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data/00/AB/C6/00ABC664036EBCB69FFF6AEA22919165.xml
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<document ID-DOI="http://dx.doi.org/10.3897/ijm.5.2090" ID-GBIF-Dataset="67fa3f59-684b-4cc3-9a57-3436ea681205" ID-GBIF-Taxon="152016727" ID-Pensoft-Pub="1875-2543-5-1" LSID="urn:lsid:zoobank.org:act:7656859C-40A3-449C-BA2C-125C0C5C5CC7" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1875-2543-5-1" ModsDocOrigin="International Journal of Myriapodology 5" ModsDocTitle="Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451252547380" checkinUser="pensoft" docAuthor="Golovatch, Sergei I. &amp; Stoev, Pavel" docDate="2011" docId="00ABC664036EBCB69FFF6AEA22919165" docLanguage="en" docName="InternJourMyriapod 5: 1-26" docOrigin="International Journal of Myriapodology 5" docSource="http://dx.doi.org/10.3897/ijm.5.2090" docTitle="Eustrongylosoma liklik Golovatch &amp; Stoev, 2011, sp. n." docType="treatment" docUuid="7656859C-40A3-449C-BA2C-125C0C5C5CC7" docUuidSource="ZooBank" docVersion="3" lastPageId="14" lastPageNumber="15" masterDocId="724E36204F7968779912FFD2FFC3892C" masterDocTitle="Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="26" masterPageNumber="1" pageId="12" pageNumber="13" updateTime="1643719993040" updateUser="ExternalLinkService">
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<mods:title>Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:title id="B485F0FDE015055EBA9B663754C9D09E">Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:namePart>Stoev, Pavel</mods:namePart>
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<treatment ID-GBIF-Taxon="152016727" LSID="urn:lsid:zoobank.org:act:7656859C-40A3-449C-BA2C-125C0C5C5CC7" httpUri="http://treatment.plazi.org/id/00ABC664036EBCB69FFF6AEA22919165" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection pageId="12" pageNumber="13" type="nomenclature">
<paragraph pageId="12" pageNumber="13">
<taxonomicName LSID="urn:lsid:zoobank.org:act:7656859C-40A3-449C-BA2C-125C0C5C5CC7" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma liklik" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="liklik">Eustrongylosoma liklik</taxonomicName>
<taxonomicNameLabel pageId="12" pageNumber="13">sp. n.</taxonomicNameLabel>
<treatment id="00ABC664036EBCB69FFF6AEA22919165" ID-GBIF-Taxon="152016727" LSID="urn:lsid:zoobank.org:act:7656859C-40A3-449C-BA2C-125C0C5C5CC7" httpUri="http://treatment.plazi.org/id/00ABC664036EBCB69FFF6AEA22919165" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
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<paragraph id="F2DB93CB037DF071EC7BBAEF75F5C45D" pageId="12" pageNumber="13">
<taxonomicName id="39BC8F751273D38D296EBCBCE6A47CEC" ID-CoL="3DBGN" LSID="urn:lsid:zoobank.org:act:7656859C-40A3-449C-BA2C-125C0C5C5CC7" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma liklik" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="liklik">Eustrongylosoma liklik</taxonomicName>
<taxonomicNameLabel id="9CAF79A3B5AA5645068679DFCE9B9F3B" pageId="12" pageNumber="13">sp. n.</taxonomicNameLabel>
Figs 41-47
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="type material">
<paragraph pageId="12" pageNumber="13">Type material.</paragraph>
<paragraph pageId="12" pageNumber="13">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin, 1700 m, 19 October 1975, B.S.E. (leg. P. Beron).</paragraph>
<subSubSection id="3F820DF67250CC9EF6E4A720A022990E" pageId="12" pageNumber="13" type="type material">
<paragraph id="0DF0675E9569069134D58DA15A513D55" pageId="12" pageNumber="13">Type material.</paragraph>
<paragraph id="3D3BB3782A98F38666D80FB69B470F4B" pageId="12" pageNumber="13">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin, 1700 m, 19 October 1975, B.S.E. (leg. P. Beron).</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="name">
<paragraph pageId="12" pageNumber="13">Name.</paragraph>
<paragraph pageId="12" pageNumber="13">
<normalizedToken originalValue="“liklik”">&quot;liklik&quot;</normalizedToken>
<subSubSection id="4DBE26C0B5B17D53AD49EE983F37677D" pageId="12" pageNumber="13" type="name">
<paragraph id="DDD271BE0CA71079CEA0697B8F449A8B" pageId="12" pageNumber="13">Name.</paragraph>
<paragraph id="40A91D47C0A7E8AE3228123932249AC7" pageId="12" pageNumber="13">
<normalizedToken id="00CFB041A88B0A6001EBC9451379EBA6" originalValue="“liklik”">&quot;liklik&quot;</normalizedToken>
means
<normalizedToken originalValue="“small”">&quot;small&quot;</normalizedToken>
<normalizedToken id="52C1C96F9C553AF41447B65B0094CC93" originalValue="“small”">&quot;small&quot;</normalizedToken>
or
<normalizedToken originalValue="“little”">&quot;little&quot;</normalizedToken>
<normalizedToken id="C88953A08CEC3AA0BF5D89B1ABAF0E61" originalValue="“little”">&quot;little&quot;</normalizedToken>
in Pidgin English. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="diagnosis">
<paragraph pageId="12" pageNumber="13">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="13">Distinguished by the small size, coupled with the short legs, the poorly developed paraterga and the contrasting infuscate distal antenno- and podomeres (see also Key below).</paragraph>
<subSubSection id="BE1BBAD44B68DEC22E236490B363719C" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="A06615ED75BAB4D76E65ECFCB6682DD1" pageId="12" pageNumber="13">Diagnosis.</paragraph>
<paragraph id="74E5D3A4AC361DED78650C162A2A239F" pageId="12" pageNumber="13">Distinguished by the small size, coupled with the short legs, the poorly developed paraterga and the contrasting infuscate distal antenno- and podomeres (see also Key below).</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph pageId="12" pageNumber="13">Description.</paragraph>
<paragraph pageId="12" pageNumber="13">Length ca 15 mm, width of pro- and metazona 1.2 and 1.4 mm, respectively. Coloration uniformly light yellowish, with distal antennomeres, as well as tibiae and, especially, tarsi contrasting dark brown.</paragraph>
<paragraph lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="24B92D283733FDBA91C2A39812949B97" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="CA65B26C8AE69DE78E3439E9BCA74D62" pageId="12" pageNumber="13">Description.</paragraph>
<paragraph id="696141DF24BAD58E49FE614072464350" pageId="12" pageNumber="13">Length ca 15 mm, width of pro- and metazona 1.2 and 1.4 mm, respectively. Coloration uniformly light yellowish, with distal antennomeres, as well as tibiae and, especially, tarsi contrasting dark brown.</paragraph>
<paragraph id="EC9DA23BAAD76018D5CB6F5E02B903BC" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
In width, head &lt;collum = segments 3 = 4 &lt;2 &lt;5-16. Head densely setose, only vertex bare. Antennae rather short, slightly clavate, reaching beyond segment 2 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate; rear halves of metaterga sometimes faintly rugulose. Paraterga of postcollum segments poorly developed (Figs 41, 42), especially weak in poreless segments, set low (at about 1/2 midbody height), sometimes with only a faint undulation in front of ozopore near caudal third, slightly more evident in segments 2 and 3, but even paraterga 2 and 3 with both anterolateral and caudal corners rounded. Caudal corners of postcollum paraterga never extended beyond rear tergal margin. Lateral calluses of paraterga always narrow, delimited by a sulcus dorsally, in most of pore-bearing segments ventrally in caudal 1/4 as well. Ozopores lateral, superficial. Tergal setae fully abraded, setation pattern vague. Axial line wanting. Transverse sulcus on metaterga rather superficial, incomplete, far from reaching bases of paraterga, visible on metaterga 5-17, barely traceable on 18th. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, complete in segments 2-4, a caudal denticle retained also in segments 5-7, thereafter entirely missing. Epiproct (Fig. 43) short, flattened dorsoventrally, digitiform; apical papillae small, tip narrowly subtrun
<pageBreakToken pageId="13" pageNumber="14" start="start">cate</pageBreakToken>
<pageBreakToken id="10FF6B6742E3E06D98FFD2D8B4ED1D48" pageId="13" pageNumber="14" start="start">cate</pageBreakToken>
; subapical papillae very small, well removed from tip. Hypoproct (Fig. 44) subtrapeziform, setigerous paramedian knobs at caudal margin well-separated and very small.
</paragraph>
<paragraph pageId="13" pageNumber="14">Sterna rather sparsely setose; a rather high, linguiform, roundly subtriangular, ventral, setose lamina only between coxae 4 (Fig. 45). Legs short, midbody ones ca 1.0-1.1 times longer than body height, apparently somewhat incrassate and longer compared to ♀; ♂evident tarsal brushes increasingly well reduced towards legs of segment 19.</paragraph>
<paragraph pageId="13" pageNumber="14">Gonopods as in Figs 46 &amp; 47. Distal half of solenophore split into a lateral lobe (k) and a narrowly bifid larger branch (j). Femorite with a large, rounded, apicolateral lobe (l).</paragraph>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
<pageBreakToken pageId="14" pageNumber="15" start="start">Figures</pageBreakToken>
<paragraph id="20CA2B6B54F44D77EF8392F8785872E6" pageId="13" pageNumber="14">Sterna rather sparsely setose; a rather high, linguiform, roundly subtriangular, ventral, setose lamina only between coxae 4 (Fig. 45). Legs short, midbody ones ca 1.0-1.1 times longer than body height, apparently somewhat incrassate and longer compared to ♀; ♂evident tarsal brushes increasingly well reduced towards legs of segment 19.</paragraph>
<paragraph id="A504F47B2FB7C4F507FBC731699DE4BA" pageId="13" pageNumber="14">Gonopods as in Figs 46 &amp; 47. Distal half of solenophore split into a lateral lobe (k) and a narrowly bifid larger branch (j). Femorite with a large, rounded, apicolateral lobe (l).</paragraph>
<caption id="71C9C11642C601AB077A1327FFE132D1" pageId="14" pageNumber="15">
<paragraph id="B2965B37E43DF1DCB7A1744677D036A8" pageId="14" pageNumber="15">
<pageBreakToken id="DB85B398D6762B21443CEDFC515ED089" pageId="14" pageNumber="15" start="start">Figures</pageBreakToken>
41-47.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma liklik" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="liklik">Eustrongylosoma liklik</taxonomicName>
<taxonomicName id="A47FD6110305388DCD46E171F53425B9" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma liklik" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="liklik">Eustrongylosoma liklik</taxonomicName>
sp. n., holotype. 41 segment 10, lateral view 42 right half of segment 10, dorsal view 43 tip of epiproct, dorsal view 44 hypoproct, ventral view 45 sternal lobe between coxae 4, caudal view 46, 47 right gonopod, submesal and sublateral views, respectively. Scale bars: 1.0 (41-45) and 0.25 mm (46, 47).
</paragraph>
</caption>

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<mods:namePart>Stoev, Pavel</mods:namePart>
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<mods:date>2011</mods:date>
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<taxonomicName LSID="urn:lsid:zoobank.org:act:A7917E87-A72C-4F9A-BC2E-BB70C6CE24CD" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma masalai" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="masalai">Eustrongylosoma masalai</taxonomicName>
<taxonomicNameLabel pageId="8" pageNumber="9">sp. n.</taxonomicNameLabel>
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<paragraph id="5948FCBE9974D836216FA408C1D7B06B" pageId="8" pageNumber="9">
<taxonomicName id="2B35E0E9EEE2AE4E52BD6F02E222013F" ID-CoL="3DBGT" LSID="urn:lsid:zoobank.org:act:A7917E87-A72C-4F9A-BC2E-BB70C6CE24CD" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma masalai" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="masalai">Eustrongylosoma masalai</taxonomicName>
<taxonomicNameLabel id="344E745726CFAA1BC9C1D06D546F9FDC" pageId="8" pageNumber="9">sp. n.</taxonomicNameLabel>
Figs 27-33
</paragraph>
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<paragraph pageId="8" pageNumber="9">Type material.</paragraph>
<paragraph pageId="8" pageNumber="9">Holotype♂(NMNHS),Papua New Guinea, Western Province, Finim Tel area, October 1975, B.S.E. (leg. P. Beron). - Paratypes: 2 ♀(NMNHS), same locality, together with holotype.</paragraph>
<subSubSection id="5A99E7FEE140F9041B9F22209C79ADB0" pageId="8" pageNumber="9" type="type material">
<paragraph id="C6C1D6957BA01B3A35E6F0B9338B9B9D" pageId="8" pageNumber="9">Type material.</paragraph>
<paragraph id="F02A21F6EE903C7FBEEB903CE1FD8FDE" pageId="8" pageNumber="9">Holotype♂(NMNHS),Papua New Guinea, Western Province, Finim Tel area, October 1975, B.S.E. (leg. P. Beron). - Paratypes: 2 ♀(NMNHS), same locality, together with holotype.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="name">
<paragraph pageId="8" pageNumber="9">Name.</paragraph>
<paragraph pageId="8" pageNumber="9">
<normalizedToken originalValue="“masalai”">&quot;masalai&quot;</normalizedToken>
<subSubSection id="E3584231FC17F9E13066481203313040" pageId="8" pageNumber="9" type="name">
<paragraph id="2059C586459AD93BF8428DDF6981592B" pageId="8" pageNumber="9">Name.</paragraph>
<paragraph id="2F2BF96D02FEE70A99D0F7FF55E2F18B" pageId="8" pageNumber="9">
<normalizedToken id="1241DA9B3F92FC2ED4BAD25959507CD5" originalValue="“masalai”">&quot;masalai&quot;</normalizedToken>
means a
<normalizedToken originalValue="“brownie”">&quot;brownie&quot;</normalizedToken>
<normalizedToken id="6A6E86321BF13DC6AA0A06465ED87A15" originalValue="“brownie”">&quot;brownie&quot;</normalizedToken>
in Pidgin English. A noun in apposition.
