FFAE1904BFAEE" box="[712,846,1337,1358]" pageId="6" pageNumber="491" value="1998-07-23">
<collectingDate id="5E0DE9F3FFECFFC8FD5FFAE1904BFAEE" box="[712,846,1337,1358]" pageId="6" pageNumber="491" value="1998-07-23">23­VII­1998</collectingDate>
</date>
, trampa luz,
<collectorName id="9702530DFFECFFC8FC4CFAE1973FFAEE" box="[987,1082,1337,1358]" pageId="6" pageNumber="491">J. García</collectorName>
(
<specimenCount id="2CF1FD52FFECFFC8FE96FA8C9245FAC9" box="[257,320,1363,1385]" pageId="6" pageNumber="491" type="female">2 ♀♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="8A9F3C86FFECFFC8FECFFA8C904CFAC9" box="[344,841,1363,1385]" collectingDate="1998-07-24" collectorName="El Terrero &amp; J. Garcia" country="Mexico" elevation="2200" location="Alt." municipality="Minatitlan" pageId="6" pageNumber="491" specimenCount="7" specimenCount-female="7" stateProvince="Colima" typeStatus="paratype">
same data except
<date id="4E49101BFFECFFC8FD8FFA8C9199FAC9" box="[536,668,1364,1385]" pageId="6" pageNumber="491" value="1998-07-24">
<collectingDate id="5E0DE9F3FFECFFC8FD8FFA8C9199FAC9" box="[536,668,1364,1385]" pageId="6" pageNumber="491" value="1998-07-24">24­VII­1998</collectingDate>
</date>
(12 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FC96FA8C903AFAC9" box="[769,831,1363,1385]" pageId="6" pageNumber="491" type="female">7 ♀♀</specimenCount>
)
</materialsCitation>
; M. López, trampa uv (♂);
<date id="4E49101BFFECFFC8FEB9FAB792B7FA24" box="[302,434,1391,1412]" pageId="6" pageNumber="491" value="1998-07-25">25­VII­1998</date>
(♂); J. García (3 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FD31FAB791E1FA23" box="[678,740,1389,1412]" pageId="6" pageNumber="491" type="female">6 ♀♀</specimenCount>
);
<date id="4E49101BFFECFFC8FD6BFAB7908CFA24" box="[764,905,1391,1412]" pageId="6" pageNumber="491" value="1998-08-20">20­VIII­1998</date>
(11 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FC7BFAB7972CFA23" box="[1004,1065,1389,1412]" pageId="6" pageNumber="491" type="female">9 ♀♀</specimenCount>
);
<materialsCitation id="8A9F3C86FFECFFC8FF6EFA519135FA3E" box="[249,560,1416,1438]" country="Mexico" location="Lopez" pageId="6" pageNumber="491" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
M.
