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<mods:title id="86F5C110F8785938E0A890505544CDC7">Ninetis subtilissima Simon, 1890 (Araneae: Pholcidae): Redescription and SEM Ultrastructure</mods:title>
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<mods:namePart id="8BEBD6E8BF03A97B18E201041FCB9C05">HUBER, BERNHARD A.</mods:namePart>
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<mods:namePart id="96EED1D71140038FCF6FD330C0F4ABC4">VAN HARTEN, ANTONIUS</mods:namePart>
<mods:affiliation id="991567A3C999B79F09380DC2DFBFA64D">General Department of Plant Protection, P. O. Box 26, Sana, Republic of Yemen.</mods:affiliation>
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<mods:date id="318D2ED34017C65032033799D686767F">2001</mods:date>
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<subSubSection id="C3656555FFA41823FF7CF232E865FCC9" box="[191,583,798,821]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BC036DEFFA41823FF7CF232E865FCC9" blockId="1.[191,583,798,821]" box="[191,583,798,821]" pageId="1" pageNumber="2">
<taxonomicName id="4C7F4D5DFFA41823FF7CF232E865FCC9" ID-CoL="47FFR" authority="Simon, 1890" authorityName="Simon" authorityYear="1890" box="[191,583,798,821]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="B90BEACCFFA41823FF7CF232EB85FCC9" box="[191,423,798,821]" italics="true" pageId="1" pageNumber="2">Ninetis subtilissima</emphasis>
<bibRefCitation id="EFEE4B2FFFA41823FE73F232E865FCC9" author="Simon, E." box="[432,583,798,821]" journalOrPublisher="Ann. Soc. Entomol. France" pageId="1" pageNumber="2" pagination="77 - 124" part="6" refId="ref1958" refString="Simon, E. 1890. Etude sur les arachnides de l'Yemen. Ann. Soc. Entomol. France 6 (10): 77 - 124." title="Etude sur les arachnides de l'Yemen" type="journal article" year="1890">Simon, 1890</bibRefCitation>
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</paragraph>
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<subSubSection id="C3656555FFA41823FE89F212EBE4FCAD" box="[330,454,830,849]" pageId="1" pageNumber="2" type="description">
<paragraph id="8BC036DEFFA41823FE89F212EBE4FCAD" blockId="1.[330,454,830,849]" box="[330,454,830,849]" pageId="1" pageNumber="2">
<heading id="D08881B2FFA41823FE89F212EBE4FCAD" box="[330,454,830,849]" centered="true" fontSize="36" level="3" pageId="1" pageNumber="2" reason="7">
<figureCitation id="13442A5BFFA41823FE89F212EBE4FCAD" box="[330,454,830,849]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStartId-0="1.[147,195,579,600]" captionStartId-1="2.[150,198,1381,1402]" captionStartId-2="3.[154,202,1576,1597]" captionStartId-3="4.[150,198,1635,1656]" captionStartId-4="5.[154,202,1317,1338]" captionTargetBox-0="[138,1223,187,558]" captionTargetBox-1="[156,1199,193,1356]" captionTargetBox-2="[136,1224,187,1555]" captionTargetBox-3="[132,1220,187,1614]" captionTargetBox-4="[134,1227,187,1296]" captionTargetId-0="figure-13@1.[138,1223,187,558]" captionTargetId-1="figure-15@2.[150,1204,187,1360]" captionTargetId-2="figure-11@3.[136,1224,187,1555]" captionTargetId-3="figure-12@4.[132,1220,187,1614]" captionTargetId-4="figure-13@5.[134,1227,187,1296]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionText-0="Figs. 1, 2. Ninetis subtilissima Simon. 1. Male 2. Female. Photos courtesy of B. Thaler­Knoflach." captionText-1="Figs. 38. Ninetis subtilissima Simon. 3. Left male palp, prolateral view. 4. Left male palp, retrolateral view. 5. Male chelicerae, frontal view. 6. Male chelicerae, lateral view. 7. Epigynum, ventral view. 8. Epigynum, dorsal view." captionText-2="Figs. 915. Ninetis subtilissima Simon. 9. Female prosoma, frontal view. 10. Female cheliceral laminae, frontal view. 11. Stridulatory files on male chelicera. 12. Location of cheliceral gland openings (arrow) near tip of fang. 13. Detail of fig. 12, showing two pores. 14. Female palpal tarsal organ (TO). 15. Male palpal tarsal organ (TO)." captionText-3="Figs. 1623. Ninetis subtilissima Simon. 16, 17. Male ALS showing widened (W), pointed (P), and cylindrically shaped (C) spigots. 18. Male PMS, showing two spigots (arrows). 19. Putative remnant of colulus (arrow) and sensillum between ALS. 20. Tip of female palp. 21. Female opisthosoma, showing epigynum and spinnerets (arrow points to pocket). 22. Hairs on anal tubercle. 23. Pocket on epigynum, posterior view." captionText-4="Figs. 2428. Ninetis subtilissima Simon. 24. Hairs on tibiae (female tibia 2). 25. Hairs on tarsi (female tarsus 2). 26. Female tarsus 2 (note the apparent absence of pseudosegments). 27. Trichobothrium base (female palpal tibia). 28. Tarsal claws (female tarsus 1)." figureDoi-0="http://doi.org/10.5281/zenodo.5370750" figureDoi-1="http://doi.org/10.5281/zenodo.5370752" figureDoi-2="http://doi.org/10.5281/zenodo.5370754" figureDoi-3="http://doi.org/10.5281/zenodo.5370756" figureDoi-4="http://doi.org/10.5281/zenodo.5370758" httpUri-0="https://zenodo.org/record/5370750/files/figure.png" httpUri-1="https://zenodo.org/record/5370752/files/figure.png" httpUri-2="https://zenodo.org/record/5370754/files/figure.png" httpUri-3="https://zenodo.org/record/5370756/files/figure.png" httpUri-4="https://zenodo.org/record/5370758/files/figure.png" pageId="1" pageNumber="2">Figures 128</figureCitation>
</heading>
</paragraph>
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<paragraph id="8BC036DEFFA41823FF43F249EB10FC03" blockId="1.[128,657,869,1023]" pageId="1" pageNumber="2">
<taxonomicName id="4C7F4D5DFFA41823FF43F249E83EFC86" authority="Simon, 1890: 96" authorityName="Simon" authorityPageNumber="96" authorityYear="1890" box="[128,540,869,890]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="B90BEACCFFA41823FF43F249EB76FC86" box="[128,340,869,890]" italics="true" pageId="1" pageNumber="2">Ninetis subtilissima</emphasis>
<bibRefCitation id="EFEE4B2FFFA41823FEA3F249E83EFC86" author="Simon, E." box="[352,540,869,890]" journalOrPublisher="Ann. Soc. Entomol. France" pageId="1" pageNumber="2" pagination="77 - 124" part="6" refId="ref1958" refString="Simon, E. 1890. Etude sur les arachnides de l'Yemen. Ann. Soc. Entomol. France 6 (10): 77 - 124." title="Etude sur les arachnides de l'Yemen" type="journal article" year="1890">Simon, 1890: 96</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFEE4B2FFFA41823FD85F249EB1DFC69" author="Simon, E." bookContentInfo="Paris" pageId="1" pageNumber="2" pagination="256 - 488" part="1" refId="ref1987" refString="1893. Histoire naturelle des araignees 1 (2): 256 - 488. Paris." title="Histoire naturelle des araignees" type="book chapter" year="1893">Simon, 1893: 486487</bibRefCitation>
, figs. 487489.
