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data/03/FF/DB/03FFDB7FE2311A35FF4B189DD085FCEB.xml
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<document id="99256D82EE76748CAB6C81F4FF3ECF52" ID-DOI="10.5281/zenodo.5398335" ID-ISSN="1638-9387" ID-Zenodo-Dep="5398335" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1630327988213" checkinUser="marcus" docAuthor="Mauriès, Jean-Paul" docDate="2001" docId="03FFDB7FE2311A35FF4B189DD085FCEB" docLanguage="fr" docName="Zoosystema.2001.23.1.131-142.pdf" docOrigin="Zoosystema 23 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Chinosphaera maculosa Attems 1935" docType="treatment" docVersion="1" lastPageNumber="134" masterDocId="FFC6A307E2321A36FFE41B79D406FFA1" masterDocTitle="Sur l’identité de Zephronia hainani Gressitt, 1941, à propos de la description d’un nouveau Prionobelum (Diplopoda, Sphaerotheriida, Sphaeropoeidae) de Haïnan, Chine" masterLastPageNumber="142" masterPageNumber="131" pageNumber="134" updateTime="1741721054631" updateUser="juliana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="74848FD90CB45051481279EA5CC61159">Sur l’identité de Zephronia hainani Gressitt, 1941, à propos de la description d’un nouveau Prionobelum (Diplopoda, Sphaerotheriida, Sphaeropoeidae) de Haïnan, Chine</mods:title>
<document id="3180F45205FF6CBA8158DCBFAAC4AB56" ID-CLB-Dataset="308554" ID-DOI="10.5281/zenodo.5398335" ID-GBIF-Dataset="433e218c-b197-4a27-b6d9-05174461c35c" ID-ISSN="1638-9387" ID-Zenodo-Dep="5398335" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1630327988213" checkinUser="marcus" docAuthor="Mauriès, Jean-Paul" docDate="2001" docId="03FFDB7FE2311A35FF4B189DD085FCEB" docLanguage="fr" docName="Zoosystema.2001.23.1.131-142.pdf" docOrigin="Zoosystema 23 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Chinosphaera maculosa Attems 1935" docType="treatment" docVersion="3" lastPageNumber="134" masterDocId="FFC6A307E2321A36FFE41B79D406FFA1" masterDocTitle="Sur l’identité de Zephronia hainani Gressitt, 1941, à propos de la description d’un nouveau Prionobelum (Diplopoda, Sphaerotheriida, Sphaeropoeidae) de Haïnan, Chine" masterLastPageNumber="142" masterPageNumber="131" pageNumber="134" updateTime="1741722043715" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5D150C1999F54FE1CEA77E6EF128E425">Sur l’identité de Zephronia hainani Gressitt, 1941, à propos de la description d’un nouveau Prionobelum (Diplopoda, Sphaerotheriida, Sphaeropoeidae) de Haïnan, Chine</mods:title>
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<mods:namePart id="AEFFF408CD940A250ACE7D6D600BB351">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="FA54BB547FD63EB87ECC7D4F4122EFD1">Laboratoire de Zoologie (Arthropodes), Muséum national d’Histoire naturelle, 61 rue de Buffon, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:namePart id="53260BBDC41E653992ECE4AB36BA2DEA">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="51669223E0073C877E7ED8D971AF77ED">Laboratoire de Zoologie (Arthropodes), Muséum national d’Histoire naturelle, 61 rue de Buffon, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:date id="44CCF54E8EFC5AC49F375E77DF1837B9">2001</mods:date>
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<treatment id="03FFDB7FE2311A35FF4B189DD085FCEB" LSID="urn:lsid:plazi:treatment:03FFDB7FE2311A35FF4B189DD085FCEB" httpUri="http://treatment.plazi.org/id/03FFDB7FE2311A35FF4B189DD085FCEB" lastPageNumber="134" pageId="3" pageNumber="134">
<treatment id="03FFDB7FE2311A35FF4B189DD085FCEB" ID-DOI="http://doi.org/10.5281/zenodo.15008580" ID-Zenodo-Dep="15008580" LSID="urn:lsid:plazi:treatment:03FFDB7FE2311A35FF4B189DD085FCEB" httpUri="http://treatment.plazi.org/id/03FFDB7FE2311A35FF4B189DD085FCEB" lastPageNumber="134" pageId="3" pageNumber="134" scope_class="Diplopoda" scope_family="Sphaeropoeidae" scope_order="Sphaerotheriida">
<subSubSection id="C34C39E2E2311A35FF4B189DD64FFC5F" box="[175,585,996,1022]" pageId="3" pageNumber="134" type="nomenclature">
<paragraph id="8BE96A69E2311A35FF4B189DD64FFC5F" blockId="3.[175,585,996,1052]" box="[175,585,996,1022]" pageId="3" pageNumber="134">
<taxonomicName id="4C5611EAE2311A35FF4B189DD64FFC5F" ID-CoL="7TVCL" authority="Attems, 1935" authorityName="Attems" authorityYear="1935" box="[175,585,996,1022]" class="Diplopoda" family="Zephroniidae" genus="Chinosphaera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="maculosa">
@ -50,7 +52,7 @@
<subSubSection id="C34C39E2E2311A35FEB01F7BD5A2FBBD" box="[340,420,1026,1052]" pageId="3" pageNumber="134" type="description">
<paragraph id="8BE96A69E2311A35FEB01F7BD5A2FBBD" blockId="3.[175,585,996,1052]" box="[340,420,1026,1052]" pageId="3" pageNumber="134">
(
<figureCitation id="136D76ECE2311A35FEBF1F7BD59AFBBD" box="[347,412,1026,1052]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." pageId="3" pageNumber="134">Fig. 2</figureCitation>
<figureCitation id="136D76ECE2311A35FEBF1F7BD59AFBBD" box="[347,412,1026,1052]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.15008574" httpUri="https://zenodo.org/record/15008574/files/figure.png" pageId="3" pageNumber="134">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
@ -78,7 +80,7 @@ déposé au
En complément et émendation de la description originale qui est, dans son ensemble, satisfaisante, il y a lieu d’attirer l’attention sur les points ci-après:
<emphasis id="B922B67BE2311A35FF951E49D481FAEB" bold="true" box="[113,135,1328,1354]" pageId="3" pageNumber="134">1.</emphasis>
une figure de la tête vue de face (
<figureCitation id="136D76ECE2311A35FE121E49D64FFAEB" box="[502,585,1328,1354]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." pageId="3" pageNumber="134">Fig. 2A</figureCitation>
<figureCitation id="136D76ECE2311A35FE121E49D64FFAEB" box="[502,585,1328,1354]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.15008574" httpUri="https://zenodo.org/record/15008574/files/figure.png" pageId="3" pageNumber="134">Fig. 2A</figureCitation>
) avec l’antenne droite montre la dense ponctuation de la face et le 7
<superScript id="7C23C721E2311A35FEEA1E12D510FADF" attach="left" box="[270,278,1387,1406]" fontSize="8" pageId="3" pageNumber="134">e</superScript>
antennomère avec ses 36 cônes sensitifs;
@ -94,16 +96,16 @@ le corselet, en vue latérale, se montre très semblable à celui de
<paragraph id="8BE96A69E2311A35FF951D29D069FEA8" blockId="3.[113,647,1200,1706]" lastBlockId="3.[677,1210,207,842]" pageId="3" pageNumber="134">
<emphasis id="B922B67BE2311A35FF951D29D481F9CB" bold="true" box="[113,135,1616,1642]" pageId="3" pageNumber="134">3.</emphasis>
il était utile de donner de nouvelles figures du paratélopode (P.22) vu sous plusieurs angles (
<figureCitation id="136D76ECE2311A35FF9D1DE9D4BBF90B" box="[121,189,1680,1706]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." pageId="3" pageNumber="134">Fig. 2</figureCitation>
<figureCitation id="136D76ECE2311A35FF9D1DE9D4BBF90B" box="[121,189,1680,1706]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.15008574" httpUri="https://zenodo.org/record/15008574/files/figure.png" pageId="3" pageNumber="134">Fig. 2</figureCitation>
C-E); la
<figureCitation id="136D76ECE2311A35FECE1DE9D59AF90B" box="[298,412,1680,1706]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." pageId="3" pageNumber="134">Figure 2E</figureCitation>
<figureCitation id="136D76ECE2311A35FECE1DE9D59AF90B" box="[298,412,1680,1706]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.15008574" httpUri="https://zenodo.org/record/15008574/files/figure.png" pageId="3" pageNumber="134">Figure 2E</figureCitation>
, comme la figure 25 d’Attems, est effectuée sous l’angle le moins démonstratif; les autres figures montrent:
</paragraph>
<paragraph id="8BE96A69E2311A35FD411A76D796FD88" blockId="3.[677,1210,207,842]" pageId="3" pageNumber="134">
– qu’il existe bien un processus préfémoral, qui est ici seulement en angle droit (
<figureCitation id="136D76ECE2311A35FBFF1A49D095FEEB" box="[1051,1171,304,330]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." pageId="3" pageNumber="134">Fig. 2C, d</figureCitation>
<figureCitation id="136D76ECE2311A35FBFF1A49D095FEEB" box="[1051,1171,304,330]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.15008574" httpUri="https://zenodo.org/record/15008574/files/figure.png" pageId="3" pageNumber="134">Fig. 2C, d</figureCitation>
) et non en dent aiguë comme chez les autres espèces; – la forme en crochet du tibia; ce dernier (
<figureCitation id="136D76ECE2311A35FB861A09D0B4FE2B" box="[1122,1202,368,394]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." pageId="3" pageNumber="134">Fig. 2B</figureCitation>
<figureCitation id="136D76ECE2311A35FB861A09D0B4FE2B" box="[1122,1202,368,394]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.15008574" httpUri="https://zenodo.org/record/15008574/files/figure.png" pageId="3" pageNumber="134">Fig. 2B</figureCitation>
) abrite, dans sa concavité, face orale, un tarse en moignon moins visible que ne l’indique le dessin d’Attems. Il est à noter la ressemblance frappante de ce tibiotarse avec celui figuré par
<bibRefCitation id="EFC71798E2311A35FBB11A89D6DBFD8B" author="VERHOEFF K. W." pageId="3" pageNumber="134" pagination="1 - 142" refId="ref5736" refString="VERHOEFF K. W. 1924. - Myriapoda: Diplopoda. Results of Dr E. Mjoberg's Swedish Scientific Expeditions to Australia 1910 - 1913. Arkiv fur Zoologi, Stockholm 16 (5): 1 - 142." type="journal article" year="1924">Verhoeff 1924</bibRefCitation>
pour
@ -115,7 +117,7 @@ pour
<paragraph id="8BE96A69E2311A35FD411949D7E7FD4B" blockId="3.[677,1210,207,842]" pageId="3" pageNumber="134">
<emphasis id="B922B67BE2311A35FD411949D6BCFDEB" bold="true" box="[677,698,560,586]" pageId="3" pageNumber="134">4.</emphasis>
les télopodes (P.23) sont figurés à nouveau ici (
<figureCitation id="136D76ECE2311A35FD481929D6FAFDCB" box="[684,764,592,618]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." pageId="3" pageNumber="134">Fig. 2F</figureCitation>
<figureCitation id="136D76ECE2311A35FD481929D6FAFDCB" box="[684,764,592,618]" captionStart="FIG" captionStartId="4.[114,125,1392,1409]" captionTargetBox="[98,1224,184,1367]" captionTargetId="figure-100@4.[66,1247,192,1407]" captionTargetPageId="4" captionText="FIG. 2. — Chinosphaera maculosa Attems, 1935, holotype; A, moitié droite de la tête et du collum, vue antéro-ventrale; B, tibiotarse du télopode (P.23) gauche, en vue orale; C, vue antéro-latérale du paratélopode (P.22) droit; D, vue orale du paratélopode (P.22) gauche; E, vue distale-caudale du paratélopode (P.22) droit; F, vue caudale du syncoxite et du télopode (P.23) droit. Abréviation: d, dent. Échelles: A, C-F, 1 mm; B, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.15008574" httpUri="https://zenodo.org/record/15008574/files/figure.png" pageId="3" pageNumber="134">Fig. 2F</figureCitation>
) bien qu’ils n’apportent que peu par rapport aux figures d’Attems. À noter toutefois: les dents de la crête du tibia sont plus petites et plus nombreuses (9/10 au lieu de 8) que sur les figures d’Attems, et de teinte foncée.
</paragraph>
<paragraph id="8BE96A69E2311A35FD411989D085FCEB" blockId="3.[677,1210,207,842]" pageId="3" pageNumber="134">Il est manifeste, à l’examen des pattes copulatrices du mâle, que cette espèce est très proche, donc congénérique des deux taxons vus plus haut.</paragraph>

374
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(
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)
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MATÉRIEL- TYPE. — Un
MATÉRIEL- TYPE. —
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Un
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, une
,
</materialsCitation>
<materialsCitation id="3B3E6034E2371A33FBF91EBED6C6FA5B" collectingDate="1995-11-19" collectorName="P. Jolivet" country="China" latitude="19.5" location="Danzhou" longLatPrecision="1271" longitude="109.55" pageId="5" pageNumber="136" specimenCount="1" stateProvince="Hainan" typeStatus="allotype">
une
<typeStatus id="54EDD4CBE2371A33FB851EB1D0B3FA41" box="[1121,1205,1480,1504]" pageId="5" pageNumber="136" type="allotype">allotype</typeStatus>
, 12 et
<specimenCount id="9D50A1E0E2371A33FCC51E9BD7D6FA58" box="[801,976,1505,1530]" pageId="5" pageNumber="136" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
sont déposés dans la collection Myriapodes du Muséum national d’Histoire naturelle de Paris (CB 054). Un et une
, 12
</materialsCitation>
<materialsCitation id="3B3E6034E2371A33FCE01E98D052F98D" collectingDate="1995-11-19" collectionCode="Museum national d'Histoire naturelle de Paris" collectorName="P. Jolivet" country="China" latitude="19.5" location="Danzhou" longLatPrecision="1271" longitude="109.55" pageId="5" pageNumber="136" specimenCount="4" stateProvince="Hainan" typeStatus="paratype">
et
<specimenCount id="9D50A1E0E2371A33FCC51E9BD728FA5B" box="[801,814,1506,1530]" pageId="5" pageNumber="136" type="generic">4</specimenCount>
<typeStatus id="54EDD4CBE2371A33FC8E1E98D7D6FA58" box="[874,976,1505,1529]" pageId="5" pageNumber="136">paratypes</typeStatus>
sont déposés dans la collection Myriapodes du
<collectionCode id="ED47F2ACE2371A33FC031E82D7DDF98D" pageId="5" pageNumber="136">Muséum national d’Histoire naturelle de Paris</collectionCode>
(CB 054).
</materialsCitation>
<materialsCitation id="3B3E6034E2371A33FBB91D6AD073F9FE" collectionCode="California Academy of Science de San Francisco" pageId="5" pageNumber="136" specimenCount="1" typeStatus="paratype">
Un et une
<typeStatus id="54EDD4CBE2371A33FD141D54D753F9E4" box="[752,853,1581,1605]" pageId="5" pageNumber="136" type="paratype">paratypes</typeStatus>
sont légués à la
<collectingRegion id="4992A48BE2371A33FBF61D57D086F9E7" box="[1042,1152,1582,1606]" country="United States of America" name="California" pageId="5" pageNumber="136">California</collectingRegion>
Academy of Science de San Francisco,
<collectingCountry id="F3412AF9E2371A33FBDE1D3ED077F9FE" box="[1082,1137,1607,1631]" name="United States of America" pageId="5" pageNumber="136">USA</collectingCountry>
, et un autre couple de
<collectionCode id="ED47F2ACE2371A33FBF61D57D02AF9FE" pageId="5" pageNumber="136">California Academy of Science de San Francisco</collectionCode>
, USA,
</materialsCitation>
<materialsCitation id="3B3E6034E2371A33FB9B1D3FD701F90A" collectionCode="l'Academie des Sciences de Russie" pageId="5" pageNumber="136" specimenCount="1" typeStatus="paratype">
et un autre couple de
<typeStatus id="54EDD4CBE2371A33FCAE1D26D7ADF9D6" box="[842,939,1631,1655]" pageId="5" pageNumber="136" type="paratype">paratypes</typeStatus>
à l’Académie des Sciences de
<collectingCountry id="F3412AF9E2371A33FD211D03D700F933" box="[709,774,1658,1682]" name="Russia" pageId="5" pageNumber="136">Russie</collectingCountry>
(Inst. Problems of Ecology &amp; Evolution,
<collectingRegion id="4992A48BE2371A33FD411DEAD6FBF90A" box="[677,765,1683,1707]" country="Russia" name="Moscow" pageId="5" pageNumber="136">Moscou</collectingRegion>
).
à
<collectionCode id="ED47F2ACE2371A33FC201D19D700F933" pageId="5" pageNumber="136">l’Académie des Sciences de Russie</collectionCode>
(Inst. Problems of Ecology &amp; Evolution, Moscou).
</materialsCitation>
</paragraph>
<paragraph id="8BE96A69E2341A30FF961BB7D0ACF8A4" pageId="6" pageNumber="137">
<table id="F95698C9E234E5C9FF961BB7D0BDF8A4" box="[114,1211,206,1797]" gridcols="2" gridrows="13" pageId="6" pageNumber="137">
<tr id="3566682BE234E5C9FF961BB7D0BDFF46" box="[114,1211,206,231]" gridrow="0" pageId="6" pageNumber="137">
<th id="76B70157E234E5C9FF961BB7D5BFFF46" box="[114,441,206,231]" gridcol="0" gridrow="0" pageId="6" pageNumber="137">A</th>
<th id="76B70157E234E5C9FE3C1BB7D0BDFF46" box="[472,1211,206,231]" gridcol="1" gridrow="0" pageId="6" pageNumber="137">B C D</th>
</tr>
<tr id="3566682BE234E5C9FF961AD5D0BDFE1C" box="[114,1211,428,445]" gridrow="1" pageId="6" pageNumber="137" rowspan-0="1">
<td id="76B70157E234E5C9FE3C1AD5D0BDFE1C" box="[472,1211,428,445]" gridcol="1" gridrow="1" pageId="6" pageNumber="137">B-D</td>
</tr>
<tr id="3566682BE234E5C9FF961A90D0BDFE5B" box="[114,1211,489,506]" gridrow="2" pageId="6" pageNumber="137" rowspan-0="1">
<td id="76B70157E234E5C9FE3C1A90D0BDFE5B" box="[472,1211,489,506]" gridcol="1" gridrow="2" pageId="6" pageNumber="137">A</td>
</tr>
<tr id="3566682BE234E5C9FF961A84D0BDFDB0" box="[114,1211,509,529]" gridrow="3" pageId="6" pageNumber="137" rowspan-0="1">
<td id="76B70157E234E5C9FE3C1A84D0BDFDB0" box="[472,1211,509,529]" gridcol="1" gridrow="3" pageId="6" pageNumber="137">d</td>
</tr>
<tr id="3566682BE234E5C9FF961961D0BDFD90" box="[114,1211,536,561]" gridrow="4" pageId="6" pageNumber="137">
<td id="76B70157E234E5C9FF961961D5BFFD90" box="[114,441,536,561]" gridcol="0" gridrow="4" pageId="6" pageNumber="137">H</td>
<td id="76B70157E234E5C9FE3C1961D0BDFD90" box="[472,1211,536,561]" gridcol="1" gridrow="4" pageId="6" pageNumber="137">E F</td>
</tr>
<tr id="3566682BE234E5C9FF9618D5D0BDFC5B" box="[114,1211,940,1018]" gridrow="5" pageId="6" pageNumber="137" rowspan-0="1">
<td id="76B70157E234E5C9FE3C18D5D0BDFC5B" box="[472,1211,940,1018]" gridcol="1" gridrow="5" pageId="6" pageNumber="137">G d O E-N</td>
</tr>
<tr id="3566682BE234E5C9FF961F17D0BDFB26" box="[114,1211,1134,1159]" gridrow="6" pageId="6" pageNumber="137" rowspan-1="1">
<td id="76B70157E234E5C9FF961F17D5BFFB26" box="[114,441,1134,1159]" gridcol="0" gridrow="6" pageId="6" pageNumber="137">I</td>
</tr>
<tr id="3566682BE234E5C9FF961E79D0BDFAB0" box="[114,1211,1280,1297]" gridrow="7" pageId="6" pageNumber="137" rowspan-0="1">
<td id="76B70157E234E5C9FE3C1E79D0BDFAB0" box="[472,1211,1280,1297]" gridcol="1" gridrow="7" pageId="6" pageNumber="137">O</td>
</tr>
<tr id="3566682BE234E5C9FF961E15D0BDFA24" box="[114,1211,1388,1413]" gridrow="8" pageId="6" pageNumber="137">
<td id="76B70157E234E5C9FF961E15D5BFFA24" box="[114,441,1388,1413]" gridcol="0" gridrow="8" pageId="6" pageNumber="137">J K</td>
<td id="76B70157E234E5C9FE3C1E15D0BDFA24" box="[472,1211,1388,1413]" gridcol="1" gridrow="8" pageId="6" pageNumber="137">L M N</td>
</tr>
<tr id="3566682BE234E5C9FF961D60D0BDF98B" box="[114,1211,1561,1578]" gridrow="9" pageId="6" pageNumber="137">
<td id="76B70157E234E5C9FF961D60D5BFF98B" box="[114,441,1561,1578]" gridcol="0" gridrow="9" pageId="6" pageNumber="137">
<caption id="DF293AE1E2341A30FF961D60D62BF90B" ID-DOI="http://doi.org/10.5281/zenodo.15008578" ID-Zenodo-Dep="15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="6" pageNumber="137" startId="6.[114,125,1561,1578]" targetBox="[114,1211,177,1549]" targetPageId="6">
<paragraph id="8BE96A69E2341A30FF961D60D62BF90B" blockId="6.[114,1211,1561,1706]" pageId="6" pageNumber="137">
FIG. 4. —
<taxonomicName id="4C5611EAE2341A30FF2E1D60D57EF98B" authority="Mauriès, 2001" authorityName="Mauriès" authorityYear="2001" box="[202,376,1561,1578]" class="Diplopoda" family="Zephroniidae" genus="Prionobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="joliveti" status="sp. nov.">
<emphasis id="B922B67BE2341A30FF2E1D60D57EF98B" box="[202,376,1561,1578]" italics="true" pageId="6" pageNumber="137">Prionobelum joliveti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2110B00E2341A30FE9B1D60D5B3F98B" box="[383,437,1561,1578]" pageId="6" pageNumber="137" rank="species">n. sp.</taxonomicNameLabel>
,
</td>
<td id="76B70157E234E5C9FE3C1D60D0BDF98B" box="[472,1211,1561,1578]" gridcol="1" gridrow="9" pageId="6" pageNumber="137">
holotype, paratypes;
, holotype, paratypes;
<emphasis id="B922B67BE2341A30FD2A1D60D6DDF98B" bold="true" box="[718,731,1561,1578]" pageId="6" pageNumber="137">A</emphasis>
, vue antéro-distale de la moitié droite de la tête d’un
</td>
</tr>
<tr id="3566682BE234E5C9FF961D56D0BDF934" box="[114,1211,1583,1685]" gridrow="10" pageId="6" pageNumber="137">
<td id="76B70157E234E5C9FF961D56D0BDF934" box="[114,1211,1583,1685]" colspan="2" colspanRight="1" gridcol="0" gridrow="10" pageId="6" pageNumber="137">
paratype;
, vue antéro-distale de la moitié droite de la tête d’un paratype;
<emphasis id="B922B67BE2341A30FF011D56D4F4F9E1" bold="true" box="[229,242,1583,1600]" pageId="6" pageNumber="137">B</emphasis>
, vue orale d’une hanche de la P.6 gauche du holotype;
<emphasis id="B922B67BE2341A30FD141D56D6F8F9E1" bold="true" box="[752,766,1583,1600]" pageId="6" pageNumber="137">C</emphasis>
@ -142,27 +127,12 @@ paratype;
<emphasis id="B922B67BE2341A30FEB61DFDD573F934" bold="true" box="[338,373,1668,1685]" pageId="6" pageNumber="137">K -N</emphasis>
, paratypes;
<emphasis id="B922B67BE2341A30FE051DFDD5F6F934" bold="true" box="[481,496,1668,1685]" pageId="6" pageNumber="137">O</emphasis>
, allotype, vue caudale de la hanche gauche avec la vulve correspondante.
<tr id="3566682BE234E5C9FF961DE0D0BDF90B" box="[114,1211,1689,1706]" gridrow="11" pageId="6" pageNumber="137">
<td id="76B70157E234E5C9FF961DE0D0BDF90B" box="[114,1211,1689,1706]" colspan="2" colspanRight="1" gridcol="0" gridrow="11" pageId="6" pageNumber="137">
Abréviation:
, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation:
<emphasis id="B922B67BE2341A30FF441DE0D4ADF90B" bold="true" box="[160,171,1689,1706]" pageId="6" pageNumber="137">d</emphasis>
, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm.