</paragraph>
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<subSubSection pageId="8" pageNumber="9" type="diagnosis">
<paragraph pageId="8" pageNumber="9">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="9">Distinguished by the rather narrow paratergal calluses, coupled with a missing colour pattern and a deeply split distal part of the solenophore (see also Key below).</paragraph>
<subSubSection id="4F0E29B21006B79CD47CF48BD833B879" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="E5CD22AA44DE488A14ED91F12D4B29CF" pageId="8" pageNumber="9">Diagnosis.</paragraph>
<paragraph id="93FDE3774EE07F8EB59EE629EFB1D67E" pageId="8" pageNumber="9">Distinguished by the rather narrow paratergal calluses, coupled with a missing colour pattern and a deeply split distal part of the solenophore (see also Key below).</paragraph>
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<paragraph pageId="8" pageNumber="9">Description.</paragraph>
<paragraph pageId="8" pageNumber="9">Length ca 18 (holotype) or 20 mm (paratypes), width of pro- and metazona 1.3 and 1.8 (holotype) or 1.7 and 2.0 mm (paratypes), respectively. Coloration uniformly castaneous brown to light brown; antennae light yellow-brown, legs yellow.</paragraph>
<paragraph lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="132FAD647E8943A9D3EAE176300CCDC0" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="4312DD4E3FA36D9CBE8999C0CA4EF41F" pageId="8" pageNumber="9">Description.</paragraph>
<paragraph id="0A8F3C2C30A40CBC42845B7B8C83654D" pageId="8" pageNumber="9">Length ca 18 (holotype) or 20 mm (paratypes), width of pro- and metazona 1.3 and 1.8 (holotype) or 1.7 and 2.0 mm (paratypes), respectively. Coloration uniformly castaneous brown to light brown; antennae light yellow-brown, legs yellow.</paragraph>
<paragraph id="F1D8B3A00E40BD0E5B9E0D2E04633DCE" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
In width, head&gt; collum = segments 5-17 &lt;2 &lt;3 = 4. Head densely setose, only vertex with 1+1 setae. Antennae rather short, slightly clavate, reaching beyond (♂) or end of segment 2 (♀) dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate; rear halves of metaterga sometimes faintly rugulose. Paraterga of postcollum segments moderately well developed (Figs 27, 28), set low (at about 1/3 midbody height), subhorizontal, evidently thicker in pore-bearing segments than in poreless ones; paratergum 2 with a small anterolateral denticle, its front margin straight; following paraterga without denticulations at lateral margin. Caudal corner of all postcollum paraterga rounded, slightly extended beyond rear tergal margin only in segments 2 and 17-19, better so in 18th. Lateral calluses of paraterga narrow, a little broader in pore-bearing segments, each delimited by a sulcus both dorsally and, in caudal 1/3, ventrally. Ozopores lateral, lying inside an ovoid groove. Tergal setae fully abraded, setation pattern untraceable. Axial line wanting. Transverse sulcus on metaterga rather superficial, far from reaching bases of paraterga, visible on metaterga 5-17, barely traceable also on 18th. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, com
<pageBreakToken pageId="9" pageNumber="10" start="start">plete</pageBreakToken>
<pageBreakToken id="9A6293A4E8310BC20D2EB2800893697D" pageId="9" pageNumber="10" start="start">plete</pageBreakToken>
or nearly so in segments 2-5, a sharp or rounded caudal denticle retained also in segments 6 and 7, thereafter entirely missing. Epiproct (Fig. 29) short, flattened dorsoventrally, digitiform; apical papillae rather large, tip narrowly emarginate; subapical papillae missing. Hypoproct (Fig. 30) subtrapeziform, setigerous paramedian knobs at caudal margin well-separated and very small.
</paragraph>
<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<paragraph id="79E5D643542C26FE686E1BE4529553DD" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Sterna rather sparsely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 31). Legs rather long and slender, mi
<pageBreakToken pageId="10" pageNumber="11" start="start">dbody</pageBreakToken>
<pageBreakToken id="7D3C0E72941855FB053E0FD5BE739746" pageId="10" pageNumber="11" start="start">dbody</pageBreakToken>
ones ca 1.5-1.6 (♂) or 1.1-1.2 (♀)times longer than body height, in ♂slightly incrassate compared to ♀; ♂tarsal brushes increasingly well reduced towards legs of segment 14.
</paragraph>
<paragraph pageId="10" pageNumber="11">Gonopods as in Figs 32 &amp; 33. Tip (j) of a slender and coiled solenophore rather deeply biramous, larger branch bearing four apical or subapical denticles. Apicolateral lobe (l) of femorite relatively small.</paragraph>
<caption pageId="10" pageNumber="11">
<paragraph pageId="10" pageNumber="11">
<paragraph id="7846EEF1E467F678725F7AA1051D07DD" pageId="10" pageNumber="11">Gonopods as in Figs 32 &amp; 33. Tip (j) of a slender and coiled solenophore rather deeply biramous, larger branch bearing four apical or subapical denticles. Apicolateral lobe (l) of femorite relatively small.</paragraph>
<caption id="87B9964467B4FC83621871D40675B76A" pageId="10" pageNumber="11">
<paragraph id="C9E50CC564873191985A1828F818CEF7" pageId="10" pageNumber="11">
Figures 27-33.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma masalai" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="masalai">Eustrongylosoma masalai</taxonomicName>
<taxonomicName id="136CA1C2CA59AFEDDF53AF190DD05B4F" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma masalai" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="masalai">Eustrongylosoma masalai</taxonomicName>
sp. n., holotype. 27 segment 10, lateral view 28 left half of segment 10, dorsal view 29 tip of epiproct, dorsal view 30 hypoproct, ventral view 31 sternal lobe between coxae 4, caudal view 32, 33 right gonopod, mesal and subventral views, respectively. Scale bars: 1.0 (27-31) and 0.3 mm (32 &amp; 33).
</paragraph>
</caption>

80
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<taxonomicName LSID="http://species-id.net/wiki/Titanoeca_sharmai" authority="Bastawade, 2008" authorityName="Bastawade" authorityYear="2008" class="Arachnida" family="Titanoecidae" genus="Titanoeca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanoeca sharmai" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="sharmai">Titanoeca sharmai (Bastawade, 2008)</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="62">comb.n.</taxonomicNameLabel>
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Bastawade, 2008: 40, f. 1-12 (♂♀).
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<paragraph pageId="7" pageNumber="62">Comments.</paragraph>
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<paragraph id="86F68F9FC3350ECB0322E30F05D03BB0" pageId="7" pageNumber="62">Comments.</paragraph>
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This species was described from the northeastern region of Himachal Pradesh, India (Fig. 17) on the basis of both sexes, but the figures and description are of rather poorly quality. The figures provided by the author, namely the tibial and
<pageBreakToken pageId="8" pageNumber="63" start="start">metatarsal</pageBreakToken>
<pageBreakToken id="20D5FCD0DB48BE66F448A59AB54E25A7" pageId="8" pageNumber="63" start="start">metatarsal</pageBreakToken>
spines on the legs in males and the structure of the palp, leaves no doubt that the species belongs in
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. Although there are three titanoecid genera in India, judging from the locality, high elevation, and the unmodified male palpal patella,
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must be placed in
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<taxonomicName id="73EFC73CDB2F0A0D13FF8FF160DC7C95" class="Arachnida" family="Titanoecidae" genus="Titanoeca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanoeca" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="genus">Titanoeca</taxonomicName>
. It is worth mentioning that this species may be a junior synonym of
<taxonomicName class="Arachnida" family="Titanoecidae" genus="Titanoeca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanoeca intermedia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="intermedia">Titanoeca intermedia</taxonomicName>
<taxonomicName id="43AE283BB13D004ACECE913B081FD3C2" class="Arachnida" family="Titanoecidae" genus="Titanoeca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanoeca intermedia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="intermedia">Titanoeca intermedia</taxonomicName>
Caporiacco, 1934 (species incorrectly synonymized with
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<taxonomicName id="7FFF5933B34F6A79496156599BF958FE" class="Arachnida" family="Titanoecidae" genus="Titanoeca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanoeca flavicoma" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="flavicoma">Titanoeca flavicoma</taxonomicName>
L. Koch, 1872), which was described from territories now belonging to northeastern Pakistan and from northern India (Jammu &amp; Kashmir).
</paragraph>
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126
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<taxonomicName LSID="http://species-id.net/wiki/Anuvinda_milloti" authority="Hubert, 1973" authorityName="Hubert" authorityYear="1973" class="Arachnida" family="Titanoecidae" genus="Anuvinda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anuvinda milloti" order="Araneae" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="milloti">Anuvinda milloti (Hubert, 1973)</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="61">comb. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="046694826F2378942C972F9B7F97B683" pageId="6" pageNumber="61">comb. n.</taxonomicNameLabel>
</paragraph>
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<paragraph pageId="6" pageNumber="61">
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Hubert, 1973a: 676, f. 1-6 (♂♀).
</paragraph>
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<paragraph pageId="6" pageNumber="61">Comments.</paragraph>
<paragraph lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61">
<subSubSection id="58D750F1FA515F99F53C583DB972F8A0" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61" type="comments">
<paragraph id="025AF6F5B27030691CC94E3D53C5CA27" pageId="6" pageNumber="61">Comments.</paragraph>
<paragraph id="2FC156AF66D952A07B3DE44227D7033A" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61">
This species is perfectly described from central and eastern Nepal (Fig. 17). Judging from the structure of the male palp, and particularly the modified patella, it undoubtedly belongs to
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Lehntinen, 1967, the type species of which,
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<pageBreakToken pageId="7" pageNumber="62" start="start">(</pageBreakToken>
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<pageBreakToken id="2874828B5A973D414806DF2FA6406FC5" pageId="7" pageNumber="62" start="start">(</pageBreakToken>
Reimoser, 1934) was recently well redescribed on the basis of both sexes by
<bibRefCitation author="Almeida-Silva, LM" journalOrPublisher="Zootaxa" pageId="8" pageNumber="63" pagination="61 - 68" title="On the poorly known genus Anuvinda Lehtinen, 1967 (Araneae: Titanoecidae)." volume="2266" year="2009">Almeida-Silva et al. (2009)</bibRefCitation>
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. Judging from the diagnosis and figures of the copulatory organs of
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<taxonomicName id="DEAA4854045C5B20BFA9BD9A3EB8AE18" class="Arachnida" family="Titanoecidae" genus="Anuvinda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anuvinda escheri" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="escheri">Anuvinda escheri</taxonomicName>
(Reimoser, 1934), it is very likely that the two names should be synonymized. An additional argument which supports their probable synonymy is the distribution of both species.
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<taxonomicName id="6DE61664127F4C92737347024584E68F" class="Arachnida" family="Titanoecidae" genus="Anuvinda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anuvinda escheri" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="escheri">Anuvinda escheri</taxonomicName>
is known from central India (type locality), Thailand, Laos and southern China (Yunnan) and
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<taxonomicName id="8AF880B05F63DB64A9D5D2FE49EAF7D0" class="Arachnida" family="Titanoecidae" genus="Anuvinda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anuvinda milloti" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="milloti">Anuvinda milloti</taxonomicName>
has been recorded from several localities in central and eastern Nepal.
</paragraph>
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<paragraph id="BFE627B347B6EDFD274C951A16A0CC4F" pageId="7" pageNumber="62">
Figure 17. Distribution of
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sp. n. (square), four species transferred here into
<taxonomicName family="Titanoecidae" lsidName="" pageId="7" pageNumber="62" rank="family">Titanoecidae</taxonomicName>
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:
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1
<taxonomicName class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava nathabhaii" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="nathabhaii">Pandava nathabhaii</taxonomicName>
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2
<taxonomicName class="Arachnida" family="Titanoecidae" genus="Titanoeca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanoeca sharmai" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="sharmai">Titanoeca sharmai</taxonomicName>
<taxonomicName id="CC661FAE1C2A6BC8EA69620BC0D35D56" class="Arachnida" family="Titanoecidae" genus="Titanoeca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Titanoeca sharmai" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="sharmai">Titanoeca sharmai</taxonomicName>
3
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<taxonomicName id="C0342AB3194563FAE1616C99A4BB13DC" class="Arachnida" family="Titanoecidae" genus="Anuvinda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anuvinda milloti" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="milloti">Anuvinda milloti</taxonomicName>
4 and four
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<taxonomicName id="8824D95917BF1F5B418B09CD21B5BF61" class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">Pandava</taxonomicName>
species recently described from India:
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<taxonomicName id="9646739B5F7A54976CDECBCCDC9FA700" class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava shiva" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="shiva">Pandava shiva</taxonomicName>
5
<taxonomicName class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava ganga" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="ganga">Pandava ganga</taxonomicName>
<taxonomicName id="D587BB8E6C0982E90AF579AEB007B337" class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava ganga" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="ganga">Pandava ganga</taxonomicName>
6
<taxonomicName class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava kama" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="kama">Pandava kama</taxonomicName>
<taxonomicName id="B6DDF077E11C8278F41394FB4214FD4F" class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava kama" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="kama">Pandava kama</taxonomicName>
7 and
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<taxonomicName id="1B62F9F2B7232A0C239975BB25214A9A" class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava ganesha" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="ganesha">Pandava ganesha</taxonomicName>
8.