<bibRefCitation id="EFEE4B2FFFA41823FDD3F2ACE8AFFC69" author="Fage, L." box="[528,653,896,917]" journalOrPublisher="Arch. Zool. Exper. Gen." pageId="1" pageNumber="2" pagination="97 - 162" part="5" refId="ref1814" refString="Fage, L. Fig. 29. Ninetis spp., diagnostic characters. Upper row: Left procursi and bulbs, retrolateral views. Middle row: Epigyna, dorsal views. Low- er row: Chelicerae, lateral views. Illustrations in each row drawn to same scale. Drawings of N. minuta and N. namibiae copied from Huber (2000). 1912. Etudes sur les araingees cavernicoles. I. Revision des Ochyroceratidae. Arch. Zool. Exper. Gen. (5) 10: 97 - 162, pls. IV - XII." title="Etudes sur les araingees cavernicoles. I. Revision des Ochyroceratidae" type="journal article" year="1912">Fage, 1912</bibRefCitation>
: figs. 3a (copied from
<bibRefCitation id="EFEE4B2FFFA41823FEBAF2B6E823FC53" author="Simon, E." bookContentInfo="Paris" box="[377,513,922,943]" pageId="1" pageNumber="2" pagination="256 - 488" part="1" refId="ref1987" refString="1893. Histoire naturelle des araignees 1 (2): 256 - 488. Paris." title="Histoire naturelle des araignees" type="book chapter" year="1893">Simon, 1893</bibRefCitation>
), 134.
<bibRefCitation id="EFEE4B2FFFA41823FD9FF2B6EB69FC36" author="Bristowe, W. S." journalOrPublisher="Proc. Zool. Soc" pageId="1" pageNumber="2" pagination="285 - 321" part="B" refId="ref1783" refString="Bristowe, W. S. 1938. The classification of spiders. Proc. Zool. Soc., Ser. B (1938): 285 - 321." title="The classification of spiders" type="journal article" year="1938">Bristowe, 1938: 310</bibRefCitation>
, figs. 2, 7 (copied from
<bibRefCitation id="EFEE4B2FFFA41823FD94F299EAEDFC19" author="Fage, L." journalOrPublisher="Arch. Zool. Exper. Gen." pageId="1" pageNumber="2" pagination="97 - 162" part="5" refId="ref1814" refString="Fage, L. Fig. 29. Ninetis spp., diagnostic characters. Upper row: Left procursi and bulbs, retrolateral views. Middle row: Epigyna, dorsal views. Low- er row: Chelicerae, lateral views. Illustrations in each row drawn to same scale. Drawings of N. minuta and N. namibiae copied from Huber (2000). 1912. Etudes sur les araingees cavernicoles. I. Revision des Ochyroceratidae. Arch. Zool. Exper. Gen. (5) 10: 97 - 162, pls. IV - XII." title="Etudes sur les araingees cavernicoles. I. Revision des Ochyroceratidae" type="journal article" year="1912">Fage, 1912</bibRefCitation>
, and
<bibRefCitation id="EFEE4B2FFFA41823FED0F2FCEB80FC19" author="Simon, E." bookContentInfo="Paris" box="[275,418,976,997]" pageId="1" pageNumber="2" pagination="256 - 488" part="1" refId="ref1987" refString="1893. Histoire naturelle des araignees 1 (2): 256 - 488. Paris." title="Histoire naturelle des araignees" type="book chapter" year="1893">Simon, 1893</bibRefCitation>
).
<bibRefCitation id="EFEE4B2FFFA41823FE16F2FCE8AEFC19" author="Huber" box="[469,652,976,997]" firstAuthor="Huber" page="82" pageId="1" pageNumber="2" pagination="23" refId="ref1110" refString="FEMALE (Ja'ar): In general very similar to male. Carapace width 0.48; tibia 1: 0.60. Palpal tarsal organ as in figure 14. Tip of palp with strong spine (fig. 20). Epigynum consisting of posterior plate and large frontal plate with median pocket opening posteriorly (figs. 7, 21, 23); dorsal view as in figure 8. See Huber (2000) for redescription of female syntypes." type="journal article" year="2000">Huber, 2000: 82</bibRefCitation>
, figs. 310314.
</paragraph>
</subSubSection>
<subSubSection id="C3656555FFA41823FF5EF538EB01FB27" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BC036DEFFA41823FF5EF538EB01FB27" blockId="1.[128,657,1044,1741]" pageId="1" pageNumber="2">
<materialsCitation id="3B173C83FFA41823FF5EF538EB3DFB27" collectionCode="MNHN" collectorName="E. Simon" country="Yemen" determinerName="Huber, 2000" location="Al'Adan" pageId="1" pageNumber="2" specimenCode="10788" specimenCount="2" specimenCount-adult="1" stateProvince="'Adan" typeStatus="syntype">
TYPES: About four to
<specimenCount id="9D79FD57FFA41823FE58F538EBE5FBD7" box="[411,455,1044,1067]" pageId="1" pageNumber="2">five</specimenCount>
adult
<specimenCount id="9D79FD57FFA41823FDD3F538E87DFBD7" box="[528,607,1044,1067]" pageId="1" pageNumber="2" type="female">female</specimenCount>
and several juvenile
<typeStatus id="54C4887CFFA41823FE9AF51DEBE4FBB4" box="[345,454,1073,1096]" pageId="1" pageNumber="2" type="syntype">syntypes</typeStatus>
from
<location id="8EA06005FFA41823FDDEF51DE8AFFBB4" LSID="urn:lsid:plazi:treatment:03D687C8FFA41824FF7CF232E918FE3F:8EA06005FFA41823FDDEF51DE8AFFBB4" box="[541,653,1073,1096]" country="Yemen" name="Al'Adan" pageId="1" pageNumber="2" stateProvince="'Adan">AlAdan</location>
,
<collectingCountry id="F368764EFFA41823FF43F562EAF7FB99" box="[128,213,1102,1125]" name="Yemen" pageId="1" pageNumber="2">Yemen</collectingCountry>
; no date (
<collectorName id="268A5308FFA41823FEADF562EBCEFB99" box="[366,492,1102,1125]" pageId="1" pageNumber="2">E. Simon</collectorName>
), in
<collectionCode id="ED6EAE1BFFA41823FDFBF562E8B3FB99" box="[568,657,1102,1125]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="1" pageNumber="2">MNHN</collectionCode>
(
<specimenCode id="DBD99EA5FFA41823FF49F540EAFCFB7F" box="[138,222,1132,1155]" pageId="1" pageNumber="2">10788</specimenCode>
); with Simons handwritten label 10788 Ninet. subtilissima E.S. Aden!, examined (see redescription of this material in
<bibRefCitation id="EFEE4B2FFFA41823FF43F5E8EB35FB27" author="Huber" box="[128,279,1220,1243]" firstAuthor="Huber" pageId="1" pageNumber="2" pagination="23" refId="ref1110" refString="FEMALE (Ja'ar): In general very similar to male. Carapace width 0.48; tibia 1: 0.60. Palpal tarsal organ as in figure 14. Tip of palp with strong spine (fig. 20). Epigynum consisting of posterior plate and large frontal plate with median pocket opening posteriorly (figs. 7, 21, 23); dorsal view as in figure 8. See Huber (2000) for redescription of female syntypes." type="journal article" year="2000">
<determinerName id="7E1CDA1AFFA41823FF43F5E8EB35FB27" box="[128,279,1220,1243]" pageId="1" pageNumber="2">Huber, 2000</determinerName>
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656555FFA41823FF5EF5CDEBB6FA54" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BC036DEFFA41823FF5EF5CDEBB6FA54" blockId="1.[128,657,1044,1741]" pageId="1" pageNumber="2">
DIAGNOSIS: This species is easily distinguished from the two known congeners (
<taxonomicName id="4C7F4D5DFFA41823FDB5F5D3EB6BFACF" authority="(Berland)" baseAuthorityName="Berland" baseAuthorityYear="1920" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B90BEACCFFA41823FDB5F5D3EAF3FACF" italics="true" pageId="1" pageNumber="2">N. minuta</emphasis>
(Berland)
</taxonomicName>
and
<taxonomicName id="4C7F4D5DFFA41823FE40F430E846FACF" authority="Huber" authorityName="Huber" authorityYear="2000" box="[387,612,1308,1331]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="namibiae">
<emphasis id="B90BEACCFFA41823FE40F430E833FACF" box="[387,529,1308,1331]" italics="true" pageId="1" pageNumber="2">N. namibiae</emphasis>
Huber
</taxonomicName>
) by the short, pointed procursus and the shapes of the bulbal projections, the shape of the male cheliceral apophyses, and the shape of the epigynum (fig. 29).
