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<mods:title>Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:title id="D131E9B67E86CEF9797398E681D2CC56">Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="D1BA20022EFE6DD25C4CA6CA52EA657F">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart>Stoev, Pavel</mods:namePart>
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<mods:date>2011</mods:date>
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<mods:number>5</mods:number>
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<mods:date id="C032AF473D4E6F278261AF4411B2A804">2011</mods:date>
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<treatment ID-GBIF-Taxon="152016718" LSID="urn:lsid:zoobank.org:act:EFBC2841-A6A2-4BED-93D4-E1EBCAADF757" httpUri="http://treatment.plazi.org/id/E29AE4FCE1F0E8A46ADF53973398BF25" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<subSubSection pageId="5" pageNumber="6" type="nomenclature">
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<paragraph pageId="5" pageNumber="6">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:EFBC2841-A6A2-4BED-93D4-E1EBCAADF757" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma maculatum" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maculatum">Eustrongylosoma maculatum</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="6">sp. n.</taxonomicNameLabel>
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<treatment id="E29AE4FCE1F0E8A46ADF53973398BF25" ID-GBIF-Taxon="152016718" LSID="urn:lsid:zoobank.org:act:EFBC2841-A6A2-4BED-93D4-E1EBCAADF757" httpUri="http://treatment.plazi.org/id/E29AE4FCE1F0E8A46ADF53973398BF25" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
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<subSubSection id="1529BEBDE70F014770E19C6E6C0832FB" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="F30298850C1D3E4B964C03B38ECF7A40" pageId="5" pageNumber="6">
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<taxonomicName id="F13F9E01D058D99F5FCDA5D69DEA8028" ID-CoL="3DBGS" LSID="urn:lsid:zoobank.org:act:EFBC2841-A6A2-4BED-93D4-E1EBCAADF757" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma maculatum" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maculatum">Eustrongylosoma maculatum</taxonomicName>
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<taxonomicNameLabel id="19DB607DE3FC3AED8BC15D2D743533A1" pageId="5" pageNumber="6">sp. n.</taxonomicNameLabel>
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Figs 15-19
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="type material">
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<paragraph pageId="5" pageNumber="6">Type material.</paragraph>
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<paragraph pageId="5" pageNumber="6">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin area, September 1975, leg. B.S.E.</paragraph>
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<subSubSection id="407F176297F92C8E58B1EBAC57AA06D5" pageId="5" pageNumber="6" type="type material">
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<paragraph id="6997F759951EA2F63C3EFD5D6C75B4D7" pageId="5" pageNumber="6">Type material.</paragraph>
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<paragraph id="BFF00CFCA4BC517304CC623129ECF3A5" pageId="5" pageNumber="6">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin area, September 1975, leg. B.S.E.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="name">
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<paragraph pageId="5" pageNumber="6">Name.</paragraph>
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<paragraph pageId="5" pageNumber="6">To reflect the wide spots/bands in the front halves of the collum, following metaterga and epiproct.</paragraph>
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<subSubSection id="06C33D0971120F142C4F1A7136AC8932" pageId="5" pageNumber="6" type="name">
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<paragraph id="9F9DCE2D3E7699C8140D7AF59B49A591" pageId="5" pageNumber="6">Name.</paragraph>
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<paragraph id="1D364AEE8CBB9ED0F7D517FC2FA558F0" pageId="5" pageNumber="6">To reflect the wide spots/bands in the front halves of the collum, following metaterga and epiproct.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="diagnosis">
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<paragraph pageId="5" pageNumber="6">Diagnosis.</paragraph>
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<paragraph pageId="5" pageNumber="6">Distinguished by the very poorly developed paraterga, coupled with a non-fringed tip of the solenophore (see also Key below).</paragraph>
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<subSubSection id="884DE8A515C305885DAE43875804C268" pageId="5" pageNumber="6" type="diagnosis">
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<paragraph id="BB63E089DE4FB97D06F22C5866CED4A6" pageId="5" pageNumber="6">Diagnosis.</paragraph>
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<paragraph id="7A7B67D34DF970E1A7E3EA793807CD5E" pageId="5" pageNumber="6">Distinguished by the very poorly developed paraterga, coupled with a non-fringed tip of the solenophore (see also Key below).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
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<paragraph pageId="5" pageNumber="6">Description.</paragraph>
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<paragraph pageId="5" pageNumber="6">Length ca 24 mm, width of pro- and metazona 2.0 and 2.4 mm, respectively. Coloration light yellow-brown with front halves of collum, following metaterga and epiproct, as well as entire head and antennae contrasting brown, thus creating a clear cingulate pattern.</paragraph>