</paragraph>
</caption>

130
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<treatment ID-GBIF-Taxon="152016717" LSID="urn:lsid:plazi:treatment:219B7BC69DA2A66E98F3220E93D14A05" httpUri="http://treatment.plazi.org/id/219B7BC69DA2A66E98F3220E93D14A05" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<paragraph pageId="2" pageNumber="3">
<treatment id="219B7BC69DA2A66E98F3220E93D14A05" ID-GBIF-Taxon="152016717" LSID="urn:lsid:plazi:treatment:219B7BC69DA2A66E98F3220E93D14A05" httpUri="http://treatment.plazi.org/id/219B7BC69DA2A66E98F3220E93D14A05" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
<subSubSection id="C7925082F0D838FCF735E24A86A76715" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="A16D259C68C80363C45EFB1289CC443F" pageId="2" pageNumber="3">
The genus
<taxonomicName authority="Silvestri, 1896" authorityName="Silvestri" authorityYear="1896" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Eustrongylosoma Silvestri, 1896</taxonomicName>
<taxonomicName id="6B1F0C9A2E76297D57DF72D2B9CEF176" ID-CoL="7PFJC" authority="Silvestri, 1896" authorityName="Silvestri" authorityYear="1896" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Eustrongylosoma Silvestri, 1896</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="diagnosis">
<paragraph pageId="2" pageNumber="3">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="3">
<subSubSection id="1557251D05D3916016E2FA7DF8345863" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="3C3CF6F0196F2F7957D7C773764D2803" pageId="2" pageNumber="3">Diagnosis.</paragraph>
<paragraph id="D48EAF58D563F1384113B4BAF4014B6A" pageId="2" pageNumber="3">
Small to medium-sized (12-33 mm long)
<taxonomicName lsidName="" pageId="2" pageNumber="3" rank="tribe" tribe="Eustrongylosomatini">Eustrongylosomatini</taxonomicName>
<taxonomicName id="0A7CC739291FF1B3536D657888490C11" lsidName="" pageId="2" pageNumber="3" rank="tribe" tribe="Eustrongylosomatini">Eustrongylosomatini</taxonomicName>
with 20 segments and a normal ozopore formula. Paraterga medium-sized to missing, never as high as dorsum. Tergal setae usually few, mostly abraded and traceable only as insertion points, metaterga only rarely abundantly setose. ♂ legs without adenostyles, but usually with tarsal (sometimes also distotibial) brushes. A sternal lobe or two cones between ♂ coxae 4.
</paragraph>
<paragraph pageId="2" pageNumber="3">Gonopod subcylindrical, setose distodorsally, rather long, but considerably shorter than telopodite; prefemoral (= densely setose) part at least twice shorter than femorite; the latter without evidence of torsion (= seminal groove running entirely mesally), but with an evident dorsomesal groove (only rarely transformed apically into a small projection) delimited by longitudinal folds or crests; distal part of femorite projected into a distinct lateral lobe or process (l) and extended considerably beyond beginning of a long and flagelliform solenomere; solenophore about as long as to only slightly shorter than femorite, coiled, directed laterad, with lamina lateralis produced basad before subtending solenomere and evidently more strongly developed than lamina medialis, both laminae being elaborate.</paragraph>
<paragraph id="58779B0D15D2A5547DA7EDF2BCDFF171" pageId="2" pageNumber="3">Gonopod subcylindrical, setose distodorsally, rather long, but considerably shorter than telopodite; prefemoral (= densely setose) part at least twice shorter than femorite; the latter without evidence of torsion (= seminal groove running entirely mesally), but with an evident dorsomesal groove (only rarely transformed apically into a small projection) delimited by longitudinal folds or crests; distal part of femorite projected into a distinct lateral lobe or process (l) and extended considerably beyond beginning of a long and flagelliform solenomere; solenophore about as long as to only slightly shorter than femorite, coiled, directed laterad, with lamina lateralis produced basad before subtending solenomere and evidently more strongly developed than lamina medialis, both laminae being elaborate.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="type-species">
<paragraph pageId="2" pageNumber="3">Type-species.</paragraph>
<paragraph pageId="2" pageNumber="3">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma fasciatum" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="fasciatum">Strongylosoma fasciatum</taxonomicName>
<subSubSection id="F461F7977F166A1EE3F6F327C4E8E8C8" pageId="2" pageNumber="3" type="type-species">
<paragraph id="22A97FABCCDDEE42D7F83207F75339C0" pageId="2" pageNumber="3">Type-species.</paragraph>
<paragraph id="D46758101387AA09062A7B36E1EF24DF" pageId="2" pageNumber="3">
<taxonomicName id="3560C66FB731EA161DC7706E8A3648D5" class="Diplopoda" family="Paradoxosomatidae" genus="Strongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Strongylosoma fasciatum" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="fasciatum">Strongylosoma fasciatum</taxonomicName>
Silvestri, 1895, by original designation by
<bibRefCitation pageId="2" pageNumber="3">Silvestri (1896)</bibRefCitation>
<bibRefCitation id="0F08B058BF3926F77821906CB0C42B94" pageId="2" pageNumber="3">Silvestri (1896)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="remarks">
<paragraph pageId="2" pageNumber="3">Remarks.</paragraph>
<paragraph lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="256F27614EF3A70C846B5E722C898442" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="remarks">
<paragraph id="F3266F4CF893A7C1FCFBB261B2B12301" pageId="2" pageNumber="3">Remarks.</paragraph>
<paragraph id="691F2B4FDF4C8B1D2AFA7E65A0389385" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Above is a slightly amended diagnosis of
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<taxonomicName id="97729F68DDA1C6A6BD8E97339A37D77F" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Eustrongylosoma</taxonomicName>
which seems to be especially topical because a number of
<taxonomicName lsidName="" pageId="2" pageNumber="3" rank="tribe" tribe="Eustrongylosomatini">Eustrongylosomatini</taxonomicName>
<taxonomicName id="642332C624F09098F9724BE019E7D5E2" lsidName="" pageId="2" pageNumber="3" rank="tribe" tribe="Eustrongylosomatini">Eustrongylosomatini</taxonomicName>
species from New
<pageBreakToken pageId="3" pageNumber="4" start="start">Guinea</pageBreakToken>
<pageBreakToken id="B4D7A0480B24886D4609DA4539525BF6" pageId="3" pageNumber="4" start="start">Guinea</pageBreakToken>
have recently been revised and/or reallocated, and a new genus proposed (
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Myriapod Memoranda" pageId="18" pageNumber="19" pagination="75 - 82" title="Records of Paradoxosomatidae from New Guinea (Diplopoda, Polydesmida)." volume="11" year="2009">Jeekel 2009</bibRefCitation>
<bibRefCitation id="B07C820BFBACD7C05CE10CFD55BCCD40" author="Jeekel, CAW" journalOrPublisher="Myriapod Memoranda" pageId="18" pageNumber="19" pagination="75 - 82" title="Records of Paradoxosomatidae from New Guinea (Diplopoda, Polydesmida)." volume="11" year="2009">Jeekel 2009</bibRefCitation>
).
</paragraph>
<paragraph pageId="3" pageNumber="4">
<paragraph id="FDE62519C097887CA0A68FF55CC2DF3E" pageId="3" pageNumber="4">
In addition to
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Eustrongylosoma</taxonomicName>
<taxonomicName id="2464D21698EE0D656093C88DD927FDFD" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Eustrongylosoma</taxonomicName>
, this tribe currently comprises the following five genera from New Guinea:
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Nothrosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothrosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Nothrosoma</taxonomicName>
<taxonomicName id="733FF81BD13B39F10D4D3E56145B3676" class="Diplopoda" family="Paradoxosomatidae" genus="Nothrosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothrosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Nothrosoma</taxonomicName>
Attems, 1929 (three species),
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Perittotresis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perittotresis" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Perittotresis</taxonomicName>
<taxonomicName id="153AE68CD34D267BA43BE156E1B0C3CE" class="Diplopoda" family="Paradoxosomatidae" genus="Perittotresis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perittotresis" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Perittotresis</taxonomicName>
Attems, 1914,
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Astromontosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Astromontosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Astromontosoma</taxonomicName>
<taxonomicName id="34F90317E222BDBFD645BAEDE8499C5E" class="Diplopoda" family="Paradoxosomatidae" genus="Astromontosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Astromontosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Astromontosoma</taxonomicName>
Hoffman, 1978,
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Selminosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selminosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Selminosoma</taxonomicName>
<taxonomicName id="576AA8B9F716F0B6AF1AD64980BC9B96" class="Diplopoda" family="Paradoxosomatidae" genus="Selminosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selminosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Selminosoma</taxonomicName>
Hoffman, 1978 (all three monobasic),
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Silvattia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silvattia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Silvattia</taxonomicName>
<taxonomicName id="51101CDB7537CDC2C1D66D6A7B5F25B2" class="Diplopoda" family="Paradoxosomatidae" genus="Silvattia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silvattia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Silvattia</taxonomicName>
Jeekel, 2009 (two species), as well as one more from the Seychelles:
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Diglossosternoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diglossosternoides" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Diglossosternoides</taxonomicName>
<taxonomicName id="7B584788A7D6F827202DA7C1645A6241" class="Diplopoda" family="Paradoxosomatidae" genus="Diglossosternoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diglossosternoides" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Diglossosternoides</taxonomicName>
Golovatch &amp;
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<normalizedToken id="CA89533187C0E9DA0DCC2999CB65BE2A" originalValue="Korsós">Korsos</normalizedToken>
, 1992 (monobasic) (
<bibRefCitation author="Hoffman, RL" journalOrPublisher="International Journal of Speleology" pageId="18" pageNumber="19" pagination="281 - 307" title="Diplopoda from Papuan caves (Zoological Results of the British Speleological Expedition to Papua New Guinea, 1975, 4)." volume="9" year="1977 / 1978">Hoffman 1977/1978</bibRefCitation>
<bibRefCitation id="0ECB1BB041493CE2A0FB775F1A5B6A84" author="Hoffman, RL" journalOrPublisher="International Journal of Speleology" pageId="18" pageNumber="19" pagination="281 - 307" title="Diplopoda from Papuan caves (Zoological Results of the British Speleological Expedition to Papua New Guinea, 1975, 4)." volume="9" year="1977 / 1978">Hoffman 1977/1978</bibRefCitation>
;
<bibRefCitation author="Golovatch, SI" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="18" pageNumber="19" pagination="1 - 31" title="Diplopoda collected by the Soviet Zoological Expedition to the Seychelles Islands in 1984." volume="38" year="1992">
<bibRefCitation id="C749DC824EE141CF3B5561B8CD0D089C" author="Golovatch, SI" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="18" pageNumber="19" pagination="1 - 31" title="Diplopoda collected by the Soviet Zoological Expedition to the Seychelles Islands in 1984." volume="38" year="1992">
Golovatch and
<normalizedToken originalValue="Korsós">Korsos</normalizedToken>
<normalizedToken id="A20F05859EC09EC39F87D3BD4F9AE812" originalValue="Korsós">Korsos</normalizedToken>
1992
</bibRefCitation>
;
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Myriapod Memoranda" pageId="18" pageNumber="19" pagination="75 - 82" title="Records of Paradoxosomatidae from New Guinea (Diplopoda, Polydesmida)." volume="11" year="2009">Jeekel 2009</bibRefCitation>
<bibRefCitation id="3728ACDA57158C257B0540FDB837E494" author="Jeekel, CAW" journalOrPublisher="Myriapod Memoranda" pageId="18" pageNumber="19" pagination="75 - 82" title="Records of Paradoxosomatidae from New Guinea (Diplopoda, Polydesmida)." volume="11" year="2009">Jeekel 2009</bibRefCitation>
). Most of them have long been keyed (
<bibRefCitation author="Hoffman, RL" journalOrPublisher="International Journal of Speleology" pageId="18" pageNumber="19" pagination="281 - 307" title="Diplopoda from Papuan caves (Zoological Results of the British Speleological Expedition to Papua New Guinea, 1975, 4)." volume="9" year="1977 / 1978">Hoffman 1977/1978</bibRefCitation>
<bibRefCitation id="6E5015C396E05B4153804B0630963C33" author="Hoffman, RL" journalOrPublisher="International Journal of Speleology" pageId="18" pageNumber="19" pagination="281 - 307" title="Diplopoda from Papuan caves (Zoological Results of the British Speleological Expedition to Papua New Guinea, 1975, 4)." volume="9" year="1977 / 1978">Hoffman 1977/1978</bibRefCitation>
). Basically,
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Eustrongylosoma</taxonomicName>
<taxonomicName id="DA46710D935214CDD64AEAD0D76C82AF" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Eustrongylosoma</taxonomicName>
is distinguished from all of them by the presence of a distinct distolateral lobe or process on a medially excavate gonopod femorite, combined with the absence of any evident outgrowths just before the beginning of a long solenomere.
