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<mods:titleInfo id="A04260BDFA9E3E686333528ADAAB9B86">
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<mods:title id="DD69011CE44640CF54CCFB36534EB3F4">Two new species of fanged frog from Southeastern Borneo, Indonesia (Amphibia: Anura: Dicroglossidae)</mods:title>
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<mods:namePart id="67C05D20A53F737F62250FC1217CA35D">Gonggoli, Ade Damara</mods:namePart>
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<mods:namePart id="8C44235A0D792BA7640CE291DE9BE459">Shimada, Tomohiko</mods:namePart>
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<mods:affiliation id="06212D14BB6871BACB365CA451DE1388">Faculty of Education, Aichi University of Education, Kariya, Aichi 448 - 8542, Japan.</mods:affiliation>
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<mods:namePart id="D315F86BE38F7683755D6B621F1A5253">Matsui, Masafumi</mods:namePart>
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<mods:affiliation id="9323A11E8C2242DAAF8B992FF664E164">Fukakusa Bo-cho 12 - 13, Fushimi, Kyoto 612 - 0871, Japan.</mods:affiliation>
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<mods:namePart id="147CC1AF7E42CA39B675B2F1D086F6F6">Nishikawa, Kanto</mods:namePart>
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<mods:affiliation id="AF918300D4E0C94032D4B7B15228E860">Graduate School of Human and Environmental Studies, Kyoto University, Yoshida-Nihonmatsu-cho, Sakyo-ku, Kyoto 606 - 8501, Japan. & Graduate School of Global Environmental Studies, Kyoto University, Yoshida-Hon-machi, Sakyo-ku, Kyoto 606 - 8501, Japan.</mods:affiliation>
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<mods:namePart id="1EBACD586617BA308C7F01C7B29FCF67">Sidik, Irvan</mods:namePart>
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<mods:affiliation id="759D6CD705E636FF8FBA7EA3F52D7B5B">Laboratory of Herpetology, Museum Zoologicum Bogoriense, Research Centre for Biosystematics and Evolution, National Research and Innovation Agency (BRIN), Gd. Widyasatwaloka, KST Soekarno, Jl. Raya Jakarta Bogor km 46, Cibinong, Bogor, West Java 16912, Indonesia.</mods:affiliation>
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<mods:namePart id="FDE5E8060AF6F703DDF0F7AFAC48079B">Kadafi, Ahmad Muammar</mods:namePart>
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<mods:affiliation id="9E8D56401D46C91E99624D4DE382DE7B">Department of Biology, Faculty of Mathematics and Natural Sciences, University of Palangka Raya, Jl. Yos Sudarso, Palangka Raya 74874, Central Kalimantan, Indonesia.</mods:affiliation>
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<mods:namePart id="749C785D42AC6DB50123552D2CA4A837">Farajallah, Achmad</mods:namePart>
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<mods:affiliation id="4DFD16B3B4EEF5ECF8493F2249DB7414">Graduate Program of Animal Bioscience, Department of Biology, Faculty of Mathematics and Natural Sciences, Bogor Agricultural University, Bogor, West Java 16680, Indonesia.</mods:affiliation>
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<mods:affiliation id="43B523DF35B5D004D3692E9B90960127">Laboratory of Herpetology, Museum Zoologicum Bogoriense, Research Centre for Biosystematics and Evolution, National Research and Innovation Agency (BRIN), Gd. Widyasatwaloka, KST Soekarno, Jl. Raya Jakarta Bogor km 46, Cibinong, Bogor, West Java 16912, Indonesia.</mods:affiliation>
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<mods:date id="E2FBAAB0726CD4D026B19B4F5C39C08B">2025</mods:date>
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<taxonomicName id="4C7F66C5FF8E8F660AF79D64CBD9F8B0" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[151,491,1839,1865]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="394" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
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<emphasis id="B90BC154FF8E8F660AF79D64CBD9F8B0" bold="true" box="[151,491,1839,1865]" italics="true" pageId="7" pageNumber="394">Limnonectes maanyanorum</emphasis>
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(
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<figureCitation id="134401C3FF8E8F660AFF9D18CB38F897" box="[159,266,1875,1902]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="8.[151,250,1598,1622]" captionStartId-1="9.[151,250,1556,1580]" captionTargetBox-0="[340,1247,453,1574]" captionTargetBox-1="[340,1247,776,1532]" captionTargetId-0="figure-137@8.[340,1247,453,1574]" captionTargetId-1="figure-288@9.[340,1247,776,1532]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 5. Dorsolateral view, dorsal view, and ventral view, respectively, of adult female holotype, MZB Amph 33530 (A–C), and unvouchered adult male (D–F) of Limnonectes maanyanorum sp. nov. Photographs by Ade Damara Gonggoli." captionText-1="FIGURE 6. Dorsal view (A), ventral view (B), ventral view of right foot (C), ventral view of right hand (D), frontal view of head with opened mouth, showing the odontoids processes, not to scale (E) of Limnonectes maanyanorum sp. nov., adult female holotype (MZB Amph 33530). Scale = 1 mm. Photographs by Ade Damara Gonggoli." figureDoi-0="http://doi.org/10.5281/zenodo.14747088" figureDoi-1="http://doi.org/10.5281/zenodo.14747090" httpUri-0="https://zenodo.org/record/14747088/files/figure.png" httpUri-1="https://zenodo.org/record/14747090/files/figure.png" pageId="7" pageNumber="394">Figs. 5–6</figureCitation>
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)
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<emphasis id="B90BC154FF8E8F660AF79DD3CB6AF856" box="[151,344,1943,1967]" italics="true" pageId="7" pageNumber="394">Limnonectes kuhlii</emphasis>
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“Lineage
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3”:
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<bibRefCitation id="EFEE60B7FF8E8F660B869DDCC89DF856" author="Matsui, M. & Kuraishi, N. & Eto, K. & Hamidy, A. & Nishikawa, K. & Shimada, T. & Yambun, P. & Vairappan, C. S. & Hossman, M. Y. B." box="[486,687,1943,1967]" pageId="7" pageNumber="394" pagination="305 - 319" refId="ref14683" refString="Matsui, M., Kuraishi, N., Eto, K., Hamidy, A., Nishikawa, K., Shimada, T., Yambun, P., Vairappan, C. S. & Hossman, M. Y. B. (2016) Unusually high genetic diversity in the Bornean Limnonectes kuhlii - like fanged frogs (Anura: Dicroglossidae). Molecular Phylogenetics and Evolution, 102, 305 - 319. https: // doi. org / 10.1016 / j. ympev. 2016.06.009" type="journal article" year="2016">
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Matsui
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<emphasis id="B90BC154FF8E8F6608539DD3C854F856" box="[563,614,1943,1967]" italics="true" pageId="7" pageNumber="394">et al.</emphasis>
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(2016)
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.
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female (MZB Amph 33530) from
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,
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,
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at -
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||||
,
|
||||
<geoCoordinate id="EE4B7B81FF818F690EE79AF1CF18FF2C" box="[1159,1322,186,213]" degrees="115.303402" direction="east" orientation="longitude" pageId="8" pageNumber="395" precision="1" value="115.303406">115.303402°E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0052FA75FF818F690F569AF0CB3FFF00" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="8" pageNumber="395" unit="m" value="91.0">
|
||||
elevation
|
||||
<quantity id="4C87B0A3FF818F690AF79A94CAE2FF00" box="[151,208,223,249]" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="8" pageNumber="395" unit="m" value="91.0">91 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
, collected by
|
||||
<collectorName id="268A7890FF818F690BC79A94C894FF00" box="[423,678,223,249]" pageId="8" pageNumber="395">Ade Damara Gonggoli</collectorName>
|
||||
and
|
||||
<collectorName id="268A7890FF818F6908BA9A94CE27FF00" box="[730,1045,223,249]" pageId="8" pageNumber="395">Adventus Robertino Rangin</collectorName>
|
||||
on
|
||||
<date id="FFC13B86FF818F690E5F9A94CECEFF00" box="[1087,1276,223,249]" pageId="8" pageNumber="395" value="2023-01-12">
|
||||
<collectingDate id="EF85C26EFF818F690E5F9A94CECEFF00" box="[1087,1276,223,249]" pageId="8" pageNumber="395" value="2023-01-12">12 January 2023</collectingDate>
|
||||
</date>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF818F690AA79B48CBF4FE54" blockId="8.[151,1437,150,429]" pageId="8" pageNumber="395">
|
||||
<materialsCitation id="3B17171BFF818F690AA79B48CBF1FE54" collectingDate="2022-02-22" collectorName="Ade Damara Gonggoli & Adventus Robertino Rangin & Tendang & Pa Nika" country="Indonesia" county="Barito Timur Regency" elevation="164" latitude="-1.918157" location="Total" longLatPrecision="1" longitude="115.308815" municipality="Awang District" pageId="8" pageNumber="395" specimenCount="16" specimenCount-adult="8" stateProvince="Kalimantan Tengah" typeStatus="paratype">
|
||||
<emphasis id="B90BC154FF818F690AA79B48CB80FEE5" bold="true" box="[199,434,258,285]" pageId="8" pageNumber="395">
|
||||
<typeStatus id="54C4A3E4FF818F690AA79B48CB0DFEE4" box="[199,319,259,285]" pageId="8" pageNumber="395" type="paratype">Paratypes</typeStatus>
|
||||
(n = 10).
|
||||
</emphasis>
|
||||
<location id="8EA04B9DFF818F690BDC9B48CBC7FEE4" LSID="urn:lsid:plazi:treatment:03D6AC50FF8E8F6A0AF79D64CE53FD7C:8EA04B9DFF818F690BDC9B48CBC7FEE4" box="[444,501,259,285]" country="Indonesia" county="Barito Timur Regency" latitude="-1.918157" longLatPrecision="1" longitude="115.308815" municipality="Awang District" name="Total" pageId="8" pageNumber="395" stateProvince="Kalimantan Tengah">Total</location>
|
||||
of
|
||||
<specimenCount id="9D79D6CFFF818F6908409B48C8E5FEE4" box="[544,727,259,285]" count="8" pageId="8" pageNumber="395" type="generic">eight specimens</specimenCount>
|
||||
from
|
||||
<location id="8EA04B9DFF818F6909419B48C9F5FEE4" LSID="urn:lsid:plazi:treatment:03D6AC50FF8E8F6A0AF79D64CE53FD7C:8EA04B9DFF818F6909419B48C9F5FEE4" box="[801,967,259,285]" country="Indonesia" county="Barito Timur Regency" latitude="-1.918157" longLatPrecision="1" longitude="115.308815" municipality="Awang District" name="Mekang River" pageId="8" pageNumber="395" stateProvince="Kalimantan Tengah">Mekang River</location>
|
||||
,
|
||||
<location id="8EA04B9DFF818F6909B59B48CE53FEE4" LSID="urn:lsid:plazi:treatment:03D6AC50FF8E8F6A0AF79D64CE53FD7C:8EA04B9DFF818F6909B59B48CE53FEE4" box="[981,1121,259,285]" country="Indonesia" county="Barito Timur Regency" latitude="-1.918157" longLatPrecision="1" longitude="115.308815" municipality="Awang District" name="Mt. Karasik" pageId="8" pageNumber="395" stateProvince="Kalimantan Tengah">Mt. Karasik</location>
|
||||
,
|
||||
<location id="8EA04B9DFF818F690E0D9B48CF70FEE4" LSID="urn:lsid:plazi:treatment:03D6AC50FF8E8F6A0AF79D64CE53FD7C:8EA04B9DFF818F690E0D9B48CF70FEE4" box="[1133,1346,259,285]" country="Indonesia" county="Barito Timur Regency" latitude="-1.918157" longLatPrecision="1" longitude="115.308815" municipality="Awang District" name="Apar Batu Village" pageId="8" pageNumber="395" stateProvince="Kalimantan Tengah">Apar Batu Village</location>
|
||||
,
|
||||
<collectingMunicipality id="6BA4873CFF818F690F2E9B48CADCFEB8" pageId="8" pageNumber="395">Awang District</collectingMunicipality>
|
||||
,
|
||||
<collectingCounty id="62A165CAFF818F690A9A9B6CCBC5FEB8" box="[250,503,295,321]" pageId="8" pageNumber="395">Barito Timur Regency</collectingCounty>
|
||||
,
|
||||
<collectingRegion id="49BBD3A4FF818F6908649B6CC97FFEB8" box="[516,845,295,321]" country="Indonesia" name="Kalimantan Tengah" pageId="8" pageNumber="395">Central Kalimantan Province</collectingRegion>
|
||||
(Borneo),
|
||||
<collectingCountry id="F3685DD6FF818F6909A89B6CCE07FEB8" box="[968,1077,295,321]" name="Indonesia" pageId="8" pageNumber="395">Indonesia</collectingCountry>
|
||||
at -
|
||||
<geoCoordinate id="EE4B7B81FF818F690E019B6DCEDBFEB8" box="[1121,1257,294,321]" degrees="1.918157" direction="south" orientation="latitude" pageId="8" pageNumber="395" precision="1" value="-1.918157">1.918157°S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4B7B81FF818F690E959B6DCFABFEB8" box="[1269,1433,294,321]" degrees="115.308813" direction="east" orientation="longitude" pageId="8" pageNumber="395" precision="1" value="115.308815">115.308813°E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0052FA75FF818F690AF79B00CBA0FE9C" box="[151,402,331,357]" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="8" pageNumber="395" unit="m" value="164.0">
|
||||
elevation
|
||||
<quantity id="4C87B0A3FF818F690B689B00CB63FE9C" box="[264,337,331,357]" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="8" pageNumber="395" unit="m" value="164.0">164 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
:
|
||||
<specimenCount id="9D79D6CFFF818F690BC19B00C839FE9C" box="[417,523,331,357]" count="1" pageId="8" pageNumber="395" type="adult">one adult</specimenCount>
|
||||
male (MZB Amph 33514) collected with holotype;
|
||||
<specimenCount id="9D79D6CFFF818F690E0A9B00CEDEFE9C" box="[1130,1260,331,357]" count="7" pageId="8" pageNumber="395" type="adult">seven adult</specimenCount>
|
||||
females (MZB Amph 33513, 33515, 33531–35) collected by
|
||||
<collectorName id="268A7890FF818F6908F59B24C9A4FE70" box="[661,918,367,393]" pageId="8" pageNumber="395">Ade Damara Gonggoli</collectorName>
|
||||
,
|
||||
<collectorName id="268A7890FF818F6909FF9B24CEE9FE70" box="[927,1243,367,393]" pageId="8" pageNumber="395">Adventus Robertino Rangin</collectorName>
|
||||
,
|
||||
<collectorName id="268A7890FF818F690E849B24CF75FE70" box="[1252,1351,367,393]" pageId="8" pageNumber="395">Tendang</collectorName>
|
||||
, and
|
||||
<collectorName id="268A7890FF818F690FE09B24CAFFFE54" pageId="8" pageNumber="395">Pa Nika</collectorName>
|
||||
on
|
||||
<date id="FFC13B86FF818F690A989BD8CBF1FE54" box="[248,451,403,429]" pageId="8" pageNumber="395" value="2022-02-22">
|
||||
<collectingDate id="EF85C26EFF818F690A989BD8CBF1FE54" box="[248,451,403,429]" pageId="8" pageNumber="395" value="2022-02-22">22 February 2022</collectingDate>
|
||||
</date>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF004DCEFF818F690AF79C75CF17F983" ID-DOI="http://doi.org/10.5281/zenodo.14747088" ID-Zenodo-Dep="14747088" httpUri="https://zenodo.org/record/14747088/files/figure.png" pageId="8" pageNumber="395" startId="8.[151,250,1598,1622]" targetBox="[340,1247,453,1574]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8BC01D46FF818F690AF79C75CF17F983" blockId="8.[151,1436,1598,1658]" pageId="8" pageNumber="395">
|
||||
<emphasis id="B90BC154FF818F690AF79C75CB20F9AF" bold="true" box="[151,274,1598,1622]" pageId="8" pageNumber="395">FIGURE 5.</emphasis>
|
||||
Dorsolateral view, dorsal view, and ventral view, respectively, of adult female holotype, MZB Amph 33530 (A–C), and unvouchered adult male (D–F) of
|
||||
<taxonomicName id="4C7F66C5FF818F69087C9C28C906F983" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[540,820,1635,1658]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="395" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF818F69087C9C28C906F983" box="[540,820,1635,1658]" italics="true" pageId="8" pageNumber="395">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A2387C2FFF818F69095B9C29C9BFF983" box="[827,909,1634,1658]" pageId="8" pageNumber="395" rank="species">
|
||||
<emphasis id="B90BC154FF818F69095B9C29C9B4F983" bold="true" box="[827,902,1634,1658]" pageId="8" pageNumber="395">sp. nov</emphasis>
|
||||
.