</paragraph>
</subSubSection>
<subSubSection id="C3656555FFA41820FF5EF482E908F9C7" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="8BC036DEFFA41823FF5EF482EE3EF931" blockId="1.[128,657,1044,1741]" lastBlockId="1.[704,1234,665,1741]" pageId="1" pageNumber="2">
MALE (Jaar): Total length 0.9; carapace width 0.45. Leg 1: 2.28 (0.61 + 0.16 + 0.57 + 0.60 + 0.33), tibia 2: 0.47, tibia 3: 0.37, tibia 4: 0.67; tibia 1 l/d: 9.5. Habitus as in figs. 1 and 2 (see also drawings of the very similar
<taxonomicName id="4C7F4D5DFFA41823FF18F76DEB72F9A4" baseAuthorityName="Berland" baseAuthorityYear="1920" box="[219,336,1601,1624]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B90BEACCFFA41823FF18F76DEB72F9A4" box="[219,336,1601,1624]" italics="true" pageId="1" pageNumber="2">N. minuta</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFEE4B2FFFA41823FEBAF76DE82DF9A4" author="Huber" box="[377,527,1601,1624]" firstAuthor="Huber" pageId="1" pageNumber="2" pagination="23" refId="ref1110" refString="FEMALE (Ja'ar): In general very similar to male. Carapace width 0.48; tibia 1: 0.60. Palpal tarsal organ as in figure 14. Tip of palp with strong spine (fig. 20). Epigynum consisting of posterior plate and large frontal plate with median pocket opening posteriorly (figs. 7, 21, 23); dorsal view as in figure 8. See Huber (2000) for redescription of female syntypes." type="journal article" year="2000">Huber, 2000</bibRefCitation>
: figs. 315 318). Prosoma and legs ochre yellow; opisthosoma gray, with white mark above spinnerets and very indistinct darker spots dorsally. Carapace without thoracic furrow (cf. female: fig. 9). Eye pattern as in female (cf. fig. 9); diameter (in µm; ± 3 µm) AME 30, ALE 65, PME 50, PLE 60; distance PME­PME 50, PME­ALE 30. Chelicerae as in figures 5 and 6, with pair of apophyses frontally and very fine stridulatory ridges laterally (fig. 11); cheliceral laminae as in female (cf. fig. 10); cheliceral glands open through pores at slightly modified fields close to tips of fangs (figs. 12, 13). Sternum with small humps near bases of coxae 1. Palps as figures 3 and 4, with simple procursus that is pointed distally, and genital bulb with two processes: embolus dorsally and pointed (slightly hooked) apophysis ventrally; palpal tarsal organ capsulate with small orifice (fig. 15). Legs without spines, without curved hairs, with very few short vertical hairs; normal mechanoreceptive hairs of variable shape (e.g., figs. 24, 25); three trichobothria on tibiae, one on metatarsi; trichobothria with simple capsulate basis (fig. 27); retrolateral trichobothrium on tibia 1 at 58%; tarsus 1 apparently with 5 pseudosegments, barely visible as slightly darker rings in light microscope, but not discernible in SEM (figs. 25, 26). Three tarsal claws: superior claws with about 12 teeth, inferior claw with two teeth (fig. 28). Opisthosoma globular, with long hair dorsally. Three pairs of spinnerets, colulus possibly represented by modified area between ALS (fig. 19); ALS with one widened, one pointed, and several cylindrically shaped spigots (figs. 16, 17); PMS with one pair of cylindrically shaped spigots (fig. 18); PLS without spigots. Anal tubercle with multibranched hairs (fig. 22).
</paragraph>
<caption id="DF006656FFA71820FF55F449EBFBFA4C" ID-DOI="http://doi.org/10.5281/zenodo.5370752" ID-Zenodo-Dep="5370752" httpUri="https://zenodo.org/record/5370752/files/figure.png" pageId="2" pageNumber="3" startId="2.[150,198,1381,1402]" targetBox="[156,1199,193,1356]" targetPageId="2">
<paragraph id="8BC036DEFFA71820FF55F449EBFBFA4C" blockId="2.[124,1229,1381,1456]" pageId="2" pageNumber="3">
Figs. 38.
<taxonomicName id="4C7F4D5DFFA71820FED8F44AE863FA86" authority="Simon." authorityName="Simon." box="[283,577,1381,1403]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="B90BEACCFFA71820FED8F44AEBCEFA87" box="[283,492,1382,1403]" italics="true" pageId="2" pageNumber="3">Ninetis subtilissima</emphasis>
Simon.
</taxonomicName>
<emphasis id="B90BEACCFFA71820FD89F449E87CFA86" bold="true" box="[586,606,1381,1402]" pageId="2" pageNumber="3">3.</emphasis>
Left male palp, prolateral view.
<emphasis id="B90BEACCFFA71820FC04F449E9F8FA86" bold="true" box="[967,986,1381,1402]" pageId="2" pageNumber="3">4.</emphasis>
Left male palp, retrolateral view.
<emphasis id="B90BEACCFFA71820FECFF4ACEB02FA69" bold="true" box="[268,288,1408,1429]" pageId="2" pageNumber="3">5.</emphasis>
Male chelicerae, frontal view.
<emphasis id="B90BEACCFFA71820FDBFF4ACE8ADFA69" bold="true" box="[636,655,1408,1429]" pageId="2" pageNumber="3">6.</emphasis>
Male chelicerae, lateral view.
<emphasis id="B90BEACCFFA71820FC24F4ACE9D8FA69" bold="true" box="[999,1018,1408,1429]" pageId="2" pageNumber="3">7.</emphasis>
Epigynum, ventral view.