</paragraph>
</subSubSection>

112
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<taxonomicName LSID="urn:lsid:zoobank.org:act:41B50F67-A72E-40CC-A304-FC871ADEE9DF" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma papua" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="papua">Eustrongylosoma papua</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="4">sp. n.</taxonomicNameLabel>
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<taxonomicName id="4DF85C938812EF4FC1F79C50CDA84715" ID-CoL="3DBGZ" LSID="urn:lsid:zoobank.org:act:41B50F67-A72E-40CC-A304-FC871ADEE9DF" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma papua" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="papua">Eustrongylosoma papua</taxonomicName>
<taxonomicNameLabel id="D3E505C616B120BF9EBDFF6F8648EF77" pageId="3" pageNumber="4">sp. n.</taxonomicNameLabel>
Figs 1-6
</paragraph>
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<paragraph pageId="3" pageNumber="4">Type material.</paragraph>
<paragraph pageId="3" pageNumber="4">Holotype♂(NMNHS),Papua New Guinea, Western Province, Finim Tel Plateau, near Cave Kumsop Tem, 16 August 1975, B.S.E. (leg. Ph. Chapman &amp; P. Beron). - Paratypes: 1 ♀(NMNHS), same locality, together with holotype. 2 ♂ (fragmented, one with only head and segments 1-8, the other lacking segments 8-12) (NMNHS), 1 ♂ (ZMUM), West Sepik Province, Telefomin area, 1700 m, September 1975, B.S.E. (leg. P. Beron).</paragraph>
<subSubSection id="96826B633E8A63F9D75D0D8358995C68" pageId="3" pageNumber="4" type="type material">
<paragraph id="AD90769A083EEEB08A5A5EE084753F61" pageId="3" pageNumber="4">Type material.</paragraph>
<paragraph id="3203C188076B4920576C5CBBB6F0B5C3" pageId="3" pageNumber="4">Holotype♂(NMNHS),Papua New Guinea, Western Province, Finim Tel Plateau, near Cave Kumsop Tem, 16 August 1975, B.S.E. (leg. Ph. Chapman &amp; P. Beron). - Paratypes: 1 ♀(NMNHS), same locality, together with holotype. 2 ♂ (fragmented, one with only head and segments 1-8, the other lacking segments 8-12) (NMNHS), 1 ♂ (ZMUM), West Sepik Province, Telefomin area, 1700 m, September 1975, B.S.E. (leg. P. Beron).</paragraph>
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<subSubSection pageId="3" pageNumber="4" type="name">
<paragraph pageId="3" pageNumber="4">Name.</paragraph>
<paragraph pageId="3" pageNumber="4">
<subSubSection id="3CD86EC99DB033737CD8268ADCBAE17A" pageId="3" pageNumber="4" type="name">
<paragraph id="D1784220E41789798B2EC49DD0885B47" pageId="3" pageNumber="4">Name.</paragraph>
<paragraph id="6C15803F03B0CAB4C77E7046AB2BFFD8" pageId="3" pageNumber="4">
Emphasizes the abundant setae on the collum and following metaterga, to reflect
<normalizedToken originalValue="“papua”">&quot;papua&quot;</normalizedToken>
<normalizedToken id="0C9288E68A591A3DCFE77548D48DA33F" originalValue="“papua”">&quot;papua&quot;</normalizedToken>
which means
<normalizedToken originalValue="“curly-headed”">&quot;curly-headed&quot;</normalizedToken>
<normalizedToken id="06C50E4624878405775022EB97925963" originalValue="“curly-headed”">&quot;curly-headed&quot;</normalizedToken>
in Tok Pisin (= Pidgin English, Neomelanesian). A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="diagnosis">
<paragraph pageId="3" pageNumber="4">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="4">
<subSubSection id="18F08296326A03C4C0FC7E9E8A5974B4" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="39A22C44C9131614DF1EBAD2E33301FB" pageId="3" pageNumber="4">Diagnosis.</paragraph>
<paragraph id="5D50AD1027949AEBED2A3D14B5759DFC" pageId="3" pageNumber="4">
Due toabundanttergal setae, comes closest tothe partly sympatric
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma beroni" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="beroni">Eustrongylosoma beroni</taxonomicName>
<taxonomicName id="D0CE3F7C5C63DEC7E83B0B0D346047A0" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma beroni" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="beroni">Eustrongylosoma beroni</taxonomicName>
Golovatch &amp; Stoev, 2009, but differs in the semi-circular hypoproct, the more elaborate solenophore tip and the stouter sternal lobe between ♂ coxae 4 (see also Key below).
</paragraph>
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<paragraph pageId="3" pageNumber="4">Description.</paragraph>
<paragraph pageId="3" pageNumber="4">Length of both holo- and complete paratypes ca 20-21 mm, width of pro- and metazona 1.4-1.6 and 1.7-1.9 mm (♂), or 1.9 and 2.2 mm (♀), respectively. Holotype ca 20 mm long, 1.6 and 1.9 mm wide on midbody pro- and metazona, respectively. Coloration rather uniformly light brown to brown, sometimes with a faint cingulated pattern due to slightly infuscate prozona and more flavous metazona; paraterga, epiproct, venter and legs light yellow-brownish.</paragraph>
<paragraph pageId="4" pageNumber="5">
<pageBreakToken pageId="4" pageNumber="5" start="start">In</pageBreakToken>
<subSubSection id="CBC4CEEBC41FB21394B40EF3BA550CA1" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="5FBDB2DC166D0C2CA032322C1C2858A1" pageId="3" pageNumber="4">Description.</paragraph>
<paragraph id="8E29EC3494FCBDBC54E7A884B1236A0A" pageId="3" pageNumber="4">Length of both holo- and complete paratypes ca 20-21 mm, width of pro- and metazona 1.4-1.6 and 1.7-1.9 mm (♂), or 1.9 and 2.2 mm (♀), respectively. Holotype ca 20 mm long, 1.6 and 1.9 mm wide on midbody pro- and metazona, respectively. Coloration rather uniformly light brown to brown, sometimes with a faint cingulated pattern due to slightly infuscate prozona and more flavous metazona; paraterga, epiproct, venter and legs light yellow-brownish.</paragraph>
<paragraph id="2A5991A551C8AF6A236BB8FBA4BB09CB" pageId="4" pageNumber="5">
<pageBreakToken id="6C6F5820C18B69DA880A91226703E2E1" pageId="4" pageNumber="5" start="start">In</pageBreakToken>
width, head&gt; segment 2 = 5-16&gt; collum&gt; 3 = 4; on segments 17-19, body gently tapering towards telson. Entire head densely setose. Antennae rather long, slightly clavate, reaching beyond segment 4 (♂) or 3 (♀) dorsally; antennomere 2 longer than 6th. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally rather smooth and shining, only below several anterior paraterga faintly microgranulate. Paraterga of postcollum segments rather poorly developed (Figs 1, 2), set low (at about 1/2-1/3 midbody height), subhorizontal to faintly declivous, thin, blade-shaped, only a little thicker in pore-bearing segments; each callus with 3-5 small, but evident denticulations at lateral margin. Only paraterga 2 with caudal corner slightly obtusangular, narrowly rounded; following paraterga acutangular and pointed, yet evidently extending beyond rear tergal margin only on segment 18. Lateral callus of paraterga narrow, delimited by a complete sulcus only dorsally, in pore-bearing segments in caudal 1/3 also ventrally. Collum and following metaterga abundantly and irregularly setose (Figs 1, 2), setae being medium-sized, not borne on knobs. Ozopores lateral, superficial, ovoid, lying in front of penultimate denticulation. Axial line nearly wanting, vaguely traceable only here and there on metaterga. Transverse sulcus on metaterga especially superficial and incomplete on metatergum 4, slightly deeper, but still somewhat not reaching bases of paraterga in metaterga 5-17. Stricture between pro- and metazona evidently beaded. Pleurosternal carinae small, complete only in segments 2-4, divided into a front swelling and a sharp caudal denticle in segments 5-7(8), thereafter entirely missing. Epiproct short, flattened dorsoventrally, digitiform; apical papillae rather evident, dentiform, directed caudad; subapical papillae small, strongly removed from tip. Hypoproct semi-circular, setigerous paramedian knobs at caudal margin well-separated and small.
</paragraph>
<paragraph pageId="4" pageNumber="5">Sterna rather densely setose; a rather high, linguiform, subtruncate, ventral, setose lamina only between ♂ coxae 4 (Fig. 3). Legs relatively long, midbody ones ca 1.5-1.7 (♂) or 1.2-1.3 times (♀) longer than body height, a little slenderer in ♀; ♂tarsal brushes increasingly reduced towards legs of segment 12.</paragraph>
<paragraph pageId="4" pageNumber="5">Gonopods as in Figs 4-6. Solenomere as usual, flagelliform, nearly entirely sheathed by well-developed lamina lateralis and lamina medialis. Distal 1/3 of solenophore with a lateral denticle (k), as well as a subapical and an apical lobule on a larger branch (j). Femorite with a large, rounded, apicolateral lobe (l).</paragraph>
<caption pageId="4" pageNumber="5">
<paragraph pageId="4" pageNumber="5">
<paragraph id="5A3B64610F61E408107AF01150C516BA" pageId="4" pageNumber="5">Sterna rather densely setose; a rather high, linguiform, subtruncate, ventral, setose lamina only between ♂ coxae 4 (Fig. 3). Legs relatively long, midbody ones ca 1.5-1.7 (♂) or 1.2-1.3 times (♀) longer than body height, a little slenderer in ♀; ♂tarsal brushes increasingly reduced towards legs of segment 12.</paragraph>
<paragraph id="6B0FD28BE6A7D4ED870BB5F0D05EEB6B" pageId="4" pageNumber="5">Gonopods as in Figs 4-6. Solenomere as usual, flagelliform, nearly entirely sheathed by well-developed lamina lateralis and lamina medialis. Distal 1/3 of solenophore with a lateral denticle (k), as well as a subapical and an apical lobule on a larger branch (j). Femorite with a large, rounded, apicolateral lobe (l).</paragraph>
<caption id="0B90A0DAA8040584884EA50EB43DB0B4" pageId="4" pageNumber="5">
<paragraph id="E66AEA6B3EAE601A0827E196682A38E3" pageId="4" pageNumber="5">
Figures 1-6.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma papua" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="papua">Eustrongylosoma papua</taxonomicName>
<taxonomicName id="17099F140AE3B9F48B5CFCD13A2D8280" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma papua" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="papua">Eustrongylosoma papua</taxonomicName>
sp. n., ♂ paratype from Telefomin area. 1 segment 9, lateral view 2 left half of segment 9, dorsal view 3 sternal lobe between coxae 4, caudal view 4-6 right gonopod, mesal, dorsal and lateral views, respectively. Scale bars: 1.0 (1-3) and 0.3 mm (4-6).
</paragraph>
</caption>

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Patel &amp; Reddy, 1990: 41, f. 1
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(♂♀).
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<paragraph pageId="7" pageNumber="62">Comments.</paragraph>
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<paragraph id="E23169000614397B08684CB92D4FBADF" pageId="7" pageNumber="62">Comments.</paragraph>
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This species was described on the basis of both sexes from Andhra Pradesh (Fig. 17), but the description and figures are of poor quality. Judging from the figures of the male palp this species belongs in
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. Judging from the colour, shape of the epigyne and its distribution, the species belongs to
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, a titanoecid genus restricted to India, Sri Lanka, southern China, Myanmar and Thailand. The type species of the genus has a broader distribution. Judging from the shape of the epigyne, this species may be a junior synonym of
<taxonomicName class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava laminata" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="laminata">Pandava laminata</taxonomicName>
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(Thorell, 1878), the type species of the genus, known from East Africa to the Philippines and Marquesas Islands.
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108
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Figs 55-61
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<paragraph pageId="14" pageNumber="15">Type material.</paragraph>
<paragraph pageId="14" pageNumber="15">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin area, 1700 m, September 1975, B.S.E. (leg. P. Beron). - Paratype: 1 ♂ fragment(NMNHS), same locality, together with holotype.</paragraph>
<subSubSection id="CDDEAF892605299D2005369625DCA932" pageId="14" pageNumber="15" type="type material">
<paragraph id="6B9F99075A84E67E99DBA0D1FC824AC0" pageId="14" pageNumber="15">Type material.</paragraph>
<paragraph id="869F38EF57D62919C4B488E9FE0D0F80" pageId="14" pageNumber="15">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin area, 1700 m, September 1975, B.S.E. (leg. P. Beron). - Paratype: 1 ♂ fragment(NMNHS), same locality, together with holotype.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="name">
<paragraph pageId="14" pageNumber="15">Name.</paragraph>
<paragraph pageId="14" pageNumber="15">To reflect the beautiful appearance.</paragraph>
<subSubSection id="B1B582A66CAD09CFBEB3EA7731772476" pageId="14" pageNumber="15" type="name">
<paragraph id="C5B026947D56D5C85F5DD091A5F935DF" pageId="14" pageNumber="15">Name.</paragraph>
<paragraph id="6FC618754D6B168C8BDA867EC6311DD8" pageId="14" pageNumber="15">To reflect the beautiful appearance.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="diagnosis">
<paragraph pageId="14" pageNumber="15">Diagnosis.</paragraph>
<paragraph pageId="14" pageNumber="15">Distinguished by the rugulose to rugose caudal halves of metaterga and the relatively thin paraterga (see also Key below).</paragraph>
<subSubSection id="6CEE8DBDD1B3FEF962B0AF0B0600CEDB" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="9732A115FF373A8703EC0F6E00181712" pageId="14" pageNumber="15">Diagnosis.</paragraph>
<paragraph id="29F407EACC905BE4FAC11ABC58431978" pageId="14" pageNumber="15">Distinguished by the rugulose to rugose caudal halves of metaterga and the relatively thin paraterga (see also Key below).</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph pageId="14" pageNumber="15">Description.</paragraph>
<paragraph pageId="14" pageNumber="15">Length ca 20 mm (holotype), width of pro- and metazona 1.5 and 2.0 mm, respectively (both holo- and paratype). Coloration uniformly light yellow-brownish to brownish.</paragraph>
<paragraph lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="E22B87095A6267E58D9B8AF5F75899D6" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="DF881359781139C579AB94A675ED788D" pageId="14" pageNumber="15">Description.</paragraph>
<paragraph id="6BA1CBA6F9A5BDBDDEDB821CCAAEFB36" pageId="14" pageNumber="15">Length ca 20 mm (holotype), width of pro- and metazona 1.5 and 2.0 mm, respectively (both holo- and paratype). Coloration uniformly light yellow-brownish to brownish.</paragraph>
<paragraph id="0C00387A93E43EF26142C329917DBF1A" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
In width, head &gt;&gt; collum = segment 3 = 4 &lt;2 &lt;5-16. Head densely setose, only vertex with 1+1 setae. Antennae rather short, slightly clavate, reaching midway of segment 3
<pageBreakToken pageId="15" pageNumber="16" start="start">dorsally</pageBreakToken>
<pageBreakToken id="842E7F1433A445AFC506BE8480307279" pageId="15" pageNumber="16" start="start">dorsally</pageBreakToken>
; antennomere 2 longer than 6th. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate; rear halves of metaterga rugulose to rugose longitudinally. Paraterga of postcollum segments relatively well developed (Figs 55, 56), narrow, only slightly thicker in pore-bearing segments than in poreless ones, set low (at about 1/2 midbody height), with a clear undulation in front of ozopore near caudal third to quarter; paraterga 2 with a small anterolateral indentation, following ones often with a minute indentation or puncture near midway. Caudal corners of postcollum paraterga mainly nearly pointed to pointed, beak-shaped, mostly not or only barely extending beyond rear tergal margin, evidently extending behind it only on segments 16-19. Lateral calluses of paraterga
<pageBreakToken pageId="16" pageNumber="17" start="start">medium-sized</pageBreakToken>
<pageBreakToken id="ACB0746B054BC2FC89B6F1B6DE13B1A6" pageId="16" pageNumber="17" start="start">medium-sized</pageBreakToken>
, somewhat narrower in poreless segments, delimited by a sulcus dorsally, in pore-bearing segments ventrally in caudal 1/3-1/4 as well. Ozopores lateral, placed inside an ovoid groove. Tergal setae mostly abraded, medium-sized (ca 1/3 metatergal length), setation pattern traceable at least as 1+1 paramedian insertion points in front row. Axial line weak, traceable only on some metaterga, especially so in caudal halves. Transverse sulcus on metaterga rather superficial, incomplete, not reaching bases of paraterga, visible on metaterga 5-18. Stricture between pro- and metazona evidently beaded. Pleurosternal carinae small, well visible only in segment 2, reduced in 3rd, thereafter entirely missing. Epiproct (Fig. 57) short, flattened dorsoventrally, digitiform; apical papillae small, tip narrowly subtruncate; subapical papillae very small, removed from tip. Hypoproct (Fig. 58) semi-circular, setigerous paramedian knobs at caudal margin well-separated and very small.