</td>
</tr>
</td>
</tr>
<tr id="3566682BE234E5C9FF961D92D0BDF8A4" box="[114,1211,1771,1797]" gridrow="12" pageId="6" pageNumber="137">
<td id="76B70157E234E5C9FF961D92D5BFF8A4" box="[114,441,1771,1797]" gridcol="0" gridrow="12" pageId="6" pageNumber="137">ZOOSYSTEMA • 2001 • 23 (1)</td>
<td id="76B70157E234E5C9FE3C1D92D0BDF8A4" box="[472,1211,1771,1797]" gridcol="1" gridrow="12" pageId="6" pageNumber="137">
<pageNumber id="BBFD61BFE2341A30FB661D94D0ACF8A4" ambiguity="1" box="[1154,1194,1773,1797]" fuzzyness="0" pageId="6" pageNumber="137" score="36.0" value="137">
<pageTitle id="CBC9B20EE2341A30FB661D94D0ACF8A4" box="[1154,1194,1773,1797]" pageId="6" pageNumber="137">137</pageTitle>
</pageNumber>
</td>
</tr>
</table>
</paragraph>
<caption id="DF293AE1E2351A31FF951BB7D5EEFF55" pageId="7" pageNumber="138" startId="7.[113,124,206,223]">
</caption>
<caption id="DF293AE1E2351A31FF951BB7D5EEFF55" ID-Table-UUID="DF293AE1E2351A31FF951BB7D5EEFF55" httpUri="http://table.plazi.org/id/DF293AE1E2351A31FF951BB7D5EEFF55" pageId="7" pageNumber="138" startId="7.[113,124,206,223]" targetBox="[113,568,291,865]" targetIsTable="true" targetPageId="7">
<paragraph id="8BE96A69E2351A31FF951BB7D5EEFF55" blockId="7.[113,647,206,244]" pageId="7" pageNumber="138">
TABLEAU 1. — Longueur et largeur du corps en mm des 22 exemplaires de
<taxonomicName id="4C5611EAE2351A31FEEB1B9AD5B4FF55" authority="Mauriès, 2001" authorityName="Mauriès" authorityYear="2001" box="[271,434,227,244]" class="Diplopoda" family="Zephroniidae" genus="Prionobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="joliveti" status="sp. nov.">
@ -171,24 +141,158 @@ TABLEAU 1. — Longueur et largeur du corps en mm des 22 exemplaires de
<taxonomicNameLabel id="A2110B00E2351A31FE5C1B9AD5EEFF55" box="[440,488,227,244]" pageId="7" pageNumber="138" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BE96A69E2351A31FEAD1A5AD62EFCC0" pageId="7" pageNumber="138">
<table id="F95698C9E235E5C9FF951A5AD63EFCC0" box="[113,568,291,865]" gridcols="3" gridrows="23" pageId="7" pageNumber="138">
<tr id="3566682BE235E5C9FF951A5AD63EFE96" box="[113,568,291,311]" gridrow="0" pageId="7" pageNumber="138" rowspan-0="1">
<th id="76B70157E235E5C9FEAD1A5AD5ACFE96" box="[329,426,291,311]" gridcol="1" gridrow="0" pageId="7" pageNumber="138">Longueur</th>
<th id="76B70157E235E5C9FE0D1A5AD63EFE96" box="[489,568,291,311]" gridcol="2" gridrow="0" pageId="7" pageNumber="138">
<emphasis id="B922B67BE2351A31FE0D1A5AD63EFE96" bold="true" box="[489,568,291,311]" pageId="7" pageNumber="138">Largeur</emphasis>
</th>
</tr>
<tr id="3566682BE235E5C9FF951A2CD63EFEC8" box="[113,568,341,361]" gridrow="1" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951A2CD4C1FEC8" box="[113,199,341,361]" gridcol="0" gridrow="1" pageId="7" pageNumber="138">Holotype</th>
<td id="76B70157E235E5C9FEAD1A2CD5ACFEC8" box="[329,426,341,361]" gridcol="1" gridrow="1" pageId="7" pageNumber="138">24,1</td>
<td id="76B70157E235E5C9FE0D1A2CD63EFEC8" box="[489,568,341,361]" gridcol="2" gridrow="1" pageId="7" pageNumber="138">12,2</td>
</tr>
<tr id="3566682BE235E5C9FF951A14D63EFE20" box="[113,568,365,385]" gridrow="2" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951A14D4C1FE20" box="[113,199,365,385]" gridcol="0" gridrow="2" pageId="7" pageNumber="138">Paratype</th>
<td id="76B70157E235E5C9FEAD1A14D5ACFE20" box="[329,426,365,385]" gridcol="1" gridrow="2" pageId="7" pageNumber="138">30,4</td>
<td id="76B70157E235E5C9FE0D1A14D63EFE20" box="[489,568,365,385]" gridcol="2" gridrow="2" pageId="7" pageNumber="138">14,6</td>
</tr>
<tr id="3566682BE235E5C9FF951AFCD63EFE38" box="[113,568,389,409]" gridrow="3" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951AFCD4C1FE38" box="[113,199,389,409]" gridcol="0" gridrow="3" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1AFCD5ACFE38" box="[329,426,389,409]" gridcol="1" gridrow="3" pageId="7" pageNumber="138">28,1</td>
<td id="76B70157E235E5C9FE0D1AFCD63EFE38" box="[489,568,389,409]" gridcol="2" gridrow="3" pageId="7" pageNumber="138">13,6</td>
</tr>
<tr id="3566682BE235E5C9FF951AE4D63EFE10" box="[113,568,413,433]" gridrow="4" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951AE4D4C1FE10" box="[113,199,413,433]" gridcol="0" gridrow="4" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1AE4D5ACFE10" box="[329,426,413,433]" gridcol="1" gridrow="4" pageId="7" pageNumber="138">27,5</td>
<td id="76B70157E235E5C9FE0D1AE4D63EFE10" box="[489,568,413,433]" gridcol="2" gridrow="4" pageId="7" pageNumber="138">13,6</td>
</tr>
<tr id="3566682BE235E5C9FF951ACCD63EFE68" box="[113,568,437,457]" gridrow="5" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951ACCD4C1FE68" box="[113,199,437,457]" gridcol="0" gridrow="5" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1ACCD5ACFE68" box="[329,426,437,457]" gridcol="1" gridrow="5" pageId="7" pageNumber="138">27,1</td>
<td id="76B70157E235E5C9FE0D1ACCD63EFE68" box="[489,568,437,457]" gridcol="2" gridrow="5" pageId="7" pageNumber="138">13,1</td>
</tr>
<tr id="3566682BE235E5C9FF951AB4D63EFE40" box="[113,568,461,481]" gridrow="6" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951AB4D4C1FE40" box="[113,199,461,481]" gridcol="0" gridrow="6" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1AB4D5ACFE40" box="[329,426,461,481]" gridcol="1" gridrow="6" pageId="7" pageNumber="138">26,7</td>
<td id="76B70157E235E5C9FE0D1AB4D63EFE40" box="[489,568,461,481]" gridcol="2" gridrow="6" pageId="7" pageNumber="138">13,6</td>
</tr>
<tr id="3566682BE235E5C9FF951A9CD63EFE58" box="[113,568,485,505]" gridrow="7" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951A9CD4C1FE58" box="[113,199,485,505]" gridcol="0" gridrow="7" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1A9CD5ACFE58" box="[329,426,485,505]" gridcol="1" gridrow="7" pageId="7" pageNumber="138">26,6</td>
<td id="76B70157E235E5C9FE0D1A9CD63EFE58" box="[489,568,485,505]" gridcol="2" gridrow="7" pageId="7" pageNumber="138">13,7</td>
</tr>
<tr id="3566682BE235E5C9FF951A84D63EFDB0" box="[113,568,509,529]" gridrow="8" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951A84D4C1FDB0" box="[113,199,509,529]" gridcol="0" gridrow="8" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1A84D5ACFDB0" box="[329,426,509,529]" gridcol="1" gridrow="8" pageId="7" pageNumber="138">26,5</td>
<td id="76B70157E235E5C9FE0D1A84D63EFDB0" box="[489,568,509,529]" gridcol="2" gridrow="8" pageId="7" pageNumber="138">14,7</td>
</tr>
<tr id="3566682BE235E5C9FF95196CD63EFD88" box="[113,568,533,553]" gridrow="9" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF95196CD4C1FD88" box="[113,199,533,553]" gridcol="0" gridrow="9" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD196CD5ACFD88" box="[329,426,533,553]" gridcol="1" gridrow="9" pageId="7" pageNumber="138">26,4</td>
<td id="76B70157E235E5C9FE0D196CD63EFD88" box="[489,568,533,553]" gridcol="2" gridrow="9" pageId="7" pageNumber="138">13,6</td>
</tr>
<tr id="3566682BE235E5C9FF951954D63EFDE0" box="[113,568,557,577]" gridrow="10" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951954D4C1FDE0" box="[113,199,557,577]" gridcol="0" gridrow="10" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1954D5ACFDE0" box="[329,426,557,577]" gridcol="1" gridrow="10" pageId="7" pageNumber="138">26,3</td>
<td id="76B70157E235E5C9FE0D1954D63EFDE0" box="[489,568,557,577]" gridcol="2" gridrow="10" pageId="7" pageNumber="138">12,2</td>
</tr>
<tr id="3566682BE235E5C9FF95193CD63EFDF8" box="[113,568,581,601]" gridrow="11" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF95193CD4C1FDF8" box="[113,199,581,601]" gridcol="0" gridrow="11" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD193CD5ACFDF8" box="[329,426,581,601]" gridcol="1" gridrow="11" pageId="7" pageNumber="138">25,6</td>
<td id="76B70157E235E5C9FE0D193CD63EFDF8" box="[489,568,581,601]" gridcol="2" gridrow="11" pageId="7" pageNumber="138">13,7</td>
</tr>
<tr id="3566682BE235E5C9FF951924D63EFDD0" box="[113,568,605,625]" gridrow="12" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951924D4C1FDD0" box="[113,199,605,625]" gridcol="0" gridrow="12" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1924D5ACFDD0" box="[329,426,605,625]" gridcol="1" gridrow="12" pageId="7" pageNumber="138">25,6</td>
<td id="76B70157E235E5C9FE0D1924D63EFDD0" box="[489,568,605,625]" gridcol="2" gridrow="12" pageId="7" pageNumber="138">12,5</td>
</tr>
<tr id="3566682BE235E5C9FF95190CD63EFD28" box="[113,568,629,649]" gridrow="13" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF95190CD4C1FD28" box="[113,199,629,649]" gridcol="0" gridrow="13" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD190CD5ACFD28" box="[329,426,629,649]" gridcol="1" gridrow="13" pageId="7" pageNumber="138">24,8</td>
<td id="76B70157E235E5C9FE0D190CD63EFD28" box="[489,568,629,649]" gridcol="2" gridrow="13" pageId="7" pageNumber="138">13</td>
</tr>
<tr id="3566682BE235E5C9FF9519F4D63EFD00" box="[113,568,653,673]" gridrow="14" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF9519F4D4C1FD00" box="[113,199,653,673]" gridcol="0" gridrow="14" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD19F4D5ACFD00" box="[329,426,653,673]" gridcol="1" gridrow="14" pageId="7" pageNumber="138">24</td>
<td id="76B70157E235E5C9FE0D19F4D63EFD00" box="[489,568,653,673]" gridcol="2" gridrow="14" pageId="7" pageNumber="138">12</td>
</tr>
<tr id="3566682BE235E5C9FF9519DCD63EFD18" box="[113,568,677,697]" gridrow="15" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF9519DCD4C1FD18" box="[113,199,677,697]" gridcol="0" gridrow="15" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD19DCD5ACFD18" box="[329,426,677,697]" gridcol="1" gridrow="15" pageId="7" pageNumber="138">23,8</td>
<td id="76B70157E235E5C9FE0D19DCD63EFD18" box="[489,568,677,697]" gridcol="2" gridrow="15" pageId="7" pageNumber="138">13,8</td>
</tr>
<tr id="3566682BE235E5C9FF9519C4D63EFD70" box="[113,568,701,721]" gridrow="16" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF9519C4D4C1FD70" box="[113,199,701,721]" gridcol="0" gridrow="16" pageId="7" pageNumber="138">Allotype</th>
<td id="76B70157E235E5C9FEAD19C4D5ACFD70" box="[329,426,701,721]" gridcol="1" gridrow="16" pageId="7" pageNumber="138">25,3</td>
<td id="76B70157E235E5C9FE0D19C4D63EFD70" box="[489,568,701,721]" gridcol="2" gridrow="16" pageId="7" pageNumber="138">13,8</td>
</tr>
<tr id="3566682BE235E5C9FF9519ACD63EFD48" box="[113,568,725,745]" gridrow="17" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF9519ACD4C1FD48" box="[113,199,725,745]" gridcol="0" gridrow="17" pageId="7" pageNumber="138">Paratype</th>
<td id="76B70157E235E5C9FEAD19ACD5ACFD48" box="[329,426,725,745]" gridcol="1" gridrow="17" pageId="7" pageNumber="138">26,2</td>
<td id="76B70157E235E5C9FE0D19ACD63EFD48" box="[489,568,725,745]" gridcol="2" gridrow="17" pageId="7" pageNumber="138">14,2</td>
</tr>
<tr id="3566682BE235E5C9FF951994D63EFCA0" box="[113,568,749,769]" gridrow="18" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951994D4C1FCA0" box="[113,199,749,769]" gridcol="0" gridrow="18" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1994D5ACFCA0" box="[329,426,749,769]" gridcol="1" gridrow="18" pageId="7" pageNumber="138">26,1</td>
<td id="76B70157E235E5C9FE0D1994D63EFCA0" box="[489,568,749,769]" gridcol="2" gridrow="18" pageId="7" pageNumber="138">14,2</td>
</tr>
<tr id="3566682BE235E5C9FF95187CD63EFCB8" box="[113,568,773,793]" gridrow="19" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF95187CD4C1FCB8" box="[113,199,773,793]" gridcol="0" gridrow="19" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD187CD5ACFCB8" box="[329,426,773,793]" gridcol="1" gridrow="19" pageId="7" pageNumber="138">25,7</td>
<td id="76B70157E235E5C9FE0D187CD63EFCB8" box="[489,568,773,793]" gridcol="2" gridrow="19" pageId="7" pageNumber="138">13</td>
</tr>
<tr id="3566682BE235E5C9FF951864D63EFC90" box="[113,568,797,817]" gridrow="20" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951864D4C1FC90" box="[113,199,797,817]" gridcol="0" gridrow="20" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1864D5ACFC90" box="[329,426,797,817]" gridcol="1" gridrow="20" pageId="7" pageNumber="138">24,6</td>
<td id="76B70157E235E5C9FE0D1864D63EFC90" box="[489,568,797,817]" gridcol="2" gridrow="20" pageId="7" pageNumber="138">12,8</td>
</tr>
<tr id="3566682BE235E5C9FF95184CD63EFCE8" box="[113,568,821,841]" gridrow="21" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF95184CD4C1FCE8" box="[113,199,821,841]" gridcol="0" gridrow="21" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD184CD5ACFCE8" box="[329,426,821,841]" gridcol="1" gridrow="21" pageId="7" pageNumber="138">23,8</td>
<td id="76B70157E235E5C9FE0D184CD63EFCE8" box="[489,568,821,841]" gridcol="2" gridrow="21" pageId="7" pageNumber="138">13,2</td>
</tr>
<tr id="3566682BE235E5C9FF951834D63EFCC0" box="[113,568,845,865]" gridrow="22" pageId="7" pageNumber="138">
<th id="76B70157E235E5C9FF951834D4C1FCC0" box="[113,199,845,865]" gridcol="0" gridrow="22" pageId="7" pageNumber="138">–</th>
<td id="76B70157E235E5C9FEAD1834D5ACFCC0" box="[329,426,845,865]" gridcol="1" gridrow="22" pageId="7" pageNumber="138">23,5</td>
<td id="76B70157E235E5C9FE0D1834D63EFCC0" box="[489,568,845,865]" gridcol="2" gridrow="22" pageId="7" pageNumber="138">12,7</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE96A69E2351A31FF9518DAD636FBF5" blockId="7.[113,647,931,1391]" pageId="7" pageNumber="138">
LOCALITÉ-
<typeStatus id="54EDD4CBE2351A31FF3E18D1D50EFC19" box="[218,264,936,952]" pageId="7" pageNumber="138">TYPE</typeStatus>
. —
<materialsCitation id="3B3E6034E2351A31FEDC18DDD636FBF5" collectingDate="1995-11-19" collectorName="P. Jolivet" country="China" latitude="19.5" location="Danzhou" longLatPrecision="1271" longitude="109.55" pageId="7" pageNumber="138" specimenCount="22" stateProvince="Hainan">
<collectingCountry id="F3412AF9E2351A31FEDC18DDD57EFC1D" box="[312,376,932,956]" name="China" pageId="7" pageNumber="138">Chine</collectingCountry>
,
<collectingRegion id="4992A48BE2351A31FE6018DAD5D4FC1B" box="[388,466,931,954]" country="China" name="Hainan" pageId="7" pageNumber="138">Haïnan</collectingRegion>
, sur une colline à l’ouest de la ville nouvelle (et universitaire) de Danzhou (
, sur une colline à l’ouest de la ville nouvelle (et universitaire) de
<location id="8E893CB2E2351A31FF9518AFD4C8FC4F" LSID="urn:lsid:plazi:treatment:03FFDB7FE2371A3EFCFA1E12D582FB2B:8E893CB2E2351A31FF9518AFD4C8FC4F" box="[113,206,982,1006]" country="China" latitude="19.5" longLatPrecision="1271" longitude="109.55" name="Danzhou" pageId="7" pageNumber="138" stateProvince="Hainan">Danzhou</location>
(
<geoCoordinate id="EE620CAEE2351A31FF3B18AFD547FC4F" box="[223,321,982,1006]" degrees="109" direction="east" minutes="33" orientation="longitude" pageId="7" pageNumber="138" precision="925" value="109.55">109°33’E</geoCoordinate>
,
<geoCoordinate id="EE620CAEE2351A31FEA918AFD5A1FC4F" box="[333,423,982,1006]" degrees="19" direction="north" minutes="30" orientation="latitude" pageId="7" pageNumber="138" precision="925" value="19.5">19°30’N</geoCoordinate>
), dans une plantation d’hévéas. 22 exemplaires (15 et 7) ont été récoltés le
<date id="FFE84CA9E2351A31FF3E1F70D54AFB80" box="[218,332,1033,1057]" pageId="7" pageNumber="138" value="1995-11-19">19.XI.1995</date>
par P. Jolivet, au pied d’hévéas où ils pullulent et creusent des petits terriers dans lesquels ils ont été capturés (P. Jolivet
), dans une plantation d’hévéas.
<specimenCount id="9D50A1E0E2351A31FF311889D4E9FBA9" box="[213,239,1008,1032]" pageId="7" pageNumber="138">22</specimenCount>
exemplaires (15 et 7) ont été récoltés le
<date id="FFE84CA9E2351A31FF3E1F70D54AFB80" box="[218,332,1033,1057]" pageId="7" pageNumber="138" value="1995-11-19">
<collectingDate id="EFACB541E2351A31FF3E1F70D54AFB80" box="[218,332,1033,1057]" pageId="7" pageNumber="138" value="1995-11-19">19.XI.1995</collectingDate>
</date>
par
<collectorName id="26A30FBFE2351A31FE9F1F71D5DEFB80" box="[379,472,1032,1057]" pageId="7" pageNumber="138">P. Jolivet</collectorName>
, au pied d’hévéas où ils pullulent et creusent des petits terriers dans lesquels ils ont été capturés (P. Jolivet
<emphasis id="B922B67BE2351A31FE021F42D61AFBF5" box="[486,540,1083,1108]" italics="true" pageId="7" pageNumber="138">in litt</emphasis>
.).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34C39E2E2351A31FF951F1DD4D6FB43" pageId="7" pageNumber="138" type="etymology">
<paragraph id="8BE96A69E2351A31FF951F1DD4D6FB43" blockId="7.[113,647,931,1391]" pageId="7" pageNumber="138">ÉTYMOLOGIE. — Espèce nouvelle bien cordialement dédiée à son inventeur, le Dr Pierre Jolivet, infatigable voyageur et grand pourvoyeur de matériel entomologique et arthropodologique des collections du Muséum.</paragraph>
</subSubSection>
<subSubSection id="C34C39E2E2351A31FF951F88D63BFACE" pageId="7" pageNumber="138" type="diagnosis">
<paragraph id="8BE96A69E2351A31FF951F88D63BFACE" blockId="7.[113,647,931,1391]" pageId="7" pageNumber="138">
DIAGNOSE. — Se distingue des deux autres espèces d’Haïnan (
<taxonomicName id="4C5611EAE2351A31FF3F1E72D538FA82" authorityName="Gressitt" authorityYear="1941" box="[219,318,1291,1315]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="hainani">
@ -200,6 +304,8 @@ et
</taxonomicName>
) par sa plus petite taille, sa coloration vert foncé, les lobes coxaux de ses pattes ambulatoires plus étroits et la dent fémorale des paratélopodes du mâle nettement plus allongée.
</paragraph>
</subSubSection>
<subSubSection id="C34C39E2E2351A3EFF951ED6D582FB2B" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138" type="description">
<paragraph id="8BE96A69E2351A31FF951ED6D504FA66" blockId="7.[113,647,1455,1706]" box="[113,258,1455,1481]" pageId="7" pageNumber="138">DESCRIPTION</paragraph>
<paragraph id="8BE96A69E2351A31FF951EB6D5DBFA48" blockId="7.[113,647,1455,1706]" box="[113,477,1487,1514]" pageId="7" pageNumber="138">
<emphasis id="B922B67BE2351A31FF951EB6D5DBFA48" box="[113,477,1487,1514]" italics="true" pageId="7" pageNumber="138">Caractères morphologiques externes</emphasis>
@ -207,18 +313,22 @@ et
<paragraph id="8BE96A69E2351A31FF951E89D550F90B" blockId="7.[113,647,1455,1706]" pageId="7" pageNumber="138">Coloration générale brun-verdâtre très foncé (presque noire), non seulement des plaques tergales et de la tête, mais aussi des appendices, tant ambulatoires que copulateurs (télopodes et paratélopodes); seuls les pleurites sont un peu plus clairs (brun-olivâtre).</paragraph>
<paragraph id="8BE96A69E2351A31FD411BA9D00AFF4B" blockId="7.[677,1210,207,810]" box="[677,1036,207,234]" pageId="7" pageNumber="138">
Mensurations: voir le
<tableCitation id="C6D45FD2E2351A31FC7C1BA9D000FF4B" box="[920,1030,208,234]" captionStart="TABLEAU" captionStartId="7.[113,124,206,223]" captionText="TABLEAU 1. — Longueur et largeur du corps en mm des 22 exemplaires de Prionobelum joliveti n. sp." pageId="7" pageNumber="138">Tableau 1</tableCitation>
<tableCitation id="C6D45FD2E2351A31FC7C1BA9D000FF4B" box="[920,1030,208,234]" captionStart="TABLEAU" captionStartId="7.[113,124,206,223]" captionTargetPageId="7" captionText="TABLEAU 1. — Longueur et largeur du corps en mm des 22 exemplaires de Prionobelum joliveti n. sp." httpUri="http://table.plazi.org/id/DF293AE1E2351A31FF951BB7D5EEFF55" pageId="7" pageNumber="138" tableUuid="DF293AE1E2351A31FF951BB7D5EEFF55">Tableau 1</tableCitation>
.
</paragraph>
<paragraph id="8BE96A69E2351A31FD411B96D028FD28" blockId="7.[677,1210,207,810]" pageId="7" pageNumber="138">
La tête (Fig. 4A) possède des yeux à nombreux ocelles (entre 40 et 50). Antennes à antennomères III à VI très courts et évasés, beaucoup plus larges distalement que basalement; antennomère subterminal renflé mais non évasé, 1,5 fois plus long que large; sa partie distale, oblique, est coiffée du très court et circulaire antennomère terminal qui est garni d’une vingtaine de cônes. La face est parsemée de grandes et profondes ponctuations et la pointe de la dent du labre, petite, est en retrait du bord de celui-ci; les bords antérolatéraux de la tête sont largement échancrés (ils sont presque droits chez
La tête (
<figureCitation id="136D76ECE2351A31FCE71B89D75CFEAB" box="[771,858,240,266]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="7" pageNumber="138">Fig. 4A</figureCitation>
) possède des yeux à nombreux ocelles (entre 40 et 50). Antennes à antennomères III à VI très courts et évasés, beaucoup plus larges distalement que basalement; antennomère subterminal renflé mais non évasé, 1,5 fois plus long que large; sa partie distale, oblique, est coiffée du très court et circulaire antennomère terminal qui est garni d’une vingtaine de cônes. La face est parsemée de grandes et profondes ponctuations et la pointe de la dent du labre, petite, est en retrait du bord de celui-ci; les bords antérolatéraux de la tête sont largement échancrés (ils sont presque droits chez
<taxonomicName id="4C5611EAE2351A31FC4B1909D018FD2B" authorityName="Gressitt" authorityYear="1941" box="[943,1054,624,650]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="hainani">
<emphasis id="B922B67BE2351A31FC4B1909D018FD2B" box="[943,1054,624,650]" italics="true" pageId="7" pageNumber="138">P. hainani</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BE96A69E2351A31FD4119F6D71DFC88" blockId="7.[677,1210,207,810]" pageId="7" pageNumber="138">
Les pattes ambulatoires (Fig. 4B, C) sont du
Les pattes ambulatoires (
<figureCitation id="136D76ECE2351A31FC4B19E9D025FD0B" box="[943,1059,656,682]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="7" pageNumber="138">Fig. 4B, C</figureCitation>
) sont du
<typeStatus id="54EDD4CBE2351A31FB6F19F6D0BFFD08" box="[1163,1209,655,681]" pageId="7" pageNumber="138">type</typeStatus>
habituel du genre; à noter que le lobe latéral des hanches des pattes du premier tiers du corps est aussi long mais nettement plus étroit que chez
<taxonomicName id="4C5611EAE2351A31FD411869D712FC8B" authorityName="Gressitt" authorityYear="1941" box="[677,788,784,810]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="hainani">
@ -231,7 +341,9 @@ habituel du genre; à noter que le lobe latéral des hanches des pattes du premi
</paragraph>
<paragraph id="8BE96A69E2351A31FD411816D765FC6B" blockId="7.[677,1210,847,1706]" pageId="7" pageNumber="138">Pores des canaux déférents petits, en ovale transverse, situés dans le tiers basal de la face caudale des coxas des P.2.</paragraph>
<paragraph id="8BE96A69E2351A31FD4118A9D7D4FBA8" blockId="7.[677,1210,847,1706]" pageId="7" pageNumber="138">
Paratélopodes (Fig. 4E-G, P.22) se distinguant des autres
Paratélopodes (
<figureCitation id="136D76ECE2351A31FCB518A9D793FC4B" box="[849,917,976,1002]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="7" pageNumber="138">Fig. 4</figureCitation>
E-G, P.22) se distinguant des autres
<taxonomicName id="4C5611EAE2351A31FCF11889D79CFBAB" authorityName="Verhoeff" authorityYear="1924" box="[789,922,1008,1034]" class="Diplopoda" family="Zephroniidae" genus="Prionobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B922B67BE2351A31FCF11889D79CFBAB" box="[789,922,1008,1034]" italics="true" pageId="7" pageNumber="138">Prionobelum</emphasis>
</taxonomicName>
@ -239,9 +351,15 @@ par:
</paragraph>
<paragraph id="8BE96A69E2351A31FD411F76D72EFB28" blockId="7.[677,1210,847,1706]" pageId="7" pageNumber="138">– la présence d’une forte dent (d) aiguë prolongeant l’angle disto-caudal du processus fémoral dont il est séparé par une minuscule mais nette échancrure;</paragraph>
<paragraph id="8BE96A69E2351A31FD411FF6D055FB68" blockId="7.[677,1210,847,1706]" pageId="7" pageNumber="138">– par la forme en crochet du tibia, dont la pointe se situe en vis-à-vis de la dent fémorale;</paragraph>
<paragraph id="8BE96A69E2351A31FD411FB6D6D8FA2B" blockId="7.[677,1210,847,1706]" pageId="7" pageNumber="138">– par la quasi disparition, au tibiotarse, du tarse (Fig. 4D, G) qui, contrairement à la plupart des autres espèces, se fond au tibia et ne se reconnaît qu’à une faible trace oblique délimitant une petite aire disto-interne qui porte deux minuscules soies.</paragraph>
<paragraph id="8BE96A69E2351A31FD411FB6D6D8FA2B" blockId="7.[677,1210,847,1706]" pageId="7" pageNumber="138">
– par la quasi disparition, au tibiotarse, du tarse (
<figureCitation id="136D76ECE2351A31FD481F89D72CFAAB" box="[684,810,1264,1290]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="7" pageNumber="138">Fig. 4D, G</figureCitation>
) qui, contrairement à la plupart des autres espèces, se fond au tibia et ne se reconnaît qu’à une faible trace oblique délimitant une petite aire disto-interne qui porte deux minuscules soies.
</paragraph>
<paragraph id="8BE96A69E2351A31FD411EE9D088F9C8" blockId="7.[677,1210,847,1706]" pageId="7" pageNumber="138">
Télopodes (Fig. 4H, I, P.23) ne se distinguant pas fondamentalement de celui des autres espèces; le nombre de dents tibiales distinctes (12) est plus important que chez
Télopodes (
<figureCitation id="136D76ECE2351A31FCCE1EE9D7C5FA0B" box="[810,963,1423,1450]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="7" pageNumber="138">Fig. 4H, I, P</figureCitation>
.23) ne se distinguant pas fondamentalement de celui des autres espèces; le nombre de dents tibiales distinctes (12) est plus important que chez
<taxonomicName id="4C5611EAE2351A31FBFF1E89D088F9AB" authorityName="Gressitt" authorityYear="1941" box="[1051,1166,1520,1546]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="hainani">
<emphasis id="B922B67BE2351A31FBFF1E89D088F9AB" box="[1051,1166,1520,1546]" italics="true" pageId="7" pageNumber="138">P. hainani</emphasis>
</taxonomicName>
@ -254,19 +372,33 @@ de
<taxonomicName id="4C5611EAE2351A31FC901DE9D7EFF90B" authorityName="Gressitt" authorityYear="1941" box="[884,1001,1680,1706]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="hainani">
<emphasis id="B922B67BE2351A31FC901DE9D7EFF90B" box="[884,1001,1680,1706]" italics="true" pageId="7" pageNumber="138">P. hainani</emphasis>
</taxonomicName>
n’est pas toujours présente et lorsqu’elle existe, elle est plus ou moins grande et plus ou moins dépigmentée; parfois, elle se traduit par une légère déformation en dépression ou en troncature (Fig. 4I, J, N) de la face interne (dorsale) de l’extrémité du processus;
n’est pas toujours présente et lorsqu’elle existe, elle est plus ou moins grande et plus ou moins dépigmentée; parfois, elle se traduit par une légère déformation en dépression ou en troncature (
<figureCitation id="136D76ECE23A1A3EFE0F1A49D678FEE8" box="[491,638,303,330]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="8" pageNumber="139">Fig. 4I, J, N</figureCitation>
) de la face interne (dorsale) de l’extrémité du processus;
</paragraph>
<paragraph id="8BE96A69E23A1A3EFF961AF6D65CFC8B" blockId="8.[114,647,207,810]" pageId="8" pageNumber="139">
– la face externe, aplanie et de couleur claire, porte le plus souvent des petites formations membraneuses de structure variable (même entre les deux télopodes d’un même individu), de couleur claire (nommés « weissen Zipfel » par
<bibRefCitation id="EFC71798E23A1A3EFDD11969D4ADFDEB" author="ATTEMS C." pageId="8" pageNumber="139" pagination="60 - 73" refId="ref5578" refString="ATTEMS C. 1943. - Neue Sphaerotheriden des Wiener Museums. Annalen des Naturhistorischen Museum Wien 53: 60 - 73." type="journal article" year="1943">Attems 1943</bibRefCitation>
); ces formations manquent sur cinq des six télopodes de trois mâles (Fig. 4K), constituent une lame allongée et basse sur trois mâles (dont l’holotype, Fig. 4J), sont réduites à une minuscule dent sur deux mâles (Fig. 4L), sont assez fortement développées en une dent simple sur sept des huit télopodes de quatre mâles (Fig. 4M) ou en une double dent sur trois mâles (Fig. 4N).