<emphasis id="B90BEACCFFA71820FF7DF4B6EAF3FA53" bold="true" box="[190,209,1434,1455]" pageId="2" pageNumber="3">8.</emphasis>
Epigynum, dorsal view.
</paragraph>
</caption>
<paragraph id="8BC036DEFFA71820FF5AF72AE908F9C7" blockId="2.[124,653,1542,1741]" lastBlockId="2.[700,1229,1542,1741]" pageId="2" pageNumber="3">
FEMALE (Jaar): In general very similar to male. Carapace width 0.48; tibia 1: 0.60. Palpal tarsal organ as in figure 14. Tip of palp with strong spine (fig. 20). Epigynum consisting of posterior plate and large frontal plate with median pocket opening posteriorly (figs. 7,
<part id="FFDB9C43FFA71820FF20F79AEB20F931" box="[227,258,1718,1741]" pageId="2" pageNumber="3" type="volume">21</part>
,
<pagination id="B2B6C82AFFA71820FED0F79AEB16F931" box="[275,308,1718,1741]" pageId="2" pageNumber="3" type="pagination">23</pagination>
); dorsal view as in figure 8.
<bibRefCitation id="EFEE4B2FFFA71820FD7FF72AE9AFF9E1" author="See Huber" box="[700,909,1542,1565]" pageId="2" pageNumber="3" pagination="23" refId="ref1110" refString="FEMALE (Ja'ar): In general very similar to male. Carapace width 0.48; tibia 1: 0.60. Palpal tarsal organ as in figure 14. Tip of palp with strong spine (fig. 20). Epigynum consisting of posterior plate and large frontal plate with median pocket opening posteriorly (figs. 7, 21, 23); dorsal view as in figure 8. See Huber (2000) for redescription of female syntypes." type="journal article" year="2000">
See Huber (
<year id="FFD7778DFFA71820FC84F72AE9A6F9E1" box="[839,900,1542,1565]" pageId="2" pageNumber="3">2000</year>
)
</bibRefCitation>
for redescription of female syntypes.
</paragraph>
</subSubSection>
<subSubSection id="C3656555FFA71820FD1AF76DE9C9F931" pageId="2" pageNumber="3" type="biology_ecology">
<paragraph id="8BC036DEFFA71820FD1AF76DE9C9F931" blockId="2.[700,1229,1542,1741]" pageId="2" pageNumber="3">
BIOLOGY: The specimens from Jaar were shaken out of old
<collectingCountry id="F368764EFFA71820FC6BF772E9D3F989" box="[936,1009,1630,1653]" name="Sudan" pageId="2" pageNumber="3">Sudan</collectingCountry>
grass plants in a garden within the town. The spiders ran rapidly when disturbed and built flimsy webs when kept in glass tubes.
</paragraph>
</subSubSection>
<caption id="DF006656FFA61821FF59F704EBD1F971" ID-DOI="http://doi.org/10.5281/zenodo.5370754" ID-Zenodo-Dep="5370754" httpUri="https://zenodo.org/record/5370754/files/figure.png" pageId="3" pageNumber="4" startId="3.[154,202,1576,1597]" targetBox="[136,1224,187,1555]" targetPageId="3">
<paragraph id="8BC036DEFFA61821FF59F704EBD1F971" blockId="3.[128,1233,1576,1677]" pageId="3" pageNumber="4">
Figs. 915.
<taxonomicName id="4C7F4D5DFFA61821FEF2F704E87EF9C1" authority="Simon." authorityName="Simon." box="[305,604,1576,1597]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="B90BEACCFFA61821FEF2F704E826F9C1" box="[305,516,1576,1597]" italics="true" pageId="3" pageNumber="4">Ninetis subtilissima</emphasis>
Simon.
</taxonomicName>
<emphasis id="B90BEACCFFA61821FDABF704E859F9C1" bold="true" box="[616,635,1576,1597]" pageId="3" pageNumber="4">9.</emphasis>
Female prosoma, frontal view.
<emphasis id="B90BEACCFFA61821FC26F704EE27F9C1" bold="true" box="[997,1029,1576,1597]" pageId="3" pageNumber="4">10.</emphasis>
Female cheliceral laminae, frontal view.
<emphasis id="B90BEACCFFA61821FEACF76EEBB2F9AB" bold="true" box="[367,400,1602,1623]" pageId="3" pageNumber="4">11.</emphasis>
Stridulatory files on male chelicera.
<emphasis id="B90BEACCFFA61821FCD9F76EE918F9AB" bold="true" box="[794,826,1602,1623]" pageId="3" pageNumber="4">12.</emphasis>
Location of cheliceral gland openings (arrow) near tip of fang.
<emphasis id="B90BEACCFFA61821FE4EF771EB8CF98E" bold="true" box="[397,430,1629,1650]" pageId="3" pageNumber="4">13.</emphasis>
Detail of fig. 12, showing two pores.
<emphasis id="B90BEACCFFA61821FC89F771E949F98E" bold="true" box="[842,875,1629,1650]" pageId="3" pageNumber="4">14.</emphasis>
Female palpal tarsal organ (TO).
<emphasis id="B90BEACCFFA61821FF43F754EA83F971" bold="true" box="[128,161,1656,1677]" pageId="3" pageNumber="4">15.</emphasis>
Male palpal tarsal organ (TO).
</paragraph>
</caption>
<caption id="DF006656FFA11826FF55F74FEB3EF91E" ID-DOI="http://doi.org/10.5281/zenodo.5370756" ID-Zenodo-Dep="5370756" httpUri="https://zenodo.org/record/5370756/files/figure.png" pageId="4" pageNumber="5" startId="4.[150,198,1635,1656]" targetBox="[132,1220,187,1614]" targetPageId="4">
<paragraph id="8BC036DEFFA11826FF55F74FEB3EF91E" blockId="4.[124,1229,1634,1762]" pageId="4" pageNumber="5">
Figs. 1623.
<taxonomicName id="4C7F4D5DFFA11826FEF7F74FE875F984" authority="Simon." authorityName="Simon." box="[308,599,1635,1656]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="B90BEACCFFA11826FEF7F74FE826F984" box="[308,516,1635,1656]" italics="true" pageId="4" pageNumber="5">Ninetis subtilissima</emphasis>
Simon.
</taxonomicName>
<emphasis id="B90BEACCFFA11826FD9CF74EE88AF98B" bold="true" box="[607,680,1634,1655]" pageId="4" pageNumber="5">16, 17.</emphasis>
Male ALS showing widened (W), pointed (P), and cylindrically shaped (C) spigots.
<emphasis id="B90BEACCFFA11826FE1DF751EBDDF96E" bold="true" box="[478,511,1661,1682]" pageId="4" pageNumber="5">18.</emphasis>
Male PMS, showing two spigots (arrows).
<emphasis id="B90BEACCFFA11826FC10F751E9D1F96E" bold="true" box="[979,1011,1661,1682]" pageId="4" pageNumber="5">19.</emphasis>
Putative remnant of colulus (arrow) and sensillum between ALS.
<emphasis id="B90BEACCFFA11826FDA0F7B4E8A6F951" bold="true" box="[611,644,1688,1709]" pageId="4" pageNumber="5">20.</emphasis>
Tip of female palp.