</paragraph>
<paragraph pageId="16" pageNumber="17">Sterna rather sparsely setose; a rather high, linguiform, roundly subtruncate, ventral, setose lamina only between coxae 4 (Fig. 59). Legs long, midbody ones ca 1.9-2.0 times longer than body height, apparently somewhat incrassate and longer compared to ♀; ♂evident tarsal and distotibial brushes increasingly well reduced towards legs of segment 19.</paragraph>
<paragraph pageId="16" pageNumber="17">Gonopods as in Figs 60 &amp; 61. Distal third of solenophore split into a lateral lobe (k) and a large branch/lobe (j) of irregular shape. Apicolateral lobe (l) of femorite rounded, rather low.</paragraph>
<caption pageId="16" pageNumber="17">
<paragraph pageId="16" pageNumber="17">
<paragraph id="43FBBD47A18186804A102CA01317A4FB" pageId="16" pageNumber="17">Sterna rather sparsely setose; a rather high, linguiform, roundly subtruncate, ventral, setose lamina only between coxae 4 (Fig. 59). Legs long, midbody ones ca 1.9-2.0 times longer than body height, apparently somewhat incrassate and longer compared to ♀; ♂evident tarsal and distotibial brushes increasingly well reduced towards legs of segment 19.</paragraph>
<paragraph id="1E54687214243BC2E544F590A3B53C99" pageId="16" pageNumber="17">Gonopods as in Figs 60 &amp; 61. Distal third of solenophore split into a lateral lobe (k) and a large branch/lobe (j) of irregular shape. Apicolateral lobe (l) of femorite rounded, rather low.</paragraph>
<caption id="59ADAFF6B25E6BB2B6A662EE2E882705" pageId="16" pageNumber="17">
<paragraph id="7A75CE77F4F93F9060D8F77E409BDF92" pageId="16" pageNumber="17">
Figures 55-61.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma mirabile" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mirabile">Eustrongylosoma mirabile</taxonomicName>
<taxonomicName id="96511597CBE80220F1604E31156471A5" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma mirabile" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mirabile">Eustrongylosoma mirabile</taxonomicName>
sp. n., holotype. 55 segment 10, lateral view 56 left half of segment 10, dorsal view 57 tip of epiproct, dorsal view 58 hypoproct, ventral view 59 sternal lobe between coxae 4, caudal view 60 &amp; 61 right gonopod, mesal and sublateral views, respectively. Scale bars: 1.0 (55-59) and 0.5 mm (60 &amp; 61).
</paragraph>
</caption>

104
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@ -1,74 +1,74 @@
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<taxonomicNameLabel pageId="14" pageNumber="15">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="A024532FDFDC8102A09A35722ED3E7B4" pageId="14" pageNumber="15">sp. n.</taxonomicNameLabel>
Figs 48-54
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="type material">
<paragraph pageId="14" pageNumber="15">Type material.</paragraph>
<paragraph pageId="14" pageNumber="15">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin, 1700 m, 19 October 1975, B.S.E. (leg. P. Beron).</paragraph>
<subSubSection id="AB88BB13F7F5E5D31201AB0BCD884B30" pageId="14" pageNumber="15" type="type material">
<paragraph id="A2104DFAB88D5B7F9D452EF685DA447D" pageId="14" pageNumber="15">Type material.</paragraph>
<paragraph id="3E02D246BDF88D6D977C291DA85E7C78" pageId="14" pageNumber="15">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin, 1700 m, 19 October 1975, B.S.E. (leg. P. Beron).</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="name">
<paragraph pageId="14" pageNumber="15">Name.</paragraph>
<paragraph pageId="14" pageNumber="15">To reflect the pallid coloration.</paragraph>
<subSubSection id="6F77AE7B81C1A7D2AC56A6BB7C9909BE" pageId="14" pageNumber="15" type="name">
<paragraph id="14FEF0843A0DF7C66B96331128E34ECA" pageId="14" pageNumber="15">Name.</paragraph>
<paragraph id="5F34FB159F36FEB26BFE84AC92F9E1EE" pageId="14" pageNumber="15">To reflect the pallid coloration.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="diagnosis">
<paragraph pageId="14" pageNumber="15">Diagnosis.</paragraph>
<paragraph pageId="14" pageNumber="15">Distinguished by the pallid coloration, coupled with very poorly developed paraterga and the barbed solenophore tip (see also Key below).</paragraph>
<subSubSection id="844986198C9B760D48B84C43CE479C16" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="A87FDF39B19E006EFE2AB225ECE22B6B" pageId="14" pageNumber="15">Diagnosis.</paragraph>
<paragraph id="209F9778C7E184F56B414F3F483BC6BF" pageId="14" pageNumber="15">Distinguished by the pallid coloration, coupled with very poorly developed paraterga and the barbed solenophore tip (see also Key below).</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="description">
<paragraph pageId="14" pageNumber="15">Description.</paragraph>
<paragraph pageId="14" pageNumber="15">Length ca 20 mm, width of pro- and metazona 1.8 and 1.95 mm, respectively. Coloration uniformly light white-yellowish, with traces of light brown on rear halves of metaterga.</paragraph>
<paragraph pageId="14" pageNumber="15">In width, head&gt; collum&gt; segment 3 = 4 &lt;2 &lt;5-16. Head densely setose, only vertex with 1+1 setae. Antennae rather short, slightly clavate, reaching beyond segment 2 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate and rugulose; rear halves of metaterga sometimes faintly rugulose as well. Paraterga of postcollum segments very poorly developed (Fig. 48), full ridges only in segments 2-4, set low (at about 1/2 midbody height), thereafter evident lateral swellings; paraterga 2 with a small, but evident anterolateral indentation, following paraterga completely smooth at lateral margin. Lateral calluses of paraterga at most only very faintly delimited by a sulcus dorsally, more often totally wanting. Ozopores lateral, superficial. Tergal setae nearly fully abraded, a few short ones retained only on collum; setation pattern vague. Axial line wanting. Transverse sulcus on metaterga rather superficial, incomplete, far from reaching bases of paraterga, visible on metaterga 6-17, barely traceable on 5th and 18th. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, complete in segments 2-4, a rounded caudal denticle retained also in segments 5-7, thereafter entirely missing. Epiproct (Fig. 49) short, flattened dorsoventrally, digitiform; apical papillae large, unciform, directed caudoventrally, tip distinctly emarginate; subapical papillae wanting. Hypoproct (Fig. 50) semi-circular, setigerous paramedian knobs at caudal margin well-separated and small.</paragraph>
<paragraph pageId="14" pageNumber="15">Sterna rather sparsely setose; a rather high, linguiform, roundly subtrapeziform, ventral, setose lamina only between coxae 4 (Fig. 51). Legs short, midbody ones ca 0.9-1.0 times as long as body height, apparently somewhat incrassate and longer compared to ♀; ♂tarsal brushes wanting.</paragraph>
<paragraph pageId="14" pageNumber="15">Gonopods as in Figs 52-54. Distal quarter of solenophore split into a small lateral lobe (k) and an abundantly denticulate larger branch (j). Femorite with a rather small apicolateral lobe (l).</paragraph>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
<subSubSection id="73590ADF03AA7204FC33AEB39681DED3" pageId="14" pageNumber="15" type="description">
<paragraph id="BE9E6F3EB384644FC3ADA50061BEEA73" pageId="14" pageNumber="15">Description.</paragraph>
<paragraph id="65F057034412EF3E878209C8B59BAA5D" pageId="14" pageNumber="15">Length ca 20 mm, width of pro- and metazona 1.8 and 1.95 mm, respectively. Coloration uniformly light white-yellowish, with traces of light brown on rear halves of metaterga.</paragraph>
<paragraph id="2EA3D5231D1383A585C0A12118E78207" pageId="14" pageNumber="15">In width, head&gt; collum&gt; segment 3 = 4 &lt;2 &lt;5-16. Head densely setose, only vertex with 1+1 setae. Antennae rather short, slightly clavate, reaching beyond segment 2 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate and rugulose; rear halves of metaterga sometimes faintly rugulose as well. Paraterga of postcollum segments very poorly developed (Fig. 48), full ridges only in segments 2-4, set low (at about 1/2 midbody height), thereafter evident lateral swellings; paraterga 2 with a small, but evident anterolateral indentation, following paraterga completely smooth at lateral margin. Lateral calluses of paraterga at most only very faintly delimited by a sulcus dorsally, more often totally wanting. Ozopores lateral, superficial. Tergal setae nearly fully abraded, a few short ones retained only on collum; setation pattern vague. Axial line wanting. Transverse sulcus on metaterga rather superficial, incomplete, far from reaching bases of paraterga, visible on metaterga 6-17, barely traceable on 5th and 18th. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, complete in segments 2-4, a rounded caudal denticle retained also in segments 5-7, thereafter entirely missing. Epiproct (Fig. 49) short, flattened dorsoventrally, digitiform; apical papillae large, unciform, directed caudoventrally, tip distinctly emarginate; subapical papillae wanting. Hypoproct (Fig. 50) semi-circular, setigerous paramedian knobs at caudal margin well-separated and small.</paragraph>
<paragraph id="2F10AF1A0CA0B1CCF741417C27CBC232" pageId="14" pageNumber="15">Sterna rather sparsely setose; a rather high, linguiform, roundly subtrapeziform, ventral, setose lamina only between coxae 4 (Fig. 51). Legs short, midbody ones ca 0.9-1.0 times as long as body height, apparently somewhat incrassate and longer compared to ♀; ♂tarsal brushes wanting.</paragraph>
<paragraph id="35C049FA359643D194AF832163AF0252" pageId="14" pageNumber="15">Gonopods as in Figs 52-54. Distal quarter of solenophore split into a small lateral lobe (k) and an abundantly denticulate larger branch (j). Femorite with a rather small apicolateral lobe (l).</paragraph>
<caption id="8D893F91B63BED5BA2FD86410DD8E5CC" pageId="14" pageNumber="15">
<paragraph id="439EC8A67C138D9D4C4E9C9721BAEA04" pageId="14" pageNumber="15">
Figures 48-54.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma pallidum" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pallidum">Eustrongylosoma pallidum</taxonomicName>
<taxonomicName id="54976943F9EE40B34A58E3A1C3F71A48" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma pallidum" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pallidum">Eustrongylosoma pallidum</taxonomicName>
sp. n., holotype. 48 segment 10, lateral view 49 tip of epiproct, dorsal view 50 hypoproct, ventral view 51 sternal lobe between coxae 4, caudal view 52-54 right gonopod, mesal, subventral and sublateral views, respectively. Scale bars: 1.0 (48-51) and 0.3 mm (52-54).
</paragraph>
</caption>

96
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<taxonomicNameLabel pageId="7" pageNumber="62">comb. n.</taxonomicNameLabel>
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<paragraph id="2A195A06B92A7E3E12E13D82CF302887" pageId="7" pageNumber="62">
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<taxonomicNameLabel id="8086B6C6873321A7A2798BB14160F582" pageId="7" pageNumber="62">comb. n.</taxonomicNameLabel>
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<paragraph pageId="7" pageNumber="62">
<taxonomicName class="Arachnida" family="Amaurobiidae" genus="Amaurobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amaurobius nathabhaii" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="nathabhaii">Amaurobius nathabhaii</taxonomicName>
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Patel &amp; Patel, 1975: 801, f. 1
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(♀).
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<paragraph pageId="7" pageNumber="62">Comments.</paragraph>
<paragraph pageId="7" pageNumber="62">
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<paragraph id="EB40EA44983B11A2753205C2F37EA72A" pageId="7" pageNumber="62">Comments.</paragraph>
<paragraph id="CD109BE62A7D8CB91074E7595F77D78A" pageId="7" pageNumber="62">
This species was described on the basis of the female sex from Gujarat (Fig. 17), but the description and figures are of very poor quality. This species is placed in
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because the other Indian species placed in
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<taxonomicName id="7C36E4EA46C40DBCD2697293D1ED662C" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amaurobius" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">Amaurobius</taxonomicName>
belong to
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. It is transferred to
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<taxonomicName id="BC9923A82189A5927F2DBBF396DF4175" class="Arachnida" family="Titanoecidae" genus="Pandava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pandava" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">Pandava</taxonomicName>
because of its southern distribution.