|
||||
</taxonomicNameLabel>
|
||||
Photographs by Ade Damara Gonggoli.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BC01D46FF818F690AA79C86CE38F8D6" blockId="8.[151,1437,1740,2019]" pageId="8" pageNumber="395">
|
||||
<materialsCitation id="3B17171BFF818F690AA79C86CE68F8F3" country="Indonesia" county="Kutai Kertanegara Regency" elevation="68" latitude="-1.05225" location="Margomulyo Village" longLatPrecision="1" longitude="116.9928" municipality="Samboja District" pageId="8" pageNumber="395" specimenCount="2" stateProvince="Kalimantan Timur">
|
||||
<specimenCount id="9D79D6CFFF818F690AA79C86CB4AF91F" box="[199,376,1740,1767]" count="2" pageId="8" pageNumber="395" type="generic">Two specimens</specimenCount>
|
||||
from
|
||||
<location id="8EA04B9DFF818F690BA39C87C880F91F" LSID="urn:lsid:plazi:treatment:03D6AC50FF8E8F6A0AF79D64CE53FD7C:8EA04B9DFF818F690BA39C87C880F91F" box="[451,690,1740,1766]" country="Indonesia" county="Kutai Kertanegara Regency" latitude="-1.05225" longLatPrecision="1" longitude="116.9928" municipality="Samboja District" name="Margomulyo Village" pageId="8" pageNumber="395" stateProvince="Kalimantan Timur">Margomulyo Village</location>
|
||||
,
|
||||
<collectingMunicipality id="6BA4873CFF818F6908A09C87C9B6F91F" box="[704,900,1740,1766]" pageId="8" pageNumber="395">Samboja District</collectingMunicipality>
|
||||
,
|
||||
<collectingCounty id="62A165CAFF818F6909F29C87CEE3F91E" box="[914,1233,1740,1767]" pageId="8" pageNumber="395">Kutai Kertanegara Regency</collectingCounty>
|
||||
,
|
||||
<collectingRegion id="49BBD3A4FF818F690EBF9C86CAC9F8F3" country="Indonesia" name="Kalimantan Timur" pageId="8" pageNumber="395">East Kalimantan Province</collectingRegion>
|
||||
(Borneo),
|
||||
<collectingCountry id="F3685DD6FF818F690B199CBBCBD4F8F3" box="[377,486,1776,1802]" name="Indonesia" pageId="8" pageNumber="395">Indonesia</collectingCountry>
|
||||
at -
|
||||
<geoCoordinate id="EE4B7B81FF818F6908749CBBC8AEF8F2" box="[532,668,1776,1803]" degrees="1.052250" direction="south" orientation="latitude" pageId="8" pageNumber="395" precision="1" value="-1.05225">1.052250°S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="EE4B7B81FF818F6908C99CBBC97EF8F3" box="[681,844,1776,1802]" degrees="116.992798" direction="east" orientation="longitude" pageId="8" pageNumber="395" precision="1" value="116.9928">116.992798°E</geoCoordinate>
|
||||
,
|
||||
<elevation id="0052FA75FF818F69093A9CBBCE68F8F3" box="[858,1114,1776,1803]" metricMagnitude="1" metricUnit="m" metricValue="6.83" pageId="8" pageNumber="395" unit="m" value="68.3">
|
||||
elevation
|
||||
<quantity id="4C87B0A3FF818F6909A99CBBCE28F8F2" box="[969,1050,1776,1803]" metricMagnitude="1" metricUnit="m" metricValue="6.83" pageId="8" pageNumber="395" unit="m" value="68.3">68.3 m</quantity>
|
||||
a.s.l.
|
||||
</elevation>
|
||||
</materialsCitation>
|
||||
:
|
||||
<materialsCitation id="3B17171BFF818F690E079CBACE38F8D6" pageId="8" pageNumber="395">
|
||||
<specimenCount id="9D79D6CFFF818F690E079CBACECDF8F3" box="[1127,1279,1776,1803]" count="2" pageId="8" pageNumber="395" type="juvenile">two juveniles</specimenCount>
|
||||
(MZB Amph 15264–65) collected by Amir Hamidy and Abran Slamet on
|
||||
<date id="FFC13B86FF818F69095B9D5ECE34F8D6" box="[827,1030,1812,1839]" pageId="8" pageNumber="395" value="2009-02-18">18 February 2009</date>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3654ECDFF818F680AA79D72C995FF2C" lastPageId="9" lastPageNumber="396" pageId="8" pageNumber="395" type="etymology">
|
||||
<paragraph id="8BC01D46FF818F690AA79D72CB12F862" blockId="8.[151,1437,1740,2019]" pageId="8" pageNumber="395">
|
||||
<emphasis id="B90BC154FF818F690AA79D72CB7CF8AA" bold="true" box="[199,334,1849,1875]" pageId="8" pageNumber="395">Etymology.</emphasis>
|
||||
The species epithet
|
||||
<taxonomicName id="4C7F66C5FF818F69084F9D72C8E3F8AB" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[559,721,1849,1874]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="8" pageNumber="395" phylum="Chordata" rank="species" species="maanyanorum">
|
||||
<emphasis id="B90BC154FF818F69084F9D72C8E3F8AB" box="[559,721,1849,1874]" italics="true" pageId="8" pageNumber="395">maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
refers to the name of the local ethnic, Dayak Maanyan, who live in
|
||||
<collectingRegion id="49BBD3A4FF818F690AD69D17CBA5F88F" box="[182,407,1884,1910]" country="Indonesia" name="Kalimantan Tengah" pageId="8" pageNumber="395">Central Kalimantan</collectingRegion>
|
||||
, from where the species is found. The species epithet is in genitive plural, meaning of the “Maanyan”.
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF818F690AA79DEEC9C4F81B" blockId="8.[151,1437,1740,2019]" pageId="8" pageNumber="395">
|
||||
<emphasis id="B90BC154FF818F690AA79DEECBF4F846" bold="true" box="[199,454,1957,1983]" pageId="8" pageNumber="395">Local common name.</emphasis>
|
||||
On Mt. Karasik, the local people of the Dayak Maanyan ethnic call this frog “Senteleng Watu”, which means “Katak Batu” in
|
||||
<collectingCountry id="F3685DD6FF818F6908219D83C89CF81B" box="[577,686,1992,2018]" name="Indonesia" pageId="8" pageNumber="395">Indonesia</collectingCountry>
|
||||
and “Rock Frog” in English.
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF808F680AA79ADDC968FF48" blockId="9.[151,1437,150,753]" box="[199,858,150,177]" pageId="9" pageNumber="396">
|
||||
<emphasis id="B90BC154FF808F680AA79ADDC86FFF49" bold="true" box="[199,605,150,176]" pageId="9" pageNumber="396">Suggested English common name.</emphasis>
|
||||
Maanyan Creek Frog.
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF808F680AA79AF1C995FF2C" blockId="9.[151,1437,150,753]" box="[199,935,186,213]" pageId="9" pageNumber="396">
|
||||
<emphasis id="B90BC154FF808F680AA79AF1C8B5FF2D" bold="true" box="[199,647,186,212]" pageId="9" pageNumber="396">Suggested Indonesian common name.</emphasis>
|
||||
Bangkong-tuli Maanyan.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3654ECDFF808F680AA79A95CB5CFE0D" pageId="9" pageNumber="396" type="diagnosis">
|
||||
<paragraph id="8BC01D46FF808F680AA79A95CB5CFE0D" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="396">
|
||||
<emphasis id="B90BC154FF808F680AA79A95CB70FF01" bold="true" box="[199,322,222,248]" pageId="9" pageNumber="396">Diagnosis.</emphasis>
|
||||
The new species clustered within the
|
||||
<taxonomicName id="4C7F66C5FF808F6808BA9A94C907FF01" box="[730,821,222,248]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="396" phylum="Chordata" rank="species" species="kuhlii">
|
||||
<emphasis id="B90BC154FF808F6808BA9A94C907FF01" box="[730,821,222,248]" italics="true" pageId="9" pageNumber="396">L. kuhlii</emphasis>
|
||||
</taxonomicName>
|
||||
complex in molecular analyses and can be distinguished from its congeners by the following combination of morphological characters: medium size (SVL
|
||||
<quantity id="4C87B0A3FF808F680E909B49CF67FEE4" box="[1264,1365,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="4.59" pageId="9" pageNumber="396" unit="mm" value="45.9">45.9 mm</quantity>
|
||||
in the only adult male,
|
||||
<quantity id="4C87B0A3FF808F680B319B6DCBC6FEB8" box="[337,500,294,321]" metricMagnitude="-2" metricUnit="m" metricValue="4.945" metricValueMax="5.85" metricValueMin="4.04" pageId="9" pageNumber="396" unit="mm" value="49.45" valueMax="58.5" valueMin="40.4">40.4–58.5 mm</quantity>
|
||||
in adult females); toes fully webbed; tympanum hidden; dorsal skin of body rugose; dorsal skin of tibia densely covered by small warts and coarsely scattered with well-elevated large tubercles; flap of skin on outer side of third finger well-developed, moveable; inner and outer webbing of fourth toe fairly well excised; disk of third finger not much wider than basal phalanx; disk of fourth toe wider than basal phalanx; male nuptial pad on first finger without minute asperities; head longer than wide; throat covered with confluent irregular dark gray blotches.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3654ECDFF808F6A0AA79BB5CF37FEB8" lastPageId="11" lastPageNumber="398" pageId="9" pageNumber="396" type="description">
|
||||
<paragraph id="8BC01D46FF808F680AA79BB5CE72FD09" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="396">
|
||||
<emphasis id="B90BC154FF808F680AA79BB5C900FDE1" bold="true" box="[199,818,510,536]" pageId="9" pageNumber="396">
|
||||
Description of the
|
||||
<typeStatus id="54C4A3E4FF808F680BC79BB5C83CFDE1" box="[423,526,510,536]" pageId="9" pageNumber="396" type="holotype">holotype</typeStatus>
|
||||
(measurements in mm).
|
||||
</emphasis>
|
||||
Adult female (SVL 57.0), habitus stocky; head longer (HL 24.1, 42.3 % SVL) than wide (HW 23.0, 40.3 % SVL); rostrum longer (RL 9.3, 16.4 % SVL) than eye (ED 5.6, 9.9 % SVL); nostril dorsolateral on canthus, closer to snout tip (SL 4.0, 7.0 % SVL) than to eye (N-EL 5.4, 9.4 % SVL); internarial distance (IND 4.6, 8.0 % SVL) longer than interorbital distance (IOD 4.2, 7.3 % SVL); snout obtusely pointed in lateral view; canthus rounded; lore concave; pineal spot distinct; tympanum indistinct; pair of odontoid processes on lower jaw, obtuse in profile; vomerine teeth in oblique groups, groups separated from each other and from choana by half length of one group; vocal sac and vocal slits absent.
|
||||
</paragraph>
|
||||
<caption id="DF004DCEFF808F680AF79C5FC9CDF98D" ID-DOI="http://doi.org/10.5281/zenodo.14747090" ID-Zenodo-Dep="14747090" httpUri="https://zenodo.org/record/14747090/files/figure.png" pageId="9" pageNumber="396" startId="9.[151,250,1556,1580]" targetBox="[340,1247,776,1532]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="8BC01D46FF808F680AF79C5FC9CDF98D" blockId="9.[151,1436,1556,1652]" pageId="9" pageNumber="396">
|
||||
<emphasis id="B90BC154FF808F680AF79C5FCB26F9D5" bold="true" box="[151,276,1556,1580]" pageId="9" pageNumber="396">FIGURE 6.</emphasis>
|
||||
Dorsal view (A), ventral view (B), ventral view of right foot (C), ventral view of right hand (D), frontal view of head with opened mouth, showing the odontoids processes, not to scale (E) of
|
||||
<taxonomicName id="4C7F66C5FF808F6809C69C72CE8CF9A9" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[934,1214,1593,1616]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="396" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF808F6809C69C72CE8CF9A9" box="[934,1214,1593,1616]" italics="true" pageId="9" pageNumber="396">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF808F680EA49C73CF28F9A9" bold="true" box="[1220,1306,1592,1616]" pageId="9" pageNumber="396">
|
||||
<taxonomicNameLabel id="A2387C2FFF808F680EA49C73CF27F9A9" box="[1220,1301,1592,1616]" pageId="9" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
,
|
||||
</emphasis>
|
||||
adult female holotype (MZB Amph 33530). Scale = 1 mm. Photographs by Ade Damara Gonggoli.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BC01D46FF808F680AA79CF0CE79F854" blockId="9.[151,1437,1723,2038]" pageId="9" pageNumber="396">Forelimb stout, relatively short (FLL 28.0, 49.2 % SVL); fingers moderately slender; finger length formula II <I <IV <III; finger webbing absent; finger tips bluntly rounded, without circummarginal grooves; disk of third finger (3TDW 1.0, 1.7 % SVL) not much wider than basal phalanx; inner palmar tubercle oval and low (IPTL 3.3, 5.7 % SVL); middle palmar tubercle oval and low; outer palmar tubercle slightly oval and elongated, contacting with middle palmar tubercle; proximal subarticular tubercle slightly rounded, elevated; distal subarticular tubercle indistinct; skin flaps on inner and outer side of second and third fingers present, more developed on outer side of third finger than inner, freely moveable; supernumerary metacarpal tubercles absent.</paragraph>
|
||||
<paragraph id="8BC01D46FF808F6B0AA79DFCCF24FE9D" blockId="9.[151,1437,1723,2038]" lastBlockId="10.[151,1437,151,2043]" lastPageId="10" lastPageNumber="397" pageId="9" pageNumber="396">Hindlimb stout, relatively short (HLL 82.0, 143.9 % SVL), about three times length of forelimb, toe length formula I <II <V <III <IV; foot (FL 25.8, 45.3 % SVL) slightly longer than tibia (TL 25.5, 44.7 % SVL); tibiotarsal articulation reaching between eye and nostril in preserved condition; toes tips blunt, rounded; disk of fourth toe (4TDW 1.5, 2.6 % SVL) wider than basal phalanx; toes fully webbed, webbing formula I 1–1 II 1–1 III 1–1 IV 1–1 V; inner and outer webbing of fourth toe fairly well excised; skin flaps on outer side of fifth and inner side of first toe present, moveable; proximal subarticular tubercle distinct, oval elongated; middle subarticular tubercle distinct, oval elongated, longer than proximal subarticular tubercle; distal subarticular tubercle distinct, slightly rounded; inner metatarsal tubercle elongated (IMTL 4.2, 7.3 % SVL); outer metatarsal tubercle absent.</paragraph>
|
||||
<paragraph id="8BC01D46FF838F6B0AA79B25CB4EFD7A" blockId="10.[151,1437,151,2043]" pageId="10" pageNumber="397">Dorsal skin of head and anterior part of the trunk rugose, wrinkled, without warts; posteriorly rugose with small warts; few small warts and wrinkles on upper eyelids; transverse strip between posterior margins of eyes; dorsolateral fold absent; short dorsolateral ridge behind eyes; strong, curved, supratympanic fold from eye to above arm base; ventral skin of abdomen and thigh smooth; cloaca rugose with small warts; ulnar side of hand with scattered translucent spinules; dorsal skin of tibia densely covered with small warts and coarsely scattered with well-elevated large warts, each wart tipped with translucent spinule; tarsus covered with small warts dorsolaterally; skin flap along outer side of fifth toe with translucent spinules; ventral metatarsal skin of fourth toe with irregular translucent spinules.</paragraph>
|
||||
<paragraph id="8BC01D46FF838F6B0AA798C7CB91FCA3" blockId="10.[151,1437,151,2043]" pageId="10" pageNumber="397">
|
||||
<emphasis id="B90BC154FF838F6B0AA798C7CB7DFD5F" bold="true" box="[199,335,652,678]" pageId="10" pageNumber="397">Coloration.</emphasis>
|
||||
Dorsal side greenish-brown with irregularly shaped, indistinct confluent dark spots; head with one dark brown interorbital bar; upper and lower lips with three dark brown bands; canthus and supratympanic fold covered by one dark gray band; throat white, with confluent irregular dark gray blotches; abdomen white; limbs marked dorsally with dark brown crossbars; thigh ventral side nearly translucent with thin yellowish blotches. In preservative, whole-body coloration turned olive-gray; dark gray blotches on throat remain unchanged; translucent spinules turned whitish.