<location id="3F286000FFECFFC8FEB5FA519263FA3E" LSID="urn:lsid:plazi:treatment:B25E87CDFFEEFFC9FF6EFAFD91BAFC99:3F286000FFECFFC8FEB5FA519263FA3E" box="[290,358,1417,1438]" country="Mexico" name="Lopez" pageId="6" pageNumber="491">López</location>
, trampa uv (♂
<emphasis id="0883EAC9FFECFFC8FD9DFA519114FA3E" bold="true" box="[522,529,1417,1438]" pageId="6" pageNumber="491">/</emphasis>
<specimenCount id="2CF1FD52FFECFFC8FD85FA509123FA3E" box="[530,550,1416,1438]" pageId="6" pageNumber="491" type="female"></specimenCount>
)
</materialsCitation>
;
<date id="4E49101BFFECFFC8FDD6FA5191CBFA3E" box="[577,718,1417,1438]" pageId="6" pageNumber="491" value="1998-08-21">21­VIII­1998</date>
(
<specimenCount id="2CF1FD52FFECFFC8FD76FA519024FA3E" box="[737,801,1416,1438]" pageId="6" pageNumber="491" type="female">3 ♀♀</specimenCount>
); J. García (
<specimenCount id="2CF1FD52FFECFFC8FC26FA5190FAFA3E" box="[945,1023,1416,1438]" pageId="6" pageNumber="491" type="female">15 ♀♀</specimenCount>
); luz fluor. (37 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FE01FA7C92E4FA19" box="[406,481,1443,1465]" pageId="6" pageNumber="491" type="female">28 ♀♀</specimenCount>
);
<date id="4E49101BFFECFFC8FE6CFA7C918FFA19" box="[507,650,1444,1465]" pageId="6" pageNumber="491" value="1998-08-22">22­VIII­1998</date>
, trampa uv (1 ♂
<specimenCount id="2CF1FD52FFECFFC8FCDEFA7C9090FA19" box="[841,917,1443,1465]" pageId="6" pageNumber="491" type="female">36 ♀♀</specimenCount>
); luz fluor. (42 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FEBDFA679273FA73" box="[298,374,1469,1492]" pageId="6" pageNumber="491" type="female">66 ♀♀</specimenCount>
);
<date id="4E49101BFFECFFC8FE18FA679108FA74" box="[399,525,1471,1492]" pageId="6" pageNumber="491" value="1998-09-24">24­IX­1998</date>
, M. López (
<specimenCount id="2CF1FD52FFECFFC8FD02FA6591ACFA73" box="[661,681,1469,1491]" pageId="6" pageNumber="491" type="female"></specimenCount>
); J. García (1 ♂
<specimenCount id="2CF1FD52FFECFFC8FCF0FA6790A0FA73" box="[871,933,1469,1492]" pageId="6" pageNumber="491" type="female">3 ♀♀</specimenCount>
);
<date id="4E49101BFFECFFC8FC29FA67973FFA74" box="[958,1082,1471,1492]" pageId="6" pageNumber="491" value="1998-09-25">25­IX­1998</date>
(
<specimenCount id="2CF1FD52FFECFFC8FE96FA019246FA4E" box="[257,323,1496,1518]" pageId="6" pageNumber="491" type="female">6 ♀♀</specimenCount>
); M. López (
<specimenCount id="2CF1FD52FFECFFC8FE49FA0092F7FA4E" box="[478,498,1496,1518]" pageId="6" pageNumber="491" type="female"></specimenCount>
); trampa uv (
<specimenCount id="2CF1FD52FFECFFC8FD06FA0191D7FA4E" box="[657,722,1496,1518]" pageId="6" pageNumber="491" type="female">3 ♀♀</specimenCount>
);
<date id="4E49101BFFECFFC8FD79FA019065FA4E" box="[750,864,1497,1518]" pageId="6" pageNumber="491" value="1998-10-22">22­X­1998</date>
(♂);
<date id="4E49101BFFECFFC8FC32FA01971DFA4E" box="[933,1048,1497,1518]" pageId="6" pageNumber="491" value="1998-10-23">23­X­1998</date>
(
<specimenCount id="2CF1FD52FFECFFC8FBBCFA019224F9A9" pageId="6" pageNumber="491" type="female">6 ♀♀</specimenCount>
);
<date id="4E49101BFFECFFC8FEAFFA2C92AEF9A9" box="[312,427,1524,1545]" pageId="6" pageNumber="491" value="1998-10-24">24­X­1998</date>
(
<specimenCount id="2CF1FD52FFECFFC8FE2CFA2B92CAF9A9" box="[443,463,1523,1545]" pageId="6" pageNumber="491" type="female"></specimenCount>
); luz fluor. (2 ♂♂
<specimenCount id="2CF1FD52FFECFFC8FD0FFA2C91D1F9A9" box="[664,724,1523,1545]" pageId="6" pageNumber="491" type="female">6 ♀♀</specimenCount>
);
<date id="4E49101BFFECFFC8FD7CFA2C906CF9A9" box="[747,873,1524,1545]" pageId="6" pageNumber="491" value="1998-11-24">24­XI­1998</date>
, trampa uv (
<specimenCount id="2CF1FD52FFECFFC8FC7AFA2C972FF9A9" box="[1005,1066,1523,1545]" pageId="6" pageNumber="491" type="female">2 ♀♀</specimenCount>
).