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110
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<taxonomicName LSID="urn:lsid:zoobank.org:act:7CC90A09-ED88-4047-9FC7-EFE46F7A95B0" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma tifalmin" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tifalmin">Eustrongylosoma tifalmin</taxonomicName>
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Figs 34-40
</paragraph>
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<paragraph pageId="10" pageNumber="11">Type material.</paragraph>
<paragraph pageId="10" pageNumber="11">Holotype♂(NMNHS),Papua New Guinea, Western Province, Finim Tel area, October 1975, B.S.E. (leg. P. Beron).</paragraph>
<subSubSection id="E21EB2D9C36A44AFF094631308908E94" pageId="10" pageNumber="11" type="type material">
<paragraph id="B3A3B813A901155CDD97D1B183285341" pageId="10" pageNumber="11">Type material.</paragraph>
<paragraph id="A2A7737E0041F059268D24FAEF7CF254" pageId="10" pageNumber="11">Holotype♂(NMNHS),Papua New Guinea, Western Province, Finim Tel area, October 1975, B.S.E. (leg. P. Beron).</paragraph>
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<subSubSection pageId="10" pageNumber="11" type="name">
<paragraph pageId="10" pageNumber="11">Name.</paragraph>
<paragraph pageId="10" pageNumber="11">
<normalizedToken originalValue="“Tifalmin”">&quot;Tifalmin&quot;</normalizedToken>
<subSubSection id="58C6B1038D87269AAACA36F4308D66BC" pageId="10" pageNumber="11" type="name">
<paragraph id="4EF9152F47E480B21A1E72093E0D9D6A" pageId="10" pageNumber="11">Name.</paragraph>
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<normalizedToken id="6AE92D5B3FDCE0B6CB4E3736573A60DE" originalValue="“Tifalmin”">&quot;Tifalmin&quot;</normalizedToken>
is the name of one of the main local tribes. A noun in apposition.
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<subSubSection pageId="10" pageNumber="11" type="diagnosis">
<paragraph pageId="10" pageNumber="11">Diagnosis.</paragraph>
<paragraph pageId="10" pageNumber="11">Distinguished by the relatively large size, coupled with a peculiar, albeit vague, colour pattern and a deeply split distal part of the solenophore (see also Key below).</paragraph>
<subSubSection id="98E854D6E49CCB61F4F2890D35DAA992" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="BA2A7C54D04B0C3020F2D43424EA8E33" pageId="10" pageNumber="11">Diagnosis.</paragraph>
<paragraph id="A28A202CA4FF56516A3BCEE7F7AC5401" pageId="10" pageNumber="11">Distinguished by the relatively large size, coupled with a peculiar, albeit vague, colour pattern and a deeply split distal part of the solenophore (see also Key below).</paragraph>
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<paragraph pageId="10" pageNumber="11">Description.</paragraph>
<paragraph pageId="10" pageNumber="11">Length ca 24 mm, width of pro- and metazona 1.9 and 2.6 mm, respectively. Coloration rather uniformly castaneous brown with a broad, rather vague, light yellow-brown, axial stripe running from collum to end of epiproct; much of prozona dorsally and dorsolaterally slightly infuscate, up to dark brown; head and antennae light brown; legs mostly light yellow-brown, only tibiae and tarsi infuscate, dark brown.</paragraph>
<paragraph lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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<paragraph id="0E2440ADC074784C19DD1EC1AF4ED737" pageId="10" pageNumber="11">Description.</paragraph>
<paragraph id="664CF1121EEE398DA6300C6D4D3C3C7A" pageId="10" pageNumber="11">Length ca 24 mm, width of pro- and metazona 1.9 and 2.6 mm, respectively. Coloration rather uniformly castaneous brown with a broad, rather vague, light yellow-brown, axial stripe running from collum to end of epiproct; much of prozona dorsally and dorsolaterally slightly infuscate, up to dark brown; head and antennae light brown; legs mostly light yellow-brown, only tibiae and tarsi infuscate, dark brown.</paragraph>
<paragraph id="4B110B98C8C5CE97D093BF516C5439DE" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
In width, head = segments 5-17&gt; 2&gt; collum&gt; 3 = 4. Head densely setose, only vertex bare. Antennae medium-sized, slightly clavate, reaching beyond segment 3 dorsally; antennomere 2 longer than 6th. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate; rear halves of metaterga sometimes faintly rugulose. Paraterga of postcollum segments moderately well developed (Figs 34, 35), set low (at about 1/2-1/3 midbody height), subhorizontal, evidently thicker in pore-bearing segments than in poreless ones; paratergum 2 with a small anterolateral denticle, its front margin straight, its caudal corner slightly obtusangular, rather narrowly rounded, not extended beyond rear tergal margin; following paraterga without denticulations at lateral margin, sometimes with only a faint undulation in front of ozopore near caudal third. Caudal corners of postcollum paraterga always rounded, at most only slightly extended beyond rear tergal margin, more evidently so only in segments 16-19. Lateral calluses of paraterga medium-sized, broader in pore-bearing segments, each delimited by a sulcus both dorsally and, in caudal 1/3, ventrally. Ozopores lateral, lying inside an ovoid groove. Tergal setae mostly abraded, a few retained only on collum, about 1/5 the length of metaterga,
<pageBreakToken pageId="11" pageNumber="12" start="start">setation</pageBreakToken>
<pageBreakToken id="E42854900934A774C58ABC7E39342661" pageId="11" pageNumber="12" start="start">setation</pageBreakToken>
pattern sometimes traceable at least as 1+1 paramedian insertion points in front row. Axial line wanting. Transverse sulcus on metaterga rather superficial, nearly reaching bases of paraterga in metaterga 5-18. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, complete in segments 2-4, a sharp or rounded caudal denticle retained also in segments 5-8, thereafter entirely missing. Epiproct (Fig. 36) short, flattened dorsoventrally, digitiform; apical papillae large, tip narrowly emarginate; subapical papillae missing. Hypoproct (Fig. 37) roundly subtrapeziform, setigerous paramedian knobs at caudal margin well-separated and small.
</paragraph>
<paragraph pageId="12" pageNumber="13">
<pageBreakToken pageId="12" pageNumber="13" start="start">Sterna</pageBreakToken>
<paragraph id="9FB096E828A3878D251F7D0C27E492A3" pageId="12" pageNumber="13">
<pageBreakToken id="6DCD90207B5130A7EFE79BEE2F7E2A6E" pageId="12" pageNumber="13" start="start">Sterna</pageBreakToken>
rather sparsely setose; a rather high, linguiform, roundly subtriangular, ventral, setose lamina only between coxae 4 (Fig. 38). Legs relatively long, midbody ones ca 1.6-1.7 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal and distotibial brushes increasingly well reduced towards legs of segment 19 and 9, respectively.
</paragraph>
<paragraph pageId="12" pageNumber="13">Gonopods as in Figs 39 &amp; 40. Tip (j) of solenophore very deeply biramous, larger (= outer) branch being faintly bidentate, smaller (= inner) one spiniform. Apicolateral lobe (l) of femorite rather small.</paragraph>
<caption pageId="12" pageNumber="13">
<paragraph pageId="12" pageNumber="13">
<paragraph id="704AF34AAF320234F27069627784410D" pageId="12" pageNumber="13">Gonopods as in Figs 39 &amp; 40. Tip (j) of solenophore very deeply biramous, larger (= outer) branch being faintly bidentate, smaller (= inner) one spiniform. Apicolateral lobe (l) of femorite rather small.</paragraph>
<caption id="0320701B8E0449F0342D70F04BEA657B" pageId="12" pageNumber="13">
<paragraph id="42FEEB1CA89342DC2ECDBEFB6CDE9B5E" pageId="12" pageNumber="13">
Figures 34-40.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma tifalmin" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tifalmin">Eustrongylosoma tifalmin</taxonomicName>
<taxonomicName id="76B553E2A033FF8AD30EF2EB0F3315E4" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma tifalmin" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tifalmin">Eustrongylosoma tifalmin</taxonomicName>
sp. n., holotype. 34 segment 10, lateral view 35 left half of segment 10, dorsal view 36 tip of epiproct, dorsal view 37 hypoproct, ventral view 38 sternal lobe between coxae 4, caudal view 39, 40 left gonopod, mesal and subdorsal views, respectively. Scale bars: 1.0 (34-38) and 0.3 mm (39 &amp; 40).
</paragraph>
</caption>

112
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<taxonomicNameLabel id="F36D1BABBA9DEB75595D479934C619EC" pageId="4" pageNumber="5">sp. n.</taxonomicNameLabel>
Figs 7-14
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="type material">
<paragraph pageId="4" pageNumber="5">Type material.</paragraph>
<paragraph pageId="4" pageNumber="5">Holotype♂(NMNHS),Papua New Guinea, Western Province, Mt Fugilil, near camp, 2980 m, 9 September 1975, B.S.E. (leg. Ph. Chapman &amp; P. Beron). - Paratype: 1 ♂(NMNHS), West Sepik Province, Telefomin area, 1700 m, September 1975, B.S.E. (leg. P. Beron).</paragraph>
<subSubSection id="AD1D32A0AF67AABE852367C8773E27FD" pageId="4" pageNumber="5" type="type material">
<paragraph id="5294DBD1DE8F2268FCD8314E02FC84AD" pageId="4" pageNumber="5">Type material.</paragraph>
<paragraph id="2F63824C199FE77B15A9E65E92015306" pageId="4" pageNumber="5">Holotype♂(NMNHS),Papua New Guinea, Western Province, Mt Fugilil, near camp, 2980 m, 9 September 1975, B.S.E. (leg. Ph. Chapman &amp; P. Beron). - Paratype: 1 ♂(NMNHS), West Sepik Province, Telefomin area, 1700 m, September 1975, B.S.E. (leg. P. Beron).</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="name">
<paragraph pageId="4" pageNumber="5">Name.</paragraph>
<paragraph pageId="4" pageNumber="5">
<normalizedToken originalValue="“tumbuna”">&quot;tumbuna&quot;</normalizedToken>
<subSubSection id="1461115E27A613C60A88F45A694D6992" pageId="4" pageNumber="5" type="name">
<paragraph id="C7C73F59541AE17CB128000960411BF4" pageId="4" pageNumber="5">Name.</paragraph>
<paragraph id="00495E75C3F69A1DA46CA65BC0AFEE65" pageId="4" pageNumber="5">
<normalizedToken id="E7E2861772822D912869CDC6A2925CB4" originalValue="“tumbuna”">&quot;tumbuna&quot;</normalizedToken>
means
<normalizedToken originalValue="“ancestor”">&quot;ancestor&quot;</normalizedToken>
<normalizedToken id="968FDFB948166BC37AA23F4A70A10D72" originalValue="“ancestor”">&quot;ancestor&quot;</normalizedToken>
or
<normalizedToken originalValue="“ancestors”">&quot;ancestors&quot;</normalizedToken>
<normalizedToken id="9A455D055D032E05809FAB416F3BD364" originalValue="“ancestors”">&quot;ancestors&quot;</normalizedToken>
in Pidgin English. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="diagnosis">
<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="5">Differs from congeners by the rather wide paratergal calluses strongly delimited not only dorsally, but also ventrally, by the distal part of the solenophore being dentate/lobulate, not split (see also Key below).</paragraph>
<subSubSection id="901D60BF860355B8294A3ECA94A84FF7" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="A4FBFC239363477753212C4536F4B8B6" pageId="4" pageNumber="5">Diagnosis.</paragraph>
<paragraph id="420C1B81353E948D2C8A89B53B44C894" pageId="4" pageNumber="5">Differs from congeners by the rather wide paratergal calluses strongly delimited not only dorsally, but also ventrally, by the distal part of the solenophore being dentate/lobulate, not split (see also Key below).</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph pageId="4" pageNumber="5">Description.</paragraph>
<paragraph pageId="4" pageNumber="5">Length of both holo- and paratype ca 20 mm, width of pro- and metazona 1.4 and 2.1 (holotype) or 1.2 and 1.8 mm (paratype), respectively. Coloration rather uniformly light brown to brown, only front 1/3 of body slightly more intensely brown; dorsal sides of prozona, calluses on paraterga, epiproct and venter light yellow-brown to nearly pallid. Sometimes (paratype) region near stricture between pro- and metazona slightly infuscate, brown, thus creating a faint cingulate pattern.</paragraph>
<paragraph pageId="5" pageNumber="6">
<pageBreakToken pageId="5" pageNumber="6" start="start">In</pageBreakToken>
<subSubSection id="58D009DF84B3F26DE9903F1803164433" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="25F604A7F9F289576866CC28944F0C7B" pageId="4" pageNumber="5">Description.</paragraph>
<paragraph id="B72E22013F91D8408592559F86856E06" pageId="4" pageNumber="5">Length of both holo- and paratype ca 20 mm, width of pro- and metazona 1.4 and 2.1 (holotype) or 1.2 and 1.8 mm (paratype), respectively. Coloration rather uniformly light brown to brown, only front 1/3 of body slightly more intensely brown; dorsal sides of prozona, calluses on paraterga, epiproct and venter light yellow-brown to nearly pallid. Sometimes (paratype) region near stricture between pro- and metazona slightly infuscate, brown, thus creating a faint cingulate pattern.</paragraph>
<paragraph id="BEE3B839BF1297070881C2071BFE6564" pageId="5" pageNumber="6">
<pageBreakToken id="1AF8B33D9B9BD3FC2D9FF326DB69B52B" pageId="5" pageNumber="6" start="start">In</pageBreakToken>
width, head &gt;&gt; collum = segment 3 = 4 &lt;2 &lt;5-16; thereafter body gently tapering towards telson. Entire head densely setose. Antennae rather long, slightly clavate, reaching beyond midlength of segment 3 or 4 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate; rear halves of metaterga sometimes faintly rugulose. Paraterga of postcollum segments moderately well developed (Figs 7, 8), set low (at about 1/3 midbody height), subhorizontal, evidently thicker in pore-bearing segments than in poreless ones; paratergum 2 with a small anterolateral denticle, its front margin straight; following paraterga without denticulations at lateral margin, sometimes with only a very faint undulation near midway. Caudal corner of all postcollum paraterga pointed to narrowly rounded, starting from 5th extending beyond rear tergal margin. Lateral calluses of paraterga rather wide, especially so on pore-bearing segments, each delimited by a sulcus both dorsally and ventrally. Ozopores lateral to dorsolateral, placed inside an ovoid groove. Tergal setae mostly abraded, retained ones about 1/5 the length of metaterga, pattern traceable at least as 1+1 paramedian insertion points in front row (Figs 7, 8). Axial line nearly wanting, vaguely traceable only here and there on metaterga. Transverse sulcus on metaterga rather superficial, nearly reaching bases of paraterga in metaterga 5-18. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, complete or nearly so in segments 2-8, a sharp or rounded caudal denticle retained also in segments 9 and 10, thereafter entirely missing. Epiproct (Fig. 9) short, flattened dorsoventrally, digitiform; apical papillae small, tip faintly concave; subapical papillae evident, removed from tip. Hypoproct subtrapeziform, setigerous paramedian knobs at caudal margin well-separated and small.