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF838F6B0AA7992FCE10FBF4" blockId="10.[151,1437,151,2043]" pageId="10" pageNumber="397">
|
||||
<emphasis id="B90BC154FF838F6B0AA7992FCB0DFC87" bold="true" box="[199,319,868,894]" pageId="10" pageNumber="397">Variation.</emphasis>
|
||||
Individuals of the
|
||||
<typeStatus id="54C4A3E4FF838F6B086A992EC808FC86" box="[522,570,869,895]" pageId="10" pageNumber="397">type</typeStatus>
|
||||
series were very similar in proportions.
|
||||
<tableCitation id="C6FD28FDFF838F6B0990992FCE72FC87" box="[1008,1088,868,894]" captionStart="TABLE 2" captionStartId="6.[152,249,654,680]" captionTargetPageId="6" captionText="TABLE 2. Morphometric variation of measurements in Limnonectes maanyanorum sp. nov. and Limnonectes nusantara sp. nov. Relative values (R) of each character to SVL (in %). The significant character ≤0.05 is indicated by the asterisk (*)." pageId="10" pageNumber="397">Table 2</tableCitation>
|
||||
shows the individual variations in morphometric values. A single male was greater than females in RHW, RIOD, R1TOEL, ROSL, and RITOL, while females were larger than males in ROW. Male with obtuse tip of odontoid processes. Throat of
|
||||
<specimenCount id="9D79D6CFFF838F6B0F7F99E7CFAEFC3F" box="[1311,1436,940,966]" count="1" pageId="10" pageNumber="397" type="female">one female</specimenCount>
|
||||
(MZB Amph 33535) almost entirely covered by dark blotches. From base to level of subarticular tubercle of male first finger brownish, thickened, forming a nuptial pad without minute asperities.
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF838F6B0AA79E5CC89DFAF0" blockId="10.[151,1437,151,2043]" pageId="10" pageNumber="397">
|
||||
<emphasis id="B90BC154FF838F6B0AA79E5CCB5FFBC8" bold="true" box="[199,365,1047,1073]" pageId="10" pageNumber="397">Comparisons.</emphasis>
|
||||
<taxonomicName id="4C7F66C5FF838F6B0B159E53C89FFBC8" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[373,685,1048,1073]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF838F6B0B159E53C89FFBC8" box="[373,685,1048,1073]" italics="true" pageId="10" pageNumber="397">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF838F6B08D59E5CC922FBC8" bold="true" box="[693,784,1047,1073]" pageId="10" pageNumber="397">
|
||||
<taxonomicNameLabel id="A2387C2FFF838F6B08D59E5CC922FBC8" box="[693,784,1047,1073]" pageId="10" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF838F6B09C89E53CE09FBC8" box="[936,1083,1047,1073]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="barioensis">
|
||||
<emphasis id="B90BC154FF838F6B09C89E53CE09FBC8" box="[936,1083,1047,1073]" italics="true" pageId="10" pageNumber="397">L. barioensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C7F66C5FF838F6B0E149E53CF3DFBC8" box="[1140,1295,1048,1073]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="tawauensis">
|
||||
<emphasis id="B90BC154FF838F6B0E149E53CF3DFBC8" box="[1140,1295,1048,1073]" italics="true" pageId="10" pageNumber="397">L. tawauensis</emphasis>
|
||||
</taxonomicName>
|
||||
by webbing on inner and outer of the fourth toes fairly well excised,
|
||||
<figureCitation id="134401C3FF838F6B09679E70C96BFBAC" box="[775,857,1083,1109]" captionStart="FIGURE 9" captionStartId="15.[151,250,1330,1354]" captionTargetBox="[190,1400,436,1278]" captionTargetId="figure-156@15.[151,1436,416,1306]" captionTargetPageId="15" captionText="FIGURE 9. The upper section of the figure shows the dorsal view of tibia tuberculation of adult female holotype, MZB Amph 33530 (A) and unvouchered adult male (B) of Limnonectes maanyanorum sp. nov.; adult male holotype, MZB Amph 33516 (C), and unvouchered adult female (D) of Limnonectes nusantara sp.nov. The lower section of the figure shows webbing differences between Limnonectes maanyanorum sp. nov. (E), and Limnonectes nusantara sp. nov. (F) with L. tawauensis (G) and L. barioensis (H). Dark arrows indicate fairly well excised on the outer and inner webbing of the fourth toe.Red arrows indicate little excised on the outer and inner webbing of the fourth toe. Photographs of A–F were taken by Ade Damara Gonggoli; G and H were retrieved from Matsui et al. (2024)." figureDoi="http://doi.org/10.5281/zenodo.14747098" httpUri="https://zenodo.org/record/14747098/files/figure.png" pageId="10" pageNumber="397">Fig. 9E</figureCitation>
|
||||
[vs. little excised,
|
||||
<figureCitation id="134401C3FF838F6B0E4A9E70CE96FBAF" box="[1066,1188,1083,1110]" captionStart="FIGURE 9" captionStartId="15.[151,250,1330,1354]" captionTargetBox="[190,1400,436,1278]" captionTargetId="figure-156@15.[151,1436,416,1306]" captionTargetPageId="15" captionText="FIGURE 9. The upper section of the figure shows the dorsal view of tibia tuberculation of adult female holotype, MZB Amph 33530 (A) and unvouchered adult male (B) of Limnonectes maanyanorum sp. nov.; adult male holotype, MZB Amph 33516 (C), and unvouchered adult female (D) of Limnonectes nusantara sp.nov. The lower section of the figure shows webbing differences between Limnonectes maanyanorum sp. nov. (E), and Limnonectes nusantara sp. nov. (F) with L. tawauensis (G) and L. barioensis (H). Dark arrows indicate fairly well excised on the outer and inner webbing of the fourth toe.Red arrows indicate little excised on the outer and inner webbing of the fourth toe. Photographs of A–F were taken by Ade Damara Gonggoli; G and H were retrieved from Matsui et al. (2024)." figureDoi="http://doi.org/10.5281/zenodo.14747098" httpUri="https://zenodo.org/record/14747098/files/figure.png" pageId="10" pageNumber="397">Fig. 9G, H</figureCitation>
|
||||
], first finger distinctly longer than second finger [vs. slightly longer:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B08CA9E14C9A1FB80" author="Matsui, M. & Nishikawa, K. & Shimada, T. & Eto, K. & Hamidy, A. & Sudin, A. & Hossman, Y. H. & Gumal, M. & Vairappan, C. S." box="[682,915,1119,1145]" pageId="10" pageNumber="397" pagination="226 - 277" refId="ref14780" refString="Matsui, M., Nishikawa, K., Shimada, T., Eto, K., Hamidy, A., Sudin, A., Hossman, Y. H., Gumal, M. & Vairappan, C. S. (2024) Revision of the Limnonectes kuhlii - like Fanged Frogs from Malaysian Borneo (Amphibia: Anura: Dicroglossidae). Current Herpetology, 43, 226 - 277. https: // doi. org / 10.5358 / hsj. 43.226" type="journal article" year="2024">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B09619E2BC909FB80" box="[769,827,1119,1145]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2024)
|
||||
</bibRefCitation>
|
||||
], skin flap on outer side of third finger freely moveable [vs. weakly moveable, except
|
||||
<taxonomicName id="4C7F66C5FF838F6B08369ECFC8DCFB64" box="[598,750,1156,1181]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="tawauensis">
|
||||
<emphasis id="B90BC154FF838F6B08369ECFC8DCFB64" box="[598,750,1156,1181]" italics="true" pageId="10" pageNumber="397">L. tawauensis</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B089C9EC8C9E9FB64" author="Matsui, M. & Nishikawa, K. & Shimada, T. & Eto, K. & Hamidy, A. & Sudin, A. & Hossman, Y. H. & Gumal, M. & Vairappan, C. S." box="[764,987,1155,1181]" pageId="10" pageNumber="397" pagination="226 - 277" refId="ref14780" refString="Matsui, M., Nishikawa, K., Shimada, T., Eto, K., Hamidy, A., Sudin, A., Hossman, Y. H., Gumal, M. & Vairappan, C. S. (2024) Revision of the Limnonectes kuhlii - like Fanged Frogs from Malaysian Borneo (Amphibia: Anura: Dicroglossidae). Current Herpetology, 43, 226 - 277. https: // doi. org / 10.5358 / hsj. 43.226" type="journal article" year="2024">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B092F9ECFC9B4FB64" box="[847,902,1155,1181]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2024)
|
||||
</bibRefCitation>
|
||||
], male having obtuse odontoid processes tip [vs. sharply pointed, except
|
||||
<taxonomicName id="4C7F66C5FF838F6B0B979EE3C8B8FB38" box="[503,650,1191,1217]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="barioensis">
|
||||
<emphasis id="B90BC154FF838F6B0B979EE3C8B8FB38" box="[503,650,1191,1217]" italics="true" pageId="10" pageNumber="397">L. barioensis</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B08F99EECC94EFB38" author="Matsui, M. & Nishikawa, K. & Shimada, T. & Eto, K. & Hamidy, A. & Sudin, A. & Hossman, Y. H. & Gumal, M. & Vairappan, C. S." box="[665,892,1191,1217]" pageId="10" pageNumber="397" pagination="226 - 277" refId="ref14780" refString="Matsui, M., Nishikawa, K., Shimada, T., Eto, K., Hamidy, A., Sudin, A., Hossman, Y. H., Gumal, M. & Vairappan, C. S. (2024) Revision of the Limnonectes kuhlii - like Fanged Frogs from Malaysian Borneo (Amphibia: Anura: Dicroglossidae). Current Herpetology, 43, 226 - 277. https: // doi. org / 10.5358 / hsj. 43.226" type="journal article" year="2024">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B088D9EE3C914FB38" box="[749,806,1191,1217]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2024)
|
||||
</bibRefCitation>
|
||||
]; and by female having longer first finger, RFFL 17.2–19.4 % SVL (vs. shorter, ≤ 15.6 % SVL:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B08C19E80C943FB1C" author="Matsui, M. & Nishikawa, K. & Shimada, T. & Eto, K. & Hamidy, A. & Sudin, A. & Hossman, Y. H. & Gumal, M. & Vairappan, C. S." box="[673,881,1227,1253]" pageId="10" pageNumber="397" pagination="226 - 277" refId="ref14780" refString="Matsui, M., Nishikawa, K., Shimada, T., Eto, K., Hamidy, A., Sudin, A., Hossman, Y. H., Gumal, M. & Vairappan, C. S. (2024) Revision of the Limnonectes kuhlii - like Fanged Frogs from Malaysian Borneo (Amphibia: Anura: Dicroglossidae). Current Herpetology, 43, 226 - 277. https: // doi. org / 10.5358 / hsj. 43.226" type="journal article" year="2024">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B08969E80C91CFB1C" box="[758,814,1227,1253]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
), and longer first toe, R1TOEL 19.4–21.9 % SVL (vs. shorter, ≤ 16.0 % SVL:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0BB29EA5C891FAF0" author="Matsui, M. & Nishikawa, K. & Shimada, T. & Eto, K. & Hamidy, A. & Sudin, A. & Hossman, Y. H. & Gumal, M. & Vairappan, C. S." box="[466,675,1262,1289]" pageId="10" pageNumber="397" pagination="226 - 277" refId="ref14780" refString="Matsui, M., Nishikawa, K., Shimada, T., Eto, K., Hamidy, A., Sudin, A., Hossman, Y. H., Gumal, M. & Vairappan, C. S. (2024) Revision of the Limnonectes kuhlii - like Fanged Frogs from Malaysian Borneo (Amphibia: Anura: Dicroglossidae). Current Herpetology, 43, 226 - 277. https: // doi. org / 10.5358 / hsj. 43.226" type="journal article" year="2024">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B08479EA4C86DFAF1" box="[551,607,1262,1288]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF838F6B0AA79F58C866F9D1" blockId="10.[151,1437,151,2043]" pageId="10" pageNumber="397">
|
||||
<taxonomicName id="4C7F66C5FF838F6B0AA79F58CBC9FAD5" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[199,507,1299,1324]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF838F6B0AA79F58CBC9FAD5" box="[199,507,1299,1324]" italics="true" pageId="10" pageNumber="397">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF838F6B08609F59C865FAD5" bold="true" box="[512,599,1298,1324]" pageId="10" pageNumber="397">
|
||||
<taxonomicNameLabel id="A2387C2FFF838F6B08609F59C865FAD5" box="[512,599,1298,1324]" pageId="10" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF838F6B08859F58C9AEFAD5" baseAuthorityName="Gunther" baseAuthorityYear="1872" box="[741,924,1298,1324]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="conspicillatus">
|
||||
<emphasis id="B90BC154FF838F6B08859F58C9AEFAD5" box="[741,924,1298,1324]" italics="true" pageId="10" pageNumber="397">L. conspicillatus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B09C79F58CE77FAD5" box="[935,1093,1298,1324]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="lambirensis">
|
||||
<emphasis id="B90BC154FF838F6B09C79F58CE77FAD5" box="[935,1093,1298,1324]" italics="true" pageId="10" pageNumber="397">L. lambirensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0E309F58CED1FAD5" box="[1104,1251,1298,1324]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="mocquardi">
|
||||
<emphasis id="B90BC154FF838F6B0E309F58CED1FAD5" box="[1104,1251,1298,1324]" italics="true" pageId="10" pageNumber="397">L. mocquardi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0E8F9F58CFA7FAD5" box="[1263,1429,1299,1324]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="penerisensis">
|
||||
<emphasis id="B90BC154FF838F6B0E8F9F58CFA7FAD5" box="[1263,1429,1299,1324]" italics="true" pageId="10" pageNumber="397">L. penerisensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0AF79F7CCB53FAA9" box="[151,353,1335,1360]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="paginatanensis">
|
||||
<emphasis id="B90BC154FF838F6B0AF79F7CCB53FAA9" box="[151,353,1335,1360]" italics="true" pageId="10" pageNumber="397">L. paginatanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0B109F7CCBCFFAA9" box="[368,509,1335,1360]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="separatus">
|
||||
<emphasis id="B90BC154FF838F6B0B109F7CCBCFFAA9" box="[368,509,1335,1360]" italics="true" pageId="10" pageNumber="397">L. separatus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B086D9F7CC898FAA9" box="[525,682,1334,1360]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="lanjakensis">
|
||||
<emphasis id="B90BC154FF838F6B086D9F7CC898FAA9" box="[525,682,1334,1360]" italics="true" pageId="10" pageNumber="397">L. lanjakensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B08DA9F7CC95CFAA9" box="[698,878,1334,1360]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="abanghamidi">
|
||||
<emphasis id="B90BC154FF838F6B08DA9F7CC95CFAA9" box="[698,878,1334,1360]" italics="true" pageId="10" pageNumber="397">L. abanghamidi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B091E9F7CCE1DFAA9" box="[894,1071,1334,1360]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="batulawensis">
|
||||
<emphasis id="B90BC154FF838F6B091E9F7CCE1DFAA9" box="[894,1071,1334,1360]" italics="true" pageId="10" pageNumber="397">L. batulawensis</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C7F66C5FF838F6B0E119F7CCF11FAA9" box="[1137,1315,1334,1360]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="paulyambuni">
|
||||
<emphasis id="B90BC154FF838F6B0E119F7CCF11FAA9" box="[1137,1315,1334,1360]" italics="true" pageId="10" pageNumber="397">L. paulyambuni</emphasis>
|
||||
</taxonomicName>
|
||||
, by dorsal skin of tibia having pattern I of warts (vs. pattern II or III, see
|
||||
<tableCitation id="C6FD28FDFF838F6B093F9F11C987FA8D" box="[863,949,1370,1396]" captionStart="TABLE 3" captionStartId="11.[152,244,692,716]" captionTargetBox="[163,1416,770,1669]" captionTargetPageId="11" captionText="TABLE 3. Morphological variation between the full webbed toes species of Bornean Limnonectes kuhlii complex (data retrieved and modified from Matsui et al. (2024)" pageId="10" pageNumber="397">Table 3</tableCitation>
|
||||
). Additionally,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0E059F10CFAEFA8D" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[1125,1436,1371,1396]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF838F6B0E059F10CFAEFA8D" box="[1125,1436,1371,1396]" italics="true" pageId="10" pageNumber="397">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF838F6B0AF79F35CAC7FA61" bold="true" box="[151,245,1406,1432]" pageId="10" pageNumber="397">
|
||||
<taxonomicNameLabel id="A2387C2FFF838F6B0AF79F35CAC7FA61" box="[151,245,1406,1432]" pageId="10" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF838F6B0BF79F34C803FA61" box="[407,561,1406,1432]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="mocquardi">
|
||||
<emphasis id="B90BC154FF838F6B0BF79F34C803FA61" box="[407,561,1406,1432]" italics="true" pageId="10" pageNumber="397">L. mocquardi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B08239F34C93DFA61" box="[579,783,1407,1432]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="paginatanensis">
|
||||
<emphasis id="B90BC154FF838F6B08239F34C93DFA61" box="[579,783,1407,1432]" italics="true" pageId="10" pageNumber="397">L. paginatanensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B09419F34C982FA61" box="[801,944,1407,1432]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="separatus">
|
||||
<emphasis id="B90BC154FF838F6B09419F34C982FA61" box="[801,944,1407,1432]" italics="true" pageId="10" pageNumber="397">L. separatus</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B09A39F34CE4BFA61" box="[963,1145,1406,1432]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="abanghamidi">
|
||||
<emphasis id="B90BC154FF838F6B09A39F34CE4BFA61" box="[963,1145,1406,1432]" italics="true" pageId="10" pageNumber="397">L. abanghamidi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0EEB9F34CF0DFA61" box="[1163,1343,1406,1432]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="batulawensis">
|
||||
<emphasis id="B90BC154FF838F6B0EEB9F34CF0DFA61" box="[1163,1343,1406,1432]" italics="true" pageId="10" pageNumber="397">L. batulawensis</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C7F66C5FF838F6B0FE69F34CB1BFA45" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="paulyambuni">
|
||||
<emphasis id="B90BC154FF838F6B0FE69F34CB1BFA45" italics="true" pageId="10" pageNumber="397">L. paulyambuni</emphasis>
|
||||
</taxonomicName>
|
||||
, by having first finger distinctly longer than second finger [vs. first finger as long as or slightly shorter than second:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0B489F8DC83AFA19" author="Matsui, M. & Nishikawa, K. & Shimada, T. & Eto, K. & Hamidy, A. & Sudin, A. & Hossman, Y. H. & Gumal, M. & Vairappan, C. S." box="[296,520,1478,1504]" pageId="10" pageNumber="397" pagination="226 - 277" refId="ref14780" refString="Matsui, M., Nishikawa, K., Shimada, T., Eto, K., Hamidy, A., Sudin, A., Hossman, Y. H., Gumal, M. & Vairappan, C. S. (2024) Revision of the Limnonectes kuhlii - like Fanged Frogs from Malaysian Borneo (Amphibia: Anura: Dicroglossidae). Current Herpetology, 43, 226 - 277. https: // doi. org / 10.5358 / hsj. 43.226" type="journal article" year="2024">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B0B1B9F8DCB80FA19" box="[379,434,1478,1504]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2024)
|
||||
</bibRefCitation>
|
||||
].