<typeStatus id="E54C8879FFECFFC8FF6EF9D7925AF984" box="[249,351,1551,1572]" pageId="6" pageNumber="491" type="paratype">Paratypes</typeStatus>
are deposited at AAHC,
<collectionCode id="5CE6AE1EFFECFFC8FDE2F9D791CDF984" box="[629,712,1551,1572]" pageId="6" pageNumber="491">AMNH</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FD42F9D79019F984" box="[725,796,1551,1572]" pageId="6" pageNumber="491">CASC</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FCBEF9D79064F984" box="[809,865,1551,1572]" pageId="6" pageNumber="491">CNC</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FCF9F9D790B9F984" box="[878,956,1551,1572]" pageId="6" pageNumber="491">FMNH</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FC5EF9D79733F984" box="[969,1078,1551,1572]" pageId="6" pageNumber="491">IBUNAM</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FF6EF9F1923FF99E" box="[249,314,1577,1598]" pageId="6" pageNumber="491">IEXA</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FED0F9F19281F99E" box="[327,388,1577,1598]" pageId="6" pageNumber="491">IMEJ</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FE07F9F192DDF99E" box="[400,472,1577,1598]" pageId="6" pageNumber="491">GNPC</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FE72F9F1913DF99E" box="[485,568,1577,1598]" pageId="6" pageNumber="491">MNHN</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FDD3F9F19191F99E" box="[580,660,1577,1598]" pageId="6" pageNumber="491">MXAL</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FD36F9F191F7F99E" box="[673,754,1577,1598]" pageId="6" pageNumber="491">NHMC</collectionCode>
,
<collectionCode id="5CE6AE1EFFECFFC8FD69F9F1904CF99E" box="[766,841,1577,1598]" pageId="6" pageNumber="491">UNSM</collectionCode>
and
<collectionCode id="5CE6AE1EFFECFFC8FC15F9F190D7F99E" box="[898,978,1577,1598]" pageId="6" pageNumber="491">ZMHU</collectionCode>
.
</paragraph>
<paragraph id="3A4836DBFFECFFC8FE84F99C9724F9D4" blockId="6.[249,1082,484,1652]" pageId="6" pageNumber="491">
<materialsCitation id="8A9F3C86FFECFFC8FE84F99C9718F9D4" country="Mexico" county="Autlan" location="Las Joyas" municipality="Biosphere Reserve Sierra de Manantlan" pageId="6" pageNumber="491" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<emphasis id="0883EAC9FFECFFC8FE84F99C92B2F9F9" bold="true" box="[275,439,1604,1625]" pageId="6" pageNumber="491">
<typeStatus id="E54C8879FFECFFC8FE84F99C9249F9F9" box="[275,332,1604,1625]" pageId="6" pageNumber="491">Type</typeStatus>
Locality.