<emphasis id="B90BEACCFFA11826FCA0F7B4E9A1F951" bold="true" box="[867,899,1688,1709]" pageId="4" pageNumber="5">21.</emphasis>
Female opisthosoma, showing epigynum and spinnerets (arrow points to pocket).
<emphasis id="B90BEACCFFA11826FD5CF79EE89DF93B" bold="true" box="[671,703,1714,1735]" pageId="4" pageNumber="5">22.</emphasis>
Hairs on anal tubercle.
<emphasis id="B90BEACCFFA11826FC00F79EE9C1F93B" bold="true" box="[963,995,1714,1735]" pageId="4" pageNumber="5">23.</emphasis>
Pocket on epigynum, posterior view.
</paragraph>
</caption>
<caption id="DF006656FFA01827FF59F409E969FA8C" ID-DOI="http://doi.org/10.5281/zenodo.5370758" ID-Zenodo-Dep="5370758" httpUri="https://zenodo.org/record/5370758/files/figure.png" pageId="5" pageNumber="6" startId="5.[154,202,1317,1338]" targetBox="[134,1227,187,1296]" targetPageId="5">
<paragraph id="8BC036DEFFA01827FF59F409E969FA8C" blockId="5.[128,1233,1317,1392]" pageId="5" pageNumber="6">
Figs. 2428.
<taxonomicName id="4C7F4D5DFFA01827FEFEF40AE845FAC6" authority="Simon." authorityName="Simon." box="[317,615,1317,1339]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="B90BEACCFFA01827FEFEF40AE832FAC7" box="[317,528,1318,1339]" italics="true" pageId="5" pageNumber="6">Ninetis subtilissima</emphasis>
Simon.
</taxonomicName>
<emphasis id="B90BEACCFFA01827FDB1F409E8B1FAC6" bold="true" box="[626,659,1317,1338]" pageId="5" pageNumber="6">24.</emphasis>
Hairs on tibiae (female tibia 2).
<emphasis id="B90BEACCFFA01827FBCEF409EE0CFAC6" bold="true" box="[1037,1070,1317,1338]" pageId="5" pageNumber="6">25.</emphasis>
Hairs on tarsi (female tarsus 2).
<emphasis id="B90BEACCFFA01827FE86F46CEB44FAA9" bold="true" box="[325,358,1344,1365]" pageId="5" pageNumber="6">26.</emphasis>
Female tarsus 2 (note the apparent absence of pseudosegments).
<emphasis id="B90BEACCFFA01827FBEFF46CEE6EFAA9" bold="true" box="[1068,1100,1344,1365]" pageId="5" pageNumber="6">27.</emphasis>
Trichobothrium base (female palpal tibia).
<emphasis id="B90BEACCFFA01827FE19F476EBD9FA93" bold="true" box="[474,507,1370,1391]" pageId="5" pageNumber="6">28.</emphasis>
Tarsal claws (female tarsus 1).
</paragraph>
</caption>
<subSubSection id="C3656555FFA01827FF5EF482EB75F989" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BC036DEFFA01827FF5EF482EB75F989" blockId="5.[128,657,1454,1741]" pageId="5" pageNumber="6">
<materialsCitation id="3B173C83FFA01827FF5EF482EBC6F9E1" collectingDate="1999-07-11" collectionCode="AMNH" collectorName="A. van Harten" country="Yemen" location="Ja'ar" municipality="Al'Adan" pageId="5" pageNumber="6" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="syntype">
MATERIAL EXAMINED:
<collectingCountry id="F368764EFFA01827FE6BF482E836FA39" box="[424,532,1454,1477]" name="Yemen" pageId="5" pageNumber="6">YEMEN</collectingCountry>
:
<collectingMunicipality id="6BA4ACA4FFA01827FDE0F482E8AFFA39" box="[547,653,1454,1477]" pageId="5" pageNumber="6">AlAdan</collectingMunicipality>
:
<typeStatus id="54C4887CFFA01827FF43F4E0EAC9FA1F" box="[128,235,1484,1507]" pageId="5" pageNumber="6">Syntypes</typeStatus>
above.
<location id="8EA06005FFA01827FE97F4E0EBB3FA1F" LSID="urn:lsid:plazi:treatment:03D687C8FFA41824FF7CF232E918FE3F:8EA06005FFA01827FE97F4E0EBB3FA1F" box="[340,401,1484,1507]" country="Yemen" municipality="Al'Adan" name="Ja'ar" pageId="5" pageNumber="6">Jaar</location>
:
<specimenCount id="9D79FD57FFA01827FE60F4E0EBF4FA1F" box="[419,470,1484,1507]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
<specimenCount id="9D79FD57FFA01827FE27F4E0E835FA1F" box="[484,535,1484,1507]" count="2" pageId="5" pageNumber="6" type="female">2 ♀</specimenCount>
,
<date id="FFC1101EFFA01827FDEEF4E0EA98F9FC" pageId="5" pageNumber="6" value="1999-07-11">
<collectingDate id="EF85E9F6FFA01827FDEEF4E0EA98F9FC" pageId="5" pageNumber="6" value="1999-07-11">July 11, 1999</collectingDate>
</date>
(
<collectorName id="268A5308FFA01827FF0EF4C5EB59F9FC" box="[205,379,1513,1536]" pageId="5" pageNumber="6">A. van Harten</collectorName>
), in American Museum of Natural History (
<collectionCode id="ED6EAE1BFFA01827FEBBF72AEBFAF9E1" box="[376,472,1542,1565]" collectionName="USA, New York, New York, American Museum of Natural History" pageId="5" pageNumber="6">AMNH</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B173C83FFA01827FE32F72AEB75F989" collectingDate="2000-05-15" collectionCode="AMNH" collectorName="A. van Harten" country="Yemen" location="Ja'ar" municipality="Al'Adan" pageId="5" pageNumber="6" specimenCount="19" specimenCount-female="17" specimenCount-male="2">
same locality and collector,
<date id="FFC1101EFFA01827FEE5F708EBEFF9C7" box="[294,461,1572,1595]" pageId="5" pageNumber="6" value="2000-05-15">
<collectingDate id="EF85E9F6FFA01827FEE5F708EBEFF9C7" box="[294,461,1572,1595]" pageId="5" pageNumber="6" value="2000-05-15">May 15, 2000</collectingDate>
</date>
,
<specimenCount id="9D79FD57FFA01827FE19F708E825F9C7" box="[474,519,1572,1595]" count="2" pageId="5" pageNumber="6" type="male">2 ♂</specimenCount>
<specimenCount id="9D79FD57FFA01827FDD3F708E86EF9C7" box="[528,588,1572,1595]" count="17" pageId="5" pageNumber="6" type="female">17 ♀</specimenCount>
, temporarily in collection of first author (eventually in
<collectionCode id="ED6EAE1BFFA01827FF2EF772EB69F989" box="[237,331,1630,1653]" collectionName="USA, New York, New York, American Museum of Natural History" pageId="5" pageNumber="6">AMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3656555FFA01824FF5EF750E918FE3F" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BC036DEFFA01827FF5EF750E9CDF9A4" blockId="5.[128,657,1454,1741]" lastBlockId="5.[704,1233,1454,1741]" pageId="5" pageNumber="6">
NOTES: In contrast to the two known congeners (
<taxonomicName id="4C7F4D5DFFA01827FF1DF7B5EBEBF94C" authority="(Berland)" baseAuthorityName="Berland" baseAuthorityYear="1920" box="[222,457,1689,1712]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B90BEACCFFA01827FF1DF7B5EB70F94C" box="[222,338,1689,1712]" italics="true" pageId="5" pageNumber="6">N. minuta</emphasis>
(Berland)
</taxonomicName>
and
<taxonomicName id="4C7F4D5DFFA01827FDC1F7B5EAEFF931" authority="Huber" authorityName="Huber" authorityYear="2000" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="namibiae">
<emphasis id="B90BEACCFFA01827FDC1F7B5E8B2F94C" box="[514,656,1689,1712]" italics="true" pageId="5" pageNumber="6">N. namibiae</emphasis>
Huber
</taxonomicName>
), but in agreement with most other ninetines,
<taxonomicName id="4C7F4D5DFFA01827FCE0F483E9F3FA3A" box="[803,977,1455,1478]" class="Arachnida" family="Pholcidae" genus="Ninetis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="subtilissima">
<emphasis id="B90BEACCFFA01827FCE0F483E9F3FA3A" box="[803,977,1455,1478]" italics="true" pageId="5" pageNumber="6">N. subtilissima</emphasis>
</taxonomicName>
has stridulatory ridges on the male chelicerae. However, the striation is extremely fine and difficult to see in light microscopy, and scanning electron micrographs of chelicerae of the other two species have not been made.