|
||||
<taxonomicName id="4C7F66C5FF838F6B08789F8DC97FFA26" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[536,845,1478,1503]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF838F6B08789F8DC97FFA26" box="[536,845,1478,1503]" italics="true" pageId="10" pageNumber="397">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF838F6B09339F8DC999FA19" bold="true" box="[851,939,1478,1504]" pageId="10" pageNumber="397">
|
||||
<taxonomicNameLabel id="A2387C2FFF838F6B09339F8DC999FA19" box="[851,939,1478,1504]" pageId="10" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF838F6B0E5A9F8DCEEAFA19" box="[1082,1240,1478,1504]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="lambirensis">
|
||||
<emphasis id="B90BC154FF838F6B0E5A9F8DCEEAFA19" box="[1082,1240,1478,1504]" italics="true" pageId="10" pageNumber="397">L. lambirensis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0E859F8DCF4BFA19" box="[1253,1401,1478,1504]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="mocquardi">
|
||||
<emphasis id="B90BC154FF838F6B0E859F8DCF4BFA19" box="[1253,1401,1478,1504]" italics="true" pageId="10" pageNumber="397">L. mocquardi</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0FE69F8DCB73F9FA" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="paginatanensis">
|
||||
<emphasis id="B90BC154FF838F6B0FE69F8DCB73F9FA" italics="true" pageId="10" pageNumber="397">L. paginatanensis</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C7F66C5FF838F6B0B1E9FA1C802F9FA" box="[382,560,1513,1539]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="abanghamidi">
|
||||
<emphasis id="B90BC154FF838F6B0B1E9FA1C802F9FA" box="[382,560,1513,1539]" italics="true" pageId="10" pageNumber="397">L. abanghamidi</emphasis>
|
||||
</taxonomicName>
|
||||
, by outer side of third finger having moveable skin flap [vs. weekly moveable or non-moveable:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0B049C46C87AF9D1" author="Matsui, M. & Nishikawa, K. & Shimada, T. & Eto, K. & Hamidy, A. & Sudin, A. & Hossman, Y. H. & Gumal, M. & Vairappan, C. S." box="[356,584,1549,1576]" pageId="10" pageNumber="397" pagination="226 - 277" refId="ref14780" refString="Matsui, M., Nishikawa, K., Shimada, T., Eto, K., Hamidy, A., Sudin, A., Hossman, Y. H., Gumal, M. & Vairappan, C. S. (2024) Revision of the Limnonectes kuhlii - like Fanged Frogs from Malaysian Borneo (Amphibia: Anura: Dicroglossidae). Current Herpetology, 43, 226 - 277. https: // doi. org / 10.5358 / hsj. 43.226" type="journal article" year="2024">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B0BD99C45CBC0F9DE" box="[441,498,1549,1575]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2024)
|
||||
</bibRefCitation>
|
||||
].
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF838F6B0AA79C79C883F8BF" blockId="10.[151,1437,151,2043]" pageId="10" pageNumber="397">
|
||||
<taxonomicName id="4C7F66C5FF838F6B0AA79C79CBCDF9B2" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[199,511,1586,1611]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF838F6B0AA79C79CBCDF9B2" box="[199,511,1586,1611]" italics="true" pageId="10" pageNumber="397">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF838F6B08689C7AC851F9B2" bold="true" box="[520,611,1585,1611]" pageId="10" pageNumber="397">
|
||||
<taxonomicNameLabel id="A2387C2FFF838F6B08689C7AC851F9B2" box="[520,611,1585,1611]" pageId="10" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<emphasis id="B90BC154FF838F6B089C9C79CE17F9B2" box="[764,1061,1585,1611]" italics="true" pageId="10" pageNumber="397">
|
||||
<taxonomicName id="4C7F66C5FF838F6B089C9C79C9B9F9B2" baseAuthorityName="Inger, Boeadi & Taufik" baseAuthorityYear="1996" box="[764,907,1586,1611]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="asperatus">L. asperatus</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B09F89C79CE10F9B2" baseAuthorityName="Inger, Boeadi & Taufik" baseAuthorityYear="1996" box="[920,1058,1585,1611]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="rhacodus">L. rhacodus</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
<taxonomicName id="4C7F66C5FF838F6B0E4E9C79CF2AF9B2" authority="Dehling" authorityName="Dehling" authorityYear="2014" box="[1070,1304,1585,1611]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="deinodon">
|
||||
<emphasis id="B90BC154FF838F6B0E4E9C79CE87F9B2" box="[1070,1205,1585,1611]" italics="true" pageId="10" pageNumber="397">L. deinodon</emphasis>
|
||||
Dehling
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0F489C79CFA7F9B2" authorityName="Matsui & Nishikawa" authorityYear="2014" box="[1320,1429,1585,1611]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="hikidai">
|
||||
<emphasis id="B90BC154FF838F6B0F489C79CFA7F9B2" box="[1320,1429,1585,1611]" italics="true" pageId="10" pageNumber="397">L. hikidai</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0AF79C1DCB0EF996" authorityName="Matsui, Nishikawa & Eto" authorityYear="2014" box="[151,316,1621,1647]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="cintalubang">
|
||||
<emphasis id="B90BC154FF838F6B0AF79C1DCB0EF996" box="[151,316,1621,1647]" italics="true" pageId="10" pageNumber="397">L. cintalubang</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0B2A9C1DC85CF996" authority="Matsui" authorityName="Matsui" authorityYear="2015" box="[330,622,1621,1647]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="sinuatodorsalis">
|
||||
<emphasis id="B90BC154FF838F6B0B2A9C1DC827F996" box="[330,533,1621,1647]" italics="true" pageId="10" pageNumber="397">L. sinuatodorsalis</emphasis>
|
||||
Matsui
|
||||
</taxonomicName>
|
||||
, and
|
||||
<emphasis id="B90BC154FF838F6B08C99C1DC937F996" box="[681,773,1621,1647]" italics="true" pageId="10" pageNumber="397">
|
||||
<taxonomicName id="4C7F66C5FF838F6B08C99C1DC933F996" box="[681,769,1621,1647]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="kong">L. kong</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
by having fully webbed toes [vs. fourth toe not webbed to base of tip:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0B7B9C31CBD8F96A" author="Inger, R. F. & Boeadi & Taufik, A." box="[283,490,1657,1684]" pageId="10" pageNumber="397" pagination="363 - 369" refId="ref13891" refString="Inger, R. F., Boeadi. & Taufik, A. (1996) New species of ranid frogs (Amphibia: Anura) from Central Kalimantan, Borneo. Raffles Bulletin of Zoology, 44, 363 - 369." type="journal article" year="1996">
|
||||
Inger
|
||||
<emphasis id="B90BC154FF838F6B0B3E9C31CBA4F96A" box="[350,406,1657,1683]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(1996)
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0B969C32C897F96A" author="Dehling, J. M." box="[502,677,1657,1683]" pageId="10" pageNumber="397" pagination="17 - 30" refId="ref13334" refString="Dehling, J. M. (2014) Eine neue Fangzahnfroschart der Gattung Limnonectes (Anura: Dicroglossidae) vom Gunung Lawit, Malaiische Halbinsel. Sauria, 36, 17 - 30." type="journal article" year="2014">Dehling (2014)</bibRefCitation>
|
||||
, Matsui & Nishikawa (2014),
|
||||
<bibRefCitation id="EFEE60B7FF838F6B099A9C32CED8F96A" author="Matsui, M. & Nishikawa, K. & Eto, K." box="[1018,1258,1657,1683]" pageId="10" pageNumber="397" pagination="679 - 687" refId="ref14514" refString="Matsui, M., Nishikawa, K. & Eto, K. (2014 b): A new burrow-utilizing fanged frog from Sarawak, East Malaysia (Anura: Dicroglossidae). Raffles Bulletin of Zoology, 62, 679 - 687." type="journal article" year="2014">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B0E2F9C31CEBAF96A" box="[1103,1160,1657,1683]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2014b)
|
||||
</bibRefCitation>
|
||||
, Matsui (2015),
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0AF79CD6CBF8F94E" author="Dehling, J. M. & Dehling, D. M." box="[151,458,1693,1719]" pageId="10" pageNumber="397" pagination="291 - 309" refId="ref13369" refString="Dehling, J. M. & Dehling, D. M. (2017): A new wide-headed fanged frog of the Limnonectes kuhlii group (Anura: Dicroglossidae) from western Borneo with a redescription of Rana conspicillata Gunther, 1872. Zootaxa, 4317 (2), 291 - 309. https: // doi. org / 10.11646 / zootaxa. 4317.2.6" type="journal article" year="2017">Dehling & Dehling (2017)</bibRefCitation>
|
||||
]; and odontoid processes tip obtuse in male [vs. pointed tusk, except
|
||||
<taxonomicName id="4C7F66C5FF838F6B0EBF9CD5CF06F94E" box="[1247,1332,1693,1719]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="kong">
|
||||
<emphasis id="B90BC154FF838F6B0EBF9CD5CF06F94E" box="[1247,1332,1693,1719]" italics="true" pageId="10" pageNumber="397">L. kong</emphasis>
|
||||
</taxonomicName>
|
||||
:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0F259CD6CB3AF922" author="Inger, R. F. & Boeadi & Taufik, A." pageId="10" pageNumber="397" pagination="363 - 369" refId="ref13891" refString="Inger, R. F., Boeadi. & Taufik, A. (1996) New species of ranid frogs (Amphibia: Anura) from Central Kalimantan, Borneo. Raffles Bulletin of Zoology, 44, 363 - 369." type="journal article" year="1996">
|
||||
Inger
|
||||
<emphasis id="B90BC154FF838F6B0FE89CD5CA86F922" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(1996)
|
||||
</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0B749C8ACBF1F922" author="Dehling, J. M." box="[276,451,1729,1755]" pageId="10" pageNumber="397" pagination="17 - 30" refId="ref13334" refString="Dehling, J. M. (2014) Eine neue Fangzahnfroschart der Gattung Limnonectes (Anura: Dicroglossidae) vom Gunung Lawit, Malaiische Halbinsel. Sauria, 36, 17 - 30." type="journal article" year="2014">Dehling (2014)</bibRefCitation>
|
||||
, Matsui & Nishikawa (2014),
|
||||
<bibRefCitation id="EFEE60B7FF838F6B09799C8ACE3BF922" author="Matsui, M. & Nishikawa, K. & Eto, K." box="[793,1033,1729,1755]" pageId="10" pageNumber="397" pagination="679 - 687" refId="ref14514" refString="Matsui, M., Nishikawa, K. & Eto, K. (2014 b): A new burrow-utilizing fanged frog from Sarawak, East Malaysia (Anura: Dicroglossidae). Raffles Bulletin of Zoology, 62, 679 - 687." type="journal article" year="2014">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF838F6B090E9C89C995F922" box="[878,935,1729,1755]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2014b)
|
||||
</bibRefCitation>
|
||||
, Matsui (2015),
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0EA39C8ACAD5F906" author="Dehling, J. M. & Dehling, D. M." pageId="10" pageNumber="397" pagination="291 - 309" refId="ref13369" refString="Dehling, J. M. & Dehling, D. M. (2017): A new wide-headed fanged frog of the Limnonectes kuhlii group (Anura: Dicroglossidae) from western Borneo with a redescription of Rana conspicillata Gunther, 1872. Zootaxa, 4317 (2), 291 - 309. https: // doi. org / 10.11646 / zootaxa. 4317.2.6" type="journal article" year="2017">Dehling & Dehling (2017)</bibRefCitation>
|
||||
]. Additionally,
|
||||
<taxonomicName id="4C7F66C5FF838F6B0BF29CAEC8FAF907" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[402,712,1765,1790]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF838F6B0BF29CAEC8FAF907" box="[402,712,1765,1790]" italics="true" pageId="10" pageNumber="397">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF838F6B08AF9CAEC91BF906" bold="true" box="[719,809,1765,1791]" pageId="10" pageNumber="397">
|
||||
<taxonomicNameLabel id="A2387C2FFF838F6B08AF9CAEC91BF906" box="[719,809,1765,1791]" pageId="10" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF838F6B09DE9CAECE23F906" box="[958,1041,1765,1791]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="kong">
|
||||
<emphasis id="B90BC154FF838F6B09DE9CAECE23F906" box="[958,1041,1765,1791]" italics="true" pageId="10" pageNumber="397">L. kong</emphasis>
|
||||
</taxonomicName>
|
||||
by absence of minute asperities on male nuptial pads [vs. present:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0B939D43C913F8DA" author="Dehling, J. M. & Dehling, D. M." box="[499,801,1800,1827]" pageId="10" pageNumber="397" pagination="291 - 309" refId="ref13369" refString="Dehling, J. M. & Dehling, D. M. (2017): A new wide-headed fanged frog of the Limnonectes kuhlii group (Anura: Dicroglossidae) from western Borneo with a redescription of Rana conspicillata Gunther, 1872. Zootaxa, 4317 (2), 291 - 309. https: // doi. org / 10.11646 / zootaxa. 4317.2.6" type="journal article" year="2017">Dehling & Dehling (2017)</bibRefCitation>
|
||||
], and moveable skin flap on the outer side of third finger [vs. non-moveable:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0B169D67C897F8BF" author="Dehling, J. M. & Dehling, D. M." box="[374,677,1836,1862]" pageId="10" pageNumber="397" pagination="291 - 309" refId="ref13369" refString="Dehling, J. M. & Dehling, D. M. (2017): A new wide-headed fanged frog of the Limnonectes kuhlii group (Anura: Dicroglossidae) from western Borneo with a redescription of Rana conspicillata Gunther, 1872. Zootaxa, 4317 (2), 291 - 309. https: // doi. org / 10.11646 / zootaxa. 4317.2.6" type="journal article" year="2017">Dehling & Dehling (2017)</bibRefCitation>
|
||||
].