</emphasis>
<location id="3F286000FFECFFC8FE57F99C9129F9F9" LSID="urn:lsid:plazi:treatment:B25E87CDFFEEFFC9FF6EFAFD91BAFC99:3F286000FFECFFC8FE57F99C9129F9F9" box="[448,556,1604,1625]" country="Mexico" county="Autlan" municipality="Biosphere Reserve Sierra de Manantlan" name="Las Joyas" pageId="6" pageNumber="491" stateProvince="Jalisco">Las Joyas</location>
,
<collectingMunicipality id="DA2CACA1FFECFFC8FDAFF99C90E0F9F9" box="[568,997,1604,1625]" pageId="6" pageNumber="491">Biosphere Reserve Sierra de Manantlán</collectingMunicipality>
,
<collectingCounty id="D3294E57FFECFFC8FC65F99C973CF9F9" box="[1010,1081,1604,1625]" pageId="6" pageNumber="491">Autlán</collectingCounty>
municipality, state of
<collectingRegion id="F833F839FFECFFC8FE74F9879128F9D4" box="[483,557,1631,1652]" country="Mexico" name="Jalisco" pageId="6" pageNumber="491">Jalisco</collectingRegion>
,
<collectingCountry id="42E0764BFFECFFC8FDADF987918FF9D4" box="[570,650,1631,1652]" name="Mexico" pageId="6" pageNumber="491">México</collectingCountry>
(approx. 19
<emphasis id="0883EAC9FFECFFC8FC99F9869013F9D3" bold="true" box="[782,790,1630,1651]" pageId="6" pageNumber="491">°</emphasis>
35
<emphasis id="0883EAC9FFECFFC8FCA7F985903DF9D3" bold="true" box="[816,824,1629,1651]" pageId="6" pageNumber="491">'</emphasis>
13
<emphasis id="0883EAC9FFECFFC8FCC5F985905FF9D3" bold="true" box="[850,858,1629,1651]" pageId="6" pageNumber="491">&quot;</emphasis>
N; 104
<emphasis id="0883EAC9FFECFFC8FC33F98690A9F9D3" bold="true" box="[932,940,1630,1651]" pageId="6" pageNumber="491">°</emphasis>
16
<emphasis id="0883EAC9FFECFFC8FC51F98590CBF9D3" bold="true" box="[966,974,1629,1651]" pageId="6" pageNumber="491">'</emphasis>
48
<emphasis id="0883EAC9FFECFFC8FC66F98590FCF9D3" bold="true" box="[1009,1017,1629,1651]" pageId="6" pageNumber="491">&quot;</emphasis>
W)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFEDFFC9FE87FED6917DFE59" pageId="7" pageNumber="492" type="biology_ecology">
<paragraph id="3A4836DBFFEDFFC9FE87FED6917DFE59" blockId="7.[246,1079,270,825]" pageId="7" pageNumber="492">
<emphasis id="0883EAC9FFEDFFC9FE87FED692C1FE83" bold="true" box="[272,452,270,291]" pageId="7" pageNumber="492">Biological Data.</emphasis>
Specimens of
<taxonomicName id="FDF74D58FFEDFFC9FDF3FED791F8FE84" box="[612,765,271,292]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="manantleca" status="sp. nov.">
<emphasis id="0883EAC9FFEDFFC9FDF3FED791F8FE84" box="[612,765,271,292]" italics="true" pageId="7" pageNumber="492">P. manantleca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEDFFC9FC92FED79032FE84" box="[773,823,271,292]" pageId="7" pageNumber="492" rank="species">n.sp.</taxonomicNameLabel>
were collected at white fluorescent and UV lights in montane cloud forest and pine forest located at
<quantity id="FD0F9B3EFFEDFFC9FF61FE9C929DFEF9" box="[246,408,324,345]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="7" pageNumber="492" unit="m" value="2050.0" valueMax="2200.0" valueMin="1900.0">
<geoCoordinate id="5FC3501CFFEDFFC9FF61FE9C9235FEF9" box="[246,304,324,345]" degrees="1,900" direction="north" orientation="latitude" pageId="7" pageNumber="492" precision="55" value="1.9">1,900</geoCoordinate>
<geoCoordinate id="5FC3501CFFEDFFC9FEA7FE9C927FFEF9" box="[304,378,324,345]" degrees="2,200" direction="west" orientation="longitude" pageId="7" pageNumber="492" precision="55" value="-2.2">2,200</geoCoordinate>