</paragraph>
<paragraph id="8BC036DEFFA01824FD1EF772E918FE3F" blockId="5.[704,1233,1454,1741]" lastBlockId="6.[700,1229,193,451]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
The figures herein differ in some respects from
<bibRefCitation id="EFEE4B2FFFA01827FCC3F750E996F96F" author="Simon, E." bookContentInfo="Paris" box="[768,948,1660,1683]" pageId="5" pageNumber="6" pagination="256 - 488" part="1" refId="ref1987" refString="1893. Histoire naturelle des araignees 1 (2): 256 - 488. Paris." title="Histoire naturelle des araignees" type="book chapter" year="1893">
Simons (
<year id="FFD7778DFFA01827FCACF750E989F96F" box="[879,939,1660,1683]" pageId="5" pageNumber="6">1893</year>
)
</bibRefCitation>
figures of the males and from
<bibRefCitation id="EFEE4B2FFFA01827FCC6F7B5E999F94C" author="Huber, B. A." bookContentInfo="348 pp." box="[773,955,1689,1712]" journalOrPublisher="Bull. Am. Mus. Nat. Hist." pageId="5" pageNumber="6" part="254" refId="ref1919" refString="Huber, B. A. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bull. Am. Mus. Nat. Hist. 254: 348 pp." title="New World pholcid spiders (Araneae: Pholcidae): a revision at generic level" type="book" year="2000">Hubers (2000)</bibRefCitation>
figures of the epigynum. Most notably, the long bulbal apophysis is strongly curved in Simons fig. 488, while it is almost straight in the new material. Also, the male cheliceral apophyses seem more slender in Simons fig. 489. In general, however, Simons illustrations are not reliable tools for species identification. The differences between the epigyna of the females from Jaar and the female syntypes are minimal and within the usual range of intraspecific variation. Therefore, we provisionally interpret the differences as resulting from either misdrawing or intraspecific variation, or from both.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D794A1DB4CB590539E30A9A1BC58F194">A new species of the genus Orsodacne Latreille, 1802 (Coleoptera: Orsodacnidae) from Iraqi Kurdistan</mods:title>
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<mods:namePart id="BDE33C9FA95AA8718DB1FAF863FA41BA">Legalov, Andrei A.</mods:namePart>
<mods:affiliation id="9ED4B8F6A6F9A4CC68679F1FD5A26C1A">Institute of Systematics and Ecology of Animals, SB RAS, Frunze street 11, Novosibirsk, 630091, Russia &amp; Altai State University, Lenina 61, Barnaul, 656049, Russia &amp; Tomsk State University, Lenina Prospekt 36, 634050 Tomsk, Russia * Corresponding author. E-mail: fossilweevils @ gmail. com</mods:affiliation>
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<mods:title id="C8B2E42CE6C06A5DC3967D234EDABB20">Ecologica Montenegrina</mods:title>
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<mods:date id="90DC7A74D71756EF5067F188E3E75A57">2022</mods:date>
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<mods:number id="C239B08416E823160706D080C4C7CE55">2022-08-09</mods:number>
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Species:
<taxonomicName id="1BFC4D5EFFCAFFDBFDFBFEBAEED4FE4E" authority="Legalov" authorityName="Legalov" authorityYear="2022" box="[551,981,364,391]" class="Insecta" family="Orsodacnidae" genus="Orsodacne" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="species" species="mesopotamicus" status="sp. nov.">
<emphasis id="EE88EACFFFCAFFDBFDFBFEBAEE6AFE4F" bold="true" box="[551,875,364,390]" italics="true" pageId="1" pageNumber="82">Orsodacne mesopotamicus</emphasis>
Legalov
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,
<taxonomicNameLabel id="F5BB57B4FFCAFFDBFC3DFEBBE93CFE4E" box="[993,1085,365,391]" pageId="1" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="44C72A58FFCAFFDBFB91FEBAE99FFE4E" box="[1101,1182,364,391]" captionStart="Figure 1" captionStartId="2.[151,226,1982,2006]" captionTargetBox="[233,1309,216,1936]" captionTargetId="figure-17@2.[225,1363,189,1976]" captionTargetPageId="2" captionText="Figure 1. Orsodacne mesopotamicus sp. nov., holotype, dorsal view: a body, b abdomen, c tergite 8, d tegmen, ventral view. Scale bar for a-b = 1.0 mm, c-d = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13240134" httpUri="https://zenodo.org/record/13240134/files/figure.png" pageId="1" pageNumber="82">Figs. 1</figureCitation>
,
<figureCitation id="44C72A58FFCAFFDBFB71FEBAE9CDFE4E" box="[1197,1228,364,391]" captionStart="Figure 2" captionStartId="3.[151,226,1035,1059]" captionTargetBox="[281,1296,200,1006]" captionTargetId="figure-8@3.[253,1335,189,1029]" captionTargetPageId="3" captionText="Figure 2. Orsodacne spp., aedeagus, dorsal view: a O. cerasi, b O. mesopotamicus sp. nov., holotype, c O. humeralis. Scale bar for a-b = 0.5 mm, c = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13240138" httpUri="https://zenodo.org/record/13240138/files/figure.png" pageId="1" pageNumber="82">2b</figureCitation>
)
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<subSubSection id="94E66556FFCAFFDBFF4BFE19E9A8FDC2" pageId="1" pageNumber="82" type="materials_examined">
<paragraph id="DC4336DDFFCAFFDBFF4BFE19E9A8FDC2" blockId="1.[151,1436,462,523]" pageId="1" pageNumber="82">
<emphasis id="EE88EACFFFCAFFDBFF4BFE19EC48FE20" bold="true" box="[151,329,463,489]" pageId="1" pageNumber="82">Type material</emphasis>
:
<materialsCitation id="6C943C80FFCAFFDBFE86FE18E9A8FDC2" collectingDate="2017-04-17" collectingDateMax="2017-04-25" collectingDateMin="2017-04-17" collectionCode="NHML" collectorName="Reuter" country="Iraq" elevation="1400" latitude="36.5" location="Soran District" longLatPrecision="1187" longitude="44.6" municipality="Erbil" pageId="1" pageNumber="82" specimenCount="1" stateProvince="Kurdistan Region" typeStatus="holotype">
<typeStatus id="0347887FFFCAFFDBFE86FE18ECCBFE20" box="[346,458,462,489]" pageId="1" pageNumber="82" type="holotype">Holotype</typeStatus>
,
<specimenCount id="CAFAFD54FFCAFFDBFE0AFE18EF0EFE20" box="[470,527,462,489]" pageId="1" pageNumber="82" type="male">male</specimenCount>
(
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),
<collectingCountry id="A4EB764DFFCAFFDBFD4DFE18EFDAFE20" box="[657,731,462,489]" name="Iraq" pageId="1" pageNumber="82">IRAQ</collectingCountry>
,
<collectingRegion id="1E38F83FFFCAFFDBFD34FE18EEBCFE20" box="[744,957,462,489]" pageId="1" pageNumber="82">Kurdistan Region</collectingRegion>
,
<collectingMunicipality id="3C27ACA7FFCAFFDBFC15FE18E905FE20" box="[969,1028,462,489]" pageId="1" pageNumber="82">Erbil</collectingMunicipality>
Province,
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, “N- Iraq, S Rawandoz, Akoian valley,
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,
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, ~
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, leg.