); ces formations manquent sur cinq des six télopodes de trois mâles (
<figureCitation id="136D76ECE23A1A3EFE711929D5F7FDCB" box="[405,497,592,618]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="8" pageNumber="139">Fig. 4K</figureCitation>
), constituent une lame allongée et basse sur trois mâles (dont l’holotype,
<figureCitation id="136D76ECE23A1A3EFF1619E9D547FD0B" box="[242,321,656,682]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="8" pageNumber="139">Fig. 4J</figureCitation>
), sont réduites à une minuscule dent sur deux mâles (
<figureCitation id="136D76ECE23A1A3EFE6D19C9D5DAFD6B" box="[393,476,688,714]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="8" pageNumber="139">Fig. 4L</figureCitation>
), sont assez fortement développées en une dent simple sur sept des huit télopodes de quatre mâles (
<figureCitation id="136D76ECE23A1A3EFE1A1989D65FFCAB" box="[510,601,752,778]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="8" pageNumber="139">Fig. 4M</figureCitation>
) ou en une double dent sur trois mâles (
<figureCitation id="136D76ECE23A1A3EFE101869D64BFC8B" box="[500,589,784,810]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="8" pageNumber="139">Fig. 4N</figureCitation>
).
</paragraph>
<paragraph id="8BE96A69E23A1A3EFF961836D56EFCCB" blockId="8.[114,647,847,1162]" box="[114,360,847,874]" pageId="8" pageNumber="139">
<emphasis id="B922B67BE23A1A3EFF961836D56EFCCB" box="[114,360,847,874]" italics="true" pageId="8" pageNumber="139">Caractères de la femelle</emphasis>
</paragraph>
<paragraph id="8BE96A69E23A1A3EFF961816D582FB2B" blockId="8.[114,647,847,1162]" pageId="8" pageNumber="139">
Les vulves (Fig. 4O) montrent une conformation typique de
<taxonomicName id="4C5611EAE23A1A3EFF0E18E9D59EFC0B" authorityName="Brolemann" authorityYear="1913" box="[234,408,912,938]" pageId="8" pageNumber="131" rank="subFamily" subFamily="Sphaeropoeinae">Sphaeropoeinae</taxonomicName>
Les vulves (
<figureCitation id="136D76ECE23A1A3EFF0A1809D543FC2B" box="[238,325,880,906]" captionStart="FIG" captionStartId="6.[114,125,1561,1578]" captionTargetBox="[114,1211,177,1549]" captionTargetId="figure-23@6.[122,1201,188,1541]" captionTargetPageId="6" captionText="FIG. 4. — Prionobelum joliveti n. sp., holotype, paratypes; A, vue antéro-distale de la moitié droite de la tête d’un paratype; B, vue orale d’une hanche de la P.6 gauche du holotype; C, extrémité du tarse d’une patte ambulatoire (P.6) du même; D, vue caudale du tibiotarse isolé du paratélopode (P. 22) droit du même; E, vue caudale du paratélopode (P.22) droit du même; F, vue orale du même; G, vue interne du même; H, vue orale de la moitié gauche du syncoxite et du télopode (P.23) gauche du même; I, vue caudale du télopode (P. 23) isolé du même; J-N, variations de la forme du processus fémoral du télopode (P.23), en vue orale; J, holotype; K-N, paratypes; O, allotype, vue caudale de la hanche gauche avec la vulve correspondante. Abréviation: d, dent. Échelles: A-C, E-O, 1 mm; D, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.15008578" httpUri="https://zenodo.org/record/15008578/files/figure.png" pageId="8" pageNumber="139">Fig. 4O</figureCitation>
) montrent une conformation typique de
<taxonomicName id="4C5611EAE23A1A3EFF0E18E9D59EFC0B" authorityName="Brolemann" authorityYear="1913" baseAuthorityName="Broelemann" baseAuthorityYear="1913" box="[234,408,912,938]" class="Diplopoda" family="Zephroniidae" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="131" phylum="Arthropoda" rank="subFamily" subFamily="Sphaeropoeinae">Sphaeropoeinae</taxonomicName>
, chez qui la bourse est embrassée par l’opercule; ce dernier présente une forme trés allongée, mais non élargie distalement, qui paraît être caractéristique du genre. Les hanches, conjointes mais séparées, ne présentent aucune particularité. Le sternite est inexistant médialement, réduit et fondu aux poches trachéennes latéralement.
</paragraph>
</subSubSection>

586
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<mods:namePart id="E7622DC15AC8A7206CD71B9FA259136D">Dellacasa, Marco</mods:namePart>
<mods:affiliation id="E4B871F3B7AB6A65135104B848A04BA0">Centro Interdipartimentale, Museo di Storia Naturale e del Territorio, Università di Pisa, via Roma 79, I- 56011 Calci, Pisa (Italy) dellacasa @ museo. unipi. it</mods:affiliation>
<mods:nameIdentifier id="9EC2604B187E19AAF84E6B67C5F4DD1E" type="email">dellacasa@museo.unipi.it</mods:nameIdentifier>
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<mods:namePart id="ED3BE272BAA19DBE663EB83FBFDD09B3">Dellacasa, Giovanni</mods:namePart>
<mods:affiliation id="E19224B360E869110725B76F81CAD994">CP 921, I- 16121 Genova (Italy)</mods:affiliation>
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<mods:typeOfResource id="EDA8287776DC4DC6F491FB129A2F6146">text</mods:typeOfResource>
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<mods:title id="C79011682FF668C026EE7D352C18B905">Zoosystema</mods:title>
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<mods:part id="D597644020A189EE90E6403A3BD07F60">
<mods:date id="4F28EEF20677DD9BD3751C62FB7F587C">2008</mods:date>
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<mods:number id="74521262C293F8625919EDB1FB96C369">2008-02-21</mods:number>
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<mods:number id="29FAD1470A6589C6E7C8CAD0EE454396">30</mods:number>
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<mods:number id="2368881BC72707BED31563D49D9C4674">3</mods:number>
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<mods:start id="7A176F0242F853825B6B7DAE99A59057">629</mods:start>
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<mods:identifier id="6405C5154420C86620EA577FC060B187" type="DOI">10.5281/zenodo.5402557</mods:identifier>
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<subSubSection id="59F3E4A2DC17FFD2BCA15135FD97D0F6" box="[138,622,1387,1415]" pageId="8" pageNumber="637" type="nomenclature">
<paragraph id="1156B729DC17FFD2BCA15135FD97D0F6" blockId="8.[138,622,1387,1447]" box="[138,622,1387,1415]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BCA15135FDFED0F6" ID-CoL="69QNB" authority="(Olivier, 1789)" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[138,519,1387,1415]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus" status="comb. nov.">
<emphasis id="239D6B3BDC17FFD2BCA15135FEA4D0F7" bold="true" box="[138,349,1387,1414]" italics="true" pageId="8" pageNumber="637">Chelotrupes momus</emphasis>
(
<bibRefCitation id="7578CAD8DC17FFD2BD475132FE06D0F6" author="OLIVIER A. G." box="[364,511,1388,1415]" pageId="8" pageNumber="637" refId="ref11485" refString="OLIVIER A. G. 1789. - Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Baudouin, Paris, 1, 190 p." type="book" year="1789">Olivier, 1789</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38AED640DC17FFD2BE255132FD97D0F6" box="[526,622,1388,1415]" pageId="8" pageNumber="637" rank="species">n. comb.</taxonomicNameLabel>
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<subSubSection id="59F3E4A2DC17FFD2BD2151D2FE14D0D6" box="[266,493,1420,1447]" pageId="8" pageNumber="637" type="description">
<paragraph id="1156B729DC17FFD2BD2151D2FE14D0D6" blockId="8.[138,622,1387,1447]" box="[266,493,1420,1447]" pageId="8" pageNumber="637">
(
<figureCitation id="89D2ABACDC17FFD2BD3A51D2FE4ED0D6" box="[273,439,1420,1447]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="8" pageNumber="637">Figs 1C, F, I, L</figureCitation>
;
<figureCitation id="89D2ABACDC17FFD2BDE851D2FE1CD0D6" box="[451,485,1420,1447]" captionStart="FIG" captionStartId="8.[113,124,732,749]" captionTargetBox="[204,1069,228,686]" captionTargetId="figure-20@8.[188,1134,199,724]" captionTargetPageId="8" captionText="FIG. 2. — Aedeagus in dorsal view: A, Chelotrupes hiostius (Gené, 1836) n. comb., Costa Verde, Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; B, C. matutinalis (Baudi, 1870) n. comb.,Costa Verde,Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; C, C. momus (Olivier, 1789) n. comb., Andalusia, Cadiz, Tarifa. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5402561" httpUri="https://zenodo.org/record/5402561/files/figure.png" pageId="8" pageNumber="637">2C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC17FFD3BC5A519BFE2AD103" lastPageId="9" lastPageNumber="638" pageId="8" pageNumber="637" type="reference_group">
<paragraph id="1156B729DC17FFD2BC5A519BFEEBD086" blockId="8.[113,647,1477,1527]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BC5A519BFE27D0AC" authority="Olivier, 1789: 60" authorityName="Olivier" authorityPageNumber="60" authorityYear="1789" box="[113,478,1477,1501]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC17FFD2BC5A519BFEDDD0AD" box="[113,292,1477,1501]" italics="true" pageId="8" pageNumber="637">Scarabaeus momus</emphasis>
<bibRefCitation id="7578CAD8DC17FFD2BD07519BFE27D0AC" author="OLIVIER A. G." box="[300,478,1477,1501]" pageId="8" pageNumber="637" refId="ref11485" refString="OLIVIER A. G. 1789. - Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Baudouin, Paris, 1, 190 p." type="book" year="1789">Olivier, 1789: 60</bibRefCitation>
</taxonomicName>
. —
<bibRefCitation id="7578CAD8DC17FFD2BE21519BFF62D086" author="ILLIGER J. K. W." pageId="8" pageNumber="637" refId="ref11152" refString="ILLIGER J. K. W. 1800. - Olivier's Entomologie oder Naturgeschichte der Insekten mit ihren Gattungs- und ArtMerkmalen, ihrer Beschreibung und Synonymie. Kafer. Uebersetzt und mit Zusatzen und Anmerkungen. K. Reichard, Braunschweig, 1, 309 p." type="book" year="1800">Illiger 1800: 152</bibRefCitation>
;
<bibRefCitation id="7578CAD8DC17FFD2BC8D5181FEF7D086" author="ILLIGER J. K. W." box="[166,270,1503,1527]" pageId="8" pageNumber="637" pagination="186 - 258" refId="ref11199" refString="ILLIGER J. K. W. 1803. - Verzeichniss der in Portugall einheimischen Kafer. Erste Lieferung. Magazin fur Insektenkunde (Illiger) 2: 186 - 258." type="journal article" year="1803">1803: 208</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BC5A5251FE0ED356" blockId="8.[113,503,1551,1575]" box="[113,503,1551,1575]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BC5A5251FED9D357" authorityName="Olivier" authorityYear="1789" box="[113,288,1551,1575]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC17FFD2BC5A5251FED9D357" box="[113,288,1551,1575]" italics="true" pageId="8" pageNumber="637">Scarabaeus momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BD125251FE0AD356" author="FABRICIUS J. C." box="[313,499,1551,1575]" pageId="8" pageNumber="637" refId="ref10952" refString="FABRICIUS J. C. 1792. - Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Hafniae, 538 p." type="book" year="1792">Fabricius 1792: 13</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BC5A5260FE0DD327" blockId="8.[113,500,1598,1622]" box="[113,500,1598,1622]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BC5A5260FED2D327" box="[113,299,1598,1622]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC17FFD2BC5A5260FED2D327" box="[113,299,1598,1622]" italics="true" pageId="8" pageNumber="637">Ceratophyus momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BD6E5260FE09D327" author="STURM J." box="[325,496,1598,1622]" pageId="8" pageNumber="637" refId="ref11809" refString="STURM J. 1843. - Catalog der Kaefer-Sammlung. Nurnberg, 386 p." type="book" year="1843">Sturm 1843: 113</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BC5A5230FDCFD3D0" blockId="8.[112,646,1646,1697]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BC5A5230FD9AD3F7" authority="Mulsant &amp; Godard, 1855: 1" authorityName="Mulsant &amp; Godard" authorityPageNumber="1" authorityYear="1855" box="[113,611,1646,1670]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="239D6B3BDC17FFD2BC5A5230FEB7D3F7" box="[113,334,1646,1670]" italics="true" pageId="8" pageNumber="637">Ceratophyus laevipennis</emphasis>
<bibRefCitation id="7578CAD8DC17FFD2BD785230FD9AD3F7" author="MULSANT E. &amp; GODARD A." box="[339,611,1646,1670]" pageId="8" pageNumber="637" pagination="1 - 8" refId="ref11458" refString="MULSANT E. &amp; GODARD A. 1855. - Description de quelques Coleopteres nouveaux ou peu connus. Opuscules entomologiques 6: 1 - 8." type="journal article" year="1855">Mulsant &amp; Godard, 1855: 1</bibRefCitation>
</taxonomicName>
. —
<bibRefCitation id="7578CAD8DC17FFD2BC5B52D7FEBAD3D0" author="BOUCOMONT A." box="[112,323,1673,1697]" pageId="8" pageNumber="637" refId="ref10730" refString="BOUCOMONT A. 1912. - Scarabaeidae: Taurocerastinae, Geotrupinae. Pars 46. Coleopterorum Catalogus auspiciis et auxilio. W. Junk, Berlin, 47 p." type="book" year="1912">Boucomont 1912: 22</bibRefCitation>
(as synonym of
<taxonomicName id="D6E9CCAADC17FFD2BDCD52D7FDD1D3D1" box="[486,552,1673,1696]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC17FFD2BDCD52D7FDD1D3D1" box="[486,552,1673,1696]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E5711FCB2D6F3" blockId="8.[675,1209,847,898]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E5711FC30D616" authorityName="Germar" authorityYear="1823" box="[677,969,847,871]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus" subGenus="Minotaurus">
<emphasis id="239D6B3BDC17FFD2BE8E5711FCF9D617" box="[677,768,847,870]" italics="true" pageId="8" pageNumber="637">Geotrupes</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF27570EFC83D616" box="[780,890,848,871]" italics="true" pageId="8" pageNumber="637">Minotaurus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BFAC570EFC30D616" box="[903,969,848,871]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BFF45711FB60D616" author="MARSEUL S. A." box="[991,1177,847,871]" pageId="8" pageNumber="637" refId="ref11347" refString="MARSEUL S. A. 1863. - Catalogue des Coleopteres d'Europe et du Bassin de la Mediterranee en Afrique &amp; en Asie. 2 nd ed. Chez l'auteur, Ach. Deyrolle, Paris, 300 p." type="book" year="1863">Marseul 1863: 122</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC17FFD2BE885734FCBED6F3" author="REITTER E." box="[675,839,874,898]" pageId="8" pageNumber="637" pagination="93 - 102" refId="ref11551" refString="REITTER E. 1883. - [Scarabaeidae], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae et Caucasi. 3 rd ed. W. Janson, London; Libraria Nicolai, Berolini; Luc. Buquet, Paris: 93 - 102." type="book chapter" year="1883">Reitter 1883: 97</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E57C7FC58D6BA" blockId="8.[677,1210,921,971]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E57C7FBFFD6C0" box="[677,1030,921,945]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="laevipennis" subGenus="Minotaurus">
<emphasis id="239D6B3BDC17FFD2BE8E57C7FCFDD6C1" box="[677,772,921,944]" italics="true" pageId="8" pageNumber="637">Geotrupes</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF3E57C7FC71D6C1" box="[789,904,921,944]" italics="true" pageId="8" pageNumber="637">Minotaurus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BFB157C7FBFFD6C0" box="[922,1030,921,945]" italics="true" pageId="8" pageNumber="637">laevipennis</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2B80F57C7FD36D6BA" author="MARSEUL S. A." pageId="8" pageNumber="637" refId="ref11347" refString="MARSEUL S. A. 1863. - Catalogue des Coleopteres d'Europe et du Bassin de la Mediterranee en Afrique &amp; en Asie. 2 nd ed. Chez l'auteur, Ach. Deyrolle, Paris, 300 p." type="book" year="1863">Marseul 1863: 122</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC17FFD2BED257EDFC64D6BA" author="REITTER E." box="[761,925,947,971]" pageId="8" pageNumber="637" pagination="93 - 102" refId="ref11551" refString="REITTER E. 1883. - [Scarabaeidae], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae et Caucasi. 3 rd ed. W. Janson, London; Libraria Nicolai, Berolini; Luc. Buquet, Paris: 93 - 102." type="book chapter" year="1883">Reitter 1883: 97</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E57BCFB64D68B" blockId="8.[677,1181,994,1018]" box="[677,1181,994,1018]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E57BCFC1BD68B" box="[677,994,994,1018]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus" subGenus="Chelotrupes">
<emphasis id="239D6B3BDC17FFD2BE8E57BCFCEFD68B" box="[677,790,994,1018]" italics="true" pageId="8" pageNumber="637">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF0F57BCFC68D68B" box="[804,913,994,1018]" italics="true" pageId="8" pageNumber="637">Chelotrupes</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BF8B57BDFC1BD68B" box="[928,994,995,1018]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BFD757BCFB60D68B" author="JEKEL H." box="[1020,1177,994,1018]" pageId="8" pageNumber="637" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">Jekel 1866: 550</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E504CFBEED15B" blockId="8.[677,1047,1042,1066]" box="[677,1047,1042,1066]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E504CFCB1D158" authorityName="Germar" authorityYear="1823" box="[677,840,1042,1065]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC17FFD2BE8E504CFCB1D158" box="[677,840,1042,1065]" italics="true" pageId="8" pageNumber="637">Geotrupes momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BF49504CFBEAD15B" author="MARSEUL S. A." box="[866,1043,1042,1066]" pageId="8" pageNumber="637" pagination="1 - 131" refId="ref11387" refString="MARSEUL S. A. 1867. - Catalogus Coleopterorum Europae et confinium. L'Abeille 4: 1 - 131." type="journal article" year="1867">Marseul 1867: 54</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E501FFBC5D128" blockId="8.[677,1084,1089,1113]" box="[677,1084,1089,1113]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E501FFC94D128" box="[677,877,1089,1113]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="239D6B3BDC17FFD2BE8E501FFC94D128" box="[677,877,1089,1113]" italics="true" pageId="8" pageNumber="637">Geotrupes laevipennis</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BFAC501FFBC1D128" author="MARSEUL S. A." box="[903,1080,1089,1113]" pageId="8" pageNumber="637" pagination="1 - 131" refId="ref11387" refString="MARSEUL S. A. 1867. - Catalogus Coleopterorum Europae et confinium. L'Abeille 4: 1 - 131." type="journal article" year="1867">Marseul 1867: 54</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E502EFBB8D1D3" blockId="8.[677,1210,1136,1186]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E502EFB9AD1F9" authority="Deyrolle, 1869: 10" authorityName="Deyrolle" authorityPageNumber="10" authorityYear="1869" box="[677,1123,1136,1160]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="subSpecies" species="andalusiacus" subSpecies="teste">
<emphasis id="239D6B3BDC17FFD2BE8E502EFC5CD1F9" box="[677,933,1136,1160]" italics="true" pageId="8" pageNumber="637">Geotrupes andalusiacus teste</emphasis>
<bibRefCitation id="7578CAD8DC17FFD2BF80502EFB9AD1F9" author="DEYROLLE E." box="[939,1123,1136,1160]" pageId="8" pageNumber="637" pagination="10" refId="ref10936" refString="DEYROLLE E. 1869. - Petites nouvelles. Petites Nouvelles entomologiques 3: 10." type="journal article" year="1869">Deyrolle, 1869: 10</bibRefCitation>
</taxonomicName>
. —
<bibRefCitation id="7578CAD8DC17FFD2B8A2502EFCB6D1D3" author="BOUCOMONT A." pageId="8" pageNumber="637" refId="ref10730" refString="BOUCOMONT A. 1912. - Scarabaeidae: Taurocerastinae, Geotrupinae. Pars 46. Coleopterorum Catalogus auspiciis et auxilio. W. Junk, Berlin, 47 p." type="book" year="1912">Boucomont 1912: 22</bibRefCitation>
(as synonym of
<taxonomicName id="D6E9CCAADC17FFD2BFDB50D5FBCBD1D3" authorityName="Germar" authorityYear="1823" box="[1008,1074,1163,1186]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC17FFD2BFDB50D5FBCBD1D3" box="[1008,1074,1163,1186]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E50E4FB7BD1A3" blockId="8.[677,1154,1209,1234]" box="[677,1154,1209,1234]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E50E4FC4AD1A0" authorityName="Germar" authorityYear="1823" box="[677,947,1209,1234]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus" subGenus="Typhoeus">
<emphasis id="239D6B3BDC17FFD2BE8E50E4FCF8D1A0" box="[677,769,1210,1233]" italics="true" pageId="8" pageNumber="637">Geotrupes</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF2450E7FC9AD1A0" box="[783,867,1209,1233]" italics="true" pageId="8" pageNumber="637">Typhoeus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BF5A50E4FC4AD1A0" box="[881,947,1210,1233]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BFE650E4FB87D1A3" author="REITTER E." box="[973,1150,1210,1234]" pageId="8" pageNumber="637" pagination="177 - 194" refId="ref11608" refString="REITTER E. 1891. - [Scarabaeidae], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. R. Friedlander &amp; Sohn, Berlin; E. Reitter, Modling; Revue d'Entomologie, Caen: 177 - 194." type="book chapter" year="1891">Reitter 1891: 184</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E50B7FB5ED070" blockId="8.[677,1191,1257,1281]" box="[677,1191,1257,1281]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E50B7FC21D070" box="[677,984,1257,1281]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="laevipennis" subGenus="Typhoeus">
<emphasis id="239D6B3BDC17FFD2BE8E50B7FCF8D071" box="[677,769,1257,1280]" italics="true" pageId="8" pageNumber="637">Geotrupes</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF2450B7FC9AD070" box="[783,867,1257,1281]" italics="true" pageId="8" pageNumber="637">Typhoeus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BF5A50B7FC21D070" box="[881,984,1257,1281]" italics="true" pageId="8" pageNumber="637">laevipennis</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BFD950B7FB5AD070" author="REITTER E." box="[1010,1187,1257,1281]" pageId="8" pageNumber="637" pagination="177 - 194" refId="ref11608" refString="REITTER E. 1891. - [Scarabaeidae], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. R. Friedlander &amp; Sohn, Berlin; E. Reitter, Modling; Revue d'Entomologie, Caen: 177 - 194." type="book chapter" year="1891">Reitter 1891: 184</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E5146FB60D041" blockId="8.[677,1177,1304,1328]" box="[677,1177,1304,1328]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E5146FC1DD05E" box="[677,996,1304,1328]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus" subGenus="Minotaurus">
<emphasis id="239D6B3BDC17FFD2BE8E5146FCEFD041" box="[677,790,1304,1328]" italics="true" pageId="8" pageNumber="637">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF0F5146FC6AD05E" box="[804,915,1304,1327]" italics="true" pageId="8" pageNumber="637">Minotaurus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BF895146FC1DD05E" box="[930,996,1304,1327]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BFD55146FB6CD041" author="REITTER E." box="[1022,1173,1304,1328]" pageId="8" pageNumber="637" pagination="1 - 109" refId="ref11666" refString="REITTER E. 1893. - Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des naturforschenden Vereines in Brunn 31: 1 - 109 (dated 1892, published 1893)." type="journal article" year="1893">Reitter 1893: 9</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E5119FD0CD008" blockId="8.[675,1210,1351,1401]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E5119FB9DD02E" box="[677,1124,1351,1375]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="variety" species="momus" subGenus="Minotaurus" variety="laevipennis">
<emphasis id="239D6B3BDC17FFD2BE8E5119FCEAD02E" box="[677,787,1351,1375]" italics="true" pageId="8" pageNumber="637">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF355116FC70D02E" box="[798,905,1352,1375]" italics="true" pageId="8" pageNumber="637">Minotaurus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BFBD5116FC2ED02E" box="[918,983,1352,1375]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
var.
<emphasis id="239D6B3BDC17FFD2B82B5119FB9DD02E" box="[1024,1124,1351,1375]" italics="true" pageId="8" pageNumber="637">laevipennis</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2B8525119FD08D008" author="REITTER E." pageId="8" pageNumber="637" pagination="1 - 109" refId="ref11666" refString="REITTER E. 1893. - Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des naturforschenden Vereines in Brunn 31: 1 - 109 (dated 1892, published 1893)." type="journal article" year="1893">Reitter 1893: 9</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E51CEFB61D0D8" blockId="8.[677,1176,1424,1449]" box="[677,1176,1424,1449]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E51CEFC30D0D9" box="[677,969,1424,1449]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="species" species="momus" subGenus="Typhoeus">
<emphasis id="239D6B3BDC17FFD2BE8E51CEFCEFD0D9" box="[677,790,1424,1448]" italics="true" pageId="8" pageNumber="637">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF0F51CEFC81D0D9" box="[804,888,1424,1448]" italics="true" pageId="8" pageNumber="637">Typhoeus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BFAC51CFFC30D0D9" box="[903,969,1425,1448]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BFC851CFFB6DD0D8" author="REITTER E." box="[995,1172,1425,1449]" pageId="8" pageNumber="637" pagination="714 - 750" refId="ref11701" refString="REITTER E. 1906. - [Lamellicornia], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. 2 nd ed. R. Friedlander &amp; Sohn, Berlin; Edmund Reitter, Paskau; Revue d'Entomologie, Caen: 714 - 750." type="book chapter" year="1906">Reitter 1906: 725</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E519EFCE8D083" blockId="8.[677,1210,1472,1522]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E519EFBA7D0A9" box="[677,1118,1472,1496]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="variety" species="momus" subGenus="Typhoeus" variety="laevipennis">
<emphasis id="239D6B3BDC17FFD2BE8E519EFCEFD0A9" box="[677,790,1472,1496]" italics="true" pageId="8" pageNumber="637">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF0F519EFC81D0A9" box="[804,888,1472,1496]" italics="true" pageId="8" pageNumber="637">Typhoeus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BFAD519EFC31D0A6" box="[902,968,1472,1495]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
var.
<emphasis id="239D6B3BDC17FFD2BFDC519EFBA7D0A9" box="[1015,1118,1472,1496]" italics="true" pageId="8" pageNumber="637">laevipennis</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2B85C519EFCF4D083" author="REITTER E." pageId="8" pageNumber="637" pagination="714 - 750" refId="ref11701" refString="REITTER E. 1906. - [Lamellicornia], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. 2 nd ed. R. Friedlander &amp; Sohn, Berlin; Edmund Reitter, Paskau; Revue d'Entomologie, Caen: 714 - 750." type="book chapter" year="1906">Reitter 1906: 725</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD2BE8E5257FCE8D34D" blockId="8.[677,1210,1545,1596]" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E5257FBA2D350" authorityName="Reitter" authorityYear="1892" box="[677,1115,1545,1569]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="637" phylum="Arthropoda" rank="variety" species="momus" subGenus="Typhoeus" variety="momoides">
<emphasis id="239D6B3BDC17FFD2BE8E5257FCEFD350" box="[677,790,1545,1569]" italics="true" pageId="8" pageNumber="637">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF0D5257FC83D350" box="[806,890,1545,1569]" italics="true" pageId="8" pageNumber="637">Typhoeus</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BFA15254FC35D350" box="[906,972,1546,1569]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
var.