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF838F6B0AA79D1AC814F803" blockId="10.[151,1437,151,2043]" pageId="10" pageNumber="397">
|
||||
<taxonomicName id="4C7F66C5FF838F6B0AA79D1ACBCFF893" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[199,509,1873,1898]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF838F6B0AA79D1ACBCFF893" box="[199,509,1873,1898]" italics="true" pageId="10" pageNumber="397">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF838F6B08649D1BC86FF893" bold="true" box="[516,605,1872,1898]" pageId="10" pageNumber="397">
|
||||
<taxonomicNameLabel id="A2387C2FFF838F6B08649D1BC86FF893" box="[516,605,1872,1898]" pageId="10" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF838F6B088F9D1AC97EF893" box="[751,844,1872,1898]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="kuhlii">
|
||||
<emphasis id="B90BC154FF838F6B088F9D1AC97EF893" box="[751,844,1872,1898]" italics="true" pageId="10" pageNumber="397">L. kuhlii</emphasis>
|
||||
</taxonomicName>
|
||||
by the absence of thickened skin on the anterior chin in male (vs. present), and nuptial pad present in males (vs. absent).
|
||||
<taxonomicName id="4C7F66C5FF838F6B09CB9D3ECED6F877" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[939,1252,1909,1934]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF838F6B09CB9D3ECED6F877" box="[939,1252,1909,1934]" italics="true" pageId="10" pageNumber="397">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF838F6B0E8E9D3FCF78F877" bold="true" box="[1262,1354,1908,1934]" pageId="10" pageNumber="397">
|
||||
<taxonomicNameLabel id="A2387C2FFF838F6B0E8E9D3FCF78F877" box="[1262,1354,1908,1934]" pageId="10" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF838F6B0AB59DD2C96DF84B" authority="McLeod, Horner, Husted, Barley & Iskandar" authorityName="McLeod, Horner, Husted, Barley & Iskandar" authorityYear="2011" box="[213,863,1944,1971]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="397" phylum="Chordata" rank="species" species="sisikdagu">
|
||||
<emphasis id="B90BC154FF838F6B0AB59DD2CB6CF84B" box="[213,350,1944,1970]" italics="true" pageId="10" pageNumber="397">L. sisikdagu</emphasis>
|
||||
McLeod, Horner, Husted, Barley & Iskandar
|
||||
</taxonomicName>
|
||||
, by absence of translucent spinules on ventral skin of anterior chin of males [vs. present:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B085F9DF7C903F82F" author="McLeod, D. S. & Horner, S. J. & Husted, C. & Barley, A. & Iskandar, D." box="[575,817,1980,2006]" pageId="10" pageNumber="397" pagination="52 - 64" refId="ref14975" refString="McLeod, D. S., Horner, S. J., Husted, C., Barley, A. & Iskandar, D. (2011) " Same-same, but different ": an unusual new species of the Limnonectes kuhlii complex from West Sumatra (Anura: Dicroglossidae). Zootaxa, 2883 (1), 52 - 64. https: // doi. org / 10.11646 / zootaxa. 2883.1.4" type="journal article" year="2011">
|
||||
McLeod
|
||||
<emphasis id="B90BC154FF838F6B08C59DF6C8EEF82F" box="[677,732,1980,2006]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2011)
|
||||
</bibRefCitation>
|
||||
], and absence of minute asperities on male nuptial pads [vs. present:
|
||||
<bibRefCitation id="EFEE60B7FF838F6B0B479DABC82EF803" author="McLeod, D. S. & Horner, S. J. & Husted, C. & Barley, A. & Iskandar, D." box="[295,540,2016,2042]" pageId="10" pageNumber="397" pagination="52 - 64" refId="ref14975" refString="McLeod, D. S., Horner, S. J., Husted, C., Barley, A. & Iskandar, D. (2011) " Same-same, but different ": an unusual new species of the Limnonectes kuhlii complex from West Sumatra (Anura: Dicroglossidae). Zootaxa, 2883 (1), 52 - 64. https: // doi. org / 10.11646 / zootaxa. 2883.1.4" type="journal article" year="2011">
|
||||
McLeod
|
||||
<emphasis id="B90BC154FF838F6B0BED9DAACBF7F803" box="[397,453,2016,2042]" italics="true" pageId="10" pageNumber="397">et al.</emphasis>
|
||||
(2011)
|
||||
</bibRefCitation>
|
||||
].
|
||||
</paragraph>
|
||||
<paragraph id="8BC01D46FF828F6A0AA79ADCCF37FEB8" blockId="11.[151,1437,151,645]" pageId="11" pageNumber="398">
|
||||
<taxonomicName id="4C7F66C5FF828F6A0AA79ADCCBCDFF49" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[199,511,151,176]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF828F6A0AA79ADCCBCDFF49" box="[199,511,151,176]" italics="true" pageId="11" pageNumber="398">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF828F6A08689ADCC851FF48" bold="true" box="[520,611,151,177]" pageId="11" pageNumber="398">
|
||||
<taxonomicNameLabel id="A2387C2FFF828F6A08689ADCC851FF48" box="[520,611,151,177]" pageId="11" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF828F6A089C9ADCCEA2FF48" authority="Matsui, Belabut & Ahmad" authorityName="Matsui, Belabut & Ahmad" authorityYear="2014" box="[764,1168,151,177]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="utara">
|
||||
<emphasis id="B90BC154FF828F6A089C9ADCC96BFF49" box="[764,857,151,176]" italics="true" pageId="11" pageNumber="398">L. utara</emphasis>
|
||||
Matsui, Belabut & Ahmad
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C7F66C5FF828F6A0EAF9ADCCB57FF2C" authority="Matsui, Belabut & Ahmad" authorityName="Matsui, Belabut & Ahmad" authorityYear="2014" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="selatan">
|
||||
<emphasis id="B90BC154FF828F6A0EAF9ADCCF0DFF48" box="[1231,1343,151,177]" italics="true" pageId="11" pageNumber="398">L. selatan</emphasis>
|
||||
Matsui, Belabut & Ahmad
|
||||
</taxonomicName>
|
||||
, by smaller of female body size, SVL
|
||||
<quantity id="4C87B0A3FF828F6A09709AF1C981FF2C" box="[784,947,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.945" metricValueMax="5.85" metricValueMin="4.04" pageId="11" pageNumber="398" unit="mm" value="49.45" valueMax="58.5" valueMin="40.4">40.4–58.5 mm</quantity>
|
||||
[vs. ≥
|
||||
<quantity id="4C87B0A3FF828F6A099F9AF1CE55FF2C" box="[1023,1127,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="5.92" pageId="11" pageNumber="398" unit="mm" value="59.2">59.2 mm</quantity>
|
||||
:
|
||||
<bibRefCitation id="EFEE60B7FF828F6A0E119AF0CF52FF2C" author="Matsui, M. & Belabut, D. M. & Ahmad, N." box="[1137,1376,187,213]" pageId="11" pageNumber="398" pagination="75 - 93" refId="ref14456" refString="Matsui, M., Belabut, D. M. & Ahmad, N. (2014 a) Two new species of fanged frogs from Peninsular Malaysia (Anura: Dicroglossidae). Zootaxa, 3881 (1), 75 - 93. https: // doi. org / 10.11646 / zootaxa. 3881.1.6" type="journal article" year="2014">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF828F6A0EA69AF0CECCFF2C" box="[1222,1278,187,213]" italics="true" pageId="11" pageNumber="398">et al.</emphasis>
|
||||
(2014a)
|
||||
</bibRefCitation>
|
||||
], few large warts on dorsal skin of tibia [vs. many large wars:
|
||||
<bibRefCitation id="EFEE60B7FF828F6A09409A94CE27FF00" author="Matsui, M. & Belabut, D. M. & Ahmad, N." box="[800,1045,223,249]" pageId="11" pageNumber="398" pagination="75 - 93" refId="ref14456" refString="Matsui, M., Belabut, D. M. & Ahmad, N. (2014 a) Two new species of fanged frogs from Peninsular Malaysia (Anura: Dicroglossidae). Zootaxa, 3881 (1), 75 - 93. https: // doi. org / 10.11646 / zootaxa. 3881.1.6" type="journal article" year="2014">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF828F6A09169A94C982FF00" box="[886,944,223,249]" italics="true" pageId="11" pageNumber="398">et al.</emphasis>
|
||||
(2014a)
|
||||
</bibRefCitation>
|
||||
], and absence of minute asperities on male nuptial pads [vs. present:
|
||||
<bibRefCitation id="EFEE60B7FF828F6A08799B48C938FEE4" author="Matsui, M. & Belabut, D. M. & Ahmad, N." box="[537,778,259,285]" pageId="11" pageNumber="398" pagination="75 - 93" refId="ref14456" refString="Matsui, M., Belabut, D. M. & Ahmad, N. (2014 a) Two new species of fanged frogs from Peninsular Malaysia (Anura: Dicroglossidae). Zootaxa, 3881 (1), 75 - 93. https: // doi. org / 10.11646 / zootaxa. 3881.1.6" type="journal article" year="2014">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF828F6A080E9B48C894FEE4" box="[622,678,259,285]" italics="true" pageId="11" pageNumber="398">et al.</emphasis>
|
||||
(2014a)
|
||||
</bibRefCitation>
|
||||
]. Additionally,
|
||||
<taxonomicName id="4C7F66C5FF828F6A09D59B48CED9FEE5" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[949,1259,259,284]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF828F6A09D59B48CED9FEE5" box="[949,1259,259,284]" italics="true" pageId="11" pageNumber="398">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF828F6A0E929B48CF7EFEE4" bold="true" box="[1266,1356,259,285]" pageId="11" pageNumber="398">
|
||||
<taxonomicNameLabel id="A2387C2FFF828F6A0E929B48CF7EFEE4" box="[1266,1356,259,285]" pageId="11" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
differs from
|
||||
<taxonomicName id="4C7F66C5FF828F6A0AB59B6CCB1DFEB9" authorityName="Matsui, Belabut & Ahmad" authorityYear="2014" box="[213,303,295,320]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="utara">
|
||||
<emphasis id="B90BC154FF828F6A0AB59B6CCB1DFEB9" box="[213,303,295,320]" italics="true" pageId="11" pageNumber="398">L. utara</emphasis>
|
||||
</taxonomicName>
|
||||
by less confluent dark markings on dorsum [vs. more confluent:
|
||||
<bibRefCitation id="EFEE60B7FF828F6A0E6A9B6CCEC8FEB8" author="Matsui, M. & Belabut, D. M. & Ahmad, N." box="[1034,1274,295,321]" pageId="11" pageNumber="398" pagination="75 - 93" refId="ref14456" refString="Matsui, M., Belabut, D. M. & Ahmad, N. (2014 a) Two new species of fanged frogs from Peninsular Malaysia (Anura: Dicroglossidae). Zootaxa, 3881 (1), 75 - 93. https: // doi. org / 10.11646 / zootaxa. 3881.1.6" type="journal article" year="2014">
|
||||
Matsui
|
||||
<emphasis id="B90BC154FF828F6A0E3F9B6CCEA5FEB8" box="[1119,1175,295,321]" italics="true" pageId="11" pageNumber="398">et al.</emphasis>
|
||||
(2014a)
|
||||
</bibRefCitation>
|
||||
].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3654ECDFF828F6A0AB19B01CE53FD7C" pageId="11" pageNumber="398" type="distribution">
|
||||
<paragraph id="8BC01D46FF828F6A0AB19B01CE53FD7C" blockId="11.[151,1437,151,645]" pageId="11" pageNumber="398">
|
||||
<emphasis id="B90BC154FF828F6A0AB19B01C86DFE9D" bold="true" box="[209,607,330,357]" pageId="11" pageNumber="398">Distribution and natural history.</emphasis>
|
||||
<taxonomicName id="4C7F66C5FF828F6A08089B00C993FE9D" authorityName="Gonggoli & Shimada & Matsui & Nishikawa & Sidik & Kadafi & Farajallah & Hamidy" authorityYear="2025" box="[616,929,331,356]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="maanyanorum" status="sp. nov.">
|
||||
<emphasis id="B90BC154FF828F6A08089B00C993FE9D" box="[616,929,331,356]" italics="true" pageId="11" pageNumber="398">Limnonectes maanyanorum</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B90BC154FF828F6A09CA9B00CE34FE9C" bold="true" box="[938,1030,331,357]" pageId="11" pageNumber="398">
|
||||
<taxonomicNameLabel id="A2387C2FFF828F6A09CA9B00CE34FE9C" box="[938,1030,331,357]" pageId="11" pageNumber="398" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is distributed within an elevational range of
|
||||
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||||
a.s.l. in the northern part of Meratus Mountain Range based on available data. The type series from Mt. Karasik were found in secondary forest, near disturbed areas along small, rocky stream with a width of up to
|
||||
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(
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<figureCitation id="134401C3FF828F6A0B7A9BFCCBB7FE28" box="[282,389,439,465]" captionStart="FIGURE 10" captionStartId="16.[152,255,1952,1976]" captionTargetBox="[340,1247,633,1929]" captionTargetId="figure-223@16.[340,1247,633,1929]" captionTargetPageId="16" captionText="FIGURE 10. Natural habitat of Limnonectes maanyanorum sp. nov. in Mt. Karasik (A), and Limnonectes nusantara sp. nov. in Loksado (B). Photographs by Ade Damara Gonggoli." figureDoi="http://doi.org/10.5281/zenodo.14747100" httpUri="https://zenodo.org/record/14747100/files/figure.png" pageId="11" pageNumber="398">Fig. 10A</figureCitation>
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||||
). The
|
||||
<typeStatus id="54C4A3E4FF828F6A0BAF9BFCC803FE28" box="[463,561,439,465]" pageId="11" pageNumber="398" type="holotype">holotype</typeStatus>
|
||||
was collected from the ground at the edge of the stream at around 20:00 h, while other type specimens have been found on leaves or rocks near the stream. The following anuran species have been found at the Mt. Karasik site with the new species:
|
||||
<taxonomicName id="4C7F66C5FF828F6A09539BB4CE96FDE0" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1920" box="[819,1188,511,537]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="picturata">
|
||||
<emphasis id="B90BC154FF828F6A09539BB4CE3DFDE1" box="[819,1039,511,537]" italics="true" pageId="11" pageNumber="398">Hylarana picturata</emphasis>
|
||||
(Boulenger)
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C7F66C5FF828F6A0ED29BB4CAFAFDC4" authority="Zug" authorityName="Zug" authorityYear="2015" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="meizon">
|
||||
<emphasis id="B90BC154FF828F6A0ED29BB4CFAEFDE1" box="[1202,1436,511,537]" italics="true" pageId="11" pageNumber="398">Kalophrynus meizon</emphasis>
|
||||
Zug
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C7F66C5FF828F6A0AB69868CBCAFDC4" authority="(Andersson)" baseAuthorityName="Andersson" baseAuthorityYear="1923" box="[214,504,547,573]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="leporinus">
|
||||
<emphasis id="B90BC154FF828F6A0AB69868CB52FDC4" box="[214,352,547,573]" italics="true" pageId="11" pageNumber="398">L. leporinus</emphasis>
|
||||
(Andersson)