m
</quantity>
altitude. They were also collected at Hg vapor lamps in oak forest located at
<quantity id="FD0F9B3EFFEDFFC9FE38FE879154FED4" box="[431,593,351,372]" metricMagnitude="3" metricUnit="m" metricValue="1.56" metricValueMax="1.65" metricValueMin="1.47" pageId="7" pageNumber="492" unit="m" value="1560.0" valueMax="1650.0" valueMin="1470.0">
<geoCoordinate id="5FC3501CFFEDFFC9FE38FE8792EFFED4" box="[431,490,351,372]" degrees="1,470" direction="north" orientation="latitude" pageId="7" pageNumber="492" precision="55" value="1.47">1,470</geoCoordinate>
<geoCoordinate id="5FC3501CFFEDFFC9FE7DFE879136FED4" box="[490,563,351,372]" degrees="1,650" direction="west" orientation="longitude" pageId="7" pageNumber="492" precision="55" value="-1.65">1,650</geoCoordinate>
m
</quantity>
altitude, where the dominant tree species is
<taxonomicName id="FDF74D58FFEDFFC9FF61FEA29153FE2E" authority="Nee. Phenology" authorityName="Nee. Phenology" box="[246,598,377,399]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="492" phylum="Tracheophyta" rank="species" species="rugosa">
<emphasis id="0883EAC9FFEDFFC9FF61FEA292A5FE2F" box="[246,416,378,399]" italics="true" pageId="7" pageNumber="492">Quercus rugosa</emphasis>
Née. Phenology
</taxonomicName>
: June (33), July (43), August (276), September (34), October (35), November (9), December (2). Other species of
<taxonomicName id="FDF74D58FFEDFFC9FC4BFE4C922FFE64" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="genus">
<emphasis id="0883EAC9FFEDFFC9FC4BFE4C922FFE64" italics="true" pageId="7" pageNumber="492">Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and place were
<taxonomicName id="FDF74D58FFEDFFC9FD4EFE7790D2FE64" box="[729,983,431,452]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="dugesiana" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFEDFFC9FD4EFE7790D2FE64" box="[729,983,431,452]" italics="true" pageId="7" pageNumber="492">P. (Phytalus) dugesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEDFFC9FC77FE779716FE64" box="[992,1043,431,452]" pageId="7" pageNumber="492" rank="species">n.sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFEDFFC9FB88FE7792DBFE7F" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="jalisciensis" status="sp. nov." subGenus="Phytalus">
<emphasis id="0883EAC9FFEDFFC9FB88FE7792DBFE7F" italics="true" pageId="7" pageNumber="492">P. (Phytalus) jalisciensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEDFFC9FE71FE11911CFE7E" box="[486,537,457,478]" pageId="7" pageNumber="492" rank="species">n.sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFEDFFC9FDB2FE1291FAFE7F" box="[549,767,457,479]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="sayloriana" status="sp. nov.">
<emphasis id="0883EAC9FFEDFFC9FDB2FE129138FE7F" box="[549,573,458,479]" italics="true" pageId="7" pageNumber="492">P.</emphasis>
(
<emphasis id="0883EAC9FFEDFFC9FDDBFE12917AFE7F" box="[588,639,458,479]" italics="true" pageId="7" pageNumber="492">s.str.</emphasis>
)
<emphasis id="0883EAC9FFEDFFC9FD18FE1291FAFE7F" box="[655,767,458,479]" italics="true" pageId="7" pageNumber="492">sayloriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEDFFC9FC91FE119044FE7E" box="[774,833,457,478]" pageId="7" pageNumber="492" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="FDF74D58FFEDFFC9FCDAFE1290F8FE7F" box="[845,1021,457,479]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="joyana" status="sp. nov.">