<emphasis id="239D6B3BDC17FFD2BFD45257FBA2D350" box="[1023,1115,1545,1569]" italics="true" pageId="8" pageNumber="637">momoides</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2B85C5257FCF4D34D" author="REITTER E." pageId="8" pageNumber="637" pagination="714 - 750" refId="ref11701" refString="REITTER E. 1906. - [Lamellicornia], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. 2 nd ed. R. Friedlander &amp; Sohn, Berlin; Edmund Reitter, Paskau; Revue d'Entomologie, Caen: 714 - 750." type="book chapter" year="1906">Reitter 1906: 725</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC17FFD3BE8E520DFE2AD103" blockId="8.[677,1212,1619,1696]" lastBlockId="9.[115,467,1114,1138]" lastPageId="9" lastPageNumber="638" pageId="8" pageNumber="637">
<taxonomicName id="D6E9CCAADC17FFD2BE8E520DFC22D31A" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[677,987,1619,1643]" class="Clitellata" family="Octochaetidae" genus="Typhoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="637" phylum="Annelida" rank="species" species="momus" subGenus="Chelotrupes">
<emphasis id="239D6B3BDC17FFD2BE8E520DFD07D31A" box="[677,766,1619,1643]" italics="true" pageId="8" pageNumber="637">Typhoeus</emphasis>
(
<emphasis id="239D6B3BDC17FFD2BF3B520DFC7DD31A" box="[784,900,1619,1643]" italics="true" pageId="8" pageNumber="637">Chelotrupes</emphasis>
)
<emphasis id="239D6B3BDC17FFD2BFBE520AFC22D31A" box="[917,987,1620,1643]" italics="true" pageId="8" pageNumber="637">momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC17FFD2BFD1520DFD3AD3F7" author="BOUCOMONT A." pageId="8" pageNumber="637" refId="ref10730" refString="BOUCOMONT A. 1912. - Scarabaeidae: Taurocerastinae, Geotrupinae. Pars 46. Coleopterorum Catalogus auspiciis et auxilio. W. Junk, Berlin, 47 p." type="book" year="1912">Boucomont 1912: 22</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC17FFD2BEC15230FC4FD3F7" author="WINKLER A." box="[746,950,1646,1670]" pageId="8" pageNumber="637" pagination="1009 - 1136" refId="ref11824" refString="WINKLER A. 1929. - Catalogus Coleopterorum regionis palaearcticae. Scarabaeidae. Albert Winkler, Wien, 9: 1009 - 1136." type="journal article" year="1929">Winkler 1929: 1036</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC17FFD2BFF65230FD37D3D1" author="BAGUENA CORELLA L." pageId="8" pageNumber="637" refId="ref10502" refString="BAGUENA CORELLA L. 1967. - Scarabaeoidea de la fauna ibero-balear y pirenaica. Consejo Superior de Investigaciones Cientificas, Instituto Espanol de Entomologia, Madrid, 576 p." type="book" year="1967">Báguena Corella 1967: 260</bibRefCitation>
. — Martín-Piera &amp; López-Colón 2000: 122.
<taxonomicName id="D6E9CCAADC16FFD3BC585004FEE9D103" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[115,272,1114,1138]" class="Clitellata" family="Octochaetidae" genus="Typhoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="638" phylum="Annelida" rank="species" species="momus">
<emphasis id="239D6B3BDC16FFD3BC585004FEE9D103" box="[115,272,1114,1138]" italics="true" pageId="9" pageNumber="638">Typhoeus momus</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC16FFD3BD035004FE36D103" author="BARAUD J." box="[296,463,1114,1138]" pageId="9" pageNumber="638" refId="ref10533" refString="BARAUD J. 1992. - Coleopteres Scarabaeoidea d'Europe. Faune de France, France et regions limitrophes, 78. Federation francaise des Societes de Sciences naturelles, Paris; Societe linneenne de Lyon, Lyon, 856 p." type="book" year="1992">Baraud 1992: 49</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="4596E7A1DC16FFD3BC5A57B5FDFAD158" ID-DOI="http://doi.org/10.5281/zenodo.5402563" ID-Zenodo-Dep="5402563" httpUri="https://zenodo.org/record/5402563/files/figure.png" pageId="9" pageNumber="638" startId="9.[113,124,1003,1020]" targetBox="[135,643,227,952]" targetPageId="9">
<paragraph id="1156B729DC16FFD3BC5A57B5FDFAD158" blockId="9.[113,646,1002,1065]" pageId="9" pageNumber="638">
FIG. 3. — Distribution in Sardinia (Italy) of
<taxonomicName id="D6E9CCAADC16FFD3BDF257B4FD7FD68A" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[473,646,1002,1019]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="638" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC16FFD3BDF257B4FD7FD68A" box="[473,646,1002,1019]" italics="true" pageId="9" pageNumber="638">Chelotrupes hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC16FFD3BC58505CFF3BD162" box="[115,194,1026,1043]" pageId="9" pageNumber="638" rank="species">n. comb.</taxonomicNameLabel>
(■) and
<taxonomicName id="D6E9CCAADC16FFD3BD24505FFE78D163" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Baudi" baseAuthorityYear="1870" box="[271,385,1025,1042]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="638" phylum="Arthropoda" rank="species" species="matutinalis" status="comb. nov.">
<emphasis id="239D6B3BDC16FFD3BD24505FFE78D163" box="[271,385,1025,1042]" italics="true" pageId="9" pageNumber="638">C matutinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC16FFD3BDAC505CFE2ED162" box="[391,471,1026,1043]" pageId="9" pageNumber="638" rank="species">n. comb.</taxonomicNameLabel>
(▲). ○, sympatry of
<taxonomicName id="D6E9CCAADC16FFD3BC585046FF32D158" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[115,203,1048,1065]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="638" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC16FFD3BC585046FF32D158" box="[115,203,1048,1065]" italics="true" pageId="9" pageNumber="638">C. hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC16FFD3BCE55046FEE0D158" box="[206,281,1048,1065]" pageId="9" pageNumber="638" rank="species">n. comb.</taxonomicNameLabel>
and
<taxonomicName id="D6E9CCAADC16FFD3BD695046FE4BD158" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Baudi" baseAuthorityYear="1870" box="[322,434,1048,1065]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="638" phylum="Arthropoda" rank="species" species="matutinalis" status="comb. nov.">
<emphasis id="239D6B3BDC16FFD3BD695046FE4BD158" box="[322,434,1048,1065]" italics="true" pageId="9" pageNumber="638">C. matutinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC16FFD3BD935046FDFAD158" box="[440,515,1048,1065]" pageId="9" pageNumber="638" rank="species">n. comb.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="59F3E4A2DC16FFD3BC5A50D4FE50D084" pageId="9" pageNumber="638" type="materials_examined">
<paragraph id="1156B729DC16FFD3BC5A50D4FDF9D07D" blockId="9.[112,647,1162,1318]" pageId="9" pageNumber="638">
<collectionCode id="77F82FECDC16FFD3BC5A50D4FF5AD1D1" box="[113,163,1162,1185]" pageId="9" pageNumber="638">TYPE</collectionCode>
MATERIAL. —
<materialsCitation id="A181BD74DC16FFD3BD1250D4FDF9D07D" collectionCode="MNHN" determinerName="Des. Dellacasa" pageId="9" pageNumber="638" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="CE52098BDC16FFD3BD1250D4FE62D1D3" box="[313,411,1162,1186]" pageId="9" pageNumber="638" type="lectotype">Lectotype</typeStatus>
(here designated):
<specimenCount id="07EF7CA0DC16FFD3BE7650D4FD7FD1D0" box="[605,646,1162,1186]" pageId="9" pageNumber="638" type="male">1 ♂</specimenCount>
(
<collectionCode id="77F82FECDC16FFD3BC5350FAFF33D1CD" box="[120,202,1188,1212]" pageId="9" pageNumber="638">MNHN</collectionCode>
), bearing the following labels: 1) blue, handwritten: “
<taxonomicName id="D6E9CCAADC16FFD3BCE750E1FE78D1A6" authorityName="Germar" authorityYear="1823" box="[204,385,1215,1239]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="638" phylum="Arthropoda" rank="species" species="momus">Geotrupes momus</taxonomicName>
/ C. Olivier / Afr. aequin.”; 2) red, printed: “
<taxonomicName id="D6E9CCAADC16FFD3BD0F5084FE2BD180" authorityName="Germar" authorityYear="1823" box="[292,466,1242,1265]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="638" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC16FFD3BD0F5084FE2BD180" box="[292,466,1242,1265]" italics="true" pageId="9" pageNumber="638">Geotrupes momus</emphasis>
</taxonomicName>
/
<bibRefCitation id="7578CAD8DC16FFD3BDC05087FD8CD180" author="OLIVIER A. G." box="[491,629,1241,1265]" pageId="9" pageNumber="638" refId="ref11485" refString="OLIVIER A. G. 1789. - Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Baudouin, Paris, 1, 190 p." type="book" year="1789">Olivier, 1789</bibRefCitation>
/ LECTOTYPUS /
<determinerName id="E48A5BEDDC16FFD3BD0150AAFE4FD07D" box="[298,438,1268,1292]" pageId="9" pageNumber="638">Des. Dellacasa</determinerName>
2005”.
</materialsCitation>
</paragraph>
<paragraph id="1156B729DC16FFD3BC5A5150FD82D057" blockId="9.[112,647,1162,1318]" box="[113,635,1294,1318]" pageId="9" pageNumber="638">
<materialsCitation id="A181BD74DC16FFD3BC5A5150FD82D057" box="[113,635,1294,1318]" collectionCode="MNHN" determinerName="Des. Dellacasa" pageId="9" pageNumber="638" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="CE52098BDC16FFD3BC5A5150FF0ED057" box="[113,247,1294,1318]" pageId="9" pageNumber="638" type="paralectotype">Paralectotype</typeStatus>
: same data as lectotype,
<specimenCount id="07EF7CA0DC16FFD3BDCF5150FDF1D054" box="[484,520,1294,1318]" pageId="9" pageNumber="638" type="female">1 ♀</specimenCount>
(
<collectionCode id="77F82FECDC16FFD3BE3D5150FD90D057" box="[534,617,1294,1318]" pageId="9" pageNumber="638">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1156B729DC16FFD3BC5A5160FE50D084" blockId="9.[112,648,1342,1525]" pageId="9" pageNumber="638">
<typeStatus id="CE52098BDC16FFD3BC5A5160FF5BD022" box="[113,162,1342,1365]" pageId="9" pageNumber="638">
<collectionCode id="77F82FECDC16FFD3BC5A5160FF5BD022" box="[113,162,1342,1365]" pageId="9" pageNumber="638">TYPE</collectionCode>
</typeStatus>
LOCALITY. — “Afr. aequin.” [Africa aequinoctialis;
<collectingCountry id="69FEF7B9DC16FFD3BC5B5106FF0FD001" box="[112,246,1368,1392]" name="Sierra Leone" pageId="9" pageNumber="638">Sierra Leone</collectingCountry>
?].
<bibRefCitation id="7578CAD8DC16FFD3BD3A5106FE5FD001" author="OLIVIER A. G." box="[273,422,1368,1392]" pageId="9" pageNumber="638" refId="ref11485" refString="OLIVIER A. G. 1789. - Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Baudouin, Paris, 1, 190 p." type="book" year="1789">Olivier (1789)</bibRefCitation>
and
<bibRefCitation id="7578CAD8DC16FFD3BDF75106FD7DD001" author="FABRICIUS J. C." box="[476,644,1368,1392]" pageId="9" pageNumber="638" refId="ref10952" refString="FABRICIUS J. C. 1792. - Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Hafniae, 538 p." type="book" year="1792">Fabricius (1792)</bibRefCitation>
recorded as “patria typica” of this taxon respectively “
<collectingCountry id="69FEF7B9DC16FFD3BC5751D3FF03D0D4" box="[124,250,1421,1445]" name="Sierra Leone" pageId="9" pageNumber="638">Sierra Leone</collectingCountry>
” and “
<collectingCountry id="69FEF7B9DC16FFD3BD1551D3FE8AD0D4" box="[318,371,1421,1445]" name="India" pageId="9" pageNumber="638">India</collectingCountry>
orientali”, and
<bibRefCitation id="7578CAD8DC16FFD3BE2451D3FD71D0D4" author="JEKEL H." box="[527,648,1421,1445]" pageId="9" pageNumber="638" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">Jekel (1866)</bibRefCitation>
“le
<collectingCountry id="69FEF7B9DC16FFD3BCBF51F6FF1FD0B1" box="[148,230,1448,1472]" name="Portugal" pageId="9" pageNumber="638">Portugal</collectingCountry>
(Algarves), l’Andalousie et la Barbarie occidentale”. Conversely the species is known only from southwestern Iberian Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC16FFD3BC5A5253FE15D34E" pageId="9" pageNumber="638" type="etymology">
<paragraph id="1156B729DC16FFD3BC5A5253FE15D34E" blockId="9.[113,647,1548,1599]" pageId="9" pageNumber="638">ETYMOLOGY. — Momus is the Greek deity of mockery, faultfinding, scoff and unfair criticism.</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC16FFD3BC5A5209FC34D69F" pageId="9" pageNumber="638" type="materials_examined">
<paragraph id="1156B729DC16FFD3BC5A5209FB6FD75D" blockId="9.[111,647,1622,1699]" lastBlockId="9.[674,1211,215,1006]" pageId="9" pageNumber="638">
MATERIAL EXAMINED. —
<emphasis id="239D6B3BDC16FFD3BD7C5208FE6BD31F" bold="true" box="[343,402,1622,1646]" pageId="9" pageNumber="638">
<collectingCountry id="69FEF7B9DC16FFD3BD7C5208FE77D31F" box="[343,398,1622,1646]" name="Spain" pageId="9" pageNumber="638">Spain</collectingCountry>
.
</emphasis>
Algeciras, leg. A Grouvelle,
<specimenCount id="07EF7CA0DC16FFD3BC44522FFF64D3F8" box="[111,157,1649,1673]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3BD1A522FFE98D3F8" box="[305,353,1649,1673]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3BD1A522FFE98D3F8" box="[305,353,1649,1673]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC16FFD3BD47522FFE28D3F8" box="[364,465,1649,1673]" pageId="9" pageNumber="638" value="1906-04-06">6.IV.1906</date>
, ex coll. Schramm,
<specimenCount id="07EF7CA0DC16FFD3BC5A52D5FF66D3D2" box="[113,159,1675,1699]" pageId="9" pageNumber="638" type="generic">2 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3BD1A52D5FE98D3D2" box="[305,353,1675,1699]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3BD1A52D5FE98D3D2" box="[305,353,1675,1699]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC16FFD3BD4752D5FE28D3D2" box="[364,465,1675,1699]" pageId="9" pageNumber="638" value="1905-04-07">7.IV.1905</date>
, ex coll. Schramm,
<specimenCount id="07EF7CA0DC16FFD3BE8E5489FD36D59E" box="[677,719,215,239]" pageId="9" pageNumber="638" type="generic">6 ex.</specimenCount>
(MNHN). — Cadiz,
<specimenCount id="07EF7CA0DC16FFD3BFB35489FC3AD59E" box="[920,963,215,239]" pageId="9" pageNumber="638" type="generic">3 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3B8675489FB80D59E" box="[1100,1145,215,239]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3B8675489FB80D59E" box="[1100,1145,215,239]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
</taxonomicName>
, 1842, leg. Gilliani,
<specimenCount id="07EF7CA0DC16FFD3BF3254AFFCBCD478" box="[793,837,241,265]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(MNHN). — Cadiz, Puerto Galiz,
<date id="655791E9DC16FFD3B8BA54AFFD0DD455" pageId="9" pageNumber="638" value="1984-04-03" valueMax="1984-04-09" valueMin="1984-04-03">3-9. IV.1984</date>
, leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3BFAA5552FC40D455" box="[897,953,268,292]" pageId="9" pageNumber="638" type="generic">11 ex.</specimenCount>
(MNHN). — Cadiz, Rota,
<date id="655791E9DC16FFD3BE885578FCEBD44F" box="[675,786,294,318]" pageId="9" pageNumber="638" value="1973-11-29">29.XI.1973</date>
, leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3BFB55578FC33D44F" box="[926,970,294,318]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(MNHN). —
<collectingRegion id="D32D79CBDC16FFD3B87C5578FB4FD44F" box="[1111,1206,294,318]" country="Spain" name="Andalucia" pageId="9" pageNumber="638">Andalusia</collectingRegion>
, El Rocio, Huelva,
<date id="655791E9DC16FFD3BF71551FFC35D428" box="[858,972,321,345]" pageId="9" pageNumber="638" value="1986-04-14">14.IV.1986</date>
, leg. G. &amp; E. Dellacasa,
<specimenCount id="07EF7CA0DC16FFD3BE8E5505FD2ED402" box="[677,727,347,371]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(coll. MD). —
<collectingRegion id="D32D79CBDC16FFD3BF565505FC1BD402" box="[893,994,347,371]" country="Spain" name="Andalucia" pageId="9" pageNumber="638">Andalusia</collectingRegion>
, Facinas,
<date id="655791E9DC16FFD3B86F5505FB4FD402" box="[1092,1206,347,371]" pageId="9" pageNumber="638" value="1984-12-04">4.XII.1984</date>
, leg. G. &amp; M. Dellacasa,
<specimenCount id="07EF7CA0DC16FFD3BFB35528FC2DD4FF" box="[920,980,374,398]" pageId="9" pageNumber="638" type="generic">41 ex.</specimenCount>
(coll. MD). — Lag. de la Janda,
<date id="655791E9DC16FFD3BED655CEFCB6D4D9" box="[765,847,400,424]" pageId="9" pageNumber="638" value="1918-04">IV.1918</date>
, leg. Brevil,
<specimenCount id="07EF7CA0DC16FFD3BFED55CEFC0DD4D9" box="[966,1012,400,424]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(MNHN). — Prov. Sevilla, Villamanrique,
<date id="655791E9DC16FFD3BF5155F5FC0CD4B2" box="[890,1013,427,451]" pageId="9" pageNumber="638" value="1960-10-01" valueMax="1960-10-11" valueMin="1960-10-01">1-11.X.1960</date>
, leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3B8A955F5FB43D4B2" box="[1154,1210,427,451]" pageId="9" pageNumber="638" type="generic">27 ex.</specimenCount>
(MNHN). —
<collectingRegion id="D32D79CBDC16FFD3BF1A559BFC6AD4AC" box="[817,915,453,477]" country="Spain" name="Andalucia" pageId="9" pageNumber="638">Andalusia</collectingRegion>
, Tarifa, Cadiz,
<date id="655791E9DC16FFD3B80A559BFB76D4AC" box="[1057,1167,453,477]" pageId="9" pageNumber="638" value="1984-12-03">3.XII.1984</date>
, leg. G. &amp; M. Dellacasa,
<specimenCount id="07EF7CA0DC16FFD3BF7255BEFC7DD489" box="[857,900,480,504]" pageId="9" pageNumber="638" type="generic">2 ex.</specimenCount>
(coll. MD). —
<taxonomicName id="D6E9CCAADC16FFD3B83F5581FBBBD486" box="[1044,1090,479,503]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3B83F5581FBBBD486" box="[1044,1090,479,503]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC16FFD3B86055BEFB4FD489" box="[1099,1206,480,504]" pageId="9" pageNumber="638" value="1984-12-07">7.XII.1984</date>
, leg. G. &amp; M. Dellacasa,
<specimenCount id="07EF7CA0DC16FFD3BFB255A4FC2FD763" box="[921,982,506,530]" pageId="9" pageNumber="638" type="generic">20 ex.</specimenCount>
(coll. MD). — Cadiz,
<date id="655791E9DC16FFD3BE8E564AFD01D75D" box="[677,760,532,556]" pageId="9" pageNumber="638" value="1972-11">XI.1972</date>
, leg. A. J. Villarreal,
<specimenCount id="07EF7CA0DC16FFD3BFE8564AFC0BD75D" box="[963,1010,532,556]" pageId="9" pageNumber="638" type="generic">5 ex.</specimenCount>
(MCSNG -PC).
</paragraph>
<paragraph id="1156B729DC16FFD3BE8E5671FC34D69F" blockId="9.[674,1211,215,1006]" pageId="9" pageNumber="638">
<emphasis id="239D6B3BDC16FFD3BE8E5671FD06D736" bold="true" box="[677,767,559,583]" pageId="9" pageNumber="638">
<collectingCountry id="69FEF7B9DC16FFD3BE8E5671FD02D736" box="[677,763,559,583]" name="Portugal" pageId="9" pageNumber="638">Portugal</collectingCountry>
.
</emphasis>
Algarve, env. de
<collectingRegion id="D32D79CBDC16FFD3BFB75671FC30D736" box="[924,969,559,583]" country="Portugal" name="Faro" pageId="9" pageNumber="638">Faro</collectingRegion>
,
<date id="655791E9DC16FFD3BFFA5671FBE5D736" box="[977,1052,559,583]" pageId="9" pageNumber="638" value="1970-10">X.1970</date>
, leg. H. Coiffait,
<specimenCount id="07EF7CA0DC16FFD3BE8E5617FD21D710" box="[677,728,585,609]" pageId="9" pageNumber="638" type="generic">2 ex.</specimenCount>
(MNHN). — Algarve, Moncique, ex Musaeo Harold,
<specimenCount id="07EF7CA0DC16FFD3BED2563AFCD1D70D" box="[761,808,612,636]" pageId="9" pageNumber="638" type="generic">4 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3BF97563AFC15D70D" box="[956,1004,612,636]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3BF97563AFC15D70D" box="[956,1004,612,636]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
</taxonomicName>
, ex. coll. Bedel,
<specimenCount id="07EF7CA0DC16FFD3B8A0563AFB43D70D" box="[1163,1210,612,636]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3BF1C5620FC9ED7E7" box="[823,871,638,662]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3BF1C5620FC9ED7E7" box="[823,871,638,662]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC16FFD3BF5F5620FC20D7E7" box="[884,985,638,662]" pageId="9" pageNumber="638" value="1967-04-02">2.IV.1967</date>
, leg. M. Coiffait,
<specimenCount id="07EF7CA0DC16FFD3B8A15620FB43D7E7" box="[1162,1210,638,662]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3BF6B56C7FC49D7C0" authority=", Foia" authorityName="Foia" box="[832,944,665,689]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3BF6B56C7FC8BD7C0" box="[832,882,665,689]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
, Foia
</taxonomicName>
,
<quantity id="D6111ACCDC16FFD3BF9656C7FBFDD7C0" box="[957,1028,665,689]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="638" unit="m" value="900.0">900 m</quantity>
,
<date id="655791E9DC16FFD3B83A56C7FB71D7C0" box="[1041,1160,665,689]" pageId="9" pageNumber="638" value="1982-04-20">20.IV.1982</date>
; leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3BF3E56EDFCBED7BA" box="[789,839,691,715]" pageId="9" pageNumber="638" type="generic">5 ex.</specimenCount>
(MNHN). — Algarve, Sta. Barbara de N.,
<date id="655791E9DC16FFD3BEC55690FCB9D797" box="[750,832,718,742]" pageId="9" pageNumber="638" value="1963-04">IV.1963</date>
, leg. M. Coiffait,
<specimenCount id="07EF7CA0DC16FFD3BFDF5690FBDCD797" box="[1012,1061,718,742]" pageId="9" pageNumber="638" type="generic">8 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3BE8D56B6FCBBD671" authority=", XII. 1963" authorityName="XII." authorityYear="1963" box="[678,834,744,768]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3BE8D56B6FD2CD671" box="[678,725,744,768]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
,
<date id="655791E9DC16FFD3BECF56B6FCBBD671" box="[740,834,744,768]" pageId="9" pageNumber="638" value="1963-12">XII.1963</date>
</taxonomicName>
,
<specimenCount id="07EF7CA0DC16FFD3BF6556B6FC79D671" box="[846,896,744,768]" pageId="9" pageNumber="638" type="generic">2 ex.</specimenCount>
(MNHN). — Algarve, Tavira,
<date id="655791E9DC16FFD3BE8E575DFCC3D66A" box="[677,826,771,795]" pageId="9" pageNumber="638" value="1965-02-14" valueMax="1965-02-25" valueMin="1965-02-14">14-25.II.1965</date>
, leg. T. Branco (excr. humain),
<specimenCount id="07EF7CA0DC16FFD3B8A3575DFB42D66A" box="[1160,1211,771,795]" pageId="9" pageNumber="638" type="generic">5 ex.</specimenCount>
(MNHN). — Ribatejo,
<quantity id="D6111ACCDC16FFD3BFAF5743FC46D644" box="[900,959,797,821]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="638" unit="km" value="16.0">16 km</quantity>
Sud Coruche,
<date id="655791E9DC16FFD3B8685743FB4FD644" box="[1091,1206,797,821]" pageId="9" pageNumber="638" value="1984-04-03" valueMax="1984-04-09" valueMin="1984-04-03">3-9.IV.1984</date>
, leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3BF1B5766FCA7D621" box="[816,862,824,848]" pageId="9" pageNumber="638" type="generic">9 ex.</specimenCount>
(MNHN). — Ribatejo,
<quantity id="D6111ACCDC16FFD3B8795766FB7DD621" box="[1106,1156,824,848]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="638" unit="km" value="6.0">6 km</quantity>
N de Lavre,
<date id="655791E9DC16FFD3BECC570CFCA5D61B" box="[743,860,850,874]" pageId="9" pageNumber="638" value="1984-03-29">29.III.1984</date>
, leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3BFD3570CFBD1D61B" box="[1016,1064,850,874]" pageId="9" pageNumber="638" type="generic">2 ex.</specimenCount>
(MNHN). — Ribatejo, Monte da Barca,
<date id="655791E9DC16FFD3BF935732FBE7D6F5" box="[952,1054,876,900]" pageId="9" pageNumber="638" value="1984-04-02">2.IV.1984</date>
, leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3BE8D57D9FD2AD6EE" box="[678,723,903,927]" pageId="9" pageNumber="638" type="generic">2 ex.</specimenCount>
(MNHN). — Ribatejo, Muge,
<date id="655791E9DC16FFD3BFD657D9FB76D6EE" box="[1021,1167,903,927]" pageId="9" pageNumber="638" value="1984-03-27" valueMax="1984-03-28" valueMin="1984-03-27">27-28.III.1984</date>
, leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3BF2757FFFCC2D6C8" box="[780,827,929,953]" pageId="9" pageNumber="638" type="generic">3 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3BFFE57FFFBFCD6C8" box="[981,1029,929,953]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3BFFE57FFFBFCD6C8" box="[981,1029,929,953]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC16FFD3B83957FFFB8FD6C8" box="[1042,1142,929,953]" pageId="9" pageNumber="638" value="1981-04-06">6.IV.1981</date>
, leg. J. Baraud,
<specimenCount id="07EF7CA0DC16FFD3BEDE57E2FCDBD6A5" box="[757,802,956,980]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC16FFD3BF9957E2FC18D6A5" box="[946,993,956,980]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="638" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC16FFD3BF9957E2FC18D6A5" box="[946,993,956,980]" italics="true" pageId="9" pageNumber="638">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC16FFD3BFC657E2FBA8D6A5" box="[1005,1105,956,980]" pageId="9" pageNumber="638" value="1986-04-07">7.IV.1986</date>
, Leg. G. &amp; E. Dellacasa,
<specimenCount id="07EF7CA0DC16FFD3BF0D5788FCACD69F" box="[806,853,982,1006]" pageId="9" pageNumber="638" type="generic">1 ex.</specimenCount>
(coll. MD).
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC16FFD3BE8E5058FC53D0FA" pageId="9" pageNumber="638" type="distribution">
<paragraph id="1156B729DC16FFD3BE8E5058FC15D1CC" blockId="9.[676,1211,1030,1213]" pageId="9" pageNumber="638">
DISTRIBUTION. — Known from southern
<collectingCountry id="69FEF7B9DC16FFD3B87F5058FB75D16F" box="[1108,1164,1030,1054]" name="Spain" pageId="9" pageNumber="638">Spain</collectingCountry>
and
<collectingCountry id="69FEF7B9DC16FFD3BE8D507EFD02D149" box="[678,763,1056,1080]" name="Portugal" pageId="9" pageNumber="638">Portugal</collectingCountry>
(Ribatejo and Algarve). We have examined specimens preserved in the Paris Museum coming from localities that need to be confirmed:
<collectingCountry id="69FEF7B9DC16FFD3B80B500BFB9DD11C" box="[1056,1124,1109,1133]" name="Morocco" pageId="9" pageNumber="638">Maroc</collectingCountry>
, ex coll. Boucomont,
<specimenCount id="07EF7CA0DC16FFD3BF0D502EFCAFD1F9" box="[806,854,1136,1160]" pageId="9" pageNumber="638" type="generic">3 ex.</specimenCount>
;
<collectingCountry id="69FEF7B9DC16FFD3BF74502EFC5FD1F9" box="[863,934,1136,1160]" name="Algeria" pageId="9" pageNumber="638">Algérie</collectingCountry>
, ex coll. Ernest André, 1914,
<specimenCount id="07EF7CA0DC16FFD3BE8E50D4FD2CD1D3" box="[677,725,1162,1186]" pageId="9" pageNumber="638" type="generic">2 ex.</specimenCount>
The record from Ibiza, Balearic Islands (
<bibRefCitation id="7578CAD8DC16FFD3B84C50D4FD20D1CC" author="CROVETTI A." pageId="9" pageNumber="638" pagination="229 - 250" refId="ref10845" refString="CROVETTI A. 1970. - Contributi alla conoscenza dei Coleotteri Scarabeidi. IV. Revisione sistematica dei Typhoeus Leach del sottogenere Chelotrupes Jekel. &quot; Studi Sassaresi &quot; - Sez. III - Annali della Facolta di Agraria dell'Universita di Sassari 18 (1): 229 - 250." type="journal article" year="1970">Crovetti 1970</bibRefCitation>
) needs to be confirmed too.