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C7F66C5FF828F6A08669868C914FDC4" authority="(Inger)" baseAuthorityName="Inger" baseAuthorityYear="1966" box="[518,806,547,573]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="paramacrodon">
|
||||
<emphasis id="B90BC154FF828F6A08669868C8F9FDC4" box="[518,715,547,573]" italics="true" pageId="11" pageNumber="398">L. paramacrodon</emphasis>
|
||||
(Inger)
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C7F66C5FF828F6A09549868CE8AFDC4" authority="(Cochran)" baseAuthorityName="Cochran" baseAuthorityYear="1926" box="[820,1208,547,573]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="abbotti">
|
||||
<emphasis id="B90BC154FF828F6A09549868CE0AFDC4" box="[820,1080,547,573]" italics="true" pageId="11" pageNumber="398">Leptobrachium abbotti</emphasis>
|
||||
(Cochran)
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C7F66C5FF828F6A0EA59868CFA7FDC4" box="[1221,1429,547,573]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="undefined-1">
|
||||
<emphasis id="B90BC154FF828F6A0EA59868CF61FDC5" box="[1221,1363,547,572]" italics="true" pageId="11" pageNumber="398">Limnonectes</emphasis>
|
||||
sp. 1
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C7F66C5FF828F6A0AF7980CCE58FD98" authority="(Munir, Hamidy, Matsui, Iskandar, Sidik, & Shimada)" baseAuthorityName="Munir, Hamidy, Matsui, Iskandar, Sidik, & Shimada" box="[151,1130,583,609]" class="Amphibia" family="Megophryidae" genus="Pelobatrachus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="kalimantanensis">
|
||||
<emphasis id="B90BC154FF828F6A0AF7980CCBCBFD98" box="[151,505,583,609]" italics="true" pageId="11" pageNumber="398">Pelobatrachus kalimantanensis</emphasis>
|
||||
(Munir, Hamidy, Matsui, Iskandar, Sidik, & Shimada)
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C7F66C5FF828F6A0E16980CCB4BFD7C" authority="(Dehling & Matsui)" baseAuthorityName="Dehling & Matsui" baseAuthorityYear="2013" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="fritinniens">
|
||||
<emphasis id="B90BC154FF828F6A0E16980CCFAEFD98" box="[1142,1436,583,609]" italics="true" pageId="11" pageNumber="398">Leptobrachella fritinniens</emphasis>
|
||||
(Dehling & Matsui)
|
||||
</taxonomicName>
|
||||
;
|
||||
<taxonomicName id="4C7F66C5FF828F6A0BE49820C8DEFD7C" authority="(Gravenhorst)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1829" box="[388,748,619,645]" class="Amphibia" family="Bufonidae" genus="Phrynoidis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="asper">
|
||||
<emphasis id="B90BC154FF828F6A0BE49820C876FD7D" box="[388,580,619,645]" italics="true" pageId="11" pageNumber="398">Phrynoidis asper</emphasis>
|
||||
(Gravenhorst)
|
||||
</taxonomicName>
|
||||
; and
|
||||
<taxonomicName id="4C7F66C5FF828F6A09479820CE6CFD7C" authority="(Gunther)" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[807,1118,619,645]" class="Amphibia" family="Ranidae" genus="Staurois" kingdom="Animalia" order="Anura" pageId="11" pageNumber="398" phylum="Chordata" rank="species" species="guttatus">
|
||||
<emphasis id="B90BC154FF828F6A09479820C9D6FD7D" box="[807,996,619,645]" italics="true" pageId="11" pageNumber="398">Staurois guttatus</emphasis>
|
||||
(Günther)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,55 +1,55 @@
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|
||||
<treatment id="361ABCDCC5CF5D3B856FAF0ABB5F7CDD" ID-DOI="http://doi.org/10.5281/zenodo.14757044" ID-Zenodo-Dep="14757044" ID-arpha="361ABCDC-C5CF-5D3B-856F-AF0ABB5F7CDD" ID-zoobank="https://zoobank.org/E78D0880-A34C-404E-9D59-5B15FD1A75C6" LSID="urn:lsid:plazi:treatment:361ABCDCC5CF5D3B856FAF0ABB5F7CDD" LSID-ZBK="urn:lsid:zoobank.org:act:E78D0880-A34C-404E-9D59-5B15FD1A75C6" httpUri="http://treatment.plazi.org/id/361ABCDCC5CF5D3B856FAF0ABB5F7CDD" scope_family="Clubionidae" scope_order="Araneae">
|
||||
<subSubSection id="E3B4ECB76BD516451B83DFCB0308B3E0" type="nomenclature">
|
||||
<paragraph id="F17CF9B112885D2EEDCB14E880AC6A3C">
|
||||
<taxonomicName id="80D8DD865F4FF31224C6CF64B30CF56C" authorityName="Guo & Li & Zhang" authorityYear="2025" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="longyangensis" status="sp. nov.">
|
||||
|
|
@ -72,13 +72,15 @@
|
|||
<heading id="DD6DC89DB6C256E2BE912FE0565783AA" reason="title">Type material.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="A78B837E785F240A1F4B0478D8148832">
|
||||
<materialsCitation id="DF08490DB5BA6EA81198402D8D91AE07" collectingDate="2017-11-24" collectorName="Zhaoyi Li." country="China" county="Baoshan City" elevation="1669" latitude="25.3016" location="Lujiang Town" longLatPrecision="7" longitude="98.7994" municipality="Longyang District" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
|
||||
<materialsCitation id="43F9EBD4A7A0E0C2971A69E83D89F874" collectingDate="2017-11-24" collectionCode="CLU" collectorName="Zhaoyi Li" country="China" county="Baoshan City" elevation="1669" latitude="25.3016" location="Baihua Ling Village" longLatPrecision="7" longitude="98.7994" municipality="Longyang District" specimenCode="CLU 847-1" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
|
||||
<emphasis id="67C95388BB59AD7E9243C36CBBF510FF" bold="true" italics="true">
|
||||
<typeStatus id="B53737B7B61855248423E0D98AAAE122" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
•
|
||||
<specimenCount id="8708B12598027E77A28DC30020B85D56" count="1" type="male">♂</specimenCount>
|
||||
(CLU 847-1);
|
||||
(
|
||||
<specimenCode id="29FA3A2935BE70E6B6A18ED732F8F25B">CLU 847-1</specimenCode>
|
||||
);
|
||||
<collectingCountry id="CF02D333B48A794D7056D63C8D79B7A2" name="China">China</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="C871EE47690D60AA8D4E3E0171A81A88" country="China" name="Yunnan">Yunnan Province</collectingRegion>
|
||||
|
|
@ -87,9 +89,9 @@
|
|||
,
|
||||
<collectingMunicipality id="C23BBD03FDC303BF7D6362ECB84C15BD">Longyang District</collectingMunicipality>
|
||||
,
|
||||
<location id="3F2EFFF8B661BF3857CF47CE19DAD479" LSID="urn:lsid:plazi:treatment:361ABCDCC5CF5D3B856FAF0ABB5F7CDD:3F2EFFF8B661BF3857CF47CE19DAD479" country="China" county="Baoshan City" latitude="25.3016" longLatPrecision="7" longitude="98.7994" municipality="Longyang District" name="Lujiang Town" stateProvince="Yunnan">Lujiang Town</location>
|
||||
<collectingMunicipality id="04A82B9CE25137AD3D8CFBDE31B0F755">Lujiang Town</collectingMunicipality>
|
||||
,
|
||||
<location id="4A672145137B4CB2A8F3A10A83AC7E64" LSID="urn:lsid:plazi:treatment:361ABCDCC5CF5D3B856FAF0ABB5F7CDD:4A672145137B4CB2A8F3A10A83AC7E64" country="China" county="Baoshan City" latitude="25.3016" longLatPrecision="7" longitude="98.7994" municipality="Longyang District" name="Baihua Ling Village" stateProvince="Yunnan">Baihua Ling Village</location>
|
||||
<location id="4A672145137B4CB2A8F3A10A83AC7E64" LSID="urn:lsid:plazi:treatment:361ABCDCC5CF5D3B856FAF0ABB5F7CDD:4A672145137B4CB2A8F3A10A83AC7E64" country="China" county="Baoshan City" latitude="25.3016" longLatPrecision="7" longitude="98.7994" municipality="Longyang District" name="Baihua Ling Village" stateProvince="Yunnan Province">Baihua Ling Village</location>
|
||||
;
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFDAE" specific-use="{"type":"Point","coordinates":[98.799400,25.301600]}">
|
||||
<geoCoordinate id="8C2D7667A7B1221B008720008FA09ED0" degrees="25.3016" direction="north" orientation="latitude" precision="5" value="25.3016">25.3016 ° N</geoCoordinate>
|
||||
|
|
@ -105,16 +107,21 @@ elev.;
|
|||
<collectingDate id="035F0D0E17C21C9C245973F8AC250AA8" value="2017-11-24">24. XI. 2017</collectingDate>
|
||||
</date>
|
||||
; leg.
|
||||
<collectorName id="6B00065661AC45903DDE261860A697C2">Zhaoyi Li.</collectorName>
|
||||
<collectorName id="CEAA05C1DA19708E8BA58579A44050EB">Zhaoyi Li</collectorName>
|
||||
</materialsCitation>
|
||||
<materialsCitation id="850EF8F22D080B65E0AAE778407EB7E4" collectingDate="2017-11-24" collectorName="Zhaoyi Li." country="China" county="Baoshan City" elevation="1669" latitude="25.3016" location="Lujiang Town" longLatPrecision="7" longitude="98.7994" municipality="Longyang District" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
|
||||
.
|
||||
<materialsCitation id="850EF8F22D080B65E0AAE778407EB7E4" collectingDate="2017-11-24" collectionCode="CLU" collectorName="Zhaoyi Li." country="China" county="Baoshan City" elevation="1669" latitude="25.3016" location="Lujiang Town" longLatPrecision="7" longitude="98.7994" municipality="Longyang District" specimenCode="CLU 847-2, CLU 847-6" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
|
||||
<emphasis id="B5708E1535C2B053553D1997F8CC2FB5" bold="true" italics="true">
|
||||
<typeStatus id="C8E8DB3067A6A245CF7260D1E79A53B4" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
•
|
||||
<specimenCount id="F724853BEDDD72D484B3BA1F22DE83CE" count="3" type="male">3 ♂</specimenCount>
|
||||
<specimenCount id="FBC13EE8CC75BE55AE44D7DD6F0CC831" count="2" type="female">2 ♀</specimenCount>
|
||||
(CLU 847-2 – CLU 847 - 6); same data as holotype
|
||||
(
|
||||
<specimenCode id="086F444F5EEA8927894946966AE943FB">CLU 847-2</specimenCode>
|
||||
–
|
||||
<specimenCode id="2DFC9AD57D31D868BE37B05D65326287">CLU 847-6</specimenCode>
|
||||
); same data as holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
|
|
@ -124,30 +131,40 @@ elev.;
|
|||
<heading id="E6E1D56C5DD46021E24526F6B59CEC20" reason="title">Other material examined.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="FEE7C68837BB28D4A933983202642E4E">
|
||||
<materialsCitation id="6FAA94749D24F6A7D67EF8C8B9AD288E" collectingDate="2017-11-24" collectionCode="CLU" collectorName="Zhaoyi Li" elevation="1983" latitude="25.2981" location="Baihua Ling Village" longLatPrecision="7" longitude="98.7863" specimenCode="CLU 845-1, CLU 845-4" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
|
||||
<specimenCount id="E4E62A9A389F4FF68987E55877FF357D" count="3" type="male">3 ♂</specimenCount>
|
||||
<specimenCount id="FB864A49C96773C6EDCBBA157579D71B" count="1" type="female">1 ♀</specimenCount>
|
||||
(CLU 845-1 – CLU 845 - 4); Baihua Ling Village;
|
||||
(
|
||||
<specimenCode id="0D1B4656DA27EAB5ED75F97EDEC68E6B">CLU 845-1</specimenCode>
|
||||
–
|
||||
<specimenCode id="B834C5C4E164E559BB8958E9F17623FF">CLU 845-4</specimenCode>
|
||||
);
|
||||
<location id="C6F9A3D7395CD6DB3DB9B9D4D971A4FC" LSID="urn:lsid:plazi:treatment:361ABCDCC5CF5D3B856FAF0ABB5F7CDD:C6F9A3D7395CD6DB3DB9B9D4D971A4FC" latitude="25.2981" longLatPrecision="7" longitude="98.7863" name="Baihua Ling Village">Baihua Ling Village</location>
|
||||
;
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWDAE" specific-use="{"type":"Point","coordinates":[98.786300,25.298100]}">
|
||||
<geoCoordinate id="2D96030995C9F9DA87C4FF1884792E63" degrees="25.2981" direction="north" orientation="latitude" precision="5" value="25.2981">25.2981 ° N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="1DAC0E1C6B131E4EE2EE28FBFEE48E06" degrees="98.7863" direction="east" orientation="longitude" precision="5" value="98.7863">98.7863 ° E</geoCoordinate>
|
||||
</named-content>
|
||||
;
|
||||
<quantity id="F9452E2FABD028129691C82FE3E4F56D" metricMagnitude="3" metricUnit="m" metricValue="1.983" unit="m" value="1983.0">1983 m</quantity>
|
||||
<quantity id="F9452E2FABD028129691C82FE3E4F56D" metricMagnitude="3" metricUnit="m" metricValue="1.983" unit="m" value="1983.0">
|
||||
<elevation id="73408337AC16E25E8B489AE9C54A90D1" metricMagnitude="3" metricUnit="m" metricValue="1.983" unit="m" value="1983.0">1983 m</elevation>
|
||||
</quantity>
|
||||
elev.;
|
||||
<date id="647CD40885F0C90DD4BD7C522C9C31E7" value="2017-11-24">24. XI. 2017</date>
|
||||
; leg. Zhaoyi Li.
|
||||
<date id="647CD40885F0C90DD4BD7C522C9C31E7" value="2017-11-24">
|
||||
<collectingDate id="DE4DBC208BCFC89D4C8F202A36863280" value="2017-11-24">24. XI. 2017</collectingDate>
|
||||
</date>
|
||||
; leg.
|
||||
<collectorName id="C30CF820ECFA5CD7A8F33C398220FE84">Zhaoyi Li</collectorName>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E2DAE" type="etymology">
|
||||
<paragraph id="178ED1C9CAFF0BD195EE7209B01A52E5">
|
||||
<heading id="0B1EAEFAE162D2C672D791C9711428D1" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="2FD0E7EBE99869832A20216B65C0AF67">
|
||||
The specific name is derived from the
|
||||
<typeStatus id="BCEBD61AED98B796E97162F850F7A73B">type</typeStatus>
|
||||
locality; an adjective.
|
||||
</paragraph>
|
||||
<paragraph id="2FD0E7EBE99869832A20216B65C0AF67">The specific name is derived from the type locality; an adjective.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EAEAE" type="diagnosis">
|
||||
<paragraph id="B665945F510BA11795E766AD7611C4FC">
|
||||
|
|
@ -376,9 +393,7 @@ same, retrolateral view. Abbreviations: C = conductor; E = embolus;
|
|||
(Fig.
|
||||
<figureCitation id="1D1D6807C40A1B726D617AEE4E3FAA16" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Clubiona longyangensis sp. nov., holotype male A left palp, ventral view B same, prolateral view C same, retrolateral view. Abbreviations: C = conductor; E = embolus; LTA = lateral tibial apophysis; PPA = prolateral patellar apophysis; RFA = retrolateral femoral apophysis; RPA = retrolateral patellar apophysis; RTA = retrolateral tibial apophysis; VTA = ventral tibial apophysis." figureDoi="10.3897/zookeys.1224.135572.figure2" httpUri="https://binary.pensoft.net/fig/1236382">2</figureCitation>
|
||||
). Femur retrolaterally with a slanting finger-like apophysis (
|
||||
<abbrev id="ABBRID0EVQAE" xlink_title="retrolateral femoral apophysis">
|
||||
<collectionCode id="9DF46BA88B2F6B53DE836D613D1E77E1" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" type="Herbarium">RFA</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EVQAE" xlink_title="retrolateral femoral apophysis">RFA</abbrev>
|
||||
), more than 1 / 2 of femur’s length.