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<paragraph lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<subSubSection id="2BC633F483682C4EEAB2065EDB7DBC9C" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
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<paragraph id="5640B7DC59B46CD682FC479A62A8B8E4" pageId="5" pageNumber="6">Description.</paragraph>
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<paragraph id="4B4D875CD4055D7556B8D823273E9D26" pageId="5" pageNumber="6">Length ca 24 mm, width of pro- and metazona 2.0 and 2.4 mm, respectively. Coloration light yellow-brown with front halves of collum, following metaterga and epiproct, as well as entire head and antennae contrasting brown, thus creating a clear cingulate pattern.</paragraph>
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<paragraph id="A03B8D95E45D6CCE019809321F7A6D86" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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In width, head> collum = segment 3 = 4 <2 <5-16; thereafter body gently tapering towards telson. Head densely setose, only vertex bare. Antennae rather short, clavate, reaching beyond segment 2 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining. Paraterga of postcollum segments very poorly developed (Fig. 15), set low (at about 1/2-1/3 midbody height); paratergum 2 a low ridge with a few wrinkles above; paraterga 3 and 4 virtually missing, traceable only as 3-4 similar wrinkles; following paraterga retained only as a dorsal sulcus/line (Fig. 15), barely vi
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<pageBreakToken pageId="6" pageNumber="7" start="start">sible</pageBreakToken>
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<pageBreakToken id="5CC4037FC29C143C37E546AF151BBFFB" pageId="6" pageNumber="7" start="start">sible</pageBreakToken>
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in segment 19. Ozopores lateral, superficial. Tergal setae mostly abraded, retained ones about 1/4 the length of metaterga; setation pattern vague, sometimes traceable as 1+1 paramedian insertion points in front row. Axial line wanting. Transverse sulcus on metaterga highly superficial, incomplete, far from reaching bases of paraterga, visible only in metaterga 9-17. Stricture between pro- and metazona rather distinctly beaded. Pleurosternal carinae very small, complete crests or nearly so only in segments 2-4. Epiproct (Fig. 16) short, flattened dorsoventrally, digitiform; apical papillae very large, spiniform, directed more caudally than ventrally; subapical papillae missing. Hypoproct subtriangular, apex rounded, setigerous paramedian knobs at caudal margin small and rather faintly separated.
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</paragraph>
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<paragraph pageId="6" pageNumber="7">Sterna densely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 17). Legs relatively long, midbody ones ca 1.5-1.6 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal brushes increasingly well reduced towards legs of segment 10.</paragraph>
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<paragraph pageId="6" pageNumber="7">Gonopods as in Figs 18 & 19. Distal part of solenophore split into a distinct frontomesal tooth (k) and a longer, somewhat curved lateral branch (j) with two subapical denticles and a lobule. Femorite with a large, rounded, apicolateral lobe (l).</paragraph>
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<caption pageId="6" pageNumber="7">
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<paragraph pageId="6" pageNumber="7">
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<paragraph id="037E537F48B8C939CDC60639D411C5C0" pageId="6" pageNumber="7">Sterna densely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 17). Legs relatively long, midbody ones ca 1.5-1.6 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal brushes increasingly well reduced towards legs of segment 10.</paragraph>
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<paragraph id="D93F571E0922561B6803BEB9A5E6D295" pageId="6" pageNumber="7">Gonopods as in Figs 18 & 19. Distal part of solenophore split into a distinct frontomesal tooth (k) and a longer, somewhat curved lateral branch (j) with two subapical denticles and a lobule. Femorite with a large, rounded, apicolateral lobe (l).</paragraph>
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<caption id="10779BBF146A17221B363592AED07EDC" pageId="6" pageNumber="7">
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<paragraph id="BD7F9B80A310D8DA7E71AE5F0184C185" pageId="6" pageNumber="7">
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Figures 15-19.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma maculatum" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maculatum">Eustrongylosoma maculatum</taxonomicName>
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<taxonomicName id="B8D8171A188754CFF65ECA467EB19ECC" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma maculatum" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maculatum">Eustrongylosoma maculatum</taxonomicName>
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sp. n., holotype. 15 segment 10, lateral view 16 tip of epiproct, dorsal view 17 sternal lobe between coxae 4, caudal view 18, 19 left gonopod, mesal and lateral views, respectively. Scale bars: 1.0 (15-17) and 0.5 mm (18 & 19).
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</paragraph>
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</caption>
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