</paragraph>
<paragraph pageId="5" pageNumber="6">Sterna rather densely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 10). Legs relatively long, midbody ones ca 1.6-1.7 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal and distotibial brushes increasingly well reduced towards legs of segment 10.</paragraph>
<paragraph pageId="5" pageNumber="6">Gonopods as in Figs 11-14. Tip of solenophore simple, faintly trifid, with three apical denticles or lobules. Femorite with a large rounded lobe (l) somewhat hollow on mesal side.</paragraph>
<caption pageId="5" pageNumber="6">
<paragraph pageId="5" pageNumber="6">
<paragraph id="B36B85350FE9C9C412D268F53F42EAE6" pageId="5" pageNumber="6">Sterna rather densely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 10). Legs relatively long, midbody ones ca 1.6-1.7 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal and distotibial brushes increasingly well reduced towards legs of segment 10.</paragraph>
<paragraph id="95F59F1E646C2A878B76DB354FF3AFCB" pageId="5" pageNumber="6">Gonopods as in Figs 11-14. Tip of solenophore simple, faintly trifid, with three apical denticles or lobules. Femorite with a large rounded lobe (l) somewhat hollow on mesal side.</paragraph>
<caption id="739E9E8EF2A3DD7D20C9968E3D9D0253" pageId="5" pageNumber="6">
<paragraph id="E53C9D7CEAA36BC343C8AA4B4EA43598" pageId="5" pageNumber="6">
Figures 7-14.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma tumbuna" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tumbuna">Eustrongylosoma tumbuna</taxonomicName>
<taxonomicName id="1BF25124CFD7998CF58133A2B7E3D393" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma tumbuna" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tumbuna">Eustrongylosoma tumbuna</taxonomicName>
sp. n., holotype. 7 segment 10, lateral view 8 left half of segment 10, dorsal view 9 tip of epiproct, dorsal view 10 sternal lobe between coxae 4, caudal view 11-14 left gonopod, mesal, ventral, sublateral and subdorsal views, respectively. Scale bars: 1.0 (7-10) and 0.3 mm (11-14).
</paragraph>
</caption>

72
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stigma
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(Loew, 1840)

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<taxonomicName id="9B7970249DAB8538299AC8FBA6702C42" ID-CoL="3DBGB" LSID="urn:lsid:zoobank.org:act:0BD40883-E0A0-40D3-AB66-8DF034269CB2" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma curtipes" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="curtipes">Eustrongylosoma curtipes</taxonomicName>
<taxonomicNameLabel id="0FCD3C81DE4C93DABEE6A9745DD49439" pageId="6" pageNumber="7">sp. n.</taxonomicNameLabel>
Figs 20-26
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="type material">
<paragraph pageId="6" pageNumber="7">Type material.</paragraph>
<paragraph pageId="6" pageNumber="7">Holotype♂(incomplete, lacking segments 8-11)(NMNHS),Papua New Guinea, Western Province, Finim Tel Plateau, rain forest, 2300 m, 8 August 1975, B.S.E. (leg. P. Beron).</paragraph>
<subSubSection id="BA29AD45CCBB8ED35E12992669EE3406" pageId="6" pageNumber="7" type="type material">
<paragraph id="7C7BAEDF0FD4A4DB72FB131AFDBD5E80" pageId="6" pageNumber="7">Type material.</paragraph>
<paragraph id="C998B1B44EADCC17C7AB4D1F400C0D8D" pageId="6" pageNumber="7">Holotype♂(incomplete, lacking segments 8-11)(NMNHS),Papua New Guinea, Western Province, Finim Tel Plateau, rain forest, 2300 m, 8 August 1975, B.S.E. (leg. P. Beron).</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="name">
<paragraph pageId="6" pageNumber="7">Name.</paragraph>
<paragraph pageId="6" pageNumber="7">To reflect the short legs.</paragraph>
<subSubSection id="803156F87200FB3211D6FD55FA8ACF51" pageId="6" pageNumber="7" type="name">
<paragraph id="3A8A0D8E7B9D2802E964EA6A6BC244E3" pageId="6" pageNumber="7">Name.</paragraph>
<paragraph id="2D1DD8F4681A7D88B8725477B5B9EECA" pageId="6" pageNumber="7">To reflect the short legs.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="diagnosis">
<paragraph pageId="6" pageNumber="7">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="7">Distinguished by the rather short legs, coupled with quite evident paraterga and a missing colour pattern (see also Key below).</paragraph>
<subSubSection id="3932EF460FAF4639E9C5836B7E48645F" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="18701B523B4747A4DAAF0809962A52F2" pageId="6" pageNumber="7">Diagnosis.</paragraph>
<paragraph id="C6590FC062C78C69556F7C7FBBBBE1CE" pageId="6" pageNumber="7">Distinguished by the rather short legs, coupled with quite evident paraterga and a missing colour pattern (see also Key below).</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph pageId="6" pageNumber="7">Description.</paragraph>
<paragraph pageId="6" pageNumber="7">Length ca 18 mm, width of pro- and metazona 1.1 and 1.3 mm, respectively. Coloration rather uniformly light brown to red-brown, only legs yellowish.</paragraph>
<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="8CC61857C98AD98198FAD188CE6B5811" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="5848D647EE0E4BE03344753007E9214E" pageId="6" pageNumber="7">Description.</paragraph>
<paragraph id="398EF49381D5DB6732F5E01C0FFF8E11" pageId="6" pageNumber="7">Length ca 18 mm, width of pro- and metazona 1.1 and 1.3 mm, respectively. Coloration rather uniformly light brown to red-brown, only legs yellowish.</paragraph>
<paragraph id="38B0018A5E7FC02A75945673001EED28" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
In width, head&gt; collum = segment 2 &lt;3 = 4 &lt;5-17; thereafter body gently tapering towards telson. Head densely setose, but vertex with 1+1 setae. Antennae short, clavate, reaching beyond segment 2 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate. Paraterga of postcollum segments nearly missing (Figs 20, 21), set low (at about 1/2 midbody height), represented by low keels only in segments 2-4, thereafter only by evident lateral swellings. Ozopores lateral, superficial. Tergal setae entirely abraded, setation pattern untraceable. Axial line wanting. Transverse sulcus on metaterga superficial, far from reaching bases of paraterga, visible on metaterga 5-17. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, complete in seg
<pageBreakToken pageId="7" pageNumber="8" start="start">ments</pageBreakToken>
<pageBreakToken id="DF78DA532D0FF70B1727E954ABA9DACF" pageId="7" pageNumber="8" start="start">ments</pageBreakToken>
2-5, only a sharp or rounded caudal denticle until segment 7, thereafter entirely missing. Epiproct (Fig. 22) short, flattened dorsoventrally, digitiform; apical papillae small, tip faintly concave; subapical papillae small, removed from tip. Hypoproct semi-circular, setigerous paramedian knobs at caudal margin well-separated and small.
</paragraph>
<paragraph pageId="8" pageNumber="9">
<pageBreakToken pageId="8" pageNumber="9" start="start">Sterna</pageBreakToken>
<paragraph id="1F985593B46E62A8484505E0C1E8E2A6" pageId="8" pageNumber="9">
<pageBreakToken id="79DB1A1E041D45A10F9F1EDD9EB8D17D" pageId="8" pageNumber="9" start="start">Sterna</pageBreakToken>
rather sparsely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between coxae 4 (Fig. 23). Legs relatively short, midbody ones ca 1.1-1.2 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal and distotibial brushes increasingly well reduced towards legs of segment 19 and 16, respectively.
</paragraph>
<paragraph pageId="8" pageNumber="9">Gonopods as in Figs 24-26. Base of solenophore and entire apicolateral lobe of femorite (l) evidently plicate. Tip of solenophore (j) rather deeply bifid, with a mesally fringed subapical margin.</paragraph>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
<paragraph id="8B43CEC5F018132BBBFC3DEADEE853F6" pageId="8" pageNumber="9">Gonopods as in Figs 24-26. Base of solenophore and entire apicolateral lobe of femorite (l) evidently plicate. Tip of solenophore (j) rather deeply bifid, with a mesally fringed subapical margin.</paragraph>
<caption id="6520103092F2D2FE0BA389430D4FBEF8" pageId="8" pageNumber="9">
<paragraph id="551CD9D3CB2595904E480B1FFD951EFB" pageId="8" pageNumber="9">
Figures 20-26.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma curtipes" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="curtipes">Eustrongylosoma curtipes</taxonomicName>
<taxonomicName id="02304EF2E891B5AB065847A1A9AEEB21" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma curtipes" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="curtipes">Eustrongylosoma curtipes</taxonomicName>
sp. n., holotype. 20 segment 10, lateral view 21 right half of segment 10, dorsal view 22 tip of epiproct, dorsal view 23 sternal lobe between coxae 4, caudal view 24-26 left gonopod, mesal, sublateral and subdorsal views, respectively. Scale bars: 0.5 (20-23) and 0.3 mm (24-26).
</paragraph>
</caption>

150
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<taxonomicName LSID="urn:lsid:ipni.org:names:77186066-1" authority="X. H. Jin, L. A. Ye &amp; A. T. Mu" family="Odontochilus" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="1" pageNumber="20" rank="species" species="putaoensis">Odontochilus putaoensis X.H. Jin, L.A. Ye &amp; A.T. Mu</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="20">sp. nov.</taxonomicNameLabel>
<treatment id="DCCE73E5643D9DE9D951813ADD6A6034" ID-GBIF-Taxon="204887169" LSID="urn:lsid:plazi:treatment:DCCE73E5643D9DE9D951813ADD6A6034" httpUri="http://treatment.plazi.org/id/DCCE73E5643D9DE9D951813ADD6A6034" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" scope_family="Orchidaceae">
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<taxonomicNameLabel id="69AE2AC7F2F30B3F676AFD21A2665EAC" pageId="1" pageNumber="20">sp. nov.</taxonomicNameLabel>
Figure 1
</paragraph>
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<subSubSection pageId="1" pageNumber="20" type="diagnosis">
<paragraph pageId="1" pageNumber="20">Diagnosis.</paragraph>
<paragraph pageId="1" pageNumber="20">
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="1" pageNumber="20" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
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<paragraph id="63E5DE06DFD0F00113021D84D821950D" pageId="1" pageNumber="20">Diagnosis.</paragraph>
<paragraph id="F4F81AE47857E14607856FEB51613619" pageId="1" pageNumber="20">
<taxonomicName id="4DDB27866040CBD1B8DCD234686CABD3" class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="1" pageNumber="20" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
is similar to
<taxonomicName lsidName="O. duplex" pageId="1" pageNumber="20" rank="species" species="duplex">O. duplex</taxonomicName>
<taxonomicName id="48C9FA7536BAA265BE4B2B410CBA9300" lsidName="O. duplex" pageId="1" pageNumber="20" rank="species" species="duplex">O. duplex</taxonomicName>
, but can be easily distinguished from the latter by having a light yellow lip composed of a bisaccate hypochile with a small, erect, blade-like and emarginate callus within each sac, a mesochile with a pair of dentate-pectinate flanges and bilobed epichile with a pair of widely diverging lobes that are erect and concave.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="materials_examined">
<paragraph pageId="2" pageNumber="21">
<pageBreakToken pageId="2" pageNumber="21" start="start">Type</pageBreakToken>
<subSubSection id="D031AC5864A61A3BE192FE2FFF8CF8B2" pageId="2" pageNumber="21" type="materials_examined">
<paragraph id="FBE632863445CFAD25EB5811586371E0" pageId="2" pageNumber="21">
<pageBreakToken id="BC2753128E5E63B6392688ED667E1DBC" pageId="2" pageNumber="21" start="start">Type</pageBreakToken>
.
</paragraph>
<paragraph pageId="2" pageNumber="21">MYANMAR. Kachin State: Putao Township, Hponkanrazi Wildlife Sanctuary, subtropical, evergreen, broad-leaved, montane forest, 2000 m a.s.l., 20 October 2014, Xiaohua Jin et al, PT-ET 959 (Holotype, PE!).</paragraph>
<paragraph id="70C3631B09491819BA0C65E95D228E8D" pageId="2" pageNumber="21">MYANMAR. Kachin State: Putao Township, Hponkanrazi Wildlife Sanctuary, subtropical, evergreen, broad-leaved, montane forest, 2000 m a.s.l., 20 October 2014, Xiaohua Jin et al, PT-ET 959 (Holotype, PE!).</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="description">
<paragraph pageId="2" pageNumber="21">Description.</paragraph>
<paragraph pageId="2" pageNumber="21">
<subSubSection id="B1710F766AF6A895F353F3E12014B5B3" pageId="2" pageNumber="21" type="description">
<paragraph id="6D24939D6D9B7A12A9A83953B0181BE5" pageId="2" pageNumber="21">Description.</paragraph>
<paragraph id="CB0BC4D93C3C1E47BBDCA911E7C9FA81" pageId="2" pageNumber="21">
Plants autotrophic, terrestrial, 40-60 cm tall. Stem ascending, pubescent, 2-6-leaved. Leaves dark green, ovate-lanceolate, 6-7.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="ED8D682F99E59FA9B71EEF964B2028DF" originalValue="×">x</normalizedToken>
3.2-4 cm, attenuate at apex, blade glabrous; petiole-like base and tubular sheath ca. 2.8 cm long. Peduncle pubescent, with 1 or 2 sheathing bracts, reddish-brown, ovate-lanceolate, 12-15
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="7B1245B49702F6EB0732E465ABBBDC5D" originalValue="×">x</normalizedToken>
4-5 mm, pubescent, long acuminate; rachis pubescent, sub-densely 16-flowered; floral bracts
<normalizedToken originalValue="reddish–brown">reddish-brown</normalizedToken>
<normalizedToken id="74CCE63A83E88E9FC84B8E8033F91F1B" originalValue="reddish–brown">reddish-brown</normalizedToken>
, ovate-lanceolate, ca. 9
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="27E9176F4E8636DD4DDDF43DE9AA9EA3" originalValue="×">x</normalizedToken>
4 mm, as long as ovary, abaxially pubescent, long acuminate at apex. Flowers resupinate; dorsal sepal forming a hood with petals, dark greenish-brown, ovate, ca. 6
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="BDCC821749B7E9FA6B53A4CE9DD63DB8" originalValue="×">x</normalizedToken>
3 mm, acute at apex, abaxially pubescent; lateral sepals greenish-brown, elliptic, oblique, ca. 8
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B5789FD36611B26E8E90A3C81063BE23" originalValue="×">x</normalizedToken>
4 mm, acute at apex, abaxially pubescent. Petals obliquely ovate-falcate, ca. 6
<normalizedToken originalValue="×">x</normalizedToken>
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3 mm, membranous, glabrous; lip light yellow, T-shaped, shallowly grooved along the mid-line, ca. 1 cm long; hypochile bisaccate, sac sub-globose, ca. 2 mm in diameter, containing a low median keel and a small, erect, blade-like and apically emarginate callus on each side; mesochile ca. 4 mm long, with a pair of dentate-pectinate flanges, each flange composed of three narrow filaments, ca. 4 mm long and one broad blade-like posterior filament, ca. 3 mm long; epichile bilobed, lobes erect, diverging at obtuse angle to each other, elliptic, ca. 5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="EE7A6EEB310380DD771EE3C33A160CE6" originalValue="×">x</normalizedToken>
3 mm, margin involute and consequently resulting in concave lobes, obtuse at apex. Column ca. 1 mm long, stout; anther acuminate in front, ca. 4 mm long; pollinia 2, clavate; stigma lobes confluent; ovary and pedicel cylindric, twisted, sparsely pubescent.