|
||||
<abbrev id="ABBRID0EZQAE" xlink_title="retrolateral femoral apophysis">RFA</abbrev>
|
||||
partly membranous, arising mesially from femur, directing retrolatero-dorsally. Patella approximately twice as long as the tibia, with two apophyses:
|
||||
|
|
@ -430,17 +445,11 @@ body length 4.39. Carapace 2.10 long, 1.55 wide; abdomen 2.29 long, 1.53 wide. C
|
|||
). Epigynal plate slightly longer than wide. Atrium (A) distinctly large, umbelliform, with a delimited, arched anterior margin (hood) and nearly invisble postterior margin, located at anterior portion of epigynal plate. Copulatory openings (
|
||||
<abbrev id="ABBRID0EITAE" xlink_title="copulatory opening">CO</abbrev>
|
||||
) indistinct, tiny, located centrally in atrium. Copulatory ducts (
|
||||
<abbrev id="ABBRID0EMTAE" xlink_title="copulatory duct">
|
||||
<collectionCode id="D05F24342CEF066F84F0C3AEE9D5A251">CD</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EMTAE" xlink_title="copulatory duct">CD</abbrev>
|
||||
), heavily sclerotised, relatively long and thick, ca 1 / 2 length of epigynal plate, descend obliquely, forming a ˄ - shaped course. Spermathecae (S) longer laterally juxtaposed at the tip represented by two ~ - shaped tubes, anterior surface with a papilliform spermathecal appendage (
|
||||
<abbrev id="ABBRID0EQTAE" xlink_title="spermathecal appendage">
|
||||
<collectionCode id="87C1B74229F3EEFCF1CE3EADA719B31A" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie">SA</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EQTAE" xlink_title="spermathecal appendage">SA</abbrev>
|
||||
), respectively. Fertilisation ducts (
|
||||
<abbrev id="ABBRID0EUTAE" xlink_title="fertilisation duct">
|
||||
<collectionCode id="4CACE97696DECE1C7DAC4652C4846A7B">FD</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EUTAE" xlink_title="fertilisation duct">FD</abbrev>
|
||||
) acicular, located terminally on spermathecae. Bursae (B) close together, spherical, weakly sclerotized, surface smooth.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,55 +1,55 @@
|
|||
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<mods:title id="E211C1381BB75024645D8ED6DCD9BC73">Two new species of the Clubiona corticalis group (Araneae, Clubionidae) from Yunnan, China</mods:title>
|
||||
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|
||||
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|
||||
<mods:name id="4E85D4F41D0671694247354E2689FBE9" type="personal">
|
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<mods:role id="6139A341008025D1EB56837AD9ED8875">
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<mods:roleTerm id="7556AB775C29AE4760688C8670023EDE">Author</mods:roleTerm>
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<mods:name id="07945354068A29D054ACE2155D3B2798" type="personal">
|
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<mods:role id="B0AEA041568E7A9000195CFF1506C447">
|
||||
<mods:roleTerm id="68BD3F00A6C322A115DD4802E9B54BD5">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="A92041E384051867EFE68446D4D56694">Guo, Minghao</mods:namePart>
|
||||
<mods:affiliation id="8167A15EACACFF6C1E066AFEFB20EA61">Hebei Vocational University of Industry and Technology, Shijiazhuang, Hebei 050091, China</mods:affiliation>
|
||||
<mods:namePart id="64F7D7455469FDE6E0AC824EE2B779D8">Guo, Minghao</mods:namePart>
|
||||
<mods:affiliation id="E508454180FB4B53D42392A2818719A0">Hebei Vocational University of Industry and Technology, Shijiazhuang, Hebei 050091, China</mods:affiliation>
|
||||
</mods:name>
|
||||
<mods:name id="CC7CBB272464A395699136EAF5E2AD59" type="personal">
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<mods:name id="7BA70054604AD3CF0FC2FCD8AF4570E6" type="personal">
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<mods:role id="CE841713F4C743549BB563A56969A5E2">
|
||||
<mods:roleTerm id="59A4903D615A84FB5AB056E028FB0335">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="ACDA47319F121BC386E63AD6DDE1E54E">Li, Zhaoyi</mods:namePart>
|
||||
<mods:nameIdentifier id="4665DA5E6B0C50E48452FA96EF441FB8" type="ORCID">0000-0001-6303-6041</mods:nameIdentifier>
|
||||
<mods:affiliation id="38F55A1B1E3A7CC587DE5C35668476F7">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
|
||||
<mods:namePart id="72788B8FC2598FFB2049ADD72E2D2D27">Li, Zhaoyi</mods:namePart>
|
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<mods:nameIdentifier id="3F3F30AA519518C9177CF47490427528" type="ORCID">0000-0001-6303-6041</mods:nameIdentifier>
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<mods:affiliation id="97905565739506DDA959C149301379A6">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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||||
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<mods:namePart id="11A37886D7C58B54EAE2FFE0DB20D58A">Zhang, Feng</mods:namePart>
|
||||
<mods:nameIdentifier id="93A763D4F46CF5822AD1123741A90813" type="ORCID">0000-0002-3347-1031</mods:nameIdentifier>
|
||||
<mods:affiliation id="03F8B135416FF3AF63828C0443D302A9">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
|
||||
<mods:namePart id="2B2E19147C5E0D034458703BCDE69C71">Zhang, Feng</mods:namePart>
|
||||
<mods:nameIdentifier id="A00B4C6920DC8951AA58DDE83E0B6A31" type="ORCID">0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation id="C8372C25276B71DD5CA321432E79E743">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:date id="22D6073EF03CF22F70EE19EE3E26A476">2025</mods:date>
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<mods:date id="02958E4D799B4091DE6DDABA65FC01E1">2025</mods:date>
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<mods:number id="9A73645D5D8D860A8E042745C74A59BE">1224</mods:number>
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<mods:classification id="B6E692BE29D5284D50B29D775329DF77">journal article</mods:classification>
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<mods:identifier id="EBD344455FD0D14151436922A24CCEAB" type="DOI">10.3897/zookeys.1224.135572</mods:identifier>
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<mods:classification id="8E33A52BB23AA4B5035C74AA0DDE04C7">journal article</mods:classification>
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<mods:identifier id="3AEE1FF460B169B0AAEB26BE88B5649E" type="DOI">10.3897/zookeys.1224.135572</mods:identifier>
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<mods:identifier id="6F7EEC88C9A2CD28BF6DB73C44FC585E" type="ZooBank">45FAE35A-4E17-4D75-8FAF-B43A3390AC8A</mods:identifier>
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<treatment id="9F72E0A29FDA57BBB05D3F0F35EB9FF3" ID-DOI="http://doi.org/10.5281/zenodo.14757047" ID-Zenodo-Dep="14757047" ID-arpha="9F72E0A2-9FDA-57BB-B05D-3F0F35EB9FF3" ID-zoobank="https://zoobank.org/86311755-A356-458D-8AA8-672B437D6514" LSID="urn:lsid:plazi:treatment:9F72E0A29FDA57BBB05D3F0F35EB9FF3" LSID-ZBK="urn:lsid:zoobank.org:act:86311755-A356-458D-8AA8-672B437D6514" httpUri="http://treatment.plazi.org/id/9F72E0A29FDA57BBB05D3F0F35EB9FF3">
|
||||
<treatment id="9F72E0A29FDA57BBB05D3F0F35EB9FF3" ID-DOI="http://doi.org/10.5281/zenodo.14757047" ID-Zenodo-Dep="14757047" ID-arpha="9F72E0A2-9FDA-57BB-B05D-3F0F35EB9FF3" ID-zoobank="https://zoobank.org/86311755-A356-458D-8AA8-672B437D6514" LSID="urn:lsid:plazi:treatment:9F72E0A29FDA57BBB05D3F0F35EB9FF3" LSID-ZBK="urn:lsid:zoobank.org:act:86311755-A356-458D-8AA8-672B437D6514" httpUri="http://treatment.plazi.org/id/9F72E0A29FDA57BBB05D3F0F35EB9FF3" scope_family="Clubionidae" scope_order="Araneae">
|
||||
<subSubSection id="4E6BC38CFE36CCA01BE52E16827EBA14" type="nomenclature">
|
||||
<paragraph id="2C139ACB56DE5436F59EA72CD49909F1">
|
||||
<taxonomicName id="1426A82CD96D3F4BD4EE3C3E444BCEA5" authorityName="Guo & Li & Zhang" authorityYear="2025" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="multiprocessa" status="sp. nov.">
|
||||
|
|
@ -72,20 +72,22 @@
|
|||
<heading id="76160C262BCF23AD98DB87928D4EB42E" reason="title">Type material.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="E91021D5FBBF7755BB3B57C4882E3B4F">
|
||||
<materialsCitation id="DAECE6C5A4144225138FAC85209D2CCC" collectingDate="2011-09-11" collectorName="Qiuju Wei." country="China" elevation="2500" location="Cang Shan" municipality="Dali Bai Autonomous Prefecture" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
|
||||
<materialsCitation id="A7CF1DD942D9E9505209A0753A377E09" collectingDate="2011-09-11" collectionCode="CLU" collectorName="Qiuju Wei" country="China" county="Dali Bai Autonomous Prefecture" elevation="2500" location="Cang Shan" municipality="Dali Bai Autonomous Prefecture" specimenCode="CLU 1440-1" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
|
||||
<emphasis id="DAEF73596DAA6A2C464234C5B1DE372B" bold="true" italics="true">
|
||||
<typeStatus id="93B20BB20FC9211D33EB4AA24E504EF1" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
•
|
||||
<specimenCount id="127469E1D528B736C30B93D1377988BF" count="1" type="male">♂</specimenCount>
|
||||
(CLU 1440-1),
|
||||
(
|
||||
<specimenCode id="68DD6E9B3710CC0950E69F55157603D7">CLU 1440-1</specimenCode>
|
||||
),
|
||||
<collectingCountry id="F66B8333F8D61CB4EF2D98BAF492B93A" name="China">China</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="1C73ED36E77971778C23A1A776D6DF73" country="China" name="Yunnan">Yunnan Province</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="F561CA26569F6DE0383FC4F72F488DF0">Dali Bai Autonomous Prefecture</collectingMunicipality>
|
||||
<collectingCounty id="4A88C6C1A5EF9E7C38DBABA4E3755A2D">Dali Bai Autonomous Prefecture</collectingCounty>
|
||||
,
|
||||
<location id="B008F7C6B5477A76AC0E16AE7672C54A" LSID="urn:lsid:plazi:treatment:9F72E0A29FDA57BBB05D3F0F35EB9FF3:B008F7C6B5477A76AC0E16AE7672C54A" country="China" municipality="Dali Bai Autonomous Prefecture" name="Cang Shan" stateProvince="Yunnan">Cang Shan</location>
|
||||
<location id="B008F7C6B5477A76AC0E16AE7672C54A" LSID="urn:lsid:plazi:treatment:9F72E0A29FDA57BBB05D3F0F35EB9FF3:B008F7C6B5477A76AC0E16AE7672C54A" country="China" county="Dali Bai Autonomous Prefecture" municipality="Dali Bai Autonomous Prefecture" name="Cang Shan" stateProvince="Yunnan Province">Cang Shan</location>
|
||||
;
|
||||
<quantity id="CA7CD4039BAE81EEBB36EB803EB6E682" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">
|
||||
<elevation id="13B9FC886D638880671DC769C213E815" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">2500 m</elevation>
|
||||
|
|
@ -95,16 +97,21 @@ elev.;
|
|||
<collectingDate id="6712AE0E6CDA88C1EF47A50BC4DDDB35" value="2011-09-11">11. IX. 2011</collectingDate>
|
||||
</date>
|
||||
; leg.
|
||||
<collectorName id="81B97589E544BAC0AE8778A571708969">Qiuju Wei.</collectorName>
|
||||
<collectorName id="0B331837CBBDD01DA8AC1B0D9959A351">Qiuju Wei</collectorName>
|
||||
</materialsCitation>
|
||||
<materialsCitation id="B899712AE5DD1E5B041EF4B0B59F2E45" collectingDate="2011-09-11" collectorName="Qiuju Wei." country="China" elevation="2500" location="Cang Shan" municipality="Dali Bai Autonomous Prefecture" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
|
||||
.
|
||||
<materialsCitation id="B899712AE5DD1E5B041EF4B0B59F2E45" collectingDate="2011-09-11" collectionCode="CLU" collectorName="Qiuju Wei." country="China" county="Dali Bai Autonomous Prefecture" elevation="2500" location="Cang Shan" municipality="Dali Bai Autonomous Prefecture" specimenCode="CLU 1440-2, CLU 1440-7" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
|
||||
<emphasis id="B75621412A5F57826B1BA7EE700C4E36" bold="true" italics="true">
|
||||
<typeStatus id="81B9B44A89BB98C8BD6E114C102F777D" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
•
|
||||
<specimenCount id="D3A0F2B433250FE29DD99A0141FE175C" count="2" type="male">2 ♂</specimenCount>
|
||||
<specimenCount id="6FFD322A6DB9F16305C15E54E4B0725D" count="4" type="female">4 ♀</specimenCount>
|
||||
(CLU 1440-2 – CLU 1440 - 7); same data as holotype
|
||||
(
|
||||
<specimenCode id="A5F777A08CC1577B0D5FE18AFF954E83">CLU 1440-2</specimenCode>
|
||||
–
|
||||
<specimenCode id="4B062C321FA8EB5AEC183D3BAF9701A9">CLU 1440-7</specimenCode>
|
||||
); same data as holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
|
|
@ -115,12 +122,15 @@ elev.;
|
|||
</paragraph>
|
||||
<paragraph id="18B4BF3692D4790935714101B2132EA1">
|
||||
•
|
||||
<materialsCitation id="F8064F14ECC6804849677C483871A406" collectingDate="2008-08-11" collectorName="Tingbang Yang" elevation="2500" location="Cang" specimenCount="10" specimenCount-female="9" specimenCount-male="1" stateProvince="Shan">
|
||||
<materialsCitation id="F8064F14ECC6804849677C483871A406" collectingDate="2008-08-11" collectionCode="CLU" collectorName="Tingbang Yang" elevation="2500" location="Cang Shan" specimenCode="CLU 1439-1, CLU 1439-10" specimenCount="10" specimenCount-female="9" specimenCount-male="1" stateProvince="Shan">
|
||||
<specimenCount id="7B2861FA01054A1F6419817DC1413494" count="1" type="male">1 ♂</specimenCount>
|
||||
<specimenCount id="742C990FBA0632C2C75F65E57F5DA53C" count="9" type="female">9 ♀</specimenCount>
|
||||
(CLU 1439-1 – CLU 1439 - 10),
|
||||
<location id="17038E02C0E548CBDE1DA5C740310DD9" LSID="urn:lsid:plazi:treatment:9F72E0A29FDA57BBB05D3F0F35EB9FF3:17038E02C0E548CBDE1DA5C740310DD9" name="Cang" stateProvince="Shan">Cang</location>
|
||||
<collectingRegion id="86526E04DFDEEEC3361CD18AB6C4ADB9" country="Myanmar" name="Shan">Shan</collectingRegion>
|
||||
(
|
||||
<specimenCode id="80ABA21F1E54025E8EFD1F33D307E6E0">CLU 1439-1</specimenCode>
|
||||
–
|
||||
<specimenCode id="8FEC6FAE4684DF823543FFDA3AF6F0C4">CLU 1439-10</specimenCode>
|
||||
),
|
||||
<location id="9C38A3438C0CE3E2E81870B4E3A0A3CE" LSID="urn:lsid:plazi:treatment:9F72E0A29FDA57BBB05D3F0F35EB9FF3:9C38A3438C0CE3E2E81870B4E3A0A3CE" name="Cang Shan" stateProvince="Shan">Cang Shan</location>
|
||||
,
|
||||
<quantity id="52FB1AA9CB96F536DF240CDBDE789084" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">
|
||||
<elevation id="A7A8F9D5102B193221FF63BF9B378B2A" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="m" value="2500.0">2500 m</elevation>
|
||||
|
|
@ -133,12 +143,15 @@ elev.;
|
|||
<collectorName id="9F08FB04F71C0AF8BFBD8904FE23CE37">Tingbang Yang</collectorName>
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation id="05A408CD742496DE304C8EAF1DB48E9F" collectingDate="2011-08-09" collectorName="Qiuju Wei." elevation="2600" location="Cang" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="Shan">
|
||||
<materialsCitation id="05A408CD742496DE304C8EAF1DB48E9F" collectingDate="2011-08-09" collectionCode="CLU" collectorName="Qiuju Wei." elevation="2600" location="Cang Shan" specimenCode="CLU 1441-1, CLU 1441-8" specimenCount="8" specimenCount-female="7" specimenCount-male="1" stateProvince="Shan">
|
||||
<specimenCount id="141C8BAB07EB3C0F23E152EDAA9F9317" count="1" type="male">1 ♂</specimenCount>
|
||||
<specimenCount id="7B2C7DF7EE213D5F820B699AC980677F" count="7" type="female">7 ♀</specimenCount>
|
||||
(CLU 1441-1 – CLU 1441 - 8);
|
||||
<location id="146BE7D3A2039A26EF8233D3BCEBE286" LSID="urn:lsid:plazi:treatment:9F72E0A29FDA57BBB05D3F0F35EB9FF3:146BE7D3A2039A26EF8233D3BCEBE286" name="Cang" stateProvince="Shan">Cang</location>
|
||||
<collectingRegion id="4102210E063E9E99AE0091B0D6A822D3" country="Myanmar" name="Shan">Shan</collectingRegion>
|
||||
(
|
||||
<specimenCode id="4E71F11E966B785FDD663BBD64347648">CLU 1441-1</specimenCode>
|
||||
–
|
||||
<specimenCode id="61F0B07C5F0B864F256B52CF71A99622">CLU 1441-8</specimenCode>
|
||||
);
|
||||
<location id="1CD24CE91451044D54D4F273FE18A04E" LSID="urn:lsid:plazi:treatment:9F72E0A29FDA57BBB05D3F0F35EB9FF3:1CD24CE91451044D54D4F273FE18A04E" name="Cang Shan" stateProvince="Shan">Cang Shan</location>
|
||||
;
|
||||
<quantity id="2E3978E310D78637C26B866E201917AF" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
|
||||
<elevation id="23EF4B6CDD3CD4B1DFB021998016DC7F" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
|
||||
|
|
@ -156,7 +169,7 @@ elev.;
|
|||
<paragraph id="2F35A7FC54792686CA7A90D6BAFF8AF7">
|
||||
<heading id="BEF5CD72EF5A76EDAD96C57AC8CD7765" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="A75D4F3F95E81A851C129C7190B9A1F2">The specific name comes from the combination of “ multi- ” and “ processus ”, referring to the multiple (six) apophysis on the male palp; an adjective.</paragraph>
|
||||
<paragraph id="A75D4F3F95E81A851C129C7190B9A1F2">The specific name comes from the combination of “multi-” and “processus”, referring to the multiple (six) apophysis on the male palp; an adjective.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ELWAE" type="diagnosis">
|
||||
<paragraph id="AF0D64946A943B685751F75DA1622A78">
|
||||
|
|
@ -255,9 +268,7 @@ wider than
|
|||
(Fig.
|
||||
<figureCitation id="0FD1CF4DD39CAC4FA8BDDBA086CA2595" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Clubiona multiprocessa sp. nov., holotype male A left palp, ventral view B same, prolateral view C same, retrolateral view. Abbreviations: C = conductor; E = embolus; LTA = lateral tibial apophysis; PPA = prolateral patellar apophysis; RFA = retrolateral femoral apophysis; RPA = retrolateral patellar apophysis; RTA = retrolateral tibial apophysis; VTA = ventral tibial apophysis." figureDoi="10.3897/zookeys.1224.135572.figure5" httpUri="https://binary.pensoft.net/fig/1236385">5</figureCitation>
|
||||
). Femur with a slanting finger-like retrolateral apophysis (
|
||||
<abbrev id="ABBRID0E42AE" xlink_title="retrolateral femoral apophysis">
|
||||
<collectionCode id="CCDDB2A392C45A42C328984E573B0BC8" country="Brazil" lsid="urn:lsid:biocol.org:col:14341" name="Universidade Federal do Rio de Janeiro" type="Herbarium">RFA</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0E42AE" xlink_title="retrolateral femoral apophysis">RFA</abbrev>
|
||||
), ca 3 / 4 of femur’s length.