<collectorName id="7109530BFFCAFFDBFB9AFE26E99AFDC2" box="[1094,1179,496,523]" pageId="1" pageNumber="82">Reuter</collectorName>
”.
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<paragraph id="DC4336DDFFCAFFDBFF4BFDE2E9A6F8C4" blockId="1.[151,1437,563,1805]" pageId="1" pageNumber="82">
<emphasis id="EE88EACFFFCAFFDBFF4BFDE2EC29FD87" bold="true" box="[151,296,564,590]" pageId="1" pageNumber="82">Description</emphasis>
. Male: Body black, covered with very sparse semierect setae. Mandibles, labrum, clypeus, pronotum, basal half of elytra, antennomeres 1-4, maxillary and labial palpi, trochanters, basal half of femora, tibiae and tarsi brown-yellow. Antennomeres 5-11, coxae, apical part of femora, apex of tibiae brownish. Head capsule hypognathous, short, weak constricted behind eyes. Labrum free, 2.0 times wider than long, finely and sparsely punctate. Mandibles medium, curved. Eyes rounded, convex, finely faceted. Forehead 1.3 times as wide as width of head at apex, weakly convex, densely punctuate, without grooves. Temples about 0.4 times as short as eye length. Gular suture double. Maxillary palpi 4-segmented. Mentum transverse. Labial palpi 3-segmented. Antennae inserted before eyes, filiform and long, reaching humeri. Antennomere 1 suboval, 1.7 times as long as wide. Antennomeres 2-10 long-conical. Antennomeres 2-4 equal in width. Antennomere 2 about 1.4 times as long as wide, about 0.6 times as long as and 0.7 times as narrow as antennomere 1. Antennomere 3 1.6 times as long as wide, 1.2 times as long as antennomere 2. Antennomere 4 about 1.8 times as long as wide, 1.1 times as long as antennomere 3. Antennomeres 5-8 equal in length. Antennomere 5 1.8 times as long as wide, 1.1 times as long as and about 1.1 times as wide as antennomere 4. Antennomere 6 1.9 times as long as wide, about 1.1 times as long as antennomere 5. Antennomere 7 2.0 times as long as wide, about 1.1 times as long as antennomere 6. Antennomere 8 equal to antennomere 7. Antennomere 9 2.0 times as long as wide, 1.1 times as long as and 1.1 times as wide as antennomere 8. Antennomere 10 2.0 times as long as wide, 1.1 times as wide as and 1.1 times as wide as antennomere 9. Antennomere 11 2.6 times as long as wide, 1.2 times as long as and 0.9 times as narrow as antennomere 10, weakly pointed at apex. Pronotum with weakly arcuate sides in apical and middle thirds, concave before middle, 1.3 times as long as wide at apex, 0.9 times as long as wide in middle, 1.3 times as long as wide at base, sparsely punctate. Distances between punctures larger than their diameters. Disc of pronotum moderately convex. Lateral margin of pronotum not carinate. Base 0.6 times as narrow as elytral base. Scutellum distinct, semi-oval. Elytra subparallel-sided, with weak humeri and distinct epipleuron, without striate, 2.1 times as long as wide at base, 1.9 times as long as wide in middle, 2.2 times as long as wide at apical fourth, 3.0 times as long as pronotum. Distances between punctures larger than their diameters. Scutellar striole present. Procoxal cavities narrowly separated. Precoxal portion of prosternum 1.2 times as long as length of procoxal cavity. Postcoxal portion about 0.2 times as long as length of procoxal cavity. Metaventrite 2.7 times as long as metacoxal length, flattened, punctate. Metanepisterna narrow, about 6.3 times as long as wide in middle. Wings developed. Abdomen with free ventrites. Ventrite 1 2.0 times as long as metacoxal cavity length. Ventrite 2 about 0.8 times as long as ventrite 1. Ventrite 3 0.8 times as long as ventrite 2. Ventrite 4 0.8 times as long as ventrite 3. Ventrite 5 0.9 times as long as ventrite 4. Legs long. Femora thickened. Tibiae quite almost straight, with two spurs, without mucros. Tarsi pseudoquadrisegmented. Tarsomere 1 wide-conical. Tarsomere 2 conical. Tarsomere 3 bilobed. Tarsomeres 1-3 with weak pulvilli on lower surface. Tarsomere 5 long. Trasal claws free and bifid. Protarsi: tarsomere 1 1.4 times as long as wide at apex; tarsomere 2 1.2 times as long as wide at apex, 0.6 times as long as tarsomere 1; tarsomere 3 0.9 times as long as wide at apex, 0.8 times as long as tarsomere 2; tarsomere 5 3.7 times as long as wide at apex, 0.5 times as long as tarsomere 3. Length of body:
<quantity id="1B049B38FFCAFFDBFB9AF924E9A2F8C4" box="[1094,1187,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="82" unit="mm" value="4.4">4.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="94E66556FFCAFFDBFF4BF8E0EF15F87C" pageId="1" pageNumber="82" type="diagnosis">
<paragraph id="DC4336DDFFCAFFDBFF4BF8E0EF15F87C" blockId="1.[151,1437,1845,1973]" pageId="1" pageNumber="82">
<emphasis id="EE88EACFFFCAFFDBFF4BF8E0ECAAF899" bold="true" box="[151,427,1846,1872]" pageId="1" pageNumber="82">Differential diagnosis</emphasis>
. The new species is close to
<taxonomicName id="1BFC4D5EFFCAFFDBFCECF8E0EEA2F886" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[816,931,1846,1872]" class="Insecta" family="Orsodacnidae" genus="Orsodacne" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="species" species="cerasi">
<emphasis id="EE88EACFFFCAFFDBFCECF8E0EE44F899" box="[816,837,1846,1872]" italics="true" pageId="1" pageNumber="82">O</emphasis>
.