</paragraph>
<paragraph id="1156B729DC16FFD3BE8E508BFC53D0FA" blockId="9.[675,1211,1236,1419]" pageId="9" pageNumber="638">
BIONOMICS. — Flightless telocoprid species utilizing several kinds of dung (sheep, horse, cow), particularly frequent in dung of rabbits (Sanchez-Piñero &amp; Avila 1991), rarely in dung of wild boar (
<bibRefCitation id="7578CAD8DC16FFD3BFD3517AFB4AD04D" author="HIDALGO J. M. &amp; BACH C. &amp; CARDENAS A. M." box="[1016,1203,1315,1340]" pageId="9" pageNumber="638" pagination="9 - 35" refId="ref11059" refString="HIDALGO J. M., BACH C. &amp; CARDENAS A. M. 1998. - I. Los Scarabaeoidea coprofagos de las comarcas naturales de la provincia de Cordoba: I. Aphodiidae. Boletin de la Asociacion espanola de Entomologia 22 (3 - 4): 9 - 35." type="journal article" year="1998">
Hidalgo
<emphasis id="239D6B3BDC16FFD3B866517AFB83D04A" box="[1101,1146,1315,1339]" italics="true" pageId="9" pageNumber="638">et al.</emphasis>
1998
</bibRefCitation>
) and of lynx (Martín-Piera &amp; López-Colón 2000), also in human excrement. Present all year round and distributed from sea level to high hills.
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC16FFD0BE8E51F2FED2D0F6" lastPageId="10" lastPageNumber="639" pageId="9" pageNumber="638" type="description">
<paragraph id="1156B729DC16FFD3BE8E51F2FCCCD0B5" blockId="9.[674,1211,1452,1700]" box="[677,821,1452,1478]" pageId="9" pageNumber="638">DESCRIPTION</paragraph>
<paragraph id="1156B729DC16FFD0BE8E5195FE2DD7DF" blockId="9.[674,1211,1452,1700]" lastBlockId="10.[111,649,216,1415]" lastPageId="10" lastPageNumber="639" pageId="9" pageNumber="638">Length 10.0-21.0 mm, moderately stout, strongly convex, shiny, glabrous. Black; antennae and legs blackish. Head with epistoma narrowly rhomboidal, glabrous in both sexes, with median longitudinal gibbosity moderately raised and more or less coarsely punctate; clypeus subtriangular, rounded at anterior angle, thinly margined, the edge sparsely ciliate and distinctly upturned distally; genal sutures cariniform, strongly raised; genae widely rounded; front smooth on disc, moderately punctate laterally. Pronotum strongly transverse, moderately convex, with distinct antero-median rounded foveola on sides; latter feebly arcuate, thinly margined, the edge sparsely ciliate; hind angles widely rounded; base thinly margined, not sinuate.Scutellum subregularly triangular, almost convex. Elytra distinctly longer than wide, strongly convex, narrowly flattened before the epipleural carina; the latter glabrous; striae almost obsolete with nearly imperceptible punctures; intervals flat, smooth. Fore tibiae apical spur feebly sinuate. First segment of hind tarsi as long as following two combined.</paragraph>
<paragraph id="1156B729DC15FFD0BCA356EDFE26D13B" blockId="10.[111,649,216,1415]" pageId="10" pageNumber="639">
Male: epistoma with median longitudinal gibbosity feeble, finely and very superficially punctate on disc, punctation coarser distally; pronotum superficially punctate on sides toward the anterior angles, anteriorly with three hornlike apophyses; the lateral ones straight, anteriorly projecting from above the anterior angles, subparallel at outer margin and with distinctly raised longitudinal carina subtuberculiform at middle and from this abruptly recumbent toward the inner margin that is sinuately widened inwardly, the groove distinctly rugulose; the central apophysis shortened, regularly acuminate, upturned; aedeagus illustrated on
<figureCitation id="89D2ABACDC15FFD0BD405071FE22D13B" box="[363,475,1071,1098]" captionStart="FIG" captionStartId="8.[113,124,732,749]" captionTargetBox="[204,1069,228,686]" captionTargetId="figure-20@8.[188,1134,199,724]" captionTargetPageId="8" captionText="FIG. 2. — Aedeagus in dorsal view: A, Chelotrupes hiostius (Gené, 1836) n. comb., Costa Verde, Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; B, C. matutinalis (Baudi, 1870) n. comb.,Costa Verde,Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; C, C. momus (Olivier, 1789) n. comb., Andalusia, Cadiz, Tarifa. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5402561" httpUri="https://zenodo.org/record/5402561/files/figure.png" pageId="10" pageNumber="639">Figure 2C</figureCitation>
.
</paragraph>
<paragraph id="1156B729DC15FFD0BCA35011FED2D0F6" blockId="10.[111,649,216,1415]" pageId="10" pageNumber="639">Female: epistoma with median longitudinal gibbosity distinctly tuberculiform at top, strongly, rugosely, confusedly punctate throughout; pronotum sparsely and superficially punctate on disc, punctation irregular and more dense on sides, with antero-lateral angles dentiform, feebly projecting anteriorly and distinctly outwardly curved; at each side foveolate and with an outer tooth sharp and distinctly raised; with short strong antero-median transverse carina.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="F38E4B773C1496B47DC5FE8ADF65FB35">Revision of the genus Chelotrupes Jekel, 1866 n. stat. (Insecta, Coleoptera, Geotrupidae)</mods:title>
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<subSubSection id="59F3E4A2DC1CFFD9BCA55489FD91D582" box="[142,616,215,243]" pageId="3" pageNumber="632" type="nomenclature">
<paragraph id="1156B729DC1CFFD9BCA55489FD91D582" blockId="3.[142,616,215,275]" box="[142,616,215,243]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BCA55489FDF8D582" ID-CoL="TWR6" authority="(Gene, 1836)" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[142,513,215,243]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1CFFD9BCA55489FE9FD583" bold="true" box="[142,358,215,242]" italics="true" pageId="3" pageNumber="632">Chelotrupes hiostius</emphasis>
(
<bibRefCitation id="7578CAD8DC1CFFD9BD5C5486FE00D582" author="GENE J." box="[375,505,216,243]" pageId="3" pageNumber="632" pagination="161 - 191" refId="ref10988" refString="GENE J. 1836. - De quibusdam insectis Sardiniae novis aut minus cognitis. Fasciculus I. Memoralia Regiae Scientiarum Academiae Taurinensis 39: 161 - 191." type="journal article" year="1836">Gené, 1836</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1CFFD9BE235486FD91D582" box="[520,616,216,243]" pageId="3" pageNumber="632" rank="species">n. comb.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1CFFD9BD2F54A6FE0AD462" box="[260,499,248,275]" pageId="3" pageNumber="632" type="description">
<paragraph id="1156B729DC1CFFD9BD2F54A6FE0AD462" blockId="3.[142,616,215,275]" box="[260,499,248,275]" pageId="3" pageNumber="632">
(
<figureCitation id="89D2ABACDC1CFFD9BD2054A6FE46D462" box="[267,447,248,275]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="3" pageNumber="632">Figs 1A, D, G, J</figureCitation>
;
<figureCitation id="89D2ABACDC1CFFD9BDE054A6FE15D462" box="[459,492,248,275]" captionStart="FIG" captionStartId="8.[113,124,732,749]" captionTargetBox="[204,1069,228,686]" captionTargetId="figure-20@8.[188,1134,199,724]" captionTargetPageId="8" captionText="FIG. 2. — Aedeagus in dorsal view: A, Chelotrupes hiostius (Gené, 1836) n. comb., Costa Verde, Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; B, C. matutinalis (Baudi, 1870) n. comb.,Costa Verde,Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; C, C. momus (Olivier, 1789) n. comb., Andalusia, Cadiz, Tarifa. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5402561" httpUri="https://zenodo.org/record/5402561/files/figure.png" pageId="3" pageNumber="632">2A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1CFFD9BC5A556EFE2FD61F" pageId="3" pageNumber="632" type="reference_group">
<paragraph id="1156B729DC1CFFD9BC5A556EFE6AD40D" blockId="3.[113,647,303,380]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A556EFE35D436" authority="Gene, 1836: 187" authorityName="Gene" authorityPageNumber="187" authorityYear="1836" box="[113,460,303,327]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius">
<emphasis id="239D6B3BDC1CFFD9BC5A556EFEE0D436" box="[113,281,303,327]" italics="true" pageId="3" pageNumber="632">Geotrupes hiostius</emphasis>
<bibRefCitation id="7578CAD8DC1CFFD9BD0A5571FE35D436" author="GENE J." box="[289,460,303,327]" pageId="3" pageNumber="632" pagination="161 - 191" refId="ref10988" refString="GENE J. 1836. - De quibusdam insectis Sardiniae novis aut minus cognitis. Fasciculus I. Memoralia Regiae Scientiarum Academiae Taurinensis 39: 161 - 191." type="journal article" year="1836">Gené, 1836: 187</bibRefCitation>
</taxonomicName>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BDDC5571FF74D413" author="MARSEUL S. A." pageId="3" pageNumber="632" pagination="1 - 131" refId="ref11387" refString="MARSEUL S. A. 1867. - Catalogus Coleopterorum Europae et confinium. L'Abeille 4: 1 - 131." type="journal article" year="1867">Marseul 1867: 54</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BC935514FE83D413" author="HAROLD E." box="[184,378,330,354]" pageId="3" pageNumber="632" pagination="979 - 1346" refId="ref11017" refString="HAROLD E. 1869. - Scarabaeidae, in GEMMINGER M. &amp; HAROLD E. (eds), Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Grummi E. H., Monachia, 4: 979 - 1346." type="journal article" year="1869">Harold 1869: 1082</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BD8F5514FD9AD413" author="BARGAGLI P." box="[420,611,330,354]" pageId="3" pageNumber="632" pagination="279 - 290" refId="ref10574" refString="BARGAGLI P. 1872. - Materiali per la fauna entomologica dell'isola di Sardegna. Coleotteri (Continuazione). Bullettino della Societa entomologica italiana 4: 279 - 290." type="journal article" year="1872">Bargagli 1872: 283</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BC5A553AFED3D40D" author="BERTOLINI S." box="[113,298,356,380]" pageId="3" pageNumber="632" refId="ref10674" refString="BERTOLINI S. 1899. - Catalogo dei Coleotteri d'Italia. Rivista Italiana di Scienze Naturali. Sordomuti di L. Lazzeri, Siena, 144 p." type="book" year="1899">Bertolini 1899: 62</bibRefCitation>
;
<bibRefCitation id="7578CAD8DC1CFFD9BD1F553AFE76D40D" author="BERTOLINI S." box="[308,399,356,380]" pageId="3" pageNumber="632" refId="ref10702" refString="BERTOLINI S. 1904. - Catalogo dei Coleotteri d'Italia. Rivista Italiana di Scienze Naturali. Sordomuti di L. Lazzeri, Siena, 127 p." type="book" year="1904">1904: 62</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A55CDFE0ED4DA" blockId="3.[113,503,403,427]" box="[113,503,403,427]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A55CDFED7D4DA" box="[113,302,403,427]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius">
<emphasis id="239D6B3BDC1CFFD9BC5A55CDFED7D4DA" box="[113,302,403,427]" italics="true" pageId="3" pageNumber="632">Ceratophyus hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BD6355CDFE0AD4DA" author="STURM J." box="[328,499,403,427]" pageId="3" pageNumber="632" refId="ref11809" refString="STURM J. 1843. - Catalog der Kaefer-Sammlung. Nurnberg, 386 p." type="book" year="1843">Sturm 1843: 113</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A559CFEE1D485" blockId="3.[112,646,449,500]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A559CFE61D4A8" authorityName="Gene" authorityYear="1836" box="[113,408,449,473]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius" subGenus="Minotaurus">
<emphasis id="239D6B3BDC1CFFD9BC5A559CFF35D4A8" box="[113,204,450,473]" italics="true" pageId="3" pageNumber="632">Geotrupes</emphasis>
(
<emphasis id="239D6B3BDC1CFFD9BCF3559CFEBFD4A8" box="[216,326,450,473]" italics="true" pageId="3" pageNumber="632">Minotaurus</emphasis>
)
<emphasis id="239D6B3BDC1CFFD9BD78559FFE61D4A8" box="[339,408,449,473]" italics="true" pageId="3" pageNumber="632">hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BD86559FFD9FD4A8" author="MARSEUL S. A." box="[429,614,449,473]" pageId="3" pageNumber="632" refId="ref11347" refString="MARSEUL S. A. 1863. - Catalogue des Coleopteres d'Europe et du Bassin de la Mediterranee en Afrique &amp; en Asie. 2 nd ed. Chez l'auteur, Ach. Deyrolle, Paris, 300 p." type="book" year="1863">Marseul 1863: 122</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BC5B5582FEEDD485" author="REITTER E." box="[112,276,476,500]" pageId="3" pageNumber="632" pagination="93 - 102" refId="ref11551" refString="REITTER E. 1883. - [Scarabaeidae], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae et Caucasi. 3 rd ed. W. Janson, London; Libraria Nicolai, Berolini; Luc. Buquet, Paris: 93 - 102." type="book chapter" year="1883">Reitter 1883: 97</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5655FDAED753" blockId="3.[113,599,522,546]" box="[113,599,522,546]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A5655FE65D753" authorityName="Gene" authorityYear="1836" box="[113,412,522,546]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius" subGenus="Chelotrupes">
<emphasis id="239D6B3BDC1CFFD9BC5A5655FF34D753" box="[113,205,523,546]" italics="true" pageId="3" pageNumber="632">Geotrupes</emphasis>
(
<emphasis id="239D6B3BDC1CFFD9BCF05654FEB1D753" box="[219,328,522,546]" italics="true" pageId="3" pageNumber="632">Chelotrupes</emphasis>
)
<emphasis id="239D6B3BDC1CFFD9BD7C5654FE65D753" box="[343,412,522,546]" italics="true" pageId="3" pageNumber="632">hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BD9D5654FDAAD753" author="JEKEL H." box="[438,595,522,546]" pageId="3" pageNumber="632" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">Jekel 1866: 550</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5667FDABD720" blockId="3.[113,594,569,593]" box="[113,594,569,593]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A5667FE7AD720" authorityName="Gene" authorityYear="1836" box="[113,387,569,593]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius" subGenus="Typhoeus">
<emphasis id="239D6B3BDC1CFFD9BC5A5667FF34D721" box="[113,205,569,592]" italics="true" pageId="3" pageNumber="632">Geotrupes</emphasis>
(
<emphasis id="239D6B3BDC1CFFD9BCF05667FED6D720" box="[219,303,569,593]" italics="true" pageId="3" pageNumber="632">Typhoeus</emphasis>
)
<emphasis id="239D6B3BDC1CFFD9BD155667FE7AD720" box="[318,387,569,593]" italics="true" pageId="3" pageNumber="632">hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BDB65667FDB7D720" author="REITTER E." box="[413,590,569,593]" pageId="3" pageNumber="632" pagination="177 - 194" refId="ref11608" refString="REITTER E. 1891. - [Scarabaeidae], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. R. Friedlander &amp; Sohn, Berlin; E. Reitter, Modling; Revue d'Entomologie, Caen: 177 - 194." type="book chapter" year="1891">Reitter 1891: 184</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5639FD90D70E" blockId="3.[113,617,615,639]" box="[113,617,615,639]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A5639FE4DD70E" box="[113,436,615,639]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius" subGenus="Minotaurus">
<emphasis id="239D6B3BDC1CFFD9BC5A5639FF1BD70E" box="[113,226,615,639]" italics="true" pageId="3" pageNumber="632">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC1CFFD9BCDA5636FE99D70E" box="[241,352,616,639]" italics="true" pageId="3" pageNumber="632">Minotaurus</emphasis>
)
<emphasis id="239D6B3BDC1CFFD9BD445639FE4DD70E" box="[367,436,615,639]" italics="true" pageId="3" pageNumber="632">hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BDE65639FD9CD70E" author="REITTER E." box="[461,613,615,639]" pageId="3" pageNumber="632" pagination="1 - 109" refId="ref11666" refString="REITTER E. 1893. - Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des naturforschenden Vereines in Brunn 31: 1 - 109 (dated 1892, published 1893)." type="journal article" year="1893">Reitter 1893: 9</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A56C8FD91D7DF" blockId="3.[113,616,662,686]" box="[113,616,662,686]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A56C8FE60D7DF" box="[113,409,662,686]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius" subGenus="Typhoeus">
<emphasis id="239D6B3BDC1CFFD9BC5A56C8FF1BD7DF" box="[113,226,662,686]" italics="true" pageId="3" pageNumber="632">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC1CFFD9BCDA56C8FEBCD7DF" box="[241,325,662,686]" italics="true" pageId="3" pageNumber="632">Typhoeus</emphasis>
)
<emphasis id="239D6B3BDC1CFFD9BD7F56C8FE60D7DF" box="[340,409,662,686]" italics="true" pageId="3" pageNumber="632">hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BD9856C8FD9DD7DF" author="REITTER E." box="[435,612,662,686]" pageId="3" pageNumber="632" pagination="714 - 750" refId="ref11701" refString="REITTER E. 1906. - [Lamellicornia], in HEYDEN L., REITTER E. &amp; WEISE J. (eds), Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. 2 nd ed. R. Friedlander &amp; Sohn, Berlin; Edmund Reitter, Paskau; Revue d'Entomologie, Caen: 714 - 750." type="book chapter" year="1906">Reitter 1906: 725</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A569AFDC8D660" blockId="3.[112,646,708,785]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A569AFE7FD7AD" authorityName="Gene" authorityYear="1836" box="[113,390,708,732]" class="Clitellata" family="Octochaetidae" genus="Typhoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="632" phylum="Annelida" rank="species" species="hiostius" subGenus="Chelotrupes">
<emphasis id="239D6B3BDC1CFFD9BC5A569AFF3BD7AD" box="[113,194,708,732]" italics="true" pageId="3" pageNumber="632">Typhoeus</emphasis>
(
<emphasis id="239D6B3BDC1CFFD9BCE5569AFECED7AD" box="[206,311,708,732]" italics="true" pageId="3" pageNumber="632">Chelotrupes</emphasis>
)
<emphasis id="239D6B3BDC1CFFD9BD68569AFE7FD7AD" box="[323,390,708,732]" italics="true" pageId="3" pageNumber="632">hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BDB1569AFD9CD7AD" author="BOUCOMONT A." box="[410,613,708,732]" pageId="3" pageNumber="632" refId="ref10730" refString="BOUCOMONT A. 1912. - Scarabaeidae: Taurocerastinae, Geotrupinae. Pars 46. Coleopterorum Catalogus auspiciis et auxilio. W. Junk, Berlin, 47 p." type="book" year="1912">Boucomont 1912: 22</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BC5B5681FED0D786" author="LUIGIONI P." box="[112,297,735,759]" pageId="3" pageNumber="632" pagination="1 - 1160" refId="ref11315" refString="LUIGIONI P. 1929. - I Coleotteri d'Italia. Catalogo sinonimico-topografico-bibliografico. Memorie della Pontificia Accademia delle Scienze - I Nuovi Lincei 13 ser. 2: 1 - 1160." type="book chapter" year="1929">Luigioni 1929: 390</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BD655681FDEBD786" author="WINKLER A." box="[334,530,735,759]" pageId="3" pageNumber="632" pagination="1009 - 1136" refId="ref11824" refString="WINKLER A. 1929. - Catalogus Coleopterorum regionis palaearcticae. Scarabaeidae. Albert Winkler, Wien, 9: 1009 - 1136." type="journal article" year="1929">Winkler 1929: 1036</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BE1D5681FF21D660" author="CROVETTI A." pageId="3" pageNumber="632" pagination="229 - 250" refId="ref10845" refString="CROVETTI A. 1970. - Contributi alla conoscenza dei Coleotteri Scarabeidi. IV. Revisione sistematica dei Typhoeus Leach del sottogenere Chelotrupes Jekel. &quot; Studi Sassaresi &quot; - Sez. III - Annali della Facolta di Agraria dell'Universita di Sassari 18 (1): 229 - 250." type="journal article" year="1970">Crovetti 1970: 235</bibRefCitation>
. —
<bibRefCitation id="7578CAD8DC1CFFD9BD2A56A7FDD4D660" author="CARPANETO G. M. &amp; PIATTELLA E." box="[257,557,761,785]" pageId="3" pageNumber="632" pagination="1 - 18" refId="ref10760" refString="CARPANETO G. M. &amp; PIATTELLA E. 1995. - Coleoptera Polyphaga V (Lucanoidea, Scarabaeoidea), in MINELLI A., RUFFO S. &amp; LA POSTA S. (eds), Checklist delle specie della fauna italiana, 50. Calderini, Bologna: 1 - 18." type="book chapter" year="1995">Carpaneto &amp; Piattella 1995: 4</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5779FD8BD631" blockId="3.[113,626,807,832]" box="[113,626,807,832]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A5779FE4BD64E" box="[113,434,807,832]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="species" species="hiostius" subGenus="Chelotrupes">
<emphasis id="239D6B3BDC1CFFD9BC5A5779FF1BD64E" box="[113,226,807,831]" italics="true" pageId="3" pageNumber="632">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC1CFFD9BCDA5779FEA7D64E" box="[241,350,807,831]" italics="true" pageId="3" pageNumber="632">Chelotrupes</emphasis>
)
<emphasis id="239D6B3BDC1CFFD9BD465779FE4BD64E" box="[365,434,807,831]" italics="true" pageId="3" pageNumber="632">hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BDE75776FD97D631" author="PORTA A." box="[460,622,808,832]" pageId="3" pageNumber="632" refId="ref11528" refString="PORTA A. 1932. - Fauna Coleopterorum italica. Rhyncophora-Lamellicornia. Stabilimento tipografico Piacentino, Piacenza, 5, 476 p." type="book" year="1932">Porta 1932: 401</bibRefCitation>
.
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5708FE2FD61F" blockId="3.[113,470,854,878]" box="[113,470,854,878]" pageId="3" pageNumber="632">
<taxonomicName id="D6E9CCAADC1CFFD9BC5A5708FEE8D61F" authorityName="Gene" authorityYear="1836" box="[113,273,854,878]" class="Clitellata" family="Octochaetidae" genus="Typhoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="632" phylum="Annelida" rank="species" species="hiostius">
<emphasis id="239D6B3BDC1CFFD9BC5A5708FEE8D61F" box="[113,273,854,878]" italics="true" pageId="3" pageNumber="632">Typhoeus hiostius</emphasis>
</taxonomicName>
–
<bibRefCitation id="7578CAD8DC1CFFD9BD005708FE2BD61F" author="BARAUD J." box="[299,466,854,878]" pageId="3" pageNumber="632" refId="ref10533" refString="BARAUD J. 1992. - Coleopteres Scarabaeoidea d'Europe. Faune de France, France et regions limitrophes, 78. Federation francaise des Societes de Sciences naturelles, Paris; Societe linneenne de Lyon, Lyon, 856 p." type="book" year="1992">Baraud 1992: 49</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1CFFD9BC5A57DBFE4DD144" pageId="3" pageNumber="632" type="materials_examined">
<paragraph id="1156B729DC1CFFD9BC5A57DBFEE4D177" blockId="3.[111,648,901,1030]" pageId="3" pageNumber="632">
<collectionCode id="77F82FECDC1CFFD9BC5A57DBFF5CD6EB" box="[113,165,901,924]" pageId="3" pageNumber="632">TYPE</collectionCode>
MATERIAL. —
<materialsCitation id="A181BD74DC1CFFD9BD6957DBFEE4D177" collectionCode="MRSNT" determinerName="Des. Dellacasa" location="Sard [inia]" municipality="Sard [inia]" pageId="3" pageNumber="632" specimenCount="1" specimenCount-male="1" typeStatus="neotype">
<specimenCount id="07EF7CA0DC1CFFD9BD6957DBFE95D6ED" box="[322,364,901,925]" pageId="3" pageNumber="632" type="male">1 ♂</specimenCount>
<typeStatus id="CE52098BDC1CFFD9BD5D57DBFE31D6EC" box="[374,456,901,925]" pageId="3" pageNumber="632" type="neotype">neotype</typeStatus>
(here designated) (
<collectionCode id="77F82FECDC1CFFD9BC5357C1FF28D6C6" box="[120,209,927,951]" pageId="3" pageNumber="632">MRSNT</collectionCode>
), bearing the following labels:1) yellow, printed: “1 / 37”; 2) white, printed: “
<collectingMunicipality id="F1322D53DC1CFFD9BD5057E4FE19D6A3" box="[379,480,954,978]" pageId="3" pageNumber="632">Sard[inia]</collectingMunicipality>
.”; 3) red, printed: “
<taxonomicName id="D6E9CCAADC1CFFD9BC57578AFF21D69A" authorityName="Latreille" authorityYear="1796" box="[124,216,980,1003]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BC57578AFF21D69A" box="[124,216,980,1003]" italics="true" pageId="3" pageNumber="632">Geotrupes</emphasis>
</taxonomicName>
/
<taxonomicName id="D6E9CCAADC1CFFD9BCC2578AFED7D69D" authorityName="Gene" authorityYear="1836" box="[233,302,980,1004]" class="Clitellata" family="Octochaetidae" genus="Typhoeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="632" phylum="Annelida" rank="species" species="hiostius">
<emphasis id="239D6B3BDC1CFFD9BCC2578AFED7D69D" box="[233,302,980,1004]" italics="true" pageId="3" pageNumber="632">hiostius</emphasis>
</taxonomicName>
/
<bibRefCitation id="7578CAD8DC1CFFD9BD6A578AFE4AD69D" author="GENE J." box="[321,435,980,1004]" pageId="3" pageNumber="632" pagination="161 - 191" refId="ref10988" refString="GENE J. 1836. - De quibusdam insectis Sardiniae novis aut minus cognitis. Fasciculus I. Memoralia Regiae Scientiarum Academiae Taurinensis 39: 161 - 191." type="journal article" year="1836">Gené, 1836</bibRefCitation>
/ NEOTYPUS /
<determinerName id="E48A5BEDDC1CFFD9BE71578AFF36D177" pageId="3" pageNumber="632">Des. Dellacasa</determinerName>
, 2005”.
</materialsCitation>
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5043FE4DD144" blockId="3.[113,436,1053,1077]" box="[113,436,1053,1077]" pageId="3" pageNumber="632">
<typeStatus id="CE52098BDC1CFFD9BC5A5043FF5BD142" box="[113,162,1053,1076]" pageId="3" pageNumber="632">
<collectionCode id="77F82FECDC1CFFD9BC5A5043FF5BD142" box="[113,162,1053,1076]" pageId="3" pageNumber="632">TYPE</collectionCode>
</typeStatus>
LOCALITY. —
<collectingRegion id="D32D79CBDC1CFFD9BD055041FE85D144" box="[302,380,1055,1077]" country="Italy" name="Sardegna" pageId="3" pageNumber="632">Sardinia</collectingRegion>
,
<collectingCountry id="69FEF7B9DC1CFFD9BDAD5041FE49D144" box="[390,432,1055,1077]" name="Italy" pageId="3" pageNumber="632">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1CFFD9BC5A5012FF31D1E9" pageId="3" pageNumber="632" type="etymology">
<paragraph id="1156B729DC1CFFD9BC5A5012FF31D1E9" blockId="3.[112,647,1099,1176]" pageId="3" pageNumber="632">ETYMOLOGY. — Named in honour of Hiostius [Hostius], son of Ampsicora, hero of Sardinian resistance to Roman invasion.</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1CFFD9BC5A50F1FC56D6F5" pageId="3" pageNumber="632" type="materials_examined">
<paragraph id="1156B729DC1CFFD9BC5A50F1FF25D067" blockId="3.[111,648,1198,1699]" pageId="3" pageNumber="632">
MATERIAL EXAMINED. —
<emphasis id="239D6B3BDC1CFFD9BD4A50F0FE41D1B7" bold="true" box="[353,440,1198,1222]" pageId="3" pageNumber="632">
<collectingRegion id="D32D79CBDC1CFFD9BD4A50F0FE4DD1B7" box="[353,436,1198,1222]" country="Italy" name="Sardegna" pageId="3" pageNumber="632">Sardinia</collectingRegion>
.
</emphasis>
Sard. [
<collectingRegion id="D32D79CBDC1CFFD9BDDC50F1FDB4D1B6" box="[503,589,1199,1223]" country="Italy" name="Sardegna" pageId="3" pageNumber="632">Sardinia</collectingRegion>
],
<specimenCount id="07EF7CA0DC1CFFD9BE7150F1FD7ED1B6" box="[602,647,1199,1223]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(MRSNT).—
<collectingRegion id="D32D79CBDC1CFFD9BCD25097FEB7D190" box="[249,334,1225,1249]" country="Italy" name="Sardegna" pageId="3" pageNumber="632">Sardegna</collectingRegion>
,
<specimenCount id="07EF7CA0DC1CFFD9BD7E5097FE86D190" box="[341,383,1225,1249]" pageId="3" pageNumber="632" type="generic">2 ex.</specimenCount>
(MCSNG-BC).—
<collectingRegion id="D32D79CBDC1CFFD9BE1F5097FD78D190" box="[564,641,1225,1249]" country="Italy" name="Sardegna" pageId="3" pageNumber="632">Sardinia</collectingRegion>
,
<date id="655791E9DC1CFFD9BC4450BAFF4BD18D" box="[111,178,1252,1276]" pageId="3" pageNumber="632" value="1837-01">I.1837</date>
,
<specimenCount id="07EF7CA0DC1CFFD9BC9750BAFF13D18D" box="[188,234,1252,1276]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(MSNTC). —
<collectingRegion id="D32D79CBDC1CFFD9BDAD50BAFE20D18D" box="[390,473,1252,1276]" country="Italy" name="Sardegna" pageId="3" pageNumber="632">Sardinia</collectingRegion>
, leg. Damry,
<specimenCount id="07EF7CA0DC1CFFD9BE7C50BAFD7CD18D" box="[599,645,1252,1276]" pageId="3" pageNumber="632" type="generic">3 ex.</specimenCount>
(MNHN).