|
||||
<abbrev id="ABBRID0EB3AE" xlink_title="retrolateral femoral apophysis">RFA</abbrev>
|
||||
partly membranous, arising mesially from femur, directing retrolatero-dorsally. Patella with two apophyses:
|
||||
|
|
@ -309,21 +320,15 @@ body length 3.75. Carapace 1.68 long, 1.33 wide; abdomen 2.07 long, 1.47 wide. C
|
|||
). Epigynal plate slightly wider than long. Atrium (A) nearly heart-shaped, located at anterior portion of epigynal plate. Copulatory openings (
|
||||
<abbrev id="ABBRID0EQ5AE" xlink_title="copulatory opening">CO</abbrev>
|
||||
) tiny, located centrally in atrium. Copulatory ducts (
|
||||
<abbrev id="ABBRID0EU5AE" xlink_title="copulatory duct">
|
||||
<collectionCode id="F5D246F958084A0D52D19F2A0234C996">CD</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EU5AE" xlink_title="copulatory duct">CD</abbrev>
|
||||
) invisible. Spermathecae (S) long, located at the anterior position of bursae, spermathecal heads (
|
||||
<abbrev id="ABBRID0EY5AE" xlink_title="spermathecal head">SH</abbrev>
|
||||
) ovate, with a papilliform spermathecal appendage (
|
||||
<abbrev id="ABBRID0E35AE" xlink_title="spermathecal appendage">
|
||||
<collectionCode id="14EE9BC63DF2BA15EF022A665662E616" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie">SA</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0E35AE" xlink_title="spermathecal appendage">SA</abbrev>
|
||||
), spermathecal bases (
|
||||
<abbrev id="ABBRID0EA6AE" xlink_title="spermathecal base">SB</abbrev>
|
||||
) tubular, with small fertilisation ducts terminally (
|
||||
<abbrev id="ABBRID0EE6AE" xlink_title="fertilisation duct">
|
||||
<collectionCode id="BB19345D2E86A68F88504F6CFC1B07C4">FD</collectionCode>
|
||||
</abbrev>
|
||||
<abbrev id="ABBRID0EE6AE" xlink_title="fertilisation duct">FD</abbrev>
|
||||
). Bursae (B) close together, nearly spherical, situated posteriorly.
|
||||
</paragraph>
|
||||
<caption id="925F0A9A79E934371FB4CD6FF8219CAF" ID-DOI="10.3897/zookeys.1224.135572.figure6" ID-arpha="F4A67286-7ECC-53CD-AFC1-75C191514CB1" httpUri="https://binary.pensoft.net/fig/1236386" startId="F6">
|
||||
|
|
|
|||
146
data/B9/BE/40/B9BE40AA7B0E58F394553984242060F6.xml
Normal file
146
data/B9/BE/40/B9BE40AA7B0E58F394553984242060F6.xml
Normal file
|
|
@ -0,0 +1,146 @@
|
|||
<document id="267B0DA10D7BB474AE9353276556B127" ID-DOI="10.3897/zookeys.1224.135572" ID-ZooBank="45FAE35A-4E17-4D75-8FAF-B43A3390AC8A" ID-publisher-id="135572" URI-arpha="04F180BC-E406-56DC-B94F-53EAA0AEFDF7" URI-zoobank="http://zoobank.org/45FAE35A-4E17-4D75-8FAF-B43A3390AC8A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="review-article" checkinTime="1738088623208" checkinUser="pensoft" docAuthor="Guo, Minghao, Li, Zhaoyi & Zhang, Feng" docDate="2025" docId="B9BE40AA7B0E58F394553984242060F6" docLanguage="en" docName="ZooKeys 1224: 175-186" docOrigin="ZooKeys 1224" docSource="https://zookeys.pensoft.net/article/135572/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Clubiona corticalis" docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="186" masterDocId="04F180BCE40656DCB94F53EAA0AEFDF7" masterDocTitle="Two new species of the Clubiona corticalis group (Araneae, Clubionidae) from Yunnan, China" masterLastPageNumber="186" masterPageNumber="175" pageNumber="175" updateTime="1738093079267" updateUser="ExternalLinkService">
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<mods:mods id="9F7D61FD94C2754B625C5DC947AB87B0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A96C0C1ECC01D4E74B44170C984149E0">
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<mods:title id="2E17EA19568D5328EA37E6CF6E0247B7">Two new species of the Clubiona corticalis group (Araneae, Clubionidae) from Yunnan, China</mods:title>
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</mods:titleInfo>
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<mods:name id="2000E464EFFAAE5866C0938C670A739C" type="personal">
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<mods:namePart id="1014944D3D4805C5E0A1AF49805F2962">Guo, Minghao</mods:namePart>
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<mods:affiliation id="6FE5B062C7AC9C8011942DD9A81BA0E8">Hebei Vocational University of Industry and Technology, Shijiazhuang, Hebei 050091, China</mods:affiliation>
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</mods:role>
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<mods:namePart id="EB20D5BE1B4B1F972A14F03203F04C99">Li, Zhaoyi</mods:namePart>
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<mods:nameIdentifier id="45267C38FA22909ABC527402C74ECE04" type="ORCID">0000-0001-6303-6041</mods:nameIdentifier>
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<mods:affiliation id="19A8A7C73F52E53CF746690E50113EC9">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="7B92B706E3599F7C6D8D5A8BAA77F66B">Zhang, Feng</mods:namePart>
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<mods:affiliation id="847F484D3515F2E26D94A3650B5E959D">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:title id="8CCC7AF749BBDD441C9FA23549333458">ZooKeys</mods:title>
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<mods:part id="E740B32A7E9ED54CAAF640B4FDFBF615">
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<mods:date id="4DB90C9A48C20DAA93DC43573A36F61F">2025</mods:date>
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<mods:number id="13EA4C3F050805620ED2308CEC431BFD">2025-01-28</mods:number>
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</mods:detail>
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<mods:detail id="96D34CC41397E3DA1E548D3DF1ACC691" type="volume">
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<mods:number id="F5BCA031BBAF679488AFBB8B0E6B5765">1224</mods:number>
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<mods:start id="AC7AAC74D809C21CB55E70F91D6F9184">175</mods:start>
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</mods:extent>
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<mods:classification id="26D0F4A9A7DC819C7ADAFAA940FF3ABE">journal article</mods:classification>
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<mods:identifier id="E60BEB54E20552249D7D52933DA0BC08" type="ZooBank">45FAE35A-4E17-4D75-8FAF-B43A3390AC8A</mods:identifier>
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<treatment id="B9BE40AA7B0E58F394553984242060F6" ID-DOI="http://doi.org/10.5281/zenodo.14757438" ID-Zenodo-Dep="14757438" ID-arpha="B9BE40AA-7B0E-58F3-9455-3984242060F6" LSID="urn:lsid:plazi:treatment:B9BE40AA7B0E58F394553984242060F6" httpUri="http://treatment.plazi.org/id/B9BE40AA7B0E58F394553984242060F6" scope_family="Clubionidae" scope_order="Araneae">
|
||||
<subSubSection id="2C13E1654FD37465BA818A2381586FB9" type="nomenclature">
|
||||
<paragraph id="EC8CA2E27DC6EA0BC8F7B520E73011D5">
|
||||
<taxonomicName id="B11326E91A3A3F70C8BE30E53ABEF7A4" ID-CoL="WC6Z" baseAuthorityName="Walckenaer" baseAuthorityYear="1802" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="corticalis">
|
||||
<emphasis id="7770F8B0C0EEEDDD0B3331946F92D1BE" italics="true">Clubiona corticalis</emphasis>
|
||||
</taxonomicName>
|
||||
group
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="64462D68A878234BCD13FCF3E58738D3" type="taxon_list">
|
||||
<paragraph id="DDE2989DD169FCCD9E407214B63FEE87">
|
||||
<treatmentCitationGroup id="517E70FB8EE4D07C751D4B8EB809E5E7">
|
||||
<treatmentCitation id="B1A9C738486CF87E4AA32A955948A759" author="Thorell T" page="54" year="1887">
|
||||
<taxonomicName id="5802778C4E637509C1C11A687B2A079B" authority="Thorell, 1887: 54" authorityName="Thorell" authorityPageNumber="54" authorityYear="1887" class="Arachnida" family="Clubionidae" genus="Atalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="39DED0BF55D4B2D11F7CEC380140F0D9" italics="true">Atalia</emphasis>
|
||||
<comment id="A4EB8BD0A3922A878054BBE81EE1A463">
|
||||
<bibRefCitation id="9A6D526C722BB69ED00146C45EF27FE5" author="Thorell T" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pagination="5-417" refId="B12" refString="Thorell T (1887) Viaggio di L. Fea in Birmania e regioni vicine. II. Primo saggio sui ragni birmani. Annali del Museo Civico di Storia Naturale di Genova 25: 5 - 417." title="Viaggio di L. Fea in Birmania e regioni vicine. II. Primo saggio sui ragni birmani." volume="25" year="1887">Thorell, 1887: 54</bibRefCitation>
|
||||
(type species
|
||||
<treatmentCitation id="47A7DF5568BAC8CA095A93940A3E9D98">
|
||||
<taxonomicName id="AD7BDF42BF8E0D540F38FE216181D613" authority="Thorell, 1887" authorityName="Thorell" authorityYear="1887" class="Arachnida" family="Clubionidae" genus="Atalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="concinna">
|
||||
<emphasis id="A8132D2375BB472DA33EE4EFAD37BD24" italics="true">Atalia concinna</emphasis>
|
||||
<bibRefCitation id="6073A22A25EEEF69ABAB15772708C407" author="Thorell T" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pagination="5-417" refId="B12" refString="Thorell T (1887) Viaggio di L. Fea in Birmania e regioni vicine. II. Primo saggio sui ragni birmani. Annali del Museo Civico di Storia Naturale di Genova 25: 5 - 417." title="Viaggio di L. Fea in Birmania e regioni vicine. II. Primo saggio sui ragni birmani." volume="25" year="1887">Thorell, 1887</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
).
|
||||
</comment>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="FC515046A94EFE2719B855EBA45DD206">
|
||||
<treatmentCitationGroup id="AB2A05D84FB4063145D31297BD6A4B37">
|
||||
<taxonomicName id="A1DF26E6D967F4646CF83779338E106D" ID-CoL="WC6Z" baseAuthorityName="Walckenaer" baseAuthorityYear="1802" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="corticalis">
|
||||
<emphasis id="95B237548304C2704E92D3BFBA1F0C47" italics="true">Clubiona corticalis</emphasis>
|
||||
</taxonomicName>
|
||||
<comment id="E38BFEB4BABCDDF513BF07DE78760BE5">
|
||||
group:
|
||||
<treatmentCitation id="84C8AE53333BD89224946074454A6D83" author="Simon E" page="905" year="1932">
|
||||
<bibRefCitation id="FF242D9A499AC2C824B22B418B232B69" author="Simon E" firstAuthor="Simon" page="905" refId="B11" refString="Simon E (1932) Les arachnides de France. Tome VI. Synopsis générale et catalogue des espèces françaises de l’ordre des Araneae; 4 e partie. Roret, Paris, 773–978." year="1932">Simon 1932: 905</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
;
|
||||
<treatmentCitation id="306DAC8F6861636A6C325634F8E02BE4" author="Mikhailov KG" page="142" year="1990">
|
||||
<bibRefCitation id="F3F5B51D591C3771CB29237C508F78F8" author="Mikhailov KG" firstAuthor="Mikhailov" journalOrPublisher="Korean Arachnology" page="142" pagination="139-175" refId="B8" refString="Mikhailov KG (1990) The spider genus Clubiona Latreille 1804 in the Soviet Far East, 1 (Arachnida, Aranei, Clubionidae). Korean Arachnology 5: 139–175." title="The spider genus Clubiona Latreille 1804 in the Soviet Far East, 1 (Arachnida, Aranei, Clubionidae)." volume="5" year="1990">Mikhailov 1990: 142</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
;
|
||||
<treatmentCitation id="C651DD772AE0AD4DC7156E6F3AB35246" author="Mikhailov KG" page="38" year="1995">
|
||||
<bibRefCitation id="967FFFD5F6090B54A326ACD636AD07B8" author="Mikhailov KG" firstAuthor="Mikhailov" issue="2" journalOrPublisher="Arthropoda Selecta" page="38" pagination="33-48" refId="B9" refString="Mikhailov KG (1995) Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach. Arthropoda Selecta 4 (2): 33–48." title="Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach." volume="4" year="1995">Mikhailov 1995: 38</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
, 42;
|
||||
<treatmentCitation id="8E63DCE143CF507F613976E8455FF977" author="Deeleman-Reinhold CL" page="90" year="2001">
|
||||
<bibRefCitation id="09C473AD8AC1AD0B0E07A69EA3C9959F" DOI="10.1163/9789004475588" author="Deeleman-Reinhold CL" firstAuthor="Deeleman-Reinhold" journalOrPublisher="Brill, Leiden" page="90" refId="B1" refString="Deeleman-Reinhold CL (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp. https://doi.org/10.1163/9789004475588" title="Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae)." year="2001">Deeleman-Reinhold 2001: 90</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
;
|
||||
<treatmentCitation id="8B45611E6DBC89445FF8ED7AE3637F4E" ID-DOI="http://doi.org/10.5281/zenodo.3797845" ID-Zenodo-Dep="3797845" author="Yu H & Li SQ" httpUri="http://treatment.plazi.org/id/995687C0FF8ECA09CDC82148FDEACD4E" page="153" year="2019">
|
||||
<bibRefCitation id="D899F60B6BB5C1309E6C6578DCF00E4E" DOI="10.11646/zootaxa.4545.2.1" author="Yu H & Li SQ" firstAuthor="Yu" issue="2" journalOrPublisher="Zootaxa" page="153" pagination="151-178" refId="B16" refString="Yu H, Li SQ (2019) Eight new species of the genus Clubiona Latreille, 1804 from Xishuangbanna Rainforest, southwestern China (Araneae, Clubionidae). Zootaxa 4545 (2): 151–178. https://doi.org/10.11646/zootaxa.4545.2.1" title="Eight new species of the genus Clubiona Latreille, 1804 from Xishuangbanna Rainforest, southwestern China (Araneae, Clubionidae)." volume="4545" year="2019">Yu and Li 2019: 153</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
.
|
||||
</comment>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="97D72A75D3E2816E387C0F9FB48FEAD0">
|
||||
<treatmentCitationGroup id="B8BCD4364A4ECEEAA3F60C17E6943245">
|
||||
<treatmentCitation id="0C25F448D66226B1EA71FBE5EEFD445E" author="Lohmander H" page="19" year="1944">
|
||||
<taxonomicName id="0A875B43FEF4E3115AE355996DBD2AA2" authority="Lohmander, 1944: 19" authorityName="Lohmander" authorityPageNumber="19" authorityYear="1944" class="Arachnida" family="Clubionidae" genus="Paraclubiona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="A30A973FDB77984E980CDE41C82A9325" italics="true">Paraclubiona</emphasis>
|
||||
<comment id="920C6BBA0D8FBA18A79BDBD99A7DC83F">
|
||||
<bibRefCitation id="D82CE2F62A76B7E092CCDF97B37AB1E9" author="Lohmander H" refId="B7" refString="Lohmander H (1944) Vorlaufige Spinnennotisen. Arkiv for Zoologi 35 (A, 16): 1 - 21." year="1944">Lohmander, 1944: 19</bibRefCitation>
|
||||
(type species
|
||||
<taxonomicName id="86A45FB631F95DC2AB1EC0B2F9272E65" authority="Walckenaer, 1802" authorityName="Walckenaer" authorityYear="1802" class="Arachnida" family="Clubionidae" genus="Aranea" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="corticalis">
|
||||
<emphasis id="6BF3240476F66F29AB5717D7AFA7D488" italics="true">Aranea corticalis</emphasis>
|
||||
Walckenaer, 1802
|
||||
</taxonomicName>
|
||||
).
|
||||
</comment>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EPAAE" type="diagnosis">
|
||||
<paragraph id="8BE24BF3BEBD8F7BFB99E936B57F5DF9">
|
||||
<heading id="FA4B4F604771C8F6BD5F6B05FC472C28" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="5FBF6BD732E7290F6EC0B9BC294F10D4">
|
||||
See
|
||||
<bibRefCitation id="ADAE3FBD973735CCD1AE9B3859430782" author="Mikhailov KG" firstAuthor="Mikhailov" issue="2" journalOrPublisher="Arthropoda Selecta" pagination="33-48" refId="B9" refString="Mikhailov KG (1995) Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach. Arthropoda Selecta 4 (2): 33–48." title="Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach." volume="4" year="1995">Mikhailov (1995)</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="098CF128AF9D294E3EF4256457D06335" DOI="10.1163/9789004475588" author="Deeleman-Reinhold CL" firstAuthor="Deeleman-Reinhold" journalOrPublisher="Brill, Leiden" refId="B1" refString="Deeleman-Reinhold CL (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, 591 pp. https://doi.org/10.1163/9789004475588" title="Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae)." year="2001">Deeleman-Reinhold (2001)</bibRefCitation>
|
||||
, and
|
||||
<bibRefCitation id="6409C3EA494FA51ED0965D913A6E23AF" DOI="10.11646/zootaxa.4545.2.1" author="Yu H & Li SQ" firstAuthor="Yu" issue="2" journalOrPublisher="Zootaxa" pagination="151-178" refId="B16" refString="Yu H, Li SQ (2019) Eight new species of the genus Clubiona Latreille, 1804 from Xishuangbanna Rainforest, southwestern China (Araneae, Clubionidae). Zootaxa 4545 (2): 151–178. https://doi.org/10.11646/zootaxa.4545.2.1" title="Eight new species of the genus Clubiona Latreille, 1804 from Xishuangbanna Rainforest, southwestern China (Araneae, Clubionidae)." volume="4545" year="2019">Yu and Li (2019)</bibRefCitation>
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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