<emphasis id="EE88EACFFFCAFFDBFC87F8E0EEA2F886" box="[859,931,1846,1871]" italics="true" pageId="1" pageNumber="82">cerasi</emphasis>
</taxonomicName>
and is distinguished by a more weakly narrowed pronotum before the base (fig. 1a) and the armament of the endophallus (figs. 2a, 2b). It differs from
<taxonomicName id="1BFC4D5EFFCAFFDBFF04F8ACEC73F85D" authorityName="Latreille" authorityYear="1804" box="[216,370,1913,1940]" class="Insecta" family="Orsodacnidae" genus="Orsodacne" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="EE88EACFFFCAFFDBFF04F8ACEC73F85D" box="[216,370,1913,1940]" italics="true" pageId="1" pageNumber="82">O. humeralis</emphasis>
</taxonomicName>
in the sparsely punctate elytra covered with very sparse semierect setae and the armament of the endophallus (figs. 2b, 2c).
</paragraph>
</subSubSection>
<subSubSection id="94E66556FFCAFFDBFF4BF808EF37F830" box="[151,566,2014,2041]" pageId="1" pageNumber="82" type="etymology">
<paragraph id="DC4336DDFFCAFFDBFF4BF808EF37F830" blockId="1.[151,566,2014,2041]" box="[151,566,2014,2041]" pageId="1" pageNumber="82">
<emphasis id="EE88EACFFFCAFFDBFF4BF808EC1EF831" bold="true" box="[151,287,2014,2040]" pageId="1" pageNumber="82">Etymology</emphasis>
. From “
<collectingCountry id="A4EB764DFFCAFFDBFEA2F808EF26F830" box="[382,551,2014,2041]" name="Iraq" pageId="1" pageNumber="82">Mesopotamia</collectingCountry>
”.
</paragraph>
</subSubSection>
<caption id="88836655FFC9FFD8FF4BF868EFEDF83D" ID-DOI="http://doi.org/10.5281/zenodo.13240134" ID-Zenodo-Dep="13240134" httpUri="https://zenodo.org/record/13240134/files/figure.png" pageId="2" pageNumber="83" startId="2.[151,226,1982,2006]" targetBox="[233,1309,216,1936]" targetPageId="2" targetType="figure">
<paragraph id="DC4336DDFFC9FFD8FF4BF868EFEDF83D" blockId="2.[151,1437,1982,2036]" pageId="2" pageNumber="83">
<emphasis id="EE88EACFFFC9FFD8FF4BF868EDF6F81F" bold="true" box="[151,247,1982,2006]" pageId="2" pageNumber="83">Figure 1</emphasis>
.
<taxonomicName id="1BFC4D5EFFC9FFD8FEDAF868EF26F81C" authorityName="Legalov" authorityYear="2022" box="[262,551,1982,2006]" class="Insecta" family="Orsodacnidae" genus="Orsodacne" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="mesopotamicus" status="sp. nov.">
<emphasis id="EE88EACFFFC9FFD8FEDAF868EF26F81C" box="[262,551,1982,2006]" italics="true" pageId="2" pageNumber="83">Orsodacne mesopotamicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F5BB57B4FFC9FFD8FDF3F868EF87F81C" box="[559,646,1982,2005]" pageId="2" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, dorsal view: a body, b abdomen, c tergite 8, d tegmen, ventral view. Scale bar for a-b = 1.0 mm, c-d = 0.5 mm.
</paragraph>
</caption>
<caption id="88836655FFC8FFD9FF4BFBDDEFBDFB88" ID-DOI="http://doi.org/10.5281/zenodo.13240138" ID-Zenodo-Dep="13240138" httpUri="https://zenodo.org/record/13240138/files/figure.png" pageId="3" pageNumber="84" startId="3.[151,226,1035,1059]" targetBox="[281,1296,200,1006]" targetPageId="3" targetType="figure">
<paragraph id="DC4336DDFFC8FFD9FF4BFBDDEFBDFB88" blockId="3.[151,1437,1035,1090]" pageId="3" pageNumber="84">
<emphasis id="EE88EACFFFC8FFD9FF4BFBDDEDFBFBEA" bold="true" box="[151,250,1035,1059]" pageId="3" pageNumber="84">Figure 2</emphasis>
.
<taxonomicName id="1BFC4D5EFFC8FFD9FED1FBDDECBAFBEA" box="[269,443,1035,1059]" class="Insecta" family="Orsodacnidae" genus="Orsodacne" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EE88EACFFFC8FFD9FED1FBDDEC83FBEA" box="[269,386,1035,1059]" italics="true" pageId="3" pageNumber="84">Orsodacne</emphasis>
spp.
</taxonomicName>
, aedeagus, dorsal view: a
<taxonomicName id="1BFC4D5EFFC8FFD9FD22FBDAEE65FBEA" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[766,868,1036,1059]" class="Insecta" family="Orsodacnidae" genus="Orsodacne" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="cerasi">
<emphasis id="EE88EACFFFC8FFD9FD22FBDAEE65FBEA" box="[766,868,1036,1059]" italics="true" pageId="3" pageNumber="84">O. cerasi</emphasis>
</taxonomicName>
, b
<taxonomicName id="1BFC4D5EFFC8FFD9FC7BFBDAE96EFBEA" authorityName="Legalov" authorityYear="2022" box="[935,1135,1036,1059]" class="Insecta" family="Orsodacnidae" genus="Orsodacne" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="mesopotamicus" status="sp. nov.">
<emphasis id="EE88EACFFFC8FFD9FC7BFBDAE96EFBEA" box="[935,1135,1036,1059]" italics="true" pageId="3" pageNumber="84">O. mesopotamicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F5BB57B4FFC8FFD9FBA7FBDAE9D4FBEA" box="[1147,1237,1036,1059]" pageId="3" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, c
<taxonomicName id="1BFC4D5EFFC8FFD9FA5EFBDAEC03FB8B" authorityName="Latreille" authorityYear="1804" class="Insecta" family="Orsodacnidae" genus="Orsodacne" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="EE88EACFFFC8FFD9FA5EFBDAEC03FB8B" italics="true" pageId="3" pageNumber="84">O. humeralis</emphasis>
</taxonomicName>
. Scale bar for a-b = 0.5 mm, c = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="94E66556FFC8FFD9FF4BFB5BEF79FB6E" box="[151,632,1164,1191]" pageId="3" pageNumber="84" type="distribution">
<paragraph id="DC4336DDFFC8FFD9FF4BFB5BEF79FB6E" blockId="3.[151,632,1164,1191]" box="[151,632,1164,1191]" pageId="3" pageNumber="84">
<emphasis id="EE88EACFFFC8FFD9FF4BFB5BEC30FB6E" bold="true" box="[151,305,1165,1191]" pageId="3" pageNumber="84">Distribution</emphasis>
. North-eastern
<collectingCountry id="A4EB764DFFC8FFD9FE34FB5BEF16FB6E" box="[488,535,1165,1191]" name="Iraq" pageId="3" pageNumber="84">Iraq</collectingCountry>
(fig. 3).
</paragraph>
</subSubSection>
</treatment>
</document>