<emphasis id="0883EAC9FFEDFFC9FCDAFE129061FE7F" box="[845,868,458,479]" italics="true" pageId="7" pageNumber="492">P.</emphasis>
(
<emphasis id="0883EAC9FFEDFFC9FCE3FE1290A2FE7F" box="[884,935,458,479]" italics="true" pageId="7" pageNumber="492">s.str.</emphasis>
)
<emphasis id="0883EAC9FFEDFFC9FC21FE1290F8FE7F" box="[950,1021,458,479]" italics="true" pageId="7" pageNumber="492">joyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEDFFC9FB92FE119733FE7E" box="[1029,1078,457,478]" pageId="7" pageNumber="492" rank="species">n.sp.</taxonomicNameLabel>
and
<taxonomicName id="FDF74D58FFEDFFC9FEB2FE3C917DFE59" authority="Moron." authorityName="Moron." box="[293,632,484,505]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="chamacayoca">
<emphasis id="0883EAC9FFEDFFC9FEB2FE3C9239FE59" box="[293,316,484,505]" italics="true" pageId="7" pageNumber="492">P.</emphasis>
(
<emphasis id="0883EAC9FFEDFFC9FEDAFE3C9285FE59" box="[333,384,484,505]" italics="true" pageId="7" pageNumber="492">s.str.</emphasis>
)
<emphasis id="0883EAC9FFEDFFC9FE06FE3C9124FE59" box="[401,545,484,505]" italics="true" pageId="7" pageNumber="492">chamacayoca</emphasis>
Morón.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFEDFFC9FE87FE269084FD6E" pageId="7" pageNumber="492" type="discussion">
<paragraph id="3A4836DBFFEDFFC9FE87FE269084FD6E" blockId="7.[246,1079,270,825]" pageId="7" pageNumber="492">
<emphasis id="0883EAC9FFEDFFC9FE87FE269278FDB3" bold="true" box="[272,381,510,531]" pageId="7" pageNumber="492">Remarks.</emphasis>
<taxonomicName id="FDF74D58FFEDFFC9FE1CFE27919CFDB4" box="[395,665,511,532]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="manantleca" status="sp. nov.">
<emphasis id="0883EAC9FFEDFFC9FE1CFE27919CFDB4" box="[395,665,511,532]" italics="true" pageId="7" pageNumber="492">Phyllophaga manantleca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13B057B2FFEDFFC9FD3FFE2791ECFDB4" box="[680,745,511,532]" pageId="7" pageNumber="492" rank="species">n. sp.</taxonomicNameLabel>
belong to the species group
<taxonomicName id="FDF74D58FFEDFFC9FE92FDC2927DFD8F" box="[261,376,538,559]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="0883EAC9FFEDFFC9FE92FDC2927DFD8F" box="[261,376,538,559]" italics="true" pageId="7" pageNumber="492">blanchardi</emphasis>
</taxonomicName>
, complex
<taxonomicName id="FDF74D58FFEDFFC9FD84FDC29184FD8F" box="[531,641,538,559]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="pubicauda">
<emphasis id="0883EAC9FFEDFFC9FD84FDC29184FD8F" box="[531,641,538,559]" italics="true" pageId="7" pageNumber="492">pubicauda</emphasis>
</taxonomicName>
(
<emphasis id="0883EAC9FFEDFFC9FD30FDC291E4FD8F" box="[679,737,538,559]" italics="true" pageId="7" pageNumber="492">sensu</emphasis>
<bibRefCitation id="5E664B2AFFEDFFC9FD78FDC1907EFD8E" author="Moron, M. A." bookContentInfo="Publ. 20, Instituto de Ecologia, Mexico. 341 pp." box="[751,891,537,558]" pageId="7" pageNumber="492" refId="ref10446" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica. (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico. 341 pp." title="El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica. (Insecta: Coleoptera)" type="book" year="1986">Morón 1986</bibRefCitation>