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5147FF5FD03A" blockId="3.[111,648,1198,1699]" pageId="3" pageNumber="632">
Cagliari province, 7-fratelli, leg. U. Lostia,
<specimenCount id="07EF7CA0DC1CFFD9BDDA5147FDE5D040" box="[497,540,1305,1329]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(MCSNG-DC).
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5110FEC2D325" blockId="3.[111,648,1198,1699]" pageId="3" pageNumber="632">
Carbonia-Iglesias province,Carloforte, leg. A.Dodero,
<specimenCount id="07EF7CA0DC1CFFD9BE765110FD71D017" box="[605,648,1358,1382]" pageId="3" pageNumber="632" type="generic">3 ex.</specimenCount>
(MCSNG -MC);
<specimenCount id="07EF7CA0DC1CFFD9BD325136FEB1D0F1" box="[281,328,1384,1408]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(MNHN). —
<taxonomicName id="D6E9CCAADC1CFFD9BDF35136FDFED0F1" box="[472,519,1384,1408]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BDF35136FDFED0F1" box="[472,519,1384,1408]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9BE395136FD7BD0F1" box="[530,642,1384,1408]" pageId="3" pageNumber="632" value="1902-04-27">27.IV.1902</date>
, leg. A.Dodero,
<specimenCount id="07EF7CA0DC1CFFD9BCD051DCFEDCD0EB" box="[251,293,1410,1434]" pageId="3" pageNumber="632" type="generic">9 ex.</specimenCount>
(MCSNG -DC). —
<taxonomicName id="D6E9CCAADC1CFFD9BDC851DCFDF6D0EB" box="[483,527,1410,1434]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BDC851DCFDF6D0EB" box="[483,527,1410,1434]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9BE3251DCFD7DD0EB" box="[537,644,1410,1434]" pageId="3" pageNumber="632" value="1902-04-27">27.IV.1902</date>
, leg. A. Dodero,
<specimenCount id="07EF7CA0DC1CFFD9BCD551C3FED0D0C4" box="[254,297,1437,1461]" pageId="3" pageNumber="632" type="generic">3 ex.</specimenCount>
(MCSNG -BC). —
<taxonomicName id="D6E9CCAADC1CFFD9BDCC51C3FDEDD0C4" box="[487,532,1437,1461]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BDCC51C3FDEDD0C4" box="[487,532,1437,1461]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9BE3451C3FD7AD0C4" box="[543,643,1437,1461]" pageId="3" pageNumber="632" value="1892-05-20">20.V.1892</date>
, leg. A. Dodero,
<specimenCount id="07EF7CA0DC1CFFD9BD3F51E9FEBFD0BE" box="[276,326,1463,1487]" pageId="3" pageNumber="632" type="generic">3 ex.</specimenCount>
(MCSNG -DC). — Sant’Anna Arresi, Cala Sa Barracca (o
<collectingRegion id="D32D79CBDC1CFFD9BDA3518CFE46D09B" box="[392,447,1490,1514]" country="Portugal" name="Porto" pageId="3" pageNumber="632">Portu</collectingRegion>
Su Trigu),
<quantity id="D6111ACCDC1CFFD9BE1E518CFD7BD09B" box="[565,642,1490,1514]" metricMagnitude="1" metricUnit="m" metricValue="2.65" metricValueMax="4.8" metricValueMin="0.5" pageId="3" pageNumber="632" unit="m" value="26.5" valueMax="48.0" valueMin="5.0">5-48 m</quantity>
,
<date id="655791E9DC1CFFD9BC5851B2FF18D375" box="[115,225,1516,1540]" pageId="3" pageNumber="632" value="1998-12-03">3.XII.1998</date>
, leg. C. Meloni,
<specimenCount id="07EF7CA0DC1CFFD9BD5051B2FE51D375" box="[379,424,1516,1540]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(coll. CM). — Spiaggia di Portixeddu (Fluminimaggiore-Buggerru),
<date id="655791E9DC1CFFD9BE365259FD79D36E" box="[541,640,1543,1567]" pageId="3" pageNumber="632" value="1984-01-17">17.I.1984</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9BCC9527FFEEBD348" box="[226,274,1569,1593]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(coll. PL). —
<taxonomicName id="D6E9CCAADC1CFFD9BD8E527FFE2CD348" box="[421,469,1569,1593]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BD8E527FFE2CD348" box="[421,469,1569,1593]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9BDC8527FFDBBD348" box="[483,578,1569,1593]" pageId="3" pageNumber="632" value="1984-02-07">7.II.1984</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9BC895262FF2BD325" box="[162,210,1596,1620]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(coll. PL).
</paragraph>
<paragraph id="1156B729DC1CFFD9BC5A5208FC64D710" blockId="3.[111,648,1198,1699]" lastBlockId="3.[675,1211,215,900]" pageId="3" pageNumber="632">
Medio Campidano province, Arbus, loc. Piscinas di Ingurtosu,
<date id="655791E9DC1CFFD9BC88522FFEFED3F8" box="[163,263,1649,1673]" pageId="3" pageNumber="632" value="2001-03-05">5.III.2001</date>
, leg. Lisa,
<specimenCount id="07EF7CA0DC1CFFD9BD48522FFE76D3F8" box="[355,399,1649,1673]" pageId="3" pageNumber="632" type="generic">3 ex.</specimenCount>
(coll. SZ). — Costa Verde,
<taxonomicName id="D6E9CCAADC1CFFD9BC4452D5FEEFD3D2" authority="di Arbus" authorityName="di Arbus" box="[111,278,1675,1699]" class="Magnoliopsida" family="Fabaceae" genus="Marina" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="632" phylum="Tracheophyta" rank="genus">Marina di Arbus</taxonomicName>
dint.,
<geoCoordinate id="74DDD1EEDC1CFFD9BD7752D5FE12D3D2" box="[348,491,1675,1699]" degrees="39" direction="north" minutes="33" orientation="latitude" pageId="3" pageNumber="632" precision="1" seconds="51.0" value="39.564167">39°33’51.0”N</geoCoordinate>
,
<geoCoordinate id="74DDD1EEDC1CFFD9BDDC52D5FD79D3D2" box="[503,640,1675,1699]" degrees="08" direction="east" minutes="27" orientation="longitude" pageId="3" pageNumber="632" precision="1" seconds="48.6" value="8.4635">08°27’48.6”E</geoCoordinate>
,
<quantity id="D6111ACCDC1CFFD9BE8E5489FD22D59E" box="[677,731,215,239]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="632" unit="m" value="50.0">50 m</quantity>
,
<date id="655791E9DC1CFFD9BECD5489FCA2D59E" box="[742,859,215,239]" pageId="3" pageNumber="632" value="2005-03-13">13.III.2005</date>
, leg. M. Dellacasa &amp; C. Fresi,
<specimenCount id="07EF7CA0DC1CFFD9B8A05489FB43D59E" box="[1163,1210,215,239]" pageId="3" pageNumber="632" type="generic">8 ex.</specimenCount>
(coll. MD). —
<taxonomicName id="D6E9CCAADC1CFFD9BF1254AFFC90D478" box="[825,873,241,265]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BF1254AFFC90D478" box="[825,873,241,265]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9BF5E54AFFC2AD478" box="[885,979,241,265]" pageId="3" pageNumber="632" value="2005-10-01">1.X.2005</date>
, leg. M. Dellacasa &amp; C. Fresi,
<specimenCount id="07EF7CA0DC1CFFD9BECA5552FCEBD455" box="[737,786,268,292]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(coll. MD). —
<taxonomicName id="D6E9CCAADC1CFFD9BF925552FC10D455" box="[953,1001,268,292]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BF925552FC10D455" box="[953,1001,268,292]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9BFD35552FB9DD455" box="[1016,1124,268,292]" pageId="3" pageNumber="632" value="2005-10-29">29.X.2005</date>
, leg. M. Dellacasa &amp; C. Fresi,
<specimenCount id="07EF7CA0DC1CFFD9BF505578FC4ED44F" box="[891,951,294,318]" pageId="3" pageNumber="632" type="generic">13 ex.</specimenCount>
(coll. MD). — Duna di is Arenas-s’Acqua e s’Ollastu (tra Punta s’Aschivoni e Torre di Flumentorgiu),
<date id="655791E9DC1CFFD9BF705505FC36D402" box="[859,975,347,371]" pageId="3" pageNumber="632" value="1993-11-11">11.XI.1993</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9B8645505FB79D402" box="[1103,1152,347,371]" pageId="3" pageNumber="632" type="generic">2 ex.</specimenCount>
(coll. PL). — Giara di Gesturi,
<date id="655791E9DC1CFFD9BF8B5528FBECD4FF" box="[928,1045,374,398]" pageId="3" pageNumber="632" value="1987-06-30">30.VI.1987</date>
, leg. M. Arnone,
<specimenCount id="07EF7CA0DC1CFFD9BE8F55CEFD28D4D9" box="[676,721,400,424]" pageId="3" pageNumber="632" type="generic">2 ex.</specimenCount>
(MCSNG). —
<taxonomicName id="D6E9CCAADC1CFFD9BF4355CEFBFCD4D9" authority="di Arbus" authorityName="di Arbus" box="[872,1029,400,424]" class="Magnoliopsida" family="Fabaceae" genus="Marina" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="632" phylum="Tracheophyta" rank="genus">Marina di Arbus</taxonomicName>
,
<date id="655791E9DC1CFFD9B82655CEFB84D4D9" box="[1037,1149,400,424]" pageId="3" pageNumber="632" value="1995-04-26">26.IV.1995</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9BEF855F5FCF8D4B2" box="[723,769,427,451]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(coll. MD). —
<taxonomicName id="D6E9CCAADC1CFFD9BFB755F4FC35D4B3" box="[924,972,426,450]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BFB755F4FC35D4B3" box="[924,972,426,450]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9BFFC55F5FBB0D4B2" box="[983,1097,427,451]" pageId="3" pageNumber="632" value="1995-04-26">26.IV.1995</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9BE8E559BFD26D4AC" box="[677,735,453,477]" pageId="3" pageNumber="632" type="generic">11 ex.</specimenCount>
(coll. SZ). —
<taxonomicName id="D6E9CCAADC1CFFD9BF43559BFBF0D4AC" authority="di Arbus" authorityName="di Arbus" box="[872,1033,453,477]" class="Magnoliopsida" family="Fabaceae" genus="Marina" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="632" phylum="Tracheophyta" rank="genus">Marina di Arbus</taxonomicName>
, foce Rio Piscinas,
<date id="655791E9DC1CFFD9BE8855BEFCF6D489" box="[675,783,480,504]" pageId="3" pageNumber="632" value="1995-04-26">26.IV.1995</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9BF5155BEFC5FD489" box="[890,934,480,504]" pageId="3" pageNumber="632" type="generic">8 ex.</specimenCount>
(coll. PL).—
<taxonomicName id="D6E9CCAADC1CFFD9B80D5581FBAAD486" box="[1062,1107,479,503]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9B80D5581FBAAD486" box="[1062,1107,479,503]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9B87555BEFB4FD489" box="[1118,1206,480,504]" pageId="3" pageNumber="632" value="1999-05-04">4.V.1999</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9BF3F55A4FCA8D763" box="[788,849,506,530]" pageId="3" pageNumber="632" type="generic">19 ex.</specimenCount>
(coll. PL). —
<taxonomicName id="D6E9CCAADC1CFFD9BFCF55A4FBEDD763" box="[996,1044,506,530]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BFCF55A4FBEDD763" box="[996,1044,506,530]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9B80A55A4FB75D763" box="[1057,1164,506,530]" pageId="3" pageNumber="632" value="1979-10-19">19.X.1979</date>
, leg. L. &amp; R. Pittino,
<specimenCount id="07EF7CA0DC1CFFD9BF60564AFC82D75D" box="[843,891,532,556]" pageId="3" pageNumber="632" type="generic">6 ex.</specimenCount>
(coll. RP). —
<taxonomicName id="D6E9CCAADC1CFFD9B826564AFBC4D75D" box="[1037,1085,532,556]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9B826564AFBC4D75D" box="[1037,1085,532,556]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9B861564AFB4FD75D" box="[1098,1206,532,556]" pageId="3" pageNumber="632" value="1994-10-24">24.X.1994</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9BF3A5671FCB9D736" box="[785,832,559,583]" pageId="3" pageNumber="632" type="generic">4 ex.</specimenCount>
(coll. PL). —
<taxonomicName id="D6E9CCAADC1CFFD9BFE15671FC03D736" box="[970,1018,559,583]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BFE15671FC03D736" box="[970,1018,559,583]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
, 8/
<date id="655791E9DC1CFFD9B8325671FB76D736" box="[1049,1167,559,583]" pageId="3" pageNumber="632" value="2002-11-17">17.XI.2002</date>
, leg. P. Leo,
<specimenCount id="07EF7CA0DC1CFFD9BEC75617FCCDD710" box="[748,820,585,609]" pageId="3" pageNumber="632" type="generic">362 ex.</specimenCount>
(coll. PL).
</paragraph>
<paragraph id="1156B729DC1CFFD9BE8E563AFBA0D7E7" blockId="3.[675,1211,215,900]" pageId="3" pageNumber="632">
Nuoro province, Macomer, Stagno di Bara,
<quantity id="D6111ACCDC1CFFD9B844563AFB4CD70D" box="[1135,1205,612,636]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="632" unit="m" value="660.0">660 m</quantity>
,
<date id="655791E9DC1CFFD9BE8D5620FCE5D7E7" box="[678,796,638,662]" pageId="3" pageNumber="632" value="1979-03-25">25.III.1979</date>
. leg. C. Meloni,
<specimenCount id="07EF7CA0DC1CFFD9BFEB5620FC16D7E7" box="[960,1007,638,662]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(coll. SZ).
</paragraph>
<paragraph id="1156B729DC1CFFD9BE8E56C7FC40D7BA" blockId="3.[675,1211,215,900]" pageId="3" pageNumber="632">
Ogliastra province, Villanova Strisaili,
<date id="655791E9DC1CFFD9B83756C7FB77D7C0" box="[1052,1166,665,689]" pageId="3" pageNumber="632" value="1872-04-29">29.IV.1872</date>
, leg. R. Gestro,
<specimenCount id="07EF7CA0DC1CFFD9BF3B56EDFCC6D7BA" box="[784,831,691,715]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(MCSNG).
</paragraph>
<paragraph id="1156B729DC1CFFD9BE8E5690FC56D6F5" blockId="3.[675,1211,215,900]" pageId="3" pageNumber="632">
Oristano province, Monte Arci,
<date id="655791E9DC1CFFD9BFE65690FBC1D797" box="[973,1080,718,742]" pageId="3" pageNumber="632" value="1983-12-06">6.XII.1983</date>
, leg. Fancello,
<specimenCount id="07EF7CA0DC1CFFD9BE8856B6FD2DD671" box="[675,724,744,768]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(MCSNG). — Oristano, Monte Ferru,
<quantity id="D6111ACCDC1CFFD9B84456B6FB4AD671" box="[1135,1203,744,768]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="632" unit="m" value="350.0">350 m</quantity>
,
<date id="655791E9DC1CFFD9BE8D575DFCE2D66A" box="[678,795,771,795]" pageId="3" pageNumber="632" value="1979-04-30">30.IV.1979</date>
, leg. S. Riese,
<specimenCount id="07EF7CA0DC1CFFD9BF98575DFC1CD66A" box="[947,997,771,795]" pageId="3" pageNumber="632" type="generic">2 ex.</specimenCount>
(MCSNG). — Oristano, Pen. del Sinis,
<date id="655791E9DC1CFFD9BFAB5743FC0AD644" box="[896,1011,797,821]" pageId="3" pageNumber="632" value="1978-04-26">26.IV.1978</date>
, leg. S. Riese,
<specimenCount id="07EF7CA0DC1CFFD9B8A25743FB43D644" box="[1161,1210,797,821]" pageId="3" pageNumber="632" type="generic">2 ex.</specimenCount>
(MCSNG). — San Giovanni di Sinis,
<date id="655791E9DC1CFFD9B8315766FB77D621" box="[1050,1166,824,848]" pageId="3" pageNumber="632" value="1988-11-17">17.XI.1988</date>
, leg. C. Meloni,
<specimenCount id="07EF7CA0DC1CFFD9BF33570CFCB0D61B" box="[792,841,850,874]" pageId="3" pageNumber="632" type="generic">1 ex.</specimenCount>
(coll. SZ). —
<taxonomicName id="D6E9CCAADC1CFFD9BFF0570CFBF2D61B" box="[987,1035,850,874]" class="Arachnida" family="Brachychthoniidae" genus="Idem" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="632" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1CFFD9BFF0570CFBF2D61B" box="[987,1035,850,874]" italics="true" pageId="3" pageNumber="632">Idem</emphasis>
</taxonomicName>
,
<date id="655791E9DC1CFFD9B833570CFB75D61B" box="[1048,1164,850,874]" pageId="3" pageNumber="632" value="1986-11-21">21.XI.1986</date>
, leg. C. Meloni,
<specimenCount id="07EF7CA0DC1CFFD9BF3D5732FCBFD6F5" box="[790,838,876,900]" pageId="3" pageNumber="632" type="generic">3 ex.</specimenCount>
(coll. SZ).
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1CFFD9BE8E57C2FB41D142" pageId="3" pageNumber="632" type="distribution">
<paragraph id="1156B729DC1CFFD9BE8E57C2FC0ED698" blockId="3.[675,1210,924,1001]" pageId="3" pageNumber="632">
DISTRIBUTION. — Known only from Sardinia (
<figureCitation id="89D2ABACDC1CFFD9B85957C2FB55D6C5" box="[1138,1196,924,948]" captionStart="FIG" captionStartId="9.[113,124,1003,1020]" captionTargetBox="[135,643,227,952]" captionTargetId="figure-20@9.[135,668,209,978]" captionTargetPageId="9" captionText="FIG. 3. — Distribution in Sardinia (Italy) of Chelotrupes hiostius n. comb. (■) and C matutinalis n. comb. (▲). ○, sympatry of C. hiostius n. comb. and C. matutinalis n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402563" httpUri="https://zenodo.org/record/5402563/files/figure.png" pageId="3" pageNumber="632">Fig. 3</figureCitation>
). We examined
<specimenCount id="07EF7CA0DC1CFFD9BF0257E9FC50D6BE" box="[809,937,951,975]" pageId="3" pageNumber="632" type="generic">one specimen</specimenCount>
in MNHN labelled “
<collectingCountry id="69FEF7B9DC1CFFD9B85A57E9FB54D6BE" box="[1137,1197,951,975]" name="Greece" pageId="3" pageNumber="632">Grèce</collectingCountry>
”, the locality needs to be confirmed.
</paragraph>
<paragraph id="1156B729DC1CFFD9BE8E505FFB41D142" blockId="3.[677,1210,1025,1075]" pageId="3" pageNumber="632">BIONOMICS. — Flightless telocoprid species present all year round, distributed from the sea level to high hills.</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1CFFDFBE8E5031FDAAD71E" lastPageId="5" lastPageNumber="634" pageId="3" pageNumber="632" type="description">
<paragraph id="1156B729DC1CFFD9BE8E5031FCCCD1F6" blockId="3.[674,1212,1135,1700]" box="[677,821,1135,1161]" pageId="3" pageNumber="632">DESCRIPTION</paragraph>
<paragraph id="1156B729DC1CFFDEBE8E50D0FDDFD3D5" blockId="3.[674,1212,1135,1700]" lastBlockId="4.[111,649,1483,1700]" lastPageId="4" lastPageNumber="633" pageId="3" pageNumber="632">Length 13.5-26.0 mm, very stout, strongly convex, almost glabrous, upper face entirely with metallic shine. Blackish, antennae and legs black.Head with epistoma broadly rhomboidal, with median longitudinal gibbosity more or less raised and coarsely, densely punctate; clypeus broadly triangular, widely rounded at anterior angle, very thinly margined, the edge almost sparsely ciliate and distinctly raised distally; genal sutures sharply cariniform,strongly raised;genae widely rounded; front smooth or sparsely punctate on disc, distinctly punctate laterally. Pronotum strongly transverse, moderately convex, with deep anteromedian rounded foveola on sides; the latter arcuate, thinly margined, the edge almost densely ciliate; hind angles widely rounded; base thinly margined, slightly inwardly sinuate.Scutellum broadly triangular, slightly grooved at middle. Elytra very short, strongly convex, narrowly flattened before the epipleural carina; the latter sparsely and almost elongately ciliate; humeral callus strongly gibbous; striae very superficial, nearly obsolete, almost imperceptibly punctate; intervals flat, smooth. Fore tibiae apical spur distinctly curved apically. First segment of hind tarsi as long as following three combined.</paragraph>
<caption id="4596E7A1DC1BFFDEBC5A5115FDF6D0D1" ID-DOI="http://doi.org/10.5281/zenodo.5402559" ID-Zenodo-Dep="5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="4" pageNumber="633" startId="4.[113,124,1355,1372]" targetBox="[182,1142,216,1290]" targetPageId="4">
<paragraph id="1156B729DC1BFFDEBC5A5115FDF6D0D1" blockId="4.[113,1211,1354,1440]" pageId="4" pageNumber="633">
FIG. 1. —
<emphasis id="239D6B3BDC1BFFDEBCEA5114FF1AD02A" bold="true" box="[193,227,1354,1372]" pageId="4" pageNumber="633">A -C</emphasis>
, heads;
<emphasis id="239D6B3BDC1BFFDEBD065114FEC3D02A" bold="true" box="[301,314,1354,1371]" pageId="4" pageNumber="633">A</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBD6F5114FDA5D02D" authority="(Gene, 1836)" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[324,604,1354,1372]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBD6F5114FE17D02A" box="[324,494,1354,1371]" italics="true" pageId="4" pageNumber="633">Chelotrupes hiostius</emphasis>
(
<bibRefCitation id="7578CAD8DC1BFFDEBDDC5115FDA0D02D" author="GENE J." box="[503,601,1355,1372]" pageId="4" pageNumber="633" pagination="161 - 191" refId="ref10988" refString="GENE J. 1836. - De quibusdam insectis Sardiniae novis aut minus cognitis. Fasciculus I. Memoralia Regiae Scientiarum Academiae Taurinensis 39: 161 - 191." type="journal article" year="1836">Gené, 1836</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBE495115FD49D02D" box="[610,688,1355,1372]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBE9D5114FD3AD02A" bold="true" box="[694,707,1354,1371]" pageId="4" pageNumber="633">B</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBEE65114FC48D02D" authority="(Baudi, 1870)" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Baudi" baseAuthorityYear="1870" box="[717,945,1354,1372]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="matutinalis" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBEE65114FCC7D02A" box="[717,830,1354,1371]" italics="true" pageId="4" pageNumber="633">C. matutinalis</emphasis>
(
<bibRefCitation id="7578CAD8DC1BFFDEBF635115FC54D02D" author="BAUDI F." box="[840,941,1355,1372]" pageId="4" pageNumber="633" pagination="49 - 90" refId="ref10605" refString="BAUDI F. 1870. - Celeopterorum messis in insula Cypro et Asia minore ab EugenioTruqui congregata recensitio: de Europaeis notis quibusdam additis. Pars tertia. Berliner entomologische Zeitschrift 14: 49 - 90." type="journal article" year="1870">Baudi, 1870</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBF9D5115FBFDD02D" box="[950,1028,1355,1372]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEB8215114FBE1D02A" bold="true" box="[1034,1048,1354,1371]" pageId="4" pageNumber="633">C</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEB8095114FF66D003" authority="(Olivier, 1789)" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Olivier" baseAuthorityYear="1789" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="momus" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEB8095114FB81D02A" box="[1058,1144,1354,1371]" italics="true" pageId="4" pageNumber="633">C. momus</emphasis>
(
<bibRefCitation id="7578CAD8DC1BFFDEB8AA5115FF62D003" author="OLIVIER A. G." pageId="4" pageNumber="633" refId="ref11485" refString="OLIVIER A. G. 1789. - Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Baudouin, Paris, 1, 190 p." type="book" year="1789">Olivier, 1789</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBC8E513FFF0DD003" box="[165,244,1377,1394]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBCD1513FFEE3D003" bold="true" box="[250,282,1377,1394]" pageId="4" pageNumber="633">D -F</emphasis>
, pronotum of the male;
<emphasis id="239D6B3BDC1BFFDEBDCC513FFE0CD003" bold="true" box="[487,501,1377,1394]" pageId="4" pageNumber="633">D</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBE2B513FFDA1D003" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[512,600,1377,1394]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBE2B513FFDA1D003" box="[512,600,1377,1394]" italics="true" pageId="4" pageNumber="633">C. hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBE76513FFD55D003" box="[605,684,1377,1394]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBE99513FFD47D003" bold="true" box="[690,702,1377,1394]" pageId="4" pageNumber="633">E</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBEE2513FFCC3D003" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Baudi" baseAuthorityYear="1870" box="[713,826,1377,1394]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="matutinalis" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBEE2513FFCC3D003" box="[713,826,1377,1394]" italics="true" pageId="4" pageNumber="633">C. matutinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBF6B513FFC76D003" box="[832,911,1377,1394]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBFBE513FFC59D003" bold="true" box="[917,928,1377,1394]" pageId="4" pageNumber="633">F</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBF80513FFBF8D003" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[939,1025,1377,1394]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="momus" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBF80513FFBF8D003" box="[939,1025,1377,1394]" italics="true" pageId="4" pageNumber="633">C. momus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEB82C513FFBACD003" box="[1031,1109,1377,1394]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEB877513FFB8ED003" bold="true" box="[1116,1143,1377,1394]" pageId="4" pageNumber="633">G -I</emphasis>
, pronotum of the female;
<emphasis id="239D6B3BDC1BFFDEBD3E5126FEDAD0F8" bold="true" box="[277,291,1400,1417]" pageId="4" pageNumber="633">G</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBD055126FE7ED0F8" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[302,391,1400,1417]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBD055126FE7ED0F8" box="[302,391,1400,1417]" italics="true" pageId="4" pageNumber="633">C. hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBDA65126FE24D0F8" box="[397,477,1400,1417]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBDCF5126FE0BD0F8" bold="true" box="[484,498,1400,1417]" pageId="4" pageNumber="633">H</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBDD65126FD96D0F8" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Baudi" baseAuthorityYear="1870" box="[509,623,1400,1417]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="matutinalis" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBDD65126FD96D0F8" box="[509,623,1400,1417]" italics="true" pageId="4" pageNumber="633">C. matutinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBE5D5126FD3CD0F8" box="[630,709,1400,1417]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBEE75126FD2BD0F8" bold="true" box="[716,722,1400,1417]" pageId="4" pageNumber="633">I</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBEF65126FCCDD0F8" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[733,820,1400,1417]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="momus" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBEF65126FCCDD0F8" box="[733,820,1400,1417]" italics="true" pageId="4" pageNumber="633">C. momus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBF115126FC73D0F8" box="[826,906,1400,1417]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBFBA5126FC54D0F8" bold="true" box="[913,941,1400,1417]" pageId="4" pageNumber="633">J -L</emphasis>
, elytra;
<emphasis id="239D6B3BDC1BFFDEBFDE5126FC06D0F8" bold="true" box="[1013,1023,1400,1417]" pageId="4" pageNumber="633">J</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEB8215126FB9AD0F8" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[1034,1123,1400,1417]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEB8215126FB9AD0F8" box="[1034,1123,1400,1417]" italics="true" pageId="4" pageNumber="633">C. hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEB8425126FB40D0F8" box="[1129,1209,1400,1417]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBC5A51D0FF87D0EE" bold="true" box="[113,126,1422,1439]" pageId="4" pageNumber="633">K</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBCA251D0FF03D0EE" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Baudi" baseAuthorityYear="1870" box="[137,250,1422,1439]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="matutinalis" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBCA251D0FF03D0EE" box="[137,250,1422,1439]" italics="true" pageId="4" pageNumber="633">C. matutinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBCD451D1FEB7D0D1" box="[255,334,1423,1440]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
;
<emphasis id="239D6B3BDC1BFFDEBD7F51D0FEA6D0EE" bold="true" box="[340,351,1422,1439]" pageId="4" pageNumber="633">L</emphasis>
,
<taxonomicName id="D6E9CCAADC1BFFDEBD4251D0FE46D0EE" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[361,447,1422,1439]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="633" phylum="Arthropoda" rank="species" species="momus" status="comb. nov.">
<emphasis id="239D6B3BDC1BFFDEBD4251D0FE46D0EE" box="[361,447,1422,1439]" italics="true" pageId="4" pageNumber="633">C. momus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1BFFDEBDEF51D1FDF6D0D1" box="[452,527,1423,1440]" pageId="4" pageNumber="633" rank="species">n. comb.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="1156B729DC1BFFDFBE905195FE81D423" blockId="4.[675,1211,1483,1700]" lastBlockId="5.[111,648,216,623]" lastPageId="5" lastPageNumber="634" pageId="4" pageNumber="633">
Male: epistoma glabrous, with feeble median longitudinal gibbosity, regularly punctate on disc, punctation distally more coarse and subrugose; pronotum laterally with few very fine punctures toward front angles, anteriorly with three hornlike apophyses, the lateral ones straight, subparallel, somewhat curved apically only, narrowly rugose from apex almost to base, anteriorly projected from the anterior angles; the central apophysis almost elongate, acuminate, distinctly upturned; aedeagus illustrated on
<figureCitation id="89D2ABACDC1AFFDFBD2E5569FE8AD423" box="[261,371,311,338]" captionStart="FIG" captionStartId="8.[113,124,732,749]" captionTargetBox="[204,1069,228,686]" captionTargetId="figure-20@8.[188,1134,199,724]" captionTargetPageId="8" captionText="FIG. 2. — Aedeagus in dorsal view: A, Chelotrupes hiostius (Gené, 1836) n. comb., Costa Verde, Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; B, C. matutinalis (Baudi, 1870) n. comb.,Costa Verde,Marina di Arbus dint., 39°33’51.0”N, 08°27’48.6”E; C, C. momus (Olivier, 1789) n. comb., Andalusia, Cadiz, Tarifa. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5402561" httpUri="https://zenodo.org/record/5402561/files/figure.png" pageId="5" pageNumber="634">Figure 2A</figureCitation>
.