</paragraph>
<caption pageId="2" pageNumber="21">
<paragraph pageId="2" pageNumber="21">
<caption id="7E278AC0681954EA6631DEBA69DF7EDB" pageId="2" pageNumber="21">
<paragraph id="7C959DE55806F35F39D1A5F3E1D0C093" pageId="2" pageNumber="21">
Figure 1.
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
<taxonomicName id="4B70A5DA729A5BEF19668566F7DB33E5" class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
X.H.Jin, L.A.Ye &amp; A.T.Mu. A Habit of
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
<taxonomicName id="CE44AC3A0C68E85A8D50014ED87C2503" class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
B Front view of flower, showing lip epichile with a pair of erect and concave lobes C Hypochile of
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
<taxonomicName id="0ACFBC3195492D134805F6867C26F3FE" class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
, indicating small, erect, blade-like, emarginate callus within each sac D Dissected flower, showing pedicel and ovary, column, sepals, petals, lip and a pair of clavate pollinia E Dorsal view of flower, showing dorsal sepal forming a hood with petals. Photographed by X.H. Jin.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="etymology">
<paragraph pageId="2" pageNumber="21">Etymology.</paragraph>
<paragraph pageId="2" pageNumber="21">The new species is named after Putao, the northernmost town of Myanmar, near which it was discovered in a vast area of undisturbed mountain forest.</paragraph>
<subSubSection id="E05F4001836EC1F17BBCC8F5B3F09621" pageId="2" pageNumber="21" type="etymology">
<paragraph id="B1D3C0D562DA9FC0DB844DF13F4A7443" pageId="2" pageNumber="21">Etymology.</paragraph>
<paragraph id="B4B84AFC31029FC47DE51074BE50E0AF" pageId="2" pageNumber="21">The new species is named after Putao, the northernmost town of Myanmar, near which it was discovered in a vast area of undisturbed mountain forest.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="distribution">
<paragraph pageId="2" pageNumber="21">Distribution and habitat.</paragraph>
<paragraph pageId="2" pageNumber="21">
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
<subSubSection id="E05D9E857FD13E014201FCA1549D908F" pageId="2" pageNumber="21" type="distribution">
<paragraph id="830C3C7EFBD2F405F6FEEE38B293B88C" pageId="2" pageNumber="21">Distribution and habitat.</paragraph>
<paragraph id="C7E7FBA777DFDA53D996D14EA1E3B85E" pageId="2" pageNumber="21">
<taxonomicName id="B068736582B2C058D5C89DCAF46D095D" class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
grows in shaded and damp humus in humid, broad-leaved, evergreen forest, at an elevation of about 1500-2000 m. At present,
<taxonomicName lsidName="O. putaoensis" pageId="2" pageNumber="21" rank="species" species="putaoensis">O. putaoensis</taxonomicName>
<taxonomicName id="1F735BF14F5333C35CFEAE2B2CB58C06" lsidName="O. putaoensis" pageId="2" pageNumber="21" rank="species" species="putaoensis">O. putaoensis</taxonomicName>
is only known from the type locality.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="21" type="conservation status">
<paragraph pageId="2" pageNumber="21">Conservation status.</paragraph>
<paragraph pageId="2" pageNumber="21">
<subSubSection id="08E06B9E1A019061BCA00A588A3A244C" pageId="2" pageNumber="21" type="conservation status">
<paragraph id="2453610276CBAAC9FA7F09DBDC257EFE" pageId="2" pageNumber="21">Conservation status.</paragraph>
<paragraph id="B232923938C1D05292F095272F66A0DC" pageId="2" pageNumber="21">
Least Concern (LC).
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
<taxonomicName id="60AE9B23635A9E1F32577A4B9749B27A" class="Liliopsida" family="Orchidaceae" genus="Odontochilus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Odontochilus putaoensis" order="Asparagales" pageId="2" pageNumber="21" phylum="Tracheophyta" rank="species" species="putaoensis">Odontochilus putaoensis</taxonomicName>
was collected in the Hponkanrazi Wildlife Sanctuary, Putao Township, Kachin State, northern Myanmar. Until now, only one population, consisting of ca. 200 individuals, has been discovered in the vast reserve of 2704 km2. As there is no threat currently affecting the quality of its habitat and there is also a considerable number of mature individuals, the species is here preliminarily assigned a status of Least Concern (LC) according to the guidelines for using the IUCN Red List Categories and Criteria (
<bibRefCitation author="Geissmann, T" journalOrPublisher="American Journal of Primatology" pageId="4" pageNumber="23" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2017">IUCN Standards and Petitions Subcommittee, 2017</bibRefCitation>
<bibRefCitation id="039CC54153D7FB16AE97E00AAAA39C09" author="Geissmann, T" journalOrPublisher="American Journal of Primatology" pageId="4" pageNumber="23" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2017">IUCN Standards and Petitions Subcommittee, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>

106
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<paragraph pageId="5" pageNumber="6">
<taxonomicName LSID="urn:lsid:zoobank.org:act:EFBC2841-A6A2-4BED-93D4-E1EBCAADF757" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma maculatum" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maculatum">Eustrongylosoma maculatum</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="6">sp. n.</taxonomicNameLabel>
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<paragraph id="F30298850C1D3E4B964C03B38ECF7A40" pageId="5" pageNumber="6">
<taxonomicName id="F13F9E01D058D99F5FCDA5D69DEA8028" ID-CoL="3DBGS" LSID="urn:lsid:zoobank.org:act:EFBC2841-A6A2-4BED-93D4-E1EBCAADF757" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma maculatum" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maculatum">Eustrongylosoma maculatum</taxonomicName>
<taxonomicNameLabel id="19DB607DE3FC3AED8BC15D2D743533A1" pageId="5" pageNumber="6">sp. n.</taxonomicNameLabel>
Figs 15-19
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="type material">
<paragraph pageId="5" pageNumber="6">Type material.</paragraph>
<paragraph pageId="5" pageNumber="6">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin area, September 1975, leg. B.S.E.</paragraph>
<subSubSection id="407F176297F92C8E58B1EBAC57AA06D5" pageId="5" pageNumber="6" type="type material">
<paragraph id="6997F759951EA2F63C3EFD5D6C75B4D7" pageId="5" pageNumber="6">Type material.</paragraph>
<paragraph id="BFF00CFCA4BC517304CC623129ECF3A5" pageId="5" pageNumber="6">Holotype♂(NMNHS),Papua New Guinea, West Sepik Province, Telefomin area, September 1975, leg. B.S.E.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="name">
<paragraph pageId="5" pageNumber="6">Name.</paragraph>
<paragraph pageId="5" pageNumber="6">To reflect the wide spots/bands in the front halves of the collum, following metaterga and epiproct.</paragraph>
<subSubSection id="06C33D0971120F142C4F1A7136AC8932" pageId="5" pageNumber="6" type="name">
<paragraph id="9F9DCE2D3E7699C8140D7AF59B49A591" pageId="5" pageNumber="6">Name.</paragraph>
<paragraph id="1D364AEE8CBB9ED0F7D517FC2FA558F0" pageId="5" pageNumber="6">To reflect the wide spots/bands in the front halves of the collum, following metaterga and epiproct.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="diagnosis">
<paragraph pageId="5" pageNumber="6">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="6">Distinguished by the very poorly developed paraterga, coupled with a non-fringed tip of the solenophore (see also Key below).</paragraph>
<subSubSection id="884DE8A515C305885DAE43875804C268" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="BB63E089DE4FB97D06F22C5866CED4A6" pageId="5" pageNumber="6">Diagnosis.</paragraph>
<paragraph id="7A7B67D34DF970E1A7E3EA793807CD5E" pageId="5" pageNumber="6">Distinguished by the very poorly developed paraterga, coupled with a non-fringed tip of the solenophore (see also Key below).</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph pageId="5" pageNumber="6">Description.</paragraph>
<paragraph pageId="5" pageNumber="6">Length ca 24 mm, width of pro- and metazona 2.0 and 2.4 mm, respectively. Coloration light yellow-brown with front halves of collum, following metaterga and epiproct, as well as entire head and antennae contrasting brown, thus creating a clear cingulate pattern.</paragraph>
<paragraph lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="2BC633F483682C4EEAB2065EDB7DBC9C" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="5640B7DC59B46CD682FC479A62A8B8E4" pageId="5" pageNumber="6">Description.</paragraph>
<paragraph id="4B4D875CD4055D7556B8D823273E9D26" pageId="5" pageNumber="6">Length ca 24 mm, width of pro- and metazona 2.0 and 2.4 mm, respectively. Coloration light yellow-brown with front halves of collum, following metaterga and epiproct, as well as entire head and antennae contrasting brown, thus creating a clear cingulate pattern.</paragraph>
<paragraph id="A03B8D95E45D6CCE019809321F7A6D86" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
In width, head&gt; collum = segment 3 = 4 &lt;2 &lt;5-16; thereafter body gently tapering towards telson. Head densely setose, only vertex bare. Antennae rather short, clavate, reaching beyond segment 2 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining. Paraterga of postcollum segments very poorly developed (Fig. 15), set low (at about 1/2-1/3 midbody height); paratergum 2 a low ridge with a few wrinkles above; paraterga 3 and 4 virtually missing, traceable only as 3-4 similar wrinkles; following paraterga retained only as a dorsal sulcus/line (Fig. 15), barely vi
<pageBreakToken pageId="6" pageNumber="7" start="start">sible</pageBreakToken>
<pageBreakToken id="5CC4037FC29C143C37E546AF151BBFFB" pageId="6" pageNumber="7" start="start">sible</pageBreakToken>
in segment 19. Ozopores lateral, superficial. Tergal setae mostly abraded, retained ones about 1/4 the length of metaterga; setation pattern vague, sometimes traceable as 1+1 paramedian insertion points in front row. Axial line wanting. Transverse sulcus on metaterga highly superficial, incomplete, far from reaching bases of paraterga, visible only in metaterga 9-17. Stricture between pro- and metazona rather distinctly beaded. Pleurosternal carinae very small, complete crests or nearly so only in segments 2-4. Epiproct (Fig. 16) short, flattened dorsoventrally, digitiform; apical papillae very large, spiniform, directed more caudally than ventrally; subapical papillae missing. Hypoproct subtriangular, apex rounded, setigerous paramedian knobs at caudal margin small and rather faintly separated.
</paragraph>
<paragraph pageId="6" pageNumber="7">Sterna densely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 17). Legs relatively long, midbody ones ca 1.5-1.6 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal brushes increasingly well reduced towards legs of segment 10.</paragraph>
<paragraph pageId="6" pageNumber="7">Gonopods as in Figs 18 &amp; 19. Distal part of solenophore split into a distinct frontomesal tooth (k) and a longer, somewhat curved lateral branch (j) with two subapical denticles and a lobule. Femorite with a large, rounded, apicolateral lobe (l).</paragraph>
<caption pageId="6" pageNumber="7">
<paragraph pageId="6" pageNumber="7">
<paragraph id="037E537F48B8C939CDC60639D411C5C0" pageId="6" pageNumber="7">Sterna densely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 17). Legs relatively long, midbody ones ca 1.5-1.6 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal brushes increasingly well reduced towards legs of segment 10.</paragraph>
<paragraph id="D93F571E0922561B6803BEB9A5E6D295" pageId="6" pageNumber="7">Gonopods as in Figs 18 &amp; 19. Distal part of solenophore split into a distinct frontomesal tooth (k) and a longer, somewhat curved lateral branch (j) with two subapical denticles and a lobule. Femorite with a large, rounded, apicolateral lobe (l).</paragraph>
<caption id="10779BBF146A17221B363592AED07EDC" pageId="6" pageNumber="7">
<paragraph id="BD7F9B80A310D8DA7E71AE5F0184C185" pageId="6" pageNumber="7">
Figures 15-19.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma maculatum" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maculatum">Eustrongylosoma maculatum</taxonomicName>
<taxonomicName id="B8D8171A188754CFF65ECA467EB19ECC" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma maculatum" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maculatum">Eustrongylosoma maculatum</taxonomicName>
sp. n., holotype. 15 segment 10, lateral view 16 tip of epiproct, dorsal view 17 sternal lobe between coxae 4, caudal view 18, 19 left gonopod, mesal and lateral views, respectively. Scale bars: 1.0 (15-17) and 0.5 mm (18 &amp; 19).
</paragraph>
</caption>