). The diagnostic combination of characters for both sexes include: anterior border of clypeus widely sinuated; frons and pronotum with dense vestiture of erect, long setae; abdominal sternites 2
<emphasis id="0883EAC9FFEDFFC9FE77FDB192EDFDDE" bold="true" box="[480,488,617,638]" pageId="7" pageNumber="492">°</emphasis>
to 4
<emphasis id="0883EAC9FFEDFFC9FDB7FDB1912DFDDE" bold="true" box="[544,552,617,638]" pageId="7" pageNumber="492">°</emphasis>
with deep mesial sulcus; abdominal sternite 5
<emphasis id="0883EAC9FFEDFFC9FBB9FDB19733FDDE" bold="true" box="[1070,1078,617,638]" pageId="7" pageNumber="492">°</emphasis>
with granulose­setiferous patch; and pygidium finely punctate and moderately setiferous. Shape of paramera is similar to
<taxonomicName id="FDF74D58FFEDFFC9FD51FD4790B3FD14" authority="Moser" authorityName="Moser" box="[710,950,671,692]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="inflativentris">
<emphasis id="0883EAC9FFEDFFC9FD51FD47906FFD14" box="[710,874,671,692]" italics="true" pageId="7" pageNumber="492">P. inflativentris</emphasis>
Moser
</taxonomicName>
and
<taxonomicName id="FDF74D58FFEDFFC9FC7AFD479289FD6E" authority="Saylor" authorityName="Saylor" authorityYear="1941" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="species" species="hintonella">
<emphasis id="0883EAC9FFEDFFC9FC7AFD47923AFD6F" italics="true" pageId="7" pageNumber="492">P. hintonella</emphasis>
Saylor
</taxonomicName>
, but the proportions of each part are different.
</paragraph>
</subSubSection>
<subSubSection id="72ED6550FFEDFFC9FE87FD0C91BAFC99" pageId="7" pageNumber="492" type="etymology">
<paragraph id="3A4836DBFFEDFFC9FE87FD0C91BAFC99" blockId="7.[246,1079,270,825]" pageId="7" pageNumber="492">
<emphasis id="0883EAC9FFEDFFC9FE87FD0C928AFD49" bold="true" box="[272,399,724,745]" pageId="7" pageNumber="492">Etymology.</emphasis>
Derived from Sierra de Manantlán, old Aztec name
<emphasis id="0883EAC9FFEDFFC9FB9CFD0C9265FCA4" italics="true" pageId="7" pageNumber="492">Manauantlan</emphasis>
meaning, near the site where defence is developed or near defended site, near fortification (
<emphasis id="0883EAC9FFEDFFC9FD04FCD291F4FCBF" box="[659,753,778,799]" italics="true" pageId="7" pageNumber="492">Manauia</emphasis>
<emphasis id="0883EAC9FFEDFFC9FD69FCD19017FCBE" bold="true" box="[766,786,777,798]" pageId="7" pageNumber="492">=</emphasis>
defended,
<emphasis id="0883EAC9FFEDFFC9FC06FCD290BDFCBF" box="[913,952,778,799]" italics="true" pageId="7" pageNumber="492">ana</emphasis>
<emphasis id="0883EAC9FFEDFFC9FC52FCD190DCFCBE" bold="true" box="[965,985,777,798]" pageId="7" pageNumber="492">=</emphasis>
growth, development,
<emphasis id="0883EAC9FFEDFFC9FE1BFCFC92B0FC99" box="[396,437,804,825]" italics="true" pageId="7" pageNumber="492">tlan</emphasis>
<emphasis id="0883EAC9FFEDFFC9FE29FCFC92D7FC99" bold="true" box="[446,466,804,825]" pageId="7" pageNumber="492">=</emphasis>
near)
<bibRefCitation id="5E664B2AFFEDFFC9FD8EFCFC91BEFC99" author="Simeon, R." bookContentInfo="Siglo XXI Editores, Mexico. 782 pp." box="[537,699,804,825]" pageId="7" pageNumber="492" refId="ref10587" refString="Simeon, R. 1988. Diccionario de la lengua nahuatl o mexicana. Siglo XXI Editores, Mexico. 782 pp." title="Diccionario de la lengua nahuatl o mexicana" type="book" year="1988">Simeón (1988)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>