</paragraph>
<paragraph id="1156B729DC1AFFDFBCA35509FDAAD71E" blockId="5.[111,648,216,623]" pageId="5" pageNumber="634">Female: epistoma shortly pubescent on disc, with median longitudinal gibbosity subtuberculiform at top, confusedly, coarsely, granulosely punctate; pronotum irregularly moderately coarsely punctate on sides, smooth on disc; strongly excavate anteriorly; front angles very distinct, obtusely rounded, anteriorly projected, not outwardly curved; at each side foveolate and with an outer tooth moderately raised; antero-median carina short, distinctly raised.</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1AFFDFBC5A56CAFDD1D016" pageId="5" pageNumber="634" type="discussion">
<paragraph id="1156B729DC1AFFDFBC5A56CAFF2CD7DD" blockId="5.[111,649,660,1383]" box="[113,213,660,686]" pageId="5" pageNumber="634">REMARKS</paragraph>
<paragraph id="1156B729DC1AFFDFBC5A56EDFDD1D016" blockId="5.[111,649,660,1383]" pageId="5" pageNumber="634">
<bibRefCitation id="7578CAD8DC1AFFDFBC5A56EDFE96D7BF" author="HORN W. &amp; KAHLE J." box="[113,367,691,718]" pageId="5" pageNumber="634" pagination="1 - 160" refId="ref11114" refString="HORN W. &amp; KAHLE J. 1935. - Uber entomologische Sammlungen, Entomologen &amp; Entomo-Museologie. (Ein Beitrag zur Geschichte der Entomologie). I Teil. Entomologische Beihefte 2: 1 - 160." type="journal article" year="1935">Horn &amp; Kahle (1935)</bibRefCitation>
as well as
<bibRefCitation id="7578CAD8DC1AFFDFBDC756EDFD7FD7BF" author="CONCI C." box="[492,646,691,718]" pageId="5" pageNumber="634" pagination="817 - 1069" refId="ref10815" refString="CONCI C. 1969. - Repertorio delle biografie e bibliografie degli scrittori e cultori italiani di entomologia. Memorie della Societa Entomologica italiana 48: 817 - 1069." type="journal article" year="1969">Conci (1969)</bibRefCitation>
reported that the Gené’s
<taxonomicName id="D6E9CCAADC1AFFDFBD51568DFE16D79F" authorityName="Linnaeus" authorityYear="1758" box="[378,495,723,750]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="5" pageNumber="634" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
from
<collectingRegion id="D32D79CBDC1AFFDFBE1A568DFD71D79F" box="[561,648,723,750]" country="Italy" name="Sardegna" pageId="5" pageNumber="634">Sardinia</collectingRegion>
were preserved in the Zoological Museum of Turin University then aggregated to the “general collection” of Museo regionale di Scienze naturali of Turin. Checking the
<taxonomicName id="D6E9CCAADC1AFFDFBD29570FFE7CD61D" authorityName="Latreille" authorityYear="1802" box="[258,389,849,876]" class="Insecta" family="Geotrupidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="634" phylum="Arthropoda" rank="family">Geotrupidae</taxonomicName>
material at present stored in this collection, we have not found any specimen of
<taxonomicName id="D6E9CCAADC1AFFDFBC4457CFFED6D6DA" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[111,303,913,939]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="634" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1AFFDFBC4457CFFED6D6DA" box="[111,303,913,939]" italics="true" pageId="5" pageNumber="634">Chelotrupes hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1AFFDFBD1E57CFFE69D6DD" box="[309,400,913,940]" pageId="5" pageNumber="634" rank="species">n. comb.</taxonomicNameLabel>
In the P. Savi’s collection of the Museo di Storia Naturale e del Territorio of Pisa University are present several specimens of Gené’s collection but no specimens of
<taxonomicName id="D6E9CCAADC1AFFDFBD9357AEFDDBD17B" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[440,546,1008,1034]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="634" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1AFFDFBD9357AEFDDBD17B" box="[440,546,1008,1034]" italics="true" pageId="5" pageNumber="634">C. hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1AFFDFBE0357AEFD7ED17A" box="[552,647,1008,1035]" pageId="5" pageNumber="634" rank="species">n. comb.</taxonomicNameLabel>
can be found. Since the Gené type material has to be considered lost, we have designated as
<typeStatus id="CE52098BDC1AFFDFBE0A5071FD8CD13B" box="[545,629,1071,1098]" pageId="5" pageNumber="634" type="neotype">neotype</typeStatus>
a specimen from the Baudi collection. In the original description Gené recorded as type locality “in viis atque in sabuletis frequentissimus, praesertim in peninsula
<emphasis id="239D6B3BDC1AFFDFBCFD50F0FE9FD1B9" box="[214,358,1198,1224]" italics="true" pageId="5" pageNumber="634">di Sant’Antioco</emphasis>
, m. aprili”.Notwithstanding the specimens preserved in Baudi collection bear a label reporting “Sard.” only, we have chosen among them the
<typeStatus id="CE52098BDC1AFFDFBCE45153FED8D059" box="[207,289,1293,1320]" pageId="5" pageNumber="634" type="neotype">neotype</typeStatus>
because they were utilized by Baudi when describing his species
<taxonomicName id="D6E9CCAADC1AFFDFBDA05173FD9DD036" authorityName="Baudi" authorityYear="1870" box="[395,612,1325,1351]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="634" phylum="Arthropoda" rank="species" species="matutinalis">
<emphasis id="239D6B3BDC1AFFDFBDA05173FD9DD036" box="[395,612,1325,1351]" italics="true" pageId="5" pageNumber="634">Geotrupes matutinalis</emphasis>
</taxonomicName>
for discriminating it from the taxon of Gené.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0A52ACCD3685DA6AAC943424614BB598">Revision of the genus Chelotrupes Jekel, 1866 n. stat. (Insecta, Coleoptera, Geotrupidae)</mods:title>
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Genus
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<bibRefCitation id="7578CAD8DC1DFFD8BD565486FE0FD582" author="JEKEL H." box="[381,502,216,243]" pageId="2" pageNumber="631" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">Jekel, 1866</bibRefCitation>
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n. stat.
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<emphasis id="239D6B3BDC1DFFD8BC5A554DFF25D45A" box="[113,220,275,299]" italics="true" pageId="2" pageNumber="631">Chelotrupes</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BCC8554AFE78D45D" author="JEKEL H." box="[227,385,276,300]" pageId="2" pageNumber="631" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">Jekel, 1866: 549</bibRefCitation>
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(as subgenus of
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).
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<taxonomicName id="D6E9CCAADC1DFFD8BC5A5518FE92D42F" authority="" box="[113,363,326,350]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="subGenus" subGenus="Chelotrupes">
<emphasis id="239D6B3BDC1DFFD8BC5A5518FF1DD42F" box="[113,228,326,350]" italics="true" pageId="2" pageNumber="631">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC1DFFD8BCDE5518FE9AD42F" box="[245,355,326,350]" italics="true" pageId="2" pageNumber="631">Chelotrupes</emphasis>
)
</taxonomicName>
–
<treatmentCitation id="90489138DC1DFFD8BDA35518FDDDD42F" author="REITTER E." box="[392,548,326,350]" page="8" pageId="2" pageNumber="631" year="1893">
<bibRefCitation id="7578CAD8DC1DFFD8BDA35518FDDDD42F" author="REITTER E." box="[392,548,326,350]" pageId="2" pageNumber="631" pagination="1 - 109" refId="ref11666" refString="REITTER E. 1893. - Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des naturforschenden Vereines in Brunn 31: 1 - 109 (dated 1892, published 1893)." type="journal article" year="1893">Reitter 1893: 8</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="90489138DC1DFFD8BE795518FF22D409" author="PORTA A." page="401" pageId="2" pageNumber="631" year="1932">
<bibRefCitation id="7578CAD8DC1DFFD8BE795518FF22D409" author="PORTA A." pageId="2" pageNumber="631" refId="ref11528" refString="PORTA A. 1932. - Fauna Coleopterorum italica. Rhyncophora-Lamellicornia. Stabilimento tipografico Piacentino, Piacenza, 5, 476 p." type="book" year="1932">Porta 1932: 401</bibRefCitation>
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.
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<emphasis id="239D6B3BDC1DFFD8BC5A55CCFF38D4DB" box="[113,193,402,426]" italics="true" pageId="2" pageNumber="631">Typhaeus</emphasis>
(
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)
</taxonomicName>
–
<treatmentCitation id="90489138DC1DFFD8BD6555CDFDECD4DA" author="BOUCOMONT A." box="[334,533,403,427]" page="22" pageId="2" pageNumber="631" year="1912">
<bibRefCitation id="7578CAD8DC1DFFD8BD6555CDFDECD4DA" author="BOUCOMONT A." box="[334,533,403,427]" pageId="2" pageNumber="631" refId="ref10730" refString="BOUCOMONT A. 1912. - Scarabaeidae: Taurocerastinae, Geotrupinae. Pars 46. Coleopterorum Catalogus auspiciis et auxilio. W. Junk, Berlin, 47 p." type="book" year="1912">Boucomont 1912: 22</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="90489138DC1DFFD8BE1355CDFF2CD4B4" author="LUIGIONI P." page="390" pageId="2" pageNumber="631" year="1929">
<bibRefCitation id="7578CAD8DC1DFFD8BE1355CDFF2CD4B4" author="LUIGIONI P." pageId="2" pageNumber="631" pagination="1 - 1160" refId="ref11315" refString="LUIGIONI P. 1929. - I Coleotteri d'Italia. Catalogo sinonimico-topografico-bibliografico. Memorie della Pontificia Accademia delle Scienze - I Nuovi Lincei 13 ser. 2: 1 - 1160." type="book chapter" year="1929">Luigioni 1929: 390</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="90489138DC1DFFD8BCD155F3FE38D4B4" author="WINKLER A." box="[250,449,429,453]" page="1036" pageId="2" pageNumber="631" year="1929">
<bibRefCitation id="7578CAD8DC1DFFD8BCD155F3FE38D4B4" author="WINKLER A." box="[250,449,429,453]" pageId="2" pageNumber="631" pagination="1009 - 1136" refId="ref11824" refString="WINKLER A. 1929. - Catalogus Coleopterorum regionis palaearcticae. Scarabaeidae. Albert Winkler, Wien, 9: 1009 - 1136." type="journal article" year="1929">Winkler 1929: 1036</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="90489138DC1DFFD8BDCC55F3FF2AD4AE" author="BAGUENA CORELLA L." page="260" pageId="2" pageNumber="631" year="1967">
<bibRefCitation id="7578CAD8DC1DFFD8BDCC55F3FF2AD4AE" author="BAGUENA CORELLA L." pageId="2" pageNumber="631" refId="ref10502" refString="BAGUENA CORELLA L. 1967. - Scarabaeoidea de la fauna ibero-balear y pirenaica. Consejo Superior de Investigaciones Cientificas, Instituto Espanol de Entomologia, Madrid, 576 p." type="book" year="1967">Báguena Corella 1967: 260</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="90489138DC1DFFD8BCD35599FDEDD4AE" author="CARPANETO G. M. &amp; PIATTELLA E." box="[248,532,455,479]" page="4" pageId="2" pageNumber="631" year="1995">
<bibRefCitation id="7578CAD8DC1DFFD8BCD35599FDEDD4AE" author="CARPANETO G. M. &amp; PIATTELLA E." box="[248,532,455,479]" pageId="2" pageNumber="631" pagination="1 - 18" refId="ref10760" refString="CARPANETO G. M. &amp; PIATTELLA E. 1995. - Coleoptera Polyphaga V (Lucanoidea, Scarabaeoidea), in MINELLI A., RUFFO S. &amp; LA POSTA S. (eds), Checklist delle specie della fauna italiana, 50. Calderini, Bologna: 1 - 18." type="book chapter" year="1995">Carpaneto &amp; Piattella 1995: 4</bibRefCitation>
</treatmentCitation>
. — Martín- Piera &amp; López-Colón 2000: 122.
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SPECIES
</collectionCode>
. —
<taxonomicName id="D6E9CCAADC1DFFD8BD0C564AFD7FD75D" authority="Fabricius, 1792" authorityName="Fabricius" authorityYear="1792" box="[295,646,532,556]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC1DFFD8BD0C564AFE27D75D" box="[295,478,532,556]" italics="true" pageId="2" pageNumber="631">Scarabaeus momus</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BDCC564AFD7FD75D" author="FABRICIUS J. C." box="[487,646,532,556]" pageId="2" pageNumber="631" refId="ref10952" refString="FABRICIUS J. C. 1792. - Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Hafniae, 538 p." type="book" year="1792">Fabricius, 1792</bibRefCitation>
</taxonomicName>
(junior homonym and synonym of
<taxonomicName id="D6E9CCAADC1DFFD8BDFE5670FF0CD710" authority="Olivier, 1789" authorityName="Olivier" authorityYear="1789" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC1DFFD8BDFE5670FD7ED737" box="[469,647,558,582]" italics="true" pageId="2" pageNumber="631">Scarabaeus momus</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BC5A5617FF0CD710" author="OLIVIER A. G." box="[113,245,585,609]" pageId="2" pageNumber="631" refId="ref11485" refString="OLIVIER A. G. 1789. - Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Baudouin, Paris, 1, 190 p." type="book" year="1789">Olivier, 1789</bibRefCitation>
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) (original designation).
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DISTRIBUTION. —
<collectingRegion id="D32D79CBDC1DFFD8BD045625FE7AD7E2" box="[303,387,635,659]" country="Italy" name="Sardegna" pageId="2" pageNumber="631">Sardinia</collectingRegion>
, southwestern Iberian Peninsula
</paragraph>
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<subSubSection id="59F3E4A2DC1DFFD8BC5A5696FB4ED6C5" pageId="2" pageNumber="631" type="diagnosis">
<paragraph id="1156B729DC1DFFD8BC5A5696FCE6D61B" blockId="2.[111,649,712,948]" lastBlockId="2.[675,1211,215,874]" pageId="2" pageNumber="631">DIAGNOSIS. — Species with joined elytral sutural margin (flightless species), length 10.0-22.0 mm, glabrous or almost glabrous, blackish, entirely shiny, sometimes with metallic shine or with elytra dull. Head rhomboidal, strongly narrowed and elongate posteriorly; epistoma with median longitudinal gibbosity more or less elevated and more or less coarsely, diffusedly punctate; clypeus subtriangular with anterior angles rounded, finely margined, the edge ciliate, distally upturned; genal sutures cariniform, usually acutely protruded laterally beyond clypeal margin; genae widely rounded, densely elongately ciliate; front sparsely punctate or smooth; second antennal segment cylindrical, distinctly longer than wide, the club oblong, rather slender, with the median segment as large as the others. Pronotum strongly transverse, moderately convex, at sides with an antero-median rounded foveola more or less deep; sides more or less feebly arcuate, more or less widely flattened, thinly margined, the edge ciliate; hind angles rounded, base margined, not or faintly sinuate. Scutellum triangularly more or less broadly transverse. Elytra short or very short, at most two times longer than the pronotal length; truncate at base, strongly convex, flattened laterally before the epipleural carina; the latter glabrous or almost sparsely ciliate; humeral callus more or less strongly gibbous; striae fine, often obsolete; very superficially, usually almost indistinctly punctate; intervals flat, not punctate, smooth or with feeble transverse rugulae. Fore tibiae denticulate at outer margin; the apical tooth simple. Middle tibiae with four transverse carinae on outer face. Hind tibiae with five transverse carinae on outer face. Sexual dimorphism shown in males mainly by the exceptionally different shape of pronotum and by strongly different punctation of epistoma.</paragraph>
<paragraph id="1156B729DC1DFFD8BE8E57DCFB4ED6C5" blockId="2.[677,1210,898,948]" pageId="2" pageNumber="631">BIONOMICS. — Flightless telocoprid species present all year round and distributed from sea level to high hills.</paragraph>
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<mods:affiliation id="298247315D6C4263DE6B08025A89A559">Centro Interdipartimentale, Museo di Storia Naturale e del Territorio, Università di Pisa, via Roma 79, I- 56011 Calci, Pisa (Italy) dellacasa @ museo. unipi. it</mods:affiliation>
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<mods:namePart id="59E50361844F36F08E74AC76355A2D21">Dellacasa, Giovanni</mods:namePart>
<mods:affiliation id="349A060B836D1AAC8BDC3F363A32F842">CP 921, I- 16121 Genova (Italy)</mods:affiliation>
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<mods:title id="331E35D60734F5CA3FFB0BE091EBB69D">Zoosystema</mods:title>
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<mods:date id="E8F331D04A923E313583520BACE41EF1">2008</mods:date>
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<mods:number id="83492DC91B33FC1B17DC90723D60AA3A">30</mods:number>
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<mods:identifier id="F1A4BB954E4711C2919E68D1A9DDD174" type="DOI">10.5281/zenodo.5402557</mods:identifier>
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Genus
<taxonomicName id="D6E9CCAADC1DFFD8BCD55486FE0FD582" ID-CoL="3MQG" authority="Jekel, 1866" authorityName="Jekel" authorityYear="1866" box="[254,502,216,243]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="genus" status="n. stat.">
<emphasis id="239D6B3BDC1DFFD8BCD55486FE8ED583" box="[254,375,216,242]" italics="true" pageId="2" pageNumber="631">Chelotrupes</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BD565486FE0FD582" author="JEKEL H." box="[381,502,216,243]" pageId="2" pageNumber="631" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">Jekel, 1866</bibRefCitation>
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<taxonomicNameLabel id="38AED640DC1DFFD8BDD65486FDBED582" box="[509,583,216,243]" pageId="2" pageNumber="631">n. stat.</taxonomicNameLabel>
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<paragraph id="1156B729DC1DFFD8BC5A554DFD7FD45D" blockId="2.[113,646,275,300]" box="[113,646,275,300]" pageId="2" pageNumber="631">
<treatmentCitationGroup id="31F99007DC1DFFD8BC5A554DFD7FD45D" box="[113,646,275,300]" pageId="2" pageNumber="631">
<treatmentCitation id="90489138DC1DFFD8BC5A554DFE78D45D" author="JEKEL H." box="[113,385,275,300]" page="549" pageId="2" pageNumber="631" year="1866">
<taxonomicName id="D6E9CCAADC1DFFD8BC5A554DFE78D45D" ID-CoL="3MQG" authority="Jekel, 1866: 549" authorityName="Jekel" authorityPageNumber="549" authorityYear="1866" box="[113,385,275,300]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1DFFD8BC5A554DFF25D45A" box="[113,220,275,299]" italics="true" pageId="2" pageNumber="631">Chelotrupes</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BCC8554AFE78D45D" author="JEKEL H." box="[227,385,276,300]" pageId="2" pageNumber="631" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">Jekel, 1866: 549</bibRefCitation>
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(as subgenus of
<taxonomicName id="D6E9CCAADC1DFFD8BE35554AFD81D45A" ID-CoL="4N3G" authorityName="Latreille" authorityYear="1796" box="[542,632,276,299]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1DFFD8BE35554AFD81D45A" box="[542,632,276,299]" italics="true" pageId="2" pageNumber="631">Geotrupes</emphasis>
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).
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</paragraph>
<paragraph id="1156B729DC1DFFD8BC5A5518FF26D409" blockId="2.[113,646,326,376]" pageId="2" pageNumber="631">
<treatmentCitationGroup id="31F99007DC1DFFD8BC5A5518FF26D409" pageId="2" pageNumber="631">
<taxonomicName id="D6E9CCAADC1DFFD8BC5A5518FE92D42F" authority="" box="[113,363,326,350]" class="Insecta" family="Geotrupidae" genus="Ceratophyus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="subGenus" subGenus="Chelotrupes">
<emphasis id="239D6B3BDC1DFFD8BC5A5518FF1DD42F" box="[113,228,326,350]" italics="true" pageId="2" pageNumber="631">Ceratophyus</emphasis>
(
<emphasis id="239D6B3BDC1DFFD8BCDE5518FE9AD42F" box="[245,355,326,350]" italics="true" pageId="2" pageNumber="631">Chelotrupes</emphasis>
)
</taxonomicName>
–
<treatmentCitation id="90489138DC1DFFD8BDA35518FDDDD42F" author="REITTER E." box="[392,548,326,350]" page="8" pageId="2" pageNumber="631" year="1893">
<bibRefCitation id="7578CAD8DC1DFFD8BDA35518FDDDD42F" author="REITTER E." box="[392,548,326,350]" pageId="2" pageNumber="631" pagination="1 - 109" refId="ref11666" refString="REITTER E. 1893. - Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des naturforschenden Vereines in Brunn 31: 1 - 109 (dated 1892, published 1893)." type="journal article" year="1893">Reitter 1893: 8</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="90489138DC1DFFD8BE795518FF22D409" author="PORTA A." page="401" pageId="2" pageNumber="631" year="1932">
<bibRefCitation id="7578CAD8DC1DFFD8BE795518FF22D409" author="PORTA A." pageId="2" pageNumber="631" refId="ref11528" refString="PORTA A. 1932. - Fauna Coleopterorum italica. Rhyncophora-Lamellicornia. Stabilimento tipografico Piacentino, Piacenza, 5, 476 p." type="book" year="1932">Porta 1932: 401</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="1156B729DC1DFFD8BC5A55CCFE4ED48B" blockId="2.[111,647,402,506]" pageId="2" pageNumber="631">
<treatmentCitationGroup id="31F99007DC1DFFD8BC5A55CCFE4ED48B" pageId="2" pageNumber="631">
<taxonomicName id="D6E9CCAADC1DFFD8BC5A55CCFEC3D4DA" authority="" box="[113,314,402,427]" class="Insecta" family="Geotrupidae" genus="Typhaeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="subGenus" subGenus="Chelotrupes">
<emphasis id="239D6B3BDC1DFFD8BC5A55CCFF38D4DB" box="[113,193,402,426]" italics="true" pageId="2" pageNumber="631">Typhaeus</emphasis>
(
<emphasis id="239D6B3BDC1DFFD8BCE655CCFECAD4DB" box="[205,307,402,426]" italics="true" pageId="2" pageNumber="631">Chelotrupes</emphasis>
)
</taxonomicName>
–
<treatmentCitation id="90489138DC1DFFD8BD6555CDFDECD4DA" author="BOUCOMONT A." box="[334,533,403,427]" page="22" pageId="2" pageNumber="631" year="1912">
<bibRefCitation id="7578CAD8DC1DFFD8BD6555CDFDECD4DA" author="BOUCOMONT A." box="[334,533,403,427]" pageId="2" pageNumber="631" refId="ref10730" refString="BOUCOMONT A. 1912. - Scarabaeidae: Taurocerastinae, Geotrupinae. Pars 46. Coleopterorum Catalogus auspiciis et auxilio. W. Junk, Berlin, 47 p." type="book" year="1912">Boucomont 1912: 22</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="90489138DC1DFFD8BE1355CDFF2CD4B4" author="LUIGIONI P." page="390" pageId="2" pageNumber="631" year="1929">
<bibRefCitation id="7578CAD8DC1DFFD8BE1355CDFF2CD4B4" author="LUIGIONI P." pageId="2" pageNumber="631" pagination="1 - 1160" refId="ref11315" refString="LUIGIONI P. 1929. - I Coleotteri d'Italia. Catalogo sinonimico-topografico-bibliografico. Memorie della Pontificia Accademia delle Scienze - I Nuovi Lincei 13 ser. 2: 1 - 1160." type="book chapter" year="1929">Luigioni 1929: 390</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="90489138DC1DFFD8BCD155F3FE38D4B4" author="WINKLER A." box="[250,449,429,453]" page="1036" pageId="2" pageNumber="631" year="1929">
<bibRefCitation id="7578CAD8DC1DFFD8BCD155F3FE38D4B4" author="WINKLER A." box="[250,449,429,453]" pageId="2" pageNumber="631" pagination="1009 - 1136" refId="ref11824" refString="WINKLER A. 1929. - Catalogus Coleopterorum regionis palaearcticae. Scarabaeidae. Albert Winkler, Wien, 9: 1009 - 1136." type="journal article" year="1929">Winkler 1929: 1036</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="90489138DC1DFFD8BDCC55F3FF2AD4AE" author="BAGUENA CORELLA L." page="260" pageId="2" pageNumber="631" year="1967">
<bibRefCitation id="7578CAD8DC1DFFD8BDCC55F3FF2AD4AE" author="BAGUENA CORELLA L." pageId="2" pageNumber="631" refId="ref10502" refString="BAGUENA CORELLA L. 1967. - Scarabaeoidea de la fauna ibero-balear y pirenaica. Consejo Superior de Investigaciones Cientificas, Instituto Espanol de Entomologia, Madrid, 576 p." type="book" year="1967">Báguena Corella 1967: 260</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="90489138DC1DFFD8BCD35599FDEDD4AE" author="CARPANETO G. M. &amp; PIATTELLA E." box="[248,532,455,479]" page="4" pageId="2" pageNumber="631" year="1995">
<bibRefCitation id="7578CAD8DC1DFFD8BCD35599FDEDD4AE" author="CARPANETO G. M. &amp; PIATTELLA E." box="[248,532,455,479]" pageId="2" pageNumber="631" pagination="1 - 18" refId="ref10760" refString="CARPANETO G. M. &amp; PIATTELLA E. 1995. - Coleoptera Polyphaga V (Lucanoidea, Scarabaeoidea), in MINELLI A., RUFFO S. &amp; LA POSTA S. (eds), Checklist delle specie della fauna italiana, 50. Calderini, Bologna: 1 - 18." type="book chapter" year="1995">Carpaneto &amp; Piattella 1995: 4</bibRefCitation>
</treatmentCitation>
. — Martín-Piera &amp; López-Colón 2000: 122.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1DFFD8BC5A564BFE22D710" pageId="2" pageNumber="631" type="materials_examined">
<paragraph id="1156B729DC1DFFD8BC5A564BFE22D710" blockId="2.[113,647,532,609]" pageId="2" pageNumber="631">
<collectionCode id="77F82FECDC1DFFD8BC5A564BFF0ED75A" box="[113,247,533,556]" pageId="2" pageNumber="631">
<typeStatus id="CE52098BDC1DFFD8BC5A564BFF5DD75B" box="[113,164,533,556]" pageId="2" pageNumber="631">TYPE</typeStatus>
SPECIES
</collectionCode>
. —
<taxonomicName id="D6E9CCAADC1DFFD8BD0C564AFD7FD75D" authority="Fabricius, 1792" authorityName="Fabricius" authorityYear="1792" box="[295,646,532,556]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC1DFFD8BD0C564AFE27D75D" box="[295,478,532,556]" italics="true" pageId="2" pageNumber="631">Scarabaeus momus</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BDCC564AFD7FD75D" author="FABRICIUS J. C." box="[487,646,532,556]" pageId="2" pageNumber="631" refId="ref10952" refString="FABRICIUS J. C. 1792. - Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Hafniae, 538 p." type="book" year="1792">Fabricius, 1792</bibRefCitation>
</taxonomicName>
(junior homonym and synonym of
<taxonomicName id="D6E9CCAADC1DFFD8BDFE5670FF0CD710" authority="Olivier, 1789" authorityName="Olivier" authorityYear="1789" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="momus">
<emphasis id="239D6B3BDC1DFFD8BDFE5670FD7ED737" box="[469,647,558,582]" italics="true" pageId="2" pageNumber="631">Scarabaeus momus</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BC5A5617FF0CD710" author="OLIVIER A. G." box="[113,245,585,609]" pageId="2" pageNumber="631" refId="ref11485" refString="OLIVIER A. G. 1789. - Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Baudouin, Paris, 1, 190 p." type="book" year="1789">Olivier, 1789</bibRefCitation>
</taxonomicName>
) (original designation).
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1DFFD8BC5A5622FF41D7DF" pageId="2" pageNumber="631" type="distribution">
<paragraph id="1156B729DC1DFFD8BC5A5622FF41D7DF" blockId="2.[113,646,635,686]" pageId="2" pageNumber="631">
DISTRIBUTION. —
<collectingRegion id="D32D79CBDC1DFFD8BD045625FE7AD7E2" box="[303,387,635,659]" country="Italy" name="Sardegna" pageId="2" pageNumber="631">Sardinia</collectingRegion>
, southwestern Iberian Peninsula
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1DFFD8BC5A5696FB4ED6C5" pageId="2" pageNumber="631" type="diagnosis">
<paragraph id="1156B729DC1DFFD8BC5A5696FCE6D61B" blockId="2.[111,649,712,948]" lastBlockId="2.[675,1211,215,874]" pageId="2" pageNumber="631">DIAGNOSIS. — Species with joined elytral sutural margin (flightless species), length 10.0-22.0 mm, glabrous or almost glabrous, blackish, entirely shiny, sometimes with metallic shine or with elytra dull. Head rhomboidal, strongly narrowed and elongate posteriorly; epistoma with median longitudinal gibbosity more or less elevated and more or less coarsely, diffusedly punctate; clypeus subtriangular with anterior angles rounded, finely margined, the edge ciliate, distally upturned; genal sutures cariniform, usually acutely protruded laterally beyond clypeal margin; genae widely rounded, densely elongately ciliate; front sparsely punctate or smooth; second antennal segment cylindrical, distinctly longer than wide, the club oblong, rather slender, with the median segment as large as the others. Pronotum strongly transverse, moderately convex, at sides with an antero-median rounded foveola more or less deep; sides more or less feebly arcuate, more or less widely flattened, thinly margined, the edge ciliate; hind angles rounded, base margined, not or faintly sinuate. Scutellum triangularly more or less broadly transverse. Elytra short or very short, at most two times longer than the pronotal length; truncate at base, strongly convex, flattened laterally before the epipleural carina; the latter glabrous or almost sparsely ciliate; humeral callus more or less strongly gibbous; striae fine, often obsolete; very superficially, usually almost indistinctly punctate; intervals flat, not punctate, smooth or with feeble transverse rugulae. Fore tibiae denticulate at outer margin; the apical tooth simple. Middle tibiae with four transverse carinae on outer face. Hind tibiae with five transverse carinae on outer face. Sexual dimorphism shown in males mainly by the exceptionally different shape of pronotum and by strongly different punctation of epistoma.</paragraph>
<paragraph id="1156B729DC1DFFD8BE8E57DCFB4ED6C5" blockId="2.[677,1210,898,948]" pageId="2" pageNumber="631">BIONOMICS. — Flightless telocoprid species present all year round and distributed from sea level to high hills.</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1DFFD8BDA3505FFB96D3D2" pageId="2" pageNumber="631" type="key">
<paragraph id="1156B729DC1DFFD8BDA3505FFC5AD16D" blockId="2.[392,931,1025,1052]" box="[392,931,1025,1052]" pageId="2" pageNumber="631">
KEY TO THE SPECIES OF
<taxonomicName id="D6E9CCAADC1DFFD8BEA2505FFC5AD16D" ID-CoL="3MQG" authority="JEKEL, 1866" authorityName="Jekel" authorityYear="1866" box="[649,931,1025,1052]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1DFFD8BEA2505FFCE3D168" box="[649,794,1025,1051]" italics="true" pageId="2" pageNumber="631">CHELOTRUPES</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BF0A505FFC5AD16D" author="JEKEL H." box="[801,931,1025,1052]" pageId="2" pageNumber="631" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">JEKEL, 1866</bibRefCitation>
</taxonomicName>
</paragraph>
<keyStep id="AA1DAF8CDC1DFFD8BC965068FB97D045" pageId="2" pageNumber="631">
<paragraph id="1156B729DC1DFFD8BC965068FB96D1B0" blockId="2.[189,1135,1078,1332]" pageId="2" pageNumber="631">
<keyLead id="AA18141CDC1DFFD8BC965068FB96D1B0" pageId="2" pageNumber="631">
1. Pronotum, in males, with antero-lateral hornlike apophyses anteriorly projected from above the anterior angles (
<figureCitation id="89D2ABACDC1DFFD8BDD5500DFDB7D11A" box="[510,590,1107,1131]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1A</figureCitation>
); pronotum, in females anteriorly truncate (
<figureCitation id="89D2ABACDC1DFFD8B831500DFB99D11A" box="[1050,1120,1107,1131]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1I</figureCitation>
); epistoma glabrous in both sexes; scutellum subregularly tringular (
<figureCitation id="89D2ABACDC1DFFD8BFA45031FC24D1F6" box="[911,989,1135,1159]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1L</figureCitation>
). Shiny black. Length 10.0-21.0 mm.
<collectingRegion id="D32D79CBDC1DFFD8BDF450D2FD79D1D5" box="[479,640,1164,1188]" country="Spain" name="Illes Baleares" pageId="2" pageNumber="631">Balearic Islands</collectingRegion>
?, southern
<collectingCountry id="69FEF7B9DC1DFFD8BEDE50D2FCC8D1D5" box="[757,817,1164,1188]" name="Spain" pageId="2" pageNumber="631">Spain</collectingCountry>
, central and southern
<collectingCountry id="69FEF7B9DC1DFFD8B83350D2FB97D1D5" box="[1048,1134,1164,1188]" name="Portugal" pageId="2" pageNumber="631">Portugal</collectingCountry>
.......................................................................................................
<taxonomicName id="D6E9CCAADC1DFFD8BF8C50F7FBF5D1B0" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[935,1036,1193,1217]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="momus" status="comb. nov.">
<emphasis id="239D6B3BDC1DFFD8BF8C50F7FBF5D1B0" box="[935,1036,1193,1217]" italics="true" pageId="2" pageNumber="631">C. momus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1DFFD8B83950F7FB96D1B0" box="[1042,1135,1193,1217]" pageId="2" pageNumber="631" rank="species">n. comb.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="1156B729DC1DFFD8BC965098FB97D045" blockId="2.[189,1135,1078,1332]" pageId="2" pageNumber="631">
<keyLead id="AA18141CDC1DFFD8BC965098FB97D045" pageId="2" pageNumber="631">
— Pronotum, in males, with antero-lateral hornlike apophyses projected anteriorly from the anterior angles (
<figureCitation id="89D2ABACDC1DFFD8BDA450BCFDF9D18B" box="[399,512,1250,1274]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1D, E</figureCitation>
), pronotum, in females, anteriorly deeply excavate (
<figureCitation id="89D2ABACDC1DFFD8B83C50BCFEFBD066" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1G, H</figureCitation>
); epistoma glabrous in males, pubescent on disc in females; scutellum broadly transverse (
<figureCitation id="89D2ABACDC1DFFD8BCDB5142FEACD045" box="[240,341,1308,1332]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1J, K</figureCitation>
) ................................................................................................................. 2
</keyLead>
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</keyStep>
<keyStep id="AA1DAF8CDC1DFFD8BC965111FB96D3D2" pageId="2" pageNumber="631">
<paragraph id="1156B729DC1DFFD8BC965111FB96D086" blockId="2.[189,1137,1359,1699]" pageId="2" pageNumber="631">
<keyLead id="AA18141CDC1DFFD8BC965111FB96D086" pageId="2" pageNumber="631">
2. Upper face entirely with metallic shine; first segment of hind tarsi usually as long as the following three combined; pronotum, in males, with antero-lateral hornlike apophyses straight, subparallel, only faintly inwardly curved apically (
<figureCitation id="89D2ABACDC1DFFD8BF4951D7FC4ED0D0" box="[866,951,1417,1441]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1D</figureCitation>
), the central one upturned; pronotum, in females, smooth on disc, moderately irregularly punctate on sides, with anterior angles obtusely rounded and anteriorly projected (
<figureCitation id="89D2ABACDC1DFFD8BF9B519CFBF9D0AB" box="[944,1024,1474,1498]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1G</figureCitation>
).
<location id="1436E1F2DC1DFFD8B838519CFECAD086" LSID="urn:lsid:plazi:treatment:9940063FDC1DFFD8BC9A5486FB96D3D2:1436E1F2DC1DFFD8B838519CFECAD086" name="Blackish. Length" pageId="2" pageNumber="631" stateProvince="Sardegna">Blackish. Length</location>
13.5-26.0 mm.
<collectingRegion id="D32D79CBDC1DFFD8BDF55181FDCBD086" box="[478,562,1503,1527]" country="Italy" name="Sardegna" pageId="2" pageNumber="631">Sardinia</collectingRegion>
....................................................
<taxonomicName id="D6E9CCAADC1DFFD8BF8F5181FBF5D086" ID-CoL="TWR6" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[932,1036,1503,1527]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1DFFD8BF8F5181FBF5D086" box="[932,1036,1503,1527]" italics="true" pageId="2" pageNumber="631">C. hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1DFFD8B8395181FB96D086" box="[1042,1135,1503,1527]" pageId="2" pageNumber="631" rank="species">n. comb.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="1156B729DC1DFFD8BC9651A2FB96D3D2" blockId="2.[189,1137,1359,1699]" pageId="2" pageNumber="631">
<keyLead id="AA18141CDC1DFFD8BC9651A2FB96D3D2" pageId="2" pageNumber="631">
— Pronotum with metallic shine, elytra dull; first segment of hind tarsi usually as long as the following two combined; pronotum, in males, with antero-lateral hornlike apophyses curved and inwardly convergent (
<figureCitation id="89D2ABACDC1DFFD8BE15526BFD73D33C" box="[574,650,1589,1613]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1E</figureCitation>
), the central one straightly anteriorly projected; pronotum, in females, sparsely punctate on disc, coarsely densely punctate on sides, with anterior angles subdenticulate and outwardly protruding (
<figureCitation id="89D2ABACDC1DFFD8BF645230FC5CD3F7" box="[847,933,1646,1670]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1H</figureCitation>
).
<location id="1436E1F2DC1DFFD8BF975230FB94D3F7" LSID="urn:lsid:plazi:treatment:9940063FDC1DFFD8BC9A5486FB96D3D2:1436E1F2DC1DFFD8BF975230FB94D3F7" box="[956,1133,1646,1670]" name="Blackish. Length" pageId="2" pageNumber="631" stateProvince="Sardegna">Blackish. Length</location>
11.0-20.0 mm.
<collectingRegion id="D32D79CBDC1DFFD8BDBB52D5FE1DD3D2" box="[400,484,1675,1699]" country="Italy" name="Sardegna" pageId="2" pageNumber="631">Sardinia</collectingRegion>
..........................................................
<taxonomicName id="D6E9CCAADC1DFFD8BF5752D5FBF2D3D2" ID-CoL="TWR9" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Baudi" baseAuthorityYear="1870" box="[892,1035,1675,1699]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="matutinalis" status="comb. nov.">
<emphasis id="239D6B3BDC1DFFD8BF5752D5FBF2D3D2" box="[892,1035,1675,1699]" italics="true" pageId="2" pageNumber="631">C. matutinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1DFFD8B83952D2FB96D3D2" box="[1042,1135,1675,1699]" pageId="2" pageNumber="631" rank="species">n. comb.</taxonomicNameLabel>
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137
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KEY TO THE SPECIES OF
<taxonomicName id="D6E9CCAADC1DFFD8BEA2505FFC5AD16D" ID-CoL="3MQG" authority="JEKEL, 1866" authorityName="Jekel" authorityYear="1866" box="[649,931,1025,1052]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1DFFD8BEA2505FFCE3D168" box="[649,794,1025,1051]" italics="true" pageId="2" pageNumber="631">CHELOTRUPES</emphasis>
<bibRefCitation id="7578CAD8DC1DFFD8BF0A505FFC5AD16D" author="JEKEL H." box="[801,931,1025,1052]" pageId="2" pageNumber="631" pagination="513 - 618" refId="ref11231" refString="JEKEL H. 1866. - Essai sur la classification naturelle des Geotrupes Latreille et descriptions d'especes nouvelles. Annales de la Societe entomologique de France ser. 4, 5: 513 - 618." type="journal article" year="1866">JEKEL, 1866</bibRefCitation>
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</paragraph>
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<paragraph id="1156B729DC1DFFD8BC965068FB96D1B0" blockId="2.[189,1135,1078,1332]" pageId="2" pageNumber="631">
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1. Pronotum, in males, with antero-lateral hornlike apophyses anteriorly projected from above the anterior angles (
<figureCitation id="89D2ABACDC1DFFD8BDD5500DFDB7D11A" box="[510,590,1107,1131]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1A</figureCitation>
); pronotum, in females anteriorly truncate (
<figureCitation id="89D2ABACDC1DFFD8B831500DFB99D11A" box="[1050,1120,1107,1131]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1I</figureCitation>
); epistoma glabrous in both sexes; scutellum subregularly tringular (
<figureCitation id="89D2ABACDC1DFFD8BFA45031FC24D1F6" box="[911,989,1135,1159]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1L</figureCitation>
). Shiny black. Length 10.0-21.0 mm.
<collectingRegion id="D32D79CBDC1DFFD8BDF450D2FD79D1D5" box="[479,640,1164,1188]" country="Spain" name="Illes Baleares" pageId="2" pageNumber="631">Balearic Islands</collectingRegion>
?, southern
<collectingCountry id="69FEF7B9DC1DFFD8BEDE50D2FCC8D1D5" box="[757,817,1164,1188]" name="Spain" pageId="2" pageNumber="631">Spain</collectingCountry>
, central and southern
<collectingCountry id="69FEF7B9DC1DFFD8B83350D2FB97D1D5" box="[1048,1134,1164,1188]" name="Portugal" pageId="2" pageNumber="631">Portugal</collectingCountry>
.......................................................................................................
<taxonomicName id="D6E9CCAADC1DFFD8BF8C50F7FBF5D1B0" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[935,1036,1193,1217]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="momus" status="comb. nov.">
<emphasis id="239D6B3BDC1DFFD8BF8C50F7FBF5D1B0" box="[935,1036,1193,1217]" italics="true" pageId="2" pageNumber="631">C. momus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1DFFD8B83950F7FB96D1B0" box="[1042,1135,1193,1217]" pageId="2" pageNumber="631" rank="species">n. comb.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="1156B729DC1DFFD8BC965098FB97D045" blockId="2.[189,1135,1078,1332]" pageId="2" pageNumber="631">
<keyLead id="AA18141CDC1DFFD8BC965098FB97D045" pageId="2" pageNumber="631">
— Pronotum, in males, with antero-lateral hornlike apophyses projected anteriorly from the anterior angles (
<figureCitation id="89D2ABACDC1DFFD8BDA450BCFDF9D18B" box="[399,512,1250,1274]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1D, E</figureCitation>
), pronotum, in females, anteriorly deeply excavate (
<figureCitation id="89D2ABACDC1DFFD8B83C50BCFEFBD066" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1G, H</figureCitation>
); epistoma glabrous in males, pubescent on disc in females; scutellum broadly transverse (
<figureCitation id="89D2ABACDC1DFFD8BCDB5142FEACD045" box="[240,341,1308,1332]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1J, K</figureCitation>
) ................................................................................................................. 2
</keyLead>
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<keyStep id="AA1DAF8CDC1DFFD8BC965111FB96D3D2" pageId="2" pageNumber="631">
<paragraph id="1156B729DC1DFFD8BC965111FB96D086" blockId="2.[189,1137,1359,1699]" pageId="2" pageNumber="631">
<keyLead id="AA18141CDC1DFFD8BC965111FB96D086" pageId="2" pageNumber="631">
2. Upper face entirely with metallic shine; first segment of hind tarsi usually as long as the following three combined; pronotum, in males, with antero-lateral hornlike apophyses straight, subparallel, only faintly inwardly curved apically (
<figureCitation id="89D2ABACDC1DFFD8BF4951D7FC4ED0D0" box="[866,951,1417,1441]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1D</figureCitation>
), the central one upturned;
<materialsCitation id="A181BD74DC1DFFD8BD7651F8FB92D086" location="Blackish. Length" pageId="2" pageNumber="631" specimenCount="1" stateProvince="Sardegna">
pronotum, in females, smooth on disc, moderately irregularly punctate on sides, with anterior angles obtusely rounded and anteriorly projected (
<figureCitation id="89D2ABACDC1DFFD8BF9B519CFBF9D0AB" box="[944,1024,1474,1498]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1G</figureCitation>
).
<location id="1436E1F2DC1DFFD8B838519CFECAD086" LSID="urn:lsid:plazi:treatment:9940063FDC1DFFD8BDA3505FFB96D3D2:1436E1F2DC1DFFD8B838519CFECAD086" name="Blackish. Length" pageId="2" pageNumber="631" stateProvince="Sardegna">Blackish. Length</location>
13.5-26.0 mm.
<collectingRegion id="D32D79CBDC1DFFD8BDF55181FDCBD086" box="[478,562,1503,1527]" country="Italy" name="Sardegna" pageId="2" pageNumber="631">Sardinia</collectingRegion>
....................................................
<taxonomicName id="D6E9CCAADC1DFFD8BF8F5181FBF5D086" ID-CoL="TWR6" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Gene" baseAuthorityYear="1836" box="[932,1036,1503,1527]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="hiostius" status="comb. nov.">
<emphasis id="239D6B3BDC1DFFD8BF8F5181FBF5D086" box="[932,1036,1503,1527]" italics="true" pageId="2" pageNumber="631">C. hiostius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1DFFD8B8395181FB92D086" box="[1042,1131,1503,1527]" pageId="2" pageNumber="631" rank="species">n. comb</taxonomicNameLabel>
</materialsCitation>
.
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<paragraph id="1156B729DC1DFFD8BC9651A2FB96D3D2" blockId="2.[189,1137,1359,1699]" pageId="2" pageNumber="631">
<keyLead id="AA18141CDC1DFFD8BC9651A2FB96D3D2" pageId="2" pageNumber="631">
— Pronotum with metallic shine, elytra dull; first segment of hind tarsi usually as long as the following two combined; pronotum, in males, with antero-lateral hornlike apophyses curved and inwardly convergent (
<figureCitation id="89D2ABACDC1DFFD8BE15526BFD73D33C" box="[574,650,1589,1613]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1E</figureCitation>
), the central one straightly anteriorly projected;
<materialsCitation id="A181BD74DC1DFFD8BCC1520CFB92D3D2" location="Blackish. Length" pageId="2" pageNumber="631" specimenCount="1" stateProvince="Sardegna">
pronotum, in females, sparsely punctate on disc, coarsely densely punctate on sides, with anterior angles subdenticulate and outwardly protruding (
<figureCitation id="89D2ABACDC1DFFD8BF645230FC5CD3F7" box="[847,933,1646,1670]" captionStart="FIG" captionStartId="4.[113,124,1355,1372]" captionTargetBox="[182,1142,216,1290]" captionTargetId="figure-20@4.[162,1161,206,1315]" captionTargetPageId="4" captionText="FIG. 1. — A-C, heads; A, Chelotrupes hiostius (Gené, 1836) n. comb.; B, C. matutinalis (Baudi, 1870) n. comb.; C, C. momus (Olivier, 1789) n. comb.; D-F, pronotum of the male; D, C. hiostius n. comb.; E, C. matutinalis n. comb.; F, C. momus n. comb.; G-I, pronotum of the female; G, C. hiostius n. comb.; H, C. matutinalis n. comb.; I, C. momus n. comb.; J-L, elytra; J, C. hiostius n. comb.; K, C. matutinalis n. comb.; L, C. momus n. comb." figureDoi="http://doi.org/10.5281/zenodo.5402559" httpUri="https://zenodo.org/record/5402559/files/figure.png" pageId="2" pageNumber="631">Fig. 1H</figureCitation>
).
<location id="1436E1F2DC1DFFD8BF975230FB94D3F7" LSID="urn:lsid:plazi:treatment:9940063FDC1DFFD8BDA3505FFB96D3D2:1436E1F2DC1DFFD8BF975230FB94D3F7" box="[956,1133,1646,1670]" name="Blackish. Length" pageId="2" pageNumber="631" stateProvince="Sardegna">Blackish. Length</location>
11.0-20.0 mm.
<collectingRegion id="D32D79CBDC1DFFD8BDBB52D5FE1DD3D2" box="[400,484,1675,1699]" country="Italy" name="Sardegna" pageId="2" pageNumber="631">Sardinia</collectingRegion>
..........................................................
<taxonomicName id="D6E9CCAADC1DFFD8BF5752D5FBF2D3D2" ID-CoL="TWR9" authorityName="Dellacasa &amp; Dellacasa" authorityYear="2008" baseAuthorityName="Baudi" baseAuthorityYear="1870" box="[892,1035,1675,1699]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="631" phylum="Arthropoda" rank="species" species="matutinalis" status="comb. nov.">
<emphasis id="239D6B3BDC1DFFD8BF5752D5FBF2D3D2" box="[892,1035,1675,1699]" italics="true" pageId="2" pageNumber="631">C. matutinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38AED640DC1DFFD8B83952D2FB92D3D2" box="[1042,1131,1675,1699]" pageId="2" pageNumber="631" rank="species">n. comb</taxonomicNameLabel>
</materialsCitation>
.
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97
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<mods:date id="2D00861BCB17A70688249F0823FF6ADF">2008</mods:date>
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<mods:number id="58544897F63ADD56493392A269467B90">2008-02-21</mods:number>
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<mods:number id="5E9C5DDC5B4ED2EA9C3DB1C31776667E">30</mods:number>
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<mods:number id="6133791A2CA2ED2C800E566CC6EA1191">3</mods:number>
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<mods:identifier id="FEA058A71577F7124B00FDCC75FB1864" type="DOI">10.5281/zenodo.5402557</mods:identifier>
<mods:identifier id="C26FECE916A67EF69BF7641BB6D3D266" type="ISSN">1638-9387</mods:identifier>
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<treatment id="9940063FDC1EFFDBBEE65090FB97D3D2" LSID="urn:lsid:plazi:treatment:9940063FDC1EFFDBBEE65090FB97D3D2" httpUri="http://treatment.plazi.org/id/9940063FDC1EFFDBBEE65090FB97D3D2" lastPageNumber="630" pageId="1" pageNumber="630">
<subSubSection id="59F3E4A2DC1EFFDBBEE65090FB68D198" box="[717,1169,1230,1257]" pageId="1" pageNumber="630" type="nomenclature">
<paragraph id="1156B729DC1EFFDBBEE65090FB68D198" blockId="1.[717,1169,1230,1257]" box="[717,1169,1230,1257]" pageId="1" pageNumber="630">
<heading id="4A1E0045DC1EFFDBBEE65090FB68D198" box="[717,1169,1230,1257]" centered="true" fontSize="36" level="2" pageId="1" pageNumber="630" reason="6">
Tribe
<taxonomicName id="D6E9CCAADC1EFFDBBF275091FB68D198" ID-CoL="8RYSL" authority="Zunino, 1984" authorityName="Zunino" authorityYear="1984" box="[780,1169,1230,1257]" class="Insecta" family="Geotrupidae" genus="Chromogeotrupini" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="630" phylum="Arthropoda" rank="genus" tribe="Chromogeotrupini">CHROMOGEOTRUPINI Zunino, 1984</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="59F3E4A2DC1EFFDBBE8E5153FBF5D016" pageId="1" pageNumber="630" type="description">
<paragraph id="1156B729DC1EFFDBBE8E5153FBF5D016" blockId="1.[677,1210,1293,1383]" pageId="1" pageNumber="630">
In the western Palaearctic fauna, only two genera belong to
<taxonomicName id="D6E9CCAADC1EFFDBBF325173FB43D039" authority="(Lobl et al. 2006)" baseAuthorityName="Lobl" baseAuthorityYear="2006" box="[793,1210,1325,1352]" class="Insecta" family="Geotrupidae" genus="Chromogeotrupini" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="630" phylum="Arthropoda" rank="genus">
Chromogeotrupini (
<bibRefCitation id="7578CAD8DC1EFFDBBFD05173FB48D039" author="LOBL I. &amp; NIKOLAJEV G. V. &amp; KRAL D." box="[1019,1201,1325,1352]" pageId="1" pageNumber="630" pagination="84 - 92" refId="ref11267" refString="LOBL I., NIKOLAJEV G. V. &amp; KRAL D. 2006. - Geotrupinae, in LOBL I. &amp; SMETANA A. (eds), Catalogue of Palaearctic Coleoptera, Vol. 3. Apollo Books, Stenstrup: 84 - 92." type="book chapter" year="2006">
Löbl
<emphasis id="239D6B3BDC1EFFDBB8135173FB96D036" box="[1080,1135,1325,1351]" italics="true" pageId="1" pageNumber="630">et al.</emphasis>
2006
</bibRefCitation>
)
</taxonomicName>
and they can be keyed as follows.
</paragraph>
</subSubSection>
<subSubSection id="59F3E4A2DC1EFFDBBCF851D3FB97D3D2" pageId="1" pageNumber="630" type="key">
<paragraph id="1156B729DC1EFFDBBCF851D3FBA1D0D9" blockId="1.[211,1112,1421,1448]" box="[211,1112,1421,1448]" pageId="1" pageNumber="630">
KEY TO THE GENERA OF WESTERN PALEARCTIC
<taxonomicName id="D6E9CCAADC1EFFDBBEE151D0FBA1D0D9" authority="ZUNINO, 1984" authorityName="Zunino" authorityYear="1984" box="[714,1112,1421,1448]" class="Insecta" family="Geotrupidae" genus="Chromogeotrupini" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="630" phylum="Arthropoda" rank="genus" tribe="Chromogeotrupini">CHROMOGEOTRUPINI ZUNINO, 1984</taxonomicName>
</paragraph>
<key id="6578E018DC1EFFDBBC96519CFB97D3D2" pageId="1" pageNumber="630">
<paragraph id="1156B729DC1EFFDBBC96519CFB97D341" blockId="1.[189,1136,1474,1699]" pageId="1" pageNumber="630">
<keyLead id="AA18141CDC1EFFDBBC96519CFB97D341" pageId="1" pageNumber="630">
— Second segment of antennae globous, slightly longer than wide; head, in males, feebly narrowed and feebly elongate posteriorly; base of pronotum strongly sinuate; scutellum triangularly elongate; elytra not fused (winged species) with striae deep and intervals convex ............................................................................................................
<taxonomicName id="D6E9CCAADC1EFFDBB83F5246FB97D341" authorityName="Leach" authorityYear="1815" box="[1044,1134,1560,1584]" class="Insecta" family="Geotrupidae" genus="Typhaeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="630" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1EFFDBB83F5246FB97D341" box="[1044,1134,1560,1584]" italics="true" pageId="1" pageNumber="630">Typhaeus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1156B729DC1EFFDBBC96526BFB97D3D2" blockId="1.[189,1136,1474,1699]" pageId="1" pageNumber="630">
<keyLead id="AA18141CDC1EFFDBBC96526BFB97D3D2" pageId="1" pageNumber="630">
— Second segment of antennae cylindrical, distinctly longer than wide; head, in males, strongly narrowed and very elongate posteriorly; base of pronotum not or, at most, faintly sinuate; scutellum triangularly transverse; elytra fused (flightless species) with striae fine, sometimes obsolete, and intervals flat .........................................................
<taxonomicName id="D6E9CCAADC1EFFDBBFD052D5FB97D3D2" authorityName="Jekel" authorityYear="1866" box="[1019,1134,1675,1699]" class="Insecta" family="Geotrupidae" genus="Chelotrupes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="630" phylum="Arthropoda" rank="genus">
<emphasis id="239D6B3BDC1EFFDBBFD052D5FB97D3D2" box="[1019,1134,1675,1699]" italics="true" pageId="1" pageNumber="630">Chelotrupes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</key>
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